|
Bivalvia | Unionoida | Unionoidea | Unionidae | Gonideinae tribe CONTRADENTINI Modell, 1942: cladomics
6 genera
Cladomics Summary
number of species = 35
number of subtaxa (rank: genus) = 6
analyzed in 56 cladograms in 33 publications
maximum number of terminals in an analysis = 23, mean = 7.21
maximum number of subtaxa (genus) in an analysis = 5, mean = 2.68
Basic Monophyly
number of cladograms that test basic monophyly (i.e., more than 1 included species) = 47
maximum number of included terminals = 23, mean = 8.4
maximum number of subtaxa (genus) = 5, mean = 3.0
proportion of cladograms that support basic monophyly = 0.96
Rigorous Monophyly
number of cladograms that test rigorous monophyly = 42
(Rigorous monophyly refers to cladograms with terminal species representing more than one subtaxon (genus) taxon and at least one rigorous outgroup (tribe).
maximum number of included terminals = 23, mean = 8.21
maximum number of subtaxa (genus) = 5, mean = 3.1
proportion of cladograms that support rigorous monophyly = 0.95
Pairwise Cladogram Clustering
The graphs below depict in two dimensions all pair-wise ingroup distances within the cladome. See the Cladmoics FARQ for more information.
Pfeiffer et al., 2019 [Publ 3937], fig. 4-5
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 5 |
5 |
3 |
5 |
52 |
12 |
5 |
0.0 |
*** Top Monophyly Cladogram ***
+----------------- Lamprotula_cornuumlunae
+--+
| | +-------------- Gonidea_angulata
| +--+
| | +----------- Indopseudodon_crebristriatus
| +--+
| | +-------- Bineurus_anodontinum
| +--+
| | +----- Pilsbryoconcha_exilis
| +--+
--+ | +-- Pseudodon_vondembuschianus
| +--+
| +-- Pseudodon_inoscularis
|
| +----- Ensidens_ingallsianus
| +-----+
| | | +-- Hyriopsis_bialata
| | +--+
| | +-- Rectidens_sumatrensis
+--------+
| +-- Trapezoideus_foliaceus **
| +-----+
| | +-- Physunio_superbus **
+--+
| +----- Yaukthwa_paiensis **
+--+
| +-- Lens_contradens **
+--+
+-- Solenaia_emarginata **
Wu et al., 2024 [Publ 4350], fig. 5a
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 7 |
7 |
5 |
4 |
31 |
29 |
7 |
4.0 |
*** Top Sister-Group Cladogram ***
+-- Khairuloconcha_lunbawangorum
+--------------+
| +-- Khairuloconcha_sahanae
|
| +-------- Ensidens_sagittarius
--+ +-----+
| | | +----- Rectidens_sumatrensis
| | +--+
| | | +-- Hyriopsis_bialata
| | +--+
+--+ +-- Hyriopsis_desowitzi
|
| +-- Physunio_superbus **
| +--------+
| | +-- Trapezoideus_foliaceus **
| |
+--+ +-- Lens_contradens **
| +-----+
| | +-- Lens_eximius **
+--+
| +----- Yaukthwa_elongatula **
+--+
| +-- Yaukthwa_inlenensis **
+--+
+-- Yaukthwa_paiensis **
Bolotov et al., 2020 [Publ 4100], fig. 1
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 23 |
23 |
21 |
4 |
97 |
51 |
5 |
20.0 |
+----------- Hyriopsis_desowitzi
|
+--+ +-------- Rectidens_sumatrensis
| | |
| +--+ +-- Hyriopsis_khoratensis
| | +--+
| | | +-- Hyriopsis_bialata
| +--+
| | +-- Hyriopsis_myersiana
+--------+ +--+
| | +-- Hyriopsis_panhai
| |
| | +-- Ensidens_telus
| | +-----+
| | | +-- Ensidens_ingallsianus
| | |
| +-----+ +-- Ensidens_jaculus
| | +--+
| | | +-- Ensidens_spiculus
| +--+
| | +-- Ensidens_dugasti
| +--+
| +-- Ensidens_sagittarius
+-----+
| | +-- Trapezoideus_lenya **
| | +-----+
| | | +-- Trapezoideus_foliaceus **
| | +-----------+
| | | | +----- Physunio_superbus **
| | | +--+
| | | | +-- Physunio_modelli **
| | | +--+
| | | +-- Physunio **
| | |
| | | +-- Yaukthwa_elongatula **
| | | +-----------+
| | | | +-- Yaukthwa_baniensis **
| +--+ |
| | +--+ +----- Yaukthwa_inlenensis **
| | | | +-----+
| | | | | | +-- Yaukthwa_avaensis **
| | | | | +--+
| | | +--+ +-- Yaukthwa_paiensis **
| | | |
| | | | +-------- Yaukthwa_panhai **
| | | +--+
| | | | +----- Yaukthwa_rectangularis **
| +--+ +--+
| | | +-- Yaukthwa_zayleymanensis **
| | +--+
| | +-- Yaukthwa_nesemanni **
| |
| | +----- Lens_misellus **
| | +-----+
| | | | +-- Lens_contradens **
| | | +--+
| | | +-- Lens_micropterus **
| +-----+
| | +----- Lens_pallegoixi **
| | +--+
--+ | | | +-- Lens_comptus **
| | | +--+
| +--+ +-- Lens_novoselovi **
| |
| | +----- Lens_maenamensis **
| +--+
| | +-- Lens_eximius **
| +--+
| +-- Lens_rolfbrandti **
|
| +-- Sinosolenaia_iridinea
| |
| +-- Sinosolenaia_recognita[1]
| |
| +-- Sinosolenaia_carinata
| |
| +-- Leguminaia_wheatleyi
| |
| +-- Microcondylaea_bonellii
| |
| +-- Gonidea_angulata
| |
| +-- Ptychorhynchus_pfisteri
| |
| +-- Parvasolenaia_rivularis
| |
| +-----------------------+-- Lamprotula_leaii
| | |
| | +-- Lamprotula
| | |
| | +-- Lamprotula_caveata
| | |
| | +-- Potomida_littoralis
| | |
| | +-- Pronodularia_japanensis
| | |
| | +-- Sinohyriopsis_cumingii
| | |
+--+ +-- Chamberlainia_somsakpanhai
| |
| +-- Sinosolenaia_recognita[2]
| |
| +-- Sinosolenaia_oleivora
|
| +-------- Indopseudodon_kayinensis
| +--------------+
| | | +----- Indopseudodon_indawgyiensis
| | +--+
| | | +-- Indopseudodon_salwenianus
| | +--+
| | +-- Indopseudodon_bogani
+--+
| +----- Bineurus_loeiensis
| +--------------+
| | | +-- Bineurus_exilis
| | +--+
| | +-- Bineurus_mouhotii
| |
+--+ +----------------- Nyeinchanconcha_nyeinchani
| |
| | +-- Thaiconcha_callifera
| | +-----+
| | | +-- Thaiconcha_munelliptica
+--+ +-----+
| | | +----- Pilsbryoconcha_exilis[2]
| | +--+
| | | +-- Pilsbryoconcha_exilis[1]
| | +--+
| | +-- Pilsbryoconcha
+--+
| +-------- Pseudodon_lenyanensis
| +--+
| | | +----- Pseudodon_vagulus
| | +--+
| | | +-- Pseudodon_vondembuschianus
| | +--+
+--+ +-- Pseudodon_mekongi
|
| +-- Sundadontina_taskaevi
| +-----+
| | +-- Sundadontina_brandti
+--+
| +----- Sundadontina_tumida
+--+
| +-- Sundadontina_cumingii
+--+
+-- Sundadontina_tanintharyiensis
Bolotov et al., 2020 [Publ 4100], fig. S1
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 23 |
23 |
21 |
4 |
91 |
51 |
5 |
22.0 |
+-------------- Hyriopsis_desowitzi
+--+
| | +----------- Rectidens_sumatrensis
| +--+
| | +-------- Hyriopsis_panhai
| +--+
| | +----- Hyriopsis_myersiana
| +--+
+--------+ | +-- Hyriopsis_khoratensis
| | +--+
| | +-- Hyriopsis_bialata
| |
| | +-- Ensidens_telus
| | +-----+
| | | +-- Ensidens_ingallsianus
| | |
| +--------+ +-- Ensidens_dugasti
| | +--+
| | | +-- Ensidens_sagittarius
| +--+
+--+ | +-- Ensidens_spiculus
| | +--+
| | +-- Ensidens_jaculus
| |
| | +----- Physunio **
| | +-----------------+
| | | | +-- Physunio_modelli **
| | | +--+
| | | +-- Physunio_superbus **
| | |
| | | +-- Trapezoideus_lenya **
| | | +-----------------+
| +--+ | +-- Trapezoideus_foliaceus **
| | |
| | | +----- Lens_misellus **
| | | +-----+
| | | | | +-- Lens_contradens **
| | | | +--+
| | | | +-- Lens_micropterus **
| +--+ +-----+
| | | | +----- Lens_maenamensis **
| | | | +--+
| | | | | | +-- Lens_eximius **
| | | | | +--+
| | | +--+ +-- Lens_rolfbrandti **
| | | |
| | | | +----- Lens_pallegoixi **
| +--+ +--+
| | | +-- Lens_novoselovi **
| | +--+
| | +-- Lens_comptus **
| |
| | +-- Yaukthwa_elongatula **
| | +-----------+
| | | +-- Yaukthwa_baniensis **
| | |
| +--+ +----- Yaukthwa_inlenensis **
| | +-----+
--+ | | | +-- Yaukthwa_avaensis **
| | | +--+
| +--+ +-- Yaukthwa_paiensis **
| |
| | +-------- Yaukthwa_panhai **
| +--+
| | +----- Yaukthwa_rectangularis **
| +--+
| | +-- Yaukthwa_zayleymanensis **
| +--+
| +-- Yaukthwa_nesemanni **
|
| +-- Sinohyriopsis_cumingii
| +--+
| | +-- Chamberlainia_somsakpanhai
| |
| +--+ +-- Lamprotula_leaii
| | | |
| | | +-- Lamprotula
| | | |
| | +--+-- Lamprotula_caveata
| | |
| | +-- Potomida_littoralis
| | |
| | +-- Pronodularia_japanensis
| +-----------------+
| | | +-- Sinosolenaia_iridinea
| | | |
| | | +-- Sinosolenaia_recognita[1]
| | | |
| | | +-- Sinosolenaia_carinata
| | | |
| | | +-- Leguminaia_wheatleyi
| | | |
| | | +-- Microcondylaea_bonellii
| | +-----+
| | +-- Gonidea_angulata
| | |
| | +-- Ptychorhynchus_pfisteri
+--+ |
| +-- Parvasolenaia_rivularis
| |
| +-- Sinosolenaia_recognita[2]
| |
| +-- Sinosolenaia_oleivora
|
| +-------- Indopseudodon_kayinensis
| +--------------+
| | | +----- Indopseudodon_indawgyiensis
| | +--+
| | | +-- Indopseudodon_salwenianus
| | +--+
| | +-- Indopseudodon_bogani
+--+
| +----- Bineurus_loeiensis
| +--------------+
| | | +-- Bineurus_exilis
| | +--+
| | +-- Bineurus_mouhotii
| |
+--+ +----------------- Nyeinchanconcha_nyeinchani
| |
| | +-- Thaiconcha_callifera
| | +-----+
| | | +-- Thaiconcha_munelliptica
+--+ +-----+
| | | +----- Pilsbryoconcha_exilis[2]
| | +--+
| | | +-- Pilsbryoconcha_exilis[1]
| | +--+
| | +-- Pilsbryoconcha
+--+
| +-------- Pseudodon_lenyanensis
| +--+
| | | +----- Pseudodon_vagulus
| | +--+
| | | +-- Pseudodon_vondembuschianus
| | +--+
+--+ +-- Pseudodon_mekongi
|
| +-- Sundadontina_tanintharyiensis
| +-----+
| | +-- Sundadontina_cumingii
+--+
| +----- Sundadontina_tumida
+--+
| +-- Sundadontina_taskaevi
+--+
+-- Sundadontina_brandti
Jeratthitikul et al., 2019 [Publ 3911], fig. 4
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 19 |
19 |
11 |
2 |
3 |
2 |
2 |
18.0 |
+-------------------------- Ensidens_ingallsianus
|
| +-- Physunio_modelli **
--+ +--------------------+
| | +-- Physunio_superbus **
| |
| | +-- Lens_maenamensis **
+--+ +-----------------+
| | +-- Lens_rolfbrandti **
| |
| | +----------------- Lens_contradens[12] **
+--+ |
| | +----- Lens_micropterus[3] **
| | |
| +-----------+ +-- Lens_contradens[10] **
| | | |
+--+ +--+-- Lens_micropterus[2] **
| |
| +-- Lens_contradens[11] **
|
| +-------------- Lens_contradens[9] **
| |
+--+ +----------- Lens_contradens[8] **
| |
+--+ +-------- Lens_contradens[7] **
| |
+--+ +----- Lens_contradens[6] **
| |
| | +-- Lens_contradens[1] **
+--+ |
| +-- Lens_contradens[2] **
| |
| +-- Lens_contradens[3] **
+--+
+-- Lens_contradens[4] **
|
+-- Lens_micropterus[1] **
|
+-- Lens_contradens[5] **
Konopleva et al., 2019 [Publ 4041], fig. 3
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 19 |
19 |
17 |
4 |
16 |
16 |
2 |
18.0 |
+-- Indopseudodon_indawgyiensis
|
+--------------------+-- Indopseudodon_bogani
| |
| +-- Indopseudodon_salwenianus
|
| +-- Ensidens_ingallsianus
| |
| +-- Ensidens_telus
| |
| +-- Ensidens_dugasti
| |
| +-- Hyriopsis_khoratensis
| |
| +-- Hyriopsis_bialata
| |
--+ +-- Hyriopsis_myersiana
| |
| +-----------------+-- Hyriopsis_desowitzi
| | |
| | +-- Rectidens_sumatrensis
| | |
| | +-- Hyriopsis_panhai
| | |
| | +-- Ensidens_jaculus[1]
| | |
| | +-- Ensidens_jaculus[2]
| | |
| | +-- Ensidens_spiculus
| | |
+--+ +-- Ensidens_sagittarius
|
| +----------- Yaukthwa_elongatula **
| |
| +-----+ +-- Yaukthwa_inlenensis **
| | | +-----+
| | | | +-- Yaukthwa_paiensis **
| | +--+
| | | +----- Yaukthwa_panhai **
| | +--+
| | | +-- Yaukthwa_nesemanni **
| | +--+
+--+ +-- Yaukthwa_rectangularis **
|
| +-------- Trapezoideus_foliaceus **
| +-----+
| | | +----- Physunio_superbus **
| | +--+
| | | +-- Physunio_modelli **
| | +--+
| | +-- Physunio **
+--+
| +----- Lens_misellus **
| +-----+
| | | +-- Lens_contradens **
| | +--+
| | +-- Lens_micropterus **
+--+
| +----- Lens_pallegoixi **
| +--+
| | | +-- Lens_novoselovi **
| | +--+
+--+ +-- Lens_comptus **
|
| +----- Lens_maenamensis **
+--+
| +-- Lens_eximius **
+--+
+-- Lens_rolfbrandti **
Konopleva et al., 2023 [Publ 4262], fig. 8
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 19 |
19 |
17 |
4 |
65 |
51 |
4 |
17.0 |
+-- Gonidea_angulata
+--+
| +-- Leguminaia_wheatleyi
+-----------------------+
| | +-- Lamprotula_leaii
| +--+
| +-- Potomida_littoralis
|
| +-- Ensidens_ingallsianus
| +-----+
| | +-- Ensidens_telus
| |
| +--------------+ +-- Ensidens_jaculus
| | | +--+
| | | | +-- Ensidens_spiculus[1]
| | +--+
| | | +-- Ensidens_spiculus[2]
| +--+ +--+
| | | +-- Ensidens_dugasti
--+ | |
| | | +-- Hyriopsis_desowitzi
| | | +-----------------+
| | | | +-- Hyriopsis_delaportei
| | +--+
| | | +----------------- Rectidens_sumatrensis
| | | |
| | +--+ +-------------- Hyriopsis_panhai
| | | |
| | +--+ +----------- Hyriopsis_myersiana
| | | |
| | +--+ +-- Hyriopsis_khoratensis
| | | +-----+
| | | | +-- Hyriopsis_bialata
| | +--+
| | | +----- Hyriopsis_sakhonensis
+--+ +--+
| | +-- Hyriopsis_phuphaniensis
| +--+
| +-- Hyriopsis_kratiensis
|
| +----- Physunio_modelli **
| +-----+
| | | +-- Physunio_superbus **
| | +--+
| | +-- Physunio_massini **
| +-----+
| | | +-- Trapezoideus_thachiadensis **
| | | +-----+
| | | | +-- Trapezoideus_subviridis **
| | +--+
| | | +----- Trapezoideus_mitanensis **
| | +--+
| | | +-- Trapezoideus_foliaceus **
+--------+ +--+
| +-- Trapezoideus_lenya **
|
| +-- Yaukthwa_nesemanni **
| +-----------+
| | +-- Yaukthwa_elongatula **
| |
| | +----- Lens_misellus **
+--+ +-----+
| | | +-- Lens_contradens **
| | +--+
| | +-- Lens_micropterus **
+--+
| +----- Lens_pallegoixi **
| +--+
| | | +-- Lens_novoselovi **
| | +--+
+--+ +-- Lens_comptus **
|
| +----- Lens_maenamensis **
+--+
| +-- Lens_eximius **
+--+
+-- Lens_rolfbrandti **
Konopleva et al., 2019 [Publ 3900], fig. 1
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 16 |
16 |
14 |
4 |
36 |
22 |
2 |
17.0 |
+-- Ensidens_ingallsianus
+-----+
| +-- Ensidens_telus
+-----+
| | +----- Ensidens_sagittarius
| +--+
| | +-- Ensidens_jaculus
+-----------+ +--+
| | +-- Ensidens_spiculus
| |
| | +----------- Hyriopsis_desowitzi
| | |
| +--+----------- Rectidens_sumatrensis
| |
| | +-------- Hyriopsis_panhai
| +--+
| | +----- Hyriopsis_myersiana
| +--+
--+ | +-- Hyriopsis_khoratensis
| +--+
| +-- Hyriopsis_bialata
|
| +-- Yaukthwa_paiensis **
| +-----+
| | +-- Yaukthwa_inlenensis **
| +--------------+
| | | +----- Yaukthwa_panhai **
| | +--+
| | | +-- Yaukthwa_nesemanni **
| | +--+
+--+ +-- Yaukthwa_rectangularis **
|
| +----- Physunio_superbus **
| +--------------+
| | | +-- Physunio_modelli **
| | +--+
+--+ +-- Physunio **
|
| +----------------- Trapezoideus_foliaceus **
| |
+--+ +-- Lens_contradens **
| +-----------+
| | +-- Lens_micropterus **
+--+
| +----------- Lens_misellus **
+--+
| +-------- Lens_eximius **
+--+
| +----- Lens_pallegoixi **
+--+
| +-- Lens_comptus **
+--+
+-- Lens_novoselovi **
Pfeiffer et al., 2021 [Publ 4126], fig. 9
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 15 |
15 |
13 |
5 |
14 |
14 |
2 |
10.0 |
+-------------------------- Lamprotula_cornuumlunae
|
| +-- Ensidens_ingallsianus
| +--------+
| | +-- Ensidens_telus
| +--------+
| | | +-------- Ensidens_spiculus
| | +--+
| | | +----- Ensidens_dugasti
| | +--+
| +--+ | +-- Ensidens_jaculus
--+ | | +--+
| | | +-- Ensidens_sagittarius
| | |
| | | +----------------- Rectidens_sumatrensis
| | +--+
| | | +-------------- Hyriopsis_delaportei
| | +--+
| | | +----------- Hyriopsis_myersiana
| | +--+
| | | +-------- Hyriopsis_bialata
| | +--+
| | | +----- Hyriopsis_khoratensis
+--+ +--+
| | +-- Hyriopsis_kratiensis
| +--+
| +-- Hyriopsis_sakhonensis
|
| +-- Trapezoideus_subviridis **
| +-----+
| | +-- Trapezoideus_foliaceus **
| +-----------+
| | | +----- Physunio_superbus **
| | +--+
| | | +-- Physunio_modelli **
| | +--+
+--+ +-- Physunio_pellucidus **
|
| +----------------- Yaukthwa_paiensis **
| |
| | +-------------- Solenaia_emarginata **
+--+ |
| | +-------- Lens_misellus **
| | +--+
+--+ | | +----- Lens_inornatus **
| | +--+
| | | +-- Lens_contradens **
| | +--+
+--+ +-- Lens_micropterus **
|
| +-- Lens_maenamensis **
| +--+
| | +-- Lens_eximius **
+-----+
| +-- Lens_pallegoixi **
+--+
+-- Lens_comptus **
Bolotov et al., 2022 [Publ 4250], fig. S5
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 15 |
15 |
12 |
3 |
0 |
0 |
0 |
14.0 |
+-- Physunio_modelli **
+-----+
| +-- Physunio_superbus **
+--------+
| | +----- Trapezoideus_foliaceus **
| +--+
| | +-- Trapezoideus_lenya **
| +--+
| +-- Trapezoideus_mitanensis **
--+
| +-- Yaukthwa_elongatula **
| +-----------+
| | +-- Yaukthwa_baniensis **
| |
| | +-------- Yaukthwa_panhai **
+--+ +--+
| | | +----- Yaukthwa_rectangularis **
| | +--+
| | | +-- Yaukthwa_nesemanni **
+--+ +--+
| +-- Yaukthwa_zayleymanensis **
|
| +----- Yaukthwa_dalliana **
| |
+-----+ +-- Yaukthwa_paiensis **
| |
+--+-- Yaukthwa_inlenensis **
|
+-- Yaukthwa_avaensis **
Zieritz et al., 2020 [Publ 4102], fig. 4
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 14 |
14 |
12 |
4 |
17 |
17 |
6 |
15.0 |
+-- Pilsbryoconcha_carinifera
+-----------------------+
| +-- Indopseudodon_indawgyiensis
|
| +-------------- Khairuloconcha_lunbawangorum
| |
| | +-- Ensidens_telus
| | +-----+
| | | +-- Ensidens_ingallsianus
| +--------+ +--+
| | | | | +----- Ensidens_sagittarius
--+ | | | +--+
| | | | | +-- Ensidens_spiculus
| | | | +--+
| | +--+ +-- Ensidens_jaculus
| | |
| | | +-------- Hyriopsis_desowitzi
| | | |
| | | +-------- Rectidens_sumatrensis
| | +--+
| | | +----- Hyriopsis_panhai
+--+ | |
| +--+ +-- Hyriopsis_khoratensis
| | |
| +--+-- Hyriopsis_bialata
| |
| +-- Hyriopsis_myersiana
|
| +-- Physunio_modelli **
| +-----------------+
| | +-- Physunio_superbus **
| |
| | +-- Yaukthwa_paiensis **
| | +-----+
+--+ | +-- Yaukthwa_inlenensis **
| +--------+
| | | +----- Yaukthwa_panhai **
| | +--+
| | | +-- Yaukthwa_rectangularis **
+--+ +--+
| +-- Yaukthwa_nesemanni **
|
| +-------------- Trapezoideus_foliaceus **
| |
+--+ +-- Lens_contradens **
| +--------+
| | +-- Lens_misellus **
+--+
| +-------- Lens_eximius **
+--+
| +----- Lens_pallegoixi **
+--+
| +-- Lens_comptus **
+--+
+-- Lens_novoselovi **
Bolotov et al., 2017 [Publ 3761], fig. S1
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 12 |
12 |
10 |
3 |
129 |
39 |
5 |
12.0 |
+-- Ensidens_telus
+-----+
| +-- Ensidens_ingallsianus
+-----+
| | +----- Ensidens_sagittarius
| +--+
| | +-- Ensidens_spiculus
| +--+
+--+ +-- Ensidens_jaculus
| |
| | +-- Hyriopsis_bialata
| | +--------+
| | | +-- Hyriopsis_khoratensis
| +--+
| | +-------- Hyriopsis_panhai
| +--+
| | +----- Hyriopsis_myersiana
| +--+
--+ | +-- Rectidens_sumatrensis
| +--+
| +-- Hyriopsis_desowitzi
|
| +----- Physunio **
| +--------+
| | | +-- Physunio_modelli **
| | +--+
| | +-- Physunio_superbus **
| |
+--+ +-------- Yaukthwa_inlenensis **
| +--+
| | | +----- Yaukthwa_panhai **
| | +--+
| | | +-- Yaukthwa_rectangularis **
| | +--+
+--+ +-- Yaukthwa_nesemanni **
|
| +-- Lens_contradens **
| +-----+
| | +-- Lens_micropterus **
+--+
| +----- Lens_eximius **
+--+
| +-- Lens_comptus **
+--+
+-- Lens_novoselovi **
Bolotov et al., 2017 [Publ 3764], fig. S1
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 12 |
12 |
10 |
3 |
119 |
41 |
5 |
12.0 |
+-- Ensidens_telus
+-----+
| +-- Ensidens_ingallsianus
+--+
| | +----- Ensidens_jaculus
| +--+
| | +-- Ensidens_spiculus
| +--+
+-----+ +-- Ensidens_sagittarius
| |
| | +-------- Hyriopsis_desowitzi
| | |
| | +-------- Rectidens_sumatrensis
| +--+
| | +----- Hyriopsis_panhai
| | |
| +--+ +-- Hyriopsis_myersiana
| | |
--+ +--+-- Hyriopsis_bialata
| |
| +-- Hyriopsis_khoratensis
|
| +----- Physunio **
| +--------+
| | | +-- Physunio_superbus **
| | +--+
| | +-- Physunio_modelli **
| |
+--+ +-------- Yaukthwa_inlenensis **
| +--+
| | | +----- Yaukthwa_panhai **
| | +--+
| | | +-- Yaukthwa_nesemanni **
| | +--+
+--+ +-- Yaukthwa_rectangularis **
|
| +-- Lens_contradens **
| +-----+
| | +-- Lens_micropterus **
+--+
| +----- Lens_eximius **
+--+
| +-- Lens_comptus **
+--+
+-- Lens_novoselovi **
Huang et al., 2019 [Publ 3843], fig. 2
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 12 |
12 |
10 |
3 |
192 |
47 |
5 |
12.0 |
+----------- Hyriopsis_desowitzi
|
+----------- Rectidens_sumatrensis
|
| +-- Hyriopsis_bialata
| |
| +-- Hyriopsis_myersiana
+-----+--------+
| | +-- Hyriopsis_panhai
| | |
| | +-- Hyriopsis_khoratensis
| |
| | +-- Ensidens_ingallsianus
| | +-----+
| | | +-- Ensidens_telus
| +--+
| | +----- Ensidens_sagittarius
--+ +--+
| | +-- Ensidens_spiculus
| +--+
| +-- Ensidens_jaculus
|
| +----- Physunio **
| +--------+
| | | +-- Physunio_modelli **
| | +--+
| | +-- Physunio_superbus **
| |
+--+ +-------- Yaukthwa_inlenensis **
| +--+
| | | +----- Yaukthwa_panhai **
| | +--+
| | | +-- Yaukthwa_nesemanni **
| | +--+
+--+ +-- Yaukthwa_rectangularis **
|
| +-- Lens_contradens **
| +-----+
| | +-- Lens_micropterus **
+--+
| +----- Lens_eximius **
+--+
| +-- Lens_comptus **
+--+
+-- Lens_novoselovi **
Huang et al., 2019 [Publ 3843], fig. S3
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 12 |
12 |
10 |
3 |
189 |
46 |
5 |
12.0 |
+-- Ensidens_telus
+-----+
| +-- Ensidens_ingallsianus
+-----+
| | +----- Ensidens_sagittarius
| +--+
| | +-- Ensidens_spiculus
+--+ +--+
| | +-- Ensidens_jaculus
| |
| | +----------- Hyriopsis_desowitzi
| | |
| +--+ +-------- Rectidens_sumatrensis
| | |
| +--+ +-- Hyriopsis_panhai
| | +--+
| | | +-- Hyriopsis_khoratensis
--+ +--+
| | +-- Hyriopsis_bialata
| +--+
| +-- Hyriopsis_myersiana
|
| +----- Physunio **
| +--------+
| | | +-- Physunio_superbus **
| | +--+
| | +-- Physunio_modelli **
| |
+--+ +-------- Yaukthwa_inlenensis **
| +--+
| | | +----- Yaukthwa_panhai **
| | +--+
| | | +-- Yaukthwa_nesemanni **
| | +--+
+--+ +-- Yaukthwa_rectangularis **
|
| +-- Lens_contradens **
| +-----+
| | +-- Lens_micropterus **
+--+
| +----- Lens_eximius **
+--+
| +-- Lens_novoselovi **
+--+
+-- Lens_comptus **
Huang et al., 2019 [Publ 3843], fig. S4
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 12 |
12 |
7 |
3 |
190 |
46 |
5 |
13.0 |
+-------------- Hyriopsis_desowitzi
|
+-------------- Hyriopsis_myersiana
|
+-------------- Hyriopsis_panhai
|
+--+ +-- Hyriopsis_bialata
| +-----------+
| | +-- Hyriopsis_khoratensis
| |
| | +----------- Rectidens_sumatrensis
| | |
| +--+ +-- Ensidens_telus
| | +-----+
| | | +-- Ensidens_ingallsianus
| +--+
| | +----- Ensidens_sagittarius
| +--+
--+ | +-- Ensidens_spiculus
| +--+
| +-- Ensidens_jaculus
|
| +----------- Physunio_modelli **
| |
| +----------- Lens **
| |
| +----------- Physunio_superbus **
| |
| | +-------- Yaukthwa_inlenensis **
| +--+
+-----+ | +----- Yaukthwa_panhai **
| +--+
| | +-- Yaukthwa_nesemanni **
| +--+
| +-- Yaukthwa_rectangularis **
|
| +-- Lens_comptus **
| +-----+
| | +-- Lens_novoselovi **
+--+
| +----- Lens_eximius **
+--+
| +-- Lens_contradens **
+--+
+-- Lens_micropterus **
Bolotov et al., 2022 [Publ 4250], fig. 8
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 12 |
12 |
10 |
2 |
37 |
11 |
3 |
13.0 |
+-- Pseudodon_lenyanensis
+--------+
| +-- Sundadontina_tanintharyiensis
+--------+
| | +-------- Indopseudodon_kayinensis
| +--+
| | +----- Indopseudodon_indawgyiensis
| +--+
| | +-- Indopseudodon_salwenianus
| +--+
--+ +-- Indopseudodon_bogani
|
| +----- Trapezoideus_mitanensis **
| +-----------+
| | | +-- Trapezoideus_foliaceus **
| | +--+
| | +-- Trapezoideus_lenya **
+--+
| +-- Yaukthwa_elongatula **
| +-----------+
| | +-- Yaukthwa_baniensis **
| |
+--+ +----- Yaukthwa_inlenensis **
| +-----+
| | | +-- Yaukthwa_paiensis **
| | +--+
+--+ +-- Yaukthwa_avaensis **
|
| +-------- Yaukthwa_panhai **
+--+
| +----- Yaukthwa_rectangularis **
+--+
| +-- Yaukthwa_nesemanni **
+--+
+-- Yaukthwa_zayleymanensis **
Bolotov et al., 2017 [Publ 3761], fig. S2
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 11 |
11 |
9 |
3 |
130 |
40 |
5 |
13.0 |
+-------- Lens_contradens **
+-----+
| | +----- Lens_eximius **
| +--+
| | +-- Lens_comptus **
| +--+
+--+ +-- Lens_novoselovi **
| |
| | +----- Physunio **
| | +-----+
| | | | +-- Physunio_modelli **
| | | +--+
| +--+ +-- Physunio_superbus **
| |
| | +-------- Yaukthwa_inlenensis **
| +--+
| | +----- Yaukthwa_panhai **
--+ +--+
| | +-- Yaukthwa_rectangularis **
| +--+
| +-- Yaukthwa_nesemanni **
|
| +----- Hyriopsis_panhai
| |
| +--------+----- Hyriopsis_khoratensis
| | |
| | | +-- Hyriopsis_myersiana
| | +--+
+--+ +-- Hyriopsis_bialata
|
| +----------- Hyriopsis_desowitzi
| |
| +----------- Rectidens_sumatrensis
+--+
| +-- Ensidens_telus
| +-----+
| | +-- Ensidens_ingallsianus
+--+
| +----- Ensidens_sagittarius
+--+
| +-- Ensidens_spiculus
+--+
+-- Ensidens_jaculus
Bolotov et al., 2018 [Publ 3826], fig. 2
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 11 |
11 |
0 |
3 |
74 |
25 |
4 |
11.0 |
+-- Ensidens_telus
|
+-- Ensidens_ingallsianus
|
+-- Ensidens_jaculus
|
+-- Hyriopsis_desowitzi
+--+
| +-- Hyriopsis_myersiana
| |
| +-- Hyriopsis_khoratensis
| |
| +-- Ensidens_spiculus
| |
| +-- Ensidens_sagittarius
|
--+ +-- Lens_eximius **
| |
| +-- Physunio_modelli[1] **
| |
| +-- Yaukthwa_nesemanni **
| |
| +-- Yaukthwa_panhai **
| |
| +-- Lens_comptus **
| |
+--+-- Lens_novoselovi **
|
+-- Physunio_modelli[2] **
|
+-- Physunio **
|
+-- Yaukthwa_inlenensis **
|
+-- Lens_contradens **
|
+-- Lens_micropterus **
Pfeiffer et al., 2021 [Publ 4126], fig. 12
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 11 |
11 |
8 |
2 |
0 |
0 |
0 |
12.0 |
+-------------- Solenaia_emarginata **
|
| +-- Lens_contradens **
| +--------+
--+ | +-- Lens_micropterus **
| |
| | +-- Lens_inornatus **
| +--------+
+--+ +-- Lens_misellus **
|
| +----- Lens_maenamensis **
| +--+
| | | +-- Lens_rolfbrandti **
| | +--+
+--+ +-- Lens_eximius **
|
| +----- Lens_pallegoixi **
+--+
| +-- Lens_comptus **
+--+
+-- Lens_novoselovi **
Pfeiffer et al., 2021 [Publ 4126], fig. 18
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 10 |
10 |
7 |
3 |
0 |
0 |
0 |
8.0 |
+-------------- Yaukthwa_paiensis **
|
| +----- Trapezoideus_foliaceus **
--+ +-----+
| | | +-- Trapezoideus_lenya **
| | +--+
| | +-- Trapezoideus_subviridis **
+--+
| +-- Physunio_massini **
| +-----+
| | +-- Physunio_superbus **
+--+
| +----- Physunio_modelli[3] **
| |
+--+ +-- Physunio_modelli[1] **
| |
+--+-- Physunio_modelli[2] **
|
+-- Physunio_pellucidus **
Jeratthitkul et al., 2021 [Publ 4212], fig. 4
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 10 |
10 |
8 |
4 |
11 |
11 |
1 |
8.0 |
+-------- Trapezoideus_foliaceus **
+-----+
| | +----- Physunio_superbus **
| +--+
| | +-- Physunio_pellucidus **
| +--+
+-----+ +-- Physunio_modelli **
| |
| | +-- Yaukthwa_nesemanni **
| | +--------+
| | | +-- Yaukthwa_panhai **
| +--+
| | +-------- Lens_contradens **
| +--+
| | +----- Lens_novoselovi **
--+ +--+
| | +-- Lens_rolfbrandti **
| +--+
| +-- Lens_maenamensis **
|
| +-- Ensidens_ingallsianus
| +--------------+
| | +-- Ensidens_jaculus
| |
+--+ +-- Hyriopsis_desowitzi
| +-----------+
| | +-- Hyriopsis_delaportei
| |
+--+ +----------- Rectidens_sumatrensis
| |
| | +----- Hyriopsis_myersiana
+--+ +--+
| | | +-- Hyriopsis_bialata
| | +--+
+--+ +-- Hyriopsis_panhai
|
| +----- Hyriopsis_khoratensis
+--+
| +-- Hyriopsis_phuphaniensis
+--+
+-- Hyriopsis_sakhonensis
Pfeiffer et al., 2021 [Publ 4126], fig. 25
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 9 |
9 |
7 |
2 |
0 |
0 |
0 |
10.0 |
+-------------- Solenaia_emarginata **
|
| +-- Yaukthwa_elongatula **
--+ +--------+
| | +-- Yaukthwa_baniensis **
| |
+--+ +----- Yaukthwa_panhai **
| +--+
| | | +-- Yaukthwa_rectangularis **
| | +--+
+--+ +-- Yaukthwa_nesemanni **
|
| +----- Yaukthwa_avaensis **
+--+
| +-- Yaukthwa_inlenensis **
+--+
+-- Yaukthwa_paiensis **
Zieritz et al., 2021 [Publ 4218], fig. 5
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 7 |
7 |
5 |
4 |
25 |
21 |
7 |
6.0 |
+----- Ctenodesma_borneensis
+--------------+
| | +-- Khairuloconcha_lunbawangorum
| +--+
| +-- Khairuloconcha_sahanae
|
--+ +-------- Ensidens_sagittarius
| +--------+
| | | +----- Rectidens_sumatrensis
| | +--+
| | | +-- Hyriopsis_bialata
+--+ +--+
| +-- Hyriopsis_desowitzi
|
| +-------------- Physunio_superbus **
| |
+--+ +----------- Trapezoideus_foliaceus **
| |
+--+ +-- Lens_contradens **
| +-----+
| | +-- Lens_eximius **
+--+
| +----- Yaukthwa_elongatula **
+--+
| +-- Yaukthwa_inlenensis **
+--+
+-- Yaukthwa_paiensis **
Wu et al., 2024 [Publ 4350], fig. 5b
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 7 |
7 |
5 |
4 |
31 |
29 |
7 |
6.0 |
+-------------------- Ensidens_sagittarius
|
--+ +----------------- Hyriopsis_desowitzi
| |
+--+ +-------------- Yaukthwa_elongatula **
| |
+--+ +----------- Trapezoideus_foliaceus **
| |
+--+ +-- Physunio_superbus **
| +-----+
| | +-- Yaukthwa_inlenensis **
+--+
| +----- Yaukthwa_paiensis **
+--+
| +-- Lens_contradens **
+--+
+-- Lens_eximius **
Wu et al., 2024 [Publ 4350], fig. 5c
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 7 |
7 |
1 |
4 |
31 |
29 |
7 |
6.0 |
+-------- Ensidens_sagittarius
|
+-------- Hyriopsis_desowitzi
|
+-------- Rectidens_sumatrensis
|
+-------- Hyriopsis_bialata
|
+--------------+ +-- Khairuloconcha_lunbawangorum
| +-----+
| | +-- Khairuloconcha_sahanae
| |
| | +----- Physunio_superbus **
| | |
| | +----- Yaukthwa_paiensis **
| | |
| | +----- Yaukthwa_inlenensis **
| +--+
| +----- Trapezoideus_foliaceus **
--+ |
| +----- Yaukthwa_elongatula **
| |
| | +-- Lens_contradens **
| +--+
| +-- Lens_eximius **
|
| +-- Inversidens_rentianensis
| +-----------------+
| | +-- Inversidens_brandtii
| |
| | +-- Chamberlainia_somsakpanhai
+--+ +--+
| | +-- Potomida_littoralis
| +-----------+
| | | +-- Lamprotula_caveata
| | +--+
+--+ +-- Lamprotula_leaii
|
| +-------------- Sinosolenaia_carinata
| |
+--+ +----------- Gonidea_angulata
| |
+--+----------- Microcondylaea_bonellii
|
| +-------- Koreosolenaia_sitgyensis
+--+
| +----- Parvasolenaia_rivularis
+--+
| +-- Postolata_guangxiensis
+--+
+-- Ptychorhynchus_pfisteri
Wu et al., 2024 [Publ 4350], fig. 6-7
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 7 |
7 |
5 |
4 |
31 |
29 |
7 |
4.0 |
+-- Khairuloconcha_lunbawangorum
+--------------+
| +-- Khairuloconcha_sahanae
|
| +-------- Ensidens_sagittarius
--+ +-----+
| | | +----- Rectidens_sumatrensis
| | +--+
| | | +-- Hyriopsis_bialata
| | +--+
+--+ +-- Hyriopsis_desowitzi
|
| +-- Physunio_superbus **
| +--------+
| | +-- Trapezoideus_foliaceus **
| |
+--+ +-- Lens_contradens **
| +-----+
| | +-- Lens_eximius **
+--+
| +----- Yaukthwa_elongatula **
+--+
| +-- Yaukthwa_inlenensis **
+--+
+-- Yaukthwa_paiensis **
Pfeiffer et al., 2019 [Publ 3937], fig. 6
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 5 |
5 |
3 |
5 |
52 |
12 |
5 |
0.0 |
+-- Gonidea_angulata
+-----------+
| +-- Lamprotula_cornuumlunae
+--+
| | +----------- Indopseudodon_crebristriatus
| +--+
| | +-------- Pilsbryoconcha_exilis
| +--+
| | +----- Bineurus_anodontinum
| +--+
| | +-- Pseudodon_vondembuschianus
--+ +--+
| +-- Pseudodon_inoscularis
|
| +----- Ensidens_ingallsianus
| +-----+
| | | +-- Hyriopsis_bialata
| | +--+
| | +-- Rectidens_sumatrensis
+-----+
| +-- Trapezoideus_foliaceus **
| +-----+
| | +-- Physunio_superbus **
+--+
| +----- Yaukthwa_paiensis **
+--+
| +-- Lens_contradens **
+--+
+-- Solenaia_emarginata **
Bolotov et al., 2019 [Publ 3974], fig. 1
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 5 |
5 |
3 |
1 |
13 |
0 |
0 |
9.0 |
+----------- Yaukthwa_inlenensis **
--+
| +-------- Yaukthwa_elongatula **
+--+
| +----- Yaukthwa_panhai **
+--+
| +-- Yaukthwa_nesemanni **
+--+
+-- Yaukthwa_rectangularis **
Pfeiffer et al., 2018 [Publ 3827], fig. 1
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 4 |
4 |
1 |
3 |
4 |
4 |
1 |
5.0 |
+----- Physunio **
|
+--+----- Trapezoideus **
| |
| | +-- Lens_comptus **
| +--+
--+ +-- Lens_contradens **
|
| +-- Ensidens
| |
| +-- Hyriopsis[1]
+-----+
+-- Hyriopsis[2]
|
+-- Rectidens
Pfeiffer & Graf, 2015 [Publ 3620], fig. 2
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 3 |
3 |
1 |
2 |
37 |
11 |
4 |
5.5 |
+-------- Ensidens_telus
+--------+
| | +----- Lens_micropterus **
| +--+
| | +-- Physunio **
| +--+
| +-- Physunio_modelli **
--+
| +-------- Gonidea_angulata
| +-----+
| | | +----- Potomida_littoralis
| | +--+
| | | +-- Pronodularia_japanensis
+--+ +--+
| +-- Lamprotula_caveata
|
| +----------- Pilsbryoconcha_carinifera
| |
+--+ +-- Pseudodon_vondembuschianus
| +-----+
| | +-- Pseudodon_vagulus
+--+
| +----- Sundadontina_tumida
+--+
| +-- Nyeinchanconcha_nyeinchani
+--+
+-- Bineurus_exilis
Lopes-Lima et al., 2017 [Publ 3745], fig. 1-2
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 3 |
4 |
2 |
2 |
78 |
24 |
5 |
6.0 |
+-- Lens_contradens **
+--+
| +-- Lens_comptus **
+--+
| | +-- Physunio_modelli **
| +--+
| +-- unionoid_species
|
--+ +----- Hyriopsis_desowitzi
| |
| +----- Rectidens_sumatrensis
| |
| | +-- Hyriopsis_khoratensis
+--+--+
| +-- Hyriopsis_myersiana
|
| +-- Ensidens_ingallsianus
| |
+--+-- Ensidens_telus
|
+-- Ensidens_jaculus
Konopleva et al., 2017 [Publ 3807], fig. 6
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 3 |
3 |
1 |
2 |
28 |
7 |
2 |
5.5 |
+-------- Ensidens_sagittarius
+-----+
| | +----- Lens_micropterus **
| +--+
| | +-- Physunio **
--+ +--+
| +-- Physunio_modelli **
|
| +----------- Sundadontina_tumida
| |
+--+ +-- Nyeinchanconcha_nyeinchani
| +-----+
| | +-- Bineurus_exilis
+--+
| +----- Pilsbryoconcha_carinifera
+--+
| +-- Pseudodon_vagulus
+--+
+-- Pseudodon_vondembuschianus
Lopes-Lima et al., 2020 [Publ 4080], fig. 2-4
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 3 |
5 |
0 |
3 |
119 |
32 |
5 |
7.0 |
+-- Lens_contradens **
|
+-- Ensidens_sagittarius
|
--+-- Physunio_modelli **
|
+-- Rectidens_sumatrensis
|
+-- Yaukthwa_nesemanni **
Jeratthitikul et al., 2021 [Publ 4219], fig. 1
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 3 |
3 |
1 |
3 |
33 |
33 |
5 |
2.0 |
+----- Lens_contradens **
+--+
| | +-- Trapezoideus_foliaceus **
| +--+
+--------------------+ +-- Physunio_superbus **
| |
| | +----- Ensidens_ingallsianus
| +--+
| | +-- Hyriopsis_bialata
| +--+
| +-- Rectidens_sumatrensis
|
| +----- Sinosolenaia_carinata
| +-----+
| | | +-- Leguminaia_wheatleyi
--+ | +--+
| | +-- Gonidea_angulata
| +--------------+
| | | +-- Chamberlainia_somsakpanhai
| | | +-----+
| | | | +-- Sinohyriopsis_cumingii
| | +--+
| | | +----- Potomida_littoralis
| | +--+
| | | +-- Pronodularia_japanensis
| | +--+
+--+ +-- Lamprotula_leaii
|
| +-------- Indopseudodon_kayinensis
| +--------------+
| | | +----- Indopseudodon_indawgyiensis
| | +--+
| | | +-- Indopseudodon_salwenianus
| | +--+
| | +-- Indopseudodon_bogani
| |
+--+ +----- Pilsbryoconcha_exilis
| +--------------+
| | | +-- Pilsbryoconcha_linguaeformis
| | +--+
| | +-- Pilsbryoconcha
| |
| | +-------- Nyeinchanconcha_nyeinchani
| | |
+--+ +--------+ +----- Thaiconcha_callifera
| | | |
| | +--+ +-- Bineurus_panhai
| | | |
| | +--+-- Bineurus_exilis
| | |
| | +-- Bineurus_mouhotii
+--+
| +-- Sundadontina_brandti
| +--+
| | +-- Sundadontina_taskaevi
| +--------+
| | | +-- Sundadontina_tumida
| | +--+
| | +-- Sundadontina_cumingii
+--+
| +-- Namkongnaia_lemeslei
| +--------+
| | +-- Namkongnaia_inkhavilayi
+--+
| +-------- Pseudodon_lenyanensis
+--+
| +----- Pseudodon_vagulus
+--+
| +-- Pseudodon_vondembuschianus
+--+
+-- Pseudodon_mekongi
Jeratthitikul et al., 2021 [Publ 4219], fig. 2
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 3 |
3 |
1 |
3 |
33 |
33 |
5 |
2.0 |
+----- Lens_contradens **
+--+
| | +-- Trapezoideus_foliaceus **
| +--+
+--------------------+ +-- Physunio_superbus **
| |
| | +----- Ensidens_ingallsianus
| +--+
| | +-- Hyriopsis_bialata
| +--+
| +-- Rectidens_sumatrensis
|
| +----- Sinosolenaia_carinata
| +-----+
| | | +-- Gonidea_angulata
--+ | +--+
| | +-- Leguminaia_wheatleyi
| +--------------+
| | | +-- Chamberlainia_somsakpanhai
| | | +-----+
| | | | +-- Sinohyriopsis_cumingii
| | +--+
| | | +----- Potomida_littoralis
| | +--+
| | | +-- Lamprotula_leaii
| | +--+
+--+ +-- Pronodularia_japanensis
|
| +-------- Indopseudodon_kayinensis
| +--------------+
| | | +----- Indopseudodon_indawgyiensis
| | +--+
| | | +-- Indopseudodon_salwenianus
| | +--+
| | +-- Indopseudodon_bogani
| |
+--+ +----- Pilsbryoconcha_exilis
| +--------------+
| | | +-- Pilsbryoconcha_linguaeformis
| | +--+
| | +-- Pilsbryoconcha
| |
| | +-- Nyeinchanconcha_nyeinchani
| | +--------+
+--+ | +-- Sundadontina_cumingii
| +-----+
| | | +-------- Thaiconcha_callifera
| | +--+
| | | +----- Bineurus_mouhotii
| | +--+
| | | +-- Bineurus_panhai
| | +--+
+--+ +-- Bineurus_exilis
|
| +----- Sundadontina_tumida
| +--------+
| | | +-- Sundadontina_brandti
| | +--+
| | +-- Sundadontina_taskaevi
+--+
| +-- Namkongnaia_lemeslei
| +--------+
| | +-- Namkongnaia_inkhavilayi
+--+
| +-------- Pseudodon_lenyanensis
+--+
| +----- Pseudodon_vagulus
+--+
| +-- Pseudodon_vondembuschianus
+--+
+-- Pseudodon_mekongi
Jeratthitikul et al., 2022 [Publ 4260], fig. S1, 1
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 3 |
3 |
1 |
3 |
73 |
28 |
5 |
2.0 |
+----- Lens_contradens **
+--+
| | +-- Trapezoideus_foliaceus **
| +--+
--+ +-- Physunio_superbus **
|
| +-- Ensidens_ingallsianus
| |
+-----+-- Hyriopsis_bialata
|
+-- Rectidens_sumatrensis
Jeratthitikul et al., 2022 [Publ 4260], fig. S2, 2
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 3 |
3 |
1 |
3 |
73 |
28 |
5 |
2.0 |
+----- Ensidens_ingallsianus
+--+
| | +-- Rectidens_sumatrensis
| +--+
--+ +-- Hyriopsis_bialata
|
| +----- Lens_contradens **
+--+
| +-- Trapezoideus_foliaceus **
+--+
+-- Physunio_superbus **
Dai et al., 2023 [Publ 4285], fig. 3-BI
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 3 |
3 |
1 |
2 |
42 |
40 |
5 |
4.0 |
+-- Hyriopsis_bialata
+-----+
| +-- Ensidens_ingallsianus
+-----------+
| | +----- Physunio_superbus **
| +--+
| | +-- Lens_contradens **
| +--+
| +-- Lens_comptus **
|
| +-- Inversidens_brandtii
| +--------------+
| | +-- Inversidens_rentianensis
--+ |
| | +----- Chamberlainia_somsakpanhai
| | +-----+
| | | | +-- Sinohyriopsis_cumingii
| | | +--+
| +-----+ +-- Sinohyriopsis_schlegelii
| | |
| | | +-------- Potomida_littoralis
| | +--+
| | | +----- Pronodularia_japanensis
+--+ +--+
| | +-- Lamprotula_leaii
| +--+
| +-- Lamprotula_caveata
|
| +-------- Sinosolenaia_carinata
| +-----+
| | | +----- Sinosolenaia_recognita
| | +--+
| | | +-- Sinosolenaia_iridinea
| | +--+
| | +-- Sinosolenaia_oleivora
+--+
| +----- Gonidea_angulata
| +-----+
| | | +-- Leguminaia_wheatleyi
| | +--+
| | +-- Microcondylaea_bonellii
+--+
| +-- Koreosolenaia_sitgyensis
| +-----+
| | +-- Parvasolenaia_rivularis
+--+
| +----- Postolata_guangxiensis
+--+
| +-- Ptychorhynchus_pfisteri
+--+
+-- Obovalis_omiensis
Dai et al., 2023 [Publ 4285], fig. 3-ML
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 3 |
3 |
1 |
2 |
42 |
40 |
5 |
4.0 |
+-- Hyriopsis_bialata
+-----+
| +-- Ensidens_ingallsianus
+-----------+
| | +----- Physunio_superbus **
| +--+
| | +-- Lens_contradens **
| +--+
| +-- Lens_comptus **
|
| +-- Inversidens_brandtii
| +-----------+
| | +-- Inversidens_rentianensis
--+ |
| +--+ +----- Chamberlainia_somsakpanhai
| | | +-----+
| | | | | +-- Sinohyriopsis_cumingii
| | | | +--+
| | +--+ +-- Sinohyriopsis_schlegelii
| | |
| | | +-------- Potomida_littoralis
| | +--+
| | | +----- Pronodularia_japanensis
| | +--+
+--+ | +-- Lamprotula_leaii
| +--+
| +-- Lamprotula_caveata
|
| +-------- Sinosolenaia_carinata
| +-----+
| | | +----- Sinosolenaia_recognita
| | +--+
| | | +-- Sinosolenaia_iridinea
| | +--+
| | +-- Sinosolenaia_oleivora
+--+
| +----- Gonidea_angulata
| +-----+
| | | +-- Leguminaia_wheatleyi
| | +--+
| | +-- Microcondylaea_bonellii
+--+
| +-- Koreosolenaia_sitgyensis
| +-----+
| | +-- Parvasolenaia_rivularis
+--+
| +----- Postolata_guangxiensis
+--+
| +-- Ptychorhynchus_pfisteri
+--+
+-- Obovalis_omiensis
Zieritz et al., 2021 [Publ 4231], fig. 3-F, 5
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 2 |
2 |
0 |
2 |
47 |
15 |
5 |
-- |
+-- Lens_contradens **
+--+
| +-- Physunio_superbus **
+-----------+
| | +-- Hyriopsis_bialata
| +--+
| +-- Rectidens_sumatrensis
|
--+ +-------- Potomida_littoralis
| +-----+
| | | +----- Pronodularia_japanensis
| | +--+
| | | +-- Lamprotula_caveata
| | +--+
+--+ +-- Lamprotula_leaii
|
| +-- Pilsbryoconcha_exilis
| +--------+
| | +-- Pseudodon_vondembuschianus
+--+
| +-------- Ptychorhynchus_pfisteri
+--+
| +----- Microcondylaea_bonellii
+--+
| +-- Sinosolenaia_recognita
+--+
+-- Sinosolenaia_carinata
Zieritz et al., 2021 [Publ 4231], fig. 3-M, 4
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 2 |
2 |
0 |
2 |
21 |
10 |
5 |
-- |
+-- Lens_contradens **
+--+
| +-- Physunio_superbus **
+-----+
| | +-- Hyriopsis_bialata
| +--+
| +-- Rectidens_sumatrensis
--+
| +-- Pilsbryoconcha_exilis
| +-----+
| | +-- Pseudodon_vondembuschianus
| |
+--+ +-- Pronodularia_japanensis
| +--+
| | +-- Potomida_littoralis
+--+
| +-- Microcondylaea_bonellii
+--+
+-- Sinosolenaia_carinata
Dai et al., 2023 [Publ 4285], fig. 5
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 2 |
2 |
0 |
2 |
51 |
17 |
5 |
-- |
+-- Hyriopsis_bialata
+--+
| +-- Rectidens_sumatrensis
+-----------+
| | +-- Lens_contradens **
| +--+
| +-- Physunio_superbus **
|
--+ +-------- Potomida_littoralis
| +-----+
| | | +----- Pronodularia_japanensis
| | +--+
| | | +-- Lamprotula_caveata
| | +--+
+--+ +-- Lamprotula_leaii
|
| +-- Pseudodon_vondembuschianus
| +--------+
| | +-- Pilsbryoconcha_exilis
| |
+--+ +----- Parvasolenaia_rivularis
| +--+
| | | +-- Postolata_guangxiensis
| | +--+
+--+ +-- Ptychorhynchus_pfisteri
|
| +----- Microcondylaea_bonellii
+--+
| +-- Sinosolenaia_oleivora
+--+
+-- Sinosolenaia_carinata
Wu et al., 2023 [Publ 4339], fig. 4, S5-aa
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 2 |
2 |
0 |
2 |
94 |
18 |
5 |
-- |
+-- Hyriopsis_bialata
+--+
| +-- Rectidens_sumatrensis
+--------------+
| | +-- Lens_contradens **
| +--+
| +-- Physunio_superbus **
|
| +----------- Potomida_littoralis
--+ +-----+
| | | +-------- Pronodularia_japanensis
| | +--+
| | | +----- Lamprotula_leaii
| | +--+
| | | +-- Lamprotula_caveata
+--+ +--+
| +-- Lamprotula_cornuumlunae
|
| +-- Pseudodon_vondembuschianus
| +-----------+
| | +-- Pilsbryoconcha_exilis
+--+
| +----------- Microcondylaea_bonellii
| |
+--+ +-- Parvasolenaia_rivularis
| +-----+
| | +-- Ptychorhynchus_pfisteri
+--+
| +----- Sinosolenaia_carinata
+--+
| +-- Sinosolenaia_iridinea
+--+
+-- Sinosolenaia_oleivora
Wu et al., 2023 [Publ 4339], fig. 4, S5-nt
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 2 |
2 |
0 |
2 |
95 |
18 |
5 |
-- |
+-- Hyriopsis_bialata
+--+
| +-- Rectidens_sumatrensis
+--------------+
| | +-- Lens_contradens **
| +--+
| +-- Physunio_superbus **
|
| +----------- Potomida_littoralis
--+ +-----+
| | | +-------- Pronodularia_japanensis
| | +--+
| | | +----- Lamprotula_leaii
| | +--+
| | | +-- Lamprotula_caveata
+--+ +--+
| +-- Lamprotula_cornuumlunae
|
| +-- Pseudodon_vondembuschianus
| +-----------+
| | +-- Pilsbryoconcha_exilis
| |
+--+ +-- Ptychorhynchus_pfisteri
| +--------+
| | +-- Microcondylaea_bonellii
+--+
| +-------- Parvasolenaia_rivularis
+--+
| +----- Sinosolenaia_carinata
+--+
| +-- Sinosolenaia_iridinea
+--+
+-- Sinosolenaia_oleivora
Wu et al., 2024 [Publ 4350], fig. 8a, S1
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 2 |
2 |
0 |
2 |
90 |
19 |
5 |
-- |
+-- Hyriopsis_bialata
+--+
| +-- Rectidens_sumatrensis
+-----------------+
| | +-- Lens_contradens **
| +--+
| +-- Physunio_superbus **
|
| +----------- Potomida_littoralis
--+ +--------+
| | | +-------- Pronodularia_japanensis
| | +--+
| | | +----- Lamprotula_leaii
| | +--+
| | | +-- Lamprotula_caveata
+--+ +--+
| +-- Lamprotula_cornuumlunae
|
| +-- Pseudodon_vondembuschianus
| +--------------+
| | +-- Pilsbryoconcha_exilis
+--+
| +-------------- Inversidens_rentianensis
| |
+--+ +-- Parvasolenaia_rivularis
| +--------+
| | +-- Ptychorhynchus_pfisteri
+--+
| +-------- Microcondylaea_bonellii
+--+
| +----- Sinosolenaia_carinata
+--+
| +-- Sinosolenaia_iridinea
+--+
+-- Sinosolenaia_oleivora
Wu et al., 2024 [Publ 4350], fig. 8b, S2
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 2 |
2 |
0 |
2 |
90 |
19 |
5 |
-- |
+-- Hyriopsis_bialata
+--+
| +-- Rectidens_sumatrensis
+-----------------+
| | +-- Lens_contradens **
| +--+
| +-- Physunio_superbus **
|
| +----------- Potomida_littoralis
--+ +--------+
| | | +-------- Pronodularia_japanensis
| | +--+
| | | +----- Lamprotula_leaii
| | +--+
| | | +-- Lamprotula_caveata
+--+ +--+
| +-- Lamprotula_cornuumlunae
|
| +-- Pseudodon_vondembuschianus
| +--------------+
| | +-- Pilsbryoconcha_exilis
+--+
| +-------------- Inversidens_rentianensis
| |
+--+ +-- Parvasolenaia_rivularis
| +--------+
| | +-- Ptychorhynchus_pfisteri
+--+
| +-------- Microcondylaea_bonellii
+--+
| +----- Sinosolenaia_carinata
+--+
| +-- Sinosolenaia_iridinea
+--+
+-- Sinosolenaia_oleivora
Graf & Cummings, 2006 [Publ 1968], fig. 2
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 1 |
1 |
0 |
1 |
46 |
3 |
2 |
-- |
-- Lens_contradens
Graf, 2002 [Publ 2651], fig. 2
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 1 |
1 |
0 |
1 |
17 |
3 |
2 |
-- |
-- Lens_contradens
Hoeh et al., 2009 [Publ 2716], fig. 3
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 1 |
1 |
0 |
1 |
46 |
3 |
2 |
-- |
-- Lens_contradens
Hoeh et al., 2009 [Publ 2716], fig. 4
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 1 |
1 |
0 |
1 |
45 |
3 |
2 |
-- |
-- Lens_contradens
Hoeh et al., 2009 [Publ 2716], fig. 5
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 1 |
1 |
0 |
1 |
45 |
3 |
2 |
-- |
-- Lens_contradens
Whelan et al., 2011 [Publ 3380], fig. 1 ML
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 1 |
1 |
0 |
1 |
36 |
5 |
3 |
-- |
+-- Pseudodon_vondembuschianus
--+
+-- Lens_contradens **
Whelan et al., 2011 [Publ 3380], fig. 1 MP
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 1 |
1 |
0 |
1 |
36 |
5 |
3 |
-- |
-- Lens_contradens
Pfeiffer & Graf, 2013 [Publ 3531], fig. 2
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 1 |
1 |
0 |
1 |
34 |
11 |
4 |
-- |
-- Physunio
Smith, 2021 [Publ 4331], fig. 1
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 1 |
1 |
0 |
1 |
19 |
2 |
2 |
-- |
+----- Sinohyriopsis_cumingii
--+
| +-- Lens_contradens **
+--+
+-- Lamprotula_leaii
| 
