|
Bivalvia | Unionoida | Unionoidea | Unionidae subfamily AMBLEMINAE Rafinesque, 1820: cladomics
5 tribes
Cladomics Summary
number of species = 413
number of subtaxa (rank: tribe) = 5
analyzed in 280 cladograms in 167 publications
maximum number of terminals in an analysis = 182, mean = 12.31
maximum number of subtaxa (tribe) in an analysis = 5, mean = 2.58
Basic Monophyly
number of cladograms that test basic monophyly (i.e., more than 1 included species) = 241
maximum number of included terminals = 182, mean = 14.14
maximum number of subtaxa (tribe) = 5, mean = 2.83
proportion of cladograms that support basic monophyly = 0.87
Rigorous Monophyly
number of cladograms that test rigorous monophyly = 169
(Rigorous monophyly refers to cladograms with terminal species representing more than one subtaxon (tribe) taxon and at least one rigorous outgroup (subfamily).
maximum number of included terminals = 182, mean = 11.93
maximum number of subtaxa (tribe) = 5, mean = 3.04
proportion of cladograms that support rigorous monophyly = 0.82
Pairwise Cladogram Clustering
The graphs below depict in two dimensions all pair-wise ingroup distances within the cladome. See the Cladmoics FARQ for more information.
Wu et al., 2023 [Publ 4339], fig. 4, S5-nt
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 17 |
17 |
15 |
5 |
80 |
69 |
3 |
0.0 |
*** Top Monophyly Cladogram ***
+-- Quadrula_quadrula **
+--------------------------+
| +-- Uniomerus_tetralasmus **
|
+-----------+ +-- Elliptio_complanata **
| | +-----------------------+
| | | +-- Pleurobema_oviforme **
| +--+
| | +----------------------- Amblema_plicata **
| | |
| +--+ +-------------------- Popenaias_popeii **
| | |
| +--+ +----------------- Toxolasma_parvum **
| | |
| | | +----- Potamilus_leptodon **
| +--+ +--------+
| | | | +-- Potamilus_alatus **
| | | +--+
| | | +-- Potamilus_streckersoni **
| +--+
| | +----------- Leaunio_lienosus **
| | |
| | | +----- Lampsilis_ornata **
| +--+ +--+
| | | | +-- Lampsilis_cardium **
| | | +--+
| +--+ +-- Lampsilis_cariosa **
| |
| | +----- Venustaconcha_ellipsiformis **
| +--+
| | +-- Lampsilis_powellii **
| +--+
| +-- Lampsilis_siliquoidea **
--+
| +----- Chamberlainia_somsakpanhai
| +-----------------+
| | | +-- Sinohyriopsis_cumingii
| | +--+
| | +-- Sinohyriopsis_schlegelii
| |
| | +-- Hyriopsis_bialata
| | +--+
| +--------------+ | +-- Rectidens_sumatrensis
| | | +--------------+
| | | | | +-- Lens_contradens
| | | | +--+
| | | | +-- Physunio_superbus
| | | |
| | | | +----------- Potomida_littoralis
| | +--+ +-----+
| | | | | +-------- Pronodularia_japanensis
| | | | +--+
| | | | | +----- Lamprotula_leaii
| | | | +--+
| | | | | +-- Lamprotula_caveata
| | +--+ +--+
| | | +-- Lamprotula_cornuumlunae
| | |
| | | +-- Pseudodon_vondembuschianus
| | | +-----------+
| | | | +-- Pilsbryoconcha_exilis
| | | |
| | +--+ +-- Ptychorhynchus_pfisteri
+--+ | +--------+
| | | +-- Microcondylaea_bonellii
| +--+
| | +-------- Parvasolenaia_rivularis
| +--+
| | +----- Sinosolenaia_carinata
| +--+
| | +-- Sinosolenaia_iridinea
| +--+
| +-- Sinosolenaia_oleivora
|
| +----------------------------------- Lamellidens_marginalis
| |
| | +-- Lepidodesma_aligera
| | +-----------------------------+
| | | +-- Lepidodesma_languilati
| | |
| | | +-------- Acuticosta_chinensis
| | | +--------------+
| | | | | +----- Lanceolaria_gladiola
| | | | +--+
+--+ | | | +-- Lanceolaria_grayii
| | | +--+
| | | +-- Lanceolaria_lanceolata
| | |
| | | +-------------- Cristaria_plicata
| | +-----+ |
| | | | +-----+ +-- Anemina_arcaeformis
| | | | | | +--------+
| | | | | | | +-- Anemina_euscaphys
| | | | | +--+
+--+ | | | | +-------- Beringiana_fukuharai
| | | | +--+
| | | | | +----- Sinanodonta_lucida
| | | | +--+
| | +--+ | +-- Sinanodonta_tumens
| | | +--+
| | | +-- Sinanodonta_woodiana
| | |
| | | +----------------- Anodonta_nuttalliana
| | | |
| | | +----------------- Pseudanodonta_complanata
| | | |
| | +--+ +-------------- Anodonta_cygnea
| | | |
| | | | +-- Anodonta_exulcerata
| | | +-----------+
| | | | +-- Anodonta_anatina
| | +--+
+--+ | +----- Pyganodon_grandis
| | +-----+
| | | | +-- Utterbackia_imbecillis
| | | +--+
| +--+ +-- Utterbackia_peninsularis
| |
| | +-------- Lasmigona_complanata
| +--+
| | +----- Prolasmidonta_heterodon
| +--+
| | +-- Alasmidonta_varicosa
| +--+
| +-- Platynaias_compressa
|
| +-------- Aculamprotula_tortuosa
| |
| +-----------------+ +-- Aculamprotula_coreana
| | | +--+
| | | | +-- Aculamprotula_tientsinensis
| | +--+
| | | +-- Aculamprotula_polysticta
| | +--+
| | +-- Aculamprotula_scripta
| |
+--+ +----------- Unio_tumidus
| |
| +-----------+ +-------- Unio_crassus
| | | |
| | +--+ +----- Unio_elongatulus
| | | |
| | +--+ +-- Unio_delphinus
| | | |
| | +--+-- Unio_mancus
+--+ |
| +-- Unio_pictorum
|
| +-- Pseudocuneopsis_capitata
| +-----------------+
| | +-- Pseudocuneopsis_sichuanensis
| |
+--+ +-- Nodularia_breviconcha
| +--------------+
| | +-- Nodularia_nipponensis
| |
+--+ +----- Tchangsinaia_piscicula
| +--------+
| | | +-- Schistodesmus_lampreyanus[2]
| | +--+
+--+ +-- Schistodesmus_spinosus
|
| +----------- Schistodesmus_lampreyanus[1]
+--+
| +-------- Cuneopsis_rufescens
+--+
| +----- Cuneopsis_celtiformis
+--+
| +-- Cuneopsis_demangei
+--+
+-- Cuneopsis_heudei
Huang et al., 2019 [Publ 3843], fig. 2
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 25 |
25 |
20 |
4 |
179 |
140 |
3 |
23.0 |
*** Top Sister-Group Cladogram ***
+----------- Hyriopsis_desowitzi
|
+----------- Rectidens_sumatrensis
|
| +-- Hyriopsis_bialata
| |
| +-- Hyriopsis_myersiana
+-----+--------+
| | +-- Hyriopsis_panhai
| | |
| | +-- Hyriopsis_khoratensis
| |
| | +-- Ensidens_ingallsianus
| | +-----+
| | | +-- Ensidens_telus
| +--+
| | +----- Ensidens_sagittarius
+-----------------+ +--+
| | | +-- Ensidens_spiculus
| | +--+
| | +-- Ensidens_jaculus
| |
| | +----- Physunio
| | +--------+
| | | | +-- Physunio_modelli
| | | +--+
| | | +-- Physunio_superbus
| | |
| +--+ +-------- Yaukthwa_inlenensis
| | +--+
| | | | +----- Yaukthwa_panhai
| | | +--+
| | | | +-- Yaukthwa_nesemanni
| | | +--+
| +--+ +-- Yaukthwa_rectangularis
| |
| | +-- Lens_contradens
| | +-----+
| | | +-- Lens_micropterus
| +--+
| | +----- Lens_eximius
| +--+
| | +-- Lens_comptus
| +--+
| +-- Lens_novoselovi
|
| +-- Megalonaias_nervosa **
| +--+
--+ | +-- Uniomerus_carolinianus **
| +-----------------------+
| | | +-- Quadrula_quadrula **
| | +--+
| | +-- Quadrula_verrucosa **
| |
| | +-------- Elliptoideus_sloatianus **
| +--+ |
| | | +-----------------+ +-- Elliptio_complanata **
| | | | | +--+
| | | | | | +-- Eurynia_dilatata **
| | | | +--+
| | | | | +-- Fusconaia_flava **
| | +--+ +--+
| | | +-- Pleurobema_sintoxia **
| | |
| | | +----------------------- Amblema_plicata **
| | | |
| | | | +----- Toxolasma_parvum **
| | +--+ +--------------+
| | | | | +-- Obliquaria_reflexa **
| | | | +--+
| | | | +-- Plectomerus_dombeyanus **
| | +--+
| | | +----------------- Potamilus_leptodon **
| | | |
| | | | +-------------- Potamilus_alatus **
| | +--+ |
| | | | +----------- Venustaconcha_ellipsiformis **
| | | | |
| | +--+ +----------- Leaunio_lienosus **
| | | |
+--+ | | +-- Ellipsaria_lineolata **
| | +--------+
| +--+ +-- Truncilla_truncata **
| |
| | +-- Medionidus_conradicus **
| +--------+
| | +-- Ptychobranchus_subtentus **
| |
| | +-------- Cambarunio_iris **
| +--+
| | +----- Ortmanniana_ligamentina **
| +--+
| | +-- Lampsilis_cardium **
| +--+
| +-- Lampsilis_ornata **
|
| +----- Chamberlainia_somsakpanhai
| +-----------------------+
| | | +-- Sinohyriopsis_cumingii
| | +--+
| | +-- Sinohyriopsis_schlegelii
| |
| | +-- Potomida_littoralis
| | +-----+
| | | +-- Pronodularia_japanensis
| | +-----------------+
+--+ | | +----- Lamprotula_leaii
| | +--+
| | | +-- Lamprotula_caveata
| | +--+
| | +-- Lamprotula_cornuumlunae
| |
| | +-------- Sinosolenaia_carinata
| | +--+
| | | | +----- Sinosolenaia_recognita
| | | +--+
+--+ | | +-- Sinosolenaia_oleivora
| | +--+
| +-----------+ +-- Sinosolenaia_iridinea
| | |
| | | +----- Gonidea_angulata
| | | +--+
| | | | | +-- Leguminaia_wheatleyi
| | | | +--+
| | +--+ +-- Microcondylaea_bonellii
| | |
| | | +-- Ptychorhynchus_pfisteri
| | | |
+--+ +-----+-- Parvasolenaia_rivularis
| |
| +-- Koreosolenaia_sitgyensis
|
| +----- Indopseudodon_salwenianus
| +--------------+
| | | +-- Indopseudodon_indawgyiensis
| | +--+
| | +-- Indopseudodon_bogani
| |
| | +-------- Pilsbryoconcha_exilis[3]
+--+ +--------+
| | | +----- Pilsbryoconcha_exilis[2]
| | +--+
| | | +-- Pilsbryoconcha_exilis[1]
| | +--+
| | +-- Pilsbryoconcha_carinifera
| |
+--+ +----------- Nyeinchanconcha_nyeinchani
| +--+
| | | +-------- Thaiconcha_callifera
| | +--+
| | | +----- Bineurus_loeiensis
| | +--+
| | | +-- Bineurus_mouhotii
+--+ +--+
| +-- Bineurus_exilis
|
| +----- Pseudodon_vagulus
| +--+
| | | +-- Pseudodon_mekongi
| | +--+
+-----+ +-- Pseudodon_vondembuschianus
|
| +-- Sundadontina_cumingii
| |
+-----+-- Sundadontina_tumida[1]
|
+-- Sundadontina_tumida[2]
Inoue et al., 2018 [Publ 3858], fig. 1, 3a-c
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 182 |
182 |
62 |
5 |
23 |
22 |
2 |
187.0 |
+-- Alasmidonta_marginata
|
+-- Prolasmidonta_heterodon
|
+-- Alasmidonta_undulata
|
+-- Beringiana_beringiana
|
+-- Anodonta_kennerlyi
|
+-- Anodonta_nuttalliana
|
+-- Anodonta_oregonensis
|
+-- Arcidens_confragosus
|
+-- Arcidens_wheeleri
|
+-- Lasmigona_complanata
|
+--------------------------------------------------+-- Platynaias_compressa
| |
| +-- Lasmigona_costata
| |
| +-- Platynaias_decorata
| |
| +-- Platynaias_subviridis
| |
| +-- Pyganodon_grandis
| |
| +-- Pyganodon_lacustris
| |
| +-- Pseudodontoideus_subvexus
| |
| +-- Strophitus_undulatus
| |
| +-- Utterbackia_imbecillis
| |
--+ +-- Utterbackia_peninsularis[1]
| |
| +-- Utterbackia_peninsularis[2]
|
| +-------------------------------------------------- Gonidea_angulata
| |
| | +-- Reginaia_ebenus **
| | +--------------------------------------------+
| | | +-- Reginaia_rotulata **
| | |
| | | +-- Amblema_plicata **
| | +--------------------------------------------+
| | | +-- Amblema_elliottii **
| | |
| | | +----- Popenaias_popeii **
| | | |
| | | | +-- Dromus_dromas **
| | | | |
+--+ | | +-- Epioblasma_rangiana **
| | | |
| +-----------------------------------------+ +-- Glebula_rotundata **
| | | |
| | | +-- Lampsilis_ovata **
| | | |
| | | +-- Lampsilis_virescens **
| | | |
| | | +-- Lemiox_rimosus **
| | +--+
| | +-- Obliquaria_reflexa **
| | |
| | +-- Obovaria_olivaria **
| | |
+--+ +-- Obovaria_unicolor **
| |
| +-- Plectomerus_dombeyanus **
| |
| +-- Potamilus_alatus **
| |
| +-- Potamilus_capax **
|
| +-- Uniomerus_declivis **
| +--+
| | +-- Uniomerus_tetralasmus **
| |
| | +-- Cyclonaias_tuberculata **
| +--------------------------------------+ |
| | | +-- Pustulosa_pustulosa[1] **
| | | |
| | | +-- Theliderma_metanevra **
| | +--+
| | +-- Pustulosa_pustulosa[2] **
| | |
| | +-- Quadrula_quadrula **
| | |
| | +-- Quadrula_verrucosa **
| |
| | +----------------------------------------- Elliptoideus_sloatianus **
+--+ |
| | +-- Elliptio_lanceolata **
| | +-----------------------------------+
| | | +-- Parvaspina_collina **
| | |
| | | +----------------------------------- Plethobasus_cyphyus **
| | | |
| | | +----------------------------------- Eurynia_dilatata **
| | | |
| | | +----------------------------------- Hemistena_lata **
| | | |
| | | | +-- Elliptio_complanata **
+--+ | | |
| | | +-- Elliptio_hopetonensis **
| | | |
| | | +-- Elliptio_producta[1] **
| | | |
| | | +-- Elliptio_shepardiana **
| | | |
| | | +-- Elliptio_angustata **
| | | |
| | | +-- Elliptio_arca **
| | | |
| | | +-- Elliptio_arctata **
| | | |
+--+ | +-- Elliptio_jayensis **
| | |
| | +-- Elliptio_crassidens **
| | |
| +--------------------------------+-- Elliptio_dariensis **
| | |
| | +-- Elliptio_fisheriana **
| | |
| | +-- Elliptio_folliculata **
| | |
| | +-- Elliptio_fumata **
| | |
| | +-- Elliptio_icterina **
| | |
| | +-- Elliptio_mcmichaeli **
| | |
| | +-- Elliptio_producta[2] **
| | |
| | +-- Elliptio_pullata **
| | |
| | +-- Elliptio_purpurella **
| | |
| | +-- Elliptio_congaraea **
| |
+--+ +-- Pleuronaia_barnesiana **
| |
| +-----------------------------+-- Pleuronaia_dolabelloides **
| | |
| | +-- Pleuronaia_gibber **
| |
| | +-------------------------- Pleurobema_plenum **
| | |
| | | +----------------------- Pleurobema_stabile **
| | | |
| | | | +----- Pleurobema_pyriforme **
| | +--+ | +--+
| | | | | | | +-- Pleurobema_georgianum[2] **
| | | | | | +--+
| | | | | +-----------+ +-- Pleurobema_athearni **
| | | | | | |
| | | +--+ | | +----- Pleurobema_decisum **
| | | | | | |
| | | | | +--+ +-- Pleurobema_georgianum[1] **
| | | | | | |
| | | | | +--+-- Pleurobema_hanleyianum[1] **
| | | | | |
| | | | | +-- Pleurobema_hanleyianum[2] **
| | | | |
| | | +--+ +-------------- Pleurobema_beadleianum[2] **
| | | | |
| | | | +--+ +----------- Pleurobema_strodeanum[2] **
| | | | | | |
| | | | | +--+----------- Pleurobema_rubellum **
| | | | | |
| | | | | | +-------- Pleurobema_beadleianum[1] **
| | | | | +--+
| | | | | | +----- Pleurobema_taitianum **
| | | | | +--+
| | | | | | +-- Pleurobema_perovatum **
| | | +--+ +--+
| | | | +-- Pleurobema_strodeanum[1] **
| | | |
| | | | +----- Pleurobema_oviforme[6] **
| | | | |
| | | | +--+----- Pleurobema_clava[3] **
| | | | | |
| | | | | | +-- Pleurobema_oviforme[5] **
| | | | | +--+
| | | +--------+ +-- Pleurobema_clava[2] **
+--+ | |
| | | +----- Pleurobema_oviforme[2] **
| | | |
| | | +----- Pleurobema_oviforme[3] **
| | +--+
| | +----- Pleurobema_oviforme[4] **
| | |
| | | +-- Pleurobema_clava[1] **
| | +--+
| | +-- Pleurobema_oviforme[1] **
| |
| | +-------------------------- Pleurobema_cordatum **
| | |
| | | +----- Fusconaia_subrotunda **
| | | +-----------------+
| +--+ | | +-- Fusconaia_cor **
| | | | +--+
| | | | +-- Fusconaia_cuneolus **
| | | |
| | | | +-------------------- Fusconaia_ozarkensis **
| | +--+ |
| | | | +-- Fusconaia_escambia **
| | | | +--------------+
| | | | | +-- Fusconaia_burkei **
| | | | |
| | | | | +----- Fusconaia_flava[27] **
| | | | | |
| | +--+ | | +-- Fusconaia_iheringi **
| | | | +--+
| | | | | +-- Fusconaia_mitchelli **
| | | +-----------+
| | | | | +-- Fusconaia_askewi[1] **
| | | | | |
| | | | | +-- Fusconaia_askewi[2] **
| | | | | |
| | +--+ | +-- Fusconaia_askewi[3] **
| | | +--+
| | | +-- Fusconaia_askewi[4] **
| | | |
| | | +-- Fusconaia_askewi[5] **
| | | |
| | | +-- Fusconaia_chunii **
| | |
+--+ | +-------------- Fusconaia_masoni **
| | |
| | | +----------- Fusconaia_flava[26] **
| | | |
| | | | +-- Fusconaia_cerina[8] **
| +--+ | +-----+
| | | | +-- Fusconaia_flava[25] **
| | | |
| | | | +----- Fusconaia_flava[5] **
| | | | |
| | | | +----- Fusconaia_flava[6] **
| +--+ | |
| | | +----- Fusconaia_flava[7] **
| | | |
| | | +----- Fusconaia_flava[8] **
| | | |
| | | +----- Fusconaia_cerina[4] **
| | | |
| | | +----- Fusconaia_flava[9] **
| | | |
| +--+ +----- Fusconaia_flava[10] **
| | |
| | +----- Fusconaia_flava[11] **
| | |
| | +----- Fusconaia_flava[12] **
| | |
| | +----- Fusconaia_flava[13] **
| | |
| | +----- Fusconaia_flava[14] **
| | |
| | +----- Fusconaia_flava[15] **
| | |
| | +----- Fusconaia_flava[16] **
| | |
| | +----- Fusconaia_flava[17] **
| +--+
| +----- Fusconaia_flava[18] **
| |
| +----- Fusconaia_flava[19] **
| |
| +----- Fusconaia_flava[20] **
| |
| +----- Fusconaia_flava[21] **
| |
| +----- Fusconaia_flava[22] **
| |
| +----- Fusconaia_flava[23] **
| |
| +----- Fusconaia_flava[24] **
| |
| +----- Fusconaia_cerina[5] **
| |
| +----- Fusconaia_cerina[6] **
| |
| +----- Fusconaia_cerina[7] **
| |
| | +-- Fusconaia_flava[1] **
| | |
| | +-- Fusconaia_flava[2] **
| | |
| | +-- Fusconaia_flava[3] **
| | |
| +--+-- Fusconaia_flava[4] **
| |
| +-- Fusconaia_cerina[1] **
| |
| +-- Fusconaia_cerina[2] **
| |
| +-- Fusconaia_cerina[3] **
|
| +----------------- Pleurobema_riddellii **
| |
| | +-------------- Pleurobema_sintoxia[21] **
| | |
| | | +----------- Pleurobema_sintoxia[19] **
| | | |
| | | +----------- Pleurobema_sintoxia[20] **
| | | |
+-----------+ | | +----- Pleurobema_sintoxia[18] **
| | | |
| | | +----- Pleurobema_rubrum[19] **
| | | |
| | | +----- Pleurobema_rubrum[20] **
| | +-----+
| | | +----- Pleurobema_rubrum[21] **
| | | |
| | | | +-- Pleurobema_rubrum[17] **
+--+ | | |
| | | +-- Pleurobema_sintoxia[16] **
| | +--+
| | +-- Pleurobema_sintoxia[17] **
| | |
| | +-- Pleurobema_rubrum[18] **
| |
| | +-------- Pleurobema[4] **
| | |
| | +-------- Pleurobema[5] **
| | |
| | +-------- Pleurobema_sintoxia[1] **
| | |
| | +-------- Pleurobema[8] **
| | |
| | +-------- Pleurobema[9] **
+--+ |
| +-------- Pleurobema[10] **
| |
| +-------- Pleurobema_sintoxia[3] **
| |
| +-------- Pleurobema_sintoxia[4] **
| |
| +-------- Pleurobema_sintoxia[5] **
| |
| +-------- Pleurobema_sintoxia[6] **
| |
| +-------- Pleurobema_sintoxia[7] **
| |
| +-------- Pleurobema_sintoxia[8] **
| |
| +-------- Pleurobema_sintoxia[9] **
| |
| +-------- Pleurobema_sintoxia[10] **
| |
| +-------- Pleurobema_sintoxia[11] **
| |
| +-------- Pleurobema_sintoxia[12] **
| |
| +-------- Pleurobema_sintoxia[13] **
| |
| +-------- Pleurobema_sintoxia[14] **
| |
| +-------- Pleurobema_sintoxia[15] **
| |
+--+-------- Pleurobema_rubrum[4] **
|
+-------- Pleurobema_rubrum[5] **
|
+-------- Pleurobema_rubrum[6] **
|
+-------- Pleurobema_rubrum[7] **
|
+-------- Pleurobema_rubrum[8] **
|
+-------- Pleurobema_rubrum[9] **
|
+-------- Pleurobema_rubrum[10] **
|
+-------- Pleurobema_rubrum[11] **
|
+-------- Pleurobema_rubrum[12] **
|
+-------- Pleurobema_rubrum[13] **
|
+-------- Pleurobema_rubrum[14] **
|
+-------- Pleurobema_rubrum[15] **
|
+-------- Pleurobema_rubrum[16] **
|
| +-- Pleurobema[6] **
+-----+
| +-- Pleurobema_rubrum[1] **
|
| +-- Pleurobema_sintoxia[2] **
+-----+
| +-- Pleurobema_rubrum[2] **
|
| +-- Pleurobema[7] **
+-----+
| +-- Pleurobema_rubrum[3] **
|
| +----- Pleurobema[3] **
+--+
| +-- Pleurobema[1] **
+--+
+-- Pleurobema[2] **
Campbell et al., 2005 [Publ 2660], fig. 1
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 105 |
105 |
93 |
5 |
12 |
10 |
2 |
103.0 |
+-------------------------------------------------- Sinohyriopsis_cumingii
|
| +----------- Anodonta_oregonensis
| |
+--------------------------------------+ +-- Anodonta_cygnea
| | +-----+
| | | +-- Alasmidonta_holstonia
| +--+
| | +----- Pyganodon_grandis
| +--+
| | +-- Lasmigona_costata
| +--+
| +-- Pseudodontoideus_subvexus
|
| +----- Gonidea_angulata
| +-----------------------------------------+
| | | +-- Potomida_littoralis
--+ | +--+
| | +-- Pronodularia_japanensis
| |
| | +-- Megalonaias_nervosa **
| | +-----------+
| | | +-- Uniomerus_declivis **
| | |
| | +-----------------------------+ +----------- Theliderma_metanevra **
| | | | |
| | | | | +----- Quadrula_verrucosa **
| | | +--+ +--+
| | | | | | +-- Quadrula_quadrula **
| | | | | +--+
| | | | | +-- Quadrula_nobilis **
| | | +--+
| | | | +----- Pustulosa_kieneriana **
| | | | |
| | | | | +-- Cyclonaias_tuberculata **
+--+ | +--+--+
| | | +-- Pustulosa_succissa **
| | |
| | | +-- Pustulosa_infucata **
| | +--+
| | +-- Pustulosa_kleiniana **
| |
| | +----------------------------------- Elliptoideus_sloatianus **
| | |
| | | +-------------------------------- Hemistena_lata **
| | | |
| | +-----+ | +----- Plethobasus_cyphyus **
| | | | | +-----------------------+
| | | | | | | +-- Elliptio_arca **
| | | | | | +--+
| | | | | | +-- Elliptio_crassidens **
| | | +--+ |
| | | | | +-- Eurynia_dilatata **
| | | | | +-----------------------+
| | | | | | +-- Parvaspina_collina **
+--+ | | | |
| | | | | +----- Pleuronaia_gibber **
| | | | +--------------------+
| | | | | | +-- Pleuronaia_barnesiana **
| | +--+ | +--+
| | | | +-- Pleuronaia_dolabelloides **
| | | |
| | | | +----------- Fusconaia_subrotunda **
| | | | |
| | | | | +-- Fusconaia_cor **
| | | +--------------+ +-----+
| | | | | | +-- Fusconaia_cuneolus **
| | | | | |
| | | | | | +-- Fusconaia_cerina[1] **
| | | | +--+ |
| | | | | +--+-- Fusconaia_cerina[2] **
| | +--+ | | |
| | | | | +-- Fusconaia_flava **
| | | +--+
| | | | +-- Fusconaia_burkei[1] **
| | | | |
| | | | +-- Fusconaia_burkei[2] **
| | | +--+
| | | +-- Fusconaia_burkei[3] **
| | | |
| | | +-- Fusconaia_escambia **
| | |
| | | +-- Pleurobema_rubrum[2] **
| | | +--+
| | | | +-- Pleurobema_cordatum **
| | | +-----------------+
| | | | | +-- Pleurobema_rubrum[1] **
+--+ | | +--+
| | | +-- Pleurobema_sintoxia **
| +--+
| | +-------------------- Pleurobema_pyriforme **
| | |
| | | +-- Pleurobema_georgianum[2] **
| | | |
| +--+ +--------------+-- Pleurobema_georgianum[3] **
| | | |
| | | +-- Pleurobema_athearni **
| | |
| | | +----- Pleurobema_decisum **
| +--+ |
| | +--------+ +-- Pleurobema_hanleyianum[1] **
| | | | |
| | | +--+-- Pleurobema_hanleyianum[2] **
| | | |
| | | +-- Pleurobema_georgianum[1] **
| +--+
| | +-- Pleurobema_oviforme[1] **
| | |
| | +--------+-- Pleurobema_oviforme[2] **
| | | |
| +--+ +-- Pleurobema_clava **
| |
| | +-------- Pleurobema_strodeanum **
| +--+
| | +----- Pleurobema_rubellum **
| +--+
| | +-- Pleurobema_perovatum **
| +--+
| +-- Pleurobema_taitianum **
|
| +----- Plectomerus_dombeyanus **
| +--------------------------------+
| | | +-- Reginaia_ebenus **
| | +--+
| | +-- Reginaia_rotulata **
| |
| | +----- Popenaias_popeii **
| | +-----------------------------+
| | | | +-- Amblema_elliottii **
+--+ | +--+
| | +-- Amblema_plicata **
| |
| | +-- Toxolasma_parvum **
| | +-----+
| | | +-- Toxolasma_texasiense **
| | +-----------------------+
| | | | +----- Obliquaria_reflexa **
+--+ | +--+
| | | +-- Cyrtonaias_tampicoensis **
| | +--+
| | +-- Glebula_rotundata **
| |
| | +-- Potamilus_fragilis **
| | +--+
| | | +-- Potamilus_leptodon **
| | +-----------+
| | | | +-- Potamilus_alatus **
| | | +--+
+--+ | +-- Potamilus_purpuratus **
| |
| +-----------+ +-- Sagittunio_nasutus **
| | | +--+
| | | | +-- Medionidus_conradicus **
| | | +--------+
| | | | | +-- Ptychobranchus_fasciolaris **
| | | | +--+
| | +--+ +-- Venustaconcha_ellipsiformis[2] **
| | |
| | | +----------- Medionidus_acutissimus **
| | +--+
| | | +-------- Lemiox_rimosus **
| | +--+
+--+ | +----- Dromus_dromas **
| +--+
| | +-- Cyprogenia_aberti **
| +--+
| +-- Cyprogenia_stegaria **
|
| +-- Ellipsaria_lineolata **
| +-----------------------+
| | +-- Truncilla_truncata **
| |
| | +----------------------- Villosa_villosa **
| | |
| | | +-- Obovaria_olivaria **
+--+ | +--------------+
| | | +-- Venustaconcha_ellipsiformis[1] **
| | |
| | +--+ +----- Obovaria_unicolor **
| | | | +--------+
| | | | | | +-- Obovaria_arkansasensis **
+--+ | | | +--+
| | +--+ +-- Obovaria_subrotunda **
| | |
| | | +----------- Epioblasma_brevidens **
| | +--+
| | | +-------- Epioblasma_triquetra **
| | +--+
| | | +----- Epioblasma_walkeri[2] **
| | +--+
| | | +-- Epioblasma_capsaeformis **
+--+ +--+
| +-- Epioblasma_walkeri[1] **
|
| +-- Ligumia_recta **
| +--------+
| | +-- Cambarunio_iris **
| +--+
| | | +-------- Lampsilis_teres **
| | +--+
| | | +----- Ortmanniana_ligamentina **
| | +--+
| | | +-- Lampsilis_siliquoidea **
| | +--+
| | +-- Leaunio_vanuxemensis **
+-----+
| +-- Lampsilis_cardium **
| +--+
| | +-- Lampsilis_ovata **
| +-----+
| | | +-- Ortmanniana_pectorosa **
| | +--+
| | +-- Lampsilis_ornata **
| |
+--+ +----- Hamiota_australis[2] **
| +--+
| | | +-- Hamiota_australis[1] **
| | +--+
| | +-- Hamiota_subangulata **
+--+
| +----- Hamiota_altilis[2] **
| |
| | +-- Hamiota_altilis[1] **
+--+--+
| +-- Hamiota_perovalis[1] **
|
| +-- Hamiota_altilis[3] **
+--+
+-- Hamiota_perovalis[2] **
Campbell et al., 2005 [Publ 2660], fig. 2
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 103 |
103 |
82 |
5 |
12 |
10 |
2 |
100.0 |
+----------- Sinohyriopsis_cumingii
|
+--------------------------------+ +-------- Pyganodon_grandis
| | |
| | | +-- Lasmigona_costata
| +--+-----+
| | +-- Pseudodontoideus_subvexus
| |
| | +----- Alasmidonta_holstonia
| +--+
| | +-- Anodonta_cygnea
| +--+
| +-- Anodonta_oregonensis
|
| +----- Gonidea_angulata
| +-----------------------------------+
| | | +-- Potomida_littoralis
| | +--+
--+ | +-- Pronodularia_japanensis
| |
| | +-------------------- Megalonaias_nervosa **
| | |
| | +-----------------+ +----------------- Uniomerus_declivis **
| | | | |
| | | +--+ +-------------- Theliderma_metanevra **
| | | | |
| | | +--+ +----------- Quadrula_quadrula **
| | | | |
| | | | | +-- Quadrula_nobilis **
| | | +--+ +-----+
| | | | | +-- Quadrula_verrucosa **
| | | | |
| | | +--+ +----- Pustulosa_kieneriana **
| | | | |
+--+ | | | +-- Cyclonaias_tuberculata **
| | +--+--+
| | | +-- Pustulosa_succissa **
| | |
| | | +-- Pustulosa_infucata **
| | +--+
| | +-- Pustulosa_kleiniana **
| |
| | +-------------------------------- Elliptoideus_sloatianus **
| | |
| | | +----------------------------- Plethobasus_cyphyus **
| | | |
| | | +----------------------------- Parvaspina_collina **
| | +--+ |
| | | | | +-- Elliptio_arca **
| | | | +--------------------------+
| | | | | +-- Elliptio_crassidens **
| | | | |
| | | | | +-------------------------- Eurynia_dilatata **
| | | +--+ |
+--+ | | +-------------------------- Hemistena_lata **
| | | |
| | | | +-- Pleurobema_rubrum[1] **
| | | | +--+
| | | | | +-- Pleurobema_sintoxia **
| | | | +-----------------+
| | | | | | +-- Pleurobema_rubrum[2] **
| | | | | +--+
| | +--+ | +-- Pleurobema_cordatum **
| | | |
| | | | +-------------- Fusconaia_subrotunda **
| | | | |
| | | | | +-- Fusconaia_cor **
| | | +--------+ +--------+
| | | | | | +-- Fusconaia_cuneolus **
| | | | | |
| | | | +--+ +-- Fusconaia_cerina **
| | | | | +-----+
| | | | | | +-- Fusconaia_flava **
| | +--+ +--+
| | | | +----- Fusconaia_burkei[2] **
| | | +--+
| | | | +-- Fusconaia_escambia **
| | | +--+
| | | +-- Fusconaia_burkei[1] **
| | |
| | | +----- Pleuronaia_gibber **
| | | +--------------+
| | | | | +-- Pleuronaia_barnesiana **
| | | | +--+
| | | | +-- Pleuronaia_dolabelloides **
| | | |
| | | | +-------- Pleurobema_rubellum **
+--+ +--+ +--------+
| | | | +----- Pleurobema_strodeanum **
| | | +--+
| | | | +-- Pleurobema_perovatum **
| | | +--+
| | | +-- Pleurobema_taitianum **
| +--+
| | +-- Pleurobema_oviforme[1] **
| | |
| | +-----------+-- Pleurobema_oviforme[2] **
| | | |
| | | +-- Pleurobema_clava **
| +--+
| | +----------- Pleurobema_pyriforme **
| | |
| +--+ +-- Pleurobema_athearni **
| | +-----+
| | | +-- Pleurobema_georgianum[2] **
| +--+
| | +----- Pleurobema_decisum **
| | |
| +--+ +-- Pleurobema_hanleyianum[1] **
| | |
| +--+-- Pleurobema_hanleyianum[2] **
| |
| +-- Pleurobema_georgianum[1] **
|
| +-------- Popenaias_popeii **
| |
| +--------------+ +-- Amblema_elliottii **
| | | +--+
| | | | +-- Amblema_plicata **
| | +--+
| | | +-- Reginaia_ebenus **
| | | |
| | +--+-- Reginaia_rotulata[1] **
| | |
| | +-- Reginaia_rotulata[2] **
| |
| | +----- Plectomerus_dombeyanus **
| | +--------------+
+-----------+ | | +-- Toxolasma_parvum **
| | +--+
| | +-- Toxolasma_texasiense **
| |
| | +----- Obliquaria_reflexa **
| | +-----------+
| | | | +-- Cyrtonaias_tampicoensis **
| | | +--+
| | | +-- Glebula_rotundata **
| | |
| | | +-------------- Sagittunio_nasutus **
| | | |
+--+ | +-------------- Leaunio_vanuxemensis **
| | |
| | | +-- Ellipsaria_lineolata **
| | +-----------+
| | | +-- Truncilla_truncata **
| | |
| | | +----- Lampsilis_teres **
| | +--------+
| | | | +-- Ortmanniana_ligamentina **
| | | +--+
| | | +-- Lampsilis_siliquoidea **
| | |
| | | +----- Villosa_villosa **
| | +--------+
+--+ | | +-- Ligumia_recta **
| | +--+
| | +-- Cambarunio_iris **
| |
| | +-------- Ortmanniana_pectorosa **
| +-----+
| | | +----- Lampsilis_ornata **
| | +--+
| | | +-- Lampsilis_cardium **
| | +--+
| | +-- Lampsilis_ovata **
| |
| | +-- Potamilus_fragilis **
| | +--+
| | | +-- Potamilus_leptodon **
| +--------+
| | | +-- Potamilus_alatus **
| | +--+
| | +-- Potamilus_purpuratus **
| |
| | +-- Ptychobranchus_fasciolaris **
+--+ +--------+
| | +-- Venustaconcha_ellipsiformis[1] **
| |
+--+ +-------- Medionidus_conradicus **
| | |
| | | +-- Lemiox_rimosus **
| +--+-----+
| | +-- Medionidus_acutissimus **
| |
| | +----- Dromus_dromas **
| +--+
| | +-- Cyprogenia_aberti **
| +--+
| +-- Cyprogenia_stegaria **
|
| +----- Hamiota_australis[2] **
| +-----+
| | | +-- Hamiota_australis[1] **
| | +--+
| | +-- Hamiota_subangulata **
+--+
| | +-- Hamiota_altilis[3] **
| | +-----+
| | | +-- Hamiota_perovalis[2] **
| +--+
| | +----- Hamiota_altilis[2] **
| +--+
| | +-- Hamiota_altilis[1] **
| +--+
| +-- Hamiota_perovalis[1] **
|
| +----------- Obovaria_olivaria **
| |
| +----------- Venustaconcha_ellipsiformis[2] **
| |
| | +----- Obovaria_unicolor **
| +-----+
+--+ | +-- Obovaria_arkansasensis **
| +--+
| +-- Obovaria_subrotunda **
|
| +-- Epioblasma_brevidens **
| +-----+
| | +-- Epioblasma_triquetra **
+--+
| +----- Epioblasma_walkeri[1] **
+--+
| +-- Epioblasma_walkeri[2] **
+--+
+-- Epioblasma_capsaeformis **
Campbell & Lydeard, 2012 [Publ 3309], fig. 1-2
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 74 |
74 |
64 |
5 |
0 |
0 |
0 |
76.0 |
+-- Uniomerus_declivis **
+-----------------+
| +-- Uniomerus_tetralasmus **
+--------------------+
| | +----------------- Theliderma_metanevra **
| | |
| +--+ +-------------- Quadrula_verrucosa **
| | |
| | | +-- Quadrula_quadrula **
| +--+ +--------+
| | | +-- Quadrula_nobilis **
| | |
| +--+ +-- Pustulosa_infucata **
| | +-----+
| | | +-- Pustulosa_kleiniana **
| +--+
| | +----- Cyclonaias_tuberculata **
| +--+
| | +-- Pustulosa_pustulosa **
| +--+
| +-- Pustulosa_succissa **
|
| +----- Plectomerus_dombeyanus **
| +-----------------+
| | | +-- Reginaia_ebenus **
| | +--+
--+ | +-- Reginaia_rotulata **
| |
| | +----- Popenaias_popeii **
| +--------------+ +--------------+
| | | | | +-- Amblema_elliottii **
| | | | +--+
| | | | +-- Amblema_plicata **
| | | |
| | | | +----------------- Potamilus_alatus **
| | +--+ |
| | | +----------------- Potamilus_capax **
| | | |
| | | +----------------- Glebula_rotundata **
| | | |
| | | | +-- Toxolasma_lividum **
| | | +--------------+
| | +--+ +-- Paetulunio_fabalis **
| | |
| | | +-- Dromus_dromas **
| | +--------------+
| | | +-- Lemiox_rimosus **
| | |
| | | +-- Lampsilis_ovata **
+--+ | +-----------+
| | | +-- Obliquaria_reflexa **
| +--+
| | +----------- Lampsilis_virescens **
| | |
| +--+ +-- Epioblasma_rangiana **
| | +-----+
| | | +-- Obovaria_arkansasensis **
| +--+
| | +----- Obovaria_olivaria **
| +--+
| | +-- Venustaconcha_ellipsiformis **
| +--+
| +-- Venustaconcha_pleasii **
|
| +----------------------------------- Elliptoideus_sloatianus **
| |
| | +-------------------------------- Pleurobema_stabile **
| | |
| | | +-- Pleurobema_clava **
| | | +-----------+
+--+ | | +-- Pleurobema_oviforme **
| | |
| | +--------------+ +----------- Pleurobema_pyriforme **
| | | | |
| | | | | +-- Pleurobema_athearni **
| | | | | +--+
+--+ | +--+ | +-- Pleurobema_georgianum **
| | | +--+
| | | | | +-- Pleurobema_decisum **
| | | | +--+
| | | | +-- Pleurobema_hanleyianum **
| | +--+
| | | +----- Pleurobema_beadleianum **
| | | |
| | | +----- Pleurobema_strodeanum **
| | +--+
| | +----- Pleurobema_rubellum **
+--+ |
| | +-- Pleurobema_perovatum **
| +--+
| +-- Pleurobema_taitianum **
|
| +-------- Pleurobema_riddellii **
| +-----------------+
| | | +----- Pleurobema_cordatum **
| | +--+
| | | +-- Pleurobema_rubrum **
| | +--+
| | +-- Pleurobema_sintoxia **
| |
| | +----------------- Fusconaia_subrotunda **
| | |
+--+ +-----+ +-- Fusconaia_cor **
| | | +-----------+
| | | | +-- Fusconaia_cuneolus **
| | +--+
| | | +----------- Fusconaia_ozarkensis **
| | | |
| | +--+ +-- Fusconaia_escambia **
| | | +-----+
| | | | +-- Fusconaia_burkei **
| | +--+
+--+ | +----- Fusconaia_masoni **
| +--+
| | +-- Fusconaia_flava **
| +--+
| +-- Fusconaia_cerina **
|
| +----- Pleuronaia_gibber **
| +--------------+
| | | +-- Pleuronaia_barnesiana **
| | +--+
| | +-- Pleuronaia_dolabelloides **
+--+
| +-- Eurynia_dilatata **
| +--------------+
| | +-- Parvaspina_collina **
| |
+--+ +-- Hemistena_lata **
| +-----------+
| | +-- Plethobasus_cyphyus **
| |
+--+ +----------- Elliptio_shepardiana **
| |
| | +-------- Elliptio_complanata **
+--+ |
| | +----- Elliptio_purpurella **
| +--+
| | | +-- Elliptio_arca **
+--+ +--+
| +-- Elliptio_arctata **
|
| +-- Elliptio_mcmichaeli **
| |
+-----+-- Elliptio_pullata **
|
+-- Elliptio_crassidens **
Campbell & Lydeard, 2012 [Publ 3309], fig. 3
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 74 |
74 |
68 |
5 |
0 |
0 |
0 |
73.0 |
+-- Uniomerus_declivis **
+--------------+
| +-- Uniomerus_tetralasmus **
|
+--------------------+ +-- Quadrula_verrucosa **
| | +-----------+
| | | +-- Quadrula_nobilis **
| | |
| +--+ +----------- Quadrula_quadrula **
| | |
| | +----------- Theliderma_metanevra **
| +--+
| | +-- Pustulosa_infucata **
| | +-----+
| | | +-- Pustulosa_kleiniana **
| +--+
| | +----- Cyclonaias_tuberculata **
| +--+
| | +-- Pustulosa_pustulosa **
| +--+
| +-- Pustulosa_succissa **
|
| +-- Amblema_elliottii **
| +--+
| | +-- Amblema_plicata **
| +--------------------------+
| | | +-- Reginaia_ebenus **
--+ | +--+
| | +-- Reginaia_rotulata **
| +--+
| | | +----------------------------- Popenaias_popeii **
| | | |
| | | | +-- Glebula_rotundata **
| | +--+ +-----------------------+
| | | | +-- Paetulunio_fabalis **
| | | |
| | +--+ +-- Toxolasma_lividum **
| | | +--------------------+
| | | | +-- Plectomerus_dombeyanus **
| | | |
| | +--+ +-------------------- Obliquaria_reflexa **
| | | |
| | | | +-- Dromus_dromas **
| | | | +--+
| | +--+ | +-- Lemiox_rimosus **
| | | +-----------+
| | | | | +-- Potamilus_alatus **
| | | | +--+
| | +--+ +-- Potamilus_capax **
| | |
| | | +-------------- Lampsilis_ovata **
+--+ | |
| +--+ +----------- Lampsilis_virescens **
| | |
| +--+ +-- Epioblasma_rangiana **
| | +-----+
| | | +-- Obovaria_arkansasensis **
| +--+
| | +----- Obovaria_olivaria **
| +--+
| | +-- Venustaconcha_ellipsiformis **
| +--+
| +-- Venustaconcha_pleasii **
|
| +-------------------------------- Elliptoideus_sloatianus **
| |
| | +-- Eurynia_dilatata **
| | +--------------------------+
| | | +-- Parvaspina_collina **
| | |
| | | +-- Hemistena_lata **
| | +--------------------------+
| | | +-- Plethobasus_cyphyus **
+--+ |
| | +----------- Elliptio_shepardiana **
| | |
| | | +----- Elliptio_crassidens **
| +-----------------+ +--+
| | | | | +-- Elliptio_mcmichaeli **
| | | | +--+
| | +--+ +-- Elliptio_pullata **
| | |
| | | +----- Elliptio_complanata **
+--+ | |
| +--+ +-- Elliptio_purpurella **
| | |
| +--+-- Elliptio_arca **
| |
| +-- Elliptio_arctata **
|
| +----- Pleuronaia_gibber **
| +--------------------+
| | | +-- Pleuronaia_barnesiana **
| | +--+
| | +-- Pleuronaia_dolabelloides **
| |
| | +-------------- Fusconaia_ozarkensis **
| | |
| | | +----- Fusconaia_subrotunda **
| | +--------+ +-----+
+--+ | | | | +-- Fusconaia_cor **
| | | | +--+
| | | | +-- Fusconaia_cuneolus **
| | +--+
| | | +-- Fusconaia_escambia **
| | | +-----+
| | | | +-- Fusconaia_burkei **
| | +--+
| | | +----- Fusconaia_masoni **
+--+ +--+
| | +-- Fusconaia_cerina **
| +--+
| +-- Fusconaia_flava **
|
| +-------- Pleurobema_riddellii **
| +-----------+
| | | +----- Pleurobema_cordatum **
| | +--+
| | | +-- Pleurobema_rubrum **
| | +--+
+--+ +-- Pleurobema_sintoxia **
|
| +----------------- Pleurobema_stabile **
| |
| | +----------- Pleurobema_strodeanum **
| | +--+
+--+ | | +-------- Pleurobema_rubellum **
| | +--+
| | | +----- Pleurobema_beadleianum **
| | +--+
| | | +-- Pleurobema_perovatum **
+--+ +--+
| +-- Pleurobema_taitianum **
|
| +-- Pleurobema_clava **
| +--------+
| | +-- Pleurobema_oviforme **
+--+
| +-------- Pleurobema_pyriforme **
| |
+--+ +-- Pleurobema_athearni **
| +--+
| | +-- Pleurobema_georgianum **
+--+
| +-- Pleurobema_decisum **
+--+
+-- Pleurobema_hanleyianum **
Zanatta & Murphy, 2006 [Publ 2672], fig. 2
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 55 |
55 |
47 |
4 |
2 |
1 |
1 |
55.0 |
+-------- Pleuronaia_dolabelloides **
|
+-----------------------------------------+-------- Pleurobema_clava **
| |
| | +----- Fusconaia_flava **
| +--+
| | +-- Hemistena_lata **
| +--+
| +-- Eurynia_dilatata **
--+
| +----------------------------------------------- Amblema_plicata **
| |
| +----------------------------------------------- Popenaias_popeii **
| |
| | +----- Paetulunio_fabalis **
| | |
| | +--------------------------------------+ +-- Toxolasma_parvum **
+--+ | | |
| | +--+-- Toxolasma_lividum **
| | |
| | +-- Toxolasma_texasiense **
| |
| | +----- Obliquaria_reflexa **
| | +-----------------------------------+
+--+ | | +-- Glebula_rotundata **
| | +--+
| | +-- Cyrtonaias_tampicoensis **
| |
| | +----- Ptychobranchus_fasciolaris **
| | +--+
| | | | +-- Dromus_dromas **
| | | +--+
+--+ +-----------------------------+ +-- Cyprogenia_stegaria **
| | |
| | | +----- Medionidus_conradicus **
| | +--+
| | | +-- Medionidus_acutissimus **
| | +--+
| | +-- Lemiox_rimosus **
| |
| | +----------------------------------- Sagittunio_nasutus **
| | |
+--+ | +-------------- Potamilus_leptodon **
| | |
| +--------------------+ +----------- Potamilus_fragilis **
| | | |
| | +--+ +-- Potamilus_alatus **
| | | +-----+
| | | | +-- Potamilus_purpuratus **
| | +--+
| | | +----- Potamilus_inflatus **
| | +--+
+--+ | +-- Potamilus_ohiensis **
| +--+
| +-- Potamilus_amphichaenus **
|
| +-- Truncilla_truncata **
| +-----------------------------+
| | +-- Ellipsaria_lineolata **
| |
| | +-- Ortmanniana_pectorosa **
| | +-----+
| | | +-- Lampsilis_fasciola **
| | +--------------------+
+--+ | | +----- Lampsilis_ornata **
| | +--+
| | | +-- Lampsilis_ovata **
| | +--+
| | +-- Lampsilis_cardium **
| |
| | +-- Ligumia_recta **
+--+ +--------+
| | +-- Cambarunio_iris **
| +--------------+
| | | +-------- Lampsilis_teres **
| | | |
| | +--+ +-- Hamiota_australis **
| | | +--+
| | | | +-- Hamiota_subangulata **
+--+ +--+
| | +-- Hamiota_altilis **
| +--+
| +-- Hamiota_perovalis **
|
| +----------------------- Leaunio_vanuxemensis **
| |
+--+ +-------------------- Ortmanniana_ligamentina **
| |
+--+ +----------------- Lampsilis_siliquoidea **
| |
| | +-------------- Obovaria_olivaria **
+--+ |
| +-------------- Venustaconcha_ellipsiformis **
| |
| | +----- Obovaria_unicolor **
+--+--------+
| | +-- Obovaria_subrotunda **
| +--+
| +-- Obovaria_arkansasensis **
|
| +----------- Epioblasma_brevidens **
+--+
| +-------- Epioblasma_triquetra **
+--+
| +----- Epioblasma_rangiana **
+--+
| +-- Epioblasma_walkeri **
+--+
+-- Epioblasma_capsaeformis **
Zanatta & Murphy, 2006 [Publ 2672], fig. 1
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 54 |
54 |
49 |
4 |
2 |
1 |
1 |
55.0 |
+-------------------------------------------------- Pyganodon_grandis
|
| +-- Amblema_plicata **
| +--+
| | +-- Popenaias_popeii **
| |
--+ +-----------------------------------------+ +-- Pleurobema_clava **
| | | |
| | | +-- Eurynia_dilatata **
| | +--+
| | +-- Pleuronaia_dolabelloides **
| | |
+--+ +-- Fusconaia_flava **
|
| +-- Cyrtonaias_tampicoensis **
| +-----------------------------------------+
| | +-- Glebula_rotundata **
| |
| | +----------------------------------------- Paetulunio_fabalis **
+--+ |
| | +-------- Obliquaria_reflexa **
| | +-----------------------------+
| | | | +----- Toxolasma_texasiense **
| | | +--+
+--+ | | +-- Toxolasma_parvum **
| | +--+
| | +-- Toxolasma_lividum **
| |
| | +----------------------------------- Sagittunio_nasutus **
| | |
+--+ | +-- Medionidus_conradicus **
| | +-----+
| | | +-- Ptychobranchus_fasciolaris **
| | +--------+
| | | | +----- Medionidus_acutissimus **
| | | | |
| | | +--+----- Lemiox_rimosus **
| | | |
| | +--------------+ | +-- Dromus_dromas **
+--+ | | +--+
| | | +-- Cyprogenia_stegaria **
| | |
| | | +-------------- Potamilus_leptodon **
| | | |
| | +--+ +----------- Potamilus_fragilis **
| | | |
| | +--+ +-- Potamilus_alatus **
| | | +-----+
| | | | +-- Potamilus_purpuratus **
| | +--+
+--+ | +----- Potamilus_inflatus **
| +--+
| | +-- Potamilus_ohiensis **
| +--+
| +-- Potamilus_amphichaenus **
|
| +----- Lampsilis_teres **
| +-----------------------+
| | | +-- Truncilla_truncata **
| | +--+
| | +-- Ellipsaria_lineolata **
| |
| | +-------------------------- Leaunio_vanuxemensis **
+--+ |
| | +-- Hamiota_australis **
| | +--+
| | | +-- Hamiota_subangulata **
| | +-----+
| | | | +-- Hamiota_altilis **
| | | +--+
+--+ | +-- Hamiota_perovalis **
| +-----------+
| | | +-- Lampsilis_fasciola **
| | | +-----+
| | | | +-- Ortmanniana_pectorosa **
| | +--+
| | | +----- Lampsilis_ornata **
| | +--+
| | | +-- Lampsilis_ovata **
+--+ +--+
| +-- Lampsilis_cardium **
|
| +-------- Lampsilis_siliquoidea **
| +-----------+
| | | +----- Ortmanniana_ligamentina **
| | +--+
| | | +-- Cambarunio_iris **
| | +--+
+--+ +-- Ligumia_recta **
|
| +----------------- Venustaconcha_ellipsiformis **
| |
| | +-------- Obovaria_olivaria **
+--+ +-----+
| | | +----- Obovaria_unicolor **
| | +--+
| | | +-- Obovaria_subrotunda **
+--+ +--+
| +-- Obovaria_arkansasensis **
|
| +----------- Epioblasma_brevidens **
+--+
| +-------- Epioblasma_triquetra **
+--+
| +----- Epioblasma_rangiana **
+--+
| +-- Epioblasma_walkeri **
+--+
+-- Epioblasma_capsaeformis **
Campbell & Lydeard, 2012 [Publ 3308], fig. 3
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 51 |
51 |
42 |
5 |
0 |
0 |
0 |
59.0 |
+-------------- Uniomerus_declivis **
|
+--------------------------------+ +-- Pustulosa_infucata **
| | +--------+
| | | +-- Pustulosa_kleiniana **
| +--+
| | +-------- Quadrula_quadrula **
| +--+
| | +----- Cyclonaias_tuberculata **
| +--+
| | +-- Pustulosa_succissa **
| +--+
| +-- Pustulosa_pustulosa **
--+
| +-------------- Elliptoideus_sloatianus **
| +-----------------------------+
| | | +----------- Plectomerus_dombeyanus **
| | +--+
| | | +-------- Obovaria_unicolor **
| | +--+
| | | +----- Obovaria_olivaria **
| | +--+
| | | +-- Lampsilis_ovata **
| | +--+
| | +-- Lemiox_rimosus **
+--+
| +----------- Cyrtonaias_tampicoensis **
| |
| | +----- Popenaias_popeii **
| +-----------------------------+ +--+
| | | | | +-- Amblema_elliottii **
| | | | +--+
| | +--+ +-- Amblema_plicata **
| | |
| | | +-- Reginaia_ebenus[1] **
| | | |
+--+ +-----+-- Reginaia_ebenus[2] **
| |
| +-- Reginaia_rotulata **
|
| +-- Elliptio_arca **
| +-----------------------------------+
| | +-- Elliptio_crassidens **
| |
+--+ +----------------------------------- Eurynia_dilatata **
| |
| | +-------------------------------- Parvaspina_collina **
| | |
+--+ | +-- Hemistena_lata **
| +-----------------------------+
| | +-- Plethobasus_cyphyus **
| |
+--+ +----- Pleuronaia_barnesiana **
| +-----------------------+
| | | +-- Pleuronaia_dolabelloides **
| | +--+
| | +-- Pleuronaia_gibber **
| |
| | +-- Pleurobema_stabile **
+--+ +-----+
| | +-- Pleurobema_sintoxia **
| +-----------------+
| | | +----- Pleurobema_decisum **
| | +--+
| | | +-- Pleurobema_clava **
+--+ +--+
| +-- Pleurobema_pyriforme **
|
| +----------------------- Pleurobema_cordatum **
| |
+--+ +-------------------- Fusconaia_subrotunda **
| |
| | +-- Fusconaia_cor **
+--+ +--------------+
| | +-- Fusconaia_cuneolus **
| |
+--+ +-------------- Fusconaia_ozarkensis **
| |
| | +-- Fusconaia_flava[6] **
| | +--------+
+--+ | +-- Fusconaia_masoni **
| |
| | +-- Fusconaia_escambia **
| | |
+--+ +-- Fusconaia_burkei[1] **
| +-----+
| | +-- Fusconaia_burkei[2] **
| | |
| | +-- Fusconaia_burkei[3] **
+--+
| +----- Fusconaia_flava[5] **
| |
| | +-- Fusconaia_cerina **
+--+ |
| +-- Fusconaia_flava[1] **
| |
+--+-- Fusconaia_flava[2] **
|
+-- Fusconaia_flava[3] **
|
+-- Fusconaia_flava[4] **
Burlakova et al., 2012 [Publ 3486], fig. 1
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 44 |
44 |
42 |
4 |
0 |
0 |
0 |
57.0 |
+-- Amblema_elliottii **
+--------------------------------------------+
| +-- Amblema_plicata **
|
--+ +-- Lampsilis_ovata **
| +-----------------------------------------+
| | +-- Plectomerus_dombeyanus **
| |
+--+ +----------------------------------------- Popenaias_popeii **
| |
| | +-------------------------------------- Cyrtonaias_tampicoensis **
+--+ |
| | +----------------------------------- Parvaspina_collina **
| | |
+--+ | +----- Plethobasus_cyphyus **
| | +--------------------------+
| | | | +-- Elliptio_arca **
| | | +--+
| | | +-- Elliptio_crassidens **
+--+ |
| | +-------------------- Fusconaia_subrotunda **
| | |
| | | +-- Fusconaia_cor **
| | +--------+ +--------------+
| | | | | +-- Fusconaia_cuneolus **
| | | | |
+--+ | +--+ +-------------- Fusconaia_ozarkensis **
| | | |
| | | | +-------- Fusconaia_askewi[3] **
| | | | +--+
| | +--+ | | +----- Fusconaia_askewi[2] **
| | | | +--+
| | | | | +-- Fusconaia_lananensis **
| | | | +--+
| | +--+ +-- Fusconaia_askewi[1] **
| | |
| | | +-- Fusconaia_burkei **
| | | +-----+
+--+ | | +-- Fusconaia_escambia **
| +--+
| | +----- Fusconaia_masoni **
| +--+
| | +-- Fusconaia_flava **
| +--+
| +-- Fusconaia_cerina **
|
| +-- Eurynia_dilatata **
| +-----------------------+
| | +-- Hemistena_lata **
| |
| | +----- Pleuronaia_gibber **
| | +-----+
| | | | +-- Pleuronaia_dolabelloides **
+--+ | +--+
| +-----------+ +-- Pleuronaia_barnesiana **
| | |
| | | +-------- Pleurobema_riddellii **
| | +--+
| | | +----- Pleurobema_sintoxia **
| | +--+
+--+ | +-- Pleurobema_rubrum **
| +--+
| +-- Pleurobema_cordatum **
|
| +-------------------- Pleurobema_stabile **
| |
+--+ +----------------- Pleurobema_pyriforme **
| |
| | +----- Pleurobema_georgianum **
+--+ +--------+
| | | +-- Pleurobema_hanleyianum **
| | +--+
| | +-- Pleurobema_decisum **
+--+
| +-- Pleurobema_clava **
| +--------+
| | +-- Pleurobema_oviforme **
+--+
| +-------- Pleurobema_beadleianum **
| |
+--+ +-- Pleurobema_rubellum **
| +--+
| | +-- Pleurobema_strodeanum **
+--+
| +-- Pleurobema_perovatum **
+--+
+-- Pleurobema_taitianum **
Campbell & Lydeard, 2012 [Publ 3308], fig. 1
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 43 |
43 |
41 |
5 |
0 |
0 |
0 |
52.0 |
+----------- Quadrula_quadrula **
|
+-----------------------------------+ +-- Pustulosa_infucata **
| | +-----+
| | | +-- Pustulosa_kleiniana **
| +--+
| | +----- Cyclonaias_tuberculata **
| +--+
| | +-- Pustulosa_succissa **
| +--+
| +-- Pustulosa_pustulosa **
|
| +----- Popenaias_popeii **
| +-----------+
--+ | | +-- Amblema_elliottii **
| | +--+
| | +-- Amblema_plicata **
| +--------------------------+
| | | +----- Cyrtonaias_tampicoensis **
| | | +--------+
| | | | | +-- Reginaia_ebenus **
| | | | +--+
| | +--+ +-- Reginaia_rotulata **
| | |
| | | +----------- Plectomerus_dombeyanus **
| | +--+
| | | +-------- Lemiox_rimosus **
+--+ +--+
| | +----- Lampsilis_ovata **
| +--+
| | +-- Obovaria_unicolor **
| +--+
| +-- Obovaria_olivaria **
|
| +----------------------------------------- Elliptoideus_sloatianus **
| |
| | +-------------------------------------- Pleurobema_stabile **
+--+ |
| | +----- Pleurobema_pyriforme **
| | +-----------------------------+
+--+ | | +-- Pleurobema_clava **
| | +--+
| | +-- Pleurobema_decisum **
| |
+--+ +-- Pleurobema_cordatum **
| +-----------------------------+
| | +-- Pleurobema_sintoxia **
| |
| | +-- Eurynia_dilatata **
| | +-----+
+--+ | +-- Hemistena_lata **
| +--------------------+
| | | +----- Plethobasus_cyphyus **
| | +--+
| | | +-- Elliptio_arca **
| | +--+
+--+ +-- Elliptio_crassidens **
|
| +----- Pleuronaia_gibber **
| +--------------------+
| | | +-- Pleuronaia_barnesiana **
| | +--+
+--+ +-- Pleuronaia_dolabelloides **
|
| +----------------------- Parvaspina_collina **
| |
+--+ +-------------------- Fusconaia_subrotunda **
| |
+--+ +-- Fusconaia_cor **
| +--------------+
| | +-- Fusconaia_cuneolus **
+--+
| +-------------- Fusconaia_ozarkensis **
| |
+--+ +-- Fusconaia_burkei **
| +--------+
| | +-- Fusconaia_escambia **
+--+
| +-------- Fusconaia_masoni **
+--+
| +----- Fusconaia_cerina[2] **
+--+
| +-- Fusconaia_flava **
+--+
+-- Fusconaia_cerina[1] **
Campbell & Lydeard, 2012 [Publ 3308], fig. 2
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 43 |
43 |
36 |
5 |
0 |
0 |
0 |
50.0 |
+----------- Quadrula_quadrula **
|
+--------------------+ +-- Pustulosa_infucata **
| | +-----+
| | | +-- Pustulosa_kleiniana **
| +--+
| | +----- Cyclonaias_tuberculata **
| +--+
| | +-- Pustulosa_succissa **
| +--+
| +-- Pustulosa_pustulosa **
|
| +-- Reginaia_ebenus **
| +--+
| | +-- Reginaia_rotulata **
--+ +-----------+
| | | +-- Amblema_elliottii **
| | +--+
| +-----------+ +-- Amblema_plicata **
| | |
| | | +-------------- Popenaias_popeii **
| | | |
| | +--+ +----------- Cyrtonaias_tampicoensis **
| | | |
| | +--+----------- Plectomerus_dombeyanus **
| | |
| | | +-------- Lemiox_rimosus **
| | +--+
| | | +----- Lampsilis_ovata **
+--+ +--+
| | +-- Obovaria_unicolor **
| +--+
| +-- Obovaria_olivaria **
|
| +-------------------------- Elliptoideus_sloatianus **
| |
| | +----------------------- Hemistena_lata **
| | |
| | +----------------------- Plethobasus_cyphyus **
| | |
+--+ | +-- Elliptio_arca **
| +--------------------+
| | +-- Elliptio_crassidens **
| |
| | +-- Eurynia_dilatata **
| +--------------------+
| | +-- Parvaspina_collina **
+--+
| +----- Pleuronaia_gibber **
| +--------------+
| | | +-- Pleuronaia_barnesiana **
| | +--+
| | +-- Pleuronaia_dolabelloides **
| |
| | +-- Pleurobema_cordatum **
| | +--------+
| | | +-- Pleurobema_sintoxia **
| +--------+
| | | +-------- Pleurobema_stabile **
+--+ +--+
| | +----- Pleurobema_pyriforme **
| +--+
| | +-- Pleurobema_clava **
| +--+
| +-- Pleurobema_decisum **
|
| +----- Fusconaia_subrotunda **
| +-----------+
| | | +-- Fusconaia_cor **
| | +--+
+--+ +-- Fusconaia_cuneolus **
|
| +-------------- Fusconaia_ozarkensis **
| |
+--+ +-- Fusconaia_burkei **
| +--------+
| | +-- Fusconaia_escambia **
+--+
| +-------- Fusconaia_masoni **
+--+
| +----- Fusconaia_cerina[2] **
+--+
| +-- Fusconaia_flava **
+--+
+-- Fusconaia_cerina[1] **
Campbell & Lydeard, 2012 [Publ 3308], fig. 4
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 43 |
43 |
40 |
5 |
0 |
0 |
0 |
52.0 |
+----------- Quadrula_quadrula **
|
+--------------------------------------+ +-- Pustulosa_infucata **
| | +-----+
| | | +-- Pustulosa_kleiniana **
| +--+
| | +----- Cyclonaias_tuberculata **
| +--+
| | +-- Pustulosa_succissa **
| +--+
| +-- Pustulosa_pustulosa **
|
| +----- Popenaias_popeii **
| +--------+
--+ | | +-- Amblema_elliottii **
| | +--+
| | +-- Amblema_plicata **
| +--------------------------------+
| | | +----------- Plectomerus_dombeyanus **
| | | |
| | | | +----- Cyrtonaias_tampicoensis **
| | | +-----+
| | +--+ | +-- Reginaia_ebenus **
| | | +--+
| | | +-- Reginaia_rotulata **
| | |
+--+ | +-------- Lemiox_rimosus **
| +--+
| | +----- Lampsilis_ovata **
| +--+
| | +-- Obovaria_unicolor **
| +--+
| +-- Obovaria_olivaria **
|
| +-------------------------------------------- Elliptoideus_sloatianus **
| |
+--+ +----------------------------------------- Pleurobema_stabile **
| |
+--+ +-------------------------------------- Hemistena_lata **
| |
+--+ +----------------------------------- Eurynia_dilatata **
| |
| | +-- Pleurobema_cordatum **
+--+ +-----------------------------+
| | +-- Pleurobema_sintoxia **
| |
| | +----- Pleurobema_clava **
+--+ +-----------------------+
| | | +-- Pleurobema_decisum **
| | +--+
| | +-- Pleurobema_pyriforme **
| |
+--+ +-------- Parvaspina_collina **
| +-----------------+
| | | +----- Pleuronaia_gibber **
| | +--+
| | | +-- Pleuronaia_barnesiana **
| | +--+
+--+ +-- Pleuronaia_dolabelloides **
|
| +----- Plethobasus_cyphyus **
| +-----------------+
| | | +-- Elliptio_arca **
| | +--+
+--+ +-- Elliptio_crassidens **
|
| +-------------------- Fusconaia_subrotunda **
| |
+--+ +-- Fusconaia_cor **
| +--------------+
| | +-- Fusconaia_cuneolus **
+--+
| +-------------- Fusconaia_ozarkensis **
| |
+--+ +-- Fusconaia_burkei **
| +--------+
| | +-- Fusconaia_escambia **
+--+
| +-------- Fusconaia_masoni **
+--+
| +----- Fusconaia_cerina[2] **
+--+
| +-- Fusconaia_flava **
+--+
+-- Fusconaia_cerina[1] **
Burlakova et al., 2012 [Publ 3486], fig. 3
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 43 |
43 |
26 |
2 |
0 |
0 |
0 |
51.0 |
+----------------------------------- Plectomerus_dombeyanus **
|
| +-------------------------------- Hemistena_lata **
| |
--+ +-------------------------------- Pleurobema_stabile **
| |
| | +----------------------------- Eurynia_dilatata **
| | |
+--+ | +-------- Pleurobema_riddellii **
| | |
| | +-----------------+ +----- Pleurobema_sintoxia **
| | | | |
| | | +--+----- Pleurobema_rubrum[2] **
| | | |
+--+ | | +-- Pleurobema_cordatum **
| | +--+
| | +-- Pleurobema_rubrum[1] **
| |
| | +----------------------- Parvaspina_collina **
| | |
| | | +----- Pleuronaia_gibber **
| | +-----------------+
| | | | +-- Pleuronaia_dolabelloides **
+--+ | +--+
| | +-- Pleuronaia_barnesiana **
| |
| | +----- Plethobasus_cyphyus **
| +-----------------+
| | | +-- Elliptio_arca **
| | +--+
| | +-- Elliptio_crassidens **
| |
| | +-- Pleurobema_clava **
| | +--------+
| | | +-- Pleurobema_oviforme **
| | |
| | | +-------- Pleurobema_decisum **
+--+-----------+ |
| | +-------- Pleurobema_pyriforme **
| | |
| | +-------- Pleurobema_hanleyianum **
| +--+
| +-------- Pleurobema_georgianum **
| |
| | +----- Pleurobema_rubellum **
| | |
| | +----- Pleurobema_beadleianum **
| +--+
| +----- Pleurobema_strodeanum **
| |
| | +-- Pleurobema_perovatum **
| +--+
| +-- Pleurobema_taitianum **
|
| +-------------------- Fusconaia_subrotunda **
| |
| | +-- Fusconaia_cor **
+--+ +--------------+
| | +-- Fusconaia_cuneolus **
| |
| | +-------------- Fusconaia_ozarkensis **
+--+ |
| | +-------- Fusconaia_masoni **
| | |
| | | +-- Fusconaia_burkei **
| | +--+-----+
+--+ | | +-- Fusconaia_escambia **
| | |
| | | +----- Fusconaia_cerina **
| | +--+
| | | +-- Fusconaia_askewi[1] **
| | +--+
+--+ +-- Fusconaia_flava **
|
| +-- Fusconaia_askewi[2] **
| |
| +-- Fusconaia_askewi[3] **
| |
| +-- Fusconaia_askewi[4] **
+--------+
+-- Fusconaia_lananensis[1] **
|
+-- Fusconaia_lananensis[2] **
|
+-- Fusconaia_askewi[5] **
Inoue et al., 2020 [Publ 3941], fig. 3
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 43 |
43 |
41 |
4 |
1 |
1 |
1 |
43.0 |
+----------------------------------------------- Arcidens_confragosus
|
| +-- Pustulosa_petrina **
| +-----------------------------------------+
| | +-- Quadrula_quadrula **
--+ |
| | +-------------- Parvaspina_collina **
| | |
| | | +-- Eurynia_dilatata **
| | +--------------------------+ +--------+
| | | | | +-- Hemistena_lata **
+--+ | | |
| | +--+ +-- Pleurobema_clava **
| | | +-----+
| | | | +-- Pleurobema_sintoxia **
| | +--+
| | | +----- Pleuronaia_dolabelloides **
| | +--+
| | | +-- Fusconaia_flava **
+--+ +--+
| +-- Fusconaia_subrotunda **
|
| +-- Amblema_plicata **
| +--+
| | +-- Amblema_elliottii **
| +--------------------------------+
| | | +-- Reginaia_ebenus **
| | +--+
| | +-- Reginaia_rotulata **
| |
+--+ +----------------------------------- Toxolasma_lividum **
| |
| | +-------- Potamilus_fragilis **
| | |
| | +-----------------------+ +-- Potamilus_ohiensis **
+--+ | | +--+
| | | | +-- Potamilus_amphichaenus **
| | +--+
| | | +-- Potamilus_metnecktayi **
| | +--+
| | +-- Potamilus_purpuratus **
+--+
| +-------- Ellipsaria_lineolata **
| +--------------------+
| | | +----- Truncilla_macrodon **
| | +--+
| | | +-- Truncilla_donaciformis **
| | +--+
| | +-- Truncilla_cognata **
+--+
| +-------------------------- Leaunio_lienosus **
| |
| | +-------------------- Lampsilis_bracteata **
| | |
| | +--+ +----------------- Lampsilis_fasciola **
| | | | |
+--+ | +--+ +-------------- Lampsilis_ornata **
| | | |
| | +--+ +-- Lampsilis_ovata **
| | | +--------+
| | | | +-- Lampsilis_cariosa **
| | +--+
| | | +-------- Lampsilis_cardium[3] **
+--+ +--+
| | +----- Lampsilis_cardium[2] **
| +--+
| | +-- Lampsilis_satura **
| +--+
| +-- Lampsilis_cardium[1] **
|
| +-- Lampsilis_higginsii **
| +--------------+
| | +-- Ortmanniana_ligamentina **
+-----+
| +-------------- Lampsilis_teres **
| |
+--+ +-- Lampsilis_radiata **
| +--------+
| | +-- Lampsilis_siliquoidea **
+--+
| +-------- Lampsilis_bergmanni **
+--+
| +----- Lampsilis_hydiana **
+--+
| +-- Lampsilis_virescens **
+--+
+-- Lampsilis_straminea **
Inoue et al., 2020 [Publ 3941], fig. 4
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 42 |
42 |
40 |
4 |
1 |
1 |
1 |
44.0 |
+-------------------------------------------------- Arcidens_confragosus
|
| +-- Pustulosa_petrina **
| +--------------------------------------------+
--+ | +-- Quadrula_quadrula **
| |
| | +-- Reginaia_ebenus **
| | +-----------------------------------------+
+--+ | +-- Reginaia_rotulata **
| |
| | +-------------- Parvaspina_collina **
| | |
| | | +-- Eurynia_dilatata **
| | +--------------------------+ +--------+
+--+ | | | +-- Hemistena_lata **
| | | |
| | +--+ +-- Pleurobema_clava **
| | | +-----+
| | | | +-- Pleurobema_sintoxia **
| | +--+
| | | +----- Pleuronaia_dolabelloides **
+--+ +--+
| | +-- Fusconaia_flava **
| +--+
| +-- Fusconaia_subrotunda **
|
| +-- Amblema_plicata **
| +-----------------------------------+
| | +-- Amblema_elliottii **
| |
+--+ +----------------------------------- Toxolasma_lividum **
| |
| | +-------------------------------- Ellipsaria_lineolata **
+--+ |
| | +----- Truncilla_cognata **
| | +-----------------------+
| | | | +-- Truncilla_macrodon **
+--+ | +--+
| | +-- Truncilla_donaciformis **
| |
| | +-------- Potamilus_fragilis **
| | |
+--+ +-----------------+ +-- Potamilus_ohiensis **
| | | +--+
| | | | +-- Potamilus_amphichaenus **
| | +--+
| | | +-- Potamilus_metnecktayi **
| | +--+
| | +-- Potamilus_purpuratus **
+--+
| +----------------------- Leaunio_lienosus **
| |
| | +----------------- Lampsilis_bracteata **
| | |
| | +--+ +-- Lampsilis_ornata **
| | | | +-----------+
+--+ | | | +-- Lampsilis_fasciola **
| | +--+
| | | +----------- Lampsilis_ovata **
| | +--+
| | | +-------- Lampsilis_cardium[3] **
| | +--+
+--+ | +----- Lampsilis_cardium[2] **
| +--+
| | +-- Lampsilis_satura **
| +--+
| +-- Lampsilis_cardium[1] **
|
| +-------------- Lampsilis_teres **
| |
| | +-- Lampsilis_higginsii **
+-----+ +--------+
| | +-- Ortmanniana_ligamentina **
| |
+--+ +-- Lampsilis_radiata **
| +-----+
| | +-- Lampsilis_siliquoidea **
| |
+--+ +-- Lampsilis_hydiana **
| +--+
| | +-- Lampsilis_bergmanni **
+--+
| +-- Lampsilis_virescens **
+--+
+-- Lampsilis_straminea **
Smith et al., 2019 [Publ 3951], fig. 2
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 42 |
42 |
40 |
4 |
0 |
0 |
0 |
43.0 |
+----------------------------------------------- Quadrula_quadrula **
|
| +-- Amblema_plicata **
--+ +-----------------------------------------+
| | +-- Reginaia_ebenus **
| |
+--+ +----------------------------------------- Plectomerus_dombeyanus **
| |
| | +-------------------------------------- Toxolasma_lividum **
+--+ |
| | +----- Popenaias_popeii **
| | +-----------------------------+
+--+ | | +-- Glebula_rotundata **
| | +--+
| | +-- Cyrtonaias_tampicoensis **
| |
| | +-------------------------------- Obliquaria_reflexa **
+--+ |
| | +-- Medionidus_conradicus **
| | +--+
| | | +-- Lemiox_rimosus **
| | +-----------------------+
| | | | +-- Cyprogenia_stegaria **
| | | +--+
+--+ | +-- Ptychobranchus_fasciolaris **
| |
| | +-- Hamiota_subangulata **
| | +--------------------+
| | | +-- Villosa_villosa **
| | |
| | +--+ +-------------------- Ligumia_recta **
| | | | |
+--+ | | | +----- Venustaconcha_ellipsiformis **
| | +--+ +-----------+
| | | | | +-- Obovaria_subrotunda **
| | | | +--+
| | +--+ +-- Epioblasma_rangiana **
| | |
| | | +-------------- Ortmanniana_pectorosa **
| | +--+
| | | +----------- Ortmanniana_ligamentina **
| | +--+
| | | +-------- Lampsilis_ovata **
| | +--+
+--+ | +----- Lampsilis_dolabraeformis **
| +--+
| | +-- Lampsilis_satura **
| +--+
| +-- Lampsilis_cardium **
|
| +-------------- Atlanticoncha_ochracea **
| +-----+
| | | +----------- Ellipsaria_lineolata **
| | +--+
| | | +-------- Truncilla_truncata **
| | +--+
| | | +----- Truncilla_cognata **
| | +--+
+-----+ | +-- Truncilla_macrodon **
| +--+
| +-- Truncilla_donaciformis **
|
| +----------------- Potamilus_capax **
| |
| | +----- Potamilus_inflatus **
+--+ +--------+
| | | +-- Potamilus_fragilis **
| | +--+
| | +-- Potamilus_leptodon **
+--+
| +----- Potamilus_ohiensis **
| +-----+
| | | +-- Potamilus_amphichaenus **
| | +--+
+--+ +-- Potamilus_streckersoni **
|
| +-------- Potamilus_metnecktayi **
+--+
| +----- Potamilus_purpuratus[1] **
+--+
| +-- Potamilus_purpuratus[2] **
+--+
+-- Potamilus_alatus **
Burlakova et al., 2012 [Publ 3486], fig. 2
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 40 |
40 |
30 |
1 |
0 |
0 |
0 |
53.0 |
+----------------------------------- Parvaspina_collina **
|
| +-------------------------------- Pleurobema_plenum **
| |
--+ | +-- Plethobasus_cyphyus **
| | +-----+
| | | +-- Hemistena_lata **
| | +--------------------+
+--+ | | +----- Eurynia_dilatata **
| | +--+
| | | +-- Elliptio_arca **
| | +--+
| | +-- Elliptio_crassidens **
| |
| | +-------------------------- Pleurobema_cordatum[2] **
+--+ |
| | +-------------- Fusconaia_subrotunda **
| | |
| | | +-- Fusconaia_cor **
| | | +--------+
| | +--------+ | +-- Fusconaia_cuneolus **
| | | | |
| | | | | +-------- Fusconaia_flava[2] **
| | | | | |
+--+ | +--+ +-------- Fusconaia_ozarkensis **
| | | |
| | | +-------- Fusconaia_masoni **
| | | |
| | | | +----- Fusconaia_askewi **
| | +--+--+
| | | | +-- Fusconaia_burkei **
| | | +--+
| | | +-- Fusconaia_escambia **
| | |
+--+ | +-- Fusconaia_flava[1] **
| | |
| +-----+-- Fusconaia_cerina[1] **
| |
| +-- Fusconaia_cerina[2] **
|
| +-------- Pleurobema_stabile **
| |
| +-----------+ +----- Pleurobema_riddellii **
| | | |
| | +--+ +-- Pleurobema_cordatum[1] **
| | | |
| | | +-- Pleurobema_rubrum **
| | +--+
| | +-- Pleurobema_sintoxia[1] **
+--+ |
| +-- Pleurobema_sintoxia[2] **
|
| +----- Pleuronaia_barnesiana **
| +-----------+
| | | +-- Pleuronaia_dolabelloides **
| | +--+
| | +-- Pleuronaia_gibber **
+--+
| +-- Pleurobema_georgianum **
| +-----------+
| | +-- Pleurobema_pyriforme **
| |
+--+ +-- Pleurobema_decisum **
| +--------+
| | +-- Pleurobema_hanleyianum **
| |
+--+ +-- Pleurobema_clava **
| +-----+
| | +-- Pleurobema_oviforme **
| |
+--+ +----- Pleurobema_rubellum **
| |
| +----- Pleurobema_beadleianum **
+--+
+----- Pleurobema_strodeanum **
|
| +-- Pleurobema_perovatum **
+--+
+-- Pleurobema_taitianum **
Burdick & White, 2007 [Publ 2679], fig. 3
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 38 |
38 |
31 |
3 |
0 |
0 |
0 |
46.0 |
+----------------------------------------- Lampsilis_ornata **
|
--+ +-------------------------------------- Parvaspina_collina **
| |
| | +----------------------------------- Reginaia_ebenus **
+--+ |
| | +----- Eurynia_dilatata **
| | +--------------------------+
| | | | +-- Plethobasus_cyphyus **
+--+ | +--+
| | +-- Elliptio_crassidens **
| |
| | +----------------------------- Pleurobema_plenum **
| | |
| | | +-- Pleuronaia_barnesiana **
+--+ +--------------------------+
| | +-- Pleuronaia_gibber **
| |
| | +-------------- Pleurobema_stabile **
| | |
| | | +-- Pleurobema_rubellum **
| | | +-----+
| | | | +-- Pleurobema_strodeanum **
+--+ +-----------+ +--+
| | | | | +----- Pleurobema_beadleianum **
| | | | +--+
| | | | | +-- Pleurobema_perovatum **
| | | | +--+
| | +--+ +-- Pleurobema_taitianum **
| | |
| | | +-- Pleurobema_clava **
| | | +-----+
| | | | +-- Pleurobema_oviforme **
| | | |
| | +--+ +-- Pleurobema_decisum **
+--+ | +--+
| | | +-- Pleurobema_hanleyianum **
| +--+
| | +-- Pleurobema_pyriforme **
| +--+
| +-- Pleurobema_georgianum **
|
| +----- Pleurobema_sintoxia[1] **
| |
| +-----------------+ +-- Pleurobema_cordatum[2] **
| | | |
| | +--+-- Pleurobema_sintoxia[2] **
| | |
+--+ +-- Pleurobema_rubrum **
|
| +-------------------- Pleurobema_cordatum[1] **
| |
+--+ +----------------- Fusconaia_subrotunda **
| |
| | +-- Fusconaia_cor **
+--+ +-----------+
| | +-- Fusconaia_cuneolus **
| |
+--+ +----------- Fusconaia_ozarkensis **
| |
| | +-- Fusconaia_masoni **
+--+ +-----+
| | +-- Fusconaia_flava[3] **
| |
+--+ +----- Fusconaia_flava[1] **
| |
| +----- Fusconaia_cerina[4] **
+--+
| +-- Fusconaia_cerina[1] **
| |
| +-- Fusconaia_cerina[2] **
+--+
+-- Fusconaia_cerina[3] **
|
+-- Fusconaia_flava[2] **
Pfeiffer et al., 2016 [Publ 3660], fig. 2
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 38 |
38 |
33 |
4 |
1 |
1 |
1 |
47.0 |
+-------------------------------------- Utterbackiana_suborbiculata
|
| +-- Amblema_plicata **
| +--+
| | +-- Amblema_neislerii **
| +-----------------------------+
--+ | | +-- Reginaia_rotulata **
| | +--+
| | +-- Reginaia_ebenus **
| |
| | +-- Cyrtonaias_tampicoensis **
| | +-----+
| | | +-- Glebula_rotundata **
+--+ |
| +-----------------------+ +-- Lampsilis_teres **
| | | +--+
| | | | +-- Leaunio_lienosus **
| | +--+
| | | +-- Truncilla_macrodon **
| | +--+
| | +-- Potamilus_fragilis **
+--+
| +----------------------------- Elliptoideus_sloatianus **
| |
| | +-------------- Uniomerus_declivis **
| | +-----------+
| | | | +----------- Quadrula_quadrula **
| | | +--+
+--+ | | +-------- Quadrula_verrucosa **
| | +--+
| | | +----- Pustulosa_pustulosa **
| | +--+
| | | +-- Pustulosa_petrina **
| | +--+
+--+ +-- Cyclonaias_tuberculata **
|
| +-- Elliptio_fumata **
| +--------------------+
| | +-- Elliptio_crassidens **
| |
| | +-------------------- Parvaspina_collina **
| | |
| | +-------------------- Plethobasus_cyphyus **
+--+ |
| | +-- Hemistena_lata **
| +-----------------+
| | +-- Eurynia_dilatata **
| |
| | +----- Pleurobema_riddellii **
+--+ +--+
| | | +-- Pleurobema_sintoxia **
| | +--+
| +--------+ +-- Pleurobema_cordatum **
| | |
| | | +----- Pleuronaia_dolabelloides **
| | | |
| | +--+ +-- Pleurobema_pyriforme **
| | | |
+--+ +--+-- Pleurobema_strodeanum **
| |
| +-- Pleurobema_clava **
|
| +-------------- Fusconaia_askewi **
| |
| | +-- Fusconaia_mitchelli **
+--+ +--------+
| | +-- Fusconaia_iheringi **
| |
+--+ +-- Fusconaia_escambia **
| +-----+
| | +-- Fusconaia_burkei **
+--+
| +----- Fusconaia_flava[2] **
+--+
| +-- Fusconaia_cerina **
+--+
+-- Fusconaia_flava[1] **
Pfeiffer et al., 2019 [Publ 3866], fig. 1
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 38 |
38 |
33 |
5 |
1 |
1 |
1 |
42.0 |
+-------------------------------------- Gonidea_angulata
|
| +----------- Megalonaias_nervosa **
| +-----------------------+
| | | +-------- Uniomerus_declivis **
| | +--+
--+ | | +----- Quadrula_verrucosa **
| | +--+
| | | +-- Cyclonaias_tuberculata **
| | +--+
| | +-- Quadrula_quadrula **
| |
| | +-------------- Elliptoideus_sloatianus **
+--+ +-----------------+
| | | +----------- Elliptio_fumata **
| | +--+
| | | +-------- Pleuronaia_barnesiana **
| | +--+
| | | +----- Pleurobema_riddellii **
| | +--+
| | | +-- Fusconaia_flava **
| | +--+
| | +-- Pleurobema_strodeanum **
| |
+--+ +-- Amblema_plicata **
| +--+
| | +-- Amblema_neislerii **
| +-----------------------+
| | | +-- Reginaia_ebenus **
| | +--+
| | +-- Reginaia_rotulata **
| |
| | +----------------- Popenaias_popeii **
| | |
| | +--------+ +-------------- Psoronaias_guatemalensis[2] **
| | | | |
+--+ | +--+ +----------- Psoronaias_guatemalensis[1] **
| | | |
| | +--+ +-------- Psoronaias_semigranosa[3] **
| | | |
| | | | +----- Psoronaias_semigranosa[2] **
| | +--+ |
| | | +----- Psoronaias_usumasintae[2] **
| | | |
| | +--+----- Psoronaias_percompressa **
+--+ |
| | +-- Psoronaias_semigranosa[1] **
| | |
| +--+-- Psoronaias_profunda **
| |
| +-- Psoronaias_usumasintae[1] **
|
| +-- Cyrtonaias_tampicoensis **
| +--------------------+
| | +-- Glebula_rotundata **
| |
+--+ +-------------------- Potamilus_alatus **
| |
| | +-- Truncilla_truncata **
+--+ +--------------+
| | +-- Ptychobranchus_fasciolaris **
| |
+--+ +-------------- Pachynaias_spheniopsis **
| |
| | +-- Cyrtonaias_explicata **
+--+ +--------+
| | +-- Sphenonaias_microdon **
| |
+--+ +-- Lampsilis_cardium **
| +-----+
| | +-- Ortmanniana_ligamentina **
+--+
| +----- Cambarunio_iris **
+--+
| +-- Epioblasma_triquetra **
+--+
+-- Ligumia_recta **
Perkins et al., 2017 [Publ 3777], fig. 2
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 37 |
37 |
34 |
2 |
0 |
0 |
0 |
48.0 |
+----------------------------------------- Amblema_plicata **
|
+----------------------------------------- Plectomerus_dombeyanus **
|
| +-- Pleurobema_clava **
| +-----------+
| | +-- Pleurobema_oviforme **
| |
| | +-------- Pleurobema_pyriforme **
| +-----------------------+ +--+
--+ | | | | +----- Pleurobema_georgianum **
| | | | +--+
| | | | | +-- Pleurobema_hanleyianum **
| | +--+ +--+
| | | +-- Pleurobema_decisum **
| | |
| | | +-------- Pleurobema_rubellum **
| | +--+
| | | +----- Pleurobema_perovatum **
| | +--+
+--+ | +-- Pleurobema_beadleianum **
| +--+
| +-- Pleurobema_strodeanum **
|
| +----- Pleuronaia_gibber **
| +-----------------------------+
| | | +-- Pleuronaia_barnesiana **
| | +--+
| | +-- Pleuronaia_dolabelloides **
| |
| | +-- Fusconaia_masoni **
+--+ +--+
| | +-- Fusconaia_flava **
| +--------------------------+
| | | +-- Fusconaia_subrotunda **
| | +--+
| | +-- Fusconaia_cor **
+--+
| +-- Parvaspina_collina **
| +--------------------------+
| | +-- Parvaspina_steinstansana **
| |
+--+ +-- Eurynia_dilatata **
| +-----------------------+
| | +-- Hemistena_lata **
+--+
| +----------------------- Elliptio_producta[2] **
| |
+--+ +-------------------- Elliptio_producta[1] **
| |
+--+ +----------------- Elliptio_spinosa **
| |
| | +-------------- Elliptio_icterina **
+--+ |
| | +-------- Elliptio_roanokensis **
| | +--+
+--+ | | +----- Elliptio_complanata **
| | +--+
| | | +-- Elliptio_crassidens **
| | +--+
+--+ +-- Elliptio_pullata **
|
| +-- Elliptio_congaraea **
| +-----+
| | +-- Elliptio_hopetonensis **
| |
+--+ +-- Elliptio_folliculata **
| +--+
| | +-- Elliptio_jayensis **
+--+
| +-- Elliptio_arca **
+--+
+-- Elliptio_arctata **
Pieri et al., 2018 [Publ 3867], fig. 3
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 36 |
36 |
22 |
2 |
1 |
1 |
1 |
38.0 |
+----------------------------------------- Arcidens_confragosus
|
| +-- Quadrula_quadrula **
| +-----------------------------------+
--+ | +-- Pustulosa_petrina **
| |
| | +----------------------------------- Eurynia_dilatata **
| | |
+--+ | +----- Pleurobema_clava **
| +-----------------------------+
| | | +-- Pleurobema_riddellii **
| | +--+
| | +-- Pleurobema_sintoxia **
+--+
| +----- Fusconaia_subrotunda **
| +--------------------------+
| | | +-- Fusconaia_cor **
| | +--+
| | +-- Fusconaia_cuneolus **
| |
+--+ +----------------------------- Fusconaia_ozarkensis **
| |
| | +----------- Fusconaia_cerina[3] **
| | |
| | +--------------+ +-------- Fusconaia_flava[1] **
+--+ | | |
| | +--+ +----- Fusconaia_flava[3] **
| | | |
| | +--+ +-- Fusconaia_cerina[1] **
| | | |
| | +--+-- Fusconaia_cerina[2] **
+--+ |
| +-- Fusconaia_flava[2] **
|
| +----------------------- Fusconaia_masoni **
| |
| | +-- Fusconaia_burkei **
| +--------------------+
+--+ +-- Fusconaia_escambia **
|
| +-------------------- Fusconaia_mitchelli **
| |
+--+ +----------------- Fusconaia_chunii **
| |
| | +-------------- Fusconaia_askewi[9] **
+--+ |
| | +----------- Fusconaia_askewi[6] **
| | |
+--+ +----------- Fusconaia_askewi[7] **
| |
| +----------- Fusconaia_askewi[8] **
| |
| +----------- Fusconaia_lananensis[5] **
+--+
+----------- Fusconaia_lananensis[6] **
|
| +-------- Fusconaia_askewi[5] **
| |
| +-------- Fusconaia_lananensis[4] **
| |
+--+ +----- Fusconaia_askewi[2] **
| |
| +----- Fusconaia_askewi[3] **
| |
+--+----- Fusconaia_askewi[4] **
|
| +-- Fusconaia_askewi[1] **
| |
| +-- Fusconaia_lananensis[1] **
+--+
+-- Fusconaia_lananensis[2] **
|
+-- Fusconaia_lananensis[3] **
Burlakova et al., 2019 [Publ 3899], fig. 1
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 33 |
33 |
21 |
5 |
0 |
0 |
0 |
38.0 |
+-- Quadrula_quadrula **
+-----------------------+
| +-- Uniomerus_declivis **
|
| +----------------------- Plectomerus_dombeyanus **
--+ |
| | +-------------------- Popenaias_popeii **
| | |
| | | +-- Elliptio_crassidens **
| | +-----------------+
+--+ | +-- Pleurobema_rubellum **
| |
| | +-- Cyrtonaias_tampicoensis **
| | +--+
| | | +-- Glebula_rotundata **
| +--------------+
| | | +-- Reginaia_ebenus **
+--+ +--+
| +-- Obliquaria_reflexa **
|
| +----------------- Toxolasma_lividum **
| |
| | +-- Potamilus_fragilis **
| | +-----------+
| | | +-- Potamilus_metnecktayi **
| | |
| | | +----- Medionidus_acutissimus **
| | +--------+
+--+ | | +-- Lemiox_rimosus **
| | +--+
| | +-- Dromus_dromas **
| |
| | +----- Ellipsaria_lineolata **
| | |
| +--------+ +-- Truncilla_cognata **
| | | |
+--+ | +-- Truncilla_donaciformis **
| +--+
| +-- Truncilla_macrodon **
| |
| +-- Truncilla_truncata **
|
| +----------- Cambarunio_simus **
| |
| +----------- Leaunio_lienosus **
| |
| +----------- Villosa_villosa **
| |
| | +-- Sagittunio_nasutus **
| +--------+
+--+ +-- Lampsilis_ovata **
|
| +----- Venustaconcha_pleasii **
+-----+
| | +-- Epioblasma_rangiana **
| +--+
| +-- Obovaria_unicolor **
|
| +-------- Lampsilis_teres **
+--+
| +----- Lampsilis_siliquoidea **
+--+
| +-- Lampsilis_hydiana **
+--+
+-- Lampsilis_straminea **
Campbell et al., 2008 [Publ 2675], fig. 5
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 30 |
30 |
7 |
4 |
0 |
0 |
0 |
45.0 |
+-------------- Uniomerus_declivis **
|
+-------------- Plectomerus_dombeyanus **
|
| +-- Elliptio_arctata **
| +--------+
| | +-- Elliptio_crassidens **
| |
| | +-------- Pleurobema_decisum **
--+ | |
| | +-------- Pleurobema_rubellum **
| | |
| | +-------- Pleurobema_georgianum[2] **
| | |
| | +-------- Pleurobema_stabile **
| | |
| | +-------- Pleurobema_pyriforme **
| | |
+--+ +-------- Pleurobema_oviforme[2] **
| |
| +-------- Pleurobema_hanleyianum[2] **
| |
| +-------- Pleuronaia_gibber **
| |
| +-------- Pleurobema_georgianum[3] **
| |
| +-------- Pleurobema_cordatum **
| |
| +-------- Pleurobema_beadleianum **
| |
| +-------- Hemistena_lata **
+--+
+-------- Fusconaia_flava **
|
+-------- Fusconaia_cerina **
|
+-------- Eurynia_dilatata **
|
+-------- Elliptio_arca **
|
+-------- Cyrtonaias_tampicoensis **
|
| +-- Plethobasus_cyphyus **
+-----+
| +-- Parvaspina_collina **
|
| +-- Pleurobema_athearni **
| |
+-----+-- Pleurobema_georgianum[1] **
| |
| +-- Pleurobema_hanleyianum[1] **
|
| +----- Pleurobema_oviforme[1] **
| |
+--+----- Pleurobema_clava **
|
| +-- Pleurobema_rubrum **
+--+
+-- Pleurobema_sintoxia **
Serb et al., 2003 [Publ 2659], fig. 2
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 29 |
29 |
20 |
4 |
0 |
0 |
0 |
38.0 |
+-- Lampsilis_siliquoidea **
+--------------------+
| +-- Lampsilis_ornata **
|
| +-------------------- Obliquaria_reflexa **
| |
| +-------------------- Amblema_plicata **
| |
| +-------------------- Reginaia_rotulata **
| |
--+ +-------------------- Potamilus_alatus **
| |
| | +-- Elliptoideus_sloatianus **
| +-----------------+
| | +-- Plectomerus_dombeyanus **
| |
| | +----- Medionidus_conradicus **
| +--------------+
| | | +-- Dromus_dromas **
+--+ +--+
| +-- Cyprogenia_aberti **
|
| +-------- Hemistena_lata **
+-----------+
| | +----- Elliptio_complanata **
| +--+
| | +-- Fusconaia_burkei **
| +--+
| +-- Fusconaia_flava **
|
| +----------------- Megalonaias_nervosa **
| |
| | +-------- Theliderma_cylindrica **
| | +-----+
+--+ | | +----- Theliderma_intermedia **
| | +--+
| | | +-- Theliderma_sparsa **
| | +--+
| | +-- Theliderma_metanevra **
+--+
| +----- Quadrula_verrucosa **
| +-----+
| | | +-- Quadrula_quadrula **
| | +--+
| | +-- Quadrula_nobilis **
+--+
| +-------- Pustulosa_kieneriana **
| |
| +-------- Pustulosa_infucata **
+--+
| +-- Pustulosa_pustulosa **
| +--+
| | +-- Pustulosa_succissa **
+--+
| +-- Pustulosa_nodulata **
+--+
+-- Pustulosa_petrina **
Serb et al., 2003 [Publ 2659], fig. 3
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 29 |
29 |
27 |
4 |
0 |
0 |
0 |
40.0 |
+-- Lampsilis_siliquoidea **
+-----------------------------------------+
| +-- Lampsilis_ornata **
|
--+ +----------------------------------------- Potamilus_alatus **
| |
| | +----- Medionidus_conradicus **
+--+ +--------------------------------+
| | | +-- Dromus_dromas **
| | +--+
| | +-- Cyprogenia_aberti **
+--+
| +----------------------------------- Obliquaria_reflexa **
| |
| | +-------------------------------- Amblema_plicata **
+--+ |
| | +----- Reginaia_rotulata **
| | +-----------------------+
+--+ | | +-- Elliptoideus_sloatianus **
| | +--+
| | +-- Plectomerus_dombeyanus **
| |
+--+ +-------- Hemistena_lata **
| +-----------------+
| | | +----- Elliptio_complanata **
| | +--+
| | | +-- Fusconaia_burkei **
| | +--+
+--+ +-- Fusconaia_flava **
|
| +----------------------- Megalonaias_nervosa **
| |
| | +-- Quadrula_verrucosa **
+--+ +-----------------+
| | +-- Quadrula_nobilis **
| |
+--+ +----------------- Quadrula_quadrula **
| |
| | +-------- Theliderma_cylindrica **
+--+ +-----+
| | | +----- Theliderma_sparsa **
| | +--+
| | | +-- Theliderma_intermedia **
+--+ +--+
| +-- Theliderma_metanevra **
|
| +----------- Pustulosa_kieneriana **
| |
+--+ +-------- Pustulosa_infucata **
| |
+--+ +-- Pustulosa_succissa **
| +--+
| | +-- Pustulosa_pustulosa **
+--+
| +-- Pustulosa_nodulata **
+--+
+-- Pustulosa_petrina **
Campbell et al., 2008 [Publ 2675], fig. 4
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 29 |
29 |
10 |
4 |
0 |
0 |
0 |
42.0 |
+----------------------- Uniomerus_declivis **
|
+----------------------- Plectomerus_dombeyanus **
|
| +-------------------- Cyrtonaias_tampicoensis **
| |
| +-------------------- Eurynia_dilatata **
| |
| +-------------------- Fusconaia_cerina **
| |
--+ +-------------------- Fusconaia_flava **
| |
| +-------------------- Hemistena_lata **
| |
| +-------------------- Plethobasus_cyphyus **
| |
| +-------------------- Parvaspina_collina **
| |
| +-------------------- Pleurobema_cordatum **
| |
| +-------------------- Pleuronaia_gibber **
+--+
+-------------------- Pleurobema_oviforme[2] **
|
+-------------------- Pleurobema_stabile **
|
| +----- Elliptio_arca **
+--------------+
| | +-- Elliptio_crassidens **
| +--+
| +-- Elliptio_arctata **
|
| +----------------- Pleurobema_decisum **
| |
| +----------------- Pleurobema_clava **
| |
| +----------------- Pleurobema_oviforme[1] **
| |
+--+ +-- Pleurobema_rubrum **
+--------------+
| +-- Pleurobema_sintoxia **
|
| +-------------- Pleurobema_beadleianum **
| |
+--+ +----------- Pleurobema_pyriforme **
| |
+--+ +-------- Pleurobema_rubellum **
| |
| +-------- Pleurobema_hanleyianum[2] **
+--+
+-------- Pleurobema_georgianum[2] **
|
| +----- Pleurobema_hanleyianum[1] **
+--+
| +-- Pleurobema_georgianum[1] **
+--+
+-- Pleurobema_athearni **
Campbell et al., 2008 [Publ 2675], fig. 6
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 29 |
29 |
24 |
3 |
0 |
0 |
0 |
41.0 |
+----------------------------------------- Cyrtonaias_tampicoensis **
|
+----------------------------------------- Plectomerus_dombeyanus **
--+
| +-------------------------------------- Parvaspina_collina **
| |
| | +----------------------------------- Hemistena_lata **
+--+ |
| | +----- Plethobasus_cyphyus **
| | +--------------------------+
+--+ | | +-- Elliptio_arca **
| | +--+
| | +-- Elliptio_crassidens **
| |
+--+ +-- Fusconaia_cerina **
| +--------------------------+
| | +-- Fusconaia_flava **
| |
| | +-- Eurynia_dilatata **
+--+ +-----------------------+
| | +-- Pleuronaia_gibber **
| |
| | +-- Pleurobema_cordatum **
+--+ +--+
| | +-- Pleurobema_rubrum[1] **
| +-----------------+
| | | +-- Pleurobema_rubrum[2] **
| | +--+
+--+ +-- Pleurobema_sintoxia **
|
| +-------------------- Pleurobema_stabile **
| |
+--+ +----------------- Pleurobema_pyriforme **
| |
| | +----- Pleurobema_decisum **
+--+ +--------+
| | | +-- Pleurobema_hanleyianum **
| | +--+
| | +-- Pleurobema_georgianum[2] **
+--+
| +-- Pleurobema_georgianum[1] **
| +--------+
| | +-- Pleurobema_athearni **
| |
+--+ +-- Pleurobema_oviforme **
| +-----+
| | +-- Pleurobema_clava **
| |
+--+ +----- Pleurobema_beadleianum **
| |
| +----- Pleurobema_rubellum **
+--+
+----- Pleurobema_strodeanum **
|
| +-- Pleurobema_perovatum **
+--+
+-- Pleurobema_taitianum **
Boyer et al., 2011 [Publ 2718], fig. 3
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 29 |
29 |
22 |
4 |
6 |
6 |
1 |
35.0 |
+----- Alasmidonta_marginata
|
+----- Pyganodon_grandis
|
+-----------------------+ +-- Lasmigona_complanata
| +--+
| | +-- Lasmigona_costata
| |
| | +-- Platynaias_compressa
| +--+
| +-- Strophitus_undulatus
|
--+ +-------------------------- Reginaia_ebenus **
| |
| | +-- Fusconaia_flava **
| | +--+
| | | +-- Pleurobema_sintoxia **
| | +--------------+
| | | | +-- Eurynia_dilatata **
| | | +--+
+--+ +--+ +-- Elliptio_crassidens **
| | |
| | | +----------------- Megalonaias_nervosa **
| | +--+
| | | +-------------- Quadrula_quadrula **
| | +--+
| | | +----------- Quadrula_verrucosa **
| | +--+
| | | +-------- Theliderma_metanevra **
+--+ +--+
| | +----- Pustulosa_pustulosa **
| +--+
| | +-- Cyclonaias_tuberculata **
| +--+
| +-- Pustulosa_nodulata **
|
| +----------------- Amblema_plicata **
| |
| | +-------------- Toxolasma_parvum **
+-----+ |
| | +----------- Obliquaria_reflexa **
| | |
+--+ | +-------- Obovaria_olivaria **
| | |
| | +-------- Lampsilis_cardium **
| | |
+--+ | +-- Epioblasma_triquetra **
| +-----+
| | +-- Lampsilis_siliquoidea **
| |
| | +-- Ortmanniana_ligamentina **
| +-----+
| | +-- Lampsilis_higginsii **
+--+
| +-- Ligumia_recta **
+-----+
| +-- Lampsilis_teres **
|
| +----- Potamilus_fragilis **
+--+
| | +-- Potamilus_ohiensis **
| +--+
| +-- Potamilus_alatus **
|
| +----- Ellipsaria_lineolata **
+--+
| +-- Truncilla_truncata **
+--+
+-- Truncilla_donaciformis **
Boyer et al., 2011 [Publ 2718], fig. 4
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 29 |
29 |
25 |
4 |
9 |
9 |
1 |
36.0 |
+-------------- Strophitus_undulatus
|
| +----- Lasmigona_complanata
+--------------------+ +-----+
| | | | +-- Lasmigona_costata
| | | +--+
| +--+ +-- Arcidens_confragosus
| |
| | +-------- Utterbackia_imbecillis
| | |
| +--+-------- Alasmidonta_marginata
| |
| | +----- Pyganodon_grandis
| +--+
--+ | +-- Anodontoides_ferussacianus
| +--+
| +-- Platynaias_compressa
|
| +----------- Theliderma_metanevra **
| |
| +--------------------+ +-- Quadrula_verrucosa **
| | | +-----+
| | | | +-- Quadrula_quadrula **
| | +--+
| | | +----- Pustulosa_pustulosa **
| | +--+
+--+ | +-- Cyclonaias_tuberculata **
| +--+
| +-- Pustulosa_nodulata **
|
| +----------------------------- Megalonaias_nervosa **
| |
+--+ +-------------------------- Toxolasma_parvum **
| |
| | +----------------------- Amblema_plicata **
+--+ |
| | +-- Reginaia_ebenus **
| | +-----------------+
| | | +-- Obliquaria_reflexa **
+--+ |
| | +-------- Pleurobema_sintoxia **
| +-----------+
| | | +----- Elliptio_crassidens **
| | +--+
| | | +-- Fusconaia_flava **
+--+ +--+
| +-- Eurynia_dilatata **
|
| +----- Potamilus_fragilis **
| +-----------+
| | | +-- Potamilus_ohiensis **
| | +--+
| | +-- Potamilus_alatus **
| |
+--+ +----- Ellipsaria_lineolata **
| +--------+
| | | +-- Truncilla_donaciformis **
| | +--+
| | +-- Truncilla_truncata **
+--+
| +----------- Lampsilis_cardium **
| |
| | +----- Ligumia_recta **
| +-----+
+--+ | +-- Obovaria_olivaria **
| +--+
| +-- Epioblasma_triquetra **
|
| +-------- Lampsilis_teres **
+--+
| +----- Lampsilis_siliquoidea **
+--+
| +-- Ortmanniana_ligamentina **
+--+
+-- Lampsilis_higginsii **
Inoue et al., 2020 [Publ 4128], fig. 2
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 29 |
29 |
17 |
4 |
0 |
0 |
0 |
36.0 |
+-------------------------- Quadrula_quadrula **
|
| +-- Reginaia_ebenus **
| +--------------------+
| | +-- Reginaia_rotulata **
--+ |
| | +----- Amblema_plicata **
| | +--------------+
| | | | +-- Amblema_elliottii **
| | | +--+
+--+ | +-- Amblema_neislerii **
| |
| | +----- Popenaias_popeii **
| | +--+
| | | | +-- Popenaias_berezai **
| | | +--+
| | | +-- Popenaias_semirasa **
+--+ |
| +--------+ +-- Psoronaias_profunda **
| | | |
| | | +-- Psoronaias_semigranosa[1] **
| | | |
| | | +-- Psoronaias_semigranosa[2] **
| | | |
| | +-----+-- Psoronaias_guatemalensis[1] **
| | |
| | +-- Psoronaias_guatemalensis[2] **
+--+ |
| +-- Psoronaias_percompressa **
| |
| +-- Psoronaias_semigranosa[3] **
|
| +-- Glebula_rotundata **
| +-----------+
| | +-- Cyrtonaias_tampicoensis **
| |
+--+ +----------- Potamilus_alatus **
| |
| | +-- Ptychobranchus_fasciolaris **
+--+ +-----+
| | +-- Truncilla_truncata **
| |
| | +----- Ortmanniana_ligamentina **
+--+ |
| +----- Epioblasma_triquetra **
| |
| +----- Lampsilis_cardium **
| |
+--+----- Ligumia_recta **
|
+----- Cambarunio_iris **
|
+----- Pachynaias_spheniopsis **
|
| +-- Cyrtonaias_explicata **
+--+
+-- Sphenonaias_microdon **
Inoue et al., 2020 [Publ 4128], fig. 3
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 29 |
29 |
27 |
4 |
0 |
0 |
0 |
35.0 |
+----------------------------------------- Quadrula_quadrula **
|
| +-- Reginaia_ebenus **
| +-----------------------------------+
--+ | +-- Reginaia_rotulata **
| |
| | +----- Amblema_neislerii **
| | +-----------------------------+
| | | | +-- Amblema_plicata **
+--+ | +--+
| | +-- Amblema_elliottii **
| |
| | +----- Popenaias_popeii **
| | +--------+
| | | | +-- Popenaias_berezai **
| | | +--+
+--+ | +-- Popenaias_semirasa **
| +-----------------+
| | | +----------- Psoronaias_profunda **
| | | |
| | | | +----- Psoronaias_guatemalensis[1] **
| | +--+ +--+
| | | | | +-- Psoronaias_semigranosa[1] **
| | | | +--+
| | +--+ +-- Psoronaias_percompressa **
+--+ |
| | +----- Psoronaias_semigranosa[3] **
| +--+
| | +-- Psoronaias_guatemalensis[2] **
| +--+
| +-- Psoronaias_semigranosa[2] **
|
| +----------------------------- Cyrtonaias_tampicoensis **
| |
+--+ +-------------------------- Ptychobranchus_fasciolaris **
| |
+--+ +----------------------- Ligumia_recta **
| |
+--+ +-------------------- Potamilus_alatus **
| |
+--+ +----------------- Pachynaias_spheniopsis **
| |
| | +-- Epioblasma_triquetra **
+--+ +-----------+
| | +-- Truncilla_truncata **
| |
+--+ +-- Ortmanniana_ligamentina **
| +--------+
| | +-- Lampsilis_cardium **
+--+
| +-------- Cambarunio_iris **
+--+
| +----- Sphenonaias_microdon[2] **
+--+
| +-- Cyrtonaias_explicata **
+--+
+-- Sphenonaias_microdon[1] **
Campbell et al., 2008 [Publ 2675], fig. 2
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 28 |
28 |
18 |
3 |
0 |
0 |
0 |
41.0 |
+----------------------- Cyrtonaias_tampicoensis **
|
+----------------------- Plectomerus_dombeyanus **
|
| +-- Pleurobema_cordatum **
| |
| +-----------+-- Pleurobema_rubrum **
| | |
| | +-- Pleurobema_sintoxia **
| |
--+ +-----+ +-- Fusconaia_cerina **
| | | +--------+
| | | | +-- Fusconaia_flava **
| | | |
| | +--+ +-------- Pleuronaia_gibber **
| | | |
| | | | +----- Eurynia_dilatata **
| | +--+ |
| | | +----- Hemistena_lata **
| | | |
| | +--+----- Plethobasus_cyphyus **
+--+ |
| +----- Parvaspina_collina **
| |
| | +-- Elliptio_arca **
| +--+
| +-- Elliptio_crassidens **
|
| +----------------- Pleurobema_stabile **
| |
| | +-------------- Pleurobema_pyriforme **
| | |
+--+ | +-- Pleurobema_georgianum[2] **
| +-----------+
| | +-- Pleurobema_athearni **
| |
| | +----- Pleurobema_decisum **
+--+--------+
| | +-- Pleurobema_hanleyianum **
| +--+
| +-- Pleurobema_georgianum[1] **
|
| +-- Pleurobema_clava **
| +--------+
| | +-- Pleurobema_oviforme **
+--+
| +-------- Pleurobema_rubellum **
| |
+--+-------- Pleurobema_strodeanum **
|
| +----- Pleurobema_beadleianum **
+--+
| +-- Pleurobema_taitianum **
+--+
+-- Pleurobema_perovatum **
Perkins et al., 2017 [Publ 3777], fig. 3
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 28 |
28 |
11 |
3 |
0 |
0 |
0 |
41.0 |
+----------------------- Plectomerus_dombeyanus **
|
+----------------------- Uniomerus_declivis **
|
| +-- Parvaspina_steinstansana **
| |
| +-- Parvaspina_collina[1] **
--+ |
| +-----------------+-- Parvaspina_collina[2] **
| | |
| | +-- Parvaspina_collina[3] **
| | |
| | +-- Parvaspina_collina[4] **
| |
+--+ +----------------- Elliptio_arca **
| |
| +----------------- Hemistena_lata **
| |
| | +-- Elliptio_arctata **
| +--------------+
| | +-- Elliptio_crassidens **
+--+
| +-------------- Eurynia_dilatata **
| |
| | +----------- Pleurobema_oviforme **
| | |
| | +----------- Pleurobema_clava **
| | |
+--+ +----------- Pleurobema_pyriforme **
| |
| +----------- Pleurobema_georgianum **
| |
| +----------- Pleurobema_beadleianum **
| |
| +----------- Pleurobema_hanleyianum **
+--+
+----------- Pleurobema_decisum **
|
| +----- Pleurobema_rubellum **
+-----+
| | +-- Pleurobema_sintoxia **
| +--+
| +-- Pleurobema_rubrum **
|
| +-------- Elliptio_spinosa **
| |
+--+ +----- Fusconaia_escambia **
| |
| +----- Fusconaia_flava **
+--+
+----- Fusconaia_cerina **
|
| +-- Pleuronaia_gibber **
+--+
+-- Pleuronaia_dolabelloides **
Smith et al., 2020 [Publ 4129], fig. 1
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 28 |
28 |
26 |
5 |
0 |
0 |
0 |
32.0 |
+-------------------------------------- Quadrula_quadrula **
|
| +-- Pleurobema_strodeanum **
--+ +--------------------------------+
| | +-- Elliptoideus_sloatianus **
| |
| | +----- Reginaia_ebenus **
+--+ +--------------------------+
| | | +-- Amblema_plicata **
| | +--+
| | +-- Plectomerus_dombeyanus **
+--+
| +-- Popenaias_popeii **
| +--------------------------+
| | +-- Psoronaias_semigranosa **
| |
+--+ +-------------------------- Obliquaria_reflexa **
| |
| | +----------------------- Cyrtonaias_tampicoensis **
+--+ |
| | +-- Pachynaias_spheniopsis **
| | +-----------------+
+--+ | +-- Lampsilis_cardium **
| |
| | +----------- Ellipsaria_lineolata **
| | +-----+
+--+ | | +-------- Truncilla_truncata **
| | +--+
| | | +----- Truncilla_donaciformis **
| | +--+
| | | +-- Truncilla_cognata **
| | +--+
+--+ +-- Truncilla_macrodon **
|
| +-------------- Atlanticoncha_ochracea **
| |
| | +-- Potamilus_fragilis **
| | +-----+
| | | +-- Potamilus_leptodon **
+--+ +--+
| | | +----- Potamilus_metnecktayi **
| | +--+
| | | +-- Potamilus_alatus **
| | +--+
+--+ +-- Potamilus_purpuratus **
|
| +-- Potamilus_capax **
| +-----+
| | +-- Potamilus_inflatus **
+--+
| +----- Potamilus_ohiensis **
+--+
| +-- Potamilus_streckersoni **
+--+
+-- Potamilus_amphichaenus **
Smith et al., 2020 [Publ 4129], fig. 2-3
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 28 |
28 |
26 |
5 |
0 |
0 |
0 |
32.0 |
+----------------------------------------- Quadrula_quadrula **
|
| +-- Pleurobema_strodeanum **
--+ +-----------------------------------+
| | +-- Elliptoideus_sloatianus **
| |
| | +----- Reginaia_ebenus **
+--+ +-----------------------------+
| | | +-- Amblema_plicata **
| | +--+
| | +-- Plectomerus_dombeyanus **
+--+
| +-- Popenaias_popeii **
| +-----------------------------+
| | +-- Psoronaias_semigranosa **
| |
+--+ +----------------------------- Obliquaria_reflexa **
| |
| | +-------------------------- Cyrtonaias_tampicoensis **
+--+ |
| | +-- Pachynaias_spheniopsis **
| | +--------------------+
+--+ | +-- Lampsilis_cardium **
| |
| | +----------- Ellipsaria_lineolata **
| | +--------+
+--+ | | +-------- Truncilla_truncata **
| | +--+
| | | +----- Truncilla_donaciformis **
| | +--+
| | | +-- Truncilla_cognata **
+--+ +--+
| +-- Truncilla_macrodon **
|
| +----------------- Atlanticoncha_ochracea **
| |
| | +-- Potamilus_capax **
+--+ +-----------+
| | +-- Potamilus_inflatus **
| |
| | +----- Potamilus_ohiensis **
+--+ +-----+
| | | +-- Potamilus_streckersoni **
| | +--+
| | +-- Potamilus_amphichaenus **
+--+
| +-- Potamilus_fragilis **
| +-----+
| | +-- Potamilus_leptodon **
+--+
| +----- Potamilus_metnecktayi **
+--+
| +-- Potamilus_alatus **
+--+
+-- Potamilus_purpuratus **
Neemuchwala et al., 2023 [Publ 4292], fig. 2
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 26 |
26 |
24 |
5 |
1 |
1 |
1 |
32.0 |
+-------------------------------- Gonidea_angulata
|
| +-------- Pleurobema_strodeanum **
| +--------------------+
--+ | | +----- Amblema_plicata **
| | +--+
| | | +-- Lampsilis_cardium **
| | +--+
+--+ +-- Popenaias_popeii **
|
| +-------------------------- Megalonaias_nervosa **
| |
| | +-------- Uniomerus_carolinianus **
| | +--------------+
+--+ | | +----- Uniomerus_columbensis **
| | +--+
| | | +-- Uniomerus_tetralasmus **
| | +--+
| | +-- Uniomerus_declivis **
+--+
| +-------- Theliderma_cylindrica **
| +-----------+
| | | +----- Theliderma_metanevra **
| | +--+
| | | +-- Theliderma_sparsa **
| | +--+
+--+ +-- Theliderma_johnsoni **
|
| +----------------- Cyclonaias_tuberculata **
| |
| | +-------- Quadrula_nobilis **
| | +-----+
+--+ | | +----- Quadrula_verrucosa **
| | +--+
| | | +-- Quadrula_fragosa **
| | +--+
| | +-- Quadrula_quadrula **
+--+
| +----- Pustulosa_nodulata **
| +-----+
| | | +-- Pustulosa_necki **
| | +--+
| | +-- Pustulosa_petrina **
+--+
| +-- Pustulosa_kieneriana **
| +-----+
| | +-- Pustulosa_pustulosa **
+--+
| +----- Pustulosa_succissa **
+--+
| +-- Pustulosa_infucata **
+--+
+-- Pustulosa_kleiniana **
Huang et al., 2019 [Publ 3843], fig. S3
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 25 |
25 |
23 |
4 |
176 |
137 |
3 |
22.0 |
+----- Chamberlainia_somsakpanhai
+--------------------------------------+
| | +-- Sinohyriopsis_cumingii
| +--+
| +-- Sinohyriopsis_schlegelii
|
| +-- Potomida_littoralis
| +-----+
| | +-- Pronodularia_japanensis
| +-----+
| | | +----- Lamprotula_leaii
| | +--+
| | | +-- Lamprotula_caveata
| | +--+
| | +-- Lamprotula_cornuumlunae
--+ +--------------------------+
| | | +----- Sinosolenaia_recognita
| | | +-----+
| | | | | +-- Sinosolenaia_oleivora
| | | | +--+
| | | | +-- Sinosolenaia_iridinea
| | +--+
| | | +----- Gonidea_angulata
| | | +--+
| | | | | +-- Leguminaia_wheatleyi
| | | | +--+
| | +--+ +-- Microcondylaea_bonellii
| | |
| | | +----- Ptychorhynchus_pfisteri
+--+ +--+
| | +-- Parvasolenaia_rivularis
| +--+
| +-- Koreosolenaia_sitgyensis
|
| +-- Megalonaias_nervosa **
| +--+
| | +-- Uniomerus_carolinianus **
| +--------------------------------+
| | | +-- Quadrula_quadrula **
| | +--+
| | +-- Quadrula_verrucosa **
| |
| | +-------- Elliptoideus_sloatianus **
+--+ |
| +--------------------------+ +-- Elliptio_complanata **
| | | +--+
| | | | +-- Eurynia_dilatata **
| | +--+
| | | +-- Fusconaia_flava **
+--+ +--+
| +-- Pleurobema_sintoxia **
|
| +-------------------------------- Amblema_plicata **
| |
+--+ +----------------------------- Toxolasma_parvum **
| |
+--+ +-- Obliquaria_reflexa **
| +-----------------------+
| | +-- Plectomerus_dombeyanus **
+--+
| +----------------------- Potamilus_leptodon **
| |
+--+ +-------------------- Potamilus_alatus **
| |
| | +-- Medionidus_conradicus **
+--+ +--------------+
| | +-- Ptychobranchus_subtentus **
| |
+--+ +-- Ellipsaria_lineolata **
| +-----------+
| | +-- Truncilla_truncata **
+--+
| +----------- Leaunio_lienosus **
| |
+--+ +-- Venustaconcha_ellipsiformis **
| +-----+
| | +-- Cambarunio_iris **
+--+
| +----- Ortmanniana_ligamentina **
+--+
| +-- Lampsilis_cardium **
+--+
+-- Lampsilis_ornata **
Huang et al., 2019 [Publ 3843], fig. S4
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 25 |
127 |
70 |
4 |
177 |
138 |
3 |
128.0 |
+-- Parreysia_keralaensis
|
+--------------------+-- Parreysia_nagpoorensis
| |
| +-- Parreysia_plagiosoma
|
| +----- Trapezidens_exolescens
| +--------+
| | | +-- Trapezidens_angustior
| | +--+
+--------+ +-- Trapezidens_dolichorhynchus
| |
| | +----------- Lamellidens_corrianus
| | |
| +--+ +-------- Lamellidens_savadiensis
| | |
| | | +----- Lamellidens_marginalis
| +--+ |
| | +----- Lamellidens_ferrugineus
| | |
+--+ +--+ +-- Lamellidens_mainwaringi[1]
| | | |
| | | +-- Lamellidens_mainwaringi[2]
| | | |
| | +--+-- Lamellidens_lamellatus
| | |
| | +-- Lamellidens_generosus
| | |
| | +-- Lamellidens_jenkinsianus
| |
| | +----- Leoparreysia_tavoyensis
| | +--------------+
| | | | +-- Leoparreysia_canefrii
| | | +--+
| | | +-- Leoparreysia_olivacea
| | |
| | | +-------- Coelatura_aegyptiaca[2]
| | | |
| | | +-------- Nitia_teretiuscula
| +--+ +--------+
| | | | +----- Coelatura_aegyptiaca[1]
| | | | |
| | | +--+ +-- Coelatura_gabonensis
| | | | |
| | | +--+-- Coelatura_kunenensis
| | | |
| | | +-- Prisodontopsis_mweruensis
| +--+
| | +----- Indonaia_shurtleffiana
| | |
| | | +-- Indonaia_bonneaudii
| | +--------+--+
| | | | +-- Indonaia_rugosa
| | | |
| | | | +-- Indonaia_andersoniana
| | | +--+
| +--+ +-- Indonaia_caerulea
| |
| | +----------- Indochinella_pugio
| | |
| | | +----- Scabies_humilis
| +--+ +--+
| | | | +-- Scabies_phaselus
| | | +--+
| +--+ +-- Scabies_mandarinus
| |
| | +----- Radiatula_mouhoti
| +--+
| | +-- Radiatula_myitkyinae
| +--+
| +-- Radiatula_chaudhurii
|
| +-------------------- Chamberlainia_somsakpanhai
| |
| +-------------------- Megalonaias_nervosa **
| |
| +-------------------- Uniomerus_carolinianus **
| |
| | +-- Quadrula_quadrula **
| +-----------------+
| | +-- Quadrula_verrucosa **
| |
| | +-- Sinohyriopsis_cumingii
| +-----------------+
| | +-- Sinohyriopsis_schlegelii
| |
| | +-- Potomida_littoralis
| | +-----+
| | | +-- Pronodularia_japanensis
| +-----------+
| | | +----- Lamprotula_leaii
| | +--+
| | | +-- Lamprotula_caveata
| | +--+
| | +-- Lamprotula_cornuumlunae
| |
| | +-------- Sinosolenaia_carinata
--+ | |
| | | +----- Gonidea_angulata
| | +--+
| | | | +-- Leguminaia_wheatleyi
| | | +--+
| | | +-- Microcondylaea_bonellii
| +-----------+
| | | +----- Ptychorhynchus_pfisteri
| | +--+
| | | | +-- Parvasolenaia_rivularis
| | | +--+
| | | +-- Koreosolenaia_sitgyensis
| | |
| | | +----- Sinosolenaia_recognita
| | +--+
| | | +-- Sinosolenaia_iridinea
| | +--+
| | +-- Sinosolenaia_oleivora
| |
| | +----- Indopseudodon_salwenianus
| | +-----+
| | | | +-- Indopseudodon_indawgyiensis
| | | +--+
| | | +-- Indopseudodon_bogani
| | |
| | | +-------- Nyeinchanconcha_nyeinchani
| | | |
| | | +-------- Thaiconcha_callifera
| +--------+ |
| | | +-------- Sundadontina_cumingii
| | | |
| | | +-------- Sundadontina_tumida[1]
| | | |
| | | +-------- Sundadontina_tumida[2]
| | | |
| | | | +----- Pilsbryoconcha_exilis[2]
| | +--+--+
| | | | +-- Pilsbryoconcha_carinifera
| | | +--+
| | | +-- Pilsbryoconcha_exilis[1]
| | |
| | | +----- Pseudodon_vagulus
| | +--+
| | | | +-- Pseudodon_mekongi
| | | +--+
| | | +-- Pseudodon_vondembuschianus
| | |
| | | +----- Bineurus_loeiensis
| | +--+
| | | +-- Bineurus_mouhotii
| | +--+
| | +-- Bineurus_exilis
| |
| | +-------------- Amblema_plicata **
| | |
| | +-------------- Toxolasma_parvum **
| | |
| | | +-- Obliquaria_reflexa **
| | +-----------+
| | | +-- Plectomerus_dombeyanus **
| | |
| | | +----------- Elliptoideus_sloatianus **
| | +--+
| | | | +-------- Elliptio_complanata **
| | | +--+
| | | | +----- Eurynia_dilatata **
| +-----+ +--+
| | | | +-- Fusconaia_flava **
| | | +--+
| | | +-- Pleurobema_sintoxia **
| | |
| | | +-------- Ellipsaria_lineolata **
| | | |
| | | +-------- Potamilus_leptodon **
| | | |
| | | +-------- Medionidus_conradicus **
| | | |
+-----+ | +-------- Potamilus_alatus **
| | |
| | +-------- Ptychobranchus_subtentus **
| +-----+
| +-------- Truncilla_truncata **
| |
| +-------- Venustaconcha_ellipsiformis **
| |
| +-------- Cambarunio_iris **
| |
| +-------- Leaunio_lienosus **
| |
| | +----- Ortmanniana_ligamentina **
| +--+
| | +-- Lampsilis_cardium **
| +--+
| +-- Lampsilis_ornata **
|
| +-------------- Hyriopsis_desowitzi
| |
| +-------------- Hyriopsis_myersiana
| |
| +-------------- Hyriopsis_panhai
| |
| +--+ +-- Hyriopsis_bialata
| | +-----------+
| | | +-- Hyriopsis_khoratensis
| | |
| | | +----------- Rectidens_sumatrensis
| | | |
| | +--+ +-- Ensidens_telus
| | | +-----+
| | | | +-- Ensidens_ingallsianus
| | +--+
| | | +----- Ensidens_sagittarius
| | +--+
+--+ | +-- Ensidens_spiculus
| | +--+
| | +-- Ensidens_jaculus
| |
| | +----------- Physunio_modelli
| | |
| | +----------- Lens
| | |
| | +----------- Physunio_superbus
| | |
| | | +-------- Yaukthwa_inlenensis
| | +--+
| +-----+ | +----- Yaukthwa_panhai
| | +--+
| | | +-- Yaukthwa_nesemanni
| | +--+
| | +-- Yaukthwa_rectangularis
| |
| | +-- Lens_comptus
| | +-----+
| | | +-- Lens_novoselovi
| +--+
| | +----- Lens_eximius
| +--+
| | +-- Lens_contradens
| +--+
| +-- Lens_micropterus
|
| +----------------- Acuticosta_chinensis
| |
| +----------------- Alasmidonta_marginata
| |
| +----------------- Anodonta_anatina
| |
| +----------------- Anemina_arcaeformis
| |
| +----------------- Anodonta_cygnea
| |
| +----------------- Cristaria_plicata
| |
| +----------------- Anodonta_nuttalliana
| |
| +----------------- Lasmigona_costata
| |
| +----------------- Lepidodesma_languilati
| |
| +----------------- Pseudanodonta_complanata
| |
| +----------------- Pyganodon_grandis
| |
| +----------------- Simpsonaias_ambigua
| |
| +----------------- Strophitus_undulatus
| |
| | +-- Schistodesmus_lampreyanus
| +--------------+
| | +-- Schistodesmus_spinosus
| |
| | +-- Utterbackia_imbecillis
| +--------------+
| | +-- Utterbackia_peninsularis
| |
| | +-------- Lanceolaria_gladiola
| +--------+
| | | +----- Lanceolaria_grayii
| | +--+
+--+ | +-- Lanceolaria_lanceolata
| +--+
| +-- Lanceolaria_triformis
|
| +-------- Unio_tumidus
+--------+
| | +----- Unio_crassus
| +--+
| | +-- Unio_delphinus
| +--+
| +-- Unio_pictorum
|
| +-------- Aculamprotula_tortuosa
| |
+--------+ +-- Aculamprotula_coreana
| | +--+
| | | +-- Aculamprotula_scripta
| +--+
| | +-- Aculamprotula_fibrosa
| +--+
| +-- Aculamprotula_tientsinensis
|
| +-------- Cuneopsis_heudei
| |
| +-------- Cuneopsis_celtiformis
| |
+--------+-------- Tchangsinaia_piscicula
| |
| +-------- Cuneopsis_rufescens
| |
| | +----- Nodularia_douglasiae[2]
| +--+
| | +-- Nodularia_douglasiae[1]
| +--+
| +-- Nodularia_nuxpersicae
|
| +-------------- Pletholophus_tenuis
| |
+--+ +-- Sinanodonta_angula
| +--------+
| | +-- Sinanodonta_lucida
+--+
| +-------- Sinanodonta_woodiana[1]
+--+
| +----- Sinanodonta_jourdyi
+--+
| +-- Sinanodonta_elliptica
+--+
+-- Sinanodonta_woodiana[2]
Lopes-Lima et al., 2019 [Publ 3901], fig. 3
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 25 |
25 |
23 |
4 |
4 |
2 |
1 |
30.0 |
+-- Anodonta_nuttalliana
+--------------------------------+
| +-- Pyganodon_grandis
|
| +-------------------------------- Megalonaias_nervosa **
--+ |
| | +-- Pleurobema_oviforme **
| | +-----+
| | | +-- Pleurobema_riddellii **
| | |
+--+ +--------------------+ +-- Lampsilis_ornata **
| | | +--+
| | | | +-- Potamilus_leptodon **
| | +--+
| | | +-- Amblema_elliottii **
| | +--+
+--+ +-- Amblema_plicata **
|
| +-------------------------- Uniomerus_tetralasmus **
| |
| | +----------- Theliderma_cylindrica **
| | +-----------+
| | | | +-------- Theliderma_sparsa **
+--+ | +--+
| | | +----- Theliderma_intermedia **
| | +--+
| | | +-- Theliderma_metanevra **
| | +--+
+--+ +-- Theliderma_johnsoni **
|
| +----- Quadrula_quadrula **
| +--------------+
| | | +-- Quadrula_nobilis **
| | +--+
| | +-- Quadrula_verrucosa **
+--+
| +----------------- Pustulosa_kieneriana **
| |
| | +-- Pustulosa_infucata **
+--+ +-----------+
| | +-- Pustulosa_kleiniana **
| |
+--+ +-- Pustulosa_pustulosa **
| +--------+
| | +-- Pustulosa_succissa **
+--+
| +-------- Cyclonaias_tuberculata **
+--+
| +----- Pustulosa_necki **
+--+
| +-- Pustulosa_petrina **
+--+
+-- Pustulosa_nodulata **
Lopes-Lima et al., 2019 [Publ 3901], fig. 2
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 24 |
24 |
21 |
4 |
4 |
2 |
1 |
33.0 |
+-- Anodonta_nuttalliana
+-----------------------------------+
| +-- Pyganodon_grandis
|
| +-- Amblema_elliottii **
--+ +--------------------------------+
| | +-- Amblema_plicata **
| |
| | +-- Lampsilis_ornata **
+--+ +-----------------------------+
| | +-- Potamilus_leptodon **
| |
| | +-- Pleurobema_riddellii **
+--+ +--------------------------+
| | +-- Elliptio_crassidens **
| |
| | +----- Megalonaias_nervosa **
+--+ |
| +--------------------+ +-- Uniomerus_carolinianus **
| | | |
| | +--+-- Uniomerus_declivis **
| | |
+--+ +-- Uniomerus_tetralasmus **
|
| +----------------------- Quadrula_quadrula **
| |
| | +-- Quadrula_verrucosa **
+--+ +-----------------+
| | +-- Quadrula_nobilis **
| |
+--+ +-- Theliderma_metanevra **
| +--------------+
| | +-- Theliderma_johnsoni **
| |
+--+ +----- Pustulosa_kieneriana **
| +--------+
| | | +-- Pustulosa_infucata **
| | +--+
+--+ +-- Pustulosa_kleiniana **
|
| +----------- Cyclonaias_tuberculata **
| |
+--+ +-- Pustulosa_pustulosa **
| +-----+
| | +-- Pustulosa_succissa **
+--+
| +----- Pustulosa_nodulata **
+--+
| +-- Pustulosa_necki **
+--+
+-- Pustulosa_petrina **
Krebs et al., 2003 [Publ 2691], fig. 3
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 22 |
22 |
20 |
4 |
12 |
10 |
1 |
31.0 |
+----------- Utterbackiana_couperiana
|
+----------- Platynaias_compressa
|
+----------- Alasmidonta_triangulata
|
+-----------------+ +-- Lasmigona_complanata
| +--------+
| | +-- Lasmigona_costata
| |
| | +-- Strophitus_radiatus
| | +-----+
| | | +-- Strophitus_undulatus
| +--+
| | +----- Utterbackia_imbecillis
| +--+
| | +-- Pyganodon_grandis
--+ +--+
| +-- Anodonta
|
| +-------- Megalonaias_nervosa **
| +-----------------+
| | | +----- Quadrula_quadrula[2] **
| | +--+
| | | +-- Quadrula_quadrula[1] **
| | +--+
| | +-- Theliderma_metanevra **
| |
+--+ +----------- Elliptoideus_sloatianus **
| +-----------+
| | | +-------- Pleurobema_pyriforme **
| | +--+
| | | +----- Uniomerus_carolinianus **
| | +--+
| | | +-- Fusconaia_flava **
+--+ +--+
| +-- Eurynia_dilatata **
|
| +-------------------- Amblema_plicata **
| |
| | +----- Toxolasma_parvum **
| | +-----------+
+--+ | | +-- Plectomerus_dombeyanus **
| | +--+
| | +-- Obliquaria_reflexa **
| |
+--+ +----- Ptychobranchus_subtentus **
| +--------+
| | | +-- Potamilus_fragilis **
| | +--+
| | +-- Medionidus_conradicus **
+--+
| +-- Villosa_delumbis **
| +--------+
| | +-- Ellipsaria_lineolata **
+--+
| +-------- Lampsilis_teres **
+--+
| +----- Lampsilis_siliquoidea **
+--+
| +-- Potamilus_purpuratus **
+--+
+-- Potamilus_alatus **
Lopes-Lima et al., 2019 [Publ 3901], fig. 4
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 22 |
22 |
20 |
4 |
4 |
2 |
1 |
29.0 |
+-- Anodonta_nuttalliana
+--------------------------------+
| +-- Pyganodon_grandis
|
| +-- Amblema_elliottii **
| +-----+
| | +-- Amblema_plicata **
--+ |
| +-----------------------+ +-- Pleurobema_riddellii **
| | | +--+
| | | | +-- Pleurobema_oviforme **
| | +--+
| | | +-- Lampsilis_ornata **
+--+ +--+
| +-- Potamilus_leptodon **
|
| +----------------------------- Megalonaias_nervosa **
| |
+--+ +-------------------------- Uniomerus_tetralasmus **
| |
| | +-- Theliderma_metanevra **
+--+ +--------------------+
| | +-- Theliderma_johnsoni **
| |
+--+ +-------------------- Quadrula_quadrula **
| |
| | +-- Quadrula_verrucosa **
+--+ +--------------+
| | +-- Quadrula_nobilis **
| |
| | +----- Pustulosa_kieneriana **
+--+ +--------+
| | | +-- Pustulosa_infucata **
| | +--+
| | +-- Pustulosa_kleiniana **
+--+
| +-- Pustulosa_pustulosa **
| +--------+
| | +-- Pustulosa_succissa **
+--+
| +-------- Cyclonaias_tuberculata **
+--+
| +----- Pustulosa_necki **
+--+
| +-- Pustulosa_petrina **
+--+
+-- Pustulosa_nodulata **
Chapman et al., 2008 [Publ 2667], fig. 1
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 20 |
20 |
18 |
5 |
1 |
1 |
1 |
26.0 |
+-------------------------------------------- Pronodularia_japanensis
|
| +-- Quadrula_quadrula **
--+ +--------------------------------------+
| | +-- Pustulosa_pustulosa **
| |
+--+ +-- Pleurobema_sintoxia **
| +-----------------------------------+
| | +-- Fusconaia_flava **
+--+
| +----------------------------------- Amblema_plicata **
| |
+--+ +-------------------------------- Plectomerus_dombeyanus **
| |
+--+ +----------------------------- Popenaias_popeii **
| |
+--+ +-------------------------- Toxolasma_lividum **
| |
+--+ +----------------------- Obliquaria_reflexa **
| |
| | +-- Cyrtonaias_tampicoensis **
+--+ +-----------------+
| | +-- Glebula_rotundata **
| |
+--+ +-- Ptychobranchus_fasciolaris **
| +--------------+
| | +-- Lemiox_rimosus **
| |
+--+ +-- Venustaconcha_ellipsiformis **
| +-----------+
| | +-- Obovaria_olivaria **
+--+
| +----------- Hamiota_subangulata **
+--+
| +-------- Lampsilis_ovata **
+--+
| +----- Ortmanniana_ligamentina **
+--+
| +-- Lampsilis_straminea **
+--+
+-- Lampsilis_hydiana **
Chapman et al., 2008 [Publ 2744], fig. 2
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 20 |
20 |
18 |
5 |
1 |
1 |
1 |
26.0 |
+-------------------------------------------- Pronodularia_japanensis
|
| +-- Quadrula_quadrula **
--+ +--------------------------------------+
| | +-- Pustulosa_pustulosa **
| |
+--+ +-- Pleurobema_sintoxia **
| +-----------------------------------+
| | +-- Fusconaia_flava **
+--+
| +----------------------------------- Amblema_plicata **
| |
+--+ +-------------------------------- Plectomerus_dombeyanus **
| |
+--+ +----------------------------- Popenaias_popeii **
| |
+--+ +-------------------------- Toxolasma_lividum **
| |
+--+ +----------------------- Obliquaria_reflexa **
| |
| | +-- Cyrtonaias_tampicoensis **
+--+ +-----------------+
| | +-- Glebula_rotundata **
| |
+--+ +-- Ptychobranchus_fasciolaris **
| +--------------+
| | +-- Lemiox_rimosus **
| |
+--+ +-- Venustaconcha_ellipsiformis **
| +-----------+
| | +-- Obovaria_olivaria **
+--+
| +----------- Hamiota_subangulata **
+--+
| +-------- Lampsilis_ovata **
+--+
| +----- Ortmanniana_ligamentina **
+--+
| +-- Lampsilis_straminea **
+--+
+-- Lampsilis_hydiana **
Porto-Hannes et al., 2019 [Publ 3969], fig. 2
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 20 |
20 |
16 |
2 |
0 |
0 |
0 |
27.0 |
+-------------------------- Amblema_plicata **
|
| +----------------------- Obliquaria_reflexa **
| |
--+ | +-- Cambarunio_iris **
| | +-----------------+
| | | +-- Leaunio_lienosus **
| | |
+--+ | +----------- Lampsilis_teres **
| | |
| +--------+ +-------- Ortmanniana_ligamentina **
| | | |
| | +--+ +-- Lampsilis_hydiana **
| | | +--+
+--+ | | +-- Lampsilis_virescens **
| +--+
| | +-- Lampsilis_siliquoidea **
| +--+
| +-- Lampsilis_radiata **
|
| +----- Obovaria_arkansasensis **
| +-----------+
| | | +-- Obovaria_olivaria **
| | +--+
+--+ +-- Obovaria_subrotunda **
|
| +-------------- Lampsilis_bracteata **
| |
+--+ +----------- Lampsilis_ornata **
| |
+--+ +-------- Lampsilis_ovata **
| |
+--+ +----- Lampsilis_cardium[3] **
| |
+--+ +-- Lampsilis_satura **
| |
+--+-- Lampsilis_cardium[1] **
|
+-- Lampsilis_cardium[2] **
Porto-Hannes et al., 2019 [Publ 3969], fig. 3
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 20 |
20 |
13 |
2 |
0 |
0 |
0 |
27.0 |
+-------------------- Obliquaria_reflexa **
|
+-------------------- Amblema_plicata **
|
| +----------------- Lampsilis_teres **
| |
| | +-- Leaunio_lienosus **
| +--------------+
--+ | +-- Cambarunio_iris **
| |
| | +----- Obovaria_arkansasensis **
| +-----------+
| | | +-- Obovaria_olivaria **
| | +--+
| | +-- Obovaria_subrotunda **
| |
+--+ +-------- Ortmanniana_ligamentina **
| |
+--------+ +-- Lampsilis_hydiana **
| | +--+
| | | +-- Lampsilis_virescens **
| +--+
| | +-- Lampsilis_siliquoidea **
| +--+
| +-- Lampsilis_radiata **
|
| +-------------- Lampsilis_bracteata **
| |
+--+ +----------- Lampsilis_ornata **
| |
+--+ +-------- Lampsilis_ovata **
| |
+--+ +----- Lampsilis_cardium[3] **
| |
+--+ +-- Lampsilis_satura **
| |
+--+-- Lampsilis_cardium[1] **
|
+-- Lampsilis_cardium[2] **
Smith et al., 2020 [Publ 4136], fig. 1
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 20 |
20 |
13 |
2 |
0 |
0 |
0 |
33.0 |
+-------------------------- Quadrula_quadrula **
|
--+ +----------------------- Elliptoideus_sloatianus **
| |
| | +-------------------- Elliptio_crassidens **
+--+ |
| +-------------------- Plethobasus_cyphyus **
| |
+--+ +----------------- Parvaspina_steinstansana **
| |
| | +-- Hemistena_lata **
| +--------------+
| | +-- Eurynia_dilatata **
+--+
| +-- Pleuronaia_barnesiana **
| +-----------+
| | +-- Pleurobema_clava **
| |
| | +----------- Fusconaia_subrotunda **
+--+ |
| | +-- Fusconaia_cor **
| +--------+
| | +-- Fusconaia_cuneolus **
+--+
| +-------- Fusconaia_flava **
| |
| +-------- Fusconaia_masoni **
| |
| | +-- Fusconaia_burkei **
+--+-----+
| +-- Fusconaia_escambia **
|
| +-- Fusconaia_iheringi **
| +--+
| | +-- Fusconaia_mitchelli **
+--+
| +-- Fusconaia_askewi **
+--+
+-- Fusconaia_chunii **
Breton et al., 2011 [Publ 3376], fig. 3
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 19 |
34 |
31 |
4 |
20 |
15 |
2 |
38.0 |
+----- Quadrula_quadrula **
+--------------------------+
| | +-- Cyclonaias_tuberculata **
| +--+
| +-- Pustulosa_pustulosa **
|
| +-------------------------- Pronodularia_japanensis
| |
| +--+ +----------------------- Sinohyriopsis_cumingii
| | | |
| | +--+ +-------------------- Cristaria_plicata
| | | |
--+ | | | +-- Lasmigona_complanata
| | +--+ +--------------+
| | | | +-- Lasmigona_costata
| | | |
| | +--+ +-------------- Strophitus_undulatus
| | | |
| | | | +-- Platynaias_compressa
| | | | +--+
| | +--+ | +-- Platynaias_subviridis
| | | +-----+
| | | | | +-- Alasmidonta_marginata
+--+ | | +--+
| | | +-- Alasmidonta_undulata
| +--+
| | +-- Pyganodon_lacustris
| | +-----+
| | | +-- Pyganodon_grandis
| +--+
| | +----- Utterbackia_peninsularis[2]
| +--+
| | +-- Utterbackia_peninsularis[1]
| +--+
| +-- Utterbackia_imbecillis
|
| +-------------------------- Megalonaias_nervosa **
| |
+--+ +----------------------- Fusconaia_flava **
| |
+--+ +-------------------- Reginaia_ebenus **
| |
| | +-------- Toxolasma_paulum[2] **
+--+ +--------+
| | | +----- Toxolasma_lividum **
| | +--+
| | | +-- Toxolasma_parvum **
+--+ +--+
| +-- Toxolasma_paulum[1] **
|
| +-------------- Ellipsaria_lineolata **
| |
+--+ +----------- Truncilla_macrodon **
| |
| | +-- Lemiox_rimosus **
+--+ +-----+
| | +-- Potamilus_metnecktayi **
| |
+--+ +-- Cambarunio_iris **
| +--+
| | +-- Venustaconcha_ellipsiformis **
+--+
| +-- Lampsilis_powellii **
| |
+--+-- Lampsilis_ornata **
|
+-- Lampsilis_siliquoidea **
Lydeard et al., 1996 [Publ 2656], fig. 3
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 18 |
18 |
14 |
4 |
9 |
6 |
1 |
30.0 |
+----- Utterbackia_imbecillis
+--+
| | +-- Utterbackiana_couperiana
| +--+
+--------------------+ +-- Utterbackiana
| |
| | +----- Strophitus_radiatus
| +--+
| | +-- Lasmigona_costata
| +--+
| +-- Alasmidonta_triangulata
--+
| +----- Megalonaias_nervosa **
| +--------------------+
| | | +-- Quadrula_quadrula **
| | +--+
| | +-- Theliderma_metanevra **
| |
+--+ +----- Elliptoideus_sloatianus **
| |
| +-----------------+ +-- Eurynia_dilatata **
| | | |
| | +--+-- Pleurobema_pyriforme **
| | |
+--+ +-- Uniomerus_carolinianus **
|
| +-------------------- Amblema_plicata **
| |
+--+ +-- Plectomerus_dombeyanus **
| +--------------+
| | +-- Obliquaria_reflexa **
+--+
| +-------------- Potamilus_fragilis **
| |
+--+ +----------- Ptychobranchus_subtentus **
| |
+--+ +-------- Medionidus_conradicus **
| |
+--+ +----- Potamilus_purpuratus **
| |
| | +-- Ellipsaria_lineolata **
+--+--+
| +-- Villosa_delumbis **
|
| +-- Lampsilis_teres **
+--+
+-- Lampsilis_siliquoidea **
Lydeard et al., 1996 [Publ 2656], fig. 4
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 18 |
18 |
16 |
4 |
10 |
6 |
1 |
30.0 |
+----- Utterbackia_imbecillis
+--+
| | +-- Utterbackiana_couperiana
| +--+
+-----------------------------+ +-- Anodonta
| |
| | +----- Strophitus_radiatus
| +--+
| | +-- Lasmigona_costata
| +--+
| +-- Alasmidonta_triangulata
--+
| +----- Megalonaias_nervosa **
| +-----------------------------+
| | | +-- Quadrula_quadrula **
| | +--+
| | +-- Theliderma_metanevra **
| |
+--+ +-------- Elliptoideus_sloatianus **
| +-----------------------+
| | | +----- Uniomerus_carolinianus **
| | +--+
| | | +-- Eurynia_dilatata **
+--+ +--+
| +-- Pleurobema_pyriforme **
|
| +----------------------------- Amblema_plicata **
+--+
| +-------------------------- Plectomerus_dombeyanus **
+--+
| +----------------------- Obliquaria_reflexa **
+--+
| +-------------------- Medionidus_conradicus **
+--+
| +----------------- Ptychobranchus_subtentus **
+--+
| +-------------- Potamilus_fragilis **
+--+
| +----------- Potamilus_purpuratus **
+--+
| +-------- Ellipsaria_lineolata **
+--+
| +----- Villosa_delumbis **
+--+
| +-- Lampsilis_teres **
+--+
+-- Lampsilis_siliquoidea **
Graf & Ó Foighil, 2000 [Publ 2648], fig. 2A
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 17 |
17 |
11 |
4 |
13 |
8 |
1 |
28.0 |
+-- Unio_pictorum
+--------------------+
| +-- Unio_caffer
|
| +----------- Anodonta_cygnea
| |
| +--------+ +-- Alasmidonta_marginata
--+ | | +-----+
| | | | +-- Lasmigona_costata
| | +--+
| | | +----- Strophitus_undulatus
| | +--+
| | | +-- Pyganodon_grandis
| | +--+
+--+ +-- Pyganodon_fragilis
|
| +-------- Quadrula_quadrula **
| +--------+
| | | +----- Eurynia_dilatata **
| | +--+
| | | +-- Pleurobema_sintoxia **
| | +--+
+--+ +-- Fusconaia_flava **
|
| +-------------- Amblema_plicata **
| |
| | +----------- Ligumia_recta **
| | |
+--+ +----------- Truncilla_truncata **
| |
| | +-- Ptychobranchus_fasciolaris **
| +--------+
+--+ +-- Sagittunio_nasutus **
|
| +----- Lampsilis_siliquoidea **
| +--+
| | | +-- Epioblasma_triquetra **
| | +--+
| | +-- Epioblasma_brevidens **
+--+
| +----- Ortmanniana_ligamentina **
| |
| +----- Cambarunio_iris **
+--+
+----- Leaunio_vanuxemensis **
|
| +-- Lampsilis_cardium **
+--+
+-- Lampsilis_fasciola **
Graf & Ó Foighil, 2000 [Publ 2648], fig. 2B
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 17 |
17 |
15 |
4 |
13 |
8 |
1 |
28.0 |
+-- Unio_pictorum
+-----------------------------+
| +-- Unio_caffer
|
| +----------- Anodonta_cygnea
| |
| +-----------------+ +-- Alasmidonta_marginata
--+ | | +-----+
| | | | +-- Lasmigona_costata
| | +--+
| | | +----- Strophitus_undulatus
| | +--+
| | | +-- Pyganodon_grandis
+--+ +--+
| +-- Pyganodon_fragilis
|
| +-------- Quadrula_quadrula **
| +-----------------+
| | | +----- Eurynia_dilatata **
| | +--+
| | | +-- Pleurobema_sintoxia **
+--+ +--+
| +-- Fusconaia_flava **
|
| +----------------------- Amblema_plicata **
| |
+--+ +-------------------- Truncilla_truncata **
| |
+--+ +----------------- Ligumia_recta **
| |
| | +-- Ptychobranchus_fasciolaris **
+--+ +-----------+
| | +-- Sagittunio_nasutus **
| |
+--+ +----------- Lampsilis_siliquoidea **
| |
| | +----- Ortmanniana_ligamentina **
+--+ +--+
| | | +-- Lampsilis_cardium **
| | +--+
| | +-- Lampsilis_fasciola **
+--+
| +-- Epioblasma_brevidens **
| +--+
| | +-- Epioblasma_triquetra **
+--+
| +-- Cambarunio_iris **
+--+
+-- Leaunio_vanuxemensis **
Grobler et al., 2011 [Publ 3359], fig. 2
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 17 |
17 |
13 |
2 |
0 |
0 |
0 |
26.0 |
+----------------- Amblema_plicata **
|
| +----------- Lemiox_rimosus **
| +--+
--+ | | +-------- Medionidus_conradicus **
| | +--+
| | | +----- Dromus_dromas **
| | +--+
| | | +-- Cyprogenia_stegaria **
+--+ +--+
| +-- Cyprogenia_aberti **
|
| +----------- Potamilus_alatus **
| |
| | +----- Lampsilis_siliquoidea **
| | +--+
+--+ | | +-- Lampsilis_higginsii **
| | +--+
| | +-- Ortmanniana_ligamentina **
| |
| | +----- Lampsilis_fasciola **
+--+--+
| | +-- Lampsilis_cardium **
| +--+
| +-- Lampsilis_ovata **
|
| +----- Cambarunio_iris **
| |
+--+----- Ligumia_recta **
|
| +-- Epioblasma_brevidens **
+--+
+-- Epioblasma_capsaeformis **
Chase et al., 2018 [Publ 3847], fig. 4
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 17 |
18 |
16 |
4 |
22 |
17 |
1 |
29.0 |
+-- Unio_delphinus
+--------------------+
| +-- Unio_tumidus
|
+--+ +-------------------- Sinanodonta_woodiana
| | |
| | | +-- Anodonta_cygnea
| | | +--------------+
| +--+ | +-- Anodonta_anatina
| | |
| | | +-- Pyganodon_lacustris
| | | +-----+
| +--+ | +-- Pyganodon_grandis
| | +-----+
| | | | +----- Utterbackia_peninsularis[2]
| | | +--+
| | | | +-- Utterbackia_peninsularis[1]
| | | +--+
| +--+ +-- Utterbackia_imbecillis
| |
| | +-- Lasmigona_costata
| | +--------+
| | | +-- Lasmigona_complanata
| | |
--+ +--+ +-- Alasmidonta_undulata
| | +-----+
| | | +-- Alasmidonta_marginata
| +--+
| | +----- Strophitus_undulatus
| +--+
| | +-- Platynaias_compressa
| +--+
| +-- Platynaias_subviridis
|
| +-- Echyridella_menziesii
| +--------+
| | +-- Reginaia_ebenus **
| +--------+
| | | +-------- Fusconaia_flava **
| | +--+
| | | +----- Megalonaias_nervosa **
| | +--+
| | | +-- Cyclonaias_tuberculata **
| | +--+
+-----+ +-- Pustulosa_pustulosa **
|
| +-------- Toxolasma_parvum **
| +--------+
| | | +----- Toxolasma_lividum **
| | +--+
| | | +-- Toxolasma_paulum **
| | +--+
+--+ +-- Toxolasma_texasiense **
|
| +-------------- Venustaconcha_ellipsiformis **
| |
| | +----- Lemiox_rimosus **
+--+ +-----+
| | | +-- Lampsilis_powellii **
| | +--+
+--+ +-- Lampsilis_siliquoidea **
|
| +-------- Potamilus_metnecktayi **
+--+
| +----- Cambarunio_iris **
+--+
| +-- Ellipsaria_lineolata **
+--+
+-- Truncilla_macrodon **
Wu et al., 2023 [Publ 4339], fig. 4, S5-aa
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 17 |
17 |
15 |
5 |
79 |
68 |
3 |
0.0 |
+-- Quadrula_quadrula **
+--------------------------+
| +-- Uniomerus_tetralasmus **
|
+--------+ +-- Elliptio_complanata **
| | +-----------------------+
| | | +-- Pleurobema_oviforme **
| +--+
| | +----------------------- Amblema_plicata **
| | |
| +--+ +-------------------- Popenaias_popeii **
| | |
| +--+ +----------------- Toxolasma_parvum **
| | |
| | | +----- Potamilus_leptodon **
| +--+ +--------+
| | | | +-- Potamilus_alatus **
| | | +--+
| | | +-- Potamilus_streckersoni **
| +--+
| | +----------- Leaunio_lienosus **
| | |
| | | +----- Lampsilis_ornata **
| +--+ +--+
| | | | +-- Lampsilis_cardium **
| | | +--+
| +--+ +-- Lampsilis_cariosa **
| |
| | +----- Venustaconcha_ellipsiformis **
| +--+
| | +-- Lampsilis_powellii **
| +--+
| +-- Lampsilis_siliquoidea **
--+
| +----- Chamberlainia_somsakpanhai
| +-----------------+
| | | +-- Sinohyriopsis_cumingii
| | +--+
| | +-- Sinohyriopsis_schlegelii
| |
| | +-- Hyriopsis_bialata
| | +--+
| +-----------+ | +-- Rectidens_sumatrensis
| | | +--------------+
| | | | | +-- Lens_contradens
| | | | +--+
| | | | +-- Physunio_superbus
| | | |
| | | | +----------- Potomida_littoralis
| | +--+ +-----+
| | | | | +-------- Pronodularia_japanensis
| | | | +--+
| | | | | +----- Lamprotula_leaii
| | | | +--+
| | | | | +-- Lamprotula_caveata
| | +--+ +--+
| | | +-- Lamprotula_cornuumlunae
| | |
| | | +-- Pseudodon_vondembuschianus
| | | +-----------+
| | | | +-- Pilsbryoconcha_exilis
| | +--+
| | | +----------- Microcondylaea_bonellii
+--+ | |
| +--+ +-- Parvasolenaia_rivularis
| | +-----+
| | | +-- Ptychorhynchus_pfisteri
| +--+
| | +----- Sinosolenaia_carinata
| +--+
| | +-- Sinosolenaia_iridinea
| +--+
| +-- Sinosolenaia_oleivora
|
| +-------------------------------- Lamellidens_marginalis
| |
| | +-- Lepidodesma_aligera
| | +--------------------------+
| | | +-- Lepidodesma_languilati
| | |
| | | +-------- Aculamprotula_tortuosa
| | | |
| | | +--------------+ +-- Aculamprotula_coreana
| | | | | +--+
+--+ | | | | +-- Aculamprotula_tientsinensis
| | | +--+
| | | | +-- Aculamprotula_polysticta
| | | +--+
| | | +-- Aculamprotula_scripta
| | |
| | +--+ +-------------- Unio_tumidus
| | | | +-----+
| | | | | | +----------- Unio_crassus
| | | | | +--+
+--+ | | | | +-------- Unio_elongatulus
| | | | +--+
| | | | | +----- Unio_pictorum
| | | | +--+
| | | | | +-- Unio_delphinus
| | +--+ +--+
| | | +-- Unio_mancus
| | |
| | | +-- Nodularia_breviconcha
| | | +--+
| | | | +-- Nodularia_nipponensis
| | | +-----------+
| | | | | +-- Pseudocuneopsis_capitata
| | | | +--+
| | +--+ +-- Pseudocuneopsis_sichuanensis
| | |
| | | +-------------- Tchangsinaia_piscicula
| | | |
+--+ +--+ +-- Schistodesmus_lampreyanus
| | +--------+
| | | +-- Schistodesmus_spinosus
| +--+
| | +-------- Cuneopsis_rufescens
| +--+
| | +----- Cuneopsis_celtiformis
| +--+
| | +-- Cuneopsis_demangei
| +--+
| +-- Cuneopsis_heudei
|
| +-------- Acuticosta_chinensis
| +--------+
| | | +----- Lanceolaria_gladiola
| | +--+
| | | +-- Lanceolaria_grayii
| | +--+
| | +-- Lanceolaria_lanceolata
| |
| | +----- Cristaria_plicata
| | +-----+
| | | | +-- Anemina_arcaeformis
+--------+ | +--+
| +--+ +-- Anemina_euscaphys
| | |
| | | +-------- Beringiana_fukuharai
| | +--+
| | | +----- Sinanodonta_lucida
| | +--+
| | | +-- Sinanodonta_tumens
| | +--+
| | +-- Sinanodonta_woodiana
+--+
| +-- Anodonta_nuttalliana
| +-----+
| | +-- Pseudanodonta_complanata
| +--+
| | | +----- Anodonta_anatina
| | +--+
| | | +-- Anodonta_cygnea
| | +--+
| | +-- Anodonta_exulcerata
+--+
| +----- Pyganodon_grandis
| +--+
| | | +-- Utterbackia_peninsularis
| | +--+
| | +-- Utterbackia_imbecillis
+--+
| +-- Alasmidonta_varicosa
| +--+
| | +-- Platynaias_compressa
+--+
| +-- Prolasmidonta_heterodon
+--+
+-- Lasmigona_complanata
Wu et al., 2024 [Publ 4350], fig. 8a, S1
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 17 |
17 |
15 |
5 |
75 |
69 |
3 |
0.0 |
+-- Quadrula_quadrula **
+--------------------------+
| +-- Uniomerus_tetralasmus **
|
+--------+ +-- Elliptio_complanata **
| | +-----------------------+
| | | +-- Pleurobema_oviforme **
| +--+
| | +----------------------- Amblema_plicata **
| | |
| +--+ +-------------------- Popenaias_popeii **
| | |
| +--+ +----------------- Toxolasma_parvum **
| | |
| | | +----- Potamilus_leptodon **
| +--+ +--------+
| | | | +-- Potamilus_alatus **
| | | +--+
| | | +-- Potamilus_streckersoni **
| +--+
| | +----------- Leaunio_lienosus **
| | |
| | | +----- Lampsilis_ornata **
| +--+ +--+
| | | | +-- Lampsilis_cardium **
| | | +--+
| +--+ +-- Lampsilis_cariosa **
| |
| | +----- Venustaconcha_ellipsiformis **
| +--+
| | +-- Lampsilis_powellii **
| +--+
| +-- Lampsilis_siliquoidea **
|
--+ +----- Chamberlainia_somsakpanhai
| +--------------------+
| | | +-- Sinohyriopsis_cumingii
| | +--+
| | +-- Sinohyriopsis_schlegelii
| |
| | +-- Hyriopsis_bialata
| | +--+
| +--------+ | +-- Rectidens_sumatrensis
| | | +-----------------+
| | | | | +-- Lens_contradens
| | | | +--+
| | | | +-- Physunio_superbus
| | | |
| | | | +----------- Potomida_littoralis
| | +--+ +--------+
| | | | | +-------- Pronodularia_japanensis
| | | | +--+
| | | | | +----- Lamprotula_leaii
| | | | +--+
| | | | | +-- Lamprotula_caveata
| | +--+ +--+
| | | +-- Lamprotula_cornuumlunae
| | |
| | | +-- Pseudodon_vondembuschianus
| | | +--------------+
| | | | +-- Pilsbryoconcha_exilis
| | +--+
| | | +-------------- Inversidens_rentianensis
| | | |
+--+ +--+ +-- Parvasolenaia_rivularis
| | +--------+
| | | +-- Ptychorhynchus_pfisteri
| +--+
| | +-------- Microcondylaea_bonellii
| +--+
| | +----- Sinosolenaia_carinata
| +--+
| | +-- Sinosolenaia_iridinea
| +--+
| +-- Sinosolenaia_oleivora
|
| +-------------------------------- Lamellidens_marginalis
| |
| | +-- Lepidodesma_aligera
| | +--------------------------+
| | | +-- Lepidodesma_languilati
| | |
| | | +-------- Aculamprotula_tortuosa
| | | |
| | | +--------------+ +-- Aculamprotula_coreana
| | | | | +--+
+--+ | | | | +-- Aculamprotula_tientsinensis
| | | +--+
| | | | +-- Aculamprotula_polysticta
| | | +--+
| | | +-- Aculamprotula_scripta
| | |
| | +--+ +-------------- Unio_tumidus
| | | | +-----+
| | | | | | +----------- Unio_crassus
| | | | | +--+
+--+ | | | | +-------- Unio_elongatulus
| | | | +--+
| | | | | +----- Unio_pictorum
| | | | +--+
| | | | | +-- Unio_delphinus
| | +--+ +--+
| | | +-- Unio_mancus
| | |
| | | +-- Nodularia_breviconcha
| | | +--------------+
| | | | +-- Nodularia_douglasiae
| | | |
| | | | +-- Pseudocuneopsis_capitata
| | | +--------------+
| | +--+ +-- Pseudocuneopsis_sichuanensis
| | |
| | | +-- Schistodesmus_lampreyanus
| | | +-----------+
+--+ | | +-- Schistodesmus_spinosus
| +--+
| | +----------- Tchangsinaia_piscicula
| +--+
| | +-------- Cuneopsis_rufescens
| +--+
| | +----- Cuneopsis_celtiformis
| +--+
| | +-- Cuneopsis_demangei
| +--+
| +-- Cuneopsis_heudei
|
| +-------- Acuticosta_chinensis
| +--------------+
| | | +----- Lanceolaria_gladiola
| | +--+
| | | +-- Lanceolaria_grayii
| | +--+
| | +-- Lanceolaria_lanceolata
| |
| | +-------------- Cristaria_plicata
| | |
+--+ +-----+ +-- Anemina_arcaeformis
| | | +--------+
| | | | +-- Anemina_euscaphys
| | +--+
| | | +-------- Beringiana_fukuharai
| | +--+
| | | +----- Sinanodonta_lucida
| | +--+
| | | +-- Sinanodonta_tumens
+--+ +--+
| +-- Sinanodonta_woodiana
|
| +-------- Pseudanodonta_complanata
| +--------+
| | | +----- Anodonta_anatina
| | +--+
| | | +-- Anodonta_cygnea
| | +--+
+--+ +-- Anodonta_exulcerata
|
| +-------------- Anodonta_nuttalliana
| |
| | +----- Pyganodon_grandis
+--+ +-----+
| | | +-- Utterbackia_imbecillis
| | +--+
+--+ +-- Utterbackia_peninsularis
|
| +-------- Lasmigona_complanata
+--+
| +----- Prolasmidonta_heterodon
+--+
| +-- Alasmidonta_varicosa
+--+
+-- Platynaias_compressa
Wu et al., 2024 [Publ 4350], fig. 8b, S2
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 17 |
17 |
14 |
5 |
75 |
69 |
3 |
1.0 |
+-- Quadrula_quadrula **
+--------------------------+
| +-- Uniomerus_tetralasmus **
|
+--------+ +-- Elliptio_complanata **
| | +-----------------------+
| | | +-- Pleurobema_oviforme **
| +--+
| | +----------------------- Amblema_plicata **
| | |
| +--+ +-------------------- Popenaias_popeii **
| | |
| +--+ +----------------- Toxolasma_parvum **
| | |
| | | +----- Potamilus_leptodon **
| +--+ +--------+
| | | | +-- Potamilus_alatus **
| | | +--+
| +--+ +-- Potamilus_streckersoni **
| |
| | +----------- Leaunio_lienosus **
| | |
| +--+ +-------- Venustaconcha_ellipsiformis **
| | |
| | | +-- Lampsilis_powellii **
| +--+-----+
| | +-- Lampsilis_siliquoidea **
| |
| | +----- Lampsilis_ornata **
| +--+
| | +-- Lampsilis_cardium **
| +--+
| +-- Lampsilis_cariosa **
|
--+ +----- Chamberlainia_somsakpanhai
| +--------------------+
| | | +-- Sinohyriopsis_cumingii
| | +--+
| | +-- Sinohyriopsis_schlegelii
| |
| | +-- Hyriopsis_bialata
| | +--+
| +--------+ | +-- Rectidens_sumatrensis
| | | +-----------------+
| | | | | +-- Lens_contradens
| | | | +--+
| | | | +-- Physunio_superbus
| | | |
| | | | +----------- Potomida_littoralis
| | +--+ +--------+
| | | | | +-------- Pronodularia_japanensis
| | | | +--+
| | | | | +----- Lamprotula_leaii
| | | | +--+
| | | | | +-- Lamprotula_caveata
| | +--+ +--+
| | | +-- Lamprotula_cornuumlunae
| | |
| | | +-- Pseudodon_vondembuschianus
| | | +--------------+
| | | | +-- Pilsbryoconcha_exilis
| | +--+
| | | +-------------- Inversidens_rentianensis
| | | |
+--+ +--+ +-- Parvasolenaia_rivularis
| | +--------+
| | | +-- Ptychorhynchus_pfisteri
| +--+
| | +-------- Microcondylaea_bonellii
| +--+
| | +----- Sinosolenaia_carinata
| +--+
| | +-- Sinosolenaia_iridinea
| +--+
| +-- Sinosolenaia_oleivora
|
| +-------------------------------- Lamellidens_marginalis
| |
| | +-- Lepidodesma_aligera
| | +--------------------------+
| | | +-- Lepidodesma_languilati
| | |
| | | +-------- Aculamprotula_tortuosa
| | | |
| | | +--------------+ +-- Aculamprotula_coreana
| | | | | +--+
+--+ | | | | +-- Aculamprotula_tientsinensis
| | | +--+
| | | | +-- Aculamprotula_polysticta
| | | +--+
| | | +-- Aculamprotula_scripta
| | |
| | +--+ +-------------- Unio_tumidus
| | | | +-----+
| | | | | | +----------- Unio_crassus
| | | | | +--+
+--+ | | | | +-------- Unio_elongatulus
| | | | +--+
| | | | | +----- Unio_pictorum
| | | | +--+
| | | | | +-- Unio_delphinus
| | +--+ +--+
| | | +-- Unio_mancus
| | |
| | | +-- Nodularia_breviconcha
| | | +--------------+
| | | | +-- Nodularia_douglasiae
| | | |
| | | | +-- Pseudocuneopsis_capitata
| | | +--------------+
| | +--+ +-- Pseudocuneopsis_sichuanensis
| | |
| | | +-- Schistodesmus_lampreyanus
+--+ | +-----------+
| | | +-- Schistodesmus_spinosus
| +--+
| | +----------- Tchangsinaia_piscicula
| +--+
| | +-------- Cuneopsis_rufescens
| +--+
| | +----- Cuneopsis_celtiformis
| +--+
| | +-- Cuneopsis_demangei
| +--+
| +-- Cuneopsis_heudei
|
| +-------- Acuticosta_chinensis
| +--------------+
| | | +----- Lanceolaria_gladiola
| | +--+
| | | +-- Lanceolaria_grayii
| | +--+
| | +-- Lanceolaria_lanceolata
| |
| | +-------------- Cristaria_plicata
+--+ |
| +-----+ +-- Anemina_arcaeformis
| | | +--------+
| | | | +-- Anemina_euscaphys
| | +--+
| | | +-------- Beringiana_fukuharai
| | +--+
| | | +----- Sinanodonta_lucida
| | +--+
+--+ | +-- Sinanodonta_tumens
| +--+
| +-- Sinanodonta_woodiana
|
| +----------------- Anodonta_nuttalliana
| |
| | +-------- Pseudanodonta_complanata
| | +-----+
+--+ | | +----- Anodonta_anatina
| | +--+
| | | +-- Anodonta_cygnea
| | +--+
+--+ +-- Anodonta_exulcerata
|
| +----- Pyganodon_grandis
| +-----+
| | | +-- Utterbackia_imbecillis
| | +--+
+--+ +-- Utterbackia_peninsularis
|
| +-------- Lasmigona_complanata
+--+
| +----- Prolasmidonta_heterodon
+--+
| +-- Alasmidonta_varicosa
+--+
+-- Platynaias_compressa
Roe et al., 2001 [Publ 2689], fig. 3a
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 16 |
16 |
8 |
1 |
0 |
0 |
0 |
25.0 |
+-------------- Ligumia_recta **
|
+-------------- Obliquaria_reflexa **
|
| +----- Lampsilis_teres **
+--------+
--+ | +-- Lampsilis_ovata **
| +--+
| +-- Lampsilis_ornata **
|
| +----------- Villosa_villosa **
| |
| | +----- Hamiota_australis[2] **
+--+ +--+
| | | +-- Hamiota_subangulata **
| | +--+
| | +-- Hamiota_australis[1] **
+--+
| +----- Hamiota_altilis[3] **
| |
| | +-- Hamiota_perovalis[1] **
+--+ |
| +-- Hamiota_perovalis[2] **
| |
| +-- Hamiota_altilis[1] **
+--+
+-- Hamiota_altilis[2] **
|
+-- Hamiota_altilis[4] **
|
+-- Hamiota_altilis[5] **
Szumowski et al., 2012 [Publ 3569], fig. 4
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 16 |
16 |
12 |
2 |
0 |
0 |
0 |
23.0 |
+-------------------------- Amblema_plicata **
|
--+ +----------------------- Ellipsaria_lineolata **
| |
+--+ +-------------------- Truncilla_truncata **
| |
+--+ +----------------- Obliquaria_reflexa **
| |
| | +-------------- Potamilus_alatus **
+--+ |
| | +----- Lampsilis_ornata **
| | +-----+
| | | | +-- Lampsilis_cardium **
+--+ | +--+
| | +-- Lampsilis_ovata **
| |
| | +-------- Lampsilis_teres **
| +--+
+--+ | +----- Lampsilis_siliquoidea **
| +--+
| | +-- Ortmanniana_ligamentina **
| +--+
| +-- Lampsilis_higginsii **
|
| +----- Hamiota_altilis **
| |
+-----+ +-- Epioblasma_triquetra **
| |
+--+-- Cambarunio_iris **
|
+-- Ligumia_recta **
Johnson et al., 2018 [Publ 3844], fig. 2
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 16 |
16 |
14 |
3 |
0 |
0 |
0 |
28.0 |
+-------------------------------- Amblema_plicata **
|
--+ +----------------------------- Elliptio_crassidens **
| |
+--+ +-------------------------- Megalonaias_nervosa **
| |
+--+ +----------------------- Uniomerus_tetralasmus **
| |
+--+ +-------------------- Theliderma_metanevra **
| |
| | +-- Quadrula_quadrula **
+--+ +--------------+
| | +-- Quadrula_verrucosa **
| |
| | +----- Pustulosa_kieneriana **
+--+ +--------+
| | | +-- Pustulosa_infucata **
| | +--+
| | +-- Pustulosa_kleiniana **
+--+
| +-- Pustulosa_pustulosa **
| +--------+
| | +-- Pustulosa_succissa **
+--+
| +-------- Cyclonaias_tuberculata **
+--+
| +----- Pustulosa_necki **
+--+
| +-- Pustulosa_nodulata **
+--+
+-- Pustulosa_petrina **
Inoue et al., 2020 [Publ 3941], fig. 5
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 16 |
16 |
14 |
1 |
0 |
0 |
0 |
25.0 |
+-------------------- Truncilla_macrodon **
|
| +-------------- Lampsilis_bracteata **
| |
| +--+ +-- Lampsilis_ornata **
--+ | | +--------+
| | | | +-- Lampsilis_fasciola **
| | +--+
| | | +-------- Lampsilis_ovata **
| | +--+
| | | +----- Lampsilis_cariosa **
+--+ +--+
| | +-- Lampsilis_cardium **
| +--+
| +-- Lampsilis_satura **
|
| +-------------- Lampsilis_higginsii **
| |
+--+ +----------- Lampsilis_teres **
| |
| | +-- Lampsilis_radiata **
+--+ +-----+
| | +-- Lampsilis_siliquoidea **
| |
+--+ +-- Lampsilis_hydiana **
| +--+
| | +-- Lampsilis_bergmanni **
+--+
| +-- Lampsilis_straminea **
+--+
+-- Lampsilis_virescens **
Porto-Hannes et al., 2021 [Publ 4227], fig. 3
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 16 |
16 |
12 |
4 |
0 |
0 |
0 |
23.0 |
+-------------------- Quadrula_quadrula **
|
+-------------------- Quadrula_verrucosa **
|
--+ +-- Eurynia_dilatata **
| +--------------+
| | +-- Pleurobema_sintoxia **
| |
| | +-- Amblema_plicata **
+--+ +-----------+
| | +-- Amblema_elliottii **
| |
| | +-- Obovaria_olivaria **
+--+ +--------+
| | +-- Ortmanniana_ligamentina **
| |
| | +-- Obliquaria_reflexa **
| +--------+
+--+ +-- Lampsilis_ornata **
|
| +----- Lampsilis_radiata[2] **
| +--+
| | | +-- Lampsilis_siliquoidea **
| | +--+
+--+ +-- Lampsilis_radiata[1] **
|
| +----- Lampsilis_virescens **
+--+
| +-- Lampsilis_hydiana **
+--+
+-- Lampsilis_straminea **
Lydeard et al., 2000 [Publ 2654], fig. 2a
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 15 |
17 |
11 |
4 |
2 |
1 |
1 |
31.0 |
+-------------------- Anodonta_cygnea
|
+-------------------- Cumberlandia_monodonta
|
| +----- Lampsilis_teres **
+--------------+
| | +-- Obovaria_unicolor **
| +--+
| +-- Obovaria_olivaria **
|
--+ +-- Reginaia_rotulata[1] **
| |
+-----------------+-- Reginaia_rotulata[2] **
| |
| +-- Reginaia_ebenus **
|
| +----- Quadrula_nobilis **
| +-----------+
| | | +-- Pustulosa_infucata **
| | +--+
+--+ +-- Pustulosa_succissa **
|
| +-------------- Pleurobema_decisum **
+--+
| +----------- Fusconaia_flava **
+--+
| +-------- Fusconaia_burkei[1] **
+--+
| +----- Fusconaia_burkei[2] **
+--+
| +-- Fusconaia_burkei[3] **
+--+
+-- Fusconaia_escambia **
Lydeard et al., 2000 [Publ 2654], fig. 2b
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 15 |
15 |
11 |
4 |
2 |
1 |
1 |
30.0 |
+----- Lampsilis_teres **
+-----------------+
| | +-- Obovaria_unicolor **
| +--+
| +-- Obovaria_olivaria **
|
--+ +-- Reginaia_rotulata[1] **
| |
| +-----------------+-- Reginaia_rotulata[2] **
| | |
| | +-- Reginaia_ebenus **
| |
+--+ +-- Quadrula_nobilis **
| |
| +--------------+-- Pustulosa_infucata **
| | |
+--+ +-- Pustulosa_succissa **
|
| +-------------- Pleurobema_decisum **
+--+
| +----------- Fusconaia_flava **
+--+
| +-------- Fusconaia_burkei[1] **
+--+
| +----- Fusconaia_burkei[2] **
+--+
| +-- Fusconaia_burkei[3] **
+--+
+-- Fusconaia_escambia **
Inoue et al., 2013 [Publ 3528], fig. 3
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 15 |
15 |
12 |
4 |
2 |
2 |
2 |
26.0 |
+-------------------------- Gonidea_angulata
|
+-------------------------- Pyganodon_grandis
|
--+ +-- Eurynia_dilatata **
| +--------------------+
| | +-- Quadrula_quadrula **
| |
| | +-------- Paetulunio_fabalis **
+--+ +-----------+
| | | +----- Amblema_plicata **
| | +--+
| | | +-- Reginaia_ebenus **
+--+ +--+
| +-- Reginaia_rotulata **
|
| +----------------- Lampsilis_ornata **
| |
+--+ +-------------- Leaunio_vanuxemensis **
| |
| | +-- Villosa_villosa **
+--+-----------+
| +-- Cambarunio_iris **
|
| +----------- Obovaria_olivaria **
+--+
| +-------- Obovaria_arkansasensis **
+--+
| +----- Obovaria_retusa **
+--+
| +-- Obovaria_unicolor **
+--+
+-- Obovaria_subrotunda **
Szumowski et al., 2012 [Publ 3569], fig. 5
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 15 |
15 |
12 |
1 |
0 |
0 |
0 |
30.0 |
+----------------- Megalonaias_nervosa **
|
| +-------- Theliderma_cylindrica **
+--------+
| | +----- Theliderma_intermedia **
| +--+
| | +-- Theliderma_metanevra **
--+ +--+
| +-- Theliderma_sparsa **
|
| +-------- Quadrula_quadrula[1] **
| +-----+
| | | +----- Quadrula_verrucosa **
| | +--+
| | | +-- Quadrula_nobilis **
+--+ +--+
| +-- Quadrula_quadrula[2] **
|
| +----------- Pustulosa_kieneriana **
| |
+--+ +-------- Pustulosa_infucata **
| |
+--+ +-- Pustulosa_pustulosa **
| +--+
| | +-- Pustulosa_succissa **
+--+
| +-- Pustulosa_nodulata **
+--+
+-- Pustulosa_petrina **
Pfeiffer et al., 2019 [Publ 3937], fig. 4-5
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 15 |
15 |
13 |
5 |
42 |
35 |
3 |
17.0 |
+-- Quadrula_quadrula **
+--------------------+
| +-- Uniomerus_tetralasmus **
|
| +----- Elliptoideus_sloatianus **
+-----+ +--------------+
| | | | +-- Elliptio_complanata **
| | | +--+
| | | +-- Pleurobema_strodeanum **
| | |
| +--+ +----- Reginaia_ebenus **
| | +-----------+
| | | | +-- Plectomerus_dombeyanus **
| | | +--+
| | | +-- Amblema_plicata **
| +--+
| | +-- Psoronaias_semigranosa **
| | +-----------+
| | | +-- Popenaias_popeii **
| +--+
| | +----------- Obliquaria_reflexa **
| +--+
| | +-------- Cyrtonaias_tampicoensis **
| +--+
| | +----- Leaunio_lienosus **
| +--+
| | +-- Lampsilis_cardium **
| +--+
| +-- Pachynaias_spheniopsis **
|
--+ +-- Chamberlainia_hainesiana
| +--------------------+
| | +-- Sinohyriopsis_cumingii
| |
| | +----------------- Lamprotula_cornuumlunae
| | +--+
| | | | +-------------- Gonidea_angulata
| +--+ | +--+
| | | | | +----------- Indopseudodon_crebristriatus
| | | | +--+
| | | | | +-------- Bineurus_anodontinum
| | | | +--+
| | | | | +----- Pilsbryoconcha_exilis
| | | | +--+
| | +--+ | +-- Pseudodon_vondembuschianus
| | | +--+
| | | +-- Pseudodon_inoscularis
| | |
| | | +----- Ensidens_ingallsianus
| | | +-----+
| | | | | +-- Hyriopsis_bialata
| | | | +--+
| | | | +-- Rectidens_sumatrensis
| | +--------+
| | | +-- Trapezoideus_foliaceus
| | | +-----+
+--+ | | +-- Physunio_superbus
| +--+
| | +----- Yaukthwa_paiensis
| +--+
| | +-- Lens_contradens
| +--+
| +-- Solenaia_emarginata
|
| +----------- Lanceolaria_lanceolata
| |
| | +----- Unio_gibbus
| +--------+ +--+
| | | | | +-- Nodularia_douglasiae
| | | | +--+
| | | | +-- Tchangsinaia_piscicula
| | +--+
| | | +-- Cristaria_plicata
| | | +--+
| | | | +-- Sinanodonta_woodiana
+-----+ +--+
| | +-- Anodonta_cygnea
| +--+
| +-- Pyganodon_grandis
|
| +-- Trapezidens_dolichorhynchus
| +--------------+
| | +-- Lamellidens_savadiensis
| |
+--+ +-- Leoparreysia_tavoyensis
| +-----------+
| | +-- Leoparreysia_olivacea
| |
+--+ +-- Coelatura_choziensis
| +--------+
| | +-- Prisodontopsis_aviculaeformis
+--+
| +-------- Indonaia_caerulea
+--+
| +----- Indochinella_pugio
+--+
| +-- Scabies_phaselus
+--+
+-- Harmandia_somboriensis
Pfeiffer et al., 2019 [Publ 3937], fig. 6
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 15 |
15 |
13 |
5 |
42 |
35 |
3 |
19.0 |
+-- Quadrula_quadrula **
+-----------------+
| +-- Uniomerus_tetralasmus **
|
| +----- Elliptoideus_sloatianus **
| +--+
| | | +-- Elliptio_complanata **
+--+ | +--+
| | +--------+ +-- Pleurobema_strodeanum **
| | | |
| | | | +----- Reginaia_ebenus **
| | | +--+
| | | | +-- Plectomerus_dombeyanus **
| +--+ +--+
| | +-- Amblema_plicata **
| |
| | +-- Psoronaias_semigranosa **
| | +-----------+
| | | +-- Popenaias_popeii **
| +--+
| | +----------- Obliquaria_reflexa **
| +--+
| | +-------- Cyrtonaias_tampicoensis **
| +--+
| | +----- Leaunio_lienosus **
| +--+
| | +-- Lampsilis_cardium **
| +--+
| +-- Pachynaias_spheniopsis **
|
| +-- Chamberlainia_hainesiana
| +-----------------+
| | +-- Sinohyriopsis_cumingii
| |
| | +-- Gonidea_angulata
| | +-----------+
| | | +-- Lamprotula_cornuumlunae
| | +--+
+--+ | | +----------- Indopseudodon_crebristriatus
| | | +--+
| | | | +-------- Pilsbryoconcha_exilis
| | | +--+
--+ | | | +----- Bineurus_anodontinum
| | | +--+
| | | | +-- Pseudodon_vondembuschianus
| +--+ +--+
| | +-- Pseudodon_inoscularis
| |
| | +----- Ensidens_ingallsianus
| | +-----+
| | | | +-- Hyriopsis_bialata
| | | +--+
| | | +-- Rectidens_sumatrensis
| +-----+
| | +-- Trapezoideus_foliaceus
| | +-----+
| | | +-- Physunio_superbus
| +--+
| | +----- Yaukthwa_paiensis
| +--+
| | +-- Lens_contradens
| +--+
| +-- Solenaia_emarginata
|
| +-------- Lanceolaria_lanceolata
| +--+
| | | +----- Unio_gibbus
| | +--+
| | | +-- Tchangsinaia_piscicula
| | +--+
| +--------+ +-- Nodularia_douglasiae
| | |
| | | +-- Cristaria_plicata
| | | +--+
| | | | +-- Sinanodonta_woodiana
| | +-----+
| | | +-- Anodonta_cygnea
+--+ +--+
| +-- Pyganodon_grandis
|
| +-- Trapezidens_dolichorhynchus
| +--------------+
| | +-- Lamellidens_savadiensis
| |
+--+ +-- Leoparreysia_tavoyensis
| +-----------+
| | +-- Leoparreysia_olivacea
| |
+--+ +-- Coelatura_choziensis
| +--------+
| | +-- Prisodontopsis_aviculaeformis
+--+
| +-------- Indonaia_caerulea
+--+
| +----- Indochinella_pugio
+--+
| +-- Scabies_phaselus
+--+
+-- Harmandia_somboriensis
Wu et al., 2022 [Publ 4238], fig. 3
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 15 |
15 |
13 |
5 |
59 |
50 |
2 |
2.0 |
+-- Uniomerus_tetralasmus **
+--------------------------+
| +-- Quadrula_quadrula **
|
+--------+ +-- Pleurobema_oviforme **
| | +-----------------------+
| | | +-- Elliptio_complanata **
| +--+
| | +----------------------- Amblema_plicata **
| | |
| +--+ +-------------------- Popenaias_popeii **
| | |
| +--+ +----------------- Toxolasma_parvum **
| | |
| +--+ +-- Potamilus_alatus **
| | +-----------+
| | | +-- Potamilus_leptodon **
| +--+
| | +----------- Leaunio_lienosus **
| | |
| +--+ +-- Lampsilis_cardium **
| | +-----+
| | | +-- Lampsilis_ornata **
| +--+
| | +----- Venustaconcha_ellipsiformis **
| +--+
| | +-- Lampsilis_siliquoidea **
| +--+
--+ +-- Lampsilis_powellii **
|
| +----- Chamberlainia_somsakpanhai
| +-----------+
| | | +-- Sinohyriopsis_schlegelii
| | +--+
| | +-- Sinohyriopsis_cumingii
| |
| +-----------------+ +-------- Potomida_littoralis
| | | +-----+
| | | | | +----- Pronodularia_japanensis
| | | | +--+
| | | | | +-- Lamprotula_caveata
| | | | +--+
| | +--+ +-- Lamprotula_leaii
| | |
| | | +-- Pilsbryoconcha_exilis
| | | +--------+
| | | | +-- Pseudodon_vondembuschianus
| | | |
| | +--+ +-- Ptychorhynchus_pfisteri
| | | +-----+
| | | | +-- Parvasolenaia_rivularis
| | +--+
| | | +----- Microcondylaea_bonellii
+--+ +--+
| | +-- Sinosolenaia_carinata
| +--+
| +-- Sinosolenaia_recognita
|
| +-------------------------------- Lepidodesma_languilati
| |
| | +-------- Acuticosta_chinensis
| | +--------+
| | | | +----- Lanceolaria_gladiola
| | | +--+
| | | | +-- Lanceolaria_grayii
| | | +--+
| | | +-- Lanceolaria_lanceolata
| | |
| | +-----------+ +-------- Cristaria_plicata
| | | | |
+--+ | | +-----+ +-- Sinanodonta_woodiana
| | | | | +--+
| | | | | | +-- Sinanodonta_lucida
| | | | +--+
| | | | | +-- Anemina_arcaeformis
| | +--+ +--+
| | | +-- Anemina_euscaphys
| | |
| | | +-- Anodonta_cygnea
| | | +--------+
| | | | +-- Anodonta_anatina
| | +--+
| | | +-------- Platynaias_compressa
+--+ +--+
| | +----- Pyganodon_grandis
| +--+
| | +-- Utterbackia_imbecillis
| +--+
| +-- Utterbackia_peninsularis
|
| +-------- Aculamprotula_tortuosa
| |
| +-----------------+ +-- Aculamprotula_tientsinensis
| | | +--+
| | | | +-- Aculamprotula_coreana
| | +--+
| | | +-- Aculamprotula_polysticta
| | +--+
| | +-- Aculamprotula_scripta
+--+
| +-------- Unio_tumidus
| +--------------+
| | | +----- Unio_crassus
| | +--+
| | | +-- Unio_pictorum
| | +--+
+--+ +-- Unio_delphinus
|
| +-- Pseudocuneopsis_capitata
| +-----------------+
| | +-- Pseudocuneopsis_sichuanensis
| |
+--+ +-- Nodularia_douglasiae
| +--------------+
| | +-- Nodularia_breviconcha
| |
+--+ +-- Schistodesmus_lampreyanus
| +-----------+
| | +-- Schistodesmus_spinosus
+--+
| +----------- Tchangsinaia_piscicula
+--+
| +-------- Cuneopsis_rufescens
+--+
| +----- Cuneopsis_celtiformis
+--+
| +-- Cuneopsis_heudei
+--+
+-- Cuneopsis_demangei
Mejia-Trujillo & Smith, 2022 [Publ 4274], fig. 2-F
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 15 |
15 |
13 |
5 |
0 |
0 |
0 |
10.0 |
+-------------- Toxolasma_parvum **
|
+--+ +----------- Popenaias_popeii **
| | |
| +--+ +-------- Amblema_plicata **
| | |
| +--+ +-- Quadrula_quadrula **
| | +--+
| | | +-- Uniomerus_tetralasmus **
| +--+
--+ | +-- Pleurobema_oviforme **
| +--+
| +-- Elliptio_complanata **
|
| +----- Potamilus_leptodon **
| +-----+
| | | +-- Potamilus_streckersoni **
| | +--+
| | +-- Potamilus_alatus **
+-----+
| +-- Lampsilis_cardium **
| +-----+
| | +-- Lampsilis_ornata **
+--+
| +----- Venustaconcha_ellipsiformis **
+--+
| +-- Lampsilis_siliquoidea **
+--+
+-- Lampsilis_powellii **
Dai et al., 2023 [Publ 4285], fig. 5
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 15 |
15 |
13 |
5 |
38 |
36 |
2 |
2.0 |
+-- Quadrula_quadrula **
+--------------------------+
| +-- Uniomerus_tetralasmus **
|
+--+ +-- Elliptio_complanata **
| | +-----------------------+
| | | +-- Pleurobema_oviforme **
| +--+
| | +----------------------- Amblema_plicata **
| | |
| +--+ +-------------------- Popenaias_popeii **
| | |
| +--+ +----------------- Toxolasma_parvum **
| | |
| +--+ +-- Potamilus_leptodon **
| | +-----------+
| | | +-- Potamilus_alatus **
| +--+
| | +----------- Leaunio_lienosus **
| | |
| +--+ +-- Lampsilis_cardium **
| | +-----+
| | | +-- Lampsilis_ornata **
| +--+
| | +----- Venustaconcha_ellipsiformis **
| +--+
| | +-- Lampsilis_powellii **
| +--+
--+ +-- Lampsilis_siliquoidea **
|
| +-------------------- Lepidodesma_languilati
| |
| | +-------------- Aculamprotula_tientsinensis
| | |
| | +--+ +----------- Unio_crassus
| | | | |
| +--+ | +--+ +-- Pseudocuneopsis_capitata
| | | | | +-----+
| | | | | | +-- Nodularia_douglasiae
| | | | +--+
| | | | | +----- Schistodesmus_lampreyanus
| | | | +--+
| | +--+ | +-- Cuneopsis_celtiformis
| | | +--+
| | | +-- Tchangsinaia_piscicula
| | |
| | | +-- Acuticosta_chinensis
| | | +-----------+
| | | | +-- Lanceolaria_grayii
| | | |
| | +--+ +----- Cristaria_plicata
| | | +-----+
| | | | | +-- Anemina_arcaeformis
+--------+ | | +--+
| +--+ +-- Sinanodonta_woodiana
| |
| | +-------- Anodonta_cygnea
| +--+
| | +----- Platynaias_compressa
| +--+
| | +-- Pyganodon_grandis
| +--+
| +-- Utterbackia_imbecillis
|
| +----- Chamberlainia_somsakpanhai
| +--------------+
| | | +-- Sinohyriopsis_cumingii
| | +--+
| | +-- Sinohyriopsis_schlegelii
| |
| | +-- Hyriopsis_bialata
+--+ +--+
| | +-- Rectidens_sumatrensis
| +-----------+
| | | +-- Lens_contradens
| | +--+
| | +-- Physunio_superbus
| |
+--+ +-------- Potomida_littoralis
| +-----+
| | | +----- Pronodularia_japanensis
| | +--+
| | | +-- Lamprotula_caveata
| | +--+
+--+ +-- Lamprotula_leaii
|
| +-- Pseudodon_vondembuschianus
| +--------+
| | +-- Pilsbryoconcha_exilis
| |
+--+ +----- Parvasolenaia_rivularis
| +--+
| | | +-- Postolata_guangxiensis
| | +--+
+--+ +-- Ptychorhynchus_pfisteri
|
| +----- Microcondylaea_bonellii
+--+
| +-- Sinosolenaia_oleivora
+--+
+-- Sinosolenaia_carinata
Graf & Cummings, 2006 [Publ 1968], fig. 2
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 14 |
14 |
10 |
4 |
33 |
11 |
3 |
26.0 |
+----------------------------- Coelatura_aegyptiaca
|
+----------------------------- Lens_contradens
|
| +-- Unio_pictorum
| +--------+
| | +-- Unio_caffer
+-----------------+
| | +-------- Lasmigona_costata
--+ +--+
| | +----- Alasmidonta_marginata
| +--+
| | +-- Pyganodon_grandis
| +--+
| +-- Strophitus_undulatus
|
| +-------------------------- Gonidea_angulata
| |
| | +-- Quadrula_verrucosa **
+--+ +--------------------+
| | +-- Quadrula_quadrula **
| |
+--+ +-------------------- Amblema_plicata **
| |
| | +----- Eurynia_dilatata **
+--+ +-----------+
| | | +-- Fusconaia_flava **
| | +--+
| | +-- Pleurobema_sintoxia **
+--+
| +-------------- Obliquaria_reflexa **
| |
| +-------------- Truncilla_truncata **
+--+
+-------------- Ptychobranchus_fasciolaris **
|
| +----------- Ligumia_recta **
+--+
| +-------- Cambarunio_iris **
+--+
| +----- Epioblasma_triquetra **
+--+
| +-- Lampsilis_cardium **
+--+
+-- Ortmanniana_ligamentina **
Lydeard et al., 2000 [Publ 2654], fig. 2c
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 14 |
16 |
9 |
4 |
2 |
1 |
1 |
30.0 |
+-------------- Anodonta_cygnea
|
+-------------- Cumberlandia_monodonta
|
| +----- Quadrula_nobilis **
+--------+
| | +-- Pustulosa_infucata **
| +--+
| +-- Pustulosa_succissa **
--+
| +----- Obovaria_unicolor **
+--------+
| | +-- Obovaria_olivaria **
| +--+
| +-- Lampsilis_teres **
|
| +-- Reginaia_ebenus **
| +--------+
| | +-- Reginaia_rotulata **
+--+
| +-------- Pleurobema_decisum **
| |
+--+ +----- Fusconaia_flava **
| |
+--+ +-- Fusconaia_burkei[1] **
| |
| +-- Fusconaia_burkei[2] **
+--+
+-- Fusconaia_burkei[3] **
|
+-- Fusconaia_escambia **
Lydeard et al., 2000 [Publ 2654], fig. 2d
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 14 |
14 |
11 |
4 |
2 |
1 |
1 |
29.0 |
+-- Reginaia_ebenus **
+--------------------+
| +-- Reginaia_rotulata **
|
| +----- Lampsilis_teres **
--+ +--------------+
| | | +-- Obovaria_unicolor **
| | +--+
| | +-- Obovaria_olivaria **
+--+
| +-- Quadrula_nobilis **
| |
| +--------------+-- Pustulosa_infucata **
| | |
+--+ +-- Pustulosa_succissa **
|
| +-------------- Pleurobema_decisum **
+--+
| +----------- Fusconaia_flava **
+--+
| +-------- Fusconaia_burkei[1] **
+--+
| +----- Fusconaia_burkei[2] **
+--+
| +-- Fusconaia_burkei[3] **
+--+
+-- Fusconaia_escambia **
Huang et al., 2002 [Publ 2657], fig. 1
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 14 |
14 |
12 |
4 |
20 |
18 |
2 |
27.0 |
+----------------------------------- Sinohyriopsis_cumingii
|
--+ +-------------------------------- Ptychorhynchus_pfisteri
| |
+--+ +----------------------------- Megalonaias_nervosa **
| |
| | +----- Elliptoideus_sloatianus **
+--+ +--------------------+
| | | +-- Uniomerus_carolinianus **
| | +--+
+--+ +-- Pleurobema_pyriforme **
|
| +----------------------- Amblema_plicata **
| |
+--+ +-- Obliquaria_reflexa **
| +-----------------+
| | +-- Plectomerus_dombeyanus **
+--+
| +----------------- Medionidus_conradicus **
+--+
| +-------------- Potamilus_fragilis **
+--+
| +----------- Ptychobranchus_subtentus **
+--+
| +-------- Potamilus_purpuratus **
+--+
| +----- Lampsilis_teres **
+--+
| +-- Villosa_delumbis **
+--+
+-- Ellipsaria_lineolata **
Huang et al., 2002 [Publ 2657], fig. 2
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 14 |
14 |
6 |
4 |
20 |
18 |
2 |
27.0 |
+-------- Strophitus_radiatus
|
+-------- Schistodesmus_lampreyanus
|
+-------- Lepidodesma_languilati
|
+-------- Acuticosta_ovata
|
| +-- Sinanodonta_woodiana
+-----+
| +-- Cristaria_plicata
+--------+
| | +-- Lanceolaria_grayii
| +-----+
| | +-- Lanceolaria_lanceolata
| |
| | +-- Nodularia_douglasiae
| +-----+
| | +-- Tchangsinaia_piscicula
| |
| | +----- Utterbackia_imbecillis
| | |
--+ +--+----- Utterbackiana_couperiana
| |
| | +-- Alasmidonta_triangulata
| +--+
| +-- Lasmigona_costata
|
| +-------------- Ptychorhynchus_pfisteri
| |
| +-------------- Sinohyriopsis_cumingii
| |
| +-------------- Lamprotula_leaii
| |
+--+-------------- Sinosolenaia_oleivora
|
| +----------- Megalonaias_nervosa **
| |
| | +----- Elliptoideus_sloatianus **
| | +--+
+--+ | | +-- Uniomerus_carolinianus **
| | +--+
| | +-- Pleurobema_pyriforme **
| |
| | +----- Potamilus_fragilis **
+--+ |
| +----- Medionidus_conradicus **
| |
| +----- Amblema_plicata **
| |
| | +-- Obliquaria_reflexa **
+--+--+
| +-- Plectomerus_dombeyanus **
|
| +-- Villosa_delumbis **
| |
| +-- Ellipsaria_lineolata **
| |
+--+-- Lampsilis_teres **
|
+-- Potamilus_purpuratus **
|
+-- Ptychobranchus_subtentus **
Hoeh et al., 2009 [Publ 2716], fig. 3
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 14 |
14 |
12 |
4 |
33 |
11 |
3 |
28.0 |
+-------------------------- Lens_contradens
|
| +----- Gonidea_angulata
| +-----------------+
| | | +-- Pilsbryoconcha_exilis
--+ | +--+
| | +-- Pseudodon_vondembuschianus
| |
| | +----------- Amblema_plicata **
| | |
+--+ | +----- Obliquaria_reflexa **
| +--------+ +--+
| | | | | +-- Quadrula_verrucosa **
| | | | +--+
| | +--+ +-- Quadrula_quadrula **
| | |
| | | +----- Eurynia_dilatata **
+--+ +--+
| | +-- Pleurobema_sintoxia **
| +--+
| +-- Fusconaia_flava **
|
| +----------------- Ptychobranchus_fasciolaris **
+--+
| +-------------- Truncilla_truncata **
+--+
| +----------- Ortmanniana_ligamentina **
+--+
| +-------- Lampsilis_cardium **
+--+
| +----- Cambarunio_iris **
+--+
| +-- Ligumia_recta **
+--+
+-- Epioblasma_triquetra **
Turner et al., 2000 [Publ 2722], fig. 2
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 14 |
15 |
11 |
4 |
2 |
1 |
1 |
22.0 |
+-- Quadrula_quadrula **
+-----------------------+
| +-- Utterbackiana_couperiana
|
--+ +-- Amblema_plicata **
| +--------------------+
| | +-- Eurynia_dilatata **
+--+
| +-------------------- Medionidus_conradicus **
| |
+--+ +----------------- Lampsilis_teres **
| |
| +----------------- Villosa_delumbis **
+--+
| +-- Lampsilis_satura **
| +-----------+
| | +-- Lampsilis_cardium **
+--+
| +----------- Lampsilis_reeveiana **
| |
+--+ +-------- Lampsilis_hydiana[1] **
| |
+--+-------- Lampsilis_hydiana[2] **
|
| +----- Lampsilis_siliquoidea[2] **
+--+
| +-- Lampsilis_siliquoidea[1] **
+--+
+-- Lampsilis_powellii **
Kandl et al., 2001 [Publ 3384], fig. 5
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 14 |
14 |
7 |
3 |
0 |
0 |
0 |
21.0 |
+-------------------- Amblema_plicata **
|
+-------------------- Quadrula_quadrula **
--+
| +----------------- Elliptio_crassidens **
| |
+--+ +-------------- Fusconaia_flava **
| |
+--+ +----------- Pleurobema_sintoxia **
| |
| | +-- Pleurobema_clava **
+--+ +-----+
| | +-- Pleurobema_strodeanum **
| |
+--+ +----- Pleurobema_pyriforme[6] **
| |
| | +-- Pleurobema_pyriforme[1] **
+--+ |
| +-- Pleurobema **
| |
| +-- Pleurobema_pyriforme[2] **
+--+
+-- Pleurobema_pyriforme[3] **
|
+-- Pleurobema_pyriforme[4] **
|
+-- Pleurobema_pyriforme[5] **
Hewitt et al., 2019 [Publ 3962], fig. 2
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 14 |
14 |
9 |
2 |
2 |
1 |
1 |
19.0 |
+-------------------------- Pyganodon_grandis
|
--+ +----------------------- Fusconaia_flava **
| |
+--+ +-------------------- Lampsilis_siliquoidea **
| |
| | +----------------- Lampsilis_fasciola **
+--+ |
| | +-- Epioblasma_rangiana **
| +--------------+
+--+ +-- Lampsilis_radiata **
|
| +-------------- Lampsilis_ornata **
| |
+--+ +-- Lampsilis_cardium[1] **
| +--------+
| | +-- Lampsilis_ovata[1] **
+--+
| +-------- Lampsilis_cardium[6] **
| |
+--+ +----- Lampsilis_cardium[5] **
| |
+--+ +-- Lampsilis_cardium[2] **
| |
| +-- Lampsilis_ovata[2] **
+--+
+-- Lampsilis_cardium[3] **
|
+-- Lampsilis_cardium[4] **
Teiga-Teixeira et al., 2020 [Publ 4173], fig. 1
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 14 |
14 |
12 |
5 |
5 |
3 |
2 |
3.0 |
+----- Lanceolaria_grayii
+-----------------------+
| | +-- Chamberlainia_somsakpanhai
| +--+
| +-- Lamprotula_leaii
--+
| +-- Uniomerus_tetralasmus **
| +-----------------------+
| | +-- Quadrula_quadrula **
| |
+--+ +-- Elliptio_complanata **
| +--------------------+
| | +-- Pleurobema_oviforme **
+--+
| +-------------------- Amblema_plicata **
| |
+--+ +----------------- Popenaias_popeii **
| |
+--+ +-------------- Toxolasma_parvum **
| |
| | +-- Potamilus_alatus **
+--+ +--------+
| | +-- Potamilus_leptodon **
| |
+--+ +-- Lampsilis_cardium **
| +-----+
| | +-- Lampsilis_ornata **
+--+
| +----- Venustaconcha_ellipsiformis **
+--+
| +-- Lampsilis_powellii **
+--+
+-- Lampsilis_siliquoidea **
Serb & Barnhart, 2008 [Publ 2674], fig. 3
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 13 |
13 |
10 |
1 |
0 |
0 |
0 |
20.0 |
+-- Lampsilis_siliquoidea **
+-----------------------+
| +-- Lampsilis_ornata **
--+
| +----------------------- Obliquaria_reflexa **
| |
+--+ +-------------------- Ptychobranchus_fasciolaris **
| |
+--+ +-- Cyprogenia_aberti[6] **
| +--------------+
| | +-- Dromus_dromas **
+--+
| +-------------- Cyprogenia_aberti[4] **
| |
+--+ +----------- Cyprogenia_aberti[3] **
| |
+--+ +-------- Cyprogenia_aberti[5] **
| |
+--+ +----- Cyprogenia_stegaria[2] **
| |
+--+----- Cyprogenia_aberti[2] **
|
| +-- Cyprogenia_stegaria[1] **
+--+
+-- Cyprogenia_aberti[1] **
Serb & Barnhart, 2008 [Publ 2674], fig. 4
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 13 |
13 |
9 |
1 |
0 |
0 |
0 |
20.0 |
+----------------------- Obliquaria_reflexa **
|
| +-- Lampsilis_ornata **
+--------------------+
--+ +-- Lampsilis_siliquoidea **
|
| +-------------------- Ptychobranchus_fasciolaris **
| |
+--+ +-- Cyprogenia_aberti[6] **
| +--------------+
| | +-- Dromus_dromas **
+--+
| +-------------- Cyprogenia_aberti[5] **
| |
+--+ +----------- Cyprogenia_aberti[4] **
| |
+--+ +-------- Cyprogenia_aberti[3] **
| |
+--+ +----- Cyprogenia_stegaria[2] **
| |
+--+----- Cyprogenia_aberti[2] **
|
| +-- Cyprogenia_stegaria[1] **
+--+
+-- Cyprogenia_aberti[1] **
Serb, 2006 [Publ 2678], fig. 2
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 13 |
13 |
7 |
1 |
0 |
0 |
0 |
20.0 |
+-------------- Obliquaria_reflexa **
|
| +----- Ptychobranchus_fasciolaris **
+--------+
| | +-- Lampsilis_ornata **
--+ +--+
| +-- Lampsilis_siliquoidea **
|
| +-- Cyprogenia_aberti[6] **
| +--------+
| | +-- Dromus_dromas **
+--+
| +-------- Cyprogenia_aberti[5] **
| |
+--+ +----- Cyprogenia_aberti[4] **
| |
| | +-- Cyprogenia_stegaria[1] **
+--+ |
| +-- Cyprogenia_stegaria[2] **
| |
+--+-- Cyprogenia_aberti[1] **
|
+-- Cyprogenia_aberti[2] **
|
+-- Cyprogenia_aberti[3] **
Hoeh et al., 2009 [Publ 2716], fig. 4
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 13 |
13 |
10 |
3 |
33 |
11 |
3 |
27.0 |
+-- Unio_pictorum
+--------+
| +-- Unio_caffer
+--------------+
| | +-------- Lasmigona_costata
| +--+
| | +----- Alasmidonta_marginata
| +--+
| | +-- Strophitus_undulatus
| +--+
--+ +-- Pyganodon_grandis
|
| +----------------------- Gonidea_angulata
| |
| | +-- Eurynia_dilatata **
| | |
| | +-----+-- Pleurobema_sintoxia **
+--+ | |
| +-----------+ +-- Fusconaia_flava **
| | |
| | | +----- Obliquaria_reflexa **
| | +--+
+--+ | +-- Quadrula_verrucosa **
| +--+
| +-- Quadrula_quadrula **
|
| +----------------- Ptychobranchus_fasciolaris **
+--+
| +-------------- Truncilla_truncata **
+--+
| +----------- Ortmanniana_ligamentina **
+--+
| +-------- Lampsilis_cardium **
+--+
| +----- Cambarunio_iris **
+--+
| +-- Ligumia_recta **
+--+
+-- Epioblasma_triquetra **
Hoeh et al., 2009 [Publ 2716], fig. 5
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 13 |
13 |
7 |
4 |
33 |
11 |
3 |
25.0 |
+-- Unio_pictorum
+--------+
| +-- Unio_caffer
+-----+
| | +-------- Lasmigona_costata
| +--+
| | +----- Alasmidonta_marginata
| +--+
| | +-- Strophitus_undulatus
--+ +--+
| +-- Pyganodon_grandis
|
| +-------------- Gonidea_angulata
| |
| | +----- Obliquaria_reflexa **
| | +-----+
+--+ | | +-- Quadrula_verrucosa **
| | +--+
| | +-- Quadrula_quadrula **
| |
+--+ +-------- Amblema_plicata **
| |
| | +----- Eurynia_dilatata **
| +--+
| | | +-- Pleurobema_sintoxia **
+--+ +--+
| +-- Fusconaia_flava **
|
| +----- Truncilla_truncata **
| |
| +----- Ptychobranchus_fasciolaris **
+--+
| +-- Ortmanniana_ligamentina **
| |
| +-- Cambarunio_iris **
+--+
+-- Ligumia_recta **
|
+-- Epioblasma_triquetra **
Kandl et al., 2001 [Publ 3384], fig. 4
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 13 |
13 |
4 |
2 |
0 |
0 |
0 |
22.5 |
+-------------- Elliptio **
|
+-------------- Uniomerus_carolinianus **
--+
| +----------- Pleurobema_sintoxia **
| |
+--+ +-------- Pleurobema_strodeanum **
| |
| | +----- Pleurobema_pyriforme[7] **
+--+ |
| | +-- Pleurobema_pyriforme[1] **
| | |
+--+ +-- Pleurobema_pyriforme[2] **
| |
| +-- Pleurobema_pyriforme[3] **
| |
| +-- Pleurobema_pyriforme[4] **
+--+
+-- Pleurobema_pyriforme[5] **
|
+-- Pleurobema[1] **
|
+-- Pleurobema_pyriforme[6] **
|
+-- Pleurobema[2] **
Jadhav & Jamkhedkar, 2009 [Publ 3697], fig. 11
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 13 |
14 |
12 |
4 |
13 |
13 |
3 |
26.0 |
+-------------------- Nodularia_douglasiae
|
+--------+ +----------------- Anodonta_cygnea
| | |
| +--+ +-------------- Unio_pictorum[2]
| | |
| +--+ +-- Unio_pictorum[1]
| | +--------+
| | | +-- Unio_caffer
| +--+
| | +-------- Anodonta
| | |
--+ +--+ +----- Lasmigona_costata
| | |
| +--+ +-- Strophitus_undulatus
| | |
| +--+-- Alasmidonta_marginata
| |
| +-- Pyganodon_grandis
|
| +-------------------------- Pronodularia_japanensis
| |
| | +-- Quadrula_quadrula **
+--+ +--------------------+
| | +-- Quadrula_verrucosa **
| |
+--+ +-------------------- Gonidea_angulata
| |
| | +----- Eurynia_dilatata **
+--+ +-----------+
| | | +-- Cambarunio_iris **
| | +--+
| | +-- Ptychobranchus_fasciolaris **
+--+
| +-- Amblema_plicata **
| +-----------+
| | +-- Pleurobema_sintoxia **
+--+
| +----------- Truncilla_truncata **
| |
+--+ +-- Lampsilis_cardium **
| +-----+
| | +-- Epioblasma_triquetra **
+--+
| +----- Ligumia_recta **
+--+
| +-- Obliquaria_reflexa **
+--+
+-- Ortmanniana_ligamentina **
Serb, 2006 [Publ 2678], fig. 3
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 12 |
12 |
8 |
1 |
0 |
0 |
0 |
20.0 |
+-------------------- Obliquaria_reflexa **
|
| +-- Lampsilis_ornata **
+-----------------+
--+ +-- Lampsilis_siliquoidea **
|
| +----------------- Ptychobranchus_fasciolaris **
| |
+--+ +-- Cyprogenia_aberti[5] **
| +-----------+
| | +-- Dromus_dromas **
+--+
| +----------- Cyprogenia_aberti[4] **
| |
+--+ +-------- Cyprogenia_aberti[3] **
| |
+--+ +----- Cyprogenia_stegaria[2] **
| |
+--+----- Cyprogenia_aberti[2] **
|
| +-- Cyprogenia_stegaria[1] **
+--+
+-- Cyprogenia_aberti[1] **
Serb, 2006 [Publ 2678], fig. 4
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 12 |
12 |
8 |
1 |
0 |
0 |
0 |
20.0 |
+-------------------- Obliquaria_reflexa **
|
| +-- Lampsilis_ornata **
+-----------------+
--+ +-- Lampsilis_siliquoidea **
|
| +----------------- Ptychobranchus_fasciolaris **
| |
+--+ +-- Cyprogenia_aberti[5] **
| +-----------+
| | +-- Dromus_dromas **
+--+
| +----------- Cyprogenia_aberti[4] **
| |
+--+ +-------- Cyprogenia_aberti[3] **
| |
+--+ +----- Cyprogenia_stegaria[2] **
| |
+--+----- Cyprogenia_aberti[2] **
|
| +-- Cyprogenia_stegaria[1] **
+--+
+-- Cyprogenia_aberti[1] **
Rosenberg et al., 1997 [Publ 2720], fig. 2A
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 12 |
17 |
0 |
4 |
11 |
6 |
2 |
28.0 |
+----- Anodonta_anatina
|
| +-- Gonidea_angulata
| |
| +-- Pyganodon_grandis
| |
| +-- Utterbackia_imbecillis
| |
| +-- Pyganodon_cataracta
--+ |
| +-- Lampsilis_straminea **
| |
| +-- Lampsilis_teres **
| |
| +-- Megalonaias_nervosa **
| |
| +-- Obliquaria_reflexa **
| |
+--+-- Theliderma_cylindrica **
|
+-- Quadrula_quadrula **
|
+-- Uniomerus_tetralasmus **
|
+-- Amblema_plicata **
|
+-- Elliptio_complanata **
|
+-- Fusconaia_cerina **
|
+-- Plectomerus_dombeyanus **
|
+-- Pleurobema_cordatum **
|
+-- Unio_pictorum
Roe, 2013 [Publ 3571], fig. 4
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 12 |
12 |
9 |
1 |
0 |
0 |
0 |
24.0 |
+----------------------- Obliquaria_reflexa **
|
| +-- Lampsilis_siliquoidea **
+--------------------+
--+ +-- Lampsilis_ornata **
|
| +-------------------- Medionidus_conradicus **
| |
+--+ +-- Cyprogenia_aberti **
| +--------------+
| | +-- Dromus_dromas **
+--+
| +-------------- Ptychobranchus_subtentus **
+--+
| +----------- Ptychobranchus_jonesi **
+--+
| +-------- Ptychobranchus_greenii **
+--+
| +----- Ptychobranchus_occidentalis[2] **
+--+
| +-- Ptychobranchus_occidentalis[1] **
+--+
+-- Ptychobranchus_fasciolaris **
Smith et al., 2020 [Publ 4136], fig. 3
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 12 |
12 |
10 |
1 |
0 |
0 |
0 |
28.0 |
+-------------- Pleurobema_clava **
|
| +----- Fusconaia_subrotunda **
| +-----+
--+ | | +-- Fusconaia_cor **
| | +--+
| | +-- Fusconaia_cuneolus **
| |
| | +-- Fusconaia_iheringi **
+--+ +--+
| | +-- Fusconaia_mitchelli **
| +--+
| | | +-- Fusconaia_askewi **
| | +--+
| | +-- Fusconaia_chunii **
+--+
| +-- Fusconaia_flava **
| +--+
| | +-- Fusconaia_masoni **
+--+
| +-- Fusconaia_burkei **
+--+
+-- Fusconaia_escambia **
Porto-Hannes et al., 2021 [Publ 4227], fig. 4
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 12 |
12 |
6 |
4 |
0 |
0 |
0 |
21.0 |
+-------------------- Quadrula_quadrula **
|
+-------------------- Pustulosa_pustulosa **
--+
| +----------------- Pleurobema_sintoxia **
| |
+--+ +-------------- Amblema_plicata **
| |
+--+ +----------- Obovaria_olivaria **
| |
+--+ +-------- Ortmanniana_ligamentina **
| |
+--+ +----- Lampsilis_hydiana **
| |
| +----- Lampsilis_straminea **
+--+
| +-- Lampsilis_siliquoidea **
| |
| +-- Lampsilis_radiata **
+--+
+-- Lampsilis[1] **
|
+-- Lampsilis[2] **
Stodola et al., 2021 [Publ 4266], fig. 4
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 12 |
12 |
7 |
1 |
0 |
0 |
0 |
25.0 |
+-- Ortmanniana_ligamentina **
|
+-----------+-- Lampsilis_abrupta **
| |
| +-- Lampsilis_higginsii **
--+
| +----------- Lampsilis_sietmani **
| |
| | +-- Lampsilis_siliquoidea **
+--+ +-----+
| | +-- Lampsilis_radiata **
| |
+--+ +----- Lampsilis_hydiana[2] **
| |
| +----- Lampsilis_straminea[2] **
| |
+--+ +-- Lampsilis_hydiana[1] **
+--+
| +-- Lampsilis_bergmanni **
|
| +-- Lampsilis_straminea[1] **
+--+
+-- Lampsilis_virescens **
Dai et al., 2024 [Publ 4358], fig. 4
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 12 |
12 |
10 |
5 |
40 |
38 |
2 |
5.0 |
+-- Quadrula_quadrula **
+--------------------+
| +-- Uniomerus_tetralasmus **
|
+--------------+ +-- Elliptio_complanata **
| | +-----------------+
| | | +-- Pleurobema_oviforme **
| +--+
| | +----------------- Amblema_plicata **
| | |
| +--+ +-------------- Popenaias_popeii **
| | |
| +--+ +----------- Toxolasma_parvum **
| | |
| +--+ +-- Potamilus_leptodon **
| | +-----+
| | | +-- Potamilus_alatus **
| +--+
| | +----- Leaunio_lienosus **
| +--+
| | +-- Lampsilis_cardium **
| +--+
--+ +-- Venustaconcha_ellipsiformis **
|
| +-- Chamberlainia_somsakpanhai
| +--------------+
| | +-- Sinohyriopsis_cumingii
| |
| | +----- Potomida_littoralis
| +-----------------+ +--------+
| | | | | +-- Lamprotula_leaii
| | | | +--+
| | | | +-- Pronodularia_japanensis
| | +--+
| | | +-- Pseudodon_vondembuschianus
| | | +--------+
| | | | +-- Pilsbryoconcha_exilis
| | +--+
| | | +-------- Microcondylaea_bonellii
| | +--+
| | | +----- Sinosolenaia_oleivora
+--+ +--+
| | +-- Ptychorhynchus_pfisteri
| +--+
| +-- Parvasolenaia_rivularis
|
| +-------------------------------- Lepidodesma_languilati
| |
| | +----------------- Lanceolaria_gladiola
| | |
| | +-----------+ +-------------- Acuticosta_chinensis
| | | | |
| | | | | +----- Cristaria_plicata
+--+ | +--+ +-----+
| | | | | +-- Anemina_arcaeformis
| | | | +--+
| | +--+ +-- Sinanodonta_woodiana
| | |
| | | +-------- Anodonta_anatina
| | +--+
+--+ | +----- Platynaias_compressa
| +--+
| | +-- Pyganodon_grandis
| +--+
| +-- Utterbackia_imbecillis
|
| +-- Aculamprotula_scripta
| +-----------------------+
| | +-- Aculamprotula_tientsinensis
| |
| | +-- Unio_crassus
+--+ +--------------------+
| | +-- Unio_delphinus
| |
| | +-------- Pseudocuneopsis_capitata
+--+ +-----------+
| | | +----- Pseudocuneopsis_yangshuoensis
| | +--+
| | | +-- Pseudocuneopsis_yemaoi
| | +--+
+--+ +-- Pseudocuneopsis_sichuanensis
|
| +-- Nodularia_breviconcha
| +--------------+
| | +-- Nodularia_douglasiae
| |
+--+ +-- Schistodesmus_lampreyanus
| +-----------+
| | +-- Schistodesmus_spinosus
+--+
| +----------- Tchangsinaia_piscicula
+--+
| +-------- Cuneopsis_rufescens
+--+
| +----- Cuneopsis_celtiformis
+--+
| +-- Cuneopsis_heudei
+--+
+-- Cuneopsis_demangei
Bogan & Hoeh, 2000 [Publ 2650], fig. 1
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 11 |
11 |
5 |
4 |
23 |
7 |
3 |
24.0 |
+-------- Toxolasma_lividum **
|
+-------- Amblema_plicata **
|
| +-- Glebula_rotundata **
+-----+
| +-- Cyrtonaias_tampicoensis **
|
--+ +----- Potamilus_alatus **
+--+
| | +-- Ortmanniana_ligamentina **
| +--+
| +-- Ligumia_recta **
|
| +----- Quadrula_quadrula **
| |
+--+ +-- Eurynia_dilatata **
| |
+--+-- Pleurobema_clava **
|
+-- Fusconaia_flava **
Hoeh et al., 2001 [Publ 2652], fig. 14.2
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 11 |
28 |
16 |
4 |
20 |
7 |
3 |
39.0 |
+-------------------- Amblema_plicata **
|
+-------------------- Coelatura_aegyptiaca
|
+-------------------- Eurynia_dilatata **
|
+-------------------- Fusconaia_flava **
|
+-------------------- Gonidea_angulata
|
+-------------------- Pleurobema_clava **
|
+-------------------- Quadrula_quadrula **
|
--+ +----------- Ortmanniana_ligamentina **
+--------+
| | +-------- Glebula_rotundata **
| +--+
| | +----- Potamilus_alatus **
| +--+
| | +-- Ligumia_recta **
| +--+
| +-- Toxolasma_lividum **
|
| +----------------- Cyrtonaias_tampicoensis **
| |
| | +-- Unio_pictorum
| | +-----------+
+--+ | +-- Unio_tumidus
| |
| | +----- Pyganodon_grandis
| | +-----+
+--+ | | +-- Anodonta_cygnea
| | +--+
| | +-- Strophitus_undulatus
| |
| | +----- Diplodon_gratus
+--+ |
| +----- Echyridella_menziesii
| +--+
| | +----- Lortiella_rugata
| | |
| | | +-- Castalia_stevensi
| | +--+
+--+ +-- Velesunio_angasi
|
| +-- Mutela_dubia
| +--+
| | +-- Mutela_rostrata
+--+
| +-- Anodontites_guanarensis
| |
+--+-- Anodontites_trigona
|
+-- Monocondylaea_minuana
Hoeh et al., 2001 [Publ 2652], fig. 14.4
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 11 |
11 |
9 |
4 |
20 |
7 |
3 |
24.0 |
+----------------------- Gonidea_angulata
|
| +-------- Unio_pictorum
| |
| +-----------+-------- Unio_tumidus
--+ | |
| | | +----- Anodonta_cygnea
| | +--+
| | | +-- Strophitus_undulatus
| | +--+
+--+ +-- Pyganodon_grandis
|
| +----------------- Amblema_plicata **
| |
| | +-------- Quadrula_quadrula **
| | +-----+
+--+ | | +----- Eurynia_dilatata **
| | +--+
| | | +-- Fusconaia_flava **
| | +--+
+--+ +-- Pleurobema_clava **
|
| +-- Cyrtonaias_tampicoensis **
| +--------+
| | +-- Glebula_rotundata **
+--+
| +-------- Toxolasma_lividum **
+--+
| +----- Potamilus_alatus **
+--+
| +-- Ligumia_recta **
+--+
+-- Ortmanniana_ligamentina **
Hoeh et al., 2002 [Publ 2653], fig. 2
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 11 |
11 |
8 |
4 |
20 |
7 |
3 |
25.0 |
+-------------- Gonidea_angulata
+-----+
| | +----------- Unio_pictorum
| +--+
| | +-------- Unio_tumidus
| +--+
| | +----- Anodonta_cygnea
| +--+
--+ | +-- Pyganodon_grandis
| +--+
| +-- Strophitus_undulatus
|
| +-- Glebula_rotundata **
| +--------------+
| | +-- Cyrtonaias_tampicoensis **
| |
+--+ +-------------- Toxolasma_lividum **
| |
| | +----- Potamilus_alatus **
| +--------+
+--+ | +-- Ligumia_recta **
| +--+
| +-- Ortmanniana_ligamentina **
|
| +----------- Amblema_plicata **
+--+
| +-------- Quadrula_quadrula **
+--+
| +----- Eurynia_dilatata **
+--+
| +-- Pleurobema_clava **
+--+
+-- Fusconaia_flava **
Roe & Hoeh, 2003 [Publ 2655], fig. 4.2
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 11 |
16 |
12 |
4 |
20 |
7 |
3 |
29.0 |
+----------------------------- Gonidea_angulata
|
--+ +-------------------------- Coelatura_aegyptiaca
| |
+--+ +----------------------- Fusconaia_flava **
| |
| +----------------------- Amblema_plicata **
+--+
| +-------------------- Quadrula_quadrula **
| |
| | +----------------- Pleurobema_clava **
+--+ |
| | +-------------- Eurynia_dilatata **
| | |
+--+ | +----------- Ortmanniana_ligamentina **
| +--+
| | | +-------- Glebula_rotundata **
| | +--+
| | | +----- Potamilus_alatus **
+--+ +--+
| | +-- Ligumia_recta **
| +--+
| +-- Toxolasma_lividum **
|
| +----------- Cyrtonaias_tampicoensis **
| |
+--+ +-- Unio_pictorum
| +-----+
| | +-- Unio_tumidus
+--+
| +----- Pyganodon_grandis
+--+
| +-- Anodonta_cygnea
+--+
+-- Strophitus_undulatus
Roe & Hoeh, 2003 [Publ 2655], fig. 4.3
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 11 |
11 |
9 |
4 |
20 |
7 |
3 |
24.0 |
+-- Unio_pictorum
+-----+
| +-- Unio_tumidus
+--------------+
| | +----- Anodonta_cygnea
| +--+
| | +-- Pyganodon_grandis
| +--+
--+ +-- Strophitus_undulatus
|
| +-------------------- Gonidea_angulata
| |
| | +----------------- Amblema_plicata **
+--+ |
| | +-------- Quadrula_quadrula **
| | +-----+
+--+ | | +----- Eurynia_dilatata **
| | +--+
| | | +-- Fusconaia_flava **
| | +--+
+--+ +-- Pleurobema_clava **
|
| +-- Cyrtonaias_tampicoensis **
| +--------+
| | +-- Glebula_rotundata **
+--+
| +-------- Toxolasma_lividum **
+--+
| +----- Ortmanniana_ligamentina **
+--+
| +-- Ligumia_recta **
+--+
+-- Potamilus_alatus **
Roe & Lydeard, 1998 [Publ 2693], fig. 3
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 11 |
11 |
5 |
2 |
0 |
0 |
0 |
22.0 |
+-------------- Fusconaia_cerina **
|
+-------------- Obliquaria_reflexa **
--+
| +----------- Potamilus_capax **
| |
| +----------- Lampsilis_ornata **
+--+
| +-------- Potamilus_fragilis **
| |
| | +----- Potamilus_inflatus **
+--+ |
| | +-- Potamilus_ohiensis **
| +--+
+--+ +-- Potamilus_amphichaenus **
|
| +-- Potamilus_purpuratus[1] **
| |
+--+-- Potamilus_purpuratus[2] **
|
+-- Potamilus_alatus **
Rosenberg et al., 1994 [Publ 2719], fig. 3
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 11 |
15 |
2 |
4 |
8 |
5 |
2 |
29.0 |
+-------- Gonidea_angulata
|
| +-- Elliptio_complanata **
| |
| +-- Fusconaia_cerina **
| |
--+ +--+-- Plectomerus_dombeyanus **
| | |
| | +-- Pleurobema_cordatum **
| | |
| | +-- Unio_pictorum
| |
| | +-- Pyganodon_cataracta
+--+ |
| +-- Pyganodon_grandis
| |
| +-- Utterbackia_imbecillis
| |
| +-- Lampsilis_teres **
| |
| +-- Lampsilis_straminea **
+--+
+-- Megalonaias_nervosa **
|
+-- Obliquaria_reflexa **
|
+-- Theliderma_cylindrica **
|
+-- Quadrula_quadrula **
|
+-- Uniomerus_tetralasmus **
Rosenberg et al., 1997 [Publ 2720], fig. 2B
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 11 |
17 |
0 |
4 |
12 |
7 |
2 |
28.0 |
+----- Anodonta
|
| +-- Gonidea_angulata
| |
| +-- Pyganodon_grandis
| |
| +-- Utterbackia_imbecillis
| |
| +-- Pyganodon_cataracta
--+ |
| +-- Lampsilis_straminea **
| |
| +-- Lampsilis_teres **
| |
| +-- Megalonaias_nervosa **
| |
| +-- Obliquaria_reflexa **
| |
+--+-- Theliderma_cylindrica **
|
+-- Quadrula_quadrula **
|
+-- Uniomerus_tetralasmus **
|
+-- Amblema_plicata **
|
+-- Elliptio_complanata **
|
+-- Fusconaia_cerina **
|
+-- Plectomerus_dombeyanus **
|
+-- Aculamprotula_tortuosa
|
+-- Unio_pictorum
Fukata & Iigo, 2020 [Publ 4106], fig. 1
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 11 |
11 |
9 |
3 |
43 |
40 |
2 |
6.0 |
+-- Quadrula_quadrula **
+-----------------+
| +-- Uniomerus_tetralasmus **
+--+
| | +----------------- Elliptio_complanata **
| | |
| +--+ +-- Potamilus_leptodon **
| | +-----------+
| | | +-- Potamilus_alatus **
| +--+
| | +----------- Leaunio_lienosus **
| | |
| +--+ +-- Lampsilis_cardium **
| | +-----+
| | | +-- Lampsilis_ornata **
+--+ +--+
| | | +----- Venustaconcha_ellipsiformis **
| | +--+
| | | +-- Lampsilis_powellii **
| | +--+
| | +-- Lampsilis_siliquoidea **
| |
| | +-- Sinohyriopsis_cumingii
| | +-----------+
| | | +-- Sinohyriopsis_schlegelii
| | |
| | | +-- Pilsbryoconcha_exilis
| +--------+ +--------+
| | | +-- Pseudodon_vondembuschianus
| | |
| | | +-- Lamprotula_caveata
| | | +--+
| +--+ | +-- Lamprotula_leaii
| | +--+
--+ | | | +-- Pronodularia_japanensis
| | | +--+
| | | +-- Potomida_littoralis
| +--+
| | +-- Sinosolenaia_carinata
| | +--+
| | | +-- Sinosolenaia_oleivora
| +--+
| | +-- Parvasolenaia_rivularis
| +--+
| +-- Ptychorhynchus_pfisteri
|
| +----------------------- Lepidodesma_languilati
| |
| | +----- Lanceolaria_gladiola
| | +--------------+
| | | | +-- Lanceolaria_grayii
| | | +--+
| | | +-- Lanceolaria_lanceolata
+--+ |
| | +-------------- Acuticosta_chinensis
| | |
| | | +-------- Cristaria_plicata
| | | |
| | | +--+ +-- Sinanodonta_woodiana
| | +--+ | | +--+
+--+ | | | | | +-- Sinanodonta_lucida
| | | | +--+
| | | | | +-- Anemina_arcaeformis
| | | | +--+
| | +--+ +-- Anemina_euscaphys
| | |
| | | +-- Anodonta_anatina
| | | +-----+
| | | | +-- Anodonta_cygnea
| | | |
| | +--+ +-- Pyganodon_grandis
+--+ | +--+
| | | +-- Utterbackia_peninsularis
| +--+
| | +-- Alasmidonta_varicosa
| +--+
| +-- Prolasmidonta_heterodon
|
| +----- Aculamprotula_tortuosa
| +--------+
| | | +-- Aculamprotula_tientsinensis
| | +--+
| | +-- Aculamprotula_coreana
| |
+--+ +-------- Unio_tumidus
| +--+
| | | +----- Unio_crassus
| | +--+
| | | +-- Unio_pictorum
| | +--+
+--+ +-- Unio_delphinus
|
| +-- Schistodesmus_lampreyanus
| +-----+
| | +-- Cuneopsis_heudei
+--+
| +----- Pseudocuneopsis_capitata
+--+
| +-- Nodularia_douglasiae
+--+
+-- Tchangsinaia_piscicula
Buhay et al., 2002 [Publ 2661], fig. 2A
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 10 |
10 |
3 |
1 |
0 |
0 |
0 |
22.0 |
+-------- Lampsilis_siliquoidea **
|
| +-- Cambarunio_taeniatus **
+-----+
| +-- Villosa_villosa **
--+
| +----- Epioblasma_triquetra **
| |
| +----- Epioblasma_brevidens **
| |
+--+ +-- Epioblasma_capsaeformis[1] **
| |
| +-- Epioblasma_capsaeformis[2] **
| |
+--+-- Epioblasma_capsaeformis[3] **
|
+-- Epioblasma_walkeri[1] **
|
+-- Epioblasma_walkeri[2] **
Buhay et al., 2002 [Publ 2661], fig. 2B
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 10 |
10 |
5 |
1 |
0 |
0 |
0 |
22.0 |
+-------------- Lampsilis_siliquoidea **
|
| +-- Cambarunio_taeniatus **
+-----------+
--+ +-- Villosa_villosa **
|
| +----------- Epioblasma_brevidens **
| |
+--+ +-------- Epioblasma_triquetra **
| |
+--+ +----- Epioblasma_capsaeformis[3] **
| |
+--+ +-- Epioblasma_capsaeformis[1] **
| |
| +-- Epioblasma_capsaeformis[2] **
+--+
+-- Epioblasma_walkeri[1] **
|
+-- Epioblasma_walkeri[2] **
Huang et al., 2019 [Publ 3843], fig. 3
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 10 |
10 |
8 |
3 |
47 |
37 |
2 |
7.0 |
+-- Sinohyriopsis_cumingii
+-----------+
| +-- Sinohyriopsis_schlegelii
|
| +-------- Potomida_littoralis
+--------+ +--+
| | | | +----- Pronodularia_japanensis
| | | +--+
| | | | +-- Lamprotula_caveata
| | | +--+
| +--+ +-- Lamprotula_leaii
| |
| | +-- Ptychorhynchus_pfisteri
| | +--+
+--------+ | | +-- Parvasolenaia_rivularis
| | +-----+
| | | +-- Sinosolenaia_oleivora
| | +--+
| | +-- Sinosolenaia_carinata
| |
| | +-- Quadrula_quadrula **
| | +-----------------+
| | | +-- Uniomerus_tetralasmus **
| +--+
| | +----------------- Elliptio_complanata **
| | |
| +--+ +-------------- Toxolasma_parvum **
| | |
| +--+ +-- Potamilus_leptodon **
| | +--------+
| | | +-- Potamilus_alatus **
--+ +--+
| | +-------- Leaunio_lienosus **
| +--+
| | +----- Venustaconcha_ellipsiformis **
| +--+
| | +-- Lampsilis_cardium **
| +--+
| +-- Lampsilis_ornata **
|
| +----------------------------- Lepidodesma_languilati
| |
| | +-------- Aculamprotula_tortuosa
| | +-----+
| | | | +----- Aculamprotula_scripta
| | | +--+
| | | | +-- Aculamprotula_tientsinensis
| | | +--+
| | +-----------+ +-- Aculamprotula_coreana
+--+ | |
| | | +----- Nodularia_douglasiae
| | | +-----+
| | | | | +-- Tchangsinaia_piscicula
| | | | +--+
| | +--+ +-- Schistodesmus_lampreyanus
| | |
| | | +-------- Unio_tumidus
| | +--+
+--+ | +----- Unio_crassus
| +--+
| | +-- Unio_delphinus
| +--+
| +-- Unio_pictorum
|
| +----- Lanceolaria_gladiola
| +-----------------+
| | | +-- Lanceolaria_lanceolata
| | +--+
| | +-- Lanceolaria_grayii
+--+
| +-------------------- Acuticosta_chinensis
| |
| | +-------- Cristaria_plicata
| | |
+--+ +--------+ +-- Sinanodonta_lucida
| | | +--+
| | | | +-- Sinanodonta_woodiana
| | +--+
+--+ | +-- Anemina_arcaeformis
| +--+
| +-- Anemina_euscaphys
|
| +-------------- Anodonta_cygnea
+--+
| +----------- Anodonta_anatina
+--+
| +-------- Lasmigona_costata
+--+
| +----- Pyganodon_grandis
+--+
| +-- Utterbackia_imbecillis
+--+
+-- Utterbackia_peninsularis
Keogh & Simons, 2019 [Publ 3935], fig. S1
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 10 |
10 |
8 |
3 |
0 |
0 |
0 |
21.0 |
+-- Amblema_plicata **
+-----------------+
| +-- Fusconaia_mitchelli **
--+
| +----------------- Potamilus_fragilis **
| |
+--+ +-------------- Leaunio_lienosus **
| |
+--+ +-- Lampsilis_floridensis **
| +--------+
| | +-- Lampsilis_teres **
+--+
| +-------- Lampsilis_sietmani **
+--+
| +----- Lampsilis_siliquoidea **
+--+
| +-- Lampsilis_hydiana **
+--+
+-- Lampsilis_straminea **
Curole & Kocher, 2005 [Publ 2682], fig. 1
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 9 |
9 |
7 |
3 |
2 |
1 |
1 |
22.0 |
+----------------- Glebula_rotundata **
|
| +----- Amblema_plicata **
--+ +--------+
| | | +-- Quadrula_quadrula **
| | +--+
+--+ +-- Pustulosa_pustulosa **
|
| +----------- Plectomerus_dombeyanus **
+--+
| +-------- Leaunio_lienosus **
+--+
| +----- Lampsilis_straminea **
+--+
| +-- Lampsilis_teres **
+--+
+-- Potamilus_purpuratus **
Curole & Kocher, 2005 [Publ 2682], fig. 1-M
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 9 |
9 |
7 |
3 |
3 |
2 |
1 |
18.0 |
+-- Utterbackiana_implicata
+-----------------+
| +-- Pyganodon_grandis
|
--+ +-- Quadrula_quadrula **
| +--------------+
| | +-- Pustulosa_pustulosa **
+--+
| +-------------- Plectomerus_dombeyanus **
| |
+--+ +----------- Amblema_plicata **
| |
+--+ +-- Potamilus_purpuratus **
| +-----+
| | +-- Glebula_rotundata **
+--+
| +----- Leaunio_lienosus **
+--+
| +-- Lampsilis_straminea **
+--+
+-- Lampsilis_teres **
Lopes-Lima et al., 2017 [Publ 3745], fig. 1-2
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 9 |
9 |
7 |
4 |
72 |
60 |
3 |
19.0 |
+-------- Amblema_plicata **
+--+
| | +----- Cambarunio_iris **
| +--+
| | +-- Lampsilis_cardium **
| +--+
+--------+ +-- Ortmanniana_ligamentina **
| |
| | +-- Quadrula_quadrula **
| | +-----+
| | | +-- Quadrula_verrucosa **
| +--+
| | +----- Elliptio_complanata **
| +--+
| | +-- Pleurobema_sintoxia **
| +--+
| +-- Eurynia_dilatata **
--+
| +-- Chamberlainia_somsakpanhai
| +-----+
| | +-- Sinohyriopsis_cumingii
| |
| +--------+ +-- Lamprotula_caveata
| | | +--+
| | | | +-- Lamprotula_leaii
| | +--+
| | | +-- Potomida_littoralis
| | +--+
| | +-- Pronodularia_japanensis
+--+
| +-- Sinosolenaia_carinata
| +-----+
| | +-- Sinosolenaia_oleivora
| +-----+
| | | +----- Gonidea_angulata
| | +--+
| | | +-- Microcondylaea_bonellii
+--+ +--+
| +-- Leguminaia_wheatleyi
|
| +----------- Parvasolenaia_rivularis
| |
+--+ +-- Pilsbryoconcha_exilis
| +-----+
| | +-- Pilsbryoconcha_acuta
+--+
| +----- Pseudodon_vagulus
+--+
| +-- Bineurus_exilis
+--+
+-- Nyeinchanconcha_nyeinchani
Bolotov et al., 2017 [Publ 3761], fig. S1
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 9 |
9 |
7 |
4 |
132 |
107 |
3 |
19.0 |
+-------- Amblema_plicata **
+--+
| | +----- Ortmanniana_ligamentina **
| +--+
| | +-- Lampsilis_cardium **
| +--+
+-----+ +-- Cambarunio_iris **
| |
| | +-- Quadrula_quadrula **
| | +-----+
| | | +-- Quadrula_verrucosa **
| +--+
| | +----- Pleurobema_sintoxia **
| +--+
| | +-- Eurynia_dilatata **
| +--+
| +-- Elliptio_complanata **
+--------+
| | +-- Sinohyriopsis_cumingii
| | +-----+
| | | +-- Chamberlainia_somsakpanhai
| | |
| | +-----+ +-- Potomida_littoralis
| | | | +--+
| | | | | +-- Pronodularia_japanensis
| | | +--+
| | | | +-- Lamprotula_caveata
| | | +--+
| +--+ +-- Lamprotula_leaii
| |
| | +----- Sinosolenaia_carinata
| | +-----+
| | | | +-- Sinosolenaia_oleivora
| | | +--+
| +--+ +-- Sinosolenaia_recognita
| |
| | +-------- Gonidea_angulata
| +--+
--+ | +----- Microcondylaea_bonellii
| +--+
| | +-- Leguminaia_wheatleyi
| +--+
| +-- Leguminaia
|
| +----- Unio_tumidus
| +--------+
| | | +-- Unio_pictorum
| | +--+
| | +-- Unio_crassus
| +--------+
| | | +----------- Aculamprotula_tortuosa
| | | |
| | | | +-- Nodularia_douglasiae
| | +--+ +-----+
| | | | +-- Nodularia_nuxpersicae
| | | |
| | +--+ +-- Cuneopsis_rufescens
| | | +--+
| | | | +-- Cuneopsis_heudei
| | +--+
+--+ | +-- Tchangsinaia_piscicula
| +--+
| +-- Schistodesmus_lampreyanus
|
| +-- Lanceolaria_grayii
| +-----------------+
| | +-- Lanceolaria_gladiola
| |
| | +-- Cristaria_plicata
| | +--------+
| | | +-- Cristaria_radiata
+--+ +-----+
| | | +-------- Pletholophus_tenuis
| | +--+
| | | +----- Sinanodonta_lucida
| | +--+
| | | +-- Sinanodonta_jourdyi
+--+ +--+
| +-- Sinanodonta_lauta
|
| +-------------- Anodonta_anatina
| |
| | +----- Pseudanodonta_complanata
+--+ +-----+
| | | +-- Anodonta_cygnea
| | +--+
| | +-- Anodonta_nuttalliana
+--+
| +-- Lasmigona_costata
| +-----+
| | +-- Alasmidonta_marginata
+--+
| +----- Simpsonaias_ambigua
+--+
| +-- Pyganodon_grandis
+--+
+-- Strophitus_undulatus
Bolotov et al., 2017 [Publ 3761], fig. S2
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 9 |
9 |
7 |
4 |
132 |
107 |
3 |
19.0 |
+-------- Lens_contradens
+-----+
| | +----- Lens_eximius
| +--+
| | +-- Lens_comptus
| +--+
+--+ +-- Lens_novoselovi
| |
| | +----- Physunio
| | +-----+
| | | | +-- Physunio_modelli
| | | +--+
| +--+ +-- Physunio_superbus
| |
| | +-------- Yaukthwa_inlenensis
| +--+
| | +----- Yaukthwa_panhai
+-----+ +--+
| | | +-- Yaukthwa_rectangularis
| | +--+
| | +-- Yaukthwa_nesemanni
| |
| | +----- Hyriopsis_panhai
| | |
| | +--------+----- Hyriopsis_khoratensis
| | | |
| | | | +-- Hyriopsis_myersiana
| | | +--+
| +--+ +-- Hyriopsis_bialata
| |
| | +----------- Hyriopsis_desowitzi
| | |
| | +----------- Rectidens_sumatrensis
| +--+
| | +-- Ensidens_telus
| | +-----+
| | | +-- Ensidens_ingallsianus
| +--+
| | +----- Ensidens_sagittarius
| +--+
| | +-- Ensidens_spiculus
--+ +--+
| +-- Ensidens_jaculus
|
| +-------- Amblema_plicata **
| +--+
| | | +----- Cambarunio_iris **
| | +--+
| | | +-- Ortmanniana_ligamentina **
| | +--+
| +--------+ +-- Lampsilis_cardium **
| | |
| | | +-- Quadrula_quadrula **
| | | +-----+
| | | | +-- Quadrula_verrucosa **
| | +--+
| | | +----- Pleurobema_sintoxia **
| | +--+
| | | +-- Elliptio_complanata **
| | +--+
| | +-- Eurynia_dilatata **
| |
| | +-------- Sinohyriopsis_cumingii
| | |
| | +-----+ +-- Lamprotula_caveata
+--+ | | +--+
| | | | +-- Lamprotula_leaii
| | +--+
| | | +-- Potomida_littoralis
| +--+ +--+
| | | +-- Pronodularia_japanensis
| | |
| | | +----------- Sinosolenaia_carinata
| | | |
| | | | +-- Sinosolenaia_oleivora
| | +--+--------+
| | | +-- Sinosolenaia_recognita
| | |
| | | +-------- Gonidea_angulata
| | +--+
| | | +----- Microcondylaea_bonellii
+--+ +--+
| | +-- Leguminaia_wheatleyi
| +--+
| +-- Leguminaia
|
| +-------- Pilsbryoconcha_exilis[3]
| +-----+
| | | +----- Pilsbryoconcha_exilis[2]
| | +--+
| | | +-- Pilsbryoconcha_exilis[1]
| | +--+
| | +-- Pilsbryoconcha_carinifera
| |
+--+ +----------- Indopseudodon_indawgyiensis
| |
| | +-------- Thaiconcha_callifera
| +--+
| | | +----- Bineurus_exilis
| | +--+
| | | +-- Bineurus_mouhotii
+--+ +--+
| +-- Bineurus_loeiensis
|
| +----- Pseudodon_vagulus
| +--+
| | | +-- Pseudodon_mekongi
| | +--+
| | +-- Pseudodon_vondembuschianus
+--+
| +-- Pseudodon_inoscularis
| +--+
| | +-- Sundadontina_tumida
+--+
| +-- Nyeinchanconcha_nyeinchani
+--+
+-- Sundadontina_cumingii
Bolotov et al., 2017 [Publ 3764], fig. S1
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 9 |
9 |
0 |
4 |
122 |
122 |
3 |
20.0 |
+-- Amblema_plicata **
|
+-- Ortmanniana_ligamentina **
|
+-- Lampsilis_cardium **
|
+-- Cambarunio_iris **
|
+--+-- Elliptio_complanata **
| |
| +-- Eurynia_dilatata **
| |
| +-- Pleurobema_sintoxia **
| |
| +-- Quadrula_quadrula **
| |
| +-- Quadrula_verrucosa **
|
| +-- Chamberlainia_somsakpanhai
+--+ |
| | +-- Sinohyriopsis_cumingii
| | |
| | +-- Lamprotula_caveata
| | |
| | +-- Lamprotula_leaii
| | |
| | +-- Potomida_littoralis
| | |
| | +-- Pronodularia_japanensis
| | |
| +--+-- Gonidea_angulata
| |
| +-- Leguminaia_wheatleyi
| |
| +-- Leguminaia
| |
| +-- Microcondylaea_bonellii
| |
| +-- Sinosolenaia_carinata
| |
| +-- Sinosolenaia_oleivora
| |
| +-- Sinosolenaia_recognita
--+
| +-- Unio_crassus
| |
| +-- Unio_pictorum
| |
| +-- Unio_tumidus
| |
| +-- Aculamprotula_tortuosa
| |
| +-- Cuneopsis_heudei
| +--+
| | +-- Tchangsinaia_piscicula
| | |
| | +-- Cuneopsis_rufescens
| | |
| | +-- Nodularia_douglasiae
| | |
| | +-- Nodularia_nuxpersicae
| | |
| | +-- Schistodesmus_lampreyanus
| |
| | +-- Alasmidonta_marginata
| | |
| | +-- Anodonta_anatina
+--+ |
| +-- Anodonta_cygnea
| |
| +-- Anodonta_nuttalliana
| |
| +-- Lasmigona_costata
| |
| +-- Pseudanodonta_complanata
| |
| +-- Pyganodon_grandis
| |
| +-- Simpsonaias_ambigua
| |
| +-- Strophitus_undulatus
+--+
+-- Sinanodonta_lauta
|
+-- Cristaria_plicata
|
+-- Cristaria_radiata
|
+-- Pletholophus_tenuis
|
+-- Sinanodonta_lucida
|
+-- Sinanodonta_jourdyi
|
+-- Lanceolaria_gladiola
|
+-- Lanceolaria_grayii[1]
|
+-- Lanceolaria_grayii[2]
Wu et al., 2018 [Publ 3836], fig. 2
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 9 |
51 |
43 |
4 |
44 |
42 |
2 |
61.0 |
+-------------------------------- Quadrula_quadrula **
|
| +-- Amblema_plicata **
+-----------------------------+
| +-- Obovaria_arkansasensis **
|
| +----------- Pleurobema_sintoxia **
| |
+--------------------+ +-- Elliptio_complanata **
| | +-----+
| | | +-- Eurynia_dilatata **
| +--+
| | +----- Fusconaia_askewi **
| +--+
| | +-- Fusconaia_cerina **
| +--+
--+ +-- Fusconaia_burkei **
|
| +----- Potomida_littoralis
| +-----+
| | | +-- Lamprotula_caveata
| | +--+
| +-----------------+ +-- Lamprotula_leaii
| | |
| | | +-------- Sinosolenaia_carinata
| | | |
| | +--+ +----- Gonidea_angulata
| | | |
| | +--+----- Sinosolenaia_oleivora
| | |
| | | +-- Parvasolenaia_rivularis
| | +--+
| | +-- Ptychorhynchus_pfisteri
+--+
| +-------------------------- Sinohyriopsis_cumingii
| |
| | +----------------------- Lepidodesma_languilati
| | |
| | | +----------- Aculamprotula_tortuosa
| | | +-----+
| | | | | +-------- Aculamprotula_scripta
| | | | +--+
| | | | | +----- Aculamprotula_coreana
| | | | +--+
+--+ | | | +-- Aculamprotula_fibrosa
| +-----+ +--+
| | | +-- Aculamprotula_tientsinensis
| | |
| | | +-- Unio_crassus
| | | +-----------+
| | | | +-- Unio_pictorum
| | | |
| | +--+ +----- Nodularia_douglasiae
| | | +-----+
| | | | | +-- Schistodesmus_lampreyanus
+--+ | | +--+
| +--+ +-- Schistodesmus_spinosus
| |
| | +-------- Tchangsinaia_piscicula
| +--+
| | +----- Cuneopsis_rufescens
| +--+
| | +-- Cuneopsis_celtiformis
| +--+
| +-- Cuneopsis_heudei
|
| +----------- Acuticosta_chinensis
| +--------+
| | | +-------- Lanceolaria_gladiola
| | +--+
| | | +----- Lanceolaria_grayii
| | +--+
| | | +-- Lanceolaria_lanceolata
| | +--+
| | +-- Lanceolaria_triformis
+--+
| +-------- Anemina_arcaeformis
| |
| +-------- Cristaria_plicata
| +--------+
| | | +-- Sinanodonta_woodiana
| | | +--+
| | | | +-- Sinanodonta_elliptica
| | +--+
+--+ | +-- Sinanodonta_lucida
| +--+
| +-- Sinanodonta_angula
|
| +-------------- Anodonta_cygnea
| |
+--+ +----------- Anodonta_anatina
| |
+--+ +-------- Strophitus_undulatus
| |
+--+ +----- Platynaias_compressa
| |
+--+----- Alasmidonta_marginata
|
| +-- Pyganodon_cataracta
+--+
+-- Pyganodon_grandis
Jeratthitikul et al., 2022 [Publ 4260], fig. S1, 1
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 9 |
9 |
7 |
4 |
67 |
57 |
2 |
19.0 |
+-- Chamberlainia_somsakpanhai
+-----+
| +-- Sinohyriopsis_cumingii
+-----------------+
| | +----- Lamprotula_leaii
| +--+
| | +-- Potomida_littoralis
| +--+
| +-- Pronodularia_japanensis
|
| +-------- Amblema_plicata **
| +--+
| | | +----- Ortmanniana_ligamentina **
| | +--+
| | | +-- Cambarunio_iris **
| | +--+
--+ +-----------+ +-- Lampsilis_cardium **
| | |
| | | +-- Quadrula_quadrula **
| | | +-----+
| | | | +-- Quadrula_verrucosa **
| | +--+
| | | +----- Elliptio_complanata **
| | +--+
| | | +-- Pleurobema_sintoxia **
| | +--+
| | +-- Eurynia_dilatata **
| |
| | +----- Unio_tumidus
+--+ +-----------+
| | | +-- Unio_pictorum
| | +--+
| +--+ +-- Unio_crassus
| | |
| | | +-------------- Aculamprotula_tortuosa
| | | |
| | +--+ +-- Nodularia_douglasiae
| | | +--------+
| | | | +-- Nodularia_nuxpersicae
| | +--+
| | | +-------- Schistodesmus_lampreyanus
| | +--+
| | | +----- Cuneopsis_heudei
+--+ +--+
| | +-- Tchangsinaia_piscicula
| +--+
| +-- Cuneopsis_rufescens
|
| +-- Lanceolaria_grayii
| +--------------+
| | +-- Lanceolaria_gladiola
| |
| | +----------- Cristaria_plicata
| | +--+
+--+ | | +-------- Pletholophus_tenuis
| | +--+
| | | +----- Sinanodonta_lucida
| | +--+
| | | +-- Sinanodonta_lauta
| | +--+
+--+ +-- Sinanodonta_jourdyi
|
| +-------- Pseudanodonta_complanata
| +--+
| | | +----- Anodonta_nuttalliana
| | +--+
| | | +-- Anodonta_anatina
| | +--+
+--+ +-- Anodonta_cygnea
|
| +-- Pyganodon_grandis
| +-----+
| | +-- Simpsonaias_ambigua
+--+
| +----- Strophitus_undulatus
+--+
| +-- Alasmidonta_marginata
+--+
+-- Lasmigona_costata
Jeratthitikul et al., 2022 [Publ 4260], fig. S2, 2
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 9 |
9 |
7 |
4 |
67 |
57 |
2 |
19.0 |
+-- Chamberlainia_somsakpanhai
+-----+
| +-- Sinohyriopsis_cumingii
+-----------------+
| | +----- Potomida_littoralis
| +--+
| | +-- Lamprotula_leaii
| +--+
| +-- Pronodularia_japanensis
|
| +-------- Amblema_plicata **
| +--+
| | | +----- Ortmanniana_ligamentina **
| | +--+
| | | +-- Lampsilis_cardium **
| | +--+
--+ +-----------+ +-- Cambarunio_iris **
| | |
| | | +-- Quadrula_quadrula **
| | | +-----+
| | | | +-- Quadrula_verrucosa **
| | +--+
| | | +----- Pleurobema_sintoxia **
| | +--+
| | | +-- Eurynia_dilatata **
| | +--+
| | +-- Elliptio_complanata **
| |
| | +----- Unio_tumidus
+--+ +-----------+
| | | +-- Unio_pictorum
| | +--+
| +--+ +-- Unio_crassus
| | |
| | | +-------------- Aculamprotula_tortuosa
| | | |
| | +--+ +-- Nodularia_douglasiae
| | | +--------+
| | | | +-- Nodularia_nuxpersicae
| | +--+
| | | +-------- Schistodesmus_lampreyanus
| | +--+
| | | +----- Cuneopsis_heudei
+--+ +--+
| | +-- Tchangsinaia_piscicula
| +--+
| +-- Cuneopsis_rufescens
|
| +-- Lanceolaria_grayii
| +--------------+
| | +-- Lanceolaria_gladiola
| |
| | +----------- Cristaria_plicata
| | +--+
+--+ | | +-------- Pletholophus_tenuis
| | +--+
| | | +----- Sinanodonta_lucida
| | +--+
| | | +-- Sinanodonta_jourdyi
| | +--+
+--+ +-- Sinanodonta_lauta
|
| +-------- Pseudanodonta_complanata
| +--+
| | | +----- Anodonta_anatina
| | +--+
| | | +-- Anodonta_nuttalliana
| | +--+
+--+ +-- Anodonta_cygnea
|
| +-- Alasmidonta_marginata
| +-----+
| | +-- Lasmigona_costata
+--+
| +----- Pyganodon_grandis
+--+
| +-- Strophitus_undulatus
+--+
+-- Simpsonaias_ambigua
Smith, 2021 [Publ 4331], fig. 1
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 9 |
9 |
7 |
4 |
11 |
7 |
3 |
14.0 |
+----- Sinohyriopsis_cumingii
+-----+
| | +-- Lens_contradens
| +--+
+--------+ +-- Lamprotula_leaii
| |
| | +-------- Scabies_phaselus
| +--+
| | +----- Cristaria_plicata
| +--+
| | +-- Pyganodon_grandis
--+ +--+
| +-- Utterbackia_peninsularis
|
| +-- Uniomerus_tetralasmus **
| +--------------+
| | +-- Megalonaias_nervosa **
| |
+--+ +-- Elliptio_crassidens **
| +-----------+
| | +-- Elliptio_complanata **
+--+
| +----------- Amblema_plicata **
+--+
| +-------- Potamilus_streckersoni **
+--+
| +----- Venustaconcha_ellipsiformis **
+--+
| +-- Lampsilis_siliquoidea **
+--+
+-- Lampsilis_cardium **
Keogh & Simons, 2019 [Publ 3935], fig. 4
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 8 |
8 |
6 |
1 |
0 |
0 |
0 |
21.0 |
+----------------- Lampsilis_sietmani **
|
--+ +-- Lampsilis_floridensis **
| +-----------+
| | +-- Lampsilis_teres **
+--+
| +----------- Lampsilis_siliquoidea **
+--+
| +-------- Lampsilis_hydiana[2] **
+--+
| +----- Lampsilis_straminea[2] **
+--+
| +-- Lampsilis_hydiana[1] **
+--+
+-- Lampsilis_straminea[1] **
Jones & Neves, 2010 [Publ 2646], fig. 16
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 7 |
7 |
5 |
1 |
0 |
0 |
0 |
23.0 |
+-- Epioblasma_triquetra **
+-----------+
| +-- Epioblasma_brevidens **
--+
| +----------- Epioblasma_rangiana **
+--+
| +-------- Epioblasma_capsaeformis **
+--+
| +----- Epioblasma_ahlstedti **
+--+
| +-- Epioblasma_aureola **
+--+
+-- Epioblasma_walkeri **
Jones & Neves, 2010 [Publ 2646], fig. 17
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 7 |
7 |
1 |
1 |
0 |
0 |
0 |
23.0 |
+----- Epioblasma_triquetra **
|
+----- Epioblasma_brevidens **
|
--+ +-- Epioblasma_capsaeformis **
| |
| +-- Epioblasma_aureola **
| |
+--+-- Epioblasma_walkeri **
|
+-- Epioblasma_rangiana **
|
+-- Epioblasma_ahlstedti **
Zhou et al., 2007 [Publ 2658], fig. 2
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 7 |
7 |
5 |
1 |
14 |
13 |
2 |
22.0 |
+-- Theliderma_intermedia **
+-----------+
| +-- Theliderma_sparsa **
--+
| +----------- Quadrula_quadrula **
+--+
| +-------- Pustulosa_infucata **
+--+
| +----- Quadrula_verrucosa **
+--+
| +-- Pustulosa_kieneriana **
+--+
+-- Pustulosa_succissa **
Buhay et al., 2002 [Publ 2661], fig. 1
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 7 |
7 |
3 |
1 |
0 |
0 |
0 |
21.0 |
+----------- Lampsilis_siliquoidea[1] **
|
+----------- Lampsilis_siliquoidea[2] **
--+
| +-------- Epioblasma_brevidens **
| |
+--+ +----- Epioblasma_triquetra **
| |
+--+ +-- Epioblasma_capsaeformis **
| |
+--+-- Epioblasma_walkeri[1] **
|
+-- Epioblasma_walkeri[2] **
Amaro et al., 2017 [Publ 3791], fig. 2
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 7 |
9 |
7 |
3 |
11 |
7 |
2 |
19.0 |
+-- Lasmigona_costata
+-----+
| +-- Platynaias_compressa
+-----+
| | +----- Pyganodon_grandis
| +--+
| | +-- Utterbackia_peninsularis
| +--+
| +-- Utterbackia_imbecillis
--+
| +----- Fusconaia_flava **
| +-----+
| | | +-- Pronodularia_japanensis
| | +--+
| | +-- Sinosolenaia_carinata
+--+
| +----- Toxolasma_parvum **
| +--+
| | | +-- Venustaconcha_ellipsiformis **
| | +--+
+--+ +-- Lampsilis_siliquoidea **
|
| +----- Quadrula_quadrula[2] **
+--+
| +-- Quadrula_quadrula[1] **
+--+
+-- Quadrula_nobilis **
Bolotov et al., 2018 [Publ 3826], fig. 2
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 7 |
13 |
0 |
4 |
78 |
78 |
3 |
24.0 |
+-- Ensidens_telus
|
+-- Ensidens_ingallsianus
|
+-- Ensidens_jaculus
|
+-- Hyriopsis_desowitzi
+--+
| +-- Hyriopsis_myersiana
| |
| +-- Hyriopsis_khoratensis
| |
| +-- Ensidens_spiculus
| |
| +-- Ensidens_sagittarius
|
+-----------------------+ +-- Lens_eximius
| | |
| | +-- Physunio_modelli[1]
| | |
| | +-- Yaukthwa_nesemanni
| | |
| | +-- Yaukthwa_panhai
| | |
| | +-- Lens_comptus
| | |
| +--+-- Lens_novoselovi
| |
| +-- Physunio_modelli[2]
| |
| +-- Physunio
| |
| +-- Yaukthwa_inlenensis
| |
| +-- Lens_contradens
| |
| +-- Lens_micropterus
|
| +-- Lamprotula_leaii
--+ |
| +-- Potomida_littoralis
| |
| +-- Pronodularia_japanensis
| |
| +-- Gonidea_angulata
| |
| +-- Leguminaia_wheatleyi
| |
| +-- Microcondylaea_bonellii
| |
| +-----------------------+-- Amblema_plicata **
| | |
| | +-- Ortmanniana_ligamentina **
| | |
| | +-- Lampsilis_cardium **
| | |
| | +-- Cambarunio_iris **
| | |
| | +-- Elliptio_complanata **
| | |
| | +-- Pleurobema_sintoxia **
| | |
+--+ +-- Quadrula_quadrula **
|
| +----- Unio_tumidus
| +--+
| | | +-- Unio_crassus
| | +--+
| +--------------+ +-- Unio_pictorum
| | |
| | | +----- Nodularia_douglasiae[1]
| | +--+
| | | +-- Nodularia_nuxpersicae
| | +--+
+--+ +-- Nodularia_douglasiae[2]
|
| +-- Lanceolaria_gladiola
| +-----------------+
| | +-- Lanceolaria_grayii
| |
| | +-------- Cristaria_plicata
+--+ +--------+
| | | +----- Pletholophus_tenuis
| | +--+
| | | +-- Sinanodonta_jourdyi
+--+ +--+
| +-- Sinanodonta_lauta
|
| +-------------- Anodonta_anatina
+--+
| +----------- Pyganodon_grandis
+--+
| +-------- Simpsonaias_ambigua
+--+
| +----- Strophitus_undulatus
+--+
| +-- Alasmidonta_marginata
+--+
+-- Lasmigona_costata
Bolotov et al., 2020 [Publ 4084], fig. 1
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 7 |
7 |
0 |
4 |
29 |
25 |
2 |
18.0 |
+-- Amblema_plicata **
|
+-- Ortmanniana_ligamentina **
|
+-- Lampsilis_cardium **
|
--+-- Cambarunio_iris **
|
+-- Elliptio_complanata **
|
+-- Pleurobema_sintoxia **
|
+-- Quadrula_quadrula **
Chen et al., 2020 [Publ 4170], fig. 1
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 7 |
16 |
14 |
2 |
27 |
26 |
2 |
27.0 |
+----- Prolasmidonta_heterodon
+--+
| | +-- Alasmidonta_varicosa
| +--+
| +-- Anemina_euscaphys
+-----------------+
| | +-- Lamprotula_caveata
| | +--+
| | | +-- Chamberlainia_somsakpanhai
| +--+
| | +-- Pseudodon_vondembuschianus
| +--+
| +-- Pilsbryoconcha_exilis
|
| +-------- Schistodesmus_lampreyanus
| +-----+
| | | +----- Lepidodesma_languilati
| | +--+
--+ | | +-- Aculamprotula_coreana
| | +--+
| +--------+ +-- Aculamprotula_tortuosa
| | |
| | | +----- Platynaias_compressa
| | | +-----+
| | | | | +-- Utterbackia_imbecillis
| | | | +--+
| | +--+ +-- Pyganodon_grandis
| | |
| | | +-------- Sinanodonta_lucida
| | +--+
| | | +----- Cristaria_plicata
| | +--+
+--+ | +-- Anodonta_cygnea
| +--+
| +-- Anodonta_anatina
|
| +----------------- Gibbosula_crassa
| +--+
| | | +-------------- Quadrula_quadrula **
| | +--+
| | | +----------- Toxolasma_parvum **
| | +--+
| | | +-------- Lampsilis_ornata **
| | +--+
| | | +----- Potamilus_leptodon **
+--+ +--+
| | +-- Lampsilis_siliquoidea **
| +--+
| +-- Lampsilis_powellii **
|
| +-- Utterbackia_peninsularis
| +--------------+
| | +-- Potamilus_alatus **
+--+
| +-------------- Lamprotula_leaii
| |
+--+ +-- Sinohyriopsis_cumingii
| +--------+
| | +-- Sinohyriopsis_schlegelii
+--+
| +-------- Parvasolenaia_rivularis
+--+
| +----- Sinosolenaia_carinata
+--+
| +-- Sinosolenaia_iridinea
+--+
+-- Sinosolenaia_oleivora
Pavan-Kumar et al., 2022 [Publ 4263], fig. 4
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 7 |
7 |
5 |
2 |
29 |
27 |
3 |
10.0 |
+-------------- Quadrula_quadrula **
|
+--------------+ +----------- Toxolasma_parvum **
| | |
| +--+ +-------- Potamilus_leptodon **
| | |
| +--+ +-- Lampsilis_cardium **
| | +--+
| | | +-- Lampsilis_ornata **
| +--+
| | +-- Lampsilis_siliquoidea **
| +--+
| +-- Venustaconcha_ellipsiformis **
|
| +----- Chamberlainia_somsakpanhai
--+ +-----------+
| | | +-- Sinohyriopsis_cumingii
| | +--+
| | +-- Sinohyriopsis_schlegelii
| +--------+
| | | +-- Lamprotula_leaii
| | | +-----------+
| | | | +-- Potomida_littoralis
| | | |
| | +--+ +-- Pseudodon_vondembuschianus
| | | +--------+
| | | | +-- Pilsbryoconcha_exilis
| | +--+
| | | +-------- Parvasolenaia_rivularis
+--+ +--+
| | +----- Microcondylaea_bonellii
| +--+
| | +-- Sinosolenaia_carinata
| +--+
| +-- Sinosolenaia_oleivora
|
| +----------------------- Lamellidens_marginalis
| |
| | +-------------------- Lepidodesma_languilati
+--+ |
| | +-- Aculamprotula_tientsinensis
| | +--------+
| | | +-- Aculamprotula_tortuosa
+--+ +-----+
| | | +-------- Unio_pictorum
| | +--+
| | | +----- Schistodesmus_lampreyanus
| | +--+
| | | +-- Nodularia_douglasiae
+--+ +--+
| +-- Tchangsinaia_piscicula
|
| +-- Lanceolaria_lanceolata
| +-----------+
| | +-- Lanceolaria_gladiola
+--+
| +----------- Acuticosta_chinensis
| |
+--+ +-- Anodonta_anatina
| +-----+
| | +-- Pyganodon_grandis
+--+
| +----- Cristaria_plicata
+--+
| +-- Sinanodonta_lucida
+--+
+-- Anemina_euscaphys
Pavan-Kumar et al., 2022 [Publ 4263], fig. S5
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 7 |
7 |
5 |
2 |
32 |
30 |
3 |
10.0 |
+-------------------- Lamellidens_marginalis
|
| +-- Anodonta_anatina
| +--------+
| | +-- Pyganodon_grandis
| +-----+
+--+ | | +-------- Cristaria_plicata
| | | | |
| | | +--+ +-- Anemina_arcaeformis
| | | | +--+
| | | | | +-- Anemina_euscaphys
| | | +--+
| | | | +-- Sinanodonta_lucida
| +--+ +--+
| | +-- Sinanodonta_woodiana
| |
| | +----- Acuticosta_chinensis
| | +--------+
| | | | +-- Lanceolaria_lanceolata
| | | +--+
| | | +-- Lanceolaria_gladiola
| +--+
| | +----- Lepidodesma_languilati
| | +-----+
| | | | +-- Aculamprotula_tientsinensis
| | | +--+
--+ +--+ +-- Aculamprotula_tortuosa
| |
| | +-------- Unio_pictorum
| +--+
| | +----- Schistodesmus_lampreyanus
| +--+
| | +-- Nodularia_douglasiae
| +--+
| +-- Tchangsinaia_piscicula
|
| +-------------- Quadrula_quadrula **
| |
| +--+ +----------- Toxolasma_parvum **
| | | |
| | +--+ +-------- Potamilus_leptodon **
| | | |
| | +--+ +-- Lampsilis_cardium **
| | | +--+
| | | | +-- Lampsilis_ornata **
| | +--+
| | | +-- Lampsilis_siliquoidea **
+-----+ +--+
| +-- Venustaconcha_ellipsiformis **
|
| +----- Chamberlainia_somsakpanhai
| +--------+
| | | +-- Sinohyriopsis_schlegelii
| | +--+
| | +-- Sinohyriopsis_cumingii
| |
| | +-- Sinosolenaia_carinata
+--+ +--+
| | +-- Sinosolenaia_oleivora
| +-----+
| | | +-- Parvasolenaia_rivularis
| | +--+
| | +-- Microcondylaea_bonellii
+--+
| +-- Pseudodon_vondembuschianus
| +-----+
| | +-- Pilsbryoconcha_exilis
+--+
| +----- Potomida_littoralis
+--+
| +-- Lamprotula_leaii
+--+
+-- Pronodularia_japanensis
Walker et al., 2006 [Publ 2666], fig. 2
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 6 |
7 |
5 |
2 |
9 |
6 |
2 |
21.0 |
+-------------------- Pronodularia_japanensis
|
| +-- Gonidea_angulata
--+ +--------------+
| | +-- Pseudodon_vondembuschianus
| |
+--+ +-- Fusconaia_flava **
| +-----------+
| | +-- Cumberlandia_monodonta
+--+
| +----------- Cyrtonaias_tampicoensis **
+--+
| +-------- Potamilus_purpuratus **
+--+
| +----- Ligumia_recta **
+--+
| +-- Ortmanniana_ligamentina **
+--+
+-- Lampsilis_teres **
Walker et al., 2006 [Publ 2666], fig. 3
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 6 |
6 |
4 |
2 |
9 |
6 |
2 |
20.0 |
+-------------------- Inversidens
|
| +----- Anodonta
--+ +-----------+
| | | +-- Pyganodon_fragilis
| | +--+
+--+ +-- Pyganodon_grandis
|
| +-------------- Fusconaia_flava **
+--+
| +----------- Potamilus_purpuratus **
+--+
| +-------- Cyrtonaias_tampicoensis **
+--+
| +----- Ligumia_recta **
+--+
| +-- Ortmanniana_ligamentina **
+--+
+-- Lampsilis_teres **
Walker et al., 2006 [Publ 2666], fig. 4
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 6 |
6 |
4 |
2 |
9 |
6 |
2 |
20.0 |
+-------------------- Inversidens
|
| +----- Anodonta
--+ +-----------+
| | | +-- Pyganodon_fragilis
| | +--+
+--+ +-- Pyganodon_grandis
|
| +-------------- Fusconaia_flava **
+--+
| +----------- Cyrtonaias_tampicoensis **
+--+
| +-------- Potamilus_purpuratus **
+--+
| +----- Ligumia_recta **
+--+
| +-- Ortmanniana_ligamentina **
+--+
+-- Lampsilis_teres **
Walker et al., 2006 [Publ 2666], fig. 6
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 6 |
6 |
3 |
2 |
32 |
4 |
2 |
20.0 |
+-- Gonidea_angulata
+-----------+
| +-- Sundadontina_tumida
--+
| +----------- Cyrtonaias_tampicoensis **
| |
+--+----------- Fusconaia_flava **
|
| +-------- Potamilus_purpuratus **
+--+
| +----- Lampsilis_teres **
+--+
| +-- Ligumia_recta **
+--+
+-- Ortmanniana_ligamentina **
Walker et al., 2006 [Publ 2666], fig. 6-M
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 6 |
6 |
4 |
2 |
9 |
4 |
2 |
20.0 |
+-- Gonidea_angulata
+-----------------+
| +-- Sundadontina_tumida
|
--+ +-- Pyganodon_fragilis
| +--------------+
| | +-- Pyganodon_grandis
+--+
| +-------------- Fusconaia_flava **
+--+
| +----------- Potamilus_purpuratus **
+--+
| +-------- Cyrtonaias_tampicoensis **
+--+
| +----- Ligumia_recta **
+--+
| +-- Lampsilis_teres **
+--+
+-- Ortmanniana_ligamentina **
Hoeh et al., 2002 [Publ 2673], fig. 1-F
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 6 |
6 |
4 |
2 |
3 |
2 |
1 |
20.0 |
+-- Pyganodon_fragilis
+--------------+
| +-- Pyganodon_grandis
--+
| +-------------- Fusconaia_flava **
+--+
| +----------- Cyrtonaias_tampicoensis **
+--+
| +-------- Ortmanniana_ligamentina **
+--+
| +----- Lampsilis_teres **
+--+
| +-- Potamilus_purpuratus **
+--+
+-- Ligumia_recta **
Hoeh et al., 2002 [Publ 2673], fig. 1-M
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 6 |
6 |
4 |
2 |
3 |
2 |
1 |
20.0 |
+-- Pyganodon_fragilis
+--------------+
| +-- Pyganodon_grandis
--+
| +-------------- Fusconaia_flava **
+--+
| +----------- Potamilus_purpuratus **
+--+
| +-------- Cyrtonaias_tampicoensis **
+--+
| +----- Ligumia_recta **
+--+
| +-- Lampsilis_teres **
+--+
+-- Ortmanniana_ligamentina **
Jones et al., 2006 [Publ 2681], fig. 2
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 6 |
6 |
4 |
1 |
0 |
0 |
0 |
21.0 |
+-- Epioblasma_triquetra **
+--------+
| +-- Epioblasma_brevidens **
--+
| +-------- Epioblasma_capsaeformis[2] **
+--+
| +----- Epioblasma_capsaeformis[1] **
+--+
| +-- Epioblasma_walkeri **
+--+
+-- Epioblasma_rangiana **
Lane et al., 2016 [Publ 3735], fig. 3
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 6 |
6 |
1 |
1 |
0 |
0 |
0 |
20.0 |
+----- Paetulunio_fabalis **
|
+----- Epioblasma_capsaeformis **
--+
| +-- Venustaconcha_trabalis[1] **
| |
| +-- Venustaconcha_trabalis[2] **
+--+
+-- Venustaconcha_ellipsiformis **
|
+-- Venustaconcha_pleasii **
Wen et al., 2017 [Publ 3795], fig. 3
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 6 |
6 |
4 |
2 |
10 |
8 |
2 |
11.0 |
+----------- Quadrula_quadrula **
|
+--+ +-------- Toxolasma_parvum **
| | |
| +--+ +-- Lampsilis_ornata **
| | +--+
| | | +-- Venustaconcha_ellipsiformis **
| +--+
--+ | +-- Potamilus_leptodon **
| +--+
| +-- Potamilus_alatus **
|
| +----------- Lamprotula_leaii
| |
| | +----- Anodonta_anatina
+--+ +--+
| | | +-- Sinanodonta_woodiana
| | +--+
| | +-- Cristaria_plicata
+--+
| +-- Aculamprotula_coreana
| +--+
| | +-- Aculamprotula_tortuosa
+--+
| +-- Lepidodesma_languilati
+--+
+-- Unio_pictorum
Huang et al., 2018 [Publ 3828], fig. 6
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 6 |
6 |
4 |
2 |
37 |
27 |
2 |
11.0 |
+----------- Quadrula_quadrula **
|
+-----+ +-------- Toxolasma_parvum **
| | |
| +--+ +-- Lampsilis_ornata **
| | +--+
| | | +-- Venustaconcha_ellipsiformis **
| +--+
| | +-- Potamilus_leptodon **
+--+ +--+
| | +-- Potamilus_alatus **
| |
| | +-- Sinohyriopsis_cumingii
| | +-----------+
| | | +-- Sinohyriopsis_schlegelii
| | |
| +--+ +-- Sinosolenaia_oleivora
| | +--------+
| | | +-- Sinosolenaia_carinata
| +--+
| | +-------- Potomida_littoralis
| +--+
| | +----- Pronodularia_japanensis
| +--+
| | +-- Lamprotula_caveata
--+ +--+
| +-- Lamprotula_leaii
|
| +----- Aculamprotula_tortuosa
| +--------+
| | | +-- Aculamprotula_coreana
| | +--+
| | +-- Aculamprotula_tientsinensis
| +--+
| | | +-- Tchangsinaia_piscicula
| | | +--------+
| | | | +-- Nodularia_douglasiae
| | +--+
| | | +-------- Unio_tumidus
| | +--+
| | | +----- Unio_crassus
| | +--+
+--+ | +-- Unio_delphinus
| +--+
| +-- Unio_pictorum
|
| +----------- Anodonta_anatina
| +--+
| | | +-------- Platynaias_compressa
| | +--+
| | | +----- Pyganodon_grandis
| | +--+
+--+ | +-- Utterbackia_imbecillis
| +--+
| +-- Utterbackia_peninsularis
|
| +-------- Cristaria_plicata
| |
+-----+ +-- Anemina_arcaeformis
| +--+
| | +-- Anemina_euscaphys
+--+
| +-- Sinanodonta_woodiana
+--+
+-- Sinanodonta_lucida
Wang et al., 2018 [Publ 3846], fig. 2
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 6 |
6 |
4 |
2 |
26 |
26 |
2 |
11.0 |
+----------- Quadrula_quadrula **
|
+--+ +-------- Toxolasma_parvum **
| | |
| +--+ +-- Venustaconcha_ellipsiformis **
| | +--+
| | | +-- Lampsilis_ornata **
| +--+
| | +-- Potamilus_leptodon **
+--------+ +--+
| | +-- Potamilus_alatus **
| |
| | +-- Sinohyriopsis_cumingii
| | +--------+
| | | +-- Sinohyriopsis_schlegelii
| | |
| +--+ +----- Potomida_littoralis
| | +--+
| | | | +-- Lamprotula_caveata
| | | +--+
| +--+ +-- Lamprotula_leaii
| |
| | +----- Ptychorhynchus_pfisteri
| +--+
--+ | +-- Sinosolenaia_carinata
| +--+
| +-- Sinosolenaia_oleivora
|
| +----- Anodonta_anatina
| +-----+
| | | +-- Pyganodon_grandis
| | +--+
| +--------+ +-- Platynaias_compressa
| | |
| | | +-------- Cristaria_plicata
| | | |
| | +--+ +-- Anemina_euscaphys
| | | +--+
| | | | +-- Anemina_arcaeformis
| | +--+
+--+ | +-- Sinanodonta_woodiana
| +--+
| +-- Sinanodonta_lucida
|
| +-- Lanceolaria_grayii
| +--------------+
| | +-- Lanceolaria_lanceolata
| |
+--+ +-------------- Aculamprotula_tientsinensis
| |
| | +----- Nodularia_douglasiae
+--+ +-----+
| | | +-- Tchangsinaia_piscicula
| | +--+
+--+ +-- Cuneopsis_heudei
|
| +-------- Unio_tumidus
+--+
| +----- Unio_crassus
+--+
| +-- Unio_pictorum
+--+
+-- Unio_delphinus
Wu et al., 2019 [Publ 3864], fig. 3
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 6 |
6 |
4 |
2 |
41 |
35 |
2 |
11.0 |
+----------- Quadrula_quadrula **
|
+-----------------+ +-------- Toxolasma_parvum **
| | |
| +--+ +-- Lampsilis_ornata **
| | +--+
| | | +-- Venustaconcha_ellipsiformis **
| +--+
| | +-- Potamilus_leptodon **
| +--+
| +-- Potamilus_alatus **
|
| +-- Sinohyriopsis_cumingii
| +--------+
--+ | +-- Sinohyriopsis_schlegelii
| |
| +--------------+ +----- Potomida_littoralis
| | | +--+
| | | | | +-- Lamprotula_leaii
| | | | +--+
| | +--+ +-- Pronodularia_japanensis
| | |
| | | +----- Ptychorhynchus_pfisteri
| | +--+
| | | +-- Sinosolenaia_oleivora
| | +--+
| | +-- Sinosolenaia_carinata
+--+
| +----------------------- Lepidodesma_languilati
| |
| | +-------- Aculamprotula_tortuosa
| | +--------+
| | | | +----- Aculamprotula_scripta
| | | +--+
| | | | +-- Aculamprotula_tientsinensis
| | | +--+
| | | +-- Aculamprotula_coreana
| | +--+
+--+ | | +-------- Unio_tumidus
| | | +-----+
| | | | | +----- Unio_crassus
| | | | +--+
| | | | | +-- Unio_delphinus
| | +--+ +--+
| | | +-- Unio_pictorum
| | |
| | | +----------- Pseudocuneopsis_capitata
| | +--+
| | | +-------- Nodularia_douglasiae
+--+ +--+
| | +----- Tchangsinaia_piscicula
| +--+
| | +-- Cuneopsis_heudei
| +--+
| +-- Schistodesmus_lampreyanus
|
| +----- Acuticosta_chinensis
| +-----------+
| | | +-- Lanceolaria_grayii
| | +--+
| | +-- Lanceolaria_lanceolata
| |
+--+ +----------- Anodonta_anatina
| +--+
| | | +-------- Platynaias_compressa
| | +--+
| | | +----- Pyganodon_grandis
| | +--+
+--+ | +-- Utterbackia_peninsularis
| +--+
| +-- Utterbackia_imbecillis
|
| +-------- Cristaria_plicata
| |
+-----+ +-- Anemina_arcaeformis
| +--+
| | +-- Anemina_euscaphys
+--+
| +-- Sinanodonta_lucida
+--+
+-- Sinanodonta_woodiana
Wang et al., 2018 [Publ 3865], fig. 2
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 6 |
6 |
4 |
2 |
26 |
26 |
2 |
11.0 |
+----------- Quadrula_quadrula **
|
+--+ +-------- Toxolasma_parvum **
| | |
| +--+ +-- Potamilus_leptodon **
| | +--+
| | | +-- Potamilus_alatus **
| +--+
| | +-- Venustaconcha_ellipsiformis **
+-----+ +--+
| | +-- Lampsilis_ornata **
| |
| | +-- Sinohyriopsis_schlegelii
| | +--------+
| | | +-- Sinohyriopsis_cumingii
| | |
| +--+ +----- Ptychorhynchus_pfisteri
| | +--+
| | | | +-- Sinosolenaia_oleivora
| | | +--+
| +--+ +-- Sinosolenaia_carinata
| |
| | +----- Potomida_littoralis
--+ +--+
| | +-- Lamprotula_caveata
| +--+
| +-- Lamprotula_leaii
|
| +-------------- Aculamprotula_tientsinensis
| |
| +--+ +-- Tchangsinaia_piscicula
| | | +--------+
| | | | +-- Nodularia_douglasiae
| | +--+
| | | +-------- Unio_tumidus
| | +--+
| | | +----- Unio_crassus
| | +--+
+--+ | +-- Unio_pictorum
| +--+
| +-- Unio_delphinus
|
| +----- Acuticosta_chinensis
| +--------+
| | | +-- Lanceolaria_grayii
| | +--+
| | +-- Lanceolaria_lanceolata
+--+
| +----- Anodonta_anatina
| +-----+
| | | +-- Pyganodon_grandis
| | +--+
+--+ +-- Platynaias_compressa
|
| +-------- Cristaria_plicata
| |
+--+ +-- Anemina_euscaphys
| +--+
| | +-- Anemina_arcaeformis
+--+
| +-- Sinanodonta_woodiana
+--+
+-- Sinanodonta_lucida
Froufe et al., 2019 [Publ 3936], fig. 3
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 6 |
6 |
4 |
2 |
39 |
36 |
2 |
11.0 |
+----------- Quadrula_quadrula **
|
+-----------------+ +-------- Toxolasma_parvum **
| | |
| +--+ +-- Lampsilis_ornata **
| | +--+
| | | +-- Venustaconcha_ellipsiformis **
| +--+
| | +-- Potamilus_leptodon **
| +--+
| +-- Potamilus_alatus **
|
| +----- Chamberlainia_somsakpanhai
| +-----------+
| | | +-- Sinohyriopsis_cumingii
| | +--+
| | +-- Sinohyriopsis_schlegelii
--+ |
| +--------+ +-------- Potomida_littoralis
| | | +-----+
| | | | | +----- Pronodularia_japanensis
| | | | +--+
| | | | | +-- Lamprotula_leaii
| | | | +--+
| | +--+ +-- Lamprotula_caveata
| | |
| | | +-- Pilsbryoconcha_exilis
| | | +--------+
| | | | +-- Pseudodon_vondembuschianus
| | +--+
| | | +-------- Ptychorhynchus_pfisteri
| | +--+
+--+ | +----- Microcondylaea_bonellii
| +--+
| | +-- Sinosolenaia_carinata
| +--+
| +-- Sinosolenaia_oleivora
|
| +----------------------- Lepidodesma_languilati
| |
| | +----- Aculamprotula_tortuosa
| | +--------+
| | | | +-- Aculamprotula_tientsinensis
| | | +--+
| | | +-- Aculamprotula_coreana
| | +-----+
+--+ | | +----- Nodularia_douglasiae
| | | +-----+
| | | | | +-- Tchangsinaia_piscicula
| | | | +--+
| | +--+ +-- Schistodesmus_lampreyanus
| | |
| | | +-------- Unio_tumidus
| | +--+
| | | +----- Unio_crassus
+--+ +--+
| | +-- Unio_pictorum
| +--+
| +-- Unio_delphinus
|
| +-- Lanceolaria_lanceolata
| +--------------+
| | +-- Lanceolaria_grayii
| |
| | +-------- Cristaria_plicata
+--+ |
| +-----+ +-- Anemina_arcaeformis
| | | +--+
| | | | +-- Anemina_euscaphys
| | +--+
+--+ | +-- Sinanodonta_lucida
| +--+
| +-- Sinanodonta_woodiana
|
| +----------- Anodonta_anatina
+--+
| +-------- Lasmigona_complanata
+--+
| +----- Pyganodon_grandis
+--+
| +-- Utterbackia_peninsularis
+--+
+-- Utterbackia_imbecillis
Froufe et al., 2019 [Publ 3936], fig. 6
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 6 |
6 |
4 |
2 |
39 |
36 |
2 |
11.0 |
+----------- Quadrula_quadrula **
|
+-----------------+ +-------- Toxolasma_parvum **
| | |
| +--+ +-- Potamilus_leptodon **
| | +--+
| | | +-- Potamilus_alatus **
| +--+
| | +-- Lampsilis_ornata **
| +--+
| +-- Venustaconcha_ellipsiformis **
|
| +----- Chamberlainia_somsakpanhai
| +-----------+
| | | +-- Sinohyriopsis_cumingii
| | +--+
| | +-- Sinohyriopsis_schlegelii
--+ |
| +--------+ +-------- Potomida_littoralis
| | | +-----+
| | | | | +----- Pronodularia_japanensis
| | | | +--+
| | | | | +-- Lamprotula_leaii
| | | | +--+
| | +--+ +-- Lamprotula_caveata
| | |
| | | +-- Pilsbryoconcha_exilis
| | | +--------+
| | | | +-- Pseudodon_vondembuschianus
| | +--+
| | | +-------- Ptychorhynchus_pfisteri
| | +--+
+--+ | +----- Microcondylaea_bonellii
| +--+
| | +-- Sinosolenaia_carinata
| +--+
| +-- Sinosolenaia_oleivora
|
| +----------------------- Lepidodesma_languilati
| |
| | +----- Aculamprotula_tortuosa
| | +--------+
| | | | +-- Aculamprotula_tientsinensis
| | | +--+
| | | +-- Aculamprotula_coreana
| | +-----+
+--+ | | +----- Nodularia_douglasiae
| | | +-----+
| | | | | +-- Tchangsinaia_piscicula
| | | | +--+
| | +--+ +-- Schistodesmus_lampreyanus
| | |
| | | +-------- Unio_tumidus
| | +--+
| | | +----- Unio_crassus
+--+ +--+
| | +-- Unio_pictorum
| +--+
| +-- Unio_delphinus
|
| +-- Lanceolaria_lanceolata
| +--------------+
| | +-- Lanceolaria_grayii
| |
| | +----------- Anodonta_anatina
+--+ +--+
| | | +-------- Lasmigona_complanata
| | +--+
| | | +----- Pyganodon_grandis
| | +--+
+--+ | +-- Utterbackia_peninsularis
| +--+
| +-- Utterbackia_imbecillis
|
| +-------- Cristaria_plicata
| |
+-----+ +-- Sinanodonta_lucida
| +--+
| | +-- Sinanodonta_woodiana
+--+
| +-- Anemina_arcaeformis
+--+
+-- Anemina_euscaphys
Wu et al., 2020 [Publ 4121], fig. 3
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 6 |
6 |
4 |
2 |
44 |
36 |
2 |
11.0 |
+----------- Quadrula_quadrula **
|
+-----------------+ +-------- Toxolasma_parvum **
| | |
| +--+ +-- Venustaconcha_ellipsiformis **
| | +--+
| | | +-- Lampsilis_ornata **
| +--+
| | +-- Potamilus_alatus **
| +--+
| +-- Potamilus_leptodon **
|
| +-- Sinohyriopsis_schlegelii
| +--------+
--+ | +-- Sinohyriopsis_cumingii
| |
| +--------------+ +----- Potomida_littoralis
| | | +--+
| | | | | +-- Pronodularia_japanensis
| | | | +--+
| | +--+ +-- Lamprotula_leaii
| | |
| | | +----- Ptychorhynchus_pfisteri
| | +--+
| | | +-- Sinosolenaia_oleivora
| | +--+
| | +-- Sinosolenaia_carinata
+--+
| +----------------------- Lepidodesma_languilati
| |
| | +----- Acuticosta_chinensis
| | +-----------+
| | | | +-- Lanceolaria_lanceolata
| | | +--+
| | | +-- Lanceolaria_grayii
| | |
| | +--+ +----------- Anodonta_anatina
| | | | +--+
| | | | | | +-------- Platynaias_compressa
+--+ | | | +--+
| | | | | +----- Pyganodon_grandis
| | | | +--+
| | +--+ | +-- Utterbackia_peninsularis
| | | +--+
| | | +-- Utterbackia_imbecillis
| | |
| | | +-------- Cristaria_plicata
| | | |
| | +-----+ +-- Sinanodonta_woodiana
| | | +--+
+--+ | | +-- Sinanodonta_lauta
| +--+
| | +-- Anemina_arcaeformis
| +--+
| +-- Anemina_euscaphys
|
| +-------- Aculamprotula_tortuosa
| |
| +--------+ +-- Aculamprotula_coreana
| | | +--+
| | | | +-- Aculamprotula_tientsinensis
| | +--+
| | | +-- Aculamprotula_polysticta
| | +--+
+--+ +-- Aculamprotula_scripta
|
| +-------- Unio_tumidus
| +-----+
| | | +----- Unio_crassus
| | +--+
| | | +-- Unio_delphinus
+--+ +--+
| +-- Unio_pictorum
|
| +----------- Pseudocuneopsis_capitata
+--+
| +-------- Nodularia_douglasiae
+--+
| +----- Tchangsinaia_piscicula
+--+
| +-- Cuneopsis_heudei
+--+
+-- Schistodesmus_lampreyanus
Zieritz et al., 2021 [Publ 4231], fig. 3-F, 5
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 6 |
6 |
4 |
2 |
43 |
40 |
2 |
11.0 |
+----------- Quadrula_quadrula **
|
+-----------------+ +-------- Toxolasma_parvum **
| | |
| +--+ +-- Potamilus_alatus **
| | +--+
| | | +-- Potamilus_leptodon **
| +--+
| | +-- Venustaconcha_ellipsiformis **
| +--+
| +-- Lampsilis_ornata **
|
| +----- Chamberlainia_somsakpanhai
| +--------------+
| | | +-- Sinohyriopsis_cumingii
| | +--+
| | +-- Sinohyriopsis_schlegelii
| |
| | +-- Lens_contradens
--+ +-----+ +--+
| | | | +-- Physunio_superbus
| | | +-----------+
| | | | | +-- Hyriopsis_bialata
| | | | +--+
| | | | +-- Rectidens_sumatrensis
| | | |
| | +--+ +-------- Potomida_littoralis
| | | +-----+
| | | | | +----- Pronodularia_japanensis
| | | | +--+
| | | | | +-- Lamprotula_caveata
| | | | +--+
| | +--+ +-- Lamprotula_leaii
| | |
| | | +-- Pilsbryoconcha_exilis
| | | +--------+
+--+ | | +-- Pseudodon_vondembuschianus
| +--+
| | +-------- Ptychorhynchus_pfisteri
| +--+
| | +----- Microcondylaea_bonellii
| +--+
| | +-- Sinosolenaia_recognita
| +--+
| +-- Sinosolenaia_carinata
|
| +----------------------- Lepidodesma_languilati
| |
| | +----- Aculamprotula_tortuosa
| | +--------+
| | | | +-- Aculamprotula_tientsinensis
| | | +--+
| | | +-- Aculamprotula_coreana
| | +-----+
+--+ | | +----- Nodularia_douglasiae
| | | +-----+
| | | | | +-- Schistodesmus_lampreyanus
| | | | +--+
| | +--+ +-- Tchangsinaia_piscicula
| | |
| | | +-------- Unio_tumidus
| | +--+
| | | +----- Unio_crassus
+--+ +--+
| | +-- Unio_pictorum
| +--+
| +-- Unio_delphinus
|
| +-- Lanceolaria_grayii
| +--------------+
| | +-- Lanceolaria_lanceolata
| |
| | +----------- Anodonta_anatina
+--+ +--+
| | | +-------- Platynaias_compressa
| | +--+
| | | +----- Pyganodon_grandis
| | +--+
+--+ | +-- Utterbackia_imbecillis
| +--+
| +-- Utterbackia_peninsularis
|
| +-------- Cristaria_plicata
| |
+-----+ +-- Sinanodonta_lucida
| +--+
| | +-- Sinanodonta_woodiana
+--+
| +-- Anemina_arcaeformis
+--+
+-- Anemina_euscaphys
Mejia-Trujillo & Smith, 2022 [Publ 4274], fig. 2-M
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 6 |
6 |
4 |
2 |
0 |
0 |
0 |
11.0 |
+----------- Quadrula_quadrula **
|
--+ +-- Potamilus_streckersoni **
| +-----+
| | +-- Potamilus_alatus **
+--+
| +----- Venustaconcha_ellipsiformis **
+--+
| +-- Lampsilis_powellii **
+--+
+-- Lampsilis_siliquoidea **
Wu et al., 2023 [Publ 4339], fig. 5
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 6 |
6 |
4 |
3 |
13 |
11 |
1 |
17.0 |
+-- Elliptio_complanata **
+--------+
| +-- Quadrula_quadrula **
+-----+
| | +-------- Obliquaria_reflexa **
| +--+
| | +----- Truncilla_truncata **
| +--+
| | +-- Lampsilis_cardium **
| +--+
--+ +-- Potamilus_alatus **
|
| +-- Lepidodesma_languilati
| +-----------+
| | +-- Lepidodesma_aligera
| |
| | +-------- Anemina_arcaeformis
+--+ +--+
| | | +----- Cristaria_plicata
| | +--+
| | | +-- Sinanodonta_tumens
| | +--+
+--+ +-- Sinanodonta_woodiana
|
| +-- Platynaias
| +-----+
| | +-- Platynaias_compressa
+--+
| +----- Anodonta_anatina
+--+
| +-- Pseudanodonta_complanata
+--+
+-- Anodonta_cygnea
Manendo et al., 2008 [Publ 2669], fig. 1
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 5 |
5 |
3 |
3 |
8 |
8 |
1 |
19.0 |
+-- Lampsilis_siliquoidea **
+-----+
| +-- Sagittunio_nasutus **
+-----+
| | +----- Amblema_plicata **
| +--+
| | +-- Pleuronaia_dolabelloides **
| +--+
| +-- Fusconaia_flava **
--+
| +----- Unio_crassus
| +-----+
| | | +-- Unio_pictorum
| | +--+
| | +-- Unio_tumidus
+--+
| +-- Anodonta_cygnea
| +-----+
| | +-- Anodonta_anatina
+--+
| +----- Pseudanodonta_complanata
+--+
| +-- Platynaias_subviridis
+--+
+-- Pyganodon_grandis
Whelan et al., 2011 [Publ 3380], fig. 1 ML
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 5 |
5 |
3 |
4 |
32 |
20 |
3 |
17.0 |
+----------------- Gonidea_angulata
|
| +----- Pronodularia_japanensis
--+ +--------+
| | | +-- Potomida_littoralis
| | +--+
+--+ +-- Potomida_semirugata
|
| +----------- Pleurobema_sintoxia **
+--+
| +-------- Quadrula_quadrula **
+--+
| +----- Amblema_plicata **
+--+
| +-- Truncilla_truncata **
+--+
+-- Lampsilis_cardium **
Whelan et al., 2011 [Publ 3380], fig. 1 MP
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 5 |
5 |
2 |
4 |
32 |
20 |
3 |
16.0 |
+-------------- Gonidea_angulata
|
| +----- Pronodularia_japanensis
--+ +-----+
| | | +-- Potomida_littoralis
| | +--+
+--+ +-- Potomida_semirugata
|
| +-------- Quadrula_quadrula **
| |
+--+-------- Pleurobema_sintoxia **
|
| +----- Amblema_plicata **
+--+
| +-- Lampsilis_cardium **
+--+
+-- Truncilla_truncata **
Krebs et al., 2013 [Publ 3515], fig. 2A
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 5 |
5 |
3 |
1 |
0 |
0 |
0 |
21.0 |
+-------- Lampsilis_radiata **
|
--+ +-- Lampsilis_ovata **
| +--+
| | +-- Ortmanniana_ligamentina **
+--+
| +-- Lampsilis_hydiana **
+--+
+-- Lampsilis_straminea **
Pfeiffer & Graf, 2013 [Publ 3531], fig. 2
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 5 |
5 |
3 |
4 |
30 |
27 |
3 |
15.0 |
+----------- Quadrula_quadrula **
--+
| +-------- Eurynia_dilatata **
+--+
| +----- Amblema_plicata **
+--+
| +-- Lampsilis_ornata **
+--+
+-- Truncilla_truncata **
Pfeiffer & Graf, 2015 [Publ 3620], fig. 2
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 5 |
5 |
3 |
3 |
35 |
25 |
3 |
14.0 |
+----- Unio_pictorum
+-----------------+
| | +-- Pyganodon_grandis
| +--+
| +-- Lasmigona_costata
|
| +----------- Quadrula_quadrula **
| +--------+
--+ | | +-------- Amblema_plicata **
| | +--+
| | | +----- Potamilus_alatus **
| | +--+
| | | +-- Lampsilis_cardium **
| | +--+
| | +-- Truncilla_truncata **
+--+
| +-------- Ensidens_telus
| +--------+
| | | +----- Lens_micropterus
| | +--+
| | | +-- Physunio
| | +--+
| | +-- Physunio_modelli
+--+
| +-------- Gonidea_angulata
| +-----+
| | | +----- Potomida_littoralis
| | +--+
| | | +-- Pronodularia_japanensis
+--+ +--+
| +-- Lamprotula_caveata
|
| +----------- Pilsbryoconcha_carinifera
| |
+--+ +-- Pseudodon_vondembuschianus
| +-----+
| | +-- Pseudodon_vagulus
+--+
| +----- Sundadontina_tumida
+--+
| +-- Nyeinchanconcha_nyeinchani
+--+
+-- Bineurus_exilis
Plazzi et al., 2011 [Publ 3676], fig. 5
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 5 |
5 |
2 |
3 |
13 |
12 |
2 |
16.0 |
+-------- Quadrula_quadrula **
|
--+-------- Parvaspina_collina **
|
| +----- Lampsilis_ornata **
+--+
| +-- Epioblasma_rangiana **
+--+
+-- Venustaconcha_ellipsiformis **
Cha et al., 2018 [Publ 3916], fig. 1
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 5 |
5 |
3 |
2 |
34 |
31 |
2 |
12.0 |
+----------- Quadrula_quadrula **
+-----------------+
| | +-------- Toxolasma_parvum **
| +--+
| | +----- Lampsilis_ornata **
| +--+
| | +-- Potamilus_alatus **
| +--+
| +-- Potamilus_leptodon **
|
| +-- Sinohyriopsis_cumingii
| +-----------+
| | +-- Sinohyriopsis_schlegelii
--+ |
| +-----------+ +----- Ptychorhynchus_pfisteri
| | | +-----+
| | | | | +-- Sinosolenaia_carinata
| | | | +--+
| | +--+ +-- Sinosolenaia_oleivora
| | |
| | | +-------- Potomida_littoralis
| | +--+
| | | +----- Pronodularia_japanensis
| | +--+
| | | +-- Lamprotula_caveata
+--+ +--+
| +-- Lamprotula_leaii
|
| +----------------------- Lepidodesma_languilati
| |
| | +----- Aculamprotula_tortuosa
| | +--------+
| | | | +-- Aculamprotula_coreana
| | | +--+
| | | +-- Aculamprotula_tientsinensis
| | +-----+
+--+ | | +----- Schistodesmus_lampreyanus
| | | +-----+
| | | | | +-- Nodularia_douglasiae
| | | | +--+
| | +--+ +-- Tchangsinaia_piscicula
| | |
| | | +-------- Unio_tumidus
| | +--+
+--+ | +----- Unio_crassus
| +--+
| | +-- Unio_pictorum
| +--+
| +-- Unio_delphinus
|
| +----------------- Lanceolaria_grayii
| |
| | +----------- Anodonta_anatina
| | +--+
+--+ | | +-------- Platynaias_compressa
| | +--+
| | | +----- Pyganodon_grandis
| | +--+
+--+ | +-- Utterbackia_peninsularis
| +--+
| +-- Utterbackia_imbecillis
|
| +-------- Cristaria_plicata
| |
+-----+ +-- Anemina_arcaeformis
| +--+
| | +-- Anemina_euscaphys
+--+
| +-- Sinanodonta_lucida
+--+
+-- Sinanodonta_woodiana
Saxon et al., 2023 [Publ 4338], fig. 4
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 5 |
5 |
3 |
1 |
0 |
0 |
0 |
21.0 |
+----------- Ellipsaria_lineolata **
--+
| +-------- Truncilla_truncata **
+--+
| +----- Truncilla_macrodon **
+--+
| +-- Truncilla_cognata **
+--+
+-- Truncilla_donaciformis **
Graf, 2002 [Publ 2651], fig. 2
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 4 |
4 |
2 |
3 |
14 |
8 |
2 |
18.0 |
+-- Unio_pictorum
+--+
| +-- Unio_caffer
+-----+
| | +-- Alasmidonta_marginata
| +--+
--+ +-- Pyganodon_grandis
|
| +-------- Gonidea_angulata
| |
+--+ +-- Lampsilis_cardium **
| +--+
| | +-- Obliquaria_reflexa **
+--+
| +-- Eurynia_dilatata **
+--+
+-- Amblema_plicata **
Moyer & Díaz-Ferguson, 2012 [Publ 3375], fig. 3
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 4 |
4 |
2 |
1 |
3 |
3 |
1 |
19.0 |
+----- Lasmigona_costata
+-----+
| | +-- Platynaias_compressa
| +--+
--+ +-- Platynaias_subviridis
|
| +-------- Lampsilis_ovata[2] **
+--+
| +----- Lampsilis_ovata[1] **
+--+
| +-- Lampsilis_virescens **
+--+
+-- Lampsilis_fasciola **
Mulvey et al., 1997 [Publ 3390], fig. 1, 3
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 4 |
4 |
2 |
2 |
1 |
1 |
1 |
18.0 |
+----------- Utterbackia_imbecillis
--+
| +-------- Megalonaias_nervosa **
+--+
| +----- Amblema_plicata **
+--+
| +-- Amblema_neislerii **
+--+
+-- Amblema_elliottii **
Zanatta & Harris, 2013 [Publ 3530], fig. 2
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 4 |
4 |
2 |
1 |
1 |
1 |
1 |
17.0 |
+----------- Pyganodon_grandis
--+
| +-------- Lampsilis_cardium **
+--+
| +----- Epioblasma_rangiana **
+--+
| +-- Venustaconcha_pleasii **
+--+
+-- Venustaconcha_ellipsiformis **
Gangloff et al., 2013 [Publ 3551], fig. 2
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 4 |
4 |
2 |
1 |
0 |
0 |
0 |
19.0 |
+-- Ligumia_recta **
+--+
| +-- Sagittunio_nasutus **
--+
| +-- Venustaconcha_ellipsiformis **
+--+
+-- Venustaconcha_pleasii **
Huang et al., 2013 [Publ 3629], fig. 3
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 4 |
4 |
2 |
2 |
12 |
11 |
2 |
13.0 |
+-------------- Unio_pictorum
|
+--+ +-- Sinanodonta_woodiana
| | +--------+
| | | +-- Cristaria_plicata
| +--+
| | +-------- Platynaias_compressa
| +--+
| | +----- Pyganodon_grandis
| +--+
| | +-- Utterbackia_imbecillis
--+ +--+
| +-- Utterbackia_peninsularis
|
| +-------- Quadrula_quadrula **
| +--+
| | | +----- Toxolasma_parvum **
| | +--+
| | | +-- Lampsilis_ornata **
| | +--+
+-----+ +-- Venustaconcha_ellipsiformis **
|
| +-- Sinosolenaia_carinata
| +--+
| | +-- Pronodularia_japanensis
+-----+
| +-- Sinohyriopsis_cumingii
+--+
+-- Sinohyriopsis_schlegelii
Inoue et al., 2014 [Publ 3635], fig. 3A
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 4 |
20 |
16 |
4 |
17 |
16 |
2 |
31.0 |
+-------------------------- Amblema_plicata **
|
+-------------------------- Eurynia_dilatata **
|
--+-------------------------- Lampsilis_ornata **
|
| +----------------------- Quadrula_quadrula **
| |
| | +-------------------- Gonidea_angulata
+--+ |
| | +----------- Pseudodontoideus_subvexus
| | +-----+
+--+ | | +-------- Lasmigona_costata
| | +--+
| | | +----- Lasmigona_complanata
| | +--+
| | | +-- Arcidens_confragosus
+--+ +--+
| +-- Arcidens_wheeleri
|
| +-------------- Strophitus_undulatus
| |
| | +-------- Pyganodon_grandis
| | +--+
+--+ | | +----- Utterbackia_peninsularis[2]
| | +--+
| | | +-- Utterbackia_imbecillis
| | +--+
+--+ +-- Utterbackia_peninsularis[1]
|
| +-- Platynaias_compressa
| +-----+
| | +-- Platynaias_subviridis
+--+
| +----- Alasmidonta_marginata
+--+
| +-- Alasmidonta_undulata
+--+
+-- Alasmidonta_varicosa
Zhou et al., 2016 [Publ 3710], fig. 1
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 4 |
4 |
2 |
2 |
16 |
15 |
2 |
13.0 |
+-------- Quadrula_quadrula **
+--+
| | +----- Toxolasma_parvum **
| +--+
| | +-- Venustaconcha_ellipsiformis **
| +--+
--+ +-- Lampsilis_ornata **
|
| +-- Sinohyriopsis_cumingii
| +-----+
| | +-- Sinohyriopsis_schlegelii
| |
+--+ +-- Lamprotula_leaii
| +--+
| | +-- Pronodularia_japanensis
+--+
| +-- Sinosolenaia_carinata
+--+
+-- Sinosolenaia_oleivora
Guerra et al., 2017 [Publ 3772], fig. 2a
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 4 |
4 |
2 |
2 |
21 |
15 |
2 |
13.0 |
+-------- Quadrula_quadrula **
+--+
| | +----- Toxolasma_parvum **
| +--+
| | +-- Venustaconcha_ellipsiformis **
| +--+
+-----------+ +-- Lampsilis_ornata **
| |
| | +-- Sinohyriopsis_schlegelii
| | +-----+
| | | +-- Sinohyriopsis_cumingii
| +--+
| | +----- Pronodularia_japanensis
--+ +--+
| | +-- Sinosolenaia_oleivora
| +--+
| +-- Sinosolenaia_carinata
|
| +-- Aculamprotula_tortuosa
| +-----------------+
| | +-- Unio_pictorum
+--+
| +----------------- Sinanodonta_woodiana
| |
+--+ +-------------- Cristaria_plicata
| |
+--+ +----------- Anodonta_anatina
| |
+--+ +-- Platynaias_subviridis
| +-----+
| | +-- Platynaias_compressa
+--+
| +----- Pyganodon_grandis
+--+
| +-- Utterbackia_peninsularis
+--+
+-- Utterbackia_imbecillis
Guerra et al., 2017 [Publ 3772], fig. 2b
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 4 |
4 |
2 |
2 |
21 |
15 |
2 |
13.0 |
+-------- Quadrula_quadrula **
+-----------------+
| | +----- Toxolasma_parvum **
| +--+
| | +-- Venustaconcha_ellipsiformis **
| +--+
| +-- Lampsilis_ornata **
|
--+ +-- Sinohyriopsis_schlegelii
| +-----+
| | +-- Sinohyriopsis_cumingii
| +--------------+
| | | +----- Pronodularia_japanensis
| | +--+
| | | +-- Sinosolenaia_oleivora
+--+ +--+
| +-- Sinosolenaia_carinata
|
| +-------------------- Unio_pictorum
| |
+--+ +----------------- Aculamprotula_tortuosa
| |
+--+ +-- Cristaria_plicata
| +-----------+
| | +-- Sinanodonta_woodiana
+--+
| +----------- Anodonta_anatina
| |
+--+ +-- Platynaias_subviridis
| +-----+
| | +-- Platynaias_compressa
+--+
| +----- Pyganodon_grandis
+--+
| +-- Utterbackia_peninsularis
+--+
+-- Utterbackia_imbecillis
Fonseca et al., 2016 [Publ 3775], fig. 1
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 4 |
4 |
2 |
2 |
28 |
26 |
2 |
13.0 |
+-------- Quadrula_quadrula **
+--------------+
| | +----- Toxolasma_parvum **
| +--+
| | +-- Lampsilis_ornata **
| +--+
| +-- Venustaconcha_ellipsiformis **
|
| +-- Sinohyriopsis_cumingii
| +--------+
| | +-- Sinohyriopsis_schlegelii
| |
--+ +--------+ +-- Sinosolenaia_oleivora
| | | +-----+
| | | | +-- Sinosolenaia_carinata
| | +--+
| | | +----- Potomida_littoralis
| | +--+
| | | +-- Lamprotula_leaii
| | +--+
| | +-- Pronodularia_japanensis
| |
| | +-- Lanceolaria_lanceolata
| | +-----------+
+--+ | +-- Lanceolaria_grayii
| |
| +--+ +-- Aculamprotula_tortuosa
| | | +--------+
| | | | +-- Aculamprotula_coreana
| | | |
| | +--+ +-- Unio_pictorum
| | | +-----+
| | | | +-- Unio_delphinus
| | +--+
| | | +----- Schistodesmus_lampreyanus
| | +--+
+--+ | +-- Nodularia_douglasiae
| +--+
| +-- Tchangsinaia_piscicula
|
| +-------- Cristaria_plicata
| |
| +-----+ +-- Anemina_euscaphys
| | | +--+
| | | | +-- Anemina_arcaeformis
| | +--+
+--+ | +-- Sinanodonta_woodiana
| +--+
| +-- Sinanodonta_lucida
|
| +----------- Anodonta_anatina
+--+
| +-------- Platynaias_compressa
+--+
| +----- Pyganodon_grandis
+--+
| +-- Utterbackia_peninsularis
+--+
+-- Utterbackia_imbecillis
Amaro et al., 2017 [Publ 3791], fig. 2-M
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 4 |
4 |
2 |
3 |
6 |
6 |
2 |
16.0 |
+-- Fusconaia_flava **
+--+
| +-- Quadrula_quadrula **
+-----+
| | +-- Lampsilis_siliquoidea **
| +--+
| +-- Venustaconcha_ellipsiformis **
--+
| +-- Pronodularia_japanensis
| +-----+
| | +-- Sinosolenaia_carinata
+--+
| +----- Pyganodon_grandis
| |
+--+ +-- Lasmigona_costata
| |
+--+-- Sinanodonta_woodiana
|
+-- Utterbackia_peninsularis
Lopes-Lima et al., 2017 [Publ 3792], fig. 2
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 4 |
4 |
2 |
2 |
29 |
26 |
2 |
13.0 |
+-------- Quadrula_quadrula **
+--------------+
| | +----- Toxolasma_parvum **
| +--+
| | +-- Lampsilis_ornata **
| +--+
| +-- Venustaconcha_ellipsiformis **
|
| +-- Sinohyriopsis_cumingii
| +--------+
| | +-- Sinohyriopsis_schlegelii
| |
--+ +--------+ +-- Sinosolenaia_oleivora
| | | +-----+
| | | | +-- Sinosolenaia_carinata
| | +--+
| | | +----- Potomida_littoralis
| | +--+
| | | +-- Pronodularia_japanensis
| | +--+
| | +-- Lamprotula_leaii
| |
| | +-- Lanceolaria_lanceolata
| | +-----------+
+--+ | +-- Lanceolaria_grayii
| |
| +--+ +-- Aculamprotula_tortuosa
| | | +--------+
| | | | +-- Aculamprotula_coreana
| | | |
| | +--+ +-- Unio_delphinus
| | | +-----+
| | | | +-- Unio_pictorum
| | +--+
| | | +----- Schistodesmus_lampreyanus
| | +--+
+--+ | +-- Tchangsinaia_piscicula
| +--+
| +-- Nodularia_douglasiae
|
| +----------- Anodonta_anatina
| +--+
| | | +-------- Platynaias_compressa
| | +--+
| | | +----- Pyganodon_grandis
| | +--+
+--+ | +-- Utterbackia_peninsularis
| +--+
| +-- Utterbackia_imbecillis
|
| +-------- Cristaria_plicata
| |
+-----+ +-- Sinanodonta_lucida
| +--+
| | +-- Sinanodonta_woodiana
+--+
| +-- Anemina_arcaeformis
+--+
+-- Anemina_euscaphys
Pfeiffer et al., 2018 [Publ 3835], fig. 2
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 4 |
4 |
2 |
1 |
0 |
0 |
0 |
18.0 |
+-------- Quadrula_quadrula **
--+
| +----- Uniomerus_declivis **
+--+
| +-- Megalonaias_nervosa **
+--+
+-- Megalonaias_nickliniana **
Aunins et al., 2018 [Publ 3863], fig. 1
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 4 |
4 |
2 |
2 |
37 |
34 |
2 |
13.0 |
+-------- Quadrula_quadrula **
+--------------------+
| | +----- Toxolasma_parvum **
| +--+
| | +-- Lampsilis_ornata **
| +--+
| +-- Venustaconcha_ellipsiformis **
|
| +-- Sinohyriopsis_cumingii
| +--------+
| | +-- Sinohyriopsis_schlegelii
--+ |
| +--------------+ +-- Lamprotula_caveata
| | | +-----+
| | | | +-- Potomida_littoralis
| | +--+
| | | +----- Sinosolenaia_oleivora
| | +--+
| | | +-- Ptychorhynchus_pfisteri
| | +--+
| | +-- Sinosolenaia_carinata
+--+
| +----------------------- Lepidodesma_languilati
| |
| | +----- Aculamprotula_tortuosa
| | +--------+
| | | | +-- Aculamprotula_coreana
| | | +--+
| | | +-- Aculamprotula_tientsinensis
| | |
| | +-----+ +-------- Nodularia_douglasiae
+--+ | | +--+
| | | | | +----- Tchangsinaia_piscicula
| | | | +--+
| | | | | +-- Cuneopsis_heudei
| | +--+ +--+
| | | +-- Schistodesmus_lampreyanus
| | |
| | | +-------- Unio_tumidus
| | +--+
| | | +----- Unio_crassus
+--+ +--+
| | +-- Unio_pictorum
| +--+
| +-- Unio_delphinus
|
| +----- Acuticosta_chinensis
| +-----------+
| | | +-- Lanceolaria_grayii
| | +--+
| | +-- Lanceolaria_lanceolata
| |
| | +-------- Cristaria_plicata
+--+ |
| +-----+ +-- Anemina_arcaeformis
| | | +--+
| | | | +-- Anemina_euscaphys
| | +--+
| | | +-- Sinanodonta_woodiana
| | +--+
+--+ +-- Sinanodonta_lucida
|
| +-- Anodonta_anatina
| +--------+
| | +-- Anodonta_cygnea
| |
+--+ +-- Prolasmidonta_heterodon
| +-----+
| | +-- Alasmidonta_varicosa
+--+
| +----- Pyganodon_grandis
+--+
| +-- Utterbackia_imbecillis
+--+
+-- Utterbackia_peninsularis
Lopes-Lima et al., 2020 [Publ 4080], fig. 2-4
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 4 |
4 |
0 |
4 |
118 |
107 |
3 |
15.0 |
+-- Lens_contradens
|
+-- Ensidens_sagittarius
|
+-----------------------+-- Physunio_modelli
| |
| +-- Rectidens_sumatrensis
| |
| +-- Yaukthwa_nesemanni
|
| +-- Amblema_plicata **
| |
| +-- Elliptio_complanata **
| +--------------------+
--+ | +-- Lampsilis_cardium **
| | |
| | +-- Quadrula_quadrula **
| |
| | +----- Chamberlainia_somsakpanhai
| | +-----------+
| | | | +-- Sinohyriopsis_cumingii
| | | +--+
| | +--+ +-- Sinohyriopsis_schlegelii
| | | |
| | | | +-------------- Pronodularia_japanensis
+--+ | | |
| | +--+ +----------- Potomida_littoralis
| | | |
| | +--+ +-------- Lamprotula_caveata
| | | |
| | +--+ +----- Lamprotula_leaii[2]
| | | |
| | +--+----- Lamprotula_leaii[3]
| | |
| | | +-- Lamprotula_seomjinensis
| | +--+
| | +-- Lamprotula_leaii[1]
+--+
| +-- Indopseudodon_indawgyiensis
| +--------+
| | +-- Indopseudodon_salwenianus
| |
| +-----+ +-- Pilsbryoconcha_exilis
| | | +-----+
| | | | +-- Pilsbryoconcha_carinifera
| | +--+
| | | +----- Pseudodon_mekongi
| | | |
| | +--+ +-- Nyeinchanconcha_nyeinchani
| | | |
| | +--+-- Bineurus_mouhotii
| | |
+--+ +-- Sundadontina_tumida
|
| +-------- Inversidens_brandtii
| |
| +-----+ +-- Obovalis_omiensis
| | | +--+
| | | | +-- Ptychorhynchus_pfisteri
| | +--+
| | | +-- Parvasolenaia_rivularis
| | +--+
+--+ +-- Koreosolenaia_sitgyensis
|
| +----- Gonidea_angulata
| +-----+
| | | +-- Leguminaia_wheatleyi
| | +--+
+--+ +-- Microcondylaea_bonellii
|
| +-------- Sinosolenaia_carinata
+--+
| +----- Sinosolenaia_recognita
+--+
| +-- Sinosolenaia_iridinea
+--+
+-- Sinosolenaia_oleivora
Guerra et al., 2019 [Publ 4168], fig. 1
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 4 |
4 |
2 |
2 |
37 |
17 |
2 |
13.0 |
+-------- Quadrula_quadrula **
+--------------+
| | +----- Toxolasma_parvum **
| +--+
| | +-- Venustaconcha_ellipsiformis **
| +--+
| +-- Potamilus_alatus **
|
--+ +-------- Sinosolenaia_carinata
| +-----------+
| | | +----- Pronodularia_japanensis
| | +--+
| | | +-- Lamprotula_leaii
| | +--+
| | +-- Potomida_littoralis
| |
+--+ +-------------- Lanceolaria_lanceolata
| +--+
| | | +----------- Aculamprotula_tortuosa
| | +--+
| | | +-------- Schistodesmus_lampreyanus
| | +--+
| | | +----- Unio_tumidus
+--+ +--+
| | +-- Unio_crassus
| +--+
| +-- Unio_delphinus
|
| +-------------- Sinanodonta_woodiana
| |
+--+ +----------- Anodonta_anatina
| |
+--+ +-- Platynaias_compressa
| +-----+
| | +-- Platynaias_subviridis
+--+
| +----- Pyganodon_grandis
+--+
| +-- Utterbackia_peninsularis
+--+
+-- Utterbackia_imbecillis
Guerra et al., 2019 [Publ 4168], fig. 2
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 4 |
4 |
2 |
2 |
37 |
17 |
2 |
13.0 |
+-------- Quadrula_quadrula **
+--------------------+
| | +----- Toxolasma_parvum **
| +--+
| | +-- Venustaconcha_ellipsiformis **
| +--+
| +-- Potamilus_alatus **
--+
| +-------- Sinosolenaia_carinata
| +-----------------+
| | | +----- Pronodularia_japanensis
| | +--+
| | | +-- Lamprotula_leaii
| | +--+
+--+ +-- Potomida_littoralis
|
| +----------------------- Aculamprotula_tortuosa
| |
| | +-------- Schistodesmus_lampreyanus
+--+ +-----------+
| | | +----- Unio_tumidus
| | +--+
| | | +-- Unio_crassus
+--+ +--+
| +-- Unio_delphinus
|
| +----------------- Lanceolaria_lanceolata
| |
+--+ +-------------- Sinanodonta_woodiana
| |
+--+ +----------- Anodonta_anatina
| |
+--+ +-- Platynaias_compressa
| +-----+
| | +-- Platynaias_subviridis
+--+
| +----- Pyganodon_grandis
+--+
| +-- Utterbackia_peninsularis
+--+
+-- Utterbackia_imbecillis
Patterson et al., 2021 [Publ 4222], fig. 3
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 4 |
4 |
2 |
2 |
0 |
0 |
0 |
18.0 |
+-------- Villosa_villosa **
--+
| +----- Amblema_neislerii **
+--+
| +-- Amblema_plicata **
+--+
+-- Amblema_elliottii **
Johnson et al., 2023 [Publ 4323], fig. 3
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 4 |
4 |
2 |
1 |
0 |
0 |
0 |
20.0 |
+-------- Fusconaia_cerina **
--+
| +----- Pleurobema_riddellii **
+--+
| +-- Pleurobema_beadleianum **
+--+
+-- Pleurobema_clava **
Krebs, 2004 [Publ 2692], fig. 2
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 3 |
5 |
3 |
3 |
2 |
2 |
1 |
17.0 |
+-------- Quadrula_quadrula **
|
--+ +-- Lasmigona_costata
| +--+
| | +-- Pyganodon_grandis
+--+
| +-- Lampsilis_siliquoidea **
+--+
+-- Fusconaia_flava **
Curole & Kocher, 2002 [Publ 2712], fig. 1-F
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 3 |
3 |
1 |
2 |
1 |
0 |
0 |
17.0 |
+----- Lampsilis_teres **
--+
| +-- Quadrula_quadrula **
+--+
+-- Pustulosa_pustulosa **
Curole & Kocher, 2002 [Publ 2712], fig. 1a-M
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 3 |
4 |
2 |
2 |
1 |
1 |
1 |
18.0 |
+-- Utterbackiana_implicata
+--+
| +-- Lampsilis_teres **
--+
| +-- Quadrula_quadrula **
+--+
+-- Pustulosa_pustulosa **
Curole & Kocher, 2002 [Publ 2712], fig. 1b-M
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 3 |
4 |
2 |
2 |
1 |
1 |
1 |
18.0 |
+-------- Lampsilis_teres **
--+
| +----- Utterbackiana_implicata
+--+
| +-- Quadrula_quadrula **
+--+
+-- Pustulosa_pustulosa **
Chen et al., 2012 [Publ 3624], fig. 4
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 3 |
3 |
1 |
2 |
4 |
4 |
2 |
14.0 |
+----- Quadrula_quadrula **
+--+
| | +-- Lampsilis_ornata **
| +--+
| +-- Venustaconcha_ellipsiformis **
--+
| +-- Pyganodon_grandis
| +--+
| | +-- Cristaria_plicata
+--+
| +-- Sinohyriopsis_cumingii
+--+
+-- Lamprotula_leaii
Sheng et al., 2014 [Publ 3704], fig. 2
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 3 |
6 |
4 |
2 |
5 |
5 |
2 |
19.0 |
+----------------- Cristaria_plicata
|
--+ +-------------- Pyganodon_grandis
| |
+--+ +----------- Lampsilis_ornata **
| |
+--+ +-- Venustaconcha_ellipsiformis **
| +-----+
| | +-- Quadrula_quadrula **
+--+
| +----- Lamprotula_leaii
+--+
| +-- Sinohyriopsis_cumingii
+--+
+-- Sinohyriopsis_schlegelii
Wu et al., 2016 [Publ 3711], fig. 1
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 3 |
3 |
1 |
2 |
18 |
18 |
2 |
14.0 |
+----- Quadrula_quadrula **
+-----+
| | +-- Venustaconcha_ellipsiformis **
| +--+
| +-- Lampsilis_ornata **
+-----------+
| | +-- Sinohyriopsis_schlegelii
| | +-----+
| | | +-- Sinohyriopsis_cumingii
| | |
| +--+ +-- Sinosolenaia_carinata
| | +--+
| | | +-- Sinosolenaia_oleivora
| +--+
| | +-- Lamprotula_leaii
--+ +--+
| +-- Pronodularia_japanensis
|
| +-- Unio_pictorum
| +-----+
| | +-- Nodularia_douglasiae
| +-----------+
| | | +----- Aculamprotula_tortuosa
| | +--+
| | | +-- Aculamprotula_coreana
+--+ +--+
| +-- Aculamprotula_tientsinensis
|
| +----------------- Sinanodonta_lucida
+--+
| +-------------- Anemina_arcaeformis
+--+
| +----------- Cristaria_plicata
+--+
| +-------- Anodonta_anatina
+--+
| +----- Pyganodon_grandis
+--+
| +-- Utterbackia_peninsularis
+--+
+-- Utterbackia_imbecillis
Zhou et al., 2017 [Publ 3788], fig. 2
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 3 |
3 |
1 |
2 |
14 |
14 |
2 |
14.0 |
+-------- Lanceolaria_lanceolata
|
+--------+ +-- Nodularia_douglasiae
| | +--+
| | | +-- Tchangsinaia_piscicula
| +--+
| | +-- Aculamprotula_tientsinensis
| +--+
--+ +-- Aculamprotula_tortuosa
|
| +----- Quadrula_quadrula **
| +--------+
| | | +-- Potamilus_leptodon **
| | +--+
| | +-- Lampsilis_ornata **
+--+
| +-- Sinohyriopsis_cumingii
| +--------+
| | +-- Sinohyriopsis_schlegelii
| |
+--+ +-- Lamprotula_caveata
| +-----+
| | +-- Lamprotula_leaii
+--+
| +----- Ptychorhynchus_pfisteri
+--+
| +-- Sinosolenaia_carinata
+--+
+-- Sinosolenaia_oleivora
Amaro et al., 2017 [Publ 3791], fig. 3A
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 3 |
3 |
1 |
2 |
9 |
6 |
2 |
14.0 |
+-------- Platynaias_compressa
+--+
| | +----- Pyganodon_grandis
| +--+
| | +-- Utterbackia_imbecillis
| +--+
--+ +-- Utterbackia_peninsularis
|
| +-- Pronodularia_japanensis
| +-----+
| | +-- Sinosolenaia_carinata
+--+
| +----- Quadrula_quadrula **
+--+
| +-- Toxolasma_parvum **
+--+
+-- Venustaconcha_ellipsiformis **
Amaro et al., 2017 [Publ 3791], fig. 3B
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 3 |
5 |
3 |
2 |
9 |
6 |
2 |
16.0 |
+-------- Platynaias_compressa
+--+
| | +----- Pyganodon_grandis
| +--+
| | +-- Utterbackia_imbecillis
| +--+
--+ +-- Utterbackia_peninsularis
|
| +-- Toxolasma_parvum **
| +-----+
| | +-- Venustaconcha_ellipsiformis **
+--+
| +----- Quadrula_quadrula **
+--+
| +-- Pronodularia_japanensis
+--+
+-- Sinosolenaia_carinata
Wen et al., 2017 [Publ 3795], fig. 3-M
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 3 |
3 |
1 |
2 |
3 |
3 |
1 |
14.0 |
+----- Anodonta_anatina
+--+
| | +-- Utterbackia_peninsularis
| +--+
--+ +-- Pyganodon_grandis
|
| +----- Quadrula_quadrula **
+--+
| +-- Potamilus_alatus **
+--+
+-- Venustaconcha_ellipsiformis **
Feng et al., 2016 [Publ 3800], fig. 2
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 3 |
3 |
1 |
2 |
12 |
10 |
2 |
14.0 |
+-- Sinosolenaia_carinata
+-----+
| +-- Sinosolenaia_oleivora
+-----+
| | +----- Quadrula_quadrula **
| +--+
| | +-- Potamilus_leptodon **
--+ +--+
| +-- Lampsilis_ornata **
|
| +----------- Tchangsinaia_piscicula
| |
+--+ +-------- Pyganodon_grandis
| |
+--+ +-- Anemina_arcaeformis
| +--+
| | +-- Anemina_euscaphys
+--+
| +-- Sinanodonta_lucida
+--+
+-- Sinanodonta_woodiana
Huang et al., 2018 [Publ 3828], fig. 6-M
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 3 |
3 |
1 |
2 |
14 |
12 |
2 |
14.0 |
+----- Quadrula_quadrula **
+-----------+
| | +-- Venustaconcha_ellipsiformis **
| +--+
| +-- Potamilus_alatus **
|
| +-------- Sinosolenaia_carinata
--+ +-----+
| | | +----- Potomida_littoralis
| | +--+
| | | +-- Lamprotula_leaii
| | +--+
| | +-- Pronodularia_japanensis
+--+
| +-------- Sinanodonta_woodiana
| +--+
| | | +----- Anodonta_anatina
| | +--+
| | | +-- Pyganodon_grandis
+--+ +--+
| +-- Utterbackia_peninsularis
|
| +-------- Aculamprotula_tortuosa
+--+
| +----- Unio_tumidus
+--+
| +-- Unio_crassus
+--+
+-- Unio_delphinus
Huang et al., 2019 [Publ 3843], fig. 3-M
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 3 |
3 |
1 |
2 |
20 |
17 |
2 |
14.0 |
+----- Quadrula_quadrula **
+-----------------+
| | +-- Venustaconcha_ellipsiformis **
| +--+
| +-- Potamilus_alatus **
|
| +----------- Sinohyriopsis_cumingii
| |
| | +----- Potomida_littoralis
--+ +--------+ +--+
| | | | | +-- Lamprotula_leaii
| | | | +--+
| | +--+ +-- Pronodularia_japanensis
| | |
| | | +----- Parvasolenaia_rivularis
| | +--+
| | | +-- Sinosolenaia_oleivora
+--+ +--+
| +-- Sinosolenaia_carinata
|
| +-------- Sinanodonta_woodiana
| +--------+
| | | +----- Anodonta_anatina
| | +--+
| | | +-- Pyganodon_grandis
+--+ +--+
| +-- Utterbackia_peninsularis
|
| +-------------- Lanceolaria_lanceolata
+--+
| +----------- Schistodesmus_lampreyanus
+--+
| +-------- Aculamprotula_tortuosa
+--+
| +----- Unio_tumidus
+--+
| +-- Unio_crassus
+--+
+-- Unio_delphinus
Chase et al., 2018 [Publ 3847], fig. 5
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 3 |
3 |
1 |
1 |
7 |
5 |
2 |
14.0 |
+----------- Sinosolenaia_carinata
|
| +----- Potamilus_alatus **
--+ +--+
| | | +-- Lampsilis_siliquoidea **
| | +--+
| | +-- Lampsilis_powellii **
+--+
| +-- Anodonta_anatina
| +--+
| | +-- Utterbackia_peninsularis
+--+
| +-- Unio_delphinus
+--+
+-- Unio_tumidus
Chase et al., 2018 [Publ 3861], fig. 1
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 3 |
3 |
1 |
2 |
8 |
6 |
2 |
14.0 |
+----- Quadrula_quadrula **
+--------+
| | +-- Lampsilis_powellii **
| +--+
--+ +-- Lampsilis_siliquoidea **
|
| +----------- Lamprotula_leaii
| |
+--+ +-- Unio_delphinus
| +-----+
| | +-- Unio_tumidus
+--+
| +----- Anodonta_anatina
+--+
| +-- Utterbackia_peninsularis
+--+
+-- Pyganodon_grandis
Chase et al., 2018 [Publ 3861], fig. 1-M
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 3 |
3 |
1 |
2 |
8 |
6 |
2 |
14.0 |
+-------- Lamprotula_leaii
+--+
| | +----- Quadrula_quadrula **
| +--+
| | +-- Lampsilis_powellii **
| +--+
--+ +-- Lampsilis_siliquoidea **
|
| +-- Unio_delphinus
| +-----+
| | +-- Unio_tumidus
+--+
| +----- Anodonta_anatina
+--+
| +-- Utterbackia_peninsularis
+--+
+-- Pyganodon_grandis
Burzynski & Soroka, 2018 [Publ 3862], fig. 5a
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 3 |
3 |
1 |
2 |
17 |
15 |
2 |
14.0 |
+----- Quadrula_quadrula **
+-----------------+
| | +-- Potamilus_alatus **
| +--+
| +-- Venustaconcha_ellipsiformis **
|
| +----------- Sinohyriopsis_cumingii
--+ +--------+
| | | +-------- Sinosolenaia_carinata
| | +--+
| | | +----- Potomida_littoralis
| | +--+
| | | +-- Lamprotula_leaii
| | +--+
+--+ +-- Pronodularia_japanensis
|
| +----- Sinanodonta_woodiana
| |
| +-----------+----- Anodonta_anatina
| | |
| | | +-- Utterbackia_peninsularis
+--+ +--+
| +-- Pyganodon_grandis
|
| +-------------- Lanceolaria_lanceolata
+--+
| +----------- Aculamprotula_tortuosa
+--+
| +-------- Unio_tumidus
+--+
| +----- Unio_crassus
+--+
| +-- Unio_pictorum
+--+
+-- Unio_delphinus
Froufe et al., 2019 [Publ 3936], fig. 3-M
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 3 |
3 |
1 |
2 |
16 |
15 |
2 |
14.0 |
+----- Quadrula_quadrula **
+-----------+
| | +-- Potamilus_alatus **
| +--+
| +-- Venustaconcha_ellipsiformis **
|
| +----- Unio_tumidus
| +-----+
| | | +-- Unio_crassus
--+ | +--+
| +--+ +-- Unio_delphinus
| | |
| | | +-------- Sinanodonta_woodiana
| | +--+
| | | +----- Anodonta_anatina
| | +--+
| | | +-- Utterbackia_peninsularis
+--+ +--+
| +-- Pyganodon_grandis
|
| +-- Chamberlainia_somsakpanhai
| +--------+
| | +-- Sinohyriopsis_cumingii
| |
+--+ +-- Potomida_littoralis
| +-----+
| | +-- Pronodularia_japanensis
| |
+--+ +-- Microcondylaea_bonellii
| +--+
| | +-- Sinosolenaia_carinata
+--+
| +-- Pilsbryoconcha_exilis
+--+
+-- Pseudodon_vondembuschianus
Froufe et al., 2019 [Publ 3936], fig. 4
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 3 |
3 |
1 |
2 |
16 |
15 |
2 |
14.0 |
+----- Quadrula_quadrula **
+-----------+
| | +-- Potamilus_alatus **
| +--+
| +-- Venustaconcha_ellipsiformis **
|
| +----- Unio_tumidus
| +-----+
| | | +-- Unio_crassus
--+ | +--+
| +--+ +-- Unio_delphinus
| | |
| | | +-------- Sinanodonta_woodiana
| | +--+
| | | +----- Anodonta_anatina
| | +--+
| | | +-- Utterbackia_peninsularis
+--+ +--+
| +-- Pyganodon_grandis
|
| +-- Chamberlainia_somsakpanhai
| +--------+
| | +-- Sinohyriopsis_cumingii
| |
+--+ +-- Potomida_littoralis
| +-----+
| | +-- Pronodularia_japanensis
| |
+--+ +-- Microcondylaea_bonellii
| +--+
| | +-- Sinosolenaia_carinata
+--+
| +-- Pilsbryoconcha_exilis
+--+
+-- Pseudodon_vondembuschianus
Wu et al., 2018 [Publ 4143], fig. 2
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 3 |
3 |
1 |
2 |
19 |
19 |
2 |
14.0 |
+----- Quadrula_quadrula **
+-----+
| | +-- Venustaconcha_ellipsiformis **
| +--+
| +-- Lampsilis_ornata **
+-----------+
| | +-- Sinohyriopsis_schlegelii
| | +-----+
| | | +-- Sinohyriopsis_cumingii
| | |
| +--+ +-- Sinosolenaia_oleivora
| | +--+
| | | +-- Sinosolenaia_carinata
| +--+
| | +-- Pronodularia_japanensis
--+ +--+
| +-- Lamprotula_leaii
|
| +-- Unio_pictorum
| +--------+
| | +-- Nodularia_douglasiae
| +--------+
| | | +-------- Aculamprotula_tortuosa
| | +--+
| | | +----- Aculamprotula_scripta
| | +--+
+--+ | +-- Aculamprotula_coreana
| +--+
| +-- Aculamprotula_tientsinensis
|
| +----------------- Sinanodonta_lucida
+--+
| +-------------- Anemina_arcaeformis
+--+
| +----------- Cristaria_plicata
+--+
| +-------- Anodonta_anatina
+--+
| +----- Pyganodon_grandis
+--+
| +-- Utterbackia_peninsularis
+--+
+-- Utterbackia_imbecillis
Wu et al., 2020 [Publ 4147], fig. 3
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 3 |
3 |
1 |
2 |
27 |
26 |
2 |
14.0 |
+----- Quadrula_quadrula **
+--------------------+
| | +-- Toxolasma_parvum **
| +--+
| +-- Venustaconcha_ellipsiformis **
|
| +-- Sinohyriopsis_schlegelii
| +--------------+
| | +-- Sinohyriopsis_cumingii
| |
| +-----+ +-- Sinosolenaia_oleivora
--+ | | +-----------+
| | | | +-- Sinosolenaia_carinata
| | +--+
| | | +----------- Pronodularia_japanensis
| | +--+
| | | +-------- Lamprotula_leaii
| | +--+
| | | +----- Lamprotula_caveata[1]
| | +--+
| | | +-- Lamprotula_cornuumlunae
| | +--+
+--+ +-- Lamprotula_caveata[2]
|
| +----- Cristaria_plicata
| +-----+
| | | +-- Sinanodonta_lucida
| | +--+
| +--------+ +-- Anemina_arcaeformis
| | |
| | | +-------- Anodonta_anatina
| | +--+
| | | +----- Pyganodon_grandis
| | +--+
| | | +-- Utterbackia_peninsularis
+--+ +--+
| +-- Utterbackia_imbecillis
|
| +-------- Aculamprotula_tortuosa
| +--------+
| | | +----- Aculamprotula_scripta
| | +--+
| | | +-- Aculamprotula_tientsinensis
+--+ +--+
| +-- Aculamprotula_coreana
|
| +-------------- Unio_pictorum
+--+
| +----------- Pseudocuneopsis_capitata
+--+
| +-------- Nodularia_douglasiae
+--+
| +----- Schistodesmus_lampreyanus
+--+
| +-- Cuneopsis_heudei
+--+
+-- Tchangsinaia_piscicula
Guerra et al., 2019 [Publ 4168], fig. 1-M
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 3 |
3 |
1 |
2 |
19 |
14 |
2 |
14.0 |
+----- Quadrula_quadrula **
+-----------------+
| | +-- Venustaconcha_ellipsiformis **
| +--+
| +-- Potamilus_alatus **
|
| +-------- Sinosolenaia_carinata
--+ +-----------+
| | | +----- Potomida_littoralis
| | +--+
| | | +-- Lamprotula_leaii
| | +--+
| | +-- Pronodularia_japanensis
+--+
| +-------- Sinanodonta_woodiana
| +--------+
| | | +----- Anodonta_anatina
| | +--+
| | | +-- Utterbackia_peninsularis
+--+ +--+
| +-- Pyganodon_grandis
|
| +-------------- Lanceolaria_lanceolata
+--+
| +----------- Schistodesmus_lampreyanus
+--+
| +-------- Aculamprotula_tortuosa
+--+
| +----- Unio_tumidus
+--+
| +-- Unio_crassus
+--+
+-- Unio_delphinus
Guerra et al., 2019 [Publ 4168], fig. 2-M
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 3 |
3 |
1 |
2 |
19 |
14 |
2 |
14.0 |
+----- Quadrula_quadrula **
+-----------------+
| | +-- Venustaconcha_ellipsiformis **
| +--+
| +-- Potamilus_alatus **
|
| +-------- Sinosolenaia_carinata
--+ +-----------+
| | | +----- Potomida_littoralis
| | +--+
| | | +-- Lamprotula_leaii
| | +--+
| | +-- Pronodularia_japanensis
+--+
| +-- Utterbackia_peninsularis
| +--+
| | +-- Pyganodon_grandis
| +-----------+
| | | +-- Anodonta_anatina
| | +--+
+--+ +-- Sinanodonta_woodiana
|
| +-------------- Lanceolaria_lanceolata
+--+
| +----------- Aculamprotula_tortuosa
+--+
| +-------- Schistodesmus_lampreyanus
+--+
| +----- Unio_tumidus
+--+
| +-- Unio_crassus
+--+
+-- Unio_delphinus
Smith & Johnson, 2020 [Publ 4229], fig. 2
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 3 |
3 |
1 |
1 |
0 |
0 |
0 |
19.0 |
+----- Potamilus_inflatus **
--+
| +-- Potamilus_purpuratus **
+--+
+-- Potamilus_fragilis **
Zieritz et al., 2021 [Publ 4231], fig. 3-M, 4
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 3 |
3 |
1 |
2 |
20 |
19 |
2 |
14.0 |
+----- Quadrula_quadrula **
+--------------+
| | +-- Venustaconcha_ellipsiformis **
| +--+
| +-- Potamilus_alatus **
|
| +----- Unio_tumidus
| +-----+
| | | +-- Unio_delphinus
--+ | +--+
| +-----+ +-- Unio_crassus
| | |
| | | +-------- Sinanodonta_woodiana
| | +--+
| | | +----- Anodonta_anatina
| | +--+
| | | +-- Utterbackia_peninsularis
| | +--+
+--+ +-- Pyganodon_grandis
|
| +-- Chamberlainia_somsakpanhai
| +-----------+
| | +-- Sinohyriopsis_cumingii
| |
| | +-- Lens_contradens
| | +--+
+--+ | +-- Physunio_superbus
| +-----+
| | | +-- Hyriopsis_bialata
| | +--+
| | +-- Rectidens_sumatrensis
+--+
| +-- Pilsbryoconcha_exilis
| +-----+
| | +-- Pseudodon_vondembuschianus
| |
+--+ +-- Pronodularia_japanensis
| +--+
| | +-- Potomida_littoralis
+--+
| +-- Microcondylaea_bonellii
+--+
+-- Sinosolenaia_carinata
Graf, 2000 [Publ 2647], fig. 1
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 2 |
3 |
0 |
2 |
16 |
4 |
2 |
17.0 |
+-------- Unio
--+
| +----- Elliptio **
+--+
| +-- Pyganodon
+--+
+-- Lampsilis **
Graf & Ó Foighil, 2000 [Publ 2649], fig. 1, 2
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 2 |
2 |
0 |
2 |
10 |
2 |
1 |
-- |
+-- Unio_pictorum
+--+
| +-- Pyganodon_grandis
--+
| +-- Amblema_plicata **
+--+
+-- Lampsilis_cardium **
Krebs, 2004 [Publ 2692], fig. 2-M
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 2 |
2 |
0 |
2 |
2 |
2 |
1 |
-- |
+-- Lasmigona_costata
+--+
| +-- Pyganodon_grandis
--+
| +-- Lampsilis_siliquoidea **
+--+
+-- Fusconaia_flava **
Reis et al., 2013 [Publ 3580], fig. 4
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 2 |
2 |
0 |
2 |
12 |
10 |
2 |
-- |
+-- Amblema_plicata **
+-----------------+
| +-- Quadrula_nobilis **
|
--+ +----------------- Potomida_littoralis
| |
| | +----- Anodonta_cygnea
+--+ +--------+
| | | +-- Anodonta_anatina
| | +--+
+--+ +-- Pseudanodonta_complanata
|
| +----------- Unio_tumidus
| |
+--+ +-- Unio_tumidiformis
| +-----+
| | +-- Unio_crassus
+--+
| +----- Unio_delphinus
+--+
| +-- Unio_mancus
+--+
+-- Unio_pictorum
Reis et al., 2013 [Publ 3580], fig. 5
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 2 |
2 |
0 |
2 |
14 |
12 |
2 |
-- |
+-- Amblema_plicata **
+--------------------+
| +-- Theliderma_metanevra **
|
--+ +-------------------- Potomida_littoralis
| |
| | +-- Anodonta_anatina
+--+ |
| +--------------+-- Anodonta_cygnea
| | |
| | +-- Pseudanodonta_complanata
+--+
| +-------------- Unio_tumidus
| |
+--+ +-- Unio_tumidiformis
| +--------+
| | +-- Unio_crassus
+--+
| +-------- Unio
| |
+--+ +-- Unio_delphinus
| +--+
| | +-- Unio_mancus
+--+
| +-- Unio_pictorum
+--+
+-- Unio_caffer
Huang et al., 2013 [Publ 3629], fig. 3-M
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 2 |
2 |
0 |
2 |
4 |
4 |
2 |
-- |
+-- Pyganodon_grandis
+-----+
| +-- Utterbackia_peninsularis
|
--+ +-- Sinosolenaia_carinata
| +--+
| | +-- Pronodularia_japanensis
+--+
| +-- Quadrula_quadrula **
+--+
+-- Venustaconcha_ellipsiformis **
Xue et al., 2016 [Publ 3659], fig. 1
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 2 |
2 |
0 |
1 |
14 |
14 |
2 |
-- |
+-- Sinohyriopsis_cumingii
+--+
| +-- Lamprotula_leaii
+-----------+
| | +-- Lampsilis_ornata **
| +--+
| +-- Venustaconcha_ellipsiformis **
|
| +-- Cristaria_plicata
| +-----+
--+ | +-- Platynaias_compressa
| |
| +-----+ +-- Anemina_arcaeformis
| | | +--+
| | | | +-- Anemina_euscaphys
| | +--+
| | | +-- Sinanodonta_lucida
+--+ +--+
| +-- Sinanodonta_woodiana
|
| +-- Lanceolaria_lanceolata
| +--------+
| | +-- Lanceolaria_grayii
+--+
| +-------- Aculamprotula_tortuosa
+--+
| +----- Schistodesmus_lampreyanus
+--+
| +-- Nodularia_douglasiae
+--+
+-- Unio_pictorum
Xue et al., 2016 [Publ 3659], fig. 1-M
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 2 |
2 |
0 |
2 |
0 |
0 |
0 |
-- |
+-- Quadrula_quadrula **
--+
+-- Venustaconcha_ellipsiformis **
Froufe et al., 2015 [Publ 3664], fig. 1
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 2 |
2 |
0 |
2 |
10 |
10 |
2 |
-- |
+-- Sinohyriopsis_cumingii
+--------+
| +-- Sinohyriopsis_schlegelii
|
--+ +-- Venustaconcha_ellipsiformis **
| +-----+
| | +-- Quadrula_quadrula **
+--+
| +----- Anodonta_anatina
+--+
| +-- Utterbackia_peninsularis
+--+
+-- Pyganodon_grandis
Froufe et al., 2015 [Publ 3664], fig. 1-M
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 2 |
2 |
0 |
2 |
6 |
6 |
2 |
-- |
+----- Anodonta_anatina
+-----+
| | +-- Utterbackia_peninsularis
| +--+
| +-- Pyganodon_grandis
--+
| +-- Venustaconcha_ellipsiformis **
| +-----+
| | +-- Quadrula_quadrula **
+--+
| +----- Sinosolenaia_carinata
+--+
| +-- Potomida_littoralis
+--+
+-- Pronodularia_japanensis
Wei et al., 2016 [Publ 3665], fig. 0
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 2 |
4 |
0 |
2 |
4 |
4 |
2 |
17.0 |
+-- Sinohyriopsis_cumingii
+--+
| +-- Sinohyriopsis_schlegelii
|
--+ +-- Cristaria_plicata
| |
| +-- Lampsilis_ornata **
+--+
+-- Pyganodon_grandis
|
+-- Quadrula_quadrula **
Giribet & Distel, 2003 [Publ 3670], fig. 3.5
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 2 |
3 |
0 |
2 |
4 |
1 |
1 |
16.0 |
+-- Elliptio_complanata **
|
--+-- Lampsilis_cardium **
|
+-- Unio_ravoisieri
Hewitt et al., 2018 [Publ 3758], fig. 5
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 2 |
2 |
0 |
1 |
7 |
6 |
1 |
-- |
+-- Fusconaia_flava **
+--------+
| +-- Pleurobema_rubellum **
|
--+ +-------- Pyganodon_grandis
| |
| | +-- Platynaias_subviridis
+--+-----+
| +-- Strophitus_undulatus
|
| +----- Lasmigona_costata
+--+
| +-- Arcidens_wheeleri
+--+
+-- Lasmigona_complanata
Soroka & Burzynski, 2018 [Publ 3762], fig. 3
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 2 |
2 |
0 |
2 |
5 |
5 |
1 |
-- |
+-- Venustaconcha_ellipsiformis **
+--------+
| +-- Quadrula_quadrula **
|
--+ +-- Unio_tumidus
| +-----+
| | +-- Aculamprotula_tortuosa
+--+
| +----- Anodonta_anatina
+--+
| +-- Pyganodon_grandis
+--+
+-- Utterbackia_peninsularis
Soroka & Burzynski, 2018 [Publ 3762], fig. 3-M
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 2 |
2 |
0 |
2 |
5 |
5 |
1 |
-- |
+-- Venustaconcha_ellipsiformis **
+--------+
| +-- Quadrula_quadrula **
|
--+ +-- Unio_tumidus
| +-----+
| | +-- Aculamprotula_tortuosa
+--+
| +----- Anodonta_anatina
+--+
| +-- Pyganodon_grandis
+--+
+-- Utterbackia_peninsularis
Guerra et al., 2017 [Publ 3772], fig. 2a-M
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 2 |
2 |
0 |
2 |
6 |
4 |
2 |
-- |
+-- Sinosolenaia_carinata
+--+
| +-- Pronodularia_japanensis
|
| +-- Utterbackia_peninsularis
--+--+
| +-- Pyganodon_grandis
|
| +-- Venustaconcha_ellipsiformis **
+--+
+-- Quadrula_quadrula **
Guerra et al., 2017 [Publ 3772], fig. 2b-M
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 2 |
2 |
0 |
2 |
6 |
4 |
2 |
-- |
+-- Sinosolenaia_carinata
+--+
| +-- Pronodularia_japanensis
|
| +-- Utterbackia_peninsularis
--+--+
| +-- Pyganodon_grandis
|
| +-- Venustaconcha_ellipsiformis **
+--+
+-- Quadrula_quadrula **
Burzynski et al., 2017 [Publ 3774], fig. 1 bottom
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 2 |
2 |
0 |
2 |
11 |
11 |
2 |
-- |
+----- Aculamprotula_tortuosa
+--+
| | +-- Unio_crassus
| +--+
+--------+ +-- Unio_tumidus
| |
| | +----- Anodonta_anatina
| +--+
| | +-- Utterbackia_peninsularis
--+ +--+
| +-- Pyganodon_grandis
|
| +-- Quadrula_quadrula **
| +-----------+
| | +-- Venustaconcha_ellipsiformis **
+--+
| +----------- Sinohyriopsis_cumingii
+--+
| +-------- Sinosolenaia_carinata
+--+
| +----- Potomida_littoralis
+--+
| +-- Lamprotula_leaii
+--+
+-- Pronodularia_japanensis
Burzynski et al., 2017 [Publ 3774], fig. 1 top
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 2 |
2 |
0 |
2 |
11 |
11 |
2 |
-- |
+----- Aculamprotula_tortuosa
+--+
| | +-- Unio_crassus
| +--+
+--------+ +-- Unio_tumidus
| |
| | +----- Anodonta_anatina
| +--+
| | +-- Utterbackia_peninsularis
--+ +--+
| +-- Pyganodon_grandis
|
| +-- Quadrula_quadrula **
| +-----------+
| | +-- Venustaconcha_ellipsiformis **
+--+
| +----------- Sinohyriopsis_cumingii
+--+
| +-------- Sinosolenaia_carinata
+--+
| +----- Lamprotula_leaii
+--+
| +-- Potomida_littoralis
+--+
+-- Pronodularia_japanensis
Fonseca et al., 2016 [Publ 3775], fig. 1-M
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 2 |
2 |
0 |
2 |
7 |
7 |
2 |
-- |
+-- Venustaconcha_ellipsiformis **
+-----------+
| +-- Quadrula_quadrula **
|
--+ +----- Sinosolenaia_carinata
| +-----+
| | | +-- Potomida_littoralis
| | +--+
+--+ +-- Pronodularia_japanensis
|
| +-------- Unio_delphinus
+--+
| +----- Anodonta_anatina
+--+
| +-- Utterbackia_peninsularis
+--+
+-- Pyganodon_grandis
Amaro et al., 2017 [Publ 3791], fig. 3A-M
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 2 |
6 |
0 |
2 |
5 |
4 |
2 |
19.0 |
+----------- Quadrula_quadrula **
|
--+ +-- Pronodularia_japanensis
| +-----+
| | +-- Sinosolenaia_carinata
+--+
| +----- Venustaconcha_ellipsiformis **
+--+
| +-- Pyganodon_grandis
+--+
+-- Utterbackia_peninsularis
Amaro et al., 2017 [Publ 3791], fig. 3B-M
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 2 |
6 |
0 |
2 |
5 |
4 |
2 |
19.0 |
+----------- Quadrula_quadrula **
|
--+ +-------- Venustaconcha_ellipsiformis **
| |
+--+ +-- Utterbackia_peninsularis
| +--+
| | +-- Pyganodon_grandis
+--+
| +-- Pronodularia_japanensis
+--+
+-- Sinosolenaia_carinata
Lopes-Lima et al., 2017 [Publ 3792], fig. 2-M
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 2 |
2 |
0 |
2 |
8 |
7 |
2 |
-- |
+-- Venustaconcha_ellipsiformis **
+-----------+
| +-- Quadrula_quadrula **
|
--+ +----- Sinosolenaia_carinata
| +-----+
| | | +-- Potomida_littoralis
| | +--+
+--+ +-- Pronodularia_japanensis
|
| +-------- Unio_delphinus
+--+
| +----- Anodonta_anatina
+--+
| +-- Utterbackia_peninsularis
+--+
+-- Pyganodon_grandis
Araujo et al., 2017 [Publ 3801], fig. 2
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 2 |
2 |
0 |
2 |
12 |
11 |
3 |
-- |
+-- Lamellidens_savadiensis
+--------------+
| +-- Coelatura_aegyptiaca
|
| +-- Amblema_plicata **
--+ +-----------+
| | +-- Lampsilis_teres **
| |
| | +-- Schistodesmus_lampreyanus
| | +--+
+--+ | +-- Aculamprotula_tortuosa
| +-----+
| | | +-- Anodonta_anatina
| | +--+
| | +-- Unio_delphinus
+--+
| +-- Gonidea_angulata
| +-----+
| | +-- Microcondylaea_bonellii
+--+
| +----- Potomida_littoralis
+--+
| +-- Lamprotula_leaii
+--+
+-- Pronodularia_japanensis
Metzger et al., 2018 [Publ 3952], fig. 1C
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 2 |
2 |
0 |
2 |
1 |
1 |
1 |
-- |
+----- Lasmigona_costata
--+
| +-- Elliptio_complanata **
+--+
+-- Lampsilis_siliquoidea **
Zieritz et al., 2021 [Publ 4218], fig. 5
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 2 |
2 |
0 |
2 |
30 |
28 |
1 |
-- |
+-- Amblema_plicata **
+--------------------------------+
| +-- Lampsilis_ovata **
|
| +-------------------------------- Chamberlainia_somsakpanhai
--+ |
| | +----------------------------- Potomida_littoralis
| | |
| | | +-- Gonidea_angulata
+--+ | +--------------------+
| | | +-- Microcondylaea_bonellii
| | +--+
| | | | +-------------------- Schepmania
| | | | |
| | | +--+ +-- Indopseudodon_bogani
+--+ | | +--------------+
| | | | +-- Indopseudodon_crebristriatus
| | +--+
| | | +-------------- Pilsbryoconcha_exilis
| | | |
| | +--+ +-- Pseudodon_mekongi
| | | +--------+
| | | | +-- Pseudodon_vondembuschianus
| | +--+
+--+ | +-------- Sundadontina
| +--+
| | +----- Thaiconcha_callifera
| +--+
| | +-- Bineurus_loeiensis
| +--+
| +-- Bineurus_anodontinum
|
| +----- Ctenodesma_borneensis
| +--------------+
| | | +-- Khairuloconcha_lunbawangorum
| | +--+
| | +-- Khairuloconcha_sahanae
| |
+-----+ +-------- Ensidens_sagittarius
| +--------+
| | | +----- Rectidens_sumatrensis
| | +--+
| | | +-- Hyriopsis_bialata
+--+ +--+
| +-- Hyriopsis_desowitzi
|
| +-------------- Physunio_superbus
| |
+--+ +----------- Trapezoideus_foliaceus
| |
+--+ +-- Lens_contradens
| +-----+
| | +-- Lens_eximius
+--+
| +----- Yaukthwa_elongatula
+--+
| +-- Yaukthwa_inlenensis
+--+
+-- Yaukthwa_paiensis
Sonowal et al., 2021 [Publ 4221], fig. 3a
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 2 |
6 |
0 |
2 |
13 |
7 |
3 |
19.0 |
+----- Elliptio_complanata **
|
+----- Lampsilis_cardium **
--+
| +-- Anodonta_cygnea
| |
| +-- Unio_ravoisieri
+--+
+-- Unio_pictorum
|
+-- Aculamprotula_tientsinensis
Sonowal et al., 2021 [Publ 4221], fig. 3b
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 2 |
9 |
0 |
2 |
13 |
7 |
3 |
22.0 |
+-- Aculamprotula_tientsinensis
|
+-- Lamprotula_leaii
|
+-- Lampsilis_cardium **
|
+-- Potomida_littoralis
|
--+-- Unio_pictorum
|
+-- Anodonta_cygnea
|
+-- Unio_ravoisieri
|
+-- Lamellidens_mainwaringi
|
+-- Elliptio_complanata **
Sonowal et al., 2021 [Publ 4221], fig. 4
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 2 |
7 |
0 |
2 |
14 |
8 |
3 |
20.0 |
+----------------------- Lamprotula_leaii
--+
| +-------------------- Potomida_littoralis
+--+
| +----------------- Lampsilis_cardium **
+--+
| +-------------- Elliptio_complanata **
+--+
| +----------- Aculamprotula_tientsinensis
+--+
| +-------- Unio_pictorum
+--+
| +----- Anodonta_cygnea
+--+
| +-- Unio_ravoisieri
+--+
+-- Lamellidens_corrianus
Konopleva et al., 2023 [Publ 4269], fig. S1, 4
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 2 |
2 |
0 |
2 |
54 |
53 |
3 |
-- |
+-- Lampsilis_ovata **
+--------------+
| +-- Amblema_plicata **
|
| +----------- Sinosolenaia_recognita
| |
| +--+ +-- Ptychorhynchus_pfisteri
+-----------+ | | +-----+
| | | | | +-- Parvasolenaia_rivularis
| | | +--+
| | | | +----- Gonidea_angulata
| | | +--+
| | | | +-- Microcondylaea_bonellii
| +--+ +--+
| | +-- Leguminaia_wheatleyi
| |
| | +----- Chamberlainia_somsakpanhai
| | +-----+
| | | | +-- Sinohyriopsis_schlegelii
| | | +--+
| +--+ +-- Sinohyriopsis_cumingii
| |
| | +-------- Potomida_littoralis
| +--+
| | +----- Pronodularia_japanensis
| +--+
| | +-- Lamprotula_caveata
--+ +--+
| +-- Lamprotula_leaii
|
| +----------------- Middendorffinaia_mongolica
| |
| | +-------- Aculamprotula_tortuosa
| | +-----+
| +--------+ | | +----- Aculamprotula_scripta
| | | | +--+
| | | | | +-- Aculamprotula_tientsinensis
| | | | +--+
| | +--+ +-- Aculamprotula_coreana
| | |
| | | +----- Unio_tumidus
| | | +-----+
| | | | | +-- Unio_pictorum
| | | | +--+
| | +--+ +-- Unio_crassus
| | |
| | | +-------- Nodularia_douglasiae
+--+ +--+
| | +----- Tchangsinaia_piscicula
| +--+
| | +-- Cuneopsis_rufescens
| +--+
| +-- Cuneopsis_heudei
|
| +-------- Lepidodesma_languilati
| +--------------+
| | | +----- Lanceolaria_gladiola
| | +--+
| | | +-- Lanceolaria_grayii
| | +--+
+--+ +-- Lanceolaria_triformis
|
| +-------------------- Acuticosta_chinensis
| |
| | +----- Anodonta_anatina
| | +-----------+
+--+ | | +-- Pseudanodonta_complanata
| | +--+
| | +-- Anodonta_cygnea
| |
| | +-------- Anemina_arcaeformis
+--+ +-----+
| | | +----- Amuranodonta_kijaensis
| | +--+
| | | +-- Buldowskia_suifunica
| | +--+
| | +-- Buldowskia_shadini
+--+
| +-------- Beringiana_beringiana
| +--+
| | | +----- Sinanodonta_schrenkii
| | +--+
| | | +-- Sinanodonta_woodiana
+--+ +--+
| +-- Sinanodonta_lauta
|
| +-------- Cristaria_radiata
| |
+--+ +-- Cristaria_truncata
| +--+
| | +-- Cristaria_plicata
+--+
| +-- Cristaria_bellua
+--+
+-- Cristaria_clessini
Chen et al., 2023 [Publ 4302], fig. 1
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 2 |
2 |
0 |
2 |
38 |
36 |
3 |
-- |
+-- Pronodularia_japanensis
+--+
| +-- Lamprotula_leaii
+-----------------------------+
| | +-- Lampsilis_siliquoidea **
| +--+
| +-- Quadrula_quadrula **
|
| +-- Coelatura_aegyptiaca
--+ |
| +-----------------------------+-- Indonaia_andersoniana
| | |
| | +-- Parreysia_nagpoorensis
| |
| | +-- Anemina_arcaeformis
+--+ +--------------------------+
| | +-- Amuranodonta_kijaensis
| |
| | +----- Pletholophus_tenuis
| | +--------------------+
| | | | +-- Sinanodonta_jourdyi
+--+ | +--+
| | +-- Beringiana_beringiana
| |
| | +----------------------- Lanceolaria_gladiola
| | |
| | +----------------------- Acuticosta_chinensis
+--+ |
| +----------------------- Lepidodesma_languilati
| |
| +----------------------- Lanceolaria_oxyrhyncha
| |
| +----------------------- Cristaria_plicata
| |
| | +-- Alasmidonta_marginata
+--+ +--+
| | +-- Lasmigona_costata
+-----------------+
| | +-- Anodonta_anatina
| +--+
| +-- Pseudanodonta_complanata
|
| +-------------------- Aculamprotula_fibrosa
| |
| | +----- Middendorffinaia_mongolica
| | +-----------+
+--+ | | +-- Nodularia_douglasiae
| | +--+
| | +-- Nodularia_breviconcha
| |
| | +-- Cuneopsis_celtiformis
+--+ +-----------+
| | +-- Cuneopsis_heudei
| |
| | +-- Diaurora_aurorea
| | +--+
| | | +-- Diaurora_laeve
| +--------+
+--+ | +-- Schistodesmus_lampreyanus
| +--+
| +-- Schistodesmus_spinosus
|
| +-- Pseudocuneopsis_sichuanensis
| +--------+
| | +-- Pseudocuneopsis_capitata
| |
+--+ +-- Inversiunio_yanagawensis
| +-----+
| | +-- Inversiunio_reinianus
+--+
| +----- Tchangsinaia_piscicula
+--+
| +-- Unio_pictorum
+--+
+-- Unio_crassus
Chen et al., 2023 [Publ 4302], fig. 2
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 2 |
2 |
0 |
2 |
38 |
36 |
3 |
-- |
+-- Pronodularia_japanensis
+--+
| +-- Lamprotula_leaii
+--------------------------+
| | +-- Lampsilis_siliquoidea **
| +--+
| +-- Quadrula_quadrula **
|
| +----- Parreysia_nagpoorensis
--+ +-----------------------+
| | | +-- Coelatura_aegyptiaca
| | +--+
| | +-- Indonaia_andersoniana
| |
| | +-------------------------- Acuticosta_chinensis
+--+ |
| | +----------------------- Lepidodesma_languilati
| | |
| | | +-- Lanceolaria_gladiola
| | | +--------------+
| | | | +-- Lanceolaria_oxyrhyncha
| | | |
+--+ | | +-- Alasmidonta_marginata
| +-----+ +--+
| | | | +-- Lasmigona_costata
| | | +--------+
| | | | | +-- Anodonta_anatina
| | | | +--+
| | +--+ +-- Pseudanodonta_complanata
| | |
| | | +----------- Cristaria_plicata
+--+ | |
| +--+ +-- Anemina_arcaeformis
| | +-----+
| | | +-- Amuranodonta_kijaensis
| +--+
| | +----- Pletholophus_tenuis
| +--+
| | +-- Sinanodonta_jourdyi
| +--+
| +-- Beringiana_beringiana
|
| +-------------------- Aculamprotula_fibrosa
| |
| | +----- Middendorffinaia_mongolica
| | +-----------+
+--+ | | +-- Nodularia_douglasiae
| | +--+
| | +-- Nodularia_breviconcha
| |
| | +-- Cuneopsis_celtiformis
+--+ +-----------+
| | +-- Cuneopsis_heudei
| |
| | +-- Diaurora_aurorea
| | +--+
| | | +-- Diaurora_laeve
| +--------+
+--+ | +-- Schistodesmus_lampreyanus
| +--+
| +-- Schistodesmus_spinosus
|
| +-- Pseudocuneopsis_sichuanensis
| +--------+
| | +-- Pseudocuneopsis_capitata
| |
+--+ +-- Inversiunio_yanagawensis
| +-----+
| | +-- Inversiunio_reinianus
+--+
| +----- Tchangsinaia_piscicula
+--+
| +-- Unio_pictorum
+--+
+-- Unio_crassus
Araujo et al., 2009 [Publ 2589], fig. 7A
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 1 |
1 |
0 |
1 |
11 |
10 |
2 |
-- |
+----------------- Amblema_plicata **
|
--+ +-------------- Potomida_littoralis
| |
+--+ +----------- Nodularia_douglasiae
| |
| | +-- Unio_gibbus
+--+ |
| +-----+-- Unio_tumidus
| | |
| | +-- Anodonta_cygnea
+--+
| +----- Unio_pictorum[1]
| |
| +----- Unio_pictorum[2]
+--+
+----- Unio_mancus
|
| +-- Unio_tumidiformis
+--+
+-- Unio_crassus
Araujo et al., 2009 [Publ 2589], fig. 7B
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 1 |
1 |
0 |
1 |
11 |
10 |
2 |
-- |
+----------------------- Potomida_littoralis
|
--+ +-------------------- Amblema_plicata **
| |
+--+ +----------------- Anodonta_cygnea
| |
+--+ +-------------- Unio_gibbus
| |
+--+ +----------- Unio_tumidus
| |
| | +-- Unio_tumidiformis
+--+ +-----+
| | +-- Unio_crassus
| |
+--+ +-- Unio_pictorum[1]
| +--+
| | +-- Unio_mancus
+--+
| +-- Unio_pictorum[2]
+--+
+-- Unio_caffer
Araujo et al., 2009 [Publ 2589], fig. 7C
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 1 |
1 |
0 |
1 |
10 |
9 |
2 |
-- |
+----------------- Potomida_littoralis
|
--+ +-------------- Amblema_plicata **
| |
+--+ +----------- Anodonta_cygnea
| |
| | +-- Unio_gibbus
+--+ +-----+
| | +-- Unio_tumidus
| |
+--+ +-- Unio_tumidiformis
| +--+
| | +-- Unio_crassus
+--+
| +-- Unio_pictorum[1]
| |
+--+-- Unio_pictorum[2]
|
+-- Unio_mancus
Baker et al., 2004 [Publ 2685], fig. 2-3
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 1 |
1 |
0 |
1 |
11 |
0 |
0 |
-- |
-- Epioblasma_triquetra
Baker et al., 2003 [Publ 2690], fig. 5
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 1 |
1 |
0 |
1 |
7 |
0 |
0 |
-- |
-- Epioblasma_triquetra
Hoeh et al., 1996 [Publ 2694], fig. 2
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 1 |
1 |
0 |
1 |
4 |
2 |
1 |
-- |
+-- None[1]
+-----+
| +-- None[2]
--+
| +----- Fusconaia_flava **
+--+
| +-- Pyganodon_fragilis
+--+
+-- Pyganodon_grandis
Hoeh et al., 1998 [Publ 2695], fig. 3
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 1 |
1 |
0 |
1 |
4 |
1 |
1 |
-- |
-- Amblema_plicata
Hoeh et al., 1998 [Publ 2695], fig. 4
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 1 |
1 |
0 |
1 |
4 |
1 |
1 |
-- |
-- Amblema_plicata
Huff et al., 2004 [Publ 2697], fig. 2
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 1 |
1 |
0 |
1 |
13 |
2 |
1 |
-- |
+----- Lampsilis_cardium **
--+
| +-- Unio_ravoisieri
+--+
+-- Anodonta
Huff et al., 2004 [Publ 2697], fig. 4
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 1 |
1 |
0 |
1 |
10 |
2 |
1 |
-- |
+----- Lampsilis_cardium **
--+
| +-- Unio_ravoisieri
+--+
+-- Anodonta
Huff et al., 2004 [Publ 2697], fig. 5-6
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 1 |
1 |
0 |
1 |
11 |
2 |
1 |
-- |
+-- Unio_ravoisieri
|
--+-- Anodonta
|
+-- Lampsilis_cardium **
Campbell, 2000 [Publ 2724], fig. 1-5
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 1 |
1 |
0 |
1 |
2 |
1 |
1 |
-- |
+-- Utterbackia_imbecillis
--+
+-- Elliptio_complanata **
Cyr et al., 2007 [Publ 2745], fig. 2A
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 1 |
1 |
0 |
1 |
5 |
5 |
1 |
-- |
+----------- Lampsilis_ornata **
|
--+ +-------- Pyganodon_grandis
| |
+--+-------- Pyganodon[2]
|
| +----- Pyganodon[1]
+--+
| +-- Pyganodon_cataracta
+--+
+-- Pyganodon_fragilis
Khalloufi et al., 2011 [Publ 2748], fig. 2
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 1 |
1 |
0 |
1 |
10 |
10 |
2 |
-- |
+----------------- Amblema_plicata **
|
--+ +-------------- Potomida_littoralis
| |
| | +----- Unio_tumidus
+--+ +-----+
| | | +-- Unio_gibbus
| | +--+
| | +-- Unio_durieui
+--+
| +-- Unio_tumidiformis
| +-----+
| | +-- Unio_crassus
+--+
| +----- Unio
| |
+--+----- Unio_pictorum
|
| +-- Unio_delphinus
+--+
+-- Unio_mancus
Plazzi & Passamonti, 2010 [Publ 3354], fig. 1
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 1 |
1 |
0 |
1 |
1 |
1 |
1 |
-- |
+-- Pronodularia_japanensis
--+
+-- Lampsilis_ornata **
Plazzi & Passamonti, 2010 [Publ 3354], fig. 2-3
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 1 |
1 |
0 |
1 |
3 |
3 |
2 |
-- |
+----- Sinanodonta_woodiana
|
--+----- Lampsilis_ornata **
|
| +-- Sinohyriopsis_cumingii
+--+
+-- Pronodularia_japanensis
Jiang et al., 2010 [Publ 3438], fig. 5
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 1 |
1 |
0 |
1 |
3 |
3 |
2 |
-- |
+----- Lampsilis_ornata **
|
--+----- Cristaria_plicata
|
| +-- Pronodularia_japanensis
+--+
+-- Sinohyriopsis_cumingii
Prié & Puillandre, 2014 [Publ 3529], fig. 2
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 1 |
1 |
0 |
1 |
16 |
15 |
2 |
-- |
+-------------------------- Amblema_plicata **
|
| +----------------------- Potomida_littoralis
--+ |
| | +----- Anodonta_anatina
| | +--------------+
+--+ | | +-- Pseudanodonta_complanata
| | +--+
| | +-- Anodonta_cygnea
| |
+--+ +----- Unio_tumidus
| +-----------+
| | | +-- Unio_gibbus
| | +--+
| | +-- Unio_durieui
+--+
| +----- Unio_tumidiformis
| +--------+
| | | +-- Unio_crassus
| | +--+
+--+ +-- Unio_courtillieri
|
| +----------- Unio_delphinus
+--+
| +-------- Unio_elongatulus
+--+
| +----- Unio_mancus
+--+
| +-- Unio_pictorum
+--+
+-- Unio_ravoisieri
Marshall et al., 2014 [Publ 3593], fig. 1
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 1 |
1 |
0 |
1 |
13 |
2 |
1 |
-- |
-- Amblema_plicata
Inoue et al., 2014 [Publ 3635], fig. 3B
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 1 |
1 |
0 |
1 |
7 |
7 |
1 |
-- |
+----------- Amblema_plicata **
|
| +-------- Pyganodon_grandis
--+ |
| +-------- Pyganodon_cataracta
| |
+--+-------- Pyganodon_fragilis
|
+-------- Pyganodon
|
| +----- Platynaias_subviridis
+--+
| +-- Arcidens_confragosus
+--+
+-- Arcidens_wheeleri
Takeuchi et al., 2015 [Publ 3661], fig. 2
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 1 |
1 |
0 |
1 |
9 |
0 |
0 |
-- |
-- Lampsilis_cardium
Takeuchi et al., 2015 [Publ 3661], fig. 3
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 1 |
1 |
0 |
1 |
10 |
0 |
0 |
-- |
-- Lampsilis_cardium
Soroka, 2008 [Publ 3695], fig. 1
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 1 |
1 |
0 |
1 |
3 |
3 |
1 |
-- |
+-------- Fusconaia_flava **
--+
| +----- Sinanodonta_woodiana
+--+
| +-- Anodonta_anatina
+--+
+-- Pyganodon_grandis
Soroka, 2008 [Publ 3695], fig. 1-M
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 1 |
1 |
0 |
1 |
3 |
3 |
1 |
-- |
+-------- Fusconaia_flava **
--+
| +----- Sinanodonta_woodiana
+--+
| +-- Anodonta_anatina
+--+
+-- Pyganodon_grandis
Bolotov et al., 2016 [Publ 3743], fig. 2
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 1 |
1 |
0 |
1 |
29 |
3 |
1 |
-- |
+-------- Lampsilis_cardium **
--+
| +----- Anodonta_anatina
+--+
| +-- Unio_pictorum
+--+
+-- Unio_ravoisieri
Bolotov et al., 2016 [Publ 3743], fig. 4
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 1 |
1 |
0 |
1 |
29 |
3 |
1 |
-- |
+-------- Lampsilis_cardium **
--+
| +----- Anodonta_anatina
+--+
| +-- Unio_pictorum
+--+
+-- Unio_ravoisieri
Combosch et al., 2017 [Publ 3806], fig. 1
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 1 |
1 |
0 |
1 |
14 |
2 |
2 |
-- |
+----- Unio_pictorum
--+
| +-- Lampsilis_cardium **
+--+
+-- Scabies_crispata
Combosch et al., 2017 [Publ 3806], fig. 2
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 1 |
1 |
0 |
1 |
14 |
2 |
2 |
-- |
+----- Unio_pictorum
--+
| +-- Scabies_crispata
+--+
+-- Lampsilis_cardium **
Vikhrev et al., 2018 [Publ 3812], fig. 3
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 1 |
1 |
0 |
1 |
12 |
1 |
1 |
-- |
+-- Unio_ravoisieri
--+
+-- Lampsilis_cardium **
Huang et al., 2018 [Publ 3828], fig. 3, 5
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 1 |
1 |
0 |
1 |
28 |
5 |
3 |
-- |
+-- Lampsilis_cardium **
+--+
| +-- Scabies_crispata
--+
| +-- Unio_pictorum
+--+
+-- Aculamprotula_tientsinensis
Lopes-Lima et al., 2018 [Publ 3829], fig. 1
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 1 |
1 |
0 |
1 |
27 |
2 |
2 |
-- |
+----- Potomida_littoralis
--+
| +-- Lampsilis_cardium **
+--+
+-- Unio_pictorum
Lopes-Lima et al., 2018 [Publ 3829], fig. 3
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 1 |
1 |
0 |
1 |
27 |
2 |
2 |
-- |
+----- Potomida_littoralis
--+
| +-- Lampsilis_cardium **
+--+
+-- Unio_pictorum
Lemer et al., 2019 [Publ 4093], fig. 1
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 1 |
1 |
0 |
1 |
5 |
0 |
0 |
-- |
-- Lampsilis_cardium
Wu et al., 2020 [Publ 4121], fig. 2
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 1 |
1 |
0 |
1 |
33 |
10 |
3 |
-- |
+-- Lampsilis_cardium **
+--------------+
| +-- Scabies_crispata
|
| +-------- Sinosolenaia_carinata
--+ +-----+
| | | +----- Potomida_littoralis
| | +--+
| | | +-- Lamprotula_caveata
+--+ +--+
| +-- Lamprotula_leaii
|
| +----------- Unio_pictorum
+--+
| +-------- Aculamprotula_tortuosa
+--+
| +----- Aculamprotula_polysticta
+--+
| +-- Aculamprotula_fibrosa
+--+
+-- Aculamprotula_tientsinensis
Wu et al., 2020 [Publ 4121], fig. S1
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 1 |
1 |
0 |
1 |
33 |
10 |
3 |
-- |
+-- Lampsilis_cardium **
+--------------+
| +-- Scabies_crispata
|
| +-------- Sinosolenaia_carinata
--+ +-----+
| | | +----- Potomida_littoralis
| | +--+
| | | +-- Lamprotula_caveata
+--+ +--+
| +-- Lamprotula_leaii
|
| +----------- Unio_pictorum
+--+
| +-------- Aculamprotula_tortuosa
+--+
| +----- Aculamprotula_polysticta
+--+
| +-- Aculamprotula_tientsinensis
+--+
+-- Aculamprotula_fibrosa
Sano et al., 2020 [Publ 4138], fig. 2
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 1 |
1 |
0 |
1 |
72 |
49 |
3 |
-- |
+----------------- Lampsilis_cardium **
|
| +----- Sinohyriopsis_cumingii[2]
+--------------------+ +--------+
| | | | +-- Sinohyriopsis_schlegelii
| | | +--+
| +--+ +-- Sinohyriopsis_cumingii[1]
| |
| | +----------- Inversidens_brandtii
| | |
| +--+ +-- Ptychorhynchus_pfisteri
| | +-----+
| | | +-- Obovalis_omiensis
| +--+
| | +----- Lamprotula_leaii
| +--+
--+ | +-- Pronodularia_japanensis
| +--+
| +-- Potomida_littoralis
|
| +----- Buldowskia_iwakawai
| +-----------------------------+
| | | +-- Anemina_arcaeformis
| | +--+
| | +-- Anemina_euscaphys
| |
| | +-------------- Pletholophus_tenuis
| | |
| | | +----- Beringiana_fukuharai
| | +-----------------+ +-----+
+--+ | | | | +-- Beringiana_japonica
| | | | +--+
| | +--+ +-- Beringiana_beringiana
| | |
| | | +-------- Sinanodonta_lucida
| | +--+
| | | +----- Sinanodonta_tumens
| | +--+
| | | +-- Sinanodonta_lauta
+--+ +--+
| +-- Sinanodonta_woodiana
|
| +----------------------------- Cristaria_plicata
| |
| | +-- Anodonta_cygnea
| | +--------+
| | | +-- Anodonta_anatina
| | |
| | +--------------+ +----- Alasmidonta_varicosa
+--+ | | +--+
| | | | | +-- Prolasmidonta_heterodon
| | | | +--+
| | +--+ +-- Platynaias_compressa
| | |
| | | +----- Pyganodon_grandis
| | +--+
+--+ | +-- Utterbackia_imbecillis
| +--+
| +-- Utterbackia_peninsularis
|
| +----- Lanceolaria_grayii[1]
| +-----------------+
| | | +-- Lanceolaria_lanceolata
| | +--+
| | +-- Lanceolaria_grayii[2]
+--+
| +-------------------- Lepidodesma_languilati
| |
| | +----- Aculamprotula_tortuosa
| | +-----------+
+--+ | | +-- Aculamprotula_tientsinensis
| | +--+
| | +-- Aculamprotula_coreana
| |
+--+ +----- Unio_crassus
| +--------+
| | | +-- Unio_delphinus
| | +--+
| | +-- Unio_pictorum
| |
+--+ +-------- Inversiunio_jokohamensis
| +--+
| | | +----- Inversiunio_yanagawensis[2]
| | +--+
| | | +-- Inversiunio_reinianus
| | +--+
+--+ +-- Inversiunio_yanagawensis[1]
|
| +-- Tchangsinaia_piscicula
| +-----+
| | +-- Schistodesmus_lampreyanus
+--+
| +----- Nodularia_douglasiae[2]
+--+
| +-- Nodularia_nipponensis
+--+
+-- Nodularia_douglasiae[1]
Sano et al., 2020 [Publ 4138], fig. 3
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 1 |
1 |
0 |
1 |
71 |
48 |
3 |
-- |
+----- Sinohyriopsis_cumingii[2]
+--------+
| | +-- Sinohyriopsis_schlegelii
| +--+
| +-- Sinohyriopsis_cumingii[1]
--+
| +----------- Lampsilis_cardium **
| |
| | +----- Lamprotula_leaii
+--+ +--+
| | | +-- Potomida_littoralis
| | +--+
+--+ +-- Pronodularia_japanensis
|
| +----- Inversidens_brandtii
+--+
| +-- Obovalis_omiensis
+--+
+-- Ptychorhynchus_pfisteri
Sano et al., 2020 [Publ 4138], fig. 4
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 1 |
1 |
0 |
1 |
71 |
48 |
3 |
-- |
+----------------- Lampsilis_cardium **
|
| +----- Sinohyriopsis_cumingii[2]
+-----------------------+ +--------+
| | | | +-- Sinohyriopsis_schlegelii
| | | +--+
| +--+ +-- Sinohyriopsis_cumingii[1]
| |
| | +----------- Inversidens_brandtii
| | |
| +--+ +-- Obovalis_omiensis
| | +-----+
| | | +-- Ptychorhynchus_pfisteri
| +--+
| | +----- Lamprotula_leaii
| +--+
--+ | +-- Pronodularia_japanensis
| +--+
| +-- Potomida_littoralis
|
| +----- Buldowskia_iwakawai
| +--------------------------------+
| | | +-- Anemina_arcaeformis
| | +--+
| | +-- Anemina_euscaphys
| |
| | +-------- Sinanodonta_lucida
| | +--+
| | | | +----- Sinanodonta_tumens
| | | +--+
+--+ | | +-- Sinanodonta_woodiana
| +-----------------------+ +--+
| | | +-- Sinanodonta_lauta
| | |
| | | +-------- Pletholophus_tenuis
| | +--+
| | | +----- Beringiana_fukuharai
| | +--+
| | | +-- Beringiana_japonica
| | +--+
+--+ +-- Beringiana_beringiana
|
| +-------------------------------- Cristaria_plicata
| |
| | +-- Anodonta_cygnea
| | +--------+
| | | +-- Anodonta_anatina
| | |
| | +-----------------+ +----- Alasmidonta_varicosa
+--+ | | +--+
| | | | | +-- Prolasmidonta_heterodon
| | | | +--+
| | +--+ +-- Platynaias_compressa
| | |
| | | +----- Pyganodon_grandis
| | +--+
+--+ | +-- Utterbackia_imbecillis
| +--+
| +-- Utterbackia_peninsularis
|
| +-------------------------- Lepidodesma_languilati
| |
| | +----- Lanceolaria_grayii[1]
| | +-----------------+
+--+ | | +-- Lanceolaria_lanceolata
| | +--+
| | +-- Lanceolaria_grayii[2]
| |
+--+ +----- Aculamprotula_tortuosa
| +--------------+
| | | +-- Aculamprotula_tientsinensis
| | +--+
| | +-- Aculamprotula_coreana
| |
+--+ +----- Unio_crassus
| +-----------+
| | | +-- Unio_delphinus
| | +--+
| | +-- Unio_pictorum
+--+
| +-- Nodularia_douglasiae
| +-----------+
| | +-- Nodularia_nipponensis
| |
+--+ +-- Schistodesmus_lampreyanus
| +--------+
| | +-- Tchangsinaia_piscicula
+--+
| +-------- Inversiunio_jokohamensis
+--+
| +----- Inversiunio_yanagawensis[2]
+--+
| +-- Inversiunio_reinianus
+--+
+-- Inversiunio_yanagawensis[1]
Wu et al., 2020 [Publ 4147], fig. 4
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 1 |
1 |
0 |
1 |
13 |
10 |
2 |
-- |
+----------------- Lampsilis_cardium **
|
| +----------- Sinosolenaia_carinata
| +--+
--+ | | +-------- Potomida_littoralis
| | +--+
| | | +----- Lamprotula_leaii
| | +--+
+--+ | +-- Lamprotula_caveata
| +--+
| +-- Lamprotula_cornuumlunae
|
| +----------- Unio_pictorum
+--+
| +-------- Aculamprotula_tortuosa
+--+
| +----- Aculamprotula_scripta
+--+
| +-- Aculamprotula_tientsinensis
+--+
+-- Aculamprotula_fibrosa
| 
