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Page last updated
Sun 31 Oct 2021

The Freshwater Mussels (Unionoida) of the World (and other less consequential bivalves)

FM(U)otW(aolcb) is the web version of the MUSSEL Project Database. Follow the links to browse the data or use the custom Google search field. Either way, you win!

The nominal species (available and otherwise) in this synonymy are listed by usage rather than strictly by publication date. The primary sorting order is by the most recent year (since 1969) that the species name was treated as valid, and then nominal species are sorted secondarily by publication year.

The taxonomic history of the valid species (since 1900) is listed at the bottom of the page.

Search the database pages:

Unionoida | Unionoidea | Unionidae | Ambleminae | Pleurobemini | Pleurobema

species Pleurobema clava (Lamarck, 1819): nominal species
     8 specimens available.

Unio clava Lamarck, 1819
Lamarck, 1819, Hist. Natur. Anim. sans Vert. 6: 74, sp. 18.
Type(s): TYPE MNHN_MPL_0015, Amer. Sept.

Unio clavaLamarck (1819), Menke (1828), Menke (1830), Conrad (1835), Ferussac (1835), Küster (1848), Conrad (1853), Sowerby (1868)
Unio scaleniaLea (1834)
Margarita (Unio) clavusLea (1836), Lea (1838)
Unio clavusHanley (1843), B.H. Wright (1888), Call (1895)
Pleurobema clavaAgassiz (1852), Ortmann (1909), Ortmann (1909), Ortmann (1909), ..., Pieri et al. (2018), Inoue et al. (2019), Smith et al. (2020), Graf & Cummings (2021)
Margaron (Unio) clavusLea (1852), Lea (1870)
Unio (Lampsilis) clavusAdams & Adams (1857)
Unio (Limnium sec. Scalenaria) clavusFischer (1887)
Pleurobema (s.s.) clavaSimpson (1900), Simpson (1914)
Unio (s.s.) clavusCall (1900)
Pleurobema (s.s.) mytiloidesFrierson (1927)
Pleurobema mytiloidesModell (1942)
Pleurobema clava clavaHaas (1969)

Unio bournianus Lea, 1840
Lea, 1840, Proc. Amer. Phil. Soc. 1: 288.
Lea, 1842, Trans. Amer. Phil. Soc. 8 [O. 3]: 213 [51], pl. 15, fig. 28.
Type(s): HOLOTYPE USNM_84683, Scioto R., Chillicothe, Ohio.

Unio bournianusLea (1840), Conrad (1853), B.H. Wright (1888)
Margaron (Unio) bournianusLea (1852), Lea (1870)
Unio (Lampsilis) bournianusAdams & Adams (1857)
Unio (sec. Lampsilis) bournianusPaetel (1890)
Pleurobema (s.s.) bournianusSimpson (1900)
Pleurobema (s.s.) bournianumSimpson (1914)
Quadrula (Pleuronaia) corFrierson (1927), Haas (1969)
Pleurobema bournianumTurgeon et al. (1988), Williams et al. (1993), Turgeon et al. (1998), IUCN (2000), Graf & Cummings (2007), Cummings & Graf (2010), ITIS (2012)
Pleurobema clavaWilliams et al. (2008), Watters et al. (2009), Williams et al. (2017), Graf & Cummings (2021)

Pleurobema mytiloides Rafinesque, 1820
Rafinesque, 1819, Jour. Phys. Chim. Hist. Nat. 88: 427 [as ‘Unio (Pleurobema) mytiloïdes’]. [nomen nudum]
Rafinesque, 1820, Ann. Gén. Sci. Phys. 5 (13): 313, pl. 82, figs. 8-10 [also as ‘U. mytiloides’].
Rafinesque, 1864, [in] Binney & Tryon, reprint: 29, 56.
Type(s): LECTOTYPE ANSP_20223, [no locality available - North America].

Pleurobema mytiloidesRafinesque (1820), Agassiz (1852), Modell (1942), Haas (1969)
Unio (Pleurobema) mytiloidesRafinesque (1819)
Unio mytiloidesLea (1829), Ravenel (1834), Jay (1835), Ferussac (1835), Conrad (1836), Küster (1852), Lapham (1860), Küster (1862), Lewis (1871), B.H. Wright (1888)
Margarita (Unio) mytiloidesLea (1836), Lea (1838)
Unio (s.s.) mytiloidesSwainson (1840)
Unio pyramidatusHanley (1843)
Margaron (Unio) mytiloidesLea (1852), Lea (1870)
Unio clavaConrad (1853)
Unio (Obovaria) mytiloidesMörch (1853)
Unio (Lampsilis) mytiloidesAdams & Adams (1857)
Unio (Limnium sec. Pleurobema) mytiloidesFischer (1887)
Pleurobema clavaVanatta (1915), Ortmann & Walker (1922), F.C. Baker (1928), Thiele (1934), ..., Williams et al. (2008), Watters et al. (2009), Campbell & Lydeard (2012), Graf & Cummings (2021)
Pleurobema obliqua rubrumOrtmann (1918)
Pleurobema (s.s.) mytiloidesFrierson (1927), Haas (1930)
Pleurobema clava clavaHaas (1969)

Unio patulus Lea, 1829
Lea, 1829, Trans. Amer. Phil. Soc. 3 [O. 1]: 441, pl. 12, fig. 20.
Type(s): HOLOTYPE USNM_84760, Ohio ?

Unio patulusLea (1829), Jay (1835), Conrad (1838), Hanley (1843), Küster (1862), Reeve (1864), Lewis (1871), B.H. Wright (1888), de Gregorio (1914)
Unio cuneatusFerussac (1835)
Margarita (Unio) patulusLea (1836), Lea (1838)
Unio (Cunicula) patulaSwainson (1840)
Margaron (Unio) patulusLea (1852), Lea (1870)
Unio (Lampsilis) patulusAdams & Adams (1857)
Unio clavusCall (1895)
Pleurobema (s.s.) clavaSimpson (1900), Simpson (1914)
Unio (s.s.) clavusCall (1900)
Pleurobema clavaOrtmann & Walker (1922), Burch (1975), Strayer & Jirka (1997), Parmalee & Bogan (1998), Williams et al. (2008), Watters et al. (2009)
Pleurobema (s.s.) clava-patulumFrierson (1927)
Pleurobema clava patulusHaas (1969)

Obliquaria (Scalenaria) scalenia Rafinesque, 1820
Rafinesque, 1820, Ann. Gén. Sci. Phys. 5 (13): 309, pl. 81, figs. 24-25 [also as ‘U. scalenia’].
Rafinesque, 1864, [in] Binney & Tryon, reprint: 53.
Type(s): NEOTYPE ANSP_20229, Ohio.

Obliquaria (Scalenaria) scaleniaRafinesque (1820)
Unio scaleniusLea (1829), Ravenel (1834), Jay (1835)
Unio scaleniaLea (1834)
Unio clavaFerussac (1835), Conrad (1835), Küster (1848), Conrad (1853)
Margarita (Unio) clavusLea (1836), Lea (1838)
Unio clavusHanley (1843)
Margaron (Unio) clavusLea (1852), Lea (1870)
Unio (Scalenaria) scalenariusMörch (1853)
Pleurobema clavaVanatta (1915), Ortmann & Walker (1922), Burch (1975), Parmalee & Bogan (1998), Williams et al. (2008), Watters et al. (2009), Campbell & Lydeard (2012)

Pleurobema cuneata Rafinesque, 1820
Rafinesque, 1820, Ann. Gén. Sci. Phys. 5 (13): 313 [also as ‘Unio cuneata’].
Rafinesque, 1864, [in] Binney & Tryon, reprint: 56.
Type(s): LECTOTYPE ANSP_20228, Riv. Ohio.

Pleurobema cuneataRafinesque (1820)
Unio cuneatusFerussac (1835), Reeve (1864)
Unio (Cunicula) cuneataSwainson (1840)
Unio clavaConrad (1853)
Pleurobema clavaVanatta (1915), Ortmann & Walker (1922), Burch (1975), Parmalee & Bogan (1998), Williams et al. (2008), Watters et al. (2009)
Pleurobema (s.s.) mytiloidesFrierson (1927)

Pleurobema cuneata var. maculata Rafinesque, 1820
Rafinesque, 1820, Ann. Gén. Sci. Phys. 5 (13): 313.
Rafinesque, 1864, [in] Binney & Tryon, reprint: 56.
(intro'd as intra-sp. of Pleurobema cuneata Rafinesque, 1820)

Pleurobema cuneata var. maculataRafinesque (1820)

Pleurobema cuneata var. sulcata Rafinesque, 1820
Rafinesque, 1820, Ann. Gén. Sci. Phys. 5 (13): 313.
Rafinesque, 1864, [in] Binney & Tryon, reprint: 56.
(intro'd as intra-sp. of Pleurobema cuneata Rafinesque, 1820)

Pleurobema cuneata var. sulcataRafinesque (1820)

Unio (Aximedia) elliptica Rafinesque, 1820
Rafinesque, 1818, Amer. Mon. Mag. 3: 355 [as ‘Potamilus ellipticus’]. [nomen nudum]
Rafinesque, 1820, Ann. Gén. Sci. Phys. 5 (13): 296 [also as ‘Elliptio elliptica’].
Rafinesque, 1864, [in] Binney & Tryon, reprint: 22, 42.
Type(s): LECTOTYPE ANSP_20213, [no locality available - North America].

Unio (Aximedia) ellipticaRafinesque (1820)
Potamilus ellipticusRafinesque (1818)
Obliquaria (Aximedia) ellipticaRafinesque (1831)
Unio ellipticaFerussac (1835)
Unio (Limnium sec. Aximedia) ellipticusFischer (1887)
Pleurobema clavaVanatta (1915), Ortmann & Walker (1922), Burch (1975), Parmalee & Bogan (1998), Williams et al. (2008), Watters et al. (2009)
Obovaria (s.s.) subrotundaFrierson (1927), Haas (1969)

Unio cuneatus Say, 1834
‘Rafinesque’ Say, 1834, Amer. Conch. 6: ??.
(reference to Pleurobema cuneata Rafinesque, 1820)

Unio cuneatusSay (1834)
Pleurobema (s.s.) clavaSimpson (1900), Simpson (1914)
Pleurobema clavaParmalee & Bogan (1998), Watters et al. (2009)

Unio modioliformis Lea, 1834
‘Say’ of Authors.
Lea, 1834, Trans. Amer. Phil. Soc. 5 [O. 1]: 89.
Conrad, 1835, Monogr. Fam. Unionidae 1(1): 5.
Ferussac, 1835, Mag. Zool. 5: 28.
Conrad, 1853, Proc. Acad. Nat. Sci. 6: 246.
(intro'd as syn of Unio clava Lamarck, 1819)

Unio modioliformisLea (1834)
Unio scaleniaLea (1834)
Unio clavaFerussac (1835), Conrad (1835), Küster (1848), Conrad (1853)
Margarita (Unio) clavusLea (1836), Lea (1838)
Unio clavusHanley (1843)
Margaron (Unio) clavusLea (1852), Lea (1870)
Unio (s.s.) clavusCall (1900)

Unio mytilloides Conrad, 1834
Conrad, 1834, Amer. J. Sci. 25(2): 343, pl. 1, fig. 7.
(error for Pleurobema mytiloides Rafinesque, 1820)

Unio mytilloidesConrad (1834)
Pleurobema (s.s.) corSimpson (1900), Simpson (1914)

Unio clavus Conrad, 1835
‘Lamarck’ Conrad, 1835, Monogr. Fam. Unionidae 1(1): 5, pl. 3, fig. 1.
(reference to Unio clava Lamarck, 1819)

Unio clavusConrad (1835), Hanley (1843)
Margarita (Unio) clavusLea (1838)
Unio clavaKüster (1848), Conrad (1853)
Margaron (Unio) clavusLea (1852), Lea (1870)
Pleurobema (s.s.) clavaSimpson (1900), Simpson (1914)
Unio (s.s.) clavusCall (1900)
Pleurobema (s.s.) mytiloidesFrierson (1927)
Pleurobema mytiloidesModell (1942)

Unio gibbosus Küster, 1848
‘Rafinesque’ Küster, 1848, Conchyl. Cab. 9(2): 40.
(error for Obliquaria (Scalenaria) scalenia Rafinesque, 1820)

Unio gibbosusKüster (1848)

Unio (Scalenaria) scalenarius Mörch, 1853
‘Raf.’ Mörch, 1853, Cat. Conch. Yoldi 2: 44.
(error for Obliquaria (Scalenaria) scalenia Rafinesque, 1820)

Unio (Scalenaria) scalenariusMörch (1853)

Unio consanguineus de Gregorio, 1914
de Gregorio, 1914, Il Nat. Siciliano 2: 46.
(homonym of Unio consanguineus Lea, 1861)

Unio consanguineusde Gregorio (1914)
Pleurobema clavaParmalee & Bogan (1998), Williams et al. (2008), Watters et al. (2009)

Unio anaticulus var. ohiensis de Gregorio, 1914
de Gregorio, 1914, Il Nat. Siciliano 2: 51, pl. 10, figs. 3a-d.
(intro'd as intra-sp. of Unio anaticulus Lea, 1861)

Unio anaticulus var. ohiensisde Gregorio (1914)
Pleurobema clavaParmalee & Bogan (1998), Watters et al. (2009)

Taxonomic History (since 1900)

Pleurobema (s.s.) clavaSimpson (1900), Simpson (1914)
Unio (s.s.) clavusCall (1900)
Pleurobema (s.s.) corSimpson (1900), Simpson (1914)
Pleurobema (s.s.) bournianusSimpson (1900)
Pleurobema clavaOrtmann (1909), Ortmann (1909), Ortmann (1909), Ortmann (1910), ..., Pieri et al. (2018), Inoue et al. (2019), Smith et al. (2020), Graf & Cummings (2021)
Unio patulusde Gregorio (1914)
Pleurobema (s.s.) bournianumSimpson (1914)
Unio consanguineusde Gregorio (1914)
Unio anaticulus var. ohiensisde Gregorio (1914)
Pleurobema obliqua rubrumOrtmann (1918)
Pleurobema (s.s.) mytiloidesFrierson (1927), Haas (1930)
Obovaria (s.s.) subrotundaFrierson (1927), Haas (1969)
Pleurobema (s.s.) clava-patulumFrierson (1927)
Quadrula (Pleuronaia) corFrierson (1927), Haas (1969)
Pleurobema mytiloidesModell (1942), Haas (1969)
Pleurobema clava clavaHaas (1969)
Pleurobema clava patulusHaas (1969)
Pleurobema bournianumTurgeon et al. (1988), Williams et al. (1993), Turgeon et al. (1998), IUCN (2000), Graf & Cummings (2007), Cummings & Graf (2010), ITIS (2012)

Page updated: 17:27:24 Sun 31 Oct 2021

 
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