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9 August
2018

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The Freshwater Mussels (Unionoida) of the World
(and other less consequential bivalves)

FM(U)otW(aolcb) is the web version of the MUSSEL Project Database. Follow the links to browse the data or use the search fields. Either way, you win!

Taxa known only as fossils are highlighted in gray. Those without representatives in fresh waters are highlighted in gold.

 

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Publication Information

Haas, F. 1940. A tentative classification of the Palearctic unionids. Zoological Series of Field Museum of Natural History 24: 115-141.
Availably electronically? yes.

Species Described
Anodonta quelleneci Haas, 1940.
‘Drouet’ Haas, 1940, Zool. Ser. Field Mus. Nat. Hist.: 137.
(error for Anodonta quellenneci Drouet, 1892)
= Anodonta cygnea
 
Margarita (Unio) egyptiacus Jay, 1836.
‘Cailliaud’ Of Authors.
Lea, 1836, Synopsis Fam. Naiades: 29.
Jay, 1836, Cat. Rec. Shells: 20.
Lea, 1838, Synopsis Fam. Naiades: 21.
Hanley, 1843, Illustr. Descr. Cat. Rec. Bivalve Shells: 198.
Lea, 1852, Synopsis Fam. Naiades: 32.
Conrad, 1853, Proc. Acad. Nat. Sci.: 267.
Adams & Adams, 1857, Gen. Rec. Moll.: 494.
Lea, 1870, Synopsis Fam. Naiades: 50.
Haas, 1940, Zool. Ser. Field Mus. Nat. Hist.: 140 [as ‘Caelatura egyptiaca’].
Araujo et al., 2017, Zool. J. Linn. Soc.: 280.
(error for Unio aegyptiacus Cailliaud, 1827)
= Coelatura aegyptiaca
 
Margaritana durrowensis Haas, 1940.
‘Phillips’ Haas, 1940, Zool. Ser. Field Mus. Nat. Hist.: 120.
(error for Margaritana durrovensis Phillips, 1928)
= Margaritifera margaritifera
 
Margaritifera maroccana Haas, 1940.
Haas, 1940, Zool. Ser. Field Mus. Nat. Hist.: 119.
(error for Margaritana marocana Pallary, 1918)
= Pseudunio marocana
 
Microcondylus crassus Haas, 1940.
‘Drouët’ Haas, 1940, Zool. Ser. Field Mus. Nat. Hist.: 133.
(error for Microcondylus crassulus Drouët, 1879)
= Microcondylaea bonellii
 
Unio albovicensis Haas, 1940.
‘Locard’ Haas, 1940, Zool. Ser. Field Mus. Nat. Hist.: 129.
(error for Unio elboviensis Locard, 1893)
= Unio tumidus
 
Unio brevieri Locard, 1882.
‘Bourguignat in Locard’ Of Authors.
Locard, 1882, Prodr. Malac. Franç.: 286.
Germain, 1904, Bull. Soc. Sci. Nat. Ouest France: 265.
Germain, 1931, Faune de France: 732.
Haas, 1940, Zool. Ser. Field Mus. Nat. Hist.: 130.
Haas, 1969, Das Tierreich: 49.
(incorrect original spelling of Unio brevierei Locard, 1882)
= Unio crassus
 
Unio bythinicus Simpson, 1900.
‘Kobelt’ Of Authors.
Simpson, 1900, Proc. U.S. Nat. Mus.: 695.
Simpson, 1914, Descr. Cat. Naiades: 565.
Haas, 1940, Zool. Ser. Field Mus. Nat. Hist.: 141.
(error for Unio bithynicus Kobelt, 1893)
= Elongaria orientalis
 
Unio caravellus Haas, 1940.
‘Servain’ Haas, 1940, Zool. Ser. Field Mus. Nat. Hist.: 121.
‘Servain’ Haas, 1969, Das Tierreich: 23.
(error for Unio cavarellus Servain, 1887)
= Unio pictorum
 
Unio diarbekrianus Haas, 1940.
‘Kobelt’ Haas, 1940, Zool. Ser. Field Mus. Nat. Hist.: 132.
(error for Unio diarbekirianus Kobelt, 1913)
= Unio crassus
 
Unio gangraenosus Küster, 1855.
‘Ziegler’ Of Authors.
Küster, 1855, Conchyl. Cab.: 124.
Westerlund, 1890, Fauna Paläarct. Binnenconch.: 81.
Haas, 1940, Zool. Ser. Field Mus. Nat. Hist.: 130.
Haas, 1969, Das Tierreich: 46.
(error for Unio gangrenosus J.F. Schmidt, 1840)
= Unio crassus
 
Unio hattmanni Haas, 1940.
‘Bourguignat’ Haas, 1940, Zool. Ser. Field Mus. Nat. Hist.: 131.
(error for Unio hattemani Locard, 1889)
= Unio crassus
 
Unio hospitali Locard, 1893.
‘Locard’ Of Authors.
Locard, 1893, Coqu. Eau Douce Saumâtre France: 186.
Germain, 1931, Faune de France: 735.
Haas, 1940, Zool. Ser. Field Mus. Nat. Hist.: 121.
(error for Unio hopitali Locard, 1889)
= Unio pictorum
 
Unio hydrellus Haas, 1940.
‘Locard’ Haas, 1940, Zool. Ser. Field Mus. Nat. Hist.: 122.
(error for Unio hydrelus Locard, 1889)
= Unio mancus
 
Unio jonicus Kobelt, 1886.
‘Drouët’ Of Authors.
Kobelt, 1886, Icon. Land- und Süssw.-Moll.: 43.
Westerlund, 1890, Fauna Paläarct. Binnenconch.: 153.
Kobelt, 1893, Icon. Land- und Süssw.-Moll.: 87.
Haas, 1940, Zool. Ser. Field Mus. Nat. Hist.: 131.
Haas, 1969, Das Tierreich: 52.
(error for Unio ionicus Drouët, 1879)
= Unio ionicus
 
Unio jousseaumi Haas, 1940.
‘Bourguignat’ Haas, 1940, Zool. Ser. Field Mus. Nat. Hist.: 121.
(error for Unio jousseaumei Locard, 1882)
= Unio pictorum
 
Unio lamboltei Locard, 1893.
‘Malzine’ Of Authors.
Locard, 1893, Coqu. Eau Douce Saumâtre France: 177.
Germain, 1931, Faune de France: 732.
Haas, 1940, Zool. Ser. Field Mus. Nat. Hist.: 130.
(error for Unio lambottei Malzine, 1867)
= Unio crassus
 
Unio lawleyanus Kobelt, 1884.
‘Gentiluomo’ Of Authors.
Kobelt, 1884, Icon. Land- und Süssw.-Moll.: 65.
Simpson, 1900, Proc. U.S. Nat. Mus.: 683.
Simpson, 1914, Descr. Cat. Naiades: 533.
Haas, 1940, Zool. Ser. Field Mus. Nat. Hist.: 127.
(error for Unio lawleyianus Gentiluomo, 1868)
= Unio mancus
 
Unio pincianus Haas, 1940.
Haas, 1940, Zool. Ser. Field Mus. Nat. Hist.: 121.
Haas, 1969, Das Tierreich: 23.
(error for Unio pinciacus Locard, 1882)
= Unio mancus
 
Unio platyodon Haas, 1940.
‘Jordan’ Haas, 1940, Zool. Ser. Field Mus. Nat. Hist.: 121.
(error for Unio pachyodon Jordan, 1879)
= Unio pictorum
 
Unio platyrhynchoideus Locard, 1882.
‘Dupuy’ Of Authors.
Locard, 1882, Prodr. Malac. Franç.: 297.
Locard, 1889, Contr. Faune Malacol. Franc.: 154 [48].
Westerlund, 1890, Fauna Paläarct. Binnenconch.: 121.
Locard, 1893, Coqu. Eau Douce Saumâtre France: 187.
Germain, 1931, Faune de France: 739.
Haas, 1940, Zool. Ser. Field Mus. Nat. Hist.: 123.
Haas, 1969, Das Tierreich: 27.
Fauna Europaea, 2012, Fauna Europa: [web].
(error for Unio platyrinchoideus Dupuy, 1849)
= Unio pictorum
 
Unio quelleneci Kobelt, 1896.
‘Drouët’ Kobelt, 1896, Icon. Land- und Süssw.-Moll.: 100.
‘Drouet’ Haas, 1940, Zool. Ser. Field Mus. Nat. Hist.: 125.
(error for Unio quellenneci Drouët, 1895)
= Unio mancus
 
Unio ricconianus Haas, 1940.
‘Monterosato’ Haas, 1940, Zool. Ser. Field Mus. Nat. Hist.: 127.
(intro'd as syn of Unio gargottae Philippi, 1836)
= Unio mancus
 
Unio sabinensis Haas, 1940.
Haas, 1940, Zool. Ser. Field Mus. Nat. Hist.: 127.
(error for Unio sebinensis Bourguignat, 1883)
= Unio mancus
 
Unio schrenckianus Haas, 1940.
‘Clessin’ Haas, 1940, Zool. Ser. Field Mus. Nat. Hist.: 122.
(error for Unio schrenkianus Clessin, 1880)
= Unio pictorum
 
Unio souzanus Locard, 1893.
‘Castro’ Of Authors.
Locard, 1893, Coqu. Eau Douce Saumâtre France: 197.
Germain, 1904, Bull. Soc. Sci. Nat. Ouest France: 275 [37].
Germain, 1931, Faune de France: 737.
Haas, 1940, Zool. Ser. Field Mus. Nat. Hist.: 126.
(error for Unio sousanus Locard, 1889)
= Unio delphinus
 
Unio zoasthenicus Haas, 1940.
‘Locard’ Haas, 1940, Zool. Ser. Field Mus. Nat. Hist.: 131.
(error for Unio zoasthenus Locard, 1889)
= Unio mancus
 
Unio (cilicicus var.) berythensis Haas, 1940.
‘Kobelt & Rolle’ Haas, 1940, Zool. Ser. Field Mus. Nat. Hist.: 128.
Haas, 1969, Das Tierreich: 42.
(error for Unio berytensis Kobelt & Rolle, 1895)
= Unio mancus
 
Unio kungurensis sylvensis Haas, 1940.
‘Kobelt’ Haas, 1940, Zool. Ser. Field Mus. Nat. Hist.: 132.
‘Kobelt’ Haas, 1969, Das Tierreich: 53.
(error for Unio sylwensis Kobelt, 1912)
= Unio crassus
 
Unio raddei var. kutaisanus Haas, 1940.
‘Kobelt’ Haas, 1940, Zool. Ser. Field Mus. Nat. Hist.: 131.
(error for Unio koutaisiana Kobelt, 1886)
= Unio crassus
 
Taxonomic Opinions
Anodonta apollonica Bourguignat, 1880. — Haas (1940: 138).
Anodonta cygnea (Linnaeus, 1758). — Haas (1940: 136).
Anodonta taurica Bourguignat, 1880. — Haas (1940: 138).
Anodonta vescoiana Bourguignat, 1856. — Haas (1940: 138).
Caelatura [sic] egyptiaca [sic] (Cailliaud, 1827). — Haas (1940: 140).
Diplodon parallelipipedon [sic] (Lea, 1834). — Haas (1940: 141).
Elongaria orientalis (Lea, 1840). — Haas (1940: 141).
Gabillotia pseudodopsis (Locard, 1883). — Haas (1940: 140).
Lampsilis (Disconaias) discus (Lea, 1838). — Haas (1940: 141).
Leguminaia saulcyi (Bourguignat, 1852). — Haas (1940: 133).
Leguminaia wheatleyi (Lea, 1862). — Haas (1940: 133).
Margaritifera auricularia auricularia (Spengler, 1793). — Haas (1940: 119).
Margaritifera auricularia maroccana [sic] (Pallary, 1918). — Haas (1940: 119).
Margaritifera margaritifera dahurica (Middendorff, 1850). — Haas (1940: 120).
Margaritifera margaritifera laevis (Haas, 1910). — Haas (1940: 120).
Margaritifera margaritifera margaritifera (Linnaeus, 1758). — Haas (1940: 120).
Margaritifera margaritifera middendorffi (Rósen, 1926). — Haas (1940: 120).
Microcondylaea compressa (Menke, 1830). — Haas (1940: 133).
Pseudanodonta complanata (Rossmässler, 1835). — Haas (1940: 139).
Pseudanodonta elongata (Holandre, 1836). — Haas (1940: 139).
Pseudanodonta middendorffi compacta (Zelebor in Haidinger, 1851). — Haas (1940: 140).
Pseudanodonta middendorffi middendorffi (Siemaschko, 1848). — Haas (1940: 140).
Pseudodontopsis euphraticus (Bourguignat, 1852). — Haas (1940: 133).
Psilunio acarnanicus (Kobelt, 1879). — Haas (1940: 134).
Psilunio littoralis delesserti (Bourguignat, 1852). — Haas (1940: 135).
Psilunio littoralis fellmanni (DeShayes, 1848). — Haas (1940: 135).
Psilunio littoralis homsensis (Lea, 1864). — Haas (1940: 135).
Psilunio littoralis komarowi (Boettger, O., 1880). — Haas (1940: 135).
Psilunio littoralis littoralis (Lamarck, 1801). — Haas (1940: 134).
Psilunio littoralis semirugatus (Lamarck, 1819). — Haas (1940: 135).
Psilunio littoralis umbonatus (Rossmässler, 1844). — Haas (1940: 134).
Unio abyssinicus von Martens, 1866. — Haas (1940: 124).
Unio crassus batavus (Maton & Rackett, 1807). — Haas (1940: 130).
Unio crassus bruguierianus Bourguignat, 1853. — Haas (1940: 132).
Unio crassus carneus Küster, 1854. — Haas (1940: 131).
Unio crassus crassus Retzius, 1788. — Haas (1940: 129).
Unio crassus cytherea Küster, 1833. — Haas (1940: 129).
Unio crassus gontieri Bourguignat, 1856. — Haas (1940: 131).
Unio crassus jonicus [sic] Drouet, 1879. — Haas (1940: 131).
Unio crassus mongolicus Middendorff, 1851. — Haas (1940: 132).
Unio elongatulus aleroni Companyo & Massot, 1845. — Haas (1940: 126).
Unio elongatulus bourgeticus Bourguignat in Locard, 1882. — Haas (1940: 125).
Unio elongatulus decipiens Drouet, 1881. — Haas (1940: 125).
Unio elongatulus dembeae Sowerby, 1865. — Haas (1940: 128).
Unio elongatulus durieui DeShayes, 1847. — Haas (1940: 128).
Unio elongatulus elongatulus Pfeiffer, C., 1825. — Haas (1940: 124).
Unio elongatulus eucirrus Bourguignat, 1857. — Haas (1940: 128).
Unio elongatulus fiscallianus Kleciach, 1872. — Haas (1940: 125).
Unio elongatulus gargottae Philippi, 1836. — Haas (1940: 127).
Unio elongatulus glaucinus Porro, 1838. — Haas (1940: 127).
Unio elongatulus kruperi Drouet, 1879. — Haas (1940: 125).
Unio elongatulus lawleyanus [sic] Gentiluomo, 1868. — Haas (1940: 127).
Unio elongatulus mancus Lamarck, 1819. — Haas (1940: 126).
Unio elongatulus moquinianus Dupuy, 1843. — Haas (1940: 126).
Unio elongatulus pallens Rossmässler, 1842. — Haas (1940: 124).
Unio elongatulus penchinatianus Bourguignat, 1865. — Haas (1940: 126).
Unio elongatulus quelleneci [sic] Drouet, 1895. — Haas (1940: 125).
Unio elongatulus requienii Michaud, 1831. — Haas (1940: 126).
Unio elongatulus rousii Dupuy, 1849. — Haas (1940: 126).
Unio elongatulus sandrii Rossmässler, 1844. — Haas (1940: 125).
Unio elongatulus turtoni Payraudeau, 1826. — Haas (1940: 125).
Unio elongatulus valentinus Rossmässler, 1854. — Haas (1940: 127).
Unio pictorum ascanius Kobelt, 1913. — Haas (1940: 122).
Unio pictorum delphinus Spengler, 1793. — Haas (1940: 123).
Unio pictorum gaudioni Drouet, 1881. — Haas (1940: 122).
Unio pictorum latirostris Küster, 1854. — Haas (1940: 121).
Unio pictorum middendorffi Westerlund, 1890. — Haas (1940: 122).
Unio pictorum moussonianus Clessin, 1887. — Haas (1940: 122).
Unio pictorum mucidus Morelet, 1845. — Haas (1940: 123).
Unio pictorum pictorum (Linnaeus, 1758). — Haas (1940: 121).
Unio pictorum platyrhynchoideus [sic] Dupuy, 1849. — Haas (1940: 123).
Unio pictorum platyrhynchus Rossmässler, 1835. — Haas (1940: 122).
Unio pictorum praeposterus Küster, 1854. — Haas (1940: 121).
Unio pictorum proëchistus Bourguignat, 1870. — Haas (1940: 122).
Unio pictorum proëchus Bourguignat, 1862. — Haas (1940: 122).
Unio pictorum ravoisieri DeShayes, 1848. — Haas (1940: 123).
Unio pictorum rostratus Lamarck, 1819. — Haas (1940: 122).
Unio pictorum schrenckianus [sic] Clessin, 1880. — Haas (1940: 122).
Unio tigridis terminalis Bourguignat, 1852. — Haas (1940: 124).
Unio tigridis tigridis Bourguignat, 1852. — Haas (1940: 123).
Unio tumidus borysthenensis Kobelt, 1879. — Haas (1940: 129).
Unio tumidus tumidus Retzius, 1788. — Haas (1940: 128).
 
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