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Page last updated
Tue 04 Jan 2022

The Freshwater Mussels (Unionoida) of the World (and other less consequential bivalves)

FM(U)otW(aolcb) is the web version of the MUSSEL Project Database. Follow the links to browse the data or use the custom Google search field. Either way, you win!

The nominal species (available and otherwise) in this synonymy are listed by usage rather than strictly by publication date. The primary sorting order is by the most recent year (since 1969) that the species name was treated as valid, and then nominal species are sorted secondarily by publication year.

The taxonomic history of the valid species (since 1900) is listed at the bottom of the page.

Search the database pages:

Unionoida | Unionoidea | Unionidae | Ambleminae | Lampsilini | Epioblasma

species Epioblasma torulosa (Rafinesque, 1820): nominal species
     5 specimens available.

Amblema torulosa Rafinesque, 1820
Rafinesque, 1820, Ann. Gén. Sci. Phys. 5 (13): 314, pl. 82, fig. 11-12 [also as ‘Unio torulosa’].
Rafinesque, 1864, [in] Binney & Tryon, reprint: 57.
Type(s): HOLOTYPE ANSP_20218, [no locality available - North America].

Amblema torulosaRafinesque (1820)
Epioblasma torulosaRafinesque (1831), Cummings & Mayer (1992), Parmalee & Bogan (1998), Graf (2002), ..., Bogan (2010), Haag (2012), Williams et al. (2017), Graf & Cummings (2021)
Unio torulosusFerussac (1835), Conrad (1853)
Unio gibbosusConrad (1836)
Truncilla torulosaVanatta (1915), Ortmann (1918)
Dysnomia (Pilea) torulosaOrtmann & Walker (1922), Ortmann (1925)
Dysnomia (s.s.) torulosaOrtmann (1924)
Dysnomia (Torulosa) torulosaFrierson (1927), Haas (1930), Haas (1969)
Epioblasma (sec. Torulosa) torulosumThiele (1934)
Dysnomia (Torulosa) torulosa torulosaHaas (1969)
Dysnomia torulosaStansbery (1970)
Dysnomia torulosa torulosaBurch (1975)
Plagiola (Torulosa) torulosaR.I. Johnson (1978)
Plagiola torulosaR.I. Johnson (1980)
Epioblasma torulosa torulosaTurgeon et al. (1988), Williams et al. (1993), Bogan (1993), Turgeon et al. (1998), Lydeard et al. (1999), Cicerello & Schuster (2003), Watters et al. (2009), ITIS (2012), IUCN (2012)

Amblema gibbosa Rafinesque, 1820
Rafinesque, 1818, Amer. Mon. Mag. 3: 355 [as ‘Potamilus gibbosus’]. [nomen nudum]
Rafinesque, 1820, Ann. Gén. Sci. Phys. 5 (13): 315 [also as ‘Unio gibbosa’].
Rafinesque, 1864, [in] Binney & Tryon, reprint: 23, 58.
Type(s): LECTOTYPE ANSP_20232, Riv. Ohio.

Amblema gibbosaRafinesque (1820)
Potamilus gibbosusRafinesque (1818)
Unio gibbosusFerussac (1835), Conrad (1836), Küster (1848), Reeve (1864)
Margarita (Unio) perplexusLea (1836), Lea (1838)
Dysnomia gibbosaAgassiz (1852)
Margaron (Unio) perplexusLea (1852), Lea (1870)
Unio torulosusConrad (1853)
Unio (Dysnomia) gibbosusMörch (1853)
Truncilla torulosaVanatta (1915), Ortmann (1918)
Dysnomia (Pilea) torulosaOrtmann & Walker (1922)
Dysnomia (Torulosa) torulosaFrierson (1927)
Dysnomia (Torulosa) torulosa torulosaHaas (1969)
Dysnomia torulosa torulosaBurch (1975)
Plagiola (Torulosa) torulosaR.I. Johnson (1978)
Epioblasma torulosaParmalee & Bogan (1998), Williams et al. (2008)
Epioblasma torulosa torulosaWatters et al. (2009)

Amblema gibbosa var. difformis Rafinesque, 1820
Rafinesque, 1820, Ann. Gén. Sci. Phys. 5 (13): 315.
Rafinesque, 1864, [in] Binney & Tryon, reprint: 58.
(intro'd as intra-sp. of Amblema gibbosa Rafinesque, 1820)

Amblema gibbosa var. difformisRafinesque (1820)
Dysnomia (Torulosa) torulosa torulosaHaas (1969)
Epioblasma torulosaParmalee & Bogan (1998), Williams et al. (2008)
Epioblasma torulosa torulosaWatters et al. (2009)

Amblema gibbosa var. olivacea Rafinesque, 1820
Rafinesque, 1820, Ann. Gén. Sci. Phys. 5 (13): 315.
Rafinesque, 1864, [in] Binney & Tryon, reprint: 58.
(intro'd as intra-sp. of Amblema gibbosa Rafinesque, 1820)

Amblema gibbosa var. olivaceaRafinesque (1820)
Dysnomia (Torulosa) torulosa torulosaHaas (1969)
Epioblasma torulosaParmalee & Bogan (1998), Williams et al. (2008)
Epioblasma torulosa torulosaWatters et al. (2009)

Amblema gibbosa var. radiata Rafinesque, 1820
Rafinesque, 1820, Ann. Gén. Sci. Phys. 5 (13): 315.
Rafinesque, 1864, [in] Binney & Tryon, reprint: 58.
(intro'd as intra-sp. of Amblema gibbosa Rafinesque, 1820)

Amblema gibbosa var. radiataRafinesque (1820)
Dysnomia (Torulosa) torulosa torulosaHaas (1969)
Epioblasma torulosaParmalee & Bogan (1998), Williams et al. (2008)
Epioblasma torulosa torulosaWatters et al. (2009)

Amblema torulosa var. angulata Rafinesque, 1820
Rafinesque, 1820, Ann. Gén. Sci. Phys. 5 (13): 315.
Rafinesque, 1864, [in] Binney & Tryon, reprint: 57.
(intro'd as intra-sp. of Amblema torulosa Rafinesque, 1820)

Amblema torulosa var. angulataRafinesque (1820)
Dysnomia (Torulosa) torulosa torulosaHaas (1969)
Plagiola (Torulosa) torulosaR.I. Johnson (1978)
Epioblasma torulosaParmalee & Bogan (1998), Williams et al. (2008)
Epioblasma torulosa torulosaWatters et al. (2009)

Unio perplexus Lea, 1831
Lea, 1831, Trans. Amer. Phil. Soc. 4 [O. 1]: 112, pl. 17, fig. 42.
(homonym of Unio perplexus Say, 1829)
Type(s): SYNTYPE USNM_84324, Cincinnati, Ohio.

Unio perplexusLea (1831), Ravenel (1834), Jay (1835), Hanley (1843), Lewis (1871), B.H. Wright (1888), de Gregorio (1914)
Unio gibbosusFerussac (1835), Conrad (1836), Küster (1848), Reeve (1864)
Margarita (Unio) perplexusLea (1836), Lea (1838)
Dysnomia gibbosaAgassiz (1852)
Margaron (Unio) perplexusLea (1852), Lea (1870)
Unio torulosusConrad (1853)
Unio (Dysnomia) gibbosusMörch (1853)
Unio (Rotundaria) perplexusAdams & Adams (1857)
Truncilla (Pilea) perplexaSimpson (1900), Simpson (1914)
Unio (s.s.) perplexusCall (1900)
Truncilla perplexaOrtmann (1909)
Truncilla torulosaVanatta (1915), Ortmann (1918)
Dysnomia (Pilea) torulosaOrtmann & Walker (1922)
Dysnomia (Torulosa) torulosaFrierson (1927)
Dysnomia (Torulosa) torulosa torulosaHaas (1969)
Dysnomia torulosa torulosaBurch (1975)
Plagiola (Torulosa) torulosaR.I. Johnson (1978)
Epioblasma torulosaStrayer & Jirka (1997), Parmalee & Bogan (1998), Williams et al. (2008)

Unio torulosus Conrad, 1834
‘Rafinesque’ Conrad, 1834, New Fresh-Water Shells: 72.
‘Raf.’ Conrad, 1853, Proc. Acad. Nat. Sci. 6: 259.
(reference to Amblema torulosa Rafinesque, 1820)

Unio torulosusConrad (1834)
Truncilla (Pilea) perplexaSimpson (1900), Simpson (1914)

Unio gibbosus Ferussac, 1835
‘Rafinesque’ of Authors.
Ferussac, 1835, Mag. Zool. 5: 28.
Conrad, 1836, Monogr. Fam. Unionidae 1(6): 50, pl. 27, fig. 1.
Reeve, 1864, Conch. Icon. 16: pl. 11, sp. 41, figs. 41a-b.
(reference to Amblema gibbosa Rafinesque, 1820)

Unio gibbosusFerussac (1835), Küster (1848)
Unio torulosusFerussac (1835)
Margarita (Unio) perplexusLea (1836), Lea (1838)
Unio perplexusHanley (1843)
Margaron (Unio) perplexusLea (1852), Lea (1870)
Truncilla (Pilea) perplexaSimpson (1900), Simpson (1914)
Dysnomia torulosa torulosaBurch (1975)

Unio obliquus Potiez & Michaud, 1844
‘Lamarck’ Potiez & Michaud, 1844, Galérie Moll. at. Mus. Douai 2: 153, pl. 58, figs. 3-4.
(misidentified reference to Unio obliqua Lamarck, 1819)

Unio obliquusPotiez & Michaud (1844)
Unio (Dysnomia) gibbosusMörch (1853)
Truncilla (Pilea) perplexaSimpson (1900), Simpson (1914)
Dysnomia (Torulosa) torulosaFrierson (1927)
Dysnomia (Torulosa) torulosa torulosaHaas (1969)
Dysnomia torulosa torulosaBurch (1975)
Plagiola (Torulosa) torulosaR.I. Johnson (1978)
Epioblasma torulosaParmalee & Bogan (1998)

Unio phillipsii Reeve, 1864
‘Conrad’ Reeve, 1864, Conch. Icon. 16: pl. 4, sp. 15.
(misidentified reference to Unio phillipsii Conrad, 1835)

Unio phillipsiiReeve (1864)
Truncilla (Pilea) perplexa var. cincinnatiensisSimpson (1900), Simpson (1914)
Plagiola (Torulosa) torulosaR.I. Johnson (1978)

Unio abnormis Haas, 1930
‘Say’ Haas, 1930, Senckenbergiana 12(6): 329. [MS]
(intro'd as syn of Amblema torulosa Rafinesque, 1820)

Unio abnormisHaas (1930)
Dysnomia (Torulosa) torulosaHaas (1930)
Epioblasma torulosaParmalee & Bogan (1998)

Taxonomic History (since 1900)

Truncilla (Pilea) perplexaSimpson (1900), Simpson (1914)
Unio (s.s.) perplexusCall (1900)
Truncilla (Pilea) perplexa var. cincinnatiensisSimpson (1900), Simpson (1914)
Truncilla perplexaOrtmann (1909)
Unio perplexusde Gregorio (1914)
Truncilla torulosaVanatta (1915), Ortmann (1918)
Dysnomia (Pilea) torulosaOrtmann & Walker (1922), Ortmann (1925)
Dysnomia (s.s.) torulosaOrtmann (1924)
Dysnomia (Torulosa) torulosaFrierson (1927), Haas (1930), Haas (1969)
Unio abnormisHaas (1930)
Epioblasma (sec. Torulosa) torulosumThiele (1934)
Dysnomia (Torulosa) torulosa torulosaHaas (1969)
Dysnomia torulosaStansbery (1970)
Dysnomia torulosa torulosaBurch (1975)
Plagiola (Torulosa) torulosaR.I. Johnson (1978)
Plagiola torulosaR.I. Johnson (1980)
Epioblasma torulosa torulosaTurgeon et al. (1988), Williams et al. (1993), Bogan (1993), Turgeon et al. (1998), Lydeard et al. (1999), Cicerello & Schuster (2003), Watters et al. (2009), ITIS (2012), IUCN (2012)
Epioblasma torulosaCummings & Mayer (1992), Strayer & Jirka (1997), Parmalee & Bogan (1998), Graf (2002), ..., Bogan (2010), Haag (2012), Williams et al. (2017), Graf & Cummings (2021)

Page updated: 09:26:36 Tue 04 Jan 2022

 
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