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Page last updated
Thu 04 Apr 2024

The Freshwater Mussels (Unionoida) of the World (and other less consequential bivalves)

FM(U)otW(aolcb) is the web version of the MUSSEL Project Database. Follow the links to browse the data or use the custom Google search field. Either way, you win!

Cladomics pages are experimental — even more so than the rest of this site!

The purpose of the cladomics pages for each genus and family-group level taxon is to provide statistics and references indicating the extent to which published phylogenetic work supports the classification used here.

The Cladomics Summary provides an overview of the number of studies that have included terminal species representing this taxon as well as an indication of how frequently it has been recovered as monophyletic.

Following the summary, the available published clades representing the taxon are listed.

For more information, visit the Cladomics FARQ (frequently asked rhetorical questions) page. Abbreviations for the various statistics are explained on the FARQ!

Search the database pages:

Unionoida | Unionoidea | Unionidae | Gonideinae | Pseudodontini | Pseudodontina

genus Pilsbryoconcha Simpson, 1900: cladomics
     9 valid species

Cladomics Summary

number of species = 9

analyzed in 59 cladograms in 35 publications
maximum number of terminals in an analysis = 10, mean = 2.25

Basic Monophyly

number of cladograms that test basic monophyly (i.e., more than 1 included species) = 23
maximum number of included terminals = 10, mean = 4.22
proportion of cladograms that support basic monophyly = 0.87

Pairwise Cladogram Clustering

The graphs below depict in two dimensions all pair-wise ingroup distances within the cladome. See the Cladmoics FARQ for more information.


PCoA Ordination


NJ tree

Bolotov et al., 2023 [Publ 4312], fig. S2

iT tT bi riS oT roT roS iD
10 10 8 10 32 27 7 0.0

*** Top Monophyly Cladogram *** *** Top Sister-Group Cladogram ***

                          +-- Namkongnaia_inkhavilayi        
           +--------------+                                  
           |              +-- Namkongnaia_lemeslei           
           |                                                 
  +--------+              +-- Thaiconcha_callifera           
  |        |              |                                  
  |        |  +-----------+-- Thaiconcha_thaiensis           
  |        |  |           |                                  
  |        +--+           +-- Thaiconcha_munelliptica        
  |           |                                              
  |           |  +----------- Bineurus_loeiensis             
  |           +--+                                           
  |              |  +-------- Bineurus_panhai                
  |              +--+                                        
  |                 |  +----- Bineurus_anodontinum           
  |                 +--+                                     
--+                    |  +-- Bineurus_exilis                
  |                    +--+                                  
  |                       +-- Bineurus_mouhotii              
  |                                                          
  |              +----------- Pseudodon_vagulus              
  |              |                                           
  |  +-----------+        +-- Pseudodon_lenyanensis          
  |  |           |  +-----+                                  
  |  |           |  |     +-- Pseudodon_inoscularis          
  |  |           +--+                                        
  |  |              |  +----- Pseudodon_cambodjensis         
  |  |              +--+                                     
  +--+                 |  +-- Pseudodon_vondembuschianus     
     |                 +--+                                  
     |                    +-- Pseudodon_mekongi              
     |                                                       
     |  +-------------------- Pilsbryoconcha_carinifera **      
     |  |                                                    
     +--+                 +-- Pilsbryoconcha_linguaeformis[2] **
        |  +--------------+                                  
        |  |              +-- Pilsbryoconcha **                 
        +--+                                                 
           |  +-------------- Pilsbryoconcha_linguaeformis[1] **
           |  |                                              
           +--+           +-- Pilsbryoconcha_schomburgki **     
              |  +--------+                                  
              |  |        +-- Pilsbryoconcha_exilis **          
              +--+                                           
                 |  +-------- Pilsbryoconcha_hoikaab **         
                 +--+                                        
                    |  +----- Pilsbryoconcha_mekongiana **      
                    +--+                                     
                       |  +-- Pilsbryoconcha_acuta **           
                       +--+                                  
                          +-- Pilsbryoconcha_kittitati **

Jeratthitikul et al., 2022 [Publ 4260], fig. S1, 1

iT tT bi riS oT roT roS iD
8 8 6 8 68 7 6 6.0 
                    +-- Nyeinchanconcha_nyeinchani  
              +-----+                               
              |     +-- Sundadontina_cumingii       
  +-----------+                                     
  |           |  +----- Thaiconcha_callifera        
  |           +--+                                  
  |              |  +-- Bineurus_panhai             
  |              +--+                               
--+                 +-- Bineurus_mouhotii           
  |                                                 
  |                 +-- Namkongnaia_inkhavilayi     
  |  +--------------+                               
  |  |              +-- Pseudodon_vondembuschianus  
  |  |                                              
  +--+              +-- Pilsbryoconcha_exilis **       
     |  +-----------+                               
     |  |           +-- Pilsbryoconcha_schomburgki **  
     |  |                                           
     +--+           +-- Pilsbryoconcha_carinifera **   
        |  +--------+                               
        |  |        +-- Pilsbryoconcha_linguaeformis **
        +--+                                        
           |  +-------- Pilsbryoconcha_hoikaab **      
           +--+                                     
              |  +----- Pilsbryoconcha_mekongiana **   
              +--+                                  
                 |  +-- Pilsbryoconcha_acuta **        
                 +--+                               
                    +-- Pilsbryoconcha_kittitati **

Jeratthitikul et al., 2022 [Publ 4260], fig. S2, 2

iT tT bi riS oT roT roS iD
8 8 6 8 68 7 6 6.0 
                    +-- Nyeinchanconcha_nyeinchani  
              +-----+                               
              |     +-- Sundadontina_cumingii       
  +-----------+                                     
  |           |  +----- Thaiconcha_callifera        
  |           +--+                                  
  |              |  +-- Bineurus_panhai             
  |              +--+                               
--+                 +-- Bineurus_mouhotii           
  |                                                 
  |                 +-- Namkongnaia_inkhavilayi     
  |  +--------------+                               
  |  |              +-- Pseudodon_vondembuschianus  
  |  |                                              
  +--+              +-- Pilsbryoconcha_exilis **       
     |  +-----------+                               
     |  |           +-- Pilsbryoconcha_schomburgki **  
     |  |                                           
     +--+           +-- Pilsbryoconcha_carinifera **   
        |  +--------+                               
        |  |        +-- Pilsbryoconcha_linguaeformis **
        +--+                                        
           |  +-------- Pilsbryoconcha_hoikaab **      
           +--+                                     
              |  +----- Pilsbryoconcha_mekongiana **   
              +--+                                  
                 |  +-- Pilsbryoconcha_acuta **        
                 +--+                               
                    +-- Pilsbryoconcha_kittitati **

Bolotov et al., 2023 [Publ 4312], fig. 2

iT tT bi riS oT roT roS iD
8 8 6 8 32 27 7 10.0 
                          +-- Pilsbryoconcha_exilis **        
              +-----------+                                
              |           +-- Pilsbryoconcha_schomburgki **   
              |                                            
  +-----------+           +-- Pilsbryoconcha_carinifera **    
  |           |  +--------+                                
  |           |  |        +-- Pilsbryoconcha_linguaeformis ** 
  |           +--+                                         
  |              |  +-------- Pilsbryoconcha_acuta **         
  |              +--+                                      
  |                 |  +----- Pilsbryoconcha_kittitati **     
  |                 +--+                                   
  |                    |  +-- Pilsbryoconcha_mekongiana **    
  |                    +--+                                
--+                       +-- Pilsbryoconcha_hoikaab **       
  |                                                        
  |  +----------------------- Songkhlanaia_tamodienica     
  |  |                                                     
  |  |                    +-- Namkongnaia_inkhavilayi      
  |  |           +--------+                                
  |  |           |        +-- Namkongnaia_lemeslei         
  |  |           |                                         
  |  |           |     +----- Pseudodon_vondembuschianus[2]
  |  |  +--------+  +--+                                   
  |  |  |        |  |  |  +-- Pseudodon_vondembuschianus[1]
  +--+  |        |  |  +--+                                
     |  |        |  |     +-- Pseudodon_mekongi            
     |  |        +--+                                      
     |  |           |     +-- Pseudodon_vagulus            
     |  |           |  +--+                                
     |  |           |  |  +-- Pseudodon_cambodjensis       
     |  |           +--+                                   
     |  |              |  +-- Pseudodon_lenyanensis        
     |  |              +--+                                
     |  |                 +-- Pseudodon_inoscularis        
     +--+                                                  
        |     +-------------- Nyeinchanconcha_nyeinchani   
        |     |                                            
        |     |        +----- Thaiconcha_munelliptica      
        |  +--+  +-----+                                   
        |  |  |  |     |  +-- Thaiconcha_callifera         
        |  |  |  |     +--+                                
        |  |  +--+        +-- Thaiconcha_thaiensis         
        |  |     |                                         
        |  |     |  +-------- Bineurus_loeiensis           
        |  |     +--+                                      
        |  |        |  +----- Bineurus_mouhotii            
        |  |        +--+                                   
        +--+           |  +-- Bineurus_anodontinum         
           |           +--+                                
           |              +-- Bineurus_exilis              
           |                                               
           |        +-------- Sundadontina_sulcata         
           |  +-----+                                      
           |  |     |  +----- Sundadontina_harmandi        
           |  |     +--+                                   
           |  |        |  +-- Sundadontina_brandti         
           +--+        +--+                                
              |           +-- Sundadontina_taskaevi        
              |                                            
              |  +----------- Sundadontina_tumida          
              +--+                                         
                 |  +-------- Sundadontina                 
                 +--+                                      
                    |  +----- Sundadontina_tanintharyiensis
                    +--+                                   
                       |  +-- Sundadontina_cumingii        
                       +--+                                
                          +-- Sundadontina_plugpomenica

Bolotov et al., 2023 [Publ 4312], fig. S1

iT tT bi riS oT roT roS iD
8 8 6 8 31 26 7 10.0 
                          +-- Pilsbryoconcha_exilis **        
              +-----------+                                
              |           +-- Pilsbryoconcha_schomburgki **   
              |                                            
  +-----------+           +-- Pilsbryoconcha_carinifera **    
  |           |  +--------+                                
  |           |  |        +-- Pilsbryoconcha_linguaeformis ** 
  |           +--+                                         
  |              |  +-------- Pilsbryoconcha_kittitati **     
  |              +--+                                      
  |                 |  +----- Pilsbryoconcha_acuta **         
  |                 +--+                                   
  |                    |  +-- Pilsbryoconcha_hoikaab **       
  |                    +--+                                
  |                       +-- Pilsbryoconcha_mekongiana **    
  |                                                        
--+           +-------------- Nyeinchanconcha_nyeinchani   
  |           |                                            
  |           |        +----- Thaiconcha_thaiensis         
  |  +--------+  +-----+                                   
  |  |        |  |     |  +-- Thaiconcha_callifera         
  |  |        |  |     +--+                                
  |  |        +--+        +-- Thaiconcha_munelliptica      
  |  |           |                                         
  |  |           |  +-------- Bineurus_loeiensis           
  |  |           +--+                                      
  |  |              |  +----- Bineurus_mouhotii            
  |  |              +--+                                   
  +--+                 |  +-- Bineurus_anodontinum         
     |                 +--+                                
     |                    +-- Bineurus_exilis              
     |                                                     
     |  +-------------------- Songkhlanaia_tamodienica     
     |  |                                                  
     |  |                 +-- Namkongnaia_lemeslei         
     |  |        +--------+                                
     |  |        |        +-- Namkongnaia_inkhavilayi      
     |  |        |                                         
     +--+  +-----+        +-- Pseudodon_inoscularis        
        |  |     |  +-----+                                
        |  |     |  |     +-- Pseudodon_lenyanensis        
        |  |     |  |                                      
        |  |     +--+     +-- Pseudodon_vagulus            
        |  |        |  +--+                                
        |  |        |  |  +-- Pseudodon_cambodjensis       
        +--+        +--+                                   
           |           |  +-- Pseudodon_vondembuschianus   
           |           +--+                                
           |              +-- Pseudodon_mekongi            
           |                                               
           |  +-------------- Sundadontina_tumida          
           |  |                                            
           |  |     +-------- Sundadontina_sulcata         
           +--+  +--+                                      
              |  |  |  +----- Sundadontina_harmandi        
              |  |  +--+                                   
              |  |     |  +-- Sundadontina_brandti         
              +--+     +--+                                
                 |        +-- Sundadontina_taskaevi        
                 |                                         
                 |  +-------- Sundadontina_tanintharyiensis
                 +--+                                      
                    |  +----- Sundadontina                 
                    +--+                                   
                       |  +-- Sundadontina_cumingii        
                       +--+                                
                          +-- Sundadontina_plugpomenica

Konopleva et al., 2023 [Publ 4262], fig. 8

iT tT bi riS oT roT roS iD
6 6 4 6 78 21 7 12.0 
                             +-- Pilsbryoconcha_schomburgki **   
                    +--------+                                
                    |        +-- Pilsbryoconcha_exilis[1] **     
  +-----------------+                                         
  |                 |  +-------- Pilsbryoconcha **               
  |                 +--+                                      
  |                    |  +----- Pilsbryoconcha_linguaeformis ** 
  |                    +--+                                   
  |                       |  +-- Pilsbryoconcha_exilis[2] **     
--+                       +--+                                
  |                          +-- Pilsbryoconcha_carinifera **    
  |                                                           
  |  +-------------------------- Songkhlanaia_tamodienica     
  |  |                                                        
  |  |                       +-- Namkongnaia_inkhavilayi      
  |  |  +--------------------+                                
  +--+  |                    +-- Namkongnaia_lemeslei         
     |  |                                                     
     |  |              +-------- Pseudodon_lenyanensis        
     |  |  +-----------+                                      
     |  |  |           |  +----- Pseudodon_vagulus            
     +--+  |           +--+                                   
        |  |              |  +-- Pseudodon_vondembuschianus   
        |  |              +--+                                
        |  |                 +-- Pseudodon_mekongi            
        |  |                                                  
        |  |                 +-- Sundadontina_taskaevi        
        +--+        +--------+                                
           |        |        +-- Sundadontina_brandti         
           |  +-----+                                         
           |  |     |  +-------- Sundadontina_tumida          
           |  |     +--+                                      
           |  |        |  +----- Sundadontina_tanintharyiensis
           |  |        +--+                                   
           |  |           |  +-- Sundadontina_plugpomenica    
           +--+           +--+                                
              |              +-- Sundadontina_cumingii        
              |                                               
              |  +-------------- Nyeinchanconcha_nyeinchani   
              |  |                                            
              |  |        +----- Thaiconcha_thaiensis         
              +--+  +-----+                                   
                 |  |     |  +-- Thaiconcha_munelliptica      
                 |  |     +--+                                
                 +--+        +-- Thaiconcha_callifera         
                    |                                         
                    |  +-------- Bineurus_loeiensis           
                    +--+                                      
                       |  +----- Bineurus_mouhotii            
                       +--+                                   
                          |  +-- Bineurus_anodontinum         
                          +--+                                
                             +-- Bineurus_exilis

Bolotov et al., 2017 [Publ 3761], fig. S1

iT tT bi riS oT roT roS iD
4 4 2 4 137 10 5 9.0 
           +-------- Pilsbryoconcha_exilis[1] **  
  +--------+                                   
  |        |  +----- Pilsbryoconcha_exilis[3] **  
  |        +--+                                
  |           |  +-- Pilsbryoconcha_exilis[2] **  
  |           +--+                             
  |              +-- Pilsbryoconcha_carinifera ** 
--+                                            
  |        +-------- Sundadontina_tumida       
  |  +-----+                                   
  |  |     |  +----- Pseudodon_vagulus         
  |  |     +--+                                
  |  |        |  +-- Pseudodon_mekongi         
  |  |        +--+                             
  +--+           +-- Pseudodon_vondembuschianus
     |                                         
     |           +-- Nyeinchanconcha_nyeinchani
     |  +--------+                             
     |  |        +-- Sundadontina_cumingii     
     +--+                                      
        |  +-------- Thaiconcha_callifera      
        +--+                                   
           |  +----- Bineurus_exilis           
           +--+                                
              |  +-- Bineurus_mouhotii         
              +--+                             
                 +-- Bineurus_loeiensis

Bolotov et al., 2017 [Publ 3761], fig. S2

iT tT bi riS oT roT roS iD
4 4 2 4 137 11 5 9.0 
  +-------- Pilsbryoconcha_exilis[3] ** 
--+                                  
  |  +----- Pilsbryoconcha_exilis[2] ** 
  +--+                               
     |  +-- Pilsbryoconcha_exilis[1] ** 
     +--+                            
        +-- Pilsbryoconcha_carinifera **

Bolotov et al., 2017 [Publ 3764], fig. S1

iT tT bi riS oT roT roS iD
4 4 2 4 127 10 5 9.0 
              +-- Nyeinchanconcha_nyeinchani
  +-----------+                             
  |           +-- Sundadontina_cumingii     
  |                                         
  |        +----- Bineurus_exilis           
  +--------+                                
  |        |  +-- Bineurus_mouhotii         
  |        +--+                             
  |           +-- Bineurus_loeiensis        
  |                                         
--+     +-------- Sundadontina_tumida       
  +-----+                                   
  |     |  +----- Pseudodon_vagulus         
  |     +--+                                
  |        |  +-- Pseudodon_mekongi         
  |        +--+                             
  |           +-- Pseudodon_vondembuschianus
  |                                         
  |  +----------- Thaiconcha_callifera      
  +--+                                      
     |  +-------- Pilsbryoconcha_exilis[3] **  
     +--+                                   
        |  +----- Pilsbryoconcha_exilis[2] **  
        +--+                                
           |  +-- Pilsbryoconcha_exilis[1] **  
           +--+                             
              +-- Pilsbryoconcha_carinifera **

Huang et al., 2019 [Publ 3843], fig. 2

iT tT bi riS oT roT roS iD
4 4 2 4 200 11 5 9.0 
           +-------- Pilsbryoconcha_exilis[3] **  
  +--------+                                   
  |        |  +----- Pilsbryoconcha_exilis[2] **  
  |        +--+                                
  |           |  +-- Pilsbryoconcha_exilis[1] **  
  |           +--+                             
  |              +-- Pilsbryoconcha_carinifera ** 
  |                                            
--+     +----------- Nyeinchanconcha_nyeinchani
  |  +--+                                      
  |  |  |  +-------- Thaiconcha_callifera      
  |  |  +--+                                   
  |  |     |  +----- Bineurus_loeiensis        
  |  |     +--+                                
  |  |        |  +-- Bineurus_mouhotii         
  +--+        +--+                             
     |           +-- Bineurus_exilis           
     |                                         
     |        +----- Pseudodon_vagulus         
     |     +--+                                
     |     |  |  +-- Pseudodon_mekongi         
     |     |  +--+                             
     +-----+     +-- Pseudodon_vondembuschianus
           |                                   
           |     +-- Sundadontina_cumingii     
           |     |                             
           +-----+-- Sundadontina_tumida[1]    
                 |                             
                 +-- Sundadontina_tumida[2]

Huang et al., 2019 [Publ 3843], fig. S3

iT tT bi riS oT roT roS iD
4 4 2 4 197 11 5 9.0 
           +-------- Pilsbryoconcha_exilis[3] **     
  +--------+                                      
  |        |  +----- Pilsbryoconcha_exilis[2] **     
  |        +--+                                   
  |           |  +-- Pilsbryoconcha_carinifera **    
  |           +--+                                
  |              +-- Pilsbryoconcha_exilis[1] **     
  |                                               
--+     +----------- Sundadontina_tumida          
  |  +--+                                         
  |  |  |  +-------- Pseudodon_vagulus            
  |  |  +--+                                      
  |  |     |  +----- Pseudodon_vondembuschianus[1]
  |  |     +--+                                   
  |  |        |  +-- Pseudodon_mekongi            
  +--+        +--+                                
     |           +-- Pseudodon_vondembuschianus[2]
     |                                            
     |           +-- Nyeinchanconcha_nyeinchani   
     |  +--------+                                
     |  |        +-- Sundadontina_cumingii        
     +--+                                         
        |  +-------- Thaiconcha_callifera         
        +--+                                      
           |  +----- Bineurus_loeiensis           
           +--+                                   
              |  +-- Bineurus_mouhotii            
              +--+                                
                 +-- Bineurus_exilis

Huang et al., 2019 [Publ 3843], fig. S4

iT tT bi riS oT roT roS iD
3 3 1 3 199 11 5 9.0 
  +-------- Nyeinchanconcha_nyeinchani
  |                                   
  +-------- Thaiconcha_callifera      
  |                                   
  +-------- Sundadontina_cumingii     
  |                                   
  +-------- Sundadontina_tumida[1]    
  |                                   
  +-------- Sundadontina_tumida[2]    
  |                                   
  |  +----- Pilsbryoconcha_exilis[2] **  
--+--+                                
  |  |  +-- Pilsbryoconcha_carinifera ** 
  |  +--+                             
  |     +-- Pilsbryoconcha_exilis[1] **  
  |                                   
  |  +----- Pseudodon_vagulus         
  +--+                                
  |  |  +-- Pseudodon_mekongi         
  |  +--+                             
  |     +-- Pseudodon_vondembuschianus
  |                                   
  |  +----- Bineurus_loeiensis        
  +--+                                
     |  +-- Bineurus_mouhotii         
     +--+                             
        +-- Bineurus_exilis

Bolotov et al., 2020 [Publ 4100], fig. 1

iT tT bi riS oT roT roS iD
3 3 1 3 117 15 5 11.0 
              +-- Thaiconcha_callifera         
        +-----+                                
        |     +-- Thaiconcha_munelliptica      
  +-----+                                      
  |     |  +----- Pilsbryoconcha_exilis[2] **     
  |     +--+                                   
  |        |  +-- Pilsbryoconcha_exilis[1] **     
  |        +--+                                
  |           +-- Pilsbryoconcha **               
--+                                            
  |     +-------- Pseudodon_lenyanensis        
  |  +--+                                      
  |  |  |  +----- Pseudodon_vagulus            
  |  |  +--+                                   
  |  |     |  +-- Pseudodon_vondembuschianus   
  |  |     +--+                                
  +--+        +-- Pseudodon_mekongi            
     |                                         
     |        +-- Sundadontina_taskaevi        
     |  +-----+                                
     |  |     +-- Sundadontina_brandti         
     +--+                                      
        |  +----- Sundadontina_tumida          
        +--+                                   
           |  +-- Sundadontina_cumingii        
           +--+                                
              +-- Sundadontina_tanintharyiensis

Bolotov et al., 2020 [Publ 4100], fig. S1

iT tT bi riS oT roT roS iD
3 3 1 3 111 15 5 11.0 
              +-- Thaiconcha_callifera         
        +-----+                                
        |     +-- Thaiconcha_munelliptica      
  +-----+                                      
  |     |  +----- Pilsbryoconcha_exilis[2] **     
  |     +--+                                   
  |        |  +-- Pilsbryoconcha_exilis[1] **     
  |        +--+                                
  |           +-- Pilsbryoconcha **               
--+                                            
  |     +-------- Pseudodon_lenyanensis        
  |  +--+                                      
  |  |  |  +----- Pseudodon_vagulus            
  |  |  +--+                                   
  |  |     |  +-- Pseudodon_vondembuschianus   
  |  |     +--+                                
  +--+        +-- Pseudodon_mekongi            
     |                                         
     |        +-- Sundadontina_tanintharyiensis
     |  +-----+                                
     |  |     +-- Sundadontina_cumingii        
     +--+                                      
        |  +----- Sundadontina_tumida          
        +--+                                   
           |  +-- Sundadontina_taskaevi        
           +--+                                
              +-- Sundadontina_brandti

Jeratthitikul et al., 2021 [Publ 4219], fig. 1

iT tT bi riS oT roT roS iD
3 3 1 3 33 15 6 9.0 
                 +----- Pilsbryoconcha_exilis **       
  +--------------+                                  
  |              |  +-- Pilsbryoconcha_linguaeformis **
  |              +--+                               
  |                 +-- Pilsbryoconcha **              
  |                                                 
  |           +-------- Nyeinchanconcha_nyeinchani  
  |           |                                     
--+  +--------+  +----- Thaiconcha_callifera        
  |  |        |  |                                  
  |  |        +--+  +-- Bineurus_panhai             
  |  |           |  |                               
  |  |           +--+-- Bineurus_exilis             
  |  |              |                               
  |  |              +-- Bineurus_mouhotii           
  +--+                                              
     |              +-- Sundadontina_brandti        
     |           +--+                               
     |           |  +-- Sundadontina_taskaevi       
     |  +--------+                                  
     |  |        |  +-- Sundadontina_tumida         
     |  |        +--+                               
     |  |           +-- Sundadontina_cumingii       
     +--+                                           
        |           +-- Namkongnaia_lemeslei        
        |  +--------+                               
        |  |        +-- Namkongnaia_inkhavilayi     
        +--+                                        
           |  +-------- Pseudodon_lenyanensis       
           +--+                                     
              |  +----- Pseudodon_vagulus           
              +--+                                  
                 |  +-- Pseudodon_vondembuschianus  
                 +--+                               
                    +-- Pseudodon_mekongi

Jeratthitikul et al., 2021 [Publ 4219], fig. 2

iT tT bi riS oT roT roS iD
3 3 1 3 33 15 6 9.0 
                 +----- Pilsbryoconcha_exilis **       
  +--------------+                                  
  |              |  +-- Pilsbryoconcha_linguaeformis **
  |              +--+                               
  |                 +-- Pilsbryoconcha **              
  |                                                 
  |                 +-- Nyeinchanconcha_nyeinchani  
  |        +--------+                               
--+        |        +-- Sundadontina_cumingii       
  |  +-----+                                        
  |  |     |  +-------- Thaiconcha_callifera        
  |  |     +--+                                     
  |  |        |  +----- Bineurus_mouhotii           
  |  |        +--+                                  
  |  |           |  +-- Bineurus_panhai             
  |  |           +--+                               
  +--+              +-- Bineurus_exilis             
     |                                              
     |           +----- Sundadontina_tumida         
     |  +--------+                                  
     |  |        |  +-- Sundadontina_brandti        
     |  |        +--+                               
     |  |           +-- Sundadontina_taskaevi       
     +--+                                           
        |           +-- Namkongnaia_lemeslei        
        |  +--------+                               
        |  |        +-- Namkongnaia_inkhavilayi     
        +--+                                        
           |  +-------- Pseudodon_lenyanensis       
           +--+                                     
              |  +----- Pseudodon_vagulus           
              +--+                                  
                 |  +-- Pseudodon_vondembuschianus  
                 +--+                               
                    +-- Pseudodon_mekongi

Lopes-Lima et al., 2017 [Publ 3745], fig. 1-2

iT tT bi riS oT roT roS iD
2 2 0 2 79 3 3 -- 
        +-- Pilsbryoconcha_exilis **     
  +-----+                             
  |     +-- Pilsbryoconcha_acuta **      
--+                                   
  |  +----- Pseudodon_vagulus         
  +--+                                
     |  +-- Bineurus_exilis           
     +--+                             
        +-- Nyeinchanconcha_nyeinchani

Bolotov et al., 2018 [Publ 3826], fig. 2

iT tT bi riS oT roT roS iD
2 8 0 2 83 6 2 14.0 
  +-- Bineurus_mouhotii         
  |                             
  +-- Pseudodon_vagulus         
  |                             
  +-- Pseudodon_mekongi         
  |                             
  +-- Pilsbryoconcha_exilis **     
--+                             
  +-- Pilsbryoconcha_carinifera ** 
  |                             
  +-- Bineurus_loeiensis        
  |                             
  +-- Bineurus_exilis           
  |                             
  +-- Pseudodon_vondembuschianus

Konopleva et al., 2019 [Publ 3900], fig. 1

iT tT bi riS oT roT roS iD
2 8 0 2 50 6 2 14.0 
  +-- Bineurus_mouhotii         
  |                             
  +-- Pseudodon_vagulus         
  |                             
  +-- Pseudodon_mekongi         
  |                             
  +-- Pilsbryoconcha_exilis **     
--+                             
  +-- Pilsbryoconcha_carinifera ** 
  |                             
  +-- Bineurus_loeiensis        
  |                             
  +-- Bineurus_exilis           
  |                             
  +-- Pseudodon_vondembuschianus

Lopes-Lima et al., 2020 [Publ 4080], fig. 2-4

iT tT bi riS oT roT roS iD
2 2 0 2 120 4 4 -- 
        +-- Pilsbryoconcha_exilis **     
  +-----+                             
  |     +-- Pilsbryoconcha_carinifera ** 
--+                                   
  |  +----- Pseudodon_mekongi         
  |  |                                
  +--+  +-- Nyeinchanconcha_nyeinchani
     |  |                             
     +--+-- Bineurus_mouhotii         
        |                             
        +-- Sundadontina_tumida

Konopleva et al., 2021 [Publ 4213], fig. 1

iT tT bi riS oT roT roS iD
2 3 0 2 26 18 6 11.0 
                 +----- Pilsbryoconcha_exilis[2] **     
  +--------------+                                   
  |              |  +-- Pilsbryoconcha_exilis[1] **     
  |              +--+                                
  |                 +-- Namkongnaia_lemeslei         
  |                                                  
  |     +-------------- Nyeinchanconcha_nyeinchani   
  |     |                                            
  |     |           +-- Thaiconcha_munelliptica      
--+  +--+           |                                
  |  |  |  +--------+-- Thaiconcha_thaiensis         
  |  |  |  |        |                                
  |  |  +--+        +-- Thaiconcha_callifera         
  |  |     |                                         
  |  |     |  +-------- Bineurus_loeiensis           
  |  |     +--+                                      
  |  |        |  +----- Bineurus_mouhotii            
  |  |        +--+                                   
  +--+           |  +-- Bineurus_anodontinum         
     |           +--+                                
     |              +-- Bineurus_exilis              
     |                                               
     |        +-------- Pseudodon_lenyanensis        
     |     +--+                                      
     |     |  |  +----- Pseudodon_vagulus            
     |     |  +--+                                   
     |     |     |  +-- Pseudodon_vondembuschianus   
     |     |     +--+                                
     +-----+        +-- Pseudodon_mekongi            
           |                                         
           |        +-- Sundadontina_taskaevi        
           |  +-----+                                
           |  |     +-- Sundadontina_brandti         
           +--+                                      
              |  +----- Sundadontina_tumida          
              +--+                                   
                 |  +-- Sundadontina_cumingii        
                 +--+                                
                    +-- Sundadontina_tanintharyiensis

Dai et al., 2023 [Publ 4285], fig. 3-BI

iT tT bi riS oT roT roS iD
2 2 0 2 43 12 5 -- 
     +-------- Nyeinchanconcha_nyeinchani   
  +--+                                      
  |  |  +----- Bineurus_loeiensis           
  |  +--+                                   
  |     |  +-- Bineurus_mouhotii            
  |     +--+                                
  |        +-- Bineurus_exilis              
  |                                         
  |        +-- Thaiconcha_callifera         
  |  +-----+                                
  |  |     +-- Thaiconcha_munelliptica      
--+--+                                      
  |  |  +----- Sundadontina_tanintharyiensis
  |  +--+                                   
  |     |  +-- Sundadontina_brandti         
  |     +--+                                
  |        +-- Sundadontina_taskaevi        
  |                                         
  |        +-- Pilsbryoconcha_carinifera **    
  |  +-----+                                
  |  |     +-- Pilsbryoconcha_exilis **        
  +--+                                      
     |  +----- Pseudodon_lenyanensis        
     +--+                                   
        |  +-- Pseudodon_mekongi            
        +--+                                
           +-- Pseudodon_vagulus

Dai et al., 2023 [Publ 4285], fig. 3-ML

iT tT bi riS oT roT roS iD
2 2 0 2 43 12 5 -- 
           +-- Thaiconcha_callifera         
     +-----+                                
     |     +-- Thaiconcha_munelliptica      
  +--+                                      
  |  |  +----- Sundadontina_tanintharyiensis
  |  +--+                                   
  |     |  +-- Sundadontina_brandti         
  |     +--+                                
--+        +-- Sundadontina_taskaevi        
  |                                         
  |        +-- Pilsbryoconcha_carinifera **    
  |  +-----+                                
  |  |     +-- Pilsbryoconcha_exilis **        
  +--+                                      
     |  +----- Pseudodon_lenyanensis        
     +--+                                   
        |  +-- Pseudodon_mekongi            
        +--+                                
           +-- Pseudodon_vagulus

Graf & Cummings, 2006 [Publ 1968], fig. 2

iT tT bi riS oT roT roS iD
1 1 0 1 46 1 1 -- 
  +-- Pilsbryoconcha_exilis **     
--+                             
  +-- Pseudodon_vondembuschianus

Graf, 2002 [Publ 2651], fig. 2

iT tT bi riS oT roT roS iD
1 1 0 1 17 1 1 -- 
  +-- Pilsbryoconcha_exilis **     
--+                             
  +-- Pseudodon_vondembuschianus

Hoeh et al., 2009 [Publ 2716], fig. 3

iT tT bi riS oT roT roS iD
1 1 0 1 46 1 1 -- 
  +-- Pilsbryoconcha_exilis **     
--+                             
  +-- Pseudodon_vondembuschianus

Hoeh et al., 2009 [Publ 2716], fig. 4

iT tT bi riS oT roT roS iD
1 1 0 1 45 1 1 -- 
  +-- Pilsbryoconcha_exilis **     
--+                             
  +-- Pseudodon_vondembuschianus

Hoeh et al., 2009 [Publ 2716], fig. 5

iT tT bi riS oT roT roS iD
1 1 0 1 45 1 1 -- 
  +-- Pilsbryoconcha_exilis **     
--+                             
  +-- Pseudodon_vondembuschianus

Pfeiffer & Graf, 2015 [Publ 3620], fig. 2

iT tT bi riS oT roT roS iD
1 1 0 1 39 5 4 -- 
  +----------- Pilsbryoconcha_carinifera ** 
  |                                      
--+        +-- Pseudodon_vondembuschianus
  |  +-----+                             
  |  |     +-- Pseudodon_vagulus         
  +--+                                   
     |  +----- Sundadontina_tumida       
     +--+                                
        |  +-- Nyeinchanconcha_nyeinchani
        +--+                             
           +-- Bineurus_exilis

Konopleva et al., 2017 [Publ 3807], fig. 6

iT tT bi riS oT roT roS iD
1 1 0 1 30 5 4 -- 
        +-- Nyeinchanconcha_nyeinchani
  +-----+                             
  |     +-- Bineurus_exilis           
--+                                   
  |  +----- Pilsbryoconcha_carinifera ** 
  +--+                                
     |  +-- Pseudodon_vagulus         
     +--+                             
        +-- Pseudodon_vondembuschianus

Jeratthitikul et al., 2019 [Publ 3911], fig. 4

iT tT bi riS oT roT roS iD
1 1 0 1 21 0 0 -- 
-- Pilsbryoconcha_exilis

Froufe et al., 2019 [Publ 3936], fig. 3

iT tT bi riS oT roT roS iD
1 1 0 1 44 1 1 -- 
  +-- Pilsbryoconcha_exilis **     
--+                             
  +-- Pseudodon_vondembuschianus

Froufe et al., 2019 [Publ 3936], fig. 3-M

iT tT bi riS oT roT roS iD
1 1 0 1 18 1 1 -- 
  +-- Pilsbryoconcha_exilis **     
--+                             
  +-- Pseudodon_vondembuschianus

Froufe et al., 2019 [Publ 3936], fig. 4

iT tT bi riS oT roT roS iD
1 1 0 1 18 1 1 -- 
  +-- Pilsbryoconcha_exilis **     
--+                             
  +-- Pseudodon_vondembuschianus

Froufe et al., 2019 [Publ 3936], fig. 6

iT tT bi riS oT roT roS iD
1 1 0 1 44 1 1 -- 
  +-- Pilsbryoconcha_exilis **     
--+                             
  +-- Pseudodon_vondembuschianus

Pfeiffer et al., 2019 [Publ 3937], fig. 4-5

iT tT bi riS oT roT roS iD
1 1 0 1 56 3 2 -- 
  +-------- Bineurus_anodontinum      
--+                                   
  |  +----- Pilsbryoconcha_exilis **     
  +--+                                
     |  +-- Pseudodon_vondembuschianus
     +--+                             
        +-- Pseudodon_inoscularis

Pfeiffer et al., 2019 [Publ 3937], fig. 6

iT tT bi riS oT roT roS iD
1 1 0 1 56 3 2 -- 
  +-------- Pilsbryoconcha_exilis **     
--+                                   
  |  +----- Bineurus_anodontinum      
  +--+                                
     |  +-- Pseudodon_vondembuschianus
     +--+                             
        +-- Pseudodon_inoscularis

Zieritz et al., 2020 [Publ 4102], fig. 4

iT tT bi riS oT roT roS iD
1 1 0 1 30 0 0 -- 
-- Pilsbryoconcha_carinifera

Fukata & Iigo, 2020 [Publ 4106], fig. 1

iT tT bi riS oT roT roS iD
1 1 0 1 53 1 1 -- 
  +-- Pilsbryoconcha_exilis **     
--+                             
  +-- Pseudodon_vondembuschianus

Chen et al., 2020 [Publ 4170], fig. 1

iT tT bi riS oT roT roS iD
1 1 0 1 33 1 1 -- 
  +-- Pseudodon_vondembuschianus
--+                             
  +-- Pilsbryoconcha_exilis **

Wu et al., 2021 [Publ 4215], fig. 3

iT tT bi riS oT roT roS iD
1 1 0 1 23 2 2 -- 
  +-- Pseudodon_vagulus        
--+                            
  +-- Pilsbryoconcha_carinifera **

Zieritz et al., 2021 [Publ 4218], fig. 5

iT tT bi riS oT roT roS iD
1 1 0 1 31 6 4 -- 
  +-------------- Pilsbryoconcha_exilis **     
  |                                         
--+           +-- Pseudodon_mekongi         
  |  +--------+                             
  |  |        +-- Pseudodon_vondembuschianus
  +--+                                      
     |  +-------- Sundadontina              
     +--+                                   
        |  +----- Thaiconcha_callifera      
        +--+                                
           |  +-- Bineurus_loeiensis        
           +--+                             
              +-- Bineurus_anodontinum

Zieritz et al., 2021 [Publ 4231], fig. 3-F, 5

iT tT bi riS oT roT roS iD
1 1 0 1 48 1 1 -- 
  +-- Pilsbryoconcha_exilis **     
--+                             
  +-- Pseudodon_vondembuschianus

Zieritz et al., 2021 [Publ 4231], fig. 3-M, 4

iT tT bi riS oT roT roS iD
1 1 0 1 22 1 1 -- 
  +-- Pilsbryoconcha_exilis **     
--+                             
  +-- Pseudodon_vondembuschianus

Wu et al., 2022 [Publ 4238], fig. 3

iT tT bi riS oT roT roS iD
1 1 0 1 73 1 1 -- 
  +-- Pilsbryoconcha_exilis **     
--+                             
  +-- Pseudodon_vondembuschianus

Pavan-Kumar et al., 2022 [Publ 4263], fig. 4

iT tT bi riS oT roT roS iD
1 1 0 1 35 1 1 -- 
  +-- Pseudodon_vondembuschianus
--+                             
  +-- Pilsbryoconcha_exilis **

Pavan-Kumar et al., 2022 [Publ 4263], fig. S5

iT tT bi riS oT roT roS iD
1 1 0 1 38 1 1 -- 
  +-- Pseudodon_vondembuschianus
--+                             
  +-- Pilsbryoconcha_exilis **

Dai et al., 2023 [Publ 4285], fig. 5

iT tT bi riS oT roT roS iD
1 1 0 1 52 1 1 -- 
  +-- Pseudodon_vondembuschianus
--+                             
  +-- Pilsbryoconcha_exilis **

Jeratthitikul & Sutcharit, 2023 [Publ 4289], fig. 1

iT tT bi riS oT roT roS iD
1 1 0 1 12 10 6 -- 
  +----- Pseudodon_vondembuschianus
--+                                
  |  +-- Pilsbryoconcha_exilis **     
  +--+                             
     +-- Thaiconcha_callifera

Kongim et al., 2023 [Publ 4290], fig. 1

iT tT bi riS oT roT roS iD
1 1 0 1 14 1 1 -- 
  +-- Namkongnaia_inkhavilayi
--+                          
  +-- Pilsbryoconcha_exilis **

Wu et al., 2023 [Publ 4339], fig. 4, S5-aa

iT tT bi riS oT roT roS iD
1 1 0 1 95 1 1 -- 
  +-- Pseudodon_vondembuschianus
--+                             
  +-- Pilsbryoconcha_exilis **

Wu et al., 2023 [Publ 4339], fig. 4, S5-nt

iT tT bi riS oT roT roS iD
1 1 0 1 96 1 1 -- 
  +-- Pseudodon_vondembuschianus
--+                             
  +-- Pilsbryoconcha_exilis **

Wu et al., 2024 [Publ 4350], fig. 5a

iT tT bi riS oT roT roS iD
1 1 0 1 37 5 3 -- 
  +----------- Pilsbryoconcha_exilis **     
  |                                      
--+        +-- Pseudodon_mekongi         
  |  +-----+                             
  |  |     +-- Pseudodon_vondembuschianus
  +--+                                   
     |  +----- Thaiconcha_callifera      
     +--+                                
        |  +-- Bineurus_anodontinum      
        +--+                             
           +-- Bineurus_loeiensis

Wu et al., 2024 [Publ 4350], fig. 5b

iT tT bi riS oT roT roS iD
1 1 0 1 37 5 3 -- 
  +----------- Pilsbryoconcha_exilis **     
  |                                      
--+        +-- Bineurus_anodontinum      
  |  +-----+                             
  |  |     +-- Bineurus_loeiensis        
  +--+                                   
     |  +----- Thaiconcha_callifera      
     +--+                                
        |  +-- Pseudodon_mekongi         
        +--+                             
           +-- Pseudodon_vondembuschianus

Wu et al., 2024 [Publ 4350], fig. 5c

iT tT bi riS oT roT roS iD
1 1 0 1 37 5 3 -- 
  +----------- Pilsbryoconcha_exilis **     
  |                                      
--+  +-------- Bineurus_anodontinum      
  |  |                                   
  +--+-------- Bineurus_loeiensis        
     |                                   
     |  +----- Thaiconcha_callifera      
     +--+                                
        |  +-- Pseudodon_mekongi         
        +--+                             
           +-- Pseudodon_vondembuschianus

Wu et al., 2024 [Publ 4350], fig. 6-7

iT tT bi riS oT roT roS iD
1 1 0 1 37 5 3 -- 
  +----------- Pilsbryoconcha_exilis **     
  |                                      
--+        +-- Pseudodon_mekongi         
  |  +-----+                             
  |  |     +-- Pseudodon_vondembuschianus
  +--+                                   
     |  +----- Thaiconcha_callifera      
     +--+                                
        |  +-- Bineurus_anodontinum      
        +--+                             
           +-- Bineurus_loeiensis

Wu et al., 2024 [Publ 4350], fig. 8a, S1

iT tT bi riS oT roT roS iD
1 1 0 1 91 1 1 -- 
  +-- Pseudodon_vondembuschianus
--+                             
  +-- Pilsbryoconcha_exilis **

Wu et al., 2024 [Publ 4350], fig. 8b, S2

iT tT bi riS oT roT roS iD
1 1 0 1 91 1 1 -- 
  +-- Pseudodon_vondembuschianus
--+                             
  +-- Pilsbryoconcha_exilis **

Dai et al., 2024 [Publ 4358], fig. 4

iT tT bi riS oT roT roS iD
1 1 0 1 51 1 1 -- 
  +-- Pseudodon_vondembuschianus
--+                             
  +-- Pilsbryoconcha_exilis **

Page updated: 07:04:00 Thu 04 Apr 2024
 
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