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Page last updated
Mon 06 May 2024

The Freshwater Mussels (Unionoida) of the World (and other less consequential bivalves)

FM(U)otW(aolcb) is the web version of the MUSSEL Project Database. Follow the links to browse the data or use the custom Google search field. Either way, you win!

Cladomics pages are experimental — even more so than the rest of this site!

The purpose of the cladomics pages for each genus and family-group level taxon is to provide statistics and references indicating the extent to which published phylogenetic work supports the classification used here.

The Cladomics Summary provides an overview of the number of studies that have included terminal species representing this taxon as well as an indication of how frequently it has been recovered as monophyletic.

Following the summary, the available published clades representing the taxon are listed.

For more information, visit the Cladomics FARQ (frequently asked rhetorical questions) page. Abbreviations for the various statistics are explained on the FARQ!

Search the database pages:

Bivalvia | Unionoida | Unionoidea | Unionidae | Unioninae

tribe ACUTICOSTINI Starobogatov, 1967: cladomics
     1 genus

Cladomics Summary

number of species = 5
number of subtaxa (rank: genus) = 1

analyzed in 42 cladograms in 29 publications
maximum number of terminals in an analysis = 10, mean = 1.24
maximum number of subtaxa (genus) in an analysis = 1, mean = 1.0

Basic Monophyly

number of cladograms that test basic monophyly (i.e., more than 1 included species) = 2
maximum number of included terminals = 10, mean = 6.0
maximum number of subtaxa (genus) = 1, mean = 1.0
proportion of cladograms that support basic monophyly = 1.0

Huang et al., 2019 [Publ 3843], fig. S1

iT tT bi riS oT roT roS iD
10 10 5 1 72 52 4 0.0

*** Top Monophyly Cladogram *** 

                             +-- Sinanodonta_angula         
                 +-----------+                              
                 |           +-- Sinanodonta_lucida         
                 |                                          
                 |     +-------- Sinanodonta_jourdyi        
  +--------------+  +--+                                    
  |              |  |  |  +----- Sinanodonta_elliptica      
  |              |  |  +--+                                 
  |              |  |     |  +-- Sinanodonta_woodiana[5]    
  |              +--+     +--+                              
  |                 |        +-- Sinanodonta_lauta          
  |                 |                                       
  |                 |  +-------- Sinanodonta_woodiana[4]    
  |                 |  |                                    
  |                 +--+  +----- Sinanodonta_pacifica[3]    
  |                    |  |                                 
  |                    |  |  +-- Sinanodonta_woodiana[1]    
  |                    +--+  |                              
  |                       |  +-- Sinanodonta_pacifica[1]    
  |                       |  |                              
  |                       |  +-- Sinanodonta_pacifica[2]    
  |                       +--+                              
  |                          +-- Sinanodonta_woodiana[2]    
  |                          |                              
  |                          +-- Sinanodonta_woodiana[3]    
--+                          |                              
  |                          +-- Sinanodonta_woodiana[6]    
  |                                                         
  |              +-------------- Acuticosta_ovata[1] **        
  |              |                                          
  |              |  +----------- Acuticosta_chinensis[3] **    
  |  +-----------+  |                                       
  |  |           |  +----------- Acuticosta_chinensis[4] **    
  |  |           |  |                                       
  |  |           +--+----------- Acuticosta_chinensis[6] **    
  |  |              |                                       
  |  |              |        +-- Acuticosta_chinensis[1] **    
  |  |              |  +-----+                              
  |  |              |  |     +-- Acuticosta_ovata[2] **        
  |  |              +--+                                    
  |  |                 |  +----- Acuticosta_chinensis[5] **    
  |  |                 |  |                                 
  |  |                 +--+----- Acuticosta_chinensis[7] **    
  |  |                    |                                 
  |  |                    |  +-- Acuticosta_chinensis[2] **    
  +--+                    +--+                              
     |                       +-- Acuticosta_ovata[3] **        
     |                                                      
     |                       +-- Lepidodesma_languilati     
     |  +--------------------+                              
     |  |                    +-- Cristaria_plicata          
     |  |                                                   
     |  |           +----------- Lanceolaria_gladiola       
     |  |           |                                       
     |  |           |  +-------- Lanceolaria_grayii[5]      
     |  |  +--------+  |                                    
     |  |  |        |  |     +-- Lanceolaria_lanceolata     
     |  |  |        |  +-----+                              
     |  |  |        |  |     +-- Lanceolaria_triformis      
     +--+  |        +--+                                    
        |  |           |  +----- Lanceolaria_grayii[4]      
        |  |           |  |                                 
        |  |           |  |  +-- Lanceolaria_eucylindrica[1]
        |  |           |  |  |                              
        |  |           +--+  +-- Lanceolaria_eucylindrica[2]
        |  |              |  |                              
        |  |              |  +-- Lanceolaria_eucylindrica[3]
        |  |              |  |                              
        |  |              +--+-- Lanceolaria_eucylindrica[4]
        +--+                 |                              
           |                 +-- Lanceolaria_grayii[1]      
           |                 |                              
           |                 +-- Lanceolaria_grayii[2]      
           |                 |                              
           |                 +-- Lanceolaria_grayii[3]      
           |                                                
           |        +----------- Aculamprotula_tortuosa     
           |  +-----+                                       
           |  |     |  +-------- Aculamprotula_tientsinensis
           |  |     +--+                                    
           |  |        |  +----- Aculamprotula_fibrosa      
           |  |        +--+                                 
           |  |           |  +-- Aculamprotula_coreana      
           +--+           +--+                              
              |              +-- Aculamprotula_scripta      
              |                                             
              |           +----- Nodularia_douglasiae[1]    
              |  +--------+                                 
              |  |        |  +-- Nodularia_nuxpersicae      
              |  |        +--+                              
              +--+           +-- Nodularia_douglasiae[2]    
                 |                                          
                 |  +----------- Tchangsinaia_piscicula     
                 |  |                                       
                 +--+        +-- Schistodesmus_spinosus     
                    |  +-----+                              
                    |  |     +-- Schistodesmus_lampreyanus  
                    +--+                                    
                       |  +----- Cuneopsis_rufescens        
                       +--+                                 
                          |  +-- Cuneopsis_heudei           
                          +--+                              
                             +-- Cuneopsis_celtiformis

Lopes-Lima et al., 2020 [Publ 4080], fig. 2-4

iT tT bi riS oT roT roS iD
1 1 0 1 121 68 4 --

*** Top Sister-Group Cladogram ***

  +-------------------------------- Acuticosta_chinensis **       
  |                                                            
  +-------------------------------- Lepidodesma_languilati     
  |                                                            
  |                    +----------- Aculamprotula_tortuosa     
  |     +--------------+                                       
  |     |              |  +-------- Aculamprotula_scripta      
  |     |              +--+                                    
  |     |                 |  +----- Aculamprotula_coreana      
  |     |                 +--+                                 
  |     |                    |  +-- Aculamprotula_fibrosa      
  |     |                    +--+                              
  +-----+                       +-- Aculamprotula_tientsinensis
  |     |                                                      
  |     |  +----------------------- Middendorffinaia_mongolica 
  |     |  |                                                   
  |     |  |                 +----- Unio_gibbus                
  |     |  |                 |                                 
  |     |  |  +--------------+----- Unio_tumidus               
  |     +--+  |              |                                 
  |        |  |              |  +-- Unio_crassus               
  |        |  |              +--+                              
  |        |  |                 +-- Unio_pictorum              
  |        |  |                                                
  |        |  |     +-------------- Nodularia_breviconcha      
  |        +--+  +--+                                          
  |           |  |  |  +----------- Nodularia[2]               
  |           |  |  +--+                                       
  |           |  |     |  +-------- Nodularia_nuxpersicae      
  |           |  |     +--+                                    
  |           |  |        |  +----- Nodularia_nipponensis      
  |           |  |        +--+                                 
  |           |  |           |  +-- Nodularia_douglasiae       
  |           +--+           +--+                              
  |              |              +-- Nodularia[1]               
  |              |                                             
--+              |              +-- Schistodesmus_lampreyanus  
  |              |  +-----------+                              
  |              |  |           +-- Schistodesmus_spinosus     
  |              |  |                                          
  |              |  |        +----- Cuneopsis_rufescens        
  |              |  +--------+                                 
  |              +--+        |  +-- Cuneopsis_heudei           
  |                 |        +--+                              
  |                 |           +-- Cuneopsis_celtiformis      
  |                 |                                          
  |                 |  +----------- Tchangsinaia_piscicula     
  |                 +--+                                       
  |                    |  +-------- Inversiunio_jokohamensis   
  |                    +--+                                    
  |                       |  +----- Inversiunio_verrucosus     
  |                       +--+                                 
  |                          |  +-- Inversiunio_reinianus      
  |                          +--+                              
  |                             +-- Inversiunio_yanagawensis   
  |                                                            
  |              +----------------- Lanceolaria_oxyrhyncha     
  |              |                                             
  |  +-----------+  +-------------- Lanceolaria_gladiola       
  |  |           |  |                                          
  |  |           +--+           +-- Lanceolaria_bilirata       
  |  |              |  +--------+                              
  |  |              |  |        +-- Lanceolaria_grayii         
  |  |              +--+                                       
  |  |                 |  +-------- Lanceolaria_acrorrhyncha   
  |  |                 +--+                                    
  |  |                    |  +----- Lanceolaria_kihirai        
  |  |                    +--+                                 
  |  |                       |  +-- Lanceolaria_lanceolata     
  |  |                       +--+                              
  |  |                          +-- Lanceolaria_triformis      
  |  |                                                         
  |  |                    +-------- Pseudanodonta_complanata   
  +--+                    |                                    
     |                    |  +----- Anodonta_cygnea            
     |                    +--+                                 
     |                    |  |  +-- Anodonta_anatina           
     |  +-----------------+  +--+                              
     |  |                 |     +-- Anodonta_nuttalliana       
     |  |                 |                                    
     |  |                 |  +----- Pyganodon_grandis          
     |  |                 |  |                                 
     |  |                 |  +----- Simpsonaias_ambigua        
     |  |                 +--+                                 
     |  |                    +----- Strophitus_undulatus       
     |  |                    |                                 
     |  |                    |  +-- Alasmidonta_marginata      
     |  |                    +--+                              
     |  |                       +-- Lasmigona_costata          
     +--+                                                      
        |                       +-- Cristaria_plicata          
        |  +--------------------+                              
        |  |                    +-- Cristaria_clessini         
        |  |                                                   
        |  |        +-------------- Anemina_arcaeformis        
        |  |        |                                          
        |  +--------+  +----------- Amuranodonta_kijaensis     
        |  |        |  |                                       
        |  |        +--+  +-------- Buldowskia_suifunica       
        |  |           |  |                                    
        |  |           +--+     +-- Buldowskia_flavotincta     
        +--+              |  +--+                              
           |              |  |  +-- Buldowskia_shadini         
           |              +--+                                 
           |                 |  +-- Buldowskia_iwakawai        
           |                 +--+                              
           |                    +-- Buldowskia_kamiyai         
           |                                                   
           |  +-------------------- Pletholophus_tenuis        
           |  |                                                
           |  |           +-------- Beringiana_japonica        
           |  |  +--------+                                    
           +--+  |        |  +----- Beringiana_beringiana      
              |  |        +--+                                 
              |  |           |  +-- Beringiana_fukuharai       
              |  |           +--+                              
              +--+              +-- Beringiana_gosannensis     
                 |                                             
                 |              +-- Sinanodonta_angula         
                 |  +-----------+                              
                 |  |           +-- Sinanodonta_lucida         
                 |  |                                          
                 +--+  +----------- Sinanodonta_schrenkii      
                    |  |                                       
                    |  |        +-- Sinanodonta_calipygos      
                    |  +--------+                              
                    +--+        +-- Sinanodonta_tumens         
                       |                                       
                       |  +-------- Sinanodonta_woodiana[2]    
                       |  |                                    
                       +--+  +----- Sinanodonta_elliptica      
                          |  |                                 
                          +--+----- Sinanodonta_jourdyi        
                             |                                 
                             |  +-- Sinanodonta_lauta          
                             +--+                              
                                +-- Sinanodonta_woodiana[1]

Zhou et al., 2007 [Publ 2658], fig. 1

iT tT bi riS oT roT roS iD
2 2 0 1 21 13 3 -- 
        +-- Schistodesmus_lampreyanus
  +-----+                            
  |     +-- Schistodesmus_spinosus   
  |                                  
--+     +-- Lanceolaria_triformis    
  |  +--+                            
  |  |  +-- Lanceolaria_lanceolata   
  +--+                               
     |  +-- Acuticosta_ovata **         
     +--+                            
        +-- Acuticosta_chinensis **

Huang et al., 2002 [Publ 2657], fig. 1

iT tT bi riS oT roT roS iD
1 1 0 1 33 13 4 -- 
     +----- Acuticosta_ovata **         
  +--+                               
  |  |  +-- Nodularia_douglasiae     
  |  +--+                            
  |     +-- Tchangsinaia_piscicula   
--+                                  
  |     +-- Lanceolaria_grayii       
  |  +--+                            
  |  |  +-- Lanceolaria_lanceolata   
  +--+                               
     |  +-- Schistodesmus_lampreyanus
     +--+                            
        +-- Lepidodesma_languilati

Huang et al., 2002 [Publ 2657], fig. 2

iT tT bi riS oT roT roS iD
1 1 0 1 33 13 4 -- 
  +-------- Strophitus_radiatus      
  |                                  
  +-------- Schistodesmus_lampreyanus
  |                                  
  +-------- Lepidodesma_languilati   
  |                                  
  +-------- Acuticosta_ovata **         
  |                                  
  |     +-- Sinanodonta_woodiana     
  +-----+                            
  |     +-- Cristaria_plicata        
--+                                  
  |     +-- Lanceolaria_grayii       
  +-----+                            
  |     +-- Lanceolaria_lanceolata   
  |                                  
  |     +-- Nodularia_douglasiae     
  +-----+                            
  |     +-- Tchangsinaia_piscicula   
  |                                  
  |  +----- Utterbackia_imbecillis   
  |  |                               
  +--+----- Utterbackiana_couperiana 
     |                               
     |  +-- Alasmidonta_triangulata  
     +--+                            
        +-- Lasmigona_costata

Zhou et al., 2007 [Publ 2658], fig. 2

iT tT bi riS oT roT roS iD
1 1 0 1 20 9 2 -- 
           +----- Acuticosta_chinensis **       
  +--------+                                 
  |        |  +-- Nodularia_douglasiae       
  |        +--+                              
--+           +-- Cuneopsis_celtiformis      
  |                                          
  |  +----------- Aculamprotula_tortuosa     
  +--+                                       
     |  +-------- Aculamprotula_scripta      
     +--+                                    
        |  +----- Aculamprotula_fibrosa      
        +--+                                 
           |  +-- Aculamprotula_zonata       
           +--+                              
              +-- Aculamprotula_tientsinensis

Zhang et al., 2004 [Publ 2735], fig. 1

iT tT bi riS oT roT roS iD
1 1 0 1 13 11 3 -- 
                 +----- Acuticosta_chinensis **     
  +--------------+                               
  |              |  +-- Lanceolaria_grayii       
  |              +--+                            
--+                 +-- Lanceolaria_lanceolata   
  |                                              
  |  +----------------- Tchangsinaia_piscicula   
  +--+                                           
     |  +-------------- Cuneopsis_heudei         
     +--+                                        
        |  +----------- Nodularia_douglasiae     
        +--+                                     
           |  +-------- Pseudocuneopsis_capitata 
           +--+                                  
              |  +----- Cuneopsis_rufescens      
              +--+                               
                 |  +-- Schistodesmus_lampreyanus
                 +--+                            
                    +-- Schistodesmus_spinosus

Ouyang et al., 2011 [Publ 3408], fig. 1

iT tT bi riS oT roT roS iD
1 1 0 1 12 8 3 -- 
        +-- Nodularia_douglasiae  
  +-----+                         
  |     +-- Cuneopsis_heudei      
--+                               
  |  +----- Lanceolaria_grayii    
  |  |                            
  +--+----- Acuticosta_chinensis **  
     |                            
     |  +-- Lanceolaria_gladiola  
     +--+                         
        +-- Aculamprotula_tortuosa

Pfeiffer & Graf, 2013 [Publ 3531], fig. 2

iT tT bi riS oT roT roS iD
1 1 0 1 34 12 2 -- 
                 +-- Cristaria_plicata          
           +-----+                              
           |     +-- Sinanodonta_woodiana       
  +--------+                                    
  |        |  +----- Strophitus_undulatus       
  |        +--+                                 
  |           |  +-- Platynaias_compressa       
--+           +--+                              
  |              +-- Pyganodon_grandis          
  |                                             
  |  +-------------- Acuticosta_chinensis **       
  |  |                                          
  +--+           +-- Unio_pictorum              
     |  +--------+                              
     |  |        +-- Nodularia_douglasiae       
     +--+                                       
        |  +-------- Aculamprotula_tortuosa     
        |  |                                    
        +--+  +----- Aculamprotula_scripta      
           |  |                                 
           +--+  +-- Aculamprotula_tientsinensis
              |  |                              
              +--+-- Aculamprotula_fibrosa      
                 |                              
                 +-- Aculamprotula_zonata

Sano et al., 2017 [Publ 3799], fig. 2

iT tT bi riS oT roT roS iD
1 1 0 1 35 23 4 -- 
              +-- Schistodesmus_lampreyanus  
  +-----------+                              
  |           +-- Acuticosta_ovata **           
  |                                          
  |     +-------- Lepidodesma_languilati     
--+  +--+                                    
  |  |  |  +----- Lanceolaria_grayii[1]      
  |  |  +--+                                 
  |  |     |  +-- Lanceolaria_lanceolata     
  +--+     +--+                              
     |        +-- Lanceolaria_grayii[2]      
     |                                       
     |  +-------- Inversiunio_jokohamensis   
     |  |                                    
     +--+  +----- Inversiunio_yanagawensis[3]
        |  |                                 
        +--+  +-- Inversiunio_yanagawensis[1]
           |  |                              
           +--+-- Inversiunio_yanagawensis[2]
              |                              
              +-- Inversiunio_reinianus

Wu et al., 2018 [Publ 3836], fig. 2

iT tT bi riS oT roT roS iD
1 1 0 1 52 32 4 -- 
           +----------- Acuticosta_chinensis **  
  +--------+                                  
  |        |  +-------- Lanceolaria_gladiola  
  |        +--+                               
  |           |  +----- Lanceolaria_grayii    
  |           +--+                            
  |              |  +-- Lanceolaria_lanceolata
  |              +--+                         
  |                 +-- Lanceolaria_triformis 
--+                                           
  |           +-------- Anemina_arcaeformis   
  |           |                               
  |           +-------- Cristaria_plicata     
  |  +--------+                               
  |  |        |     +-- Sinanodonta_woodiana  
  |  |        |  +--+                         
  |  |        |  |  +-- Sinanodonta_elliptica 
  |  |        +--+                            
  +--+           |  +-- Sinanodonta_lucida    
     |           +--+                         
     |              +-- Sinanodonta_angula    
     |                                        
     |  +-------------- Anodonta_cygnea       
     |  |                                     
     +--+  +----------- Anodonta_anatina      
        |  |                                  
        +--+  +-------- Strophitus_undulatus  
           |  |                               
           +--+  +----- Platynaias_compressa  
              |  |                            
              +--+----- Alasmidonta_marginata 
                 |                            
                 |  +-- Pyganodon_cataracta   
                 +--+                         
                    +-- Pyganodon_grandis

Huang et al., 2019 [Publ 3843], fig. 1

iT tT bi riS oT roT roS iD
1 1 0 1 43 27 4 -- 
  +-------------------------------- Anemina_arcaeformis        
  |                                                            
  |  +----------------------------- Acuticosta_chinensis **       
  |  |                                                         
--+  |                          +-- Sinanodonta_angula         
  |  |                 +--------+                              
  |  |                 |        +-- Sinanodonta_lucida         
  |  |  +--------------+                                       
  |  |  |              |  +-------- Sinanodonta_jourdyi        
  +--+  |              +--+                                    
     |  |                 |  +----- Sinanodonta_woodiana[2]    
     |  |                 +--+                                 
     |  |                    |  +-- Sinanodonta_elliptica      
     |  |                    +--+                              
     |  |                       +-- Sinanodonta_woodiana[1]    
     |  |                                                      
     +--+              +----------- Aculamprotula_tortuosa     
        |  +-----------+                                       
        |  |           |  +-------- Aculamprotula_scripta      
        |  |           +--+                                    
        |  |              |  +----- Aculamprotula_coreana      
        |  |              +--+                                 
        |  |                 |  +-- Aculamprotula_fibrosa      
        |  |                 +--+                              
        |  |                    +-- Aculamprotula_tientsinensis
        +--+                                                   
           |           +----------- Cristaria_plicata          
           |  +--------+                                       
           |  |        |  +-------- Lanceolaria_gladiola       
           |  |        +--+                                    
           |  |           |  +----- Lanceolaria_grayii         
           |  |           +--+                                 
           |  |              |  +-- Lanceolaria_lanceolata     
           +--+              +--+                              
              |                 +-- Lanceolaria_triformis      
              |                                                
              |  +----------------- Lepidodesma_languilati     
              |  |                                             
              |  |              +-- Schistodesmus_lampreyanus  
              +--+  +-----------+                              
                 |  |           +-- Schistodesmus_spinosus     
                 |  |                                          
                 +--+        +----- Nodularia_douglasiae[1]    
                    |  +-----+                                 
                    |  |     |  +-- Nodularia_douglasiae[2]    
                    |  |     +--+                              
                    +--+        +-- Nodularia_nuxpersicae      
                       |                                       
                       |  +-------- Cuneopsis_rufescens        
                       +--+                                    
                          |  +----- Tchangsinaia_piscicula     
                          +--+                                 
                             |  +-- Cuneopsis_heudei           
                             +--+                              
                                +-- Cuneopsis_celtiformis

Huang et al., 2019 [Publ 3843], fig. 2

iT tT bi riS oT roT roS iD
1 1 0 1 203 44 4 -- 
  +-------------------------------- Acuticosta_chinensis **       
  |                                                            
  |  +----------------------------- Lepidodesma_languilati     
  |  |                                                         
  |  |                    +-------- Aculamprotula_tortuosa     
  |  |                    |                                    
  |  |        +-----------+  +----- Aculamprotula_coreana      
  |  |        |           |  |                                 
  |  |        |           +--+----- Aculamprotula_scripta      
  |  |        |              |                                 
  |  |        |              |  +-- Aculamprotula_fibrosa      
  |  |        |              +--+                              
  |  |        |                 +-- Aculamprotula_tientsinensis
--+  +--------+                                                
  |  |        |           +-------- Unio_tumidus               
  |  |        |  +--------+                                    
  |  |        |  |        |  +----- Unio_crassus               
  |  |        |  |        +--+                                 
  |  |        |  |           |  +-- Unio_delphinus             
  |  |        |  |           +--+                              
  |  |        +--+              +-- Unio_pictorum              
  |  |           |                                             
  |  |           |              +-- Schistodesmus_lampreyanus  
  |  |           |  +-----------+                              
  |  |           |  |           +-- Schistodesmus_spinosus     
  |  |           |  |                                          
  |  |           +--+        +----- Nodularia_douglasiae[1]    
  +--+              |  +-----+                                 
     |              |  |     |  +-- Nodularia_douglasiae[2]    
     |              |  |     +--+                              
     |              +--+        +-- Nodularia_nuxpersicae      
     |                 |                                       
     |                 |  +-------- Tchangsinaia_piscicula     
     |                 +--+                                    
     |                    |  +----- Cuneopsis_rufescens        
     |                    +--+                                 
     |                       |  +-- Cuneopsis_heudei           
     |                       +--+                              
     |                          +-- Cuneopsis_celtiformis      
     |                                                         
     |                    +-------- Lanceolaria_gladiola       
     |                    |                                    
     |  +-----------------+  +----- Lanceolaria_grayii[1]      
     |  |                 |  |                                 
     |  |                 +--+----- Lanceolaria_grayii[2]      
     |  |                    |                                 
     |  |                    |  +-- Lanceolaria_lanceolata     
     |  |                    +--+                              
     |  |                       +-- Lanceolaria_triformis      
     |  |                                                      
     |  |     +-------------------- Anemina_arcaeformis        
     |  |     |                                                
     |  |  +--+  +----------------- Cristaria_plicata          
     +--+  |  |  |                                             
        |  |  +--+  +-------------- Pletholophus_tenuis        
        |  |     |  |                                          
        |  |     +--+           +-- Sinanodonta_angula         
        |  |        |  +--------+                              
        |  |        |  |        +-- Sinanodonta_lucida         
        |  |        +--+                                       
        |  |           |  +-------- Sinanodonta_woodiana[2]    
        |  |           +--+                                    
        |  |              |  +----- Sinanodonta_jourdyi        
        |  |              +--+                                 
        +--+                 |  +-- Sinanodonta_elliptica      
           |                 +--+                              
           |                    +-- Sinanodonta_woodiana[1]    
           |                                                   
           |              +-------- Alasmidonta_marginata      
           |              |                                    
           |              +-------- Lasmigona_costata          
           |              |                                    
           |              +-------- Pseudanodonta_complanata   
           |              |                                    
           |              +-------- Simpsonaias_ambigua        
           |              |                                    
           +--------------+-------- Strophitus_undulatus       
                          |                                    
                          |  +----- Anodonta_anatina           
                          +--+                                 
                          |  |  +-- Anodonta_cygnea            
                          |  +--+                              
                          |     +-- Anodonta_nuttalliana       
                          |                                    
                          |  +----- Pyganodon_grandis          
                          +--+                                 
                             |  +-- Utterbackia_imbecillis     
                             +--+                              
                                +-- Utterbackia_peninsularis

Huang et al., 2019 [Publ 3843], fig. 3

iT tT bi riS oT roT roS iD
1 1 0 1 56 26 4 -- 
                    +----- Lanceolaria_gladiola    
  +-----------------+                              
  |                 |  +-- Lanceolaria_lanceolata  
  |                 +--+                           
  |                    +-- Lanceolaria_grayii      
--+                                                
  |  +-------------------- Acuticosta_chinensis **    
  |  |                                             
  |  |           +-------- Cristaria_plicata       
  |  |           |                                 
  +--+  +--------+     +-- Sinanodonta_lucida      
     |  |        |  +--+                           
     |  |        |  |  +-- Sinanodonta_woodiana    
     |  |        +--+                              
     +--+           |  +-- Anemina_arcaeformis     
        |           +--+                           
        |              +-- Anemina_euscaphys       
        |                                          
        |  +-------------- Anodonta_cygnea         
        +--+                                       
           |  +----------- Anodonta_anatina        
           +--+                                    
              |  +-------- Lasmigona_costata       
              +--+                                 
                 |  +----- Pyganodon_grandis       
                 +--+                              
                    |  +-- Utterbackia_imbecillis  
                    +--+                           
                       +-- Utterbackia_peninsularis

Huang et al., 2019 [Publ 3843], fig. S3

iT tT bi riS oT roT roS iD
1 1 0 1 200 42 4 -- 
  +----------------------------------- Acuticosta_chinensis **       
  |                                                               
  |                          +-------- Aculamprotula_tortuosa     
  |                          |                                    
  |              +-----------+     +-- Aculamprotula_coreana      
  |              |           |  +--+                              
  |              |           |  |  +-- Aculamprotula_scripta      
  |              |           +--+                                 
  |              |              |  +-- Aculamprotula_fibrosa      
  |              |              +--+                              
  |              |                 +-- Aculamprotula_tientsinensis
  |  +-----------+                                                
  |  |           |           +-------- Unio_tumidus               
--+  |           |  +--------+                                    
  |  |           |  |        |  +----- Unio_crassus               
  |  |           |  |        +--+                                 
  |  |           |  |           |  +-- Unio_delphinus             
  |  |           |  |           +--+                              
  |  |           +--+              +-- Unio_pictorum              
  |  |              |                                             
  |  |              |           +----- Nodularia_douglasiae[1]    
  |  |              |  +--------+                                 
  |  |              |  |        |  +-- Nodularia_douglasiae[2]    
  |  |              |  |        +--+                              
  |  |              +--+           +-- Nodularia_nuxpersicae      
  |  |                 |                                          
  +--+                 |  +----------- Schistodesmus_lampreyanus  
     |                 +--+                                       
     |                    |  +-------- Tchangsinaia_piscicula     
     |                    +--+                                    
     |                       |  +----- Cuneopsis_rufescens        
     |                       +--+                                 
     |                          |  +-- Cuneopsis_heudei           
     |                          +--+                              
     |                             +-- Cuneopsis_celtiformis      
     |                                                            
     |  +----------------------------- Lepidodesma_languilati     
     |  |                                                         
     |  |                    +-------- Lanceolaria_gladiola       
     |  |  +-----------------+                                    
     |  |  |                 |  +----- Lanceolaria_grayii         
     |  |  |                 +--+                                 
     +--+  |                    |  +-- Lanceolaria_lanceolata     
        |  |                    +--+                              
        |  |                       +-- Lanceolaria_triformis      
        |  |                                                      
        |  |     +-------------------- Anemina_arcaeformis        
        |  |     |                                                
        |  |  +--+  +----------------- Cristaria_plicata          
        +--+  |  |  |                                             
           |  |  +--+  +-------------- Pletholophus_tenuis        
           |  |     |  |                                          
           |  |     +--+           +-- Sinanodonta_angula         
           |  |        |  +--------+                              
           |  |        |  |        +-- Sinanodonta_lucida         
           |  |        +--+                                       
           |  |           |  +-------- Sinanodonta_woodiana[2]    
           |  |           +--+                                    
           |  |              |  +----- Sinanodonta_jourdyi        
           +--+              +--+                                 
              |                 |  +-- Sinanodonta_elliptica      
              |                 +--+                              
              |                    +-- Sinanodonta_woodiana[1]    
              |                                                   
              |                    +-- Alasmidonta_marginata      
              |           +--------+                              
              |           |        +-- Lasmigona_costata          
              |           |                                       
              |           |     +----- Anodonta_anatina           
              |           +-----+                                 
              |           |     |  +-- Anodonta_cygnea            
              +-----------+     +--+                              
                          |        +-- Anodonta_nuttalliana       
                          |                                       
                          |     +----- Pseudanodonta_complanata   
                          |  +--+                                 
                          |  |  |  +-- Simpsonaias_ambigua        
                          |  |  +--+                              
                          +--+     +-- Strophitus_undulatus       
                             |                                    
                             |  +----- Pyganodon_grandis          
                             +--+                                 
                                |  +-- Utterbackia_imbecillis     
                                +--+                              
                                   +-- Utterbackia_peninsularis

Huang et al., 2019 [Publ 3843], fig. S4

iT tT bi riS oT roT roS iD
1 1 0 1 201 43 4 -- 
  +----------------- Acuticosta_chinensis **       
  |                                             
  +----------------- Alasmidonta_marginata      
  |                                             
  +----------------- Anodonta_anatina           
  |                                             
  +----------------- Anemina_arcaeformis        
  |                                             
  +----------------- Anodonta_cygnea            
  |                                             
  +----------------- Cristaria_plicata          
  |                                             
  +----------------- Anodonta_nuttalliana       
  |                                             
  +----------------- Lasmigona_costata          
  |                                             
  +----------------- Lepidodesma_languilati     
  |                                             
  +----------------- Pseudanodonta_complanata   
  |                                             
  +----------------- Pyganodon_grandis          
  |                                             
  +----------------- Simpsonaias_ambigua        
  |                                             
  +----------------- Strophitus_undulatus       
  |                                             
  |              +-- Schistodesmus_lampreyanus  
  +--------------+                              
  |              +-- Schistodesmus_spinosus     
  |                                             
  |              +-- Utterbackia_imbecillis     
  +--------------+                              
  |              +-- Utterbackia_peninsularis   
  |                                             
  |        +-------- Lanceolaria_gladiola       
  +--------+                                    
  |        |  +----- Lanceolaria_grayii         
  |        +--+                                 
--+           |  +-- Lanceolaria_lanceolata     
  |           +--+                              
  |              +-- Lanceolaria_triformis      
  |                                             
  |        +-------- Unio_tumidus               
  +--------+                                    
  |        |  +----- Unio_crassus               
  |        +--+                                 
  |           |  +-- Unio_delphinus             
  |           +--+                              
  |              +-- Unio_pictorum              
  |                                             
  |        +-------- Aculamprotula_tortuosa     
  |        |                                    
  +--------+     +-- Aculamprotula_coreana      
  |        |  +--+                              
  |        |  |  +-- Aculamprotula_scripta      
  |        +--+                                 
  |           |  +-- Aculamprotula_fibrosa      
  |           +--+                              
  |              +-- Aculamprotula_tientsinensis
  |                                             
  |        +-------- Cuneopsis_heudei           
  |        |                                    
  |        +-------- Cuneopsis_celtiformis      
  |        |                                    
  +--------+-------- Tchangsinaia_piscicula     
  |        |                                    
  |        +-------- Cuneopsis_rufescens        
  |        |                                    
  |        |  +----- Nodularia_douglasiae[2]    
  |        +--+                                 
  |           |  +-- Nodularia_douglasiae[1]    
  |           +--+                              
  |              +-- Nodularia_nuxpersicae      
  |                                             
  |  +-------------- Pletholophus_tenuis        
  |  |                                          
  +--+           +-- Sinanodonta_angula         
     |  +--------+                              
     |  |        +-- Sinanodonta_lucida         
     +--+                                       
        |  +-------- Sinanodonta_woodiana[1]    
        +--+                                    
           |  +----- Sinanodonta_jourdyi        
           +--+                                 
              |  +-- Sinanodonta_elliptica      
              +--+                              
                 +-- Sinanodonta_woodiana[2]

Aunins et al., 2018 [Publ 3863], fig. 1

iT tT bi riS oT roT roS iD
1 1 0 1 40 26 4 -- 
              +----- Acuticosta_chinensis **    
  +-----------+                              
  |           |  +-- Lanceolaria_grayii      
  |           +--+                           
  |              +-- Lanceolaria_lanceolata  
  |                                          
  |        +-------- Cristaria_plicata       
--+        |                                 
  |  +-----+     +-- Anemina_arcaeformis     
  |  |     |  +--+                           
  |  |     |  |  +-- Anemina_euscaphys       
  |  |     +--+                              
  |  |        |  +-- Sinanodonta_woodiana    
  |  |        +--+                           
  +--+           +-- Sinanodonta_lucida      
     |                                       
     |           +-- Anodonta_anatina        
     |  +--------+                           
     |  |        +-- Anodonta_cygnea         
     |  |                                    
     +--+        +-- Prolasmidonta_heterodon 
        |  +-----+                           
        |  |     +-- Alasmidonta_varicosa    
        +--+                                 
           |  +----- Pyganodon_grandis       
           +--+                              
              |  +-- Utterbackia_imbecillis  
              +--+                           
                 +-- Utterbackia_peninsularis

Wu et al., 2019 [Publ 3864], fig. 3

iT tT bi riS oT roT roS iD
1 1 0 1 46 26 4 -- 
              +----- Acuticosta_chinensis **    
  +-----------+                              
  |           |  +-- Lanceolaria_grayii      
  |           +--+                           
  |              +-- Lanceolaria_lanceolata  
  |                                          
--+     +----------- Anodonta_anatina        
  |  +--+                                    
  |  |  |  +-------- Platynaias_compressa    
  |  |  +--+                                 
  |  |     |  +----- Pyganodon_grandis       
  |  |     +--+                              
  +--+        |  +-- Utterbackia_peninsularis
     |        +--+                           
     |           +-- Utterbackia_imbecillis  
     |                                       
     |     +-------- Cristaria_plicata       
     |     |                                 
     +-----+     +-- Anemina_arcaeformis     
           |  +--+                           
           |  |  +-- Anemina_euscaphys       
           +--+                              
              |  +-- Sinanodonta_lucida      
              +--+                           
                 +-- Sinanodonta_woodiana

Wang et al., 2018 [Publ 3865], fig. 2

iT tT bi riS oT roT roS iD
1 1 0 1 31 17 3 -- 
           +----- Acuticosta_chinensis **  
  +--------+                            
  |        |  +-- Lanceolaria_grayii    
  |        +--+                         
  |           +-- Lanceolaria_lanceolata
--+                                     
  |        +----- Anodonta_anatina      
  |  +-----+                            
  |  |     |  +-- Pyganodon_grandis     
  |  |     +--+                         
  +--+        +-- Platynaias_compressa  
     |                                  
     |  +-------- Cristaria_plicata     
     |  |                               
     +--+     +-- Anemina_euscaphys     
        |  +--+                         
        |  |  +-- Anemina_arcaeformis   
        +--+                            
           |  +-- Sinanodonta_woodiana  
           +--+                         
              +-- Sinanodonta_lucida

Liu et al., 2017 [Publ 3939], fig. 2

iT tT bi riS oT roT roS iD
1 1 0 1 16 15 3 -- 
-- Acuticosta_chinensis

Fukata & Iigo, 2020 [Publ 4106], fig. 1

iT tT bi riS oT roT roS iD
1 1 0 1 53 27 4 -- 
     +-------------- Acuticosta_chinensis **       
     |                                          
     |     +-------- Cristaria_plicata          
     |     |                                    
     |  +--+     +-- Sinanodonta_woodiana       
  +--+  |  |  +--+                              
  |  |  |  |  |  +-- Sinanodonta_lucida         
  |  |  |  +--+                                 
  |  |  |     |  +-- Anemina_arcaeformis        
  |  |  |     +--+                              
  |  +--+        +-- Anemina_euscaphys          
  |     |                                       
  |     |        +-- Anodonta_anatina           
  |     |  +-----+                              
  |     |  |     +-- Anodonta_cygnea            
  |     |  |                                    
  |     +--+     +-- Pyganodon_grandis          
--+        |  +--+                              
  |        |  |  +-- Utterbackia_peninsularis   
  |        +--+                                 
  |           |  +-- Alasmidonta_varicosa       
  |           +--+                              
  |              +-- Prolasmidonta_heterodon    
  |                                             
  |           +----- Aculamprotula_tortuosa     
  |  +--------+                                 
  |  |        |  +-- Aculamprotula_tientsinensis
  |  |        +--+                              
  |  |           +-- Aculamprotula_coreana      
  |  |                                          
  +--+     +-------- Unio_tumidus               
     |  +--+                                    
     |  |  |  +----- Unio_crassus               
     |  |  +--+                                 
     |  |     |  +-- Unio_pictorum              
     |  |     +--+                              
     +--+        +-- Unio_delphinus             
        |                                       
        |        +-- Schistodesmus_lampreyanus  
        |  +-----+                              
        |  |     +-- Cuneopsis_heudei           
        +--+                                    
           |  +----- Pseudocuneopsis_capitata   
           +--+                                 
              |  +-- Nodularia_douglasiae       
              +--+                              
                 +-- Tchangsinaia_piscicula

Fukata & Iigo, 2019 [Publ 4108], fig. 1

iT tT bi riS oT roT roS iD
1 1 0 1 13 13 2 -- 
  +----------- Aculamprotula_tientsinensis
--+                                       
  |  +-------- Acuticosta_chinensis **       
  +--+                                    
     |  +----- Lanceolaria_gladiola       
     +--+                                 
        |  +-- Lanceolaria_grayii         
        +--+                              
           +-- Lanceolaria_lanceolata

Wu et al., 2020 [Publ 4121], fig. 3

iT tT bi riS oT roT roS iD
1 1 0 1 49 27 4 -- 
              +----- Acuticosta_chinensis **    
  +-----------+                              
  |           |  +-- Lanceolaria_lanceolata  
  |           +--+                           
  |              +-- Lanceolaria_grayii      
  |                                          
--+     +----------- Anodonta_anatina        
  |  +--+                                    
  |  |  |  +-------- Platynaias_compressa    
  |  |  +--+                                 
  |  |     |  +----- Pyganodon_grandis       
  |  |     +--+                              
  +--+        |  +-- Utterbackia_peninsularis
     |        +--+                           
     |           +-- Utterbackia_imbecillis  
     |                                       
     |     +-------- Cristaria_plicata       
     |     |                                 
     +-----+     +-- Sinanodonta_woodiana    
           |  +--+                           
           |  |  +-- Sinanodonta_lauta       
           +--+                              
              |  +-- Anemina_arcaeformis     
              +--+                           
                 +-- Anemina_euscaphys

Sano et al., 2020 [Publ 4138], fig. 1

iT tT bi riS oT roT roS iD
1 1 0 1 47 33 4 -- 
  +-------- Lanceolaria_grayii[1]      
  |                                    
  +-------- Lanceolaria_grayii[2]      
  |                                    
  +-------- Cristaria_plicata[1]       
  |                                    
  +-------- Cristaria_plicata[2]       
  |                                    
  +-------- Sinanodonta_lauta          
  |                                    
  +-------- Beringiana_japonica        
  |                                    
  +-------- Beringiana_fukuharai[1]    
  |                                    
  +-------- Beringiana_beringiana      
  |                                    
  +-------- Beringiana_fukuharai[2]    
  |                                    
  +-------- Beringiana_fukuharai[3]    
  |                                    
  +-------- Buldowskia_iwakawai[1]     
  |                                    
  +-------- Buldowskia_iwakawai[2]     
  |                                    
  +-------- Sinanodonta_tumens[1]      
  |                                    
  +-------- Sinanodonta_tumens[2]      
  |                                    
--+-------- Pletholophus_tenuis        
  |                                    
  +-------- Anemina_arcaeformis        
  |                                    
  +-------- Lanceolaria_lanceolata     
  |                                    
  +-------- Cristaria_plicata[3]       
  |                                    
  +-------- Lanceolaria_gladiola       
  |                                    
  +-------- Sinanodonta_angula         
  |                                    
  +-------- Lepidodesma_languilati     
  |                                    
  |     +-- Aculamprotula_tortuosa     
  +-----+                              
  |     +-- Acuticosta_chinensis **       
  |                                    
  |  +----- Cuneopsis_celtiformis      
  |  |                                 
  +--+  +-- Nodularia_douglasiae[1]    
  |  |  |                              
  |  +--+-- Nodularia_douglasiae[2]    
  |     |                              
  |     +-- Nodularia_nipponensis      
  |                                    
  |  +----- Schistodesmus_lampreyanus  
  |  |                                 
  |  |  +-- Inversiunio_reinianus[1]   
  +--+  |                              
     |  +-- Inversiunio_reinianus[2]   
     |  |                              
     |  +-- Inversiunio_jokohamensis[1]
     +--+                              
        +-- Inversiunio_jokohamensis[2]
        |                              
        +-- Inversiunio_jokohamensis[3]
        |                              
        +-- Inversiunio_jokohamensis[4]

Kim et al., 2021 [Publ 4197], fig. 1

iT tT bi riS oT roT roS iD
1 1 0 1 14 13 3 -- 
        +-------- Acuticosta_chinensis **     
  +-----+                                  
  |     |  +----- Lanceolaria_gladiola     
  |     +--+                               
  |        |  +-- Lanceolaria_grayii       
  |        +--+                            
  |           +-- Lanceolaria_lanceolata   
--+                                        
  |     +-------- Unio_tumidus             
  |  +--+                                  
  |  |  |  +----- Unio_crassus             
  |  |  +--+                               
  |  |     |  +-- Unio_delphinus           
  |  |     +--+                            
  +--+        +-- Unio_pictorum            
     |                                     
     |        +-- Nodularia_douglasiae     
     |     +--+                            
     |     |  +-- Nodularia_breviconcha    
     +-----+                               
           |  +-- Schistodesmus_lampreyanus
           +--+                            
              +-- Cuneopsis_heudei

Wu et al., 2021 [Publ 4215], fig. 3

iT tT bi riS oT roT roS iD
1 1 0 1 23 3 1 -- 
  +-------- Acuticosta_chinensis **    
--+                                 
  |  +----- Nodularia_douglasiae    
  +--+                              
     |  +-- Inversiunio_jokohamensis
     +--+                           
        +-- Inversiunio_reinianus

Wu et al., 2022 [Publ 4238], fig. 2d

iT tT bi riS oT roT roS iD
1 1 0 1 38 38 4 -- 
                 +----- Lanceolaria_gladiola    
  +--------------+                              
  |              |  +-- Lanceolaria_grayii      
  |              +--+                           
  |                 +-- Lanceolaria_lanceolata  
--+                                             
  |  +----------------- Acuticosta_chinensis **    
  |  |                                          
  |  |        +-------- Cristaria_plicata       
  |  |        |                                 
  +--+  +-----+     +-- Sinanodonta_woodiana    
     |  |     |  +--+                           
     |  |     |  |  +-- Sinanodonta_lucida      
     |  |     +--+                              
     |  |        |  +-- Anemina_arcaeformis     
     +--+        +--+                           
        |           +-- Anemina_euscaphys       
        |                                       
        |           +-- Anodonta_cygnea         
        |  +--------+                           
        |  |        +-- Anodonta_anatina        
        +--+                                    
           |  +-------- Platynaias_compressa    
           +--+                                 
              |  +----- Pyganodon_grandis       
              +--+                              
                 |  +-- Utterbackia_imbecillis  
                 +--+                           
                    +-- Utterbackia_peninsularis

Wu et al., 2022 [Publ 4238], fig. 3

iT tT bi riS oT roT roS iD
1 1 0 1 73 35 4 -- 
           +-------- Acuticosta_chinensis **    
  +--------+                                 
  |        |  +----- Lanceolaria_gladiola    
  |        +--+                              
  |           |  +-- Lanceolaria_grayii      
  |           +--+                           
  |              +-- Lanceolaria_lanceolata  
  |                                          
--+        +-------- Cristaria_plicata       
  |        |                                 
  |  +-----+     +-- Sinanodonta_woodiana    
  |  |     |  +--+                           
  |  |     |  |  +-- Sinanodonta_lucida      
  |  |     +--+                              
  |  |        |  +-- Anemina_arcaeformis     
  +--+        +--+                           
     |           +-- Anemina_euscaphys       
     |                                       
     |           +-- Anodonta_cygnea         
     |  +--------+                           
     |  |        +-- Anodonta_anatina        
     +--+                                    
        |  +-------- Platynaias_compressa    
        +--+                                 
           |  +----- Pyganodon_grandis       
           +--+                              
              |  +-- Utterbackia_imbecillis  
              +--+                           
                 +-- Utterbackia_peninsularis

Pavan-Kumar et al., 2022 [Publ 4263], fig. 4

iT tT bi riS oT roT roS iD
1 1 0 1 35 14 4 -- 
              +-- Lanceolaria_lanceolata
  +-----------+                         
  |           +-- Lanceolaria_gladiola  
--+                                     
  |  +----------- Acuticosta_chinensis **  
  |  |                                  
  +--+        +-- Anodonta_anatina      
     |  +-----+                         
     |  |     +-- Pyganodon_grandis     
     +--+                               
        |  +----- Cristaria_plicata     
        +--+                            
           |  +-- Sinanodonta_lucida    
           +--+                         
              +-- Anemina_euscaphys

Pavan-Kumar et al., 2022 [Publ 4263], fig. S5

iT tT bi riS oT roT roS iD
1 1 0 1 38 16 4 -- 
           +----- Acuticosta_chinensis **       
  +--------+                                 
  |        |  +-- Lanceolaria_lanceolata     
  |        +--+                              
  |           +-- Lanceolaria_gladiola       
--+                                          
  |        +----- Lepidodesma_languilati     
  |  +-----+                                 
  |  |     |  +-- Aculamprotula_tientsinensis
  |  |     +--+                              
  +--+        +-- Aculamprotula_tortuosa     
     |                                       
     |  +-------- Unio_pictorum              
     +--+                                    
        |  +----- Schistodesmus_lampreyanus  
        +--+                                 
           |  +-- Nodularia_douglasiae       
           +--+                              
              +-- Tchangsinaia_piscicula

Konopleva et al., 2023 [Publ 4269], fig. S1, 4

iT tT bi riS oT roT roS iD
1 1 0 1 55 32 4 -- 
                 +-------- Lepidodesma_languilati  
  +--------------+                                 
  |              |  +----- Lanceolaria_gladiola    
  |              +--+                              
  |                 |  +-- Lanceolaria_grayii      
  |                 +--+                           
--+                    +-- Lanceolaria_triformis   
  |                                                
  |  +-------------------- Acuticosta_chinensis **    
  |  |                                             
  |  |              +----- Anodonta_anatina        
  |  |  +-----------+                              
  +--+  |           |  +-- Pseudanodonta_complanata
     |  |           +--+                           
     |  |              +-- Anodonta_cygnea         
     |  |                                          
     |  |        +-------- Anemina_arcaeformis     
     +--+  +-----+                                 
        |  |     |  +----- Amuranodonta_kijaensis  
        |  |     +--+                              
        |  |        |  +-- Buldowskia_suifunica    
        |  |        +--+                           
        |  |           +-- Buldowskia_shadini      
        +--+                                       
           |     +-------- Beringiana_beringiana   
           |  +--+                                 
           |  |  |  +----- Sinanodonta_schrenkii   
           |  |  +--+                              
           |  |     |  +-- Sinanodonta_woodiana    
           +--+     +--+                           
              |        +-- Sinanodonta_lauta       
              |                                    
              |  +-------- Cristaria_radiata       
              |  |                                 
              +--+     +-- Cristaria_truncata      
                 |  +--+                           
                 |  |  +-- Cristaria_plicata       
                 +--+                              
                    |  +-- Cristaria_bellua        
                    +--+                           
                       +-- Cristaria_clessini

Dai et al., 2023 [Publ 4285], fig. 5

iT tT bi riS oT roT roS iD
1 1 0 1 52 16 4 -- 
              +-- Acuticosta_chinensis **  
  +-----------+                         
  |           +-- Lanceolaria_grayii    
  |                                     
--+        +----- Cristaria_plicata     
  |  +-----+                            
  |  |     |  +-- Anemina_arcaeformis   
  |  |     +--+                         
  +--+        +-- Sinanodonta_woodiana  
     |                                  
     |  +-------- Anodonta_cygnea       
     +--+                               
        |  +----- Platynaias_compressa  
        +--+                            
           |  +-- Pyganodon_grandis     
           +--+                         
              +-- Utterbackia_imbecillis

Wu et al., 2023 [Publ 4296], fig. 4

iT tT bi riS oT roT roS iD
1 1 0 1 22 20 3 -- 
  +-------------------------------- Acuticosta_chinensis **         
  |                                                              
  |                          +----- Lanceolaria_gladiola         
  |  +-----------------------+                                   
--+  |                       |  +-- Lanceolaria_lanceolata       
  |  |                       +--+                                
  |  |                          +-- Lanceolaria_grayii           
  |  |                                                           
  +--+                    +-------- Pseudanodonta_complanata     
     |  +-----------------+                                      
     |  |                 |  +----- Lasmigona_costata            
     |  |                 +--+                                   
     |  |                    |  +-- Strophitus_undulatus         
     |  |                    +--+                                
     +--+                       +-- Pyganodon_grandis            
        |                                                        
        |  +----------------------- Unio_tumidus                 
        |  |                                                     
        |  |                    +-- Aculamprotula_scripta        
        +--+  +-----------------+                                
           |  |                 +-- Aculamprotula_tientsinensis  
           |  |                                                  
           +--+                 +-- Cuneopsis_heudei             
              |  +--------------+                                
              |  |              +-- Cuneopsis_rufescens          
              |  |                                               
              +--+           +----- Nodularia_douglasiae         
                 |  +--------+                                   
                 |  |        |  +-- Schistodesmus_lampreyanus    
                 |  |        +--+                                
                 +--+           +-- Schistodesmus_spinosus       
                    |                                            
                    |  +----------- Tchangsinaia_piscicula       
                    +--+                                         
                       |  +-------- Inversiunio_yanagawensis     
                       +--+                                      
                          |  +----- Pseudocuneopsis_capitata     
                          +--+                                   
                             |  +-- Pseudocuneopsis_yangshuoensis
                             +--+                                
                                +-- Pseudocuneopsis_sichuanensis

Chen et al., 2023 [Publ 4302], fig. 1

iT tT bi riS oT roT roS iD
1 1 0 1 39 30 4 -- 
                       +----- Pletholophus_tenuis         
  +--------------------+                                  
  |                    |  +-- Sinanodonta_jourdyi         
  |                    +--+                               
  |                       +-- Beringiana_beringiana       
  |                                                       
  |  +----------------------- Lanceolaria_gladiola        
  |  |                                                    
  |  +----------------------- Acuticosta_chinensis **        
--+  |                                                    
  |  +----------------------- Lepidodesma_languilati      
  |  |                                                    
  |  +----------------------- Lanceolaria_oxyrhyncha      
  |  |                                                    
  |  +----------------------- Cristaria_plicata           
  |  |                                                    
  |  |                    +-- Alasmidonta_marginata       
  +--+                 +--+                               
     |                 |  +-- Lasmigona_costata           
     +-----------------+                                  
     |                 |  +-- Anodonta_anatina            
     |                 +--+                               
     |                    +-- Pseudanodonta_complanata    
     |                                                    
     |  +-------------------- Aculamprotula_fibrosa       
     |  |                                                 
     |  |              +----- Middendorffinaia_mongolica  
     |  |  +-----------+                                  
     +--+  |           |  +-- Nodularia_douglasiae        
        |  |           +--+                               
        |  |              +-- Nodularia_breviconcha       
        |  |                                              
        |  |              +-- Cuneopsis_celtiformis       
        +--+  +-----------+                               
           |  |           +-- Cuneopsis_heudei            
           |  |                                           
           |  |           +-- Diaurora_aurorea            
           |  |        +--+                               
           |  |        |  +-- Diaurora_laeve              
           |  +--------+                                  
           +--+        |  +-- Schistodesmus_lampreyanus   
              |        +--+                               
              |           +-- Schistodesmus_spinosus      
              |                                           
              |           +-- Pseudocuneopsis_sichuanensis
              |  +--------+                               
              |  |        +-- Pseudocuneopsis_capitata    
              |  |                                        
              +--+        +-- Inversiunio_yanagawensis    
                 |  +-----+                               
                 |  |     +-- Inversiunio_reinianus       
                 +--+                                     
                    |  +----- Tchangsinaia_piscicula      
                    +--+                                  
                       |  +-- Unio_pictorum               
                       +--+                               
                          +-- Unio_crassus

Chen et al., 2023 [Publ 4302], fig. 2

iT tT bi riS oT roT roS iD
1 1 0 1 39 30 4 -- 
  +-------------------------- Acuticosta_chinensis **        
  |                                                       
  |  +----------------------- Lepidodesma_languilati      
  |  |                                                    
  |  |                    +-- Lanceolaria_gladiola        
  |  |     +--------------+                               
  |  |     |              +-- Lanceolaria_oxyrhyncha      
  |  |     |                                              
--+  |     |              +-- Alasmidonta_marginata       
  |  +-----+           +--+                               
  |  |     |           |  +-- Lasmigona_costata           
  |  |     |  +--------+                                  
  |  |     |  |        |  +-- Anodonta_anatina            
  |  |     |  |        +--+                               
  |  |     +--+           +-- Pseudanodonta_complanata    
  |  |        |                                           
  |  |        |  +----------- Cristaria_plicata           
  +--+        |  |                                        
     |        +--+        +-- Anemina_arcaeformis         
     |           |  +-----+                               
     |           |  |     +-- Amuranodonta_kijaensis      
     |           +--+                                     
     |              |  +----- Pletholophus_tenuis         
     |              +--+                                  
     |                 |  +-- Sinanodonta_jourdyi         
     |                 +--+                               
     |                    +-- Beringiana_beringiana       
     |                                                    
     |  +-------------------- Aculamprotula_fibrosa       
     |  |                                                 
     |  |              +----- Middendorffinaia_mongolica  
     |  |  +-----------+                                  
     +--+  |           |  +-- Nodularia_douglasiae        
        |  |           +--+                               
        |  |              +-- Nodularia_breviconcha       
        |  |                                              
        |  |              +-- Cuneopsis_celtiformis       
        +--+  +-----------+                               
           |  |           +-- Cuneopsis_heudei            
           |  |                                           
           |  |           +-- Diaurora_aurorea            
           |  |        +--+                               
           |  |        |  +-- Diaurora_laeve              
           |  +--------+                                  
           +--+        |  +-- Schistodesmus_lampreyanus   
              |        +--+                               
              |           +-- Schistodesmus_spinosus      
              |                                           
              |           +-- Pseudocuneopsis_sichuanensis
              |  +--------+                               
              |  |        +-- Pseudocuneopsis_capitata    
              |  |                                        
              +--+        +-- Inversiunio_yanagawensis    
                 |  +-----+                               
                 |  |     +-- Inversiunio_reinianus       
                 +--+                                     
                    |  +----- Tchangsinaia_piscicula      
                    +--+                                  
                       |  +-- Unio_pictorum               
                       +--+                               
                          +-- Unio_crassus

Liu et al., 2023 [Publ 4310], fig. 4

iT tT bi riS oT roT roS iD
1 1 0 1 23 21 3 -- 
  +-------------------------------------- Acuticosta_chinensis **         
  |                                                                    
  |                                   +-- Lanceolaria_lanceolata       
--+  +--------------------------------+                                
  |  |                                +-- Lanceolaria_grayii           
  |  |                                                                 
  +--+  +-------------------------------- Lanceolaria_gladiola         
     |  |                                                              
     |  |                       +-------- Pseudanodonta_complanata     
     |  |  +--------------------+                                      
     +--+  |                    |  +----- Lasmigona_costata            
        |  |                    +--+                                   
        |  |                       |  +-- Strophitus_undulatus         
        |  |                       +--+                                
        +--+                          +-- Pyganodon_grandis            
           |                                                           
           |  +-------------------------- Unio_tumidus                 
           |  |                                                        
           |  |                       +-- Aculamprotula_scripta        
           +--+  +--------------------+                                
              |  |                    +-- Aculamprotula_tientsinensis  
              |  |                                                     
              +--+                    +-- Cuneopsis_heudei             
                 |  +-----------------+                                
                 |  |                 +-- Cuneopsis_rufescens          
                 |  |                                                  
                 +--+              +----- Nodularia_douglasiae         
                    |  +-----------+                                   
                    |  |           |  +-- Schistodesmus_lampreyanus    
                    |  |           +--+                                
                    +--+              +-- Schistodesmus_spinosus       
                       |                                               
                       |  +-------------- Tchangsinaia_piscicula       
                       +--+                                            
                          |  +----------- Inversiunio_yanagawensis     
                          +--+                                         
                             |  +-------- Pseudocuneopsis_capitata     
                             +--+                                      
                                |  +----- Pseudocuneopsis_sichuanensis 
                                +--+                                   
                                   |  +-- Pseudocuneopsis_wuana        
                                   +--+                                
                                      +-- Pseudocuneopsis_yangshuoensis

Liu et al., 2023 [Publ 4310], fig. 5

iT tT bi riS oT roT roS iD
1 1 0 1 23 21 3 -- 
  +-------------------------- Acuticosta_chinensis **         
  |                                                        
  |  +----------------------- Lanceolaria_grayii           
  |  |                                                     
--+  +----------------------- Lanceolaria_gladiola         
  |  |                                                     
  |  +----------------------- Lanceolaria_lanceolata       
  |  |                                                     
  +--+              +-------- Lasmigona_costata            
     |  +-----------+                                      
     |  |           |  +----- Pseudanodonta_complanata     
     |  |           +--+                                   
     |  |              |  +-- Pyganodon_grandis            
     |  |              +--+                                
     |  |                 +-- Strophitus_undulatus         
     +--+                                                  
        |  +----------------- Unio_tumidus                 
        |  |                                               
        |  |              +-- Aculamprotula_tientsinensis  
        |  +--------------+                                
        |  |              +-- Aculamprotula_scripta        
        |  |                                               
        +--+              +-- Cuneopsis_rufescens          
           |  +-----------+                                
           |  |           +-- Cuneopsis_heudei             
           |  |                                            
           |  |        +----- Nodularia_douglasiae         
           |  +--------+                                   
           +--+        |  +-- Schistodesmus_lampreyanus    
              |        +--+                                
              |           +-- Schistodesmus_spinosus       
              |                                            
              |  +----------- Tchangsinaia_piscicula       
              |  |                                         
              +--+----------- Inversiunio_yanagawensis     
                 |                                         
                 |  +-------- Pseudocuneopsis_capitata     
                 +--+                                      
                    |  +----- Pseudocuneopsis_sichuanensis 
                    +--+                                   
                       |  +-- Pseudocuneopsis_wuana        
                       +--+                                
                          +-- Pseudocuneopsis_yangshuoensis

Wu et al., 2023 [Publ 4339], fig. 4, S5-aa

iT tT bi riS oT roT roS iD
1 1 0 1 95 46 4 -- 
           +-------- Acuticosta_chinensis **    
  +--------+                                 
  |        |  +----- Lanceolaria_gladiola    
  |        +--+                              
  |           |  +-- Lanceolaria_grayii      
  |           +--+                           
  |              +-- Lanceolaria_lanceolata  
  |                                          
  |           +----- Cristaria_plicata       
  |     +-----+                              
  |     |     |  +-- Anemina_arcaeformis     
--+     |     +--+                           
  |  +--+        +-- Anemina_euscaphys       
  |  |  |                                    
  |  |  |  +-------- Beringiana_fukuharai    
  |  |  +--+                                 
  |  |     |  +----- Sinanodonta_lucida      
  |  |     +--+                              
  |  |        |  +-- Sinanodonta_tumens      
  |  |        +--+                           
  |  |           +-- Sinanodonta_woodiana    
  +--+                                       
     |           +-- Anodonta_nuttalliana    
     |     +-----+                           
     |     |     +-- Pseudanodonta_complanata
     |  +--+                                 
     |  |  |  +----- Anodonta_anatina        
     |  |  +--+                              
     |  |     |  +-- Anodonta_cygnea         
     |  |     +--+                           
     |  |        +-- Anodonta_exulcerata     
     +--+                                    
        |     +----- Pyganodon_grandis       
        |  +--+                              
        |  |  |  +-- Utterbackia_peninsularis
        |  |  +--+                           
        |  |     +-- Utterbackia_imbecillis  
        +--+                                 
           |     +-- Alasmidonta_varicosa    
           |  +--+                           
           |  |  +-- Platynaias_compressa    
           +--+                              
              |  +-- Prolasmidonta_heterodon 
              +--+                           
                 +-- Lasmigona_complanata

Wu et al., 2023 [Publ 4339], fig. 4, S5-nt

iT tT bi riS oT roT roS iD
1 1 0 1 96 47 4 -- 
                 +-------- Acuticosta_chinensis **    
  +--------------+                                 
  |              |  +----- Lanceolaria_gladiola    
  |              +--+                              
  |                 |  +-- Lanceolaria_grayii      
  |                 +--+                           
  |                    +-- Lanceolaria_lanceolata  
  |                                                
  |        +-------------- Cristaria_plicata       
--+        |                                       
  |  +-----+           +-- Anemina_arcaeformis     
  |  |     |  +--------+                           
  |  |     |  |        +-- Anemina_euscaphys       
  |  |     +--+                                    
  |  |        |  +-------- Beringiana_fukuharai    
  |  |        +--+                                 
  |  |           |  +----- Sinanodonta_lucida      
  |  |           +--+                              
  +--+              |  +-- Sinanodonta_tumens      
     |              +--+                           
     |                 +-- Sinanodonta_woodiana    
     |                                             
     |  +----------------- Anodonta_nuttalliana    
     |  |                                          
     |  +----------------- Pseudanodonta_complanata
     |  |                                          
     +--+  +-------------- Anodonta_cygnea         
        |  |                                       
        |  |           +-- Anodonta_exulcerata     
        |  +-----------+                           
        |  |           +-- Anodonta_anatina        
        +--+                                       
           |        +----- Pyganodon_grandis       
           |  +-----+                              
           |  |     |  +-- Utterbackia_imbecillis  
           |  |     +--+                           
           +--+        +-- Utterbackia_peninsularis
              |                                    
              |  +-------- Lasmigona_complanata    
              +--+                                 
                 |  +----- Prolasmidonta_heterodon 
                 +--+                              
                    |  +-- Alasmidonta_varicosa    
                    +--+                           
                       +-- Platynaias_compressa

Wu et al., 2024 [Publ 4350], fig. 8a, S1

iT tT bi riS oT roT roS iD
1 1 0 1 91 46 4 -- 
                 +-------- Acuticosta_chinensis **    
  +--------------+                                 
  |              |  +----- Lanceolaria_gladiola    
  |              +--+                              
  |                 |  +-- Lanceolaria_grayii      
  |                 +--+                           
  |                    +-- Lanceolaria_lanceolata  
  |                                                
  |        +-------------- Cristaria_plicata       
  |        |                                       
--+  +-----+           +-- Anemina_arcaeformis     
  |  |     |  +--------+                           
  |  |     |  |        +-- Anemina_euscaphys       
  |  |     +--+                                    
  |  |        |  +-------- Beringiana_fukuharai    
  |  |        +--+                                 
  |  |           |  +----- Sinanodonta_lucida      
  |  |           +--+                              
  |  |              |  +-- Sinanodonta_tumens      
  +--+              +--+                           
     |                 +-- Sinanodonta_woodiana    
     |                                             
     |           +-------- Pseudanodonta_complanata
     |  +--------+                                 
     |  |        |  +----- Anodonta_anatina        
     |  |        +--+                              
     |  |           |  +-- Anodonta_cygnea         
     |  |           +--+                           
     +--+              +-- Anodonta_exulcerata     
        |                                          
        |  +-------------- Anodonta_nuttalliana    
        |  |                                       
        |  |        +----- Pyganodon_grandis       
        +--+  +-----+                              
           |  |     |  +-- Utterbackia_imbecillis  
           |  |     +--+                           
           +--+        +-- Utterbackia_peninsularis
              |                                    
              |  +-------- Lasmigona_complanata    
              +--+                                 
                 |  +----- Prolasmidonta_heterodon 
                 +--+                              
                    |  +-- Alasmidonta_varicosa    
                    +--+                           
                       +-- Platynaias_compressa

Wu et al., 2024 [Publ 4350], fig. 8b, S2

iT tT bi riS oT roT roS iD
1 1 0 1 91 46 4 -- 
                 +-------- Acuticosta_chinensis **    
  +--------------+                                 
  |              |  +----- Lanceolaria_gladiola    
  |              +--+                              
  |                 |  +-- Lanceolaria_grayii      
  |                 +--+                           
  |                    +-- Lanceolaria_lanceolata  
  |                                                
  |        +-------------- Cristaria_plicata       
--+        |                                       
  |  +-----+           +-- Anemina_arcaeformis     
  |  |     |  +--------+                           
  |  |     |  |        +-- Anemina_euscaphys       
  |  |     +--+                                    
  |  |        |  +-------- Beringiana_fukuharai    
  |  |        +--+                                 
  |  |           |  +----- Sinanodonta_lucida      
  |  |           +--+                              
  +--+              |  +-- Sinanodonta_tumens      
     |              +--+                           
     |                 +-- Sinanodonta_woodiana    
     |                                             
     |  +----------------- Anodonta_nuttalliana    
     |  |                                          
     |  |        +-------- Pseudanodonta_complanata
     |  |  +-----+                                 
     +--+  |     |  +----- Anodonta_anatina        
        |  |     +--+                              
        |  |        |  +-- Anodonta_cygnea         
        |  |        +--+                           
        +--+           +-- Anodonta_exulcerata     
           |                                       
           |        +----- Pyganodon_grandis       
           |  +-----+                              
           |  |     |  +-- Utterbackia_imbecillis  
           |  |     +--+                           
           +--+        +-- Utterbackia_peninsularis
              |                                    
              |  +-------- Lasmigona_complanata    
              +--+                                 
                 |  +----- Prolasmidonta_heterodon 
                 +--+                              
                    |  +-- Alasmidonta_varicosa    
                    +--+                           
                       +-- Platynaias_compressa

Dai et al., 2024 [Publ 4358], fig. 4

iT tT bi riS oT roT roS iD
1 1 0 1 51 26 4 -- 
  +----------------- Lanceolaria_gladiola  
  |                                        
--+  +-------------- Acuticosta_chinensis **  
  |  |                                     
  |  |        +----- Cristaria_plicata     
  +--+  +-----+                            
     |  |     |  +-- Anemina_arcaeformis   
     |  |     +--+                         
     +--+        +-- Sinanodonta_woodiana  
        |                                  
        |  +-------- Anodonta_anatina      
        +--+                               
           |  +----- Platynaias_compressa  
           +--+                            
              |  +-- Pyganodon_grandis     
              +--+                         
                 +-- Utterbackia_imbecillis

Page updated: 08:51:02 Mon 06 May 2024
 
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