|
Bivalvia | Unionoida superfamily UNIONOIDEA Rafinesque, 1820: cladomics
2 families
Cladomics Summary
number of species = 1078
number of subtaxa (rank: family) = 2
analyzed in 495 cladograms in 277 publications
maximum number of terminals in an analysis = 205, mean = 22.45
maximum number of subtaxa (family) in an analysis = 2, mean = 1.46
Basic Monophyly
number of cladograms that test basic monophyly (i.e., more than 1 included species) = 493
maximum number of included terminals = 205, mean = 22.54
maximum number of subtaxa (family) = 2, mean = 1.46
proportion of cladograms that support basic monophyly = 0.94
Rigorous Monophyly
number of cladograms that test rigorous monophyly = 83
(Rigorous monophyly refers to cladograms with terminal species representing more than one subtaxon (family) taxon and at least one rigorous outgroup (superfamily).
maximum number of included terminals = 178, mean = 34.45
maximum number of subtaxa (family) = 2, mean = 2.0
proportion of cladograms that support rigorous monophyly = 0.67
Pairwise Cladogram Clustering
The graphs below depict in two dimensions all pair-wise ingroup distances within the cladome. See the Cladmoics FARQ for more information.
Lopes-Lima et al., 2017 [Publ 3745], fig. 1-2
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 71 |
78 |
71 |
2 |
10 |
9 |
3 |
0.0 |
*** Top Monophyly Cladogram *** *** Top Sister-Group Cladogram ***
+-- Hyridella_depressa
+-----------------------------------+
| +-- Velesunio_ambiguus
|
| +-- Cumberlandia_monodonta **
| +--------------------------------+
| | +-- Margaritifera_margaritifera **
| |
| | +----------- Mutela_rostrata
--+ | |
| | +--------------------+ +-- Aspatharia_pfeifferiana
| | | | +-----+
| | | | | +-- Chambardia_wahlbergi
| | | +--+
| | | | +----- Etheria_elliptica
| | | +--+
| | | | +-- Tamsiella_schroeteriana
+--+ | +--+
| | +-- Mycetopoda_siliquosa
| |
| | +-------- Trapezidens_dolichorhynchus **
| | +--------------------+
| | | | +----- Leoparreysia_tavoyensis **
| | | +--+
| | | | +-- Coelatura_aegyptiaca **
| | | +--+
| | | +-- Indochinella_pugio **
| | |
| | | +----- Unio_gibbus **
+--+ | |
| | +-----------+----- Unio_tumidus **
| | | |
| | | | +-- Unio_crassus **
| | | +--+
| | +--+ +-- Unio_pictorum **
| | | |
| | | | +-------------- Aculamprotula_tortuosa **
| | | | |
| | | +--+ +-- Nodularia_nuxpersicae **
| | | | +--------+
| | | | | +-- Nodularia_douglasiae **
| | | +--+
| | | | +-------- Schistodesmus_lampreyanus **
| | | +--+
| | | | +----- Tchangsinaia_piscicula **
| | | +--+
| | +-----+ | +-- Cuneopsis_heudei **
| | | | +--+
| | | | +-- Cuneopsis_rufescens **
+--+ | |
| | | +-------- Lanceolaria_gladiola **
| | | +--------+
| | | | | +----- Lanceolaria_grayii[2] **
| | | | +--+
| | | | | +-- Lanceolaria_lanceolata **
| | | | +--+
| | | | +-- Lanceolaria_grayii[1] **
| | | |
| | +--+ +----------- Cristaria_plicata **
| | | +--+
| | | | | +-------- Anemina_arcaeformis **
| | | | +--+
| | | | | +----- Pletholophus_tenuis **
| | | | +--+
| | | | | +-- Sinanodonta_jourdyi **
| | | | +--+
| | +--+ +-- Sinanodonta_lucida **
| | |
| | | +----- Pseudanodonta_complanata **
| | | |
| | | +-----+ +-- Anodonta_anatina **
| | | | | |
| | | | +--+-- Anodonta_cygnea **
| | | | |
| | +--+ +-- Anodonta_nuttalliana **
| | |
| | | +-- Pyganodon_grandis **
+--+ | +-----+
| | | +-- Simpsonaias_ambigua **
| +--+
| | +----- Strophitus_undulatus **
| +--+
| | +-- Alasmidonta_marginata **
| +--+
| +-- Lasmigona_costata **
|
| +-- Lens_contradens **
| +--+
| | +-- Lens_comptus **
| +--+
| | | +-- Physunio_modelli **
| | +--+
| | +-- unionoid_species
| |
| +--------------+ +----- Hyriopsis_desowitzi **
| | | |
| | | +----- Rectidens_sumatrensis **
| | | |
| | | | +-- Hyriopsis_khoratensis **
| | +--+--+
| | | +-- Hyriopsis_myersiana **
| | |
| | | +-- Ensidens_ingallsianus **
| | | |
| | +--+-- Ensidens_telus **
| | |
| | +-- Ensidens_jaculus **
| |
| | +-------- Amblema_plicata **
+--+ +--+
| | | +----- Cambarunio_iris **
| | +--+
| | | +-- Lampsilis_cardium **
| | +--+
| +--------+ +-- Ortmanniana_ligamentina **
| | |
| | | +-- Quadrula_quadrula **
| | | +-----+
| | | | +-- Quadrula_verrucosa **
| | +--+
| | | +----- Elliptio_complanata **
| | +--+
| | | +-- Pleurobema_sintoxia **
| | +--+
| | +-- Eurynia_dilatata **
+--+
| +-- Chamberlainia_somsakpanhai **
| +-----+
| | +-- Sinohyriopsis_cumingii **
| |
| +--------+ +-- Lamprotula_caveata **
| | | +--+
| | | | +-- Lamprotula_leaii **
| | +--+
| | | +-- Potomida_littoralis **
| | +--+
| | +-- Pronodularia_japanensis **
+--+
| +-- Sinosolenaia_carinata **
| +-----+
| | +-- Sinosolenaia_oleivora **
| +-----+
| | | +----- Gonidea_angulata **
| | +--+
| | | +-- Microcondylaea_bonellii **
+--+ +--+
| +-- Leguminaia_wheatleyi **
|
| +----------- Parvasolenaia_rivularis **
| |
+--+ +-- Pilsbryoconcha_exilis **
| +-----+
| | +-- Pilsbryoconcha_acuta **
+--+
| +----- Pseudodon_vagulus **
+--+
| +-- Bineurus_exilis **
+--+
+-- Nyeinchanconcha_nyeinchani **
Inoue et al., 2018 [Publ 3858], fig. 1, 3a-c
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 205 |
205 |
66 |
2 |
0 |
0 |
0 |
224.0 |
+-------------------------------------------------------- Cumberlandia_monodonta **
|
| +-- Alasmidonta_marginata **
| |
| +-- Prolasmidonta_heterodon **
| |
| +-- Alasmidonta_undulata **
| |
| +-- Beringiana_beringiana **
| |
| +-- Anodonta_kennerlyi **
| |
| +-- Anodonta_nuttalliana **
| |
| +-- Anodonta_oregonensis **
| |
| +-- Arcidens_confragosus **
| |
| +-- Arcidens_wheeleri **
| |
--+ +-- Lasmigona_complanata **
| |
| +--------------------------------------------------+-- Platynaias_compressa **
| | |
| | +-- Lasmigona_costata **
| | |
| | +-- Platynaias_decorata **
| | |
| | +-- Platynaias_subviridis **
| | |
| | +-- Pyganodon_grandis **
| | |
| | +-- Pyganodon_lacustris **
| | |
| | +-- Pseudodontoideus_subvexus **
| | |
| | +-- Strophitus_undulatus **
| | |
| | +-- Utterbackia_imbecillis **
| | |
+--+ +-- Utterbackia_peninsularis[1] **
| |
| +-- Utterbackia_peninsularis[2] **
|
| +-------------------------------------------------- Gonidea_angulata **
| |
| | +-- Reginaia_ebenus **
| | +--------------------------------------------+
| | | +-- Reginaia_rotulata **
| | |
| | | +-- Amblema_plicata **
| | +--------------------------------------------+
| | | +-- Amblema_elliottii **
| | |
| | | +----- Popenaias_popeii **
| | | |
| | | | +-- Dromus_dromas **
| | | | |
+--+ | | +-- Epioblasma_rangiana **
| | | |
| +-----------------------------------------+ +-- Glebula_rotundata **
| | | |
| | | +-- Lampsilis_ovata **
| | | |
| | | +-- Lampsilis_virescens **
| | | |
| | | +-- Lemiox_rimosus **
| | +--+
| | +-- Obliquaria_reflexa **
| | |
| | +-- Obovaria_olivaria **
| | |
+--+ +-- Obovaria_unicolor **
| |
| +-- Plectomerus_dombeyanus **
| |
| +-- Potamilus_alatus **
| |
| +-- Potamilus_capax **
|
| +-- Uniomerus_declivis **
| +--+
| | +-- Uniomerus_tetralasmus **
| |
| | +-- Cyclonaias_tuberculata **
| +--------------------------------------+ |
| | | +-- Pustulosa_pustulosa[1] **
| | | |
| | | +-- Theliderma_metanevra **
| | +--+
| | +-- Pustulosa_pustulosa[2] **
| | |
| | +-- Quadrula_quadrula **
| | |
| | +-- Quadrula_verrucosa **
| |
| | +----------------------------------------- Elliptoideus_sloatianus **
+--+ |
| | +-- Elliptio_lanceolata **
| | +-----------------------------------+
| | | +-- Parvaspina_collina **
| | |
| | | +----------------------------------- Plethobasus_cyphyus **
| | | |
| | | +----------------------------------- Eurynia_dilatata **
| | | |
| | | +----------------------------------- Hemistena_lata **
| | | |
| | | | +-- Elliptio_complanata **
+--+ | | |
| | | +-- Elliptio_hopetonensis **
| | | |
| | | +-- Elliptio_producta[1] **
| | | |
| | | +-- Elliptio_shepardiana **
| | | |
| | | +-- Elliptio_angustata **
| | | |
| | | +-- Elliptio_arca **
| | | |
| | | +-- Elliptio_arctata **
| | | |
+--+ | +-- Elliptio_jayensis **
| | |
| | +-- Elliptio_crassidens **
| | |
| +--------------------------------+-- Elliptio_dariensis **
| | |
| | +-- Elliptio_fisheriana **
| | |
| | +-- Elliptio_folliculata **
| | |
| | +-- Elliptio_fumata **
| | |
| | +-- Elliptio_icterina **
| | |
| | +-- Elliptio_mcmichaeli **
| | |
| | +-- Elliptio_producta[2] **
| | |
| | +-- Elliptio_pullata **
| | |
| | +-- Elliptio_purpurella **
| | |
| | +-- Elliptio_congaraea **
| |
+--+ +-- Pleuronaia_barnesiana **
| |
| +-----------------------------+-- Pleuronaia_dolabelloides **
| | |
| | +-- Pleuronaia_gibber **
| |
| | +-------------------------- Pleurobema_plenum **
| | |
| | | +----------------------- Pleurobema_stabile **
| | | |
| | | | +----- Pleurobema_pyriforme **
| | +--+ | +--+
| | | | | | | +-- Pleurobema_georgianum[2] **
| | | | | | +--+
| | | | | +-----------+ +-- Pleurobema_athearni **
| | | | | | |
| | | +--+ | | +----- Pleurobema_decisum **
| | | | | | |
| | | | | +--+ +-- Pleurobema_georgianum[1] **
| | | | | | |
| | | | | +--+-- Pleurobema_hanleyianum[1] **
| | | | | |
| | | | | +-- Pleurobema_hanleyianum[2] **
| | | | |
| | | +--+ +-------------- Pleurobema_beadleianum[2] **
| | | | |
| | | | +--+ +----------- Pleurobema_strodeanum[2] **
| | | | | | |
| | | | | +--+----------- Pleurobema_rubellum **
| | | | | |
| | | | | | +-------- Pleurobema_beadleianum[1] **
| | | | | +--+
| | | | | | +----- Pleurobema_taitianum **
| | | | | +--+
| | | | | | +-- Pleurobema_perovatum **
| | | +--+ +--+
| | | | +-- Pleurobema_strodeanum[1] **
| | | |
| | | | +----- Pleurobema_oviforme[6] **
| | | | |
| | | | +--+----- Pleurobema_clava[3] **
| | | | | |
| | | | | | +-- Pleurobema_oviforme[5] **
| | | | | +--+
| | | +--------+ +-- Pleurobema_clava[2] **
+--+ | |
| | | +----- Pleurobema_oviforme[2] **
| | | |
| | | +----- Pleurobema_oviforme[3] **
| | +--+
| | +----- Pleurobema_oviforme[4] **
| | |
| | | +-- Pleurobema_clava[1] **
| | +--+
| | +-- Pleurobema_oviforme[1] **
| |
| | +-------------------------- Pleurobema_cordatum **
| | |
| | | +----- Fusconaia_subrotunda **
| | | +-----------------+
| +--+ | | +-- Fusconaia_cor **
| | | | +--+
| | | | +-- Fusconaia_cuneolus **
| | | |
| | | | +-------------------- Fusconaia_ozarkensis **
| | +--+ |
| | | | +-- Fusconaia_escambia **
| | | | +--------------+
| | | | | +-- Fusconaia_burkei **
| | | | |
| | | | | +----- Fusconaia_flava[27] **
| | | | | |
| | +--+ | | +-- Fusconaia_iheringi **
| | | | +--+
| | | | | +-- Fusconaia_mitchelli **
| | | +-----------+
| | | | | +-- Fusconaia_askewi[1] **
| | | | | |
| | | | | +-- Fusconaia_askewi[2] **
| | | | | |
| | +--+ | +-- Fusconaia_askewi[3] **
| | | +--+
| | | +-- Fusconaia_askewi[4] **
| | | |
| | | +-- Fusconaia_askewi[5] **
| | | |
| | | +-- Fusconaia_chunii **
| | |
+--+ | +-------------- Fusconaia_masoni **
| | |
| | | +----------- Fusconaia_flava[26] **
| | | |
| | | | +-- Fusconaia_cerina[8] **
| +--+ | +-----+
| | | | +-- Fusconaia_flava[25] **
| | | |
| | | | +----- Fusconaia_flava[5] **
| | | | |
| | | | +----- Fusconaia_flava[6] **
| +--+ | |
| | | +----- Fusconaia_flava[7] **
| | | |
| | | +----- Fusconaia_flava[8] **
| | | |
| | | +----- Fusconaia_cerina[4] **
| | | |
| | | +----- Fusconaia_flava[9] **
| | | |
| +--+ +----- Fusconaia_flava[10] **
| | |
| | +----- Fusconaia_flava[11] **
| | |
| | +----- Fusconaia_flava[12] **
| | |
| | +----- Fusconaia_flava[13] **
| | |
| | +----- Fusconaia_flava[14] **
| | |
| | +----- Fusconaia_flava[15] **
| | |
| | +----- Fusconaia_flava[16] **
| | |
| | +----- Fusconaia_flava[17] **
| +--+
| +----- Fusconaia_flava[18] **
| |
| +----- Fusconaia_flava[19] **
| |
| +----- Fusconaia_flava[20] **
| |
| +----- Fusconaia_flava[21] **
| |
| +----- Fusconaia_flava[22] **
| |
| +----- Fusconaia_flava[23] **
| |
| +----- Fusconaia_flava[24] **
| |
| +----- Fusconaia_cerina[5] **
| |
| +----- Fusconaia_cerina[6] **
| |
| +----- Fusconaia_cerina[7] **
| |
| | +-- Fusconaia_flava[1] **
| | |
| | +-- Fusconaia_flava[2] **
| | |
| | +-- Fusconaia_flava[3] **
| | |
| +--+-- Fusconaia_flava[4] **
| |
| +-- Fusconaia_cerina[1] **
| |
| +-- Fusconaia_cerina[2] **
| |
| +-- Fusconaia_cerina[3] **
|
| +----------------- Pleurobema_riddellii **
| |
| | +-------------- Pleurobema_sintoxia[21] **
| | |
| | | +----------- Pleurobema_sintoxia[19] **
| | | |
| | | +----------- Pleurobema_sintoxia[20] **
| | | |
+-----------+ | | +----- Pleurobema_sintoxia[18] **
| | | |
| | | +----- Pleurobema_rubrum[19] **
| | | |
| | | +----- Pleurobema_rubrum[20] **
| | +-----+
| | | +----- Pleurobema_rubrum[21] **
| | | |
| | | | +-- Pleurobema_rubrum[17] **
+--+ | | |
| | | +-- Pleurobema_sintoxia[16] **
| | +--+
| | +-- Pleurobema_sintoxia[17] **
| | |
| | +-- Pleurobema_rubrum[18] **
| |
| | +-------- Pleurobema[4] **
| | |
| | +-------- Pleurobema[5] **
| | |
| | +-------- Pleurobema_sintoxia[1] **
| | |
| | +-------- Pleurobema[8] **
| | |
| | +-------- Pleurobema[9] **
+--+ |
| +-------- Pleurobema[10] **
| |
| +-------- Pleurobema_sintoxia[3] **
| |
| +-------- Pleurobema_sintoxia[4] **
| |
| +-------- Pleurobema_sintoxia[5] **
| |
| +-------- Pleurobema_sintoxia[6] **
| |
| +-------- Pleurobema_sintoxia[7] **
| |
| +-------- Pleurobema_sintoxia[8] **
| |
| +-------- Pleurobema_sintoxia[9] **
| |
| +-------- Pleurobema_sintoxia[10] **
| |
| +-------- Pleurobema_sintoxia[11] **
| |
| +-------- Pleurobema_sintoxia[12] **
| |
| +-------- Pleurobema_sintoxia[13] **
| |
| +-------- Pleurobema_sintoxia[14] **
| |
| +-------- Pleurobema_sintoxia[15] **
| |
+--+-------- Pleurobema_rubrum[4] **
|
+-------- Pleurobema_rubrum[5] **
|
+-------- Pleurobema_rubrum[6] **
|
+-------- Pleurobema_rubrum[7] **
|
+-------- Pleurobema_rubrum[8] **
|
+-------- Pleurobema_rubrum[9] **
|
+-------- Pleurobema_rubrum[10] **
|
+-------- Pleurobema_rubrum[11] **
|
+-------- Pleurobema_rubrum[12] **
|
+-------- Pleurobema_rubrum[13] **
|
+-------- Pleurobema_rubrum[14] **
|
+-------- Pleurobema_rubrum[15] **
|
+-------- Pleurobema_rubrum[16] **
|
| +-- Pleurobema[6] **
+-----+
| +-- Pleurobema_rubrum[1] **
|
| +-- Pleurobema_sintoxia[2] **
+-----+
| +-- Pleurobema_rubrum[2] **
|
| +-- Pleurobema[7] **
+-----+
| +-- Pleurobema_rubrum[3] **
|
| +----- Pleurobema[3] **
+--+
| +-- Pleurobema[1] **
+--+
+-- Pleurobema[2] **
Huang et al., 2019 [Publ 3843], fig. 2
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 178 |
178 |
153 |
2 |
26 |
24 |
2 |
142.0 |
+----- Alathyria_jacksoni
+--+
| | +-- Alathyria_profuga
| +--+
+--------+ +-- Velesunio_ambiguus
| |
| | +----- Alathyria_pertexta
| +--+
| | +-- Lortiella_froggatti
+--------------------------------+ +--+
| | +-- Microdontia_anodontaeformis
| |
| | +-------------- Westralunio_carteri
| | |
| +--+ +----------- Echyridella_menziesii
| | |
| | | +----- Diplodon_suavidicus
| +--+ +--+
| | | | +-- Castalia_ambigua
| | | +--+
| +--+ +-- Triplodon_corrugatus
| |
| | +----- Hyridella_australis
| +--+
| | +-- Hyridella_depressa
| +--+
| +-- Hyridella_drapeta
|
| +-------- Anodontites_trapesialis
| +--+
| | | +----- Mutela_rostrata
| | +--+
| | | +-- Mutela_dubia
| | +--+
| +-----------------------------------+ +-- Mutela_hargeri
| | |
| | | +----- Mycetopoda_siliquosa
| | | +--+
--+ | | | | +-- Tamsiella_schroeteriana
| | | | +--+
| | +--+ +-- Lamproscapha_ensiformis
| | |
| | | +----- Etheria_elliptica
| | +--+
| | | +-- Aspatharia_pfeifferiana
| | +--+
| | +-- Chambardia_wahlbergi
| |
| | +----------------- Gibbosula_rochechouartii **
| | |
| | | +-------------- Gibbosula_laosensis **
| | +--------------------------+ |
| | | | | +----------- Cumberlandia_monodonta **
| | | | | |
| | | +--+ | +----- Pseudunio_marocanus **
| | | | +-----+
| | | | | | +-- Pseudunio_auricularius **
| | | | | +--+
| | | +--+ +-- Pseudunio_homsensis **
| | | |
| | | | +-- Margaritifera_dahurica **
| | | | +-----+
| | | | | +-- Margaritifera_margaritifera **
| | | | |
| | | +--+ +----- Margaritifera_laevis **
| | | | |
| | | | | +-- Margaritifera_falcata **
| | | +--+--+
+--+ | | +-- Margaritifera_middendorffi **
| | |
| | | +-- Margaritifera_hembeli **
| | +--+
| | +-- Margaritifera_marrianae **
| |
| | +----- Trapezidens_exolescens **
| | +-----------------+
| | | | +-- Trapezidens_dolichorhynchus **
| | | +--+
| | +--+ +-- Trapezidens_angustior **
| | | |
| | | | +-------------------- Lamellidens_corrianus **
| | | | |
| | | +--+ +----------------- Lamellidens_savadiensis **
| | | | |
| | | +--+ +-- Lamellidens_lamellatus **
| | | | +-----------+
| | | | | +-- Lamellidens_jenkinsianus **
| | | +--+
| | | | +----------- Lamellidens_generosus **
| | | +--+
| | | | +-------- Lamellidens_mainwaringi[2] **
| | | +--+
| | +--------------+ | +----- Lamellidens_mainwaringi[1] **
| | | | +--+
| | | | | +-- Lamellidens_ferrugineus **
| | | | +--+
| | | | +-- Lamellidens_marginalis **
| | | |
| | | | +----- Parreysia_keralaensis **
| | | | +-----------------+
| | | | | | +-- Parreysia_nagpoorensis **
| | | | | +--+
| | | | | +-- Parreysia_plagiosoma **
+--+ | | |
| | | | +-- Leoparreysia_tavoyensis **
| | | | |
| | +--+ +-----------------+-- Leoparreysia_canefrii **
| | | | |
| | | | +-- Leoparreysia_olivacea **
| | | |
| | | | +-------- Coelatura_aegyptiaca[2] **
| | | | |
| | | | +--------+ +----- Coelatura_aegyptiaca[1] **
| | +--+ | | |
| | | | | +----- Nitia_teretiuscula **
| | | | +--+
| | | | | +-- Coelatura_gabonensis **
| | | | | |
| | | | +--+-- Coelatura_kunenensis **
| | | | |
| | +--+ +-- Prisodontopsis_mweruensis **
| | |
| | | +-------- Indonaia_shurtleffiana **
| | | |
| | | +-----+ +----- Indonaia_bonneaudii **
| | | | | |
| | | | +--+----- Indonaia_rugosa **
| | | | |
| | | | | +-- Indonaia_andersoniana **
| | +--+ +--+
| | | +-- Indonaia_caerulea **
| | |
| | | +----------- Indochinella_pugio **
| | | |
| | | | +----- Scabies_humilis **
| | +--+ +--+
| | | | | +-- Scabies_phaselus **
| | | | +--+
| | +--+ +-- Scabies_mandarinus **
| | |
| | | +----- Radiatula_myitkyinae **
| | +--+
| | | +-- Radiatula_chaudhurii **
| | +--+
| | +-- Radiatula_mouhoti **
| |
| | +-------------------------------- Acuticosta_chinensis **
| | |
| | | +----------------------------- Lepidodesma_languilati **
| | | |
| | | | +-------- Aculamprotula_tortuosa **
| | | | |
| | | | +-----------+ +----- Aculamprotula_coreana **
| | | | | | |
| | | | | +--+----- Aculamprotula_scripta **
| | | | | |
+--+ | | | | +-- Aculamprotula_fibrosa **
| | | | +--+
| | | | +-- Aculamprotula_tientsinensis **
| +-----+ +--------+
| | | | | +-------- Unio_tumidus **
| | | | | +--------+
| | | | | | | +----- Unio_crassus **
| | | | | | +--+
| | | | | | | +-- Unio_delphinus **
| | | | | | +--+
| | | | +--+ +-- Unio_pictorum **
| | | | |
| | | | | +-- Schistodesmus_lampreyanus **
| | | | | +-----------+
| | | | | | +-- Schistodesmus_spinosus **
| | | | | |
| | | | +--+ +----- Nodularia_douglasiae[1] **
| | +--+ | +-----+
| | | | | | +-- Nodularia_douglasiae[2] **
| | | | | +--+
| | | +--+ +-- Nodularia_nuxpersicae **
| | | |
| | | | +-------- Tchangsinaia_piscicula **
| | | +--+
| | | | +----- Cuneopsis_rufescens **
| | | +--+
| | | | +-- Cuneopsis_heudei **
| | | +--+
| | | +-- Cuneopsis_celtiformis **
| | |
| | | +-------- Lanceolaria_gladiola **
| | | |
| | | +-----------------+ +----- Lanceolaria_grayii[1] **
| | | | | |
| | | | +--+----- Lanceolaria_grayii[2] **
| | | | |
| | | | | +-- Lanceolaria_lanceolata **
| | | | +--+
| | | | +-- Lanceolaria_triformis **
| | | |
| | | | +-------------------- Anemina_arcaeformis **
| | | | |
| | | | +--+ +----------------- Cristaria_plicata **
| | +--+ | | |
| | | | +--+ +-------------- Pletholophus_tenuis **
| | | | | |
| | | | +--+ +-- Sinanodonta_angula **
| | | | | +--------+
| | | | | | +-- Sinanodonta_lucida **
| | | | +--+
| | | | | +-------- Sinanodonta_woodiana[2] **
| | | | +--+
| | | | | +----- Sinanodonta_jourdyi **
| | | | +--+
| | +--+ | +-- Sinanodonta_elliptica **
| | | +--+
| | | +-- Sinanodonta_woodiana[1] **
| | |
| | | +-------- Alasmidonta_marginata **
| | | |
| | | +-------- Lasmigona_costata **
| | | |
| | | +-------- Pseudanodonta_complanata **
| | | |
| | | +-------- Simpsonaias_ambigua **
+--+ | |
| +--------------+-------- Strophitus_undulatus **
| |
| | +----- Anodonta_anatina **
| +--+
| | | +-- Anodonta_cygnea **
| | +--+
| | +-- Anodonta_nuttalliana **
| |
| | +----- Pyganodon_grandis **
| +--+
| | +-- Utterbackia_imbecillis **
| +--+
| +-- Utterbackia_peninsularis **
|
| +----------- Hyriopsis_desowitzi **
| |
| +----------- Rectidens_sumatrensis **
| |
| | +-- Hyriopsis_bialata **
| | |
| | +-- Hyriopsis_myersiana **
| +-----+--------+
| | | +-- Hyriopsis_panhai **
| | | |
| | | +-- Hyriopsis_khoratensis **
| | |
| | | +-- Ensidens_ingallsianus **
| | | +-----+
| | | | +-- Ensidens_telus **
| | +--+
| | | +----- Ensidens_sagittarius **
| +-----------------+ +--+
| | | | +-- Ensidens_spiculus **
| | | +--+
| | | +-- Ensidens_jaculus **
| | |
| | | +----- Physunio **
| | | +--------+
| | | | | +-- Physunio_modelli **
| | | | +--+
| | | | +-- Physunio_superbus **
| | | |
| | +--+ +-------- Yaukthwa_inlenensis **
| | | +--+
| | | | | +----- Yaukthwa_panhai **
| | | | +--+
| | | | | +-- Yaukthwa_nesemanni **
| | | | +--+
| | +--+ +-- Yaukthwa_rectangularis **
| | |
| | | +-- Lens_contradens **
| | | +-----+
| | | | +-- Lens_micropterus **
| | +--+
| | | +----- Lens_eximius **
| | +--+
| | | +-- Lens_comptus **
| | +--+
| | +-- Lens_novoselovi **
| |
| | +-- Megalonaias_nervosa **
| | +--+
+--+ | +-- Uniomerus_carolinianus **
| +-----------------------+
| | | +-- Quadrula_quadrula **
| | +--+
| | +-- Quadrula_verrucosa **
| |
| | +-------- Elliptoideus_sloatianus **
| +--+ |
| | | +-----------------+ +-- Elliptio_complanata **
| | | | | +--+
| | | | | | +-- Eurynia_dilatata **
| | | | +--+
| | | | | +-- Fusconaia_flava **
| | +--+ +--+
| | | +-- Pleurobema_sintoxia **
| | |
| | | +----------------------- Amblema_plicata **
| | | |
| | | | +----- Toxolasma_parvum **
| | +--+ +--------------+
| | | | | +-- Obliquaria_reflexa **
| | | | +--+
| | | | +-- Plectomerus_dombeyanus **
| | +--+
| | | +----------------- Potamilus_leptodon **
| | | |
| | | | +-------------- Potamilus_alatus **
| | +--+ |
| | | | +----------- Venustaconcha_ellipsiformis **
| | | | |
| | +--+ +----------- Leaunio_lienosus **
| | | |
+--+ | | +-- Ellipsaria_lineolata **
| | +--------+
| +--+ +-- Truncilla_truncata **
| |
| | +-- Medionidus_conradicus **
| +--------+
| | +-- Ptychobranchus_subtentus **
| |
| | +-------- Cambarunio_iris **
| +--+
| | +----- Ortmanniana_ligamentina **
| +--+
| | +-- Lampsilis_cardium **
| +--+
| +-- Lampsilis_ornata **
|
| +----- Chamberlainia_somsakpanhai **
| +-----------------------+
| | | +-- Sinohyriopsis_cumingii **
| | +--+
| | +-- Sinohyriopsis_schlegelii **
| |
| | +-- Potomida_littoralis **
| | +-----+
| | | +-- Pronodularia_japanensis **
| | +-----------------+
+--+ | | +----- Lamprotula_leaii **
| | +--+
| | | +-- Lamprotula_caveata **
| | +--+
| | +-- Lamprotula_cornuumlunae **
| |
| | +-------- Sinosolenaia_carinata **
| | +--+
| | | | +----- Sinosolenaia_recognita **
| | | +--+
+--+ | | +-- Sinosolenaia_oleivora **
| | +--+
| +-----------+ +-- Sinosolenaia_iridinea **
| | |
| | | +----- Gonidea_angulata **
| | | +--+
| | | | | +-- Leguminaia_wheatleyi **
| | | | +--+
| | +--+ +-- Microcondylaea_bonellii **
| | |
| | | +-- Ptychorhynchus_pfisteri **
| | | |
+--+ +-----+-- Parvasolenaia_rivularis **
| |
| +-- Koreosolenaia_sitgyensis **
|
| +----- Indopseudodon_salwenianus **
| +--------------+
| | | +-- Indopseudodon_indawgyiensis **
| | +--+
| | +-- Indopseudodon_bogani **
| |
| | +-------- Pilsbryoconcha_exilis[3] **
+--+ +--------+
| | | +----- Pilsbryoconcha_exilis[2] **
| | +--+
| | | +-- Pilsbryoconcha_exilis[1] **
| | +--+
| | +-- Pilsbryoconcha_carinifera **
| |
+--+ +----------- Nyeinchanconcha_nyeinchani **
| +--+
| | | +-------- Thaiconcha_callifera **
| | +--+
| | | +----- Bineurus_loeiensis **
| | +--+
| | | +-- Bineurus_mouhotii **
+--+ +--+
| +-- Bineurus_exilis **
|
| +----- Pseudodon_vagulus **
| +--+
| | | +-- Pseudodon_mekongi **
| | +--+
+-----+ +-- Pseudodon_vondembuschianus **
|
| +-- Sundadontina_cumingii **
| |
+-----+-- Sundadontina_tumida[1] **
|
+-- Sundadontina_tumida[2] **
Huang et al., 2019 [Publ 3843], fig. S4
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 176 |
176 |
108 |
2 |
26 |
24 |
2 |
155.0 |
+----------- Etheria_elliptica
|
| +-- Aspatharia_pfeifferiana
+--------+
| +-- Chambardia_wahlbergi
|
+-----------------------+ +----- Mycetopoda_siliquosa
| +-----+
| | | +-- Tamsiella_schroeteriana
| | +--+
| | +-- Lamproscapha_ensiformis
| |
| | +-------- Anodontites_trapesialis
| +--+
| | +----- Mutela_hargeri
| +--+
| | +-- Mutela_dubia
| +--+
| +-- Mutela_rostrata
|
| +----------- Echyridella_menziesii
| |
| +----------- Westralunio_carteri
| |
| | +----- Diplodon_suavidicus
| +-----+
| | | +-- Castalia_ambigua
| | +--+
| | +-- Triplodon_corrugatus
| |
| +--------------------+ +----- Hyridella_australis
| | +-----+
| | | | +-- Hyridella_depressa
| | | +--+
| | | +-- Hyridella_drapeta
| | |
| | | +----- Velesunio_ambiguus
| | | +--+
--+ | | | | +-- Alathyria_jacksoni
| | | | +--+
| | +--+ +-- Alathyria_profuga
| | |
| | | +----- Alathyria_pertexta
| | +--+
| | | +-- Lortiella_froggatti
| | +--+
| | +-- Microdontia_anodontaeformis
| |
| | +-------------------- Gibbosula_rochechouartii **
| | |
| | +--------+ +----------------- Gibbosula_laosensis **
| | | | |
| | | +--+ +-------------- Cumberlandia_monodonta **
| | | | |
| | | | | +----- Pseudunio_marocanus **
| | | +--+ +-----+
| | | | | | +-- Pseudunio_auricularius **
| | | | | +--+
| | | | | +-- Pseudunio_homsensis **
| | | +--+
| | | | +-------- Margaritifera_falcata **
| | | | |
| | | | | +-- Margaritifera_dahurica **
| | | | +-----+
| | | +--+ +-- Margaritifera_margaritifera **
| | | |
| | | | +----- Margaritifera_laevis **
| | | | |
| | | +--+----- Margaritifera_middendorffi **
| | | |
+--+ | | +-- Margaritifera_hembeli **
| | +--+
| | +-- Margaritifera_marrianae **
| |
| | +-- Parreysia_keralaensis **
| | |
| | +--------------------+-- Parreysia_nagpoorensis **
| | | |
| | | +-- Parreysia_plagiosoma **
| | |
| | | +----- Trapezidens_exolescens **
| | | +--------+
| | | | | +-- Trapezidens_angustior **
| | | | +--+
| | +--------+ +-- Trapezidens_dolichorhynchus **
| | | |
| | | | +----------- Lamellidens_corrianus **
| | | | |
| | | +--+ +-------- Lamellidens_savadiensis **
| | | | |
| | | | | +----- Lamellidens_marginalis **
| | | +--+ |
| | | | +----- Lamellidens_ferrugineus **
| | | | |
| | +--+ +--+ +-- Lamellidens_mainwaringi[1] **
| | | | | |
| | | | | +-- Lamellidens_mainwaringi[2] **
| | | | | |
| | | | +--+-- Lamellidens_lamellatus **
| | | | |
| | | | +-- Lamellidens_generosus **
| | | | |
| | | | +-- Lamellidens_jenkinsianus **
| | | |
| | | | +----- Leoparreysia_tavoyensis **
| | | | +--------------+
| | | | | | +-- Leoparreysia_canefrii **
| | | | | +--+
| | | | | +-- Leoparreysia_olivacea **
| | | | |
+--+ | | | +-------- Coelatura_aegyptiaca[2] **
| | | | |
| | | | +-------- Nitia_teretiuscula **
| | +--+ +--------+
| | | | | +----- Coelatura_aegyptiaca[1] **
| | | | | |
| | | | +--+ +-- Coelatura_gabonensis **
| | | | | |
| | | | +--+-- Coelatura_kunenensis **
| | | | |
| | | | +-- Prisodontopsis_mweruensis **
| | +--+
| | | +----- Indonaia_shurtleffiana **
| | | |
| | | | +-- Indonaia_bonneaudii **
| | | +--------+--+
| | | | | +-- Indonaia_rugosa **
| | | | |
| | | | | +-- Indonaia_andersoniana **
| | | | +--+
| | +--+ +-- Indonaia_caerulea **
| | |
| | | +----------- Indochinella_pugio **
| | | |
| | | | +----- Scabies_humilis **
| | +--+ +--+
| | | | | +-- Scabies_phaselus **
| | | | +--+
| | +--+ +-- Scabies_mandarinus **
| | |
| | | +----- Radiatula_mouhoti **
| | +--+
| | | +-- Radiatula_myitkyinae **
| | +--+
| | +-- Radiatula_chaudhurii **
| |
| | +-------------------- Chamberlainia_somsakpanhai **
| | |
| | +-------------------- Megalonaias_nervosa **
| | |
| | +-------------------- Uniomerus_carolinianus **
| | |
| | | +-- Quadrula_quadrula **
| | +-----------------+
| | | +-- Quadrula_verrucosa **
| | |
| | | +-- Sinohyriopsis_cumingii **
| | +-----------------+
| | | +-- Sinohyriopsis_schlegelii **
| | |
| | | +-- Potomida_littoralis **
| | | +-----+
| | | | +-- Pronodularia_japanensis **
| | +-----------+
| | | | +----- Lamprotula_leaii **
| | | +--+
| | | | +-- Lamprotula_caveata **
| | | +--+
| | | +-- Lamprotula_cornuumlunae **
| | |
| | | +-------- Sinosolenaia_carinata **
+--+ | |
| | | +----- Gonidea_angulata **
| | +--+
| | | | +-- Leguminaia_wheatleyi **
| | | +--+
| | | +-- Microcondylaea_bonellii **
| +-----------+
| | | +----- Ptychorhynchus_pfisteri **
| | +--+
| | | | +-- Parvasolenaia_rivularis **
| | | +--+
| | | +-- Koreosolenaia_sitgyensis **
| | |
| | | +----- Sinosolenaia_recognita **
| | +--+
| | | +-- Sinosolenaia_iridinea **
| | +--+
| | +-- Sinosolenaia_oleivora **
| |
| | +----- Indopseudodon_salwenianus **
| | +-----+
| | | | +-- Indopseudodon_indawgyiensis **
| | | +--+
| | | +-- Indopseudodon_bogani **
| | |
| | | +-------- Nyeinchanconcha_nyeinchani **
| | | |
| | | +-------- Thaiconcha_callifera **
| +--------+ |
| | | +-------- Sundadontina_cumingii **
| | | |
| | | +-------- Sundadontina_tumida[1] **
| | | |
| | | +-------- Sundadontina_tumida[2] **
| | | |
| | | | +----- Pilsbryoconcha_exilis[2] **
| | +--+--+
| | | | +-- Pilsbryoconcha_carinifera **
| | | +--+
| | | +-- Pilsbryoconcha_exilis[1] **
| | |
| | | +----- Pseudodon_vagulus **
| | +--+
| | | | +-- Pseudodon_mekongi **
| | | +--+
| | | +-- Pseudodon_vondembuschianus **
| | |
| | | +----- Bineurus_loeiensis **
| | +--+
| | | +-- Bineurus_mouhotii **
| | +--+
| | +-- Bineurus_exilis **
| |
| | +-------------- Amblema_plicata **
| | |
| | +-------------- Toxolasma_parvum **
| | |
| | | +-- Obliquaria_reflexa **
| | +-----------+
| | | +-- Plectomerus_dombeyanus **
| | |
| | | +----------- Elliptoideus_sloatianus **
| | +--+
| | | | +-------- Elliptio_complanata **
| | | +--+
| | | | +----- Eurynia_dilatata **
| +-----+ +--+
| | | | +-- Fusconaia_flava **
| | | +--+
| | | +-- Pleurobema_sintoxia **
| | |
| | | +-------- Ellipsaria_lineolata **
| | | |
| | | +-------- Potamilus_leptodon **
| | | |
| | | +-------- Medionidus_conradicus **
| | | |
+-----+ | +-------- Potamilus_alatus **
| | |
| | +-------- Ptychobranchus_subtentus **
| +-----+
| +-------- Truncilla_truncata **
| |
| +-------- Venustaconcha_ellipsiformis **
| |
| +-------- Cambarunio_iris **
| |
| +-------- Leaunio_lienosus **
| |
| | +----- Ortmanniana_ligamentina **
| +--+
| | +-- Lampsilis_cardium **
| +--+
| +-- Lampsilis_ornata **
|
| +-------------- Hyriopsis_desowitzi **
| |
| +-------------- Hyriopsis_myersiana **
| |
| +-------------- Hyriopsis_panhai **
| |
| +--+ +-- Hyriopsis_bialata **
| | +-----------+
| | | +-- Hyriopsis_khoratensis **
| | |
| | | +----------- Rectidens_sumatrensis **
| | | |
| | +--+ +-- Ensidens_telus **
| | | +-----+
| | | | +-- Ensidens_ingallsianus **
| | +--+
| | | +----- Ensidens_sagittarius **
| | +--+
+--+ | +-- Ensidens_spiculus **
| | +--+
| | +-- Ensidens_jaculus **
| |
| | +----------- Physunio_modelli **
| | |
| | +----------- Lens **
| | |
| | +----------- Physunio_superbus **
| | |
| | | +-------- Yaukthwa_inlenensis **
| | +--+
| +-----+ | +----- Yaukthwa_panhai **
| | +--+
| | | +-- Yaukthwa_nesemanni **
| | +--+
| | +-- Yaukthwa_rectangularis **
| |
| | +-- Lens_comptus **
| | +-----+
| | | +-- Lens_novoselovi **
| +--+
| | +----- Lens_eximius **
| +--+
| | +-- Lens_contradens **
| +--+
| +-- Lens_micropterus **
|
| +----------------- Acuticosta_chinensis **
| |
| +----------------- Alasmidonta_marginata **
| |
| +----------------- Anodonta_anatina **
| |
| +----------------- Anemina_arcaeformis **
| |
| +----------------- Anodonta_cygnea **
| |
| +----------------- Cristaria_plicata **
| |
| +----------------- Anodonta_nuttalliana **
| |
| +----------------- Lasmigona_costata **
| |
| +----------------- Lepidodesma_languilati **
| |
| +----------------- Pseudanodonta_complanata **
| |
| +----------------- Pyganodon_grandis **
| |
| +----------------- Simpsonaias_ambigua **
| |
| +----------------- Strophitus_undulatus **
| |
| | +-- Schistodesmus_lampreyanus **
| +--------------+
| | +-- Schistodesmus_spinosus **
| |
| | +-- Utterbackia_imbecillis **
| +--------------+
| | +-- Utterbackia_peninsularis **
| |
| | +-------- Lanceolaria_gladiola **
| +--------+
| | | +----- Lanceolaria_grayii **
| | +--+
+--+ | +-- Lanceolaria_lanceolata **
| +--+
| +-- Lanceolaria_triformis **
|
| +-------- Unio_tumidus **
+--------+
| | +----- Unio_crassus **
| +--+
| | +-- Unio_delphinus **
| +--+
| +-- Unio_pictorum **
|
| +-------- Aculamprotula_tortuosa **
| |
+--------+ +-- Aculamprotula_coreana **
| | +--+
| | | +-- Aculamprotula_scripta **
| +--+
| | +-- Aculamprotula_fibrosa **
| +--+
| +-- Aculamprotula_tientsinensis **
|
| +-------- Cuneopsis_heudei **
| |
| +-------- Cuneopsis_celtiformis **
| |
+--------+-------- Tchangsinaia_piscicula **
| |
| +-------- Cuneopsis_rufescens **
| |
| | +----- Nodularia_douglasiae[2] **
| +--+
| | +-- Nodularia_douglasiae[1] **
| +--+
| +-- Nodularia_nuxpersicae **
|
| +-------------- Pletholophus_tenuis **
| |
+--+ +-- Sinanodonta_angula **
| +--------+
| | +-- Sinanodonta_lucida **
+--+
| +-------- Sinanodonta_woodiana[1] **
+--+
| +----- Sinanodonta_jourdyi **
+--+
| +-- Sinanodonta_elliptica **
+--+
+-- Sinanodonta_woodiana[2] **
Huang et al., 2019 [Publ 3843], fig. S3
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 175 |
175 |
168 |
2 |
26 |
24 |
2 |
152.0 |
+----- Velesunio_ambiguus
+--+
| | +-- Alathyria_jacksoni
| +--+
+--------+ +-- Alathyria_profuga
| |
| | +----- Alathyria_pertexta
| +--+
| | +-- Lortiella_froggatti
+-----------------------------------------------+ +--+
| | +-- Microdontia_anodontaeformis
| |
| | +-------------- Westralunio_carteri
| | |
| +--+ +----------- Echyridella_menziesii
| | |
| | | +----- Diplodon_suavidicus
| +--+ +--+
| | | | +-- Castalia_ambigua
| | | +--+
| +--+ +-- Triplodon_corrugatus
| |
| | +----- Hyridella_australis
| +--+
| | +-- Hyridella_depressa
| +--+
| +-- Hyridella_drapeta
|
| +-------- Anodontites_trapesialis
| +--+
| | | +----- Mutela_rostrata
| | +--+
| | | +-- Mutela_dubia
| | +--+
| +--------------------------------------------------+ +-- Mutela_hargeri
| | |
| | | +----- Mycetopoda_siliquosa
--+ | | +--+
| | | | | +-- Tamsiella_schroeteriana
| | | | +--+
| | +--+ +-- Lamproscapha_ensiformis
| | |
| | | +----- Etheria_elliptica
| | +--+
| | | +-- Aspatharia_pfeifferiana
| | +--+
| | +-- Chambardia_wahlbergi
| |
| | +-------------------- Gibbosula_rochechouartii **
| | |
| | | +----------------- Gibbosula_laosensis **
| | +--------------------------------------+ |
| | | | | +-------- Cumberlandia_monodonta **
| | | | | +-----+
| | | +--+ | | +----- Pseudunio_marocanus **
| | | | | +--+
| | | | | | +-- Pseudunio_auricularius **
| | | | | +--+
| | | +--+ +-- Pseudunio_homsensis **
| | | |
| | | | +-- Margaritifera_dahurica **
| | | | +--------+
| | | | | +-- Margaritifera_margaritifera **
| | | +--+
| | | | +-------- Margaritifera_falcata **
+--+ | | |
| | +--+ +-- Margaritifera_hembeli **
| | | +--+
| | | | +-- Margaritifera_marrianae **
| | +--+
| | | +-- Margaritifera_laevis **
| | +--+
| | +-- Margaritifera_middendorffi **
| |
| | +----- Trapezidens_exolescens **
| | +-----+
| | | | +-- Trapezidens_angustior **
| | | +--+
| | | +-- Trapezidens_dolichorhynchus **
| | |
| | +--------------+ +-- Lamellidens_corrianus **
| | | | +-----+
| | | | | +-- Lamellidens_savadiensis **
| | | | |
| | | | | +----- Lamellidens_lamellatus **
| | | +--+ |
| | | | +----- Lamellidens_jenkinsianus **
| | | | |
| | | | | +-- Lamellidens_marginalis **
| | | +--+--+
| | | | +-- Lamellidens_ferrugineus **
| | | |
| | +-----------------------------+ | +-- Lamellidens_mainwaringi[1] **
| | | | | |
| | | | +--+-- Lamellidens_mainwaringi[2] **
| | | | |
| | | | +-- Lamellidens_generosus **
+--+ | |
| | | +----- Parreysia_keralaensis **
| | | +-----------------+
| | | | | +-- Parreysia_nagpoorensis **
| | | | +--+
| | | | +-- Parreysia_plagiosoma **
| | | |
| | | | +----- Leoparreysia_canefrii **
| | +--+ +--------------+
| | | | | +-- Leoparreysia_tavoyensis **
| | | | +--+
| | | | +-- Leoparreysia_olivacea **
| | | |
| | | | +----------- Coelatura_aegyptiaca[2] **
| | | | |
| | +--+ +-----+ +-- Coelatura_aegyptiaca[1] **
| | | | | +-----+
| | | | | | +-- Nitia_teretiuscula **
| | | | +--+
| | | | | +----- Coelatura_kunenensis **
| | | | +--+
| | | | | +-- Coelatura_gabonensis **
| | | | +--+
| | +--+ +-- Prisodontopsis_mweruensis **
| | |
| | | +-------- Indonaia_shurtleffiana **
| | | |
| | | +-----+ +----- Indonaia_bonneaudii **
| | | | | |
| | | | +--+----- Indonaia_rugosa **
| | | | |
| | | | | +-- Indonaia_andersoniana **
| | +--+ +--+
| | | +-- Indonaia_caerulea **
| | |
| | | +----------- Indochinella_pugio **
| | | |
| | | | +----- Scabies_phaselus **
| | +--+ +--+
| | | | | +-- Scabies_humilis **
| | | | +--+
| | +--+ +-- Scabies_mandarinus **
| | |
| | | +----- Radiatula_mouhoti **
| | +--+
| | | +-- Radiatula_myitkyinae **
| | +--+
+--+ +-- Radiatula_chaudhurii **
|
| +----- Indopseudodon_indawgyiensis **
| +--------------+
| | | +-- Indopseudodon_bogani **
| | +--+
| | +-- Indopseudodon_salwenianus **
| |
| | +-------- Pilsbryoconcha_exilis[3] **
| +--------------------------------+ +--------+
| | | | | +----- Pilsbryoconcha_exilis[2] **
| | | | +--+
| | | | | +-- Pilsbryoconcha_carinifera **
| | | | +--+
| | | | +-- Pilsbryoconcha_exilis[1] **
| | | |
| | +--+ +----------- Sundadontina_tumida **
| | | +--+
| | | | | +-------- Pseudodon_vagulus **
| | | | +--+
| | | | | +----- Pseudodon_vondembuschianus[1] **
| | | | +--+
| | | | | +-- Pseudodon_mekongi **
| | +--+ +--+
| | | +-- Pseudodon_vondembuschianus[2] **
| | |
| | | +-- Nyeinchanconcha_nyeinchani **
| | | +--------+
| | | | +-- Sundadontina_cumingii **
| | +--+
| | | +-------- Thaiconcha_callifera **
| | +--+
| | | +----- Bineurus_loeiensis **
| | +--+
| | | +-- Bineurus_mouhotii **
| | +--+
| | +-- Bineurus_exilis **
| |
| | +-- Ensidens_telus **
| | +-----+
| | | +-- Ensidens_ingallsianus **
| | +-----+
| | | | +----- Ensidens_sagittarius **
| | | +--+
| | | | +-- Ensidens_spiculus **
| | +--+ +--+
| | | | +-- Ensidens_jaculus **
| | | |
| | | | +----------- Hyriopsis_desowitzi **
| | | | |
| | | +--+ +-------- Rectidens_sumatrensis **
| | | | |
| | | +--+ +-- Hyriopsis_panhai **
+--+ | | +--+
| | | | +-- Hyriopsis_khoratensis **
| +--------------------------------+ +--+
| | | | +-- Hyriopsis_bialata **
| | | +--+
| | | +-- Hyriopsis_myersiana **
| | |
| | | +----- Physunio **
| | | +--------+
| | | | | +-- Physunio_superbus **
| | | | +--+
| | | | +-- Physunio_modelli **
| | | |
| | +--+ +-------- Yaukthwa_inlenensis **
| | | +--+
| | | | | +----- Yaukthwa_panhai **
| | | | +--+
| | | | | +-- Yaukthwa_nesemanni **
| | | | +--+
| | +--+ +-- Yaukthwa_rectangularis **
| | |
| | | +-- Lens_contradens **
| | | +-----+
| | | | +-- Lens_micropterus **
| | +--+
| | | +----- Lens_eximius **
| | +--+
| | | +-- Lens_novoselovi **
| | +--+
| | +-- Lens_comptus **
| |
| | +----- Chamberlainia_somsakpanhai **
| | +--------------------------------------+
| | | | +-- Sinohyriopsis_cumingii **
| | | +--+
| | | +-- Sinohyriopsis_schlegelii **
| | |
| | | +-- Potomida_littoralis **
| | | +-----+
| | | | +-- Pronodularia_japanensis **
| | | +-----+
| | | | | +----- Lamprotula_leaii **
| | | | +--+
| | | | | +-- Lamprotula_caveata **
+--+ | | +--+
| | | +-- Lamprotula_cornuumlunae **
| +--+ +--------------------------+
| | | | | +----- Sinosolenaia_recognita **
| | | | | +-----+
| | | | | | | +-- Sinosolenaia_oleivora **
| | | | | | +--+
| | | | | | +-- Sinosolenaia_iridinea **
| | | | +--+
| | | | | +----- Gonidea_angulata **
| | | | | +--+
| | | | | | | +-- Leguminaia_wheatleyi **
| | | | | | +--+
| | | | +--+ +-- Microcondylaea_bonellii **
| | | | |
| | | | | +----- Ptychorhynchus_pfisteri **
| | +--+ +--+
| | | | +-- Parvasolenaia_rivularis **
| | | +--+
| | | +-- Koreosolenaia_sitgyensis **
| | |
| | | +-- Megalonaias_nervosa **
| | | +--+
| | | | +-- Uniomerus_carolinianus **
| | | +--------------------------------+
| | | | | +-- Quadrula_quadrula **
| | | | +--+
| | | | +-- Quadrula_verrucosa **
| | | |
| | | | +-------- Elliptoideus_sloatianus **
| | +--+ |
| | | +--------------------------+ +-- Elliptio_complanata **
| | | | | +--+
| | | | | | +-- Eurynia_dilatata **
| | | | +--+
| | | | | +-- Fusconaia_flava **
| | +--+ +--+
| | | +-- Pleurobema_sintoxia **
| | |
| | | +-------------------------------- Amblema_plicata **
| | | |
| | +--+ +----------------------------- Toxolasma_parvum **
| | | |
+--+ +--+ +-- Obliquaria_reflexa **
| | +-----------------------+
| | | +-- Plectomerus_dombeyanus **
| +--+
| | +----------------------- Potamilus_leptodon **
| | |
| +--+ +-------------------- Potamilus_alatus **
| | |
| | | +-- Medionidus_conradicus **
| +--+ +--------------+
| | | +-- Ptychobranchus_subtentus **
| | |
| +--+ +-- Ellipsaria_lineolata **
| | +-----------+
| | | +-- Truncilla_truncata **
| +--+
| | +----------- Leaunio_lienosus **
| | |
| +--+ +-- Venustaconcha_ellipsiformis **
| | +-----+
| | | +-- Cambarunio_iris **
| +--+
| | +----- Ortmanniana_ligamentina **
| +--+
| | +-- Lampsilis_cardium **
| +--+
| +-- Lampsilis_ornata **
|
| +----------------------------------- Acuticosta_chinensis **
| |
| | +-------- Aculamprotula_tortuosa **
| | |
| | +-----------+ +-- Aculamprotula_coreana **
| | | | +--+
| | | | | +-- Aculamprotula_scripta **
| | | +--+
| | | | +-- Aculamprotula_fibrosa **
| | | +--+
| | | +-- Aculamprotula_tientsinensis **
| | +-----------+
| | | | +-------- Unio_tumidus **
+-----------+ | | +--------+
| | | | | +----- Unio_crassus **
| | | | +--+
| | | | | +-- Unio_delphinus **
| | | | +--+
| | +--+ +-- Unio_pictorum **
| | |
| | | +----- Nodularia_douglasiae[1] **
| | | +--------+
| | | | | +-- Nodularia_douglasiae[2] **
| | | | +--+
| | +--+ +-- Nodularia_nuxpersicae **
| | |
+--+ | +----------- Schistodesmus_lampreyanus **
| +--+
| | +-------- Tchangsinaia_piscicula **
| +--+
| | +----- Cuneopsis_rufescens **
| +--+
| | +-- Cuneopsis_heudei **
| +--+
| +-- Cuneopsis_celtiformis **
|
| +----------------------------- Lepidodesma_languilati **
| |
| | +-------- Lanceolaria_gladiola **
| | +-----------------+
| | | | +----- Lanceolaria_grayii **
| | | +--+
+--+ | | +-- Lanceolaria_lanceolata **
| | +--+
| | +-- Lanceolaria_triformis **
| |
| | +-------------------- Anemina_arcaeformis **
| | |
| | +--+ +----------------- Cristaria_plicata **
+--+ | | |
| | +--+ +-------------- Pletholophus_tenuis **
| | | |
| | +--+ +-- Sinanodonta_angula **
| | | +--------+
| | | | +-- Sinanodonta_lucida **
| | +--+
| | | +-------- Sinanodonta_woodiana[2] **
| | +--+
| | | +----- Sinanodonta_jourdyi **
+--+ +--+
| | +-- Sinanodonta_elliptica **
| +--+
| +-- Sinanodonta_woodiana[1] **
|
| +-- Alasmidonta_marginata **
| +--------+
| | +-- Lasmigona_costata **
| |
| | +----- Anodonta_anatina **
| +-----+
| | | +-- Anodonta_cygnea **
+-----------+ +--+
| +-- Anodonta_nuttalliana **
|
| +----- Pseudanodonta_complanata **
| +--+
| | | +-- Simpsonaias_ambigua **
| | +--+
+--+ +-- Strophitus_undulatus **
|
| +----- Pyganodon_grandis **
+--+
| +-- Utterbackia_imbecillis **
+--+
+-- Utterbackia_peninsularis **
Bolotov et al., 2017 [Publ 3764], fig. S1
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 131 |
131 |
72 |
1 |
0 |
0 |
0 |
130.0 |
+----- Trapezidens_exolescens **
+--------+
| | +-- Trapezidens_dolichorhynchus **
| +--+
+-----------+ +-- Trapezidens_angustior **
| |
| | +----------- Lamellidens_lamellatus **
| | |
| +--+ +-------- Lamellidens_jenkinsianus **
| | |
| | | +----- Lamellidens_generosus **
| +--+ |
| | +----- Lamellidens_mainwaringi **
| | |
| +--+ +-- Lamellidens_ferrugineus **
| +--+
| | +-- Lamellidens_marginalis **
| |
+--+ | +-- Lamellidens_savadiensis **
| | +--+
| | +-- Lamellidens_corrianus[1] **
| |
| | +-------- Parreysia_keralaensis **
| | +--------------+
| | | | +----- Parreysia_plagiosoma **
| | | +--+
| | | | +-- Parreysia_corrugata **
| | | +--+
| | | +-- Parreysia_nagpoorensis **
| | |
| | | +-------- Leoparreysia_olivacea **
| | | +-----------+
| | | | | +----- Leoparreysia_tavoyensis[2] **
| +--+ | +--+
| | | | +-- Leoparreysia_canefrii **
| | | +--+
| | | +-- Leoparreysia_tavoyensis[1] **
| | |
| | | +-- Nitia_teretiuscula **
| | | |
| | | +-- Coelatura_gabonensis **
| | | |
| | | +-- Prisodontopsis_mweruensis **
| +--+ |
| | +-- Coelatura_kunenensis **
| | |
| | +--------------+-- Coelatura_aegyptiaca[1] **
| | | |
| | | +-- Coelatura_aegyptiaca[2] **
| | | |
| | | +-- Prisodontopsis_aviculaeformis **
| | | |
| | | +-- Coelatura_luapulaensis **
| | | |
| | | +-- Coelatura_choziensis **
| +--+
| | +-------- Indonaia_shurtleffiana **
| | |
| | +-----+ +----- Indonaia_rugosa **
| | | | |
| | | | | +-- Indonaia_andersoniana **
| | | +--+--+
| | | | +-- Indonaia_caerulea **
| | | |
| +--+ | +-- Indonaia_bonneaudii **
| | +--+
| | +-- Lamellidens_corrianus[2] **
| |
| | +----------- Indochinella_pugio **
| | |
| | | +-- Scabies_humilis **
| +--+ |
| | +-----+-- Scabies_phaselus **
--+ | | |
| +--+ +-- Scabies_mandarinus **
| |
| | +----- Radiatula_mouhoti **
| +--+
| | +-- Radiatula_myitkyinae **
| +--+
| +-- Radiatula_chaudhurii **
|
| +----- Indopseudodon_indawgyiensis **
| +-----------+
| | | +-- Indopseudodon_salwenianus **
| | +--+
| | +-- Indopseudodon_bogani **
| |
| | +-- Nyeinchanconcha_nyeinchani **
| | +-----------+
| +-----+ | +-- Sundadontina_cumingii **
| | | |
| | | | +----- Bineurus_exilis **
| | | +--------+
| | | | | +-- Bineurus_mouhotii **
| | | | +--+
| | | | +-- Bineurus_loeiensis **
| | | |
| | +--+ +-------- Sundadontina_tumida **
| | +-----+
| | | | +----- Pseudodon_vagulus **
| | | +--+
| | | | +-- Pseudodon_mekongi **
| | | +--+
| | | +-- Pseudodon_vondembuschianus **
| | |
| | | +----------- Thaiconcha_callifera **
| | +--+
| | | +-------- Pilsbryoconcha_exilis[3] **
| | +--+
| | | +----- Pilsbryoconcha_exilis[2] **
| | +--+
| | | +-- Pilsbryoconcha_exilis[1] **
| | +--+
| | +-- Pilsbryoconcha_carinifera **
| |
| | +-- Ensidens_telus **
| | +-----+
| | | +-- Ensidens_ingallsianus **
| | +--+
| | | | +----- Ensidens_jaculus **
| | | +--+
| | | | +-- Ensidens_spiculus **
| | | +--+
| | +-----+ +-- Ensidens_sagittarius **
| | | |
| | | | +-------- Hyriopsis_desowitzi **
| | | | |
| | | | +-------- Rectidens_sumatrensis **
+-----+ | +--+
| | | +----- Hyriopsis_panhai **
| | | |
| | +--+ +-- Hyriopsis_myersiana **
| | | |
| +--+ +--+-- Hyriopsis_bialata **
| | | |
| | | +-- Hyriopsis_khoratensis **
| | |
| | | +----- Physunio **
| | | +--------+
| | | | | +-- Physunio_superbus **
| | | | +--+
| | | | +-- Physunio_modelli **
| | | |
| | +--+ +-------- Yaukthwa_inlenensis **
| | | +--+
| | | | | +----- Yaukthwa_panhai **
| | | | +--+
| | | | | +-- Yaukthwa_nesemanni **
| | | | +--+
| | +--+ +-- Yaukthwa_rectangularis **
| | |
| | | +-- Lens_contradens **
| | | +-----+
| | | | +-- Lens_micropterus **
| | +--+
| | | +----- Lens_eximius **
| | +--+
| | | +-- Lens_comptus **
| | +--+
| | +-- Lens_novoselovi **
| |
| | +-- Amblema_plicata **
| | |
| | +-- Ortmanniana_ligamentina **
| | |
| | +-- Lampsilis_cardium **
| | |
| | +-- Cambarunio_iris **
+--+ |
| +--+-- Elliptio_complanata **
| | |
| | +-- Eurynia_dilatata **
| | |
| | +-- Pleurobema_sintoxia **
| | |
| | +-- Quadrula_quadrula **
| | |
| | +-- Quadrula_verrucosa **
| |
| | +-- Chamberlainia_somsakpanhai **
| +--+ |
| | | +-- Sinohyriopsis_cumingii **
| | | |
| | | +-- Lamprotula_caveata **
| | | |
| | | +-- Lamprotula_leaii **
| | | |
| | | +-- Potomida_littoralis **
| | | |
| | | +-- Pronodularia_japanensis **
| | | |
| | +--+-- Gonidea_angulata **
| | |
| | +-- Leguminaia_wheatleyi **
| | |
| | +-- Leguminaia **
| | |
| | +-- Microcondylaea_bonellii **
| | |
| | +-- Sinosolenaia_carinata **
| | |
| | +-- Sinosolenaia_oleivora **
| | |
| | +-- Sinosolenaia_recognita **
+-----------+
| +-- Unio_crassus **
| |
| +-- Unio_pictorum **
| |
| +-- Unio_tumidus **
| |
| +-- Aculamprotula_tortuosa **
| |
| +-- Cuneopsis_heudei **
| +--+
| | +-- Tchangsinaia_piscicula **
| | |
| | +-- Cuneopsis_rufescens **
| | |
| | +-- Nodularia_douglasiae **
| | |
| | +-- Nodularia_nuxpersicae **
| | |
| | +-- Schistodesmus_lampreyanus **
| |
| | +-- Alasmidonta_marginata **
| | |
| | +-- Anodonta_anatina **
+--+ |
| +-- Anodonta_cygnea **
| |
| +-- Anodonta_nuttalliana **
| |
| +-- Lasmigona_costata **
| |
| +-- Pseudanodonta_complanata **
| |
| +-- Pyganodon_grandis **
| |
| +-- Simpsonaias_ambigua **
| |
| +-- Strophitus_undulatus **
+--+
+-- Sinanodonta_lauta **
|
+-- Cristaria_plicata **
|
+-- Cristaria_radiata **
|
+-- Pletholophus_tenuis **
|
+-- Sinanodonta_lucida **
|
+-- Sinanodonta_jourdyi **
|
+-- Lanceolaria_gladiola **
|
+-- Lanceolaria_grayii[1] **
|
+-- Lanceolaria_grayii[2] **
Bolotov et al., 2017 [Publ 3761], fig. S1
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 126 |
139 |
137 |
2 |
15 |
13 |
2 |
109.0 |
+----------------- Cumberlandia_monodonta **
|
| +----- Gibbosula_laosensis **
+-----------------------+ +--------+
| | | | +-- Pseudunio_auricularius **
| | | +--+
| | | +-- Pseudunio_marocanus **
| +--+
| | +-- Margaritifera_dahurica **
| | +--------+
| | | +-- Margaritifera_margaritifera **
| +--+
| | +-------- Margaritifera_falcata **
| +--+
| | +----- Margaritifera_middendorffi **
| +--+
| | +-- Margaritifera_laevis **
| +--+
| +-- Margaritifera_marrianae **
|
| +-------- Tamsiella_schroeteriana
| +-----+
| | | +----- Etheria_elliptica
| | +--+
| | | +-- Chambardia_wahlbergi
| | +--+
| | +-- Aspatharia_pfeifferiana
| +-----------------------+
| | | +----- Diplodon_suavidicus
| | | +-----+
| | | | | +-- Castalia_ambigua
| | | | +--+
| | | | +-- Triplodon_corrugatus
| | +--+
--+ | | +----- Alathyria_pertexta
| | | +--+
| | | | | +-- Microdontia_anodontaeformis
| | | | +--+
| | +--+ +-- Lortiella_froggatti
| | |
| | | +----- Velesunio_ambiguus
| | +--+
| | | +-- Alathyria_jacksoni
| | +--+
| | +-- Alathyria_profuga
| |
| | +-- Trapezidens_dolichorhynchus **
| | +-----------+
| | | +-- Trapezidens_exolescens **
| | +-----------+
| | | | +----------- Lamellidens_corrianus **
| | | | |
| | | +--+ +-------- Lamellidens_savadiensis **
| | | | |
| | | +--+ +-- Lamellidens_jenkinsianus **
| | | | +--+
| | | | | +-- Lamellidens_mainwaringi **
| | | +--+
| | +--------+ | +-- Lamellidens_ferrugineus **
| | | | +--+
| | | | +-- Lamellidens_marginalis **
| | | |
| | | | +-- Parreysia_nagpoorensis **
| | | | +--------------------+
| | | | | +-- Parreysia_plagiosoma **
| | | | |
+--+ | | | +-- Leoparreysia_tavoyensis **
| | | | +-----------------+
| | +--+ | +-- Leoparreysia_olivacea **
| | | |
| | | | +-------------- Nitia_teretiuscula **
| | | | +--+
| | | | | | +----------- Coelatura_aegyptiaca[2] **
| | +--+ | +--+
| | | | | +-------- Coelatura_aegyptiaca[1] **
| | | | +--+
| | | | | +----- Prisodontopsis_mweruensis **
| | | | +--+
| | | | | +-- Coelatura_gabonensis **
| | +--+ +--+
| | | +-- Coelatura_kunenensis **
| | |
| | | +-------- Indonaia_shurtleffiana **
| | | +-----+
| | | | | +----- Indonaia_andersoniana **
| | | | +--+
| | | | | +-- Indonaia_rugosa **
| | | | +--+
| | +--+ +-- Indonaia_bonneaudii **
| | |
| | | +-- Indochinella_pugio **
| | | +--------+
| | | | +-- Scabies_crispata **
| | | |
| | +--+ +-- Radiatula_myitkyinae **
| | | +-----+
| | | | +-- Radiatula_chaudhurii **
| | +--+
| | | +----- Scabies_mandarinus **
| | +--+
| | | +-- Scabies_humilis **
| | +--+
| | +-- Scabies_phaselus **
| |
| | +-- Ensidens_telus **
| | +-----+
+--+ | +-- Ensidens_ingallsianus **
| +-----+
| | | +----- Ensidens_sagittarius **
| | +--+
| | | +-- Ensidens_spiculus **
| | +--+
| +--+ +-- Ensidens_jaculus **
| | |
| | | +-- Hyriopsis_bialata **
| | | +--------+
| | | | +-- Hyriopsis_khoratensis **
| | +--+
| | | +-------- Hyriopsis_panhai **
| | +--+
| | | +----- Hyriopsis_myersiana **
| | +--+
| +--------------+ | +-- Rectidens_sumatrensis **
| | | +--+
| | | +-- Hyriopsis_desowitzi **
| | |
| | | +----- Physunio **
| | | +--------+
| | | | | +-- Physunio_modelli **
| | | | +--+
| | | | +-- Physunio_superbus **
| | | |
| | +--+ +-------- Yaukthwa_inlenensis **
| | | +--+
| | | | | +----- Yaukthwa_panhai **
| | | | +--+
| | | | | +-- Yaukthwa_rectangularis **
| | | | +--+
| | +--+ +-- Yaukthwa_nesemanni **
| | |
| | | +-- Lens_contradens **
| | | +-----+
| | | | +-- Lens_micropterus **
| | +--+
| | | +----- Lens_eximius **
| | +--+
| | | +-- Lens_comptus **
| | +--+
| | +-- Lens_novoselovi **
| |
| | +-------------------- Indopseudodon_indawgyiensis **
| | |
| | | +-------- Pilsbryoconcha_exilis[1] **
| | | +--------+
+--+ +--------+ | | +----- Pilsbryoconcha_exilis[3] **
| | | | +--+
| | | | | +-- Pilsbryoconcha_exilis[2] **
| | | | +--+
| | | | +-- Pilsbryoconcha_carinifera **
| | +--+
| | | +-------- Sundadontina_tumida **
| | | +-----+
| | | | | +----- Pseudodon_vagulus **
| | | | +--+
| | | | | +-- Pseudodon_mekongi **
| | | | +--+
| | +--+ +-- Pseudodon_vondembuschianus **
| | |
| | | +-- Nyeinchanconcha_nyeinchani **
| | | +--------+
| | | | +-- Sundadontina_cumingii **
| | +--+
| | | +-------- Thaiconcha_callifera **
| | +--+
| | | +----- Bineurus_exilis **
| | +--+
| | | +-- Bineurus_mouhotii **
| | +--+
| | +-- Bineurus_loeiensis **
| |
| | +-------- Amblema_plicata **
| | +--+
| | | | +----- Ortmanniana_ligamentina **
| | | +--+
| | | | +-- Lampsilis_cardium **
| | | +--+
+--+ +-----+ +-- Cambarunio_iris **
| | |
| | | +-- Quadrula_quadrula **
| | | +-----+
| | | | +-- Quadrula_verrucosa **
| | +--+
| | | +----- Pleurobema_sintoxia **
| | +--+
| | | +-- Eurynia_dilatata **
| | +--+
| | +-- Elliptio_complanata **
| +--------+
| | | +-- Sinohyriopsis_cumingii **
| | | +-----+
| | | | +-- Chamberlainia_somsakpanhai **
| | | |
| | | +-----+ +-- Potomida_littoralis **
| | | | | +--+
| | | | | | +-- Pronodularia_japanensis **
| | | | +--+
| | | | | +-- Lamprotula_caveata **
| | | | +--+
| | +--+ +-- Lamprotula_leaii **
| | |
| | | +----- Sinosolenaia_carinata **
| | | +-----+
| | | | | +-- Sinosolenaia_oleivora **
| | | | +--+
| | +--+ +-- Sinosolenaia_recognita **
| | |
| | | +-------- Gonidea_angulata **
| | +--+
+--+ | +----- Microcondylaea_bonellii **
| +--+
| | +-- Leguminaia_wheatleyi **
| +--+
| +-- Leguminaia **
|
| +----- Unio_tumidus **
| +--------+
| | | +-- Unio_pictorum **
| | +--+
| | +-- Unio_crassus **
| +--------+
| | | +----------- Aculamprotula_tortuosa **
| | | |
| | | | +-- Nodularia_douglasiae **
| | +--+ +-----+
| | | | +-- Nodularia_nuxpersicae **
| | | |
| | +--+ +-- Cuneopsis_rufescens **
| | | +--+
| | | | +-- Cuneopsis_heudei **
| | +--+
+--+ | +-- Tchangsinaia_piscicula **
| +--+
| +-- Schistodesmus_lampreyanus **
|
| +-- Lanceolaria_grayii **
| +-----------------+
| | +-- Lanceolaria_gladiola **
| |
| | +-- Cristaria_plicata **
| | +--------+
| | | +-- Cristaria_radiata **
+--+ +-----+
| | | +-------- Pletholophus_tenuis **
| | +--+
| | | +----- Sinanodonta_lucida **
| | +--+
| | | +-- Sinanodonta_jourdyi **
+--+ +--+
| +-- Sinanodonta_lauta **
|
| +-------------- Anodonta_anatina **
| |
| | +----- Pseudanodonta_complanata **
+--+ +-----+
| | | +-- Anodonta_cygnea **
| | +--+
| | +-- Anodonta_nuttalliana **
+--+
| +-- Lasmigona_costata **
| +-----+
| | +-- Alasmidonta_marginata **
+--+
| +----- Simpsonaias_ambigua **
+--+
| +-- Pyganodon_grandis **
+--+
+-- Strophitus_undulatus **
Bolotov et al., 2017 [Publ 3761], fig. S2
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 126 |
139 |
123 |
2 |
15 |
13 |
2 |
113.0 |
+-------------------- Chamberlainia_somsakpanhai **
|
+-----------+ +----------------- Cumberlandia_monodonta **
| | |
| | | +-------------- Gibbosula_laosensis **
| +--+ |
| | | +-- Pseudunio_auricularius **
| | +-----------+
| +--+ +-- Pseudunio_marocanus **
| |
| | +-- Margaritifera_dahurica **
| | +--------+
| | | +-- Margaritifera_margaritifera **
| +--+
| | +-------- Margaritifera_falcata **
| +--+
| | +----- Margaritifera_middendorffi **
| +--+
| | +-- Margaritifera_laevis **
| +--+
| +-- Margaritifera_marrianae **
|
| +-- Tamsiella_schroeteriana
| +--+
| | +-- Etheria_elliptica
| +--------+
| | | +-- Aspatharia_pfeifferiana
| | +--+
| | +-- Chambardia_wahlbergi
| |
+-----------------+ +----- Diplodon_suavidicus
| | +-----+
| | | | +-- Castalia_ambigua
| | | +--+
| | | +-- Triplodon_corrugatus
| +--+
| | +----- Velesunio_ambiguus
| | +--+
| | | | +-- Alathyria_jacksoni
| | | +--+
| +--+ +-- Alathyria_profuga
| |
| | +----- Microdontia_anodontaeformis
| +--+
| | +-- Alathyria_pertexta
| +--+
| +-- Lortiella_froggatti
|
| +-- Parreysia_nagpoorensis **
| +-----------+
| | +-- Parreysia_plagiosoma **
| |
| +--------+ +-- Trapezidens_dolichorhynchus **
| | | +--------+
| | | | +-- Trapezidens_exolescens **
| | | |
| | +--+ +-------- Lamellidens_savadiensis **
--+ | | |
| | | +-------- Lamellidens_corrianus **
| | +--+
| | | +-- Lamellidens_marginalis **
| | | +--+
| | | | +-- Lamellidens_ferrugineus **
| +-----+ +--+
| | | | +-- Lamellidens_jenkinsianus **
| | | +--+
| | | +-- Lamellidens_mainwaringi **
| | |
| | | +-- Leoparreysia_olivacea **
| | | +-----------------+
| | | | +-- Leoparreysia_tavoyensis **
| | | |
| | | | +-------------- Nitia_teretiuscula **
| | | | +--+
| | | | | | +----------- Coelatura_aegyptiaca[1] **
| | +--+ | +--+
| | | | | +-------- Coelatura_aegyptiaca[2] **
| | | | +--+
| | | | | +----- Coelatura_gabonensis **
| | | | +--+
| | | | | +-- Coelatura_kunenensis **
| | +--+ +--+
| | | +-- Prisodontopsis_mweruensis **
| | |
| | | +-------- Indonaia_shurtleffiana **
| | | +-----+
| | | | | +----- Indonaia_andersoniana **
| | | | +--+
| | | | | +-- Indonaia_rugosa **
| | | | +--+
| | +--+ +-- Indonaia_bonneaudii **
| | |
| | | +-- Indochinella_pugio **
| | | +--------+
| | | | +-- Scabies_crispata **
| | | |
| | +--+ +-- Radiatula_myitkyinae **
| | | +-----+
| | | | +-- Radiatula_chaudhurii **
| | +--+
| | | +----- Scabies_mandarinus **
| | +--+
| | | +-- Scabies_humilis **
| | +--+
| | +-- Scabies_phaselus **
| |
| | +----- Unio_tumidus **
| | +-----+
| | | | +-- Unio_crassus **
| | | +--+
| | | +-- Unio_pictorum **
| | +-----+
+--+ | | +-------- Aculamprotula_tortuosa **
| | | |
| | | +-------- Schistodesmus_lampreyanus **
| | | |
| | +--+ +-- Nodularia_douglasiae **
| | +-----+
| | | +-- Nodularia_nuxpersicae **
| | |
| | | +----- Cuneopsis_heudei **
| | +--+
| | | +-- Tchangsinaia_piscicula **
| | +--+
| +--------+ +-- Cuneopsis_rufescens **
| | |
| | | +-- Lanceolaria_grayii **
| | | +-----------+
| | | | +-- Lanceolaria_gladiola **
| | | |
| | | | +-- Cristaria_plicata **
| | | | +--------+
| | | | | +-- Cristaria_radiata **
| | | +--+
| | | | | +-------- Pletholophus_tenuis **
| | | | +--+
| | | | | +----- Sinanodonta_lucida **
| | +--+ +--+
| | | | +-- Sinanodonta_lauta **
| | | +--+
| | | +-- Sinanodonta_jourdyi **
| | |
| | | +-------- Anodonta_anatina **
| | | |
| | | +-------- Pseudanodonta_complanata **
| | | |
| | | | +-- Anodonta_cygnea **
| | +-----+-----+
| | | +-- Anodonta_nuttalliana **
| | |
| | | +----- Pyganodon_grandis **
| | | |
| | | +----- Simpsonaias_ambigua **
| | +--+
| | +----- Strophitus_undulatus **
| | |
| | | +-- Alasmidonta_marginata **
| | +--+
| | +-- Lasmigona_costata **
| |
| | +-------- Lens_contradens **
| | +-----+
+--+ | | +----- Lens_eximius **
| | +--+
| | | +-- Lens_comptus **
| | +--+
| +--+ +-- Lens_novoselovi **
| | |
| | | +----- Physunio **
| | | +-----+
| | | | | +-- Physunio_modelli **
| | | | +--+
| | +--+ +-- Physunio_superbus **
| | |
| | | +-------- Yaukthwa_inlenensis **
| | +--+
| | | +----- Yaukthwa_panhai **
| +-----+ +--+
| | | | +-- Yaukthwa_rectangularis **
| | | +--+
| | | +-- Yaukthwa_nesemanni **
| | |
| | | +----- Hyriopsis_panhai **
| | | |
| | | +--------+----- Hyriopsis_khoratensis **
| | | | |
| | | | | +-- Hyriopsis_myersiana **
| | | | +--+
| | +--+ +-- Hyriopsis_bialata **
| | |
| | | +----------- Hyriopsis_desowitzi **
| | | |
| | | +----------- Rectidens_sumatrensis **
| | +--+
| | | +-- Ensidens_telus **
| | | +-----+
| | | | +-- Ensidens_ingallsianus **
| | +--+
| | | +----- Ensidens_sagittarius **
| | +--+
| | | +-- Ensidens_spiculus **
+--+ +--+
| +-- Ensidens_jaculus **
|
| +-------- Amblema_plicata **
| +--+
| | | +----- Cambarunio_iris **
| | +--+
| | | +-- Ortmanniana_ligamentina **
| | +--+
| +--------+ +-- Lampsilis_cardium **
| | |
| | | +-- Quadrula_quadrula **
| | | +-----+
| | | | +-- Quadrula_verrucosa **
| | +--+
| | | +----- Pleurobema_sintoxia **
| | +--+
| | | +-- Elliptio_complanata **
| | +--+
| | +-- Eurynia_dilatata **
| |
| | +-------- Sinohyriopsis_cumingii **
| | |
| | +-----+ +-- Lamprotula_caveata **
+--+ | | +--+
| | | | +-- Lamprotula_leaii **
| | +--+
| | | +-- Potomida_littoralis **
| +--+ +--+
| | | +-- Pronodularia_japanensis **
| | |
| | | +----------- Sinosolenaia_carinata **
| | | |
| | | | +-- Sinosolenaia_oleivora **
| | +--+--------+
| | | +-- Sinosolenaia_recognita **
| | |
| | | +-------- Gonidea_angulata **
| | +--+
| | | +----- Microcondylaea_bonellii **
+--+ +--+
| | +-- Leguminaia_wheatleyi **
| +--+
| +-- Leguminaia **
|
| +-------- Pilsbryoconcha_exilis[3] **
| +-----+
| | | +----- Pilsbryoconcha_exilis[2] **
| | +--+
| | | +-- Pilsbryoconcha_exilis[1] **
| | +--+
| | +-- Pilsbryoconcha_carinifera **
| |
+--+ +----------- Indopseudodon_indawgyiensis **
| |
| | +-------- Thaiconcha_callifera **
| +--+
| | | +----- Bineurus_exilis **
| | +--+
| | | +-- Bineurus_mouhotii **
+--+ +--+
| +-- Bineurus_loeiensis **
|
| +----- Pseudodon_vagulus **
| +--+
| | | +-- Pseudodon_mekongi **
| | +--+
| | +-- Pseudodon_vondembuschianus **
+--+
| +-- Pseudodon_inoscularis **
| +--+
| | +-- Sundadontina_tumida **
+--+
| +-- Nyeinchanconcha_nyeinchani **
+--+
+-- Sundadontina_cumingii **
Bolotov et al., 2020 [Publ 4100], fig. 1
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 120 |
120 |
100 |
2 |
0 |
0 |
0 |
154.0 |
+-- Gibbosula_laosensis **
|
+-- Margaritifera_dahurica **
|
+-----------------------------------------+-- Margaritifera_margaritifera **
| |
| +-- Margaritifera_laevis **
| |
| +-- Margaritifera_middendorffi **
|
| +-------- Trapezidens_exolescens **
| +--+
| | | +----- Trapezidens_scutum **
| | +--+
| | | +-- Trapezidens_dolichorhynchus **
| +--------------------------+ +--+
| | | +-- Trapezidens_angustior **
| | |
| | | +-------- Lamellidens_savadiensis **
| | +--+
| | | +----- Lamellidens_generosus **
| | +--+
| | | +-- Lamellidens_ferrugineus **
| | +--+
| | +-- Lamellidens_mainwaringi **
| |
| | +----------- Parreysia_keralaensis **
| | +--+
| +--+ | | +-------- Parreysia_plagiosoma **
| | | | +--+
| | | | | +----- Parreysia_nagpoorensis[2] **
| | | | +--+
| | | | | +-- Parreysia_nagpoorensis[1] **
| | | +--------------------+ +--+
| | | | | +-- Parreysia_rakhinensis **
| | | | |
| | | | | +-- Leoparreysia_burmana **
| | | | | +-----+
| | | | | | +-- Leoparreysia_whitteni **
| | | | +-----+
| | | | | +----- Leoparreysia_tavoyensis **
| | | | +--+
| | +--+ | +-- Leoparreysia_olivacea **
| | | +--+
| | | +-- Leoparreysia_canefrii **
| | |
| | | +-- Coelatura_aegyptiaca **
--+ | | +-----------------------------+
| | | | +-- Nitia_teretiuscula **
| | | |
| | | | +----------- Indonaia_shurtleffiana **
| | | | +-----------------+
| | +--+ | | +-------- Indonaia_rugosa **
| | | | +--+
| | | | | +----- Indonaia_andersoniana **
| | | | +--+
| | | | | +-- Indonaia_subclathrata **
| | +--+ +--+
| | | +-- Indonaia_caerulea **
| | |
| | | +-------------------------- Indochinella_pugio **
| | | |
| | | | +-------- Radiatula_crispisulcata **
| | | | +--------------+
| | +--+ | | +----- Radiatula_mouhoti **
| | | | +--+
| | | | | +-- Radiatula_chaudhurii **
| | | | +--+
| | +--+ +-- Radiatula_myitkyinae **
| | |
| | | +-------------------- Scabiellus_songkramensis **
| | | |
| | | | +-- Unionetta_fabagina **
| | +--+ +--------------+
| | | | +-- Harmandia_somboriensis **
| | | |
| | +--+ +-------------- Scabies_nucleus **
| | | |
| | | | +-- Scabies_crispata **
| | +--+ +--------+
| | | | +-- Scabies_mandarinus **
| | | |
| | +--+ +-- Scabies_humilis **
| | | +-----+
| | | | +-- Scabies_pilata **
| | +--+
| | | +----- Scabies_anceps **
| | +--+
| | | +-- Scabies_phaselus **
| | +--+
+--+ +-- Scabies_scobinatus **
|
| +----------- Hyriopsis_desowitzi **
| |
| +--+ +-------- Rectidens_sumatrensis **
| | | |
| | +--+ +-- Hyriopsis_khoratensis **
| | | +--+
| | | | +-- Hyriopsis_bialata **
| | +--+
| | | +-- Hyriopsis_myersiana **
| +--------+ +--+
| | | +-- Hyriopsis_panhai **
| | |
| | | +-- Ensidens_telus **
| | | +-----+
| | | | +-- Ensidens_ingallsianus **
| | | |
| | +-----+ +-- Ensidens_jaculus **
| | | +--+
| | | | +-- Ensidens_spiculus **
| | +--+
| | | +-- Ensidens_dugasti **
| | +--+
| | +-- Ensidens_sagittarius **
| +-----+
| | | +-- Trapezoideus_lenya **
| | | +-----+
| | | | +-- Trapezoideus_foliaceus **
| | | +-----------+
| | | | | +----- Physunio_superbus **
| | | | +--+
| | | | | +-- Physunio_modelli **
| | | | +--+
| | | | +-- Physunio **
| | | |
| | | | +-- Yaukthwa_elongatula **
| | | | +-----------+
| | | | | +-- Yaukthwa_baniensis **
| | +--+ |
| | | +--+ +----- Yaukthwa_inlenensis **
| | | | | +-----+
| | | | | | | +-- Yaukthwa_avaensis **
| | | | | | +--+
| | | | +--+ +-- Yaukthwa_paiensis **
| | | | |
| | | | | +-------- Yaukthwa_panhai **
| | | | +--+
| | | | | +----- Yaukthwa_rectangularis **
| | +--+ +--+
| | | | +-- Yaukthwa_zayleymanensis **
| | | +--+
| | | +-- Yaukthwa_nesemanni **
| | |
| | | +----- Lens_misellus **
| | | +-----+
| | | | | +-- Lens_contradens **
| | | | +--+
| | | | +-- Lens_micropterus **
| | +-----+
| | | +----- Lens_pallegoixi **
| | | +--+
+-----------+ | | | +-- Lens_comptus **
| | | +--+
| +--+ +-- Lens_novoselovi **
| |
| | +----- Lens_maenamensis **
| +--+
| | +-- Lens_eximius **
| +--+
| +-- Lens_rolfbrandti **
|
| +-- Sinosolenaia_iridinea **
| |
| +-- Sinosolenaia_recognita[1] **
| |
| +-- Sinosolenaia_carinata **
| |
| +-- Leguminaia_wheatleyi **
| |
| +-- Microcondylaea_bonellii **
| |
| +-- Gonidea_angulata **
| |
| +-- Ptychorhynchus_pfisteri **
| |
| +-- Parvasolenaia_rivularis **
| |
| +-----------------------+-- Lamprotula_leaii **
| | |
| | +-- Lamprotula **
| | |
| | +-- Lamprotula_caveata **
| | |
| | +-- Potomida_littoralis **
| | |
| | +-- Pronodularia_japanensis **
| | |
| | +-- Sinohyriopsis_cumingii **
| | |
+--+ +-- Chamberlainia_somsakpanhai **
| |
| +-- Sinosolenaia_recognita[2] **
| |
| +-- Sinosolenaia_oleivora **
|
| +-------- Indopseudodon_kayinensis **
| +--------------+
| | | +----- Indopseudodon_indawgyiensis **
| | +--+
| | | +-- Indopseudodon_salwenianus **
| | +--+
| | +-- Indopseudodon_bogani **
+--+
| +----- Bineurus_loeiensis **
| +--------------+
| | | +-- Bineurus_exilis **
| | +--+
| | +-- Bineurus_mouhotii **
| |
+--+ +----------------- Nyeinchanconcha_nyeinchani **
| |
| | +-- Thaiconcha_callifera **
| | +-----+
| | | +-- Thaiconcha_munelliptica **
+--+ +-----+
| | | +----- Pilsbryoconcha_exilis[2] **
| | +--+
| | | +-- Pilsbryoconcha_exilis[1] **
| | +--+
| | +-- Pilsbryoconcha **
+--+
| +-------- Pseudodon_lenyanensis **
| +--+
| | | +----- Pseudodon_vagulus **
| | +--+
| | | +-- Pseudodon_vondembuschianus **
| | +--+
+--+ +-- Pseudodon_mekongi **
|
| +-- Sundadontina_taskaevi **
| +-----+
| | +-- Sundadontina_brandti **
+--+
| +----- Sundadontina_tumida **
+--+
| +-- Sundadontina_cumingii **
+--+
+-- Sundadontina_tanintharyiensis **
Campbell et al., 2005 [Publ 2660], fig. 1
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 117 |
117 |
104 |
2 |
0 |
0 |
0 |
158.0 |
+-- Margaritifera_margaritifera **
+--------------------------------------------------+
| +-- Cumberlandia_monodonta **
|
| +-------------------------------------------------- Sinohyriopsis_cumingii **
| |
| | +----------- Anodonta_oregonensis **
| | |
| +--------------------------------------+ +-- Anodonta_cygnea **
| | | +-----+
| | | | +-- Alasmidonta_holstonia **
--+ | +--+
| | | +----- Pyganodon_grandis **
| | +--+
| | | +-- Lasmigona_costata **
| | +--+
| | +-- Pseudodontoideus_subvexus **
| |
| | +----- Gonidea_angulata **
| | +-----------------------------------------+
| | | | +-- Potomida_littoralis **
+--+ | +--+
| | +-- Pronodularia_japanensis **
| |
| | +-- Megalonaias_nervosa **
| | +-----------+
| | | +-- Uniomerus_declivis **
| | |
| | +-----------------------------+ +----------- Theliderma_metanevra **
| | | | |
| | | | | +----- Quadrula_verrucosa **
| | | +--+ +--+
| | | | | | +-- Quadrula_quadrula **
| | | | | +--+
| | | | | +-- Quadrula_nobilis **
| | | +--+
| | | | +----- Pustulosa_kieneriana **
| | | | |
| | | | | +-- Cyclonaias_tuberculata **
+--+ | +--+--+
| | | +-- Pustulosa_succissa **
| | |
| | | +-- Pustulosa_infucata **
| | +--+
| | +-- Pustulosa_kleiniana **
| |
| | +----------------------------------- Elliptoideus_sloatianus **
| | |
| | | +-------------------------------- Hemistena_lata **
| | | |
| | +-----+ | +----- Plethobasus_cyphyus **
| | | | | +-----------------------+
| | | | | | | +-- Elliptio_arca **
| | | | | | +--+
| | | | | | +-- Elliptio_crassidens **
| | | +--+ |
| | | | | +-- Eurynia_dilatata **
| | | | | +-----------------------+
| | | | | | +-- Parvaspina_collina **
+--+ | | | |
| | | | | +----- Pleuronaia_gibber **
| | | | +--------------------+
| | | | | | +-- Pleuronaia_barnesiana **
| | +--+ | +--+
| | | | +-- Pleuronaia_dolabelloides **
| | | |
| | | | +----------- Fusconaia_subrotunda **
| | | | |
| | | | | +-- Fusconaia_cor **
| | | +--------------+ +-----+
| | | | | | +-- Fusconaia_cuneolus **
| | | | | |
| | | | | | +-- Fusconaia_cerina[1] **
| | | | +--+ |
| | | | | +--+-- Fusconaia_cerina[2] **
| | +--+ | | |
| | | | | +-- Fusconaia_flava **
| | | +--+
| | | | +-- Fusconaia_burkei[1] **
| | | | |
| | | | +-- Fusconaia_burkei[2] **
| | | +--+
| | | +-- Fusconaia_burkei[3] **
| | | |
| | | +-- Fusconaia_escambia **
| | |
| | | +-- Pleurobema_rubrum[2] **
| | | +--+
| | | | +-- Pleurobema_cordatum **
| | | +-----------------+
| | | | | +-- Pleurobema_rubrum[1] **
+--+ | | +--+
| | | +-- Pleurobema_sintoxia **
| +--+
| | +-------------------- Pleurobema_pyriforme **
| | |
| | | +-- Pleurobema_georgianum[2] **
| | | |
| +--+ +--------------+-- Pleurobema_georgianum[3] **
| | | |
| | | +-- Pleurobema_athearni **
| | |
| | | +----- Pleurobema_decisum **
| +--+ |
| | +--------+ +-- Pleurobema_hanleyianum[1] **
| | | | |
| | | +--+-- Pleurobema_hanleyianum[2] **
| | | |
| | | +-- Pleurobema_georgianum[1] **
| +--+
| | +-- Pleurobema_oviforme[1] **
| | |
| | +--------+-- Pleurobema_oviforme[2] **
| | | |
| +--+ +-- Pleurobema_clava **
| |
| | +-------- Pleurobema_strodeanum **
| +--+
| | +----- Pleurobema_rubellum **
| +--+
| | +-- Pleurobema_perovatum **
| +--+
| +-- Pleurobema_taitianum **
|
| +----- Plectomerus_dombeyanus **
| +--------------------------------+
| | | +-- Reginaia_ebenus **
| | +--+
| | +-- Reginaia_rotulata **
| |
| | +----- Popenaias_popeii **
| | +-----------------------------+
| | | | +-- Amblema_elliottii **
+--+ | +--+
| | +-- Amblema_plicata **
| |
| | +-- Toxolasma_parvum **
| | +-----+
| | | +-- Toxolasma_texasiense **
| | +-----------------------+
| | | | +----- Obliquaria_reflexa **
+--+ | +--+
| | | +-- Cyrtonaias_tampicoensis **
| | +--+
| | +-- Glebula_rotundata **
| |
| | +-- Potamilus_fragilis **
| | +--+
| | | +-- Potamilus_leptodon **
| | +-----------+
| | | | +-- Potamilus_alatus **
| | | +--+
+--+ | +-- Potamilus_purpuratus **
| |
| +-----------+ +-- Sagittunio_nasutus **
| | | +--+
| | | | +-- Medionidus_conradicus **
| | | +--------+
| | | | | +-- Ptychobranchus_fasciolaris **
| | | | +--+
| | +--+ +-- Venustaconcha_ellipsiformis[2] **
| | |
| | | +----------- Medionidus_acutissimus **
| | +--+
| | | +-------- Lemiox_rimosus **
| | +--+
+--+ | +----- Dromus_dromas **
| +--+
| | +-- Cyprogenia_aberti **
| +--+
| +-- Cyprogenia_stegaria **
|
| +-- Ellipsaria_lineolata **
| +-----------------------+
| | +-- Truncilla_truncata **
| |
| | +----------------------- Villosa_villosa **
| | |
| | | +-- Obovaria_olivaria **
+--+ | +--------------+
| | | +-- Venustaconcha_ellipsiformis[1] **
| | |
| | +--+ +----- Obovaria_unicolor **
| | | | +--------+
| | | | | | +-- Obovaria_arkansasensis **
+--+ | | | +--+
| | +--+ +-- Obovaria_subrotunda **
| | |
| | | +----------- Epioblasma_brevidens **
| | +--+
| | | +-------- Epioblasma_triquetra **
| | +--+
| | | +----- Epioblasma_walkeri[2] **
| | +--+
| | | +-- Epioblasma_capsaeformis **
+--+ +--+
| +-- Epioblasma_walkeri[1] **
|
| +-- Ligumia_recta **
| +--------+
| | +-- Cambarunio_iris **
| +--+
| | | +-------- Lampsilis_teres **
| | +--+
| | | +----- Ortmanniana_ligamentina **
| | +--+
| | | +-- Lampsilis_siliquoidea **
| | +--+
| | +-- Leaunio_vanuxemensis **
+-----+
| +-- Lampsilis_cardium **
| +--+
| | +-- Lampsilis_ovata **
| +-----+
| | | +-- Ortmanniana_pectorosa **
| | +--+
| | +-- Lampsilis_ornata **
| |
+--+ +----- Hamiota_australis[2] **
| +--+
| | | +-- Hamiota_australis[1] **
| | +--+
| | +-- Hamiota_subangulata **
+--+
| +----- Hamiota_altilis[2] **
| |
| | +-- Hamiota_altilis[1] **
+--+--+
| +-- Hamiota_perovalis[1] **
|
| +-- Hamiota_altilis[3] **
+--+
+-- Hamiota_perovalis[2] **
Campbell et al., 2005 [Publ 2660], fig. 2
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 115 |
115 |
93 |
2 |
0 |
0 |
0 |
159.0 |
+-- Cumberlandia_monodonta **
+--------------------------------------------+
| +-- Margaritifera_margaritifera **
|
| +----------- Sinohyriopsis_cumingii **
| |
| +--------------------------------+ +-------- Pyganodon_grandis **
| | | |
| | | | +-- Lasmigona_costata **
| | +--+-----+
| | | +-- Pseudodontoideus_subvexus **
--+ | |
| | | +----- Alasmidonta_holstonia **
| | +--+
| | | +-- Anodonta_cygnea **
| | +--+
| | +-- Anodonta_oregonensis **
| |
| | +----- Gonidea_angulata **
| | +-----------------------------------+
| | | | +-- Potomida_littoralis **
| | | +--+
+--+ | +-- Pronodularia_japanensis **
| |
| | +-------------------- Megalonaias_nervosa **
| | |
| | +-----------------+ +----------------- Uniomerus_declivis **
| | | | |
| | | +--+ +-------------- Theliderma_metanevra **
| | | | |
| | | +--+ +----------- Quadrula_quadrula **
| | | | |
| | | | | +-- Quadrula_nobilis **
| | | +--+ +-----+
| | | | | +-- Quadrula_verrucosa **
| | | | |
| | | +--+ +----- Pustulosa_kieneriana **
| | | | |
+--+ | | | +-- Cyclonaias_tuberculata **
| | +--+--+
| | | +-- Pustulosa_succissa **
| | |
| | | +-- Pustulosa_infucata **
| | +--+
| | +-- Pustulosa_kleiniana **
| |
| | +-------------------------------- Elliptoideus_sloatianus **
| | |
| | | +----------------------------- Plethobasus_cyphyus **
| | | |
| | | +----------------------------- Parvaspina_collina **
| | +--+ |
| | | | | +-- Elliptio_arca **
| | | | +--------------------------+
| | | | | +-- Elliptio_crassidens **
| | | | |
| | | | | +-------------------------- Eurynia_dilatata **
| | | +--+ |
+--+ | | +-------------------------- Hemistena_lata **
| | | |
| | | | +-- Pleurobema_rubrum[1] **
| | | | +--+
| | | | | +-- Pleurobema_sintoxia **
| | | | +-----------------+
| | | | | | +-- Pleurobema_rubrum[2] **
| | | | | +--+
| | +--+ | +-- Pleurobema_cordatum **
| | | |
| | | | +-------------- Fusconaia_subrotunda **
| | | | |
| | | | | +-- Fusconaia_cor **
| | | +--------+ +--------+
| | | | | | +-- Fusconaia_cuneolus **
| | | | | |
| | | | +--+ +-- Fusconaia_cerina **
| | | | | +-----+
| | | | | | +-- Fusconaia_flava **
| | +--+ +--+
| | | | +----- Fusconaia_burkei[2] **
| | | +--+
| | | | +-- Fusconaia_escambia **
| | | +--+
| | | +-- Fusconaia_burkei[1] **
| | |
| | | +----- Pleuronaia_gibber **
| | | +--------------+
| | | | | +-- Pleuronaia_barnesiana **
| | | | +--+
| | | | +-- Pleuronaia_dolabelloides **
| | | |
| | | | +-------- Pleurobema_rubellum **
+--+ +--+ +--------+
| | | | +----- Pleurobema_strodeanum **
| | | +--+
| | | | +-- Pleurobema_perovatum **
| | | +--+
| | | +-- Pleurobema_taitianum **
| +--+
| | +-- Pleurobema_oviforme[1] **
| | |
| | +-----------+-- Pleurobema_oviforme[2] **
| | | |
| | | +-- Pleurobema_clava **
| +--+
| | +----------- Pleurobema_pyriforme **
| | |
| +--+ +-- Pleurobema_athearni **
| | +-----+
| | | +-- Pleurobema_georgianum[2] **
| +--+
| | +----- Pleurobema_decisum **
| | |
| +--+ +-- Pleurobema_hanleyianum[1] **
| | |
| +--+-- Pleurobema_hanleyianum[2] **
| |
| +-- Pleurobema_georgianum[1] **
|
| +-------- Popenaias_popeii **
| |
| +--------------+ +-- Amblema_elliottii **
| | | +--+
| | | | +-- Amblema_plicata **
| | +--+
| | | +-- Reginaia_ebenus **
| | | |
| | +--+-- Reginaia_rotulata[1] **
| | |
| | +-- Reginaia_rotulata[2] **
| |
| | +----- Plectomerus_dombeyanus **
| | +--------------+
+-----------+ | | +-- Toxolasma_parvum **
| | +--+
| | +-- Toxolasma_texasiense **
| |
| | +----- Obliquaria_reflexa **
| | +-----------+
| | | | +-- Cyrtonaias_tampicoensis **
| | | +--+
| | | +-- Glebula_rotundata **
| | |
| | | +-------------- Sagittunio_nasutus **
| | | |
+--+ | +-------------- Leaunio_vanuxemensis **
| | |
| | | +-- Ellipsaria_lineolata **
| | +-----------+
| | | +-- Truncilla_truncata **
| | |
| | | +----- Lampsilis_teres **
| | +--------+
| | | | +-- Ortmanniana_ligamentina **
| | | +--+
| | | +-- Lampsilis_siliquoidea **
| | |
| | | +----- Villosa_villosa **
| | +--------+
+--+ | | +-- Ligumia_recta **
| | +--+
| | +-- Cambarunio_iris **
| |
| | +-------- Ortmanniana_pectorosa **
| +-----+
| | | +----- Lampsilis_ornata **
| | +--+
| | | +-- Lampsilis_cardium **
| | +--+
| | +-- Lampsilis_ovata **
| |
| | +-- Potamilus_fragilis **
| | +--+
| | | +-- Potamilus_leptodon **
| +--------+
| | | +-- Potamilus_alatus **
| | +--+
| | +-- Potamilus_purpuratus **
| |
| | +-- Ptychobranchus_fasciolaris **
+--+ +--------+
| | +-- Venustaconcha_ellipsiformis[1] **
| |
+--+ +-------- Medionidus_conradicus **
| | |
| | | +-- Lemiox_rimosus **
| +--+-----+
| | +-- Medionidus_acutissimus **
| |
| | +----- Dromus_dromas **
| +--+
| | +-- Cyprogenia_aberti **
| +--+
| +-- Cyprogenia_stegaria **
|
| +----- Hamiota_australis[2] **
| +-----+
| | | +-- Hamiota_australis[1] **
| | +--+
| | +-- Hamiota_subangulata **
+--+
| | +-- Hamiota_altilis[3] **
| | +-----+
| | | +-- Hamiota_perovalis[2] **
| +--+
| | +----- Hamiota_altilis[2] **
| +--+
| | +-- Hamiota_altilis[1] **
| +--+
| +-- Hamiota_perovalis[1] **
|
| +----------- Obovaria_olivaria **
| |
| +----------- Venustaconcha_ellipsiformis[2] **
| |
| | +----- Obovaria_unicolor **
| +-----+
+--+ | +-- Obovaria_arkansasensis **
| +--+
| +-- Obovaria_subrotunda **
|
| +-- Epioblasma_brevidens **
| +-----+
| | +-- Epioblasma_triquetra **
+--+
| +----- Epioblasma_walkeri[1] **
+--+
| +-- Epioblasma_walkeri[2] **
+--+
+-- Epioblasma_capsaeformis **
Bolotov et al., 2020 [Publ 4100], fig. S1
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 114 |
114 |
100 |
1 |
0 |
0 |
0 |
148.0 |
+-------- Trapezidens_exolescens **
+--+
| | +----- Trapezidens_scutum **
| +--+
| | +-- Trapezidens_dolichorhynchus **
+--------------------------------+ +--+
| | +-- Trapezidens_angustior **
| |
| | +-------- Lamellidens_savadiensis **
| +--+
| | +----- Lamellidens_generosus **
| +--+
| | +-- Lamellidens_ferrugineus **
| +--+
| +-- Lamellidens_mainwaringi **
|
| +----------- Parreysia_keralaensis **
| +--+
+--+ | | +-------- Parreysia_nagpoorensis[2] **
| | | +--+
| | | | +----- Parreysia_nagpoorensis[1] **
| | | +--+
| | | | +-- Parreysia_rakhinensis **
| | +--------------------------+ +--+
| | | | +-- Parreysia_plagiosoma **
| | | |
| | | | +-- Leoparreysia_burmana **
| | | | +-----+
| | | | | +-- Leoparreysia_whitteni **
| | | +-----+
| | | | +----- Leoparreysia_olivacea **
| | | +--+
| +--+ | +-- Leoparreysia_tavoyensis **
| | +--+
| | +-- Leoparreysia_canefrii **
| |
| | +-- Coelatura_aegyptiaca **
| | +-----------------------------------+
| | | +-- Nitia_teretiuscula **
| | |
| | | +-------- Indonaia_shurtleffiana **
| +--+ +--------------------------+
| | | | +----- Indonaia_andersoniana **
| | | +--+
| | | | +-- Indonaia_caerulea **
| | | +--+
| +--+ +-- Indonaia_subclathrata **
| |
| | +-------------------------------- Indochinella_pugio **
| | |
| | | +----- Radiatula_mouhoti **
| +--+ +-----------------------+
| | | | +-- Radiatula_myitkyinae **
| | | +--+
| +--+ +-- Radiatula_chaudhurii **
| |
| | +-------------------------- Radiatula_crispisulcata **
| | |
| +--+ +----------------------- Scabiellus_songkramensis **
| | |
| +--+ +-------------------- Harmandia_somboriensis **
| | |
| +--+ +----------------- Unionetta_fabagina **
| | |
| +--+ +-------------- Scabies_mandarinus **
| | |
| +--+ +----------- Scabies_crispata **
| | |
| +--+ +-------- Scabies_nucleus **
| | |
| +--+ +-- Scabies_humilis **
| | +--+
| | | +-- Scabies_pilata **
| +--+
| | +-- Scabies_phaselus **
| | |
| +--+-- Scabies_scobinatus **
| |
--+ +-- Scabies_anceps **
|
| +-------------- Hyriopsis_desowitzi **
| +--+
| | | +----------- Rectidens_sumatrensis **
| | +--+
| | | +-------- Hyriopsis_panhai **
| | +--+
| | | +----- Hyriopsis_myersiana **
| | +--+
| +--------+ | +-- Hyriopsis_khoratensis **
| | | +--+
| | | +-- Hyriopsis_bialata **
| | |
| | | +-- Ensidens_telus **
| | | +-----+
| | | | +-- Ensidens_ingallsianus **
| | | |
| | +--------+ +-- Ensidens_dugasti **
| | | +--+
| | | | +-- Ensidens_sagittarius **
| | +--+
| +--+ | +-- Ensidens_spiculus **
| | | +--+
| | | +-- Ensidens_jaculus **
| | |
| | | +----- Physunio **
| | | +-----------------+
| | | | | +-- Physunio_modelli **
| | | | +--+
| | | | +-- Physunio_superbus **
| | | |
| | | | +-- Trapezoideus_lenya **
| | | | +-----------------+
| | +--+ | +-- Trapezoideus_foliaceus **
| | | |
| | | | +----- Lens_misellus **
| | | | +-----+
| | | | | | +-- Lens_contradens **
| | | | | +--+
| | | | | +-- Lens_micropterus **
| | +--+ +-----+
| | | | | +----- Lens_maenamensis **
| | | | | +--+
| | | | | | | +-- Lens_eximius **
| | | | | | +--+
| | | | +--+ +-- Lens_rolfbrandti **
| | | | |
| | | | | +----- Lens_pallegoixi **
| | +--+ +--+
| | | | +-- Lens_novoselovi **
| | | +--+
| | | +-- Lens_comptus **
| | |
| | | +-- Yaukthwa_elongatula **
| | | +-----------+
| | | | +-- Yaukthwa_baniensis **
| | | |
| | +--+ +----- Yaukthwa_inlenensis **
| | | +-----+
+-----------------+ | | | +-- Yaukthwa_avaensis **
| | | +--+
| +--+ +-- Yaukthwa_paiensis **
| |
| | +-------- Yaukthwa_panhai **
| +--+
| | +----- Yaukthwa_rectangularis **
| +--+
| | +-- Yaukthwa_zayleymanensis **
| +--+
| +-- Yaukthwa_nesemanni **
|
| +-- Sinohyriopsis_cumingii **
| +--+
| | +-- Chamberlainia_somsakpanhai **
| |
| +--+ +-- Lamprotula_leaii **
| | | |
| | | +-- Lamprotula **
| | | |
| | +--+-- Lamprotula_caveata **
| | |
| | +-- Potomida_littoralis **
| | |
| | +-- Pronodularia_japanensis **
| +-----------------+
| | | +-- Sinosolenaia_iridinea **
| | | |
| | | +-- Sinosolenaia_recognita[1] **
| | | |
| | | +-- Sinosolenaia_carinata **
| | | |
| | | +-- Leguminaia_wheatleyi **
| | | |
| | | +-- Microcondylaea_bonellii **
| | +-----+
| | +-- Gonidea_angulata **
| | |
| | +-- Ptychorhynchus_pfisteri **
+--+ |
| +-- Parvasolenaia_rivularis **
| |
| +-- Sinosolenaia_recognita[2] **
| |
| +-- Sinosolenaia_oleivora **
|
| +-------- Indopseudodon_kayinensis **
| +--------------+
| | | +----- Indopseudodon_indawgyiensis **
| | +--+
| | | +-- Indopseudodon_salwenianus **
| | +--+
| | +-- Indopseudodon_bogani **
+--+
| +----- Bineurus_loeiensis **
| +--------------+
| | | +-- Bineurus_exilis **
| | +--+
| | +-- Bineurus_mouhotii **
| |
+--+ +----------------- Nyeinchanconcha_nyeinchani **
| |
| | +-- Thaiconcha_callifera **
| | +-----+
| | | +-- Thaiconcha_munelliptica **
+--+ +-----+
| | | +----- Pilsbryoconcha_exilis[2] **
| | +--+
| | | +-- Pilsbryoconcha_exilis[1] **
| | +--+
| | +-- Pilsbryoconcha **
+--+
| +-------- Pseudodon_lenyanensis **
| +--+
| | | +----- Pseudodon_vagulus **
| | +--+
| | | +-- Pseudodon_vondembuschianus **
| | +--+
+--+ +-- Pseudodon_mekongi **
|
| +-- Sundadontina_tanintharyiensis **
| +-----+
| | +-- Sundadontina_cumingii **
+--+
| +----- Sundadontina_tumida **
+--+
| +-- Sundadontina_taskaevi **
+--+
+-- Sundadontina_brandti **
Lopes-Lima et al., 2020 [Publ 4080], fig. 2-4
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 113 |
118 |
98 |
2 |
9 |
7 |
2 |
99.0 |
+-- Hyridella_depressa
+--------------------------------------------+
| +-- Velesunio_ambiguus
|
| +-- Margaritifera_laevis **
| +-----------------------------------------+
| | +-- Margaritifera_middendorffi **
| |
| | +-- Lamproscapha_ensiformis
--+ | +-----+
| | | +-- Mycetopoda_siliquosa
| | +--------------------------------+
| | | | +----- Etheria_elliptica
| | | +--+
| | | | +-- Aspatharia_pfeifferiana
| | | +--+
| | | +-- Chambardia_wahlbergi
| | |
+--+ | +-- Coelatura_aegyptiaca **
| | |
| | +-----------------------------------+-- Leoparreysia_tavoyensis **
| | | |
| | | +-- Radiatula_mouhoti **
| | |
| | | +-- Lens_contradens **
| | | |
| | | +-- Ensidens_sagittarius **
| | | |
| | | +-----------------------+-- Physunio_modelli **
| | | | |
| | | | +-- Rectidens_sumatrensis **
| | | | |
+--+ | | +-- Yaukthwa_nesemanni **
| | |
| | | +-- Amblema_plicata **
| | | |
| | | +-- Elliptio_complanata **
| | | +--------------------+
| | +--------+ | +-- Lampsilis_cardium **
| | | | | |
| | | | | +-- Quadrula_quadrula **
| | | | |
| | | | | +----- Chamberlainia_somsakpanhai **
| | | | | +-----------+
| | | | | | | +-- Sinohyriopsis_cumingii **
| | | | | | +--+
| | | | | +--+ +-- Sinohyriopsis_schlegelii **
| | | | | | |
| | | | | | | +-------------- Pronodularia_japanensis **
| | | +--+ | | |
| | | | | +--+ +----------- Potomida_littoralis **
| | | | | | |
| | | | | +--+ +-------- Lamprotula_caveata **
| | | | | | |
+--+ | | | +--+ +----- Lamprotula_leaii[2] **
| | | | | |
| | | | +--+----- Lamprotula_leaii[3] **
| | | | |
| | | | | +-- Lamprotula_seomjinensis **
| | | | +--+
| | | | +-- Lamprotula_leaii[1] **
| | +--+
| | | +-- Indopseudodon_indawgyiensis **
| | | +--------+
| | | | +-- Indopseudodon_salwenianus **
| | | |
| | | +-----+ +-- Pilsbryoconcha_exilis **
| | | | | +-----+
| | | | | | +-- Pilsbryoconcha_carinifera **
| | | | +--+
| | | | | +----- Pseudodon_mekongi **
| | | | | |
| | | | +--+ +-- Nyeinchanconcha_nyeinchani **
| | | | | |
| | | | +--+-- Bineurus_mouhotii **
| | | | |
| | +--+ +-- Sundadontina_tumida **
| | |
| | | +-------- Inversidens_brandtii **
| | | |
| | | +-----+ +-- Obovalis_omiensis **
| | | | | +--+
| | | | | | +-- Ptychorhynchus_pfisteri **
| | | | +--+
| | | | | +-- Parvasolenaia_rivularis **
| | | | +--+
| | +--+ +-- Koreosolenaia_sitgyensis **
| | |
+--+ | +----- Gonidea_angulata **
| | +-----+
| | | | +-- Leguminaia_wheatleyi **
| | | +--+
| +--+ +-- Microcondylaea_bonellii **
| |
| | +-------- Sinosolenaia_carinata **
| +--+
| | +----- Sinosolenaia_recognita **
| +--+
| | +-- Sinosolenaia_iridinea **
| +--+
| +-- Sinosolenaia_oleivora **
|
| +-------------------------------- Acuticosta_chinensis **
| |
| +-------------------------------- Lepidodesma_languilati **
| |
| | +----------- Aculamprotula_tortuosa **
| | +--------------+
| | | | +-------- Aculamprotula_scripta **
| | | +--+
| | | | +----- Aculamprotula_coreana **
| | | +--+
| | | | +-- Aculamprotula_fibrosa **
| | | +--+
| +-----+ +-- Aculamprotula_tientsinensis **
| | |
| | | +----------------------- Middendorffinaia_mongolica **
| | | |
| | | | +----- Unio_gibbus **
| | | | |
| | | | +--------------+----- Unio_tumidus **
| | +--+ | |
| | | | | +-- Unio_crassus **
| | | | +--+
| | | | +-- Unio_pictorum **
| | | |
| | | | +-------------- Nodularia_breviconcha **
| | +--+ +--+
| | | | | +----------- Nodularia[2] **
| | | | +--+
| | | | | +-------- Nodularia_nuxpersicae **
| | | | +--+
| | | | | +----- Nodularia_nipponensis **
| | | | +--+
| | | | | +-- Nodularia_douglasiae **
| | +--+ +--+
| | | +-- Nodularia[1] **
| | |
+--+ | +-- Schistodesmus_lampreyanus **
| | +-----------+
| | | +-- Schistodesmus_spinosus **
| | |
| | | +----- Cuneopsis_rufescens **
| | +--------+
| +--+ | +-- Cuneopsis_heudei **
| | +--+
| | +-- Cuneopsis_celtiformis **
| |
| | +----------- Tchangsinaia_piscicula **
| +--+
| | +-------- Inversiunio_jokohamensis **
| +--+
| | +----- Inversiunio_verrucosus **
| +--+
| | +-- Inversiunio_reinianus **
| +--+
| +-- Inversiunio_yanagawensis **
|
| +----------------- Lanceolaria_oxyrhyncha **
| |
| +-----------+ +-------------- Lanceolaria_gladiola **
| | | |
| | +--+ +-- Lanceolaria_bilirata **
| | | +--------+
| | | | +-- Lanceolaria_grayii **
| | +--+
| | | +-------- Lanceolaria_acrorrhyncha **
| | +--+
| | | +----- Lanceolaria_kihirai **
| | +--+
| | | +-- Lanceolaria_lanceolata **
| | +--+
| | +-- Lanceolaria_triformis **
| |
| | +-------- Pseudanodonta_complanata **
+--+ |
| | +----- Anodonta_cygnea **
| +--+
| | | +-- Anodonta_anatina **
| +-----------------+ +--+
| | | +-- Anodonta_nuttalliana **
| | |
| | | +----- Pyganodon_grandis **
| | | |
| | | +----- Simpsonaias_ambigua **
| | +--+
| | +----- Strophitus_undulatus **
| | |
| | | +-- Alasmidonta_marginata **
| | +--+
| | +-- Lasmigona_costata **
+--+
| +-- Cristaria_plicata **
| +--------------------+
| | +-- Cristaria_clessini **
| |
| | +-------------- Anemina_arcaeformis **
| | |
| +--------+ +----------- Amuranodonta_kijaensis **
| | | |
| | +--+ +-------- Buldowskia_suifunica **
| | | |
| | +--+ +-- Buldowskia_flavotincta **
+--+ | +--+
| | | +-- Buldowskia_shadini **
| +--+
| | +-- Buldowskia_iwakawai **
| +--+
| +-- Buldowskia_kamiyai **
|
| +-------------------- Pletholophus_tenuis **
| |
| | +-------- Beringiana_japonica **
| | +--------+
+--+ | | +----- Beringiana_beringiana **
| | +--+
| | | +-- Beringiana_fukuharai **
| | +--+
+--+ +-- Beringiana_gosannensis **
|
| +-- Sinanodonta_angula **
| +-----------+
| | +-- Sinanodonta_lucida **
| |
+--+ +----------- Sinanodonta_schrenkii **
| |
| | +-- Sinanodonta_calipygos **
| +--------+
+--+ +-- Sinanodonta_tumens **
|
| +-------- Sinanodonta_woodiana[2] **
| |
+--+ +----- Sinanodonta_elliptica **
| |
+--+----- Sinanodonta_jourdyi **
|
| +-- Sinanodonta_lauta **
+--+
+-- Sinanodonta_woodiana[1] **
Wu et al., 2023 [Publ 4339], fig. 4, S5-nt
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 93 |
93 |
88 |
2 |
4 |
3 |
2 |
112.0 |
+-------------------------------------------------- Echyridella_menziesii
|
| +-- Anodontites_trapesialis
| +--------------------------------------------+
| | +-- Mutela_dubia
| |
| | +-- Gibbosula_crassa **
| | +-----------+
--+ | | +-- Gibbosula_rochechouartii **
| | +-----------------------------+
| | | | +----------- Cumberlandia_monodonta **
| | | +--+
| | | | +-------- Pseudunio_marocanus **
| | | +--+
| | | | +----- Margaritifera_falcata **
| | | +--+
| | | | +-- Margaritifera_dahurica **
+--+ | +--+
| | +-- Margaritifera_margaritifera **
| |
| | +-- Quadrula_quadrula **
| | +--------------------------+
| | | +-- Uniomerus_tetralasmus **
| | |
| | +-----------+ +-- Elliptio_complanata **
| | | | +-----------------------+
| | | | | +-- Pleurobema_oviforme **
| | | +--+
| | | | +----------------------- Amblema_plicata **
| | | | |
| | | +--+ +-------------------- Popenaias_popeii **
+--+ | | |
| | +--+ +----------------- Toxolasma_parvum **
| | | |
| | | | +----- Potamilus_leptodon **
| | +--+ +--------+
| | | | | +-- Potamilus_alatus **
| | | | +--+
| | | | +-- Potamilus_streckersoni **
| | +--+
| | | +----------- Leaunio_lienosus **
| | | |
| | | | +----- Lampsilis_ornata **
| | +--+ +--+
| | | | | +-- Lampsilis_cardium **
| | | | +--+
| | +--+ +-- Lampsilis_cariosa **
| | |
| | | +----- Venustaconcha_ellipsiformis **
| | +--+
| | | +-- Lampsilis_powellii **
| | +--+
| | +-- Lampsilis_siliquoidea **
+--+
| +----- Chamberlainia_somsakpanhai **
| +-----------------+
| | | +-- Sinohyriopsis_cumingii **
| | +--+
| | +-- Sinohyriopsis_schlegelii **
| |
| | +-- Hyriopsis_bialata **
| | +--+
| +--------------+ | +-- Rectidens_sumatrensis **
| | | +--------------+
| | | | | +-- Lens_contradens **
| | | | +--+
| | | | +-- Physunio_superbus **
| | | |
| | | | +----------- Potomida_littoralis **
| | +--+ +-----+
| | | | | +-------- Pronodularia_japanensis **
| | | | +--+
| | | | | +----- Lamprotula_leaii **
| | | | +--+
| | | | | +-- Lamprotula_caveata **
| | +--+ +--+
| | | +-- Lamprotula_cornuumlunae **
| | |
| | | +-- Pseudodon_vondembuschianus **
| | | +-----------+
| | | | +-- Pilsbryoconcha_exilis **
| | | |
| | +--+ +-- Ptychorhynchus_pfisteri **
+--+ | +--------+
| | | +-- Microcondylaea_bonellii **
| +--+
| | +-------- Parvasolenaia_rivularis **
| +--+
| | +----- Sinosolenaia_carinata **
| +--+
| | +-- Sinosolenaia_iridinea **
| +--+
| +-- Sinosolenaia_oleivora **
|
| +----------------------------------- Lamellidens_marginalis **
| |
| | +-- Lepidodesma_aligera **
| | +-----------------------------+
| | | +-- Lepidodesma_languilati **
| | |
| | | +-------- Acuticosta_chinensis **
| | | +--------------+
| | | | | +----- Lanceolaria_gladiola **
| | | | +--+
+--+ | | | +-- Lanceolaria_grayii **
| | | +--+
| | | +-- Lanceolaria_lanceolata **
| | |
| | | +-------------- Cristaria_plicata **
| | +-----+ |
| | | | +-----+ +-- Anemina_arcaeformis **
| | | | | | +--------+
| | | | | | | +-- Anemina_euscaphys **
| | | | | +--+
+--+ | | | | +-------- Beringiana_fukuharai **
| | | | +--+
| | | | | +----- Sinanodonta_lucida **
| | | | +--+
| | +--+ | +-- Sinanodonta_tumens **
| | | +--+
| | | +-- Sinanodonta_woodiana **
| | |
| | | +----------------- Anodonta_nuttalliana **
| | | |
| | | +----------------- Pseudanodonta_complanata **
| | | |
| | +--+ +-------------- Anodonta_cygnea **
| | | |
| | | | +-- Anodonta_exulcerata **
| | | +-----------+
| | | | +-- Anodonta_anatina **
| | +--+
+--+ | +----- Pyganodon_grandis **
| | +-----+
| | | | +-- Utterbackia_imbecillis **
| | | +--+
| +--+ +-- Utterbackia_peninsularis **
| |
| | +-------- Lasmigona_complanata **
| +--+
| | +----- Prolasmidonta_heterodon **
| +--+
| | +-- Alasmidonta_varicosa **
| +--+
| +-- Platynaias_compressa **
|
| +-------- Aculamprotula_tortuosa **
| |
| +-----------------+ +-- Aculamprotula_coreana **
| | | +--+
| | | | +-- Aculamprotula_tientsinensis **
| | +--+
| | | +-- Aculamprotula_polysticta **
| | +--+
| | +-- Aculamprotula_scripta **
| |
+--+ +----------- Unio_tumidus **
| |
| +-----------+ +-------- Unio_crassus **
| | | |
| | +--+ +----- Unio_elongatulus **
| | | |
| | +--+ +-- Unio_delphinus **
| | | |
| | +--+-- Unio_mancus **
+--+ |
| +-- Unio_pictorum **
|
| +-- Pseudocuneopsis_capitata **
| +-----------------+
| | +-- Pseudocuneopsis_sichuanensis **
| |
+--+ +-- Nodularia_breviconcha **
| +--------------+
| | +-- Nodularia_nipponensis **
| |
+--+ +----- Tchangsinaia_piscicula **
| +--------+
| | | +-- Schistodesmus_lampreyanus[2] **
| | +--+
+--+ +-- Schistodesmus_spinosus **
|
| +----------- Schistodesmus_lampreyanus[1] **
+--+
| +-------- Cuneopsis_rufescens **
+--+
| +----- Cuneopsis_celtiformis **
+--+
| +-- Cuneopsis_demangei **
+--+
+-- Cuneopsis_heudei **
Wu et al., 2023 [Publ 4339], fig. 4, S5-aa
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 92 |
92 |
90 |
2 |
4 |
3 |
2 |
113.0 |
+----------------------------------------------- Echyridella_menziesii
|
| +-- Anodontites_trapesialis
| +-----------------------------------------+
| | +-- Mutela_dubia
| |
| | +-- Gibbosula_crassa **
| | +-----------+
--+ | | +-- Gibbosula_rochechouartii **
| | +--------------------------+
| | | | +----------- Cumberlandia_monodonta **
| | | +--+
| | | | +-------- Pseudunio_marocanus **
| | | +--+
| | | | +----- Margaritifera_falcata **
| | | +--+
| | | | +-- Margaritifera_dahurica **
+--+ | +--+
| | +-- Margaritifera_margaritifera **
| |
| | +-- Quadrula_quadrula **
| | +--------------------------+
| | | +-- Uniomerus_tetralasmus **
| | |
| | +--------+ +-- Elliptio_complanata **
| | | | +-----------------------+
| | | | | +-- Pleurobema_oviforme **
| | | +--+
| | | | +----------------------- Amblema_plicata **
| | | | |
| | | +--+ +-------------------- Popenaias_popeii **
+--+ | | |
| | +--+ +----------------- Toxolasma_parvum **
| | | |
| | | | +----- Potamilus_leptodon **
| | +--+ +--------+
| | | | | +-- Potamilus_alatus **
| | | | +--+
| | | | +-- Potamilus_streckersoni **
| | +--+
| | | +----------- Leaunio_lienosus **
| | | |
| | | | +----- Lampsilis_ornata **
| | +--+ +--+
| | | | | +-- Lampsilis_cardium **
| | | | +--+
| | +--+ +-- Lampsilis_cariosa **
| | |
| | | +----- Venustaconcha_ellipsiformis **
| | +--+
| | | +-- Lampsilis_powellii **
| | +--+
| | +-- Lampsilis_siliquoidea **
+--+
| +----- Chamberlainia_somsakpanhai **
| +-----------------+
| | | +-- Sinohyriopsis_cumingii **
| | +--+
| | +-- Sinohyriopsis_schlegelii **
| |
| | +-- Hyriopsis_bialata **
| | +--+
| +-----------+ | +-- Rectidens_sumatrensis **
| | | +--------------+
| | | | | +-- Lens_contradens **
| | | | +--+
| | | | +-- Physunio_superbus **
| | | |
| | | | +----------- Potomida_littoralis **
| | +--+ +-----+
| | | | | +-------- Pronodularia_japanensis **
| | | | +--+
| | | | | +----- Lamprotula_leaii **
| | | | +--+
| | | | | +-- Lamprotula_caveata **
| | +--+ +--+
| | | +-- Lamprotula_cornuumlunae **
| | |
| | | +-- Pseudodon_vondembuschianus **
| | | +-----------+
| | | | +-- Pilsbryoconcha_exilis **
| | +--+
| | | +----------- Microcondylaea_bonellii **
+--+ | |
| +--+ +-- Parvasolenaia_rivularis **
| | +-----+
| | | +-- Ptychorhynchus_pfisteri **
| +--+
| | +----- Sinosolenaia_carinata **
| +--+
| | +-- Sinosolenaia_iridinea **
| +--+
| +-- Sinosolenaia_oleivora **
|
| +-------------------------------- Lamellidens_marginalis **
| |
| | +-- Lepidodesma_aligera **
| | +--------------------------+
| | | +-- Lepidodesma_languilati **
| | |
| | | +-------- Aculamprotula_tortuosa **
| | | |
| | | +--------------+ +-- Aculamprotula_coreana **
| | | | | +--+
+--+ | | | | +-- Aculamprotula_tientsinensis **
| | | +--+
| | | | +-- Aculamprotula_polysticta **
| | | +--+
| | | +-- Aculamprotula_scripta **
| | |
| | +--+ +-------------- Unio_tumidus **
| | | | +-----+
| | | | | | +----------- Unio_crassus **
| | | | | +--+
+--+ | | | | +-------- Unio_elongatulus **
| | | | +--+
| | | | | +----- Unio_pictorum **
| | | | +--+
| | | | | +-- Unio_delphinus **
| | +--+ +--+
| | | +-- Unio_mancus **
| | |
| | | +-- Nodularia_breviconcha **
| | | +--+
| | | | +-- Nodularia_nipponensis **
| | | +-----------+
| | | | | +-- Pseudocuneopsis_capitata **
| | | | +--+
| | +--+ +-- Pseudocuneopsis_sichuanensis **
| | |
| | | +-------------- Tchangsinaia_piscicula **
| | | |
+--+ +--+ +-- Schistodesmus_lampreyanus **
| | +--------+
| | | +-- Schistodesmus_spinosus **
| +--+
| | +-------- Cuneopsis_rufescens **
| +--+
| | +----- Cuneopsis_celtiformis **
| +--+
| | +-- Cuneopsis_demangei **
| +--+
| +-- Cuneopsis_heudei **
|
| +-------- Acuticosta_chinensis **
| +--------+
| | | +----- Lanceolaria_gladiola **
| | +--+
| | | +-- Lanceolaria_grayii **
| | +--+
| | +-- Lanceolaria_lanceolata **
| |
| | +----- Cristaria_plicata **
| | +-----+
| | | | +-- Anemina_arcaeformis **
+--------+ | +--+
| +--+ +-- Anemina_euscaphys **
| | |
| | | +-------- Beringiana_fukuharai **
| | +--+
| | | +----- Sinanodonta_lucida **
| | +--+
| | | +-- Sinanodonta_tumens **
| | +--+
| | +-- Sinanodonta_woodiana **
+--+
| +-- Anodonta_nuttalliana **
| +-----+
| | +-- Pseudanodonta_complanata **
| +--+
| | | +----- Anodonta_anatina **
| | +--+
| | | +-- Anodonta_cygnea **
| | +--+
| | +-- Anodonta_exulcerata **
+--+
| +----- Pyganodon_grandis **
| +--+
| | | +-- Utterbackia_peninsularis **
| | +--+
| | +-- Utterbackia_imbecillis **
+--+
| +-- Alasmidonta_varicosa **
| +--+
| | +-- Platynaias_compressa **
+--+
| +-- Prolasmidonta_heterodon **
+--+
+-- Lasmigona_complanata **
Wu et al., 2024 [Publ 4350], fig. 8a, S1
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 88 |
88 |
85 |
2 |
4 |
3 |
2 |
106.0 |
+----------------------------------------------- Echyridella_menziesii
|
| +-- Anodontites_trapesialis
| +-----------------------------------------+
| | +-- Mutela_dubia
| |
| | +-- Margaritifera_dahurica **
--+ | +--------------------------------------+
| | | +-- Margaritifera_margaritifera **
| | |
| | | +-- Quadrula_quadrula **
| | | +--------------------------+
| | | | +-- Uniomerus_tetralasmus **
| | | |
+--+ | +--------+ +-- Elliptio_complanata **
| | | | +-----------------------+
| | | | | +-- Pleurobema_oviforme **
| | | +--+
| | | | +----------------------- Amblema_plicata **
| | | | |
| | | +--+ +-------------------- Popenaias_popeii **
| | | | |
| | | +--+ +----------------- Toxolasma_parvum **
| | | | |
| | | | | +----- Potamilus_leptodon **
+--+ | +--+ +--------+
| | | | | +-- Potamilus_alatus **
| | | | +--+
| | | | +-- Potamilus_streckersoni **
| | +--+
| | | +----------- Leaunio_lienosus **
| | | |
| | | | +----- Lampsilis_ornata **
| | +--+ +--+
| | | | | +-- Lampsilis_cardium **
| | | | +--+
| | +--+ +-- Lampsilis_cariosa **
| | |
| | | +----- Venustaconcha_ellipsiformis **
| | +--+
| | | +-- Lampsilis_powellii **
| | +--+
| | +-- Lampsilis_siliquoidea **
| |
+--+ +----- Chamberlainia_somsakpanhai **
| +--------------------+
| | | +-- Sinohyriopsis_cumingii **
| | +--+
| | +-- Sinohyriopsis_schlegelii **
| |
| | +-- Hyriopsis_bialata **
| | +--+
| +--------+ | +-- Rectidens_sumatrensis **
| | | +-----------------+
| | | | | +-- Lens_contradens **
| | | | +--+
| | | | +-- Physunio_superbus **
| | | |
| | | | +----------- Potomida_littoralis **
| | +--+ +--------+
| | | | | +-------- Pronodularia_japanensis **
| | | | +--+
| | | | | +----- Lamprotula_leaii **
| | | | +--+
| | | | | +-- Lamprotula_caveata **
| | +--+ +--+
| | | +-- Lamprotula_cornuumlunae **
| | |
| | | +-- Pseudodon_vondembuschianus **
| | | +--------------+
| | | | +-- Pilsbryoconcha_exilis **
| | +--+
| | | +-------------- Inversidens_rentianensis **
| | | |
+--+ +--+ +-- Parvasolenaia_rivularis **
| | +--------+
| | | +-- Ptychorhynchus_pfisteri **
| +--+
| | +-------- Microcondylaea_bonellii **
| +--+
| | +----- Sinosolenaia_carinata **
| +--+
| | +-- Sinosolenaia_iridinea **
| +--+
| +-- Sinosolenaia_oleivora **
|
| +-------------------------------- Lamellidens_marginalis **
| |
| | +-- Lepidodesma_aligera **
| | +--------------------------+
| | | +-- Lepidodesma_languilati **
| | |
| | | +-------- Aculamprotula_tortuosa **
| | | |
| | | +--------------+ +-- Aculamprotula_coreana **
| | | | | +--+
+--+ | | | | +-- Aculamprotula_tientsinensis **
| | | +--+
| | | | +-- Aculamprotula_polysticta **
| | | +--+
| | | +-- Aculamprotula_scripta **
| | |
| | +--+ +-------------- Unio_tumidus **
| | | | +-----+
| | | | | | +----------- Unio_crassus **
| | | | | +--+
+--+ | | | | +-------- Unio_elongatulus **
| | | | +--+
| | | | | +----- Unio_pictorum **
| | | | +--+
| | | | | +-- Unio_delphinus **
| | +--+ +--+
| | | +-- Unio_mancus **
| | |
| | | +-- Nodularia_breviconcha **
| | | +--------------+
| | | | +-- Nodularia_douglasiae **
| | | |
| | | | +-- Pseudocuneopsis_capitata **
| | | +--------------+
| | +--+ +-- Pseudocuneopsis_sichuanensis **
| | |
| | | +-- Schistodesmus_lampreyanus **
| | | +-----------+
+--+ | | +-- Schistodesmus_spinosus **
| +--+
| | +----------- Tchangsinaia_piscicula **
| +--+
| | +-------- Cuneopsis_rufescens **
| +--+
| | +----- Cuneopsis_celtiformis **
| +--+
| | +-- Cuneopsis_demangei **
| +--+
| +-- Cuneopsis_heudei **
|
| +-------- Acuticosta_chinensis **
| +--------------+
| | | +----- Lanceolaria_gladiola **
| | +--+
| | | +-- Lanceolaria_grayii **
| | +--+
| | +-- Lanceolaria_lanceolata **
| |
| | +-------------- Cristaria_plicata **
| | |
+--+ +-----+ +-- Anemina_arcaeformis **
| | | +--------+
| | | | +-- Anemina_euscaphys **
| | +--+
| | | +-------- Beringiana_fukuharai **
| | +--+
| | | +----- Sinanodonta_lucida **
| | +--+
| | | +-- Sinanodonta_tumens **
+--+ +--+
| +-- Sinanodonta_woodiana **
|
| +-------- Pseudanodonta_complanata **
| +--------+
| | | +----- Anodonta_anatina **
| | +--+
| | | +-- Anodonta_cygnea **
| | +--+
+--+ +-- Anodonta_exulcerata **
|
| +-------------- Anodonta_nuttalliana **
| |
| | +----- Pyganodon_grandis **
+--+ +-----+
| | | +-- Utterbackia_imbecillis **
| | +--+
+--+ +-- Utterbackia_peninsularis **
|
| +-------- Lasmigona_complanata **
+--+
| +----- Prolasmidonta_heterodon **
+--+
| +-- Alasmidonta_varicosa **
+--+
+-- Platynaias_compressa **
Wu et al., 2024 [Publ 4350], fig. 8b, S2
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 88 |
88 |
84 |
2 |
4 |
3 |
2 |
106.0 |
+----------------------------------------------- Echyridella_menziesii
|
| +-- Anodontites_trapesialis
| +-----------------------------------------+
| | +-- Mutela_dubia
| |
| | +-- Margaritifera_dahurica **
--+ | +--------------------------------------+
| | | +-- Margaritifera_margaritifera **
| | |
| | | +-- Quadrula_quadrula **
| | | +--------------------------+
| | | | +-- Uniomerus_tetralasmus **
| | | |
+--+ | +--------+ +-- Elliptio_complanata **
| | | | +-----------------------+
| | | | | +-- Pleurobema_oviforme **
| | | +--+
| | | | +----------------------- Amblema_plicata **
| | | | |
| | | +--+ +-------------------- Popenaias_popeii **
| | | | |
| | | +--+ +----------------- Toxolasma_parvum **
| | | | |
| | | | | +----- Potamilus_leptodon **
+--+ | +--+ +--------+
| | | | | +-- Potamilus_alatus **
| | | | +--+
| | +--+ +-- Potamilus_streckersoni **
| | |
| | | +----------- Leaunio_lienosus **
| | | |
| | +--+ +-------- Venustaconcha_ellipsiformis **
| | | |
| | | | +-- Lampsilis_powellii **
| | +--+-----+
| | | +-- Lampsilis_siliquoidea **
| | |
| | | +----- Lampsilis_ornata **
| | +--+
| | | +-- Lampsilis_cardium **
| | +--+
| | +-- Lampsilis_cariosa **
| |
+--+ +----- Chamberlainia_somsakpanhai **
| +--------------------+
| | | +-- Sinohyriopsis_cumingii **
| | +--+
| | +-- Sinohyriopsis_schlegelii **
| |
| | +-- Hyriopsis_bialata **
| | +--+
| +--------+ | +-- Rectidens_sumatrensis **
| | | +-----------------+
| | | | | +-- Lens_contradens **
| | | | +--+
| | | | +-- Physunio_superbus **
| | | |
| | | | +----------- Potomida_littoralis **
| | +--+ +--------+
| | | | | +-------- Pronodularia_japanensis **
| | | | +--+
| | | | | +----- Lamprotula_leaii **
| | | | +--+
| | | | | +-- Lamprotula_caveata **
| | +--+ +--+
| | | +-- Lamprotula_cornuumlunae **
| | |
| | | +-- Pseudodon_vondembuschianus **
| | | +--------------+
| | | | +-- Pilsbryoconcha_exilis **
| | +--+
| | | +-------------- Inversidens_rentianensis **
| | | |
+--+ +--+ +-- Parvasolenaia_rivularis **
| | +--------+
| | | +-- Ptychorhynchus_pfisteri **
| +--+
| | +-------- Microcondylaea_bonellii **
| +--+
| | +----- Sinosolenaia_carinata **
| +--+
| | +-- Sinosolenaia_iridinea **
| +--+
| +-- Sinosolenaia_oleivora **
|
| +-------------------------------- Lamellidens_marginalis **
| |
| | +-- Lepidodesma_aligera **
| | +--------------------------+
| | | +-- Lepidodesma_languilati **
| | |
| | | +-------- Aculamprotula_tortuosa **
| | | |
| | | +--------------+ +-- Aculamprotula_coreana **
| | | | | +--+
+--+ | | | | +-- Aculamprotula_tientsinensis **
| | | +--+
| | | | +-- Aculamprotula_polysticta **
| | | +--+
| | | +-- Aculamprotula_scripta **
| | |
| | +--+ +-------------- Unio_tumidus **
| | | | +-----+
| | | | | | +----------- Unio_crassus **
| | | | | +--+
+--+ | | | | +-------- Unio_elongatulus **
| | | | +--+
| | | | | +----- Unio_pictorum **
| | | | +--+
| | | | | +-- Unio_delphinus **
| | +--+ +--+
| | | +-- Unio_mancus **
| | |
| | | +-- Nodularia_breviconcha **
| | | +--------------+
| | | | +-- Nodularia_douglasiae **
| | | |
| | | | +-- Pseudocuneopsis_capitata **
| | | +--------------+
| | +--+ +-- Pseudocuneopsis_sichuanensis **
| | |
| | | +-- Schistodesmus_lampreyanus **
+--+ | +-----------+
| | | +-- Schistodesmus_spinosus **
| +--+
| | +----------- Tchangsinaia_piscicula **
| +--+
| | +-------- Cuneopsis_rufescens **
| +--+
| | +----- Cuneopsis_celtiformis **
| +--+
| | +-- Cuneopsis_demangei **
| +--+
| +-- Cuneopsis_heudei **
|
| +-------- Acuticosta_chinensis **
| +--------------+
| | | +----- Lanceolaria_gladiola **
| | +--+
| | | +-- Lanceolaria_grayii **
| | +--+
| | +-- Lanceolaria_lanceolata **
| |
| | +-------------- Cristaria_plicata **
+--+ |
| +-----+ +-- Anemina_arcaeformis **
| | | +--------+
| | | | +-- Anemina_euscaphys **
| | +--+
| | | +-------- Beringiana_fukuharai **
| | +--+
| | | +----- Sinanodonta_lucida **
| | +--+
+--+ | +-- Sinanodonta_tumens **
| +--+
| +-- Sinanodonta_woodiana **
|
| +----------------- Anodonta_nuttalliana **
| |
| | +-------- Pseudanodonta_complanata **
| | +-----+
+--+ | | +----- Anodonta_anatina **
| | +--+
| | | +-- Anodonta_cygnea **
| | +--+
+--+ +-- Anodonta_exulcerata **
|
| +----- Pyganodon_grandis **
| +-----+
| | | +-- Utterbackia_imbecillis **
| | +--+
+--+ +-- Utterbackia_peninsularis **
|
| +-------- Lasmigona_complanata **
+--+
| +----- Prolasmidonta_heterodon **
+--+
| +-- Alasmidonta_varicosa **
+--+
+-- Platynaias_compressa **
Bolotov et al., 2018 [Publ 3826], fig. 2
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 85 |
85 |
50 |
1 |
0 |
0 |
0 |
96.0 |
+-- Trapezidens_exolescens **
+--------+
| +-- Trapezidens_dolichorhynchus **
+--------------+
| | +-------- Lamellidens_savadiensis **
| +--+
| | +----- Lamellidens_ferrugineus **
| +--+
| | +-- Lamellidens_generosus **
| +--+
+--------+ +-- Lamellidens_mainwaringi **
| |
| | +-- Parreysia_keralaensis **
| | +--------------------+
| | | +-- Parreysia_nagpoorensis **
| | |
| | | +----- Leoparreysia_olivacea **
| +--+ +--------------+
| | | | +-- Leoparreysia_canefrii **
| | | +--+
| | | +-- Leoparreysia_tavoyensis **
| | |
| +--+ +-- Coelatura_aegyptiaca **
| | +--------------+
| | | +-- Nitia_teretiuscula **
| | |
| | | +-------- Indonaia_shurtleffiana **
| +--+ +-----+
| | | | +----- Indonaia_rugosa **
| | | +--+
| | | | +-- Indonaia_caerulea **
| | | +--+
| +--+ +-- Indonaia_andersoniana **
| |
| | +----------- Indochinella_pugio **
| | |
| | | +----- Radiatula_myitkyinae **
| +--+ +--+
| | | | +-- Radiatula_mouhoti **
| | | +--+
| +--+ +-- Radiatula_chaudhurii **
| |
| | +----- Scabies_mandarinus **
| +--+
| | +-- Scabies_humilis **
| +--+
--+ +-- Scabies_phaselus **
|
| +-- Indopseudodon_indawgyiensis **
| |
| +--+-- Indopseudodon_bogani **
| | |
| | +-- Indopseudodon_salwenianus **
| |
| | +-- Bineurus_mouhotii **
| +--------------------------+ |
| | | +-- Pseudodon_vagulus **
| | | |
| | | +-- Pseudodon_mekongi **
| | | |
| | | +-- Pilsbryoconcha_exilis **
| | +--+
| | +-- Pilsbryoconcha_carinifera **
| | |
| | +-- Bineurus_loeiensis **
| | |
| | +-- Bineurus_exilis **
| | |
| | +-- Pseudodon_vondembuschianus **
| |
| | +-- Ensidens_telus **
| | |
| | +-- Ensidens_ingallsianus **
| | |
| | +-- Ensidens_jaculus **
| | |
| | +-- Hyriopsis_desowitzi **
| | +--+
| | | +-- Hyriopsis_myersiana **
| | | |
| | | +-- Hyriopsis_khoratensis **
| | | |
+--+ | +-- Ensidens_spiculus **
| | |
| | +-- Ensidens_sagittarius **
| |
| +-----------------------+ +-- Lens_eximius **
| | | |
| | | +-- Physunio_modelli[1] **
| | | |
| | | +-- Yaukthwa_nesemanni **
| | | |
| | | +-- Yaukthwa_panhai **
| | | |
| | | +-- Lens_comptus **
| | | |
| | +--+-- Lens_novoselovi **
| | |
| | +-- Physunio_modelli[2] **
| | |
| | +-- Physunio **
| | |
| | +-- Yaukthwa_inlenensis **
| | |
| | +-- Lens_contradens **
| | |
| | +-- Lens_micropterus **
| |
| | +-- Lamprotula_leaii **
+--+ |
| +-- Potomida_littoralis **
| |
| +-- Pronodularia_japanensis **
| |
| +-- Gonidea_angulata **
| |
| +-- Leguminaia_wheatleyi **
| |
| +-- Microcondylaea_bonellii **
| |
| +-----------------------+-- Amblema_plicata **
| | |
| | +-- Ortmanniana_ligamentina **
| | |
| | +-- Lampsilis_cardium **
| | |
| | +-- Cambarunio_iris **
| | |
| | +-- Elliptio_complanata **
| | |
| | +-- Pleurobema_sintoxia **
| | |
+--+ +-- Quadrula_quadrula **
|
| +----- Unio_tumidus **
| +--+
| | | +-- Unio_crassus **
| | +--+
| +--------------+ +-- Unio_pictorum **
| | |
| | | +----- Nodularia_douglasiae[1] **
| | +--+
| | | +-- Nodularia_nuxpersicae **
| | +--+
+--+ +-- Nodularia_douglasiae[2] **
|
| +-- Lanceolaria_gladiola **
| +-----------------+
| | +-- Lanceolaria_grayii **
| |
| | +-------- Cristaria_plicata **
+--+ +--------+
| | | +----- Pletholophus_tenuis **
| | +--+
| | | +-- Sinanodonta_jourdyi **
+--+ +--+
| +-- Sinanodonta_lauta **
|
| +-------------- Anodonta_anatina **
+--+
| +----------- Pyganodon_grandis **
+--+
| +-------- Simpsonaias_ambigua **
+--+
| +----- Strophitus_undulatus **
+--+
| +-- Alasmidonta_marginata **
+--+
+-- Lasmigona_costata **
Konopleva et al., 2023 [Publ 4262], fig. 8
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 84 |
84 |
80 |
1 |
0 |
0 |
0 |
129.0 |
+----------------------- Indonaia_andersoniana **
|
+--------------+ +-------------------- Indochinella_pugio **
| | |
| +--+ +----------------- Radiatula_mouhoti **
| | |
| +--+ +-------------- Scabiellus_songkramensis **
| | |
| | | +-- Unionetta_fabagina **
| +--+ +--------+
| | | +-- Harmandia_somboriensis **
| | |
| | | +-- Scabies_humilis **
| +--+ +-----+
| | | +-- Scabies_pilata **
| | |
| | | +----- Scabies_crispata **
| | +--+
| +--+ | +-- Scabies_nucleus **
| | +--+
| | +-- Scabies_mandarinus **
| |
| | +-- Scabies_phaselus **
| | |
| +-----+-- Scabies_anceps **
| |
| +-- Scabies_scobinatus **
|
| +-------- Indopseudodon_kayinensis **
| +-----------------------+
| | | +----- Indopseudodon_indawgyiensis **
| | +--+
| | | +-- Indopseudodon_salwenianus **
| | +--+
| | +-- Indopseudodon_bogani **
| |
| | +-- Pilsbryoconcha_schomburgki **
--+ +--+ +--------+
| | | | +-- Pilsbryoconcha_exilis[1] **
| | | +-----------------+
| | | | | +-------- Pilsbryoconcha **
| | | | +--+
| | | | | +----- Pilsbryoconcha_linguaeformis **
| | | | +--+
| | | | | +-- Pilsbryoconcha_exilis[2] **
| | +--+ +--+
| | | +-- Pilsbryoconcha_carinifera **
| | |
| | | +-------------------------- Songkhlanaia_tamodienica **
| | | |
| | | | +-- Namkongnaia_inkhavilayi **
| | | | +--------------------+
| | +--+ | +-- Namkongnaia_lemeslei **
| | | |
| | | | +-------- Pseudodon_lenyanensis **
| | | | +-----------+
| | | | | | +----- Pseudodon_vagulus **
| | +--+ | +--+
| | | | | +-- Pseudodon_vondembuschianus **
| | | | +--+
| | | | +-- Pseudodon_mekongi **
| | | |
| | | | +-- Sundadontina_taskaevi **
| | +--+ +--------+
| | | | +-- Sundadontina_brandti **
| | | +-----+
| | | | | +-------- Sundadontina_tumida **
| | | | +--+
| | | | | +----- Sundadontina_tanintharyiensis **
| | | | +--+
| | | | | +-- Sundadontina_plugpomenica **
| | +--+ +--+
| | | +-- Sundadontina_cumingii **
+--+ |
| | +-------------- Nyeinchanconcha_nyeinchani **
| | |
| | | +----- Thaiconcha_thaiensis **
| +--+ +-----+
| | | | +-- Thaiconcha_munelliptica **
| | | +--+
| +--+ +-- Thaiconcha_callifera **
| |
| | +-------- Bineurus_loeiensis **
| +--+
| | +----- Bineurus_mouhotii **
| +--+
| | +-- Bineurus_anodontinum **
| +--+
| +-- Bineurus_exilis **
|
| +-- Gonidea_angulata **
| +--+
| | +-- Leguminaia_wheatleyi **
| +-----------------------+
| | | +-- Lamprotula_leaii **
| | +--+
| | +-- Potomida_littoralis **
| |
| | +-- Ensidens_ingallsianus **
| | +-----+
| | | +-- Ensidens_telus **
| | |
| | +--------------+ +-- Ensidens_jaculus **
| | | | +--+
| | | | | +-- Ensidens_spiculus[1] **
| | | +--+
| | | | +-- Ensidens_spiculus[2] **
| | +--+ +--+
| | | | +-- Ensidens_dugasti **
+-----+ | |
| | | +-- Hyriopsis_desowitzi **
| | | +-----------------+
| | | | +-- Hyriopsis_delaportei **
| | +--+
| | | +----------------- Rectidens_sumatrensis **
| | | |
| | +--+ +-------------- Hyriopsis_panhai **
| | | |
| | +--+ +----------- Hyriopsis_myersiana **
| | | |
| | +--+ +-- Hyriopsis_khoratensis **
| | | +-----+
| | | | +-- Hyriopsis_bialata **
| | +--+
| | | +----- Hyriopsis_sakhonensis **
+--+ +--+
| | +-- Hyriopsis_phuphaniensis **
| +--+
| +-- Hyriopsis_kratiensis **
|
| +----- Physunio_modelli **
| +-----+
| | | +-- Physunio_superbus **
| | +--+
| | +-- Physunio_massini **
| +-----+
| | | +-- Trapezoideus_thachiadensis **
| | | +-----+
| | | | +-- Trapezoideus_subviridis **
| | +--+
| | | +----- Trapezoideus_mitanensis **
| | +--+
| | | +-- Trapezoideus_foliaceus **
+--------+ +--+
| +-- Trapezoideus_lenya **
|
| +-- Yaukthwa_nesemanni **
| +-----------+
| | +-- Yaukthwa_elongatula **
| |
| | +----- Lens_misellus **
+--+ +-----+
| | | +-- Lens_contradens **
| | +--+
| | +-- Lens_micropterus **
+--+
| +----- Lens_pallegoixi **
| +--+
| | | +-- Lens_novoselovi **
| | +--+
+--+ +-- Lens_comptus **
|
| +----- Lens_maenamensis **
+--+
| +-- Lens_eximius **
+--+
+-- Lens_rolfbrandti **
Jeratthitikul et al., 2022 [Publ 4260], fig. S1, 1
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 76 |
76 |
73 |
2 |
0 |
0 |
0 |
65.0 |
+-------------------- Cumberlandia_monodonta **
|
+-----------------+ +----------------- Gibbosula_laosensis **
| | |
| | | +-- Pseudunio_auricularius **
| +--+ +-----------+
| | | +-- Pseudunio_marocanus **
| | |
| +--+ +-- Margaritifera_margaritifera **
| | +--------+
| | | +-- Margaritifera_dahurica **
| +--+
| | +-------- Margaritifera_falcata **
| +--+
| | +----- Margaritifera_middendorffi **
| +--+
| | +-- Margaritifera_laevis **
| +--+
| +-- Margaritifera_marrianae **
|
| +-- Indopseudodon_indawgyiensis **
| +--------------------+
| | +-- Indopseudodon_kayinensis **
| |
--+ | +-- Nyeinchanconcha_nyeinchani **
| | +-----+
| +-----------+ | +-- Sundadontina_cumingii **
| | | +-----------+
| | | | | +----- Thaiconcha_callifera **
| | | | +--+
| | | | | +-- Bineurus_panhai **
| | | | +--+
| | +--+ +-- Bineurus_mouhotii **
| | |
| | | +-- Namkongnaia_inkhavilayi **
| | | +--------------+
| | | | +-- Pseudodon_vondembuschianus **
| | | |
| | +--+ +-- Pilsbryoconcha_exilis **
| | | +-----------+
| | | | +-- Pilsbryoconcha_schomburgki **
| | | |
| | +--+ +-- Pilsbryoconcha_carinifera **
| | | +--------+
| | | | +-- Pilsbryoconcha_linguaeformis **
| | +--+
+--+ | +-------- Pilsbryoconcha_hoikaab **
| +--+
| | +----- Pilsbryoconcha_mekongiana **
| +--+
| | +-- Pilsbryoconcha_acuta **
| +--+
| +-- Pilsbryoconcha_kittitati **
|
| +----- Lens_contradens **
| +--+
| | | +-- Trapezoideus_foliaceus **
| | +--+
| +-----------------------+ +-- Physunio_superbus **
| | |
| | | +-- Ensidens_ingallsianus **
| | | |
| | +-----+-- Hyriopsis_bialata **
| | |
| | +-- Rectidens_sumatrensis **
| |
| | +----- Sinosolenaia_carinata **
+--+ +-----------------------+
| | | +-- Leguminaia_wheatleyi **
| | +--+
| | +-- Gonidea_angulata **
| |
| | +-- Chamberlainia_somsakpanhai **
| | +-----+
| | | +-- Sinohyriopsis_cumingii **
| | +-----------------+
| | | | +----- Lamprotula_leaii **
+--+ | +--+
| | | +-- Potomida_littoralis **
| | +--+
| | +-- Pronodularia_japanensis **
| |
| | +-------- Amblema_plicata **
| | +--+
| | | | +----- Ortmanniana_ligamentina **
| | | +--+
| | | | +-- Cambarunio_iris **
| | | +--+
+--+ +-----------+ +-- Lampsilis_cardium **
| | |
| | | +-- Quadrula_quadrula **
| | | +-----+
| | | | +-- Quadrula_verrucosa **
| | +--+
| | | +----- Elliptio_complanata **
| | +--+
| | | +-- Pleurobema_sintoxia **
| | +--+
| | +-- Eurynia_dilatata **
| |
| | +----- Unio_tumidus **
+--+ +-----------+
| | | +-- Unio_pictorum **
| | +--+
| +--+ +-- Unio_crassus **
| | |
| | | +-------------- Aculamprotula_tortuosa **
| | | |
| | +--+ +-- Nodularia_douglasiae **
| | | +--------+
| | | | +-- Nodularia_nuxpersicae **
| | +--+
| | | +-------- Schistodesmus_lampreyanus **
| | +--+
| | | +----- Cuneopsis_heudei **
+--+ +--+
| | +-- Tchangsinaia_piscicula **
| +--+
| +-- Cuneopsis_rufescens **
|
| +-- Lanceolaria_grayii **
| +--------------+
| | +-- Lanceolaria_gladiola **
| |
| | +----------- Cristaria_plicata **
| | +--+
+--+ | | +-------- Pletholophus_tenuis **
| | +--+
| | | +----- Sinanodonta_lucida **
| | +--+
| | | +-- Sinanodonta_lauta **
| | +--+
+--+ +-- Sinanodonta_jourdyi **
|
| +-------- Pseudanodonta_complanata **
| +--+
| | | +----- Anodonta_nuttalliana **
| | +--+
| | | +-- Anodonta_anatina **
| | +--+
+--+ +-- Anodonta_cygnea **
|
| +-- Pyganodon_grandis **
| +-----+
| | +-- Simpsonaias_ambigua **
+--+
| +----- Strophitus_undulatus **
+--+
| +-- Alasmidonta_marginata **
+--+
+-- Lasmigona_costata **
Jeratthitikul et al., 2022 [Publ 4260], fig. S2, 2
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 76 |
76 |
74 |
2 |
0 |
0 |
0 |
74.0 |
+----------------- Cumberlandia_monodonta **
|
| +----- Gibbosula_laosensis **
+--------------------+ +--------+
| | | | +-- Pseudunio_auricularius **
| | | +--+
| | | +-- Pseudunio_marocanus **
| +--+
| | +-- Margaritifera_dahurica **
| | +--------+
| | | +-- Margaritifera_margaritifera **
| +--+
| | +-------- Margaritifera_falcata **
| +--+
| | +----- Margaritifera_marrianae **
| +--+
| | +-- Margaritifera_middendorffi **
| +--+
| +-- Margaritifera_laevis **
|
| +-- Indopseudodon_indawgyiensis **
| +--------------------+
| | +-- Indopseudodon_kayinensis **
| |
--+ | +-- Nyeinchanconcha_nyeinchani **
| | +-----+
| +-----------+ | +-- Sundadontina_cumingii **
| | | +-----------+
| | | | | +----- Thaiconcha_callifera **
| | | | +--+
| | | | | +-- Bineurus_panhai **
| | | | +--+
| | +--+ +-- Bineurus_mouhotii **
| | |
| | | +-- Namkongnaia_inkhavilayi **
| | | +--------------+
| | | | +-- Pseudodon_vondembuschianus **
| | | |
| | +--+ +-- Pilsbryoconcha_exilis **
| | | +-----------+
| | | | +-- Pilsbryoconcha_schomburgki **
| | | |
| | +--+ +-- Pilsbryoconcha_carinifera **
| | | +--------+
| | | | +-- Pilsbryoconcha_linguaeformis **
| | +--+
+--+ | +-------- Pilsbryoconcha_hoikaab **
| +--+
| | +----- Pilsbryoconcha_mekongiana **
| +--+
| | +-- Pilsbryoconcha_acuta **
| +--+
| +-- Pilsbryoconcha_kittitati **
|
| +----- Ensidens_ingallsianus **
| +--+
| | | +-- Rectidens_sumatrensis **
| | +--+
| +-----------------------+ +-- Hyriopsis_bialata **
| | |
| | | +----- Lens_contradens **
| | +--+
| | | +-- Trapezoideus_foliaceus **
| | +--+
| | +-- Physunio_superbus **
| |
| | +----- Sinosolenaia_carinata **
+--+ +-----------------------+
| | | +-- Gonidea_angulata **
| | +--+
| | +-- Leguminaia_wheatleyi **
| |
| | +-- Chamberlainia_somsakpanhai **
| | +-----+
| | | +-- Sinohyriopsis_cumingii **
| | +-----------------+
| | | | +----- Potomida_littoralis **
+--+ | +--+
| | | +-- Lamprotula_leaii **
| | +--+
| | +-- Pronodularia_japanensis **
| |
| | +-------- Amblema_plicata **
| | +--+
| | | | +----- Ortmanniana_ligamentina **
| | | +--+
| | | | +-- Lampsilis_cardium **
| | | +--+
+--+ +-----------+ +-- Cambarunio_iris **
| | |
| | | +-- Quadrula_quadrula **
| | | +-----+
| | | | +-- Quadrula_verrucosa **
| | +--+
| | | +----- Pleurobema_sintoxia **
| | +--+
| | | +-- Eurynia_dilatata **
| | +--+
| | +-- Elliptio_complanata **
| |
| | +----- Unio_tumidus **
+--+ +-----------+
| | | +-- Unio_pictorum **
| | +--+
| +--+ +-- Unio_crassus **
| | |
| | | +-------------- Aculamprotula_tortuosa **
| | | |
| | +--+ +-- Nodularia_douglasiae **
| | | +--------+
| | | | +-- Nodularia_nuxpersicae **
| | +--+
| | | +-------- Schistodesmus_lampreyanus **
| | +--+
| | | +----- Cuneopsis_heudei **
+--+ +--+
| | +-- Tchangsinaia_piscicula **
| +--+
| +-- Cuneopsis_rufescens **
|
| +-- Lanceolaria_grayii **
| +--------------+
| | +-- Lanceolaria_gladiola **
| |
| | +----------- Cristaria_plicata **
| | +--+
+--+ | | +-------- Pletholophus_tenuis **
| | +--+
| | | +----- Sinanodonta_lucida **
| | +--+
| | | +-- Sinanodonta_jourdyi **
| | +--+
+--+ +-- Sinanodonta_lauta **
|
| +-------- Pseudanodonta_complanata **
| +--+
| | | +----- Anodonta_anatina **
| | +--+
| | | +-- Anodonta_nuttalliana **
| | +--+
+--+ +-- Anodonta_cygnea **
|
| +-- Alasmidonta_marginata **
| +-----+
| | +-- Lasmigona_costata **
+--+
| +----- Pyganodon_grandis **
+--+
| +-- Strophitus_undulatus **
+--+
+-- Simpsonaias_ambigua **
Huang et al., 2019 [Publ 3843], fig. S1
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 75 |
75 |
57 |
2 |
0 |
0 |
0 |
99.0 |
+-- Gibbosula_rochechouartii **
+-----------------------------------+
| +-- Margaritifera_dahurica **
|
| +----- Sinohyriopsis_cumingii[2] **
| +--------------+
| | | +-- Sinohyriopsis_cumingii[1] **
| | +--+
| | +-- Sinohyriopsis_schlegelii **
| |
| | +----- Parvasolenaia_rivularis **
| +--------------+ +-----------+
| | | | | +-- Parvasolenaia_triangularis **
| | | | +--+
| | | | +-- Ptychorhynchus_pfisteri **
| | | |
| | | | +-------- Sinosolenaia_recognita **
| | +--+ |
--+ | | +-----+ +----- Sinosolenaia_oleivora[2] **
| | | | | |
| | | | +--+ +-- Sinosolenaia_oleivora[1] **
| | | | | |
| | | | +--+-- Sinosolenaia_iridinea **
| | +--+ |
| | | +-- Sinosolenaia_oleivora[3] **
| | |
| | | +----------- Sinosolenaia_carinata **
| | | |
| | +--+ +-------- Lamprotula_leaii **
| | | |
| | +--+ +----- Lamprotula_cornuumlunae[1] **
| | | |
| | | +----- Lamprotula_cornuumlunae[3] **
| | +--+
| | +----- Lamprotula_cornuumlunae[4] **
+--+ |
| | +-- Lamprotula_cornuumlunae[2] **
| +--+
| +-- Lamprotula_caveata **
|
| +-------------- Anemina_arcaeformis[2] **
| |
| +-----------------+ +----------- Anemina_arcaeformis[5] **
| | | |
| | | | +-------- Anemina_arcaeformis[3] **
| | +--+ |
| | | +-------- Anemina_arcaeformis[7] **
| | | |
| | +--+ +----- Anemina_arcaeformis[4] **
| | | |
| | | +----- Anemina_arcaeformis[6] **
| | | |
| | +--+----- Anemina_arcaeformis[8] **
| | |
| | +----- Anemina_arcaeformis[9] **
| | |
| | | +-- Anemina_arcaeformis[1] **
| | +--+
| | +-- Anemina_euscaphys **
+--+
| +-- Sinanodonta_angula **
| +-----------+
| | +-- Sinanodonta_lucida **
| +--------------+
| | | +----------- Sinanodonta_woodiana[2] **
| | +--+
| | | +-------- Sinanodonta_jourdyi **
| | +--+
| | | +----- Sinanodonta_elliptica **
| | +--+
| | | +-- Sinanodonta_woodiana[1] **
| | +--+
| | +-- Sinanodonta_lauta **
| |
| | +-------------- Acuticosta_ovata[1] **
| | |
| | | +----------- Acuticosta_chinensis[3] **
+--+ +-----------+ |
| | | +----------- Acuticosta_chinensis[4] **
| | | |
| | +--+----------- Acuticosta_chinensis[6] **
| | |
| | | +-- Acuticosta_chinensis[1] **
| | | +-----+
| | | | +-- Acuticosta_ovata[2] **
| | +--+
| | | +----- Acuticosta_chinensis[5] **
| | | |
| | +--+----- Acuticosta_chinensis[7] **
| | |
| | | +-- Acuticosta_chinensis[2] **
+--+ +--+
| +-- Acuticosta_ovata[3] **
|
| +-- Lepidodesma_languilati **
| +--------------------+
| | +-- Cristaria_plicata **
| |
| | +----------- Lanceolaria_gladiola **
| | |
| | | +-------- Lanceolaria_grayii[5] **
| | +--------+ |
| | | | | +-- Lanceolaria_lanceolata **
| | | | +-----+
| | | | | +-- Lanceolaria_triformis **
+--+ | +--+
| | | +----- Lanceolaria_grayii[4] **
| | | |
| | | | +-- Lanceolaria_eucylindrica[1] **
| | | | |
| | +--+ +-- Lanceolaria_eucylindrica[2] **
| | | |
| | | +-- Lanceolaria_eucylindrica[3] **
| | | |
| | +--+-- Lanceolaria_eucylindrica[4] **
+--+ |
| +-- Lanceolaria_grayii[1] **
| |
| +-- Lanceolaria_grayii[2] **
| |
| +-- Lanceolaria_grayii[3] **
|
| +----------- Aculamprotula_tortuosa **
| +-----+
| | | +-------- Aculamprotula_tientsinensis **
| | +--+
| | | +----- Aculamprotula_fibrosa **
| | +--+
| | | +-- Aculamprotula_coreana **
+--+ +--+
| +-- Aculamprotula_scripta **
|
| +----- Nodularia_douglasiae[1] **
| +--------+
| | | +-- Nodularia_nuxpersicae **
| | +--+
+--+ +-- Nodularia_douglasiae[2] **
|
| +----------- Tchangsinaia_piscicula **
| |
+--+ +-- Schistodesmus_spinosus **
| +-----+
| | +-- Schistodesmus_lampreyanus **
+--+
| +----- Cuneopsis_rufescens **
+--+
| +-- Cuneopsis_heudei **
+--+
+-- Cuneopsis_celtiformis **
Campbell & Lydeard, 2012 [Publ 3309], fig. 1-2
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 74 |
74 |
64 |
1 |
0 |
0 |
0 |
142.0 |
+-- Uniomerus_declivis **
+-----------------+
| +-- Uniomerus_tetralasmus **
+--------------------+
| | +----------------- Theliderma_metanevra **
| | |
| +--+ +-------------- Quadrula_verrucosa **
| | |
| | | +-- Quadrula_quadrula **
| +--+ +--------+
| | | +-- Quadrula_nobilis **
| | |
| +--+ +-- Pustulosa_infucata **
| | +-----+
| | | +-- Pustulosa_kleiniana **
| +--+
| | +----- Cyclonaias_tuberculata **
| +--+
| | +-- Pustulosa_pustulosa **
| +--+
| +-- Pustulosa_succissa **
|
| +----- Plectomerus_dombeyanus **
| +-----------------+
| | | +-- Reginaia_ebenus **
| | +--+
--+ | +-- Reginaia_rotulata **
| |
| | +----- Popenaias_popeii **
| +--------------+ +--------------+
| | | | | +-- Amblema_elliottii **
| | | | +--+
| | | | +-- Amblema_plicata **
| | | |
| | | | +----------------- Potamilus_alatus **
| | +--+ |
| | | +----------------- Potamilus_capax **
| | | |
| | | +----------------- Glebula_rotundata **
| | | |
| | | | +-- Toxolasma_lividum **
| | | +--------------+
| | +--+ +-- Paetulunio_fabalis **
| | |
| | | +-- Dromus_dromas **
| | +--------------+
| | | +-- Lemiox_rimosus **
| | |
| | | +-- Lampsilis_ovata **
+--+ | +-----------+
| | | +-- Obliquaria_reflexa **
| +--+
| | +----------- Lampsilis_virescens **
| | |
| +--+ +-- Epioblasma_rangiana **
| | +-----+
| | | +-- Obovaria_arkansasensis **
| +--+
| | +----- Obovaria_olivaria **
| +--+
| | +-- Venustaconcha_ellipsiformis **
| +--+
| +-- Venustaconcha_pleasii **
|
| +----------------------------------- Elliptoideus_sloatianus **
| |
| | +-------------------------------- Pleurobema_stabile **
| | |
| | | +-- Pleurobema_clava **
| | | +-----------+
+--+ | | +-- Pleurobema_oviforme **
| | |
| | +--------------+ +----------- Pleurobema_pyriforme **
| | | | |
| | | | | +-- Pleurobema_athearni **
| | | | | +--+
+--+ | +--+ | +-- Pleurobema_georgianum **
| | | +--+
| | | | | +-- Pleurobema_decisum **
| | | | +--+
| | | | +-- Pleurobema_hanleyianum **
| | +--+
| | | +----- Pleurobema_beadleianum **
| | | |
| | | +----- Pleurobema_strodeanum **
| | +--+
| | +----- Pleurobema_rubellum **
+--+ |
| | +-- Pleurobema_perovatum **
| +--+
| +-- Pleurobema_taitianum **
|
| +-------- Pleurobema_riddellii **
| +-----------------+
| | | +----- Pleurobema_cordatum **
| | +--+
| | | +-- Pleurobema_rubrum **
| | +--+
| | +-- Pleurobema_sintoxia **
| |
| | +----------------- Fusconaia_subrotunda **
| | |
+--+ +-----+ +-- Fusconaia_cor **
| | | +-----------+
| | | | +-- Fusconaia_cuneolus **
| | +--+
| | | +----------- Fusconaia_ozarkensis **
| | | |
| | +--+ +-- Fusconaia_escambia **
| | | +-----+
| | | | +-- Fusconaia_burkei **
| | +--+
+--+ | +----- Fusconaia_masoni **
| +--+
| | +-- Fusconaia_flava **
| +--+
| +-- Fusconaia_cerina **
|
| +----- Pleuronaia_gibber **
| +--------------+
| | | +-- Pleuronaia_barnesiana **
| | +--+
| | +-- Pleuronaia_dolabelloides **
+--+
| +-- Eurynia_dilatata **
| +--------------+
| | +-- Parvaspina_collina **
| |
+--+ +-- Hemistena_lata **
| +-----------+
| | +-- Plethobasus_cyphyus **
| |
+--+ +----------- Elliptio_shepardiana **
| |
| | +-------- Elliptio_complanata **
+--+ |
| | +----- Elliptio_purpurella **
| +--+
| | | +-- Elliptio_arca **
+--+ +--+
| +-- Elliptio_arctata **
|
| +-- Elliptio_mcmichaeli **
| |
+-----+-- Elliptio_pullata **
|
+-- Elliptio_crassidens **
Campbell & Lydeard, 2012 [Publ 3309], fig. 3
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 74 |
74 |
68 |
1 |
0 |
0 |
0 |
141.0 |
+-- Uniomerus_declivis **
+--------------+
| +-- Uniomerus_tetralasmus **
|
+--------------------+ +-- Quadrula_verrucosa **
| | +-----------+
| | | +-- Quadrula_nobilis **
| | |
| +--+ +----------- Quadrula_quadrula **
| | |
| | +----------- Theliderma_metanevra **
| +--+
| | +-- Pustulosa_infucata **
| | +-----+
| | | +-- Pustulosa_kleiniana **
| +--+
| | +----- Cyclonaias_tuberculata **
| +--+
| | +-- Pustulosa_pustulosa **
| +--+
| +-- Pustulosa_succissa **
|
| +-- Amblema_elliottii **
| +--+
| | +-- Amblema_plicata **
| +--------------------------+
| | | +-- Reginaia_ebenus **
--+ | +--+
| | +-- Reginaia_rotulata **
| +--+
| | | +----------------------------- Popenaias_popeii **
| | | |
| | | | +-- Glebula_rotundata **
| | +--+ +-----------------------+
| | | | +-- Paetulunio_fabalis **
| | | |
| | +--+ +-- Toxolasma_lividum **
| | | +--------------------+
| | | | +-- Plectomerus_dombeyanus **
| | | |
| | +--+ +-------------------- Obliquaria_reflexa **
| | | |
| | | | +-- Dromus_dromas **
| | | | +--+
| | +--+ | +-- Lemiox_rimosus **
| | | +-----------+
| | | | | +-- Potamilus_alatus **
| | | | +--+
| | +--+ +-- Potamilus_capax **
| | |
| | | +-------------- Lampsilis_ovata **
+--+ | |
| +--+ +----------- Lampsilis_virescens **
| | |
| +--+ +-- Epioblasma_rangiana **
| | +-----+
| | | +-- Obovaria_arkansasensis **
| +--+
| | +----- Obovaria_olivaria **
| +--+
| | +-- Venustaconcha_ellipsiformis **
| +--+
| +-- Venustaconcha_pleasii **
|
| +-------------------------------- Elliptoideus_sloatianus **
| |
| | +-- Eurynia_dilatata **
| | +--------------------------+
| | | +-- Parvaspina_collina **
| | |
| | | +-- Hemistena_lata **
| | +--------------------------+
| | | +-- Plethobasus_cyphyus **
+--+ |
| | +----------- Elliptio_shepardiana **
| | |
| | | +----- Elliptio_crassidens **
| +-----------------+ +--+
| | | | | +-- Elliptio_mcmichaeli **
| | | | +--+
| | +--+ +-- Elliptio_pullata **
| | |
| | | +----- Elliptio_complanata **
+--+ | |
| +--+ +-- Elliptio_purpurella **
| | |
| +--+-- Elliptio_arca **
| |
| +-- Elliptio_arctata **
|
| +----- Pleuronaia_gibber **
| +--------------------+
| | | +-- Pleuronaia_barnesiana **
| | +--+
| | +-- Pleuronaia_dolabelloides **
| |
| | +-------------- Fusconaia_ozarkensis **
| | |
| | | +----- Fusconaia_subrotunda **
| | +--------+ +-----+
+--+ | | | | +-- Fusconaia_cor **
| | | | +--+
| | | | +-- Fusconaia_cuneolus **
| | +--+
| | | +-- Fusconaia_escambia **
| | | +-----+
| | | | +-- Fusconaia_burkei **
| | +--+
| | | +----- Fusconaia_masoni **
+--+ +--+
| | +-- Fusconaia_cerina **
| +--+
| +-- Fusconaia_flava **
|
| +-------- Pleurobema_riddellii **
| +-----------+
| | | +----- Pleurobema_cordatum **
| | +--+
| | | +-- Pleurobema_rubrum **
| | +--+
+--+ +-- Pleurobema_sintoxia **
|
| +----------------- Pleurobema_stabile **
| |
| | +----------- Pleurobema_strodeanum **
| | +--+
+--+ | | +-------- Pleurobema_rubellum **
| | +--+
| | | +----- Pleurobema_beadleianum **
| | +--+
| | | +-- Pleurobema_perovatum **
+--+ +--+
| +-- Pleurobema_taitianum **
|
| +-- Pleurobema_clava **
| +--------+
| | +-- Pleurobema_oviforme **
+--+
| +-------- Pleurobema_pyriforme **
| |
+--+ +-- Pleurobema_athearni **
| +--+
| | +-- Pleurobema_georgianum **
+--+
| +-- Pleurobema_decisum **
+--+
+-- Pleurobema_hanleyianum **
Wu et al., 2022 [Publ 4238], fig. 3
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 71 |
71 |
69 |
2 |
3 |
3 |
2 |
91.0 |
+-- Mutela_dubia
+--------------------------------------------+
| +-- Anodontites_trapesialis
|
| +-------------------------------------------- Echyridella_menziesii
| |
| | +-------------- Gibbosula_rochechouartii **
| | +--------------------------+
--+ | | | +----------- Cumberlandia_monodonta **
| | | +--+
| | | | +-------- Pseudunio_marocanus **
| | | +--+
| | | | +----- Margaritifera_falcata **
| | | +--+
| | | | +-- Margaritifera_dahurica **
+--+ | +--+
| | +-- Margaritifera_margaritifera **
| |
| | +-- Uniomerus_tetralasmus **
| | +--------------------------+
| | | +-- Quadrula_quadrula **
| | |
| | +--------+ +-- Pleurobema_oviforme **
| | | | +-----------------------+
| | | | | +-- Elliptio_complanata **
| | | +--+
+--+ | | +----------------------- Amblema_plicata **
| | | |
| | +--+ +-------------------- Popenaias_popeii **
| | | |
| | +--+ +----------------- Toxolasma_parvum **
| | | |
| | +--+ +-- Potamilus_alatus **
| | | +-----------+
| | | | +-- Potamilus_leptodon **
| | +--+
| | | +----------- Leaunio_lienosus **
| | | |
| | +--+ +-- Lampsilis_cardium **
| | | +-----+
| | | | +-- Lampsilis_ornata **
| | +--+
| | | +----- Venustaconcha_ellipsiformis **
| | +--+
| | | +-- Lampsilis_siliquoidea **
| | +--+
+--+ +-- Lampsilis_powellii **
|
| +----- Chamberlainia_somsakpanhai **
| +-----------+
| | | +-- Sinohyriopsis_schlegelii **
| | +--+
| | +-- Sinohyriopsis_cumingii **
| |
| +-----------------+ +-------- Potomida_littoralis **
| | | +-----+
| | | | | +----- Pronodularia_japanensis **
| | | | +--+
| | | | | +-- Lamprotula_caveata **
| | | | +--+
| | +--+ +-- Lamprotula_leaii **
| | |
| | | +-- Pilsbryoconcha_exilis **
| | | +--------+
| | | | +-- Pseudodon_vondembuschianus **
| | | |
| | +--+ +-- Ptychorhynchus_pfisteri **
| | | +-----+
| | | | +-- Parvasolenaia_rivularis **
| | +--+
| | | +----- Microcondylaea_bonellii **
+--+ +--+
| | +-- Sinosolenaia_carinata **
| +--+
| +-- Sinosolenaia_recognita **
|
| +-------------------------------- Lepidodesma_languilati **
| |
| | +-------- Acuticosta_chinensis **
| | +--------+
| | | | +----- Lanceolaria_gladiola **
| | | +--+
| | | | +-- Lanceolaria_grayii **
| | | +--+
| | | +-- Lanceolaria_lanceolata **
| | |
| | +-----------+ +-------- Cristaria_plicata **
| | | | |
+--+ | | +-----+ +-- Sinanodonta_woodiana **
| | | | | +--+
| | | | | | +-- Sinanodonta_lucida **
| | | | +--+
| | | | | +-- Anemina_arcaeformis **
| | +--+ +--+
| | | +-- Anemina_euscaphys **
| | |
| | | +-- Anodonta_cygnea **
| | | +--------+
| | | | +-- Anodonta_anatina **
| | +--+
| | | +-------- Platynaias_compressa **
+--+ +--+
| | +----- Pyganodon_grandis **
| +--+
| | +-- Utterbackia_imbecillis **
| +--+
| +-- Utterbackia_peninsularis **
|
| +-------- Aculamprotula_tortuosa **
| |
| +-----------------+ +-- Aculamprotula_tientsinensis **
| | | +--+
| | | | +-- Aculamprotula_coreana **
| | +--+
| | | +-- Aculamprotula_polysticta **
| | +--+
| | +-- Aculamprotula_scripta **
+--+
| +-------- Unio_tumidus **
| +--------------+
| | | +----- Unio_crassus **
| | +--+
| | | +-- Unio_pictorum **
| | +--+
+--+ +-- Unio_delphinus **
|
| +-- Pseudocuneopsis_capitata **
| +-----------------+
| | +-- Pseudocuneopsis_sichuanensis **
| |
+--+ +-- Nodularia_douglasiae **
| +--------------+
| | +-- Nodularia_breviconcha **
| |
+--+ +-- Schistodesmus_lampreyanus **
| +-----------+
| | +-- Schistodesmus_spinosus **
+--+
| +----------- Tchangsinaia_piscicula **
+--+
| +-------- Cuneopsis_rufescens **
+--+
| +----- Cuneopsis_celtiformis **
+--+
| +-- Cuneopsis_heudei **
+--+
+-- Cuneopsis_demangei **
Sano et al., 2020 [Publ 4138], fig. 2
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 62 |
62 |
60 |
2 |
11 |
9 |
2 |
103.0 |
+----- Triplodon_corrugatus
+--------------------------------------------+
| | +-- Hyridella_australis
| +--+
| +-- Velesunio_ambiguus
|
| +----------- Mutela_hargeri
| |
| +-----------------------------------+ +-- Aspatharia_pfeifferiana
| | | +-----+
| | | | +-- Chambardia_wahlbergi
--+ | +--+
| | | +----- Etheria_elliptica
| | +--+
| | | +-- Tamsiella_schroeteriana
| | +--+
| | +-- Lamproscapha_ensiformis
| |
| | +-------------------- Cumberlandia_monodonta **
| | |
| | | +-- Gibbosula_rochechouartii **
| | | +--+
+--+ +-----------------------+ | +-- Gibbosula_laosensis **
| | | +-----------+
| | | | | +-- Pseudunio_auricularius **
| | | | +--+
| | | | +-- Pseudunio_marocanus **
| | +--+
| | | +-- Margaritifera_dahurica **
| | | +-----------+
| | | | +-- Margaritifera_margaritifera **
| | +--+
| | | +----------- Margaritifera_falcata **
| | +--+
| | | +-------- Margaritifera_middendorffi **
| | +--+
+--+ | +----- Margaritifera_laevis **
| +--+
| | +-- Margaritifera_hembeli **
| +--+
| +-- Margaritifera_marrianae **
|
| +----------------------------------------- Scabies_crispata **
| |
| | +----------------- Lampsilis_cardium **
| | |
| | | +----- Sinohyriopsis_cumingii[2] **
| | +--------------------+ +--------+
| | | | | | +-- Sinohyriopsis_schlegelii **
| | | | | +--+
| | | +--+ +-- Sinohyriopsis_cumingii[1] **
+--+ | |
| | | +----------- Inversidens_brandtii **
| | | |
| | +--+ +-- Ptychorhynchus_pfisteri **
| | | +-----+
| | | | +-- Obovalis_omiensis **
| | +--+
| | | +----- Lamprotula_leaii **
| | +--+
+--+ | +-- Pronodularia_japanensis **
| +--+
| +-- Potomida_littoralis **
|
| +----- Buldowskia_iwakawai **
| +-----------------------------+
| | | +-- Anemina_arcaeformis **
| | +--+
| | +-- Anemina_euscaphys **
| |
| | +-------------- Pletholophus_tenuis **
| | |
| | | +----- Beringiana_fukuharai **
| | +-----------------+ +-----+
+--+ | | | | +-- Beringiana_japonica **
| | | | +--+
| | +--+ +-- Beringiana_beringiana **
| | |
| | | +-------- Sinanodonta_lucida **
| | +--+
| | | +----- Sinanodonta_tumens **
| | +--+
| | | +-- Sinanodonta_lauta **
+--+ +--+
| +-- Sinanodonta_woodiana **
|
| +----------------------------- Cristaria_plicata **
| |
| | +-- Anodonta_cygnea **
| | +--------+
| | | +-- Anodonta_anatina **
| | |
| | +--------------+ +----- Alasmidonta_varicosa **
+--+ | | +--+
| | | | | +-- Prolasmidonta_heterodon **
| | | | +--+
| | +--+ +-- Platynaias_compressa **
| | |
| | | +----- Pyganodon_grandis **
| | +--+
+--+ | +-- Utterbackia_imbecillis **
| +--+
| +-- Utterbackia_peninsularis **
|
| +----- Lanceolaria_grayii[1] **
| +-----------------+
| | | +-- Lanceolaria_lanceolata **
| | +--+
| | +-- Lanceolaria_grayii[2] **
+--+
| +-------------------- Lepidodesma_languilati **
| |
| | +----- Aculamprotula_tortuosa **
| | +-----------+
+--+ | | +-- Aculamprotula_tientsinensis **
| | +--+
| | +-- Aculamprotula_coreana **
| |
+--+ +----- Unio_crassus **
| +--------+
| | | +-- Unio_delphinus **
| | +--+
| | +-- Unio_pictorum **
| |
+--+ +-------- Inversiunio_jokohamensis **
| +--+
| | | +----- Inversiunio_yanagawensis[2] **
| | +--+
| | | +-- Inversiunio_reinianus **
| | +--+
+--+ +-- Inversiunio_yanagawensis[1] **
|
| +-- Tchangsinaia_piscicula **
| +-----+
| | +-- Schistodesmus_lampreyanus **
+--+
| +----- Nodularia_douglasiae[2] **
+--+
| +-- Nodularia_nipponensis **
+--+
+-- Nodularia_douglasiae[1] **
Sano et al., 2020 [Publ 4138], fig. 3
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 61 |
61 |
59 |
2 |
11 |
9 |
2 |
104.0 |
+----- Triplodon_corrugatus
+--------------------------------------------+
| | +-- Hyridella_australis
| +--+
| +-- Velesunio_ambiguus
|
| +----------- Mutela_hargeri
| |
| +-----------------------------------+ +-- Aspatharia_pfeifferiana
| | | +-----+
| | | | +-- Chambardia_wahlbergi
--+ | +--+
| | | +----- Etheria_elliptica
| | +--+
| | | +-- Tamsiella_schroeteriana
| | +--+
| | +-- Lamproscapha_ensiformis
| |
| | +----------------------- Cumberlandia_monodonta **
| | |
| | | +-- Pseudunio_auricularius **
| | +--------------------+ +-----------------+
+--+ | | | +-- Pseudunio_marocanus **
| | | |
| | +--+ +-- Gibbosula_laosensis **
| | | +--------------+
| | | | +-- Gibbosula_rochechouartii **
| | | |
| | +--+ +-- Margaritifera_margaritifera **
| | | +-----------+
| | | | +-- Margaritifera_dahurica **
| | +--+
| | | +----------- Margaritifera_falcata **
| | +--+
| | | +-------- Margaritifera_middendorffi **
| | +--+
+--+ | +----- Margaritifera_laevis **
| +--+
| | +-- Margaritifera_hembeli **
| +--+
| +-- Margaritifera_marrianae **
|
| +----------------------------------------- Scabies_crispata **
| |
| | +----- Sinohyriopsis_cumingii[2] **
| | +--------+
| | | | +-- Sinohyriopsis_schlegelii **
| | | +--+
| | | +-- Sinohyriopsis_cumingii[1] **
| | +-----------------------+
| | | | +----------- Lampsilis_cardium **
+--+ | | |
| | | | +----- Lamprotula_leaii **
| | +--+ +--+
| | | | | +-- Potomida_littoralis **
| | | | +--+
| | +--+ +-- Pronodularia_japanensis **
| | |
| | | +----- Inversidens_brandtii **
| | +--+
| | | +-- Obovalis_omiensis **
+--+ +--+
| +-- Ptychorhynchus_pfisteri **
|
| +----- Buldowskia_iwakawai **
| +-----------------------------+
| | | +-- Anemina_arcaeformis **
| | +--+
| | +-- Anemina_euscaphys **
| |
| | +-------- Pletholophus_tenuis **
| | +--+
| | | | +----- Beringiana_fukuharai **
| | | +--+
+--+ | | +-- Beringiana_japonica **
| +--------------------+ +--+
| | | +-- Beringiana_beringiana **
| | |
| | | +-------- Sinanodonta_lucida **
| | +--+
| | | +----- Sinanodonta_tumens **
| | +--+
| | | +-- Sinanodonta_woodiana **
| | +--+
+--+ +-- Sinanodonta_lauta **
|
| +----------------------------- Cristaria_plicata **
| |
| | +-- Anodonta_anatina **
| | +--------+
| | | +-- Anodonta_cygnea **
| | |
| | +--------------+ +----- Pyganodon_grandis **
+--+ | | +--+
| | | | | +-- Utterbackia_imbecillis **
| | | | +--+
| | +--+ +-- Utterbackia_peninsularis **
| | |
| | | +----- Alasmidonta_varicosa **
| | +--+
+--+ | +-- Platynaias_compressa **
| +--+
| +-- Prolasmidonta_heterodon **
|
| +----- Lanceolaria_grayii[1] **
| +-----------------+
| | | +-- Lanceolaria_lanceolata **
| | +--+
| | +-- Lanceolaria_grayii[2] **
+--+
| +-------------------- Lepidodesma_languilati **
| |
| | +----- Aculamprotula_tortuosa **
| | +-----------+
+--+ | | +-- Aculamprotula_tientsinensis **
| | +--+
| | +-- Aculamprotula_coreana **
| |
+--+ +----- Unio_crassus **
| +--------+
| | | +-- Unio_delphinus **
| | +--+
| | +-- Unio_pictorum **
| |
+--+ +-------- Inversiunio_jokohamensis **
| +--+
| | | +----- Inversiunio_yanagawensis[2] **
| | +--+
| | | +-- Inversiunio_reinianus **
| | +--+
+--+ +-- Inversiunio_yanagawensis[1] **
|
| +-- Nodularia_nipponensis **
| +--+
| | +-- Nodularia_douglasiae **
+-----+
| +-- Schistodesmus_lampreyanus **
+--+
+-- Tchangsinaia_piscicula **
Sano et al., 2020 [Publ 4138], fig. 4
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 61 |
61 |
59 |
2 |
11 |
9 |
2 |
102.0 |
+----- Triplodon_corrugatus
+-----------------------------------------------+
| | +-- Hyridella_australis
| +--+
| +-- Velesunio_ambiguus
|
| +----------- Mutela_hargeri
| |
| +--------------------------------------+ +-- Aspatharia_pfeifferiana
| | | +-----+
| | | | +-- Chambardia_wahlbergi
--+ | +--+
| | | +----- Etheria_elliptica
| | +--+
| | | +-- Tamsiella_schroeteriana
| | +--+
| | +-- Lamproscapha_ensiformis
| |
| | +-------------------- Cumberlandia_monodonta **
| | |
| | | +-- Pseudunio_auricularius **
| | +--------------------------+ +--------------+
+--+ | | | +-- Pseudunio_marocanus **
| | | |
| | +--+ +-- Gibbosula_rochechouartii **
| | | +-----------+
| | | | +-- Gibbosula_laosensis **
| | | |
| | +--+ +-- Margaritifera_margaritifera **
| | | +--------+
| | | | +-- Margaritifera_dahurica **
| | +--+
| | | +-------- Margaritifera_falcata **
| | | |
| | +--+ +-- Margaritifera_laevis **
| | | +--+
+--+ | | +-- Margaritifera_middendorffi **
| +--+
| | +-- Margaritifera_hembeli **
| +--+
| +-- Margaritifera_marrianae **
|
| +-------------------------------------------- Scabies_crispata **
| |
| | +----------------- Lampsilis_cardium **
| | |
| | | +----- Sinohyriopsis_cumingii[2] **
| | +-----------------------+ +--------+
| | | | | | +-- Sinohyriopsis_schlegelii **
| | | | | +--+
| | | +--+ +-- Sinohyriopsis_cumingii[1] **
+--+ | |
| | | +----------- Inversidens_brandtii **
| | | |
| | +--+ +-- Obovalis_omiensis **
| | | +-----+
| | | | +-- Ptychorhynchus_pfisteri **
| | +--+
| | | +----- Lamprotula_leaii **
| | +--+
+--+ | +-- Pronodularia_japanensis **
| +--+
| +-- Potomida_littoralis **
|
| +----- Buldowskia_iwakawai **
| +--------------------------------+
| | | +-- Anemina_arcaeformis **
| | +--+
| | +-- Anemina_euscaphys **
| |
| | +-------- Sinanodonta_lucida **
| | +--+
| | | | +----- Sinanodonta_tumens **
| | | +--+
+--+ | | +-- Sinanodonta_woodiana **
| +-----------------------+ +--+
| | | +-- Sinanodonta_lauta **
| | |
| | | +-------- Pletholophus_tenuis **
| | +--+
| | | +----- Beringiana_fukuharai **
| | +--+
| | | +-- Beringiana_japonica **
| | +--+
+--+ +-- Beringiana_beringiana **
|
| +-------------------------------- Cristaria_plicata **
| |
| | +-- Anodonta_cygnea **
| | +--------+
| | | +-- Anodonta_anatina **
| | |
| | +-----------------+ +----- Alasmidonta_varicosa **
+--+ | | +--+
| | | | | +-- Prolasmidonta_heterodon **
| | | | +--+
| | +--+ +-- Platynaias_compressa **
| | |
| | | +----- Pyganodon_grandis **
| | +--+
+--+ | +-- Utterbackia_imbecillis **
| +--+
| +-- Utterbackia_peninsularis **
|
| +-------------------------- Lepidodesma_languilati **
| |
| | +----- Lanceolaria_grayii[1] **
| | +-----------------+
+--+ | | +-- Lanceolaria_lanceolata **
| | +--+
| | +-- Lanceolaria_grayii[2] **
| |
+--+ +----- Aculamprotula_tortuosa **
| +--------------+
| | | +-- Aculamprotula_tientsinensis **
| | +--+
| | +-- Aculamprotula_coreana **
| |
+--+ +----- Unio_crassus **
| +-----------+
| | | +-- Unio_delphinus **
| | +--+
| | +-- Unio_pictorum **
+--+
| +-- Nodularia_douglasiae **
| +-----------+
| | +-- Nodularia_nipponensis **
| |
+--+ +-- Schistodesmus_lampreyanus **
| +--------+
| | +-- Tchangsinaia_piscicula **
+--+
| +-------- Inversiunio_jokohamensis **
+--+
| +----- Inversiunio_yanagawensis[2] **
+--+
| +-- Inversiunio_reinianus **
+--+
+-- Inversiunio_yanagawensis[1] **
Bolotov et al., 2022 [Publ 4252], fig. 2
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 61 |
61 |
55 |
1 |
0 |
0 |
0 |
127.0 |
+-- Pseudodon_lenyanensis **
|
+-- Nyeinchanconcha_nyeinchani **
+--------------------------------------------+
| +-- Indopseudodon_kayinensis **
| |
| +-- Sundadontina_tanintharyiensis **
|
| +----------- Trapezidens_mogaungensis **
| |
| +-----+ +-------- Trapezidens_exolescens **
| | | |
| | +--+ +----- Trapezidens_scutum **
| | | |
| | +--+ +-- Trapezidens_angustior **
| | | |
| +--------------------------+ +--+-- Trapezidens_dolichorhynchus **
| | | |
--+ | | +-- Trapezidens_yeti **
| | |
| | | +-------------- Lamellidens_marginalis **
| | | |
| | | | +----------- Lamellidens_ferrugineus **
| | +--+ |
| | | | +-------- Lamellidens_generosus **
| | | +--+
| | | | | +----- Lamellidens_mainwaringi **
| | +--+ +--+
| | | | +-- Lamellidens_lamellatus **
| | | +--+
| | | +-- Lamellidens_jenkinsianus **
| | |
| | | +-------- Lamellidens_nongyangensis **
+--+ +--+
| | +----- Lamellidens_savadiensis **
| +--+
| | +-- Lamellidens_corrianus **
| +--+
| +-- Lamellidens_chindwinensis **
|
| +----------- Parreysia_keralaensis **
| +-----------------------------+
| | | +-------- Parreysia_corrugata **
| | +--+
| | | +----- Parreysia_rakhinensis **
| | +--+
| | | +-- Parreysia_nagpoorensis **
| | +--+
| | +-- Parreysia_plagiosoma **
| |
| | +-- Leoparreysia_burmana **
+--+ +-----+
| | +-- Leoparreysia_whitteni **
| +-----------------------------+
| | | +----- Leoparreysia_olivacea **
| | +--+
| | | +-- Leoparreysia_canefrii **
| | +--+
| | +-- Leoparreysia_tavoyensis **
| |
| | +-------- Nitia_teretiuscula **
+--+ +--------------------------+
| | | +----- Coelatura_aegyptiaca **
| | +--+
| | | +-- Nyassunio_nyassaensis **
| | +--+
| | +-- Coelatura_kunenensis **
| |
| | +----------------- Indonaia_cylindrica **
| | |
+--+ +--------------+ +-------------- Indonaia_gratiosa **
| | | |
| | +--+ +----------- Indonaia_andersoniana **
| | | |
| | +--+ +-- Indonaia_caerulea **
| | | +-----+
| | | | +-- Indonaia_subclathrata **
| | +--+
| | | +----- Indonaia_rugosa **
+--+ +--+
| | +-- Indonaia_bonneaudii **
| +--+
| +-- Indonaia_shurtleffiana **
|
| +----------------------------- Indochinella_pugio **
| |
| | +----------- Radiatula_crispisulcata **
| | |
| | | +----- Radiatula_chaudhurii **
+--+ +--------------+ +--+
| | | | | +-- Radiatula_myitkyinae **
| | | | +--+
| | +--+ +-- Radiatula_kachinensis **
| | |
| | | +----- Radiatula_chindwinensis **
+--+ +--+
| | +-- Radiatula_myitthanensis **
| +--+
| +-- Radiatula_mouhoti **
|
| +----------------------- Scabiellus_songkramensis **
| |
+--+ +-------------------- Harmandia_somboriensis **
| |
+--+ +----------------- Unionetta_fabagina **
| |
+--+ +-- Scabies_crispata **
| +-----------+
| | +-- Scabies_mandarinus **
+--+
| +----------- Scabies_nucleus **
| |
+--+ +-- Scabies_humilis **
| +-----+
| | +-- Scabies_pilata **
+--+
| +----- Scabies_phaselus **
+--+
| +-- Scabies_anceps **
+--+
+-- Scabies_scobinatus **
Bolotov et al., 2022 [Publ 4252], fig. S1
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 61 |
61 |
49 |
1 |
0 |
0 |
0 |
127.0 |
+-- Pseudodon_lenyanensis **
|
+-- Nyeinchanconcha_nyeinchani **
+-----------------------------------------------+
| +-- Indopseudodon_kayinensis **
| |
| +-- Sundadontina_tanintharyiensis **
|
| +-- Parreysia_keralaensis **
| |
| +-- Parreysia_corrugata **
| |
| +--------------------------------------------+-- Parreysia_plagiosoma **
| | |
| | +-- Parreysia_rakhinensis **
--+ | |
| | +-- Parreysia_nagpoorensis **
| |
| | +-------------- Trapezidens_mogaungensis **
| | +--+
| | | | +----------- Trapezidens_scutum **
| | | +--+
| | | | +-------- Trapezidens_exolescens **
| | | +--+
| | | | +----- Trapezidens_yeti **
| | | +--+
| | | | +-- Trapezidens_dolichorhynchus **
| | +--------------------------+ +--+
+--+ | | +-- Trapezidens_angustior **
| | |
| | | +-------- Lamellidens_savadiensis **
| | | +-----+
| | | | | +----- Lamellidens_chindwinensis **
| | | | +--+
| | | | | +-- Lamellidens_corrianus **
| | | | +--+
| | +--+ +-- Lamellidens_nongyangensis **
| | |
| | | +-- Lamellidens_marginalis **
| | | +--------+
| | | | +-- Lamellidens_ferrugineus **
| | +--+
| | | +-------- Lamellidens_generosus **
| | +--+
| | | +----- Lamellidens_mainwaringi **
+--+ +--+
| | +-- Lamellidens_jenkinsianus **
| +--+
| +-- Lamellidens_lamellatus **
|
| +-- Leoparreysia_burmana **
| |
| +-- Leoparreysia_canefrii **
| |
| +--------------------------------------+-- Leoparreysia_olivacea **
| | |
| | +-- Leoparreysia_tavoyensis **
| | |
| | +-- Leoparreysia_whitteni **
| |
| | +-- Coelatura_aegyptiaca **
| | |
| | +-- Coelatura_kunenensis **
+--+ +-----------------------------------+
| | +-- Nitia_teretiuscula **
| | |
| | +-- Nyassunio_nyassaensis **
| |
| | +-- Indonaia_cylindrica **
| | +--------+
| | | +-- Indonaia_gratiosa **
| | |
+--+ +-----------------------+ +----- Indonaia_shurtleffiana **
| | | +--+
| | | | | +-- Indonaia_bonneaudii **
| | | | +--+
| | +--+ +-- Indonaia_rugosa **
| | |
| | | +----- Indonaia_andersoniana **
| | +--+
| | | +-- Indonaia_caerulea **
+--+ +--+
| +-- Indonaia_subclathrata **
|
| +-------------------------------- Indochinella_pugio **
| |
| | +----- Radiatula_chaudhurii **
| | +-----+
| | | | +-- Radiatula_kachinensis **
| | | +--+
+--+ +-----------------+ +-- Radiatula_myitkyinae **
| | |
| | | +-------- Radiatula_crispisulcata **
| | +--+
| | | +----- Radiatula_mouhoti **
| | +--+
+--+ | +-- Radiatula_myitthanensis **
| +--+
| +-- Radiatula_chindwinensis **
|
| +-------------------------- Scabiellus_songkramensis **
| |
+--+ +----------------------- Harmandia_somboriensis **
| |
+--+ +-------------------- Unionetta_fabagina **
| |
+--+ +----------------- Scabies_crispata **
| |
+--+ +-- Scabies_humilis **
| +-----------+
| | +-- Scabies_mandarinus **
+--+
| +----------- Scabies_nucleus **
+--+
| +-------- Scabies_pilata **
+--+
| +----- Scabies_anceps **
+--+
| +-- Scabies_phaselus **
+--+
+-- Scabies_scobinatus **
Bolotov et al., 2022 [Publ 4239], fig. 2
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 58 |
58 |
54 |
1 |
0 |
0 |
0 |
124.0 |
+-- Pseudodon_lenyanensis **
|
+-- Nyeinchanconcha_nyeinchani **
+-----------------------------------------------+
| +-- Indopseudodon_kayinensis **
| |
| +-- Sundadontina_tanintharyiensis **
|
| +-- Trapezidens_exolescens **
| +-----+
| | +-- Trapezidens_scutum **
| +-----------+
| | | +----- Trapezidens_angustior **
| | +--+
| | | +-- Trapezidens_dolichorhynchus **
| +--------------------------+ +--+
| | | +-- Trapezidens_yeti **
| | |
--+ | | +----------------- Lamellidens_corrianus **
| | | |
| | +--+ +-------------- Lamellidens_nongyangensis **
| | | |
| | +--+ +----------- Lamellidens_savadiensis **
| | | |
| | | | +-- Lamellidens_ferrugineus **
| | +--+ +-----+
| | | | +-- Lamellidens_marginalis **
| | | |
| | +--+ +-- Lamellidens_lamellatus **
| | | +--+
| | | | +-- Lamellidens_jenkinsianus **
| | +--+
| | | +-- Lamellidens_generosus **
| | +--+
+--+ +-- Lamellidens_mainwaringi **
|
| +----------- Parreysia_keralaensis **
| +--+
| | | +-------- Parreysia_plagiosoma **
| | +--+
| | | +----- Parreysia_nagpoorensis **
| | +--+
| | | +-- Parreysia_corrugata **
| +-----------------------------+ +--+
| | | +-- Parreysia_rakhinensis **
| | |
| | | +-- Leoparreysia_burmana **
| | | +-----+
| | | | +-- Leoparreysia_whitteni **
| | +-----+
| | | +----- Leoparreysia_olivacea **
| | +--+
| | | +-- Leoparreysia_canefrii **
| | +--+
+--+ +-- Leoparreysia_tavoyensis **
|
| +-------- Nitia_teretiuscula **
| +--------------------------------+
| | | +----- Coelatura_aegyptiaca **
| | +--+
| | | +-- Coelatura_kunenensis **
| | +--+
| | +-- Nyassunio_nyassaensis **
| |
| | +-- Indonaia_cylindrica **
| | +--------+
+--+ | +-- Indonaia_gratiosa **
| |
| +--------------------------+ +----- Indonaia_andersoniana **
| | | +--+
| | | | | +-- Indonaia_caerulea **
| | | | +--+
| | +--+ +-- Indonaia_subclathrata **
| | |
| | | +----- Indonaia_shurtleffiana **
| | +--+
+--+ | +-- Indonaia_bonneaudii **
| +--+
| +-- Indonaia_rugosa **
|
| +----------------------------------- Indochinella_pugio **
| |
| | +----------- Radiatula_crispisulcata **
| | |
+--+ +--------------------+ +-- Radiatula_chaudhurii **
| | | +-----+
| | | | +-- Radiatula_myitkyinae **
| | +--+
| | | +----- Radiatula_mouhoti **
+--+ +--+
| | +-- Radiatula_chindwinensis **
| +--+
| +-- Radiatula_myitthanensis **
|
| +----------------------------- Scabiellus_songkramensis **
+--+
| +-------------------------- Harmandia_somboriensis **
+--+
| +----------------------- Unionetta_fabagina **
+--+
| +-------------------- Scabies_crispata **
+--+
| +----------------- Scabies_mandarinus **
+--+
| +-------------- Scabies_nucleus **
+--+
| +----------- Scabies_pilata **
+--+
| +-------- Scabies_humilis **
+--+
| +----- Scabies_anceps **
+--+
| +-- Scabies_phaselus **
+--+
+-- Scabies_scobinatus **
Bolotov et al., 2022 [Publ 4239], fig. S1
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 58 |
58 |
52 |
1 |
0 |
0 |
0 |
124.0 |
+-- Pseudodon_lenyanensis **
|
+-- Nyeinchanconcha_nyeinchani **
+--------------------------------------------+
| +-- Indopseudodon_kayinensis **
| |
| +-- Sundadontina_tanintharyiensis **
|
| +-------- Trapezidens_exolescens **
| |
| +--------+ +----- Trapezidens_scutum **
| | | |
| | +--+ +-- Trapezidens_angustior **
| | | |
| | +--+-- Trapezidens_dolichorhynchus **
| | |
| +--------------------------+ +-- Trapezidens_yeti **
--+ | |
| | | +----- Lamellidens_savadiensis **
| | | +--------+
| | | | | +-- Lamellidens_corrianus **
| | | | +--+
| | | | +-- Lamellidens_nongyangensis **
| | +--+
| | | +-- Lamellidens_ferrugineus **
| | | +--------+
| | | | +-- Lamellidens_marginalis **
| | +--+
| | | +-------- Lamellidens_lamellatus **
| | +--+
| | | +----- Lamellidens_jenkinsianus **
+--+ +--+
| | +-- Lamellidens_generosus **
| +--+
| +-- Lamellidens_mainwaringi **
|
| +----------- Parreysia_keralaensis **
| +-----------------------------+
| | | +-------- Parreysia_corrugata **
| | +--+
| | | +----- Parreysia_rakhinensis **
| | +--+
| | | +-- Parreysia_plagiosoma **
| | +--+
| | +-- Parreysia_nagpoorensis **
| |
| | +-- Leoparreysia_burmana **
+--+ +-----+
| | +-- Leoparreysia_whitteni **
| +-----------------------------+
| | | +----- Leoparreysia_olivacea **
| | +--+
| | | +-- Leoparreysia_canefrii **
| | +--+
| | +-- Leoparreysia_tavoyensis **
| |
| | +-------- Nitia_teretiuscula **
+--+ +--------------------------+
| | | +----- Coelatura_aegyptiaca **
| | +--+
| | | +-- Coelatura_kunenensis **
| | +--+
| | +-- Nyassunio_nyassaensis **
| |
| | +----------------- Indonaia_cylindrica **
| | |
+--+ +--------------+ +-------------- Indonaia_gratiosa **
| | | |
| | +--+ +----------- Indonaia_andersoniana **
| | | |
| | +--+ +-- Indonaia_caerulea **
| | | +-----+
| | | | +-- Indonaia_subclathrata **
| | +--+
| | | +----- Indonaia_rugosa **
+--+ +--+
| | +-- Indonaia_bonneaudii **
| +--+
| +-- Indonaia_shurtleffiana **
|
| +----------------------------- Indochinella_pugio **
| |
| | +-------- Radiatula_crispisulcata **
| | |
| | +-----------------+ +-- Radiatula_chaudhurii **
+--+ | | +--+
| | | | +-- Radiatula_myitkyinae **
| | +--+
| | | +-- Radiatula_myitthanensis **
| | | |
+--+ +--+-- Radiatula_mouhoti **
| |
| +-- Radiatula_chindwinensis **
|
| +----------------------- Scabiellus_songkramensis **
| |
+--+ +-------------------- Harmandia_somboriensis **
| |
+--+ +----------------- Unionetta_fabagina **
| |
+--+ +-- Scabies_crispata **
| +-----------+
| | +-- Scabies_mandarinus **
+--+
| +----------- Scabies_nucleus **
| |
+--+ +-- Scabies_humilis **
| +-----+
| | +-- Scabies_pilata **
+--+
| +----- Scabies_phaselus **
+--+
| +-- Scabies_anceps **
+--+
+-- Scabies_scobinatus **
Bolotov et al., 2022 [Publ 4239], fig. S2
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 58 |
58 |
54 |
1 |
0 |
0 |
0 |
124.0 |
+-- Pseudodon_lenyanensis **
|
+-- Nyeinchanconcha_nyeinchani **
+-----------------------------------------------+
| +-- Indopseudodon_kayinensis **
| |
| +-- Sundadontina_tanintharyiensis **
|
| +----------- Parreysia_keralaensis **
| +-----------------------------------+
| | | +-------- Parreysia_corrugata **
| | +--+
| | | +----- Parreysia_nagpoorensis **
| | +--+
--+ | | +-- Parreysia_plagiosoma **
| | +--+
| | +-- Parreysia_rakhinensis **
| |
| | +----------- Trapezidens_scutum **
| | +-----+
| | | | +-------- Trapezidens_exolescens **
| | | +--+
| | | | +----- Trapezidens_yeti **
| | | +--+
| | | | +-- Trapezidens_dolichorhynchus **
+--+ | +--+
| +--------------------------+ +-- Trapezidens_angustior **
| | |
| | | +----- Lamellidens_savadiensis **
| | | +--------+
| | | | | +-- Lamellidens_corrianus **
| | | | +--+
| | | | +-- Lamellidens_nongyangensis **
| | +--+
| | | +-- Lamellidens_marginalis **
| | | +--------+
| | | | +-- Lamellidens_ferrugineus **
| | +--+
| | | +-------- Lamellidens_lamellatus **
| | +--+
| | | +----- Lamellidens_jenkinsianus **
+--+ +--+
| | +-- Lamellidens_mainwaringi **
| +--+
| +-- Lamellidens_generosus **
|
| +----------- Leoparreysia_burmana **
| +-----------------------------+
| | | +-------- Leoparreysia_whitteni **
| | +--+
| | | +----- Leoparreysia_olivacea **
| | +--+
| | | +-- Leoparreysia_canefrii **
| | +--+
| | +-- Leoparreysia_tavoyensis **
| |
+--+ +-------- Nitia_teretiuscula **
| +-----------------------------+
| | | +----- Coelatura_aegyptiaca **
| | +--+
| | | +-- Coelatura_kunenensis **
| | +--+
| | +-- Nyassunio_nyassaensis **
| |
| | +-- Indonaia_cylindrica **
| | +--------+
+--+ | +-- Indonaia_gratiosa **
| |
| +-----------------------+ +----- Indonaia_shurtleffiana **
| | | +--+
| | | | | +-- Indonaia_bonneaudii **
| | | | +--+
| | +--+ +-- Indonaia_rugosa **
| | |
| | | +----- Indonaia_andersoniana **
| | +--+
+--+ | +-- Indonaia_caerulea **
| +--+
| +-- Indonaia_subclathrata **
|
| +-------------------------------- Indochinella_pugio **
| |
| | +----------- Radiatula_crispisulcata **
| | |
| | +-----------------+ +-- Radiatula_chaudhurii **
+--+ | | +-----+
| | | | +-- Radiatula_myitkyinae **
| | +--+
| | | +----- Radiatula_mouhoti **
| | +--+
+--+ | +-- Radiatula_myitthanensis **
| +--+
| +-- Radiatula_chindwinensis **
|
| +-------------------------- Scabiellus_songkramensis **
| |
+--+ +----------------------- Harmandia_somboriensis **
| |
+--+ +-------------------- Unionetta_fabagina **
| |
+--+ +----------------- Scabies_crispata **
| |
+--+ +-- Scabies_humilis **
| +-----------+
| | +-- Scabies_mandarinus **
+--+
| +----------- Scabies_nucleus **
+--+
| +-------- Scabies_pilata **
+--+
| +----- Scabies_anceps **
+--+
| +-- Scabies_phaselus **
+--+
+-- Scabies_scobinatus **
Zanatta & Murphy, 2006 [Publ 2672], fig. 2
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 57 |
57 |
48 |
2 |
0 |
0 |
0 |
122.0 |
+----------------------------------------------------- Pyganodon_grandis **
|
+----------------------------------------------------- Cumberlandia_monodonta **
|
| +-------- Pleuronaia_dolabelloides **
| |
--+ +-----------------------------------------+-------- Pleurobema_clava **
| | |
| | | +----- Fusconaia_flava **
| | +--+
| | | +-- Hemistena_lata **
| | +--+
| | +-- Eurynia_dilatata **
+--+
| +----------------------------------------------- Amblema_plicata **
| |
| +----------------------------------------------- Popenaias_popeii **
| |
| | +----- Paetulunio_fabalis **
| | |
| | +--------------------------------------+ +-- Toxolasma_parvum **
+--+ | | |
| | +--+-- Toxolasma_lividum **
| | |
| | +-- Toxolasma_texasiense **
| |
| | +----- Obliquaria_reflexa **
| | +-----------------------------------+
+--+ | | +-- Glebula_rotundata **
| | +--+
| | +-- Cyrtonaias_tampicoensis **
| |
| | +----- Ptychobranchus_fasciolaris **
| | +--+
| | | | +-- Dromus_dromas **
| | | +--+
+--+ +-----------------------------+ +-- Cyprogenia_stegaria **
| | |
| | | +----- Medionidus_conradicus **
| | +--+
| | | +-- Medionidus_acutissimus **
| | +--+
| | +-- Lemiox_rimosus **
| |
| | +----------------------------------- Sagittunio_nasutus **
| | |
+--+ | +-------------- Potamilus_leptodon **
| | |
| +--------------------+ +----------- Potamilus_fragilis **
| | | |
| | +--+ +-- Potamilus_alatus **
| | | +-----+
| | | | +-- Potamilus_purpuratus **
| | +--+
| | | +----- Potamilus_inflatus **
| | +--+
+--+ | +-- Potamilus_ohiensis **
| +--+
| +-- Potamilus_amphichaenus **
|
| +-- Truncilla_truncata **
| +-----------------------------+
| | +-- Ellipsaria_lineolata **
| |
| | +-- Ortmanniana_pectorosa **
| | +-----+
| | | +-- Lampsilis_fasciola **
| | +--------------------+
+--+ | | +----- Lampsilis_ornata **
| | +--+
| | | +-- Lampsilis_ovata **
| | +--+
| | +-- Lampsilis_cardium **
| |
| | +-- Ligumia_recta **
+--+ +--------+
| | +-- Cambarunio_iris **
| +--------------+
| | | +-------- Lampsilis_teres **
| | | |
| | +--+ +-- Hamiota_australis **
| | | +--+
| | | | +-- Hamiota_subangulata **
+--+ +--+
| | +-- Hamiota_altilis **
| +--+
| +-- Hamiota_perovalis **
|
| +----------------------- Leaunio_vanuxemensis **
| |
+--+ +-------------------- Ortmanniana_ligamentina **
| |
+--+ +----------------- Lampsilis_siliquoidea **
| |
| | +-------------- Obovaria_olivaria **
+--+ |
| +-------------- Venustaconcha_ellipsiformis **
| |
| | +----- Obovaria_unicolor **
+--+--------+
| | +-- Obovaria_subrotunda **
| +--+
| +-- Obovaria_arkansasensis **
|
| +----------- Epioblasma_brevidens **
+--+
| +-------- Epioblasma_triquetra **
+--+
| +----- Epioblasma_rangiana **
+--+
| +-- Epioblasma_walkeri **
+--+
+-- Epioblasma_capsaeformis **
Zanatta & Murphy, 2006 [Publ 2672], fig. 1
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 56 |
56 |
51 |
2 |
0 |
0 |
0 |
122.0 |
+----------------------------------------------------- Cumberlandia_monodonta **
|
| +-------------------------------------------------- Pyganodon_grandis **
| |
--+ | +-- Amblema_plicata **
| | +--+
| | | +-- Popenaias_popeii **
| | |
+--+ +-----------------------------------------+ +-- Pleurobema_clava **
| | | |
| | | +-- Eurynia_dilatata **
| | +--+
| | +-- Pleuronaia_dolabelloides **
| | |
+--+ +-- Fusconaia_flava **
|
| +-- Cyrtonaias_tampicoensis **
| +-----------------------------------------+
| | +-- Glebula_rotundata **
| |
| | +----------------------------------------- Paetulunio_fabalis **
+--+ |
| | +-------- Obliquaria_reflexa **
| | +-----------------------------+
| | | | +----- Toxolasma_texasiense **
| | | +--+
+--+ | | +-- Toxolasma_parvum **
| | +--+
| | +-- Toxolasma_lividum **
| |
| | +----------------------------------- Sagittunio_nasutus **
| | |
+--+ | +-- Medionidus_conradicus **
| | +-----+
| | | +-- Ptychobranchus_fasciolaris **
| | +--------+
| | | | +----- Medionidus_acutissimus **
| | | | |
| | | +--+----- Lemiox_rimosus **
| | | |
| | +--------------+ | +-- Dromus_dromas **
+--+ | | +--+
| | | +-- Cyprogenia_stegaria **
| | |
| | | +-------------- Potamilus_leptodon **
| | | |
| | +--+ +----------- Potamilus_fragilis **
| | | |
| | +--+ +-- Potamilus_alatus **
| | | +-----+
| | | | +-- Potamilus_purpuratus **
| | +--+
+--+ | +----- Potamilus_inflatus **
| +--+
| | +-- Potamilus_ohiensis **
| +--+
| +-- Potamilus_amphichaenus **
|
| +----- Lampsilis_teres **
| +-----------------------+
| | | +-- Truncilla_truncata **
| | +--+
| | +-- Ellipsaria_lineolata **
| |
| | +-------------------------- Leaunio_vanuxemensis **
+--+ |
| | +-- Hamiota_australis **
| | +--+
| | | +-- Hamiota_subangulata **
| | +-----+
| | | | +-- Hamiota_altilis **
| | | +--+
+--+ | +-- Hamiota_perovalis **
| +-----------+
| | | +-- Lampsilis_fasciola **
| | | +-----+
| | | | +-- Ortmanniana_pectorosa **
| | +--+
| | | +----- Lampsilis_ornata **
| | +--+
| | | +-- Lampsilis_ovata **
+--+ +--+
| +-- Lampsilis_cardium **
|
| +-------- Lampsilis_siliquoidea **
| +-----------+
| | | +----- Ortmanniana_ligamentina **
| | +--+
| | | +-- Cambarunio_iris **
| | +--+
+--+ +-- Ligumia_recta **
|
| +----------------- Venustaconcha_ellipsiformis **
| |
| | +-------- Obovaria_olivaria **
+--+ +-----+
| | | +----- Obovaria_unicolor **
| | +--+
| | | +-- Obovaria_subrotunda **
+--+ +--+
| +-- Obovaria_arkansasensis **
|
| +----------- Epioblasma_brevidens **
+--+
| +-------- Epioblasma_triquetra **
+--+
| +----- Epioblasma_rangiana **
+--+
| +-- Epioblasma_walkeri **
+--+
+-- Epioblasma_capsaeformis **
Konopleva et al., 2023 [Publ 4269], fig. S1, 4
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 56 |
56 |
54 |
2 |
0 |
0 |
0 |
83.0 |
+----------------------------------- Gibbosula_laosensis **
|
| +-- Trapezidens_exolescens **
| +-----------+
| | +-- Lamellidens_savadiensis **
| +-----------------+
| | | +----------- Parreysia_rakhinensis **
| | +--+
| | | +-------- Leoparreysia_tavoyensis **
| | +--+
| | | +----- Indonaia_andersoniana **
| | +--+
--+ | | +-- Indochinella_pugio **
| | +--+
| | +-- Radiatula_mouhoti **
| |
| | +-- Lampsilis_ovata **
| | +--------------+
| | | +-- Amblema_plicata **
| | |
| | | +----------- Sinosolenaia_recognita **
| | | |
| | | +--+ +-- Ptychorhynchus_pfisteri **
| | +-----------+ | | +-----+
+--+ | | | | | +-- Parvasolenaia_rivularis **
| | | | +--+
| | | | | +----- Gonidea_angulata **
| | | | +--+
| | | | | +-- Microcondylaea_bonellii **
| | +--+ +--+
| | | +-- Leguminaia_wheatleyi **
| | |
| | | +----- Chamberlainia_somsakpanhai **
| | | +-----+
| | | | | +-- Sinohyriopsis_schlegelii **
| | | | +--+
| | +--+ +-- Sinohyriopsis_cumingii **
| | |
| | | +-------- Potomida_littoralis **
| | +--+
| | | +----- Pronodularia_japanensis **
| | +--+
| | | +-- Lamprotula_caveata **
+--+ +--+
| +-- Lamprotula_leaii **
|
| +----------------- Middendorffinaia_mongolica **
| |
| | +-------- Aculamprotula_tortuosa **
| | +-----+
| +--------+ | | +----- Aculamprotula_scripta **
| | | | +--+
| | | | | +-- Aculamprotula_tientsinensis **
| | | | +--+
| | +--+ +-- Aculamprotula_coreana **
| | |
| | | +----- Unio_tumidus **
| | | +-----+
| | | | | +-- Unio_pictorum **
| | | | +--+
| | +--+ +-- Unio_crassus **
| | |
| | | +-------- Nodularia_douglasiae **
+--+ +--+
| | +----- Tchangsinaia_piscicula **
| +--+
| | +-- Cuneopsis_rufescens **
| +--+
| +-- Cuneopsis_heudei **
|
| +-------- Lepidodesma_languilati **
| +--------------+
| | | +----- Lanceolaria_gladiola **
| | +--+
| | | +-- Lanceolaria_grayii **
| | +--+
+--+ +-- Lanceolaria_triformis **
|
| +-------------------- Acuticosta_chinensis **
| |
| | +----- Anodonta_anatina **
| | +-----------+
+--+ | | +-- Pseudanodonta_complanata **
| | +--+
| | +-- Anodonta_cygnea **
| |
| | +-------- Anemina_arcaeformis **
+--+ +-----+
| | | +----- Amuranodonta_kijaensis **
| | +--+
| | | +-- Buldowskia_suifunica **
| | +--+
| | +-- Buldowskia_shadini **
+--+
| +-------- Beringiana_beringiana **
| +--+
| | | +----- Sinanodonta_schrenkii **
| | +--+
| | | +-- Sinanodonta_woodiana **
+--+ +--+
| +-- Sinanodonta_lauta **
|
| +-------- Cristaria_radiata **
| |
+--+ +-- Cristaria_truncata **
| +--+
| | +-- Cristaria_plicata **
+--+
| +-- Cristaria_bellua **
+--+
+-- Cristaria_clessini **
Fukata & Iigo, 2020 [Publ 4106], fig. 1
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 54 |
54 |
52 |
2 |
0 |
0 |
0 |
86.0 |
+----- Gibbosula_crassa **
+-----------------------+
| | +-- Pseudunio_marocanus **
| +--+
| +-- Cumberlandia_monodonta **
|
| +-- Quadrula_quadrula **
| +-----------------+
| | +-- Uniomerus_tetralasmus **
| +--+
| | | +----------------- Elliptio_complanata **
| | | |
| | +--+ +-- Potamilus_leptodon **
| | | +-----------+
| | | | +-- Potamilus_alatus **
| | +--+
| | | +----------- Leaunio_lienosus **
| | | |
| | +--+ +-- Lampsilis_cardium **
| | | +-----+
--+ | | | +-- Lampsilis_ornata **
| +--+ +--+
| | | | +----- Venustaconcha_ellipsiformis **
| | | +--+
| | | | +-- Lampsilis_powellii **
| | | +--+
| | | +-- Lampsilis_siliquoidea **
| | |
| | | +-- Sinohyriopsis_cumingii **
| | | +-----------+
| | | | +-- Sinohyriopsis_schlegelii **
| | | |
| | | | +-- Pilsbryoconcha_exilis **
| | +--------+ +--------+
| | | | +-- Pseudodon_vondembuschianus **
| | | |
| | | | +-- Lamprotula_caveata **
| | | | +--+
| | +--+ | +-- Lamprotula_leaii **
| | | +--+
+--+ | | | +-- Pronodularia_japanensis **
| | | +--+
| | | +-- Potomida_littoralis **
| +--+
| | +-- Sinosolenaia_carinata **
| | +--+
| | | +-- Sinosolenaia_oleivora **
| +--+
| | +-- Parvasolenaia_rivularis **
| +--+
| +-- Ptychorhynchus_pfisteri **
|
| +----------------------- Lepidodesma_languilati **
| |
| | +----- Lanceolaria_gladiola **
| | +--------------+
| | | | +-- Lanceolaria_grayii **
| | | +--+
| | | +-- Lanceolaria_lanceolata **
+--+ |
| | +-------------- Acuticosta_chinensis **
| | |
| | | +-------- Cristaria_plicata **
| | | |
| | | +--+ +-- Sinanodonta_woodiana **
| | +--+ | | +--+
+--+ | | | | | +-- Sinanodonta_lucida **
| | | | +--+
| | | | | +-- Anemina_arcaeformis **
| | | | +--+
| | +--+ +-- Anemina_euscaphys **
| | |
| | | +-- Anodonta_anatina **
| | | +-----+
| | | | +-- Anodonta_cygnea **
| | | |
| | +--+ +-- Pyganodon_grandis **
+--+ | +--+
| | | +-- Utterbackia_peninsularis **
| +--+
| | +-- Alasmidonta_varicosa **
| +--+
| +-- Prolasmidonta_heterodon **
|
| +----- Aculamprotula_tortuosa **
| +--------+
| | | +-- Aculamprotula_tientsinensis **
| | +--+
| | +-- Aculamprotula_coreana **
| |
+--+ +-------- Unio_tumidus **
| +--+
| | | +----- Unio_crassus **
| | +--+
| | | +-- Unio_pictorum **
| | +--+
+--+ +-- Unio_delphinus **
|
| +-- Schistodesmus_lampreyanus **
| +-----+
| | +-- Cuneopsis_heudei **
+--+
| +----- Pseudocuneopsis_capitata **
+--+
| +-- Nodularia_douglasiae **
+--+
+-- Tchangsinaia_piscicula **
Wu et al., 2018 [Publ 3836], fig. 2
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 53 |
53 |
45 |
2 |
0 |
0 |
0 |
87.0 |
+-- Margaritifera_dahurica **
+--------------------------------+
| +-- Margaritifera_falcata **
|
| +-------------------------------- Quadrula_quadrula **
| |
| | +-- Amblema_plicata **
| +-----------------------------+
| | +-- Obovaria_arkansasensis **
| |
--+ | +----------- Pleurobema_sintoxia **
| | |
| +--------------------+ +-- Elliptio_complanata **
| | | +-----+
| | | | +-- Eurynia_dilatata **
| | +--+
| | | +----- Fusconaia_askewi **
| | +--+
| | | +-- Fusconaia_cerina **
| | +--+
+--+ +-- Fusconaia_burkei **
|
| +----- Potomida_littoralis **
| +-----+
| | | +-- Lamprotula_caveata **
| | +--+
| +-----------------+ +-- Lamprotula_leaii **
| | |
| | | +-------- Sinosolenaia_carinata **
| | | |
| | +--+ +----- Gonidea_angulata **
| | | |
| | +--+----- Sinosolenaia_oleivora **
| | |
| | | +-- Parvasolenaia_rivularis **
| | +--+
| | +-- Ptychorhynchus_pfisteri **
+--+
| +-------------------------- Sinohyriopsis_cumingii **
| |
| | +----------------------- Lepidodesma_languilati **
| | |
| | | +----------- Aculamprotula_tortuosa **
| | | +-----+
| | | | | +-------- Aculamprotula_scripta **
| | | | +--+
| | | | | +----- Aculamprotula_coreana **
| | | | +--+
+--+ | | | +-- Aculamprotula_fibrosa **
| +-----+ +--+
| | | +-- Aculamprotula_tientsinensis **
| | |
| | | +-- Unio_crassus **
| | | +-----------+
| | | | +-- Unio_pictorum **
| | | |
| | +--+ +----- Nodularia_douglasiae **
| | | +-----+
| | | | | +-- Schistodesmus_lampreyanus **
+--+ | | +--+
| +--+ +-- Schistodesmus_spinosus **
| |
| | +-------- Tchangsinaia_piscicula **
| +--+
| | +----- Cuneopsis_rufescens **
| +--+
| | +-- Cuneopsis_celtiformis **
| +--+
| +-- Cuneopsis_heudei **
|
| +----------- Acuticosta_chinensis **
| +--------+
| | | +-------- Lanceolaria_gladiola **
| | +--+
| | | +----- Lanceolaria_grayii **
| | +--+
| | | +-- Lanceolaria_lanceolata **
| | +--+
| | +-- Lanceolaria_triformis **
+--+
| +-------- Anemina_arcaeformis **
| |
| +-------- Cristaria_plicata **
| +--------+
| | | +-- Sinanodonta_woodiana **
| | | +--+
| | | | +-- Sinanodonta_elliptica **
| | +--+
+--+ | +-- Sinanodonta_lucida **
| +--+
| +-- Sinanodonta_angula **
|
| +-------------- Anodonta_cygnea **
| |
+--+ +----------- Anodonta_anatina **
| |
+--+ +-------- Strophitus_undulatus **
| |
+--+ +----- Platynaias_compressa **
| |
+--+----- Alasmidonta_marginata **
|
| +-- Pyganodon_cataracta **
+--+
+-- Pyganodon_grandis **
Huang et al., 2019 [Publ 3843], fig. 3
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 53 |
53 |
51 |
2 |
4 |
3 |
2 |
81.0 |
+-- Anodontites_trapesialis
+--------------------------------------+
| +-- Mutela_dubia
|
| +-------------------------------------- Echyridella_menziesii
| |
| | +-------------- Gibbosula_rochechouartii **
| | +--------------------+
--+ | | | +----------- Cumberlandia_monodonta **
| | | +--+
| | | | +-------- Pseudunio_marocanus **
| | | +--+
| | | | +----- Margaritifera_falcata **
| | | +--+
| | | | +-- Margaritifera_margaritifera **
| | | +--+
+--+ | +-- Margaritifera_dahurica **
| |
| | +-- Sinohyriopsis_cumingii **
| | +-----------+
| | | +-- Sinohyriopsis_schlegelii **
| | |
| | | +-------- Potomida_littoralis **
| | +--------+ +--+
| | | | | | +----- Pronodularia_japanensis **
| | | | | +--+
| | | | | | +-- Lamprotula_caveata **
| | | | | +--+
+--+ | +--+ +-- Lamprotula_leaii **
| | |
| | | +-- Ptychorhynchus_pfisteri **
| | | +--+
| +--------+ | | +-- Parvasolenaia_rivularis **
| | | +-----+
| | | | +-- Sinosolenaia_oleivora **
| | | +--+
| | | +-- Sinosolenaia_carinata **
| | |
| | | +-- Quadrula_quadrula **
| | | +-----------------+
| | | | +-- Uniomerus_tetralasmus **
| | +--+
| | | +----------------- Elliptio_complanata **
| | | |
| | +--+ +-------------- Toxolasma_parvum **
| | | |
| | +--+ +-- Potamilus_leptodon **
| | | +--------+
| | | | +-- Potamilus_alatus **
+--+ +--+
| | +-------- Leaunio_lienosus **
| +--+
| | +----- Venustaconcha_ellipsiformis **
| +--+
| | +-- Lampsilis_cardium **
| +--+
| +-- Lampsilis_ornata **
|
| +----------------------------- Lepidodesma_languilati **
| |
| | +-------- Aculamprotula_tortuosa **
| | +-----+
| | | | +----- Aculamprotula_scripta **
| | | +--+
| | | | +-- Aculamprotula_tientsinensis **
| | | +--+
| | +-----------+ +-- Aculamprotula_coreana **
+--+ | |
| | | +----- Nodularia_douglasiae **
| | | +-----+
| | | | | +-- Tchangsinaia_piscicula **
| | | | +--+
| | +--+ +-- Schistodesmus_lampreyanus **
| | |
| | | +-------- Unio_tumidus **
| | +--+
+--+ | +----- Unio_crassus **
| +--+
| | +-- Unio_delphinus **
| +--+
| +-- Unio_pictorum **
|
| +----- Lanceolaria_gladiola **
| +-----------------+
| | | +-- Lanceolaria_lanceolata **
| | +--+
| | +-- Lanceolaria_grayii **
+--+
| +-------------------- Acuticosta_chinensis **
| |
| | +-------- Cristaria_plicata **
| | |
+--+ +--------+ +-- Sinanodonta_lucida **
| | | +--+
| | | | +-- Sinanodonta_woodiana **
| | +--+
+--+ | +-- Anemina_arcaeformis **
| +--+
| +-- Anemina_euscaphys **
|
| +-------------- Anodonta_cygnea **
+--+
| +----------- Anodonta_anatina **
+--+
| +-------- Lasmigona_costata **
+--+
| +----- Pyganodon_grandis **
+--+
| +-- Utterbackia_imbecillis **
+--+
+-- Utterbackia_peninsularis **
Dai et al., 2023 [Publ 4285], fig. 5
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 53 |
53 |
51 |
2 |
0 |
0 |
0 |
87.0 |
+-- Gibbosula_rochechouartii **
+--------------------------------+
| +-- Margaritifera_dahurica **
|
| +-- Quadrula_quadrula **
| +--------------------------+
| | +-- Uniomerus_tetralasmus **
| |
| +--+ +-- Elliptio_complanata **
| | | +-----------------------+
| | | | +-- Pleurobema_oviforme **
| | +--+
| | | +----------------------- Amblema_plicata **
| | | |
| | +--+ +-------------------- Popenaias_popeii **
| | | |
--+ | +--+ +----------------- Toxolasma_parvum **
| | | |
| | +--+ +-- Potamilus_leptodon **
| | | +-----------+
| | | | +-- Potamilus_alatus **
| | +--+
| | | +----------- Leaunio_lienosus **
| | | |
| | +--+ +-- Lampsilis_cardium **
| | | +-----+
| | | | +-- Lampsilis_ornata **
| | +--+
| | | +----- Venustaconcha_ellipsiformis **
| | +--+
| | | +-- Lampsilis_powellii **
| | +--+
+--+ +-- Lampsilis_siliquoidea **
|
| +-------------------- Lepidodesma_languilati **
| |
| | +-------------- Aculamprotula_tientsinensis **
| | |
| | +--+ +----------- Unio_crassus **
| | | | |
| +--+ | +--+ +-- Pseudocuneopsis_capitata **
| | | | | +-----+
| | | | | | +-- Nodularia_douglasiae **
| | | | +--+
| | | | | +----- Schistodesmus_lampreyanus **
| | | | +--+
| | +--+ | +-- Cuneopsis_celtiformis **
| | | +--+
| | | +-- Tchangsinaia_piscicula **
| | |
| | | +-- Acuticosta_chinensis **
| | | +-----------+
| | | | +-- Lanceolaria_grayii **
| | | |
| | +--+ +----- Cristaria_plicata **
| | | +-----+
| | | | | +-- Anemina_arcaeformis **
+--------+ | | +--+
| +--+ +-- Sinanodonta_woodiana **
| |
| | +-------- Anodonta_cygnea **
| +--+
| | +----- Platynaias_compressa **
| +--+
| | +-- Pyganodon_grandis **
| +--+
| +-- Utterbackia_imbecillis **
|
| +----- Chamberlainia_somsakpanhai **
| +--------------+
| | | +-- Sinohyriopsis_cumingii **
| | +--+
| | +-- Sinohyriopsis_schlegelii **
| |
| | +-- Hyriopsis_bialata **
+--+ +--+
| | +-- Rectidens_sumatrensis **
| +-----------+
| | | +-- Lens_contradens **
| | +--+
| | +-- Physunio_superbus **
| |
+--+ +-------- Potomida_littoralis **
| +-----+
| | | +----- Pronodularia_japanensis **
| | +--+
| | | +-- Lamprotula_caveata **
| | +--+
+--+ +-- Lamprotula_leaii **
|
| +-- Pseudodon_vondembuschianus **
| +--------+
| | +-- Pilsbryoconcha_exilis **
| |
+--+ +----- Parvasolenaia_rivularis **
| +--+
| | | +-- Postolata_guangxiensis **
| | +--+
+--+ +-- Ptychorhynchus_pfisteri **
|
| +----- Microcondylaea_bonellii **
+--+
| +-- Sinosolenaia_oleivora **
+--+
+-- Sinosolenaia_carinata **
Konopleva et al., 2019 [Publ 3900], fig. 1
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 52 |
52 |
41 |
2 |
0 |
0 |
0 |
106.0 |
+-- Gibbosula_laosensis **
+--------------------------------+
| +-- Margaritifera_dahurica **
|
| +-- Indopseudodon_indawgyiensis **
| |
| +--+-- Indopseudodon_bogani **
| | |
| | +-- Indopseudodon_salwenianus **
| |
| | +-- Bineurus_mouhotii **
| +--------------------------+ |
| | | +-- Pseudodon_vagulus **
| | | |
| | | +-- Pseudodon_mekongi **
--+ | | |
| | | +-- Pilsbryoconcha_exilis **
| | +--+
| | +-- Pilsbryoconcha_carinifera **
| | |
| | +-- Bineurus_loeiensis **
| | |
| | +-- Bineurus_exilis **
| | |
| | +-- Pseudodon_vondembuschianus **
| |
| | +----------- Indonaia_shurtleffiana **
| | +--+
| | | | +-------- Indonaia_rugosa **
| | | +--+
+--+ | | +----- Indonaia_andersoniana **
| | +--+
| | | +-- Indonaia_caerulea **
| +--------------+ +--+
| | | +-- Indonaia_subclathrata **
| | |
| | | +----------- Indochinella_pugio **
| | | |
| | | | +-- Scabies_humilis **
| | +--+ |
| | | +-----+-- Scabies_mandarinus **
| | | | |
| | +--+ +-- Scabies_phaselus **
| | |
| | | +----- Radiatula_mouhoti **
| | +--+
| | | +-- Radiatula_myitkyinae **
| | +--+
| | +-- Radiatula_chaudhurii **
| |
+--+ +-- Ensidens_ingallsianus **
| +-----+
| | +-- Ensidens_telus **
| +-----+
| | | +----- Ensidens_sagittarius **
| | +--+
| | | +-- Ensidens_jaculus **
| +-----------+ +--+
| | | +-- Ensidens_spiculus **
| | |
| | | +----------- Hyriopsis_desowitzi **
| | | |
| | +--+----------- Rectidens_sumatrensis **
| | |
| | | +-------- Hyriopsis_panhai **
| | +--+
| | | +----- Hyriopsis_myersiana **
| | +--+
+--+ | +-- Hyriopsis_khoratensis **
| +--+
| +-- Hyriopsis_bialata **
|
| +-- Yaukthwa_paiensis **
| +-----+
| | +-- Yaukthwa_inlenensis **
| +--------------+
| | | +----- Yaukthwa_panhai **
| | +--+
| | | +-- Yaukthwa_nesemanni **
| | +--+
+--+ +-- Yaukthwa_rectangularis **
|
| +----- Physunio_superbus **
| +--------------+
| | | +-- Physunio_modelli **
| | +--+
+--+ +-- Physunio **
|
| +----------------- Trapezoideus_foliaceus **
| |
+--+ +-- Lens_contradens **
| +-----------+
| | +-- Lens_micropterus **
+--+
| +----------- Lens_misellus **
+--+
| +-------- Lens_eximius **
+--+
| +----- Lens_pallegoixi **
+--+
| +-- Lens_comptus **
+--+
+-- Lens_novoselovi **
Pfeiffer et al., 2019 [Publ 3937], fig. 4-5
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 52 |
52 |
50 |
2 |
5 |
4 |
2 |
96.0 |
+----- Tamsiella_schroeteriana
+--------------------------------+
| | +-- Etheria_elliptica
| +--+
| +-- Aspatharia_pfeifferiana
|
| +----------------------------------- Hyridella_australis
| |
--+ | +-- Margaritifera_margaritifera **
| | +-----------------------------+
| | | +-- Margaritifera_hembeli **
| | |
| | | +-- Quadrula_quadrula **
| | | +--------------------+
| | | | +-- Uniomerus_tetralasmus **
+--+ | |
| | | +----- Elliptoideus_sloatianus **
| | +-----+ +--------------+
| | | | | | +-- Elliptio_complanata **
| | | | | +--+
| | | | | +-- Pleurobema_strodeanum **
| | | | |
| | | +--+ +----- Reginaia_ebenus **
| | | | +-----------+
| | | | | | +-- Plectomerus_dombeyanus **
+--+ | | | +--+
| | | | +-- Amblema_plicata **
| | +--+
| | | +-- Psoronaias_semigranosa **
| | | +-----------+
| | | | +-- Popenaias_popeii **
| | +--+
| | | +----------- Obliquaria_reflexa **
| | +--+
| | | +-------- Cyrtonaias_tampicoensis **
| | +--+
| | | +----- Leaunio_lienosus **
| | +--+
| | | +-- Lampsilis_cardium **
| | +--+
| | +-- Pachynaias_spheniopsis **
| |
+--+ +-- Chamberlainia_hainesiana **
| +--------------------+
| | +-- Sinohyriopsis_cumingii **
| |
| | +----------------- Lamprotula_cornuumlunae **
| | +--+
| | | | +-------------- Gonidea_angulata **
| +--+ | +--+
| | | | | +----------- Indopseudodon_crebristriatus **
| | | | +--+
| | | | | +-------- Bineurus_anodontinum **
| | | | +--+
| | | | | +----- Pilsbryoconcha_exilis **
| | | | +--+
| | +--+ | +-- Pseudodon_vondembuschianus **
| | | +--+
| | | +-- Pseudodon_inoscularis **
| | |
| | | +----- Ensidens_ingallsianus **
| | | +-----+
| | | | | +-- Hyriopsis_bialata **
| | | | +--+
| | | | +-- Rectidens_sumatrensis **
| | +--------+
| | | +-- Trapezoideus_foliaceus **
| | | +-----+
+--+ | | +-- Physunio_superbus **
| +--+
| | +----- Yaukthwa_paiensis **
| +--+
| | +-- Lens_contradens **
| +--+
| +-- Solenaia_emarginata **
|
| +----------- Lanceolaria_lanceolata **
| |
| | +----- Unio_gibbus **
| +--------+ +--+
| | | | | +-- Nodularia_douglasiae **
| | | | +--+
| | | | +-- Tchangsinaia_piscicula **
| | +--+
| | | +-- Cristaria_plicata **
| | | +--+
| | | | +-- Sinanodonta_woodiana **
+-----+ +--+
| | +-- Anodonta_cygnea **
| +--+
| +-- Pyganodon_grandis **
|
| +-- Trapezidens_dolichorhynchus **
| +--------------+
| | +-- Lamellidens_savadiensis **
| |
+--+ +-- Leoparreysia_tavoyensis **
| +-----------+
| | +-- Leoparreysia_olivacea **
| |
+--+ +-- Coelatura_choziensis **
| +--------+
| | +-- Prisodontopsis_aviculaeformis **
+--+
| +-------- Indonaia_caerulea **
+--+
| +----- Indochinella_pugio **
+--+
| +-- Scabies_phaselus **
+--+
+-- Harmandia_somboriensis **
Pfeiffer et al., 2019 [Publ 3937], fig. 6
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 52 |
53 |
50 |
2 |
5 |
4 |
2 |
98.0 |
+----- Tamsiella_schroeteriana
+-----------------------+
| | +-- Etheria_elliptica
| +--+
| +-- Aspatharia_pfeifferiana
|
| +----- Hyridella_australis
| +--------------------+
| | | +-- Margaritifera_margaritifera **
| | +--+
| | +-- Margaritifera_hembeli **
| |
| | +-- Quadrula_quadrula **
| | +-----------------+
| | | +-- Uniomerus_tetralasmus **
--+ | |
| | | +----- Elliptoideus_sloatianus **
| | | +--+
| | | | | +-- Elliptio_complanata **
| | +--+ | +--+
| | | | +--------+ +-- Pleurobema_strodeanum **
| | | | | |
| | | | | | +----- Reginaia_ebenus **
| | | | | +--+
| | | | | | +-- Plectomerus_dombeyanus **
| | | +--+ +--+
| | | | +-- Amblema_plicata **
| | | |
| | | | +-- Psoronaias_semigranosa **
| | | | +-----------+
+--+ | | | +-- Popenaias_popeii **
| | +--+
| | | +----------- Obliquaria_reflexa **
| | +--+
| | | +-------- Cyrtonaias_tampicoensis **
| | +--+
| | | +----- Leaunio_lienosus **
| | +--+
| | | +-- Lampsilis_cardium **
| | +--+
| | +-- Pachynaias_spheniopsis **
| |
| | +-- Chamberlainia_hainesiana **
| | +-----------------+
| | | +-- Sinohyriopsis_cumingii **
| | |
| | | +-- Gonidea_angulata **
| | | +-----------+
| | | | +-- Lamprotula_cornuumlunae **
| | | +--+
| +--+ | | +----------- Indopseudodon_crebristriatus **
| | | | +--+
| | | | | +-------- Pilsbryoconcha_exilis **
| | | | +--+
+--+ | | | +----- Bineurus_anodontinum **
| | | +--+
| | | | +-- Pseudodon_vondembuschianus **
| +--+ +--+
| | +-- Pseudodon_inoscularis **
| |
| | +----- Ensidens_ingallsianus **
| | +-----+
| | | | +-- Hyriopsis_bialata **
| | | +--+
| | | +-- Rectidens_sumatrensis **
| +-----+
| | +-- Trapezoideus_foliaceus **
| | +-----+
| | | +-- Physunio_superbus **
| +--+
| | +----- Yaukthwa_paiensis **
| +--+
| | +-- Lens_contradens **
| +--+
| +-- Solenaia_emarginata **
|
| +-------- Lanceolaria_lanceolata **
| +--+
| | | +----- Unio_gibbus **
| | +--+
| | | +-- Tchangsinaia_piscicula **
| | +--+
| +--------+ +-- Nodularia_douglasiae **
| | |
| | | +-- Cristaria_plicata **
| | | +--+
| | | | +-- Sinanodonta_woodiana **
| | +-----+
| | | +-- Anodonta_cygnea **
+--+ +--+
| +-- Pyganodon_grandis **
|
| +-- Trapezidens_dolichorhynchus **
| +--------------+
| | +-- Lamellidens_savadiensis **
| |
+--+ +-- Leoparreysia_tavoyensis **
| +-----------+
| | +-- Leoparreysia_olivacea **
| |
+--+ +-- Coelatura_choziensis **
| +--------+
| | +-- Prisodontopsis_aviculaeformis **
+--+
| +-------- Indonaia_caerulea **
+--+
| +----- Indochinella_pugio **
+--+
| +-- Scabies_phaselus **
+--+
+-- Harmandia_somboriensis **
Dai et al., 2024 [Publ 4358], fig. 4
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 52 |
52 |
50 |
2 |
0 |
0 |
0 |
91.0 |
+-- Gibbosula_rochechouartii **
+--------------------------------------+
| +-- Margaritifera_dahurica **
|
| +-- Quadrula_quadrula **
| +--------------------+
| | +-- Uniomerus_tetralasmus **
| |
| +--------------+ +-- Elliptio_complanata **
| | | +-----------------+
| | | | +-- Pleurobema_oviforme **
| | +--+
| | | +----------------- Amblema_plicata **
--+ | | |
| | +--+ +-------------- Popenaias_popeii **
| | | |
| | +--+ +----------- Toxolasma_parvum **
| | | |
| | +--+ +-- Potamilus_leptodon **
| | | +-----+
| | | | +-- Potamilus_alatus **
| | +--+
| | | +----- Leaunio_lienosus **
| | +--+
| | | +-- Lampsilis_cardium **
| | +--+
+--+ +-- Venustaconcha_ellipsiformis **
|
| +-- Chamberlainia_somsakpanhai **
| +--------------+
| | +-- Sinohyriopsis_cumingii **
| |
| | +----- Potomida_littoralis **
| +-----------------+ +--------+
| | | | | +-- Lamprotula_leaii **
| | | | +--+
| | | | +-- Pronodularia_japanensis **
| | +--+
| | | +-- Pseudodon_vondembuschianus **
| | | +--------+
| | | | +-- Pilsbryoconcha_exilis **
| | +--+
| | | +-------- Microcondylaea_bonellii **
| | +--+
| | | +----- Sinosolenaia_oleivora **
+--+ +--+
| | +-- Ptychorhynchus_pfisteri **
| +--+
| +-- Parvasolenaia_rivularis **
|
| +-------------------------------- Lepidodesma_languilati **
| |
| | +----------------- Lanceolaria_gladiola **
| | |
| | +-----------+ +-------------- Acuticosta_chinensis **
| | | | |
| | | | | +----- Cristaria_plicata **
+--+ | +--+ +-----+
| | | | | +-- Anemina_arcaeformis **
| | | | +--+
| | +--+ +-- Sinanodonta_woodiana **
| | |
| | | +-------- Anodonta_anatina **
| | +--+
+--+ | +----- Platynaias_compressa **
| +--+
| | +-- Pyganodon_grandis **
| +--+
| +-- Utterbackia_imbecillis **
|
| +-- Aculamprotula_scripta **
| +-----------------------+
| | +-- Aculamprotula_tientsinensis **
| |
| | +-- Unio_crassus **
+--+ +--------------------+
| | +-- Unio_delphinus **
| |
| | +-------- Pseudocuneopsis_capitata **
+--+ +-----------+
| | | +----- Pseudocuneopsis_yangshuoensis **
| | +--+
| | | +-- Pseudocuneopsis_yemaoi **
| | +--+
+--+ +-- Pseudocuneopsis_sichuanensis **
|
| +-- Nodularia_breviconcha **
| +--------------+
| | +-- Nodularia_douglasiae **
| |
+--+ +-- Schistodesmus_lampreyanus **
| +-----------+
| | +-- Schistodesmus_spinosus **
+--+
| +----------- Tchangsinaia_piscicula **
+--+
| +-------- Cuneopsis_rufescens **
+--+
| +----- Cuneopsis_celtiformis **
+--+
| +-- Cuneopsis_heudei **
+--+
+-- Cuneopsis_demangei **
Campbell & Lydeard, 2012 [Publ 3308], fig. 3
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 51 |
51 |
42 |
1 |
0 |
0 |
0 |
114.0 |
+-------------- Uniomerus_declivis **
|
+--------------------------------+ +-- Pustulosa_infucata **
| | +--------+
| | | +-- Pustulosa_kleiniana **
| +--+
| | +-------- Quadrula_quadrula **
| +--+
| | +----- Cyclonaias_tuberculata **
| +--+
| | +-- Pustulosa_succissa **
| +--+
| +-- Pustulosa_pustulosa **
--+
| +-------------- Elliptoideus_sloatianus **
| +-----------------------------+
| | | +----------- Plectomerus_dombeyanus **
| | +--+
| | | +-------- Obovaria_unicolor **
| | +--+
| | | +----- Obovaria_olivaria **
| | +--+
| | | +-- Lampsilis_ovata **
| | +--+
| | +-- Lemiox_rimosus **
+--+
| +----------- Cyrtonaias_tampicoensis **
| |
| | +----- Popenaias_popeii **
| +-----------------------------+ +--+
| | | | | +-- Amblema_elliottii **
| | | | +--+
| | +--+ +-- Amblema_plicata **
| | |
| | | +-- Reginaia_ebenus[1] **
| | | |
+--+ +-----+-- Reginaia_ebenus[2] **
| |
| +-- Reginaia_rotulata **
|
| +-- Elliptio_arca **
| +-----------------------------------+
| | +-- Elliptio_crassidens **
| |
+--+ +----------------------------------- Eurynia_dilatata **
| |
| | +-------------------------------- Parvaspina_collina **
| | |
+--+ | +-- Hemistena_lata **
| +-----------------------------+
| | +-- Plethobasus_cyphyus **
| |
+--+ +----- Pleuronaia_barnesiana **
| +-----------------------+
| | | +-- Pleuronaia_dolabelloides **
| | +--+
| | +-- Pleuronaia_gibber **
| |
| | +-- Pleurobema_stabile **
+--+ +-----+
| | +-- Pleurobema_sintoxia **
| +-----------------+
| | | +----- Pleurobema_decisum **
| | +--+
| | | +-- Pleurobema_clava **
+--+ +--+
| +-- Pleurobema_pyriforme **
|
| +----------------------- Pleurobema_cordatum **
| |
+--+ +-------------------- Fusconaia_subrotunda **
| |
| | +-- Fusconaia_cor **
+--+ +--------------+
| | +-- Fusconaia_cuneolus **
| |
+--+ +-------------- Fusconaia_ozarkensis **
| |
| | +-- Fusconaia_flava[6] **
| | +--------+
+--+ | +-- Fusconaia_masoni **
| |
| | +-- Fusconaia_escambia **
| | |
+--+ +-- Fusconaia_burkei[1] **
| +-----+
| | +-- Fusconaia_burkei[2] **
| | |
| | +-- Fusconaia_burkei[3] **
+--+
| +----- Fusconaia_flava[5] **
| |
| | +-- Fusconaia_cerina **
+--+ |
| +-- Fusconaia_flava[1] **
| |
+--+-- Fusconaia_flava[2] **
|
+-- Fusconaia_flava[3] **
|
+-- Fusconaia_flava[4] **
Zieritz et al., 2021 [Publ 4231], fig. 3-F, 5
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 49 |
49 |
47 |
2 |
0 |
0 |
0 |
82.0 |
+----- Pseudunio_marocanus **
+--------------------------+
| | +-- Margaritifera_falcata **
| +--+
| +-- Margaritifera_dahurica **
|
| +----------- Quadrula_quadrula **
| |
| +-----------------+ +-------- Toxolasma_parvum **
| | | |
| | +--+ +-- Potamilus_alatus **
| | | +--+
| | | | +-- Potamilus_leptodon **
--+ | +--+
| | | +-- Venustaconcha_ellipsiformis **
| | +--+
| | +-- Lampsilis_ornata **
| |
| | +----- Chamberlainia_somsakpanhai **
| | +--------------+
| | | | +-- Sinohyriopsis_cumingii **
| | | +--+
| | | +-- Sinohyriopsis_schlegelii **
| | |
| | | +-- Lens_contradens **
+--+ +-----+ +--+
| | | | +-- Physunio_superbus **
| | | +-----------+
| | | | | +-- Hyriopsis_bialata **
| | | | +--+
| | | | +-- Rectidens_sumatrensis **
| | | |
| | +--+ +-------- Potomida_littoralis **
| | | +-----+
| | | | | +----- Pronodularia_japanensis **
| | | | +--+
| | | | | +-- Lamprotula_caveata **
| | | | +--+
| | +--+ +-- Lamprotula_leaii **
| | |
| | | +-- Pilsbryoconcha_exilis **
| | | +--------+
+--+ | | +-- Pseudodon_vondembuschianus **
| +--+
| | +-------- Ptychorhynchus_pfisteri **
| +--+
| | +----- Microcondylaea_bonellii **
| +--+
| | +-- Sinosolenaia_recognita **
| +--+
| +-- Sinosolenaia_carinata **
|
| +----------------------- Lepidodesma_languilati **
| |
| | +----- Aculamprotula_tortuosa **
| | +--------+
| | | | +-- Aculamprotula_tientsinensis **
| | | +--+
| | | +-- Aculamprotula_coreana **
| | +-----+
+--+ | | +----- Nodularia_douglasiae **
| | | +-----+
| | | | | +-- Schistodesmus_lampreyanus **
| | | | +--+
| | +--+ +-- Tchangsinaia_piscicula **
| | |
| | | +-------- Unio_tumidus **
| | +--+
| | | +----- Unio_crassus **
+--+ +--+
| | +-- Unio_pictorum **
| +--+
| +-- Unio_delphinus **
|
| +-- Lanceolaria_grayii **
| +--------------+
| | +-- Lanceolaria_lanceolata **
| |
| | +----------- Anodonta_anatina **
+--+ +--+
| | | +-------- Platynaias_compressa **
| | +--+
| | | +----- Pyganodon_grandis **
| | +--+
+--+ | +-- Utterbackia_imbecillis **
| +--+
| +-- Utterbackia_peninsularis **
|
| +-------- Cristaria_plicata **
| |
+-----+ +-- Sinanodonta_lucida **
| +--+
| | +-- Sinanodonta_woodiana **
+--+
| +-- Anemina_arcaeformis **
+--+
+-- Anemina_euscaphys **
Bolotov et al., 2022 [Publ 4250], fig. 8
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 49 |
49 |
42 |
2 |
0 |
0 |
0 |
119.0 |
+-- Gibbosula_laosensis **
|
+-- Margaritifera_dahurica **
|
+-- Sinosolenaia_iridinea **
|
+-----------------------------+-- Leguminaia_wheatleyi **
| |
| +-- Gonidea_angulata **
| |
| +-- Lamprotula_leaii **
| |
| +-- Potomida_littoralis **
|
| +-- Pseudodon_lenyanensis **
| +--------+
| | +-- Sundadontina_tanintharyiensis **
| +--------+
| | | +-------- Indopseudodon_kayinensis **
| | +--+
| | | +----- Indopseudodon_indawgyiensis **
| | +--+
| | | +-- Indopseudodon_salwenianus **
| | +--+
| +--------+ +-- Indopseudodon_bogani **
--+ | |
| | | +----- Trapezoideus_mitanensis **
| | | +-----------+
| | | | | +-- Trapezoideus_foliaceus **
| | | | +--+
| | | | +-- Trapezoideus_lenya **
| | +--+
| | | +-- Yaukthwa_elongatula **
| | | +-----------+
| | | | +-- Yaukthwa_baniensis **
| | | |
| | +--+ +----- Yaukthwa_inlenensis **
| | | +-----+
| | | | | +-- Yaukthwa_paiensis **
| | | | +--+
| | +--+ +-- Yaukthwa_avaensis **
| | |
| | | +-------- Yaukthwa_panhai **
| | +--+
+--+ | +----- Yaukthwa_rectangularis **
| +--+
| | +-- Yaukthwa_nesemanni **
| +--+
| +-- Yaukthwa_zayleymanensis **
|
| +-------- Lamellidens_savadiensis **
| +-----+
| | | +----- Lamellidens_ferrugineus **
| | +--+
| | | +-- Lamellidens_generosus **
| +-----------+ +--+
| | | +-- Lamellidens_mainwaringi **
| | |
| | | +----------- Trapezidens_exolescens **
| | +--+
| | | +-------- Trapezidens_scutum **
| | +--+
| | | +----- Trapezidens_yeti **
| | +--+
+--+ | +-- Trapezidens_angustior **
| +--+
| +-- Trapezidens_dolichorhynchus **
|
| +----------------------- Parreysia_rakhinensis **
| |
| | +-- Leoparreysia_whitteni **
| | +-----+
| | | +-- Leoparreysia_burmana **
+--+ +-----------+
| | | +----- Leoparreysia_olivacea **
| | +--+
| | | +-- Leoparreysia_canefrii **
| | +--+
+--+ +-- Leoparreysia_tavoyensis **
|
| +-- Indonaia_andersoniana **
| +--------------+
| | +-- Indonaia_subclathrata **
+--+
| +-------------- Indochinella_pugio **
| |
+--+ +----------- Radiatula_crispisulcata **
| |
+--+ +-- Radiatula_chaudhurii **
| +-----+
| | +-- Radiatula_myitkyinae **
+--+
| +----- Radiatula_mouhoti **
+--+
| +-- Radiatula_chindwinensis **
+--+
+-- Radiatula_myitthanensis **
Sano et al., 2020 [Publ 4138], fig. 1
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 47 |
47 |
16 |
2 |
1 |
0 |
0 |
95.0 |
+----- Gibbosula_rochechouartii **
+-----------+
| | +-- Margaritifera_middendorffi **
| +--+
| +-- Margaritifera_laevis **
|
| +-------- Sinosolenaia_iridinea **
| |
| +--+ +----- Inversidens_brandtii **
| | | |
| | +--+ +-- Obovalis_omiensis[1] **
| | | |
| | +--+-- Obovalis_omiensis[2] **
| +--+ |
| | | +-- Ptychorhynchus_pfisteri **
| | |
| | | +-- Lamprotula_caveata **
--+ | | +--+
| | | | +-- Pronodularia_japanensis **
| | +-----+
| | | +-- Sinohyriopsis_schlegelii[1] **
| | | |
| | +--+-- Sinohyriopsis_schlegelii[2] **
| | |
| | +-- Sinohyriopsis_cumingii **
| |
| | +-------- Lanceolaria_grayii[1] **
| | |
| | +-------- Lanceolaria_grayii[2] **
| | |
| | +-------- Cristaria_plicata[1] **
| | |
| | +-------- Cristaria_plicata[2] **
+--+ |
| +-------- Sinanodonta_lauta **
| |
| +-------- Beringiana_japonica **
| |
| +-------- Beringiana_fukuharai[1] **
| |
| +-------- Beringiana_beringiana **
| |
| +-------- Beringiana_fukuharai[2] **
| |
| +-------- Beringiana_fukuharai[3] **
| |
| +-------- Buldowskia_iwakawai[1] **
| |
| +-------- Buldowskia_iwakawai[2] **
| |
| +-------- Sinanodonta_tumens[1] **
| |
| +-------- Sinanodonta_tumens[2] **
| |
+-----+-------- Pletholophus_tenuis **
|
+-------- Anemina_arcaeformis **
|
+-------- Lanceolaria_lanceolata **
|
+-------- Cristaria_plicata[3] **
|
+-------- Lanceolaria_gladiola **
|
+-------- Sinanodonta_angula **
|
+-------- Lepidodesma_languilati **
|
| +-- Aculamprotula_tortuosa **
+-----+
| +-- Acuticosta_chinensis **
|
| +----- Cuneopsis_celtiformis **
| |
+--+ +-- Nodularia_douglasiae[1] **
| | |
| +--+-- Nodularia_douglasiae[2] **
| |
| +-- Nodularia_nipponensis **
|
| +----- Schistodesmus_lampreyanus **
| |
| | +-- Inversiunio_reinianus[1] **
+--+ |
| +-- Inversiunio_reinianus[2] **
| |
| +-- Inversiunio_jokohamensis[1] **
+--+
+-- Inversiunio_jokohamensis[2] **
|
+-- Inversiunio_jokohamensis[3] **
|
+-- Inversiunio_jokohamensis[4] **
Wu et al., 2020 [Publ 4121], fig. 3
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 46 |
46 |
44 |
2 |
4 |
3 |
2 |
87.0 |
+-------------------------------------- Echyridella_menziesii
|
| +-- Anodontites_trapesialis
| +--------------------------------+
| | +-- Mutela_dubia
--+ |
| | +-------- Gibbosula_rochechouartii **
| | +-----------------------+
| | | | +----- Cumberlandia_monodonta **
| | | +--+
+--+ | | +-- Margaritifera_falcata **
| | +--+
| | +-- Margaritifera_dahurica **
| |
| | +----------- Quadrula_quadrula **
| | |
| | +-----------------+ +-------- Toxolasma_parvum **
+--+ | | |
| | +--+ +-- Venustaconcha_ellipsiformis **
| | | +--+
| | | | +-- Lampsilis_ornata **
| | +--+
| | | +-- Potamilus_alatus **
| | +--+
| | +-- Potamilus_leptodon **
| |
| | +-- Sinohyriopsis_schlegelii **
| | +--------+
+--+ | +-- Sinohyriopsis_cumingii **
| |
| +--------------+ +----- Potomida_littoralis **
| | | +--+
| | | | | +-- Pronodularia_japanensis **
| | | | +--+
| | +--+ +-- Lamprotula_leaii **
| | |
| | | +----- Ptychorhynchus_pfisteri **
| | +--+
| | | +-- Sinosolenaia_oleivora **
| | +--+
| | +-- Sinosolenaia_carinata **
+--+
| +----------------------- Lepidodesma_languilati **
| |
| | +----- Acuticosta_chinensis **
| | +-----------+
| | | | +-- Lanceolaria_lanceolata **
| | | +--+
| | | +-- Lanceolaria_grayii **
| | |
| | +--+ +----------- Anodonta_anatina **
| | | | +--+
| | | | | | +-------- Platynaias_compressa **
+--+ | | | +--+
| | | | | +----- Pyganodon_grandis **
| | | | +--+
| | +--+ | +-- Utterbackia_peninsularis **
| | | +--+
| | | +-- Utterbackia_imbecillis **
| | |
| | | +-------- Cristaria_plicata **
| | | |
| | +-----+ +-- Sinanodonta_woodiana **
| | | +--+
+--+ | | +-- Sinanodonta_lauta **
| +--+
| | +-- Anemina_arcaeformis **
| +--+
| +-- Anemina_euscaphys **
|
| +-------- Aculamprotula_tortuosa **
| |
| +--------+ +-- Aculamprotula_coreana **
| | | +--+
| | | | +-- Aculamprotula_tientsinensis **
| | +--+
| | | +-- Aculamprotula_polysticta **
| | +--+
+--+ +-- Aculamprotula_scripta **
|
| +-------- Unio_tumidus **
| +-----+
| | | +----- Unio_crassus **
| | +--+
| | | +-- Unio_delphinus **
+--+ +--+
| +-- Unio_pictorum **
|
| +----------- Pseudocuneopsis_capitata **
+--+
| +-------- Nodularia_douglasiae **
+--+
| +----- Tchangsinaia_piscicula **
+--+
| +-- Cuneopsis_heudei **
+--+
+-- Schistodesmus_lampreyanus **
Wu et al., 2019 [Publ 3864], fig. 3
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 45 |
47 |
45 |
2 |
2 |
2 |
2 |
88.0 |
+-- Echyridella_menziesii
+--------+
| +-- Mutela_dubia
+--------------------+
| | +-------- Gibbosula_rochechouartii **
| +--+
| | +----- Cumberlandia_monodonta **
| +--+
| | +-- Margaritifera_dahurica **
| +--+
| +-- Margaritifera_falcata **
|
| +----------- Quadrula_quadrula **
| |
--+ +-----------------+ +-------- Toxolasma_parvum **
| | | |
| | +--+ +-- Lampsilis_ornata **
| | | +--+
| | | | +-- Venustaconcha_ellipsiformis **
| | +--+
| | | +-- Potamilus_leptodon **
| | +--+
| | +-- Potamilus_alatus **
| |
| | +-- Sinohyriopsis_cumingii **
| | +--------+
+--+ | +-- Sinohyriopsis_schlegelii **
| |
| +--------------+ +----- Potomida_littoralis **
| | | +--+
| | | | | +-- Lamprotula_leaii **
| | | | +--+
| | +--+ +-- Pronodularia_japanensis **
| | |
| | | +----- Ptychorhynchus_pfisteri **
| | +--+
| | | +-- Sinosolenaia_oleivora **
| | +--+
| | +-- Sinosolenaia_carinata **
+--+
| +----------------------- Lepidodesma_languilati **
| |
| | +-------- Aculamprotula_tortuosa **
| | +--------+
| | | | +----- Aculamprotula_scripta **
| | | +--+
| | | | +-- Aculamprotula_tientsinensis **
| | | +--+
| | | +-- Aculamprotula_coreana **
| | +--+
+--+ | | +-------- Unio_tumidus **
| | | +-----+
| | | | | +----- Unio_crassus **
| | | | +--+
| | | | | +-- Unio_delphinus **
| | +--+ +--+
| | | +-- Unio_pictorum **
| | |
| | | +----------- Pseudocuneopsis_capitata **
| | +--+
| | | +-------- Nodularia_douglasiae **
+--+ +--+
| | +----- Tchangsinaia_piscicula **
| +--+
| | +-- Cuneopsis_heudei **
| +--+
| +-- Schistodesmus_lampreyanus **
|
| +----- Acuticosta_chinensis **
| +-----------+
| | | +-- Lanceolaria_grayii **
| | +--+
| | +-- Lanceolaria_lanceolata **
| |
+--+ +----------- Anodonta_anatina **
| +--+
| | | +-------- Platynaias_compressa **
| | +--+
| | | +----- Pyganodon_grandis **
| | +--+
+--+ | +-- Utterbackia_peninsularis **
| +--+
| +-- Utterbackia_imbecillis **
|
| +-------- Cristaria_plicata **
| |
+-----+ +-- Anemina_arcaeformis **
| +--+
| | +-- Anemina_euscaphys **
+--+
| +-- Sinanodonta_lucida **
+--+
+-- Sinanodonta_woodiana **
Froufe et al., 2019 [Publ 3936], fig. 3
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 45 |
45 |
43 |
2 |
0 |
0 |
0 |
77.0 |
+----- Pseudunio_marocanus **
+--------------------------+
| | +-- Margaritifera_dahurica **
| +--+
| +-- Margaritifera_falcata **
|
| +----------- Quadrula_quadrula **
| |
| +-----------------+ +-------- Toxolasma_parvum **
| | | |
| | +--+ +-- Lampsilis_ornata **
| | | +--+
--+ | | | +-- Venustaconcha_ellipsiformis **
| | +--+
| | | +-- Potamilus_leptodon **
| | +--+
| | +-- Potamilus_alatus **
| |
| | +----- Chamberlainia_somsakpanhai **
| | +-----------+
| | | | +-- Sinohyriopsis_cumingii **
| | | +--+
| | | +-- Sinohyriopsis_schlegelii **
+--+ |
| +--------+ +-------- Potomida_littoralis **
| | | +-----+
| | | | | +----- Pronodularia_japanensis **
| | | | +--+
| | | | | +-- Lamprotula_leaii **
| | | | +--+
| | +--+ +-- Lamprotula_caveata **
| | |
| | | +-- Pilsbryoconcha_exilis **
| | | +--------+
| | | | +-- Pseudodon_vondembuschianus **
| | +--+
| | | +-------- Ptychorhynchus_pfisteri **
| | +--+
+--+ | +----- Microcondylaea_bonellii **
| +--+
| | +-- Sinosolenaia_carinata **
| +--+
| +-- Sinosolenaia_oleivora **
|
| +----------------------- Lepidodesma_languilati **
| |
| | +----- Aculamprotula_tortuosa **
| | +--------+
| | | | +-- Aculamprotula_tientsinensis **
| | | +--+
| | | +-- Aculamprotula_coreana **
| | +-----+
+--+ | | +----- Nodularia_douglasiae **
| | | +-----+
| | | | | +-- Tchangsinaia_piscicula **
| | | | +--+
| | +--+ +-- Schistodesmus_lampreyanus **
| | |
| | | +-------- Unio_tumidus **
| | +--+
| | | +----- Unio_crassus **
+--+ +--+
| | +-- Unio_pictorum **
| +--+
| +-- Unio_delphinus **
|
| +-- Lanceolaria_lanceolata **
| +--------------+
| | +-- Lanceolaria_grayii **
| |
| | +-------- Cristaria_plicata **
+--+ |
| +-----+ +-- Anemina_arcaeformis **
| | | +--+
| | | | +-- Anemina_euscaphys **
| | +--+
+--+ | +-- Sinanodonta_lucida **
| +--+
| +-- Sinanodonta_woodiana **
|
| +----------- Anodonta_anatina **
+--+
| +-------- Lasmigona_complanata **
+--+
| +----- Pyganodon_grandis **
+--+
| +-- Utterbackia_peninsularis **
+--+
+-- Utterbackia_imbecillis **
Froufe et al., 2019 [Publ 3936], fig. 6
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 45 |
45 |
43 |
2 |
0 |
0 |
0 |
77.0 |
+----- Pseudunio_marocanus **
+--------------------------+
| | +-- Margaritifera_dahurica **
| +--+
| +-- Margaritifera_falcata **
|
| +----------- Quadrula_quadrula **
| |
| +-----------------+ +-------- Toxolasma_parvum **
| | | |
| | +--+ +-- Potamilus_leptodon **
| | | +--+
--+ | | | +-- Potamilus_alatus **
| | +--+
| | | +-- Lampsilis_ornata **
| | +--+
| | +-- Venustaconcha_ellipsiformis **
| |
| | +----- Chamberlainia_somsakpanhai **
| | +-----------+
| | | | +-- Sinohyriopsis_cumingii **
| | | +--+
| | | +-- Sinohyriopsis_schlegelii **
+--+ |
| +--------+ +-------- Potomida_littoralis **
| | | +-----+
| | | | | +----- Pronodularia_japanensis **
| | | | +--+
| | | | | +-- Lamprotula_leaii **
| | | | +--+
| | +--+ +-- Lamprotula_caveata **
| | |
| | | +-- Pilsbryoconcha_exilis **
| | | +--------+
| | | | +-- Pseudodon_vondembuschianus **
| | +--+
| | | +-------- Ptychorhynchus_pfisteri **
| | +--+
+--+ | +----- Microcondylaea_bonellii **
| +--+
| | +-- Sinosolenaia_carinata **
| +--+
| +-- Sinosolenaia_oleivora **
|
| +----------------------- Lepidodesma_languilati **
| |
| | +----- Aculamprotula_tortuosa **
| | +--------+
| | | | +-- Aculamprotula_tientsinensis **
| | | +--+
| | | +-- Aculamprotula_coreana **
| | +-----+
+--+ | | +----- Nodularia_douglasiae **
| | | +-----+
| | | | | +-- Tchangsinaia_piscicula **
| | | | +--+
| | +--+ +-- Schistodesmus_lampreyanus **
| | |
| | | +-------- Unio_tumidus **
| | +--+
| | | +----- Unio_crassus **
+--+ +--+
| | +-- Unio_pictorum **
| +--+
| +-- Unio_delphinus **
|
| +-- Lanceolaria_lanceolata **
| +--------------+
| | +-- Lanceolaria_grayii **
| |
| | +----------- Anodonta_anatina **
+--+ +--+
| | | +-------- Lasmigona_complanata **
| | +--+
| | | +----- Pyganodon_grandis **
| | +--+
+--+ | +-- Utterbackia_peninsularis **
| +--+
| +-- Utterbackia_imbecillis **
|
| +-------- Cristaria_plicata **
| |
+-----+ +-- Sinanodonta_lucida **
| +--+
| | +-- Sinanodonta_woodiana **
+--+
| +-- Anemina_arcaeformis **
+--+
+-- Anemina_euscaphys **
Dai et al., 2023 [Publ 4285], fig. 3-BI
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 45 |
45 |
41 |
2 |
0 |
0 |
0 |
93.0 |
+-- Gibbosula_laosensis **
+-----------------------+
| +-- Margaritifera_margaritifera **
|
| +----- Indopseudodon_indawgyiensis **
| +--------+
| | | +-- Indopseudodon_bogani **
| | +--+
| | +-- Indopseudodon_salwenianus **
| |
| | +-------- Nyeinchanconcha_nyeinchani **
| | +--+
| | | | +----- Bineurus_loeiensis **
| +--------+ | +--+
| | | | | +-- Bineurus_mouhotii **
| | | | +--+
--+ | | | +-- Bineurus_exilis **
| | | |
| | | | +-- Thaiconcha_callifera **
| | | | +-----+
| | | | | +-- Thaiconcha_munelliptica **
| | +--+--+
| | | | +----- Sundadontina_tanintharyiensis **
| | | +--+
| | | | +-- Sundadontina_brandti **
| | | +--+
| | | +-- Sundadontina_taskaevi **
| | |
| | | +-- Pilsbryoconcha_carinifera **
| | | +-----+
| | | | +-- Pilsbryoconcha_exilis **
| | +--+
+--+ | +----- Pseudodon_lenyanensis **
| +--+
| | +-- Pseudodon_mekongi **
| +--+
| +-- Pseudodon_vagulus **
|
| +-- Hyriopsis_bialata **
| +-----+
| | +-- Ensidens_ingallsianus **
| +-----------+
| | | +----- Physunio_superbus **
| | +--+
| | | +-- Lens_contradens **
| | +--+
| | +-- Lens_comptus **
| |
| | +-- Inversidens_brandtii **
| | +--------------+
| | | +-- Inversidens_rentianensis **
+--+ |
| | +----- Chamberlainia_somsakpanhai **
| | +-----+
| | | | +-- Sinohyriopsis_cumingii **
| | | +--+
| +-----+ +-- Sinohyriopsis_schlegelii **
| | |
| | | +-------- Potomida_littoralis **
| | +--+
| | | +----- Pronodularia_japanensis **
+--+ +--+
| | +-- Lamprotula_leaii **
| +--+
| +-- Lamprotula_caveata **
|
| +-------- Sinosolenaia_carinata **
| +-----+
| | | +----- Sinosolenaia_recognita **
| | +--+
| | | +-- Sinosolenaia_iridinea **
| | +--+
| | +-- Sinosolenaia_oleivora **
+--+
| +----- Gonidea_angulata **
| +-----+
| | | +-- Leguminaia_wheatleyi **
| | +--+
| | +-- Microcondylaea_bonellii **
+--+
| +-- Koreosolenaia_sitgyensis **
| +-----+
| | +-- Parvasolenaia_rivularis **
+--+
| +----- Postolata_guangxiensis **
+--+
| +-- Ptychorhynchus_pfisteri **
+--+
+-- Obovalis_omiensis **
Dai et al., 2023 [Publ 4285], fig. 3-ML
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 45 |
45 |
43 |
2 |
0 |
0 |
0 |
93.0 |
+-- Gibbosula_laosensis **
+-----------------------+
| +-- Margaritifera_margaritifera **
|
| +----- Indopseudodon_indawgyiensis **
| +-----------+
| | | +-- Indopseudodon_bogani **
| | +--+
| | +-- Indopseudodon_salwenianus **
| |
| | +-------- Nyeinchanconcha_nyeinchani **
| +-----+ +-----+
| | | | | +----- Bineurus_loeiensis **
| | | | +--+
| | | | | +-- Bineurus_mouhotii **
| | | | +--+
--+ | | | +-- Bineurus_exilis **
| | | |
| | +--+ +-- Thaiconcha_callifera **
| | | +-----+
| | | | +-- Thaiconcha_munelliptica **
| | | +--+
| | | | | +----- Sundadontina_tanintharyiensis **
| | | | +--+
| | | | | +-- Sundadontina_brandti **
| | | | +--+
| | +--+ +-- Sundadontina_taskaevi **
| | |
| | | +-- Pilsbryoconcha_carinifera **
| | | +-----+
| | | | +-- Pilsbryoconcha_exilis **
+--+ +--+
| | +----- Pseudodon_lenyanensis **
| +--+
| | +-- Pseudodon_mekongi **
| +--+
| +-- Pseudodon_vagulus **
|
| +-- Hyriopsis_bialata **
| +-----+
| | +-- Ensidens_ingallsianus **
| +-----------+
| | | +----- Physunio_superbus **
| | +--+
| | | +-- Lens_contradens **
| | +--+
| | +-- Lens_comptus **
| |
| | +-- Inversidens_brandtii **
| | +-----------+
| | | +-- Inversidens_rentianensis **
+--+ |
| +--+ +----- Chamberlainia_somsakpanhai **
| | | +-----+
| | | | | +-- Sinohyriopsis_cumingii **
| | | | +--+
| | +--+ +-- Sinohyriopsis_schlegelii **
| | |
| | | +-------- Potomida_littoralis **
| | +--+
| | | +----- Pronodularia_japanensis **
| | +--+
+--+ | +-- Lamprotula_leaii **
| +--+
| +-- Lamprotula_caveata **
|
| +-------- Sinosolenaia_carinata **
| +-----+
| | | +----- Sinosolenaia_recognita **
| | +--+
| | | +-- Sinosolenaia_iridinea **
| | +--+
| | +-- Sinosolenaia_oleivora **
+--+
| +----- Gonidea_angulata **
| +-----+
| | | +-- Leguminaia_wheatleyi **
| | +--+
| | +-- Microcondylaea_bonellii **
+--+
| +-- Koreosolenaia_sitgyensis **
| +-----+
| | +-- Parvasolenaia_rivularis **
+--+
| +----- Postolata_guangxiensis **
+--+
| +-- Ptychorhynchus_pfisteri **
+--+
+-- Obovalis_omiensis **
Burlakova et al., 2012 [Publ 3486], fig. 1
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 44 |
44 |
42 |
1 |
0 |
0 |
0 |
114.0 |
+-- Amblema_elliottii **
+--------------------------------------------+
| +-- Amblema_plicata **
|
--+ +-- Lampsilis_ovata **
| +-----------------------------------------+
| | +-- Plectomerus_dombeyanus **
| |
+--+ +----------------------------------------- Popenaias_popeii **
| |
| | +-------------------------------------- Cyrtonaias_tampicoensis **
+--+ |
| | +----------------------------------- Parvaspina_collina **
| | |
+--+ | +----- Plethobasus_cyphyus **
| | +--------------------------+
| | | | +-- Elliptio_arca **
| | | +--+
| | | +-- Elliptio_crassidens **
+--+ |
| | +-------------------- Fusconaia_subrotunda **
| | |
| | | +-- Fusconaia_cor **
| | +--------+ +--------------+
| | | | | +-- Fusconaia_cuneolus **
| | | | |
+--+ | +--+ +-------------- Fusconaia_ozarkensis **
| | | |
| | | | +-------- Fusconaia_askewi[3] **
| | | | +--+
| | +--+ | | +----- Fusconaia_askewi[2] **
| | | | +--+
| | | | | +-- Fusconaia_lananensis **
| | | | +--+
| | +--+ +-- Fusconaia_askewi[1] **
| | |
| | | +-- Fusconaia_burkei **
| | | +-----+
+--+ | | +-- Fusconaia_escambia **
| +--+
| | +----- Fusconaia_masoni **
| +--+
| | +-- Fusconaia_flava **
| +--+
| +-- Fusconaia_cerina **
|
| +-- Eurynia_dilatata **
| +-----------------------+
| | +-- Hemistena_lata **
| |
| | +----- Pleuronaia_gibber **
| | +-----+
| | | | +-- Pleuronaia_dolabelloides **
+--+ | +--+
| +-----------+ +-- Pleuronaia_barnesiana **
| | |
| | | +-------- Pleurobema_riddellii **
| | +--+
| | | +----- Pleurobema_sintoxia **
| | +--+
+--+ | +-- Pleurobema_rubrum **
| +--+
| +-- Pleurobema_cordatum **
|
| +-------------------- Pleurobema_stabile **
| |
+--+ +----------------- Pleurobema_pyriforme **
| |
| | +----- Pleurobema_georgianum **
+--+ +--------+
| | | +-- Pleurobema_hanleyianum **
| | +--+
| | +-- Pleurobema_decisum **
+--+
| +-- Pleurobema_clava **
| +--------+
| | +-- Pleurobema_oviforme **
+--+
| +-------- Pleurobema_beadleianum **
| |
+--+ +-- Pleurobema_rubellum **
| +--+
| | +-- Pleurobema_strodeanum **
+--+
| +-- Pleurobema_perovatum **
+--+
+-- Pleurobema_taitianum **
Huang et al., 2019 [Publ 3843], fig. 1
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 44 |
44 |
41 |
2 |
0 |
0 |
0 |
96.0 |
+-- Margaritifera_dahurica **
+--------------------------------------+
| +-- Gibbosula_rochechouartii **
|
| +----- Lamprotula_leaii **
| |
| +-----------+ +-- Lamprotula_caveata[1] **
| | | |
| | +--+-- Lamprotula_cornuumlunae **
| | |
--+ +--------------------+ +-- Lamprotula_caveata[2] **
| | |
| | | +-- Parvasolenaia_rivularis **
| | | +-----------+
| | | | +-- Parvasolenaia_triangularis **
| | +--+
| | | +----------- Ptychorhynchus_pfisteri **
| | +--+
| | | +-------- Sinosolenaia_carinata **
| | +--+
+--+ | +----- Sinosolenaia_recognita **
| +--+
| | +-- Sinosolenaia_oleivora **
| +--+
| +-- Sinosolenaia_iridinea **
|
| +----- Sinohyriopsis_cumingii[1] **
| +-----------------------------+
| | | +-- Sinohyriopsis_cumingii[2] **
| | +--+
| | +-- Sinohyriopsis_schlegelii **
+--+
| +-------------------------------- Anemina_arcaeformis **
| |
| | +----------------------------- Acuticosta_chinensis **
| | |
+--+ | +-- Sinanodonta_angula **
| | +--------+
| | | +-- Sinanodonta_lucida **
| | +--------------+
| | | | +-------- Sinanodonta_jourdyi **
+--+ | +--+
| | | +----- Sinanodonta_woodiana[2] **
| | +--+
| | | +-- Sinanodonta_elliptica **
| | +--+
| | +-- Sinanodonta_woodiana[1] **
| |
+--+ +----------- Aculamprotula_tortuosa **
| +-----------+
| | | +-------- Aculamprotula_scripta **
| | +--+
| | | +----- Aculamprotula_coreana **
| | +--+
| | | +-- Aculamprotula_fibrosa **
| | +--+
| | +-- Aculamprotula_tientsinensis **
+--+
| +----------- Cristaria_plicata **
| +--------+
| | | +-------- Lanceolaria_gladiola **
| | +--+
| | | +----- Lanceolaria_grayii **
| | +--+
| | | +-- Lanceolaria_lanceolata **
+--+ +--+
| +-- Lanceolaria_triformis **
|
| +----------------- Lepidodesma_languilati **
| |
| | +-- Schistodesmus_lampreyanus **
+--+ +-----------+
| | +-- Schistodesmus_spinosus **
| |
+--+ +----- Nodularia_douglasiae[1] **
| +-----+
| | | +-- Nodularia_douglasiae[2] **
| | +--+
+--+ +-- Nodularia_nuxpersicae **
|
| +-------- Cuneopsis_rufescens **
+--+
| +----- Tchangsinaia_piscicula **
+--+
| +-- Cuneopsis_heudei **
+--+
+-- Cuneopsis_celtiformis **
Inoue et al., 2020 [Publ 3941], fig. 3
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 44 |
44 |
42 |
1 |
0 |
0 |
0 |
110.0 |
+----------------------------------------------- Arcidens_confragosus **
|
| +-- Pustulosa_petrina **
| +-----------------------------------------+
| | +-- Quadrula_quadrula **
--+ |
| | +-------------- Parvaspina_collina **
| | |
| | | +-- Eurynia_dilatata **
| | +--------------------------+ +--------+
| | | | | +-- Hemistena_lata **
+--+ | | |
| | +--+ +-- Pleurobema_clava **
| | | +-----+
| | | | +-- Pleurobema_sintoxia **
| | +--+
| | | +----- Pleuronaia_dolabelloides **
| | +--+
| | | +-- Fusconaia_flava **
+--+ +--+
| +-- Fusconaia_subrotunda **
|
| +-- Amblema_plicata **
| +--+
| | +-- Amblema_elliottii **
| +--------------------------------+
| | | +-- Reginaia_ebenus **
| | +--+
| | +-- Reginaia_rotulata **
| |
+--+ +----------------------------------- Toxolasma_lividum **
| |
| | +-------- Potamilus_fragilis **
| | |
| | +-----------------------+ +-- Potamilus_ohiensis **
+--+ | | +--+
| | | | +-- Potamilus_amphichaenus **
| | +--+
| | | +-- Potamilus_metnecktayi **
| | +--+
| | +-- Potamilus_purpuratus **
+--+
| +-------- Ellipsaria_lineolata **
| +--------------------+
| | | +----- Truncilla_macrodon **
| | +--+
| | | +-- Truncilla_donaciformis **
| | +--+
| | +-- Truncilla_cognata **
+--+
| +-------------------------- Leaunio_lienosus **
| |
| | +-------------------- Lampsilis_bracteata **
| | |
| | +--+ +----------------- Lampsilis_fasciola **
| | | | |
+--+ | +--+ +-------------- Lampsilis_ornata **
| | | |
| | +--+ +-- Lampsilis_ovata **
| | | +--------+
| | | | +-- Lampsilis_cariosa **
| | +--+
| | | +-------- Lampsilis_cardium[3] **
+--+ +--+
| | +----- Lampsilis_cardium[2] **
| +--+
| | +-- Lampsilis_satura **
| +--+
| +-- Lampsilis_cardium[1] **
|
| +-- Lampsilis_higginsii **
| +--------------+
| | +-- Ortmanniana_ligamentina **
+-----+
| +-------------- Lampsilis_teres **
| |
+--+ +-- Lampsilis_radiata **
| +--------+
| | +-- Lampsilis_siliquoidea **
+--+
| +-------- Lampsilis_bergmanni **
+--+
| +----- Lampsilis_hydiana **
+--+
| +-- Lampsilis_virescens **
+--+
+-- Lampsilis_straminea **
Whelan et al., 2023 [Publ 4287], fig. 2
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 44 |
44 |
42 |
1 |
0 |
0 |
0 |
106.0 |
+-------- Anemina_arcaeformis **
+-----------------------------+
| | +----- Sinanodonta_lucida **
| +--+
| | +-- Sinanodonta_woodiana **
| +--+
--+ +-- Sinanodonta_tumens **
|
| +----------------------------------- Anodonta_cygnea **
| |
| | +-- Anodonta_nuttalliana **
| | +-----------------------------+
+--+ | +-- Pseudanodonta_complanata **
| |
| | +----------------------------- Anodonta_anatina **
| | |
| | | +----- Utterbackia_peninsularis[2] **
+--+ | +--+
| | | | +-- Utterbackia_peninsularis[1] **
| | | +--+
| | +-----------------+ +-- Utterbackia_imbecillis **
| | | |
| | | | +----- Pyganodon_grandis[2] **
+--+ | +--+
| | | +-- Pyganodon_grandis[1] **
| | +--+
| | +-- Pyganodon_lacustris **
| |
| | +-- Utterbackiana_couperiana **
| | +--+
| | | +-- Utterbackiana_heardi **
| | +-----+
| | | | +-- Utterbackiana_suborbiculata **
+--+ | +--+
| | +-- Utterbackiana_hartfieldorum **
| +-----------+
| | | +----- Lasmigona_costata **
| | | +--+
| | | | | +-- Pseudodontoideus_subvexus **
| | | | +--+
| | +--+ +-- Pseudodontoideus_connasaugaensis **
| | |
| | | +----- Lasmigona_complanata **
| | +--+
| | | +-- Arcidens_confragosus **
| | +--+
| | +-- Arcidens_wheeleri **
+--+
| +----------------- Pressodonta_viridis **
| |
| +--+ +-- Anodontoides_denigratus **
| | | +-----------+
| | | | +-- Anodontoides_ferussacianus **
| | +--+
| | | +----------- Strophitus_undulatus **
| | +--+
| | | +-------- Strophitus_pascagoulaensis[2] **
| | +--+
| | | +----- Strophitus_radiatus **
+--+ +--+
| | +-- Strophitus_pascagoulaensis[1] **
| +--+
| +-- Strophitus_williamsi **
|
| +----- Prolasmidonta_heterodon **
| +-----------+
| | | +-- Platynaias_subviridis **
| | +--+
| | +-- Platynaias_compressa **
+--+
| +-- Alasmidonta_holstonia **
| +-----------+
| | +-- Alasminota_etowaensis **
| |
+--+ +-- Alasmidonta_raveneliana **
| +--------+
| | +-- Alasmidonta_marginata **
| |
+--+ +-- Alasmidonta_varicosa **
| +-----+
| | +-- Alasmidonta_uwharriensis **
+--+
| +----- Alasmidonta_undulata **
+--+
| +-- Alasmidonta_triangulata **
+--+
+-- Alasmidonta_arcula **
Campbell & Lydeard, 2012 [Publ 3308], fig. 1
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 43 |
43 |
41 |
1 |
0 |
0 |
0 |
114.0 |
+----------- Quadrula_quadrula **
|
+-----------------------------------+ +-- Pustulosa_infucata **
| | +-----+
| | | +-- Pustulosa_kleiniana **
| +--+
| | +----- Cyclonaias_tuberculata **
| +--+
| | +-- Pustulosa_succissa **
| +--+
| +-- Pustulosa_pustulosa **
|
| +----- Popenaias_popeii **
| +-----------+
--+ | | +-- Amblema_elliottii **
| | +--+
| | +-- Amblema_plicata **
| +--------------------------+
| | | +----- Cyrtonaias_tampicoensis **
| | | +--------+
| | | | | +-- Reginaia_ebenus **
| | | | +--+
| | +--+ +-- Reginaia_rotulata **
| | |
| | | +----------- Plectomerus_dombeyanus **
| | +--+
| | | +-------- Lemiox_rimosus **
+--+ +--+
| | +----- Lampsilis_ovata **
| +--+
| | +-- Obovaria_unicolor **
| +--+
| +-- Obovaria_olivaria **
|
| +----------------------------------------- Elliptoideus_sloatianus **
| |
| | +-------------------------------------- Pleurobema_stabile **
+--+ |
| | +----- Pleurobema_pyriforme **
| | +-----------------------------+
+--+ | | +-- Pleurobema_clava **
| | +--+
| | +-- Pleurobema_decisum **
| |
+--+ +-- Pleurobema_cordatum **
| +-----------------------------+
| | +-- Pleurobema_sintoxia **
| |
| | +-- Eurynia_dilatata **
| | +-----+
+--+ | +-- Hemistena_lata **
| +--------------------+
| | | +----- Plethobasus_cyphyus **
| | +--+
| | | +-- Elliptio_arca **
| | +--+
+--+ +-- Elliptio_crassidens **
|
| +----- Pleuronaia_gibber **
| +--------------------+
| | | +-- Pleuronaia_barnesiana **
| | +--+
+--+ +-- Pleuronaia_dolabelloides **
|
| +----------------------- Parvaspina_collina **
| |
+--+ +-------------------- Fusconaia_subrotunda **
| |
+--+ +-- Fusconaia_cor **
| +--------------+
| | +-- Fusconaia_cuneolus **
+--+
| +-------------- Fusconaia_ozarkensis **
| |
+--+ +-- Fusconaia_burkei **
| +--------+
| | +-- Fusconaia_escambia **
+--+
| +-------- Fusconaia_masoni **
+--+
| +----- Fusconaia_cerina[2] **
+--+
| +-- Fusconaia_flava **
+--+
+-- Fusconaia_cerina[1] **
Campbell & Lydeard, 2012 [Publ 3308], fig. 2
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 43 |
43 |
36 |
1 |
0 |
0 |
0 |
113.0 |
+----------- Quadrula_quadrula **
|
+--------------------+ +-- Pustulosa_infucata **
| | +-----+
| | | +-- Pustulosa_kleiniana **
| +--+
| | +----- Cyclonaias_tuberculata **
| +--+
| | +-- Pustulosa_succissa **
| +--+
| +-- Pustulosa_pustulosa **
|
| +-- Reginaia_ebenus **
| +--+
| | +-- Reginaia_rotulata **
--+ +-----------+
| | | +-- Amblema_elliottii **
| | +--+
| +-----------+ +-- Amblema_plicata **
| | |
| | | +-------------- Popenaias_popeii **
| | | |
| | +--+ +----------- Cyrtonaias_tampicoensis **
| | | |
| | +--+----------- Plectomerus_dombeyanus **
| | |
| | | +-------- Lemiox_rimosus **
| | +--+
| | | +----- Lampsilis_ovata **
+--+ +--+
| | +-- Obovaria_unicolor **
| +--+
| +-- Obovaria_olivaria **
|
| +-------------------------- Elliptoideus_sloatianus **
| |
| | +----------------------- Hemistena_lata **
| | |
| | +----------------------- Plethobasus_cyphyus **
| | |
+--+ | +-- Elliptio_arca **
| +--------------------+
| | +-- Elliptio_crassidens **
| |
| | +-- Eurynia_dilatata **
| +--------------------+
| | +-- Parvaspina_collina **
+--+
| +----- Pleuronaia_gibber **
| +--------------+
| | | +-- Pleuronaia_barnesiana **
| | +--+
| | +-- Pleuronaia_dolabelloides **
| |
| | +-- Pleurobema_cordatum **
| | +--------+
| | | +-- Pleurobema_sintoxia **
| +--------+
| | | +-------- Pleurobema_stabile **
+--+ +--+
| | +----- Pleurobema_pyriforme **
| +--+
| | +-- Pleurobema_clava **
| +--+
| +-- Pleurobema_decisum **
|
| +----- Fusconaia_subrotunda **
| +-----------+
| | | +-- Fusconaia_cor **
| | +--+
+--+ +-- Fusconaia_cuneolus **
|
| +-------------- Fusconaia_ozarkensis **
| |
+--+ +-- Fusconaia_burkei **
| +--------+
| | +-- Fusconaia_escambia **
+--+
| +-------- Fusconaia_masoni **
+--+
| +----- Fusconaia_cerina[2] **
+--+
| +-- Fusconaia_flava **
+--+
+-- Fusconaia_cerina[1] **
Campbell & Lydeard, 2012 [Publ 3308], fig. 4
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 43 |
43 |
40 |
1 |
0 |
0 |
0 |
114.0 |
+----------- Quadrula_quadrula **
|
+--------------------------------------+ +-- Pustulosa_infucata **
| | +-----+
| | | +-- Pustulosa_kleiniana **
| +--+
| | +----- Cyclonaias_tuberculata **
| +--+
| | +-- Pustulosa_succissa **
| +--+
| +-- Pustulosa_pustulosa **
|
| +----- Popenaias_popeii **
| +--------+
--+ | | +-- Amblema_elliottii **
| | +--+
| | +-- Amblema_plicata **
| +--------------------------------+
| | | +----------- Plectomerus_dombeyanus **
| | | |
| | | | +----- Cyrtonaias_tampicoensis **
| | | +-----+
| | +--+ | +-- Reginaia_ebenus **
| | | +--+
| | | +-- Reginaia_rotulata **
| | |
+--+ | +-------- Lemiox_rimosus **
| +--+
| | +----- Lampsilis_ovata **
| +--+
| | +-- Obovaria_unicolor **
| +--+
| +-- Obovaria_olivaria **
|
| +-------------------------------------------- Elliptoideus_sloatianus **
| |
+--+ +----------------------------------------- Pleurobema_stabile **
| |
+--+ +-------------------------------------- Hemistena_lata **
| |
+--+ +----------------------------------- Eurynia_dilatata **
| |
| | +-- Pleurobema_cordatum **
+--+ +-----------------------------+
| | +-- Pleurobema_sintoxia **
| |
| | +----- Pleurobema_clava **
+--+ +-----------------------+
| | | +-- Pleurobema_decisum **
| | +--+
| | +-- Pleurobema_pyriforme **
| |
+--+ +-------- Parvaspina_collina **
| +-----------------+
| | | +----- Pleuronaia_gibber **
| | +--+
| | | +-- Pleuronaia_barnesiana **
| | +--+
+--+ +-- Pleuronaia_dolabelloides **
|
| +----- Plethobasus_cyphyus **
| +-----------------+
| | | +-- Elliptio_arca **
| | +--+
+--+ +-- Elliptio_crassidens **
|
| +-------------------- Fusconaia_subrotunda **
| |
+--+ +-- Fusconaia_cor **
| +--------------+
| | +-- Fusconaia_cuneolus **
+--+
| +-------------- Fusconaia_ozarkensis **
| |
+--+ +-- Fusconaia_burkei **
| +--------+
| | +-- Fusconaia_escambia **
+--+
| +-------- Fusconaia_masoni **
+--+
| +----- Fusconaia_cerina[2] **
+--+
| +-- Fusconaia_flava **
+--+
+-- Fusconaia_cerina[1] **
Burlakova et al., 2012 [Publ 3486], fig. 3
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 43 |
43 |
26 |
1 |
0 |
0 |
0 |
111.0 |
+----------------------------------- Plectomerus_dombeyanus **
|
| +-------------------------------- Hemistena_lata **
| |
--+ +-------------------------------- Pleurobema_stabile **
| |
| | +----------------------------- Eurynia_dilatata **
| | |
+--+ | +-------- Pleurobema_riddellii **
| | |
| | +-----------------+ +----- Pleurobema_sintoxia **
| | | | |
| | | +--+----- Pleurobema_rubrum[2] **
| | | |
+--+ | | +-- Pleurobema_cordatum **
| | +--+
| | +-- Pleurobema_rubrum[1] **
| |
| | +----------------------- Parvaspina_collina **
| | |
| | | +----- Pleuronaia_gibber **
| | +-----------------+
| | | | +-- Pleuronaia_dolabelloides **
+--+ | +--+
| | +-- Pleuronaia_barnesiana **
| |
| | +----- Plethobasus_cyphyus **
| +-----------------+
| | | +-- Elliptio_arca **
| | +--+
| | +-- Elliptio_crassidens **
| |
| | +-- Pleurobema_clava **
| | +--------+
| | | +-- Pleurobema_oviforme **
| | |
| | | +-------- Pleurobema_decisum **
+--+-----------+ |
| | +-------- Pleurobema_pyriforme **
| | |
| | +-------- Pleurobema_hanleyianum **
| +--+
| +-------- Pleurobema_georgianum **
| |
| | +----- Pleurobema_rubellum **
| | |
| | +----- Pleurobema_beadleianum **
| +--+
| +----- Pleurobema_strodeanum **
| |
| | +-- Pleurobema_perovatum **
| +--+
| +-- Pleurobema_taitianum **
|
| +-------------------- Fusconaia_subrotunda **
| |
| | +-- Fusconaia_cor **
+--+ +--------------+
| | +-- Fusconaia_cuneolus **
| |
| | +-------------- Fusconaia_ozarkensis **
+--+ |
| | +-------- Fusconaia_masoni **
| | |
| | | +-- Fusconaia_burkei **
| | +--+-----+
+--+ | | +-- Fusconaia_escambia **
| | |
| | | +----- Fusconaia_cerina **
| | +--+
| | | +-- Fusconaia_askewi[1] **
| | +--+
+--+ +-- Fusconaia_flava **
|
| +-- Fusconaia_askewi[2] **
| |
| +-- Fusconaia_askewi[3] **
| |
| +-- Fusconaia_askewi[4] **
+--------+
+-- Fusconaia_lananensis[1] **
|
+-- Fusconaia_lananensis[2] **
|
+-- Fusconaia_askewi[5] **
Inoue et al., 2020 [Publ 3941], fig. 4
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 43 |
43 |
41 |
1 |
0 |
0 |
0 |
109.0 |
+-------------------------------------------------- Arcidens_confragosus **
|
| +-- Pustulosa_petrina **
| +--------------------------------------------+
--+ | +-- Quadrula_quadrula **
| |
| | +-- Reginaia_ebenus **
| | +-----------------------------------------+
+--+ | +-- Reginaia_rotulata **
| |
| | +-------------- Parvaspina_collina **
| | |
| | | +-- Eurynia_dilatata **
| | +--------------------------+ +--------+
+--+ | | | +-- Hemistena_lata **
| | | |
| | +--+ +-- Pleurobema_clava **
| | | +-----+
| | | | +-- Pleurobema_sintoxia **
| | +--+
| | | +----- Pleuronaia_dolabelloides **
+--+ +--+
| | +-- Fusconaia_flava **
| +--+
| +-- Fusconaia_subrotunda **
|
| +-- Amblema_plicata **
| +-----------------------------------+
| | +-- Amblema_elliottii **
| |
+--+ +----------------------------------- Toxolasma_lividum **
| |
| | +-------------------------------- Ellipsaria_lineolata **
+--+ |
| | +----- Truncilla_cognata **
| | +-----------------------+
| | | | +-- Truncilla_macrodon **
+--+ | +--+
| | +-- Truncilla_donaciformis **
| |
| | +-------- Potamilus_fragilis **
| | |
+--+ +-----------------+ +-- Potamilus_ohiensis **
| | | +--+
| | | | +-- Potamilus_amphichaenus **
| | +--+
| | | +-- Potamilus_metnecktayi **
| | +--+
| | +-- Potamilus_purpuratus **
+--+
| +----------------------- Leaunio_lienosus **
| |
| | +----------------- Lampsilis_bracteata **
| | |
| | +--+ +-- Lampsilis_ornata **
| | | | +-----------+
+--+ | | | +-- Lampsilis_fasciola **
| | +--+
| | | +----------- Lampsilis_ovata **
| | +--+
| | | +-------- Lampsilis_cardium[3] **
| | +--+
+--+ | +----- Lampsilis_cardium[2] **
| +--+
| | +-- Lampsilis_satura **
| +--+
| +-- Lampsilis_cardium[1] **
|
| +-------------- Lampsilis_teres **
| |
| | +-- Lampsilis_higginsii **
+-----+ +--------+
| | +-- Ortmanniana_ligamentina **
| |
+--+ +-- Lampsilis_radiata **
| +-----+
| | +-- Lampsilis_siliquoidea **
| |
+--+ +-- Lampsilis_hydiana **
| +--+
| | +-- Lampsilis_bergmanni **
+--+
| +-- Lampsilis_virescens **
+--+
+-- Lampsilis_straminea **
Smith et al., 2019 [Publ 3951], fig. 2
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 42 |
42 |
40 |
1 |
0 |
0 |
0 |
111.0 |
+----------------------------------------------- Quadrula_quadrula **
|
| +-- Amblema_plicata **
--+ +-----------------------------------------+
| | +-- Reginaia_ebenus **
| |
+--+ +----------------------------------------- Plectomerus_dombeyanus **
| |
| | +-------------------------------------- Toxolasma_lividum **
+--+ |
| | +----- Popenaias_popeii **
| | +-----------------------------+
+--+ | | +-- Glebula_rotundata **
| | +--+
| | +-- Cyrtonaias_tampicoensis **
| |
| | +-------------------------------- Obliquaria_reflexa **
+--+ |
| | +-- Medionidus_conradicus **
| | +--+
| | | +-- Lemiox_rimosus **
| | +-----------------------+
| | | | +-- Cyprogenia_stegaria **
| | | +--+
+--+ | +-- Ptychobranchus_fasciolaris **
| |
| | +-- Hamiota_subangulata **
| | +--------------------+
| | | +-- Villosa_villosa **
| | |
| | +--+ +-------------------- Ligumia_recta **
| | | | |
+--+ | | | +----- Venustaconcha_ellipsiformis **
| | +--+ +-----------+
| | | | | +-- Obovaria_subrotunda **
| | | | +--+
| | +--+ +-- Epioblasma_rangiana **
| | |
| | | +-------------- Ortmanniana_pectorosa **
| | +--+
| | | +----------- Ortmanniana_ligamentina **
| | +--+
| | | +-------- Lampsilis_ovata **
| | +--+
+--+ | +----- Lampsilis_dolabraeformis **
| +--+
| | +-- Lampsilis_satura **
| +--+
| +-- Lampsilis_cardium **
|
| +-------------- Atlanticoncha_ochracea **
| +-----+
| | | +----------- Ellipsaria_lineolata **
| | +--+
| | | +-------- Truncilla_truncata **
| | +--+
| | | +----- Truncilla_cognata **
| | +--+
+-----+ | +-- Truncilla_macrodon **
| +--+
| +-- Truncilla_donaciformis **
|
| +----------------- Potamilus_capax **
| |
| | +----- Potamilus_inflatus **
+--+ +--------+
| | | +-- Potamilus_fragilis **
| | +--+
| | +-- Potamilus_leptodon **
+--+
| +----- Potamilus_ohiensis **
| +-----+
| | | +-- Potamilus_amphichaenus **
| | +--+
+--+ +-- Potamilus_streckersoni **
|
| +-------- Potamilus_metnecktayi **
+--+
| +----- Potamilus_purpuratus[1] **
+--+
| +-- Potamilus_purpuratus[2] **
+--+
+-- Potamilus_alatus **
Bolotov et al., 2023 [Publ 4312], fig. S2
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 42 |
42 |
39 |
1 |
0 |
0 |
0 |
113.0 |
+----------- Indopseudodon_kayinensis **
+-----------------------+
| | +-------- Indopseudodon_indawgyiensis **
| +--+
| | +----- Indopseudodon_bogani **
| +--+
| | +-- Indopseudodon_salwenianus **
| +--+
| +-- Indopseudodon_crebristriatus **
|
--+ +-------------------------------- Songkhlanaia_tamodienica **
| |
| | +----------------- Nyeinchanconcha_nyeinchani **
| | |
| | +-----------+ +-------------- Sundadontina_tumida **
| | | | |
| | | | | +-------- Sundadontina **
| | | +--+ +--+
| | | | | | +----- Sundadontina_tanintharyiensis **
| | | | | +--+
+--+ | | | | +-- Sundadontina_plugpomenica **
| | +--+ +--+
| | | +-- Sundadontina_cumingii **
| | |
| | | +-------- Sundadontina_sulcata **
| | +--+
| | | +----- Sundadontina_harmandi **
| | +--+
| | | +-- Sundadontina_taskaevi **
| | +--+
| | +-- Sundadontina_brandti **
+--+
| +-- Namkongnaia_inkhavilayi **
| +--------------+
| | +-- Namkongnaia_lemeslei **
| |
| +--------+ +-- Thaiconcha_callifera **
| | | |
| | | +-----------+-- Thaiconcha_thaiensis **
| | | | |
| | +--+ +-- Thaiconcha_munelliptica **
| | |
| | | +----------- Bineurus_loeiensis **
| | +--+
| | | +-------- Bineurus_panhai **
| | +--+
| | | +----- Bineurus_anodontinum **
| | +--+
+--+ | +-- Bineurus_exilis **
| +--+
| +-- Bineurus_mouhotii **
|
| +----------- Pseudodon_vagulus **
| |
| +-----------+ +-- Pseudodon_lenyanensis **
| | | +-----+
| | | | +-- Pseudodon_inoscularis **
| | +--+
| | | +----- Pseudodon_cambodjensis **
| | +--+
+--+ | +-- Pseudodon_vondembuschianus **
| +--+
| +-- Pseudodon_mekongi **
|
| +-------------------- Pilsbryoconcha_carinifera **
| |
+--+ +-- Pilsbryoconcha_linguaeformis[2] **
| +--------------+
| | +-- Pilsbryoconcha **
+--+
| +-------------- Pilsbryoconcha_linguaeformis[1] **
| |
+--+ +-- Pilsbryoconcha_schomburgki **
| +--------+
| | +-- Pilsbryoconcha_exilis **
+--+
| +-------- Pilsbryoconcha_hoikaab **
+--+
| +----- Pilsbryoconcha_mekongiana **
+--+
| +-- Pilsbryoconcha_acuta **
+--+
+-- Pilsbryoconcha_kittitati **
Aunins et al., 2018 [Publ 3863], fig. 1
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 41 |
41 |
39 |
2 |
0 |
0 |
0 |
78.0 |
+----- Cumberlandia_monodonta **
+--------------------------+
| | +-- Margaritifera_falcata **
| +--+
| +-- Margaritifera_dahurica **
|
| +-------- Quadrula_quadrula **
| +--------------------+
| | | +----- Toxolasma_parvum **
--+ | +--+
| | | +-- Lampsilis_ornata **
| | +--+
| | +-- Venustaconcha_ellipsiformis **
| |
| | +-- Sinohyriopsis_cumingii **
| | +--------+
| | | +-- Sinohyriopsis_schlegelii **
+--+ |
| +--------------+ +-- Lamprotula_caveata **
| | | +-----+
| | | | +-- Potomida_littoralis **
| | +--+
| | | +----- Sinosolenaia_oleivora **
| | +--+
| | | +-- Ptychorhynchus_pfisteri **
| | +--+
| | +-- Sinosolenaia_carinata **
+--+
| +----------------------- Lepidodesma_languilati **
| |
| | +----- Aculamprotula_tortuosa **
| | +--------+
| | | | +-- Aculamprotula_coreana **
| | | +--+
| | | +-- Aculamprotula_tientsinensis **
| | |
| | +-----+ +-------- Nodularia_douglasiae **
+--+ | | +--+
| | | | | +----- Tchangsinaia_piscicula **
| | | | +--+
| | | | | +-- Cuneopsis_heudei **
| | +--+ +--+
| | | +-- Schistodesmus_lampreyanus **
| | |
| | | +-------- Unio_tumidus **
| | +--+
| | | +----- Unio_crassus **
+--+ +--+
| | +-- Unio_pictorum **
| +--+
| +-- Unio_delphinus **
|
| +----- Acuticosta_chinensis **
| +-----------+
| | | +-- Lanceolaria_grayii **
| | +--+
| | +-- Lanceolaria_lanceolata **
| |
| | +-------- Cristaria_plicata **
+--+ |
| +-----+ +-- Anemina_arcaeformis **
| | | +--+
| | | | +-- Anemina_euscaphys **
| | +--+
| | | +-- Sinanodonta_woodiana **
| | +--+
+--+ +-- Sinanodonta_lucida **
|
| +-- Anodonta_anatina **
| +--------+
| | +-- Anodonta_cygnea **
| |
+--+ +-- Prolasmidonta_heterodon **
| +-----+
| | +-- Alasmidonta_varicosa **
+--+
| +----- Pyganodon_grandis **
+--+
| +-- Utterbackia_imbecillis **
+--+
+-- Utterbackia_peninsularis **
Burlakova et al., 2012 [Publ 3486], fig. 2
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 40 |
40 |
30 |
1 |
0 |
0 |
0 |
112.0 |
+----------------------------------- Parvaspina_collina **
|
| +-------------------------------- Pleurobema_plenum **
| |
--+ | +-- Plethobasus_cyphyus **
| | +-----+
| | | +-- Hemistena_lata **
| | +--------------------+
+--+ | | +----- Eurynia_dilatata **
| | +--+
| | | +-- Elliptio_arca **
| | +--+
| | +-- Elliptio_crassidens **
| |
| | +-------------------------- Pleurobema_cordatum[2] **
+--+ |
| | +-------------- Fusconaia_subrotunda **
| | |
| | | +-- Fusconaia_cor **
| | | +--------+
| | +--------+ | +-- Fusconaia_cuneolus **
| | | | |
| | | | | +-------- Fusconaia_flava[2] **
| | | | | |
+--+ | +--+ +-------- Fusconaia_ozarkensis **
| | | |
| | | +-------- Fusconaia_masoni **
| | | |
| | | | +----- Fusconaia_askewi **
| | +--+--+
| | | | +-- Fusconaia_burkei **
| | | +--+
| | | +-- Fusconaia_escambia **
| | |
+--+ | +-- Fusconaia_flava[1] **
| | |
| +-----+-- Fusconaia_cerina[1] **
| |
| +-- Fusconaia_cerina[2] **
|
| +-------- Pleurobema_stabile **
| |
| +-----------+ +----- Pleurobema_riddellii **
| | | |
| | +--+ +-- Pleurobema_cordatum[1] **
| | | |
| | | +-- Pleurobema_rubrum **
| | +--+
| | +-- Pleurobema_sintoxia[1] **
+--+ |
| +-- Pleurobema_sintoxia[2] **
|
| +----- Pleuronaia_barnesiana **
| +-----------+
| | | +-- Pleuronaia_dolabelloides **
| | +--+
| | +-- Pleuronaia_gibber **
+--+
| +-- Pleurobema_georgianum **
| +-----------+
| | +-- Pleurobema_pyriforme **
| |
+--+ +-- Pleurobema_decisum **
| +--------+
| | +-- Pleurobema_hanleyianum **
| |
+--+ +-- Pleurobema_clava **
| +-----+
| | +-- Pleurobema_oviforme **
| |
+--+ +----- Pleurobema_rubellum **
| |
| +----- Pleurobema_beadleianum **
+--+
+----- Pleurobema_strodeanum **
|
| +-- Pleurobema_perovatum **
+--+
+-- Pleurobema_taitianum **
Chen et al., 2023 [Publ 4302], fig. 1
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 40 |
40 |
31 |
2 |
0 |
0 |
0 |
90.0 |
+-- Gibbosula_rochechouartii **
+-----------------------------------+
| +-- Margaritifera_dahurica **
|
| +-- Pronodularia_japanensis **
| +--+
| | +-- Lamprotula_leaii **
--+ +-----------------------------+
| | | +-- Lampsilis_siliquoidea **
| | +--+
| | +-- Quadrula_quadrula **
| |
| | +-- Coelatura_aegyptiaca **
+--+ |
| +-----------------------------+-- Indonaia_andersoniana **
| | |
| | +-- Parreysia_nagpoorensis **
| |
| | +-- Anemina_arcaeformis **
+--+ +--------------------------+
| | +-- Amuranodonta_kijaensis **
| |
| | +----- Pletholophus_tenuis **
| | +--------------------+
| | | | +-- Sinanodonta_jourdyi **
+--+ | +--+
| | +-- Beringiana_beringiana **
| |
| | +----------------------- Lanceolaria_gladiola **
| | |
| | +----------------------- Acuticosta_chinensis **
+--+ |
| +----------------------- Lepidodesma_languilati **
| |
| +----------------------- Lanceolaria_oxyrhyncha **
| |
| +----------------------- Cristaria_plicata **
| |
| | +-- Alasmidonta_marginata **
+--+ +--+
| | +-- Lasmigona_costata **
+-----------------+
| | +-- Anodonta_anatina **
| +--+
| +-- Pseudanodonta_complanata **
|
| +-------------------- Aculamprotula_fibrosa **
| |
| | +----- Middendorffinaia_mongolica **
| | +-----------+
+--+ | | +-- Nodularia_douglasiae **
| | +--+
| | +-- Nodularia_breviconcha **
| |
| | +-- Cuneopsis_celtiformis **
+--+ +-----------+
| | +-- Cuneopsis_heudei **
| |
| | +-- Diaurora_aurorea **
| | +--+
| | | +-- Diaurora_laeve **
| +--------+
+--+ | +-- Schistodesmus_lampreyanus **
| +--+
| +-- Schistodesmus_spinosus **
|
| +-- Pseudocuneopsis_sichuanensis **
| +--------+
| | +-- Pseudocuneopsis_capitata **
| |
+--+ +-- Inversiunio_yanagawensis **
| +-----+
| | +-- Inversiunio_reinianus **
+--+
| +----- Tchangsinaia_piscicula **
+--+
| +-- Unio_pictorum **
+--+
+-- Unio_crassus **
Chen et al., 2023 [Publ 4302], fig. 2
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 40 |
40 |
36 |
2 |
0 |
0 |
0 |
84.0 |
+-- Gibbosula_rochechouartii **
+--------------------------------+
| +-- Margaritifera_dahurica **
|
| +-- Pronodularia_japanensis **
| +--+
| | +-- Lamprotula_leaii **
--+ +--------------------------+
| | | +-- Lampsilis_siliquoidea **
| | +--+
| | +-- Quadrula_quadrula **
| |
| | +----- Parreysia_nagpoorensis **
+--+ +-----------------------+
| | | +-- Coelatura_aegyptiaca **
| | +--+
| | +-- Indonaia_andersoniana **
| |
| | +-------------------------- Acuticosta_chinensis **
+--+ |
| | +----------------------- Lepidodesma_languilati **
| | |
| | | +-- Lanceolaria_gladiola **
| | | +--------------+
| | | | +-- Lanceolaria_oxyrhyncha **
| | | |
+--+ | | +-- Alasmidonta_marginata **
| +-----+ +--+
| | | | +-- Lasmigona_costata **
| | | +--------+
| | | | | +-- Anodonta_anatina **
| | | | +--+
| | +--+ +-- Pseudanodonta_complanata **
| | |
| | | +----------- Cristaria_plicata **
+--+ | |
| +--+ +-- Anemina_arcaeformis **
| | +-----+
| | | +-- Amuranodonta_kijaensis **
| +--+
| | +----- Pletholophus_tenuis **
| +--+
| | +-- Sinanodonta_jourdyi **
| +--+
| +-- Beringiana_beringiana **
|
| +-------------------- Aculamprotula_fibrosa **
| |
| | +----- Middendorffinaia_mongolica **
| | +-----------+
+--+ | | +-- Nodularia_douglasiae **
| | +--+
| | +-- Nodularia_breviconcha **
| |
| | +-- Cuneopsis_celtiformis **
+--+ +-----------+
| | +-- Cuneopsis_heudei **
| |
| | +-- Diaurora_aurorea **
| | +--+
| | | +-- Diaurora_laeve **
| +--------+
+--+ | +-- Schistodesmus_lampreyanus **
| +--+
| +-- Schistodesmus_spinosus **
|
| +-- Pseudocuneopsis_sichuanensis **
| +--------+
| | +-- Pseudocuneopsis_capitata **
| |
+--+ +-- Inversiunio_yanagawensis **
| +-----+
| | +-- Inversiunio_reinianus **
+--+
| +----- Tchangsinaia_piscicula **
+--+
| +-- Unio_pictorum **
+--+
+-- Unio_crassus **
Bolotov et al., 2023 [Publ 4312], fig. 2
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 40 |
40 |
38 |
1 |
0 |
0 |
0 |
107.0 |
+----------- Indopseudodon_kayinensis **
+-----------------+
| | +-------- Indopseudodon_indawgyiensis **
| +--+
| | +----- Indopseudodon_bogani **
| +--+
| | +-- Indopseudodon_salwenianus **
| +--+
| +-- Indopseudodon_crebristriatus **
|
| +-- Pilsbryoconcha_exilis **
| +-----------+
--+ | +-- Pilsbryoconcha_schomburgki **
| |
| +-----------+ +-- Pilsbryoconcha_carinifera **
| | | +--------+
| | | | +-- Pilsbryoconcha_linguaeformis **
| | +--+
| | | +-------- Pilsbryoconcha_acuta **
| | +--+
| | | +----- Pilsbryoconcha_kittitati **
| | +--+
| | | +-- Pilsbryoconcha_mekongiana **
| | +--+
+--+ +-- Pilsbryoconcha_hoikaab **
|
| +----------------------- Songkhlanaia_tamodienica **
| |
| | +-- Namkongnaia_inkhavilayi **
| | +--------+
| | | +-- Namkongnaia_lemeslei **
| | |
| | | +----- Pseudodon_vondembuschianus[2] **
| | +--------+ +--+
| | | | | | +-- Pseudodon_vondembuschianus[1] **
+--+ | | | +--+
| | | | +-- Pseudodon_mekongi **
| | +--+
| | | +-- Pseudodon_vagulus **
| | | +--+
| | | | +-- Pseudodon_cambodjensis **
| | +--+
| | | +-- Pseudodon_lenyanensis **
| | +--+
| | +-- Pseudodon_inoscularis **
+--+
| +-------------- Nyeinchanconcha_nyeinchani **
| |
| | +----- Thaiconcha_munelliptica **
| +--+ +-----+
| | | | | +-- Thaiconcha_callifera **
| | | | +--+
| | +--+ +-- Thaiconcha_thaiensis **
| | |
| | | +-------- Bineurus_loeiensis **
| | +--+
| | | +----- Bineurus_mouhotii **
| | +--+
+--+ | +-- Bineurus_anodontinum **
| +--+
| +-- Bineurus_exilis **
|
| +-------- Sundadontina_sulcata **
| +-----+
| | | +----- Sundadontina_harmandi **
| | +--+
| | | +-- Sundadontina_brandti **
+--+ +--+
| +-- Sundadontina_taskaevi **
|
| +----------- Sundadontina_tumida **
+--+
| +-------- Sundadontina **
+--+
| +----- Sundadontina_tanintharyiensis **
+--+
| +-- Sundadontina_cumingii **
+--+
+-- Sundadontina_plugpomenica **
Pfeiffer et al., 2016 [Publ 3660], fig. 2
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 39 |
39 |
34 |
1 |
0 |
0 |
0 |
105.0 |
+-------------------------------------- Utterbackiana_suborbiculata **
|
| +-- Amblema_plicata **
| +--+
| | +-- Amblema_neislerii **
| +-----------------------------+
--+ | | +-- Reginaia_rotulata **
| | +--+
| | +-- Reginaia_ebenus **
| |
| | +-- Cyrtonaias_tampicoensis **
| | +-----+
| | | +-- Glebula_rotundata **
+--+ |
| +-----------------------+ +-- Lampsilis_teres **
| | | +--+
| | | | +-- Leaunio_lienosus **
| | +--+
| | | +-- Truncilla_macrodon **
| | +--+
| | +-- Potamilus_fragilis **
+--+
| +----------------------------- Elliptoideus_sloatianus **
| |
| | +-------------- Uniomerus_declivis **
| | +-----------+
| | | | +----------- Quadrula_quadrula **
| | | +--+
+--+ | | +-------- Quadrula_verrucosa **
| | +--+
| | | +----- Pustulosa_pustulosa **
| | +--+
| | | +-- Pustulosa_petrina **
| | +--+
+--+ +-- Cyclonaias_tuberculata **
|
| +-- Elliptio_fumata **
| +--------------------+
| | +-- Elliptio_crassidens **
| |
| | +-------------------- Parvaspina_collina **
| | |
| | +-------------------- Plethobasus_cyphyus **
+--+ |
| | +-- Hemistena_lata **
| +-----------------+
| | +-- Eurynia_dilatata **
| |
| | +----- Pleurobema_riddellii **
+--+ +--+
| | | +-- Pleurobema_sintoxia **
| | +--+
| +--------+ +-- Pleurobema_cordatum **
| | |
| | | +----- Pleuronaia_dolabelloides **
| | | |
| | +--+ +-- Pleurobema_pyriforme **
| | | |
+--+ +--+-- Pleurobema_strodeanum **
| |
| +-- Pleurobema_clava **
|
| +-------------- Fusconaia_askewi **
| |
| | +-- Fusconaia_mitchelli **
+--+ +--------+
| | +-- Fusconaia_iheringi **
| |
+--+ +-- Fusconaia_escambia **
| +-----+
| | +-- Fusconaia_burkei **
+--+
| +----- Fusconaia_flava[2] **
+--+
| +-- Fusconaia_cerina **
+--+
+-- Fusconaia_flava[1] **
Huang et al., 2018 [Publ 3828], fig. 6
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 39 |
39 |
37 |
2 |
4 |
3 |
2 |
80.0 |
+-- Anodontites_trapesialis
+--------------------------+
| +-- Mutela_dubia
|
| +-------------------------- Echyridella_menziesii
| |
| | +-------------- Gibbosula_rochechouartii **
--+ | +--------+
| | | | +----------- Cumberlandia_monodonta **
| | | +--+
| | | | +-------- Pseudunio_marocanus **
| | | +--+
| | | | +----- Margaritifera_falcata **
| | | +--+
+--+ | | +-- Margaritifera_margaritifera **
| | +--+
| | +-- Margaritifera_dahurica **
| |
| | +----------- Quadrula_quadrula **
| | |
| | +-----+ +-------- Toxolasma_parvum **
| | | | |
| | | +--+ +-- Lampsilis_ornata **
| | | | +--+
| | | | | +-- Venustaconcha_ellipsiformis **
+--+ | +--+
| | | +-- Potamilus_leptodon **
| +--+ +--+
| | | +-- Potamilus_alatus **
| | |
| | | +-- Sinohyriopsis_cumingii **
| | | +-----------+
| | | | +-- Sinohyriopsis_schlegelii **
| | | |
| | +--+ +-- Sinosolenaia_oleivora **
| | | +--------+
| | | | +-- Sinosolenaia_carinata **
| | +--+
| | | +-------- Potomida_littoralis **
| | +--+
| | | +----- Pronodularia_japanensis **
| | +--+
| | | +-- Lamprotula_caveata **
+--+ +--+
| +-- Lamprotula_leaii **
|
| +----- Aculamprotula_tortuosa **
| +--------+
| | | +-- Aculamprotula_coreana **
| | +--+
| | +-- Aculamprotula_tientsinensis **
| +--+
| | | +-- Tchangsinaia_piscicula **
| | | +--------+
| | | | +-- Nodularia_douglasiae **
| | +--+
| | | +-------- Unio_tumidus **
| | +--+
| | | +----- Unio_crassus **
| | +--+
+--+ | +-- Unio_delphinus **
| +--+
| +-- Unio_pictorum **
|
| +----------- Anodonta_anatina **
| +--+
| | | +-------- Platynaias_compressa **
| | +--+
| | | +----- Pyganodon_grandis **
| | +--+
+--+ | +-- Utterbackia_imbecillis **
| +--+
| +-- Utterbackia_peninsularis **
|
| +-------- Cristaria_plicata **
| |
+-----+ +-- Anemina_arcaeformis **
| +--+
| | +-- Anemina_euscaphys **
+--+
| +-- Sinanodonta_woodiana **
+--+
+-- Sinanodonta_lucida **
Pfeiffer et al., 2019 [Publ 3866], fig. 1
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 39 |
39 |
34 |
1 |
0 |
0 |
0 |
99.0 |
+-------------------------------------- Gonidea_angulata **
|
| +----------- Megalonaias_nervosa **
| +-----------------------+
| | | +-------- Uniomerus_declivis **
| | +--+
--+ | | +----- Quadrula_verrucosa **
| | +--+
| | | +-- Cyclonaias_tuberculata **
| | +--+
| | +-- Quadrula_quadrula **
| |
| | +-------------- Elliptoideus_sloatianus **
+--+ +-----------------+
| | | +----------- Elliptio_fumata **
| | +--+
| | | +-------- Pleuronaia_barnesiana **
| | +--+
| | | +----- Pleurobema_riddellii **
| | +--+
| | | +-- Fusconaia_flava **
| | +--+
| | +-- Pleurobema_strodeanum **
| |
+--+ +-- Amblema_plicata **
| +--+
| | +-- Amblema_neislerii **
| +-----------------------+
| | | +-- Reginaia_ebenus **
| | +--+
| | +-- Reginaia_rotulata **
| |
| | +----------------- Popenaias_popeii **
| | |
| | +--------+ +-------------- Psoronaias_guatemalensis[2] **
| | | | |
+--+ | +--+ +----------- Psoronaias_guatemalensis[1] **
| | | |
| | +--+ +-------- Psoronaias_semigranosa[3] **
| | | |
| | | | +----- Psoronaias_semigranosa[2] **
| | +--+ |
| | | +----- Psoronaias_usumasintae[2] **
| | | |
| | +--+----- Psoronaias_percompressa **
+--+ |
| | +-- Psoronaias_semigranosa[1] **
| | |
| +--+-- Psoronaias_profunda **
| |
| +-- Psoronaias_usumasintae[1] **
|
| +-- Cyrtonaias_tampicoensis **
| +--------------------+
| | +-- Glebula_rotundata **
| |
+--+ +-------------------- Potamilus_alatus **
| |
| | +-- Truncilla_truncata **
+--+ +--------------+
| | +-- Ptychobranchus_fasciolaris **
| |
+--+ +-------------- Pachynaias_spheniopsis **
| |
| | +-- Cyrtonaias_explicata **
+--+ +--------+
| | +-- Sphenonaias_microdon **
| |
+--+ +-- Lampsilis_cardium **
| +-----+
| | +-- Ortmanniana_ligamentina **
+--+
| +----- Cambarunio_iris **
+--+
| +-- Epioblasma_triquetra **
+--+
+-- Ligumia_recta **
Cha et al., 2018 [Publ 3916], fig. 1
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 39 |
39 |
37 |
2 |
0 |
0 |
0 |
83.0 |
+----- Pseudunio_marocanus **
+--------------------------+
| | +-- Margaritifera_falcata **
| +--+
| +-- Margaritifera_dahurica **
|
| +----------- Quadrula_quadrula **
| +-----------------+
| | | +-------- Toxolasma_parvum **
| | +--+
--+ | | +----- Lampsilis_ornata **
| | +--+
| | | +-- Potamilus_alatus **
| | +--+
| | +-- Potamilus_leptodon **
| |
| | +-- Sinohyriopsis_cumingii **
| | +-----------+
| | | +-- Sinohyriopsis_schlegelii **
+--+ |
| +-----------+ +----- Ptychorhynchus_pfisteri **
| | | +-----+
| | | | | +-- Sinosolenaia_carinata **
| | | | +--+
| | +--+ +-- Sinosolenaia_oleivora **
| | |
| | | +-------- Potomida_littoralis **
| | +--+
| | | +----- Pronodularia_japanensis **
| | +--+
| | | +-- Lamprotula_caveata **
+--+ +--+
| +-- Lamprotula_leaii **
|
| +----------------------- Lepidodesma_languilati **
| |
| | +----- Aculamprotula_tortuosa **
| | +--------+
| | | | +-- Aculamprotula_coreana **
| | | +--+
| | | +-- Aculamprotula_tientsinensis **
| | +-----+
+--+ | | +----- Schistodesmus_lampreyanus **
| | | +-----+
| | | | | +-- Nodularia_douglasiae **
| | | | +--+
| | +--+ +-- Tchangsinaia_piscicula **
| | |
| | | +-------- Unio_tumidus **
| | +--+
+--+ | +----- Unio_crassus **
| +--+
| | +-- Unio_pictorum **
| +--+
| +-- Unio_delphinus **
|
| +----------------- Lanceolaria_grayii **
| |
| | +----------- Anodonta_anatina **
| | +--+
+--+ | | +-------- Platynaias_compressa **
| | +--+
| | | +----- Pyganodon_grandis **
| | +--+
+--+ | +-- Utterbackia_peninsularis **
| +--+
| +-- Utterbackia_imbecillis **
|
| +-------- Cristaria_plicata **
| |
+-----+ +-- Anemina_arcaeformis **
| +--+
| | +-- Anemina_euscaphys **
+--+
| +-- Sinanodonta_lucida **
+--+
+-- Sinanodonta_woodiana **
Wu et al., 2022 [Publ 4238], fig. 2d
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 39 |
39 |
37 |
1 |
0 |
0 |
0 |
83.0 |
+-------------------------------- Lepidodesma_languilati **
|
| +----- Lanceolaria_gladiola **
| +--------------+
| | | +-- Lanceolaria_grayii **
| | +--+
| | +-- Lanceolaria_lanceolata **
| +--------+
| | | +----------------- Acuticosta_chinensis **
| | | |
| | | | +-------- Cristaria_plicata **
| | | | |
--+ | +--+ +-----+ +-- Sinanodonta_woodiana **
| | | | | +--+
| | | | | | +-- Sinanodonta_lucida **
| | | | +--+
| | | | | +-- Anemina_arcaeformis **
| | +--+ +--+
| | | +-- Anemina_euscaphys **
| | |
| | | +-- Anodonta_cygnea **
| | | +--------+
| | | | +-- Anodonta_anatina **
| | +--+
+--+ | +-------- Platynaias_compressa **
| +--+
| | +----- Pyganodon_grandis **
| +--+
| | +-- Utterbackia_imbecillis **
| +--+
| +-- Utterbackia_peninsularis **
|
| +-------- Aculamprotula_tortuosa **
| |
| +-----------------+ +-- Aculamprotula_tientsinensis **
| | | +--+
| | | | +-- Aculamprotula_coreana **
| | +--+
| | | +-- Aculamprotula_polysticta **
| | +--+
| | +-- Aculamprotula_scripta **
+--+
| +-------- Unio_tumidus **
| +--------------+
| | | +----- Unio_crassus **
| | +--+
| | | +-- Unio_pictorum **
| | +--+
| | +-- Unio_delphinus **
+--+
| +-- Pseudocuneopsis_capitata **
| +-----------------+
| | +-- Pseudocuneopsis_sichuanensis **
| |
| | +-- Nodularia_breviconcha **
+--+ +--------------+
| | +-- Nodularia_douglasiae **
| |
| | +-- Schistodesmus_lampreyanus **
+--+ +-----------+
| | +-- Schistodesmus_spinosus **
| |
| | +-------- Cuneopsis_rufescens **
+--+ +--+
| | | +----- Cuneopsis_celtiformis **
| | +--+
| | | +-- Cuneopsis_heudei **
+--+ +--+
| +-- Cuneopsis_demangei **
|
| +-------- Tchangsinaia_piscicula **
+--+
| +----- Inversiunio_jokohamensis **
+--+
| +-- Inversiunio_reinianus **
+--+
+-- Inversiunio_yanagawensis **
Pavan-Kumar et al., 2022 [Publ 4263], fig. S5
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 39 |
39 |
37 |
2 |
0 |
0 |
0 |
84.0 |
+-- Margaritifera_dahurica **
+-----------------------+
| +-- Margaritifera_falcata **
|
| +-------------------- Lamellidens_marginalis **
| |
| | +-- Anodonta_anatina **
| | +--------+
| | | +-- Pyganodon_grandis **
| | +-----+
| +--+ | | +-------- Cristaria_plicata **
| | | | | |
| | | | +--+ +-- Anemina_arcaeformis **
| | | | | +--+
| | | | | | +-- Anemina_euscaphys **
--+ | | | +--+
| | | | | +-- Sinanodonta_lucida **
| | +--+ +--+
| | | +-- Sinanodonta_woodiana **
| | |
| | | +----- Acuticosta_chinensis **
| | | +--------+
| | | | | +-- Lanceolaria_lanceolata **
| | | | +--+
| | | | +-- Lanceolaria_gladiola **
| | +--+
| | | +----- Lepidodesma_languilati **
| | | +-----+
| | | | | +-- Aculamprotula_tientsinensis **
| | | | +--+
+--+ +--+ +-- Aculamprotula_tortuosa **
| |
| | +-------- Unio_pictorum **
| +--+
| | +----- Schistodesmus_lampreyanus **
| +--+
| | +-- Nodularia_douglasiae **
| +--+
| +-- Tchangsinaia_piscicula **
|
| +-------------- Quadrula_quadrula **
| |
| +--+ +----------- Toxolasma_parvum **
| | | |
| | +--+ +-------- Potamilus_leptodon **
| | | |
| | +--+ +-- Lampsilis_cardium **
| | | +--+
| | | | +-- Lampsilis_ornata **
| | +--+
| | | +-- Lampsilis_siliquoidea **
+-----+ +--+
| +-- Venustaconcha_ellipsiformis **
|
| +----- Chamberlainia_somsakpanhai **
| +--------+
| | | +-- Sinohyriopsis_schlegelii **
| | +--+
| | +-- Sinohyriopsis_cumingii **
| |
| | +-- Sinosolenaia_carinata **
+--+ +--+
| | +-- Sinosolenaia_oleivora **
| +-----+
| | | +-- Parvasolenaia_rivularis **
| | +--+
| | +-- Microcondylaea_bonellii **
+--+
| +-- Pseudodon_vondembuschianus **
| +-----+
| | +-- Pilsbryoconcha_exilis **
+--+
| +----- Potomida_littoralis **
+--+
| +-- Lamprotula_leaii **
+--+
+-- Pronodularia_japanensis **
Bolotov et al., 2023 [Publ 4312], fig. S1
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 39 |
39 |
37 |
1 |
0 |
0 |
0 |
107.0 |
+----------- Indopseudodon_kayinensis **
+-----------------+
| | +-------- Indopseudodon_indawgyiensis **
| +--+
| | +----- Indopseudodon_bogani **
| +--+
| | +-- Indopseudodon_salwenianus **
| +--+
| +-- Indopseudodon_crebristriatus **
|
| +-- Pilsbryoconcha_exilis **
| +-----------+
| | +-- Pilsbryoconcha_schomburgki **
--+ |
| +-----------+ +-- Pilsbryoconcha_carinifera **
| | | +--------+
| | | | +-- Pilsbryoconcha_linguaeformis **
| | +--+
| | | +-------- Pilsbryoconcha_kittitati **
| | +--+
| | | +----- Pilsbryoconcha_acuta **
| | +--+
| | | +-- Pilsbryoconcha_hoikaab **
| | +--+
| | +-- Pilsbryoconcha_mekongiana **
| |
+--+ +-------------- Nyeinchanconcha_nyeinchani **
| |
| | +----- Thaiconcha_thaiensis **
| +--------+ +-----+
| | | | | +-- Thaiconcha_callifera **
| | | | +--+
| | +--+ +-- Thaiconcha_munelliptica **
| | |
| | | +-------- Bineurus_loeiensis **
| | +--+
| | | +----- Bineurus_mouhotii **
| | +--+
+--+ | +-- Bineurus_anodontinum **
| +--+
| +-- Bineurus_exilis **
|
| +-------------------- Songkhlanaia_tamodienica **
| |
| | +-- Namkongnaia_lemeslei **
| | +--------+
| | | +-- Namkongnaia_inkhavilayi **
| | |
+--+ +-----+ +-- Pseudodon_inoscularis **
| | | +-----+
| | | | +-- Pseudodon_lenyanensis **
| | | |
| | +--+ +-- Pseudodon_vagulus **
| | | +--+
| | | | +-- Pseudodon_cambodjensis **
+--+ +--+
| | +-- Pseudodon_vondembuschianus **
| +--+
| +-- Pseudodon_mekongi **
|
| +-------------- Sundadontina_tumida **
| |
| | +-------- Sundadontina_sulcata **
+--+ +--+
| | | +----- Sundadontina_harmandi **
| | +--+
| | | +-- Sundadontina_brandti **
+--+ +--+
| +-- Sundadontina_taskaevi **
|
| +-------- Sundadontina_tanintharyiensis **
+--+
| +----- Sundadontina **
+--+
| +-- Sundadontina_cumingii **
+--+
+-- Sundadontina_plugpomenica **
Burdick & White, 2007 [Publ 2679], fig. 3
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 38 |
38 |
31 |
1 |
0 |
0 |
0 |
107.0 |
+----------------------------------------- Lampsilis_ornata **
|
--+ +-------------------------------------- Parvaspina_collina **
| |
| | +----------------------------------- Reginaia_ebenus **
+--+ |
| | +----- Eurynia_dilatata **
| | +--------------------------+
| | | | +-- Plethobasus_cyphyus **
+--+ | +--+
| | +-- Elliptio_crassidens **
| |
| | +----------------------------- Pleurobema_plenum **
| | |
| | | +-- Pleuronaia_barnesiana **
+--+ +--------------------------+
| | +-- Pleuronaia_gibber **
| |
| | +-------------- Pleurobema_stabile **
| | |
| | | +-- Pleurobema_rubellum **
| | | +-----+
| | | | +-- Pleurobema_strodeanum **
+--+ +-----------+ +--+
| | | | | +----- Pleurobema_beadleianum **
| | | | +--+
| | | | | +-- Pleurobema_perovatum **
| | | | +--+
| | +--+ +-- Pleurobema_taitianum **
| | |
| | | +-- Pleurobema_clava **
| | | +-----+
| | | | +-- Pleurobema_oviforme **
| | | |
| | +--+ +-- Pleurobema_decisum **
+--+ | +--+
| | | +-- Pleurobema_hanleyianum **
| +--+
| | +-- Pleurobema_pyriforme **
| +--+
| +-- Pleurobema_georgianum **
|
| +----- Pleurobema_sintoxia[1] **
| |
| +-----------------+ +-- Pleurobema_cordatum[2] **
| | | |
| | +--+-- Pleurobema_sintoxia[2] **
| | |
+--+ +-- Pleurobema_rubrum **
|
| +-------------------- Pleurobema_cordatum[1] **
| |
+--+ +----------------- Fusconaia_subrotunda **
| |
| | +-- Fusconaia_cor **
+--+ +-----------+
| | +-- Fusconaia_cuneolus **
| |
+--+ +----------- Fusconaia_ozarkensis **
| |
| | +-- Fusconaia_masoni **
+--+ +-----+
| | +-- Fusconaia_flava[3] **
| |
+--+ +----- Fusconaia_flava[1] **
| |
| +----- Fusconaia_cerina[4] **
+--+
| +-- Fusconaia_cerina[1] **
| |
| +-- Fusconaia_cerina[2] **
+--+
+-- Fusconaia_cerina[3] **
|
+-- Fusconaia_flava[2] **
Boyer et al., 2011 [Publ 2718], fig. 4
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 38 |
38 |
33 |
1 |
0 |
0 |
0 |
103.0 |
+-------------- Strophitus_undulatus **
|
| +----- Lasmigona_complanata **
+--------------------+ +-----+
| | | | +-- Lasmigona_costata **
| | | +--+
| +--+ +-- Arcidens_confragosus **
| |
| | +-------- Utterbackia_imbecillis **
| | |
| +--+-------- Alasmidonta_marginata **
| |
| | +----- Pyganodon_grandis **
| +--+
--+ | +-- Anodontoides_ferussacianus **
| +--+
| +-- Platynaias_compressa **
|
| +----------- Theliderma_metanevra **
| |
| +--------------------+ +-- Quadrula_verrucosa **
| | | +-----+
| | | | +-- Quadrula_quadrula **
| | +--+
| | | +----- Pustulosa_pustulosa **
| | +--+
+--+ | +-- Cyclonaias_tuberculata **
| +--+
| +-- Pustulosa_nodulata **
|
| +----------------------------- Megalonaias_nervosa **
| |
+--+ +-------------------------- Toxolasma_parvum **
| |
| | +----------------------- Amblema_plicata **
+--+ |
| | +-- Reginaia_ebenus **
| | +-----------------+
| | | +-- Obliquaria_reflexa **
+--+ |
| | +-------- Pleurobema_sintoxia **
| +-----------+
| | | +----- Elliptio_crassidens **
| | +--+
| | | +-- Fusconaia_flava **
+--+ +--+
| +-- Eurynia_dilatata **
|
| +----- Potamilus_fragilis **
| +-----------+
| | | +-- Potamilus_ohiensis **
| | +--+
| | +-- Potamilus_alatus **
| |
+--+ +----- Ellipsaria_lineolata **
| +--------+
| | | +-- Truncilla_donaciformis **
| | +--+
| | +-- Truncilla_truncata **
+--+
| +----------- Lampsilis_cardium **
| |
| | +----- Ligumia_recta **
| +-----+
+--+ | +-- Obovaria_olivaria **
| +--+
| +-- Epioblasma_triquetra **
|
| +-------- Lampsilis_teres **
+--+
| +----- Lampsilis_siliquoidea **
+--+
| +-- Ortmanniana_ligamentina **
+--+
+-- Lampsilis_higginsii **
Breton et al., 2011 [Publ 3376], fig. 3
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 38 |
38 |
35 |
2 |
1 |
1 |
1 |
101.0 |
+-------------------------------------- Echyridella_menziesii
|
| +-------- Cumberlandia_monodonta **
| +--------------------------+
| | | +----- Margaritifera_margaritifera **
| | +--+
--+ | | +-- Margaritifera_falcata **
| | +--+
| | +-- Margaritifera_marrianae **
| |
| | +----- Quadrula_quadrula **
| | +--------------------------+
+--+ | | +-- Cyclonaias_tuberculata **
| | +--+
| | +-- Pustulosa_pustulosa **
| |
| | +-------------------------- Pronodularia_japanensis **
| | |
| | +--+ +----------------------- Sinohyriopsis_cumingii **
| | | | |
| | | +--+ +-------------------- Cristaria_plicata **
| | | | |
+--+ | | | +-- Lasmigona_complanata **
| | +--+ +--------------+
| | | | +-- Lasmigona_costata **
| | | |
| | +--+ +-------------- Strophitus_undulatus **
| | | |
| | | | +-- Platynaias_compressa **
| | | | +--+
| | +--+ | +-- Platynaias_subviridis **
| | | +-----+
| | | | | +-- Alasmidonta_marginata **
+--+ | | +--+
| | | +-- Alasmidonta_undulata **
| +--+
| | +-- Pyganodon_lacustris **
| | +-----+
| | | +-- Pyganodon_grandis **
| +--+
| | +----- Utterbackia_peninsularis[2] **
| +--+
| | +-- Utterbackia_peninsularis[1] **
| +--+
| +-- Utterbackia_imbecillis **
|
| +-------------------------- Megalonaias_nervosa **
| |
+--+ +----------------------- Fusconaia_flava **
| |
+--+ +-------------------- Reginaia_ebenus **
| |
| | +-------- Toxolasma_paulum[2] **
+--+ +--------+
| | | +----- Toxolasma_lividum **
| | +--+
| | | +-- Toxolasma_parvum **
+--+ +--+
| +-- Toxolasma_paulum[1] **
|
| +-------------- Ellipsaria_lineolata **
| |
+--+ +----------- Truncilla_macrodon **
| |
| | +-- Lemiox_rimosus **
+--+ +-----+
| | +-- Potamilus_metnecktayi **
| |
+--+ +-- Cambarunio_iris **
| +--+
| | +-- Venustaconcha_ellipsiformis **
+--+
| +-- Lampsilis_powellii **
| |
+--+-- Lampsilis_ornata **
|
+-- Lampsilis_siliquoidea **
Chase et al., 2018 [Publ 3847], fig. 4
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 38 |
39 |
37 |
2 |
1 |
1 |
1 |
95.0 |
+-------- Cumberlandia_monodonta **
+--------------------+
| | +----- Margaritifera_falcata **
| +--+
| | +-- Margaritifera_marrianae **
| +--+
| +-- Margaritifera_margaritifera **
|
| +-- Unio_delphinus **
| +--------------------+
| | +-- Unio_tumidus **
| |
| +--+ +-------------------- Sinanodonta_woodiana **
| | | |
| | | | +-- Anodonta_cygnea **
| | | | +--------------+
--+ | +--+ | +-- Anodonta_anatina **
| | | |
| | | | +-- Pyganodon_lacustris **
| | | | +-----+
| | +--+ | +-- Pyganodon_grandis **
| | | +-----+
| | | | | +----- Utterbackia_peninsularis[2] **
| | | | +--+
| | | | | +-- Utterbackia_peninsularis[1] **
| | | | +--+
| | +--+ +-- Utterbackia_imbecillis **
| | |
| | | +-- Lasmigona_costata **
| | | +--------+
| | | | +-- Lasmigona_complanata **
| | | |
+--+ +--+ +-- Alasmidonta_undulata **
| | +-----+
| | | +-- Alasmidonta_marginata **
| +--+
| | +----- Strophitus_undulatus **
| +--+
| | +-- Platynaias_compressa **
| +--+
| +-- Platynaias_subviridis **
|
| +-- Echyridella_menziesii
| +--------+
| | +-- Reginaia_ebenus **
| +--------+
| | | +-------- Fusconaia_flava **
| | +--+
| | | +----- Megalonaias_nervosa **
| | +--+
| | | +-- Cyclonaias_tuberculata **
| | +--+
+-----+ +-- Pustulosa_pustulosa **
|
| +-------- Toxolasma_parvum **
| +--------+
| | | +----- Toxolasma_lividum **
| | +--+
| | | +-- Toxolasma_paulum **
| | +--+
+--+ +-- Toxolasma_texasiense **
|
| +-------------- Venustaconcha_ellipsiformis **
| |
| | +----- Lemiox_rimosus **
+--+ +-----+
| | | +-- Lampsilis_powellii **
| | +--+
+--+ +-- Lampsilis_siliquoidea **
|
| +-------- Potamilus_metnecktayi **
+--+
| +----- Cambarunio_iris **
+--+
| +-- Ellipsaria_lineolata **
+--+
+-- Truncilla_macrodon **
Wu et al., 2024 [Publ 4350], fig. 5a
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 38 |
38 |
36 |
2 |
0 |
0 |
0 |
96.0 |
+-- Margaritifera_dahurica **
+--------------------------+
| +-- Margaritifera_margaritifera **
|
| +-------------------------- Chamberlainia_somsakpanhai **
| |
| | +-- Khairuloconcha_lunbawangorum **
| | +--------------+
--+ | | +-- Khairuloconcha_sahanae **
| | |
| | | +-------- Ensidens_sagittarius **
| | +-----+ +-----+
| | | | | | +----- Rectidens_sumatrensis **
| | | | | +--+
| | | | | | +-- Hyriopsis_bialata **
| | | | | +--+
+--+ | +--+ +-- Hyriopsis_desowitzi **
| | |
| | | +-- Physunio_superbus **
| | | +--------+
| | | | +-- Trapezoideus_foliaceus **
| | | |
| | +--+ +-- Lens_contradens **
| | | +-----+
| | | | +-- Lens_eximius **
| | +--+
| | | +----- Yaukthwa_elongatula **
| | +--+
| | | +-- Yaukthwa_inlenensis **
+--+ +--+
| +-- Yaukthwa_paiensis **
|
| +-- Schepmania[1] **
| +--------------+
| | +-- Schepmania[2] **
| |
| +--+ +-- Indopseudodon_bogani **
| | | +-----------+
| | | | +-- Indopseudodon_kayinensis **
| | +--+
| | | +----------- Pilsbryoconcha_exilis **
| | | |
| | +--+ +-- Pseudodon_mekongi **
| | | +-----+
| | | | +-- Pseudodon_vondembuschianus **
| | +--+
| | | +----- Thaiconcha_callifera **
+--+ +--+
| | +-- Bineurus_anodontinum **
| +--+
| +-- Bineurus_loeiensis **
|
| +-- Inversidens_brandtii **
| +-----+
| | +-- Inversidens_rentianensis **
| +--------+
| | | +----- Potomida_littoralis **
| | +--+
| | | +-- Lamprotula_caveata **
+--+ +--+
| +-- Lamprotula_leaii **
|
| +-------------- Sinosolenaia_carinata **
| |
+--+ +-- Gonidea_angulata **
| +--------+
| | +-- Microcondylaea_bonellii **
+--+
| +-------- Parvasolenaia_rivularis **
+--+
| +----- Koreosolenaia_sitgyensis **
+--+
| +-- Postolata_guangxiensis **
+--+
+-- Ptychorhynchus_pfisteri **
Wu et al., 2024 [Publ 4350], fig. 5b
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 38 |
38 |
36 |
2 |
0 |
0 |
0 |
98.0 |
+-- Margaritifera_dahurica **
+-----------------------------------+
| +-- Margaritifera_margaritifera **
|
| +-- Schepmania[1] **
| +--------------------------------+
| | +-- Schepmania[2] **
| |
--+ | +-- Indopseudodon_bogani **
| | +-----------+
| | | +-- Indopseudodon_kayinensis **
| | +-----------------+
| | | | +----------- Pilsbryoconcha_exilis **
| | | | |
| | | +--+ +-- Bineurus_anodontinum **
+--+ | | +-----+
| | | | +-- Bineurus_loeiensis **
| | +--+
| | | +----- Thaiconcha_callifera **
| | +--+
| | | +-- Pseudodon_mekongi **
| | +--+
| | +-- Pseudodon_vondembuschianus **
| |
| | +-------------------------- Hyriopsis_bialata **
| | |
+--+ | +----- Rectidens_sumatrensis **
| +--+ +-----------------+
| | | | | +-- Khairuloconcha_lunbawangorum **
| | | | +--+
| | +--+ +-- Khairuloconcha_sahanae **
| | |
| | | +-------------------- Ensidens_sagittarius **
| | | |
| | +--+ +----------------- Hyriopsis_desowitzi **
| | | |
| | +--+ +-------------- Yaukthwa_elongatula **
| | | |
| | +--+ +----------- Trapezoideus_foliaceus **
| | | |
| | +--+ +-- Physunio_superbus **
+--+ | +-----+
| | | +-- Yaukthwa_inlenensis **
| +--+
| | +----- Yaukthwa_paiensis **
| +--+
| | +-- Lens_contradens **
| +--+
| +-- Lens_eximius **
|
| +-- Inversidens_brandtii **
| +-----------------+
| | +-- Inversidens_rentianensis **
| |
| | +-- Chamberlainia_somsakpanhai **
+--------+ +--+
| | +-- Potomida_littoralis **
| +-----------+
| | | +-- Lamprotula_caveata **
| | +--+
+--+ +-- Lamprotula_leaii **
|
| +-------------- Sinosolenaia_carinata **
| |
+--+ +-- Gonidea_angulata **
| +--------+
| | +-- Microcondylaea_bonellii **
+--+
| +-------- Koreosolenaia_sitgyensis **
+--+
| +----- Parvasolenaia_rivularis **
+--+
| +-- Postolata_guangxiensis **
+--+
+-- Ptychorhynchus_pfisteri **
Wu et al., 2024 [Publ 4350], fig. 5c
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 38 |
38 |
25 |
2 |
0 |
0 |
0 |
97.0 |
+-- Margaritifera_dahurica **
+-----------------------------+
| +-- Margaritifera_margaritifera **
|
| +----------------------------- Schepmania[1] **
| |
| +----------------------------- Schepmania[2] **
| |
--+ | +-- Indopseudodon_bogani **
| | +-----------+
| | | +-- Indopseudodon_kayinensis **
| | +-----------+
| | | | +----------- Pilsbryoconcha_exilis **
| | | | |
| | | +--+ +-------- Bineurus_anodontinum **
+--+ | | |
| | +--+-------- Bineurus_loeiensis **
| | |
| | | +----- Thaiconcha_callifera **
| | +--+
| | | +-- Pseudodon_mekongi **
| | +--+
| | +-- Pseudodon_vondembuschianus **
| |
| | +-------- Ensidens_sagittarius **
| | |
| | +-------- Hyriopsis_desowitzi **
+--+ |
| +-------- Rectidens_sumatrensis **
| |
| +-------- Hyriopsis_bialata **
| |
| +--------------+ +-- Khairuloconcha_lunbawangorum **
| | +-----+
| | | +-- Khairuloconcha_sahanae **
| | |
| | | +----- Physunio_superbus **
| | | |
| | | +----- Yaukthwa_paiensis **
| | | |
| | | +----- Yaukthwa_inlenensis **
| | +--+
| | +----- Trapezoideus_foliaceus **
+--+ |
| +----- Yaukthwa_elongatula **
| |
| | +-- Lens_contradens **
| +--+
| +-- Lens_eximius **
|
| +-- Inversidens_rentianensis **
| +-----------------+
| | +-- Inversidens_brandtii **
| |
| | +-- Chamberlainia_somsakpanhai **
+--+ +--+
| | +-- Potomida_littoralis **
| +-----------+
| | | +-- Lamprotula_caveata **
| | +--+
+--+ +-- Lamprotula_leaii **
|
| +-------------- Sinosolenaia_carinata **
| |
+--+ +----------- Gonidea_angulata **
| |
+--+----------- Microcondylaea_bonellii **
|
| +-------- Koreosolenaia_sitgyensis **
+--+
| +----- Parvasolenaia_rivularis **
+--+
| +-- Postolata_guangxiensis **
+--+
+-- Ptychorhynchus_pfisteri **
Wu et al., 2024 [Publ 4350], fig. 6-7
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 38 |
38 |
36 |
2 |
0 |
0 |
0 |
96.0 |
+-- Margaritifera_dahurica **
+-----------------------------+
| +-- Margaritifera_margaritifera **
|
| +----------------------------- Chamberlainia_somsakpanhai **
| |
| | +-- Khairuloconcha_lunbawangorum **
| | +--------------+
--+ | | +-- Khairuloconcha_sahanae **
| | |
| | | +-------- Ensidens_sagittarius **
| | +--------+ +-----+
| | | | | | +----- Rectidens_sumatrensis **
| | | | | +--+
| | | | | | +-- Hyriopsis_bialata **
| | | | | +--+
+--+ | +--+ +-- Hyriopsis_desowitzi **
| | |
| | | +-- Physunio_superbus **
| | | +--------+
| | | | +-- Trapezoideus_foliaceus **
| | | |
| | +--+ +-- Lens_contradens **
| | | +-----+
| | | | +-- Lens_eximius **
| | +--+
| | | +----- Yaukthwa_elongatula **
| | +--+
+--+ | +-- Yaukthwa_inlenensis **
| +--+
| +-- Yaukthwa_paiensis **
|
| +-- Schepmania[1] **
| +--------------+
| | +-- Schepmania[2] **
| |
| +-----+ +-- Indopseudodon_bogani **
| | | +-----------+
| | | | +-- Indopseudodon_kayinensis **
| | +--+
| | | +----------- Pilsbryoconcha_exilis **
| | | |
| | +--+ +-- Pseudodon_mekongi **
| | | +-----+
| | | | +-- Pseudodon_vondembuschianus **
| | +--+
+--+ | +----- Thaiconcha_callifera **
| +--+
| | +-- Bineurus_anodontinum **
| +--+
| +-- Bineurus_loeiensis **
|
| +----- Potomida_littoralis **
| +--------------+
| | | +-- Lamprotula_caveata **
| | +--+
| | +-- Lamprotula_leaii **
+--+
| +-- Inversidens_brandtii **
| +--------------+
| | +-- Inversidens_rentianensis **
+--+
| +-------------- Sinosolenaia_carinata **
| |
+--+ +-- Gonidea_angulata **
| +--------+
| | +-- Microcondylaea_bonellii **
+--+
| +-------- Koreosolenaia_sitgyensis **
+--+
| +----- Parvasolenaia_rivularis **
+--+
| +-- Postolata_guangxiensis **
+--+
+-- Ptychorhynchus_pfisteri **
Perkins et al., 2017 [Publ 3777], fig. 2
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 37 |
37 |
34 |
1 |
0 |
0 |
0 |
109.0 |
+----------------------------------------- Amblema_plicata **
|
+----------------------------------------- Plectomerus_dombeyanus **
|
| +-- Pleurobema_clava **
| +-----------+
| | +-- Pleurobema_oviforme **
| |
| | +-------- Pleurobema_pyriforme **
| +-----------------------+ +--+
--+ | | | | +----- Pleurobema_georgianum **
| | | | +--+
| | | | | +-- Pleurobema_hanleyianum **
| | +--+ +--+
| | | +-- Pleurobema_decisum **
| | |
| | | +-------- Pleurobema_rubellum **
| | +--+
| | | +----- Pleurobema_perovatum **
| | +--+
+--+ | +-- Pleurobema_beadleianum **
| +--+
| +-- Pleurobema_strodeanum **
|
| +----- Pleuronaia_gibber **
| +-----------------------------+
| | | +-- Pleuronaia_barnesiana **
| | +--+
| | +-- Pleuronaia_dolabelloides **
| |
| | +-- Fusconaia_masoni **
+--+ +--+
| | +-- Fusconaia_flava **
| +--------------------------+
| | | +-- Fusconaia_subrotunda **
| | +--+
| | +-- Fusconaia_cor **
+--+
| +-- Parvaspina_collina **
| +--------------------------+
| | +-- Parvaspina_steinstansana **
| |
+--+ +-- Eurynia_dilatata **
| +-----------------------+
| | +-- Hemistena_lata **
+--+
| +----------------------- Elliptio_producta[2] **
| |
+--+ +-------------------- Elliptio_producta[1] **
| |
+--+ +----------------- Elliptio_spinosa **
| |
| | +-------------- Elliptio_icterina **
+--+ |
| | +-------- Elliptio_roanokensis **
| | +--+
+--+ | | +----- Elliptio_complanata **
| | +--+
| | | +-- Elliptio_crassidens **
| | +--+
+--+ +-- Elliptio_pullata **
|
| +-- Elliptio_congaraea **
| +-----+
| | +-- Elliptio_hopetonensis **
| |
+--+ +-- Elliptio_folliculata **
| +--+
| | +-- Elliptio_jayensis **
+--+
| +-- Elliptio_arca **
+--+
+-- Elliptio_arctata **
Pieri et al., 2018 [Publ 3867], fig. 3
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 37 |
37 |
23 |
1 |
0 |
0 |
0 |
91.0 |
+----------------------------------------- Arcidens_confragosus **
|
| +-- Quadrula_quadrula **
| +-----------------------------------+
--+ | +-- Pustulosa_petrina **
| |
| | +----------------------------------- Eurynia_dilatata **
| | |
+--+ | +----- Pleurobema_clava **
| +-----------------------------+
| | | +-- Pleurobema_riddellii **
| | +--+
| | +-- Pleurobema_sintoxia **
+--+
| +----- Fusconaia_subrotunda **
| +--------------------------+
| | | +-- Fusconaia_cor **
| | +--+
| | +-- Fusconaia_cuneolus **
| |
+--+ +----------------------------- Fusconaia_ozarkensis **
| |
| | +----------- Fusconaia_cerina[3] **
| | |
| | +--------------+ +-------- Fusconaia_flava[1] **
+--+ | | |
| | +--+ +----- Fusconaia_flava[3] **
| | | |
| | +--+ +-- Fusconaia_cerina[1] **
| | | |
| | +--+-- Fusconaia_cerina[2] **
+--+ |
| +-- Fusconaia_flava[2] **
|
| +----------------------- Fusconaia_masoni **
| |
| | +-- Fusconaia_burkei **
| +--------------------+
+--+ +-- Fusconaia_escambia **
|
| +-------------------- Fusconaia_mitchelli **
| |
+--+ +----------------- Fusconaia_chunii **
| |
| | +-------------- Fusconaia_askewi[9] **
+--+ |
| | +----------- Fusconaia_askewi[6] **
| | |
+--+ +----------- Fusconaia_askewi[7] **
| |
| +----------- Fusconaia_askewi[8] **
| |
| +----------- Fusconaia_lananensis[5] **
+--+
+----------- Fusconaia_lananensis[6] **
|
| +-------- Fusconaia_askewi[5] **
| |
| +-------- Fusconaia_lananensis[4] **
| |
+--+ +----- Fusconaia_askewi[2] **
| |
| +----- Fusconaia_askewi[3] **
| |
+--+----- Fusconaia_askewi[4] **
|
| +-- Fusconaia_askewi[1] **
| |
| +-- Fusconaia_lananensis[1] **
+--+
+-- Fusconaia_lananensis[2] **
|
+-- Fusconaia_lananensis[3] **
Mabuchi & Nishida, 2020 [Publ 4120], fig. 1
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 37 |
37 |
12 |
1 |
0 |
0 |
0 |
82.0 |
+-------- Obovalis_omiensis **
+-----------+
| | +----- Inversidens_brandtii **
| +--+
| | +-- Sinohyriopsis_schlegelii **
| +--+
| +-- Pronodularia_japanensis **
--+
| +----- Nodularia_douglasiae **
| +-----------+
| | | +-- Lanceolaria_grayii **
| | +--+
| | +-- Inversiunio_reinianus **
| |
+--+ +-------------- Cristaria_plicata **
| |
| | +----------- Sinanodonta_tumens **
| | |
| | | +-- Sinanodonta_lauta[5] **
| | | |
+--+ | +-- Sinanodonta_lauta[6] **
| | |
| | +-- Beringiana_japonica[15] **
| | |
| | +-- Beringiana_japonica[16] **
| | +-----+
| | | +-- Beringiana_japonica[17] **
+--+ | |
| | +-- Beringiana_japonica[18] **
| | |
| | +-- Beringiana_japonica[19] **
| | |
| | +-- Beringiana_japonica[20] **
| |
| | +----- Beringiana_japonica[1] **
| | |
| | +----- Beringiana_japonica[2] **
| | |
+--+ +----- Beringiana_japonica[3] **
| |
| +----- Beringiana_japonica[4] **
| |
| +----- Beringiana_japonica[5] **
| |
| +----- Sinanodonta_lauta[1] **
| |
| +----- Beringiana_japonica[6] **
| |
| +----- Sinanodonta_lauta[2] **
| |
| +----- Sinanodonta_calipygos[1] **
| |
+--+----- Beringiana_japonica[7] **
|
+----- Sinanodonta_lauta[4] **
|
+----- Beringiana_japonica[8] **
|
+----- Beringiana_japonica[9] **
|
+----- Beringiana_japonica[10] **
|
+----- Beringiana_japonica[11] **
|
+----- Beringiana_japonica[12] **
|
+----- Beringiana_japonica[13] **
|
+----- Beringiana_japonica[14] **
|
| +-- Sinanodonta_lauta[3] **
+--+
+-- Sinanodonta_calipygos[2] **
Bolotov et al., 2020 [Publ 4084], fig. 1
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 36 |
36 |
22 |
2 |
0 |
0 |
0 |
81.0 |
+-- Margaritifera_dahurica **
|
+-- Margaritifera_margaritifera **
+--------------------------+
| +-- Margaritifera_laevis **
| |
| +-- Margaritifera_middendorffi **
|
| +-- Amblema_plicata **
| |
| +-- Ortmanniana_ligamentina **
| |
| +-- Lampsilis_cardium **
| |
+--------------------------+-- Cambarunio_iris **
| |
| +-- Elliptio_complanata **
| |
--+ +-- Pleurobema_sintoxia **
| |
| +-- Quadrula_quadrula **
|
| +-- Lamprotula_leaii **
| |
| +-- Potomida_littoralis **
| |
| +-- Pronodularia_japanensis **
| +-----------------------+
| | +-- Gonidea_angulata **
| | |
| | +-- Leguminaia_wheatleyi **
| | |
| | +-- Microcondylaea_bonellii **
| |
+--+ +----------- Middendorffinaia_mongolica **
| +-----------+
| | | +-------- Nodularia_douglasiae **
| | +--+
| | | +----- Unio_tumidus **
| | +--+
| | | +-- Unio_pictorum **
| | +--+
+--+ +-- Unio_crassus **
|
| +-- Lanceolaria_grayii **
| +-----------------+
| | +-- Lanceolaria **
| |
| | +----- Anodonta_anatina **
+--+ +-----------+
| | | +-- Pseudanodonta_complanata **
| | +--+
| | +-- Anodonta_cygnea **
| |
+--+ +-------- Amuranodonta_kijaensis **
| +-----+
| | | +----- Anemina_arcaeformis **
| | +--+
| | | +-- Buldowskia_suifunica **
+--+ +--+
| +-- Buldowskia_shadini **
|
| +----------- Cristaria_plicata **
+--+
| +-------- Beringiana_beringiana **
+--+
| +----- Sinanodonta_schrenkii **
+--+
| +-- Sinanodonta_lauta **
+--+
+-- Sinanodonta_woodiana **
Ortiz-Sepulveda et al., 2020 [Publ 4101], fig. 1
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 36 |
36 |
22 |
1 |
0 |
0 |
0 |
108.0 |
+-------------------------------------- Leoparreysia_tavoyensis **
|
| +----------- Indonaia_andersoniana **
| |
| +-----------------------+ +-------- Indochinella_pugio **
| | | |
--+ | | | +----- Radiatula_mouhoti **
| | +--+ |
| | | +----- Radiatula_myitkyinae **
| | | |
| | +--+----- Radiatula_chaudhurii **
| | |
| | | +-- Scabies_humilis **
+--+ | |
| +--+-- Scabies_phaselus **
| |
| +-- Scabies_mandarinus **
|
| +-------------------------------- Grandidieria_burtoni **
| |
| | +-- Pseudospatha_tanganyicensis **
| | +--------------------------+
+--+ | +-- Moncetia_anceyi **
| |
| | +-- Nitia_acuminata **
| | |
+--+ +-----------------------+-- Nitia_monceti **
| | |
| | +-- Nitia_teretiuscula **
| |
| | +-- Coelatura_aegyptiaca **
| | |
+--+ +-- Coelatura_alluaudi **
| |
| +--------------------+-- Coelatura_bakeri **
| | |
| | +-- Coelatura_hauttecoeuri **
| | |
| | +-- Coelatura_stuhlmanni **
+--+
| +-- Coelatura_leopoldvillensis **
| +-----------------+
| | +-- Coelatura_mesafricana **
| |
| | +-- Coelatura_horei **
+--+ |
| +--------------+-- Coelatura_ratidota **
| | |
| | +-- Coelatura **
+--+
| +-------------- Coelatura_gabonensis **
| |
| +-------------- Coelatura_stanleyvillensis **
+--+
| +----------- Coelatura_kunenensis **
| |
| | +----- Prisodontopsis_aviculaeformis **
+--+ +--+
| | | +-- Coelatura_briarti **
| | +--+
| | +-- Prisodontopsis_mweruensis **
+--+
| +-- Coelatura_choziensis **
| |
| +-- Coelatura_hypsiprymna **
| |
+-----+-- Coelatura_luapulaensis **
|
+-- Coelatura_mossambicensis **
|
+-- Nyassunio_nyassaensis **
Ortiz-Sepulveda et al., 2020 [Publ 4101], fig. 2
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 36 |
36 |
20 |
1 |
0 |
0 |
0 |
108.0 |
+-------------------------------- Leoparreysia_tavoyensis **
|
| +----- Radiatula_mouhoti **
| |
| +----- Radiatula_myitkyinae **
| |
| +----- Radiatula_chaudhurii **
| |
--+ +-----------------------+----- Indochinella_pugio **
| | |
| | +----- Indonaia_andersoniana **
| | |
| | | +-- Scabies_humilis **
| | | |
| | +--+-- Scabies_phaselus **
| | |
+--+ +-- Scabies_mandarinus **
|
| +----- Grandidieria_burtoni **
| +--------------------+
| | | +-- Pseudospatha_tanganyicensis **
| | +--+
| | +-- Moncetia_anceyi **
| |
| | +-- Nitia_acuminata **
+--+ |
| +--------------------+-- Nitia_monceti **
| | |
| | +-- Nitia_teretiuscula **
| |
| | +-- Coelatura_aegyptiaca **
| | |
+--+ +-- Coelatura_alluaudi **
| |
| +-----------------+-- Coelatura_bakeri **
| | |
| | +-- Coelatura_hauttecoeuri **
| | |
| | +-- Coelatura_stuhlmanni **
+--+
| +-- Coelatura_leopoldvillensis **
| +--------------+
| | +-- Coelatura_mesafricana **
| |
| | +-- Coelatura_horei **
+--+ |
| +-----------+-- Coelatura_ratidota **
| | |
| | +-- Coelatura **
| |
+--+ +-- Coelatura_gabonensis **
| +--------+
| | +-- Coelatura_stanleyvillensis **
| |
+--+ +-------- Coelatura_kunenensis **
| |
| | +-- Coelatura_briarti **
| | |
+--+ +--+-- Prisodontopsis_mweruensis **
| | |
| | +-- Prisodontopsis_aviculaeformis **
| |
+--+ +-- Coelatura_choziensis **
| |
| +-- Coelatura_hypsiprymna **
| |
+--+-- Coelatura_luapulaensis **
|
+-- Coelatura_mossambicensis **
|
+-- Nyassunio_nyassaensis **
Jeratthitikul et al., 2021 [Publ 4219], fig. 1
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 36 |
36 |
33 |
1 |
0 |
0 |
0 |
87.0 |
+----- Lens_contradens **
+--+
| | +-- Trapezoideus_foliaceus **
| +--+
+--------------------+ +-- Physunio_superbus **
| |
| | +----- Ensidens_ingallsianus **
| +--+
| | +-- Hyriopsis_bialata **
| +--+
| +-- Rectidens_sumatrensis **
|
| +----- Sinosolenaia_carinata **
| +-----+
| | | +-- Leguminaia_wheatleyi **
--+ | +--+
| | +-- Gonidea_angulata **
| +--------------+
| | | +-- Chamberlainia_somsakpanhai **
| | | +-----+
| | | | +-- Sinohyriopsis_cumingii **
| | +--+
| | | +----- Potomida_littoralis **
| | +--+
| | | +-- Pronodularia_japanensis **
| | +--+
+--+ +-- Lamprotula_leaii **
|
| +-------- Indopseudodon_kayinensis **
| +--------------+
| | | +----- Indopseudodon_indawgyiensis **
| | +--+
| | | +-- Indopseudodon_salwenianus **
| | +--+
| | +-- Indopseudodon_bogani **
| |
+--+ +----- Pilsbryoconcha_exilis **
| +--------------+
| | | +-- Pilsbryoconcha_linguaeformis **
| | +--+
| | +-- Pilsbryoconcha **
| |
| | +-------- Nyeinchanconcha_nyeinchani **
| | |
+--+ +--------+ +----- Thaiconcha_callifera **
| | | |
| | +--+ +-- Bineurus_panhai **
| | | |
| | +--+-- Bineurus_exilis **
| | |
| | +-- Bineurus_mouhotii **
+--+
| +-- Sundadontina_brandti **
| +--+
| | +-- Sundadontina_taskaevi **
| +--------+
| | | +-- Sundadontina_tumida **
| | +--+
| | +-- Sundadontina_cumingii **
+--+
| +-- Namkongnaia_lemeslei **
| +--------+
| | +-- Namkongnaia_inkhavilayi **
+--+
| +-------- Pseudodon_lenyanensis **
+--+
| +----- Pseudodon_vagulus **
+--+
| +-- Pseudodon_vondembuschianus **
+--+
+-- Pseudodon_mekongi **
Jeratthitikul et al., 2021 [Publ 4219], fig. 2
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 36 |
36 |
34 |
1 |
0 |
0 |
0 |
87.0 |
+----- Lens_contradens **
+--+
| | +-- Trapezoideus_foliaceus **
| +--+
+--------------------+ +-- Physunio_superbus **
| |
| | +----- Ensidens_ingallsianus **
| +--+
| | +-- Hyriopsis_bialata **
| +--+
| +-- Rectidens_sumatrensis **
|
| +----- Sinosolenaia_carinata **
| +-----+
| | | +-- Gonidea_angulata **
--+ | +--+
| | +-- Leguminaia_wheatleyi **
| +--------------+
| | | +-- Chamberlainia_somsakpanhai **
| | | +-----+
| | | | +-- Sinohyriopsis_cumingii **
| | +--+
| | | +----- Potomida_littoralis **
| | +--+
| | | +-- Lamprotula_leaii **
| | +--+
+--+ +-- Pronodularia_japanensis **
|
| +-------- Indopseudodon_kayinensis **
| +--------------+
| | | +----- Indopseudodon_indawgyiensis **
| | +--+
| | | +-- Indopseudodon_salwenianus **
| | +--+
| | +-- Indopseudodon_bogani **
| |
+--+ +----- Pilsbryoconcha_exilis **
| +--------------+
| | | +-- Pilsbryoconcha_linguaeformis **
| | +--+
| | +-- Pilsbryoconcha **
| |
| | +-- Nyeinchanconcha_nyeinchani **
| | +--------+
+--+ | +-- Sundadontina_cumingii **
| +-----+
| | | +-------- Thaiconcha_callifera **
| | +--+
| | | +----- Bineurus_mouhotii **
| | +--+
| | | +-- Bineurus_panhai **
| | +--+
+--+ +-- Bineurus_exilis **
|
| +----- Sundadontina_tumida **
| +--------+
| | | +-- Sundadontina_brandti **
| | +--+
| | +-- Sundadontina_taskaevi **
+--+
| +-- Namkongnaia_lemeslei **
| +--------+
| | +-- Namkongnaia_inkhavilayi **
+--+
| +-------- Pseudodon_lenyanensis **
+--+
| +----- Pseudodon_vagulus **
+--+
| +-- Pseudodon_vondembuschianus **
+--+
+-- Pseudodon_mekongi **
Pavan-Kumar et al., 2022 [Publ 4263], fig. 4
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 36 |
36 |
34 |
2 |
0 |
0 |
0 |
78.0 |
+-- Margaritifera_dahurica **
+-----------------------------+
| +-- Margaritifera_falcata **
|
| +-------------- Quadrula_quadrula **
| |
| +--------------+ +----------- Toxolasma_parvum **
| | | |
| | +--+ +-------- Potamilus_leptodon **
| | | |
--+ | +--+ +-- Lampsilis_cardium **
| | | +--+
| | | | +-- Lampsilis_ornata **
| | +--+
| | | +-- Lampsilis_siliquoidea **
| | +--+
| | +-- Venustaconcha_ellipsiformis **
| |
| | +----- Chamberlainia_somsakpanhai **
+--+ +-----------+
| | | +-- Sinohyriopsis_cumingii **
| | +--+
| | +-- Sinohyriopsis_schlegelii **
| +--------+
| | | +-- Lamprotula_leaii **
| | | +-----------+
| | | | +-- Potomida_littoralis **
| | | |
| | +--+ +-- Pseudodon_vondembuschianus **
| | | +--------+
| | | | +-- Pilsbryoconcha_exilis **
| | +--+
| | | +-------- Parvasolenaia_rivularis **
+--+ +--+
| | +----- Microcondylaea_bonellii **
| +--+
| | +-- Sinosolenaia_carinata **
| +--+
| +-- Sinosolenaia_oleivora **
|
| +----------------------- Lamellidens_marginalis **
| |
| | +-------------------- Lepidodesma_languilati **
+--+ |
| | +-- Aculamprotula_tientsinensis **
| | +--------+
| | | +-- Aculamprotula_tortuosa **
+--+ +-----+
| | | +-------- Unio_pictorum **
| | +--+
| | | +----- Schistodesmus_lampreyanus **
| | +--+
| | | +-- Nodularia_douglasiae **
+--+ +--+
| +-- Tchangsinaia_piscicula **
|
| +-- Lanceolaria_lanceolata **
| +-----------+
| | +-- Lanceolaria_gladiola **
+--+
| +----------- Acuticosta_chinensis **
| |
+--+ +-- Anodonta_anatina **
| +-----+
| | +-- Pyganodon_grandis **
+--+
| +----- Cristaria_plicata **
+--+
| +-- Sinanodonta_lucida **
+--+
+-- Anemina_euscaphys **
Boyer et al., 2011 [Publ 2718], fig. 3
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 35 |
35 |
26 |
1 |
0 |
0 |
0 |
94.0 |
+----- Alasmidonta_marginata **
|
+----- Pyganodon_grandis **
|
+-----------------------+ +-- Lasmigona_complanata **
| +--+
| | +-- Lasmigona_costata **
| |
| | +-- Platynaias_compressa **
| +--+
| +-- Strophitus_undulatus **
|
--+ +-------------------------- Reginaia_ebenus **
| |
| | +-- Fusconaia_flava **
| | +--+
| | | +-- Pleurobema_sintoxia **
| | +--------------+
| | | | +-- Eurynia_dilatata **
| | | +--+
+--+ +--+ +-- Elliptio_crassidens **
| | |
| | | +----------------- Megalonaias_nervosa **
| | +--+
| | | +-------------- Quadrula_quadrula **
| | +--+
| | | +----------- Quadrula_verrucosa **
| | +--+
| | | +-------- Theliderma_metanevra **
+--+ +--+
| | +----- Pustulosa_pustulosa **
| +--+
| | +-- Cyclonaias_tuberculata **
| +--+
| +-- Pustulosa_nodulata **
|
| +----------------- Amblema_plicata **
| |
| | +-------------- Toxolasma_parvum **
+-----+ |
| | +----------- Obliquaria_reflexa **
| | |
+--+ | +-------- Obovaria_olivaria **
| | |
| | +-------- Lampsilis_cardium **
| | |
+--+ | +-- Epioblasma_triquetra **
| +-----+
| | +-- Lampsilis_siliquoidea **
| |
| | +-- Ortmanniana_ligamentina **
| +-----+
| | +-- Lampsilis_higginsii **
+--+
| +-- Ligumia_recta **
+-----+
| +-- Lampsilis_teres **
|
| +----- Potamilus_fragilis **
+--+
| | +-- Potamilus_ohiensis **
| +--+
| +-- Potamilus_alatus **
|
| +----- Ellipsaria_lineolata **
+--+
| +-- Truncilla_truncata **
+--+
+-- Truncilla_donaciformis **
Konopleva et al., 2019 [Publ 4041], fig. 3
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 35 |
35 |
21 |
1 |
0 |
0 |
0 |
94.0 |
+-- Indopseudodon_indawgyiensis **
|
+--------------------+-- Indopseudodon_bogani **
| |
| +-- Indopseudodon_salwenianus **
|
| +-- Ensidens_ingallsianus **
| |
| +-- Ensidens_telus **
| |
| +-- Ensidens_dugasti **
| |
| +-- Hyriopsis_khoratensis **
| |
| +-- Hyriopsis_bialata **
| |
--+ +-- Hyriopsis_myersiana **
| |
| +-----------------+-- Hyriopsis_desowitzi **
| | |
| | +-- Rectidens_sumatrensis **
| | |
| | +-- Hyriopsis_panhai **
| | |
| | +-- Ensidens_jaculus[1] **
| | |
| | +-- Ensidens_jaculus[2] **
| | |
| | +-- Ensidens_spiculus **
| | |
+--+ +-- Ensidens_sagittarius **
|
| +----------- Yaukthwa_elongatula **
| |
| +-----+ +-- Yaukthwa_inlenensis **
| | | +-----+
| | | | +-- Yaukthwa_paiensis **
| | +--+
| | | +----- Yaukthwa_panhai **
| | +--+
| | | +-- Yaukthwa_nesemanni **
| | +--+
+--+ +-- Yaukthwa_rectangularis **
|
| +-------- Trapezoideus_foliaceus **
| +-----+
| | | +----- Physunio_superbus **
| | +--+
| | | +-- Physunio_modelli **
| | +--+
| | +-- Physunio **
+--+
| +----- Lens_misellus **
| +-----+
| | | +-- Lens_contradens **
| | +--+
| | +-- Lens_micropterus **
+--+
| +----- Lens_pallegoixi **
| +--+
| | | +-- Lens_novoselovi **
| | +--+
+--+ +-- Lens_comptus **
|
| +----- Lens_maenamensis **
+--+
| +-- Lens_eximius **
+--+
+-- Lens_rolfbrandti **
Huang et al., 2002 [Publ 2657], fig. 1
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 34 |
34 |
31 |
2 |
0 |
0 |
0 |
91.0 |
+----------------------------------------------- Cumberlandia_monodonta **
|
+----------------------------------------------- Margaritifera_margaritifera **
|
| +-- Sinanodonta_woodiana **
| +-----------+
| | +-- Cristaria_plicata **
| +--+
| | | +----------- Utterbackiana_couperiana **
| | +--+
| | | +-------- Utterbackia_imbecillis **
| | +--+
--+ | | +----- Strophitus_radiatus **
| | +--+
| | | +-- Alasmidonta_triangulata **
| +--------------------------+ +--+
| | | +-- Lasmigona_costata **
| | |
| | | +----- Acuticosta_ovata **
| | | +--+
| | | | | +-- Nodularia_douglasiae **
| | | | +--+
| | | | +-- Tchangsinaia_piscicula **
| | +--------+
+--+ | +-- Lanceolaria_grayii **
| | +--+
| | | +-- Lanceolaria_lanceolata **
| +--+
| | +-- Schistodesmus_lampreyanus **
| +--+
| +-- Lepidodesma_languilati **
|
| +----------------------------------------- Sinosolenaia_oleivora **
| |
+--+ +-------------------------------------- Lamprotula_leaii **
| |
+--+ +----------------------------------- Sinohyriopsis_cumingii **
| |
+--+ +-------------------------------- Ptychorhynchus_pfisteri **
| |
+--+ +----------------------------- Megalonaias_nervosa **
| |
| | +----- Elliptoideus_sloatianus **
+--+ +--------------------+
| | | +-- Uniomerus_carolinianus **
| | +--+
+--+ +-- Pleurobema_pyriforme **
|
| +----------------------- Amblema_plicata **
| |
+--+ +-- Obliquaria_reflexa **
| +-----------------+
| | +-- Plectomerus_dombeyanus **
+--+
| +----------------- Medionidus_conradicus **
+--+
| +-------------- Potamilus_fragilis **
+--+
| +----------- Ptychobranchus_subtentus **
+--+
| +-------- Potamilus_purpuratus **
+--+
| +----- Lampsilis_teres **
+--+
| +-- Villosa_delumbis **
+--+
+-- Ellipsaria_lineolata **
Huang et al., 2002 [Publ 2657], fig. 2
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 34 |
34 |
15 |
2 |
0 |
0 |
0 |
89.0 |
+-------------------- Cumberlandia_monodonta **
|
+-------------------- Margaritifera_margaritifera **
|
| +-------- Strophitus_radiatus **
| |
| +-------- Schistodesmus_lampreyanus **
| |
| +-------- Lepidodesma_languilati **
| |
| +-------- Acuticosta_ovata **
| |
| | +-- Sinanodonta_woodiana **
--+ +-----+
| | +-- Cristaria_plicata **
| +--------+
| | | +-- Lanceolaria_grayii **
| | +-----+
| | | +-- Lanceolaria_lanceolata **
| | |
| | | +-- Nodularia_douglasiae **
| | +-----+
| | | +-- Tchangsinaia_piscicula **
| | |
| | | +----- Utterbackia_imbecillis **
| | | |
+--+ +--+----- Utterbackiana_couperiana **
| |
| | +-- Alasmidonta_triangulata **
| +--+
| +-- Lasmigona_costata **
|
| +-------------- Ptychorhynchus_pfisteri **
| |
| +-------------- Sinohyriopsis_cumingii **
| |
| +-------------- Lamprotula_leaii **
| |
+--+-------------- Sinosolenaia_oleivora **
|
| +----------- Megalonaias_nervosa **
| |
| | +----- Elliptoideus_sloatianus **
| | +--+
+--+ | | +-- Uniomerus_carolinianus **
| | +--+
| | +-- Pleurobema_pyriforme **
| |
| | +----- Potamilus_fragilis **
+--+ |
| +----- Medionidus_conradicus **
| |
| +----- Amblema_plicata **
| |
| | +-- Obliquaria_reflexa **
+--+--+
| +-- Plectomerus_dombeyanus **
|
| +-- Villosa_delumbis **
| |
| +-- Ellipsaria_lineolata **
| |
+--+-- Lampsilis_teres **
|
+-- Potamilus_purpuratus **
|
+-- Ptychobranchus_subtentus **
Krebs et al., 2003 [Publ 2691], fig. 3
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 34 |
34 |
29 |
2 |
0 |
0 |
0 |
100.5 |
+-- Margaritifera_margaritifera **
+-----------------------------+
| +-- Margaritifera_falcata **
|
| +----------- Utterbackiana_couperiana **
| |
| +----------- Platynaias_compressa **
| |
| +----------- Alasmidonta_triangulata **
| |
| +-----------------+ +-- Lasmigona_complanata **
--+ | +--------+
| | | +-- Lasmigona_costata **
| | |
| | | +-- Strophitus_radiatus **
| | | +-----+
| | | | +-- Strophitus_undulatus **
| | +--+
| | | +----- Utterbackia_imbecillis **
| | +--+
| | | +-- Pyganodon_grandis **
+--+ +--+
| +-- Anodonta **
|
| +-------- Megalonaias_nervosa **
| +-----------------+
| | | +----- Quadrula_quadrula[2] **
| | +--+
| | | +-- Quadrula_quadrula[1] **
| | +--+
| | +-- Theliderma_metanevra **
| |
+--+ +----------- Elliptoideus_sloatianus **
| +-----------+
| | | +-------- Pleurobema_pyriforme **
| | +--+
| | | +----- Uniomerus_carolinianus **
| | +--+
| | | +-- Fusconaia_flava **
+--+ +--+
| +-- Eurynia_dilatata **
|
| +-------------------- Amblema_plicata **
| |
| | +----- Toxolasma_parvum **
| | +-----------+
+--+ | | +-- Plectomerus_dombeyanus **
| | +--+
| | +-- Obliquaria_reflexa **
| |
+--+ +----- Ptychobranchus_subtentus **
| +--------+
| | | +-- Potamilus_fragilis **
| | +--+
| | +-- Medionidus_conradicus **
+--+
| +-- Villosa_delumbis **
| +--------+
| | +-- Ellipsaria_lineolata **
+--+
| +-------- Lampsilis_teres **
+--+
| +----- Lampsilis_siliquoidea **
+--+
| +-- Potamilus_purpuratus **
+--+
+-- Potamilus_alatus **
Pfeiffer & Graf, 2013 [Publ 3531], fig. 2
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 34 |
35 |
30 |
2 |
1 |
1 |
1 |
84.5 |
+----------------------------------- Margaritifera_margaritifera **
|
+----------------------------------- Velesunio_ambiguus
|
| +-- Lamellidens_savadiensis **
| +-----------------------------+
| | +-- Coelatura_aegyptiaca **
| |
--+ | +-- Cristaria_plicata **
| | +-----+
| | | +-- Sinanodonta_woodiana **
| | +--------+
| | | | +----- Strophitus_undulatus **
| | | +--+
| | | | +-- Platynaias_compressa **
| | +-----------+ +--+
+--+ | | +-- Pyganodon_grandis **
| | |
| | | +-------------- Acuticosta_chinensis **
| | | |
| | +--+ +-- Unio_pictorum **
| | | +--------+
| | | | +-- Nodularia_douglasiae **
| | +--+
| | | +-------- Aculamprotula_tortuosa **
| | | |
| | +--+ +----- Aculamprotula_scripta **
| | | |
+--+ +--+ +-- Aculamprotula_tientsinensis **
| | |
| +--+-- Aculamprotula_fibrosa **
| |
| +-- Aculamprotula_zonata **
|
| +----------- Quadrula_quadrula **
| +--------------+
| | | +-------- Eurynia_dilatata **
| | +--+
| | | +----- Amblema_plicata **
| | +--+
| | | +-- Lampsilis_ornata **
+--+ +--+
| +-- Truncilla_truncata **
|
| +-- Sinohyriopsis_cumingii **
| +--------------------+
| | +-- Sinohyriopsis_schlegelii **
| |
+--+ +----- Gonidea_angulata **
| +--------------+
| | | +-- Pseudodon_vagulus **
| | +--+
+--+ +-- Sundadontina_tumida **
|
| +----------------- Physunio **
| |
+--+ +-------------- Pronodularia_japanensis **
| |
+--+ +----------- Potomida_littoralis **
| |
+--+ +-------- Lamprotula_leaii **
| |
+--+ +----- Lamprotula_caveata[2] **
| |
+--+ +-- Lamprotula_cornuumlunae **
| |
+--+-- Gibbosula_rochechouartii **
|
+-- Lamprotula_caveata[1] **
Sano et al., 2017 [Publ 3799], fig. 2
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 34 |
34 |
31 |
2 |
2 |
0 |
0 |
91.0 |
+-- Margaritifera_laevis **
+-----------------------+
| +-- Margaritifera_middendorffi **
|
| +-- Obovalis_omiensis **
| +--------+
| | +-- Ptychorhynchus_pfisteri **
| |
| +-----------+ +-- Inversidens_brandtii **
| | | +-----+
| | | | +-- Sinosolenaia_oleivora **
--+ | | |
| | +--+ +-- Pronodularia_japanensis **
| | | +--+
| | | | +-- Lamprotula_leaii **
| | +--+
| | | +-- Sinohyriopsis_schlegelii **
| | +--+
| | +-- Sinohyriopsis_cumingii **
| |
| | +-- Cristaria_plicata **
+--+ +-----------+
| | +-- Buldowskia_iwakawai **
| |
| +-----+ +----- Beringiana_fukuharai **
| | | +-----+
| | | | | +-- Beringiana_japonica **
| | | | +--+
| | +--+ +-- Beringiana_beringiana **
| | |
| | | +-------- Pletholophus_tenuis **
| | +--+
| | | +----- Sinanodonta_woodiana **
| | +--+
+--+ | +-- Sinanodonta_lauta **
| +--+
| +-- Sinanodonta_tumens **
|
| +-------- Tchangsinaia_piscicula **
| +--------+
| | | +----- Nodularia_nipponensis **
| | +--+
| | | +-- Nodularia_douglasiae **
| | +--+
| | +-- Nodularia_breviconcha **
+--+
| +-- Schistodesmus_lampreyanus **
| +-----------+
| | +-- Acuticosta_ovata **
| |
| | +-------- Lepidodesma_languilati **
+--+ +--+
| | | +----- Lanceolaria_grayii[1] **
| | +--+
| | | +-- Lanceolaria_lanceolata **
+--+ +--+
| +-- Lanceolaria_grayii[2] **
|
| +-------- Inversiunio_jokohamensis **
| |
+--+ +----- Inversiunio_yanagawensis[3] **
| |
+--+ +-- Inversiunio_yanagawensis[1] **
| |
+--+-- Inversiunio_yanagawensis[2] **
|
+-- Inversiunio_reinianus **
Chen et al., 2020 [Publ 4170], fig. 1
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 34 |
34 |
32 |
2 |
0 |
0 |
0 |
103.0 |
+----- Prolasmidonta_heterodon **
+--+
| | +-- Alasmidonta_varicosa **
| +--+
| +-- Anemina_euscaphys **
+-----------------+
| | +-- Lamprotula_caveata **
| | +--+
| | | +-- Chamberlainia_somsakpanhai **
| +--+
| | +-- Pseudodon_vondembuschianus **
| +--+
| +-- Pilsbryoconcha_exilis **
|
| +-------- Schistodesmus_lampreyanus **
| +-----+
| | | +----- Lepidodesma_languilati **
| | +--+
--+ | | +-- Aculamprotula_coreana **
| | +--+
| +--------+ +-- Aculamprotula_tortuosa **
| | |
| | | +----- Platynaias_compressa **
| | | +-----+
| | | | | +-- Utterbackia_imbecillis **
| | | | +--+
| | +--+ +-- Pyganodon_grandis **
| | |
| | | +-------- Sinanodonta_lucida **
| | +--+
| | | +----- Cristaria_plicata **
| | +--+
+--+ | +-- Anodonta_cygnea **
| +--+
| +-- Anodonta_anatina **
|
| +----------------- Gibbosula_crassa **
| +--+
| | | +-------------- Quadrula_quadrula **
| | +--+
| | | +----------- Toxolasma_parvum **
| | +--+
| | | +-------- Lampsilis_ornata **
| | +--+
| | | +----- Potamilus_leptodon **
+--+ +--+
| | +-- Lampsilis_siliquoidea **
| +--+
| +-- Lampsilis_powellii **
|
| +-- Utterbackia_peninsularis **
| +--------------+
| | +-- Potamilus_alatus **
+--+
| +-------------- Lamprotula_leaii **
| |
+--+ +-- Sinohyriopsis_cumingii **
| +--------+
| | +-- Sinohyriopsis_schlegelii **
+--+
| +-------- Parvasolenaia_rivularis **
+--+
| +----- Sinosolenaia_carinata **
+--+
| +-- Sinosolenaia_iridinea **
+--+
+-- Sinosolenaia_oleivora **
Lopes-Lima et al., 2017 [Publ 3792], fig. 2
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 33 |
33 |
31 |
2 |
0 |
0 |
0 |
83.0 |
+----- Pseudunio_marocanus **
+--------------------+
| | +-- Margaritifera_falcata **
| +--+
| +-- Margaritifera_dahurica **
|
| +-------- Quadrula_quadrula **
| +--------------+
| | | +----- Toxolasma_parvum **
--+ | +--+
| | | +-- Lampsilis_ornata **
| | +--+
| | +-- Venustaconcha_ellipsiformis **
| |
| | +-- Sinohyriopsis_cumingii **
| | +--------+
| | | +-- Sinohyriopsis_schlegelii **
| | |
+--+ +--------+ +-- Sinosolenaia_oleivora **
| | | +-----+
| | | | +-- Sinosolenaia_carinata **
| | +--+
| | | +----- Potomida_littoralis **
| | +--+
| | | +-- Pronodularia_japanensis **
| | +--+
| | +-- Lamprotula_leaii **
| |
| | +-- Lanceolaria_lanceolata **
| | +-----------+
+--+ | +-- Lanceolaria_grayii **
| |
| +--+ +-- Aculamprotula_tortuosa **
| | | +--------+
| | | | +-- Aculamprotula_coreana **
| | | |
| | +--+ +-- Unio_delphinus **
| | | +-----+
| | | | +-- Unio_pictorum **
| | +--+
| | | +----- Schistodesmus_lampreyanus **
| | +--+
+--+ | +-- Tchangsinaia_piscicula **
| +--+
| +-- Nodularia_douglasiae **
|
| +----------- Anodonta_anatina **
| +--+
| | | +-------- Platynaias_compressa **
| | +--+
| | | +----- Pyganodon_grandis **
| | +--+
+--+ | +-- Utterbackia_peninsularis **
| +--+
| +-- Utterbackia_imbecillis **
|
| +-------- Cristaria_plicata **
| |
+-----+ +-- Sinanodonta_lucida **
| +--+
| | +-- Sinanodonta_woodiana **
+--+
| +-- Anemina_arcaeformis **
+--+
+-- Anemina_euscaphys **
Burlakova et al., 2019 [Publ 3899], fig. 1
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 33 |
33 |
21 |
1 |
0 |
0 |
0 |
107.0 |
+-- Quadrula_quadrula **
+-----------------------+
| +-- Uniomerus_declivis **
|
| +----------------------- Plectomerus_dombeyanus **
--+ |
| | +-------------------- Popenaias_popeii **
| | |
| | | +-- Elliptio_crassidens **
| | +-----------------+
+--+ | +-- Pleurobema_rubellum **
| |
| | +-- Cyrtonaias_tampicoensis **
| | +--+
| | | +-- Glebula_rotundata **
| +--------------+
| | | +-- Reginaia_ebenus **
+--+ +--+
| +-- Obliquaria_reflexa **
|
| +----------------- Toxolasma_lividum **
| |
| | +-- Potamilus_fragilis **
| | +-----------+
| | | +-- Potamilus_metnecktayi **
| | |
| | | +----- Medionidus_acutissimus **
| | +--------+
+--+ | | +-- Lemiox_rimosus **
| | +--+
| | +-- Dromus_dromas **
| |
| | +----- Ellipsaria_lineolata **
| | |
| +--------+ +-- Truncilla_cognata **
| | | |
+--+ | +-- Truncilla_donaciformis **
| +--+
| +-- Truncilla_macrodon **
| |
| +-- Truncilla_truncata **
|
| +----------- Cambarunio_simus **
| |
| +----------- Leaunio_lienosus **
| |
| +----------- Villosa_villosa **
| |
| | +-- Sagittunio_nasutus **
| +--------+
+--+ +-- Lampsilis_ovata **
|
| +----- Venustaconcha_pleasii **
+-----+
| | +-- Epioblasma_rangiana **
| +--+
| +-- Obovaria_unicolor **
|
| +-------- Lampsilis_teres **
+--+
| +----- Lampsilis_siliquoidea **
+--+
| +-- Lampsilis_hydiana **
+--+
+-- Lampsilis_straminea **
Pfeiffer & Graf, 2015 [Publ 3620], fig. 2
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 32 |
32 |
30 |
2 |
8 |
7 |
2 |
72.0 |
+-------- Velesunio_ambiguus
+--------------------------+
| | +----- Hyridella_australis
| +--+
| | +-- Triplodon_corrugatus
| +--+
| +-- Castalia_ambigua
|
--+ +----- Etheria_elliptica
| +--------------------------+
| | | +-- Lamproscapha_ensiformis
| | +--+
| | +-- Chambardia_wahlbergi
| |
| | +-- Margaritifera_margaritifera **
| | +--------------------------+
+--+ | +-- Cumberlandia_monodonta **
| |
| | +-- Trapezidens_dolichorhynchus **
| | +-----------+
| | | +-- Lamellidens_savadiensis **
| | |
| | +-----------+ +-- Leoparreysia_olivacea **
+--+ | | +--------+
| | | | +-- Leoparreysia_tavoyensis **
| | +--+
| | | +-------- Coelatura_aegyptiaca **
| | +--+
| | | +----- Indochinella_pugio **
| | +--+
| | | +-- Radiatula_chaudhurii **
| | +--+
+--+ +-- Scabies_mandarinus **
|
| +----- Unio_pictorum **
| +-----------------+
| | | +-- Pyganodon_grandis **
| | +--+
| | +-- Lasmigona_costata **
| |
| | +----------- Quadrula_quadrula **
| | +--------+
+--+ | | +-------- Amblema_plicata **
| | +--+
| | | +----- Potamilus_alatus **
| | +--+
| | | +-- Lampsilis_cardium **
| | +--+
| | +-- Truncilla_truncata **
+--+
| +-------- Ensidens_telus **
| +--------+
| | | +----- Lens_micropterus **
| | +--+
| | | +-- Physunio **
| | +--+
| | +-- Physunio_modelli **
+--+
| +-------- Gonidea_angulata **
| +-----+
| | | +----- Potomida_littoralis **
| | +--+
| | | +-- Pronodularia_japanensis **
+--+ +--+
| +-- Lamprotula_caveata **
|
| +----------- Pilsbryoconcha_carinifera **
| |
+--+ +-- Pseudodon_vondembuschianus **
| +-----+
| | +-- Pseudodon_vagulus **
+--+
| +----- Sundadontina_tumida **
+--+
| +-- Nyeinchanconcha_nyeinchani **
+--+
+-- Bineurus_exilis **
Fonseca et al., 2016 [Publ 3775], fig. 1
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 32 |
32 |
30 |
2 |
0 |
0 |
0 |
82.0 |
+-- Margaritifera_falcata **
+-----------------------+
| +-- Margaritifera_dahurica **
|
| +-------- Quadrula_quadrula **
| +--------------+
| | | +----- Toxolasma_parvum **
| | +--+
--+ | | +-- Lampsilis_ornata **
| | +--+
| | +-- Venustaconcha_ellipsiformis **
| |
| | +-- Sinohyriopsis_cumingii **
| | +--------+
| | | +-- Sinohyriopsis_schlegelii **
| | |
+--+ +--------+ +-- Sinosolenaia_oleivora **
| | | +-----+
| | | | +-- Sinosolenaia_carinata **
| | +--+
| | | +----- Potomida_littoralis **
| | +--+
| | | +-- Lamprotula_leaii **
| | +--+
| | +-- Pronodularia_japanensis **
| |
| | +-- Lanceolaria_lanceolata **
| | +-----------+
+--+ | +-- Lanceolaria_grayii **
| |
| +--+ +-- Aculamprotula_tortuosa **
| | | +--------+
| | | | +-- Aculamprotula_coreana **
| | | |
| | +--+ +-- Unio_pictorum **
| | | +-----+
| | | | +-- Unio_delphinus **
| | +--+
| | | +----- Schistodesmus_lampreyanus **
| | +--+
+--+ | +-- Nodularia_douglasiae **
| +--+
| +-- Tchangsinaia_piscicula **
|
| +-------- Cristaria_plicata **
| |
| +-----+ +-- Anemina_euscaphys **
| | | +--+
| | | | +-- Anemina_arcaeformis **
| | +--+
+--+ | +-- Sinanodonta_woodiana **
| +--+
| +-- Sinanodonta_lucida **
|
| +----------- Anodonta_anatina **
+--+
| +-------- Platynaias_compressa **
+--+
| +----- Pyganodon_grandis **
+--+
| +-- Utterbackia_peninsularis **
+--+
+-- Utterbackia_imbecillis **
Wang et al., 2018 [Publ 3846], fig. 2
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 32 |
32 |
30 |
1 |
0 |
0 |
0 |
74.0 |
+----------- Quadrula_quadrula **
|
+--+ +-------- Toxolasma_parvum **
| | |
| +--+ +-- Venustaconcha_ellipsiformis **
| | +--+
| | | +-- Lampsilis_ornata **
| +--+
| | +-- Potamilus_leptodon **
+--------+ +--+
| | +-- Potamilus_alatus **
| |
| | +-- Sinohyriopsis_cumingii **
| | +--------+
| | | +-- Sinohyriopsis_schlegelii **
| | |
| +--+ +----- Potomida_littoralis **
| | +--+
| | | | +-- Lamprotula_caveata **
| | | +--+
| +--+ +-- Lamprotula_leaii **
| |
| | +----- Ptychorhynchus_pfisteri **
| +--+
--+ | +-- Sinosolenaia_carinata **
| +--+
| +-- Sinosolenaia_oleivora **
|
| +----- Anodonta_anatina **
| +-----+
| | | +-- Pyganodon_grandis **
| | +--+
| +--------+ +-- Platynaias_compressa **
| | |
| | | +-------- Cristaria_plicata **
| | | |
| | +--+ +-- Anemina_euscaphys **
| | | +--+
| | | | +-- Anemina_arcaeformis **
| | +--+
+--+ | +-- Sinanodonta_woodiana **
| +--+
| +-- Sinanodonta_lucida **
|
| +-- Lanceolaria_grayii **
| +--------------+
| | +-- Lanceolaria_lanceolata **
| |
+--+ +-------------- Aculamprotula_tientsinensis **
| |
| | +----- Nodularia_douglasiae **
+--+ +-----+
| | | +-- Tchangsinaia_piscicula **
| | +--+
+--+ +-- Cuneopsis_heudei **
|
| +-------- Unio_tumidus **
+--+
| +----- Unio_crassus **
+--+
| +-- Unio_pictorum **
+--+
+-- Unio_delphinus **
Wang et al., 2018 [Publ 3865], fig. 2
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 32 |
32 |
30 |
1 |
0 |
0 |
0 |
74.0 |
+----------- Quadrula_quadrula **
|
+--+ +-------- Toxolasma_parvum **
| | |
| +--+ +-- Potamilus_leptodon **
| | +--+
| | | +-- Potamilus_alatus **
| +--+
| | +-- Venustaconcha_ellipsiformis **
+-----+ +--+
| | +-- Lampsilis_ornata **
| |
| | +-- Sinohyriopsis_schlegelii **
| | +--------+
| | | +-- Sinohyriopsis_cumingii **
| | |
| +--+ +----- Ptychorhynchus_pfisteri **
| | +--+
| | | | +-- Sinosolenaia_oleivora **
| | | +--+
| +--+ +-- Sinosolenaia_carinata **
| |
| | +----- Potomida_littoralis **
--+ +--+
| | +-- Lamprotula_caveata **
| +--+
| +-- Lamprotula_leaii **
|
| +-------------- Aculamprotula_tientsinensis **
| |
| +--+ +-- Tchangsinaia_piscicula **
| | | +--------+
| | | | +-- Nodularia_douglasiae **
| | +--+
| | | +-------- Unio_tumidus **
| | +--+
| | | +----- Unio_crassus **
| | +--+
+--+ | +-- Unio_pictorum **
| +--+
| +-- Unio_delphinus **
|
| +----- Acuticosta_chinensis **
| +--------+
| | | +-- Lanceolaria_grayii **
| | +--+
| | +-- Lanceolaria_lanceolata **
+--+
| +----- Anodonta_anatina **
| +-----+
| | | +-- Pyganodon_grandis **
| | +--+
+--+ +-- Platynaias_compressa **
|
| +-------- Cristaria_plicata **
| |
+--+ +-- Anemina_euscaphys **
| +--+
| | +-- Anemina_arcaeformis **
+--+
| +-- Sinanodonta_woodiana **
+--+
+-- Sinanodonta_lucida **
Zieritz et al., 2021 [Publ 4218], fig. 5
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 32 |
32 |
30 |
2 |
0 |
0 |
0 |
92.0 |
+-- Margaritifera_margaritifera **
+-----------------------------------+
| +-- Pseudunio_auricularius **
|
| +-- Amblema_plicata **
--+ +--------------------------------+
| | +-- Lampsilis_ovata **
| |
| | +-------------------------------- Chamberlainia_somsakpanhai **
+--+ |
| | +----------------------------- Potomida_littoralis **
| | |
| | | +-- Gonidea_angulata **
+--+ | +--------------------+
| | | +-- Microcondylaea_bonellii **
| | +--+
| | | | +-------------------- Schepmania **
| | | | |
| | | +--+ +-- Indopseudodon_bogani **
+--+ | | +--------------+
| | | | +-- Indopseudodon_crebristriatus **
| | +--+
| | | +-------------- Pilsbryoconcha_exilis **
| | | |
| | +--+ +-- Pseudodon_mekongi **
| | | +--------+
| | | | +-- Pseudodon_vondembuschianus **
| | +--+
+--+ | +-------- Sundadontina **
| +--+
| | +----- Thaiconcha_callifera **
| +--+
| | +-- Bineurus_loeiensis **
| +--+
| +-- Bineurus_anodontinum **
|
| +----- Ctenodesma_borneensis **
| +--------------+
| | | +-- Khairuloconcha_lunbawangorum **
| | +--+
| | +-- Khairuloconcha_sahanae **
| |
+-----+ +-------- Ensidens_sagittarius **
| +--------+
| | | +----- Rectidens_sumatrensis **
| | +--+
| | | +-- Hyriopsis_bialata **
+--+ +--+
| +-- Hyriopsis_desowitzi **
|
| +-------------- Physunio_superbus **
| |
+--+ +----------- Trapezoideus_foliaceus **
| |
+--+ +-- Lens_contradens **
| +-----+
| | +-- Lens_eximius **
+--+
| +----- Yaukthwa_elongatula **
+--+
| +-- Yaukthwa_inlenensis **
+--+
+-- Yaukthwa_paiensis **
Zieritz et al., 2020 [Publ 4102], fig. 4
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 31 |
31 |
27 |
1 |
0 |
0 |
0 |
90.0 |
+----------------------------------- Chamberlainia_somsakpanhai **
|
| +-------------------------------- Lamprotula_leaii **
--+ |
| | +----------------------------- Gonidea_angulata **
| | |
+--+ | +-- Pilsbryoconcha_carinifera **
| | +-----------------------+
| | | +-- Indopseudodon_indawgyiensis **
| | |
+--+ | +-------------- Khairuloconcha_lunbawangorum **
| | |
| | | +-- Ensidens_telus **
| | | +-----+
| | | | +-- Ensidens_ingallsianus **
| | +--------+ +--+
| | | | | | +----- Ensidens_sagittarius **
+--+ | | | +--+
| | | | | +-- Ensidens_spiculus **
| | | | +--+
| | +--+ +-- Ensidens_jaculus **
| | |
| | | +-------- Hyriopsis_desowitzi **
| | | |
| | | +-------- Rectidens_sumatrensis **
| | +--+
| | | +----- Hyriopsis_panhai **
+--+ | |
| +--+ +-- Hyriopsis_khoratensis **
| | |
| +--+-- Hyriopsis_bialata **
| |
| +-- Hyriopsis_myersiana **
|
| +-- Physunio_modelli **
| +-----------------+
| | +-- Physunio_superbus **
| |
| | +-- Yaukthwa_paiensis **
| | +-----+
+--+ | +-- Yaukthwa_inlenensis **
| +--------+
| | | +----- Yaukthwa_panhai **
| | +--+
| | | +-- Yaukthwa_rectangularis **
+--+ +--+
| +-- Yaukthwa_nesemanni **
|
| +-------------- Trapezoideus_foliaceus **
| |
+--+ +-- Lens_contradens **
| +--------+
| | +-- Lens_misellus **
+--+
| +-------- Lens_eximius **
+--+
| +----- Lens_pallegoixi **
+--+
| +-- Lens_comptus **
+--+
+-- Lens_novoselovi **
Lyubas et al., 2022 [Publ 4255], fig. 7
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 31 |
31 |
28 |
1 |
0 |
0 |
0 |
92.0 |
+-- Parreysia_corrugata **
+-----------------------------------+
| +-- Parreysia_keralaensis **
|
| +----------------------------------- Microcondylaea_bonellii **
--+ |
| | +----------- Pseudanodonta_complanata **
| | +--------------------+
| | | | +-------- Anodonta_anatina **
+--+ | +--+
| | | +----- Anodonta_cygnea **
| | +--+
| | | +-- Anodonta_exulcerata **
| | +--+
+--+ +-- Anodonta_seddoni **
|
| +----------------------------- Unio_tumidus **
| |
| | +-- Unio_durieui **
| | +-----------------------+
| | | +-- Unio_gibbus **
+--+ |
| | +-------------- Unio_damascensis **
| | |
| | +-------------- Unio_sesirmensis **
| | +--------+
+--+ | | +----------- Unio_ionicus **
| | | |
| | +--+ +-- Unio_bruguierianus **
| | | +-----+
| | | | +-- Unio_tumidiformis **
| | +--+
| | | +----- Unio_crassus[2] **
+--+ +--+
| | +-- Unio_carneus **
| +--+
| +-- Unio_crassus[1] **
|
| +----- Unio_caffer **
| +--------------+
| | | +-- Unio_terminalis **
| | +--+
+--+ +-- Unio_delicatus **
|
| +----------------- Unio_pictorum **
| |
+--+ +-------------- Unio_eucirrus **
| |
+--+ +-- Unio_elongatulus **
| +--------+
| | +-- Unio_mancus **
+--+
| +-------- Unio_ravoisieri **
| |
+--+ +-- Unio_delphinus **
| +--+
| | +-- Unio_foucauldianus **
+--+
| +-- Unio_hueti **
+--+
+-- Unio_tigridis **
Lyubas et al., 2022 [Publ 4255], fig. S1
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 31 |
31 |
22 |
1 |
0 |
0 |
0 |
93.0 |
+-- Parreysia_corrugata **
+-----------------------+
| +-- Parreysia_keralaensis **
|
| +----------------------- Microcondylaea_bonellii **
--+ |
| | +----------- Anodonta_anatina **
| | +--------+
| | | | +-------- Pseudanodonta_complanata **
| | | +--+
+--+ | | +----- Anodonta_exulcerata **
| | +--+
| | | +-- Anodonta_seddoni **
| | +--+
| | +-- Anodonta_cygnea **
| |
+--+ +----------------- Unio_tumidus **
| |
| | +-- Unio_durieui **
| +--------------+
| | +-- Unio_gibbus **
| |
| | +----------- Unio_sesirmensis **
| | |
| | +--+ +-------- Unio_damascensis **
+--+ | | |
| | | | +----- Unio_ionicus **
| | +--+ |
| | | | +-- Unio_bruguierianus **
| | | +--+
| | +--+ +-- Unio_tumidiformis **
| | |
| | | +-- Unio_carneus **
| | | |
| | +--+-- Unio_crassus[1] **
+--+ |
| +-- Unio_crassus[2] **
|
| +-------- Unio_elongatulus **
| |
| +-------- Unio_mancus **
| |
| | +-- Unio_pictorum **
| +-----+
| | +-- Unio_eucirrus **
| |
+-----+ +----- Unio_caffer **
+--+
| | +-- Unio_terminalis **
| +--+
| +-- Unio_delicatus **
|
| +----- Unio_ravoisieri **
| |
| | +-- Unio_delphinus **
+--+--+
| +-- Unio_foucauldianus **
|
| +-- Unio_hueti **
+--+
+-- Unio_tigridis **
Bogan et al., 2023 [Publ 4295], fig. 3
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 31 |
31 |
28 |
2 |
0 |
0 |
0 |
92.0 |
+-- Gibbosula_crassa **
+-----------------------------------+
| +-- Margaritifera_laevis **
|
--+ +----------------------------------- Schistodesmus_lampreyanus **
| |
| | +-------------------------------- Anodonta_cygnea **
+--+ |
| | +-------- Cristaria_bellua **
| | +--------------------+
| | | | +----- Cristaria_clessini **
+--+ | +--+
| | | +-- Cristaria_plicata **
| | +--+
| | +-- Cristaria_truncata **
| |
| | +-------------- Amuranodonta_kijaensis **
+--+ |
| +-----------+ +----------- Anemina_arcaeformis **
| | | |
| | +--+ +-------- Buldowskia_suifunica **
| | | |
| | +--+ +-- Buldowskia_flavotincta **
| | | +--+
| | | | +-- Buldowskia_shadini **
| | +--+
+--+ | +-- Buldowskia_iwakawai **
| +--+
| +-- Buldowskia_kamiyai **
|
| +-- Pletholophus_honglinhensis **
| +--------------------+
| | +-- Pletholophus_tenuis **
| |
| | +-------- Beringiana_japonica **
+--+ +-----------+
| | | +----- Beringiana_beringiana **
| | +--+
| | | +-- Beringiana_fukuharai **
| | +--+
+--+ +-- Beringiana_gosannensis **
|
| +-- Sinanodonta_angula **
| +--------------+
| | +-- Sinanodonta_lucida **
| |
+--+ +-------------- Sinanodonta_schrenkii **
| |
| | +-- Sinanodonta_calipygos **
+--+-----------+
| +-- Sinanodonta_tumens **
|
| +----------- Sinanodonta_woodiana[2] **
+--+
| +-------- Sinanodonta_jourdyi **
+--+
| +----- Sinanodonta_elliptica **
+--+
| +-- Sinanodonta_lauta **
+--+
+-- Sinanodonta_woodiana[1] **
Campbell et al., 2008 [Publ 2675], fig. 5
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 30 |
30 |
7 |
1 |
0 |
0 |
0 |
102.0 |
+-------------- Uniomerus_declivis **
|
+-------------- Plectomerus_dombeyanus **
|
| +-- Elliptio_arctata **
| +--------+
| | +-- Elliptio_crassidens **
| |
| | +-------- Pleurobema_decisum **
--+ | |
| | +-------- Pleurobema_rubellum **
| | |
| | +-------- Pleurobema_georgianum[2] **
| | |
| | +-------- Pleurobema_stabile **
| | |
| | +-------- Pleurobema_pyriforme **
| | |
+--+ +-------- Pleurobema_oviforme[2] **
| |
| +-------- Pleurobema_hanleyianum[2] **
| |
| +-------- Pleuronaia_gibber **
| |
| +-------- Pleurobema_georgianum[3] **
| |
| +-------- Pleurobema_cordatum **
| |
| +-------- Pleurobema_beadleianum **
| |
| +-------- Hemistena_lata **
+--+
+-------- Fusconaia_flava **
|
+-------- Fusconaia_cerina **
|
+-------- Eurynia_dilatata **
|
+-------- Elliptio_arca **
|
+-------- Cyrtonaias_tampicoensis **
|
| +-- Plethobasus_cyphyus **
+-----+
| +-- Parvaspina_collina **
|
| +-- Pleurobema_athearni **
| |
+-----+-- Pleurobema_georgianum[1] **
| |
| +-- Pleurobema_hanleyianum[1] **
|
| +----- Pleurobema_oviforme[1] **
| |
+--+----- Pleurobema_clava **
|
| +-- Pleurobema_rubrum **
+--+
+-- Pleurobema_sintoxia **
Wu et al., 2020 [Publ 4147], fig. 3
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 30 |
30 |
28 |
2 |
0 |
0 |
0 |
76.0 |
+----------------------------- Margaritifera_falcata **
|
| +----- Quadrula_quadrula **
| +--------------------+
| | | +-- Toxolasma_parvum **
| | +--+
--+ | +-- Venustaconcha_ellipsiformis **
| |
| | +-- Sinohyriopsis_schlegelii **
| | +--------------+
| | | +-- Sinohyriopsis_cumingii **
| | |
| | +-----+ +-- Sinosolenaia_oleivora **
+--+ | | +-----------+
| | | | +-- Sinosolenaia_carinata **
| | +--+
| | | +----------- Pronodularia_japanensis **
| | +--+
| | | +-------- Lamprotula_leaii **
| | +--+
| | | +----- Lamprotula_caveata[1] **
| | +--+
| | | +-- Lamprotula_cornuumlunae **
| | +--+
+--+ +-- Lamprotula_caveata[2] **
|
| +----- Cristaria_plicata **
| +-----+
| | | +-- Sinanodonta_lucida **
| | +--+
| +--------+ +-- Anemina_arcaeformis **
| | |
| | | +-------- Anodonta_anatina **
| | +--+
| | | +----- Pyganodon_grandis **
| | +--+
| | | +-- Utterbackia_peninsularis **
+--+ +--+
| +-- Utterbackia_imbecillis **
|
| +-------- Aculamprotula_tortuosa **
| +--------+
| | | +----- Aculamprotula_scripta **
| | +--+
| | | +-- Aculamprotula_tientsinensis **
+--+ +--+
| +-- Aculamprotula_coreana **
|
| +-------------- Unio_pictorum **
+--+
| +----------- Pseudocuneopsis_capitata **
+--+
| +-------- Nodularia_douglasiae **
+--+
| +----- Schistodesmus_lampreyanus **
+--+
| +-- Cuneopsis_heudei **
+--+
+-- Tchangsinaia_piscicula **
Serb et al., 2003 [Publ 2659], fig. 2
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 29 |
29 |
20 |
1 |
0 |
0 |
0 |
101.0 |
+-- Lampsilis_siliquoidea **
+--------------------+
| +-- Lampsilis_ornata **
|
| +-------------------- Obliquaria_reflexa **
| |
| +-------------------- Amblema_plicata **
| |
| +-------------------- Reginaia_rotulata **
| |
--+ +-------------------- Potamilus_alatus **
| |
| | +-- Elliptoideus_sloatianus **
| +-----------------+
| | +-- Plectomerus_dombeyanus **
| |
| | +----- Medionidus_conradicus **
| +--------------+
| | | +-- Dromus_dromas **
+--+ +--+
| +-- Cyprogenia_aberti **
|
| +-------- Hemistena_lata **
+-----------+
| | +----- Elliptio_complanata **
| +--+
| | +-- Fusconaia_burkei **
| +--+
| +-- Fusconaia_flava **
|
| +----------------- Megalonaias_nervosa **
| |
| | +-------- Theliderma_cylindrica **
| | +-----+
+--+ | | +----- Theliderma_intermedia **
| | +--+
| | | +-- Theliderma_sparsa **
| | +--+
| | +-- Theliderma_metanevra **
+--+
| +----- Quadrula_verrucosa **
| +-----+
| | | +-- Quadrula_quadrula **
| | +--+
| | +-- Quadrula_nobilis **
+--+
| +-------- Pustulosa_kieneriana **
| |
| +-------- Pustulosa_infucata **
+--+
| +-- Pustulosa_pustulosa **
| +--+
| | +-- Pustulosa_succissa **
+--+
| +-- Pustulosa_nodulata **
+--+
+-- Pustulosa_petrina **
Serb et al., 2003 [Publ 2659], fig. 3
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 29 |
29 |
27 |
1 |
0 |
0 |
0 |
100.0 |
+-- Lampsilis_siliquoidea **
+-----------------------------------------+
| +-- Lampsilis_ornata **
|
--+ +----------------------------------------- Potamilus_alatus **
| |
| | +----- Medionidus_conradicus **
+--+ +--------------------------------+
| | | +-- Dromus_dromas **
| | +--+
| | +-- Cyprogenia_aberti **
+--+
| +----------------------------------- Obliquaria_reflexa **
| |
| | +-------------------------------- Amblema_plicata **
+--+ |
| | +----- Reginaia_rotulata **
| | +-----------------------+
+--+ | | +-- Elliptoideus_sloatianus **
| | +--+
| | +-- Plectomerus_dombeyanus **
| |
+--+ +-------- Hemistena_lata **
| +-----------------+
| | | +----- Elliptio_complanata **
| | +--+
| | | +-- Fusconaia_burkei **
| | +--+
+--+ +-- Fusconaia_flava **
|
| +----------------------- Megalonaias_nervosa **
| |
| | +-- Quadrula_verrucosa **
+--+ +-----------------+
| | +-- Quadrula_nobilis **
| |
+--+ +----------------- Quadrula_quadrula **
| |
| | +-------- Theliderma_cylindrica **
+--+ +-----+
| | | +----- Theliderma_sparsa **
| | +--+
| | | +-- Theliderma_intermedia **
+--+ +--+
| +-- Theliderma_metanevra **
|
| +----------- Pustulosa_kieneriana **
| |
+--+ +-------- Pustulosa_infucata **
| |
+--+ +-- Pustulosa_succissa **
| +--+
| | +-- Pustulosa_pustulosa **
+--+
| +-- Pustulosa_nodulata **
+--+
+-- Pustulosa_petrina **
Campbell et al., 2008 [Publ 2675], fig. 4
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 29 |
29 |
10 |
1 |
0 |
0 |
0 |
101.0 |
+----------------------- Uniomerus_declivis **
|
+----------------------- Plectomerus_dombeyanus **
|
| +-------------------- Cyrtonaias_tampicoensis **
| |
| +-------------------- Eurynia_dilatata **
| |
| +-------------------- Fusconaia_cerina **
| |
--+ +-------------------- Fusconaia_flava **
| |
| +-------------------- Hemistena_lata **
| |
| +-------------------- Plethobasus_cyphyus **
| |
| +-------------------- Parvaspina_collina **
| |
| +-------------------- Pleurobema_cordatum **
| |
| +-------------------- Pleuronaia_gibber **
+--+
+-------------------- Pleurobema_oviforme[2] **
|
+-------------------- Pleurobema_stabile **
|
| +----- Elliptio_arca **
+--------------+
| | +-- Elliptio_crassidens **
| +--+
| +-- Elliptio_arctata **
|
| +----------------- Pleurobema_decisum **
| |
| +----------------- Pleurobema_clava **
| |
| +----------------- Pleurobema_oviforme[1] **
| |
+--+ +-- Pleurobema_rubrum **
+--------------+
| +-- Pleurobema_sintoxia **
|
| +-------------- Pleurobema_beadleianum **
| |
+--+ +----------- Pleurobema_pyriforme **
| |
+--+ +-------- Pleurobema_rubellum **
| |
| +-------- Pleurobema_hanleyianum[2] **
+--+
+-------- Pleurobema_georgianum[2] **
|
| +----- Pleurobema_hanleyianum[1] **
+--+
| +-- Pleurobema_georgianum[1] **
+--+
+-- Pleurobema_athearni **
Campbell et al., 2008 [Publ 2675], fig. 6
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 29 |
29 |
24 |
1 |
0 |
0 |
0 |
99.0 |
+----------------------------------------- Cyrtonaias_tampicoensis **
|
+----------------------------------------- Plectomerus_dombeyanus **
--+
| +-------------------------------------- Parvaspina_collina **
| |
| | +----------------------------------- Hemistena_lata **
+--+ |
| | +----- Plethobasus_cyphyus **
| | +--------------------------+
+--+ | | +-- Elliptio_arca **
| | +--+
| | +-- Elliptio_crassidens **
| |
+--+ +-- Fusconaia_cerina **
| +--------------------------+
| | +-- Fusconaia_flava **
| |
| | +-- Eurynia_dilatata **
+--+ +-----------------------+
| | +-- Pleuronaia_gibber **
| |
| | +-- Pleurobema_cordatum **
+--+ +--+
| | +-- Pleurobema_rubrum[1] **
| +-----------------+
| | | +-- Pleurobema_rubrum[2] **
| | +--+
+--+ +-- Pleurobema_sintoxia **
|
| +-------------------- Pleurobema_stabile **
| |
+--+ +----------------- Pleurobema_pyriforme **
| |
| | +----- Pleurobema_decisum **
+--+ +--------+
| | | +-- Pleurobema_hanleyianum **
| | +--+
| | +-- Pleurobema_georgianum[2] **
+--+
| +-- Pleurobema_georgianum[1] **
| +--------+
| | +-- Pleurobema_athearni **
| |
+--+ +-- Pleurobema_oviforme **
| +-----+
| | +-- Pleurobema_clava **
| |
+--+ +----- Pleurobema_beadleianum **
| |
| +----- Pleurobema_rubellum **
+--+
+----- Pleurobema_strodeanum **
|
| +-- Pleurobema_perovatum **
+--+
+-- Pleurobema_taitianum **
Pfeiffer et al., 2018 [Publ 3827], fig. 2
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 29 |
29 |
26 |
2 |
0 |
0 |
0 |
96.0 |
+----------------------------------------------- Margaritifera_margaritifera **
|
| +-- Trapezidens_dolichorhynchus **
--+ +-----------------------------------------+
| | +-- Lamellidens_savadiensis **
| |
| | +----- Parreysia_keralaensis **
+--+ +-----------------------------------+
| | | +-- Parreysia_nagpoorensis **
| | +--+
| | +-- Parreysia_plagiosoma **
| |
+--+ +-- Leoparreysia_tavoyensis **
| +-----------------------------------+
| | +-- Leoparreysia_olivacea **
| |
| | +----- Nitia_teretiuscula **
+--+ +-----------------------------+
| | | +-- Coelatura_aegyptiaca **
| | +--+
| | +-- Prisodontopsis_aviculaeformis **
+--+
| +----- Indonaia_bonneaudii **
| +--------------------------+
| | | +-- Indonaia_andersoniana **
| | +--+
+--+ +-- Indonaia_caerulea **
|
| +----------------------------- Indochinella_pugio **
| |
+--+ +-- Radiatula_chaudhurii **
| +-----------------------+
| | +-- Radiatula_myitkyinae **
+--+
| +----------------------- Scabiellus_songkramensis **
| |
+--+ +-------------------- Unionetta_fabagina[2] **
| |
+--+ +----------------- Harmandia_somboriensis **
| |
+--+ +-------------- Unionetta_fabagina[1] **
| |
+--+ +-- Scabies_crispata **
| +--------+
| | +-- Scabies_mandarinus **
+--+
| +-------- Scabies_nucleus **
| |
+--+ +-- Scabies_pilata **
| +--+
| | +-- Scabies_humilis **
+--+
| +-- Scabies_phaselus **
| |
+--+-- Scabies_anceps **
|
+-- Scabies_scobinatus **
Lopes-Lima et al., 2019 [Publ 3901], fig. 3
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 29 |
29 |
27 |
2 |
0 |
0 |
0 |
94.0 |
+-- Cumberlandia_monodonta **
+-----------------------------------+
| +-- Margaritifera_falcata **
|
| +-- Anodonta_nuttalliana **
--+ +--------------------------------+
| | +-- Pyganodon_grandis **
| |
| | +-------------------------------- Megalonaias_nervosa **
+--+ |
| | +-- Pleurobema_oviforme **
| | +-----+
| | | +-- Pleurobema_riddellii **
| | |
+--+ +--------------------+ +-- Lampsilis_ornata **
| | | +--+
| | | | +-- Potamilus_leptodon **
| | +--+
| | | +-- Amblema_elliottii **
| | +--+
+--+ +-- Amblema_plicata **
|
| +-------------------------- Uniomerus_tetralasmus **
| |
| | +----------- Theliderma_cylindrica **
| | +-----------+
| | | | +-------- Theliderma_sparsa **
+--+ | +--+
| | | +----- Theliderma_intermedia **
| | +--+
| | | +-- Theliderma_metanevra **
| | +--+
+--+ +-- Theliderma_johnsoni **
|
| +----- Quadrula_quadrula **
| +--------------+
| | | +-- Quadrula_nobilis **
| | +--+
| | +-- Quadrula_verrucosa **
+--+
| +----------------- Pustulosa_kieneriana **
| |
| | +-- Pustulosa_infucata **
+--+ +-----------+
| | +-- Pustulosa_kleiniana **
| |
+--+ +-- Pustulosa_pustulosa **
| +--------+
| | +-- Pustulosa_succissa **
+--+
| +-------- Cyclonaias_tuberculata **
+--+
| +----- Pustulosa_necki **
+--+
| +-- Pustulosa_petrina **
+--+
+-- Pustulosa_nodulata **
Pfeiffer et al., 2021 [Publ 4126], fig. 9
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 29 |
29 |
27 |
1 |
0 |
0 |
0 |
90.0 |
+-------------------------- Lamprotula_cornuumlunae **
|
| +-- Ensidens_ingallsianus **
| +--------+
| | +-- Ensidens_telus **
| +--------+
| | | +-------- Ensidens_spiculus **
| | +--+
| | | +----- Ensidens_dugasti **
| | +--+
| +--+ | +-- Ensidens_jaculus **
--+ | | +--+
| | | +-- Ensidens_sagittarius **
| | |
| | | +----------------- Rectidens_sumatrensis **
| | +--+
| | | +-------------- Hyriopsis_delaportei **
| | +--+
| | | +----------- Hyriopsis_myersiana **
| | +--+
| | | +-------- Hyriopsis_bialata **
| | +--+
| | | +----- Hyriopsis_khoratensis **
+--+ +--+
| | +-- Hyriopsis_kratiensis **
| +--+
| +-- Hyriopsis_sakhonensis **
|
| +-- Trapezoideus_subviridis **
| +-----+
| | +-- Trapezoideus_foliaceus **
| +-----------+
| | | +----- Physunio_superbus **
| | +--+
| | | +-- Physunio_modelli **
| | +--+
+--+ +-- Physunio_pellucidus **
|
| +----------------- Yaukthwa_paiensis **
| |
| | +-------------- Solenaia_emarginata **
+--+ |
| | +-------- Lens_misellus **
| | +--+
+--+ | | +----- Lens_inornatus **
| | +--+
| | | +-- Lens_contradens **
| | +--+
+--+ +-- Lens_micropterus **
|
| +-- Lens_maenamensis **
| +--+
| | +-- Lens_eximius **
+-----+
| +-- Lens_pallegoixi **
+--+
+-- Lens_comptus **
Inoue et al., 2020 [Publ 4128], fig. 2
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 29 |
29 |
17 |
1 |
0 |
0 |
0 |
95.0 |
+-------------------------- Quadrula_quadrula **
|
| +-- Reginaia_ebenus **
| +--------------------+
| | +-- Reginaia_rotulata **
--+ |
| | +----- Amblema_plicata **
| | +--------------+
| | | | +-- Amblema_elliottii **
| | | +--+
+--+ | +-- Amblema_neislerii **
| |
| | +----- Popenaias_popeii **
| | +--+
| | | | +-- Popenaias_berezai **
| | | +--+
| | | +-- Popenaias_semirasa **
+--+ |
| +--------+ +-- Psoronaias_profunda **
| | | |
| | | +-- Psoronaias_semigranosa[1] **
| | | |
| | | +-- Psoronaias_semigranosa[2] **
| | | |
| | +-----+-- Psoronaias_guatemalensis[1] **
| | |
| | +-- Psoronaias_guatemalensis[2] **
+--+ |
| +-- Psoronaias_percompressa **
| |
| +-- Psoronaias_semigranosa[3] **
|
| +-- Glebula_rotundata **
| +-----------+
| | +-- Cyrtonaias_tampicoensis **
| |
+--+ +----------- Potamilus_alatus **
| |
| | +-- Ptychobranchus_fasciolaris **
+--+ +-----+
| | +-- Truncilla_truncata **
| |
| | +----- Ortmanniana_ligamentina **
+--+ |
| +----- Epioblasma_triquetra **
| |
| +----- Lampsilis_cardium **
| |
+--+----- Ligumia_recta **
|
+----- Cambarunio_iris **
|
+----- Pachynaias_spheniopsis **
|
| +-- Cyrtonaias_explicata **
+--+
+-- Sphenonaias_microdon **
Inoue et al., 2020 [Publ 4128], fig. 3
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 29 |
29 |
27 |
1 |
0 |
0 |
0 |
93.0 |
+----------------------------------------- Quadrula_quadrula **
|
| +-- Reginaia_ebenus **
| +-----------------------------------+
--+ | +-- Reginaia_rotulata **
| |
| | +----- Amblema_neislerii **
| | +-----------------------------+
| | | | +-- Amblema_plicata **
+--+ | +--+
| | +-- Amblema_elliottii **
| |
| | +----- Popenaias_popeii **
| | +--------+
| | | | +-- Popenaias_berezai **
| | | +--+
+--+ | +-- Popenaias_semirasa **
| +-----------------+
| | | +----------- Psoronaias_profunda **
| | | |
| | | | +----- Psoronaias_guatemalensis[1] **
| | +--+ +--+
| | | | | +-- Psoronaias_semigranosa[1] **
| | | | +--+
| | +--+ +-- Psoronaias_percompressa **
+--+ |
| | +----- Psoronaias_semigranosa[3] **
| +--+
| | +-- Psoronaias_guatemalensis[2] **
| +--+
| +-- Psoronaias_semigranosa[2] **
|
| +----------------------------- Cyrtonaias_tampicoensis **
| |
+--+ +-------------------------- Ptychobranchus_fasciolaris **
| |
+--+ +----------------------- Ligumia_recta **
| |
+--+ +-------------------- Potamilus_alatus **
| |
+--+ +----------------- Pachynaias_spheniopsis **
| |
| | +-- Epioblasma_triquetra **
+--+ +-----------+
| | +-- Truncilla_truncata **
| |
+--+ +-- Ortmanniana_ligamentina **
| +--------+
| | +-- Lampsilis_cardium **
+--+
| +-------- Cambarunio_iris **
+--+
| +----- Sphenonaias_microdon[2] **
+--+
| +-- Cyrtonaias_explicata **
+--+
+-- Sphenonaias_microdon[1] **
Whelan et al., 2023 [Publ 4287], fig. 3
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 29 |
29 |
27 |
1 |
0 |
0 |
0 |
97.0 |
+-------------------------------- Anodonta_anatina **
|
| +-- Utterbackiana_heardi **
| +--+
| | +-- Utterbackiana_couperiana **
| +-----------------------+
--+ | | +-- Utterbackiana_hartfieldorum **
| | +--+
| | +-- Utterbackiana_suborbiculata **
| |
| | +-- Utterbackia_peninsularis **
| | +-----------------------+
+--+ | +-- Pyganodon_grandis **
| |
| | +-------------------- Alasminota_etowaensis **
| | |
| | | +----- Pressodonta_viridis **
| | +--+ +-----------+
| | | | | | +-- Platynaias_subviridis **
| | | | | +--+
+--+ | | | +-- Prolasmidonta_heterodon **
| | +--+
| | | +-- Anodontoides_ferussacianus **
| | | +-----------+
| | | | +-- Anodontoides_denigratus **
| | +--+
| | | +----------- Strophitus_undulatus **
| | +--+
| | | +-------- Strophitus_pascagoulaensis[2] **
| | +--+
+--+ | +----- Strophitus_pascagoulaensis[1] **
| +--+
| | +-- Strophitus_radiatus **
| +--+
| +-- Strophitus_williamsi **
|
| +-- Arcidens_confragosus **
| +--+
| | +-- Arcidens_wheeleri **
| +--------+
| | | +-- Pseudodontoideus_connasaugaensis **
| | +--+
| | +-- Pseudodontoideus_subvexus **
+--------+
| +-- Alasmidonta_marginata **
| +--------+
| | +-- Alasmidonta_raveneliana **
| |
+--+ +-- Alasmidonta_varicosa **
| +-----+
| | +-- Alasmidonta_uwharriensis **
+--+
| +----- Alasmidonta_undulata **
+--+
| +-- Alasmidonta_triangulata **
+--+
+-- Alasmidonta_arcula **
Whelan et al., 2023 [Publ 4287], fig. 4
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 29 |
29 |
25 |
1 |
0 |
0 |
0 |
97.0 |
+-- Anodonta_anatina **
+-----------------------+
| +-- Anodonta_cygnea **
|
| +----------------------- Pressodonta_viridis **
| |
| | +----- Utterbackiana_couperiana **
--+ | |
| | +-----------+ +-- Utterbackiana_heardi **
| | | | |
| | | +--+-- Utterbackiana_hartfieldorum **
| | | |
| | +--+ +-- Utterbackiana_suborbiculata **
| | | |
+--+ | | +-------------- Alasminota_etowaensis **
| | | |
| | +--+ +-- Arcidens_confragosus **
| | | +--------+
| | | | +-- Arcidens_wheeleri **
| | +--+
| | | +-------- Pseudodontoideus_connasaugaensis[3] **
| | +--+
| | | +----- Pseudodontoideus_connasaugaensis[2] **
| | +--+
+--+ | +-- Pseudodontoideus_subvexus **
| +--+
| +-- Pseudodontoideus_connasaugaensis[1] **
|
| +----- Alasmidonta_raveneliana **
| |
| | +-- Alasmidonta_uwharriensis **
| +-----------+--+
| | | +-- Alasmidonta_varicosa **
| | |
| | | +-- Alasmidonta_arcula **
| | +--+
+--+ +-- Alasmidonta_triangulata **
|
| +-------------- Platynaias_subviridis **
| |
| | +-------- Strophitus_pascagoulaensis **
| | +--+
+--+ | | +----- Strophitus_undulatus **
| | +--+
| | | +-- Strophitus_radiatus **
| | +--+
+--+ +-- Strophitus_williamsi **
|
| +-- Pyganodon_grandis **
| +-----+
| | +-- Pyganodon_cataracta **
+--+
| +----- Utterbackia_peninsularis **
+--+
| +-- Anodontoides_ferussacianus **
+--+
+-- Anodontoides_denigratus **
Campbell et al., 2008 [Publ 2675], fig. 2
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 28 |
28 |
18 |
1 |
0 |
0 |
0 |
101.0 |
+----------------------- Cyrtonaias_tampicoensis **
|
+----------------------- Plectomerus_dombeyanus **
|
| +-- Pleurobema_cordatum **
| |
| +-----------+-- Pleurobema_rubrum **
| | |
| | +-- Pleurobema_sintoxia **
| |
--+ +-----+ +-- Fusconaia_cerina **
| | | +--------+
| | | | +-- Fusconaia_flava **
| | | |
| | +--+ +-------- Pleuronaia_gibber **
| | | |
| | | | +----- Eurynia_dilatata **
| | +--+ |
| | | +----- Hemistena_lata **
| | | |
| | +--+----- Plethobasus_cyphyus **
+--+ |
| +----- Parvaspina_collina **
| |
| | +-- Elliptio_arca **
| +--+
| +-- Elliptio_crassidens **
|
| +----------------- Pleurobema_stabile **
| |
| | +-------------- Pleurobema_pyriforme **
| | |
+--+ | +-- Pleurobema_georgianum[2] **
| +-----------+
| | +-- Pleurobema_athearni **
| |
| | +----- Pleurobema_decisum **
+--+--------+
| | +-- Pleurobema_hanleyianum **
| +--+
| +-- Pleurobema_georgianum[1] **
|
| +-- Pleurobema_clava **
| +--------+
| | +-- Pleurobema_oviforme **
+--+
| +-------- Pleurobema_rubellum **
| |
+--+-------- Pleurobema_strodeanum **
|
| +----- Pleurobema_beadleianum **
+--+
| +-- Pleurobema_taitianum **
+--+
+-- Pleurobema_perovatum **
Perkins et al., 2017 [Publ 3777], fig. 3
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 28 |
28 |
11 |
1 |
0 |
0 |
0 |
99.0 |
+----------------------- Plectomerus_dombeyanus **
|
+----------------------- Uniomerus_declivis **
|
| +-- Parvaspina_steinstansana **
| |
| +-- Parvaspina_collina[1] **
--+ |
| +-----------------+-- Parvaspina_collina[2] **
| | |
| | +-- Parvaspina_collina[3] **
| | |
| | +-- Parvaspina_collina[4] **
| |
+--+ +----------------- Elliptio_arca **
| |
| +----------------- Hemistena_lata **
| |
| | +-- Elliptio_arctata **
| +--------------+
| | +-- Elliptio_crassidens **
+--+
| +-------------- Eurynia_dilatata **
| |
| | +----------- Pleurobema_oviforme **
| | |
| | +----------- Pleurobema_clava **
| | |
+--+ +----------- Pleurobema_pyriforme **
| |
| +----------- Pleurobema_georgianum **
| |
| +----------- Pleurobema_beadleianum **
| |
| +----------- Pleurobema_hanleyianum **
+--+
+----------- Pleurobema_decisum **
|
| +----- Pleurobema_rubellum **
+-----+
| | +-- Pleurobema_sintoxia **
| +--+
| +-- Pleurobema_rubrum **
|
| +-------- Elliptio_spinosa **
| |
+--+ +----- Fusconaia_escambia **
| |
| +----- Fusconaia_flava **
+--+
+----- Fusconaia_cerina **
|
| +-- Pleuronaia_gibber **
+--+
+-- Pleuronaia_dolabelloides **
Lopes-Lima et al., 2019 [Publ 3901], fig. 2
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 28 |
28 |
25 |
2 |
0 |
0 |
0 |
92.0 |
+-- Margaritifera_falcata **
+--------------------------------------+
| +-- Cumberlandia_monodonta **
|
| +-- Anodonta_nuttalliana **
--+ +-----------------------------------+
| | +-- Pyganodon_grandis **
| |
| | +-- Amblema_elliottii **
+--+ +--------------------------------+
| | +-- Amblema_plicata **
| |
| | +-- Lampsilis_ornata **
+--+ +-----------------------------+
| | +-- Potamilus_leptodon **
| |
| | +-- Pleurobema_riddellii **
+--+ +--------------------------+
| | +-- Elliptio_crassidens **
| |
| | +----- Megalonaias_nervosa **
+--+ |
| +--------------------+ +-- Uniomerus_carolinianus **
| | | |
| | +--+-- Uniomerus_declivis **
| | |
+--+ +-- Uniomerus_tetralasmus **
|
| +----------------------- Quadrula_quadrula **
| |
| | +-- Quadrula_verrucosa **
+--+ +-----------------+
| | +-- Quadrula_nobilis **
| |
+--+ +-- Theliderma_metanevra **
| +--------------+
| | +-- Theliderma_johnsoni **
| |
+--+ +----- Pustulosa_kieneriana **
| +--------+
| | | +-- Pustulosa_infucata **
| | +--+
+--+ +-- Pustulosa_kleiniana **
|
| +----------- Cyclonaias_tuberculata **
| |
+--+ +-- Pustulosa_pustulosa **
| +-----+
| | +-- Pustulosa_succissa **
+--+
| +----- Pustulosa_nodulata **
+--+
| +-- Pustulosa_necki **
+--+
+-- Pustulosa_petrina **
Smith et al., 2020 [Publ 4129], fig. 1
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 28 |
28 |
26 |
1 |
0 |
0 |
0 |
101.0 |
+-------------------------------------- Quadrula_quadrula **
|
| +-- Pleurobema_strodeanum **
--+ +--------------------------------+
| | +-- Elliptoideus_sloatianus **
| |
| | +----- Reginaia_ebenus **
+--+ +--------------------------+
| | | +-- Amblema_plicata **
| | +--+
| | +-- Plectomerus_dombeyanus **
+--+
| +-- Popenaias_popeii **
| +--------------------------+
| | +-- Psoronaias_semigranosa **
| |
+--+ +-------------------------- Obliquaria_reflexa **
| |
| | +----------------------- Cyrtonaias_tampicoensis **
+--+ |
| | +-- Pachynaias_spheniopsis **
| | +-----------------+
+--+ | +-- Lampsilis_cardium **
| |
| | +----------- Ellipsaria_lineolata **
| | +-----+
+--+ | | +-------- Truncilla_truncata **
| | +--+
| | | +----- Truncilla_donaciformis **
| | +--+
| | | +-- Truncilla_cognata **
| | +--+
+--+ +-- Truncilla_macrodon **
|
| +-------------- Atlanticoncha_ochracea **
| |
| | +-- Potamilus_fragilis **
| | +-----+
| | | +-- Potamilus_leptodon **
+--+ +--+
| | | +----- Potamilus_metnecktayi **
| | +--+
| | | +-- Potamilus_alatus **
| | +--+
+--+ +-- Potamilus_purpuratus **
|
| +-- Potamilus_capax **
| +-----+
| | +-- Potamilus_inflatus **
+--+
| +----- Potamilus_ohiensis **
+--+
| +-- Potamilus_streckersoni **
+--+
+-- Potamilus_amphichaenus **
Smith et al., 2020 [Publ 4129], fig. 2-3
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 28 |
28 |
26 |
1 |
0 |
0 |
0 |
101.0 |
+----------------------------------------- Quadrula_quadrula **
|
| +-- Pleurobema_strodeanum **
--+ +-----------------------------------+
| | +-- Elliptoideus_sloatianus **
| |
| | +----- Reginaia_ebenus **
+--+ +-----------------------------+
| | | +-- Amblema_plicata **
| | +--+
| | +-- Plectomerus_dombeyanus **
+--+
| +-- Popenaias_popeii **
| +-----------------------------+
| | +-- Psoronaias_semigranosa **
| |
+--+ +----------------------------- Obliquaria_reflexa **
| |
| | +-------------------------- Cyrtonaias_tampicoensis **
+--+ |
| | +-- Pachynaias_spheniopsis **
| | +--------------------+
+--+ | +-- Lampsilis_cardium **
| |
| | +----------- Ellipsaria_lineolata **
| | +--------+
+--+ | | +-------- Truncilla_truncata **
| | +--+
| | | +----- Truncilla_donaciformis **
| | +--+
| | | +-- Truncilla_cognata **
+--+ +--+
| +-- Truncilla_macrodon **
|
| +----------------- Atlanticoncha_ochracea **
| |
| | +-- Potamilus_capax **
+--+ +-----------+
| | +-- Potamilus_inflatus **
| |
| | +----- Potamilus_ohiensis **
+--+ +-----+
| | | +-- Potamilus_streckersoni **
| | +--+
| | +-- Potamilus_amphichaenus **
+--+
| +-- Potamilus_fragilis **
| +-----+
| | +-- Potamilus_leptodon **
+--+
| +----- Potamilus_metnecktayi **
+--+
| +-- Potamilus_alatus **
+--+
+-- Potamilus_purpuratus **
Guerra et al., 2019 [Publ 4168], fig. 1
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 28 |
28 |
26 |
2 |
13 |
12 |
2 |
84.0 |
+----------------- Mutela_dubia
+--------------+
| | +-------------- Chambardia_rubens
| +--+
| | +----------- Anodontites_trapesialis
| +--+
| | +-------- Monocondylaea_parchappii
| +--+
| | +----- Lamproscapha_ensiformis
| +--+
| | +-- Tamsiella_schroeteriana
| +--+
| +-- Fossula_fossiculifera
--+
| +-- Echyridella_menziesii
| +-----+
| | +-- Westralunio_carteri
| +--------------------+
| | | +----- Diplodon_suavidicus
| | +--+
| | | +-- Castalia_ambigua
| | +--+
| | +-- Prisodon_obliquus
| |
| | +-------------- Gibbosula_crassa **
| | |
+--+ +-----------+ +----------- Cumberlandia_monodonta **
| | | |
| | +--+ +-- Pseudunio_marocanus **
| | | +-----+
| | | | +-- Pseudunio_auricularius **
| | +--+
| | | +----- Margaritifera_falcata **
| | +--+
| | | +-- Margaritifera_dahurica **
| | +--+
+--+ +-- Margaritifera_margaritifera **
|
| +-------- Quadrula_quadrula **
| +--------------+
| | | +----- Toxolasma_parvum **
| | +--+
| | | +-- Venustaconcha_ellipsiformis **
| | +--+
| | +-- Potamilus_alatus **
| |
+--+ +-------- Sinosolenaia_carinata **
| +-----------+
| | | +----- Pronodularia_japanensis **
| | +--+
| | | +-- Lamprotula_leaii **
| | +--+
| | +-- Potomida_littoralis **
| |
+--+ +-------------- Lanceolaria_lanceolata **
| +--+
| | | +----------- Aculamprotula_tortuosa **
| | +--+
| | | +-------- Schistodesmus_lampreyanus **
| | +--+
| | | +----- Unio_tumidus **
+--+ +--+
| | +-- Unio_crassus **
| +--+
| +-- Unio_delphinus **
|
| +-------------- Sinanodonta_woodiana **
| |
+--+ +----------- Anodonta_anatina **
| |
+--+ +-- Platynaias_compressa **
| +-----+
| | +-- Platynaias_subviridis **
+--+
| +----- Pyganodon_grandis **
+--+
| +-- Utterbackia_peninsularis **
+--+
+-- Utterbackia_imbecillis **
Guerra et al., 2019 [Publ 4168], fig. 2
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 28 |
28 |
26 |
2 |
13 |
12 |
2 |
84.0 |
+----------------- Chambardia_rubens
+--------------------+
| | +-------------- Mutela_dubia
| +--+
| | +----------- Anodontites_trapesialis
| +--+
| | +-------- Monocondylaea_parchappii
| +--+
| | +----- Lamproscapha_ensiformis
| +--+
| | +-- Tamsiella_schroeteriana
| +--+
| +-- Fossula_fossiculifera
--+
| +----------- Westralunio_carteri
| +-----------------------+
| | | +-------- Echyridella_menziesii
| | +--+
| | | +----- Diplodon_suavidicus
| | +--+
| | | +-- Castalia_ambigua
| | +--+
| | +-- Prisodon_obliquus
| |
| | +-------------- Gibbosula_crassa **
+--+ |
| +-----------------+ +----------- Cumberlandia_monodonta **
| | | |
| | +--+ +-- Pseudunio_marocanus **
| | | +-----+
| | | | +-- Pseudunio_auricularius **
| | +--+
| | | +----- Margaritifera_falcata **
| | +--+
| | | +-- Margaritifera_dahurica **
+--+ +--+
| +-- Margaritifera_margaritifera **
|
| +-------- Quadrula_quadrula **
| +--------------------+
| | | +----- Toxolasma_parvum **
| | +--+
| | | +-- Venustaconcha_ellipsiformis **
| | +--+
| | +-- Potamilus_alatus **
+--+
| +-------- Sinosolenaia_carinata **
| +-----------------+
| | | +----- Pronodularia_japanensis **
| | +--+
| | | +-- Lamprotula_leaii **
| | +--+
+--+ +-- Potomida_littoralis **
|
| +----------------------- Aculamprotula_tortuosa **
| |
| | +-------- Schistodesmus_lampreyanus **
+--+ +-----------+
| | | +----- Unio_tumidus **
| | +--+
| | | +-- Unio_crassus **
+--+ +--+
| +-- Unio_delphinus **
|
| +----------------- Lanceolaria_lanceolata **
| |
+--+ +-------------- Sinanodonta_woodiana **
| |
+--+ +----------- Anodonta_anatina **
| |
+--+ +-- Platynaias_compressa **
| +-----+
| | +-- Platynaias_subviridis **
+--+
| +----- Pyganodon_grandis **
+--+
| +-- Utterbackia_peninsularis **
+--+
+-- Utterbackia_imbecillis **
Konopleva et al., 2021 [Publ 4213], fig. 1
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 28 |
28 |
22 |
1 |
0 |
0 |
0 |
93.0 |
+-------------------------- Lamprotula_leaii **
|
+-------------------------- Leguminaia_wheatleyi **
|
+-------------------------- Potomida_littoralis **
|
+-------------------------- Gonidea_angulata **
|
--+ +-------- Indopseudodon_kayinensis **
| +--------------+
| | | +----- Indopseudodon_indawgyiensis **
| | +--+
| | | +-- Indopseudodon_salwenianus **
| | +--+
| | +-- Indopseudodon_bogani **
| |
| | +----- Pilsbryoconcha_exilis[2] **
+--+ +--------------+
| | | +-- Pilsbryoconcha_exilis[1] **
| | +--+
| | +-- Namkongnaia_lemeslei **
| |
| | +-------------- Nyeinchanconcha_nyeinchani **
| | |
| | | +-- Thaiconcha_munelliptica **
+--+ +--+ |
| | | +--------+-- Thaiconcha_thaiensis **
| | | | |
| | +--+ +-- Thaiconcha_callifera **
| | |
| | | +-------- Bineurus_loeiensis **
| | +--+
| | | +----- Bineurus_mouhotii **
| | +--+
+--+ | +-- Bineurus_anodontinum **
| +--+
| +-- Bineurus_exilis **
|
| +-------- Pseudodon_lenyanensis **
| +--+
| | | +----- Pseudodon_vagulus **
| | +--+
| | | +-- Pseudodon_vondembuschianus **
| | +--+
+-----+ +-- Pseudodon_mekongi **
|
| +-- Sundadontina_taskaevi **
| +-----+
| | +-- Sundadontina_brandti **
+--+
| +----- Sundadontina_tumida **
+--+
| +-- Sundadontina_cumingii **
+--+
+-- Sundadontina_tanintharyiensis **
Graf & Cummings, 2006 [Publ 1968], fig. 2
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 27 |
27 |
21 |
2 |
20 |
19 |
2 |
80.0 |
+----- Velesunio_ambiguus
+--------+
| | +-- Velesunio_angasi
| +--+
+--+ +-- Lortiella_rugata
| |
| | +----------- Echyridella_menziesii
| | |
| +--+ +-- Hyridella_depressa
| | +-----+
| | | +-- Hyridella_australis
| +--+
| | +----- Castalia
| +--+
+--------------------+ | +-- Diplodon_chilensis
| | +--+
| | +-- Diplodon_gratus
| |
| | +----- Etheria_elliptica
| | +-----+
| | | | +-- Mulleria_rivoli
| | | +--+
| | | +-- Pseudomulleria_dalyi
| | |
| +-----+ +----- Monocondylaea_minuana
| | +--+
| | | | +-- Anodontites_trigona
| | | +--+
| | | +-- Anodontites_guanarensis
--+ +--+
| | +-- Chambardia_wahlbergi
| | +--+
| | | +-- Aspatharia_rugifera
| +--+
| | +-- Mutela_dubia
| +--+
| +-- Mutela_rostrata
|
| +-- Margaritifera_margaritifera **
| +--------------------------------+
| | +-- Cumberlandia_monodonta **
| |
| | +-- Pilsbryoconcha_exilis **
| | +-----------------------------+
+--+ | +-- Pseudodon_vondembuschianus **
| |
| | +----------------------------- Coelatura_aegyptiaca **
| | |
| | +----------------------------- Lens_contradens **
+--+ |
| | +-- Unio_pictorum **
| | +--------+
| | | +-- Unio_caffer **
| +-----------------+
| | | +-------- Lasmigona_costata **
+--+ +--+
| | +----- Alasmidonta_marginata **
| +--+
| | +-- Pyganodon_grandis **
| +--+
| +-- Strophitus_undulatus **
|
| +-------------------------- Gonidea_angulata **
| |
| | +-- Quadrula_verrucosa **
+--+ +--------------------+
| | +-- Quadrula_quadrula **
| |
+--+ +-------------------- Amblema_plicata **
| |
| | +----- Eurynia_dilatata **
+--+ +-----------+
| | | +-- Fusconaia_flava **
| | +--+
| | +-- Pleurobema_sintoxia **
+--+
| +-------------- Obliquaria_reflexa **
| |
| +-------------- Truncilla_truncata **
+--+
+-------------- Ptychobranchus_fasciolaris **
|
| +----------- Ligumia_recta **
+--+
| +-------- Cambarunio_iris **
+--+
| +----- Epioblasma_triquetra **
+--+
| +-- Lampsilis_cardium **
+--+
+-- Ortmanniana_ligamentina **
Graf & Ó Foighil, 2000 [Publ 2648], fig. 2A
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 27 |
27 |
21 |
2 |
3 |
2 |
2 |
80.0 |
+-------------------------------- Mutela_rostrata
|
| +----------------------------- Hyridella_depressa
--+ |
| | +-- Margaritifera_margaritifera **
| | +-----------------------+
+--+ | +-- Cumberlandia_monodonta **
| |
| | +-- Unio_pictorum **
| | +--------------------+
+--+ | +-- Unio_caffer **
| |
| | +----------- Anodonta_cygnea **
| | |
| | +--------+ +-- Alasmidonta_marginata **
+--+ | | +-----+
| | | | +-- Lasmigona_costata **
| | +--+
| | | +----- Strophitus_undulatus **
| | +--+
| | | +-- Pyganodon_grandis **
| | +--+
+--+ +-- Pyganodon_fragilis **
|
| +-------- Quadrula_quadrula **
| +--------+
| | | +----- Eurynia_dilatata **
| | +--+
| | | +-- Pleurobema_sintoxia **
| | +--+
+--+ +-- Fusconaia_flava **
|
| +-------------- Amblema_plicata **
| |
| | +----------- Ligumia_recta **
| | |
+--+ +----------- Truncilla_truncata **
| |
| | +-- Ptychobranchus_fasciolaris **
| +--------+
+--+ +-- Sagittunio_nasutus **
|
| +----- Lampsilis_siliquoidea **
| +--+
| | | +-- Epioblasma_triquetra **
| | +--+
| | +-- Epioblasma_brevidens **
+--+
| +----- Ortmanniana_ligamentina **
| |
| +----- Cambarunio_iris **
+--+
+----- Leaunio_vanuxemensis **
|
| +-- Lampsilis_cardium **
+--+
+-- Lampsilis_fasciola **
Graf & Ó Foighil, 2000 [Publ 2648], fig. 2B
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 27 |
27 |
25 |
2 |
3 |
2 |
2 |
79.0 |
+----------------------------------------- Mutela_rostrata
|
| +-------------------------------------- Hyridella_depressa
--+ |
| | +-- Margaritifera_margaritifera **
| | +--------------------------------+
+--+ | +-- Cumberlandia_monodonta **
| |
| | +-- Unio_pictorum **
| | +-----------------------------+
+--+ | +-- Unio_caffer **
| |
| | +----------- Anodonta_cygnea **
| | |
| | +-----------------+ +-- Alasmidonta_marginata **
+--+ | | +-----+
| | | | +-- Lasmigona_costata **
| | +--+
| | | +----- Strophitus_undulatus **
| | +--+
| | | +-- Pyganodon_grandis **
+--+ +--+
| +-- Pyganodon_fragilis **
|
| +-------- Quadrula_quadrula **
| +-----------------+
| | | +----- Eurynia_dilatata **
| | +--+
| | | +-- Pleurobema_sintoxia **
+--+ +--+
| +-- Fusconaia_flava **
|
| +----------------------- Amblema_plicata **
| |
+--+ +-------------------- Truncilla_truncata **
| |
+--+ +----------------- Ligumia_recta **
| |
| | +-- Ptychobranchus_fasciolaris **
+--+ +-----------+
| | +-- Sagittunio_nasutus **
| |
+--+ +----------- Lampsilis_siliquoidea **
| |
| | +----- Ortmanniana_ligamentina **
+--+ +--+
| | | +-- Lampsilis_cardium **
| | +--+
| | +-- Lampsilis_fasciola **
+--+
| +-- Epioblasma_brevidens **
| +--+
| | +-- Epioblasma_triquetra **
+--+
| +-- Cambarunio_iris **
+--+
+-- Leaunio_vanuxemensis **
Lydeard et al., 1996 [Publ 2656], fig. 3
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 27 |
27 |
23 |
2 |
0 |
0 |
0 |
91.0 |
+----- Cumberlandia_monodonta **
+--------------------------+
| | +-- Margaritifera_margaritifera **
| +--+
| +-- Margaritifera_falcata **
|
| +----- Utterbackia_imbecillis **
| +--+
| | | +-- Utterbackiana_couperiana **
--+ | +--+
| +--------------------+ +-- Utterbackiana **
| | |
| | | +----- Strophitus_radiatus **
| | +--+
| | | +-- Lasmigona_costata **
| | +--+
| | +-- Alasmidonta_triangulata **
+--+
| +----- Megalonaias_nervosa **
| +--------------------+
| | | +-- Quadrula_quadrula **
| | +--+
| | +-- Theliderma_metanevra **
| |
+--+ +----- Elliptoideus_sloatianus **
| |
| +-----------------+ +-- Eurynia_dilatata **
| | | |
| | +--+-- Pleurobema_pyriforme **
| | |
+--+ +-- Uniomerus_carolinianus **
|
| +-------------------- Amblema_plicata **
| |
+--+ +-- Plectomerus_dombeyanus **
| +--------------+
| | +-- Obliquaria_reflexa **
+--+
| +-------------- Potamilus_fragilis **
| |
+--+ +----------- Ptychobranchus_subtentus **
| |
+--+ +-------- Medionidus_conradicus **
| |
+--+ +----- Potamilus_purpuratus **
| |
| | +-- Ellipsaria_lineolata **
+--+--+
| +-- Villosa_delumbis **
|
| +-- Lampsilis_teres **
+--+
+-- Lampsilis_siliquoidea **
Lydeard et al., 1996 [Publ 2656], fig. 4
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 27 |
27 |
25 |
2 |
1 |
0 |
0 |
89.5 |
+----- Cumberlandia_monodonta **
+-----------------------------------+
| | +-- Margaritifera_margaritifera **
| +--+
| +-- Margaritifera_falcata **
|
| +----- Utterbackia_imbecillis **
| +--+
| | | +-- Utterbackiana_couperiana **
--+ | +--+
| +-----------------------------+ +-- Anodonta **
| | |
| | | +----- Strophitus_radiatus **
| | +--+
| | | +-- Lasmigona_costata **
| | +--+
| | +-- Alasmidonta_triangulata **
+--+
| +----- Megalonaias_nervosa **
| +-----------------------------+
| | | +-- Quadrula_quadrula **
| | +--+
| | +-- Theliderma_metanevra **
| |
+--+ +-------- Elliptoideus_sloatianus **
| +-----------------------+
| | | +----- Uniomerus_carolinianus **
| | +--+
| | | +-- Eurynia_dilatata **
+--+ +--+
| +-- Pleurobema_pyriforme **
|
| +----------------------------- Amblema_plicata **
+--+
| +-------------------------- Plectomerus_dombeyanus **
+--+
| +----------------------- Obliquaria_reflexa **
+--+
| +-------------------- Medionidus_conradicus **
+--+
| +----------------- Ptychobranchus_subtentus **
+--+
| +-------------- Potamilus_fragilis **
+--+
| +----------- Potamilus_purpuratus **
+--+
| +-------- Ellipsaria_lineolata **
+--+
| +----- Villosa_delumbis **
+--+
| +-- Lampsilis_teres **
+--+
+-- Lampsilis_siliquoidea **
Hoeh et al., 2009 [Publ 2716], fig. 3
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 27 |
28 |
26 |
2 |
20 |
19 |
2 |
76.0 |
+----- Velesunio_ambiguus
+--------+
| | +-- Velesunio_angasi
| +--+
+-----------------------------+ +-- Lortiella_rugata
| |
| | +----------- Echyridella_menziesii
| | |
| +--+ +-- Hyridella_depressa
| | +-----+
| | | +-- Hyridella_australis
| +--+
| | +----- Castalia
| +--+
| | +-- Diplodon_chilensis
| +--+
| +-- Diplodon_gratus
--+
| +----- Chambardia_wahlbergi
| |
| +--------+----- Aspatharia_rugifera
| | |
| | | +-- Mutela_rostrata
| +--------------------------+ +--+
| | | +-- Mutela_dubia
| | |
| | | +----------- Etheria_elliptica
| | +--+
| | | +-------- Mulleria_rivoli
| | +--+
| | | +----- Monocondylaea_minuana
+--+ +--+
| | +-- Anodontites_trigona
| +--+
| +-- Anodontites_guanarensis
|
| +-- Margaritifera_margaritifera **
| +-----------------------------------+
| | +-- Cumberlandia_monodonta **
| |
+--+ +----------------------------------- Coelatura_aegyptiaca **
| |
| | +-------------------------------- Pseudomulleria_dalyi
| | |
+--+ | +-- Unio_pictorum **
| | +--------+
| | | +-- Unio_caffer **
| | +-----------------+
+--+ | | +-------- Lasmigona_costata **
| | +--+
| | | +----- Alasmidonta_marginata **
| | +--+
| | | +-- Strophitus_undulatus **
| | +--+
+--+ +-- Pyganodon_grandis **
|
| +-------------------------- Lens_contradens **
| |
| | +----- Gonidea_angulata **
| | +-----------------+
| | | | +-- Pilsbryoconcha_exilis **
+--+ | +--+
| | +-- Pseudodon_vondembuschianus **
| |
| | +----------- Amblema_plicata **
| | |
+--+ | +----- Obliquaria_reflexa **
| +--------+ +--+
| | | | | +-- Quadrula_verrucosa **
| | | | +--+
| | +--+ +-- Quadrula_quadrula **
| | |
| | | +----- Eurynia_dilatata **
+--+ +--+
| | +-- Pleurobema_sintoxia **
| +--+
| +-- Fusconaia_flava **
|
| +----------------- Ptychobranchus_fasciolaris **
+--+
| +-------------- Truncilla_truncata **
+--+
| +----------- Ortmanniana_ligamentina **
+--+
| +-------- Lampsilis_cardium **
+--+
| +----- Cambarunio_iris **
+--+
| +-- Ligumia_recta **
+--+
+-- Epioblasma_triquetra **
Whelan et al., 2011 [Publ 3380], fig. 1 ML
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 27 |
32 |
30 |
2 |
10 |
9 |
2 |
77.0 |
+-------- Velesunio_ambiguus
+--------------------------+
| | +----- Hyridella_australis
| +--+
| | +-- Castalia_ambigua
| +--+
| +-- Triplodon_corrugatus
|
| +-- Margaritifera_margaritifera **
--+ +-----------------------------+
| | +-- Cumberlandia_monodonta **
| |
| | +-- Aspatharia_pfeifferiana
| | +-----+
| | | +-- Chambardia_wahlbergi
| | +--------------------+
| | | | +----- Mutela_rostrata
+--+ | +--+
| | | +-- Mycetopoda_siliquosa
| | +--+
| | +-- Tamsiella_schroeteriana
| |
| | +-- Lamellidens_savadiensis **
| | +-----------------+
| | | +-- Trapezidens_dolichorhynchus **
+--+ +-----+
| | | +----------------- Leoparreysia_tavoyensis **
| | | |
| | +--+ +-- Radiatula_chaudhurii **
| | | +-----------+
| | | | +-- Indochinella_pugio **
| | +--+
| | | +----------- Nitia_teretiuscula **
| | +--+
| | | +-------- Coelatura_aegyptiaca[2] **
| | +--+
+--+ | +----- Coelatura_aegyptiaca[1] **
| +--+
| | +-- Coelatura_gabonensis **
| +--+
| +-- Prisodontopsis_aviculaeformis **
|
| +-- Pseudodon_vondembuschianus **
| +--------------------+
| | +-- Lens_contradens **
| |
| | +-- Pyganodon_grandis **
| | +--+
+--+ | +-- Lasmigona_costata **
| +--------------+
| | | +-- Unio_pictorum **
| | +--+
| | +-- Unio_caffer **
+--+
| +----------------- Gonidea_angulata **
| |
| | +----- Pronodularia_japanensis **
+--+ +--------+
| | | +-- Potomida_littoralis **
| | +--+
+--+ +-- Potomida_semirugata **
|
| +----------- Pleurobema_sintoxia **
+--+
| +-------- Quadrula_quadrula **
+--+
| +----- Amblema_plicata **
+--+
| +-- Truncilla_truncata **
+--+
+-- Lampsilis_cardium **
Whelan et al., 2011 [Publ 3380], fig. 1 MP
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 27 |
31 |
27 |
2 |
10 |
9 |
2 |
83.0 |
+-- Aspatharia_pfeifferiana
+-----+
| +-- Chambardia_wahlbergi
+-----------------------+
| | +----- Mutela_rostrata
| +--+
| | +-- Mycetopoda_siliquosa
| +--+
| +-- Tamsiella_schroeteriana
|
| +-- Margaritifera_margaritifera **
| +--------+
| | +-- Cumberlandia_monodonta **
--+ +-----------------+
| | | +-------- Velesunio_ambiguus
| | +--+
| | | +----- Hyridella_australis
| | +--+
| | | +-- Castalia_ambigua
| | +--+
| | +-- Triplodon_corrugatus
| |
| | +-- Lamellidens_savadiensis **
| | +--------------+
+--+ | +-- Trapezidens_dolichorhynchus **
| +--------+
| | | +-------------- Leoparreysia_tavoyensis **
| | | |
| | +--+ +-- Radiatula_chaudhurii **
| | | +--------+
| | | | +-- Indochinella_pugio **
| | +--+
| | | +-------- Nitia_teretiuscula **
| | | |
| | +--+ +----- Coelatura_aegyptiaca[1] **
+--+ | |
| +--+----- Coelatura_aegyptiaca[2] **
| |
| | +-- Coelatura_gabonensis **
| +--+
| +-- Prisodontopsis_aviculaeformis **
|
| +----------------------- Lens_contradens **
| |
| | +-------------------- Pseudodon_vondembuschianus **
+--+ |
| | +-- Pyganodon_grandis **
| | +--+
+--+ | +-- Lasmigona_costata **
| +-----------+
| | | +-- Unio_pictorum **
| | +--+
| | +-- Unio_caffer **
+--+
| +-------------- Gonidea_angulata **
| |
| | +----- Pronodularia_japanensis **
+--+ +-----+
| | | +-- Potomida_littoralis **
| | +--+
+--+ +-- Potomida_semirugata **
|
| +-------- Quadrula_quadrula **
| |
+--+-------- Pleurobema_sintoxia **
|
| +----- Amblema_plicata **
+--+
| +-- Lampsilis_cardium **
+--+
+-- Truncilla_truncata **
Graf et al., 2014 [Publ 3591], fig. 3
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 27 |
27 |
13 |
2 |
0 |
0 |
0 |
83.0 |
+-------------------------------- Margaritifera_margaritifera **
|
+-------------------------------- Unio_pictorum **
--+
| +----------------------------- Leoparreysia_tavoyensis **
| |
+--+ +-------------------------- Nitia_teretiuscula **
| |
+--+ +----------------------- Coelatura_aegyptiaca[2] **
| |
+--+ +-------------------- Coelatura_aegyptiaca[1] **
| |
| | +----------------- Coelatura_gabonensis **
+--+ |
| | +-------------- Coelatura_kunenensis **
| | |
| | | +-- Coelatura_choziensis[1] **
+--+ | |
| | +-- Coelatura_choziensis[2] **
| | |
| | +-- Coelatura_luapulaensis[1] **
| | |
+--+ +--------+-- Coelatura_luapulaensis[2] **
| | |
| | +-- Coelatura_luapulaensis[3] **
| | |
| | +-- Coelatura_luapulaensis[4] **
| | |
| | +-- Coelatura_luapulaensis[5] **
| |
+--+ +----- Prisodontopsis_mweruensis[2] **
| |
| +----- Prisodontopsis_mweruensis[3] **
| +--+
| | | +-- Prisodontopsis_mweruensis[4] **
| | | |
| | +--+-- Prisodontopsis_mweruensis[5] **
| | |
+--+ +-- Prisodontopsis_aviculaeformis[7] **
|
| +----- Prisodontopsis_aviculaeformis[1] **
| |
| | +-- Prisodontopsis_aviculaeformis[2] **
+--+ |
| +-- Prisodontopsis_aviculaeformis[3] **
| |
| +-- Prisodontopsis_aviculaeformis[4] **
+--+
+-- Prisodontopsis_aviculaeformis[5] **
|
+-- Prisodontopsis_aviculaeformis[6] **
|
+-- Prisodontopsis_mweruensis[1] **
Neemuchwala et al., 2023 [Publ 4292], fig. 2
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 27 |
27 |
25 |
1 |
0 |
0 |
0 |
96.0 |
+-------------------------------- Gonidea_angulata **
|
| +-------- Pleurobema_strodeanum **
| +--------------------+
--+ | | +----- Amblema_plicata **
| | +--+
| | | +-- Lampsilis_cardium **
| | +--+
+--+ +-- Popenaias_popeii **
|
| +-------------------------- Megalonaias_nervosa **
| |
| | +-------- Uniomerus_carolinianus **
| | +--------------+
+--+ | | +----- Uniomerus_columbensis **
| | +--+
| | | +-- Uniomerus_tetralasmus **
| | +--+
| | +-- Uniomerus_declivis **
+--+
| +-------- Theliderma_cylindrica **
| +-----------+
| | | +----- Theliderma_metanevra **
| | +--+
| | | +-- Theliderma_sparsa **
| | +--+
+--+ +-- Theliderma_johnsoni **
|
| +----------------- Cyclonaias_tuberculata **
| |
| | +-------- Quadrula_nobilis **
| | +-----+
+--+ | | +----- Quadrula_verrucosa **
| | +--+
| | | +-- Quadrula_fragosa **
| | +--+
| | +-- Quadrula_quadrula **
+--+
| +----- Pustulosa_nodulata **
| +-----+
| | | +-- Pustulosa_necki **
| | +--+
| | +-- Pustulosa_petrina **
+--+
| +-- Pustulosa_kieneriana **
| +-----+
| | +-- Pustulosa_pustulosa **
+--+
| +----- Pustulosa_succissa **
+--+
| +-- Pustulosa_infucata **
+--+
+-- Pustulosa_kleiniana **
Hoeh et al., 2009 [Publ 2716], fig. 4
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 26 |
27 |
24 |
2 |
20 |
19 |
2 |
83.0 |
+----- Velesunio_ambiguus
+--------+
| | +-- Velesunio_angasi
| +--+
+--------------------------------+ +-- Lortiella_rugata
| |
| | +----------- Echyridella_menziesii
| | |
| +--+ +-- Hyridella_depressa
| | +-----+
| | | +-- Hyridella_australis
| +--+
| | +----- Castalia
| +--+
| | +-- Diplodon_chilensis
| +--+
| +-- Diplodon_gratus
--+
| +-- Etheria_elliptica
| +--------+
| | +-- Mulleria_rivoli
| |
| | +----- Monocondylaea_minuana
| +--------------------------------+ +--+
| | | | | +-- Anodontites_trigona
| | | | +--+
| | | | +-- Anodontites_guanarensis
| | +--+
| | | +-- Mutela_rostrata
| | | +--+
+--+ | | +-- Mutela_dubia
| +--+
| | +-- Chambardia_wahlbergi
| +--+
| +-- Aspatharia_rugifera
|
| +----------------------------------------- Coelatura_aegyptiaca **
| |
+--+ +-------------------------------------- Pseudomulleria_dalyi
| |
| | +-- Margaritifera_margaritifera **
+--+ +--------------------------------+
| | +-- Cumberlandia_monodonta **
| |
| | +-- Pilsbryoconcha_exilis **
+--+ +-----------------------------+
| | +-- Pseudodon_vondembuschianus **
| |
| | +----------------------------- Lens_contradens **
+--+ |
| | +-- Unio_pictorum **
| | +--------+
| | | +-- Unio_caffer **
| | +--------------+
+--+ | | +-------- Lasmigona_costata **
| | +--+
| | | +----- Alasmidonta_marginata **
| | +--+
| | | +-- Strophitus_undulatus **
| | +--+
+--+ +-- Pyganodon_grandis **
|
| +----------------------- Gonidea_angulata **
| |
| | +-- Eurynia_dilatata **
| | |
| | +-----+-- Pleurobema_sintoxia **
+--+ | |
| +-----------+ +-- Fusconaia_flava **
| | |
| | | +----- Obliquaria_reflexa **
| | +--+
+--+ | +-- Quadrula_verrucosa **
| +--+
| +-- Quadrula_quadrula **
|
| +----------------- Ptychobranchus_fasciolaris **
+--+
| +-------------- Truncilla_truncata **
+--+
| +----------- Ortmanniana_ligamentina **
+--+
| +-------- Lampsilis_cardium **
+--+
| +----- Cambarunio_iris **
+--+
| +-- Ligumia_recta **
+--+
+-- Epioblasma_triquetra **
Hoeh et al., 2009 [Publ 2716], fig. 5
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 26 |
45 |
39 |
2 |
20 |
19 |
2 |
99.0 |
+----------------------------- Coelatura_aegyptiaca **
|
| +-- Margaritifera_margaritifera **
| +--------------------+
| | +-- Cumberlandia_monodonta **
| |
| | +----- Velesunio_ambiguus
| | +--------+
| | | | +-- Velesunio_angasi
| | | +--+
| +--+ +-----+ +-- Lortiella_rugata
| | | | |
| | | | | +----------- Echyridella_menziesii
| | | | | |
--+ | | | +--+ +-- Hyridella_depressa
| | | | | +-----+
| | | | | | +-- Hyridella_australis
| | | | +--+
| | +--+ | +----- Castalia
| | | +--+
| | | | +-- Diplodon_chilensis
| | | +--+
| | | +-- Diplodon_gratus
| | |
| | | +----------------- Pseudomulleria_dalyi
| | | |
| | +--+ +-------------- Mulleria_rivoli
| | | |
| | +--+ +----------- Etheria_elliptica
+--+ | |
| | | +----- Monocondylaea_minuana
| +--+ +--+
| | | | +-- Anodontites_trigona
| | | +--+
| | | +-- Anodontites_guanarensis
| +--+
| | +-- Mutela_rostrata
| | +--+
| | | +-- Mutela_dubia
| +--+
| | +-- Chambardia_wahlbergi
| +--+
| +-- Aspatharia_rugifera
|
| +-- Pilsbryoconcha_exilis **
| +--------------------+
| | +-- Pseudodon_vondembuschianus **
| |
| | +-------------------- Lens_contradens **
+--+ |
| | +-- Unio_pictorum **
| | +--------+
| | | +-- Unio_caffer **
+--+ +-----+
| | | +-------- Lasmigona_costata **
| | +--+
| | | +----- Alasmidonta_marginata **
| | +--+
| | | +-- Strophitus_undulatus **
+--+ +--+
| +-- Pyganodon_grandis **
|
| +-------------- Gonidea_angulata **
| |
| | +----- Obliquaria_reflexa **
| | +-----+
+--+ | | +-- Quadrula_verrucosa **
| | +--+
| | +-- Quadrula_quadrula **
| |
+--+ +-------- Amblema_plicata **
| |
| | +----- Eurynia_dilatata **
| +--+
| | | +-- Pleurobema_sintoxia **
+--+ +--+
| +-- Fusconaia_flava **
|
| +----- Truncilla_truncata **
| |
| +----- Ptychobranchus_fasciolaris **
+--+
| +-- Ortmanniana_ligamentina **
| |
| +-- Cambarunio_iris **
+--+
+-- Ligumia_recta **
|
+-- Epioblasma_triquetra **
Jadhav & Jamkhedkar, 2009 [Publ 3697], fig. 11
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 26 |
26 |
23 |
1 |
0 |
0 |
0 |
88.0 |
+-------------------------------- Lamellidens_corrianus **
|
| +-------------------- Nodularia_douglasiae **
| |
| +--------+ +----------------- Anodonta_cygnea **
| | | |
| | +--+ +-------------- Unio_pictorum[2] **
--+ | | |
| | +--+ +-- Unio_pictorum[1] **
| | | +--------+
| | | | +-- Unio_caffer **
| | +--+
| | | +-------- Anodonta **
| | | |
+--+ +--+ +----- Lasmigona_costata **
| | |
| +--+ +-- Strophitus_undulatus **
| | |
| +--+-- Alasmidonta_marginata **
| |
| +-- Pyganodon_grandis **
|
| +-------------------------- Pronodularia_japanensis **
| |
| | +-- Quadrula_quadrula **
+--+ +--------------------+
| | +-- Quadrula_verrucosa **
| |
+--+ +-------------------- Gonidea_angulata **
| |
| | +----- Eurynia_dilatata **
+--+ +-----------+
| | | +-- Cambarunio_iris **
| | +--+
| | +-- Ptychobranchus_fasciolaris **
+--+
| +-- Amblema_plicata **
| +-----------+
| | +-- Pleurobema_sintoxia **
+--+
| +----------- Truncilla_truncata **
| |
+--+ +-- Lampsilis_cardium **
| +-----+
| | +-- Epioblasma_triquetra **
+--+
| +----- Ligumia_recta **
+--+
| +-- Obliquaria_reflexa **
+--+
+-- Ortmanniana_ligamentina **
Lopes-Lima et al., 2019 [Publ 3901], fig. 4
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 26 |
26 |
24 |
2 |
0 |
0 |
0 |
93.0 |
+-- Margaritifera_falcata **
+-----------------------------------+
| +-- Cumberlandia_monodonta **
|
| +-- Anodonta_nuttalliana **
| +--------------------------------+
--+ | +-- Pyganodon_grandis **
| |
| | +-- Amblema_elliottii **
| | +-----+
| | | +-- Amblema_plicata **
+--+ |
| +-----------------------+ +-- Pleurobema_riddellii **
| | | +--+
| | | | +-- Pleurobema_oviforme **
| | +--+
| | | +-- Lampsilis_ornata **
+--+ +--+
| +-- Potamilus_leptodon **
|
| +----------------------------- Megalonaias_nervosa **
| |
+--+ +-------------------------- Uniomerus_tetralasmus **
| |
| | +-- Theliderma_metanevra **
+--+ +--------------------+
| | +-- Theliderma_johnsoni **
| |
+--+ +-------------------- Quadrula_quadrula **
| |
| | +-- Quadrula_verrucosa **
+--+ +--------------+
| | +-- Quadrula_nobilis **
| |
| | +----- Pustulosa_kieneriana **
+--+ +--------+
| | | +-- Pustulosa_infucata **
| | +--+
| | +-- Pustulosa_kleiniana **
+--+
| +-- Pustulosa_pustulosa **
| +--------+
| | +-- Pustulosa_succissa **
+--+
| +-------- Cyclonaias_tuberculata **
+--+
| +----- Pustulosa_necki **
+--+
| +-- Pustulosa_petrina **
+--+
+-- Pustulosa_nodulata **
Sano et al., 2017 [Publ 3799], fig. 1
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 25 |
25 |
23 |
2 |
2 |
0 |
0 |
86.0 |
+-------- Margaritifera_laevis[2] **
+--------------------+
| | +----- Margaritifera_laevis[1] **
| +--+
| | +-- Margaritifera_middendorffi **
| +--+
| +-- Margaritifera_laevis[3] **
|
| +-------- Obovalis_omiensis **
--+ +-----------------+
| | | +----- Inversidens_brandtii **
| | +--+
| | | +-- Pronodularia_japanensis **
| | +--+
| | +-- Sinohyriopsis_schlegelii **
| |
| | +-- Cristaria_plicata **
+--+ +-----------+
| | +-- Buldowskia_iwakawai **
| +--------+
| | | +----------- Pletholophus_tenuis **
| | | |
| | +--+ +-- Sinanodonta_tumens **
| | | +-----+
| | | | +-- Sinanodonta_lauta **
| | +--+
+--+ | +----- Beringiana_fukuharai **
| +--+
| | +-- Beringiana_japonica **
| +--+
| +-- Beringiana_beringiana **
|
| +-------------------- Lanceolaria_grayii **
| |
+--+ +-- Nodularia_nipponensis **
| +--------------+
| | +-- Nodularia_douglasiae **
+--+
| +-------------- Inversiunio_jokohamensis **
+--+
| +----------- Inversiunio_yanagawensis[1] **
+--+
| +-------- Inversiunio_yanagawensis[2] **
+--+
| +----- Inversiunio_yanagawensis[3] **
+--+
| +-- Inversiunio_yanagawensis[4] **
+--+
+-- Inversiunio_reinianus **
Lopes-Lima et al., 2021 [Publ 4217], fig. 1
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 25 |
25 |
23 |
1 |
0 |
0 |
0 |
89.0 |
+----- Potomida_littoralis **
+--+
| | +-- Potomida_semirugata **
| +--+
+--------------------+ +-- Potomida_acarnanica **
| |
| | +----- Leguminaia_anatolica **
| +--+
| | +-- Leguminaia_saulcyi **
| +--+
--+ +-- Leguminaia_wheatleyi **
|
| +----- Anodonta_anatina **
| +--------------------+
| | | +-- Anodonta_cygnea **
| | +--+
| | +-- Anodonta_seddoni **
+--+
| +----------------------- Unio_tumidus **
| |
| | +-------- Unio_damascensis **
| | +-----------+
+--+ | | +----- Unio_sesirmensis **
| | +--+
| | | +-- Unio_bruguierianus **
| | +--+
+--+ +-- Unio_crassus **
|
| +-- Unio_delicatus **
| +--------------+
| | +-- Unio_terminalis **
| |
+--+ +-- Unio_tigridis **
| +-----------+
| | +-- Unio_hueti **
| |
+--+ +----------- Unio_ravoisieri **
| |
| | +-- Unio_pictorum **
+--+ +-----+
| | +-- Unio_eucirrus **
| |
+--+ +-- Unio_elongatulus **
| +--+
| | +-- Unio_mancus **
+--+
| +-- Unio_foucauldianus **
+--+
+-- Unio_delphinus **
Soroka & Burzynski, 2017 [Publ 3837], fig. 1
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 24 |
24 |
22 |
1 |
0 |
0 |
0 |
78.0 |
+-------- Cristaria_plicata **
|
+-----+ +-- Anemina_arcaeformis **
| | +--+
| | | +-- Anemina_euscaphys **
| +--+
| | +-- Sinanodonta_woodiana **
+--------+ +--+
| | +-- Sinanodonta_lucida **
| |
| | +-- Anodonta_cygnea **
| | +--------+
| | | +-- Anodonta_anatina **
| +--+
| | +-------- Platynaias_compressa **
| +--+
--+ | +----- Pyganodon_grandis **
| +--+
| | +-- Utterbackia_imbecillis **
| +--+
| +-- Utterbackia_peninsularis **
|
| +-- Lanceolaria_lanceolata **
| +-----------------+
| | +-- Lanceolaria_grayii **
| |
+--+ +----------------- Lepidodesma_languilati **
| |
| | +----- Aculamprotula_tortuosa **
+--+ +--------+
| | | +-- Aculamprotula_tientsinensis **
| | +--+
| | +-- Aculamprotula_coreana **
+--+
| +----- Schistodesmus_lampreyanus **
| +-----+
| | | +-- Tchangsinaia_piscicula **
| | +--+
+--+ +-- Nodularia_douglasiae **
|
| +-------- Unio_tumidus **
+--+
| +----- Unio_crassus **
+--+
| +-- Unio_pictorum **
+--+
+-- Unio_delphinus **
Wu et al., 2021 [Publ 4215], fig. 3
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 24 |
24 |
22 |
1 |
0 |
0 |
0 |
83.0 |
+-------------------------- Indonaia_andersoniana **
|
| +-------- Acuticosta_chinensis **
| +--------------+
| | | +----- Nodularia_douglasiae **
--+ | +--+
| | | +-- Inversiunio_jokohamensis **
| | +--+
| | +-- Inversiunio_reinianus **
| |
+--+ +----- Chamberlainia_somsakpanhai **
| +--------------+
| | | +-- Sundadontina_cumingii **
| | +--+
| | +-- Sinohyriopsis_schlegelii **
| |
| | +-- Indopseudodon_bogani **
| | +--+
+--+ | +-- Indopseudodon_salwenianus **
| +-----------+
| | | +-- Pseudodon_vagulus **
| | +--+
| | +-- Pilsbryoconcha_carinifera **
| |
| | +-------- Sinosolenaia_carinata **
+--+ |
| +-----+ +-- Lamprotula_caveata **
| | | +--+
| | | | +-- Lamprotula_leaii **
| | +--+
| | | +-- Potomida_littoralis **
| | +--+
+--+ +-- Pronodularia_japanensis **
|
| +-- Ptychorhynchus_pfisteri **
| +--------+
| | +-- Parvasolenaia_rivularis **
| |
+--+ +-- Inversidens_brandtii **
| +-----+
| | +-- Inversidens_rentianensis **
+--+
| +----- Gonidea_angulata **
+--+
| +-- Microcondylaea_bonellii **
+--+
+-- Leguminaia_wheatleyi **
Liu et al., 2023 [Publ 4310], fig. 4
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 24 |
24 |
22 |
1 |
0 |
0 |
0 |
81.0 |
+-- Sinosolenaia_oleivora **
+--------------------------------------+
| +-- Lamprotula_leaii **
|
--+ +-------------------------------------- Acuticosta_chinensis **
| |
| | +-- Lanceolaria_lanceolata **
+--+ +--------------------------------+
| | +-- Lanceolaria_grayii **
| |
+--+ +-------------------------------- Lanceolaria_gladiola **
| |
| | +-------- Pseudanodonta_complanata **
| | +--------------------+
+--+ | | +----- Lasmigona_costata **
| | +--+
| | | +-- Strophitus_undulatus **
| | +--+
+--+ +-- Pyganodon_grandis **
|
| +-------------------------- Unio_tumidus **
| |
| | +-- Aculamprotula_scripta **
+--+ +--------------------+
| | +-- Aculamprotula_tientsinensis **
| |
+--+ +-- Cuneopsis_heudei **
| +-----------------+
| | +-- Cuneopsis_rufescens **
| |
+--+ +----- Nodularia_douglasiae **
| +-----------+
| | | +-- Schistodesmus_lampreyanus **
| | +--+
+--+ +-- Schistodesmus_spinosus **
|
| +-------------- Tchangsinaia_piscicula **
+--+
| +----------- Inversiunio_yanagawensis **
+--+
| +-------- Pseudocuneopsis_capitata **
+--+
| +----- Pseudocuneopsis_sichuanensis **
+--+
| +-- Pseudocuneopsis_wuana **
+--+
+-- Pseudocuneopsis_yangshuoensis **
Liu et al., 2023 [Publ 4310], fig. 5
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 24 |
24 |
17 |
1 |
0 |
0 |
0 |
83.0 |
+-- Sinosolenaia_oleivora **
+--------------------------+
| +-- Lamprotula_leaii **
|
--+ +-------------------------- Acuticosta_chinensis **
| |
| | +----------------------- Lanceolaria_grayii **
| | |
+--+ +----------------------- Lanceolaria_gladiola **
| |
| +----------------------- Lanceolaria_lanceolata **
| |
+--+ +-------- Lasmigona_costata **
| +-----------+
| | | +----- Pseudanodonta_complanata **
| | +--+
| | | +-- Pyganodon_grandis **
| | +--+
| | +-- Strophitus_undulatus **
+--+
| +----------------- Unio_tumidus **
| |
| | +-- Aculamprotula_tientsinensis **
| +--------------+
| | +-- Aculamprotula_scripta **
| |
+--+ +-- Cuneopsis_rufescens **
| +-----------+
| | +-- Cuneopsis_heudei **
| |
| | +----- Nodularia_douglasiae **
| +--------+
+--+ | +-- Schistodesmus_lampreyanus **
| +--+
| +-- Schistodesmus_spinosus **
|
| +----------- Tchangsinaia_piscicula **
| |
+--+----------- Inversiunio_yanagawensis **
|
| +-------- Pseudocuneopsis_capitata **
+--+
| +----- Pseudocuneopsis_sichuanensis **
+--+
| +-- Pseudocuneopsis_wuana **
+--+
+-- Pseudocuneopsis_yangshuoensis **
Zhou et al., 2007 [Publ 2658], fig. 1
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 23 |
23 |
21 |
2 |
0 |
0 |
0 |
88.0 |
+-- Margaritifera_margaritifera **
+-----------------------+
| +-- Cumberlandia_monodonta **
|
| +-- Sinanodonta_woodiana **
| +--------------------+
--+ | +-- Cristaria_plicata **
| |
| | +-- Sinohyriopsis_cumingii **
| | +--------+
| | | +-- Sinohyriopsis_schlegelii **
| | +--------+
+--+ | | +-------- Lamprotula_leaii **
| | +--+
| | | +----- Lamprotula_caveata **
| | +--+
| | | +-- Gibbosula_rochechouartii **
| | +--+
| | +-- Lamprotula_cornuumlunae **
| |
+--+ +-- Schistodesmus_lampreyanus **
| +-----+
| | +-- Schistodesmus_spinosus **
| |
| +--------+ +-- Lanceolaria_triformis **
| | | +--+
| | | | +-- Lanceolaria_lanceolata **
| | +--+
| | | +-- Acuticosta_ovata **
+--+ +--+
| +-- Acuticosta_chinensis **
|
| +-- Tchangsinaia_piscicula **
| +-----------+
| | +-- Nodularia_douglasiae **
+--+
| +----------- Aculamprotula_tortuosa **
+--+
| +-------- Aculamprotula_scripta **
+--+
| +----- Aculamprotula_tientsinensis **
+--+
| +-- Aculamprotula_zonata **
+--+
+-- Aculamprotula_fibrosa **
Konopleva et al., 2017 [Publ 3807], fig. 6
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 23 |
23 |
21 |
2 |
8 |
6 |
2 |
83.0 |
+----- Diplodon_suavidicus
+--------------------+
| | +-- Triplodon_corrugatus
| +--+
| +-- Castalia_ambigua
|
| +----- Tamsiella_schroeteriana
--+ +-----------------+
| | | +-- Etheria_elliptica
| | +--+
| | +-- Chambardia_wahlbergi
| |
| | +-- Margaritifera_dahurica **
| | +-----------------+
+--+ | +-- Margaritifera_laevis **
| |
| | +-------- Ensidens_sagittarius **
| | +-----+
| | | | +----- Lens_micropterus **
| | | +--+
| | | | +-- Physunio **
| | +--+ +--+
+--+ | | +-- Physunio_modelli **
| | |
| | | +----------- Sundadontina_tumida **
| | | |
| | +--+ +-- Nyeinchanconcha_nyeinchani **
| | | +-----+
| | | | +-- Bineurus_exilis **
| | +--+
| | | +----- Pilsbryoconcha_carinifera **
| | +--+
+--+ | +-- Pseudodon_vagulus **
| +--+
| +-- Pseudodon_vondembuschianus **
|
| +-- Trapezidens_exolescens **
| +-----+
| | +-- Trapezidens_dolichorhynchus **
| +-----+
| | | +----- Lamellidens_savadiensis **
| | +--+
| | | +-- Lamellidens_jenkinsianus **
| | +--+
+--+ +-- Lamellidens_mainwaringi **
|
| +-- Leoparreysia_tavoyensis **
| +--------+
| | +-- Leoparreysia_olivacea **
+--+
| +-------- Coelatura_aegyptiaca **
+--+
| +----- Indochinella_pugio **
+--+
| +-- Radiatula_chaudhurii **
+--+
+-- Scabies_mandarinus **
Jeratthitikul et al., 2019 [Publ 3922], fig. 1
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 23 |
23 |
21 |
1 |
0 |
0 |
0 |
95.0 |
+-- Lamellidens_mainwaringi **
+-----------------------------------------------+
| +-- Trapezidens_exolescens **
--+
| +----------------------------------------------- Leoparreysia_canefrii **
| |
+--+ +-------------------------------------------- Parreysia_keralaensis **
| |
+--+ +----------------------------------------- Coelatura_aegyptiaca **
| |
| | +----- Indonaia_rugosa **
+--+ +--------------------------------+
| | | +-- Indonaia_caerulea **
| | +--+
| | +-- Indonaia_andersoniana **
+--+
| +----------------------------------- Indochinella_pugio **
| |
| | +----- Radiatula_mouhoti **
+--+ +--------------------------+
| | | +-- Radiatula_myitkyinae **
| | +--+
+--+ +-- Radiatula_chaudhurii **
|
| +----------------------------- Scabiellus_songkramensis **
+--+
| +-------------------------- Harmandia_somboriensis **
+--+
| +----------------------- Unionetta_fabagina **
+--+
| +-------------------- Scabies_crispata **
+--+
| +----------------- Scabies_mandarinus **
+--+
| +-------------- Scabies_pilata **
+--+
| +----------- Scabies_nucleus **
+--+
| +-------- Scabies_humilis **
+--+
| +----- Scabies_scobinatus **
+--+
| +-- Scabies_phaselus **
+--+
+-- Scabies_anceps **
Wu et al., 2020 [Publ 4121], fig. 2
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 23 |
23 |
21 |
2 |
11 |
9 |
2 |
86.0 |
+----- Triplodon_corrugatus
+--------------------+
| | +-- Hyridella_australis
| +--+
| +-- Velesunio_ambiguus
|
| +----------- Mutela_hargeri
| |
| +-----------+ +-- Aspatharia_pfeifferiana
| | | +-----+
--+ | | | +-- Chambardia_wahlbergi
| | +--+
| | | +----- Etheria_elliptica
| | +--+
| | | +-- Tamsiella_schroeteriana
| | +--+
| | +-- Lamproscapha_ensiformis
| |
| | +-- Lampsilis_cardium **
| | +--------------+
+--+ | +-- Scabies_crispata **
| |
| | +-------- Sinosolenaia_carinata **
| +--+ +-----+
| | | | | +----- Potomida_littoralis **
| | | | +--+
| | | | | +-- Lamprotula_caveata **
| | +--+ +--+
| | | +-- Lamprotula_leaii **
| | |
| | | +----------- Unio_pictorum **
| | +--+
| | | +-------- Aculamprotula_tortuosa **
+--+ +--+
| | +----- Aculamprotula_polysticta **
| +--+
| | +-- Aculamprotula_fibrosa **
| +--+
| +-- Aculamprotula_tientsinensis **
|
| +----------------- Cumberlandia_monodonta **
| |
| | +-- Gibbosula_laosensis **
| | +--+
+--+ | +-- Gibbosula_rochechouartii **
| +--------+
| | | +-- Pseudunio_auricularius **
| | +--+
| | +-- Pseudunio_marocanus **
+--+
| +-- Margaritifera_dahurica **
| +--------+
| | +-- Margaritifera_margaritifera **
+--+
| +-------- Margaritifera_falcata **
| |
+--+ +-- Margaritifera_hembeli **
| +--+
| | +-- Margaritifera_marrianae **
+--+
| +-- Margaritifera_laevis **
+--+
+-- Margaritifera_middendorffi **
Wu et al., 2020 [Publ 4121], fig. S1
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 23 |
23 |
21 |
2 |
11 |
9 |
2 |
86.0 |
+----- Triplodon_corrugatus
+-----------------------+
| | +-- Hyridella_australis
| +--+
| +-- Velesunio_ambiguus
|
| +----------- Mutela_hargeri
| |
| +--------------+ +-- Aspatharia_pfeifferiana
| | | +-----+
--+ | | | +-- Chambardia_wahlbergi
| | +--+
| | | +----- Etheria_elliptica
| | +--+
| | | +-- Tamsiella_schroeteriana
| | +--+
| | +-- Lamproscapha_ensiformis
| |
| | +-- Lampsilis_cardium **
| | +--------------+
+--+ | +-- Scabies_crispata **
| |
| | +-------- Sinosolenaia_carinata **
| +-----+ +-----+
| | | | | +----- Potomida_littoralis **
| | | | +--+
| | | | | +-- Lamprotula_caveata **
| | +--+ +--+
| | | +-- Lamprotula_leaii **
| | |
| | | +----------- Unio_pictorum **
| | +--+
+--+ | +-------- Aculamprotula_tortuosa **
| +--+
| | +----- Aculamprotula_polysticta **
| +--+
| | +-- Aculamprotula_tientsinensis **
| +--+
| +-- Aculamprotula_fibrosa **
|
| +-------------------- Gibbosula_rochechouartii **
| |
+--+ +----------------- Gibbosula_laosensis **
| |
| | +----- Cumberlandia_monodonta **
+--+ +--------+
| | | +-- Pseudunio_auricularius **
| | +--+
| | +-- Pseudunio_marocanus **
+--+
| +-- Margaritifera_dahurica **
| +--------+
| | +-- Margaritifera_margaritifera **
+--+
| +-------- Margaritifera_falcata **
| |
+--+ +-- Margaritifera_hembeli **
| +--+
| | +-- Margaritifera_marrianae **
+--+
| +-- Margaritifera_laevis **
+--+
+-- Margaritifera_middendorffi **
Zieritz et al., 2021 [Publ 4231], fig. 3-M, 4
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 23 |
23 |
21 |
2 |
0 |
0 |
0 |
77.0 |
+----------------------- Pseudunio_marocanus **
|
| +----- Quadrula_quadrula **
| +--------------+
| | | +-- Venustaconcha_ellipsiformis **
--+ | +--+
| | +-- Potamilus_alatus **
| |
| | +----- Unio_tumidus **
| | +-----+
| | | | +-- Unio_delphinus **
+--+ | +--+
| +-----+ +-- Unio_crassus **
| | |
| | | +-------- Sinanodonta_woodiana **
| | +--+
| | | +----- Anodonta_anatina **
| | +--+
| | | +-- Utterbackia_peninsularis **
| | +--+
+--+ +-- Pyganodon_grandis **
|
| +-- Chamberlainia_somsakpanhai **
| +-----------+
| | +-- Sinohyriopsis_cumingii **
| |
| | +-- Lens_contradens **
| | +--+
+--+ | +-- Physunio_superbus **
| +-----+
| | | +-- Hyriopsis_bialata **
| | +--+
| | +-- Rectidens_sumatrensis **
+--+
| +-- Pilsbryoconcha_exilis **
| +-----+
| | +-- Pseudodon_vondembuschianus **
| |
+--+ +-- Pronodularia_japanensis **
| +--+
| | +-- Potomida_littoralis **
+--+
| +-- Microcondylaea_bonellii **
+--+
+-- Sinosolenaia_carinata **
Wu et al., 2023 [Publ 4296], fig. 4
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 23 |
23 |
21 |
1 |
0 |
0 |
0 |
78.0 |
+-- Sinosolenaia_oleivora **
+--------------------------------+
| +-- Lamprotula_leaii **
|
--+ +-------------------------------- Acuticosta_chinensis **
| |
| | +----- Lanceolaria_gladiola **
| | +-----------------------+
+--+ | | +-- Lanceolaria_lanceolata **
| | +--+
| | +-- Lanceolaria_grayii **
| |
+--+ +-------- Pseudanodonta_complanata **
| +-----------------+
| | | +----- Lasmigona_costata **
| | +--+
| | | +-- Strophitus_undulatus **
| | +--+
+--+ +-- Pyganodon_grandis **
|
| +----------------------- Unio_tumidus **
| |
| | +-- Aculamprotula_scripta **
+--+ +-----------------+
| | +-- Aculamprotula_tientsinensis **
| |
+--+ +-- Cuneopsis_heudei **
| +--------------+
| | +-- Cuneopsis_rufescens **
| |
+--+ +----- Nodularia_douglasiae **
| +--------+
| | | +-- Schistodesmus_lampreyanus **
| | +--+
+--+ +-- Schistodesmus_spinosus **
|
| +----------- Tchangsinaia_piscicula **
+--+
| +-------- Inversiunio_yanagawensis **
+--+
| +----- Pseudocuneopsis_capitata **
+--+
| +-- Pseudocuneopsis_yangshuoensis **
+--+
+-- Pseudocuneopsis_sichuanensis **
Huang et al., 2019 [Publ 3843], fig. 3-M
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 22 |
22 |
20 |
2 |
1 |
1 |
1 |
76.0 |
+----------------------------- Echyridella_menziesii
|
| +-- Pseudunio_marocanus **
| +-----------------------+
--+ | +-- Margaritifera_margaritifera **
| |
| | +----- Quadrula_quadrula **
| | +-----------------+
| | | | +-- Venustaconcha_ellipsiformis **
+--+ | +--+
| | +-- Potamilus_alatus **
| |
| | +----------- Sinohyriopsis_cumingii **
| | |
| | | +----- Potomida_littoralis **
+--+ +--------+ +--+
| | | | | +-- Lamprotula_leaii **
| | | | +--+
| | +--+ +-- Pronodularia_japanensis **
| | |
| | | +----- Parvasolenaia_rivularis **
| | +--+
| | | +-- Sinosolenaia_oleivora **
+--+ +--+
| +-- Sinosolenaia_carinata **
|
| +-------- Sinanodonta_woodiana **
| +--------+
| | | +----- Anodonta_anatina **
| | +--+
| | | +-- Pyganodon_grandis **
+--+ +--+
| +-- Utterbackia_peninsularis **
|
| +-------------- Lanceolaria_lanceolata **
+--+
| +----------- Schistodesmus_lampreyanus **
+--+
| +-------- Aculamprotula_tortuosa **
+--+
| +----- Unio_tumidus **
+--+
| +-- Unio_crassus **
+--+
+-- Unio_delphinus **
Jeratthitikul et al., 2019 [Publ 3911], fig. 4
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 22 |
22 |
14 |
1 |
0 |
0 |
0 |
81.0 |
+-- Pilsbryoconcha_exilis **
+--------------------------+
| +-- Scabies_crispata **
|
--+ +-------------------------- Ensidens_ingallsianus **
| |
| | +-- Physunio_modelli **
+--+ +--------------------+
| | +-- Physunio_superbus **
| |
| | +-- Lens_maenamensis **
+--+ +-----------------+
| | +-- Lens_rolfbrandti **
| |
| | +----------------- Lens_contradens[12] **
+--+ |
| | +----- Lens_micropterus[3] **
| | |
| +-----------+ +-- Lens_contradens[10] **
| | | |
+--+ +--+-- Lens_micropterus[2] **
| |
| +-- Lens_contradens[11] **
|
| +-------------- Lens_contradens[9] **
| |
+--+ +----------- Lens_contradens[8] **
| |
+--+ +-------- Lens_contradens[7] **
| |
+--+ +----- Lens_contradens[6] **
| |
| | +-- Lens_contradens[1] **
+--+ |
| +-- Lens_contradens[2] **
| |
| +-- Lens_contradens[3] **
+--+
+-- Lens_contradens[4] **
|
+-- Lens_micropterus[1] **
|
+-- Lens_contradens[5] **
Wu et al., 2018 [Publ 4143], fig. 2
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 22 |
22 |
20 |
1 |
0 |
0 |
0 |
79.0 |
+----- Quadrula_quadrula **
+-----+
| | +-- Venustaconcha_ellipsiformis **
| +--+
| +-- Lampsilis_ornata **
+-----------+
| | +-- Sinohyriopsis_schlegelii **
| | +-----+
| | | +-- Sinohyriopsis_cumingii **
| | |
| +--+ +-- Sinosolenaia_oleivora **
| | +--+
| | | +-- Sinosolenaia_carinata **
| +--+
| | +-- Pronodularia_japanensis **
--+ +--+
| +-- Lamprotula_leaii **
|
| +-- Unio_pictorum **
| +--------+
| | +-- Nodularia_douglasiae **
| +--------+
| | | +-------- Aculamprotula_tortuosa **
| | +--+
| | | +----- Aculamprotula_scripta **
| | +--+
+--+ | +-- Aculamprotula_coreana **
| +--+
| +-- Aculamprotula_tientsinensis **
|
| +----------------- Sinanodonta_lucida **
+--+
| +-------------- Anemina_arcaeformis **
+--+
| +----------- Cristaria_plicata **
+--+
| +-------- Anodonta_anatina **
+--+
| +----- Pyganodon_grandis **
+--+
| +-- Utterbackia_peninsularis **
+--+
+-- Utterbackia_imbecillis **
Zhou et al., 2007 [Publ 2658], fig. 2
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 21 |
21 |
19 |
2 |
0 |
0 |
0 |
92.0 |
+-------------------------- Sinanodonta_woodiana **
|
| +----------------------- Cristaria_plicata **
| |
--+ | +----- Acuticosta_chinensis **
| | +--------+
| | | | +-- Nodularia_douglasiae **
| | | +--+
| | +-----+ +-- Cuneopsis_celtiformis **
+--+ | |
| | | +----------- Aculamprotula_tortuosa **
| | +--+
| | | +-------- Aculamprotula_scripta **
| | +--+
| | | +----- Aculamprotula_fibrosa **
| | +--+
| | | +-- Aculamprotula_zonata **
+--+ +--+
| +-- Aculamprotula_tientsinensis **
|
| +-------- Lamprotula_leaii **
| +--------+
| | | +----- Lamprotula_cornuumlunae **
| | +--+
| | | +-- Gibbosula_rochechouartii **
| | +--+
+--+ +-- Lamprotula_caveata **
|
| +-- Theliderma_intermedia **
| +-----------+
| | +-- Theliderma_sparsa **
+--+
| +----------- Quadrula_quadrula **
+--+
| +-------- Pustulosa_infucata **
+--+
| +----- Quadrula_verrucosa **
+--+
| +-- Pustulosa_kieneriana **
+--+
+-- Pustulosa_succissa **
Chapman et al., 2008 [Publ 2667], fig. 1
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 21 |
21 |
19 |
1 |
0 |
0 |
0 |
90.0 |
+-------------------------------------------- Pronodularia_japanensis **
|
| +-- Quadrula_quadrula **
--+ +--------------------------------------+
| | +-- Pustulosa_pustulosa **
| |
+--+ +-- Pleurobema_sintoxia **
| +-----------------------------------+
| | +-- Fusconaia_flava **
+--+
| +----------------------------------- Amblema_plicata **
| |
+--+ +-------------------------------- Plectomerus_dombeyanus **
| |
+--+ +----------------------------- Popenaias_popeii **
| |
+--+ +-------------------------- Toxolasma_lividum **
| |
+--+ +----------------------- Obliquaria_reflexa **
| |
| | +-- Cyrtonaias_tampicoensis **
+--+ +-----------------+
| | +-- Glebula_rotundata **
| |
+--+ +-- Ptychobranchus_fasciolaris **
| +--------------+
| | +-- Lemiox_rimosus **
| |
+--+ +-- Venustaconcha_ellipsiformis **
| +-----------+
| | +-- Obovaria_olivaria **
+--+
| +----------- Hamiota_subangulata **
+--+
| +-------- Lampsilis_ovata **
+--+
| +----- Ortmanniana_ligamentina **
+--+
| +-- Lampsilis_straminea **
+--+
+-- Lampsilis_hydiana **
Chong et al., 2008 [Publ 2668], fig. 3
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 21 |
21 |
10 |
1 |
0 |
0 |
0 |
86.0 |
+----------------- Anodonta_cygnea **
|
+----------------- Pyganodon_fragilis **
|
+----------------- Alasmidonta_holstonia **
|
+----------------- Anodonta_anatina **
|
+----------------- Unio_tumidus **
|
| +-- Anodonta_nuttalliana **
+--------------+
| +-- Anodonta_californiensis **
|
--+ +-- Beringiana_beringiana **
+--------------+
| +-- Sinanodonta_woodiana **
|
| +-- Pseudodontoideus_subvexus **
| |
+--------------+-- Alasmidonta_marginata **
| |
| +-- Lasmigona_complanata **
|
| +-------------- Pseudanodonta_complanata **
| |
| | +----- Strophitus_undulatus **
| +--------+
+--+ | +-- Prolasmidonta_heterodon **
| +--+
| +-- Platynaias_compressa **
|
| +----------- Anodonta_oregonensis[4] **
+--+
| +-------- Anodonta_oregonensis[3] **
+--+
| +----- Anodonta_oregonensis[2] **
+--+
| +-- Anodonta_oregonensis[1] **
+--+
+-- Anodonta_kennerlyi **
Rosenberg et al., 1997 [Publ 2720], fig. 2A
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 21 |
23 |
4 |
2 |
2 |
0 |
0 |
88.0 |
+----------- Cumberlandia_monodonta **
|
+----------- None[2]
|
| +-- Margaritifera_falcata **
+--------+
--+ +-- Margaritifera_margaritifera **
|
| +-------- None[1]
| |
| | +----- Anodonta_anatina **
| | |
| | | +-- Gonidea_angulata **
+--+ | |
| | +-- Pyganodon_grandis **
| | |
| | +-- Utterbackia_imbecillis **
| | |
| | +-- Pyganodon_cataracta **
+--+ |
| +-- Lampsilis_straminea **
| |
| +-- Lampsilis_teres **
| |
| +-- Megalonaias_nervosa **
| |
| +-- Obliquaria_reflexa **
| |
+--+-- Theliderma_cylindrica **
|
+-- Quadrula_quadrula **
|
+-- Uniomerus_tetralasmus **
|
+-- Amblema_plicata **
|
+-- Elliptio_complanata **
|
+-- Fusconaia_cerina **
|
+-- Plectomerus_dombeyanus **
|
+-- Pleurobema_cordatum **
|
+-- Unio_pictorum **
Rosenberg et al., 1997 [Publ 2720], fig. 2B
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 21 |
23 |
3 |
2 |
2 |
0 |
0 |
87.5 |
+-------- Cumberlandia_monodonta **
|
+-------- None[1]
|
+-------- None[2]
|
| +-- Margaritifera_falcata **
+-----+
| +-- Margaritifera_margaritifera **
--+
| +----- Anodonta **
| |
| | +-- Gonidea_angulata **
| | |
| | +-- Pyganodon_grandis **
| | |
| | +-- Utterbackia_imbecillis **
| | |
| | +-- Pyganodon_cataracta **
+--+ |
| +-- Lampsilis_straminea **
| |
| +-- Lampsilis_teres **
| |
| +-- Megalonaias_nervosa **
| |
| +-- Obliquaria_reflexa **
| |
+--+-- Theliderma_cylindrica **
|
+-- Quadrula_quadrula **
|
+-- Uniomerus_tetralasmus **
|
+-- Amblema_plicata **
|
+-- Elliptio_complanata **
|
+-- Fusconaia_cerina **
|
+-- Plectomerus_dombeyanus **
|
+-- Aculamprotula_tortuosa **
|
+-- Unio_pictorum **
Chapman et al., 2008 [Publ 2744], fig. 2
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 21 |
21 |
19 |
1 |
0 |
0 |
0 |
90.0 |
+-------------------------------------------- Pronodularia_japanensis **
|
| +-- Quadrula_quadrula **
--+ +--------------------------------------+
| | +-- Pustulosa_pustulosa **
| |
+--+ +-- Pleurobema_sintoxia **
| +-----------------------------------+
| | +-- Fusconaia_flava **
+--+
| +----------------------------------- Amblema_plicata **
| |
+--+ +-------------------------------- Plectomerus_dombeyanus **
| |
+--+ +----------------------------- Popenaias_popeii **
| |
+--+ +-------------------------- Toxolasma_lividum **
| |
+--+ +----------------------- Obliquaria_reflexa **
| |
| | +-- Cyrtonaias_tampicoensis **
+--+ +-----------------+
| | +-- Glebula_rotundata **
| |
+--+ +-- Ptychobranchus_fasciolaris **
| +--------------+
| | +-- Lemiox_rimosus **
| |
+--+ +-- Venustaconcha_ellipsiformis **
| +-----------+
| | +-- Obovaria_olivaria **
+--+
| +----------- Hamiota_subangulata **
+--+
| +-------- Lampsilis_ovata **
+--+
| +----- Ortmanniana_ligamentina **
+--+
| +-- Lampsilis_straminea **
+--+
+-- Lampsilis_hydiana **
Inoue et al., 2014 [Publ 3635], fig. 3A
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 21 |
21 |
17 |
2 |
0 |
0 |
0 |
89.0 |
+----------------------------- Cumberlandia_monodonta **
|
| +-------------------------- Amblema_plicata **
--+ |
| +-------------------------- Eurynia_dilatata **
| |
+--+-------------------------- Lampsilis_ornata **
|
| +----------------------- Quadrula_quadrula **
| |
| | +-------------------- Gonidea_angulata **
+--+ |
| | +----------- Pseudodontoideus_subvexus **
| | +-----+
+--+ | | +-------- Lasmigona_costata **
| | +--+
| | | +----- Lasmigona_complanata **
| | +--+
| | | +-- Arcidens_confragosus **
+--+ +--+
| +-- Arcidens_wheeleri **
|
| +-------------- Strophitus_undulatus **
| |
| | +-------- Pyganodon_grandis **
| | +--+
+--+ | | +----- Utterbackia_peninsularis[2] **
| | +--+
| | | +-- Utterbackia_imbecillis **
| | +--+
+--+ +-- Utterbackia_peninsularis[1] **
|
| +-- Platynaias_compressa **
| +-----+
| | +-- Platynaias_subviridis **
+--+
| +----- Alasmidonta_marginata **
+--+
| +-- Alasmidonta_undulata **
+--+
+-- Alasmidonta_varicosa **
Wu et al., 2016 [Publ 3711], fig. 1
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 21 |
21 |
19 |
1 |
0 |
0 |
0 |
78.0 |
+----- Quadrula_quadrula **
+-----+
| | +-- Venustaconcha_ellipsiformis **
| +--+
| +-- Lampsilis_ornata **
+-----------+
| | +-- Sinohyriopsis_schlegelii **
| | +-----+
| | | +-- Sinohyriopsis_cumingii **
| | |
| +--+ +-- Sinosolenaia_carinata **
| | +--+
| | | +-- Sinosolenaia_oleivora **
| +--+
| | +-- Lamprotula_leaii **
--+ +--+
| +-- Pronodularia_japanensis **
|
| +-- Unio_pictorum **
| +-----+
| | +-- Nodularia_douglasiae **
| +-----------+
| | | +----- Aculamprotula_tortuosa **
| | +--+
| | | +-- Aculamprotula_coreana **
+--+ +--+
| +-- Aculamprotula_tientsinensis **
|
| +----------------- Sinanodonta_lucida **
+--+
| +-------------- Anemina_arcaeformis **
+--+
| +----------- Cristaria_plicata **
+--+
| +-------- Anodonta_anatina **
+--+
| +----- Pyganodon_grandis **
+--+
| +-- Utterbackia_peninsularis **
+--+
+-- Utterbackia_imbecillis **
Guerra et al., 2017 [Publ 3772], fig. 2a
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 21 |
21 |
19 |
2 |
4 |
3 |
2 |
78.0 |
+-- Mutela_dubia
+-----------------------------+
| +-- Anodontites_trapesialis
|
| +----------------------------- Echyridella_menziesii
--+ |
| | +-- Margaritifera_falcata **
| | +-----------------------+
| | | +-- Cumberlandia_monodonta **
+--+ |
| | +-------- Quadrula_quadrula **
| | +--+
| | | | +----- Toxolasma_parvum **
| | | +--+
| | | | +-- Venustaconcha_ellipsiformis **
+--+ | +--+
| +-----------+ +-- Lampsilis_ornata **
| | |
| | | +-- Sinohyriopsis_schlegelii **
| | | +-----+
| | | | +-- Sinohyriopsis_cumingii **
| | +--+
| | | +----- Pronodularia_japanensis **
+--+ +--+
| | +-- Sinosolenaia_oleivora **
| +--+
| +-- Sinosolenaia_carinata **
|
| +-- Aculamprotula_tortuosa **
| +-----------------+
| | +-- Unio_pictorum **
+--+
| +----------------- Sinanodonta_woodiana **
| |
+--+ +-------------- Cristaria_plicata **
| |
+--+ +----------- Anodonta_anatina **
| |
+--+ +-- Platynaias_subviridis **
| +-----+
| | +-- Platynaias_compressa **
+--+
| +----- Pyganodon_grandis **
+--+
| +-- Utterbackia_peninsularis **
+--+
+-- Utterbackia_imbecillis **
Guerra et al., 2017 [Publ 3772], fig. 2b
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 21 |
21 |
19 |
2 |
4 |
3 |
2 |
80.0 |
+-- Mutela_dubia
+--------------------------------+
| +-- Anodontites_trapesialis
|
--+ +-------------------------------- Echyridella_menziesii
| |
| | +-- Margaritifera_falcata **
| | +--------------------------+
+--+ | +-- Cumberlandia_monodonta **
| |
| | +-------- Quadrula_quadrula **
| | +-----------------+
+--+ | | +----- Toxolasma_parvum **
| | +--+
| | | +-- Venustaconcha_ellipsiformis **
| | +--+
| | +-- Lampsilis_ornata **
| |
+--+ +-- Sinohyriopsis_schlegelii **
| +-----+
| | +-- Sinohyriopsis_cumingii **
| +--------------+
| | | +----- Pronodularia_japanensis **
| | +--+
| | | +-- Sinosolenaia_oleivora **
+--+ +--+
| +-- Sinosolenaia_carinata **
|
| +-------------------- Unio_pictorum **
| |
+--+ +----------------- Aculamprotula_tortuosa **
| |
+--+ +-- Cristaria_plicata **
| +-----------+
| | +-- Sinanodonta_woodiana **
+--+
| +----------- Anodonta_anatina **
| |
+--+ +-- Platynaias_subviridis **
| +-----+
| | +-- Platynaias_compressa **
+--+
| +----- Pyganodon_grandis **
+--+
| +-- Utterbackia_peninsularis **
+--+
+-- Utterbackia_imbecillis **
Jeratthitkul et al., 2021 [Publ 4212], fig. 4
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 21 |
21 |
19 |
1 |
0 |
0 |
0 |
85.0 |
+-------- Trapezoideus_foliaceus **
+-----+
| | +----- Physunio_superbus **
| +--+
| | +-- Physunio_pellucidus **
| +--+
+-----+ +-- Physunio_modelli **
| |
| | +-- Yaukthwa_nesemanni **
| | +--------+
| | | +-- Yaukthwa_panhai **
| +--+
| | +-------- Lens_contradens **
| +--+
| | +----- Lens_novoselovi **
--+ +--+
| | +-- Lens_rolfbrandti **
| +--+
| +-- Lens_maenamensis **
|
| +-- Ensidens_ingallsianus **
| +--------------+
| | +-- Ensidens_jaculus **
| |
+--+ +-- Hyriopsis_desowitzi **
| +-----------+
| | +-- Hyriopsis_delaportei **
| |
+--+ +----------- Rectidens_sumatrensis **
| |
| | +----- Hyriopsis_myersiana **
+--+ +--+
| | | +-- Hyriopsis_bialata **
| | +--+
+--+ +-- Hyriopsis_panhai **
|
| +----- Hyriopsis_khoratensis **
+--+
| +-- Hyriopsis_phuphaniensis **
+--+
+-- Hyriopsis_sakhonensis **
Bogan & Hoeh, 2000 [Publ 2650], fig. 1
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 20 |
21 |
14 |
2 |
14 |
13 |
2 |
81.0 |
+-- Castalia_stevensi
+-----+
| +-- Diplodon_gratus
+--------------------+
| | +----- Echyridella_menziesii
| +--+
| | +-- Velesunio_angasi
| +--+
| +-- Lortiella_rugata
|
| +-- Mutela_dubia
--+ +-----------+
| | +-- Mutela_rostrata
| +-----------+
| | | +----------- Etheria_elliptica
| | +--+
| | | +-------- Mulleria_rivoli
| | +--+
| | | +----- Monocondylaea_minuana
| | +--+
+--+ | +-- Anodontites_trigona
| +--+
| +-- Anodontites_guanarensis
|
| +----------------------- Coelatura_aegyptiaca **
| |
| | +-------------------- Pseudomulleria_dalyi
+--+ |
| | +----- Anodonta_cygnea **
| | +-----------+
+--+ | | +-- Strophitus_undulatus **
| | +--+
| | +-- Pyganodon_grandis **
| |
+--+ +-------------- Gonidea_angulata **
| |
| | +-- Unio_pictorum **
| +-----------+
| | +-- Unio_tumidus **
| |
+--+ +-- Cumberlandia_monodonta **
| +--------+
| | +-- Margaritifera_margaritifera **
| |
| | +-------- Toxolasma_lividum **
| | |
+--+ +-------- Amblema_plicata **
| |
| | +-- Glebula_rotundata **
| +-----+
| | +-- Cyrtonaias_tampicoensis **
| |
+--+ +----- Potamilus_alatus **
+--+
| | +-- Ortmanniana_ligamentina **
| +--+
| +-- Ligumia_recta **
|
| +----- Quadrula_quadrula **
| |
+--+ +-- Eurynia_dilatata **
| |
+--+-- Pleurobema_clava **
|
+-- Fusconaia_flava **
Hoeh et al., 2001 [Publ 2652], fig. 14.2
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 20 |
30 |
18 |
2 |
11 |
10 |
2 |
90.0 |
+-- Cumberlandia_monodonta **
+--------------------+
| +-- Margaritifera_margaritifera **
|
| +-------------------- Amblema_plicata **
| |
| +-------------------- Coelatura_aegyptiaca **
| |
| +-------------------- Eurynia_dilatata **
--+ |
| +-------------------- Fusconaia_flava **
| |
| +-------------------- Gonidea_angulata **
| |
| +-------------------- Pleurobema_clava **
| |
| +-------------------- Quadrula_quadrula **
| |
+--+ +----------- Ortmanniana_ligamentina **
+--------+
| | +-------- Glebula_rotundata **
| +--+
| | +----- Potamilus_alatus **
| +--+
| | +-- Ligumia_recta **
| +--+
| +-- Toxolasma_lividum **
|
| +----------------- Cyrtonaias_tampicoensis **
| |
| | +-- Unio_pictorum **
| | +-----------+
+--+ | +-- Unio_tumidus **
| |
| | +----- Pyganodon_grandis **
| | +-----+
+--+ | | +-- Anodonta_cygnea **
| | +--+
| | +-- Strophitus_undulatus **
| |
| | +----- Diplodon_gratus
+--+ |
| +----- Echyridella_menziesii
| +--+
| | +----- Lortiella_rugata
| | |
| | | +-- Castalia_stevensi
| | +--+
+--+ +-- Velesunio_angasi
|
| +-- Mutela_dubia
| +--+
| | +-- Mutela_rostrata
+--+
| +-- Anodontites_guanarensis
| |
+--+-- Anodontites_trigona
|
+-- Monocondylaea_minuana
Hoeh et al., 2001 [Publ 2652], fig. 14.4
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 20 |
20 |
17 |
2 |
11 |
10 |
2 |
78.0 |
+-- Castalia_stevensi
+-----+
| +-- Diplodon_gratus
+--------------------------+
| | +----- Echyridella_menziesii
| +--+
| | +-- Lortiella_rugata
| +--+
| +-- Velesunio_angasi
|
--+ +-- Mutela_dubia
| +-----+
| | +-- Mutela_rostrata
| +-----------------------+
| | | +----- Monocondylaea_minuana
| | +--+
| | | +-- Anodontites_guanarensis
| | +--+
+--+ +-- Anodontites_trigona
|
| +----------------------------- Coelatura_aegyptiaca **
| |
| | +-- Cumberlandia_monodonta **
+--+ +-----------------------+
| | +-- Margaritifera_margaritifera **
| |
| | +----------------------- Gonidea_angulata **
+--+ |
| | +-------- Unio_pictorum **
| | |
| | +-----------+-------- Unio_tumidus **
+--+ | |
| | | +----- Anodonta_cygnea **
| | +--+
| | | +-- Strophitus_undulatus **
| | +--+
+--+ +-- Pyganodon_grandis **
|
| +----------------- Amblema_plicata **
| |
| | +-------- Quadrula_quadrula **
| | +-----+
+--+ | | +----- Eurynia_dilatata **
| | +--+
| | | +-- Fusconaia_flava **
| | +--+
+--+ +-- Pleurobema_clava **
|
| +-- Cyrtonaias_tampicoensis **
| +--------+
| | +-- Glebula_rotundata **
+--+
| +-------- Toxolasma_lividum **
+--+
| +----- Potamilus_alatus **
+--+
| +-- Ligumia_recta **
+--+
+-- Ortmanniana_ligamentina **
Hoeh et al., 2002 [Publ 2653], fig. 2
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 20 |
20 |
17 |
2 |
11 |
10 |
2 |
79.0 |
+-- Castalia_stevensi
+-----+
| +-- Diplodon_gratus
+-----------------------+
| | +----- Echyridella_menziesii
| +--+
| | +-- Velesunio_angasi
| +--+
| +-- Lortiella
|
--+ +-- Mutela_dubia
| +-----+
| | +-- Mutela_rostrata
| +--------------------+
| | | +----- Monocondylaea_minuana
| | +--+
| | | +-- Anodontites_guanarensis
| | +--+
+--+ +-- Anodontites_trigona
|
| +-------------------------- Coelatura_aegyptiaca **
| |
| | +-- Margaritifera_margaritifera **
| | +--------------------+
+--+ | +-- Cumberlandia_monodonta **
| |
| | +-------------- Gonidea_angulata **
| | +-----+
+--+ | | +----------- Unio_pictorum **
| | +--+
| | | +-------- Unio_tumidus **
| | +--+
| | | +----- Anodonta_cygnea **
| | +--+
+--+ | +-- Pyganodon_grandis **
| +--+
| +-- Strophitus_undulatus **
|
| +-- Glebula_rotundata **
| +--------------+
| | +-- Cyrtonaias_tampicoensis **
| |
+--+ +-------------- Toxolasma_lividum **
| |
| | +----- Potamilus_alatus **
| +--------+
+--+ | +-- Ligumia_recta **
| +--+
| +-- Ortmanniana_ligamentina **
|
| +----------- Amblema_plicata **
+--+
| +-------- Quadrula_quadrula **
+--+
| +----- Eurynia_dilatata **
+--+
| +-- Pleurobema_clava **
+--+
+-- Fusconaia_flava **
Roe & Hoeh, 2003 [Publ 2655], fig. 4.2
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 20 |
20 |
16 |
2 |
11 |
10 |
2 |
84.0 |
+-- Mutela_dubia
+-----+
| +-- Mutela_rostrata
+--+
| | +----- Monocondylaea_minuana
| +--+
| | +-- Anodontites_guanarensis
| +--+
+-----------------------+ +-- Anodontites_trigona
| |
| | +-- Castalia_stevensi
| | +--+
| | | +-- Velesunio_angasi
| +-----+
| | +-- Echyridella_menziesii
--+ | |
| +--+-- Lortiella_rugata
| |
| +-- Diplodon_gratus
|
| +-- Cumberlandia_monodonta **
| +-----------------------------+
| | +-- Margaritifera_margaritifera **
+--+
| +----------------------------- Gonidea_angulata **
| |
+--+ +-------------------------- Coelatura_aegyptiaca **
| |
+--+ +----------------------- Fusconaia_flava **
| |
| +----------------------- Amblema_plicata **
+--+
| +-------------------- Quadrula_quadrula **
| |
| | +----------------- Pleurobema_clava **
+--+ |
| | +-------------- Eurynia_dilatata **
| | |
+--+ | +----------- Ortmanniana_ligamentina **
| +--+
| | | +-------- Glebula_rotundata **
| | +--+
| | | +----- Potamilus_alatus **
+--+ +--+
| | +-- Ligumia_recta **
| +--+
| +-- Toxolasma_lividum **
|
| +----------- Cyrtonaias_tampicoensis **
| |
+--+ +-- Unio_pictorum **
| +-----+
| | +-- Unio_tumidus **
+--+
| +----- Pyganodon_grandis **
+--+
| +-- Anodonta_cygnea **
+--+
+-- Strophitus_undulatus **
Roe & Hoeh, 2003 [Publ 2655], fig. 4.3
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 20 |
30 |
28 |
2 |
11 |
10 |
2 |
80.0 |
+-- Cumberlandia_monodonta **
+-----------------------------+
| +-- Margaritifera_margaritifera **
|
| +-- Mutela_dubia
| +-----+
| | +-- Mutela_rostrata
| +--+
| | | +----- Monocondylaea_minuana
| | +--+
--+ | | +-- Anodontites_guanarensis
| | +--+
| +-----------------+ +-- Anodontites_trigona
| | |
| | | +-- Castalia_stevensi
| | | +-----+
| | | | +-- Diplodon_gratus
| | +--+
| | | +----- Echyridella_menziesii
| | +--+
+--+ | +-- Velesunio_angasi
| +--+
| +-- Lortiella_rugata
|
| +-------------------------- Coelatura_aegyptiaca **
| |
| | +-- Unio_pictorum **
| | +-----+
| | | +-- Unio_tumidus **
+--+ +--------------+
| | | +----- Anodonta_cygnea **
| | +--+
| | | +-- Pyganodon_grandis **
| | +--+
+--+ +-- Strophitus_undulatus **
|
| +-------------------- Gonidea_angulata **
| |
| | +----------------- Amblema_plicata **
+--+ |
| | +-------- Quadrula_quadrula **
| | +-----+
+--+ | | +----- Eurynia_dilatata **
| | +--+
| | | +-- Fusconaia_flava **
| | +--+
+--+ +-- Pleurobema_clava **
|
| +-- Cyrtonaias_tampicoensis **
| +--------+
| | +-- Glebula_rotundata **
+--+
| +-------- Toxolasma_lividum **
+--+
| +----- Ortmanniana_ligamentina **
+--+
| +-- Ligumia_recta **
+--+
+-- Potamilus_alatus **
Zhou et al., 2016 [Publ 3710], fig. 1
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 20 |
20 |
18 |
2 |
0 |
0 |
0 |
81.0 |
+----- Lepidodesma_languilati **
+-----------+
| | +-- Schistodesmus_lampreyanus **
| +--+
+--+ +-- Unio_pictorum **
| |
| | +-------------- Sinanodonta_woodiana **
| +--+
| | +----------- Cristaria_plicata **
| +--+
| | +-------- Platynaias_compressa **
| +--+
| | +----- Pyganodon_grandis **
--+ +--+
| | +-- Utterbackia_imbecillis **
| +--+
| +-- Utterbackia_peninsularis **
|
| +-------------- Margaritifera_falcata **
| |
| | +-------- Quadrula_quadrula **
| | +--+
+-----+ | | +----- Toxolasma_parvum **
| | +--+
| | | +-- Venustaconcha_ellipsiformis **
| | +--+
+--+ +-- Lampsilis_ornata **
|
| +-- Sinohyriopsis_cumingii **
| +-----+
| | +-- Sinohyriopsis_schlegelii **
| |
+--+ +-- Lamprotula_leaii **
| +--+
| | +-- Pronodularia_japanensis **
+--+
| +-- Sinosolenaia_carinata **
+--+
+-- Sinosolenaia_oleivora **
Burzynski & Soroka, 2018 [Publ 3862], fig. 5a
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 20 |
20 |
17 |
2 |
0 |
0 |
0 |
77.0 |
+-- Pseudunio_marocanus **
+-----------------------+
| +-- Cumberlandia_monodonta **
|
| +----- Quadrula_quadrula **
| +-----------------+
--+ | | +-- Potamilus_alatus **
| | +--+
| | +-- Venustaconcha_ellipsiformis **
| |
| | +----------- Sinohyriopsis_cumingii **
+--+ +--------+
| | | +-------- Sinosolenaia_carinata **
| | +--+
| | | +----- Potomida_littoralis **
| | +--+
| | | +-- Lamprotula_leaii **
| | +--+
+--+ +-- Pronodularia_japanensis **
|
| +----- Sinanodonta_woodiana **
| |
| +-----------+----- Anodonta_anatina **
| | |
| | | +-- Utterbackia_peninsularis **
+--+ +--+
| +-- Pyganodon_grandis **
|
| +-------------- Lanceolaria_lanceolata **
+--+
| +----------- Aculamprotula_tortuosa **
+--+
| +-------- Unio_tumidus **
+--+
| +----- Unio_crassus **
+--+
| +-- Unio_pictorum **
+--+
+-- Unio_delphinus **
Porto-Hannes et al., 2019 [Publ 3969], fig. 2
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 20 |
20 |
16 |
1 |
0 |
0 |
0 |
90.0 |
+-------------------------- Amblema_plicata **
|
| +----------------------- Obliquaria_reflexa **
| |
--+ | +-- Cambarunio_iris **
| | +-----------------+
| | | +-- Leaunio_lienosus **
| | |
+--+ | +----------- Lampsilis_teres **
| | |
| +--------+ +-------- Ortmanniana_ligamentina **
| | | |
| | +--+ +-- Lampsilis_hydiana **
| | | +--+
+--+ | | +-- Lampsilis_virescens **
| +--+
| | +-- Lampsilis_siliquoidea **
| +--+
| +-- Lampsilis_radiata **
|
| +----- Obovaria_arkansasensis **
| +-----------+
| | | +-- Obovaria_olivaria **
| | +--+
+--+ +-- Obovaria_subrotunda **
|
| +-------------- Lampsilis_bracteata **
| |
+--+ +----------- Lampsilis_ornata **
| |
+--+ +-------- Lampsilis_ovata **
| |
+--+ +----- Lampsilis_cardium[3] **
| |
+--+ +-- Lampsilis_satura **
| |
+--+-- Lampsilis_cardium[1] **
|
+-- Lampsilis_cardium[2] **
Porto-Hannes et al., 2019 [Publ 3969], fig. 3
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 20 |
20 |
13 |
1 |
0 |
0 |
0 |
90.0 |
+-------------------- Obliquaria_reflexa **
|
+-------------------- Amblema_plicata **
|
| +----------------- Lampsilis_teres **
| |
| | +-- Leaunio_lienosus **
| +--------------+
--+ | +-- Cambarunio_iris **
| |
| | +----- Obovaria_arkansasensis **
| +-----------+
| | | +-- Obovaria_olivaria **
| | +--+
| | +-- Obovaria_subrotunda **
| |
+--+ +-------- Ortmanniana_ligamentina **
| |
+--------+ +-- Lampsilis_hydiana **
| | +--+
| | | +-- Lampsilis_virescens **
| +--+
| | +-- Lampsilis_siliquoidea **
| +--+
| +-- Lampsilis_radiata **
|
| +-------------- Lampsilis_bracteata **
| |
+--+ +----------- Lampsilis_ornata **
| |
+--+ +-------- Lampsilis_ovata **
| |
+--+ +----- Lampsilis_cardium[3] **
| |
+--+ +-- Lampsilis_satura **
| |
+--+-- Lampsilis_cardium[1] **
|
+-- Lampsilis_cardium[2] **
Riccardi et al., 2019 [Publ 4089], fig. 2
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 20 |
20 |
18 |
2 |
0 |
0 |
0 |
83.0 |
+-- Pseudunio_auricularius **
+-----------------------+
| +-- Pseudunio_marocanus **
|
| +-- Lanceolaria_grayii **
| +--+
| | +-- Lanceolaria_lanceolata **
--+ +-----------------+
| | | +-- Unio_mancus **
| | +--+
| | +-- Unio_elongatulus **
| |
| | +-------- Cristaria_plicata **
+--+ |
| +-----------+ +-- Anemina_euscaphys **
| | | +--+
| | | | +-- Anemina_arcaeformis **
| | +--+
| | | +-- Sinanodonta_lucida **
+--+ +--+
| +-- Sinanodonta_woodiana **
|
| +----------------- Anodonta_anatina **
| |
| | +----- Pseudanodonta_complanata **
+--+ +--------+
| | | +-- Anodonta_exulcerata **
| | +--+
+--+ +-- Anodonta_cygnea **
|
| +----------- Anodonta_nuttalliana **
+--+
| +-------- Platynaias_compressa **
+--+
| +----- Pyganodon_grandis **
+--+
| +-- Utterbackia_peninsularis **
+--+
+-- Utterbackia_imbecillis **
Smith et al., 2020 [Publ 4136], fig. 1
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 20 |
20 |
13 |
1 |
0 |
0 |
0 |
92.0 |
+-------------------------- Quadrula_quadrula **
|
--+ +----------------------- Elliptoideus_sloatianus **
| |
| | +-------------------- Elliptio_crassidens **
+--+ |
| +-------------------- Plethobasus_cyphyus **
| |
+--+ +----------------- Parvaspina_steinstansana **
| |
| | +-- Hemistena_lata **
| +--------------+
| | +-- Eurynia_dilatata **
+--+
| +-- Pleuronaia_barnesiana **
| +-----------+
| | +-- Pleurobema_clava **
| |
| | +----------- Fusconaia_subrotunda **
+--+ |
| | +-- Fusconaia_cor **
| +--------+
| | +-- Fusconaia_cuneolus **
+--+
| +-------- Fusconaia_flava **
| |
| +-------- Fusconaia_masoni **
| |
| | +-- Fusconaia_burkei **
+--+-----+
| +-- Fusconaia_escambia **
|
| +-- Fusconaia_iheringi **
| +--+
| | +-- Fusconaia_mitchelli **
+--+
| +-- Fusconaia_askewi **
+--+
+-- Fusconaia_chunii **
Guerra et al., 2019 [Publ 4168], fig. 1-M
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 20 |
20 |
18 |
2 |
2 |
2 |
1 |
76.0 |
+-- Echyridella_menziesii
+--------------------------+
| +-- Westralunio_carteri
|
| +----- Cumberlandia_monodonta **
| +--------------------+
--+ | | +-- Margaritifera_margaritifera **
| | +--+
| | +-- Pseudunio_marocanus **
| |
| | +----- Quadrula_quadrula **
+--+ +-----------------+
| | | +-- Venustaconcha_ellipsiformis **
| | +--+
| | +-- Potamilus_alatus **
| |
| | +-------- Sinosolenaia_carinata **
+--+ +-----------+
| | | +----- Potomida_littoralis **
| | +--+
| | | +-- Lamprotula_leaii **
| | +--+
| | +-- Pronodularia_japanensis **
+--+
| +-------- Sinanodonta_woodiana **
| +--------+
| | | +----- Anodonta_anatina **
| | +--+
| | | +-- Utterbackia_peninsularis **
+--+ +--+
| +-- Pyganodon_grandis **
|
| +-------------- Lanceolaria_lanceolata **
+--+
| +----------- Schistodesmus_lampreyanus **
+--+
| +-------- Aculamprotula_tortuosa **
+--+
| +----- Unio_tumidus **
+--+
| +-- Unio_crassus **
+--+
+-- Unio_delphinus **
Guerra et al., 2019 [Publ 4168], fig. 2-M
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 20 |
20 |
18 |
2 |
2 |
2 |
1 |
78.0 |
+-- Echyridella_menziesii
+--------------------------+
| +-- Westralunio_carteri
|
| +----- Pseudunio_marocanus **
| +--------------------+
--+ | | +-- Margaritifera_margaritifera **
| | +--+
| | +-- Cumberlandia_monodonta **
| |
| | +----- Quadrula_quadrula **
+--+ +-----------------+
| | | +-- Venustaconcha_ellipsiformis **
| | +--+
| | +-- Potamilus_alatus **
| |
| | +-------- Sinosolenaia_carinata **
+--+ +-----------+
| | | +----- Potomida_littoralis **
| | +--+
| | | +-- Lamprotula_leaii **
| | +--+
| | +-- Pronodularia_japanensis **
+--+
| +-- Utterbackia_peninsularis **
| +--+
| | +-- Pyganodon_grandis **
| +-----------+
| | | +-- Anodonta_anatina **
| | +--+
+--+ +-- Sinanodonta_woodiana **
|
| +-------------- Lanceolaria_lanceolata **
+--+
| +----------- Aculamprotula_tortuosa **
+--+
| +-------- Schistodesmus_lampreyanus **
+--+
| +----- Unio_tumidus **
+--+
| +-- Unio_crassus **
+--+
+-- Unio_delphinus **
Rosenberg et al., 1994 [Publ 2719], fig. 3
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 19 |
19 |
5 |
2 |
0 |
0 |
0 |
90.0 |
+----------- Cumberlandia_monodonta **
|
| +-- Margaritifera_falcata **
+--------+
| +-- Margaritifera_margaritifera **
|
--+ +-------- Gonidea_angulata **
| |
| | +-- Elliptio_complanata **
| | |
| | +-- Fusconaia_cerina **
| | |
+--+ +--+-- Plectomerus_dombeyanus **
| | |
| | +-- Pleurobema_cordatum **
| | |
| | +-- Unio_pictorum **
| |
| | +-- Pyganodon_cataracta **
+--+ |
| +-- Pyganodon_grandis **
| |
| +-- Utterbackia_imbecillis **
| |
| +-- Lampsilis_teres **
| |
| +-- Lampsilis_straminea **
+--+
+-- Megalonaias_nervosa **
|
+-- Obliquaria_reflexa **
|
+-- Theliderma_cylindrica **
|
+-- Quadrula_quadrula **
|
+-- Uniomerus_tetralasmus **
Araujo et al., 2018 [Publ 3779], fig. 1
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 19 |
19 |
14 |
2 |
0 |
0 |
0 |
81.0 |
+-------------------------- Pseudunio_auricularius **
|
| +----- Anodonta_anatina **
--+ +-----------------+
| | | +-- Pyganodon_grandis **
| | +--+
| | +-- Alasmidonta_marginata **
+--+
| +-------------------- Nodularia_douglasiae **
| |
| | +-- Unio_durieui **
+--+ +--------------+
| | +-- Unio_gibbus **
| |
+--+ +-------------- Unio_tumidus **
| |
| | +-------- Unio_tumidiformis **
| | +--+
+--+ | | +----- Unio_bruguierianus **
| | +--+
| | | +-- Unio_crassus **
| | +--+
+--+ +-- Unio_ionicus **
|
| +-- Unio_delphinus **
| +--+
| | +-- Unio_foucauldianus **
| |
+-----+ +-- Unio_pictorum **
| |
| +-- Unio_elongatulus **
| |
+--+-- Unio_mancus **
|
+-- Unio_ravoisieri **
|
+-- Unio_tigridis **
Smith et al., 2018 [Publ 3790], fig. 1
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 19 |
19 |
17 |
1 |
0 |
0 |
0 |
89.0 |
+-------------------- Cristaria_plicata **
|
| +-- Utterbackia_peninsularis **
| +--------------+
| | +-- Pyganodon_grandis **
--+ |
| | +-- Anodontoides_ferussacianus **
| | +--------+
| | | +-- Anodontoides_denigratus **
| | |
+--+ +--+ +----- Strophitus_pascagoulaensis **
| | | +--+
| | | | | +-- Strophitus_radiatus **
| | | | +--+
| | +--+ +-- Strophitus_williamsi **
| | |
| | | +----- Strophitus_howellsi[2] **
| | +--+
+--+ | +-- Strophitus_undulatus **
| +--+
| +-- Strophitus_howellsi[1] **
|
| +-- Alasmidonta_triangulata **
| +--------+
| | +-- Alasminota_etowaensis **
| |
+--+ +-- Pseudodontoideus_connasaugaensis **
| +-----+
| | +-- Pseudodontoideus_subvexus **
| |
+--+ +-- Utterbackiana_couperiana **
| +--+
| | +-- Utterbackiana_heardi **
+--+
| +-- Utterbackiana_hartfieldorum **
+--+
+-- Utterbackiana_suborbiculata **
Froufe et al., 2019 [Publ 3936], fig. 3-M
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 19 |
19 |
17 |
2 |
0 |
0 |
0 |
73.0 |
+-------------------- Pseudunio_marocanus **
|
| +----- Quadrula_quadrula **
| +-----------+
| | | +-- Potamilus_alatus **
--+ | +--+
| | +-- Venustaconcha_ellipsiformis **
| |
| | +----- Unio_tumidus **
| | +-----+
| | | | +-- Unio_crassus **
+--+ | +--+
| +--+ +-- Unio_delphinus **
| | |
| | | +-------- Sinanodonta_woodiana **
| | +--+
| | | +----- Anodonta_anatina **
| | +--+
| | | +-- Utterbackia_peninsularis **
+--+ +--+
| +-- Pyganodon_grandis **
|
| +-- Chamberlainia_somsakpanhai **
| +--------+
| | +-- Sinohyriopsis_cumingii **
| |
+--+ +-- Potomida_littoralis **
| +-----+
| | +-- Pronodularia_japanensis **
| |
+--+ +-- Microcondylaea_bonellii **
| +--+
| | +-- Sinosolenaia_carinata **
+--+
| +-- Pilsbryoconcha_exilis **
+--+
+-- Pseudodon_vondembuschianus **
Froufe et al., 2019 [Publ 3936], fig. 4
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 19 |
19 |
17 |
2 |
0 |
0 |
0 |
73.0 |
+-------------------- Pseudunio_marocanus **
|
| +----- Quadrula_quadrula **
| +-----------+
| | | +-- Potamilus_alatus **
--+ | +--+
| | +-- Venustaconcha_ellipsiformis **
| |
| | +----- Unio_tumidus **
| | +-----+
| | | | +-- Unio_crassus **
+--+ | +--+
| +--+ +-- Unio_delphinus **
| | |
| | | +-------- Sinanodonta_woodiana **
| | +--+
| | | +----- Anodonta_anatina **
| | +--+
| | | +-- Utterbackia_peninsularis **
+--+ +--+
| +-- Pyganodon_grandis **
|
| +-- Chamberlainia_somsakpanhai **
| +--------+
| | +-- Sinohyriopsis_cumingii **
| |
+--+ +-- Potomida_littoralis **
| +-----+
| | +-- Pronodularia_japanensis **
| |
+--+ +-- Microcondylaea_bonellii **
| +--+
| | +-- Sinosolenaia_carinata **
+--+
| +-- Pilsbryoconcha_exilis **
+--+
+-- Pseudodon_vondembuschianus **
Teiga-Teixeira et al., 2020 [Publ 4173], fig. 1
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 19 |
19 |
17 |
2 |
0 |
0 |
0 |
83.0 |
+-- Margaritifera_falcata **
+-----------------------------+
| +-- Cumberlandia_monodonta **
|
| +----- Lanceolaria_grayii **
--+ +-----------------------+
| | | +-- Chamberlainia_somsakpanhai **
| | +--+
| | +-- Lamprotula_leaii **
+--+
| +-- Uniomerus_tetralasmus **
| +-----------------------+
| | +-- Quadrula_quadrula **
| |
+--+ +-- Elliptio_complanata **
| +--------------------+
| | +-- Pleurobema_oviforme **
+--+
| +-------------------- Amblema_plicata **
| |
+--+ +----------------- Popenaias_popeii **
| |
+--+ +-------------- Toxolasma_parvum **
| |
| | +-- Potamilus_alatus **
+--+ +--------+
| | +-- Potamilus_leptodon **
| |
+--+ +-- Lampsilis_cardium **
| +-----+
| | +-- Lampsilis_ornata **
+--+
| +----- Venustaconcha_ellipsiformis **
+--+
| +-- Lampsilis_powellii **
+--+
+-- Lampsilis_siliquoidea **
Wu et al., 2023 [Publ 4339], fig. 5
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 19 |
19 |
17 |
2 |
0 |
0 |
0 |
81.0 |
+-- Margaritifera_margaritifera **
+-----------------+
| +-- Margaritifera_dahurica **
|
| +-- Elliptio_complanata **
| +--------+
| | +-- Quadrula_quadrula **
--+ +-----+
| | | +-------- Obliquaria_reflexa **
| | +--+
| | | +----- Truncilla_truncata **
| | +--+
| | | +-- Lampsilis_cardium **
| | +--+
+--+ +-- Potamilus_alatus **
|
| +-- Lepidodesma_languilati **
| +-----------+
| | +-- Lepidodesma_aligera **
| |
| | +-------- Anemina_arcaeformis **
+--+ +--+
| | | +----- Cristaria_plicata **
| | +--+
| | | +-- Sinanodonta_tumens **
| | +--+
+--+ +-- Sinanodonta_woodiana **
|
| +-- Platynaias **
| +-----+
| | +-- Platynaias_compressa **
+--+
| +----- Anodonta_anatina **
+--+
| +-- Pseudanodonta_complanata **
+--+
+-- Anodonta_cygnea **
Huang et al., 2018 [Publ 3828], fig. 3, 5
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 18 |
18 |
16 |
2 |
11 |
9 |
2 |
86.0 |
+----- Triplodon_corrugatus
+-----------------------+
| | +-- Velesunio_ambiguus
| +--+
| +-- Hyridella_australis
|
| +----------- Mutela_hargeri
| |
| +--------------+ +-- Aspatharia_pfeifferiana
--+ | | +-----+
| | | | +-- Chambardia_wahlbergi
| | +--+
| | | +----- Etheria_elliptica
| | +--+
| | | +-- Tamsiella_schroeteriana
| | +--+
| | +-- Lamproscapha_ensiformis
| |
+--+ +-- Potomida_littoralis **
| +-----+
| | +-- Lamprotula_leaii **
| |
| +--------------+ +-- Lampsilis_cardium **
| | | +--+
| | | | +-- Scabies_crispata **
| | +--+
| | | +-- Unio_pictorum **
| | +--+
+--+ +-- Aculamprotula_tientsinensis **
|
| +-- Gibbosula_rochechouartii **
| +-----------------+
| | +-- Gibbosula_laosensis **
| |
| | +----- Cumberlandia_monodonta **
+--+ +-----------+
| | | +-- Pseudunio_auricularius **
| | +--+
| | +-- Pseudunio_marocanus **
+--+
| +-- Margaritifera_margaritifera **
| +-----------+
| | +-- Margaritifera_dahurica **
+--+
| +----------- Margaritifera_falcata **
+--+
| +-------- Margaritifera_laevis **
+--+
| +----- Margaritifera_hembeli **
+--+
| +-- Margaritifera_middendorffi **
+--+
+-- Margaritifera_marrianae **
Bolotov et al., 2019 [Publ 3974], fig. 1
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 18 |
18 |
16 |
1 |
0 |
0 |
0 |
89.0 |
+----------- Yaukthwa_inlenensis **
+--------+
| | +-------- Yaukthwa_elongatula **
| +--+
| | +----- Yaukthwa_panhai **
| +--+
| | +-- Yaukthwa_nesemanni **
| +--+
| +-- Yaukthwa_rectangularis **
--+
| +-- Leoparreysia_whitteni **
| +-----+
| | +-- Leoparreysia_burmana **
| +--------+
| | | +----- Leoparreysia_olivacea **
| | +--+
| | | +-- Leoparreysia_tavoyensis **
| | +--+
+--+ +-- Leoparreysia_canefrii **
|
| +-- Nitia_teretiuscula **
| +-----------+
| | +-- Coelatura_aegyptiaca **
+--+
| +----------- Indochinella_pugio **
| |
+--+ +-- Radiatula_myitkyinae **
| +-----+
| | +-- Radiatula_chaudhurii **
+--+
| +----- Radiatula_chindwinensis **
+--+
| +-- Radiatula_mouhoti **
+--+
+-- Radiatula_myitthanensis **
Sano et al., 2022 [Publ 4311], fig. 3
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 18 |
18 |
16 |
1 |
0 |
0 |
0 |
89.0 |
+-------- Pletholophus_tenuis **
+--------------+
| | +----- Cristaria_plicata **
| +--+
| | +-- Buldowskia_kamiyai **
| +--+
| +-- Buldowskia_iwakawai **
--+
| +-------- Beringiana_fukuharai **
| +-----------+
| | | +----- Beringiana_gosannensis **
| | +--+
| | | +-- Beringiana_japonica **
| | +--+
+--+ +-- Beringiana_beringiana **
|
| +-- Sinanodonta_angula **
| +--------------+
| | +-- Sinanodonta_lucida **
| |
+--+ +----- Sinanodonta_schrenkii **
| +--------+
| | | +-- Sinanodonta_tumens **
| | +--+
+--+ +-- Sinanodonta_calipygos **
|
| +----------- Sinanodonta_woodiana[2] **
+--+
| +-------- Sinanodonta_jourdyi **
+--+
| +----- Sinanodonta_elliptica **
+--+
| +-- Sinanodonta_woodiana[1] **
+--+
+-- Sinanodonta_lauta **
Lydeard et al., 2000 [Publ 2654], fig. 2a
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 17 |
17 |
11 |
2 |
0 |
0 |
0 |
88.0 |
+-------------------- Anodonta_cygnea **
|
+-------------------- Cumberlandia_monodonta **
|
| +----- Lampsilis_teres **
+--------------+
| | +-- Obovaria_unicolor **
| +--+
| +-- Obovaria_olivaria **
|
--+ +-- Reginaia_rotulata[1] **
| |
+-----------------+-- Reginaia_rotulata[2] **
| |
| +-- Reginaia_ebenus **
|
| +----- Quadrula_nobilis **
| +-----------+
| | | +-- Pustulosa_infucata **
| | +--+
+--+ +-- Pustulosa_succissa **
|
| +-------------- Pleurobema_decisum **
+--+
| +----------- Fusconaia_flava **
+--+
| +-------- Fusconaia_burkei[1] **
+--+
| +----- Fusconaia_burkei[2] **
+--+
| +-- Fusconaia_burkei[3] **
+--+
+-- Fusconaia_escambia **
Lydeard et al., 2000 [Publ 2654], fig. 2b
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 17 |
17 |
12 |
2 |
0 |
0 |
0 |
88.0 |
+-------------------------- Anodonta_cygnea **
|
+-------------------------- Cumberlandia_monodonta **
|
| +----- Lampsilis_teres **
--+ +-----------------+
| | | +-- Obovaria_unicolor **
| | +--+
| | +-- Obovaria_olivaria **
| |
+--+ +-- Reginaia_rotulata[1] **
| |
| +-----------------+-- Reginaia_rotulata[2] **
| | |
| | +-- Reginaia_ebenus **
| |
+--+ +-- Quadrula_nobilis **
| |
| +--------------+-- Pustulosa_infucata **
| | |
+--+ +-- Pustulosa_succissa **
|
| +-------------- Pleurobema_decisum **
+--+
| +----------- Fusconaia_flava **
+--+
| +-------- Fusconaia_burkei[1] **
+--+
| +----- Fusconaia_burkei[2] **
+--+
| +-- Fusconaia_burkei[3] **
+--+
+-- Fusconaia_escambia **
Grobler et al., 2011 [Publ 3359], fig. 2
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 17 |
17 |
13 |
1 |
0 |
0 |
0 |
88.0 |
+----------------- Amblema_plicata **
|
| +----------- Lemiox_rimosus **
| +--+
--+ | | +-------- Medionidus_conradicus **
| | +--+
| | | +----- Dromus_dromas **
| | +--+
| | | +-- Cyprogenia_stegaria **
+--+ +--+
| +-- Cyprogenia_aberti **
|
| +----------- Potamilus_alatus **
| |
| | +----- Lampsilis_siliquoidea **
| | +--+
+--+ | | +-- Lampsilis_higginsii **
| | +--+
| | +-- Ortmanniana_ligamentina **
| |
| | +----- Lampsilis_fasciola **
+--+--+
| | +-- Lampsilis_cardium **
| +--+
| +-- Lampsilis_ovata **
|
| +----- Cambarunio_iris **
| |
+--+----- Ligumia_recta **
|
| +-- Epioblasma_brevidens **
+--+
+-- Epioblasma_capsaeformis **
Inoue et al., 2013 [Publ 3528], fig. 3
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 17 |
17 |
13 |
1 |
0 |
0 |
0 |
83.0 |
+-------------------------- Gonidea_angulata **
|
+-------------------------- Pyganodon_grandis **
|
--+ +-- Eurynia_dilatata **
| +--------------------+
| | +-- Quadrula_quadrula **
| |
| | +-------- Paetulunio_fabalis **
+--+ +-----------+
| | | +----- Amblema_plicata **
| | +--+
| | | +-- Reginaia_ebenus **
+--+ +--+
| +-- Reginaia_rotulata **
|
| +----------------- Lampsilis_ornata **
| |
+--+ +-------------- Leaunio_vanuxemensis **
| |
| | +-- Villosa_villosa **
+--+-----------+
| +-- Cambarunio_iris **
|
| +----------- Obovaria_olivaria **
+--+
| +-------- Obovaria_arkansasensis **
+--+
| +----- Obovaria_retusa **
+--+
| +-- Obovaria_unicolor **
+--+
+-- Obovaria_subrotunda **
Prié & Puillandre, 2014 [Publ 3529], fig. 2
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 17 |
17 |
15 |
2 |
0 |
0 |
0 |
81.0 |
+----------------------------- Pseudunio_auricularius **
|
--+ +-------------------------- Amblema_plicata **
| |
| | +----------------------- Potomida_littoralis **
+--+ |
| | +----- Anodonta_anatina **
| | +--------------+
+--+ | | +-- Pseudanodonta_complanata **
| | +--+
| | +-- Anodonta_cygnea **
| |
+--+ +----- Unio_tumidus **
| +-----------+
| | | +-- Unio_gibbus **
| | +--+
| | +-- Unio_durieui **
+--+
| +----- Unio_tumidiformis **
| +--------+
| | | +-- Unio_crassus **
| | +--+
+--+ +-- Unio_courtillieri **
|
| +----------- Unio_delphinus **
+--+
| +-------- Unio_elongatulus **
+--+
| +----- Unio_mancus **
+--+
| +-- Unio_pictorum **
+--+
+-- Unio_ravoisieri **
Plazzi et al., 2011 [Publ 3676], fig. 5
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 17 |
18 |
11 |
1 |
1 |
1 |
1 |
78.0 |
+----------------- Cristaria_plicata **
|
| +-------------- Lanceolaria_grayii **
| |
| | +-- Pyganodon_grandis **
| +-----------+
--+ | +-- Pseudanodonta_complanata **
| |
| | +-- Anodonta_anatina **
| +-----------+
| | +-- Anodonta_cygnea **
| |
| | +----- Unio_tumidus **
+--+--------+
| | +-- Unio_crassus **
| +--+
| +-- Unio_pictorum **
|
| +----- Alathyria_jacksoni
| |
| +-----+----- Pronodularia_japanensis **
| | |
| | | +-- Sinohyriopsis_cumingii **
| | +--+
+--+ +-- Sinohyriopsis_schlegelii **
|
| +-------- Quadrula_quadrula **
| |
+--+-------- Parvaspina_collina **
|
| +----- Lampsilis_ornata **
+--+
| +-- Epioblasma_rangiana **
+--+
+-- Venustaconcha_ellipsiformis **
Zhou et al., 2017 [Publ 3788], fig. 2
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 17 |
17 |
15 |
1 |
0 |
0 |
0 |
78.0 |
+-- Anemina_arcaeformis **
+-----------------+
| +-- Anemina_euscaphys **
|
| +-------- Lanceolaria_lanceolata **
| |
--+ +--------+ +-- Nodularia_douglasiae **
| | | +--+
| | | | +-- Tchangsinaia_piscicula **
| | +--+
| | | +-- Aculamprotula_tientsinensis **
| | +--+
+--+ +-- Aculamprotula_tortuosa **
|
| +----- Quadrula_quadrula **
| +--------+
| | | +-- Potamilus_leptodon **
| | +--+
| | +-- Lampsilis_ornata **
+--+
| +-- Sinohyriopsis_cumingii **
| +--------+
| | +-- Sinohyriopsis_schlegelii **
| |
+--+ +-- Lamprotula_caveata **
| +-----+
| | +-- Lamprotula_leaii **
+--+
| +----- Ptychorhynchus_pfisteri **
+--+
| +-- Sinosolenaia_carinata **
+--+
+-- Sinosolenaia_oleivora **
Amaro et al., 2017 [Publ 3791], fig. 2
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 17 |
17 |
15 |
2 |
1 |
0 |
0 |
85.0 |
+-------------------- Margaritifera_margaritifera[2] **
|
| +-- Margaritifera_margaritifera[1] **
| +--------------+
--+ | +-- Margaritifera_falcata **
| |
| | +-- Lasmigona_costata **
| | +-----+
| | | +-- Platynaias_compressa **
+--+ +-----+
| | | +----- Pyganodon_grandis **
| | +--+
| | | +-- Utterbackia_peninsularis **
| | +--+
| | +-- Utterbackia_imbecillis **
+--+
| +----- Fusconaia_flava **
| +-----+
| | | +-- Pronodularia_japanensis **
| | +--+
| | +-- Sinosolenaia_carinata **
+--+
| +----- Toxolasma_parvum **
| +--+
| | | +-- Venustaconcha_ellipsiformis **
| | +--+
+--+ +-- Lampsilis_siliquoidea **
|
| +----- Quadrula_quadrula[2] **
+--+
| +-- Quadrula_quadrula[1] **
+--+
+-- Quadrula_nobilis **
Lopes-Lima et al., 2018 [Publ 3829], fig. 1
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 17 |
17 |
15 |
2 |
11 |
9 |
2 |
87.0 |
+----- Triplodon_corrugatus
+--------------------+
| | +-- Velesunio_ambiguus
| +--+
| +-- Hyridella_australis
|
| +----------- Mutela_hargeri
| |
--+ +-----------+ +-- Aspatharia_pfeifferiana
| | | +-----+
| | | | +-- Chambardia_wahlbergi
| | +--+
| | | +----- Etheria_elliptica
| | +--+
| | | +-- Lamproscapha_ensiformis
| | +--+
+--+ +-- Tamsiella_schroeteriana
|
| +----- Potomida_littoralis **
| +--------------+
| | | +-- Lampsilis_cardium **
| | +--+
| | +-- Unio_pictorum **
| |
+--+ +----- Gibbosula_rochechouartii **
| +-----------+
| | | +-- Gibbosula_crassa **
| | +--+
| | +-- Gibbosula_laosensis **
| |
+--+ +-------- Cumberlandia_monodonta **
| +-----+
| | | +----- Pseudunio_marocanus **
| | +--+
| | | +-- Pseudunio_auricularius **
| | +--+
+--+ +-- Pseudunio_homsensis **
|
| +-- Margaritifera_dahurica **
| +--------+
| | +-- Margaritifera_margaritifera **
+--+
| +-------- Margaritifera_falcata **
| |
+--+ +-- Margaritifera_middendorffi **
| +--+
| | +-- Margaritifera_laevis **
+--+
| +-- Margaritifera_hembeli **
+--+
+-- Margaritifera_marrianae **
Lopes-Lima et al., 2018 [Publ 3829], fig. 3
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 17 |
17 |
15 |
2 |
11 |
9 |
2 |
87.0 |
+----- Triplodon_corrugatus
+-----------------------+
| | +-- Velesunio_ambiguus
| +--+
| +-- Hyridella_australis
|
| +----------- Mutela_hargeri
| |
--+ +--------------+ +-- Chambardia_wahlbergi
| | | +-----+
| | | | +-- Aspatharia_pfeifferiana
| | +--+
| | | +----- Etheria_elliptica
| | +--+
| | | +-- Tamsiella_schroeteriana
| | +--+
+--+ +-- Lamproscapha_ensiformis
|
| +----- Potomida_littoralis **
| +-----------------+
| | | +-- Lampsilis_cardium **
| | +--+
| | +-- Unio_pictorum **
| |
+--+ +----- Gibbosula_rochechouartii **
| +--------------+
| | | +-- Gibbosula_laosensis **
| | +--+
| | +-- Gibbosula_crassa **
| |
+--+ +-------- Cumberlandia_monodonta **
| +--------+
| | | +----- Pseudunio_marocanus **
| | +--+
| | | +-- Pseudunio_auricularius **
| | +--+
+--+ +-- Pseudunio_homsensis **
|
| +-- Margaritifera_margaritifera **
| +-----------+
| | +-- Margaritifera_dahurica **
+--+
| +----------- Margaritifera_falcata **
+--+
| +-------- Margaritifera_laevis **
+--+
| +----- Margaritifera_middendorffi **
+--+
| +-- Margaritifera_hembeli **
+--+
+-- Margaritifera_marrianae **
Liu et al., 2017 [Publ 3939], fig. 2
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 17 |
17 |
12 |
1 |
0 |
0 |
0 |
75.0 |
+----- Acuticosta_chinensis **
+-----------------+
| | +-- Anemina_arcaeformis **
| +--+
| +-- Sinosolenaia_oleivora **
--+
| +-------------------- Lanceolaria_grayii **
| |
| | +----- Unio_tumidus **
| | |
+--+ +-----------+----- Unio_gibbus **
| | |
| | | +-- Unio_crassus **
| | +--+
| | +-- Unio_pictorum **
+--+
| +-------------- Schistodesmus_lampreyanus **
| |
| | +-- Cuneopsis_heudei **
| +-----------+
+--+ +-- Cuneopsis_rufescens **
|
| +----------- Nodularia_breviconcha **
| |
+--+ +-------- Nodularia_douglasiae[3] **
| |
+--+ +----- Nodularia_douglasiae[2] **
| |
+--+ +-- Nodularia_douglasiae[1] **
| |
+--+-- Nodularia_nuxpersicae **
|
+-- Nodularia_nipponensis **
Wu et al., 2022 [Publ 4238], fig. 2e
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 17 |
17 |
15 |
1 |
0 |
0 |
0 |
80.0 |
+-------- Sinanodonta_woodiana **
+-----------------+
| | +----- Anodonta_anatina **
| +--+
| | +-- Utterbackia_peninsularis **
--+ +--+
| +-- Pyganodon_grandis **
|
| +----------------------- Lanceolaria_lanceolata **
| |
+--+ +-------------------- Aculamprotula_tortuosa **
| |
| | +----- Unio_tumidus **
+--+ +-----------+
| | | +-- Unio_crassus **
| | +--+
| | +-- Unio_delphinus **
+--+
| +-- Pseudocuneopsis_sichuanensis **
| +-----------+
| | +-- Pseudocuneopsis_capitata **
+--+
| +----------- Nodularia_douglasiae **
| |
+--+ +-------- Tchangsinaia_piscicula **
| |
+--+ +-- Cuneopsis_celtiformis **
| +--+
| | +-- Cuneopsis_heudei **
+--+
| +-- Schistodesmus_lampreyanus **
+--+
+-- Schistodesmus_spinosus **
Smith, 2021 [Publ 4331], fig. 1
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 17 |
17 |
15 |
2 |
3 |
2 |
1 |
85.0 |
+-- Etheria_elliptica
+-----------------------+
| +-- Aspatharia_pfeifferiana
|
| +----------------------- Margaritifera_margaritifera **
--+ |
| | +----- Sinohyriopsis_cumingii **
| | +-----+
| | | | +-- Lens_contradens **
| | | +--+
+--+ +--------+ +-- Lamprotula_leaii **
| | |
| | | +-------- Scabies_phaselus **
| | +--+
| | | +----- Cristaria_plicata **
| | +--+
| | | +-- Pyganodon_grandis **
+--+ +--+
| +-- Utterbackia_peninsularis **
|
| +-- Uniomerus_tetralasmus **
| +--------------+
| | +-- Megalonaias_nervosa **
| |
+--+ +-- Elliptio_crassidens **
| +-----------+
| | +-- Elliptio_complanata **
+--+
| +----------- Amblema_plicata **
+--+
| +-------- Potamilus_streckersoni **
+--+
| +----- Venustaconcha_ellipsiformis **
+--+
| +-- Lampsilis_siliquoidea **
+--+
+-- Lampsilis_cardium **
Lydeard et al., 2000 [Publ 2654], fig. 2c
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 16 |
16 |
9 |
2 |
0 |
0 |
0 |
89.0 |
+-------------- Anodonta_cygnea **
|
+-------------- Cumberlandia_monodonta **
|
| +----- Quadrula_nobilis **
+--------+
| | +-- Pustulosa_infucata **
| +--+
| +-- Pustulosa_succissa **
--+
| +----- Obovaria_unicolor **
+--------+
| | +-- Obovaria_olivaria **
| +--+
| +-- Lampsilis_teres **
|
| +-- Reginaia_ebenus **
| +--------+
| | +-- Reginaia_rotulata **
+--+
| +-------- Pleurobema_decisum **
| |
+--+ +----- Fusconaia_flava **
| |
+--+ +-- Fusconaia_burkei[1] **
| |
| +-- Fusconaia_burkei[2] **
+--+
+-- Fusconaia_burkei[3] **
|
+-- Fusconaia_escambia **
Lydeard et al., 2000 [Publ 2654], fig. 2d
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 16 |
16 |
12 |
2 |
0 |
0 |
0 |
87.0 |
+-------------------------- Anodonta_cygnea **
|
+-------------------------- Cumberlandia_monodonta **
|
--+ +-- Reginaia_ebenus **
| +--------------------+
| | +-- Reginaia_rotulata **
| |
| | +----- Lampsilis_teres **
+--+ +--------------+
| | | +-- Obovaria_unicolor **
| | +--+
| | +-- Obovaria_olivaria **
+--+
| +-- Quadrula_nobilis **
| |
| +--------------+-- Pustulosa_infucata **
| | |
+--+ +-- Pustulosa_succissa **
|
| +-------------- Pleurobema_decisum **
+--+
| +----------- Fusconaia_flava **
+--+
| +-------- Fusconaia_burkei[1] **
+--+
| +----- Fusconaia_burkei[2] **
+--+
| +-- Fusconaia_burkei[3] **
+--+
+-- Fusconaia_escambia **
Walker et al., 2006 [Publ 2666], fig. 6
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 16 |
16 |
13 |
2 |
22 |
21 |
2 |
83.0 |
+----------------- Castalia_stevensi
|
+----------------- Diplodon_gratus
+--------+
| | +-------------- Hyridella_depressa
| | |
| | +-------------- Hyridella_australis
| +--+
| | +----------- Echyridella_menziesii
| | |
| | | +-- Velesunio_ambiguus
| +--+ +-----+
| | | +-- Velesunio[3]
| | |
| +--+ +----- Velesunio[2]
| | |
| | | +-- Alathyria_jacksoni
| +--+--+
| | +-- Velesunio[1]
| |
--+ | +-- Velesunio_angasi
| +--+
| +-- Lortiella_rugata
|
| +-- Mutela_dubia
| +--------------+
| | +-- Mutela_rostrata
| +-----+
| | | +-------------- Etheria_elliptica
| | | |
| | +--+ +----------- Chambardia_rubens
| | | |
| | +--+ +-------- Mulleria_rivoli
| | | |
| | +--+ +-- Anodontites_guanarensis
| | | +--+
| | | | +-- Anodontites_trigona
+--+ +--+
| | +-- Tamsiella_tamsiana
| +--+
| +-- Monocondylaea_minuana
|
| +-- Pyganodon_fragilis **
| +-----------------+
| | +-- Pyganodon_grandis **
| |
| | +----------- Cumberlandia_monodonta **
| | |
+--+ +-----+ +-------- Pseudunio_auricularius **
| | | |
| | +--+ +-- Margaritifera_margaritifera **
| | | +--+
| | | | +-- Margaritifera_dahurica **
| | +--+
+--+ | +-- Margaritifera_falcata **
| +--+
| +-- Margaritifera_middendorffi **
|
| +-- Gonidea_angulata **
| +-----------+
| | +-- Sundadontina_tumida **
+--+
| +----------- Cyrtonaias_tampicoensis **
| |
+--+----------- Fusconaia_flava **
|
| +-------- Potamilus_purpuratus **
+--+
| +----- Lampsilis_teres **
+--+
| +-- Ligumia_recta **
+--+
+-- Ortmanniana_ligamentina **
Roe et al., 2001 [Publ 2689], fig. 3a
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 16 |
16 |
8 |
1 |
0 |
0 |
0 |
87.0 |
+-------------- Ligumia_recta **
|
+-------------- Obliquaria_reflexa **
|
| +----- Lampsilis_teres **
+--------+
--+ | +-- Lampsilis_ovata **
| +--+
| +-- Lampsilis_ornata **
|
| +----------- Villosa_villosa **
| |
| | +----- Hamiota_australis[2] **
+--+ +--+
| | | +-- Hamiota_subangulata **
| | +--+
| | +-- Hamiota_australis[1] **
+--+
| +----- Hamiota_altilis[3] **
| |
| | +-- Hamiota_perovalis[1] **
+--+ |
| +-- Hamiota_perovalis[2] **
| |
| +-- Hamiota_altilis[1] **
+--+
+-- Hamiota_altilis[2] **
|
+-- Hamiota_altilis[4] **
|
+-- Hamiota_altilis[5] **
Zanatta et al., 2007 [Publ 2698], fig. 1A
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 16 |
16 |
10 |
1 |
0 |
0 |
0 |
90.0 |
+-- Platynaias_compressa **
|
+-----------------+-- Lasmigona_costata **
| |
| +-- Lasmigona_complanata **
|
--+ +----------------- Anodonta_cygnea **
| |
| | +----------- Anodonta_kennerlyi **
| | +--+
+--+ | | +-------- Utterbackiana_implicata **
| | +--+
| | | +----- Utterbackiana_suborbiculata **
| | +--+
| | | +-- Utterbackiana_heardi **
+--+ +--+
| +-- Utterbackiana_couperiana **
|
| +-- Utterbackia_peggyae **
| +--+
| | +-- Utterbackia_imbecillis **
| |
+--------+ +-- Pyganodon_gibbosa **
| |
| +-- Pyganodon_cataracta **
| |
+--+-- Pyganodon_fragilis **
|
+-- Pyganodon_grandis **
|
+-- Pyganodon_lacustris **
Zanatta et al., 2007 [Publ 2698], fig. 1B
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 16 |
16 |
13 |
1 |
0 |
0 |
0 |
90.0 |
+-- Lasmigona_costata **
|
+-----------------+-- Platynaias_compressa **
| |
| +-- Lasmigona_complanata **
|
| +-------------- Anodonta_cygnea **
| +--+
--+ | | +----------- Anodonta_kennerlyi **
| | +--+
| | | +-------- Utterbackiana_implicata **
| | +--+
| | | +----- Utterbackiana_heardi **
| | +--+
+--+ | +-- Utterbackiana_suborbiculata **
| +--+
| +-- Utterbackiana_couperiana **
|
| +-- Utterbackia_peggyae **
| +-----------+
| | +-- Utterbackia_imbecillis **
+--+
| +----------- Pyganodon_lacustris **
+--+
| +-------- Pyganodon_grandis **
+--+
| +----- Pyganodon_fragilis **
+--+
| +-- Pyganodon_gibbosa **
+--+
+-- Pyganodon_cataracta **
Hoeh, 1990 [Publ 2717], fig. 2
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 16 |
16 |
13 |
1 |
0 |
0 |
0 |
89.0 |
+-- Lasmigona_complanata **
|
+-----------------+-- Platynaias_compressa **
| |
| +-- Lasmigona_costata **
|
| +-------------- Anodonta_cygnea **
| +--+
--+ | | +----------- Anodonta_kennerlyi **
| | +--+
| | | +-------- Utterbackiana_implicata **
| | +--+
| | | +----- Utterbackiana_couperiana[2] **
| | +--+
+--+ | +-- Utterbackiana_suborbiculata **
| +--+
| +-- Utterbackiana_couperiana[1] **
|
| +-- Utterbackia_imbecillis **
| +-----------+
| | +-- Utterbackia_peggyae **
+--+
| +----------- Pyganodon_lacustris **
+--+
| +-------- Pyganodon_grandis **
+--+
| +----- Pyganodon_fragilis **
+--+
| +-- Pyganodon_cataracta **
+--+
+-- Pyganodon_gibbosa **
Hoeh, 1990 [Publ 2717], fig. 3
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 16 |
16 |
2 |
1 |
0 |
0 |
0 |
88.0 |
+----- Pyganodon_fragilis **
|
+----- Utterbackia_imbecillis **
|
+----- Utterbackia_peggyae **
|
+----- Anodonta_cygnea **
|
+----- Anodonta_kennerlyi **
|
+----- Utterbackiana_couperiana[1] **
|
--+----- Utterbackiana_couperiana[2] **
|
+----- Utterbackiana_suborbiculata **
|
| +-- Lasmigona_costata **
+--+
| +-- Utterbackiana_implicata **
|
| +-- Lasmigona_complanata **
| |
| +-- Platynaias_compressa **
| |
| +-- Pyganodon_cataracta **
+--+
+-- Pyganodon_gibbosa **
|
+-- Pyganodon_grandis **
|
+-- Pyganodon_lacustris **
Hoeh, 1990 [Publ 2717], fig. 4
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 16 |
16 |
13 |
1 |
0 |
0 |
0 |
89.0 |
+-- Lasmigona_complanata **
|
+-----------------+-- Platynaias_compressa **
| |
| +-- Lasmigona_costata **
|
| +-------------- Anodonta_cygnea **
| +--+
--+ | | +----------- Anodonta_kennerlyi **
| | +--+
| | | +-------- Utterbackiana_implicata **
| | +--+
| | | +----- Utterbackiana_couperiana[2] **
| | +--+
+--+ | +-- Utterbackiana_suborbiculata **
| +--+
| +-- Utterbackiana_couperiana[1] **
|
| +-- Utterbackia_imbecillis **
| +--------+
| | +-- Utterbackia_peggyae **
+-----+
| +-------- Pyganodon_fragilis **
| |
+--+ +-- Pyganodon_cataracta **
| +--+
| | +-- Pyganodon_gibbosa **
+--+
| +-- Pyganodon_lacustris **
+--+
+-- Pyganodon_grandis **
Turner et al., 2000 [Publ 2722], fig. 2
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 16 |
16 |
12 |
2 |
0 |
0 |
0 |
87.0 |
+----------------------------- Margaritifera_margaritifera **
|
| +-- Quadrula_quadrula **
--+ +-----------------------+
| | +-- Utterbackiana_couperiana **
| |
+--+ +-- Amblema_plicata **
| +--------------------+
| | +-- Eurynia_dilatata **
+--+
| +-------------------- Medionidus_conradicus **
| |
+--+ +----------------- Lampsilis_teres **
| |
| +----------------- Villosa_delumbis **
+--+
| +-- Lampsilis_satura **
| +-----------+
| | +-- Lampsilis_cardium **
+--+
| +----------- Lampsilis_reeveiana **
| |
+--+ +-------- Lampsilis_hydiana[1] **
| |
+--+-------- Lampsilis_hydiana[2] **
|
| +----- Lampsilis_siliquoidea[2] **
+--+
| +-- Lampsilis_siliquoidea[1] **
+--+
+-- Lampsilis_powellii **
Szumowski et al., 2012 [Publ 3569], fig. 4
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 16 |
16 |
12 |
1 |
0 |
0 |
0 |
87.0 |
+-------------------------- Amblema_plicata **
|
--+ +----------------------- Ellipsaria_lineolata **
| |
+--+ +-------------------- Truncilla_truncata **
| |
+--+ +----------------- Obliquaria_reflexa **
| |
| | +-------------- Potamilus_alatus **
+--+ |
| | +----- Lampsilis_ornata **
| | +-----+
| | | | +-- Lampsilis_cardium **
+--+ | +--+
| | +-- Lampsilis_ovata **
| |
| | +-------- Lampsilis_teres **
| +--+
+--+ | +----- Lampsilis_siliquoidea **
| +--+
| | +-- Ortmanniana_ligamentina **
| +--+
| +-- Lampsilis_higginsii **
|
| +----- Hamiota_altilis **
| |
+-----+ +-- Epioblasma_triquetra **
| |
+--+-- Cambarunio_iris **
|
+-- Ligumia_recta **
Huang et al., 2013 [Publ 3629], fig. 3
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 16 |
16 |
14 |
2 |
0 |
0 |
0 |
79.0 |
+-------------------- Margaritifera_falcata **
|
| +-------------- Unio_pictorum **
| |
| +--+ +-- Sinanodonta_woodiana **
| | | +--------+
--+ | | | +-- Cristaria_plicata **
| | +--+
| | | +-------- Platynaias_compressa **
| | +--+
| | | +----- Pyganodon_grandis **
| | +--+
| | | +-- Utterbackia_imbecillis **
+--+ +--+
| +-- Utterbackia_peninsularis **
|
| +-------- Quadrula_quadrula **
| +--+
| | | +----- Toxolasma_parvum **
| | +--+
| | | +-- Lampsilis_ornata **
| | +--+
+-----+ +-- Venustaconcha_ellipsiformis **
|
| +-- Sinosolenaia_carinata **
| +--+
| | +-- Pronodularia_japanensis **
+-----+
| +-- Sinohyriopsis_cumingii **
+--+
+-- Sinohyriopsis_schlegelii **
Xue et al., 2016 [Publ 3659], fig. 1
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 16 |
16 |
14 |
1 |
0 |
0 |
0 |
78.0 |
+-- Sinohyriopsis_cumingii **
+--+
| +-- Lamprotula_leaii **
+-----------+
| | +-- Lampsilis_ornata **
| +--+
| +-- Venustaconcha_ellipsiformis **
|
| +-- Cristaria_plicata **
| +-----+
--+ | +-- Platynaias_compressa **
| |
| +-----+ +-- Anemina_arcaeformis **
| | | +--+
| | | | +-- Anemina_euscaphys **
| | +--+
| | | +-- Sinanodonta_lucida **
+--+ +--+
| +-- Sinanodonta_woodiana **
|
| +-- Lanceolaria_lanceolata **
| +--------+
| | +-- Lanceolaria_grayii **
+--+
| +-------- Aculamprotula_tortuosa **
+--+
| +----- Schistodesmus_lampreyanus **
+--+
| +-- Nodularia_douglasiae **
+--+
+-- Unio_pictorum **
Lopes-Lima et al., 2017 [Publ 3734], fig. 1
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 16 |
16 |
12 |
2 |
1 |
0 |
0 |
77.0 |
+-- Pseudunio_auricularius **
+-----------------+
| +-- Margaritifera_margaritifera **
|
| +-- Potomida_littoralis **
--+ +--------------+
| | +-- Microcondylaea_bonellii **
| |
| | +----- Anodonta_anatina **
+--+ +--------+
| | | +-- Pseudanodonta_complanata **
| | +--+
| | +-- Anodonta_cygnea **
+--+
| +-- Unio_tumidus **
| +--------+
| | +-- Unio_gibbus **
| |
+--+ +-- Unio_crassus **
| +-----+
| | +-- Unio_tumidiformis **
+--+
| +----- Unio_ravoisieri **
| |
+--+ +-- Unio_mancus **
| |
| +-- Unio_pictorum **
+--+
+-- Unio_delphinus **
|
+-- Unio_elongatulus **
Wen et al., 2017 [Publ 3795], fig. 3
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 16 |
16 |
14 |
2 |
0 |
0 |
0 |
81.0 |
+-- Margaritifera_dahurica **
+--------------+
| +-- Margaritifera_falcata **
|
| +----------- Quadrula_quadrula **
| |
--+ +--+ +-------- Toxolasma_parvum **
| | | |
| | +--+ +-- Lampsilis_ornata **
| | | +--+
| | | | +-- Venustaconcha_ellipsiformis **
| | +--+
+--+ | +-- Potamilus_leptodon **
| +--+
| +-- Potamilus_alatus **
|
| +----------- Lamprotula_leaii **
| |
| | +----- Anodonta_anatina **
+--+ +--+
| | | +-- Sinanodonta_woodiana **
| | +--+
| | +-- Cristaria_plicata **
+--+
| +-- Aculamprotula_coreana **
| +--+
| | +-- Aculamprotula_tortuosa **
+--+
| +-- Lepidodesma_languilati **
+--+
+-- Unio_pictorum **
Huang et al., 2018 [Publ 3828], fig. 6-M
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 16 |
16 |
14 |
2 |
1 |
1 |
1 |
74.0 |
+----------------------- Echyridella_menziesii
|
| +-------------------- Cumberlandia_monodonta **
--+ |
| | +----- Quadrula_quadrula **
| | +-----------+
+--+ | | +-- Venustaconcha_ellipsiformis **
| | +--+
| | +-- Potamilus_alatus **
| |
| | +-------- Sinosolenaia_carinata **
+--+ +-----+
| | | +----- Potomida_littoralis **
| | +--+
| | | +-- Lamprotula_leaii **
| | +--+
| | +-- Pronodularia_japanensis **
+--+
| +-------- Sinanodonta_woodiana **
| +--+
| | | +----- Anodonta_anatina **
| | +--+
| | | +-- Pyganodon_grandis **
+--+ +--+
| +-- Utterbackia_peninsularis **
|
| +-------- Aculamprotula_tortuosa **
+--+
| +----- Unio_tumidus **
+--+
| +-- Unio_crassus **
+--+
+-- Unio_delphinus **
Johnson et al., 2018 [Publ 3844], fig. 2
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 16 |
16 |
14 |
1 |
0 |
0 |
0 |
87.0 |
+-------------------------------- Amblema_plicata **
|
--+ +----------------------------- Elliptio_crassidens **
| |
+--+ +-------------------------- Megalonaias_nervosa **
| |
+--+ +----------------------- Uniomerus_tetralasmus **
| |
+--+ +-------------------- Theliderma_metanevra **
| |
| | +-- Quadrula_quadrula **
+--+ +--------------+
| | +-- Quadrula_verrucosa **
| |
| | +----- Pustulosa_kieneriana **
+--+ +--------+
| | | +-- Pustulosa_infucata **
| | +--+
| | +-- Pustulosa_kleiniana **
+--+
| +-- Pustulosa_pustulosa **
| +--------+
| | +-- Pustulosa_succissa **
+--+
| +-------- Cyclonaias_tuberculata **
+--+
| +----- Pustulosa_necki **
+--+
| +-- Pustulosa_nodulata **
+--+
+-- Pustulosa_petrina **
Inoue et al., 2020 [Publ 3941], fig. 5
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 16 |
16 |
14 |
1 |
0 |
0 |
0 |
92.0 |
+-------------------- Truncilla_macrodon **
|
| +-------------- Lampsilis_bracteata **
| |
| +--+ +-- Lampsilis_ornata **
--+ | | +--------+
| | | | +-- Lampsilis_fasciola **
| | +--+
| | | +-------- Lampsilis_ovata **
| | +--+
| | | +----- Lampsilis_cariosa **
+--+ +--+
| | +-- Lampsilis_cardium **
| +--+
| +-- Lampsilis_satura **
|
| +-------------- Lampsilis_higginsii **
| |
+--+ +----------- Lampsilis_teres **
| |
| | +-- Lampsilis_radiata **
+--+ +-----+
| | +-- Lampsilis_siliquoidea **
| |
+--+ +-- Lampsilis_hydiana **
| +--+
| | +-- Lampsilis_bergmanni **
+--+
| +-- Lampsilis_straminea **
+--+
+-- Lampsilis_virescens **
Hewitt et al., 2019 [Publ 3962], fig. 2
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 16 |
16 |
11 |
2 |
0 |
0 |
0 |
82.0 |
+----------------------------- Margaritifera_margaritifera **
|
--+ +-------------------------- Pyganodon_grandis **
| |
+--+ +----------------------- Fusconaia_flava **
| |
+--+ +-------------------- Lampsilis_siliquoidea **
| |
| | +----------------- Lampsilis_fasciola **
+--+ |
| | +-- Epioblasma_rangiana **
| +--------------+
+--+ +-- Lampsilis_radiata **
|
| +-------------- Lampsilis_ornata **
| |
+--+ +-- Lampsilis_cardium[1] **
| +--------+
| | +-- Lampsilis_ovata[1] **
+--+
| +-------- Lampsilis_cardium[6] **
| |
+--+ +----- Lampsilis_cardium[5] **
| |
+--+ +-- Lampsilis_cardium[2] **
| |
| +-- Lampsilis_ovata[2] **
+--+
+-- Lampsilis_cardium[3] **
|
+-- Lampsilis_cardium[4] **
Lyubas et al., 2019 [Publ 3963], fig. 10
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 16 |
16 |
12 |
1 |
0 |
0 |
0 |
90.0 |
+----- Gibbosula_rochechouartii **
+--------------+
| | +-- Gibbosula_laosensis **
| +--+
| +-- Gibbosula_crassa **
|
| +-------- Cumberlandia_monodonta **
--+ |
| +--------+ +----- Pseudunio_marocanus **
| | | |
| | +--+ +-- Pseudunio_auricularius **
| | | |
| | | +-- Pseudunio_flabellata **
| | +--+
+--+ +-- Pseudunio_flabellatiformis **
| |
| +-- Pseudunio_homsensis **
|
| +-- Margaritifera_margaritifera **
| +-----------+
| | +-- Margaritifera_dahurica **
+--+
| +----------- Margaritifera_falcata **
+--+
| +-------- Margaritifera_laevis **
+--+
| +----- Margaritifera_middendorffi **
+--+
| +-- Margaritifera_marrianae **
+--+
+-- Margaritifera_hembeli **
Porto-Hannes et al., 2021 [Publ 4227], fig. 3
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 16 |
16 |
12 |
1 |
0 |
0 |
0 |
83.0 |
+-------------------- Quadrula_quadrula **
|
+-------------------- Quadrula_verrucosa **
|
--+ +-- Eurynia_dilatata **
| +--------------+
| | +-- Pleurobema_sintoxia **
| |
| | +-- Amblema_plicata **
+--+ +-----------+
| | +-- Amblema_elliottii **
| |
| | +-- Obovaria_olivaria **
+--+ +--------+
| | +-- Ortmanniana_ligamentina **
| |
| | +-- Obliquaria_reflexa **
| +--------+
+--+ +-- Lampsilis_ornata **
|
| +----- Lampsilis_radiata[2] **
| +--+
| | | +-- Lampsilis_siliquoidea **
| | +--+
+--+ +-- Lampsilis_radiata[1] **
|
| +----- Lampsilis_virescens **
+--+
| +-- Lampsilis_hydiana **
+--+
+-- Lampsilis_straminea **
Szumowski et al., 2012 [Publ 3569], fig. 5
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 15 |
15 |
12 |
1 |
0 |
0 |
0 |
89.0 |
+----------------- Megalonaias_nervosa **
|
| +-------- Theliderma_cylindrica **
+--------+
| | +----- Theliderma_intermedia **
| +--+
| | +-- Theliderma_metanevra **
--+ +--+
| +-- Theliderma_sparsa **
|
| +-------- Quadrula_quadrula[1] **
| +-----+
| | | +----- Quadrula_verrucosa **
| | +--+
| | | +-- Quadrula_nobilis **
+--+ +--+
| +-- Quadrula_quadrula[2] **
|
| +----------- Pustulosa_kieneriana **
| |
+--+ +-------- Pustulosa_infucata **
| |
+--+ +-- Pustulosa_pustulosa **
| +--+
| | +-- Pustulosa_succissa **
+--+
| +-- Pustulosa_nodulata **
+--+
+-- Pustulosa_petrina **
Reis et al., 2013 [Publ 3580], fig. 5
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 15 |
15 |
12 |
2 |
1 |
0 |
0 |
77.0 |
+-------------------------- Margaritifera_margaritifera **
|
| +-- Amblema_plicata **
--+ +--------------------+
| | +-- Theliderma_metanevra **
| |
+--+ +-------------------- Potomida_littoralis **
| |
| | +-- Anodonta_anatina **
+--+ |
| +--------------+-- Anodonta_cygnea **
| | |
| | +-- Pseudanodonta_complanata **
+--+
| +-------------- Unio_tumidus **
| |
+--+ +-- Unio_tumidiformis **
| +--------+
| | +-- Unio_crassus **
+--+
| +-------- Unio **
| |
+--+ +-- Unio_delphinus **
| +--+
| | +-- Unio_mancus **
+--+
| +-- Unio_pictorum **
+--+
+-- Unio_caffer **
Feng et al., 2016 [Publ 3800], fig. 2
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 15 |
15 |
13 |
2 |
0 |
0 |
0 |
81.0 |
+-- Margaritifera_dahurica **
+-----------------+
| +-- Margaritifera_falcata **
|
| +-- Anodonta_anatina **
--+ +--------------+
| | +-- Utterbackia_peninsularis **
| |
| | +-- Sinosolenaia_carinata **
| | +-----+
+--+ | +-- Sinosolenaia_oleivora **
| +-----+
| | | +----- Quadrula_quadrula **
| | +--+
| | | +-- Potamilus_leptodon **
+--+ +--+
| +-- Lampsilis_ornata **
|
| +----------- Tchangsinaia_piscicula **
| |
+--+ +-------- Pyganodon_grandis **
| |
+--+ +-- Anemina_arcaeformis **
| +--+
| | +-- Anemina_euscaphys **
+--+
| +-- Sinanodonta_lucida **
+--+
+-- Sinanodonta_woodiana **
Lopes-Lima et al., 2020 [Publ 4080], fig. 17
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 15 |
15 |
5 |
2 |
0 |
0 |
0 |
80.0 |
+-- Margaritifera_laevis **
+-----------+
| +-- Margaritifera_middendorffi **
|
--+ +----------- Chamberlainia_somsakpanhai **
| |
| | +-------- Sinohyriopsis_cumingii[10] **
+--+ |
| | +----- Sinohyriopsis_cumingii[1] **
| | |
| | +----- Sinohyriopsis_cumingii[2] **
+--+ |
| +----- Sinohyriopsis_cumingii[3] **
| |
| +----- Sinohyriopsis_cumingii[4] **
| |
| +----- Sinohyriopsis_cumingii[5] **
+--+
+----- Sinohyriopsis_cumingii[6] **
|
+----- Sinohyriopsis_cumingii[7] **
|
+----- Sinohyriopsis_cumingii[8] **
|
+----- Sinohyriopsis_cumingii[9] **
|
| +-- Sinohyriopsis_schlegelii **
+--+
+-- Sinohyriopsis_goliath **
Kim et al., 2021 [Publ 4197], fig. 1
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 15 |
15 |
13 |
1 |
0 |
0 |
0 |
77.0 |
+-- Sinohyriopsis_schlegelii **
+-----------------+
| +-- Lepidodesma_languilati **
|
--+ +----------------- Aculamprotula_tientsinensis **
| |
| | +-------- Acuticosta_chinensis **
| | +-----+
+--+ | | +----- Lanceolaria_gladiola **
| | +--+
| | | +-- Lanceolaria_grayii **
| | +--+
| | +-- Lanceolaria_lanceolata **
+--+
| +-------- Unio_tumidus **
| +--+
| | | +----- Unio_crassus **
| | +--+
| | | +-- Unio_delphinus **
| | +--+
+--+ +-- Unio_pictorum **
|
| +-- Nodularia_douglasiae **
| +--+
| | +-- Nodularia_breviconcha **
+-----+
| +-- Schistodesmus_lampreyanus **
+--+
+-- Cuneopsis_heudei **
Bolotov et al., 2022 [Publ 4250], fig. S2
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 15 |
15 |
13 |
1 |
0 |
0 |
0 |
89.0 |
+----- Parreysia_rakhinensis **
+--------------+
| | +-- Leoparreysia_tavoyensis **
| +--+
| +-- Leoparreysia_olivacea **
|
--+ +----------- Indonaia_rugosa **
| +-----+
| | | +-------- Indonaia_shurtleffiana **
| | +--+
| | | +----- Indonaia_andersoniana **
| | +--+
+--+ | +-- Indonaia_subclathrata **
| +--+
| +-- Indonaia_caerulea **
|
| +-------------- Indochinella_pugio **
| |
+--+ +----------- Radiatula_crispisulcata **
| |
+--+ +-- Radiatula_chaudhurii **
| +-----+
| | +-- Radiatula_myitkyinae **
+--+
| +----- Radiatula_chindwinensis **
+--+
| +-- Radiatula_mouhoti **
+--+
+-- Radiatula_myitthanensis **
Bolotov et al., 2022 [Publ 4250], fig. S5
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 15 |
15 |
12 |
1 |
0 |
0 |
0 |
91.0 |
+-- Physunio_modelli **
+-----+
| +-- Physunio_superbus **
+--------+
| | +----- Trapezoideus_foliaceus **
| +--+
| | +-- Trapezoideus_lenya **
| +--+
| +-- Trapezoideus_mitanensis **
--+
| +-- Yaukthwa_elongatula **
| +-----------+
| | +-- Yaukthwa_baniensis **
| |
| | +-------- Yaukthwa_panhai **
+--+ +--+
| | | +----- Yaukthwa_rectangularis **
| | +--+
| | | +-- Yaukthwa_nesemanni **
+--+ +--+
| +-- Yaukthwa_zayleymanensis **
|
| +----- Yaukthwa_dalliana **
| |
+-----+ +-- Yaukthwa_paiensis **
| |
+--+-- Yaukthwa_inlenensis **
|
+-- Yaukthwa_avaensis **
Mejia-Trujillo & Smith, 2022 [Publ 4274], fig. 2-F
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 15 |
15 |
13 |
1 |
0 |
0 |
0 |
86.0 |
+-------------- Toxolasma_parvum **
|
+--+ +----------- Popenaias_popeii **
| | |
| +--+ +-------- Amblema_plicata **
| | |
| +--+ +-- Quadrula_quadrula **
| | +--+
| | | +-- Uniomerus_tetralasmus **
| +--+
--+ | +-- Pleurobema_oviforme **
| +--+
| +-- Elliptio_complanata **
|
| +----- Potamilus_leptodon **
| +-----+
| | | +-- Potamilus_streckersoni **
| | +--+
| | +-- Potamilus_alatus **
+-----+
| +-- Lampsilis_cardium **
| +-----+
| | +-- Lampsilis_ornata **
+--+
| +----- Venustaconcha_ellipsiformis **
+--+
| +-- Lampsilis_siliquoidea **
+--+
+-- Lampsilis_powellii **
Kongim et al., 2023 [Publ 4290], fig. 1
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 15 |
15 |
13 |
1 |
0 |
0 |
0 |
77.0 |
+----- Chamberlainia_somsakpanhai **
+--+
| | +-- Chamberlainia_hainesiana **
| +--+
+-----+ +-- Chamberlainia_duclerci **
| |
| | +----- Sinohyriopsis_cumingii **
| +--+
| | +-- Sinohyriopsis_schlegelii **
| +--+
--+ +-- Sinohyriopsis_goliath **
|
| +----- Lamprotula_leaii **
| +-----+
| | | +-- Pronodularia_japanensis **
| | +--+
| | +-- Potomida_littoralis **
+--+
| +----- Sinosolenaia_carinata **
| +--+
| | | +-- Leguminaia_wheatleyi **
| | +--+
+--+ +-- Gonidea_angulata **
|
| +----- Indopseudodon_indawgyiensis **
+--+
| +-- Namkongnaia_inkhavilayi **
+--+
+-- Pilsbryoconcha_exilis **
Walker et al., 2006 [Publ 2666], fig. 6-M
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 14 |
14 |
11 |
2 |
1 |
1 |
1 |
80.0 |
+-------------------------- Echyridella_menziesii
|
| +----- Cumberlandia_monodonta **
| |
--+ +-----------------+ +-- Margaritifera_margaritifera **
| | | |
| | +--+-- Margaritifera_dahurica[1] **
| | |
| | +-- Margaritifera_dahurica[2] **
+--+
| +-- Gonidea_angulata **
| +-----------------+
| | +-- Sundadontina_tumida **
| |
+--+ +-- Pyganodon_fragilis **
| +--------------+
| | +-- Pyganodon_grandis **
+--+
| +-------------- Fusconaia_flava **
+--+
| +----------- Potamilus_purpuratus **
+--+
| +-------- Cyrtonaias_tampicoensis **
+--+
| +----- Ligumia_recta **
+--+
| +-- Lampsilis_teres **
+--+
+-- Ortmanniana_ligamentina **
Zhang et al., 2004 [Publ 2735], fig. 1
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 14 |
14 |
12 |
1 |
0 |
0 |
0 |
79.0 |
+-- Lamprotula_caveata **
+-----------------------+
| +-- Sinohyriopsis_cumingii **
|
--+ +-- Sinanodonta_woodiana **
| +--------------------+
| | +-- Cristaria_plicata **
| |
+--+ +----- Acuticosta_chinensis **
| +--------------+
| | | +-- Lanceolaria_grayii **
| | +--+
+--+ +-- Lanceolaria_lanceolata **
|
| +----------------- Tchangsinaia_piscicula **
+--+
| +-------------- Cuneopsis_heudei **
+--+
| +----------- Nodularia_douglasiae **
+--+
| +-------- Pseudocuneopsis_capitata **
+--+
| +----- Cuneopsis_rufescens **
+--+
| +-- Schistodesmus_lampreyanus **
+--+
+-- Schistodesmus_spinosus **
Kandl et al., 2001 [Publ 3384], fig. 5
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 14 |
14 |
7 |
1 |
0 |
0 |
0 |
81.0 |
+-------------------- Amblema_plicata **
|
+-------------------- Quadrula_quadrula **
--+
| +----------------- Elliptio_crassidens **
| |
+--+ +-------------- Fusconaia_flava **
| |
+--+ +----------- Pleurobema_sintoxia **
| |
| | +-- Pleurobema_clava **
+--+ +-----+
| | +-- Pleurobema_strodeanum **
| |
+--+ +----- Pleurobema_pyriforme[6] **
| |
| | +-- Pleurobema_pyriforme[1] **
+--+ |
| +-- Pleurobema **
| |
| +-- Pleurobema_pyriforme[2] **
+--+
+-- Pleurobema_pyriforme[3] **
|
+-- Pleurobema_pyriforme[4] **
|
+-- Pleurobema_pyriforme[5] **
Froufe et al., 2016 [Publ 3738], fig. 3
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 14 |
14 |
12 |
1 |
0 |
0 |
0 |
83.0 |
+-- Nodularia_douglasiae **
+--------------------------+
| +-- Nodularia_breviconcha **
|
--+ +-- Unio_gibbus **
| +-----------------------+
| | +-- Unio_durieui **
+--+
| +----------------------- Unio_tumidus **
| |
+--+ +-- Unio_crassus **
| +-----------------+
| | +-- Unio_tumidiformis **
+--+
| +----------------- Unio_delphinus **
+--+
| +-------------- Unio_foucauldianus **
+--+
| +----------- Unio_ravoisieri **
+--+
| +-------- Unio_mancus **
+--+
| +----- Unio_elongatulus **
+--+
| +-- Unio_pictorum **
+--+
+-- Unio_caffer **
Bolotov et al., 2016 [Publ 3743], fig. 2
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 14 |
14 |
12 |
2 |
16 |
13 |
2 |
82.0 |
+----- Diplodon_suavidicus
+-----+
| | +-- Triplodon_corrugatus
| +--+
| +-- Castalia_ambigua
+--------------+
| | +----- Microdontia_anodontaeformis
| | +--+
| | | | +-- Lortiella_froggatti
| | | +--+
| +--+ +-- Alathyria_pertexta
| |
| | +----- Velesunio_ambiguus
| +--+
| | +-- Alathyria_jacksoni
--+ +--+
| +-- Alathyria_profuga
|
| +-------- Tamsiella_schroeteriana
| +--------------+
| | | +----- Etheria_elliptica
| | +--+
| | | +-- Aspatharia_pfeifferiana
| | +--+
| | +-- Chambardia_wahlbergi
+--+
| +-------- Lampsilis_cardium **
| +-----------+
| | | +----- Anodonta_anatina **
| | +--+
| | | +-- Unio_pictorum **
| | +--+
+--+ +-- Unio_ravoisieri **
|
| +-- Pseudunio_auricularius **
| +--------------+
| | +-- Pseudunio_marocanus **
| |
+--+ +-- Gibbosula_laosensis **
| +-----------+
| | +-- Cumberlandia_monodonta **
| |
+--+ +-- Margaritifera_dahurica **
| +--------+
| | +-- Margaritifera_margaritifera **
+--+
| +-------- Margaritifera_falcata **
+--+
| +----- Margaritifera_laevis **
+--+
| +-- Margaritifera_middendorffi **
+--+
+-- Margaritifera_marrianae **
Bolotov et al., 2016 [Publ 3743], fig. 4
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 14 |
14 |
12 |
2 |
16 |
13 |
2 |
82.0 |
+----- Diplodon_suavidicus
+-----+
| | +-- Triplodon_corrugatus
| +--+
| +-- Castalia_ambigua
+--------------+
| | +----- Microdontia_anodontaeformis
| | +--+
| | | | +-- Alathyria_pertexta
| | | +--+
| +--+ +-- Lortiella_froggatti
| |
| | +----- Velesunio_ambiguus
| +--+
| | +-- Alathyria_jacksoni
--+ +--+
| +-- Alathyria_profuga
|
| +-------- Tamsiella_schroeteriana
| +--------------+
| | | +----- Etheria_elliptica
| | +--+
| | | +-- Chambardia_wahlbergi
| | +--+
| | +-- Aspatharia_pfeifferiana
+--+
| +-------- Lampsilis_cardium **
| +-----------+
| | | +----- Anodonta_anatina **
| | +--+
| | | +-- Unio_pictorum **
| | +--+
+--+ +-- Unio_ravoisieri **
|
| +-- Pseudunio_auricularius **
| +--------------+
| | +-- Pseudunio_marocanus **
| |
+--+ +-- Gibbosula_laosensis **
| +-----------+
| | +-- Cumberlandia_monodonta **
| |
+--+ +-- Margaritifera_margaritifera **
| +--------+
| | +-- Margaritifera_dahurica **
+--+
| +-------- Margaritifera_falcata **
+--+
| +----- Margaritifera_laevis **
+--+
| +-- Margaritifera_marrianae **
+--+
+-- Margaritifera_middendorffi **
Araujo et al., 2017 [Publ 3801], fig. 2
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 14 |
14 |
12 |
2 |
0 |
0 |
0 |
74.0 |
+-------------------- Pseudunio_marocanus **
|
| +-- Lamellidens_savadiensis **
--+ +--------------+
| | +-- Coelatura_aegyptiaca **
| |
| | +-- Amblema_plicata **
+--+ +-----------+
| | +-- Lampsilis_teres **
| |
| | +-- Schistodesmus_lampreyanus **
| | +--+
+--+ | +-- Aculamprotula_tortuosa **
| +-----+
| | | +-- Anodonta_anatina **
| | +--+
| | +-- Unio_delphinus **
+--+
| +-- Gonidea_angulata **
| +-----+
| | +-- Microcondylaea_bonellii **
+--+
| +----- Potomida_littoralis **
+--+
| +-- Lamprotula_leaii **
+--+
+-- Pronodularia_japanensis **
Lopes-Lima et al., 2020 [Publ 4080], fig. 10
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 14 |
14 |
10 |
2 |
0 |
0 |
0 |
86.0 |
+-- Margaritifera_middendorffi **
+-----------------+
| +-- Margaritifera_laevis **
|
--+ +-- Beringiana_japonica **
| +--------------+
| | +-- Beringiana_beringiana **
| |
+--+ +-- Sinanodonta_angula **
| +-----------+
| | +-- Sinanodonta_lucida **
| |
+--+ +----------- Sinanodonta_schrenkii **
| |
| | +-- Sinanodonta_calipygos **
| +--------+
+--+ +-- Sinanodonta_tumens **
|
| +-------- Sinanodonta_woodiana[2] **
| |
+--+-------- Sinanodonta_jourdyi **
|
| +----- Sinanodonta_elliptica **
+--+
| +-- Sinanodonta_lauta **
+--+
+-- Sinanodonta_woodiana[1] **
Fukata & Iigo, 2019 [Publ 4108], fig. 1
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 14 |
14 |
12 |
1 |
0 |
0 |
0 |
79.0 |
+----------- Aculamprotula_tientsinensis **
+--+
| | +-------- Acuticosta_chinensis **
| +--+
| | +----- Lanceolaria_gladiola **
| +--+
| | +-- Lanceolaria_grayii **
| +--+
--+ +-- Lanceolaria_lanceolata **
|
| +-------- Unio_tumidus **
| +--+
| | | +----- Unio_crassus **
| | +--+
| | | +-- Unio_delphinus **
+--+ +--+
| +-- Unio_pictorum **
|
| +-------- Pseudocuneopsis_capitata **
| |
+--+ +-- Nodularia_douglasiae **
| +--+
| | +-- Tchangsinaia_piscicula **
+--+
| +-- Schistodesmus_lampreyanus **
+--+
+-- Cuneopsis_heudei **
Wu et al., 2020 [Publ 4147], fig. 4
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 14 |
14 |
12 |
2 |
0 |
0 |
0 |
78.0 |
+----- Gibbosula_rochechouartii **
+--------------+
| | +-- Margaritifera_margaritifera **
| +--+
| +-- Margaritifera_falcata **
--+
| +----------------- Lampsilis_cardium **
| |
| | +----------- Sinosolenaia_carinata **
| | +--+
+--+ | | +-------- Potomida_littoralis **
| | +--+
| | | +----- Lamprotula_leaii **
| | +--+
+--+ | +-- Lamprotula_caveata **
| +--+
| +-- Lamprotula_cornuumlunae **
|
| +----------- Unio_pictorum **
+--+
| +-------- Aculamprotula_tortuosa **
+--+
| +----- Aculamprotula_scripta **
+--+
| +-- Aculamprotula_tientsinensis **
+--+
+-- Aculamprotula_fibrosa **
Sano et al., 2022 [Publ 4311], fig. 2
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 14 |
14 |
12 |
1 |
0 |
0 |
0 |
89.0 |
+-------- Pletholophus_tenuis **
+-----------+
| | +----- Cristaria_plicata **
| +--+
| | +-- Buldowskia_iwakawai **
| +--+
| +-- Buldowskia_kamiyai **
--+
| +-------- Beringiana_fukuharai **
| +--------+
| | | +----- Beringiana_japonica **
| | +--+
| | | +-- Beringiana_gosannensis **
+--+ +--+
| +-- Beringiana_beringiana **
|
| +-------------- Sinanodonta_tumens **
+--+
| +----------- Sinanodonta[2] **
+--+
| +-------- Sinanodonta_calipygos **
+--+
| +----- Sinanodonta_lauta **
+--+
| +-- Sinanodonta_woodiana **
+--+
+-- Sinanodonta[1] **
Graf, 2002 [Publ 2651], fig. 2
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 13 |
13 |
11 |
2 |
5 |
4 |
1 |
78.0 |
+-------- Velesunio_ambiguus
+--------------+
| | +----- Hyridella_depressa
| +--+
| | +-- Diplodon_chilensis
| +--+
--+ +-- Castalia
|
| +-------------------- Cumberlandia_monodonta **
| |
| | +-- Pilsbryoconcha_exilis **
+--+ +--------------+
| | +-- Pseudodon_vondembuschianus **
| |
+--+ +-------------- Lens_contradens **
| |
| | +-- Unio_pictorum **
| | +--+
+--+ | +-- Unio_caffer **
| +-----+
| | | +-- Alasmidonta_marginata **
| | +--+
+--+ +-- Pyganodon_grandis **
|
| +-------- Gonidea_angulata **
| |
+--+ +-- Lampsilis_cardium **
| +--+
| | +-- Obliquaria_reflexa **
+--+
| +-- Eurynia_dilatata **
+--+
+-- Amblema_plicata **
Walker et al., 2006 [Publ 2666], fig. 2
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 13 |
13 |
11 |
2 |
2 |
1 |
1 |
84.5 |
+-------------------------- Hyridella
|
| +----- Anodonta **
--+ +-----------------+
| | | +-- Pyganodon_fragilis **
| | +--+
| | +-- Pyganodon_grandis **
+--+
| +-------------------- Pronodularia_japanensis **
| |
| | +-- Gonidea_angulata **
+--+ +--------------+
| | +-- Pseudodon_vondembuschianus **
| |
+--+ +-- Fusconaia_flava **
| +-----------+
| | +-- Cumberlandia_monodonta **
+--+
| +----------- Cyrtonaias_tampicoensis **
+--+
| +-------- Potamilus_purpuratus **
+--+
| +----- Ligumia_recta **
+--+
| +-- Ortmanniana_ligamentina **
+--+
+-- Lampsilis_teres **
Walker et al., 2006 [Publ 2666], fig. 3
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 13 |
13 |
11 |
2 |
2 |
1 |
1 |
82.5 |
+-------------------------------- Hyridella
|
--+ +----------------------------- Cumberlandia_monodonta **
| |
+--+ +-------------------------- Pseudodon_vondembuschianus **
| |
+--+ +----------------------- Gonidea_angulata **
| |
+--+ +-------------------- Inversidens **
| |
| | +----- Anodonta **
+--+ +-----------+
| | | +-- Pyganodon_fragilis **
| | +--+
+--+ +-- Pyganodon_grandis **
|
| +-------------- Fusconaia_flava **
+--+
| +----------- Potamilus_purpuratus **
+--+
| +-------- Cyrtonaias_tampicoensis **
+--+
| +----- Ligumia_recta **
+--+
| +-- Ortmanniana_ligamentina **
+--+
+-- Lampsilis_teres **
Walker et al., 2006 [Publ 2666], fig. 4
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 13 |
13 |
11 |
2 |
2 |
1 |
1 |
82.5 |
+-------------------------------- Hyridella
|
--+ +----------------------------- Cumberlandia_monodonta **
| |
+--+ +-------------------------- Pseudodon_vondembuschianus **
| |
+--+ +----------------------- Gonidea_angulata **
| |
+--+ +-------------------- Inversidens **
| |
| | +----- Anodonta **
+--+ +-----------+
| | | +-- Pyganodon_fragilis **
| | +--+
+--+ +-- Pyganodon_grandis **
|
| +-------------- Fusconaia_flava **
+--+
| +----------- Cyrtonaias_tampicoensis **
+--+
| +-------- Potamilus_purpuratus **
+--+
| +----- Ligumia_recta **
+--+
| +-- Ortmanniana_ligamentina **
+--+
+-- Lampsilis_teres **
Manendo et al., 2008 [Publ 2669], fig. 1
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 13 |
13 |
11 |
1 |
0 |
0 |
0 |
79.0 |
+-- Lampsilis_siliquoidea **
+-----+
| +-- Sagittunio_nasutus **
+-----+
| | +----- Amblema_plicata **
| +--+
| | +-- Pleuronaia_dolabelloides **
| +--+
| +-- Fusconaia_flava **
--+
| +----- Unio_crassus **
| +-----+
| | | +-- Unio_pictorum **
| | +--+
| | +-- Unio_tumidus **
+--+
| +-- Anodonta_cygnea **
| +-----+
| | +-- Anodonta_anatina **
+--+
| +----- Pseudanodonta_complanata **
+--+
| +-- Platynaias_subviridis **
+--+
+-- Pyganodon_grandis **
Serb & Barnhart, 2008 [Publ 2674], fig. 3
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 13 |
13 |
10 |
1 |
0 |
0 |
0 |
84.0 |
+-- Lampsilis_siliquoidea **
+-----------------------+
| +-- Lampsilis_ornata **
--+
| +----------------------- Obliquaria_reflexa **
| |
+--+ +-------------------- Ptychobranchus_fasciolaris **
| |
+--+ +-- Cyprogenia_aberti[6] **
| +--------------+
| | +-- Dromus_dromas **
+--+
| +-------------- Cyprogenia_aberti[4] **
| |
+--+ +----------- Cyprogenia_aberti[3] **
| |
+--+ +-------- Cyprogenia_aberti[5] **
| |
+--+ +----- Cyprogenia_stegaria[2] **
| |
+--+----- Cyprogenia_aberti[2] **
|
| +-- Cyprogenia_stegaria[1] **
+--+
+-- Cyprogenia_aberti[1] **
Serb & Barnhart, 2008 [Publ 2674], fig. 4
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 13 |
13 |
9 |
1 |
0 |
0 |
0 |
84.0 |
+----------------------- Obliquaria_reflexa **
|
| +-- Lampsilis_ornata **
+--------------------+
--+ +-- Lampsilis_siliquoidea **
|
| +-------------------- Ptychobranchus_fasciolaris **
| |
+--+ +-- Cyprogenia_aberti[6] **
| +--------------+
| | +-- Dromus_dromas **
+--+
| +-------------- Cyprogenia_aberti[5] **
| |
+--+ +----------- Cyprogenia_aberti[4] **
| |
+--+ +-------- Cyprogenia_aberti[3] **
| |
+--+ +----- Cyprogenia_stegaria[2] **
| |
+--+----- Cyprogenia_aberti[2] **
|
| +-- Cyprogenia_stegaria[1] **
+--+
+-- Cyprogenia_aberti[1] **
Serb, 2006 [Publ 2678], fig. 2
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 13 |
13 |
7 |
1 |
0 |
0 |
0 |
84.0 |
+-------------- Obliquaria_reflexa **
|
| +----- Ptychobranchus_fasciolaris **
+--------+
| | +-- Lampsilis_ornata **
--+ +--+
| +-- Lampsilis_siliquoidea **
|
| +-- Cyprogenia_aberti[6] **
| +--------+
| | +-- Dromus_dromas **
+--+
| +-------- Cyprogenia_aberti[5] **
| |
+--+ +----- Cyprogenia_aberti[4] **
| |
| | +-- Cyprogenia_stegaria[1] **
+--+ |
| +-- Cyprogenia_stegaria[2] **
| |
+--+-- Cyprogenia_aberti[1] **
|
+-- Cyprogenia_aberti[2] **
|
+-- Cyprogenia_aberti[3] **
Kandl et al., 2001 [Publ 3384], fig. 4
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 13 |
13 |
4 |
1 |
0 |
0 |
0 |
80.5 |
+-------------- Elliptio **
|
+-------------- Uniomerus_carolinianus **
--+
| +----------- Pleurobema_sintoxia **
| |
+--+ +-------- Pleurobema_strodeanum **
| |
| | +----- Pleurobema_pyriforme[7] **
+--+ |
| | +-- Pleurobema_pyriforme[1] **
| | |
+--+ +-- Pleurobema_pyriforme[2] **
| |
| +-- Pleurobema_pyriforme[3] **
| |
| +-- Pleurobema_pyriforme[4] **
+--+
+-- Pleurobema_pyriforme[5] **
|
+-- Pleurobema[1] **
|
+-- Pleurobema_pyriforme[6] **
|
+-- Pleurobema[2] **
Ouyang et al., 2011 [Publ 3408], fig. 1
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 13 |
13 |
10 |
1 |
0 |
0 |
0 |
78.0 |
+-------- Lamprotula_caveata **
+-----+
| | +----- Sinosolenaia_oleivora **
| +--+
| | +-- Sinosolenaia_carinata **
| +--+
| +-- Parvasolenaia_rivularis **
--+
| +----- Anemina_arcaeformis **
| +-----+
| | | +-- Sinanodonta_elliptica **
| | +--+
| | +-- Sinanodonta_woodiana **
+--+
| +-- Nodularia_douglasiae **
| +-----+
| | +-- Cuneopsis_heudei **
+--+
| +----- Lanceolaria_grayii **
| |
+--+----- Acuticosta_chinensis **
|
| +-- Lanceolaria_gladiola **
+--+
+-- Aculamprotula_tortuosa **
Reis et al., 2013 [Publ 3580], fig. 4
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 13 |
13 |
11 |
2 |
1 |
0 |
0 |
76.0 |
+----------------------- Margaritifera_margaritifera **
|
| +-- Amblema_plicata **
--+ +-----------------+
| | +-- Quadrula_nobilis **
| |
+--+ +----------------- Potomida_littoralis **
| |
| | +----- Anodonta_cygnea **
+--+ +--------+
| | | +-- Anodonta_anatina **
| | +--+
+--+ +-- Pseudanodonta_complanata **
|
| +----------- Unio_tumidus **
| |
+--+ +-- Unio_tumidiformis **
| +-----+
| | +-- Unio_crassus **
+--+
| +----- Unio_delphinus **
+--+
| +-- Unio_mancus **
+--+
+-- Unio_pictorum **
Burzynski et al., 2017 [Publ 3774], fig. 1 bottom
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 13 |
13 |
11 |
1 |
0 |
0 |
0 |
72.0 |
+----- Aculamprotula_tortuosa **
+--+
| | +-- Unio_crassus **
| +--+
+--------+ +-- Unio_tumidus **
| |
| | +----- Anodonta_anatina **
| +--+
| | +-- Utterbackia_peninsularis **
--+ +--+
| +-- Pyganodon_grandis **
|
| +-- Quadrula_quadrula **
| +-----------+
| | +-- Venustaconcha_ellipsiformis **
+--+
| +----------- Sinohyriopsis_cumingii **
+--+
| +-------- Sinosolenaia_carinata **
+--+
| +----- Potomida_littoralis **
+--+
| +-- Lamprotula_leaii **
+--+
+-- Pronodularia_japanensis **
Burzynski et al., 2017 [Publ 3774], fig. 1 top
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 13 |
13 |
11 |
1 |
0 |
0 |
0 |
70.0 |
+----- Aculamprotula_tortuosa **
+--+
| | +-- Unio_crassus **
| +--+
+--------+ +-- Unio_tumidus **
| |
| | +----- Anodonta_anatina **
| +--+
| | +-- Utterbackia_peninsularis **
--+ +--+
| +-- Pyganodon_grandis **
|
| +-- Quadrula_quadrula **
| +-----------+
| | +-- Venustaconcha_ellipsiformis **
+--+
| +----------- Sinohyriopsis_cumingii **
+--+
| +-------- Sinosolenaia_carinata **
+--+
| +----- Lamprotula_leaii **
+--+
| +-- Potomida_littoralis **
+--+
+-- Pronodularia_japanensis **
Vikhrev et al., 2018 [Publ 3812], fig. 3
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 13 |
13 |
11 |
2 |
0 |
0 |
0 |
84.0 |
+-- Unio_ravoisieri **
+-----------------+
| +-- Lampsilis_cardium **
|
--+ +----------------- Gibbosula_laosensis **
| |
| | +-------- Cumberlandia_monodonta **
| | +-----+
+--+ | | +----- Pseudunio_marocanus **
| | +--+
| | | +-- Pseudunio_auricularius **
| | +--+
+--+ +-- Pseudunio_homsensis **
|
| +-- Margaritifera_margaritifera **
| +--------+
| | +-- Margaritifera_dahurica **
+--+
| +-------- Margaritifera_falcata **
+--+
| +----- Margaritifera_laevis **
+--+
| +-- Margaritifera_middendorffi **
+--+
+-- Margaritifera_marrianae **
Sayenko et al., 2021 [Publ 4189], fig. 6
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 13 |
13 |
10 |
1 |
0 |
0 |
0 |
82.0 |
+-- Unio_pictorum **
+-----------------+
| +-- Unio_crassus **
|
| +-------- Inversiunio_jokohamensis **
--+ |
| +--------+ +----- Inversiunio_yanagawensis[2] **
| | | |
| | +--+----- Inversiunio_verrucosus **
| | |
+--+ | +-- Inversiunio_reinianus **
| +--+
| +-- Inversiunio_yanagawensis[1] **
|
| +-------------- Middendorffinaia_mongolica **
+--+
| +----------- Nodularia_breviconcha **
+--+
| +-------- Nodularia_douglasiae[2] **
+--+
| +----- Nodularia_nuxpersicae **
+--+
| +-- Nodularia_douglasiae[1] **
+--+
+-- Nodularia_nipponensis **
Jeratthitikul & Sutcharit, 2023 [Publ 4289], fig. 1
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 13 |
13 |
11 |
1 |
0 |
0 |
0 |
88.0 |
+-- Indopseudodon_indawgyiensis **
+--------------+
| +-- Indopseudodon_kayinensis **
|
| +----------- Bineurus_mouhotii **
| +--+
--+ | | +-------- Bineurus_loeiensis **
| | +--+
| | | +----- Bineurus_panhai **
| | +--+
| | | +-- Bineurus_anodontinum **
+--+ +--+
| +-- Bineurus_exilis **
|
| +----- Namkongnaia_inkhavilayi **
| +--+
| | | +-- Sundadontina_cumingii **
| | +--+
+-----+ +-- Nyeinchanconcha_nyeinchani **
|
| +----- Pseudodon_vondembuschianus **
+--+
| +-- Pilsbryoconcha_exilis **
+--+
+-- Thaiconcha_callifera **
Bogan et al., 2023 [Publ 4295], fig. 6
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 13 |
13 |
10 |
2 |
0 |
0 |
0 |
85.0 |
+-------------------------- Gibbosula_crassa **
|
--+ +----------------------- Pletholophus_honglinhensis **
| |
+--+ +-------------------- Cristaria_truncata **
| |
+--+ +-- Sinanodonta_angula **
| +--------------+
| | +-- Sinanodonta_lucida **
+--+
| +-------------- Sinanodonta_schrenkii **
| |
+--+ +-- Sinanodonta_calipygos **
| +--------+
| | +-- Sinanodonta_tumens **
+--+
| +-------- Sinanodonta_woodiana[2] **
| |
+--+-------- Sinanodonta_jourdyi **
|
| +----- Sinanodonta_elliptica **
+--+
| +-- Sinanodonta_lauta **
+--+
+-- Sinanodonta_woodiana[1] **
Araujo et al., 2009 [Publ 2589], fig. 7A
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 12 |
12 |
7 |
2 |
0 |
0 |
0 |
79.0 |
+-------------------- Pseudunio_auricularius **
|
--+ +----------------- Amblema_plicata **
| |
+--+ +-------------- Potomida_littoralis **
| |
+--+ +----------- Nodularia_douglasiae **
| |
| | +-- Unio_gibbus **
+--+ |
| +-----+-- Unio_tumidus **
| | |
| | +-- Anodonta_cygnea **
+--+
| +----- Unio_pictorum[1] **
| |
| +----- Unio_pictorum[2] **
+--+
+----- Unio_mancus **
|
| +-- Unio_tumidiformis **
+--+
+-- Unio_crassus **
Araujo et al., 2009 [Publ 2589], fig. 7B
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 12 |
12 |
10 |
2 |
0 |
0 |
0 |
78.0 |
+-------------------------- Pseudunio_auricularius **
|
--+ +----------------------- Potomida_littoralis **
| |
+--+ +-------------------- Amblema_plicata **
| |
+--+ +----------------- Anodonta_cygnea **
| |
+--+ +-------------- Unio_gibbus **
| |
+--+ +----------- Unio_tumidus **
| |
| | +-- Unio_tumidiformis **
+--+ +-----+
| | +-- Unio_crassus **
| |
+--+ +-- Unio_pictorum[1] **
| +--+
| | +-- Unio_mancus **
+--+
| +-- Unio_pictorum[2] **
+--+
+-- Unio_caffer **
Serb, 2006 [Publ 2678], fig. 3
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 12 |
12 |
8 |
1 |
0 |
0 |
0 |
84.0 |
+-------------------- Obliquaria_reflexa **
|
| +-- Lampsilis_ornata **
+-----------------+
--+ +-- Lampsilis_siliquoidea **
|
| +----------------- Ptychobranchus_fasciolaris **
| |
+--+ +-- Cyprogenia_aberti[5] **
| +-----------+
| | +-- Dromus_dromas **
+--+
| +----------- Cyprogenia_aberti[4] **
| |
+--+ +-------- Cyprogenia_aberti[3] **
| |
+--+ +----- Cyprogenia_stegaria[2] **
| |
+--+----- Cyprogenia_aberti[2] **
|
| +-- Cyprogenia_stegaria[1] **
+--+
+-- Cyprogenia_aberti[1] **
Serb, 2006 [Publ 2678], fig. 4
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 12 |
12 |
8 |
1 |
0 |
0 |
0 |
84.0 |
+-------------------- Obliquaria_reflexa **
|
| +-- Lampsilis_ornata **
+-----------------+
--+ +-- Lampsilis_siliquoidea **
|
| +----------------- Ptychobranchus_fasciolaris **
| |
+--+ +-- Cyprogenia_aberti[5] **
| +-----------+
| | +-- Dromus_dromas **
+--+
| +----------- Cyprogenia_aberti[4] **
| |
+--+ +-------- Cyprogenia_aberti[3] **
| |
+--+ +----- Cyprogenia_stegaria[2] **
| |
+--+----- Cyprogenia_aberti[2] **
|
| +-- Cyprogenia_stegaria[1] **
+--+
+-- Cyprogenia_aberti[1] **
Curole & Kocher, 2005 [Publ 2682], fig. 1-M
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 12 |
12 |
10 |
2 |
0 |
0 |
0 |
82.0 |
+----------------------- Margaritifera_hembeli **
|
| +-- Utterbackiana_implicata **
--+ +-----------------+
| | +-- Pyganodon_grandis **
| |
+--+ +-- Quadrula_quadrula **
| +--------------+
| | +-- Pustulosa_pustulosa **
+--+
| +-------------- Plectomerus_dombeyanus **
| |
+--+ +----------- Amblema_plicata **
| |
+--+ +-- Potamilus_purpuratus **
| +-----+
| | +-- Glebula_rotundata **
+--+
| +----- Leaunio_lienosus **
+--+
| +-- Lampsilis_straminea **
+--+
+-- Lampsilis_teres **
Huff et al., 2004 [Publ 2697], fig. 2
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 12 |
12 |
6 |
2 |
2 |
0 |
0 |
80.5 |
+----- Lampsilis_cardium **
+-----+
| | +-- Unio_ravoisieri **
| +--+
| +-- Anodonta **
|
--+ +-------- Margaritifera_dahurica **
| |
| +-------- Margaritifera_margaritifera[1] **
| |
| +-------- Margaritifera_margaritifera[2] **
| |
+--+-------- Margaritifera_margaritifera[3] **
|
| +-- Pseudunio_auricularius **
+-----+
| +-- Cumberlandia_monodonta **
|
| +----- Margaritifera_falcata **
+--+
| +-- Margaritifera_middendorffi **
+--+
+-- Margaritifera_marrianae **
Roe, 2013 [Publ 3571], fig. 4
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 12 |
12 |
9 |
1 |
0 |
0 |
0 |
88.0 |
+----------------------- Obliquaria_reflexa **
|
| +-- Lampsilis_siliquoidea **
+--------------------+
--+ +-- Lampsilis_ornata **
|
| +-------------------- Medionidus_conradicus **
| |
+--+ +-- Cyprogenia_aberti **
| +--------------+
| | +-- Dromus_dromas **
+--+
| +-------------- Ptychobranchus_subtentus **
+--+
| +----------- Ptychobranchus_jonesi **
+--+
| +-------- Ptychobranchus_greenii **
+--+
| +----- Ptychobranchus_occidentalis[2] **
+--+
| +-- Ptychobranchus_occidentalis[1] **
+--+
+-- Ptychobranchus_fasciolaris **
Froufe et al., 2015 [Publ 3664], fig. 1
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 12 |
12 |
10 |
1 |
0 |
0 |
0 |
79.0 |
+-- Sinosolenaia_oleivora **
+-----+
| +-- Sinosolenaia_carinata **
+-----+
| | +----- Potomida_littoralis **
| +--+
| | +-- Lamprotula_leaii **
| +--+
--+ +-- Pronodularia_japanensis **
|
| +-- Sinohyriopsis_cumingii **
| +--------+
| | +-- Sinohyriopsis_schlegelii **
| |
+--+ +-- Venustaconcha_ellipsiformis **
| +-----+
| | +-- Quadrula_quadrula **
+--+
| +----- Anodonta_anatina **
+--+
| +-- Utterbackia_peninsularis **
+--+
+-- Pyganodon_grandis **
Araujo et al., 2016 [Publ 3746], fig. 1,2b
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 12 |
12 |
10 |
2 |
1 |
0 |
0 |
84.0 |
+-------------------- Potomida_littoralis **
|
--+ +----------------- Gibbosula_laosensis **
| |
| | +----- Cumberlandia_monodonta **
+--+ +--------+
| | | +-- Pseudunio_marocanus **
| | +--+
| | +-- Pseudunio_auricularius **
+--+
| +-- Margaritifera_dahurica **
| +--------+
| | +-- Margaritifera_margaritifera **
+--+
| +-------- Margaritifera_falcata **
| |
+--+ +-- Margaritifera_middendorffi **
| +--+
| | +-- Margaritifera_laevis **
+--+
| +-- Margaritifera_marrianae **
+--+
+-- Margaritifera_hembeli **
Froufe et al., 2017 [Publ 3802], fig. 6
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 12 |
12 |
9 |
1 |
0 |
0 |
0 |
81.0 |
+-- Anodonta_anatina **
+--------------------+
| +-- Anodonta_cygnea **
--+
| +-------------------- Anemina_arcaeformis **
| |
+--+ +-- Cristaria_plicata **
| +--------------+
| | +-- Cristaria_radiata **
+--+
| +-------------- Beringiana_beringiana **
| |
+--+ +----------- Sinanodonta_lucida **
| |
+--+ +-------- Sinanodonta_schrenkii **
| |
+--+ +----- Sinanodonta_woodiana[1] **
| |
+--+ +-- Sinanodonta_lauta **
| |
+--+-- Sinanodonta_elliptica **
|
+-- Sinanodonta_woodiana[2] **
Konopleva et al., 2020 [Publ 4072], fig. 2
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 12 |
12 |
9 |
1 |
0 |
0 |
0 |
86.0 |
+-- Leoparreysia_olivacea **
|
+--------------+-- Leoparreysia_canefrii **
| |
| +-- Leoparreysia_tavoyensis **
|
--+ +-------- Lamellidens_savadiensis **
| +-----+
| | | +----- Lamellidens_generosus **
| | +--+
| | | +-- Lamellidens_mainwaringi **
+--+ +--+
| +-- Lamellidens_ferrugineus **
|
| +----------- Trapezidens_scutum **
+--+
| +-------- Trapezidens_exolescens **
+--+
| +----- Trapezidens_yeti **
+--+
| +-- Trapezidens_angustior **
+--+
+-- Trapezidens_dolichorhynchus **
Lopes-Lima et al., 2020 [Publ 4080], fig. 11
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 12 |
12 |
10 |
2 |
0 |
0 |
0 |
81.0 |
+-- Margaritifera_laevis **
+--------------------+
| +-- Margaritifera_middendorffi **
|
--+ +-- Nodularia_breviconcha **
| +-----------------+
| | +-- Nodularia_douglasiae **
+--+
| +----------------- Lanceolaria_oxyrhyncha **
| |
+--+ +-------------- Lanceolaria_gladiola **
| |
+--+ +-- Lanceolaria_grayii **
| +--------+
| | +-- Lanceolaria_bilirata **
+--+
| +-------- Lanceolaria_acrorrhyncha **
+--+
| +----- Lanceolaria_kihirai **
+--+
| +-- Lanceolaria_lanceolata **
+--+
+-- Lanceolaria_triformis **
Lopes-Lima et al., 2020 [Publ 4080], fig. 6
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 12 |
12 |
10 |
2 |
0 |
0 |
0 |
86.0 |
+-- Margaritifera_laevis **
+-----------------+
| +-- Margaritifera_middendorffi **
|
--+ +-- Sinanodonta_lauta **
| +--------------+
| | +-- Sinanodonta_woodiana **
| |
+--+ +-------------- Amuranodonta_kijaensis **
| |
| | +-- Anemina_arcaeformis **
+--+ +--------+
| | +-- Anemina_euscaphys **
| |
+--+ +-- Buldowskia_iwakawai **
| +-----+
| | +-- Buldowskia_kamiyai **
+--+
| +----- Buldowskia_shadini **
+--+
| +-- Buldowskia_suifunica **
+--+
+-- Buldowskia_flavotincta **
Smith et al., 2020 [Publ 4136], fig. 3
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 12 |
12 |
10 |
1 |
0 |
0 |
0 |
89.0 |
+-------------- Pleurobema_clava **
|
| +----- Fusconaia_subrotunda **
| +-----+
--+ | | +-- Fusconaia_cor **
| | +--+
| | +-- Fusconaia_cuneolus **
| |
| | +-- Fusconaia_iheringi **
+--+ +--+
| | +-- Fusconaia_mitchelli **
| +--+
| | | +-- Fusconaia_askewi **
| | +--+
| | +-- Fusconaia_chunii **
+--+
| +-- Fusconaia_flava **
| +--+
| | +-- Fusconaia_masoni **
+--+
| +-- Fusconaia_burkei **
+--+
+-- Fusconaia_escambia **
Porto-Hannes et al., 2021 [Publ 4227], fig. 4
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 12 |
12 |
6 |
1 |
0 |
0 |
0 |
83.0 |
+-------------------- Quadrula_quadrula **
|
+-------------------- Pustulosa_pustulosa **
--+
| +----------------- Pleurobema_sintoxia **
| |
+--+ +-------------- Amblema_plicata **
| |
+--+ +----------- Obovaria_olivaria **
| |
+--+ +-------- Ortmanniana_ligamentina **
| |
+--+ +----- Lampsilis_hydiana **
| |
| +----- Lampsilis_straminea **
+--+
| +-- Lampsilis_siliquoidea **
| |
| +-- Lampsilis_radiata **
+--+
+-- Lampsilis[1] **
|
+-- Lampsilis[2] **
Stodola et al., 2021 [Publ 4266], fig. 4
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 12 |
12 |
7 |
1 |
0 |
0 |
0 |
86.0 |
+-- Ortmanniana_ligamentina **
|
+-----------+-- Lampsilis_abrupta **
| |
| +-- Lampsilis_higginsii **
--+
| +----------- Lampsilis_sietmani **
| |
| | +-- Lampsilis_siliquoidea **
+--+ +-----+
| | +-- Lampsilis_radiata **
| |
+--+ +----- Lampsilis_hydiana[2] **
| |
| +----- Lampsilis_straminea[2] **
| |
+--+ +-- Lampsilis_hydiana[1] **
+--+
| +-- Lampsilis_bergmanni **
|
| +-- Lampsilis_straminea[1] **
+--+
+-- Lampsilis_virescens **
Pfarrer et al., 2022 [Publ 4305], fig. 2-5
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 12 |
12 |
10 |
1 |
0 |
0 |
0 |
73.0 |
+-- Potomida_littoralis **
+--------------+
| +-- Microcondylaea_bonellii **
|
| +----------- Unio_tumidus **
| +--+
--+ | | +-------- Unio_crassus **
| | +--+
| | | +----- Unio_elongatulus **
| | +--+
| | | +-- Unio_mancus **
+--+ +--+
| +-- Unio_pictorum **
|
| +-- Cristaria_plicata **
| +-----+
| | +-- Sinanodonta_woodiana **
+-----+
| +----- Anodonta_anatina **
+--+
| +-- Anodonta_exulcerata **
+--+
+-- Anodonta_cygnea **
Araujo et al., 2009 [Publ 2589], fig. 7C
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 11 |
11 |
8 |
2 |
0 |
0 |
0 |
77.0 |
+-------------------- Pseudunio_auricularius **
|
--+ +----------------- Potomida_littoralis **
| |
+--+ +-------------- Amblema_plicata **
| |
+--+ +----------- Anodonta_cygnea **
| |
| | +-- Unio_gibbus **
+--+ +-----+
| | +-- Unio_tumidus **
| |
+--+ +-- Unio_tumidiformis **
| +--+
| | +-- Unio_crassus **
+--+
| +-- Unio_pictorum[1] **
| |
+--+-- Unio_pictorum[2] **
|
+-- Unio_mancus **
Curole & Kocher, 2005 [Publ 2682], fig. 1
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 11 |
11 |
9 |
2 |
0 |
0 |
0 |
85.0 |
+-- Pyganodon_grandis **
+-----------------+
| +-- Margaritifera_hembeli **
|
--+ +----------------- Glebula_rotundata **
| |
| | +----- Amblema_plicata **
+--+ +--------+
| | | +-- Quadrula_quadrula **
| | +--+
+--+ +-- Pustulosa_pustulosa **
|
| +----------- Plectomerus_dombeyanus **
+--+
| +-------- Leaunio_lienosus **
+--+
| +----- Lampsilis_straminea **
+--+
| +-- Lampsilis_teres **
+--+
+-- Potamilus_purpuratus **
Roe & Lydeard, 1998 [Publ 2693], fig. 3
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 11 |
11 |
5 |
1 |
0 |
0 |
0 |
86.0 |
+-------------- Fusconaia_cerina **
|
+-------------- Obliquaria_reflexa **
--+
| +----------- Potamilus_capax **
| |
| +----------- Lampsilis_ornata **
+--+
| +-------- Potamilus_fragilis **
| |
| | +----- Potamilus_inflatus **
+--+ |
| | +-- Potamilus_ohiensis **
| +--+
+--+ +-- Potamilus_amphichaenus **
|
| +-- Potamilus_purpuratus[1] **
| |
+--+-- Potamilus_purpuratus[2] **
|
+-- Potamilus_alatus **
Huff et al., 2004 [Publ 2697], fig. 3
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 11 |
11 |
3 |
2 |
1 |
0 |
0 |
79.5 |
+----------- Anodonta **
|
--+ +-------- Margaritifera_margaritifera **
| |
| | +----- Cumberlandia_monodonta **
+--+ |
| | +-- Pseudunio_auricularius **
| | |
+--+ +-- Margaritifera_dahurica **
| |
| +-- Margaritifera_middendorffi[1] **
| |
| +-- Margaritifera_middendorffi[2] **
+--+
+-- Margaritifera_falcata[1] **
|
+-- Margaritifera_falcata[2] **
|
+-- Margaritifera_falcata[3] **
|
+-- Margaritifera_falcata[4] **
Khalloufi et al., 2011 [Publ 2748], fig. 2
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 11 |
11 |
8 |
1 |
0 |
0 |
0 |
80.0 |
+----------------- Amblema_plicata **
|
--+ +-------------- Potomida_littoralis **
| |
| | +----- Unio_tumidus **
+--+ +-----+
| | | +-- Unio_gibbus **
| | +--+
| | +-- Unio_durieui **
+--+
| +-- Unio_tumidiformis **
| +-----+
| | +-- Unio_crassus **
+--+
| +----- Unio **
| |
+--+----- Unio_pictorum **
|
| +-- Unio_delphinus **
+--+
+-- Unio_mancus **
Araujo et al., 2016 [Publ 3746], fig. 2a
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 11 |
11 |
9 |
1 |
0 |
0 |
0 |
85.0 |
+-------------------- Cumberlandia_monodonta **
|
--+ +-- Pseudunio_marocanus **
| +--------------+
| | +-- Pseudunio_auricularius **
+--+
| +-------------- Gibbosula_laosensis **
| |
+--+ +-- Margaritifera_dahurica **
| +--------+
| | +-- Margaritifera_margaritifera **
+--+
| +-------- Margaritifera_falcata **
| |
+--+ +-- Margaritifera_laevis **
| +--+
| | +-- Margaritifera_middendorffi **
+--+
| +-- Margaritifera_marrianae **
+--+
+-- Margaritifera_hembeli **
Amaro et al., 2017 [Publ 3791], fig. 3A
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 11 |
11 |
9 |
2 |
1 |
0 |
0 |
78.0 |
+-- Margaritifera_falcata **
+-----------+
| +-- Margaritifera_margaritifera **
|
| +-------- Platynaias_compressa **
--+ +--+
| | | +----- Pyganodon_grandis **
| | +--+
| | | +-- Utterbackia_imbecillis **
| | +--+
+--+ +-- Utterbackia_peninsularis **
|
| +-- Pronodularia_japanensis **
| +-----+
| | +-- Sinosolenaia_carinata **
+--+
| +----- Quadrula_quadrula **
+--+
| +-- Toxolasma_parvum **
+--+
+-- Venustaconcha_ellipsiformis **
Amaro et al., 2017 [Publ 3791], fig. 3B
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 11 |
11 |
9 |
2 |
1 |
0 |
0 |
80.0 |
+-- Margaritifera_falcata **
+-----------+
| +-- Margaritifera_margaritifera **
|
| +-------- Platynaias_compressa **
--+ +--+
| | | +----- Pyganodon_grandis **
| | +--+
| | | +-- Utterbackia_imbecillis **
| | +--+
+--+ +-- Utterbackia_peninsularis **
|
| +-- Toxolasma_parvum **
| +-----+
| | +-- Venustaconcha_ellipsiformis **
+--+
| +----- Quadrula_quadrula **
+--+
| +-- Pronodularia_japanensis **
+--+
+-- Sinosolenaia_carinata **
Klishko et al., 2019 [Publ 3886], fig. 9
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 11 |
11 |
8 |
2 |
0 |
0 |
0 |
74.0 |
+----------------- Margaritifera_dahurica **
|
| +-- Anodonta_anatina **
--+ +-----------+
| | +-- Anodonta_cygnea **
| |
| | +-------- Middendorffinaia_mongolica **
+--+ +--+
| | | +----- Nodularia_breviconcha **
| | +--+
| | | +-- Nodularia_douglasiae **
+--+ +--+
| +-- Nodularia **
|
| +----- Unio_tumidus **
| |
+-----+----- Unio_gibbus **
|
| +-- Unio_crassus **
+--+
+-- Unio_pictorum **
Pfeiffer et al., 2021 [Publ 4126], fig. 12
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 11 |
11 |
8 |
1 |
0 |
0 |
0 |
85.0 |
+-------------- Solenaia_emarginata **
|
| +-- Lens_contradens **
| +--------+
--+ | +-- Lens_micropterus **
| |
| | +-- Lens_inornatus **
| +--------+
+--+ +-- Lens_misellus **
|
| +----- Lens_maenamensis **
| +--+
| | | +-- Lens_rolfbrandti **
| | +--+
+--+ +-- Lens_eximius **
|
| +----- Lens_pallegoixi **
+--+
| +-- Lens_comptus **
+--+
+-- Lens_novoselovi **
Pfeiffer et al., 2021 [Publ 4126], fig. 27
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 11 |
11 |
7 |
1 |
0 |
0 |
0 |
82.0 |
+----------------- Khairuloconcha_lunbawangorum **
|
--+ +-------------- Rectidens_sumatrensis **
| |
| | +-- Hyriopsis_delaportei **
+--+ +--------+
| | +-- Hyriopsis_desowitzi **
| |
+--+ +-------- Hyriopsis_khoratensis **
| |
| | +-- Hyriopsis_kratiensis **
| +-----+
+--+ +-- Hyriopsis_phuphaniensis **
|
| +----- Hyriopsis_sakhonensis **
| |
+--+ +-- Hyriopsis_myersiana **
| |
+--+-- Hyriopsis_panhai **
|
+-- Hyriopsis_bialata **
Choi et al., 2020 [Publ 4169], fig. 2a
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 11 |
11 |
9 |
1 |
0 |
0 |
0 |
80.0 |
+-------- Schistodesmus_lampreyanus **
|
+--------+ +-- Unio_pictorum **
| | +--+
| | | +-- Tchangsinaia_piscicula **
| +--+
--+ | +-- Cuneopsis_heudei **
| +--+
| +-- Cuneopsis_rufescens **
|
| +-------------- Nodularia_breviconcha **
+--+
| +----------- Nodularia[2] **
+--+
| +-------- Nodularia[1] **
+--+
| +----- Nodularia_douglasiae **
+--+
| +-- Nodularia_nipponensis **
+--+
+-- Nodularia_nuxpersicae **
Buhay et al., 2002 [Publ 2661], fig. 2A
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 10 |
10 |
3 |
1 |
0 |
0 |
0 |
83.0 |
+-------- Lampsilis_siliquoidea **
|
| +-- Cambarunio_taeniatus **
+-----+
| +-- Villosa_villosa **
--+
| +----- Epioblasma_triquetra **
| |
| +----- Epioblasma_brevidens **
| |
+--+ +-- Epioblasma_capsaeformis[1] **
| |
| +-- Epioblasma_capsaeformis[2] **
| |
+--+-- Epioblasma_capsaeformis[3] **
|
+-- Epioblasma_walkeri[1] **
|
+-- Epioblasma_walkeri[2] **
Buhay et al., 2002 [Publ 2661], fig. 2B
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 10 |
10 |
5 |
1 |
0 |
0 |
0 |
83.0 |
+-------------- Lampsilis_siliquoidea **
|
| +-- Cambarunio_taeniatus **
+-----------+
--+ +-- Villosa_villosa **
|
| +----------- Epioblasma_brevidens **
| |
+--+ +-------- Epioblasma_triquetra **
| |
+--+ +----- Epioblasma_capsaeformis[3] **
| |
+--+ +-- Epioblasma_capsaeformis[1] **
| |
| +-- Epioblasma_capsaeformis[2] **
+--+
+-- Epioblasma_walkeri[1] **
|
+-- Epioblasma_walkeri[2] **
Huff et al., 2004 [Publ 2697], fig. 4
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 10 |
10 |
7 |
2 |
1 |
0 |
0 |
79.5 |
+----- Lampsilis_cardium **
+--------+
| | +-- Unio_ravoisieri **
| +--+
| +-- Anodonta **
--+
| +----------- Pseudunio_auricularius **
| |
| +----------- Cumberlandia_monodonta **
+--+
| +-- Margaritifera_dahurica **
| +-----+
| | +-- Margaritifera_margaritifera **
+--+
| +----- Margaritifera_falcata **
+--+
| +-- Margaritifera_middendorffi **
+--+
+-- Margaritifera_marrianae **
Huff et al., 2004 [Publ 2697], fig. 5-6
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 10 |
10 |
7 |
2 |
2 |
0 |
0 |
80.5 |
+-- Unio_ravoisieri **
|
+-----------+-- Anodonta **
| |
| +-- Lampsilis_cardium **
|
--+ +-- Pseudunio_auricularius **
| +--------+
| | +-- Cumberlandia_monodonta **
| |
+--+ +-- Margaritifera_dahurica **
| +-----+
| | +-- Margaritifera_margaritifera **
+--+
| +----- Margaritifera_falcata **
+--+
| +-- Margaritifera_middendorffi **
+--+
+-- Margaritifera_marrianae **
Bolotov et al., 2015 [Publ 3646], fig. 2A
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 10 |
10 |
6 |
2 |
0 |
0 |
0 |
81.0 |
+-------------- Unio_pictorum **
|
| +----------- Margaritifera_margaritifera **
--+ |
| | +----- Margaritifera_dahurica[1] **
| | |
| +-----+----- Margaritifera_dahurica[2] **
+--+ |
| | +-- Pseudunio_auricularius **
| +--+
| +-- Cumberlandia_monodonta **
|
| +-------- Margaritifera_falcata **
+--+
| +----- Margaritifera_laevis **
+--+
| +-- Margaritifera_middendorffi **
+--+
+-- Margaritifera_marrianae **
Takeuchi et al., 2015 [Publ 3661], fig. 3
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 10 |
10 |
7 |
2 |
1 |
0 |
0 |
81.0 |
+-------------------- Lampsilis_cardium **
|
--+ +----------------- Pseudunio_auricularius[1] **
| |
| +----------------- Pseudunio_auricularius[2] **
+--+
| +-------------- Cumberlandia_monodonta **
| |
+--+ +-- Margaritifera_margaritifera **
| +--------+
| | +-- Margaritifera_dahurica **
+--+
| +-------- Margaritifera_falcata **
+--+
| +----- Margaritifera_laevis **
+--+
| +-- Margaritifera_middendorffi **
+--+
+-- Margaritifera_marrianae **
Amaro et al., 2017 [Publ 3791], fig. 2-M
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 10 |
10 |
7 |
1 |
0 |
0 |
0 |
79.0 |
+-- Fusconaia_flava **
+--+
| +-- Quadrula_quadrula **
+-----+
| | +-- Lampsilis_siliquoidea **
| +--+
| +-- Venustaconcha_ellipsiformis **
--+
| +-- Pronodularia_japanensis **
| +-----+
| | +-- Sinosolenaia_carinata **
+--+
| +----- Pyganodon_grandis **
| |
+--+ +-- Lasmigona_costata **
| |
+--+-- Sinanodonta_woodiana **
|
+-- Utterbackia_peninsularis **
Lopes-Lima et al., 2017 [Publ 3792], fig. 2-M
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 10 |
10 |
8 |
2 |
0 |
0 |
0 |
75.0 |
+----------------- Pseudunio_marocanus **
|
| +-- Venustaconcha_ellipsiformis **
--+ +-----------+
| | +-- Quadrula_quadrula **
| |
+--+ +----- Sinosolenaia_carinata **
| +-----+
| | | +-- Potomida_littoralis **
| | +--+
+--+ +-- Pronodularia_japanensis **
|
| +-------- Unio_delphinus **
+--+
| +----- Anodonta_anatina **
+--+
| +-- Utterbackia_peninsularis **
+--+
+-- Pyganodon_grandis **
Froufe et al., 2017 [Publ 3802], fig. 7
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 10 |
10 |
8 |
2 |
0 |
0 |
0 |
75.0 |
+----------------- Margaritifera_margaritifera **
|
| +-- Microcondylaea_bonellii **
--+ +-----------+
| | +-- Potomida_littoralis **
| |
+--+ +----- Unio_mancus **
| +-----+
| | | +-- Unio_elongatulus **
| | +--+
+--+ +-- Unio_pictorum **
|
| +-------- Sinanodonta_woodiana **
+--+
| +----- Anodonta_anatina **
+--+
| +-- Anodonta_exulcerata **
+--+
+-- Anodonta_cygnea **
Vikhrev et al., 2017 [Publ 3810], fig. 4
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 10 |
10 |
8 |
2 |
0 |
0 |
0 |
80.0 |
+-- Gibbosula_laosensis **
+--------------------+
| +-- Margaritifera_dahurica **
--+
| +-------------------- Anemina_arcaeformis **
+--+
| +----------------- Sinanodonta_lucida **
+--+
| +-------------- Sinanodonta_schrenkii **
+--+
| +----------- Sinanodonta_woodiana[1] **
+--+
| +-------- Sinanodonta_jourdyi **
+--+
| +----- Sinanodonta_elliptica **
+--+
| +-- Sinanodonta_lauta **
+--+
+-- Sinanodonta_woodiana[2] **
Bespalaya et al., 2018 [Publ 3811], fig. 3
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 10 |
10 |
8 |
2 |
0 |
0 |
0 |
80.0 |
+-- Gibbosula_laosensis **
+--------------------+
| +-- Margaritifera_dahurica **
--+
| +-------------------- Anemina_arcaeformis **
+--+
| +----------------- Sinanodonta_lucida **
+--+
| +-------------- Sinanodonta_schrenkii **
+--+
| +----------- Sinanodonta_woodiana[2] **
+--+
| +-------- Sinanodonta_jourdyi **
+--+
| +----- Sinanodonta_elliptica **
+--+
| +-- Sinanodonta_woodiana[1] **
+--+
+-- Sinanodonta_lauta **
Chase et al., 2018 [Publ 3861], fig. 1
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 10 |
10 |
8 |
2 |
1 |
1 |
1 |
77.0 |
+-------------------- Echyridella_menziesii
|
--+ +----------------- Cumberlandia_monodonta **
| |
| | +----- Quadrula_quadrula **
+--+ +--------+
| | | +-- Lampsilis_powellii **
| | +--+
+--+ +-- Lampsilis_siliquoidea **
|
| +----------- Lamprotula_leaii **
| |
+--+ +-- Unio_delphinus **
| +-----+
| | +-- Unio_tumidus **
+--+
| +----- Anodonta_anatina **
+--+
| +-- Utterbackia_peninsularis **
+--+
+-- Pyganodon_grandis **
Chase et al., 2018 [Publ 3861], fig. 1-M
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 10 |
11 |
9 |
2 |
1 |
1 |
1 |
76.0 |
+-- Cumberlandia_monodonta **
+-----------+
| +-- Echyridella_menziesii
|
| +-------- Lamprotula_leaii **
--+ +--+
| | | +----- Quadrula_quadrula **
| | +--+
| | | +-- Lampsilis_powellii **
| | +--+
+--+ +-- Lampsilis_siliquoidea **
|
| +-- Unio_delphinus **
| +-----+
| | +-- Unio_tumidus **
+--+
| +----- Anodonta_anatina **
+--+
| +-- Utterbackia_peninsularis **
+--+
+-- Pyganodon_grandis **
Burzynski & Soroka, 2018 [Publ 3862], fig. 5b
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 10 |
10 |
8 |
1 |
0 |
0 |
0 |
75.0 |
+----------------- Lanceolaria_lanceolata **
|
| +-------- Sinanodonta_woodiana **
--+ +-----+
| | | +----- Anodonta_anatina **
| | +--+
| | | +-- Utterbackia_peninsularis **
+--+ +--+
| +-- Pyganodon_grandis **
|
| +----------- Aculamprotula_tortuosa **
+--+
| +-------- Unio_tumidus **
+--+
| +----- Unio_crassus **
+--+
| +-- Unio_pictorum **
+--+
+-- Unio_delphinus **
Muanta et al., 2019 [Publ 3921], fig. 2
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 10 |
10 |
8 |
1 |
0 |
0 |
0 |
77.0 |
+-------- Hyriopsis_desowitzi **
+-----+
| | +----- Hyriopsis_myersiana **
| +--+
| | +-- Hyriopsis_khoratensis **
| +--+
--+ +-- Hyriopsis_panhai **
|
| +-- Ensidens_ingallsianus **
| +--------+
| | +-- Ensidens_telus **
+--+
| +-------- Ensidens_sagittarius[2] **
+--+
| +----- Ensidens_jaculus **
+--+
| +-- Ensidens_sagittarius[1] **
+--+
+-- Ensidens_dugasti **
Keogh & Simons, 2019 [Publ 3935], fig. S1
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 10 |
10 |
8 |
1 |
0 |
0 |
0 |
86.0 |
+-- Amblema_plicata **
+-----------------+
| +-- Fusconaia_mitchelli **
--+
| +----------------- Potamilus_fragilis **
| |
+--+ +-------------- Leaunio_lienosus **
| |
+--+ +-- Lampsilis_floridensis **
| +--------+
| | +-- Lampsilis_teres **
+--+
| +-------- Lampsilis_sietmani **
+--+
| +----- Lampsilis_siliquoidea **
+--+
| +-- Lampsilis_hydiana **
+--+
+-- Lampsilis_straminea **
Gomes-dos-Santos et al., 2019 [Publ 3943], fig. 1
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 10 |
10 |
8 |
2 |
0 |
0 |
0 |
79.0 |
+----- Potomida_littoralis **
+-----------+
| | +-- Unio_delphinus **
| +--+
| +-- Anodonta_anatina **
--+
| +-- Gibbosula_crassa **
| +-----------+
| | +-- Gibbosula_rochechouartii **
+--+
| +----------- Cumberlandia_monodonta **
+--+
| +-------- Pseudunio_marocanus **
+--+
| +----- Margaritifera_falcata **
+--+
| +-- Margaritifera_margaritifera **
+--+
+-- Margaritifera_dahurica **
Lopes-Lima et al., 2020 [Publ 4080], fig. 13
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 10 |
10 |
8 |
2 |
0 |
0 |
0 |
81.0 |
+-- Margaritifera_laevis **
+--------------------+
| +-- Margaritifera_middendorffi **
--+
| +-------------------- Middendorffinaia_mongolica **
+--+
| +----------------- Nodularia_breviconcha **
+--+
| +-------------- Nodularia[3] **
+--+
| +----------- Nodularia[2] **
+--+
| +-------- Nodularia_nuxpersicae **
+--+
| +----- Nodularia_nipponensis **
+--+
| +-- Nodularia_douglasiae **
+--+
+-- Nodularia[1] **
Pfeiffer et al., 2021 [Publ 4126], fig. 18
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 10 |
10 |
7 |
1 |
0 |
0 |
0 |
85.0 |
+-------------- Yaukthwa_paiensis **
|
| +----- Trapezoideus_foliaceus **
--+ +-----+
| | | +-- Trapezoideus_lenya **
| | +--+
| | +-- Trapezoideus_subviridis **
+--+
| +-- Physunio_massini **
| +-----+
| | +-- Physunio_superbus **
+--+
| +----- Physunio_modelli[3] **
| |
+--+ +-- Physunio_modelli[1] **
| |
+--+-- Physunio_modelli[2] **
|
+-- Physunio_pellucidus **
Kano et al., 2019 [Publ 4194], fig. 6
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 10 |
10 |
8 |
2 |
0 |
0 |
0 |
82.0 |
+-------------- Margaritifera_falcata **
|
| +----- Sinanodonta_lauta **
--+ +-----+
| | | +-- Pletholophus_tenuis **
| | +--+
| | +-- Beringiana_japonica **
+--+
| +----- Inversidens_brandtii **
| +--+
| | | +-- Obovalis_omiensis **
| | +--+
+--+ +-- Pronodularia_japanensis **
|
| +----- Lanceolaria_grayii **
+--+
| +-- Inversiunio_yanagawensis **
+--+
+-- Nodularia_nipponensis **
Lopes-Lima et al., 2021 [Publ 4217], fig. 2
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 10 |
10 |
8 |
1 |
0 |
0 |
0 |
84.0 |
+----------------- Unio_tumidus **
|
--+ +-------------- Unio_bruguierianus **
| |
| | +-- Unio_delicatus **
+--+ +--------+
| | +-- Unio_terminalis **
| |
+--+ +-- Unio_tigridis **
| +-----+
| | +-- Unio_hueti **
| |
+--+ +-- Unio_pictorum **
| +--+
| | +-- Unio_eucirrus **
+--+
| +-- Unio_elongatulus **
+--+
+-- Unio_mancus **
Sonowal et al., 2021 [Publ 4221], fig. 4
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 10 |
10 |
8 |
1 |
6 |
3 |
1 |
86.0 |
+----- Triplodon_corrugatus
+-----------------------+
| | +-- Velesunio_ambiguus
| +--+
--+ +-- Hyridella_australis
|
| +-------------------------- Lamellidens_mainwaringi **
+--+
| +----------------------- Lamprotula_leaii **
+--+
| +-------------------- Potomida_littoralis **
+--+
| +----------------- Lampsilis_cardium **
+--+
| +-------------- Elliptio_complanata **
+--+
| +----------- Aculamprotula_tientsinensis **
+--+
| +-------- Unio_pictorum **
+--+
| +----- Anodonta_cygnea **
+--+
| +-- Unio_ravoisieri **
+--+
+-- Lamellidens_corrianus **
Nagel et al., 1998 [Publ 1946], fig. 1
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 9 |
9 |
7 |
1 |
0 |
0 |
0 |
75.0 |
+----------------- Microcondylaea_bonellii **
|
--+ +-------------- Potomida_littoralis **
| |
| | +----- Pseudanodonta_complanata **
+--+ +-----+
| | | +-- Anodonta_anatina **
| | +--+
+--+ +-- Anodonta_cygnea **
|
| +-------- Unio_crassus **
+--+
| +----- Unio_tumidus **
+--+
| +-- Unio_pictorum **
+--+
+-- Unio_mancus **
King et al., 1999 [Publ 2699], fig. 2
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 9 |
9 |
7 |
1 |
0 |
0 |
0 |
81.0 |
+-- Pyganodon_grandis **
+-----------+
| +-- Pyganodon_fragilis **
|
--+ +----- Lasmigona_costata **
| +-----+
| | | +-- Lasmigona_complanata **
| | +--+
+--+ +-- Lasmigona_alabamensis **
|
| +-------- Strophitus_undulatus **
+--+
| +----- Prolasmidonta_heterodon **
+--+
| +-- Platynaias_subviridis **
+--+
+-- Platynaias_compressa **
King et al., 1999 [Publ 2699], fig. 4
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 9 |
9 |
3 |
1 |
0 |
0 |
0 |
80.0 |
+-- Lasmigona_complanata **
+-----+
| +-- Lasmigona_costata **
|
--+ +----- Prolasmidonta_heterodon **
| |
| +----- Strophitus_undulatus **
| |
+--+ +-- Platynaias_compressa **
| |
| +-- Platynaias_subviridis[1] **
| |
+--+-- Platynaias_subviridis[2] **
|
+-- Platynaias_subviridis[3] **
|
+-- Platynaias_subviridis[4] **
Cyr et al., 2007 [Publ 2745], fig. 2C
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 9 |
9 |
4 |
1 |
0 |
0 |
0 |
79.0 |
+----------- Pseudanodonta_complanata **
|
| +-- Pyganodon_grandis **
--+ +-----+
| | +-- Pyganodon[1] **
| |
+--+ +----- Pyganodon[3] **
| |
| | +-- Pyganodon_cataracta[1] **
+--+ |
| +-- Pyganodon_fragilis[1] **
| |
+--+-- Pyganodon[2] **
|
+-- Pyganodon_cataracta[2] **
|
+-- Pyganodon_fragilis[2] **
Soroka, 2010 [Publ 3404], fig. 3a
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 9 |
9 |
7 |
1 |
0 |
0 |
0 |
78.0 |
+-- Sinanodonta_woodiana **
+-----------+
| +-- Sinanodonta_lauta **
|
--+ +----- Anodonta_anatina **
| +-----+
| | | +-- Anodonta_cygnea **
| | +--+
+--+ +-- Pseudanodonta_complanata **
|
| +-------- Unio_tumidus **
+--+
| +----- Unio_crassus **
+--+
| +-- Unio_pictorum **
+--+
+-- Unio_mancus **
Soroka, 2010 [Publ 3404], fig. 3b
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 9 |
9 |
5 |
1 |
0 |
0 |
0 |
78.0 |
+-------------- Anodonta_anatina **
|
+-------------- Anodonta_cygnea **
|
--+-------------- Pseudanodonta_complanata **
|
| +-- Sinanodonta_woodiana **
| +--------+
| | +-- Sinanodonta_lauta **
+--+
| +-------- Unio_tumidus **
+--+
| +----- Unio_crassus **
+--+
| +-- Unio_pictorum **
+--+
+-- Unio_mancus **
Inoue et al., 2013 [Publ 3579], fig. 3
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 9 |
9 |
7 |
2 |
0 |
0 |
0 |
81.0 |
+----------------- Pyganodon_grandis **
|
--+ +-------------- Cumberlandia_monodonta **
| |
| | +-- Pseudunio_auricularius **
+--+ +--------+
| | +-- Pseudunio_marocanus **
| |
+--+ +-- Margaritifera_dahurica **
| +-----+
| | +-- Margaritifera_margaritifera **
+--+
| +----- Margaritifera_falcata **
+--+
| +-- Margaritifera_laevis **
+--+
+-- Margaritifera_marrianae **
Bolotov et al., 2015 [Publ 3646], fig. 2B
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 9 |
9 |
7 |
2 |
0 |
0 |
0 |
81.0 |
+----------------- Unio_pictorum **
|
--+ +-------------- Cumberlandia_monodonta **
| |
| | +-- Pseudunio_auricularius **
+--+ +--------+
| | +-- Pseudunio_marocanus **
| |
+--+ +-- Margaritifera_dahurica **
| +-----+
| | +-- Margaritifera_margaritifera **
+--+
| +----- Margaritifera_falcata **
+--+
| +-- Margaritifera_middendorffi **
+--+
+-- Margaritifera_laevis **
Takeuchi et al., 2015 [Publ 3661], fig. 2
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 9 |
9 |
7 |
2 |
1 |
0 |
0 |
81.0 |
+-------------------- Lampsilis_cardium **
|
--+ +----------------- Pseudunio_marocanus **
| |
+--+ +-------------- Pseudunio_auricularius **
| |
+--+ +----------- Cumberlandia_monodonta **
| |
+--+ +-- Margaritifera_dahurica **
| +-----+
| | +-- Margaritifera_margaritifera **
+--+
| +----- Margaritifera_falcata **
+--+
| +-- Margaritifera_middendorffi **
+--+
+-- Margaritifera_laevis **
Hewitt et al., 2018 [Publ 3758], fig. 5
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 9 |
9 |
6 |
2 |
0 |
0 |
0 |
79.0 |
+-------------- Margaritifera_margaritifera **
|
| +-- Fusconaia_flava **
--+ +--------+
| | +-- Pleurobema_rubellum **
| |
+--+ +-------- Pyganodon_grandis **
| |
| | +-- Platynaias_subviridis **
+--+-----+
| +-- Strophitus_undulatus **
|
| +----- Lasmigona_costata **
+--+
| +-- Arcidens_wheeleri **
+--+
+-- Lasmigona_complanata **
Fonseca et al., 2016 [Publ 3775], fig. 1-M
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 9 |
9 |
7 |
1 |
0 |
0 |
0 |
75.0 |
+-- Venustaconcha_ellipsiformis **
+-----------+
| +-- Quadrula_quadrula **
|
--+ +----- Sinosolenaia_carinata **
| +-----+
| | | +-- Potomida_littoralis **
| | +--+
+--+ +-- Pronodularia_japanensis **
|
| +-------- Unio_delphinus **
+--+
| +----- Anodonta_anatina **
+--+
| +-- Utterbackia_peninsularis **
+--+
+-- Pyganodon_grandis **
Chase et al., 2018 [Publ 3847], fig. 5
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 9 |
10 |
8 |
2 |
1 |
1 |
1 |
80.0 |
+-- Cumberlandia_monodonta **
+-----------+
| +-- Echyridella_menziesii
|
--+ +----------- Sinosolenaia_carinata **
| |
| | +----- Potamilus_alatus **
+--+ +--+
| | | +-- Lampsilis_siliquoidea **
| | +--+
| | +-- Lampsilis_powellii **
+--+
| +-- Anodonta_anatina **
| +--+
| | +-- Utterbackia_peninsularis **
+--+
| +-- Unio_delphinus **
+--+
+-- Unio_tumidus **
Lopes-Lima et al., 2020 [Publ 4080], fig. 12
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 9 |
9 |
6 |
2 |
0 |
0 |
0 |
83.0 |
+-- Margaritifera_laevis **
+-----------+
| +-- Margaritifera_middendorffi **
|
--+ +-- Nodularia_breviconcha **
| +--------+
| | +-- Nodularia_douglasiae **
+--+
| +-------- Inversiunio_jokohamensis **
| |
+--+ +----- Inversiunio_verrucosus[1] **
| |
+--+----- Inversiunio_verrucosus[2] **
|
| +-- Inversiunio_reinianus **
+--+
+-- Inversiunio_yanagawensis **
Lopes-Lima et al., 2020 [Publ 4080], fig. 16
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 9 |
9 |
7 |
2 |
0 |
0 |
0 |
79.0 |
+-- Margaritifera_laevis **
+-----------------+
| +-- Margaritifera_middendorffi **
--+
| +----------------- Chamberlainia_somsakpanhai **
+--+
| +-------------- Sinohyriopsis_schlegelii **
+--+
| +----------- Pronodularia_japanensis **
+--+
| +-------- Lamprotula_caveata **
+--+
| +----- Lamprotula_leaii[2] **
+--+
| +-- Lamprotula_seomjinensis **
+--+
+-- Lamprotula_leaii[1] **
Lopes-Lima et al., 2020 [Publ 4080], fig. 5
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 9 |
9 |
7 |
2 |
0 |
0 |
0 |
82.0 |
+-- Margaritifera_laevis **
+--------------+
| +-- Margaritifera_middendorffi **
|
--+ +-- Nodularia_breviconcha **
| +-----------+
| | +-- Nodularia_douglasiae **
+--+
| +----------- Aculamprotula_tortuosa **
+--+
| +-------- Aculamprotula_tientsinensis **
+--+
| +----- Aculamprotula_fibrosa **
+--+
| +-- Aculamprotula_scripta **
+--+
+-- Aculamprotula_coreana **
Pfeiffer et al., 2021 [Publ 4126], fig. 25
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 9 |
9 |
7 |
1 |
0 |
0 |
0 |
86.0 |
+-------------- Solenaia_emarginata **
|
| +-- Yaukthwa_elongatula **
--+ +--------+
| | +-- Yaukthwa_baniensis **
| |
+--+ +----- Yaukthwa_panhai **
| +--+
| | | +-- Yaukthwa_rectangularis **
| | +--+
+--+ +-- Yaukthwa_nesemanni **
|
| +----- Yaukthwa_avaensis **
+--+
| +-- Yaukthwa_inlenensis **
+--+
+-- Yaukthwa_paiensis **
Choi et al., 2020 [Publ 4169], fig. 5
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 9 |
9 |
7 |
1 |
0 |
0 |
0 |
81.0 |
+----- Radiatula_mouhoti **
+--------+
| | +-- Lamellidens_generosus **
| +--+
--+ +-- Leoparreysia_canefrii **
|
| +----------- Nodularia_breviconcha **
| |
+--+ +-- Nodularia[1] **
| +-----+
| | +-- Nodularia[2] **
+--+
| +----- Nodularia_douglasiae **
+--+
| +-- Nodularia_nipponensis **
+--+
+-- Nodularia_nuxpersicae **
Lopes-Lima et al., 2021 [Publ 4217], fig. 14
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 9 |
9 |
4 |
1 |
0 |
0 |
0 |
80.0 |
+-- Gonidea_angulata **
+--------+
| +-- Microcondylaea_bonellii **
|
--+ +-------- Leguminaia_anatolica **
| |
| | +----- Leguminaia_wheatleyi[2] **
+--+ |
| +----- Leguminaia_wheatleyi[3] **
| |
+--+----- Leguminaia_wheatleyi[4] **
|
+----- Leguminaia_wheatleyi[5] **
|
| +-- Leguminaia_saulcyi **
+--+
+-- Leguminaia_wheatleyi[1] **
Sonowal et al., 2021 [Publ 4221], fig. 3a
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 9 |
15 |
7 |
1 |
6 |
3 |
1 |
84.0 |
+----------- Lamellidens_corrianus **
|
| +-- Lamprotula_leaii **
+--------+
| +-- Potomida_littoralis **
|
| +----- Elliptio_complanata **
| |
--+ +----- Lampsilis_cardium **
| +--+
| | | +-- Anodonta_cygnea **
| | | |
| | | +-- Unio_ravoisieri **
| | +--+
| | +-- Unio_pictorum **
| | |
+--+ +-- Aculamprotula_tientsinensis **
|
| +-- Triplodon_corrugatus
| |
| +--+-- Hyridella_australis
| | |
| | +-- Velesunio_ambiguus
+--+
| +-- Neotrigonia_bednalli
| |
+--+-- Neotrigonia_lamarckii
|
+-- Neotrigonia_margaritacea
Sonowal et al., 2021 [Publ 4221], fig. 3b
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 9 |
9 |
0 |
1 |
6 |
3 |
1 |
79.0 |
+-- Triplodon_corrugatus
|
+--+-- Velesunio_ambiguus
| |
| +-- Hyridella_australis
|
| +-- Aculamprotula_tientsinensis **
| |
--+ +-- Lamprotula_leaii **
| |
| +-- Lampsilis_cardium **
| |
| +-- Potomida_littoralis **
| |
+--+-- Unio_pictorum **
|
+-- Anodonta_cygnea **
|
+-- Unio_ravoisieri **
|
+-- Lamellidens_mainwaringi **
|
+-- Elliptio_complanata **
Bolotov et al., 2022 [Publ 4250], fig. S4
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 9 |
9 |
6 |
1 |
0 |
0 |
0 |
85.0 |
+-------- Lamellidens_savadiensis **
+--+
| | +----- Lamellidens_ferrugineus **
| +--+
| | +-- Lamellidens_mainwaringi **
--+ +--+
| +-- Lamellidens_generosus **
|
| +-------- Trapezidens_scutum **
| |
+--+ +----- Trapezidens_exolescens **
| |
+--+ +-- Trapezidens_dolichorhynchus **
| |
+--+-- Trapezidens_angustior **
|
+-- Trapezidens_yeti **
Araujo et al., 2009 [Publ 2395], fig. 3
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 8 |
8 |
6 |
2 |
0 |
0 |
0 |
80.5 |
+-------------- Unio **
|
--+ +----------- Cumberlandia_monodonta **
| |
| | +-- Pseudunio_marocanus **
+--+ +-----+
| | +-- Pseudunio_auricularius **
| |
+--+ +-- Margaritifera_margaritifera **
| +--+
| | +-- Margaritifera_dahurica **
+--+
| +-- Margaritifera_falcata **
+--+
+-- Margaritifera_middendorffi **
Hoeh et al., 2002 [Publ 2673], fig. 1-F
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 8 |
8 |
6 |
1 |
1 |
1 |
1 |
80.0 |
+-------------------- Echyridella_menziesii
|
--+ +-- Pyganodon_fragilis **
| +--------------+
| | +-- Pyganodon_grandis **
+--+
| +-------------- Fusconaia_flava **
+--+
| +----------- Cyrtonaias_tampicoensis **
+--+
| +-------- Ortmanniana_ligamentina **
+--+
| +----- Lampsilis_teres **
+--+
| +-- Potamilus_purpuratus **
+--+
+-- Ligumia_recta **
Hoeh et al., 2002 [Publ 2673], fig. 1-M
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 8 |
8 |
6 |
1 |
1 |
1 |
1 |
80.0 |
+-------------------- Echyridella_menziesii
|
--+ +-- Pyganodon_fragilis **
| +--------------+
| | +-- Pyganodon_grandis **
+--+
| +-------------- Fusconaia_flava **
+--+
| +----------- Potamilus_purpuratus **
+--+
| +-------- Cyrtonaias_tampicoensis **
+--+
| +----- Ligumia_recta **
+--+
| +-- Lampsilis_teres **
+--+
+-- Ortmanniana_ligamentina **
Soroka, 2010 [Publ 3404], fig. 5a
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 8 |
8 |
6 |
1 |
0 |
0 |
0 |
79.0 |
+----- Unio_crassus **
+-----+
| | +-- Unio_pictorum **
| +--+
--+ +-- Unio_tumidus **
|
| +-------- Anodonta_anatina **
| |
+--+ +-- Anodonta_cygnea **
| +--+
| | +-- Pseudanodonta_complanata **
+--+
| +-- Sinanodonta_woodiana **
+--+
+-- Sinanodonta_schrenkii **
Soroka, 2010 [Publ 3404], fig. 5b
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 8 |
8 |
5 |
1 |
0 |
0 |
0 |
78.0 |
+----------- Anodonta_anatina **
|
| +-- Anodonta_cygnea **
--+ +-----+
| | +-- Pseudanodonta_complanata **
| |
| | +-- Sinanodonta_woodiana **
+--+-----+
| +-- Sinanodonta_schrenkii **
|
| +----- Unio_tumidus **
+--+
| +-- Unio_pictorum **
+--+
+-- Unio_crassus **
Inoue et al., 2014 [Publ 3635], fig. 3B
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 8 |
8 |
3 |
1 |
0 |
0 |
0 |
82.0 |
+----------- Amblema_plicata **
|
| +-------- Pyganodon_grandis **
--+ |
| +-------- Pyganodon_cataracta **
| |
+--+-------- Pyganodon_fragilis **
|
+-------- Pyganodon **
|
| +----- Platynaias_subviridis **
+--+
| +-- Arcidens_confragosus **
+--+
+-- Arcidens_wheeleri **
Froufe et al., 2015 [Publ 3664], fig. 1-M
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 8 |
8 |
6 |
1 |
0 |
0 |
0 |
73.0 |
+----- Anodonta_anatina **
+-----+
| | +-- Utterbackia_peninsularis **
| +--+
| +-- Pyganodon_grandis **
--+
| +-- Venustaconcha_ellipsiformis **
| +-----+
| | +-- Quadrula_quadrula **
+--+
| +----- Sinosolenaia_carinata **
+--+
| +-- Potomida_littoralis **
+--+
+-- Pronodularia_japanensis **
Zieritz et al., 2012 [Publ 3673], fig. 4
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 8 |
8 |
4 |
2 |
0 |
0 |
0 |
77.0 |
+-- Pseudanodonta_complanata **
|
+--------+-- Anodonta_anatina **
| |
| +-- Anodonta_cygnea **
--+
| +-------- Sinanodonta_woodiana **
| |
+--+-------- Margaritifera_margaritifera **
|
| +----- Unio_crassus **
+--+
| +-- Unio_tumidus **
+--+
+-- Unio_pictorum **
Sheng et al., 2014 [Publ 3704], fig. 2
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 8 |
8 |
6 |
1 |
0 |
0 |
0 |
79.0 |
+----------------- Cristaria_plicata **
|
--+ +-------------- Pyganodon_grandis **
| |
+--+ +----------- Lampsilis_ornata **
| |
+--+ +-- Venustaconcha_ellipsiformis **
| +-----+
| | +-- Quadrula_quadrula **
+--+
| +----- Lamprotula_leaii **
+--+
| +-- Sinohyriopsis_cumingii **
+--+
+-- Sinohyriopsis_schlegelii **
Bolotov et al., 2016 [Publ 3732], fig. 2
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 8 |
8 |
6 |
2 |
0 |
0 |
0 |
82.0 |
+-- Gibbosula_laosensis **
+--------------+
| +-- Margaritifera_dahurica **
--+
| +-------------- Sinanodonta_lucida **
+--+
| +----------- Sinanodonta_schrenkii **
+--+
| +-------- Sinanodonta_woodiana[2] **
+--+
| +----- Sinanodonta_elliptica **
+--+
| +-- Sinanodonta_woodiana[1] **
+--+
+-- Sinanodonta_lauta **
Pfeiffer et al., 2018 [Publ 3827], fig. 1
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 8 |
8 |
3 |
1 |
0 |
0 |
0 |
76.0 |
+----- Physunio **
|
+--+----- Trapezoideus **
| |
| | +-- Lens_comptus **
| +--+
--+ +-- Lens_contradens **
|
| +-- Ensidens **
| |
| +-- Hyriopsis[1] **
+-----+
+-- Hyriopsis[2] **
|
+-- Rectidens **
Marrone et al., 2019 [Publ 3907], fig. 1
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 8 |
8 |
4 |
1 |
0 |
0 |
0 |
82.0 |
+-------------- Unio_tumidus **
|
--+ +----------- Unio_delphinus **
| |
+--+ +-------- Unio_foucauldianus **
| |
+--+ +----- Unio_mancus **
| |
| +----- Unio_elongatulus **
+--+
+----- Unio_ravoisieri **
|
| +-- Unio_pictorum **
+--+
+-- Unio_tigridis **
Kondakov et al., 2018 [Publ 3914], fig. 5
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 8 |
8 |
6 |
1 |
0 |
0 |
0 |
79.0 |
+-------------------- Anemina_arcaeformis **
--+
| +----------------- Sinanodonta_lucida **
+--+
| +-------------- Sinanodonta_schrenkii **
+--+
| +----------- Sinanodonta_woodiana[2] **
+--+
| +-------- Sinanodonta_jourdyi **
+--+
| +----- Sinanodonta_elliptica **
+--+
| +-- Sinanodonta_woodiana[1] **
+--+
+-- Sinanodonta_lauta **
Keogh & Simons, 2019 [Publ 3935], fig. 4
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 8 |
8 |
6 |
1 |
0 |
0 |
0 |
84.0 |
+----------------- Lampsilis_sietmani **
|
--+ +-- Lampsilis_floridensis **
| +-----------+
| | +-- Lampsilis_teres **
+--+
| +----------- Lampsilis_siliquoidea **
+--+
| +-------- Lampsilis_hydiana[2] **
+--+
| +----- Lampsilis_straminea[2] **
+--+
| +-- Lampsilis_hydiana[1] **
+--+
+-- Lampsilis_straminea[1] **
Klishko et al., 2018 [Publ 3945], fig. 8
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 8 |
8 |
5 |
1 |
0 |
0 |
0 |
76.0 |
+-------- Nodularia_breviconcha **
+--+
| | +----- Nodularia **
| +--+
| | +-- Nodularia_douglasiae **
--+ +--+
| +-- Nodularia_nipponensis **
|
| +----- Unio_tumidus **
| |
+-----+----- Unio_gibbus **
|
| +-- Unio_crassus **
+--+
+-- Unio_pictorum **
Lopes-Lima et al., 2020 [Publ 4080], fig. 7
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 8 |
8 |
6 |
2 |
0 |
0 |
0 |
84.0 |
+-- Margaritifera_laevis **
+-----------+
| +-- Margaritifera_dahurica **
|
--+ +-- Sinanodonta_woodiana **
| +--------+
| | +-- Sinanodonta_tumens **
+--+
| +-------- Beringiana_gosannensis **
+--+
| +----- Beringiana_fukuharai **
+--+
| +-- Beringiana_beringiana **
+--+
+-- Beringiana_japonica **
Tomilova et al., 2020 [Publ 4097], fig. 3
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 8 |
8 |
5 |
1 |
0 |
0 |
0 |
78.0 |
+-- Unio_tumidus **
+--------+
| +-- Unio_pictorum **
|
| +-- Sinanodonta_lauta **
--+--------+
| +-- Sinanodonta_woodiana **
|
| +-------- Anodonta_anatina **
+--+
| +----- Pseudanodonta_complanata **
+--+
| +-- Anodonta_cygnea **
+--+
+-- Anodonta_exulcerata **
Goncalves et al., 2022 [Publ 4253], fig. 2
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 8 |
8 |
6 |
1 |
0 |
0 |
0 |
80.0 |
+-------------- Lamprotula_caveata **
|
| +----- Chamberlainia_somsakpanhai **
--+ +-----+
| | | +-- Chamberlainia_hainesiana **
| | +--+
+--+ +-- Chamberlainia_duclerci **
|
| +-------- Sinohyriopsis_cumingii[2] **
+--+
| +----- Sinohyriopsis_cumingii[1] **
+--+
| +-- Sinohyriopsis_schlegelii **
+--+
+-- Sinohyriopsis_goliath **
Jones & Neves, 2010 [Publ 2646], fig. 16
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 7 |
7 |
5 |
1 |
0 |
0 |
0 |
84.0 |
+-- Epioblasma_triquetra **
+-----------+
| +-- Epioblasma_brevidens **
--+
| +----------- Epioblasma_rangiana **
+--+
| +-------- Epioblasma_capsaeformis **
+--+
| +----- Epioblasma_ahlstedti **
+--+
| +-- Epioblasma_aureola **
+--+
+-- Epioblasma_walkeri **
Jones & Neves, 2010 [Publ 2646], fig. 17
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 7 |
7 |
1 |
1 |
0 |
0 |
0 |
84.0 |
+----- Epioblasma_triquetra **
|
+----- Epioblasma_brevidens **
|
--+ +-- Epioblasma_capsaeformis **
| |
| +-- Epioblasma_aureola **
| |
+--+-- Epioblasma_walkeri **
|
+-- Epioblasma_rangiana **
|
+-- Epioblasma_ahlstedti **
Graf, 2000 [Publ 2647], fig. 1
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 7 |
17 |
14 |
2 |
11 |
10 |
2 |
88.0 |
+----------------------- Margaritifera **
|
| +-------- Unio **
| +-----------+
--+ | | +----- Elliptio **
| | +--+
| | | +-- Pyganodon **
| | +--+
+--+ +-- Lampsilis **
|
| +----------------- Parreysia **
| |
| | +-------------- Grandidieria **
+--+ |
| | +-------- Castalia
| | +--+
+--+ | | +----- Diplodon
| | +--+
| | | +-- Hyridella
| | +--+
+--+ +-- Velesunio
|
| +-- Mutela[1]
| +-----+
| | +-- Mutela[2]
+--+
| +----- Anodontites
| |
+--+----- Mycetopoda
|
| +-- Etheria
+--+
+-- Mulleria
Buhay et al., 2002 [Publ 2661], fig. 1
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 7 |
7 |
3 |
1 |
0 |
0 |
0 |
83.0 |
+----------- Lampsilis_siliquoidea[1] **
|
+----------- Lampsilis_siliquoidea[2] **
--+
| +-------- Epioblasma_brevidens **
| |
+--+ +----- Epioblasma_triquetra **
| |
+--+ +-- Epioblasma_capsaeformis **
| |
+--+-- Epioblasma_walkeri[1] **
|
+-- Epioblasma_walkeri[2] **
Källersjö et al., 2005 [Publ 2684], fig. 1A
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 7 |
7 |
2 |
2 |
0 |
0 |
0 |
74.0 |
+----- Margaritifera_margaritifera **
|
| +-- Anodonta_anatina **
| |
+--+-- Pseudanodonta_complanata **
--+ |
| +-- Anodonta_cygnea **
|
| +-- Unio_crassus **
| |
+--+-- Unio_pictorum **
|
+-- Unio_tumidus **
Källersjö et al., 2005 [Publ 2684], fig. 1B-F
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 7 |
7 |
4 |
2 |
0 |
0 |
0 |
74.0 |
+-------- Margaritifera_margaritifera **
|
| +----- Anodonta_anatina **
+--+
--+ | +-- Pseudanodonta_complanata **
| +--+
| +-- Anodonta_cygnea **
|
| +----- Unio_tumidus **
+--+
| +-- Unio_crassus **
+--+
+-- Unio_pictorum **
Moyer & Díaz-Ferguson, 2012 [Publ 3375], fig. 3
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 7 |
7 |
5 |
1 |
0 |
0 |
0 |
81.0 |
+----- Lasmigona_costata **
+-----+
| | +-- Platynaias_compressa **
| +--+
--+ +-- Platynaias_subviridis **
|
| +-------- Lampsilis_ovata[2] **
+--+
| +----- Lampsilis_ovata[1] **
+--+
| +-- Lampsilis_virescens **
+--+
+-- Lampsilis_fasciola **
Hoeh et al., 1995 [Publ 3391], fig. 2
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 7 |
7 |
4 |
1 |
0 |
0 |
0 |
83.0 |
+----- Utterbackia_peggyae **
+--+
| | +-- Utterbackia_imbecillis **
| +--+
--+ +-- Utterbackia_peninsularis **
|
| +----- Pyganodon_lacustris **
| |
+--+----- Pyganodon_grandis **
|
| +-- Pyganodon_cataracta **
+--+
+-- Pyganodon_fragilis **
Soroka, 2010 [Publ 3404], fig. 2
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 7 |
7 |
4 |
1 |
0 |
0 |
0 |
77.0 |
+----------- Sinanodonta_woodiana **
|
| +-- Unio_tumidus **
--+ |
| +-----+-- Unio_pictorum **
| | |
+--+ +-- Unio_crassus **
|
| +----- Anodonta_cygnea **
+--+
| +-- Anodonta_anatina **
+--+
+-- Pseudanodonta_complanata **
Soroka, 2010 [Publ 3404], fig. 4
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 7 |
7 |
5 |
1 |
0 |
0 |
0 |
78.0 |
+----- Sinanodonta_woodiana **
+-----+
| | +-- Anodonta_cygnea **
| +--+
--+ +-- Pseudanodonta_complanata **
|
| +-------- Anodonta_anatina **
+--+
| +----- Unio_tumidus **
+--+
| +-- Unio_pictorum **
+--+
+-- Unio_crassus **
Chen et al., 2012 [Publ 3624], fig. 4
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 7 |
7 |
5 |
1 |
0 |
0 |
0 |
76.0 |
+----- Quadrula_quadrula **
+--+
| | +-- Lampsilis_ornata **
| +--+
| +-- Venustaconcha_ellipsiformis **
--+
| +-- Pyganodon_grandis **
| +--+
| | +-- Cristaria_plicata **
+--+
| +-- Sinohyriopsis_cumingii **
+--+
+-- Lamprotula_leaii **
Froufe et al., 2016 [Publ 3727], fig. 3, 4
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 7 |
7 |
3 |
1 |
0 |
0 |
0 |
79.0 |
+----- Potomida_littoralis **
+--+
| | +-- Potomida_acarnanica **
| +--+
| +-- Potomida_semirugata **
--+
| +-- Pronodularia_japanensis **
| |
| +-- Lamprotula_leaii **
+-----+
+-- Leguminaia_wheatleyi **
|
+-- Microcondylaea_bonellii **
Soroka & Burzynski, 2018 [Publ 3762], fig. 3
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 7 |
7 |
5 |
1 |
0 |
0 |
0 |
74.0 |
+-- Venustaconcha_ellipsiformis **
+--------+
| +-- Quadrula_quadrula **
|
--+ +-- Unio_tumidus **
| +-----+
| | +-- Aculamprotula_tortuosa **
+--+
| +----- Anodonta_anatina **
+--+
| +-- Pyganodon_grandis **
+--+
+-- Utterbackia_peninsularis **
Soroka & Burzynski, 2018 [Publ 3762], fig. 3-M
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 7 |
7 |
5 |
1 |
0 |
0 |
0 |
74.0 |
+-- Venustaconcha_ellipsiformis **
+--------+
| +-- Quadrula_quadrula **
|
--+ +-- Unio_tumidus **
| +-----+
| | +-- Aculamprotula_tortuosa **
+--+
| +----- Anodonta_anatina **
+--+
| +-- Pyganodon_grandis **
+--+
+-- Utterbackia_peninsularis **
Guerra et al., 2017 [Publ 3772], fig. 2a-M
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 7 |
7 |
4 |
2 |
1 |
1 |
1 |
75.0 |
+----------- Echyridella_menziesii
|
--+ +-------- Cumberlandia_monodonta **
| |
| | +-- Sinosolenaia_carinata **
+--+ +--+
| | +-- Pronodularia_japanensis **
| |
| | +-- Utterbackia_peninsularis **
+--+--+
| +-- Pyganodon_grandis **
|
| +-- Venustaconcha_ellipsiformis **
+--+
+-- Quadrula_quadrula **
Guerra et al., 2017 [Publ 3772], fig. 2b-M
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 7 |
7 |
4 |
2 |
1 |
1 |
1 |
75.0 |
+----------- Echyridella_menziesii
|
--+ +-------- Cumberlandia_monodonta **
| |
| | +-- Sinosolenaia_carinata **
+--+ +--+
| | +-- Pronodularia_japanensis **
| |
| | +-- Utterbackia_peninsularis **
+--+--+
| +-- Pyganodon_grandis **
|
| +-- Venustaconcha_ellipsiformis **
+--+
+-- Quadrula_quadrula **
Amaro et al., 2017 [Publ 3791], fig. 3A-M
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 7 |
7 |
5 |
2 |
0 |
0 |
0 |
78.0 |
+-------------- Margaritifera_margaritifera **
|
--+ +----------- Quadrula_quadrula **
| |
+--+ +-- Pronodularia_japanensis **
| +-----+
| | +-- Sinosolenaia_carinata **
+--+
| +----- Venustaconcha_ellipsiformis **
+--+
| +-- Pyganodon_grandis **
+--+
+-- Utterbackia_peninsularis **
Amaro et al., 2017 [Publ 3791], fig. 3B-M
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 7 |
7 |
5 |
2 |
0 |
0 |
0 |
78.0 |
+-------------- Margaritifera_margaritifera **
|
--+ +----------- Quadrula_quadrula **
| |
+--+ +-------- Venustaconcha_ellipsiformis **
| |
+--+ +-- Utterbackia_peninsularis **
| +--+
| | +-- Pyganodon_grandis **
+--+
| +-- Pronodularia_japanensis **
+--+
+-- Sinosolenaia_carinata **
Lopes-Lima et al., 2020 [Publ 4080], fig. 8
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 7 |
7 |
5 |
2 |
0 |
0 |
0 |
80.0 |
+-- Margaritifera_laevis **
+--------+
| +-- Margaritifera_middendorffi **
|
--+ +-- Sinanodonta_woodiana **
| +-----+
| | +-- Sinanodonta_lucida **
+--+
| +----- Cristaria_clessini **
+--+
| +-- Cristaria_plicata **
+--+
+-- Cristaria_radiata **
Pfeiffer et al., 2021 [Publ 4126], fig. 30
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 7 |
7 |
5 |
1 |
0 |
0 |
0 |
78.0 |
+-------------- Rectidens_sumatrensis **
|
--+ +-- Ensidens_telus **
| +--------+
| | +-- Ensidens_ingallsianus **
+--+
| +-------- Ensidens_dugasti **
+--+
| +----- Ensidens_sagittarius **
+--+
| +-- Ensidens_spiculus **
+--+
+-- Ensidens_jaculus **
Lopes-Lima et al., 2021 [Publ 4217], fig. 6
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 7 |
7 |
5 |
1 |
0 |
0 |
0 |
79.0 |
+----------------- Unio_tumidus **
--+
| +-------------- Unio_pictorum **
+--+
| +----------- Unio_sesirmensis **
+--+
| +-------- Unio_damascensis **
+--+
| +----- Unio_courtillieri **
+--+
| +-- Unio_bruguierianus **
+--+
+-- Unio_crassus **
Bashê, 2022 [Publ 4265], fig. 1
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 7 |
7 |
5 |
2 |
0 |
0 |
0 |
82.0 |
+----------------- Pseudunio_homsensis **
--+
| +-------------- Unio_pictorum[2] **
+--+
| +----------- Unio_foucauldianus **
+--+
| +-------- Unio_delphinus **
+--+
| +----- Unio_tigridis **
+--+
| +-- Unio_elongatulus **
+--+
+-- Unio_pictorum[1] **
Smith et al., 2023 [Publ 4291], fig. 2
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 7 |
7 |
5 |
1 |
0 |
0 |
0 |
83.0 |
+-- Anodontoides_ferussacianus **
+--------+
| +-- Anodontoides_denigratus **
|
--+ +-- Strophitus_undulatus **
| +-----+
| | +-- Strophitus_howellsi **
+--+
| +----- Strophitus_pascagoulaensis **
+--+
| +-- Strophitus_radiatus **
+--+
+-- Strophitus_williamsi **
Bogan et al., 2023 [Publ 4295], fig. 4
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 7 |
7 |
5 |
2 |
0 |
0 |
0 |
79.0 |
+-------------- Gibbosula_crassa **
|
--+ +-- Sinanodonta_jourdyi **
| +--------+
| | +-- Pletholophus_honglinhensis **
+--+
| +-------- Cristaria_radiata **
+--+
| +----- Cristaria_clessini **
+--+
| +-- Cristaria_plicata **
+--+
+-- Cristaria_truncata **
Nagel et al., 1998 [Publ 1946], fig. 7
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 6 |
6 |
4 |
1 |
0 |
0 |
0 |
80.0 |
+-------------- Pseudanodonta_complanata **
--+
| +----------- Anodonta_cygnea **
+--+
| +-------- Sinanodonta_woodiana **
+--+
| +----- Anodonta[2] **
+--+
| +-- Anodonta_anatina **
+--+
+-- Anodonta[1] **
Nagel et al., 1998 [Publ 1946], fig. 9
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 6 |
6 |
4 |
2 |
0 |
0 |
0 |
77.0 |
+-------------- Margaritifera **
--+
| +----------- Potomida **
+--+
| +-------- Microcondylaea **
+--+
| +----- Unio **
+--+
| +-- Anodonta **
+--+
+-- Pseudanodonta **
Jones et al., 2006 [Publ 2681], fig. 2
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 6 |
6 |
4 |
1 |
0 |
0 |
0 |
82.0 |
+-- Epioblasma_triquetra **
+--------+
| +-- Epioblasma_brevidens **
--+
| +-------- Epioblasma_capsaeformis[2] **
+--+
| +----- Epioblasma_capsaeformis[1] **
+--+
| +-- Epioblasma_walkeri **
+--+
+-- Epioblasma_rangiana **
Cyr et al., 2007 [Publ 2745], fig. 2A
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 6 |
6 |
3 |
1 |
0 |
0 |
0 |
80.0 |
+----------- Lampsilis_ornata **
|
--+ +-------- Pyganodon_grandis **
| |
+--+-------- Pyganodon[2] **
|
| +----- Pyganodon[1] **
+--+
| +-- Pyganodon_cataracta **
+--+
+-- Pyganodon_fragilis **
Cyr et al., 2007 [Publ 2745], fig. 2B
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 6 |
6 |
4 |
1 |
0 |
0 |
0 |
79.0 |
+----------- Anodonta_nuttalliana **
|
--+ +-- Pyganodon_grandis **
| +-----+
| | +-- Pyganodon[1] **
+--+
| +----- Pyganodon[2] **
+--+
| +-- Pyganodon_cataracta **
+--+
+-- Pyganodon_fragilis **
Nagel et al., 1996 [Publ 3400], fig. 351
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 6 |
6 |
4 |
1 |
0 |
0 |
0 |
80.0 |
+-------------- Pseudanodonta_complanata **
--+
| +----------- Anodonta_cygnea **
+--+
| +-------- Sinanodonta_woodiana **
+--+
| +----- Anodonta[2] **
+--+
| +-- Anodonta_anatina **
+--+
+-- Anodonta[1] **
Soroka, 2010 [Publ 3404], fig. 3a-M
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 6 |
6 |
4 |
1 |
0 |
0 |
0 |
79.0 |
+----------- Anodonta_anatina **
|
--+ +-- Sinanodonta_woodiana **
| +-----+
| | +-- Sinanodonta_lauta **
+--+
| +----- Unio_crassus **
+--+
| +-- Unio_tumidus **
+--+
+-- Unio_pictorum **
Soroka, 2010 [Publ 3404], fig. 3b-M
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 6 |
6 |
4 |
1 |
0 |
0 |
0 |
77.0 |
+----------- Anodonta_anatina **
|
--+ +-- Sinanodonta_lauta **
| +-----+
| | +-- Sinanodonta_woodiana **
+--+
| +----- Unio_tumidus **
+--+
| +-- Unio_crassus **
+--+
+-- Unio_pictorum **
Soroka, 2010 [Publ 3404], fig. 8a
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 6 |
6 |
4 |
1 |
0 |
0 |
0 |
73.0 |
+----- Unio_tumidus **
+--+
| | +-- Unio_pictorum **
| +--+
--+ +-- Unio_crassus **
|
| +----- Sinanodonta_woodiana **
+--+
| +-- Anodonta_anatina **
+--+
+-- Anodonta_cygnea **
Soroka, 2010 [Publ 3404], fig. 8b
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 6 |
6 |
4 |
1 |
0 |
0 |
0 |
75.0 |
+----------- Sinanodonta_woodiana **
|
--+ +-- Anodonta_anatina **
| +-----+
| | +-- Anodonta_cygnea **
+--+
| +----- Unio_tumidus **
+--+
| +-- Unio_pictorum **
+--+
+-- Unio_crassus **
Prie et al., 2012 [Publ 3487], fig. 2
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 6 |
6 |
4 |
2 |
0 |
0 |
0 |
76.0 |
+-------------- Margaritifera_margaritifera **
--+
| +----------- Unio_tumidus **
+--+
| +-------- Unio_crassus **
+--+
| +----- Unio_pictorum **
+--+
| +-- Unio_mancus **
+--+
+-- Unio_elongatulus **
Huang et al., 2013 [Publ 3629], fig. 3-M
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 6 |
6 |
4 |
1 |
0 |
0 |
0 |
75.0 |
+-- Pyganodon_grandis **
+-----+
| +-- Utterbackia_peninsularis **
|
--+ +-- Sinosolenaia_carinata **
| +--+
| | +-- Pronodularia_japanensis **
+--+
| +-- Quadrula_quadrula **
+--+
+-- Venustaconcha_ellipsiformis **
Wei et al., 2016 [Publ 3665], fig. 0
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 6 |
6 |
2 |
1 |
0 |
0 |
0 |
79.0 |
+-- Sinohyriopsis_cumingii **
+--+
| +-- Sinohyriopsis_schlegelii **
|
--+ +-- Cristaria_plicata **
| |
| +-- Lampsilis_ornata **
+--+
+-- Pyganodon_grandis **
|
+-- Quadrula_quadrula **
Lane et al., 2016 [Publ 3735], fig. 3
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 6 |
6 |
1 |
1 |
0 |
0 |
0 |
83.0 |
+----- Paetulunio_fabalis **
|
+----- Epioblasma_capsaeformis **
--+
| +-- Venustaconcha_trabalis[1] **
| |
| +-- Venustaconcha_trabalis[2] **
+--+
+-- Venustaconcha_ellipsiformis **
|
+-- Venustaconcha_pleasii **
Wen et al., 2017 [Publ 3795], fig. 3-M
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 6 |
6 |
4 |
1 |
0 |
0 |
0 |
77.0 |
+----- Anodonta_anatina **
+--+
| | +-- Utterbackia_peninsularis **
| +--+
--+ +-- Pyganodon_grandis **
|
| +----- Quadrula_quadrula **
+--+
| +-- Potamilus_alatus **
+--+
+-- Venustaconcha_ellipsiformis **
Zieritz et al., 2016 [Publ 3804], fig. S2-1
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 6 |
6 |
3 |
1 |
0 |
0 |
0 |
77.0 |
+----------- Rectidens_sumatrensis **
|
--+ +-------- Hyriopsis_desowitzi **
| |
+--+ +----- Hyriopsis_khoratensis **
| |
+--+ +-- Hyriopsis_myersiana **
| |
+--+-- Hyriopsis_panhai **
|
+-- Hyriopsis_bialata **
Bogatov et al., 2018 [Publ 3859], fig. 1
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 6 |
6 |
4 |
2 |
0 |
0 |
0 |
74.0 |
+----------- Margaritifera_margaritifera **
|
--+ +-------- Unio_tumidus **
| |
+--+ +-- Anodonta_anatina **
| +--+
| | +-- Pseudanodonta_complanata **
+--+
| +-- Unio_crassus **
+--+
+-- Unio_pictorum **
Bogatov et al., 2018 [Publ 3859], fig. 2
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 6 |
6 |
4 |
2 |
0 |
0 |
0 |
76.0 |
+-------------- Margaritifera_margaritifera **
--+
| +----------- Unio_crassus **
+--+
| +-------- Unio_pictorum **
+--+
| +----- Unio_tumidus **
+--+
| +-- Anodonta_anatina **
+--+
+-- Pseudanodonta_complanata **
Gomes-dos-Santos et al., 2019 [Publ 3943], fig. 1-M
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 6 |
6 |
4 |
2 |
0 |
0 |
0 |
75.0 |
+----- Margaritifera_margaritifera **
+--+
| | +-- Pseudunio_marocanus **
| +--+
--+ +-- Cumberlandia_monodonta **
|
| +----- Potomida_littoralis **
+--+
| +-- Unio_delphinus **
+--+
+-- Anodonta_anatina **
Klishko et al., 2017 [Publ 3946], fig. 8
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 6 |
6 |
4 |
1 |
0 |
0 |
0 |
76.0 |
+-- Nodularia_douglasiae **
+--------+
| +-- Nodularia_breviconcha **
--+
| +-------- Unio_tumidus **
+--+
| +----- Unio_pictorum **
+--+
| +-- Unio_crassus **
+--+
+-- Unio_courtillieri **
Gomes-dos-Santos et al., 2019 [Publ 4042], fig. 8
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 6 |
6 |
4 |
1 |
0 |
0 |
0 |
76.0 |
+----- Anodonta_anatina **
+--+
| | +-- Anodonta_exulcerata **
| +--+
--+ +-- Anodonta_cygnea **
|
| +----- Cristaria_radiata **
+--+
| +-- Anemina_arcaeformis **
+--+
+-- Sinanodonta_woodiana **
Lopes-Lima et al., 2020 [Publ 4080], fig. 14
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 6 |
6 |
4 |
2 |
0 |
0 |
0 |
79.0 |
+-- Margaritifera_laevis **
+-----+
| +-- Margaritifera_middendorffi **
|
--+ +-- Parvasolenaia_rivularis **
| +--+
| | +-- Koreosolenaia_sitgyensis **
+--+
| +-- Lamprotula_leaii **
+--+
+-- Lamprotula_seomjinensis **
Lopes-Lima et al., 2020 [Publ 4080], fig. 9
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 6 |
6 |
4 |
2 |
0 |
0 |
0 |
78.0 |
+-- Margaritifera_laevis **
+--------+
| +-- Margaritifera_middendorffi **
--+
| +-------- Cristaria_plicata **
+--+
| +----- Sinanodonta_schrenkii **
+--+
| +-- Pletholophus_tenuis **
+--+
+-- Pletholophus_reinianus **
Riccardi et al., 2019 [Publ 4089], fig. 1
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 6 |
6 |
4 |
1 |
0 |
0 |
0 |
77.0 |
+-- Sinanodonta_woodiana **
+-----+
| +-- Anemina_arcaeformis **
|
--+ +-- Anodonta_anatina **
| +--+
| | +-- Pseudanodonta_complanata **
+--+
| +-- Anodonta_exulcerata **
+--+
+-- Anodonta_cygnea **
Mejia-Trujillo & Smith, 2022 [Publ 4274], fig. 2-M
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 6 |
6 |
4 |
1 |
0 |
0 |
0 |
81.0 |
+----------- Quadrula_quadrula **
|
--+ +-- Potamilus_streckersoni **
| +-----+
| | +-- Potamilus_alatus **
+--+
| +----- Venustaconcha_ellipsiformis **
+--+
| +-- Lampsilis_powellii **
+--+
+-- Lampsilis_siliquoidea **
Bogan et al., 2023 [Publ 4295], fig. 5
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 6 |
6 |
4 |
2 |
0 |
0 |
0 |
78.0 |
+-------------- Gibbosula_crassa **
--+
| +----------- Cristaria_truncata **
+--+
| +-------- Sinanodonta_jourdyi **
+--+
| +----- Pletholophus_honglinhensis **
+--+
| +-- Pletholophus_tenuis **
+--+
+-- Pletholophus_reinianus **
Dai et al., 2024 [Publ 4358], fig. 3
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 6 |
6 |
4 |
1 |
0 |
0 |
0 |
80.0 |
+-- Schistodesmus_lampreyanus **
+--------+
| +-- Nodularia_douglasiae **
--+
| +-------- Pseudocuneopsis_capitata **
+--+
| +----- Pseudocuneopsis_yangshuoensis **
+--+
| +-- Pseudocuneopsis_sichuanensis **
+--+
+-- Pseudocuneopsis_yemaoi **
Graf & Ó Foighil, 2000 [Publ 2649], fig. 1, 2
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 5 |
12 |
9 |
2 |
7 |
7 |
1 |
79.0 |
+----------------- Cumberlandia_monodonta **
|
| +-- Unio_pictorum **
| +--+
| | +-- Pyganodon_grandis **
+-----------+
--+ | +-- Amblema_plicata **
| +--+
| +-- Lampsilis_cardium **
|
| +-------------- Velesunio_ambiguus
| |
+--+ +-- Diplodon_chilensis
| +--------+
| | +-- Castalia
+--+
| +-------- Hyridella_australis[2]
+--+
| +----- Hyridella_australis[1]
+--+
| +-- Echyridella_menziesii
+--+
+-- Hyridella_depressa
Krebs, 2004 [Publ 2692], fig. 2
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 5 |
5 |
3 |
1 |
0 |
0 |
0 |
79.0 |
+-------- Quadrula_quadrula **
|
--+ +-- Lasmigona_costata **
| +--+
| | +-- Pyganodon_grandis **
+--+
| +-- Lampsilis_siliquoidea **
+--+
+-- Fusconaia_flava **
Soroka, 2005 [Publ 2726], fig. 3
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 5 |
5 |
3 |
1 |
0 |
0 |
0 |
75.0 |
+----------- Pronodularia_japanensis **
--+
| +-------- Unio_tumidus **
+--+
| +----- Unio_crassus **
+--+
| +-- Unio_caffer **
+--+
+-- Unio_pictorum **
Gustafson & Iwamoto, 2005 [Publ 2741], fig. 2
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 5 |
5 |
3 |
2 |
0 |
0 |
0 |
77.0 |
+----------- Margaritifera_falcata **
--+
| +-------- Gonidea_angulata **
+--+
| +----- Beringiana_beringiana **
+--+
| +-- Anodonta_oregonensis **
+--+
+-- Anodonta_nuttalliana **
Cyr et al., 2007 [Publ 2745], fig. 2D
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 5 |
5 |
2 |
1 |
0 |
0 |
0 |
80.0 |
+-------- Pyganodon_grandis **
|
--+-------- Pyganodon[2] **
|
| +----- Pyganodon[1] **
+--+
| +-- Pyganodon_cataracta **
+--+
+-- Pyganodon_fragilis **
Mulvey et al., 1997 [Publ 3390], fig. 1, 3
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 5 |
5 |
3 |
1 |
0 |
0 |
0 |
79.0 |
+----------- Utterbackia_imbecillis **
--+
| +-------- Megalonaias_nervosa **
+--+
| +----- Amblema_plicata **
+--+
| +-- Amblema_neislerii **
+--+
+-- Amblema_elliottii **
Soroka, 2010 [Publ 3404], fig. 2-M
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 5 |
5 |
3 |
1 |
0 |
0 |
0 |
78.0 |
+----------- Sinanodonta_woodiana **
--+
| +-------- Anodonta_anatina **
+--+
| +----- Unio_crassus **
+--+
| +-- Unio_tumidus **
+--+
+-- Unio_pictorum **
Soroka, 2010 [Publ 3404], fig. 9a
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 5 |
5 |
3 |
1 |
0 |
0 |
0 |
82.0 |
+----------- Buldowskia_flavotincta **
--+
| +-------- Beringiana_beringiana **
+--+
| +----- Sinanodonta_schrenkii **
+--+
| +-- Sinanodonta_lauta **
+--+
+-- Sinanodonta_woodiana **
Soroka, 2010 [Publ 3404], fig. 9c
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 5 |
5 |
3 |
1 |
0 |
0 |
0 |
81.0 |
+----------- Buldowskia_flavotincta **
--+
| +-------- Sinanodonta_woodiana[2] **
+--+
| +----- Sinanodonta_woodiana[1] **
+--+
| +-- Sinanodonta_lauta **
+--+
+-- Sinanodonta_schrenkii **
Krebs et al., 2013 [Publ 3515], fig. 2A
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 5 |
5 |
3 |
1 |
0 |
0 |
0 |
80.0 |
+-------- Lampsilis_radiata **
|
--+ +-- Lampsilis_ovata **
| +--+
| | +-- Ortmanniana_ligamentina **
+--+
| +-- Lampsilis_hydiana **
+--+
+-- Lampsilis_straminea **
Zanatta & Harris, 2013 [Publ 3530], fig. 2
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 5 |
5 |
3 |
1 |
0 |
0 |
0 |
79.0 |
+----------- Pyganodon_grandis **
--+
| +-------- Lampsilis_cardium **
+--+
| +----- Epioblasma_rangiana **
+--+
| +-- Venustaconcha_pleasii **
+--+
+-- Venustaconcha_ellipsiformis **
Konopleva et al., 2017 [Publ 3807], fig. 5
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 5 |
5 |
3 |
1 |
0 |
0 |
0 |
81.0 |
+-- Trapezidens_exolescens **
+-----+
| +-- Trapezidens_dolichorhynchus **
--+
| +----- Lamellidens_savadiensis **
+--+
| +-- Lamellidens_mainwaringi **
+--+
+-- Lamellidens_jenkinsianus **
Dyduch-Falniowska & Koziol, 1989 [Publ 3814], fig. 5, 7
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 5 |
5 |
3 |
1 |
0 |
0 |
0 |
75.0 |
+-- Pseudanodonta_complanata **
+-----+
| +-- Anodonta_anatina **
--+
| +----- Unio_pictorum **
+--+
| +-- Unio_tumidus **
+--+
+-- Unio_crassus **
Choi et al., 2020 [Publ 4169], fig. 3a
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 5 |
5 |
3 |
1 |
0 |
0 |
0 |
79.0 |
+-- Cuneopsis_heudei **
+-----+
| +-- Unio_pictorum **
--+
| +----- Nodularia_breviconcha **
+--+
| +-- Nodularia_douglasiae **
+--+
+-- Nodularia_nipponensis **
Mabuchi et al., 2021 [Publ 4233], fig. 1
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 5 |
5 |
3 |
1 |
0 |
0 |
0 |
79.0 |
+-------- Tchangsinaia_piscicula **
|
--+ +-- Nodularia_douglasiae[1] **
| +--+
| | +-- Nodularia_nipponensis **
+--+
| +-- Nodularia_douglasiae[2] **
+--+
+-- Nodularia_breviconcha **
Bolotov et al., 2022 [Publ 4250], fig. S3
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 5 |
5 |
3 |
1 |
0 |
0 |
0 |
81.0 |
+----------- Leoparreysia_burmana **
--+
| +-------- Leoparreysia_whitteni **
+--+
| +----- Leoparreysia_olivacea **
+--+
| +-- Leoparreysia_tavoyensis **
+--+
+-- Leoparreysia_canefrii **
Saxon et al., 2023 [Publ 4338], fig. 4
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 5 |
5 |
3 |
1 |
0 |
0 |
0 |
82.0 |
+----------- Ellipsaria_lineolata **
--+
| +-------- Truncilla_truncata **
+--+
| +----- Truncilla_macrodon **
+--+
| +-- Truncilla_cognata **
+--+
+-- Truncilla_donaciformis **
Nagel et al., 1998 [Publ 1946], fig. 8
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 4 |
4 |
1 |
1 |
0 |
0 |
0 |
76.0 |
+----- Unio_tumidus **
|
--+ +-- Unio_pictorum **
| |
+--+-- Unio_crassus **
|
+-- Unio_mancus **
Machordom et al., 2003 [Publ 2688], fig. 3
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 4 |
4 |
2 |
2 |
0 |
0 |
0 |
76.0 |
+-------- Potomida_littoralis **
--+
| +----- Cumberlandia_monodonta **
+--+
| +-- Margaritifera_margaritifera **
+--+
+-- Pseudunio_auricularius **
Krebs, 2004 [Publ 2692], fig. 2-M
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 4 |
4 |
2 |
1 |
0 |
0 |
0 |
76.0 |
+-- Lasmigona_costata **
+--+
| +-- Pyganodon_grandis **
--+
| +-- Lampsilis_siliquoidea **
+--+
+-- Fusconaia_flava **
Curole & Kocher, 2002 [Publ 2712], fig. 1-F
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 4 |
4 |
2 |
2 |
0 |
0 |
0 |
78.0 |
+-------- Margaritifera_hembeli **
--+
| +----- Lampsilis_teres **
+--+
| +-- Quadrula_quadrula **
+--+
+-- Pustulosa_pustulosa **
Curole & Kocher, 2002 [Publ 2712], fig. 1a-M
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 4 |
4 |
2 |
1 |
0 |
0 |
0 |
80.0 |
+-- Utterbackiana_implicata **
+--+
| +-- Lampsilis_teres **
--+
| +-- Quadrula_quadrula **
+--+
+-- Pustulosa_pustulosa **
Curole & Kocher, 2002 [Publ 2712], fig. 1b-M
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 4 |
4 |
2 |
1 |
0 |
0 |
0 |
80.0 |
+-------- Lampsilis_teres **
--+
| +----- Utterbackiana_implicata **
+--+
| +-- Quadrula_quadrula **
+--+
+-- Pustulosa_pustulosa **
Plazzi & Passamonti, 2010 [Publ 3354], fig. 2-3
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 4 |
4 |
1 |
1 |
0 |
0 |
0 |
77.0 |
+----- Sinanodonta_woodiana **
|
--+----- Lampsilis_ornata **
|
| +-- Sinohyriopsis_cumingii **
+--+
+-- Pronodularia_japanensis **
Soroka, 2010 [Publ 3404], fig. 9b
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 4 |
4 |
2 |
1 |
0 |
0 |
0 |
81.0 |
+-------- Buldowskia_flavotincta **
--+
| +----- Sinanodonta_schrenkii **
+--+
| +-- Sinanodonta_lauta **
+--+
+-- Sinanodonta_woodiana **
Jiang et al., 2010 [Publ 3438], fig. 5
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 4 |
4 |
1 |
1 |
0 |
0 |
0 |
76.0 |
+----- Lampsilis_ornata **
|
--+----- Cristaria_plicata **
|
| +-- Pronodularia_japanensis **
+--+
+-- Sinohyriopsis_cumingii **
Gangloff et al., 2013 [Publ 3551], fig. 2
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 4 |
4 |
2 |
1 |
0 |
0 |
0 |
81.0 |
+-- Ligumia_recta **
+--+
| +-- Sagittunio_nasutus **
--+
| +-- Venustaconcha_ellipsiformis **
+--+
+-- Venustaconcha_pleasii **
Marshall et al., 2014 [Publ 3593], fig. 1
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 4 |
14 |
12 |
2 |
10 |
10 |
1 |
86.0 |
+----- Cumberlandia_monodonta **
+-----------+
| | +-- Pyganodon_grandis **
| +--+
| +-- Unio_pictorum **
--+
| +-------------- Amblema_plicata **
| |
| | +----- Echyridella_aucklandica
| | +-----+
+--+ | | +-- Echyridella_menziesii
| | +--+
| | +-- Echyridella_onekaka
| |
+--+ +----- Velesunio_ambiguus
| +--+
| | | +-- Velesunio
| | +--+
| | +-- Lortiella
+--+
| +-- Cucumerunio_novaehollandiae
| +--+
| | +-- Hyridella_australis
+--+
| +-- Diplodon_chilensis
+--+
+-- Diplodon_gratus
Klishko et al., 2014 [Publ 3666], fig. 8
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 4 |
4 |
2 |
1 |
0 |
0 |
0 |
79.0 |
+-- Cristaria_plicata **
+--+
| +-- Cristaria_radiata **
--+
| +-- Beringiana_beringiana **
+--+
+-- Sinanodonta_woodiana **
Giribet & Distel, 2003 [Publ 3670], fig. 3.5
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 4 |
4 |
1 |
2 |
2 |
0 |
0 |
76.0 |
+----- Margaritifera_margaritifera **
|
--+ +-- Elliptio_complanata **
| |
+--+-- Lampsilis_cardium **
|
+-- Unio_ravoisieri **
Soroka, 2008 [Publ 3695], fig. 1
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 4 |
4 |
2 |
1 |
0 |
0 |
0 |
76.0 |
+-------- Fusconaia_flava **
--+
| +----- Sinanodonta_woodiana **
+--+
| +-- Anodonta_anatina **
+--+
+-- Pyganodon_grandis **
Soroka, 2008 [Publ 3695], fig. 1-M
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 4 |
4 |
2 |
1 |
0 |
0 |
0 |
76.0 |
+-------- Fusconaia_flava **
--+
| +----- Sinanodonta_woodiana **
+--+
| +-- Anodonta_anatina **
+--+
+-- Pyganodon_grandis **
Mezhzherin et al., 2014 [Publ 3705], fig. 2
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 4 |
4 |
2 |
1 |
0 |
0 |
0 |
75.0 |
+-------- Sinanodonta_woodiana **
--+
| +----- Pseudanodonta_complanata **
+--+
| +-- Anodonta_anatina **
+--+
+-- Anodonta_cygnea **
Mezhzherin et al., 2014 [Publ 3705], fig. 4
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 4 |
4 |
2 |
1 |
0 |
0 |
0 |
77.0 |
+-------- Sinanodonta_woodiana **
--+
| +----- Anodonta_anatina **
+--+
| +-- Anodonta_cygnea **
+--+
+-- Pseudanodonta_complanata **
Klishko et al., 2016 [Publ 3733], fig. 1
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 4 |
4 |
2 |
1 |
0 |
0 |
0 |
79.0 |
+-- Cristaria_plicata **
+--+
| +-- Cristaria_radiata **
--+
| +-- Sinanodonta_woodiana **
+--+
+-- Beringiana_beringiana **
Sayenko et al., 2017 [Publ 3760], fig. 3
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 4 |
4 |
2 |
1 |
0 |
0 |
0 |
81.0 |
+-------- Beringiana_beringiana **
--+
| +----- Sinanodonta_schrenkii **
+--+
| +-- Sinanodonta_woodiana **
+--+
+-- Sinanodonta_lauta **
Froufe et al., 2017 [Publ 3802], fig. 2
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 4 |
4 |
2 |
1 |
0 |
0 |
0 |
76.0 |
+-------- Cristaria_plicata **
--+
| +----- Anodonta_anatina **
+--+
| +-- Anodonta_cygnea **
+--+
+-- Anodonta_exulcerata **
Combosch et al., 2017 [Publ 3806], fig. 1
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 4 |
10 |
8 |
2 |
11 |
9 |
2 |
80.0 |
+----- Triplodon_corrugatus
+--------------+
| | +-- Velesunio_ambiguus
| +--+
| +-- Hyridella_australis
--+
| +----- Unio_pictorum **
| +-----------+
| | | +-- Lampsilis_cardium **
| | +--+
+--+ +-- Scabies_crispata **
|
| +-------------- Margaritifera_margaritifera **
| |
+--+ +-- Chambardia_wahlbergi
| +--------+
| | +-- Aspatharia_pfeifferiana
+--+
| +-------- Etheria_elliptica
+--+
| +----- Mutela_hargeri
+--+
| +-- Tamsiella_schroeteriana
+--+
+-- Lamproscapha_ensiformis
Combosch et al., 2017 [Publ 3806], fig. 2
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 4 |
4 |
2 |
2 |
11 |
9 |
2 |
77.0 |
+-- Chambardia_wahlbergi
+--------+
| +-- Aspatharia_pfeifferiana
+--+
| | +-------- Etheria_elliptica
| +--+
| | +----- Mutela_hargeri
| +--+
| | +-- Tamsiella_schroeteriana
--+ +--+
| +-- Lamproscapha_ensiformis
|
| +----- Velesunio_ambiguus
| +-----+
| | | +-- Triplodon_corrugatus
| | +--+
+--+ +-- Hyridella_australis
|
| +-------- Margaritifera_margaritifera **
+--+
| +----- Unio_pictorum **
+--+
| +-- Scabies_crispata **
+--+
+-- Lampsilis_cardium **
Pfeiffer et al., 2018 [Publ 3835], fig. 2
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 4 |
4 |
2 |
1 |
0 |
0 |
0 |
80.0 |
+-------- Quadrula_quadrula **
--+
| +----- Uniomerus_declivis **
+--+
| +-- Megalonaias_nervosa **
+--+
+-- Megalonaias_nickliniana **
Burzynski & Soroka, 2018 [Publ 3862], fig. 3
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 4 |
4 |
2 |
1 |
0 |
0 |
0 |
77.0 |
+-------- Sinanodonta_woodiana **
--+
| +----- Anodonta_anatina **
+--+
| +-- Utterbackia_peninsularis **
+--+
+-- Pyganodon_grandis **
Zhang et al., 2013 [Publ 4214], fig. 5
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 4 |
4 |
2 |
2 |
0 |
0 |
0 |
79.0 |
+-------- Cristaria_plicata **
--+
| +----- Sinanodonta_lauta **
+--+
| +-- Margaritifera_margaritifera **
+--+
+-- Lamprotula_leaii **
Lopes-Lima et al., 2021 [Publ 4217], fig. 11
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 4 |
4 |
2 |
1 |
0 |
0 |
0 |
78.0 |
+-------- Anodonta_anatina **
--+
| +----- Anodonta_exulcerata **
+--+
| +-- Anodonta_seddoni **
+--+
+-- Anodonta_cygnea **
Patterson et al., 2021 [Publ 4222], fig. 3
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 4 |
4 |
2 |
1 |
0 |
0 |
0 |
79.0 |
+-------- Villosa_villosa **
--+
| +----- Amblema_neislerii **
+--+
| +-- Amblema_plicata **
+--+
+-- Amblema_elliottii **
Johnson et al., 2023 [Publ 4323], fig. 3
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 4 |
4 |
2 |
1 |
0 |
0 |
0 |
81.0 |
+-------- Fusconaia_cerina **
--+
| +----- Pleurobema_riddellii **
+--+
| +-- Pleurobema_beadleianum **
+--+
+-- Pleurobema_clava **
Bulakhova et al., 2023 [Publ 4325], fig. 5
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 4 |
4 |
2 |
1 |
0 |
0 |
0 |
81.0 |
+-- Beringiana_beringiana **
+--+
| +-- Beringiana_japonica **
--+
| +-- Beringiana_gosannensis **
+--+
+-- Beringiana_fukuharai **
Machordom et al., 2003 [Publ 2688], fig. 2
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 3 |
3 |
1 |
2 |
0 |
0 |
0 |
77.0 |
+----- Potomida_littoralis **
--+
| +-- Margaritifera_margaritifera **
+--+
+-- Pseudunio_auricularius **
Hoeh et al., 1996 [Publ 2694], fig. 2
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 3 |
3 |
1 |
1 |
2 |
0 |
0 |
78.0 |
+-- None[1]
+-----+
| +-- None[2]
--+
| +----- Fusconaia_flava **
+--+
| +-- Pyganodon_fragilis **
+--+
+-- Pyganodon_grandis **
Hoeh et al., 1998 [Publ 2695], fig. 3
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 3 |
3 |
1 |
2 |
2 |
1 |
1 |
77.0 |
+-------- Mutela_rostrata
--+
| +----- Anodonta_cygnea **
+--+
| +-- Margaritifera_margaritifera **
+--+
+-- Amblema_plicata **
Hoeh et al., 1998 [Publ 2695], fig. 4
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 3 |
3 |
1 |
2 |
2 |
1 |
1 |
77.0 |
+-------- Mutela_rostrata
--+
| +----- Amblema_plicata **
+--+
| +-- Margaritifera_margaritifera **
+--+
+-- Anodonta_cygnea **
Taylor et al., 2007 [Publ 2696], fig. 1-3
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 3 |
3 |
1 |
2 |
1 |
0 |
0 |
75.0 |
+----- Margaritifera_margaritifera **
--+
| +-- Anodonta_cygnea **
+--+
+-- Unio_pictorum **
Campbell, 2000 [Publ 2724], fig. 1-5
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 3 |
3 |
1 |
2 |
0 |
0 |
0 |
76.0 |
+----- Margaritifera_margaritifera **
--+
| +-- Utterbackia_imbecillis **
+--+
+-- Elliptio_complanata **
Shirai et al., 2010 [Publ 3353], fig. 4
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 3 |
3 |
1 |
1 |
0 |
0 |
0 |
77.0 |
+----- Pronodularia_japanensis **
--+
| +-- Sinohyriopsis_cumingii **
+--+
+-- Sinohyriopsis_schlegelii **
Soroka, 2010 [Publ 3404], fig. 8a-M
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 3 |
3 |
1 |
1 |
0 |
0 |
0 |
76.0 |
+----- Sinanodonta_woodiana **
--+
| +-- Unio_pictorum **
+--+
+-- Unio_tumidus **
Soroka, 2010 [Publ 3404], fig. 8b-M
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 3 |
3 |
1 |
1 |
0 |
0 |
0 |
76.0 |
+----- Sinanodonta_woodiana **
--+
| +-- Unio_pictorum **
+--+
+-- Unio_tumidus **
Hinzmann et al., 2013 [Publ 3541], fig. 5
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 3 |
3 |
1 |
1 |
0 |
0 |
0 |
75.0 |
+----- Pseudanodonta_complanata **
--+
| +-- Anodonta_anatina **
+--+
+-- Anodonta_cygnea **
Froufe et al., 2014 [Publ 3592], fig. 2
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 3 |
3 |
1 |
1 |
0 |
0 |
0 |
77.0 |
+----- Anodonta_anatina **
--+
| +-- Pseudanodonta_complanata **
+--+
+-- Anodonta_cygnea **
Mock et al., 2013 [Publ 3598], fig. 3
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 3 |
3 |
1 |
1 |
0 |
0 |
0 |
79.0 |
+----- Margaritifera_margaritifera **
--+
| +-- Margaritifera_falcata **
+--+
+-- Margaritifera_laevis **
Peng et al., 2012 [Publ 3689], fig. 1
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 3 |
3 |
1 |
1 |
0 |
0 |
0 |
77.0 |
+----- Cristaria_plicata **
--+
| +-- Sinohyriopsis_schlegelii **
+--+
+-- Sinohyriopsis_cumingii **
Xia et al., 2016 [Publ 3712], fig. 7B
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 3 |
3 |
1 |
1 |
0 |
0 |
0 |
78.0 |
+----- Sinanodonta_woodiana **
--+
| +-- Cristaria_plicata **
+--+
+-- Sinohyriopsis_cumingii **
Wang et al., 2016 [Publ 3714], fig. 2
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 3 |
3 |
1 |
1 |
0 |
0 |
0 |
77.0 |
+----- Unio_tumidus **
--+
| +-- Sinohyriopsis_schlegelii **
+--+
+-- Sinohyriopsis_cumingii **
Lopes-Lima et al., 2016 [Publ 3757], fig. 1
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 3 |
3 |
1 |
1 |
0 |
0 |
0 |
75.0 |
+----- Pseudanodonta_complanata **
--+
| +-- Anodonta_anatina **
+--+
+-- Anodonta_cygnea **
Välilä et al., 2017 [Publ 3793], fig. 2
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 3 |
3 |
1 |
2 |
0 |
0 |
0 |
77.0 |
+----- Potomida_littoralis **
--+
| +-- Margaritifera_margaritifera **
+--+
+-- Pseudunio_auricularius **
Metzger et al., 2018 [Publ 3952], fig. 1C
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 3 |
3 |
1 |
1 |
0 |
0 |
0 |
76.0 |
+----- Lasmigona_costata **
--+
| +-- Elliptio_complanata **
+--+
+-- Lampsilis_siliquoidea **
Krebs et al., 2019 [Publ 4092], fig. 4
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 3 |
3 |
1 |
1 |
0 |
0 |
0 |
79.0 |
+----- Pyganodon_grandis **
--+
| +-- Pyganodon_fragilis **
+--+
+-- Pyganodon_cataracta **
Smith & Johnson, 2020 [Publ 4229], fig. 2
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 3 |
3 |
1 |
1 |
0 |
0 |
0 |
80.0 |
+----- Potamilus_inflatus **
--+
| +-- Potamilus_purpuratus **
+--+
+-- Potamilus_fragilis **
Mock et al., 2004 [Publ 2686], fig. 3
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 2 |
2 |
0 |
1 |
0 |
0 |
0 |
-- |
+-- Anodonta **
--+
+-- Sinanodonta_woodiana **
Mock et al., 2004 [Publ 2686], fig. 4
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 2 |
2 |
0 |
1 |
0 |
0 |
0 |
-- |
+-- Anodonta **
--+
+-- Sinanodonta_woodiana **
Plazzi & Passamonti, 2010 [Publ 3354], fig. 1
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 2 |
2 |
0 |
1 |
0 |
0 |
0 |
-- |
+-- Pronodularia_japanensis **
--+
+-- Lampsilis_ornata **
Soroka, 2010 [Publ 3404], fig. 4-M
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 2 |
2 |
0 |
1 |
0 |
0 |
0 |
-- |
+-- Anodonta_anatina **
--+
+-- Sinanodonta_woodiana **
Soroka, 2010 [Publ 3404], fig. 5a-M
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 2 |
2 |
0 |
1 |
0 |
0 |
0 |
-- |
+-- Anodonta_anatina **
--+
+-- Sinanodonta_woodiana **
Soroka, 2010 [Publ 3404], fig. 5b-M
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 2 |
2 |
0 |
1 |
0 |
0 |
0 |
-- |
+-- Anodonta_anatina **
--+
+-- Sinanodonta_woodiana **
Graf et al., 2015 [Publ 3619], fig. 1
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 2 |
2 |
0 |
2 |
18 |
17 |
2 |
-- |
+-- Unio_pictorum **
+-----+
| +-- Margaritifera_margaritifera **
+--------+
| | +----- Chambardia_wahlbergi
| +--+
| | +-- Tamsiella_schroeteriana
| +--+
| +-- Etheria_elliptica
|
| +----------- Westralunio_carteri
| |
--+ | +----- Alathyria_jacksoni
| +--+ +--+
| | | | | +-- Velesunio_ambiguus
| | | | +--+
| | +--+ +-- Alathyria_profuga
| | |
| | | +----- Alathyria_pertexta
| | +--+
| | | +-- Microdontia_anodontaeformis
+--+ +--+
| +-- Lortiella_froggatti
|
| +----- Diplodon_suavidicus
| +--+
| | | +-- Triplodon_corrugatus
| | +--+
| | +-- Castalia_ambigua
+-----+
| +-- Hyridella_depressa
| +--+
| | +-- Hyridella_drapeta
+--+
| +-- Hyridella_australis
+--+
+-- Cucumerunio_novaehollandiae
Bieler et al., 2014 [Publ 3622], fig. 26, 27, 29, 30
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 2 |
3 |
0 |
2 |
3 |
2 |
2 |
77.0 |
+-------- Velesunio_ambiguus
--+
| +----- Unio_pictorum **
+--+
| +-- Aspatharia_pfeifferiana
+--+
+-- Margaritifera_margaritifera **
Bieler et al., 2014 [Publ 3622], fig. 28
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 2 |
4 |
0 |
2 |
3 |
2 |
2 |
78.0 |
+-- Aspatharia_pfeifferiana
+--+
| +-- Margaritifera_margaritifera **
--+
| +-- Unio_pictorum **
+--+
+-- Velesunio_ambiguus
González & Giribet, 2015 [Publ 3632], fig. 2
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 2 |
4 |
0 |
2 |
4 |
2 |
2 |
78.0 |
+-- Aspatharia_pfeifferiana
+--+
| +-- Margaritifera_margaritifera **
--+
| +-- Unio_pictorum **
+--+
+-- Velesunio_ambiguus
González & Giribet, 2015 [Publ 3632], fig. 3
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 2 |
4 |
0 |
2 |
4 |
2 |
2 |
78.0 |
+----- Unio_pictorum **
|
--+----- Velesunio_ambiguus
|
| +-- Aspatharia_pfeifferiana
+--+
+-- Margaritifera_margaritifera **
González & Giribet, 2015 [Publ 3632], fig. 4
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 2 |
2 |
0 |
2 |
4 |
2 |
2 |
-- |
+-------- Aspatharia_pfeifferiana
--+
| +----- Velesunio_ambiguus
+--+
| +-- Unio_pictorum **
+--+
+-- Margaritifera_margaritifera **
González & Giribet, 2015 [Publ 3632], fig. 6
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 2 |
3 |
0 |
2 |
4 |
2 |
2 |
77.0 |
+-------- Velesunio_ambiguus
--+
| +----- Unio_pictorum **
+--+
| +-- Aspatharia_pfeifferiana
+--+
+-- Margaritifera_margaritifera **
Xue et al., 2016 [Publ 3659], fig. 1-M
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 2 |
2 |
0 |
1 |
0 |
0 |
0 |
-- |
+-- Quadrula_quadrula **
--+
+-- Venustaconcha_ellipsiformis **
Klunzinger et al., 2021 [Publ 4086], fig. 2
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 2 |
2 |
0 |
2 |
4 |
4 |
1 |
-- |
+-- Margaritifera_margaritifera **
+-----+
| +-- Unio_pictorum **
|
--+ +-- Westralunio_inbisi
| +--+
| | +-- Westralunio_carteri
+--+
| +-- Velesunio_ambiguus
+--+
+-- Cucumerunio_novaehollandiae
Lemer et al., 2019 [Publ 4093], fig. 1
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 2 |
2 |
0 |
2 |
4 |
2 |
1 |
-- |
+-- Etheria_elliptica
+--+
| +-- Aspatharia_pfeifferiana
--+
| +-- Margaritifera_margaritifera **
+--+
+-- Lampsilis_cardium **
Baker et al., 2004 [Publ 2685], fig. 2-3
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 1 |
1 |
0 |
1 |
11 |
11 |
1 |
-- |
+-------------------- Epioblasma_triquetra **
|
| +----- Hyridella_depressa
--+ +-----------+
| | | +-- Hyridella_australis
| | +--+
| | +-- Diplodon_gratus
+--+
| +-- Echyridella_menziesii
| +-----------+
| | +-- Castalia_stevensi
| |
+--+ +-- Velesunio[3]
| +--------+
| | +-- Velesunio[4]
+--+
| +-------- Velesunio[2]
+--+
| +----- Velesunio_angasi
+--+
| +-- Velesunio[1]
+--+
+-- Alathyria_jacksoni
Baker et al., 2003 [Publ 2690], fig. 5
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 1 |
1 |
0 |
1 |
7 |
7 |
1 |
-- |
+-------------- Epioblasma_triquetra **
|
--+ +-- Hyridella_australis
| +--------+
| | +-- Diplodon_gratus
+--+
| +-------- Hyridella_depressa
| |
+--+ +-- Velesunio_angasi
| +--+
| | +-- Velesunio_ambiguus
+--+
| +-- Echyridella_menziesii
+--+
+-- Castalia_stevensi
| 
