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Page last updated
Tue 03 Jun 2025

The Freshwater Mussels (Unionoida) of the World (and other less consequential bivalves)

FM(U)otW(aolcb) is the web version of the MUSSEL Project Database. Follow the links to browse the data or use the custom Google search field. Either way, you win!

Cladomics pages are experimental — even more so than the rest of this site!

The purpose of the cladomics pages for each genus and family-group level taxon is to provide statistics and references indicating the extent to which published phylogenetic work supports the classification used here.

The Cladomics Summary provides an overview of the number of studies that have included terminal species representing this taxon as well as an indication of how frequently it has been recovered as monophyletic.

Following the summary, the available published clades representing the taxon are listed.

For more information, visit the Cladomics FARQ (frequently asked rhetorical questions) page. Abbreviations for the various statistics are explained on the FARQ!

Search the database pages:

Bivalvia | Unionoida | Unionoidea | Unionidae | Unioninae | Anodontini

subtribe ALASMIDONTINA Rafinesque, 1820: cladomics
     15 genera

Cladomics Summary

number of species = 51
number of subtaxa (rank: genus) = 15

analyzed in 204 cladograms in 121 publications
maximum number of terminals in an analysis = 33, mean = 4.12
maximum number of subtaxa (genus) in an analysis = 13, mean = 3.0

Basic Monophyly

number of cladograms that test basic monophyly (i.e., more than 1 included species) = 169
maximum number of included terminals = 33, mean = 4.76
maximum number of subtaxa (genus) = 13, mean = 3.41
proportion of cladograms that support basic monophyly = 0.84

Rigorous Monophyly

number of cladograms that test rigorous monophyly = 124
(Rigorous monophyly refers to cladograms with terminal species representing more than one subtaxon (genus) taxon and at least one rigorous outgroup (subtribe).
maximum number of included terminals = 33, mean = 5.1
maximum number of subtaxa (genus) = 13, mean = 3.72
proportion of cladograms that support rigorous monophyly = 0.79

Pairwise Cladogram Clustering

The graphs below depict in two dimensions all pair-wise ingroup distances within the cladome. See the Cladmoics FARQ for more information.


PCoA Ordination


NJ tree

Whelan et al., 2023 [Publ 4287], fig. 2

iT tT bi riS oT roT roS iD
33 33 34 13 8 8 2 0.0

*** Top Monophyly Cladogram *** 

  +-------------------------- Cristariina         
  |                                               
--+  +----------------------- Anodonta_cygnea     
  |  |                                            
  |  |                    +-- Anodonta_nuttalliana
  +--+  +-----------------+                       
     |  |                 +-- Pseudanodonta       
     |  |                                         
     +--+  +----------------- Anodonta_anatina    
        |  |                                      
        |  |              +-- Utterbackia **         
        |  |  +-----------+                       
        +--+  |           +-- Pyganodon **           
           |  |                                   
           |  |     +-------- Utterbackiana **       
           |  |     |                             
           +--+  +--+     +-- Lasmigona_costata **   
              |  |  |  +--+                       
              |  |  |  |  +-- Pseudodontoideus **    
              |  |  +--+                          
              |  |     |  +-- Lasmigona_complanata **
              |  |     +--+                       
              +--+        +-- Arcidens **            
                 |                                
                 |     +----- Pressodonta **         
                 |  +--+                          
                 |  |  |  +-- Anodontoides **        
                 |  |  +--+                       
                 |  |     +-- Strophitus **          
                 +--+                             
                    |     +-- Prolasmidonta **       
                    |  +--+                       
                    |  |  +-- Platynaias **          
                    +--+                          
                       |  +-- Alasminota **          
                       +--+                       
                          +-- Alasmidonta **

Lopes-Lima et al., 2020 [Publ 4080], fig. 2-4

iT tT bi riS oT roT roS iD
5 5 1 5 116 28 2 36.0

*** Top Sister-Group Cladogram ***

  +----------- Cristariina  
  |                         
--+  +-------- Pseudanodonta
  |  |                      
  |  +-------- Anodonta     
  +--+                      
     |  +----- Pyganodon **    
     |  |                   
     |  +----- Simpsonaias **  
     +--+                   
        +----- Strophitus **   
        |                   
        |  +-- Alasmidonta **  
        +--+                
           +-- Lasmigona **

Whelan et al., 2023 [Publ 4287], fig. 3

iT tT bi riS oT roT roS iD
27 27 26 12 1 1 1 23.0 
  +----------------------- Anodontina      
  |                                        
--+  +-------------------- Utterbackiana **   
  |  |                                     
  |  |                 +-- Utterbackia **     
  +--+  +--------------+                   
     |  |              +-- Pyganodon **       
     |  |                                  
     +--+           +----- Alasmidonta **     
        |  +--------+                      
        |  |        |  +-- Arcidens **        
        |  |        +--+                   
        +--+           +-- Pseudodontoideus **
           |                               
           |  +----------- Alasminota **      
           |  |                            
           +--+        +-- Anodontoides **    
              |  +-----+                   
              |  |     +-- Strophitus **      
              +--+                         
                 |  +----- Pressodonta **     
                 +--+                      
                    |  +-- Platynaias **      
                    +--+                   
                       +-- Prolasmidonta **

Whelan et al., 2023 [Publ 4287], fig. 4

iT tT bi riS oT roT roS iD
25 25 23 11 2 2 1 38.0 
  +----------------------- Anodontina      
  |                                        
--+  +-------------------- Pressodonta **     
  |  |                                     
  |  |           +-------- Utterbackiana **   
  +--+  +--------+                         
     |  |        |  +----- Alasminota **      
     |  |        +--+                      
     |  |           |  +-- Arcidens **        
     +--+           +--+                   
        |              +-- Pseudodontoideus **
        |                                  
        |  +-------------- Alasmidonta **     
        +--+                               
           |  +----------- Platynaias **      
           +--+                            
              |  +-------- Strophitus **      
              +--+                         
                 |  +----- Pyganodon **       
                 +--+                      
                    |  +-- Utterbackia **     
                    +--+                   
                       +-- Anodontoides **

Smith et al., 2018 [Publ 3790], fig. 1

iT tT bi riS oT roT roS iD
17 17 16 8 1 1 1 25.0 
  +-------------- Cristariina     
  |                               
  |           +-- Utterbackia **     
--+  +--------+                   
  |  |        +-- Pyganodon **       
  |  |                            
  +--+        +-- Anodontoides **    
     |  +-----+                   
     |  |     +-- Strophitus **      
     |  |                         
     +--+     +-- Alasmidonta **     
        |  +--+                   
        |  |  +-- Alasminota **      
        +--+                      
           |  +-- Pseudodontoideus **
           +--+                   
              +-- Utterbackiana **

Inoue et al., 2018 [Publ 3858], fig. 1_3a-c

iT tT bi riS oT roT roS iD
16 20 0 9 88 4 2 39.0 
  +-- Alasmidonta_marginata **      
  |                              
  +-- Prolasmidonta **              
  |                              
  +-- Alasmidonta_undulata **       
  |                              
  +-- Cristariina                
  |                              
  +-- Anodonta_kennerlyi         
  |                              
  +-- Anodonta_nuttalliana       
  |                              
  +-- Anodonta_oregonensis       
  |                              
  +-- Arcidens_confragosus **       
  |                              
  +-- Arcidens_wheeleri **          
  |                              
  +-- Lasmigona_complanata **       
  |                              
--+-- Platynaias_compressa **       
  |                              
  +-- Lasmigona_costata **          
  |                              
  +-- Platynaias_decorata **        
  |                              
  +-- Platynaias_subviridis **      
  |                              
  +-- Pyganodon_grandis **          
  |                              
  +-- Pyganodon_lacustris **        
  |                              
  +-- Pseudodontoideus **           
  |                              
  +-- Strophitus **                 
  |                              
  +-- Utterbackia_imbecillis **     
  |                              
  +-- Utterbackia_peninsularis__1 **
  |                              
  +-- Utterbackia_peninsularis__2 **

Zanatta et al., 2007 [Publ 2698], fig. 1B

iT tT bi riS oT roT roS iD
14 16 13 5 2 2 1 41.0 
           +-- Lasmigona_costata **   
           |                       
  +--------+-- Platynaias **          
  |        |                       
  |        +-- Lasmigona_complanata **
--+                                
  |        +-- Utterbackia **         
  |  +-----+                       
  |  |     +-- Pyganodon **           
  +--+                             
     |  +----- Anodonta_cygnea     
     +--+                          
        |  +-- Anodonta_kennerlyi  
        +--+                       
           +-- Utterbackiana **

Inoue et al., 2014 [Publ 3635], fig. 3A

iT tT bi riS oT roT roS iD
14 14 13 8 6 0 0 28.0 
     +-------- Pseudodontoideus **    
  +--+                             
  |  |  +----- Lasmigona_costata **   
  |  +--+                          
  |     |  +-- Lasmigona_complanata **
--+     +--+                       
  |        +-- Arcidens **            
  |                                
  |  +-------- Strophitus **          
  |  |                             
  +--+     +-- Pyganodon **           
     |  +--+                       
     |  |  +-- Utterbackia **         
     +--+                          
        |  +-- Platynaias **          
        +--+                       
           +-- Alasmidonta **

Zanatta et al., 2007 [Publ 2698], fig. 1A

iT tT bi riS oT roT roS iD
14 16 10 5 2 2 1 39.0 
           +-- Platynaias **          
           |                       
  +--------+-- Lasmigona_costata **   
  |        |                       
  |        +-- Lasmigona_complanata **
--+                                
  |  +-------- Anodonta_cygnea     
  |  |                             
  +--+     +-- Anodonta_kennerlyi  
     |  +--+                       
     |  |  +-- Utterbackiana **       
     +--+                          
        |  +-- Utterbackia **         
        +--+                       
           +-- Pyganodon **

Hoeh, 1990 [Publ 2717], fig. 4

iT tT bi riS oT roT roS iD
13 15 13 5 2 2 1 41.0 
           +-- Lasmigona_complanata **
           |                       
  +--------+-- Platynaias **          
  |        |                       
  |        +-- Lasmigona_costata **   
--+                                
  |        +-- Utterbackia **         
  |  +-----+                       
  |  |     +-- Pyganodon **           
  +--+                             
     |  +----- Anodonta_cygnea     
     +--+                          
        |  +-- Anodonta_kennerlyi  
        +--+                       
           +-- Utterbackiana **

Hoeh, 1990 [Publ 2717], fig. 3

iT tT bi riS oT roT roS iD
13 15 2 5 2 2 1 43.0 
  +----- Pyganodon_fragilis **         
  |                                 
  +----- Utterbackia_imbecillis **     
  |                                 
  +----- Utterbackia_peggyae **        
  |                                 
  +----- Anodonta_cygnea            
  |                                 
  +----- Anodonta_kennerlyi         
  |                                 
  +----- Utterbackiana_couperiana__1 **
  |                                 
--+----- Utterbackiana_couperiana__2 **
  |                                 
  +----- Utterbackiana_suborbiculata **
  |                                 
  |  +-- Lasmigona_costata **          
  +--+                              
  |  +-- Utterbackiana_implicata **    
  |                                 
  |  +-- Lasmigona_complanata **       
  |  |                              
  |  +-- Platynaias **                 
  |  |                              
  |  +-- Pyganodon_cataracta **        
  +--+                              
     +-- Pyganodon_gibbosa **          
     |                              
     +-- Pyganodon_grandis **          
     |                              
     +-- Pyganodon_lacustris **

Hoeh, 1990 [Publ 2717], fig. 2

iT tT bi riS oT roT roS iD
13 15 13 5 2 2 1 41.0 
           +-- Lasmigona_complanata **
           |                       
  +--------+-- Platynaias **          
  |        |                       
  |        +-- Lasmigona_costata **   
--+                                
  |        +-- Utterbackia **         
  |  +-----+                       
  |  |     +-- Pyganodon **           
  +--+                             
     |  +----- Anodonta_cygnea     
     +--+                          
        |  +-- Anodonta_kennerlyi  
        +--+                       
           +-- Utterbackiana **

Breton et al., 2011 [Publ 3376], fig. 3

iT tT bi riS oT roT roS iD
11 11 10 6 26 1 1 29.0 
  +-------------- Cristariina
  |                          
--+  +----------- Lasmigona **  
  |  |                       
  +--+  +-------- Strophitus ** 
     |  |                    
     +--+     +-- Platynaias ** 
        |  +--+              
        |  |  +-- Alasmidonta **
        +--+                 
           |  +-- Pyganodon **  
           +--+              
              +-- Utterbackia **

Chase et al., 2018 [Publ 3847], fig. 4

iT tT bi riS oT roT roS iD
11 11 10 6 27 3 2 27.0 
  +----------------- Cristariina
  |                             
--+  +-------------- Anodontina 
  |  |                          
  +--+           +-- Pyganodon **  
     |  +--------+              
     |  |        +-- Utterbackia **
     +--+                       
        |  +-------- Lasmigona **  
        +--+                    
           |  +----- Alasmidonta **
           +--+                 
              |  +-- Strophitus ** 
              +--+              
                 +-- Platynaias **

King et al., 1999 [Publ 2699], fig. 2

iT tT bi riS oT roT roS iD
9 9 7 5 0 0 0 31.0 
  +----------- Pyganodon **    
--+                         
  |  +-------- Lasmigona **    
  +--+                      
     |  +----- Strophitus **   
     +--+                   
        |  +-- Prolasmidonta **
        +--+                
           +-- Platynaias **

Boyer et al., 2011 [Publ 2718], fig. 4

iT tT bi riS oT roT roS iD
9 9 6 8 29 0 0 38.0 
  +-------------- Strophitus **          
  |                                   
  |        +----- Lasmigona_complanata **
--+  +-----+                          
  |  |     |  +-- Lasmigona_costata **   
  |  |     +--+                       
  +--+        +-- Arcidens **            
     |                                
     |  +-------- Utterbackia **         
     |  |                             
     +--+-------- Alasmidonta **         
        |                             
        |  +----- Pyganodon **           
        +--+                          
           |  +-- Anodontoides **        
           +--+                       
              +-- Platynaias **

Krebs et al., 2003 [Publ 2691], fig. 3

iT tT bi riS oT roT roS iD
9 10 5 7 24 1 1 33.0 
  +----------- Utterbackiana **
  |                         
  +----------- Platynaias **   
  |                         
--+----------- Alasmidonta **  
  |                         
  +----------- Lasmigona **    
  |                         
  |  +-------- Strophitus **   
  +--+                      
     |  +----- Utterbackia **  
     +--+                   
        |  +-- Pyganodon **    
        +--+                
           +-- Anodontina

Chong et al., 2008 [Publ 2668], fig. 3

iT tT bi riS oT roT roS iD
8 18 10 8 10 9 2 45.0 
  +----------------- Anodonta_cygnea        
  |                                         
  +----------------- Pyganodon **              
  |                                         
  +----------------- Alasminota **             
  |                                         
  +----------------- Anodonta_anatina       
  |                                         
  +----------------- Unio                   
  |                                         
  +----------------- Cristariina            
  |                                         
  |              +-- Anodonta_nuttalliana   
--+--------------+                          
  |              +-- Anodonta_californiensis
  |                                         
  |              +-- Pseudodontoideus **       
  |              |                          
  +--------------+-- Alasmidonta **            
  |              |                          
  |              +-- Lasmigona **              
  |                                         
  |  +-------------- Pseudanodonta          
  |  |                                      
  |  |        +----- Strophitus **             
  |  +--------+                             
  +--+        |  +-- Prolasmidonta **          
     |        +--+                          
     |           +-- Platynaias **             
     |                                      
     |  +----------- Anodonta_oregonensis__4
     +--+                                   
        |  +-------- Anodonta_oregonensis__3
        +--+                                
           |  +----- Anodonta_oregonensis__2
           +--+                             
              |  +-- Anodonta_oregonensis__1
              +--+                          
                 +-- Anodonta_kennerlyi

Wu et al., 2021 [Publ 4366], fig. 3-BI

iT tT bi riS oT roT roS iD
7 7 5 5 66 6 2 31.0 
  +----------------- Cristariina     
  |                                  
--+  +-------------- Anodonta_cygnea 
  |  |                               
  +--+  +----------- Anodonta_anatina
     |  |                            
     +--+        +-- Pyganodon **       
        |  +-----+                   
        |  |     +-- Utterbackia **     
        +--+                         
           |  +----- Prolasmidonta **   
           +--+                      
              |  +-- Alasmidonta **     
              +--+                   
                 +-- Platynaias **

Wu et al., 2024 [Publ 4350], fig. 8b_S2

iT tT bi riS oT roT roS iD
7 7 5 6 84 11 2 31.0 
  +-------------------- Cristariina         
  |                                         
  |  +----------------- Anodonta_nuttalliana
--+  |                                      
  |  |        +-------- Pseudanodonta       
  |  |  +-----+                             
  +--+  |     |  +----- Anodonta_anatina    
     |  |     +--+                          
     |  |        |  +-- Anodonta_cygnea     
     |  |        +--+                       
     +--+           +-- Anodonta_exulcerata 
        |                                   
        |           +-- Pyganodon **           
        |  +--------+                       
        |  |        +-- Utterbackia **         
        +--+                                
           |  +-------- Lasmigona **           
           +--+                             
              |  +----- Prolasmidonta **       
              +--+                          
                 |  +-- Alasmidonta **         
                 +--+                       
                    +-- Platynaias **

Inoue et al., 2014 [Publ 3635], fig. 3B

iT tT bi riS oT roT roS iD
7 7 2 3 1 0 0 35.0 
  +----- Pyganodon_grandis **  
  |                         
  +----- Pyganodon_cataracta **
  |                         
--+----- Pyganodon_fragilis ** 
  |                         
  +----- Pyganodon **          
  |                         
  |  +-- Platynaias **         
  +--+                      
     +-- Arcidens **

Hoeh et al., 1995 [Publ 3391], fig. 2

iT tT bi riS oT roT roS iD
7 7 4 2 0 0 0 35.0 
  +-- Utterbackia **
--+              
  +-- Pyganodon **

Smith et al., 2023 [Publ 4291], fig. 2

iT tT bi riS oT roT roS iD
7 7 5 2 0 0 0 33.0 
  +-- Anodontoides **
--+               
  +-- Strophitus **

Huang et al., 2019 [Publ 3843], fig. S4

iT tT bi riS oT roT roS iD
7 45 22 6 192 13 2 71.0 
  +-------- Acuticosta           
  |                              
  +-------- Alasmidonta **          
  |                              
  +-------- Anodonta_anatina     
  |                              
  +-------- Anemina              
  |                              
  +-------- Anodonta_cygnea      
  |                              
  +-------- Cristaria            
  |                              
  +-------- Anodonta_nuttalliana 
  |                              
  +-------- Lasmigona **            
  |                              
  +-------- Lepidodesma          
  |                              
  +-------- Pseudanodonta        
  |                              
  +-------- Pyganodon **            
  |                              
--+-------- Simpsonaias **          
  |                              
  +-------- Strophitus **           
  |                              
  +-------- Schistodesmus        
  |                              
  +-------- Utterbackia **          
  |                              
  +-------- Lanceolaria          
  |                              
  +-------- Unio                 
  |                              
  +-------- Aculamprotula        
  |                              
  |     +-- Pletholophus         
  +-----+                        
  |     +-- Sinanodonta          
  |                              
  |  +----- Cuneopsis_celtiformis
  |  |                           
  |  +----- Tchangsinaia         
  |  |                           
  +--+----- Cuneopsis_rufescens  
     |                           
     +----- Nodularia            
     |                           
     |  +-- Cuneopsis_szechenyii 
     +--+                        
        +-- Cuneopsis_heudei

Wu et al., 2023 [Publ 4339], fig. 4_S5-nt

iT tT bi riS oT roT roS iD
7 7 5 6 88 11 2 31.0 
  +-------------------- Cristariina         
  |                                         
  |  +----------------- Anodonta_nuttalliana
--+  |                                      
  |  +----------------- Pseudanodonta       
  |  |                                      
  +--+  +-------------- Anodonta_cygnea     
     |  |                                   
     |  |           +-- Anodonta_exulcerata 
     |  +-----------+                       
     +--+           +-- Anodonta_anatina    
        |                                   
        |           +-- Pyganodon **           
        |  +--------+                       
        |  |        +-- Utterbackia **         
        +--+                                
           |  +-------- Lasmigona **           
           +--+                             
              |  +----- Prolasmidonta **       
              +--+                          
                 |  +-- Alasmidonta **         
                 +--+                       
                    +-- Platynaias **

Huang et al., 2019 [Publ 3843], fig. S3

iT tT bi riS oT roT roS iD
7 11 8 6 190 13 2 39.0 
  +-------------- Cristariina  
  |                            
  |  +----------- Anodonta     
--+  |                         
  |  |        +-- Alasmidonta **  
  |  +--------+                
  +--+        +-- Lasmigona **    
     |                         
     |        +-- Pyganodon **    
     |  +-----+                
     |  |     +-- Utterbackia **  
     +--+                      
        |  +----- Pseudanodonta
        +--+                   
           |  +-- Simpsonaias **  
           +--+                
              +-- Strophitus **

Hou et al., 2024 [Publ 4400], fig. 4

iT tT bi riS oT roT roS iD
7 7 5 6 69 11 2 31.0 
  +-------------------- Cristariina         
  |                                         
  |           +-------- Pseudanodonta       
--+  +--------+                             
  |  |        |  +----- Anodonta_anatina    
  |  |        +--+                          
  |  |           |  +-- Anodonta_cygnea     
  +--+           +--+                       
     |              +-- Anodonta_exulcerata 
     |                                      
     |  +-------------- Anodonta_nuttalliana
     |  |                                   
     +--+           +-- Pyganodon **           
        |  +--------+                       
        |  |        +-- Utterbackia **         
        +--+                                
           |  +-------- Lasmigona **           
           +--+                             
              |  +----- Prolasmidonta **       
              +--+                          
                 |  +-- Alasmidonta **         
                 +--+                       
                    +-- Platynaias **

Wu et al., 2021 [Publ 4366], fig. 3-ML

iT tT bi riS oT roT roS iD
7 7 5 5 66 6 2 31.0 
  +----------------- Cristariina     
  |                                  
--+  +-------------- Anodonta_cygnea 
  |  |                               
  +--+  +----------- Anodonta_anatina
     |  |                            
     +--+        +-- Pyganodon **       
        |  +-----+                   
        |  |     +-- Utterbackia **     
        +--+                         
           |  +----- Prolasmidonta **   
           +--+                      
              |  +-- Alasmidonta **     
              +--+                   
                 +-- Platynaias **

Huang et al., 2019 [Publ 3843], fig. 2

iT tT bi riS oT roT roS iD
7 11 4 6 192 13 2 37.0 
  +-------- Cristariina  
  |                      
  |  +----- Alasmidonta **  
  |  |                   
--+  +----- Lasmigona **    
  |  |                   
  |  +----- Pseudanodonta
  |  |                   
  +--+----- Simpsonaias **  
     |                   
     +----- Strophitus **   
     |                   
     +----- Anodonta     
     |                   
     |  +-- Pyganodon **    
     +--+                
        +-- Utterbackia **

Wu et al., 2024 [Publ 4350], fig. 8a_S1

iT tT bi riS oT roT roS iD
7 7 5 6 84 11 2 31.0 
  +-------------------- Cristariina         
  |                                         
  |           +-------- Pseudanodonta       
--+  +--------+                             
  |  |        |  +----- Anodonta_anatina    
  |  |        +--+                          
  |  |           |  +-- Anodonta_cygnea     
  +--+           +--+                       
     |              +-- Anodonta_exulcerata 
     |                                      
     |  +-------------- Anodonta_nuttalliana
     |  |                                   
     +--+           +-- Pyganodon **           
        |  +--------+                       
        |  |        +-- Utterbackia **         
        +--+                                
           |  +-------- Lasmigona **           
           +--+                             
              |  +----- Prolasmidonta **       
              +--+                          
                 |  +-- Alasmidonta **         
                 +--+                       
                    +-- Platynaias **

Wu et al., 2024 [Publ 4379], fig. 5

iT tT bi riS oT roT roS iD
7 7 5 6 46 11 2 31.0 
  +-------------------- Cristariina         
  |                                         
  |  +----------------- Anodonta_nuttalliana
--+  |                                      
  |  |        +-------- Pseudanodonta       
  |  |  +-----+                             
  +--+  |     |  +----- Anodonta_anatina    
     |  |     +--+                          
     |  |        |  +-- Anodonta_cygnea     
     |  |        +--+                       
     +--+           +-- Anodonta_exulcerata 
        |                                   
        |           +-- Pyganodon **           
        |  +--------+                       
        |  |        +-- Utterbackia **         
        +--+                                
           |  +-------- Lasmigona **           
           +--+                             
              |  +----- Prolasmidonta **       
              +--+                          
                 |  +-- Alasmidonta **         
                 +--+                       
                    +-- Platynaias **

Wu et al., 2023 [Publ 4339], fig. 4_S5-aa

iT tT bi riS oT roT roS iD
7 7 5 6 88 11 2 33.0 
  +-------------- Cristariina  
  |                            
--+  +----------- Anodontina   
  |  |                         
  |  |        +-- Pyganodon **    
  +--+  +-----+                
     |  |     +-- Utterbackia **  
     |  |                      
     +--+     +-- Alasmidonta **  
        |  +--+                
        |  |  +-- Platynaias **   
        +--+                   
           |  +-- Prolasmidonta **
           +--+                
              +-- Lasmigona **

Sano et al., 2020 [Publ 4138], fig. 4

iT tT bi riS oT roT roS iD
6 6 4 5 62 13 2 30.0 
  +----------- Anodontina   
  |                         
--+        +-- Pyganodon **    
  |  +-----+                
  |  |     +-- Utterbackia **  
  +--+                      
     |  +----- Alasmidonta **  
     +--+                   
        |  +-- Prolasmidonta **
        +--+                
           +-- Platynaias **

Sano et al., 2020 [Publ 4138], fig. 3

iT tT bi riS oT roT roS iD
6 6 4 5 62 13 2 30.0 
  +----------- Anodontina   
  |                         
--+        +-- Pyganodon **    
  |  +-----+                
  |  |     +-- Utterbackia **  
  +--+                      
     |  +----- Alasmidonta **  
     +--+                   
        |  +-- Platynaias **   
        +--+                
           +-- Prolasmidonta **

Sano et al., 2020 [Publ 4138], fig. 2

iT tT bi riS oT roT roS iD
6 6 4 5 62 13 2 30.0 
  +----------- Anodontina   
  |                         
--+        +-- Pyganodon **    
  |  +-----+                
  |  |     +-- Utterbackia **  
  +--+                      
     |  +----- Alasmidonta **  
     +--+                   
        |  +-- Prolasmidonta **
        +--+                
           +-- Platynaias **

Chen et al., 2020 [Publ 4170], fig. 1

iT tT bi riS oT roT roS iD
6 34 32 5 28 5 2 65.0 
                       +----- Prolasmidonta **           
                    +--+                              
                    |  |  +-- Alasmidonta **             
                    |  +--+                           
  +-----------------+     +-- Anemina                 
  |                 |                                 
  |                 |  +----- Pseudodontina           
  |                 +--+                              
  |                    |  +-- Lamprotula_caveata      
  |                    +--+                           
  |                       +-- Chamberlainia           
  |                                                   
  |                    +----- Schistodesmus           
  |              +-----+                              
  |              |     |  +-- Lepidodesma             
--+              |     +--+                           
  |              |        +-- Aculamprotula           
  |  +-----------+                                    
  |  |           |     +----- Platynaias **              
  |  |           |  +--+                              
  |  |           |  |  |  +-- Utterbackia_imbecillis **  
  |  |           |  |  +--+                           
  |  |           +--+     +-- Pyganodon **               
  |  |              |                                 
  |  |              |  +----- Sinanodonta             
  |  |              +--+                              
  |  |                 |  +-- Cristaria               
  +--+                 +--+                           
     |                    +-- Anodontina              
     |                                                
     |                    +-- Utterbackia_peninsularis **
     |           +--------+                           
     |           |        +-- Potamilus_alatus        
     |  +--------+                                    
     |  |        |  +-------- Lamprotula_leaii        
     |  |        +--+                                 
     |  |           |  +----- Sinohyriopsis           
     |  |           +--+                              
     +--+              |  +-- Parvasolenaia           
        |              +--+                           
        |                 +-- Sinosolenaia            
        |                                             
        |  +----------------- Gibbosula               
        +--+                                          
           |  +-------------- Quadrulina              
           +--+                                       
              |  +----------- Toxolasma               
              +--+                                    
                 |  +-------- Lampsilis_ornata        
                 +--+                                 
                    |  +----- Potamilus_leptodon      
                    +--+                              
                       |  +-- Lampsilis_siliquoidea   
                       +--+                           
                          +-- Lampsilis_powellii

Boyer et al., 2011 [Publ 2718], fig. 3

iT tT bi riS oT roT roS iD
6 6 2 5 29 0 0 34.0 
  +----- Alasmidonta **
  |                 
  +----- Pyganodon **  
--+                 
  +----- Lasmigona **  
  |                 
  |  +-- Platynaias ** 
  +--+              
     +-- Strophitus **

Lydeard et al., 1996 [Publ 2656], fig. 3

iT tT bi riS oT roT roS iD
6 6 4 5 21 0 0 38.0 
        +-- Utterbackia **  
  +-----+                
  |     +-- Utterbackiana **
--+                      
  |  +----- Strophitus **   
  +--+                   
     |  +-- Lasmigona **    
     +--+                
        +-- Alasmidonta **

Wu et al., 2021 [Publ 4366], fig. 4

iT tT bi riS oT roT roS iD
6 6 4 5 62 6 2 32.0 
  +----------------- Cristariina     
  |                                  
--+  +-------------- Anodonta_cygnea 
  |  |                               
  +--+  +----------- Anodonta_anatina
     |  |                            
     +--+        +-- Pyganodon **       
        |  +-----+                   
        |  |     +-- Utterbackia **     
        +--+                         
           |  +----- Prolasmidonta **   
           +--+                      
              |  +-- Platynaias **      
              +--+                   
                 +-- Alasmidonta **

King et al., 1999 [Publ 2699], fig. 4

iT tT bi riS oT roT roS iD
6 6 3 4 0 0 0 32.0 
  +----- Lasmigona **    
  |                   
--+  +-- Prolasmidonta **
  |  |                
  +--+-- Strophitus **   
     |                
     +-- Platynaias **

Hewitt et al., 2018 [Publ 3758], fig. 5

iT tT bi riS oT roT roS iD
6 6 3 5 3 0 0 31.0 
  +-------- Pyganodon **           
  |                             
  |     +-- Platynaias **          
--+-----+                       
  |     +-- Strophitus **          
  |                             
  |  +----- Lasmigona_costata **   
  +--+                          
     |  +-- Arcidens **            
     +--+                       
        +-- Lasmigona_complanata **

Huang et al., 2002 [Publ 2657], fig. 2

iT tT bi riS oT roT roS iD
5 14 5 5 29 2 1 44.0 
  +-------- Strophitus **   
  |                      
  +-------- Schistodesmus
  |                      
  +-------- Lepidodesma  
  |                      
  +-------- Acuticosta   
  |                      
  +-------- Cristariina  
--+                      
  +-------- Lanceolaria  
  |                      
  |     +-- Nodularia    
  +-----+                
  |     +-- Tchangsinaia 
  |                      
  |  +----- Utterbackia **  
  |  |                   
  +--+----- Utterbackiana **
     |                   
     |  +-- Alasmidonta **  
     +--+                
        +-- Lasmigona **

Lydeard et al., 1996 [Publ 2656], fig. 4

iT tT bi riS oT roT roS iD
5 6 4 5 23 1 1 38.0 
     +----- Utterbackia **  
  +--+                   
  |  |  +-- Utterbackiana **
  |  +--+                
--+     +-- Anodontina   
  |                      
  |  +----- Strophitus **   
  +--+                   
     |  +-- Lasmigona **    
     +--+                
        +-- Alasmidonta **

Guerra et al., 2019 [Publ 4168], fig. 1

iT tT bi riS oT roT roS iD
5 5 3 3 36 2 2 31.0 
  +----------- Cristariina
--+                       
  |  +-------- Anodontina 
  +--+                    
     |  +----- Platynaias ** 
     +--+                 
        |  +-- Pyganodon **  
        +--+              
           +-- Utterbackia **

Guerra et al., 2017 [Publ 3772], fig. 2a

iT tT bi riS oT roT roS iD
5 5 3 3 20 3 2 31.0 
  +-------------- Sinanodonta
--+                          
  |  +----------- Cristaria  
  +--+                       
     |  +-------- Anodontina 
     +--+                    
        |  +----- Platynaias ** 
        +--+                 
           |  +-- Pyganodon **  
           +--+              
              +-- Utterbackia **

Lopes-Lima et al., 2017 [Publ 3745], fig. 1-2

iT tT bi riS oT roT roS iD
5 5 3 5 75 9 2 35.0 
  +-------------- Cristariina
  |                          
--+  +----------- Anodontina 
  |  |                       
  +--+        +-- Pyganodon **  
     |  +-----+              
     |  |     +-- Simpsonaias **
     +--+                    
        |  +----- Strophitus ** 
        +--+                 
           |  +-- Alasmidonta **
           +--+              
              +-- Lasmigona **

Amaro et al., 2017 [Publ 3791], fig. 2

iT tT bi riS oT roT roS iD
5 5 3 4 11 0 0 31.0 
     +-- Lasmigona **  
  +--+              
  |  +-- Platynaias ** 
--+                 
  |  +-- Pyganodon **  
  +--+              
     +-- Utterbackia **

Bolotov et al., 2017 [Publ 3761], fig. S1

iT tT bi riS oT roT roS iD
5 5 3 5 134 10 2 37.0 
  +----------------- Cristariina         
  |                                      
--+  +-------------- Anodonta_anatina    
  |  |                                   
  |  |        +----- Pseudanodonta       
  +--+  +-----+                          
     |  |     |  +-- Anodonta_cygnea     
     |  |     +--+                       
     |  |        +-- Anodonta_nuttalliana
     +--+                                
        |        +-- Lasmigona **           
        |  +-----+                       
        |  |     +-- Alasmidonta **         
        +--+                             
           |  +----- Simpsonaias **         
           +--+                          
              |  +-- Pyganodon **           
              +--+                       
                 +-- Strophitus **

Bolotov et al., 2017 [Publ 3761], fig. S2

iT tT bi riS oT roT roS iD
5 5 1 5 134 10 2 36.0 
  +-------- Anodonta_anatina    
  |                             
  +-------- Pseudanodonta       
  |                             
  |     +-- Anodonta_cygnea     
--+-----+                       
  |     +-- Anodonta_nuttalliana
  |                             
  |  +----- Pyganodon **           
  |  |                          
  |  +----- Simpsonaias **         
  +--+                          
     +----- Strophitus **          
     |                          
     |  +-- Alasmidonta **         
     +--+                       
        +-- Lasmigona **

Jeratthitikul et al., 2022 [Publ 4260], fig. S1_1

iT tT bi riS oT roT roS iD
5 5 3 5 71 9 2 35.0 
  +-------------- Cristariina
  |                          
--+  +----------- Anodontina 
  |  |                       
  +--+        +-- Pyganodon **  
     |  +-----+              
     |  |     +-- Simpsonaias **
     +--+                    
        |  +----- Strophitus ** 
        +--+                 
           |  +-- Alasmidonta **
           +--+              
              +-- Lasmigona **

Wu et al., 2018 [Publ 3836], fig. 2

iT tT bi riS oT roT roS iD
5 5 2 4 48 8 2 36.0 
  +-------------- Cristariina     
  |                               
--+  +----------- Anodonta_cygnea 
  |  |                            
  +--+  +-------- Anodonta_anatina
     |  |                         
     +--+  +----- Strophitus **      
        |  |                      
        +--+  +-- Platynaias **      
           |  |                   
           +--+-- Alasmidonta **     
              |                   
              +-- Pyganodon **

Guerra et al., 2017 [Publ 3772], fig. 2b

iT tT bi riS oT roT roS iD
5 5 3 3 20 3 2 31.0 
  +----------- Cristariina
--+                       
  |  +-------- Anodontina 
  +--+                    
     |  +----- Platynaias ** 
     +--+                 
        |  +-- Pyganodon **  
        +--+              
           +-- Utterbackia **

Huang et al., 2002 [Publ 2657], fig. 1

iT tT bi riS oT roT roS iD
5 5 3 5 29 2 1 37.0 
  +-------------- Cristariina  
--+                            
  |  +----------- Utterbackiana **
  +--+                         
     |  +-------- Utterbackia **  
     +--+                      
        |  +----- Strophitus **   
        +--+                   
           |  +-- Alasmidonta **  
           +--+                
              +-- Lasmigona **

Jeratthitikul et al., 2022 [Publ 4260], fig. S2_2

iT tT bi riS oT roT roS iD
5 5 3 5 71 9 2 37.0 
  +-------------- Cristariina
  |                          
--+  +----------- Anodontina 
  |  |                       
  +--+        +-- Alasmidonta **
     |  +-----+              
     |  |     +-- Lasmigona **  
     +--+                    
        |  +----- Pyganodon **  
        +--+                 
           |  +-- Strophitus ** 
           +--+              
              +-- Simpsonaias **

Guerra et al., 2019 [Publ 4168], fig. 2

iT tT bi riS oT roT roS iD
5 5 3 3 36 2 2 31.0 
  +----------- Cristariina
--+                       
  |  +-------- Anodontina 
  +--+                    
     |  +----- Platynaias ** 
     +--+                 
        |  +-- Pyganodon **  
        +--+              
           +-- Utterbackia **

Bolotov et al., 2017 [Publ 3764], fig. S1

iT tT bi riS oT roT roS iD
5 17 0 5 120 10 2 46.0 
  +-- Alasmidonta **          
  |                        
  +-- Anodonta_anatina     
  |                        
  +-- Anodonta_cygnea      
  |                        
  +-- Anodonta_nuttalliana 
  |                        
  +-- Lasmigona **            
  |                        
  +-- Pseudanodonta        
  |                        
  +-- Pyganodon **            
  |                        
  +-- Simpsonaias **          
  |                        
  +-- Strophitus **           
--+                        
  +-- Sinanodonta_lauta    
  |                        
  +-- Cristaria_plicata    
  |                        
  +-- Cristaria_radiata    
  |                        
  +-- Pletholophus         
  |                        
  +-- Sinanodonta_lucida   
  |                        
  +-- Sinanodonta_jourdyi  
  |                        
  +-- Lanceolaria_gladiola 
  |                        
  +-- Lanceolaria_grayii__1
  |                        
  +-- Lanceolaria_grayii__2

Bolotov et al., 2018 [Publ 3826], fig. 2

iT tT bi riS oT roT roS iD
5 5 3 5 78 5 2 35.0 
  +----------------- Cristariina
--+                             
  |  +-------------- Anodontina 
  +--+                          
     |  +----------- Pyganodon **  
     +--+                       
        |  +-------- Simpsonaias **
        +--+                    
           |  +----- Strophitus ** 
           +--+                 
              |  +-- Alasmidonta **
              +--+              
                 +-- Lasmigona **

Aunins et al., 2018 [Publ 3863], fig. 1

iT tT bi riS oT roT roS iD
5 5 3 4 35 6 2 31.0 
  +----------- Cristariina  
  |                         
--+  +-------- Anodontina   
  |  |                      
  +--+     +-- Prolasmidonta **
     |  +--+                
     |  |  +-- Alasmidonta **  
     +--+                   
        |  +-- Pyganodon **    
        +--+                
           +-- Utterbackia **

Graf & Ó Foighil, 2000 [Publ 2648], fig. 2A

iT tT bi riS oT roT roS iD
5 5 3 4 25 1 1 37.0 
  +-------- Anodontina 
  |                    
--+     +-- Alasmidonta **
  |  +--+              
  |  |  +-- Lasmigona **  
  +--+                 
     |  +-- Strophitus ** 
     +--+              
        +-- Pyganodon **

Graf & Ó Foighil, 2000 [Publ 2648], fig. 2B

iT tT bi riS oT roT roS iD
5 5 3 4 25 1 1 37.0 
  +-------- Anodontina 
  |                    
--+     +-- Alasmidonta **
  |  +--+              
  |  |  +-- Lasmigona **  
  +--+                 
     |  +-- Strophitus ** 
     +--+              
        +-- Pyganodon **

Wu et al., 2022 [Publ 4238], fig. 2d

iT tT bi riS oT roT roS iD
4 4 2 3 34 6 2 32.0 
  +----------- Cristariina
--+                       
  |  +-------- Anodontina 
  +--+                    
     |  +----- Platynaias ** 
     +--+                 
        |  +-- Pyganodon **  
        +--+              
           +-- Utterbackia **

Amaro et al., 2017 [Publ 3791], fig. 3A

iT tT bi riS oT roT roS iD
4 4 2 3 8 0 0 32.0 
  +----- Platynaias ** 
--+                 
  |  +-- Pyganodon **  
  +--+              
     +-- Utterbackia **

Campbell et al., 2005 [Publ 2660], fig. 1

iT tT bi riS oT roT roS iD
4 5 3 4 98 2 1 35.0 
  +----------- Anodonta_oregonensis
  |                                
--+        +-- Anodonta_cygnea     
  |  +-----+                       
  |  |     +-- Alasminota **          
  +--+                             
     |  +----- Pyganodon **           
     +--+                          
        |  +-- Lasmigona **           
        +--+                       
           +-- Pseudodontoideus **

Dai et al., 2025 [Publ 4440], fig. 3

iT tT bi riS oT roT roS iD
4 4 2 3 84 6 2 32.0 
  +----------- Cristariina
--+                       
  |  +-------- Anodontina 
  +--+                    
     |  +----- Platynaias ** 
     +--+                 
        |  +-- Pyganodon **  
        +--+              
           +-- Utterbackia **

Cyr et al., 2007 [Publ 2745], fig. 2D

iT tT bi riS oT roT roS iD
4 4 2 1 0 0 0 38.0 
-- (Pyganodon

Cyr et al., 2007 [Publ 2745], fig. 2A

iT tT bi riS oT roT roS iD
4 4 2 1 1 0 0 38.0 
-- (Pyganodon

Fonseca et al., 2016 [Publ 3775], fig. 1

iT tT bi riS oT roT roS iD
4 4 2 3 27 5 2 32.0 
  +----------- Cristariina
--+                       
  |  +-------- Anodontina 
  +--+                    
     |  +----- Platynaias ** 
     +--+                 
        |  +-- Pyganodon **  
        +--+              
           +-- Utterbackia **

Huang et al., 2013 [Publ 3629], fig. 3

iT tT bi riS oT roT roS iD
4 4 2 3 12 2 1 32.0 
  +-------- Cristariina
--+                    
  |  +----- Platynaias ** 
  +--+                 
     |  +-- Pyganodon **  
     +--+              
        +-- Utterbackia **

Graf & Cummings, 2006 [Publ 1968], fig. 2

iT tT bi riS oT roT roS iD
4 4 2 4 43 0 0 36.0 
  +-------- Lasmigona **  
--+                    
  |  +----- Alasmidonta **
  +--+                 
     |  +-- Pyganodon **  
     +--+              
        +-- Strophitus **

Riccardi et al., 2020 [Publ 4089], fig. 2

iT tT bi riS oT roT roS iD
4 4 2 3 15 9 2 32.0 
  +----------------- Cristariina         
  |                                      
--+  +-------------- Anodonta_anatina    
  |  |                                   
  |  |        +----- Pseudanodonta       
  +--+  +-----+                          
     |  |     |  +-- Anodonta_exulcerata 
     |  |     +--+                       
     +--+        +-- Anodonta_cygnea     
        |                                
        |  +-------- Anodonta_nuttalliana
        +--+                             
           |  +----- Platynaias **          
           +--+                          
              |  +-- Pyganodon **           
              +--+                       
                 +-- Utterbackia **

Cyr et al., 2007 [Publ 2745], fig. 2B

iT tT bi riS oT roT roS iD
4 4 3 1 1 1 1 38.0 
  +-- Anodontina
--+             
  +-- Pyganodon **

Wu et al., 2020 [Publ 4121], fig. 3

iT tT bi riS oT roT roS iD
4 4 2 3 45 5 2 32.0 
  +----------- Cristariina
--+                       
  |  +-------- Anodontina 
  +--+                    
     |  +----- Platynaias ** 
     +--+                 
        |  +-- Pyganodon **  
        +--+              
           +-- Utterbackia **

Froufe et al., 2019 [Publ 3936], fig. 3

iT tT bi riS oT roT roS iD
4 4 2 3 40 5 2 32.0 
  +----------- Cristariina
--+                       
  |  +-------- Anodontina 
  +--+                    
     |  +----- Lasmigona **  
     +--+                 
        |  +-- Pyganodon **  
        +--+              
           +-- Utterbackia **

Cha et al., 2018 [Publ 3916], fig. 1

iT tT bi riS oT roT roS iD
4 4 2 3 34 5 2 32.0 
  +----------- Cristariina
--+                       
  |  +-------- Anodontina 
  +--+                    
     |  +----- Platynaias ** 
     +--+                 
        |  +-- Pyganodon **  
        +--+              
           +-- Utterbackia **

Soroka & Burzynski, 2017 [Publ 3837], fig. 1

iT tT bi riS oT roT roS iD
4 4 2 3 19 6 2 32.0 
  +----------- Cristariina
--+                       
  |  +-------- Anodontina 
  +--+                    
     |  +----- Platynaias ** 
     +--+                 
        |  +-- Pyganodon **  
        +--+              
           +-- Utterbackia **

Jadhav & Jamkhedkar, 2009 [Publ 3697], fig. 11

iT tT bi riS oT roT roS iD
4 4 1 4 21 2 1 35.0 
  +-------- Anodontina 
  |                    
--+  +----- Lasmigona **  
  |  |                 
  +--+  +-- Strophitus ** 
     |  |              
     +--+-- Alasmidonta **
        |              
        +-- Pyganodon **

Froufe et al., 2019 [Publ 3936], fig. 6

iT tT bi riS oT roT roS iD
4 4 2 3 40 5 2 32.0 
  +----------- Cristariina
--+                       
  |  +-------- Anodontina 
  +--+                    
     |  +----- Lasmigona **  
     +--+                 
        |  +-- Pyganodon **  
        +--+              
           +-- Utterbackia **

Cyr et al., 2007 [Publ 2745], fig. 2C

iT tT bi riS oT roT roS iD
4 4 3 1 1 1 1 38.0 
  +-- Anodontina
--+             
  +-- Pyganodon **

Wu et al., 2019 [Publ 3864], fig. 3

iT tT bi riS oT roT roS iD
4 4 2 3 42 5 2 32.0 
  +----------- Cristariina
--+                       
  |  +-------- Anodontina 
  +--+                    
     |  +----- Platynaias ** 
     +--+                 
        |  +-- Pyganodon **  
        +--+              
           +-- Utterbackia **

Zhou et al., 2016 [Publ 3710], fig. 1

iT tT bi riS oT roT roS iD
4 4 2 3 16 2 1 32.0 
  +----------- Sinanodonta
--+                       
  |  +-------- Cristaria  
  +--+                    
     |  +----- Platynaias ** 
     +--+                 
        |  +-- Pyganodon **  
        +--+              
           +-- Utterbackia **

Wu et al., 2022 [Publ 4238], fig. 3

iT tT bi riS oT roT roS iD
4 4 2 3 69 6 2 32.0 
  +----------- Cristariina
--+                       
  |  +-------- Anodontina 
  +--+                    
     |  +----- Platynaias ** 
     +--+                 
        |  +-- Pyganodon **  
        +--+              
           +-- Utterbackia **

Fukata & Iigo, 2020 [Publ 4106], fig. 1

iT tT bi riS oT roT roS iD
4 4 2 4 49 6 2 32.0 
  +----------- Cristariina  
  |                         
--+  +-------- Anodontina   
  |  |                      
  +--+     +-- Pyganodon **    
     |  +--+                
     |  |  +-- Utterbackia **  
     +--+                   
        |  +-- Alasmidonta **  
        +--+                
           +-- Prolasmidonta **

Huang et al., 2019 [Publ 3843], fig. 3

iT tT bi riS oT roT roS iD
4 4 2 3 52 6 2 32.0 
  +-------------- Cristariina     
--+                               
  |  +----------- Anodonta_cygnea 
  +--+                            
     |  +-------- Anodonta_anatina
     +--+                         
        |  +----- Lasmigona **       
        +--+                      
           |  +-- Pyganodon **       
           +--+                   
              +-- Utterbackia **

Campbell et al., 2005 [Publ 2660], fig. 2

iT tT bi riS oT roT roS iD
4 6 3 4 98 2 1 35.0 
  +----- Pyganodon **       
  |                      
  |  +-- Lasmigona **       
--+--+                   
  |  +-- Pseudodontoideus **
  |                      
  |  +-- Alasminota **      
  +--+                   
     +-- Anodontina

Hoeh et al., 2009 [Publ 2716], fig. 4

iT tT bi riS oT roT roS iD
4 4 2 4 42 0 0 36.0 
  +-------- Lasmigona **  
--+                    
  |  +----- Alasmidonta **
  +--+                 
     |  +-- Strophitus ** 
     +--+              
        +-- Pyganodon **

Hoeh et al., 2009 [Publ 2716], fig. 3

iT tT bi riS oT roT roS iD
4 4 2 4 43 0 0 36.0 
  +-------- Lasmigona **  
--+                    
  |  +----- Alasmidonta **
  +--+                 
     |  +-- Strophitus ** 
     +--+              
        +-- Pyganodon **

Hoeh et al., 2009 [Publ 2716], fig. 5

iT tT bi riS oT roT roS iD
4 4 2 4 42 0 0 36.0 
  +-------- Lasmigona **  
--+                    
  |  +----- Alasmidonta **
  +--+                 
     |  +-- Strophitus ** 
     +--+              
        +-- Pyganodon **

Zieritz et al., 2021 [Publ 4231], fig. 3-F_5

iT tT bi riS oT roT roS iD
4 4 2 3 44 5 2 32.0 
  +----------- Cristariina
--+                       
  |  +-------- Anodontina 
  +--+                    
     |  +----- Platynaias ** 
     +--+                 
        |  +-- Pyganodon **  
        +--+              
           +-- Utterbackia **

Amaro et al., 2017 [Publ 3791], fig. 3B

iT tT bi riS oT roT roS iD
4 4 2 3 8 0 0 32.0 
  +----- Platynaias ** 
--+                 
  |  +-- Pyganodon **  
  +--+              
     +-- Utterbackia **

Huang et al., 2018 [Publ 3828], fig. 6

iT tT bi riS oT roT roS iD
4 4 2 3 38 5 2 32.0 
  +----------- Cristariina
--+                       
  |  +-------- Anodontina 
  +--+                    
     |  +----- Platynaias ** 
     +--+                 
        |  +-- Pyganodon **  
        +--+              
           +-- Utterbackia **

Lopes-Lima et al., 2017 [Publ 3792], fig. 2

iT tT bi riS oT roT roS iD
4 4 2 3 28 5 2 32.0 
  +----------- Cristariina
--+                       
  |  +-------- Anodontina 
  +--+                    
     |  +----- Platynaias ** 
     +--+                 
        |  +-- Pyganodon **  
        +--+              
           +-- Utterbackia **

Dai et al., 2024 [Publ 4358], fig. 4

iT tT bi riS oT roT roS iD
3 3 1 3 49 4 2 33.0 
  +----------- Cristariina
--+                       
  |  +-------- Anodontina 
  +--+                    
     |  +----- Platynaias ** 
     +--+                 
        |  +-- Pyganodon **  
        +--+              
           +-- Utterbackia **

Wu et al., 2016 [Publ 3711], fig. 1

iT tT bi riS oT roT roS iD
3 3 1 2 18 4 2 33.0 
  +-------------- Sinanodonta
--+                          
  |  +----------- Anemina    
  +--+                       
     |  +-------- Cristaria  
     +--+                    
        |  +----- Anodontina 
        +--+                 
           |  +-- Pyganodon **  
           +--+              
              +-- Utterbackia **

Wu et al., 2018 [Publ 4143], fig. 2

iT tT bi riS oT roT roS iD
3 3 1 2 19 4 2 33.0 
  +-------------- Sinanodonta
--+                          
  |  +----------- Anemina    
  +--+                       
     |  +-------- Cristaria  
     +--+                    
        |  +----- Anodontina 
        +--+                 
           |  +-- Pyganodon **  
           +--+              
              +-- Utterbackia **

Pfeiffer & Graf, 2013 [Publ 3531], fig. 2

iT tT bi riS oT roT roS iD
3 3 1 3 28 2 1 35.0 
  +-------- Cristariina
--+                    
  |  +----- Strophitus ** 
  +--+                 
     |  +-- Platynaias ** 
     +--+              
        +-- Pyganodon **

Amaro et al., 2017 [Publ 3791], fig. 2-M

iT tT bi riS oT roT roS iD
3 4 1 3 7 1 1 36.0 
  +----- Pyganodon **  
  |                 
--+  +-- Lasmigona **  
  |  |              
  +--+-- Cristariina
     |              
     +-- Utterbackia **

Rosenberg et al., 1994 [Publ 2719], fig. 3

iT tT bi riS oT roT roS iD
3 10 0 2 16 0 0 42.0 
  +-- Pyganodon_cataracta **
  |                      
  +-- Pyganodon_grandis **  
  |                      
  +-- Utterbackia **        
  |                      
  +-- Lampsilis_teres    
  |                      
  +-- Lampsilis_straminea
--+                      
  +-- Megalonaias        
  |                      
  +-- Obliquaria         
  |                      
  +-- Theliderma         
  |                      
  +-- Quadrula           
  |                      
  +-- Uniomerus

Liu et al., 2023 [Publ 4310], fig. 4

iT tT bi riS oT roT roS iD
3 3 1 3 21 1 1 35.0 
  +-------- Anodontina
--+                   
  |  +----- Lasmigona ** 
  +--+                
     |  +-- Strophitus **
     +--+             
        +-- Pyganodon **

Liu et al., 2023 [Publ 4310], fig. 5

iT tT bi riS oT roT roS iD
3 4 2 3 21 1 1 36.0 
  +-------- Lasmigona ** 
--+                   
  |  +----- Anodontina
  +--+                
     |  +-- Pyganodon ** 
     +--+             
        +-- Strophitus **

Wu et al., 2021 [Publ 4366], fig. 2_I

iT tT bi riS oT roT roS iD
3 6 4 3 88 7 2 38.0 
  +-------------- Cristariina  
--+                            
  |  +----------- Lasmigona **    
  +--+                         
     |  +-------- Anodonta     
     +--+                      
        |  +----- Pseudanodonta
        +--+                   
           |  +-- Pyganodon **    
           +--+                
              +-- Strophitus **

Rosenberg et al., 1997 [Publ 2720], fig. 2A

iT tT bi riS oT roT roS iD
3 17 0 2 19 1 1 48.0 
  +----- Anodontina         
  |                         
  |  +-- Gonidea            
  |  |                      
  |  +-- Pyganodon_grandis **  
  |  |                      
  |  +-- Utterbackia **        
  |  |                      
  |  +-- Pyganodon_cataracta **
--+  |                      
  |  +-- Lampsilis_straminea
  |  |                      
  |  +-- Lampsilis_teres    
  |  |                      
  |  +-- Megalonaias        
  |  |                      
  |  +-- Obliquaria         
  |  |                      
  +--+-- Theliderma         
     |                      
     +-- Quadrula           
     |                      
     +-- Uniomerus          
     |                      
     +-- Amblema            
     |                      
     +-- Elliptio           
     |                      
     +-- Fusconaia          
     |                      
     +-- Plectomerus        
     |                      
     +-- Pleurobema         
     |                      
     +-- Unio

Moyer & Díaz-Ferguson, 2012 [Publ 3375], fig. 3

iT tT bi riS oT roT roS iD
3 3 1 2 3 0 0 33.0 
  +-- Lasmigona ** 
--+             
  +-- Platynaias **

Wu et al., 2021 [Publ 4366], fig. 6

iT tT bi riS oT roT roS iD
3 5 3 3 88 7 2 37.0 
  +-------------- Cristariina     
  |                               
--+  +----------- Anodonta_cygnea 
  |  |                            
  +--+        +-- Lasmigona **       
     |  +-----+                   
     |  |     +-- Anodonta_anatina
     +--+                         
        |  +----- Pseudanodonta   
        +--+                      
           |  +-- Pyganodon **       
           +--+                   
              +-- Strophitus **

Rosenberg et al., 1997 [Publ 2720], fig. 2B

iT tT bi riS oT roT roS iD
3 17 0 2 19 1 1 48.0 
  +----- Anodontina         
  |                         
  |  +-- Gonidea            
  |  |                      
  |  +-- Pyganodon_grandis **  
  |  |                      
  |  +-- Utterbackia **        
  |  |                      
  |  +-- Pyganodon_cataracta **
--+  |                      
  |  +-- Lampsilis_straminea
  |  |                      
  |  +-- Lampsilis_teres    
  |  |                      
  |  +-- Megalonaias        
  |  |                      
  |  +-- Obliquaria         
  |  |                      
  +--+-- Theliderma         
     |                      
     +-- Quadrula           
     |                      
     +-- Uniomerus          
     |                      
     +-- Amblema            
     |                      
     +-- Elliptio           
     |                      
     +-- Fusconaia          
     |                      
     +-- Plectomerus        
     |                      
     +-- Aculamprotula      
     |                      
     +-- Unio

Wu et al., 2020 [Publ 4147], fig. 3

iT tT bi riS oT roT roS iD
3 3 1 2 26 4 2 33.0 
  +-------- Cristariina
--+                    
  |  +----- Anodontina 
  +--+                 
     |  +-- Pyganodon **  
     +--+              
        +-- Utterbackia **

Wu et al., 2023 [Publ 4296], fig. 4

iT tT bi riS oT roT roS iD
3 3 1 3 20 1 1 35.0 
  +-------- Anodontina
--+                   
  |  +----- Lasmigona ** 
  +--+                
     |  +-- Strophitus **
     +--+             
        +-- Pyganodon **

Dai et al., 2023 [Publ 4285], fig. 5

iT tT bi riS oT roT roS iD
3 3 1 3 50 4 2 33.0 
  +----------- Cristariina
--+                       
  |  +-------- Anodontina 
  +--+                    
     |  +----- Platynaias ** 
     +--+                 
        |  +-- Pyganodon **  
        +--+              
           +-- Utterbackia **

Krebs et al., 2019 [Publ 4092], fig. 4

iT tT bi riS oT roT roS iD
3 3 1 1 0 0 0 37.0 
-- (Pyganodon

Walker et al., 2006 [Publ 2666], fig. 6-M

iT tT bi riS oT roT roS iD
2 2 0 1 12 0 0 -- 
-- (Pyganodon

Wang et al., 2018 [Publ 3865], fig. 2

iT tT bi riS oT roT roS iD
2 2 0 2 29 5 2 -- 
  +-------- Cristariina
--+                    
  |  +----- Anodontina 
  +--+                 
     |  +-- Pyganodon **  
     +--+              
        +-- Platynaias **

Guerra et al., 2019 [Publ 4168], fig. 1-M

iT tT bi riS oT roT roS iD
2 2 0 2 20 2 2 -- 
  +-------- Cristariina
--+                    
  |  +----- Anodontina 
  +--+                 
     |  +-- Utterbackia **
     +--+              
        +-- Pyganodon **

Hoeh et al., 2002 [Publ 2673], fig. 1-F

iT tT bi riS oT roT roS iD
2 2 0 1 7 0 0 -- 
-- (Pyganodon

Wu et al., 2023 [Publ 4339], fig. 5

iT tT bi riS oT roT roS iD
2 2 0 1 17 7 2 -- 
  +----- Cristariina
--+                 
  |  +-- Platynaias ** 
  +--+              
     +-- Anodontina

Walker et al., 2006 [Publ 2666], fig. 3

iT tT bi riS oT roT roS iD
2 2 0 1 13 1 1 -- 
  +-- Anodontina
--+             
  +-- Pyganodon **

Pfeiffer & Graf, 2015 [Publ 3620], fig. 2

iT tT bi riS oT roT roS iD
2 2 0 2 38 0 0 -- 
  +-- Pyganodon **
--+            
  +-- Lasmigona **

Hoeh et al., 2002 [Publ 2673], fig. 1-M

iT tT bi riS oT roT roS iD
2 2 0 1 7 0 0 -- 
-- (Pyganodon

Whelan et al., 2011 [Publ 3380], fig. 1-ML

iT tT bi riS oT roT roS iD
2 2 0 2 34 0 0 -- 
  +-- Pyganodon **
--+            
  +-- Lasmigona **

Burzynski & Soroka, 2018 [Publ 3862], fig. 5a

iT tT bi riS oT roT roS iD
2 2 0 2 18 2 2 -- 
  +----- Cristariina
  |                 
--+----- Anodontina 
  |                 
  |  +-- Utterbackia **
  +--+              
     +-- Pyganodon **

Hoeh et al., 2002 [Publ 2653], fig. 2

iT tT bi riS oT roT roS iD
2 2 0 2 29 1 1 -- 
  +----- Anodontina
--+                
  |  +-- Pyganodon ** 
  +--+             
     +-- Strophitus **

Fonseca et al., 2016 [Publ 3775], fig. 1-M

iT tT bi riS oT roT roS iD
2 2 0 2 7 1 1 -- 
  +----- Anodontina 
--+                 
  |  +-- Utterbackia **
  +--+              
     +-- Pyganodon **

Araujo et al., 2018 [Publ 3779], fig. 1

iT tT bi riS oT roT roS iD
2 2 0 2 16 1 1 -- 
  +----- Anodontina 
--+                 
  |  +-- Pyganodon **  
  +--+              
     +-- Alasmidonta **

Wang et al., 2018 [Publ 3846], fig. 2

iT tT bi riS oT roT roS iD
2 2 0 2 29 5 2 -- 
  +-------- Cristariina
--+                    
  |  +----- Anodontina 
  +--+                 
     |  +-- Pyganodon **  
     +--+              
        +-- Platynaias **

Bogan & Hoeh, 2000 [Publ 2650], fig. 1

iT tT bi riS oT roT roS iD
2 2 0 2 32 1 1 -- 
  +----- Anodontina
--+                
  |  +-- Strophitus **
  +--+             
     +-- Pyganodon **

Dai et al., 2025 [Publ 4440], fig. 2

iT tT bi riS oT roT roS iD
2 2 0 2 43 8 2 -- 
  +----- Anodontina 
--+                 
  |  +-- Alasmidonta **
  +--+              
     +-- Lasmigona **

Walker et al., 2006 [Publ 2666], fig. 2

iT tT bi riS oT roT roS iD
2 2 0 1 13 1 1 -- 
  +-- Anodontina
--+             
  +-- Pyganodon **

Walker et al., 2006 [Publ 2666], fig. 6

iT tT bi riS oT roT roS iD
2 2 0 1 34 0 0 -- 
-- (Pyganodon

Zieritz et al., 2021 [Publ 4231], fig. 3-M_4

iT tT bi riS oT roT roS iD
2 2 0 2 21 2 2 -- 
  +-------- Cristariina
--+                    
  |  +----- Anodontina 
  +--+                 
     |  +-- Utterbackia **
     +--+              
        +-- Pyganodon **

Roe & Hoeh, 2003 [Publ 2655], fig. 4f2

iT tT bi riS oT roT roS iD
2 3 0 2 29 1 1 35.0 
  +----- Pyganodon ** 
--+                
  |  +-- Anodontina
  +--+             
     +-- Strophitus **

Guerra et al., 2017 [Publ 3772], fig. 2a-M

iT tT bi riS oT roT roS iD
2 2 0 2 6 0 0 -- 
  +-- Utterbackia **
--+              
  +-- Pyganodon **

Wen et al., 2017 [Publ 3795], fig. 3-M

iT tT bi riS oT roT roS iD
2 2 0 2 4 1 1 -- 
  +----- Anodontina 
--+                 
  |  +-- Utterbackia **
  +--+              
     +-- Pyganodon **

Curole & Kocher, 2005 [Publ 2682], fig. 1-M

iT tT bi riS oT roT roS iD
2 2 0 2 10 0 0 -- 
  +-- Utterbackiana **
--+                
  +-- Pyganodon **

Krebs, 2004 [Publ 2692], fig. 2-M

iT tT bi riS oT roT roS iD
2 2 0 2 2 0 0 -- 
  +-- Lasmigona **
--+            
  +-- Pyganodon **

Manendo et al., 2008 [Publ 2669], fig. 1

iT tT bi riS oT roT roS iD
2 2 0 2 11 3 1 -- 
  +-------- Anodonta     
--+                      
  |  +----- Pseudanodonta
  +--+                   
     |  +-- Platynaias **   
     +--+                
        +-- Pyganodon **

Froufe et al., 2019 [Publ 3936], fig. 4

iT tT bi riS oT roT roS iD
2 2 0 2 17 2 2 -- 
  +-------- Cristariina
--+                    
  |  +----- Anodontina 
  +--+                 
     |  +-- Utterbackia **
     +--+              
        +-- Pyganodon **

Whelan et al., 2011 [Publ 3380], fig. 1-MP

iT tT bi riS oT roT roS iD
2 2 0 2 34 0 0 -- 
  +-- Pyganodon **
--+            
  +-- Lasmigona **

Chen et al., 2023 [Publ 4302], fig. 2

iT tT bi riS oT roT roS iD
2 2 0 2 38 8 2 -- 
  +-------- Cristariina
--+                    
  |  +----- Anodontina 
  +--+                 
     |  +-- Alasmidonta **
     +--+              
        +-- Lasmigona **

Walker et al., 2006 [Publ 2666], fig. 4

iT tT bi riS oT roT roS iD
2 2 0 1 13 1 1 -- 
  +-- Anodontina
--+             
  +-- Pyganodon **

Dai et al., 2025 [Publ 4440], fig. 1

iT tT bi riS oT roT roS iD
2 2 0 2 43 8 2 -- 
  +----- Anodontina 
--+                 
  |  +-- Alasmidonta **
  +--+              
     +-- Lasmigona **

Chase et al., 2018 [Publ 3861], fig. 1

iT tT bi riS oT roT roS iD
2 2 0 2 9 1 1 -- 
  +----- Anodontina 
--+                 
  |  +-- Utterbackia **
  +--+              
     +-- Pyganodon **

Guerra et al., 2017 [Publ 3772], fig. 2b-M

iT tT bi riS oT roT roS iD
2 2 0 2 6 0 0 -- 
  +-- Utterbackia **
--+              
  +-- Pyganodon **

Huang et al., 2013 [Publ 3629], fig. 3-M

iT tT bi riS oT roT roS iD
2 2 0 2 4 0 0 -- 
  +-- Pyganodon **  
--+              
  +-- Utterbackia **

Guerra et al., 2019 [Publ 4168], fig. 2-M

iT tT bi riS oT roT roS iD
2 2 0 2 20 2 2 -- 
     +-- Utterbackia **
  +--+              
  |  +-- Pyganodon **  
--+                 
  |  +-- Anodontina 
  +--+              
     +-- Cristariina

Graf, 2002 [Publ 2651], fig. 2

iT tT bi riS oT roT roS iD
2 2 0 2 16 0 0 -- 
  +-- Alasmidonta **
--+              
  +-- Pyganodon **

Burzynski et al., 2017 [Publ 3774], fig. 1-bottom

iT tT bi riS oT roT roS iD
2 2 0 2 11 1 1 -- 
  +----- Anodontina 
--+                 
  |  +-- Utterbackia **
  +--+              
     +-- Pyganodon **

Burzynski & Soroka, 2018 [Publ 3862], fig. 3

iT tT bi riS oT roT roS iD
2 2 0 2 2 2 2 -- 
  +-------- Cristariina
--+                    
  |  +----- Anodontina 
  +--+                 
     |  +-- Utterbackia **
     +--+              
        +-- Pyganodon **

Amaro et al., 2017 [Publ 3791], fig. 3B-M

iT tT bi riS oT roT roS iD
2 2 0 2 5 0 0 -- 
  +-- Utterbackia **
--+              
  +-- Pyganodon **

Huang et al., 2018 [Publ 3828], fig. 6-M

iT tT bi riS oT roT roS iD
2 2 0 2 15 2 2 -- 
  +-------- Cristariina
--+                    
  |  +----- Anodontina 
  +--+                 
     |  +-- Pyganodon **  
     +--+              
        +-- Utterbackia **

Krebs, 2004 [Publ 2692], fig. 2

iT tT bi riS oT roT roS iD
2 2 0 2 3 0 0 -- 
  +-- Lasmigona **
--+            
  +-- Pyganodon **

Smith, 2021 [Publ 4331], fig. 1

iT tT bi riS oT roT roS iD
2 2 0 2 18 1 1 -- 
  +----- Cristariina
--+                 
  |  +-- Pyganodon **  
  +--+              
     +-- Utterbackia **

Froufe et al., 2019 [Publ 3936], fig. 3-M

iT tT bi riS oT roT roS iD
2 2 0 2 17 2 2 -- 
  +-------- Cristariina
--+                    
  |  +----- Anodontina 
  +--+                 
     |  +-- Utterbackia **
     +--+              
        +-- Pyganodon **

Soroka & Burzynski, 2018 [Publ 3762], fig. 3

iT tT bi riS oT roT roS iD
2 2 0 2 5 1 1 -- 
  +----- Anodontina 
--+                 
  |  +-- Pyganodon **  
  +--+              
     +-- Utterbackia **

Chase et al., 2018 [Publ 3861], fig. 1-M

iT tT bi riS oT roT roS iD
2 2 0 2 9 1 1 -- 
  +----- Anodontina 
--+                 
  |  +-- Utterbackia **
  +--+              
     +-- Pyganodon **

Feng et al., 2016 [Publ 3800], fig. 2

iT tT bi riS oT roT roS iD
2 12 0 2 12 4 2 43.0 
              +-- Anodontina  
  +-----------+               
  |           +-- Utterbackia ** 
  |                           
--+        +----- Tchangsinaia
  |  +-----+                  
  |  |     |  +-- Pyganodon **   
  |  |     +--+               
  +--+        +-- Cristariina 
     |                        
     |  +-------- Sinosolenaia
     +--+                     
        |  +----- Quadrulina  
        +--+                  
           |  +-- Potamilus   
           +--+               
              +-- Lampsilis

Hoeh et al., 2001 [Publ 2652], fig. 14f4

iT tT bi riS oT roT roS iD
2 2 0 2 29 1 1 -- 
  +----- Anodontina
--+                
  |  +-- Strophitus **
  +--+             
     +-- Pyganodon **

Froufe et al., 2015 [Publ 3664], fig. 1

iT tT bi riS oT roT roS iD
2 2 0 2 10 1 1 -- 
  +----- Anodontina 
--+                 
  |  +-- Utterbackia **
  +--+              
     +-- Pyganodon **

Burzynski et al., 2017 [Publ 3774], fig. 1-top

iT tT bi riS oT roT roS iD
2 2 0 2 11 1 1 -- 
  +----- Anodontina 
--+                 
  |  +-- Utterbackia **
  +--+              
     +-- Pyganodon **

Froufe et al., 2015 [Publ 3664], fig. 1-M

iT tT bi riS oT roT roS iD
2 2 0 2 6 1 1 -- 
  +----- Anodontina 
--+                 
  |  +-- Utterbackia **
  +--+              
     +-- Pyganodon **

Chen et al., 2023 [Publ 4302], fig. 1

iT tT bi riS oT roT roS iD
2 2 0 2 38 8 2 -- 
  +----- Anodontina 
--+                 
  |  +-- Alasmidonta **
  +--+              
     +-- Lasmigona **

Hoeh et al., 2001 [Publ 2652], fig. 14f2

iT tT bi riS oT roT roS iD
2 3 0 2 29 1 1 35.0 
  +----- Pyganodon ** 
--+                
  |  +-- Anodontina
  +--+             
     +-- Strophitus **

Soroka & Burzynski, 2018 [Publ 3762], fig. 3-M

iT tT bi riS oT roT roS iD
2 2 0 2 5 1 1 -- 
  +----- Anodontina 
--+                 
  |  +-- Pyganodon **  
  +--+              
     +-- Utterbackia **

Hoeh et al., 1996 [Publ 2694], fig. 2

iT tT bi riS oT roT roS iD
2 2 0 1 2 0 0 -- 
-- (Pyganodon

Burzynski & Soroka, 2018 [Publ 3862], fig. 5b

iT tT bi riS oT roT roS iD
2 2 0 2 8 2 2 -- 
  +-------- Cristariina
--+                    
  |  +----- Anodontina 
  +--+                 
     |  +-- Utterbackia **
     +--+              
        +-- Pyganodon **

Huang et al., 2019 [Publ 3843], fig. 3-M

iT tT bi riS oT roT roS iD
2 2 0 2 21 2 2 -- 
  +-------- Cristariina
--+                    
  |  +----- Anodontina 
  +--+                 
     |  +-- Pyganodon **  
     +--+              
        +-- Utterbackia **

Amaro et al., 2017 [Publ 3791], fig. 3A-M

iT tT bi riS oT roT roS iD
2 2 0 2 5 0 0 -- 
  +-- Pyganodon **  
--+              
  +-- Utterbackia **

Wu et al., 2022 [Publ 4238], fig. 2e

iT tT bi riS oT roT roS iD
2 2 0 2 15 2 2 -- 
  +-------- Cristariina
--+                    
  |  +----- Anodontina 
  +--+                 
     |  +-- Utterbackia **
     +--+              
        +-- Pyganodon **

Roe & Hoeh, 2003 [Publ 2655], fig. 4f3

iT tT bi riS oT roT roS iD
2 2 0 2 29 1 1 -- 
  +----- Anodontina
--+                
  |  +-- Pyganodon ** 
  +--+             
     +-- Strophitus **

Dai et al., 2024 [Publ 4386], fig. 6

iT tT bi riS oT roT roS iD
2 2 0 2 38 1 1 -- 
  +----- Cristariina
--+                 
  |  +-- Pyganodon **  
  +--+              
     +-- Utterbackia **

Lopes-Lima et al., 2017 [Publ 3792], fig. 2-M

iT tT bi riS oT roT roS iD
2 2 0 2 8 1 1 -- 
  +----- Anodontina 
--+                 
  |  +-- Utterbackia **
  +--+              
     +-- Pyganodon **

Wu et al., 2024 [Publ 4388], fig. 6

iT tT bi riS oT roT roS iD
2 2 0 2 52 5 2 -- 
  +-------- Cristariina
--+                    
  |  +----- Anodontina 
  +--+                 
     |  +-- Alasmidonta **
     +--+              
        +-- Utterbackia **

Curole & Kocher, 2002 [Publ 2712], fig. 1b-M

iT tT bi riS oT roT roS iD
1 1 0 1 3 0 0 -- 
-- Utterbackiana_implicata

Pieri et al., 2018 [Publ 3867], fig. 3

iT tT bi riS oT roT roS iD
1 1 0 1 19 0 0 -- 
-- Arcidens_confragosus

Mulvey et al., 1997 [Publ 3390], fig. 1_3

iT tT bi riS oT roT roS iD
1 1 0 1 4 0 0 -- 
-- Utterbackia_imbecillis

Graf & Ó Foighil, 2000 [Publ 2649], fig. 1_2

iT tT bi riS oT roT roS iD
1 1 0 1 10 0 0 -- 
-- Pyganodon_grandis

Curole & Kocher, 2005 [Publ 2682], fig. 1

iT tT bi riS oT roT roS iD
1 1 0 1 10 0 0 -- 
-- Pyganodon_grandis

Lopes-Lima et al., 2019 [Publ 3901], fig. 4

iT tT bi riS oT roT roS iD
1 1 0 1 25 1 1 -- 
  +-- Anodontina
--+             
  +-- Pyganodon **

Lopes-Lima et al., 2019 [Publ 3901], fig. 3

iT tT bi riS oT roT roS iD
1 1 0 1 28 1 1 -- 
  +-- Anodontina
--+             
  +-- Pyganodon **

Curole & Kocher, 2002 [Publ 2712], fig. 1a-M

iT tT bi riS oT roT roS iD
1 1 0 1 3 0 0 -- 
-- Utterbackiana_implicata

Pfeiffer et al., 2019 [Publ 3937], fig. 6

iT tT bi riS oT roT roS iD
1 1 0 1 56 3 2 -- 
  +----- Cristariina
--+                 
  |  +-- Anodontina 
  +--+              
     +-- Pyganodon **

Plazzi et al., 2011 [Publ 3676], fig. 5

iT tT bi riS oT roT roS iD
1 1 0 1 17 4 2 -- 
  +-- Pyganodon ** 
--+             
  +-- Anodontina

Wei et al., 2016 [Publ 3665], fig. 0

iT tT bi riS oT roT roS iD
1 1 0 1 5 1 1 -- 
-- Pyganodon_grandis

Lopes-Lima et al., 2019 [Publ 3901], fig. 2

iT tT bi riS oT roT roS iD
1 1 0 1 27 1 1 -- 
  +-- Anodontina
--+             
  +-- Pyganodon **

Pavan-Kumar et al., 2022 [Publ 4263], fig. 4

iT tT bi riS oT roT roS iD
1 1 0 1 35 4 2 -- 
  +----- Cristariina
--+                 
  |  +-- Anodontina 
  +--+              
     +-- Pyganodon **

Pavan-Kumar et al., 2022 [Publ 4263], fig. S5

iT tT bi riS oT roT roS iD
1 1 0 1 37 5 2 -- 
  +----- Cristariina
--+                 
  |  +-- Anodontina 
  +--+              
     +-- Pyganodon **

Zanatta & Murphy, 2006 [Publ 2672], fig. 2

iT tT bi riS oT roT roS iD
1 1 0 1 56 0 0 -- 
-- Pyganodon_grandis

Marshall et al., 2014 [Publ 3593], fig. 1

iT tT bi riS oT roT roS iD
1 1 0 1 13 0 0 -- 
-- Pyganodon_grandis

Xue et al., 2016 [Publ 3659], fig. 1

iT tT bi riS oT roT roS iD
1 1 0 1 14 4 1 -- 
     +-- Cristaria  
  +--+              
  |  +-- Platynaias ** 
--+                 
  |  +-- Anemina    
  +--+              
     +-- Sinanodonta

Pfeiffer et al., 2019 [Publ 3937], fig. 4-5

iT tT bi riS oT roT roS iD
1 1 0 1 56 3 2 -- 
  +----- Cristariina
--+                 
  |  +-- Anodontina 
  +--+              
     +-- Pyganodon **

Chase et al., 2018 [Publ 3847], fig. 5

iT tT bi riS oT roT roS iD
1 1 0 1 9 1 1 -- 
  +-- Anodontina 
--+              
  +-- Utterbackia **

Soroka, 2008 [Publ 3695], fig. 1

iT tT bi riS oT roT roS iD
1 1 0 1 3 2 2 -- 
  +----- Cristariina
--+                 
  |  +-- Anodontina 
  +--+              
     +-- Pyganodon **

Soroka, 2008 [Publ 3695], fig. 1-M

iT tT bi riS oT roT roS iD
1 1 0 1 3 2 2 -- 
  +----- Cristariina
--+                 
  |  +-- Anodontina 
  +--+              
     +-- Pyganodon **

Campbell, 2000 [Publ 2724], fig. 1-5

iT tT bi riS oT roT roS iD
1 1 0 1 2 0 0 -- 
-- Utterbackia_imbecillis

Zanatta & Harris, 2013 [Publ 3530], fig. 2

iT tT bi riS oT roT roS iD
1 1 0 1 4 0 0 -- 
-- Pyganodon_grandis

Sheng et al., 2014 [Publ 3704], fig. 2

iT tT bi riS oT roT roS iD
1 1 0 1 7 1 1 -- 
-- Pyganodon_grandis

Inoue et al., 2013 [Publ 3579], fig. 3

iT tT bi riS oT roT roS iD
1 1 0 1 8 0 0 -- 
-- Pyganodon_grandis

Inoue et al., 2013 [Publ 3528], fig. 3

iT tT bi riS oT roT roS iD
1 1 0 1 16 0 0 -- 
-- Pyganodon_grandis

Zanatta & Murphy, 2006 [Publ 2672], fig. 1

iT tT bi riS oT roT roS iD
1 1 0 1 55 0 0 -- 
-- Pyganodon_grandis

Hewitt et al., 2019 [Publ 3962], fig. 2

iT tT bi riS oT roT roS iD
1 1 0 1 9 0 0 -- 
-- Pyganodon_grandis

Inoue et al., 2020 [Publ 3941], fig. 4

iT tT bi riS oT roT roS iD
1 1 0 1 40 0 0 -- 
-- Arcidens_confragosus

Chen et al., 2012 [Publ 3624], fig. 4

iT tT bi riS oT roT roS iD
1 1 0 1 6 1 1 -- 
  +-- Pyganodon **  
--+              
  +-- Cristariina

Inoue et al., 2020 [Publ 3941], fig. 3

iT tT bi riS oT roT roS iD
1 1 0 1 41 0 0 -- 
-- Arcidens_confragosus

Metzger et al., 2018 [Publ 3952], fig. 1C

iT tT bi riS oT roT roS iD
1 1 0 1 2 0 0 -- 
-- Lasmigona_costata

Pfeiffer et al., 2016 [Publ 3660], fig. 2

iT tT bi riS oT roT roS iD
1 1 0 1 37 0 0 -- 
-- Utterbackiana_suborbiculata

Graf, 2000 [Publ 2647], fig. 1

iT tT bi riS oT roT roS iD
1 1 0 1 16 0 0 -- 
-- Pyganodon

Turner et al., 2000 [Publ 2722], fig. 2

iT tT bi riS oT roT roS iD
1 1 0 1 13 0 0 -- 
-- Utterbackiana_couperiana

Page updated: 11:30:39 Tue 03 Jun 2025
 
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