Unionoida | Unionoidea | Unionidae | Unioninae | Anodontini | Cristariina genus Beringiana Starobogatov in Zatravkin, 1983: cladomics 4 valid species
Cladomics Summary
number of species = 4
analyzed in 31 cladograms in 21 publications
maximum number of terminals in an analysis = 20, mean = 2.68
Basic Monophyly
number of cladograms that test basic monophyly (i.e., more than 1 included species) = 14
maximum number of included terminals = 20, mean = 4.71
proportion of cladograms that support basic monophyly = 0.86
Pairwise Cladogram Clustering
The graphs below depict in two dimensions all pair-wise ingroup distances within the cladome. See the Cladmoics FARQ for more information.
Mabuchi & Nishida, 2020 [Publ 4120], fig. 1
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
20 |
28 |
3 |
20 |
17 |
10 |
2 |
0.0 |
*** Top Monophyly Cladogram ***
+-------------- Cristaria_plicata
|
| +----------- Sinanodonta_tumens
| |
| | +-- Sinanodonta_lauta[5]
| | |
--+ | +-- Sinanodonta_lauta[6]
| | |
| | +-- Beringiana_japonica[15] **
| | |
| | +-- Beringiana_japonica[16] **
| | +-----+
| | | +-- Beringiana_japonica[17] **
+--+ | |
| | +-- Beringiana_japonica[18] **
| | |
| | +-- Beringiana_japonica[19] **
| | |
| | +-- Beringiana_japonica[20] **
| |
| | +----- Beringiana_japonica[1] **
| | |
| | +----- Beringiana_japonica[2] **
| | |
+--+ +----- Beringiana_japonica[3] **
| |
| +----- Beringiana_japonica[4] **
| |
| +----- Beringiana_japonica[5] **
| |
| +----- Sinanodonta_lauta[1]
| |
| +----- Beringiana_japonica[6] **
| |
| +----- Sinanodonta_lauta[2]
| |
| +----- Sinanodonta_calipygos[1]
| |
+--+----- Beringiana_japonica[7] **
|
+----- Sinanodonta_lauta[4]
|
+----- Beringiana_japonica[8] **
|
+----- Beringiana_japonica[9] **
|
+----- Beringiana_japonica[10] **
|
+----- Beringiana_japonica[11] **
|
+----- Beringiana_japonica[12] **
|
+----- Beringiana_japonica[13] **
|
+----- Beringiana_japonica[14] **
|
| +-- Sinanodonta_lauta[3]
+--+
+-- Sinanodonta_calipygos[2]
Bogan et al., 2023 [Publ 4295], fig. 3
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
4 |
4 |
2 |
4 |
27 |
23 |
6 |
28.0 |
*** Top Sister-Group Cladogram ***
+-- Pletholophus_honglinhensis
+--------------------+
| +-- Pletholophus_tenuis
|
| +-------- Beringiana_japonica **
--+ +-----------+
| | | +----- Beringiana_beringiana **
| | +--+
| | | +-- Beringiana_fukuharai **
| | +--+
+--+ +-- Beringiana_gosannensis **
|
| +-- Sinanodonta_angula
| +--------------+
| | +-- Sinanodonta_lucida
| |
+--+ +-------------- Sinanodonta_schrenkii
| |
| | +-- Sinanodonta_calipygos
+--+-----------+
| +-- Sinanodonta_tumens
|
| +----------- Sinanodonta_woodiana[2]
+--+
| +-------- Sinanodonta_jourdyi
+--+
| +----- Sinanodonta_elliptica
+--+
| +-- Sinanodonta_lauta
+--+
+-- Sinanodonta_woodiana[1]
Sano et al., 2020 [Publ 4138], fig. 1
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
5 |
34 |
5 |
5 |
43 |
11 |
5 |
24.0 |
+-------- Lanceolaria_grayii[1]
|
+-------- Lanceolaria_grayii[2]
|
+-------- Cristaria_plicata[1]
|
+-------- Cristaria_plicata[2]
|
+-------- Sinanodonta_lauta
|
+-------- Beringiana_japonica **
|
+-------- Beringiana_fukuharai[1] **
|
+-------- Beringiana_beringiana **
|
+-------- Beringiana_fukuharai[2] **
|
+-------- Beringiana_fukuharai[3] **
|
+-------- Buldowskia_iwakawai[1]
|
+-------- Buldowskia_iwakawai[2]
|
+-------- Sinanodonta_tumens[1]
|
+-------- Sinanodonta_tumens[2]
|
--+-------- Pletholophus_tenuis
|
+-------- Anemina_arcaeformis
|
+-------- Lanceolaria_lanceolata
|
+-------- Cristaria_plicata[3]
|
+-------- Lanceolaria_gladiola
|
+-------- Sinanodonta_angula
|
+-------- Lepidodesma_languilati
|
| +-- Aculamprotula_tortuosa
+-----+
| +-- Acuticosta_chinensis
|
| +----- Cuneopsis_celtiformis
| |
+--+ +-- Nodularia_douglasiae[1]
| | |
| +--+-- Nodularia_douglasiae[2]
| |
| +-- Nodularia_nipponensis
|
| +----- Schistodesmus_lampreyanus
| |
| | +-- Inversiunio_reinianus[1]
+--+ |
| +-- Inversiunio_reinianus[2]
| |
| +-- Inversiunio_jokohamensis[1]
+--+
+-- Inversiunio_jokohamensis[2]
|
+-- Inversiunio_jokohamensis[3]
|
+-- Inversiunio_jokohamensis[4]
Lopes-Lima et al., 2020 [Publ 4080], fig. 2-4
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
4 |
4 |
2 |
4 |
118 |
20 |
6 |
28.0 |
+-------------------- Pletholophus_tenuis
|
| +-------- Beringiana_japonica **
| +--------+
--+ | | +----- Beringiana_beringiana **
| | +--+
| | | +-- Beringiana_fukuharai **
| | +--+
+--+ +-- Beringiana_gosannensis **
|
| +-- Sinanodonta_angula
| +-----------+
| | +-- Sinanodonta_lucida
| |
+--+ +----------- Sinanodonta_schrenkii
| |
| | +-- Sinanodonta_calipygos
| +--------+
+--+ +-- Sinanodonta_tumens
|
| +-------- Sinanodonta_woodiana[2]
| |
+--+ +----- Sinanodonta_elliptica
| |
+--+----- Sinanodonta_jourdyi
|
| +-- Sinanodonta_lauta
+--+
+-- Sinanodonta_woodiana[1]
Lopes-Lima et al., 2020 [Publ 4080], fig. 7
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
4 |
4 |
2 |
4 |
4 |
2 |
1 |
28.0 |
+-- Sinanodonta_woodiana
+--------+
| +-- Sinanodonta_tumens
--+
| +-------- Beringiana_gosannensis **
+--+
| +----- Beringiana_fukuharai **
+--+
| +-- Beringiana_beringiana **
+--+
+-- Beringiana_japonica **
Sano et al., 2022 [Publ 4311], fig. 2
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
4 |
4 |
2 |
4 |
10 |
10 |
4 |
28.0 |
+-------- Pletholophus_tenuis
+-----------+
| | +----- Cristaria_plicata
| +--+
| | +-- Buldowskia_iwakawai
| +--+
| +-- Buldowskia_kamiyai
--+
| +-------- Beringiana_fukuharai **
| +--------+
| | | +----- Beringiana_japonica **
| | +--+
| | | +-- Beringiana_gosannensis **
+--+ +--+
| +-- Beringiana_beringiana **
|
| +-------------- Sinanodonta_tumens
+--+
| +----------- Sinanodonta[2]
+--+
| +-------- Sinanodonta_calipygos
+--+
| +----- Sinanodonta_lauta
+--+
| +-- Sinanodonta_woodiana
+--+
+-- Sinanodonta[1]
Sano et al., 2022 [Publ 4311], fig. 3
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
4 |
4 |
2 |
4 |
14 |
14 |
4 |
28.0 |
+-------- Pletholophus_tenuis
+--------------+
| | +----- Cristaria_plicata
| +--+
| | +-- Buldowskia_kamiyai
| +--+
| +-- Buldowskia_iwakawai
--+
| +-------- Beringiana_fukuharai **
| +-----------+
| | | +----- Beringiana_gosannensis **
| | +--+
| | | +-- Beringiana_japonica **
| | +--+
+--+ +-- Beringiana_beringiana **
|
| +-- Sinanodonta_angula
| +--------------+
| | +-- Sinanodonta_lucida
| |
+--+ +----- Sinanodonta_schrenkii
| +--------+
| | | +-- Sinanodonta_tumens
| | +--+
+--+ +-- Sinanodonta_calipygos
|
| +----------- Sinanodonta_woodiana[2]
+--+
| +-------- Sinanodonta_jourdyi
+--+
| +----- Sinanodonta_elliptica
+--+
| +-- Sinanodonta_woodiana[1]
+--+
+-- Sinanodonta_lauta
Bulakhova et al., 2023 [Publ 4325], fig. 5
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
4 |
4 |
2 |
4 |
0 |
0 |
0 |
28.0 |
+-- Beringiana_beringiana **
+--+
| +-- Beringiana_japonica **
--+
| +-- Beringiana_gosannensis **
+--+
+-- Beringiana_fukuharai **
Sano et al., 2017 [Publ 3799], fig. 1
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
3 |
3 |
1 |
3 |
24 |
5 |
4 |
28.0 |
+----------- Pletholophus_tenuis
|
--+ +-- Sinanodonta_tumens
| +-----+
| | +-- Sinanodonta_lauta
+--+
| +----- Beringiana_fukuharai **
+--+
| +-- Beringiana_japonica **
+--+
+-- Beringiana_beringiana **
Sano et al., 2017 [Publ 3799], fig. 2
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
3 |
3 |
1 |
3 |
33 |
6 |
4 |
28.0 |
+-- Cristaria_plicata
+-----------+
| +-- Buldowskia_iwakawai
|
--+ +----- Beringiana_fukuharai **
| +-----+
| | | +-- Beringiana_japonica **
| | +--+
+--+ +-- Beringiana_beringiana **
|
| +-------- Pletholophus_tenuis
+--+
| +----- Sinanodonta_woodiana
+--+
| +-- Sinanodonta_lauta
+--+
+-- Sinanodonta_tumens
Sano et al., 2020 [Publ 4138], fig. 2
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
3 |
3 |
1 |
3 |
70 |
9 |
5 |
28.0 |
+-------------- Pletholophus_tenuis
|
| +----- Beringiana_fukuharai **
--+ +-----+
| | | +-- Beringiana_japonica **
| | +--+
+--+ +-- Beringiana_beringiana **
|
| +-------- Sinanodonta_lucida
+--+
| +----- Sinanodonta_tumens
+--+
| +-- Sinanodonta_lauta
+--+
+-- Sinanodonta_woodiana
Sano et al., 2020 [Publ 4138], fig. 3
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
3 |
3 |
1 |
3 |
69 |
9 |
5 |
28.0 |
+-------- Pletholophus_tenuis
+--+
| | +----- Beringiana_fukuharai **
| +--+
| | +-- Beringiana_japonica **
--+ +--+
| +-- Beringiana_beringiana **
|
| +-------- Sinanodonta_lucida
+--+
| +----- Sinanodonta_tumens
+--+
| +-- Sinanodonta_woodiana
+--+
+-- Sinanodonta_lauta
Sano et al., 2020 [Publ 4138], fig. 4
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
3 |
3 |
1 |
3 |
69 |
9 |
5 |
28.0 |
+-------- Sinanodonta_lucida
+--+
| | +----- Sinanodonta_tumens
| +--+
| | +-- Sinanodonta_woodiana
--+ +--+
| +-- Sinanodonta_lauta
|
| +-------- Pletholophus_tenuis
+--+
| +----- Beringiana_fukuharai **
+--+
| +-- Beringiana_japonica **
+--+
+-- Beringiana_beringiana **
Lopes-Lima et al., 2020 [Publ 4080], fig. 10
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
2 |
2 |
0 |
2 |
12 |
10 |
1 |
-- |
+-- Beringiana_japonica **
+--------------+
| +-- Beringiana_beringiana **
|
--+ +-- Sinanodonta_angula
| +-----------+
| | +-- Sinanodonta_lucida
| |
+--+ +----------- Sinanodonta_schrenkii
| |
| | +-- Sinanodonta_calipygos
| +--------+
+--+ +-- Sinanodonta_tumens
|
| +-------- Sinanodonta_woodiana[2]
| |
+--+-------- Sinanodonta_jourdyi
|
| +----- Sinanodonta_elliptica
+--+
| +-- Sinanodonta_lauta
+--+
+-- Sinanodonta_woodiana[1]
Chong et al., 2008 [Publ 2668], fig. 3
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
1 |
1 |
0 |
1 |
20 |
1 |
1 |
-- |
+-- Beringiana_beringiana **
--+
+-- Sinanodonta_woodiana
Gustafson & Iwamoto, 2005 [Publ 2741], fig. 2
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
1 |
1 |
0 |
1 |
4 |
0 |
0 |
-- |
-- Beringiana_beringiana
Soroka, 2010 [Publ 3404], fig. 9a
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
1 |
1 |
0 |
1 |
4 |
4 |
2 |
-- |
+----------- Buldowskia_flavotincta
--+
| +-------- Beringiana_beringiana **
+--+
| +----- Sinanodonta_schrenkii
+--+
| +-- Sinanodonta_lauta
+--+
+-- Sinanodonta_woodiana
Klishko et al., 2014 [Publ 3666], fig. 8
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
1 |
1 |
0 |
1 |
3 |
3 |
2 |
-- |
+-- Cristaria_plicata
+--+
| +-- Cristaria_radiata
--+
| +-- Beringiana_beringiana **
+--+
+-- Sinanodonta_woodiana
Klishko et al., 2016 [Publ 3733], fig. 1
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
1 |
1 |
0 |
1 |
3 |
3 |
2 |
-- |
+-- Cristaria_plicata
+--+
| +-- Cristaria_radiata
--+
| +-- Sinanodonta_woodiana
+--+
+-- Beringiana_beringiana **
Sayenko et al., 2017 [Publ 3760], fig. 3
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
1 |
1 |
0 |
1 |
3 |
3 |
1 |
-- |
+-------- Beringiana_beringiana **
--+
| +----- Sinanodonta_schrenkii
+--+
| +-- Sinanodonta_woodiana
+--+
+-- Sinanodonta_lauta
Froufe et al., 2017 [Publ 3802], fig. 6
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
1 |
1 |
0 |
1 |
11 |
9 |
3 |
-- |
+-- Cristaria_plicata
+--------------+
| +-- Cristaria_radiata
--+
| +-------------- Beringiana_beringiana **
| |
+--+ +----------- Sinanodonta_lucida
| |
+--+ +-------- Sinanodonta_schrenkii
| |
+--+ +----- Sinanodonta_woodiana[1]
| |
+--+ +-- Sinanodonta_lauta
| |
+--+-- Sinanodonta_elliptica
|
+-- Sinanodonta_woodiana[2]
Inoue et al., 2018 [Publ 3858], fig. 1, 3a-c
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
1 |
1 |
0 |
1 |
204 |
0 |
0 |
-- |
-- Beringiana_beringiana
Bolotov et al., 2020 [Publ 4084], fig. 1
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
1 |
1 |
0 |
1 |
35 |
8 |
5 |
-- |
+----------- Cristaria_plicata
--+
| +-------- Beringiana_beringiana **
+--+
| +----- Sinanodonta_schrenkii
+--+
| +-- Sinanodonta_lauta
+--+
+-- Sinanodonta_woodiana
Kano et al., 2019 [Publ 4194], fig. 6
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
1 |
1 |
0 |
1 |
9 |
2 |
2 |
-- |
+----- Sinanodonta_lauta
--+
| +-- Pletholophus_tenuis
+--+
+-- Beringiana_japonica **
Konopleva et al., 2023 [Publ 4269], fig. S1, 4
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
1 |
1 |
0 |
1 |
55 |
12 |
5 |
-- |
+-------- Beringiana_beringiana **
+--+
| | +----- Sinanodonta_schrenkii
| +--+
| | +-- Sinanodonta_woodiana
--+ +--+
| +-- Sinanodonta_lauta
|
| +-------- Cristaria_radiata
| |
+--+ +-- Cristaria_truncata
| +--+
| | +-- Cristaria_plicata
+--+
| +-- Cristaria_bellua
+--+
+-- Cristaria_clessini
Chen et al., 2023 [Publ 4302], fig. 1
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
1 |
1 |
0 |
1 |
39 |
5 |
5 |
-- |
+----- Pletholophus_tenuis
--+
| +-- Sinanodonta_jourdyi
+--+
+-- Beringiana_beringiana **
Chen et al., 2023 [Publ 4302], fig. 2
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
1 |
1 |
0 |
1 |
39 |
5 |
5 |
-- |
+----- Pletholophus_tenuis
--+
| +-- Sinanodonta_jourdyi
+--+
+-- Beringiana_beringiana **
Wu et al., 2023 [Publ 4339], fig. 4, S5-aa
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
1 |
1 |
0 |
1 |
95 |
6 |
3 |
-- |
+----- Cristaria_plicata
+-----+
| | +-- Anemina_arcaeformis
| +--+
--+ +-- Anemina_euscaphys
|
| +-------- Beringiana_fukuharai **
+--+
| +----- Sinanodonta_lucida
+--+
| +-- Sinanodonta_tumens
+--+
+-- Sinanodonta_woodiana
Wu et al., 2023 [Publ 4339], fig. 4, S5-nt
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
1 |
1 |
0 |
1 |
96 |
6 |
3 |
-- |
+-- Anemina_arcaeformis
+--------+
| +-- Anemina_euscaphys
--+
| +-------- Beringiana_fukuharai **
+--+
| +----- Sinanodonta_lucida
+--+
| +-- Sinanodonta_tumens
+--+
+-- Sinanodonta_woodiana
Wu et al., 2024 [Publ 4350], fig. 8a, S1
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
1 |
1 |
0 |
1 |
91 |
6 |
3 |
-- |
+-- Anemina_arcaeformis
+--------+
| +-- Anemina_euscaphys
--+
| +-------- Beringiana_fukuharai **
+--+
| +----- Sinanodonta_lucida
+--+
| +-- Sinanodonta_tumens
+--+
+-- Sinanodonta_woodiana
Wu et al., 2024 [Publ 4350], fig. 8b, S2
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
1 |
1 |
0 |
1 |
91 |
6 |
3 |
-- |
+-- Anemina_arcaeformis
+--------+
| +-- Anemina_euscaphys
--+
| +-------- Beringiana_fukuharai **
+--+
| +----- Sinanodonta_lucida
+--+
| +-- Sinanodonta_tumens
+--+
+-- Sinanodonta_woodiana
|