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Page last updated
Thu 04 Apr 2024

The Freshwater Mussels (Unionoida) of the World (and other less consequential bivalves)

FM(U)otW(aolcb) is the web version of the MUSSEL Project Database. Follow the links to browse the data or use the custom Google search field. Either way, you win!

Cladomics pages are experimental — even more so than the rest of this site!

The purpose of the cladomics pages for each genus and family-group level taxon is to provide statistics and references indicating the extent to which published phylogenetic work supports the classification used here.

The Cladomics Summary provides an overview of the number of studies that have included terminal species representing this taxon as well as an indication of how frequently it has been recovered as monophyletic.

Following the summary, the available published clades representing the taxon are listed.

For more information, visit the Cladomics FARQ (frequently asked rhetorical questions) page. Abbreviations for the various statistics are explained on the FARQ!

Search the database pages:

Bivalvia | Unionoida | Unionoidea | Unionidae | Unioninae | Unionini

genus Pseudocuneopsis Huang, Dai, Chen & Wu et al., 2022: cladomics
     5 valid species

Cladomics Summary

number of species = 5

analyzed in 21 cladograms in 14 publications
maximum number of terminals in an analysis = 4, mean = 2.1

Basic Monophyly

number of cladograms that test basic monophyly (i.e., more than 1 included species) = 14
maximum number of included terminals = 4, mean = 2.64
proportion of cladograms that support basic monophyly = 1.0

Pairwise Cladogram Clustering

The graphs below depict in two dimensions all pair-wise ingroup distances within the cladome. See the Cladmoics FARQ for more information.


PCoA Ordination


NJ tree

Liu et al., 2023 [Publ 4310], fig. 4

iT tT bi riS oT roT roS iD
4 4 2 4 20 10 7 0.0

*** Top Monophyly Cladogram *** 

  +-------------- Tchangsinaia_piscicula       
--+                                            
  |  +----------- Inversiunio_yanagawensis     
  +--+                                         
     |  +-------- Pseudocuneopsis_capitata **     
     +--+                                      
        |  +----- Pseudocuneopsis_sichuanensis ** 
        +--+                                   
           |  +-- Pseudocuneopsis_wuana **        
           +--+                                
              +-- Pseudocuneopsis_yangshuoensis **

Chen et al., 2023 [Publ 4302], fig. 1

iT tT bi riS oT roT roS iD
2 2 0 2 38 15 9 --

*** Top Sister-Group Cladogram ***

              +-- Cuneopsis_celtiformis       
  +-----------+                               
  |           +-- Cuneopsis_heudei            
  |                                           
  |           +-- Diaurora_aurorea            
  |        +--+                               
  |        |  +-- Diaurora_laeve              
  +--------+                                  
--+        |  +-- Schistodesmus_lampreyanus   
  |        +--+                               
  |           +-- Schistodesmus_spinosus      
  |                                           
  |           +-- Pseudocuneopsis_sichuanensis **
  |  +--------+                               
  |  |        +-- Pseudocuneopsis_capitata **    
  |  |                                        
  +--+        +-- Inversiunio_yanagawensis    
     |  +-----+                               
     |  |     +-- Inversiunio_reinianus       
     +--+                                     
        |  +----- Tchangsinaia_piscicula      
        +--+                                  
           |  +-- Unio_pictorum               
           +--+                               
              +-- Unio_crassus

Liu et al., 2023 [Publ 4310], fig. 5

iT tT bi riS oT roT roS iD
4 4 2 4 20 10 7 0.0 
              +-- Cuneopsis_rufescens          
  +-----------+                                
  |           +-- Cuneopsis_heudei             
  |                                            
  |        +----- Nodularia_douglasiae         
  +--------+                                   
--+        |  +-- Schistodesmus_lampreyanus    
  |        +--+                                
  |           +-- Schistodesmus_spinosus       
  |                                            
  |  +----------- Tchangsinaia_piscicula       
  |  |                                         
  +--+----------- Inversiunio_yanagawensis     
     |                                         
     |  +-------- Pseudocuneopsis_capitata **     
     +--+                                      
        |  +----- Pseudocuneopsis_sichuanensis ** 
        +--+                                   
           |  +-- Pseudocuneopsis_wuana **        
           +--+                                
              +-- Pseudocuneopsis_yangshuoensis **

Dai et al., 2024 [Publ 4358], fig. 3

iT tT bi riS oT roT roS iD
4 4 2 4 2 2 2 2.0 
           +-- Schistodesmus_lampreyanus    
  +--------+                                
  |        +-- Nodularia_douglasiae         
--+                                         
  |  +-------- Pseudocuneopsis_capitata **     
  +--+                                      
     |  +----- Pseudocuneopsis_yangshuoensis **
     +--+                                   
        |  +-- Pseudocuneopsis_sichuanensis ** 
        +--+                                
           +-- Pseudocuneopsis_yemaoi **

Dai et al., 2024 [Publ 4358], fig. 4

iT tT bi riS oT roT roS iD
4 4 2 4 48 13 6 2.0 
                       +-- Unio_crassus                 
  +--------------------+                                
  |                    +-- Unio_delphinus               
  |                                                     
  |              +-------- Pseudocuneopsis_capitata **     
--+  +-----------+                                      
  |  |           |  +----- Pseudocuneopsis_yangshuoensis **
  |  |           +--+                                   
  |  |              |  +-- Pseudocuneopsis_yemaoi **       
  |  |              +--+                                
  +--+                 +-- Pseudocuneopsis_sichuanensis ** 
     |                                                  
     |                 +-- Nodularia_breviconcha        
     |  +--------------+                                
     |  |              +-- Nodularia_douglasiae         
     |  |                                               
     +--+              +-- Schistodesmus_lampreyanus    
        |  +-----------+                                
        |  |           +-- Schistodesmus_spinosus       
        +--+                                            
           |  +----------- Tchangsinaia_piscicula       
           +--+                                         
              |  +-------- Cuneopsis_rufescens          
              +--+                                      
                 |  +----- Cuneopsis_celtiformis        
                 +--+                                   
                    |  +-- Cuneopsis_heudei             
                    +--+                                
                       +-- Cuneopsis_demangei

Wu et al., 2023 [Publ 4296], fig. 4

iT tT bi riS oT roT roS iD
3 3 1 3 20 10 7 1.0 
  +----------- Tchangsinaia_piscicula       
--+                                         
  |  +-------- Inversiunio_yanagawensis     
  +--+                                      
     |  +----- Pseudocuneopsis_capitata **     
     +--+                                   
        |  +-- Pseudocuneopsis_yangshuoensis **
        +--+                                
           +-- Pseudocuneopsis_sichuanensis **

Wu et al., 2022 [Publ 4238], fig. 2d

iT tT bi riS oT roT roS iD
2 2 0 2 37 21 7 -- 
                 +-------- Unio_tumidus                
  +--------------+                                     
  |              |  +----- Unio_crassus                
  |              +--+                                  
  |                 |  +-- Unio_pictorum               
  |                 +--+                               
  |                    +-- Unio_delphinus              
--+                                                    
  |                    +-- Pseudocuneopsis_capitata **    
  |  +-----------------+                               
  |  |                 +-- Pseudocuneopsis_sichuanensis **
  |  |                                                 
  |  |                 +-- Nodularia_breviconcha       
  +--+  +--------------+                               
     |  |              +-- Nodularia_douglasiae        
     |  |                                              
     |  |              +-- Schistodesmus_lampreyanus   
     +--+  +-----------+                               
        |  |           +-- Schistodesmus_spinosus      
        |  |                                           
        |  |     +-------- Cuneopsis_rufescens         
        +--+  +--+                                     
           |  |  |  +----- Cuneopsis_celtiformis       
           |  |  +--+                                  
           |  |     |  +-- Cuneopsis_heudei            
           +--+     +--+                               
              |        +-- Cuneopsis_demangei          
              |                                        
              |  +-------- Tchangsinaia_piscicula      
              +--+                                     
                 |  +----- Inversiunio_jokohamensis    
                 +--+                                  
                    |  +-- Inversiunio_reinianus       
                    +--+                               
                       +-- Inversiunio_yanagawensis

Wu et al., 2022 [Publ 4238], fig. 2e

iT tT bi riS oT roT roS iD
2 2 0 2 15 10 6 -- 
              +----- Unio_tumidus                
  +-----------+                                  
  |           |  +-- Unio_crassus                
  |           +--+                               
  |              +-- Unio_delphinus              
--+                                              
  |              +-- Pseudocuneopsis_sichuanensis **
  |  +-----------+                               
  |  |           +-- Pseudocuneopsis_capitata **    
  +--+                                           
     |  +----------- Nodularia_douglasiae        
     |  |                                        
     +--+  +-------- Tchangsinaia_piscicula      
        |  |                                     
        +--+     +-- Cuneopsis_celtiformis       
           |  +--+                               
           |  |  +-- Cuneopsis_heudei            
           +--+                                  
              |  +-- Schistodesmus_lampreyanus   
              +--+                               
                 +-- Schistodesmus_spinosus

Wu et al., 2022 [Publ 4238], fig. 3

iT tT bi riS oT roT roS iD
2 2 0 2 72 18 6 -- 
                 +-------- Unio_tumidus                
  +--------------+                                     
  |              |  +----- Unio_crassus                
  |              +--+                                  
  |                 |  +-- Unio_pictorum               
  |                 +--+                               
--+                    +-- Unio_delphinus              
  |                                                    
  |                    +-- Pseudocuneopsis_capitata **    
  |  +-----------------+                               
  |  |                 +-- Pseudocuneopsis_sichuanensis **
  |  |                                                 
  +--+                 +-- Nodularia_douglasiae        
     |  +--------------+                               
     |  |              +-- Nodularia_breviconcha       
     |  |                                              
     +--+              +-- Schistodesmus_lampreyanus   
        |  +-----------+                               
        |  |           +-- Schistodesmus_spinosus      
        +--+                                           
           |  +----------- Tchangsinaia_piscicula      
           +--+                                        
              |  +-------- Cuneopsis_rufescens         
              +--+                                     
                 |  +----- Cuneopsis_celtiformis       
                 +--+                                  
                    |  +-- Cuneopsis_heudei            
                    +--+                               
                       +-- Cuneopsis_demangei

Chen et al., 2023 [Publ 4302], fig. 2

iT tT bi riS oT roT roS iD
2 2 0 2 38 15 9 -- 
              +-- Cuneopsis_celtiformis       
  +-----------+                               
  |           +-- Cuneopsis_heudei            
  |                                           
  |           +-- Diaurora_aurorea            
  |        +--+                               
  |        |  +-- Diaurora_laeve              
  +--------+                                  
--+        |  +-- Schistodesmus_lampreyanus   
  |        +--+                               
  |           +-- Schistodesmus_spinosus      
  |                                           
  |           +-- Pseudocuneopsis_sichuanensis **
  |  +--------+                               
  |  |        +-- Pseudocuneopsis_capitata **    
  |  |                                        
  +--+        +-- Inversiunio_yanagawensis    
     |  +-----+                               
     |  |     +-- Inversiunio_reinianus       
     +--+                                     
        |  +----- Tchangsinaia_piscicula      
        +--+                                  
           |  +-- Unio_pictorum               
           +--+                               
              +-- Unio_crassus

Wu et al., 2023 [Publ 4339], fig. 4, S5-aa

iT tT bi riS oT roT roS iD
2 2 0 2 94 20 6 -- 
                 +-- Nodularia_breviconcha       
              +--+                               
              |  +-- Nodularia_nipponensis       
  +-----------+                                  
  |           |  +-- Pseudocuneopsis_capitata **    
  |           +--+                               
--+              +-- Pseudocuneopsis_sichuanensis **
  |                                              
  |  +-------------- Tchangsinaia_piscicula      
  |  |                                           
  +--+           +-- Schistodesmus_lampreyanus   
     |  +--------+                               
     |  |        +-- Schistodesmus_spinosus      
     +--+                                        
        |  +-------- Cuneopsis_rufescens         
        +--+                                     
           |  +----- Cuneopsis_celtiformis       
           +--+                                  
              |  +-- Cuneopsis_demangei          
              +--+                               
                 +-- Cuneopsis_heudei

Wu et al., 2023 [Publ 4339], fig. 4, S5-nt

iT tT bi riS oT roT roS iD
2 2 0 2 95 21 6 -- 
              +----------- Unio_tumidus                
              |                                        
  +-----------+  +-------- Unio_crassus                
  |           |  |                                     
  |           +--+  +----- Unio_elongatulus            
  |              |  |                                  
  |              +--+  +-- Unio_delphinus              
  |                 |  |                               
  |                 +--+-- Unio_mancus                 
--+                    |                               
  |                    +-- Unio_pictorum               
  |                                                    
  |                    +-- Pseudocuneopsis_capitata **    
  |  +-----------------+                               
  |  |                 +-- Pseudocuneopsis_sichuanensis **
  |  |                                                 
  +--+                 +-- Nodularia_breviconcha       
     |  +--------------+                               
     |  |              +-- Nodularia_nipponensis       
     |  |                                              
     +--+           +----- Tchangsinaia_piscicula      
        |  +--------+                                  
        |  |        |  +-- Schistodesmus_lampreyanus[2]
        |  |        +--+                               
        +--+           +-- Schistodesmus_spinosus      
           |                                           
           |  +----------- Schistodesmus_lampreyanus[1]
           +--+                                        
              |  +-------- Cuneopsis_rufescens         
              +--+                                     
                 |  +----- Cuneopsis_celtiformis       
                 +--+                                  
                    |  +-- Cuneopsis_demangei          
                    +--+                               
                       +-- Cuneopsis_heudei

Wu et al., 2024 [Publ 4350], fig. 8a, S1

iT tT bi riS oT roT roS iD
2 2 0 2 90 20 6 -- 
        +-------------- Unio_tumidus                
  +-----+                                           
  |     |  +----------- Unio_crassus                
  |     +--+                                        
  |        |  +-------- Unio_elongatulus            
  |        +--+                                     
  |           |  +----- Unio_pictorum               
  |           +--+                                  
  |              |  +-- Unio_delphinus              
--+              +--+                               
  |                 +-- Unio_mancus                 
  |                                                 
  |                 +-- Nodularia_breviconcha       
  |  +--------------+                               
  |  |              +-- Nodularia_douglasiae        
  |  |                                              
  |  |              +-- Pseudocuneopsis_capitata **    
  |  +--------------+                               
  +--+              +-- Pseudocuneopsis_sichuanensis **
     |                                              
     |              +-- Schistodesmus_lampreyanus   
     |  +-----------+                               
     |  |           +-- Schistodesmus_spinosus      
     +--+                                           
        |  +----------- Tchangsinaia_piscicula      
        +--+                                        
           |  +-------- Cuneopsis_rufescens         
           +--+                                     
              |  +----- Cuneopsis_celtiformis       
              +--+                                  
                 |  +-- Cuneopsis_demangei          
                 +--+                               
                    +-- Cuneopsis_heudei

Wu et al., 2024 [Publ 4350], fig. 8b, S2

iT tT bi riS oT roT roS iD
2 2 0 2 90 20 6 -- 
        +-------------- Unio_tumidus                
  +-----+                                           
  |     |  +----------- Unio_crassus                
  |     +--+                                        
  |        |  +-------- Unio_elongatulus            
  |        +--+                                     
  |           |  +----- Unio_pictorum               
  |           +--+                                  
  |              |  +-- Unio_delphinus              
--+              +--+                               
  |                 +-- Unio_mancus                 
  |                                                 
  |                 +-- Nodularia_breviconcha       
  |  +--------------+                               
  |  |              +-- Nodularia_douglasiae        
  |  |                                              
  |  |              +-- Pseudocuneopsis_capitata **    
  |  +--------------+                               
  +--+              +-- Pseudocuneopsis_sichuanensis **
     |                                              
     |              +-- Schistodesmus_lampreyanus   
     |  +-----------+                               
     |  |           +-- Schistodesmus_spinosus      
     +--+                                           
        |  +----------- Tchangsinaia_piscicula      
        +--+                                        
           |  +-------- Cuneopsis_rufescens         
           +--+                                     
              |  +----- Cuneopsis_celtiformis       
              +--+                                  
                 |  +-- Cuneopsis_demangei          
                 +--+                               
                    +-- Cuneopsis_heudei

Zhang et al., 2004 [Publ 2735], fig. 1

iT tT bi riS oT roT roS iD
1 1 0 1 13 6 4 -- 
  +----------- Nodularia_douglasiae     
--+                                     
  |  +-------- Pseudocuneopsis_capitata ** 
  +--+                                  
     |  +----- Cuneopsis_rufescens      
     +--+                               
        |  +-- Schistodesmus_lampreyanus
        +--+                            
           +-- Schistodesmus_spinosus

Wu et al., 2019 [Publ 3864], fig. 3

iT tT bi riS oT roT roS iD
1 1 0 1 46 12 6 -- 
        +-------- Unio_tumidus             
  +-----+                                  
  |     |  +----- Unio_crassus             
  |     +--+                               
  |        |  +-- Unio_delphinus           
--+        +--+                            
  |           +-- Unio_pictorum            
  |                                        
  |  +----------- Pseudocuneopsis_capitata ** 
  +--+                                     
     |  +-------- Nodularia_douglasiae     
     +--+                                  
        |  +----- Tchangsinaia_piscicula   
        +--+                               
           |  +-- Cuneopsis_heudei         
           +--+                            
              +-- Schistodesmus_lampreyanus

Fukata & Iigo, 2020 [Publ 4106], fig. 1

iT tT bi riS oT roT roS iD
1 1 0 1 53 11 6 -- 
        +-- Schistodesmus_lampreyanus
  +-----+                            
  |     +-- Cuneopsis_heudei         
--+                                  
  |  +----- Pseudocuneopsis_capitata ** 
  +--+                               
     |  +-- Nodularia_douglasiae     
     +--+                            
        +-- Tchangsinaia_piscicula

Fukata & Iigo, 2019 [Publ 4108], fig. 1

iT tT bi riS oT roT roS iD
1 1 0 1 13 9 6 -- 
     +-------- Unio_tumidus             
  +--+                                  
  |  |  +----- Unio_crassus             
  |  +--+                               
  |     |  +-- Unio_delphinus           
--+     +--+                            
  |        +-- Unio_pictorum            
  |                                     
  |  +-------- Pseudocuneopsis_capitata ** 
  |  |                                  
  +--+     +-- Nodularia_douglasiae     
     |  +--+                            
     |  |  +-- Tchangsinaia_piscicula   
     +--+                               
        |  +-- Schistodesmus_lampreyanus
        +--+                            
           +-- Cuneopsis_heudei

Wu et al., 2020 [Publ 4121], fig. 3

iT tT bi riS oT roT roS iD
1 1 0 1 49 13 6 -- 
        +-------- Unio_tumidus             
  +-----+                                  
  |     |  +----- Unio_crassus             
  |     +--+                               
  |        |  +-- Unio_delphinus           
--+        +--+                            
  |           +-- Unio_pictorum            
  |                                        
  |  +----------- Pseudocuneopsis_capitata ** 
  +--+                                     
     |  +-------- Nodularia_douglasiae     
     +--+                                  
        |  +----- Tchangsinaia_piscicula   
        +--+                               
           |  +-- Cuneopsis_heudei         
           +--+                            
              +-- Schistodesmus_lampreyanus

Wu et al., 2020 [Publ 4147], fig. 3

iT tT bi riS oT roT roS iD
1 1 0 1 29 9 6 -- 
  +-------------- Unio_pictorum            
--+                                        
  |  +----------- Pseudocuneopsis_capitata ** 
  +--+                                     
     |  +-------- Nodularia_douglasiae     
     +--+                                  
        |  +----- Schistodesmus_lampreyanus
        +--+                               
           |  +-- Cuneopsis_heudei         
           +--+                            
              +-- Tchangsinaia_piscicula

Dai et al., 2023 [Publ 4285], fig. 5

iT tT bi riS oT roT roS iD
1 1 0 1 52 6 6 -- 
        +-- Pseudocuneopsis_capitata ** 
  +-----+                            
  |     +-- Nodularia_douglasiae     
--+                                  
  |  +----- Schistodesmus_lampreyanus
  +--+                               
     |  +-- Cuneopsis_celtiformis    
     +--+                            
        +-- Tchangsinaia_piscicula

Page updated: 07:04:26 Thu 04 Apr 2024
 
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