MUSSELp
 
MUSSELpdb
 
 
Taxonomy
 
 
Geography
 
 
 

Page last updated
Mon 06 May 2024

The Freshwater Mussels (Unionoida) of the World (and other less consequential bivalves)

FM(U)otW(aolcb) is the web version of the MUSSEL Project Database. Follow the links to browse the data or use the custom Google search field. Either way, you win!

Cladomics pages are experimental — even more so than the rest of this site!

The purpose of the cladomics pages for each genus and family-group level taxon is to provide statistics and references indicating the extent to which published phylogenetic work supports the classification used here.

The Cladomics Summary provides an overview of the number of studies that have included terminal species representing this taxon as well as an indication of how frequently it has been recovered as monophyletic.

Following the summary, the available published clades representing the taxon are listed.

For more information, visit the Cladomics FARQ (frequently asked rhetorical questions) page. Abbreviations for the various statistics are explained on the FARQ!

Search the database pages:

Bivalvia | Unionoida | Unionoidea | Unionidae

subfamily AMBLEMINAE Rafinesque, 1820: cladomics
     5 tribes

Cladomics Summary

number of species = 413
number of subtaxa (rank: tribe) = 5

analyzed in 280 cladograms in 167 publications
maximum number of terminals in an analysis = 174, mean = 12.25
maximum number of subtaxa (tribe) in an analysis = 5, mean = 2.58

Basic Monophyly

number of cladograms that test basic monophyly (i.e., more than 1 included species) = 241
maximum number of included terminals = 174, mean = 14.07
maximum number of subtaxa (tribe) = 5, mean = 2.83
proportion of cladograms that support basic monophyly = 0.87

Rigorous Monophyly

number of cladograms that test rigorous monophyly = 169
(Rigorous monophyly refers to cladograms with terminal species representing more than one subtaxon (tribe) taxon and at least one rigorous outgroup (subfamily).
maximum number of included terminals = 174, mean = 11.88
maximum number of subtaxa (tribe) = 5, mean = 3.04
proportion of cladograms that support rigorous monophyly = 0.82


Pairwise Cladogram Clustering

The graphs below depict in two dimensions all pair-wise ingroup distances within the cladome. See the Cladmoics FARQ for more information.


PCoA Ordination


NJ tree


Wu et al., 2023 [Publ 4339], fig. 4, S5-nt

iT tT bi riS oT roT roS iD
17 17 15 5 80 69 3 0.0

*** Top Monophyly Cladogram ***

                                         +-- Quadrula_quadrula **           
              +--------------------------+                               
              |                          +-- Uniomerus_tetralasmus **       
              |                                                          
  +-----------+                          +-- Elliptio_complanata **         
  |           |  +-----------------------+                               
  |           |  |                       +-- Pleurobema_oviforme **         
  |           +--+                                                       
  |              |  +----------------------- Amblema_plicata **             
  |              |  |                                                    
  |              +--+  +-------------------- Popenaias_popeii **            
  |                 |  |                                                 
  |                 +--+  +----------------- Toxolasma_parvum **            
  |                    |  |                                              
  |                    |  |           +----- Potamilus_leptodon **          
  |                    +--+  +--------+                                  
  |                       |  |        |  +-- Potamilus_alatus **            
  |                       |  |        +--+                               
  |                       |  |           +-- Potamilus_streckersoni **      
  |                       +--+                                           
  |                          |  +----------- Leaunio_lienosus **            
  |                          |  |                                        
  |                          |  |     +----- Lampsilis_ornata **            
  |                          +--+  +--+                                  
  |                             |  |  |  +-- Lampsilis_cardium **           
  |                             |  |  +--+                               
  |                             +--+     +-- Lampsilis_cariosa **           
  |                                |                                     
  |                                |  +----- Venustaconcha_ellipsiformis ** 
  |                                +--+                                  
  |                                   |  +-- Lampsilis_powellii **          
  |                                   +--+                               
  |                                      +-- Lampsilis_siliquoidea **       
--+                                                                      
  |                                   +----- Chamberlainia_somsakpanhai  
  |                 +-----------------+                                  
  |                 |                 |  +-- Sinohyriopsis_cumingii      
  |                 |                 +--+                               
  |                 |                    +-- Sinohyriopsis_schlegelii    
  |                 |                                                    
  |                 |                    +-- Hyriopsis_bialata           
  |                 |                 +--+                               
  |  +--------------+                 |  +-- Rectidens_sumatrensis       
  |  |              |  +--------------+                                  
  |  |              |  |              |  +-- Lens_contradens             
  |  |              |  |              +--+                               
  |  |              |  |                 +-- Physunio_superbus           
  |  |              |  |                                                 
  |  |              |  |        +----------- Potomida_littoralis         
  |  |              +--+  +-----+                                        
  |  |                 |  |     |  +-------- Pronodularia_japanensis     
  |  |                 |  |     +--+                                     
  |  |                 |  |        |  +----- Lamprotula_leaii            
  |  |                 |  |        +--+                                  
  |  |                 |  |           |  +-- Lamprotula_caveata          
  |  |                 +--+           +--+                               
  |  |                    |              +-- Lamprotula_cornuumlunae     
  |  |                    |                                              
  |  |                    |              +-- Pseudodon_vondembuschianus  
  |  |                    |  +-----------+                               
  |  |                    |  |           +-- Pilsbryoconcha_exilis       
  |  |                    |  |                                           
  |  |                    +--+           +-- Ptychorhynchus_pfisteri     
  +--+                       |  +--------+                               
     |                       |  |        +-- Microcondylaea_bonellii     
     |                       +--+                                        
     |                          |  +-------- Parvasolenaia_rivularis     
     |                          +--+                                     
     |                             |  +----- Sinosolenaia_carinata       
     |                             +--+                                  
     |                                |  +-- Sinosolenaia_iridinea       
     |                                +--+                               
     |                                   +-- Sinosolenaia_oleivora       
     |                                                                   
     |  +----------------------------------- Lamellidens_marginalis      
     |  |                                                                
     |  |                                +-- Lepidodesma_aligera         
     |  |  +-----------------------------+                               
     |  |  |                             +-- Lepidodesma_languilati      
     |  |  |                                                             
     |  |  |                       +-------- Acuticosta_chinensis        
     |  |  |        +--------------+                                     
     |  |  |        |              |  +----- Lanceolaria_gladiola        
     |  |  |        |              +--+                                  
     +--+  |        |                 |  +-- Lanceolaria_grayii          
        |  |        |                 +--+                               
        |  |        |                    +-- Lanceolaria_lanceolata      
        |  |        |                                                    
        |  |        |        +-------------- Cristaria_plicata           
        |  |  +-----+        |                                           
        |  |  |     |  +-----+           +-- Anemina_arcaeformis         
        |  |  |     |  |     |  +--------+                               
        |  |  |     |  |     |  |        +-- Anemina_euscaphys           
        |  |  |     |  |     +--+                                        
        +--+  |     |  |        |  +-------- Beringiana_fukuharai        
           |  |     |  |        +--+                                     
           |  |     |  |           |  +----- Sinanodonta_lucida          
           |  |     |  |           +--+                                  
           |  |     +--+              |  +-- Sinanodonta_tumens          
           |  |        |              +--+                               
           |  |        |                 +-- Sinanodonta_woodiana        
           |  |        |                                                 
           |  |        |  +----------------- Anodonta_nuttalliana        
           |  |        |  |                                              
           |  |        |  +----------------- Pseudanodonta_complanata    
           |  |        |  |                                              
           |  |        +--+  +-------------- Anodonta_cygnea             
           |  |           |  |                                           
           |  |           |  |           +-- Anodonta_exulcerata         
           |  |           |  +-----------+                               
           |  |           |  |           +-- Anodonta_anatina            
           |  |           +--+                                           
           +--+              |        +----- Pyganodon_grandis           
              |              |  +-----+                                  
              |              |  |     |  +-- Utterbackia_imbecillis      
              |              |  |     +--+                               
              |              +--+        +-- Utterbackia_peninsularis    
              |                 |                                        
              |                 |  +-------- Lasmigona_complanata        
              |                 +--+                                     
              |                    |  +----- Prolasmidonta_heterodon     
              |                    +--+                                  
              |                       |  +-- Alasmidonta_varicosa        
              |                       +--+                               
              |                          +-- Platynaias_compressa        
              |                                                          
              |                    +-------- Aculamprotula_tortuosa      
              |                    |                                     
              |  +-----------------+     +-- Aculamprotula_coreana       
              |  |                 |  +--+                               
              |  |                 |  |  +-- Aculamprotula_tientsinensis 
              |  |                 +--+                                  
              |  |                    |  +-- Aculamprotula_polysticta    
              |  |                    +--+                               
              |  |                       +-- Aculamprotula_scripta       
              |  |                                                       
              +--+              +----------- Unio_tumidus                
                 |              |                                        
                 |  +-----------+  +-------- Unio_crassus                
                 |  |           |  |                                     
                 |  |           +--+  +----- Unio_elongatulus            
                 |  |              |  |                                  
                 |  |              +--+  +-- Unio_delphinus              
                 |  |                 |  |                               
                 |  |                 +--+-- Unio_mancus                 
                 +--+                    |                               
                    |                    +-- Unio_pictorum               
                    |                                                    
                    |                    +-- Pseudocuneopsis_capitata    
                    |  +-----------------+                               
                    |  |                 +-- Pseudocuneopsis_sichuanensis
                    |  |                                                 
                    +--+                 +-- Nodularia_breviconcha       
                       |  +--------------+                               
                       |  |              +-- Nodularia_nipponensis       
                       |  |                                              
                       +--+           +----- Tchangsinaia_piscicula      
                          |  +--------+                                  
                          |  |        |  +-- Schistodesmus_lampreyanus[2]
                          |  |        +--+                               
                          +--+           +-- Schistodesmus_spinosus      
                             |                                           
                             |  +----------- Schistodesmus_lampreyanus[1]
                             +--+                                        
                                |  +-------- Cuneopsis_rufescens         
                                +--+                                     
                                   |  +----- Cuneopsis_celtiformis       
                                   +--+                                  
                                      |  +-- Cuneopsis_demangei          
                                      +--+                               
                                         +-- Cuneopsis_heudei            

Huang et al., 2019 [Publ 3843], fig. 2

iT tT bi riS oT roT roS iD
25 25 20 4 179 140 3 23.0

*** Top Sister-Group Cladogram ***

                          +----------- Hyriopsis_desowitzi        
                          |                                       
                          +----------- Rectidens_sumatrensis      
                          |                                       
                          |        +-- Hyriopsis_bialata          
                          |        |                              
                          |        +-- Hyriopsis_myersiana        
                    +-----+--------+                              
                    |     |        +-- Hyriopsis_panhai           
                    |     |        |                              
                    |     |        +-- Hyriopsis_khoratensis      
                    |     |                                       
                    |     |        +-- Ensidens_ingallsianus      
                    |     |  +-----+                              
                    |     |  |     +-- Ensidens_telus             
                    |     +--+                                    
                    |        |  +----- Ensidens_sagittarius       
  +-----------------+        +--+                                 
  |                 |           |  +-- Ensidens_spiculus          
  |                 |           +--+                              
  |                 |              +-- Ensidens_jaculus           
  |                 |                                             
  |                 |           +----- Physunio                   
  |                 |  +--------+                                 
  |                 |  |        |  +-- Physunio_modelli           
  |                 |  |        +--+                              
  |                 |  |           +-- Physunio_superbus          
  |                 |  |                                          
  |                 +--+     +-------- Yaukthwa_inlenensis        
  |                    |  +--+                                    
  |                    |  |  |  +----- Yaukthwa_panhai            
  |                    |  |  +--+                                 
  |                    |  |     |  +-- Yaukthwa_nesemanni         
  |                    |  |     +--+                              
  |                    +--+        +-- Yaukthwa_rectangularis     
  |                       |                                       
  |                       |        +-- Lens_contradens            
  |                       |  +-----+                              
  |                       |  |     +-- Lens_micropterus           
  |                       +--+                                    
  |                          |  +----- Lens_eximius               
  |                          +--+                                 
  |                             |  +-- Lens_comptus               
  |                             +--+                              
  |                                +-- Lens_novoselovi            
  |                                                               
  |                                +-- Megalonaias_nervosa **        
  |                             +--+                              
--+                             |  +-- Uniomerus_carolinianus **     
  |     +-----------------------+                                 
  |     |                       |  +-- Quadrula_quadrula **          
  |     |                       +--+                              
  |     |                          +-- Quadrula_verrucosa **         
  |     |                                                         
  |     |                    +-------- Elliptoideus_sloatianus **    
  |  +--+                    |                                    
  |  |  |  +-----------------+     +-- Elliptio_complanata **        
  |  |  |  |                 |  +--+                              
  |  |  |  |                 |  |  +-- Eurynia_dilatata **           
  |  |  |  |                 +--+                                 
  |  |  |  |                    |  +-- Fusconaia_flava **            
  |  |  +--+                    +--+                              
  |  |     |                       +-- Pleurobema_sintoxia **        
  |  |     |                                                      
  |  |     |  +----------------------- Amblema_plicata **            
  |  |     |  |                                                   
  |  |     |  |                 +----- Toxolasma_parvum **           
  |  |     +--+  +--------------+                                 
  |  |        |  |              |  +-- Obliquaria_reflexa **         
  |  |        |  |              +--+                              
  |  |        |  |                 +-- Plectomerus_dombeyanus **     
  |  |        +--+                                                
  |  |           |  +----------------- Potamilus_leptodon **         
  |  |           |  |                                             
  |  |           |  |  +-------------- Potamilus_alatus **           
  |  |           +--+  |                                          
  |  |              |  |  +----------- Venustaconcha_ellipsiformis **
  |  |              |  |  |                                       
  |  |              +--+  +----------- Leaunio_lienosus **           
  |  |                 |  |                                       
  +--+                 |  |        +-- Ellipsaria_lineolata **       
     |                 |  +--------+                              
     |                 +--+        +-- Truncilla_truncata **         
     |                    |                                       
     |                    |        +-- Medionidus_conradicus **      
     |                    +--------+                              
     |                    |        +-- Ptychobranchus_subtentus **   
     |                    |                                       
     |                    |  +-------- Cambarunio_iris **            
     |                    +--+                                    
     |                       |  +----- Ortmanniana_ligamentina **    
     |                       +--+                                 
     |                          |  +-- Lampsilis_cardium **          
     |                          +--+                              
     |                             +-- Lampsilis_ornata **           
     |                                                            
     |                          +----- Chamberlainia_somsakpanhai 
     |  +-----------------------+                                 
     |  |                       |  +-- Sinohyriopsis_cumingii     
     |  |                       +--+                              
     |  |                          +-- Sinohyriopsis_schlegelii   
     |  |                                                         
     |  |                          +-- Potomida_littoralis        
     |  |                    +-----+                              
     |  |                    |     +-- Pronodularia_japanensis    
     |  |  +-----------------+                                    
     +--+  |                 |  +----- Lamprotula_leaii           
        |  |                 +--+                                 
        |  |                    |  +-- Lamprotula_caveata         
        |  |                    +--+                              
        |  |                       +-- Lamprotula_cornuumlunae    
        |  |                                                      
        |  |                 +-------- Sinosolenaia_carinata      
        |  |              +--+                                    
        |  |              |  |  +----- Sinosolenaia_recognita     
        |  |              |  +--+                                 
        +--+              |     |  +-- Sinosolenaia_oleivora      
           |              |     +--+                              
           |  +-----------+        +-- Sinosolenaia_iridinea      
           |  |           |                                       
           |  |           |     +----- Gonidea_angulata           
           |  |           |  +--+                                 
           |  |           |  |  |  +-- Leguminaia_wheatleyi       
           |  |           |  |  +--+                              
           |  |           +--+     +-- Microcondylaea_bonellii    
           |  |              |                                    
           |  |              |     +-- Ptychorhynchus_pfisteri    
           |  |              |     |                              
           +--+              +-----+-- Parvasolenaia_rivularis    
              |                    |                              
              |                    +-- Koreosolenaia_sitgyensis   
              |                                                   
              |                 +----- Indopseudodon_salwenianus  
              |  +--------------+                                 
              |  |              |  +-- Indopseudodon_indawgyiensis
              |  |              +--+                              
              |  |                 +-- Indopseudodon_bogani       
              |  |                                                
              |  |           +-------- Pilsbryoconcha_exilis[3]   
              +--+  +--------+                                    
                 |  |        |  +----- Pilsbryoconcha_exilis[2]   
                 |  |        +--+                                 
                 |  |           |  +-- Pilsbryoconcha_exilis[1]   
                 |  |           +--+                              
                 |  |              +-- Pilsbryoconcha_carinifera  
                 |  |                                             
                 +--+     +----------- Nyeinchanconcha_nyeinchani 
                    |  +--+                                       
                    |  |  |  +-------- Thaiconcha_callifera       
                    |  |  +--+                                    
                    |  |     |  +----- Bineurus_loeiensis         
                    |  |     +--+                                 
                    |  |        |  +-- Bineurus_mouhotii          
                    +--+        +--+                              
                       |           +-- Bineurus_exilis            
                       |                                          
                       |        +----- Pseudodon_vagulus          
                       |     +--+                                 
                       |     |  |  +-- Pseudodon_mekongi          
                       |     |  +--+                              
                       +-----+     +-- Pseudodon_vondembuschianus 
                             |                                    
                             |     +-- Sundadontina_cumingii      
                             |     |                              
                             +-----+-- Sundadontina_tumida[1]     
                                   |                              
                                   +-- Sundadontina_tumida[2]     

Inoue et al., 2018 [Publ 3858], fig. 1, 3a-c

iT tT bi riS oT roT roS iD
174 174 59 5 23 22 2 179.0
                                                     +-- Alasmidonta_marginata      
                                                     |                              
                                                     +-- Prolasmidonta_heterodon    
                                                     |                              
                                                     +-- Alasmidonta_undulata       
                                                     |                              
                                                     +-- Beringiana_beringiana      
                                                     |                              
                                                     +-- Anodonta_kennerlyi         
                                                     |                              
                                                     +-- Anodonta_nuttalliana       
                                                     |                              
                                                     +-- Anodonta_oregonensis       
                                                     |                              
                                                     +-- Arcidens_confragosus       
                                                     |                              
                                                     +-- Arcidens_wheeleri          
                                                     |                              
                                                     +-- Lasmigona_complanata       
                                                     |                              
  +--------------------------------------------------+-- Platynaias_compressa       
  |                                                  |                              
  |                                                  +-- Lasmigona_costata          
  |                                                  |                              
  |                                                  +-- Platynaias_decorata        
  |                                                  |                              
  |                                                  +-- Platynaias_subviridis      
  |                                                  |                              
  |                                                  +-- Pyganodon_grandis          
  |                                                  |                              
  |                                                  +-- Pyganodon_lacustris        
  |                                                  |                              
  |                                                  +-- Pseudodontoideus_subvexus  
  |                                                  |                              
  |                                                  +-- Strophitus_undulatus       
  |                                                  |                              
  |                                                  +-- Utterbackia_imbecillis     
--+                                                  |                              
  |                                                  +-- Utterbackia_peninsularis[1]
  |                                                  |                              
  |                                                  +-- Utterbackia_peninsularis[2]
  |                                                                                 
  |  +-------------------------------------------------- Gonidea_angulata           
  |  |                                                                              
  |  |                                               +-- Reginaia_ebenus **            
  |  |  +--------------------------------------------+                              
  |  |  |                                            +-- Reginaia_rotulata **          
  |  |  |                                                                           
  |  |  |                                            +-- Amblema_plicata **            
  |  |  +--------------------------------------------+                              
  |  |  |                                            +-- Amblema_elliottii **          
  |  |  |                                                                           
  |  |  |                                         +----- Popenaias_popeii **           
  |  |  |                                         |                                 
  |  |  |                                         |  +-- Dromus_dromas **              
  +--+  |                                         |  |                              
     |  |                                         |  +-- Epioblasma_rangiana **        
     |  |                                         |  |                              
     |  +-----------------------------------------+  +-- Glebula_rotundata **          
     |  |                                         |  |                              
     |  |                                         |  +-- Lampsilis_ovata **            
     |  |                                         |  |                              
     |  |                                         |  +-- Lampsilis_virescens **        
     |  |                                         |  |                              
     |  |                                         |  +-- Lemiox_rimosus **             
     |  |                                         +--+                              
     |  |                                            +-- Obliquaria_reflexa **         
     |  |                                            |                              
     |  |                                            +-- Obovaria_olivaria **          
     +--+                                            |                              
        |                                            +-- Obovaria_unicolor **          
        |                                            |                              
        |                                            +-- Plectomerus_dombeyanus **     
        |                                            |                              
        |                                            +-- Potamilus_alatus **           
        |                                            |                              
        |                                            +-- Potamilus_capax **            
        |                                                                           
        |                                            +-- Uniomerus_declivis **         
        |                                         +--+                              
        |                                         |  +-- Uniomerus_tetralasmus **      
        |                                         |                                 
        |                                         |  +-- Cyclonaias_tuberculata **     
        |  +--------------------------------------+  |                              
        |  |                                      |  +-- Pustulosa_pustulosa[1] **     
        |  |                                      |  |                              
        |  |                                      |  +-- Theliderma_metanevra **       
        |  |                                      +--+                              
        |  |                                         +-- Pustulosa_pustulosa[2] **     
        |  |                                         |                              
        |  |                                         +-- Quadrula_quadrula **          
        |  |                                         |                              
        |  |                                         +-- Quadrula_verrucosa **         
        |  |                                                                        
        +--+  +----------------------------------------- Elliptoideus_sloatianus **    
           |  |                                                                     
           |  |                                      +-- Elliptio_lanceolata **        
           |  |  +-----------------------------------+                              
           |  |  |                                   +-- Parvaspina_collina **         
           |  |  |                                                                  
           |  |  |  +----------------------------------- Plethobasus_cyphyus **        
           |  |  |  |                                                               
           |  |  |  +----------------------------------- Eurynia_dilatata **           
           |  |  |  |                                                               
           |  |  |  +----------------------------------- Hemistena_lata **             
           |  |  |  |                                                               
           +--+  |  |                                +-- Elliptio_complanata **        
              |  |  |                                |                              
              |  |  |                                +-- Elliptio_hopetonensis **      
              |  |  |                                |                              
              |  |  |                                +-- Elliptio_producta[1] **       
              |  |  |                                |                              
              |  |  |                                +-- Elliptio_shepardiana **       
              |  |  |                                |                              
              |  |  |                                +-- Elliptio_angustata **         
              |  |  |                                |                              
              |  |  |                                +-- Elliptio_arca **              
              |  |  |                                |                              
              |  |  |                                +-- Elliptio_arctata **           
              +--+  |                                |                              
                 |  |                                +-- Elliptio_jayensis **          
                 |  |                                |                              
                 |  |                                +-- Elliptio_crassidens **        
                 |  |                                |                              
                 |  +--------------------------------+-- Elliptio_dariensis **         
                 |  |                                |                              
                 |  |                                +-- Elliptio_fisheriana **        
                 |  |                                |                              
                 |  |                                +-- Elliptio_folliculata **       
                 |  |                                |                              
                 |  |                                +-- Elliptio_fumata **            
                 |  |                                |                              
                 |  |                                +-- Elliptio_icterina **          
                 |  |                                |                              
                 |  |                                +-- Elliptio_mcmichaeli **        
                 |  |                                |                              
                 |  |                                +-- Elliptio_producta[2] **       
                 |  |                                |                              
                 |  |                                +-- Elliptio_pullata **           
                 |  |                                |                              
                 |  |                                +-- Elliptio_purpurella **        
                 |  |                                |                              
                 +--+                                +-- Elliptio_congaraea **         
                    |                                                               
                    |                                +-- Pleuronaia_barnesiana **      
                    |                                |                              
                    |  +-----------------------------+-- Pleuronaia_dolabelloides **   
                    |  |                             |                              
                    |  |                             +-- Pleuronaia_gibber **          
                    |  |                                                            
                    |  |     +-------------------------- Pleurobema_plenum **          
                    |  |     |                                                      
                    |  |     |  +----------------------- Pleurobema_stabile **         
                    |  |     |  |                                                   
                    |  |     |  |                 +----- Pleurobema_pyriforme **       
                    |  |  +--+  |              +--+                                 
                    |  |  |  |  |              |  |  +-- Pleurobema_georgianum[2] **   
                    |  |  |  |  |              |  +--+                              
                    |  |  |  |  |  +-----------+     +-- Pleurobema_athearni **        
                    |  |  |  |  |  |           |                                    
                    |  |  |  +--+  |           |  +----- Pleurobema_decisum **         
                    |  |  |     |  |           |  |                                 
                    |  |  |     |  |           +--+  +-- Pleurobema_georgianum[1] **   
                    |  |  |     |  |              |  |                              
                    |  |  |     |  |              +--+-- Pleurobema_hanleyianum[1] **  
                    |  |  |     |  |                 |                              
                    |  |  |     |  |                 +-- Pleurobema_hanleyianum[2] **  
                    |  |  |     |  |                                                
                    |  |  |     +--+     +-------------- Pleurobema_beadleianum[2] **  
                    |  |  |        |     |                                          
                    |  |  |        |  +--+  +----------- Pleurobema_strodeanum[2] **   
                    |  |  |        |  |  |  |                                       
                    |  |  |        |  |  +--+----------- Pleurobema_rubellum **        
                    |  |  |        |  |     |                                       
                    |  |  |        |  |     |  +-------- Pleurobema_beadleianum[1] **  
                    |  |  |        |  |     +--+                                    
                    |  |  |        |  |        |  +----- Pleurobema_taitianum **       
                    |  |  |        |  |        +--+                                 
                    |  |  |        |  |           |  +-- Pleurobema_perovatum **       
                    |  |  |        +--+           +--+                              
                    |  |  |           |              +-- Pleurobema_strodeanum[1] **   
                    |  |  |           |                                             
                    |  |  |           |           +----- Pleurobema_oviforme[6] **     
                    |  |  |           |           |                                 
                    |  |  |           |        +--+----- Pleurobema_clava[3] **        
                    +--+  |           |        |  |                                 
                       |  |           |        |  |  +-- Pleurobema_oviforme[5] **     
                       |  |           |        |  +--+                              
                       |  |           +--------+     +-- Pleurobema_clava[2] **        
                       |  |                    |                                    
                       |  |                    |  +----- Pleurobema_oviforme[2] **     
                       |  |                    |  |                                 
                       |  |                    |  +----- Pleurobema_oviforme[3] **     
                       |  |                    +--+                                 
                       |  |                       +----- Pleurobema_oviforme[4] **     
                       |  |                       |                                 
                       |  |                       |  +-- Pleurobema_clava[1] **        
                       |  |                       +--+                              
                       |  |                          +-- Pleurobema_oviforme[1] **     
                       |  |                                                         
                       |  |           +----------------- Pleurobema_riddellii **       
                       |  |           |                                             
                       |  |           |  +-------------- Pleurobema_sintoxia[34] **    
                       |  |           |  |                                          
                       |  |           |  |  +----------- Pleurobema_sintoxia[31] **    
                       |  |           |  |  |                                       
                       |  +-----------+  |  +----------- Pleurobema_sintoxia[32] **    
                       |  |           |  |  |                                       
                       |  |           |  |  +----------- Pleurobema_sintoxia[33] **    
                       |  |           |  |  |                                       
                       |  |           |  |  |  +-------- Pleurobema[4] **              
                       |  |           |  |  |  |                                    
                       |  |           |  |  |  +-------- Pleurobema[5] **              
                       |  |           +--+  |  |                                    
                       |  |              |  |  +-------- Pleurobema_sintoxia[2] **     
                       |  |              |  |  |                                    
                       |  |              |  |  +-------- Pleurobema_sintoxia[3] **     
                       |  |              |  |  |                                    
                       |  |              |  |  +-------- Pleurobema[8] **              
                       |  |              |  |  |                                    
                       |  |              |  |  +-------- Pleurobema[9] **              
                       |  |              |  |  |                                    
                       |  |              |  |  +-------- Pleurobema[10] **             
                       |  |              |  |  |                                    
                       +--+              |  |  +-------- Pleurobema_sintoxia[5] **     
                          |              +--+  |                                    
                          |                 |  +-------- Pleurobema_sintoxia[6] **     
                          |                 |  |                                    
                          |                 |  +-------- Pleurobema_sintoxia[7] **     
                          |                 |  |                                    
                          |                 |  +-------- Pleurobema_sintoxia[8] **     
                          |                 |  |                                    
                          |                 |  +-------- Pleurobema_sintoxia[9] **     
                          |                 |  |                                    
                          |                 |  +-------- Pleurobema_sintoxia[10] **    
                          |                 |  |                                    
                          |                 |  +-------- Pleurobema_sintoxia[11] **    
                          |                 |  |                                    
                          |                 |  +-------- Pleurobema_sintoxia[12] **    
                          |                 |  |                                    
                          |                 |  +-------- Pleurobema_sintoxia[13] **    
                          |                 |  |                                    
                          |                 |  +-------- Pleurobema_sintoxia[14] **    
                          |                 |  |                                    
                          |                 |  +-------- Pleurobema_sintoxia[15] **    
                          |                 |  |                                    
                          |                 |  +-------- Pleurobema_sintoxia[16] **    
                          |                 +--+                                    
                          |                    +-------- Pleurobema_sintoxia[17] **    
                          |                    |                                    
                          |                    +-------- Pleurobema_sintoxia[18] **    
                          |                    |                                    
                          |                    +-------- Pleurobema_sintoxia[19] **    
                          |                    |                                    
                          |                    +-------- Pleurobema_sintoxia[20] **    
                          |                    |                                    
                          |                    +-------- Pleurobema_sintoxia[21] **    
                          |                    |                                    
                          |                    +-------- Pleurobema_sintoxia[22] **    
                          |                    |                                    
                          |                    +-------- Pleurobema_sintoxia[23] **    
                          |                    |                                    
                          |                    +-------- Pleurobema_sintoxia[24] **    
                          |                    |                                    
                          |                    +-------- Pleurobema_sintoxia[25] **    
                          |                    |                                    
                          |                    +-------- Pleurobema_sintoxia[26] **    
                          |                    |                                    
                          |                    +-------- Pleurobema_sintoxia[27] **    
                          |                    |                                    
                          |                    +-------- Pleurobema_sintoxia[28] **    
                          |                    |                                    
                          |                    +-------- Pleurobema_sintoxia[29] **    
                          |                    |                                    
                          |                    +-------- Pleurobema_sintoxia[30] **    
                          |                    |                                    
                          |                    |     +-- Pleurobema[6] **              
                          |                    +-----+                              
                          |                    |     +-- Pleurobema_sintoxia[1] **     
                          |                    |                                    
                          |                    |     +-- Pleurobema[7] **              
                          |                    +-----+                              
                          |                    |     +-- Pleurobema_sintoxia[4] **     
                          |                    |                                    
                          |                    |  +----- Pleurobema[3] **              
                          |                    +--+                                 
                          |                       |  +-- Pleurobema[1] **              
                          |                       +--+                              
                          |                          +-- Pleurobema[2] **              
                          |                                                         
                          |  +-------------------------- Pleurobema_cordatum **        
                          |  |                                                      
                          |  |                    +----- Fusconaia_subrotunda **       
                          |  |  +-----------------+                                 
                          +--+  |                 |  +-- Fusconaia_cor **              
                             |  |                 +--+                              
                             |  |                    +-- Fusconaia_cuneolus **         
                             |  |                                                   
                             |  |  +-------------------- Fusconaia_ozarkensis **       
                             +--+  |                                                
                                |  |                 +-- Fusconaia_escambia **         
                                |  |  +--------------+                              
                                |  |  |              +-- Fusconaia_burkei **           
                                |  |  |                                             
                                |  |  |           +----- Fusconaia_flava[27] **        
                                |  |  |           |                                 
                                +--+  |           |  +-- Fusconaia_iheringi **         
                                   |  |           +--+                              
                                   |  |           |  +-- Fusconaia_mitchelli **        
                                   |  +-----------+                                 
                                   |  |           |  +-- Fusconaia_askewi[1] **        
                                   |  |           |  |                              
                                   |  |           |  +-- Fusconaia_askewi[2] **        
                                   |  |           |  |                              
                                   +--+           |  +-- Fusconaia_askewi[3] **        
                                      |           +--+                              
                                      |              +-- Fusconaia_askewi[4] **        
                                      |              |                              
                                      |              +-- Fusconaia_askewi[5] **        
                                      |              |                              
                                      |              +-- Fusconaia_chunii **           
                                      |                                             
                                      |  +-------------- Fusconaia_masoni **           
                                      |  |                                          
                                      |  |  +----------- Fusconaia_flava[26] **        
                                      |  |  |                                       
                                      |  |  |        +-- Fusconaia_cerina[8] **        
                                      +--+  |  +-----+                              
                                         |  |  |     +-- Fusconaia_flava[25] **        
                                         |  |  |                                    
                                         |  |  |  +----- Fusconaia_flava[5] **         
                                         |  |  |  |                                 
                                         |  |  |  +----- Fusconaia_flava[6] **         
                                         +--+  |  |                                 
                                            |  |  +----- Fusconaia_flava[7] **         
                                            |  |  |                                 
                                            |  |  +----- Fusconaia_flava[8] **         
                                            |  |  |                                 
                                            |  |  +----- Fusconaia_cerina[4] **        
                                            |  |  |                                 
                                            |  |  +----- Fusconaia_flava[9] **         
                                            |  |  |                                 
                                            +--+  +----- Fusconaia_flava[10] **        
                                               |  |                                 
                                               |  +----- Fusconaia_flava[11] **        
                                               |  |                                 
                                               |  +----- Fusconaia_flava[12] **        
                                               |  |                                 
                                               |  +----- Fusconaia_flava[13] **        
                                               |  |                                 
                                               |  +----- Fusconaia_flava[14] **        
                                               |  |                                 
                                               |  +----- Fusconaia_flava[15] **        
                                               |  |                                 
                                               |  +----- Fusconaia_flava[16] **        
                                               |  |                                 
                                               |  +----- Fusconaia_flava[17] **        
                                               +--+                                 
                                                  +----- Fusconaia_flava[18] **        
                                                  |                                 
                                                  +----- Fusconaia_flava[19] **        
                                                  |                                 
                                                  +----- Fusconaia_flava[20] **        
                                                  |                                 
                                                  +----- Fusconaia_flava[21] **        
                                                  |                                 
                                                  +----- Fusconaia_flava[22] **        
                                                  |                                 
                                                  +----- Fusconaia_flava[23] **        
                                                  |                                 
                                                  +----- Fusconaia_flava[24] **        
                                                  |                                 
                                                  +----- Fusconaia_cerina[5] **        
                                                  |                                 
                                                  +----- Fusconaia_cerina[6] **        
                                                  |                                 
                                                  +----- Fusconaia_cerina[7] **        
                                                  |                                 
                                                  |  +-- Fusconaia_flava[1] **         
                                                  |  |                              
                                                  |  +-- Fusconaia_flava[2] **         
                                                  |  |                              
                                                  |  +-- Fusconaia_flava[3] **         
                                                  |  |                              
                                                  +--+-- Fusconaia_flava[4] **         
                                                     |                              
                                                     +-- Fusconaia_cerina[1] **        
                                                     |                              
                                                     +-- Fusconaia_cerina[2] **        
                                                     |                              
                                                     +-- Fusconaia_cerina[3] **        

Campbell et al., 2005 [Publ 2660], fig. 1

iT tT bi riS oT roT roS iD
104 104 92 5 12 10 2 102.0
  +-------------------------------------------------- Sinohyriopsis_cumingii        
  |                                                                                 
  |                                      +----------- Anodonta_oregonensis          
  |                                      |                                          
  +--------------------------------------+        +-- Anodonta_cygnea               
  |                                      |  +-----+                                 
  |                                      |  |     +-- Alasmidonta_holstonia         
  |                                      +--+                                       
  |                                         |  +----- Pyganodon_grandis             
  |                                         +--+                                    
  |                                            |  +-- Lasmigona_costata             
  |                                            +--+                                 
  |                                               +-- Pseudodontoideus_subvexus     
  |                                                                                 
  |                                            +----- Gonidea_angulata              
  |  +-----------------------------------------+                                    
  |  |                                         |  +-- Potomida_littoralis           
--+  |                                         +--+                                 
  |  |                                            +-- Pronodularia_japanensis       
  |  |                                                                              
  |  |                                            +-- Megalonaias_nervosa **           
  |  |                                +-----------+                                 
  |  |                                |           +-- Uniomerus_declivis **            
  |  |                                |                                             
  |  |  +-----------------------------+  +----------- Theliderma_metanevra **          
  |  |  |                             |  |                                          
  |  |  |                             |  |     +----- Quadrula_verrucosa **            
  |  |  |                             +--+  +--+                                    
  |  |  |                                |  |  |  +-- Quadrula_quadrula **             
  |  |  |                                |  |  +--+                                 
  |  |  |                                |  |     +-- Quadrula_nobilis **              
  |  |  |                                +--+                                       
  |  |  |                                   |  +----- Pustulosa_kieneriana **          
  |  |  |                                   |  |                                    
  |  |  |                                   |  |  +-- Cyclonaias_tuberculata **        
  +--+  |                                   +--+--+                                 
     |  |                                      |  +-- Pustulosa_succissa **            
     |  |                                      |                                    
     |  |                                      |  +-- Pustulosa_infucata **            
     |  |                                      +--+                                 
     |  |                                         +-- Pustulosa_kleiniana **           
     |  |                                                                           
     |  |        +----------------------------------- Elliptoideus_sloatianus **       
     |  |        |                                                                  
     |  |        |  +-------------------------------- Hemistena_lata **                
     |  |        |  |                                                               
     |  |  +-----+  |                          +----- Plethobasus_cyphyus **           
     |  |  |     |  |  +-----------------------+                                    
     |  |  |     |  |  |                       |  +-- Elliptio_arca **                 
     |  |  |     |  |  |                       +--+                                 
     |  |  |     |  |  |                          +-- Elliptio_crassidens **           
     |  |  |     +--+  |                                                            
     |  |  |        |  |                          +-- Eurynia_dilatata **              
     |  |  |        |  |  +-----------------------+                                 
     |  |  |        |  |  |                       +-- Parvaspina_collina **            
     +--+  |        |  |  |                                                         
        |  |        |  |  |                    +----- Pleuronaia_gibber **             
        |  |        |  |  +--------------------+                                    
        |  |        +--+  |                    |  +-- Pleuronaia_barnesiana **         
        |  |           |  |                    +--+                                 
        |  |           |  |                       +-- Pleuronaia_dolabelloides **      
        |  |           |  |                                                         
        |  |           |  |              +----------- Fusconaia_subrotunda **          
        |  |           |  |              |                                          
        |  |           |  |              |        +-- Fusconaia_cor **                 
        |  |           |  +--------------+  +-----+                                 
        |  |           |  |              |  |     +-- Fusconaia_cuneolus **            
        |  |           |  |              |  |                                       
        |  |           |  |              |  |     +-- Fusconaia_cerina[1] **           
        |  |           |  |              +--+     |                                 
        |  |           +--+                 |  +--+-- Fusconaia_cerina[2] **           
        |  |              |                 |  |  |                                 
        |  |              |                 |  |  +-- Fusconaia_flava **               
        |  |              |                 +--+                                    
        |  |              |                    |  +-- Fusconaia_burkei[1] **           
        |  |              |                    |  |                                 
        |  |              |                    |  +-- Fusconaia_burkei[2] **           
        |  |              |                    +--+                                 
        |  |              |                       +-- Fusconaia_burkei[3] **           
        |  |              |                       |                                 
        |  |              |                       +-- Fusconaia_escambia **            
        |  |              |                                                         
        |  |              |                    +----- Pleurobema_sintoxia[2] **        
        |  |              |  +-----------------+                                    
        |  |              |  |                 |  +-- Pleurobema_sintoxia[1] **        
        |  |              |  |                 +--+                                 
        +--+              |  |                    +-- Pleurobema_cordatum **           
           |              +--+                                                      
           |                 |  +-------------------- Pleurobema_pyriforme **          
           |                 |  |                                                   
           |                 |  |                 +-- Pleurobema_georgianum[2] **      
           |                 |  |                 |                                 
           |                 +--+  +--------------+-- Pleurobema_georgianum[3] **      
           |                    |  |              |                                 
           |                    |  |              +-- Pleurobema_athearni **           
           |                    |  |                                                
           |                    |  |           +----- Pleurobema_decisum **            
           |                    +--+           |                                    
           |                       |  +--------+  +-- Pleurobema_hanleyianum[1] **     
           |                       |  |        |  |                                 
           |                       |  |        +--+-- Pleurobema_hanleyianum[2] **     
           |                       |  |           |                                 
           |                       |  |           +-- Pleurobema_georgianum[1] **      
           |                       +--+                                             
           |                          |           +-- Pleurobema_oviforme[1] **        
           |                          |           |                                 
           |                          |  +--------+-- Pleurobema_oviforme[2] **        
           |                          |  |        |                                 
           |                          +--+        +-- Pleurobema_clava **              
           |                             |                                          
           |                             |  +-------- Pleurobema_strodeanum **         
           |                             +--+                                       
           |                                |  +----- Pleurobema_rubellum **           
           |                                +--+                                    
           |                                   |  +-- Pleurobema_perovatum **          
           |                                   +--+                                 
           |                                      +-- Pleurobema_taitianum **          
           |                                                                        
           |                                   +----- Plectomerus_dombeyanus **        
           |  +--------------------------------+                                    
           |  |                                |  +-- Reginaia_ebenus **               
           |  |                                +--+                                 
           |  |                                   +-- Reginaia_rotulata **             
           |  |                                                                     
           |  |                                +----- Popenaias_popeii **              
           |  |  +-----------------------------+                                    
           |  |  |                             |  +-- Amblema_elliottii **             
           +--+  |                             +--+                                 
              |  |                                +-- Amblema_plicata **               
              |  |                                                                  
              |  |                                +-- Toxolasma_parvum **              
              |  |                          +-----+                                 
              |  |                          |     +-- Toxolasma_texasiense **          
              |  |  +-----------------------+                                       
              |  |  |                       |  +----- Obliquaria_reflexa **            
              +--+  |                       +--+                                    
                 |  |                          |  +-- Cyrtonaias_tampicoensis **       
                 |  |                          +--+                                 
                 |  |                             +-- Glebula_rotundata **             
                 |  |                                                               
                 |  |                             +-- Potamilus_fragilis **            
                 |  |                          +--+                                 
                 |  |                          |  +-- Potamilus_leptodon **            
                 |  |              +-----------+                                    
                 |  |              |           |  +-- Potamilus_alatus **              
                 |  |              |           +--+                                 
                 +--+              |              +-- Potamilus_purpuratus **          
                    |              |                                                
                    |  +-----------+              +-- Sagittunio_nasutus **            
                    |  |           |           +--+                                 
                    |  |           |           |  +-- Medionidus_conradicus **         
                    |  |           |  +--------+                                    
                    |  |           |  |        |  +-- Ptychobranchus_fasciolaris **    
                    |  |           |  |        +--+                                 
                    |  |           +--+           +-- Venustaconcha_ellipsiformis[2] **
                    |  |              |                                             
                    |  |              |  +----------- Medionidus_acutissimus **        
                    |  |              +--+                                          
                    |  |                 |  +-------- Lemiox_rimosus **                
                    |  |                 +--+                                       
                    +--+                    |  +----- Dromus_dromas **                 
                       |                    +--+                                    
                       |                       |  +-- Cyprogenia_aberti **             
                       |                       +--+                                 
                       |                          +-- Cyprogenia_stegaria **           
                       |                                                            
                       |                          +-- Ellipsaria_lineolata **          
                       |  +-----------------------+                                 
                       |  |                       +-- Truncilla_truncata **            
                       |  |                                                         
                       |  |  +----------------------- Villosa_villosa **               
                       |  |  |                                                      
                       |  |  |                    +-- Obovaria_olivaria **             
                       +--+  |     +--------------+                                 
                          |  |     |              +-- Venustaconcha_ellipsiformis[1] **
                          |  |     |                                                
                          |  |  +--+           +----- Obovaria_unicolor **             
                          |  |  |  |  +--------+                                    
                          |  |  |  |  |        |  +-- Obovaria_arkansasensis **        
                          +--+  |  |  |        +--+                                 
                             |  |  +--+           +-- Obovaria_subrotunda **           
                             |  |     |                                             
                             |  |     |  +----------- Epioblasma_brevidens **          
                             |  |     +--+                                          
                             |  |        |  +-------- Epioblasma_triquetra **          
                             |  |        +--+                                       
                             |  |           |  +----- Epioblasma_walkeri[2] **         
                             |  |           +--+                                    
                             |  |              |  +-- Epioblasma_capsaeformis **       
                             +--+              +--+                                 
                                |                 +-- Epioblasma_walkeri[1] **         
                                |                                                   
                                |                 +-- Ligumia_recta **                 
                                |        +--------+                                 
                                |        |        +-- Cambarunio_iris **               
                                |     +--+                                          
                                |     |  |  +-------- Lampsilis_teres **               
                                |     |  +--+                                       
                                |     |     |  +----- Ortmanniana_ligamentina **       
                                |     |     +--+                                    
                                |     |        |  +-- Lampsilis_siliquoidea **         
                                |     |        +--+                                 
                                |     |           +-- Leaunio_vanuxemensis **          
                                +-----+                                             
                                      |           +-- Lampsilis_cardium **             
                                      |        +--+                                 
                                      |        |  +-- Lampsilis_ovata **               
                                      |  +-----+                                    
                                      |  |     |  +-- Ortmanniana_pectorosa **         
                                      |  |     +--+                                 
                                      |  |        +-- Lampsilis_ornata **              
                                      |  |                                          
                                      +--+     +----- Hamiota_australis[2] **          
                                         |  +--+                                    
                                         |  |  |  +-- Hamiota_australis[1] **          
                                         |  |  +--+                                 
                                         |  |     +-- Hamiota_subangulata **           
                                         +--+                                       
                                            |  +----- Hamiota_altilis[2] **            
                                            |  |                                    
                                            |  |  +-- Hamiota_altilis[1] **            
                                            +--+--+                                 
                                               |  +-- Hamiota_perovalis[1] **          
                                               |                                    
                                               |  +-- Hamiota_altilis[3] **            
                                               +--+                                 
                                                  +-- Hamiota_perovalis[2] **          

Campbell et al., 2005 [Publ 2660], fig. 2

iT tT bi riS oT roT roS iD
102 102 81 5 12 10 2 99.0
                                   +----------- Sinohyriopsis_cumingii        
                                   |                                          
  +--------------------------------+  +-------- Pyganodon_grandis             
  |                                |  |                                       
  |                                |  |     +-- Lasmigona_costata             
  |                                +--+-----+                                 
  |                                   |     +-- Pseudodontoideus_subvexus     
  |                                   |                                       
  |                                   |  +----- Alasmidonta_holstonia         
  |                                   +--+                                    
  |                                      |  +-- Anodonta_cygnea               
  |                                      +--+                                 
  |                                         +-- Anodonta_oregonensis          
  |                                                                           
  |                                      +----- Gonidea_angulata              
  |  +-----------------------------------+                                    
  |  |                                   |  +-- Potomida_littoralis           
  |  |                                   +--+                                 
--+  |                                      +-- Pronodularia_japanensis       
  |  |                                                                        
  |  |                    +-------------------- Megalonaias_nervosa **           
  |  |                    |                                                   
  |  |  +-----------------+  +----------------- Uniomerus_declivis **            
  |  |  |                 |  |                                                
  |  |  |                 +--+  +-------------- Theliderma_metanevra **          
  |  |  |                    |  |                                             
  |  |  |                    +--+  +----------- Quadrula_quadrula **             
  |  |  |                       |  |                                          
  |  |  |                       |  |        +-- Quadrula_nobilis **              
  |  |  |                       +--+  +-----+                                 
  |  |  |                          |  |     +-- Quadrula_verrucosa **            
  |  |  |                          |  |                                       
  |  |  |                          +--+  +----- Pustulosa_kieneriana **          
  |  |  |                             |  |                                    
  +--+  |                             |  |  +-- Cyclonaias_tuberculata **        
     |  |                             +--+--+                                 
     |  |                                |  +-- Pustulosa_succissa **            
     |  |                                |                                    
     |  |                                |  +-- Pustulosa_infucata **            
     |  |                                +--+                                 
     |  |                                   +-- Pustulosa_kleiniana **           
     |  |                                                                     
     |  |     +-------------------------------- Elliptoideus_sloatianus **       
     |  |     |                                                               
     |  |     |  +----------------------------- Plethobasus_cyphyus **           
     |  |     |  |                                                            
     |  |     |  +----------------------------- Parvaspina_collina **            
     |  |  +--+  |                                                            
     |  |  |  |  |                          +-- Elliptio_arca **                 
     |  |  |  |  +--------------------------+                                 
     |  |  |  |  |                          +-- Elliptio_crassidens **           
     |  |  |  |  |                                                            
     |  |  |  +--+  +-------------------------- Eurynia_dilatata **              
     |  |  |     |  |                                                         
     +--+  |     |  +-------------------------- Hemistena_lata **                
        |  |     |  |                                                         
        |  |     |  |                    +----- Pleurobema_sintoxia[2] **        
        |  |     |  |  +-----------------+                                    
        |  |     |  |  |                 |  +-- Pleurobema_sintoxia[1] **        
        |  |     |  |  |                 +--+                                 
        |  |     |  |  |                    +-- Pleurobema_cordatum **           
        |  |     +--+  |                                                      
        |  |        |  |        +-------------- Fusconaia_subrotunda **          
        |  |        |  |        |                                             
        |  |        |  |        |           +-- Fusconaia_cor **                 
        |  |        |  +--------+  +--------+                                 
        |  |        |  |        |  |        +-- Fusconaia_cuneolus **            
        |  |        |  |        |  |                                          
        |  |        |  |        +--+        +-- Fusconaia_cerina **              
        |  |        |  |           |  +-----+                                 
        |  |        |  |           |  |     +-- Fusconaia_flava **               
        |  |        +--+           +--+                                       
        |  |           |              |  +----- Fusconaia_burkei[2] **           
        |  |           |              +--+                                    
        |  |           |                 |  +-- Fusconaia_escambia **            
        |  |           |                 +--+                                 
        |  |           |                    +-- Fusconaia_burkei[1] **           
        |  |           |                                                      
        |  |           |                 +----- Pleuronaia_gibber **             
        |  |           |  +--------------+                                    
        |  |           |  |              |  +-- Pleuronaia_barnesiana **         
        |  |           |  |              +--+                                 
        |  |           |  |                 +-- Pleuronaia_dolabelloides **      
        |  |           |  |                                                   
        |  |           |  |           +-------- Pleurobema_rubellum **           
        |  |           +--+  +--------+                                       
        +--+              |  |        |  +----- Pleurobema_strodeanum **         
           |              |  |        +--+                                    
           |              |  |           |  +-- Pleurobema_perovatum **          
           |              |  |           +--+                                 
           |              |  |              +-- Pleurobema_taitianum **          
           |              +--+                                                
           |                 |              +-- Pleurobema_oviforme[1] **        
           |                 |              |                                 
           |                 |  +-----------+-- Pleurobema_oviforme[2] **        
           |                 |  |           |                                 
           |                 |  |           +-- Pleurobema_clava **              
           |                 +--+                                             
           |                    |  +----------- Pleurobema_pyriforme **          
           |                    |  |                                          
           |                    +--+        +-- Pleurobema_athearni **           
           |                       |  +-----+                                 
           |                       |  |     +-- Pleurobema_georgianum[2] **      
           |                       +--+                                       
           |                          |  +----- Pleurobema_decisum **            
           |                          |  |                                    
           |                          +--+  +-- Pleurobema_hanleyianum[1] **     
           |                             |  |                                 
           |                             +--+-- Pleurobema_hanleyianum[2] **     
           |                                |                                 
           |                                +-- Pleurobema_georgianum[1] **      
           |                                                                  
           |                          +-------- Popenaias_popeii **              
           |                          |                                       
           |           +--------------+     +-- Amblema_elliottii **             
           |           |              |  +--+                                 
           |           |              |  |  +-- Amblema_plicata **               
           |           |              +--+                                    
           |           |                 |  +-- Reginaia_ebenus **               
           |           |                 |  |                                 
           |           |                 +--+-- Reginaia_rotulata[1] **          
           |           |                    |                                 
           |           |                    +-- Reginaia_rotulata[2] **          
           |           |                                                      
           |           |                 +----- Plectomerus_dombeyanus **        
           |           |  +--------------+                                    
           +-----------+  |              |  +-- Toxolasma_parvum **              
                       |  |              +--+                                 
                       |  |                 +-- Toxolasma_texasiense **          
                       |  |                                                   
                       |  |              +----- Obliquaria_reflexa **            
                       |  |  +-----------+                                    
                       |  |  |           |  +-- Cyrtonaias_tampicoensis **       
                       |  |  |           +--+                                 
                       |  |  |              +-- Glebula_rotundata **             
                       |  |  |                                                
                       |  |  |  +-------------- Sagittunio_nasutus **            
                       |  |  |  |                                             
                       +--+  |  +-------------- Leaunio_vanuxemensis **          
                          |  |  |                                             
                          |  |  |           +-- Ellipsaria_lineolata **          
                          |  |  +-----------+                                 
                          |  |  |           +-- Truncilla_truncata **            
                          |  |  |                                             
                          |  |  |        +----- Lampsilis_teres **               
                          |  |  +--------+                                    
                          |  |  |        |  +-- Ortmanniana_ligamentina **       
                          |  |  |        +--+                                 
                          |  |  |           +-- Lampsilis_siliquoidea **         
                          |  |  |                                             
                          |  |  |        +----- Villosa_villosa **               
                          |  |  +--------+                                    
                          +--+  |        |  +-- Ligumia_recta **                 
                             |  |        +--+                                 
                             |  |           +-- Cambarunio_iris **               
                             |  |                                             
                             |  |     +-------- Ortmanniana_pectorosa **         
                             |  +-----+                                       
                             |  |     |  +----- Lampsilis_ornata **              
                             |  |     +--+                                    
                             |  |        |  +-- Lampsilis_cardium **             
                             |  |        +--+                                 
                             |  |           +-- Lampsilis_ovata **               
                             |  |                                             
                             |  |           +-- Potamilus_fragilis **            
                             |  |        +--+                                 
                             |  |        |  +-- Potamilus_leptodon **            
                             |  +--------+                                    
                             |  |        |  +-- Potamilus_alatus **              
                             |  |        +--+                                 
                             |  |           +-- Potamilus_purpuratus **          
                             |  |                                             
                             |  |           +-- Ptychobranchus_fasciolaris **    
                             +--+  +--------+                                 
                                |  |        +-- Venustaconcha_ellipsiformis[1] **
                                |  |                                          
                                +--+  +-------- Medionidus_conradicus **         
                                |  |  |                                       
                                |  |  |     +-- Lemiox_rimosus **                
                                |  +--+-----+                                 
                                |     |     +-- Medionidus_acutissimus **        
                                |     |                                       
                                |     |  +----- Dromus_dromas **                 
                                |     +--+                                    
                                |        |  +-- Cyprogenia_aberti **             
                                |        +--+                                 
                                |           +-- Cyprogenia_stegaria **           
                                |                                             
                                |        +----- Hamiota_australis[2] **          
                                |  +-----+                                    
                                |  |     |  +-- Hamiota_australis[1] **          
                                |  |     +--+                                 
                                |  |        +-- Hamiota_subangulata **           
                                +--+                                          
                                |  |        +-- Hamiota_altilis[3] **            
                                |  |  +-----+                                 
                                |  |  |     +-- Hamiota_perovalis[2] **          
                                |  +--+                                       
                                |     |  +----- Hamiota_altilis[2] **            
                                |     +--+                                    
                                |        |  +-- Hamiota_altilis[1] **            
                                |        +--+                                 
                                |           +-- Hamiota_perovalis[1] **          
                                |                                             
                                |  +----------- Obovaria_olivaria **             
                                |  |                                          
                                |  +----------- Venustaconcha_ellipsiformis[2] **
                                |  |                                          
                                |  |     +----- Obovaria_unicolor **             
                                |  +-----+                                    
                                +--+     |  +-- Obovaria_arkansasensis **        
                                   |     +--+                                 
                                   |        +-- Obovaria_subrotunda **           
                                   |                                          
                                   |        +-- Epioblasma_brevidens **          
                                   |  +-----+                                 
                                   |  |     +-- Epioblasma_triquetra **          
                                   +--+                                       
                                      |  +----- Epioblasma_walkeri[1] **         
                                      +--+                                    
                                         |  +-- Epioblasma_walkeri[2] **         
                                         +--+                                 
                                            +-- Epioblasma_capsaeformis **       

Campbell & Lydeard, 2012 [Publ 3309], fig. 1-2

iT tT bi riS oT roT roS iD
73 73 63 5 0 0 0 75.0
                                         +-- Uniomerus_declivis **         
                       +-----------------+                              
                       |                 +-- Uniomerus_tetralasmus **      
  +--------------------+                                                
  |                    |  +----------------- Theliderma_metanevra **       
  |                    |  |                                             
  |                    +--+  +-------------- Quadrula_verrucosa **         
  |                       |  |                                          
  |                       |  |           +-- Quadrula_quadrula **          
  |                       +--+  +--------+                              
  |                          |  |        +-- Quadrula_nobilis **           
  |                          |  |                                       
  |                          +--+        +-- Pustulosa_infucata **         
  |                             |  +-----+                              
  |                             |  |     +-- Pustulosa_kleiniana **        
  |                             +--+                                    
  |                                |  +----- Cyclonaias_tuberculata **     
  |                                +--+                                 
  |                                   |  +-- Pustulosa_pustulosa **        
  |                                   +--+                              
  |                                      +-- Pustulosa_succissa **         
  |                                                                     
  |                                   +----- Plectomerus_dombeyanus **     
  |                 +-----------------+                                 
  |                 |                 |  +-- Reginaia_ebenus **            
  |                 |                 +--+                              
--+                 |                    +-- Reginaia_rotulata **          
  |                 |                                                   
  |                 |                 +----- Popenaias_popeii **           
  |  +--------------+  +--------------+                                 
  |  |              |  |              |  +-- Amblema_elliottii **          
  |  |              |  |              +--+                              
  |  |              |  |                 +-- Amblema_plicata **            
  |  |              |  |                                                
  |  |              |  |  +----------------- Potamilus_alatus **           
  |  |              +--+  |                                             
  |  |                 |  +----------------- Potamilus_capax **            
  |  |                 |  |                                             
  |  |                 |  +----------------- Glebula_rotundata **          
  |  |                 |  |                                             
  |  |                 |  |              +-- Toxolasma_lividum **          
  |  |                 |  +--------------+                              
  |  |                 +--+              +-- Paetulunio_fabalis **         
  |  |                    |                                             
  |  |                    |              +-- Dromus_dromas **              
  |  |                    +--------------+                              
  |  |                    |              +-- Lemiox_rimosus **             
  |  |                    |                                             
  |  |                    |              +-- Lampsilis_ovata **            
  +--+                    |  +-----------+                              
     |                    |  |           +-- Obliquaria_reflexa **         
     |                    +--+                                          
     |                       |  +----------- Lampsilis_virescens **        
     |                       |  |                                       
     |                       +--+        +-- Epioblasma_rangiana **        
     |                          |  +-----+                              
     |                          |  |     +-- Obovaria_arkansasensis **     
     |                          +--+                                    
     |                             |  +----- Obovaria_olivaria **          
     |                             +--+                                 
     |                                |  +-- Venustaconcha_ellipsiformis **
     |                                +--+                              
     |                                   +-- Venustaconcha_pleasii **      
     |                                                                  
     |  +----------------------------------- Elliptoideus_sloatianus **    
     |  |                                                               
     |  |  +-------------------------------- Pleurobema_stabile **         
     |  |  |                                                            
     |  |  |                             +-- Pleurobema_clava **           
     |  |  |                 +-----------+                              
     +--+  |                 |           +-- Pleurobema_oviforme **        
        |  |                 |                                          
        |  |  +--------------+  +----------- Pleurobema_pyriforme **       
        |  |  |              |  |                                       
        |  |  |              |  |        +-- Pleurobema_athearni **        
        |  |  |              |  |     +--+                              
        +--+  |              +--+     |  +-- Pleurobema_georgianum **      
           |  |                 |  +--+                                 
           |  |                 |  |  |  +-- Pleurobema_decisum **         
           |  |                 |  |  +--+                              
           |  |                 |  |     +-- Pleurobema_hanleyianum **     
           |  |                 +--+                                    
           |  |                    |  +----- Pleurobema_beadleianum **     
           |  |                    |  |                                 
           |  |                    |  +----- Pleurobema_strodeanum **      
           |  |                    +--+                                 
           +--+                       +----- Pleurobema_rubellum **        
              |                       |                                 
              |                       |  +-- Pleurobema_perovatum **       
              |                       +--+                              
              |                          +-- Pleurobema_taitianum **       
              |                                                         
              |                       +----- Pleurobema_riddellii **       
              |  +--------------------+                                 
              |  |                    |  +-- Pleurobema_sintoxia **        
              |  |                    +--+                              
              |  |                       +-- Pleurobema_cordatum **        
              |  |                                                      
              |  |        +----------------- Fusconaia_subrotunda **       
              |  |        |                                             
              |  |  +-----+              +-- Fusconaia_cor **              
              +--+  |     |  +-----------+                              
                 |  |     |  |           +-- Fusconaia_cuneolus **         
                 |  |     +--+                                          
                 |  |        |  +----------- Fusconaia_ozarkensis **       
                 |  |        |  |                                       
                 |  |        +--+        +-- Fusconaia_escambia **         
                 |  |           |  +-----+                              
                 |  |           |  |     +-- Fusconaia_burkei **           
                 |  |           +--+                                    
                 +--+              |  +----- Fusconaia_masoni **           
                    |              +--+                                 
                    |                 |  +-- Fusconaia_flava **            
                    |                 +--+                              
                    |                    +-- Fusconaia_cerina **           
                    |                                                   
                    |                 +----- Pleuronaia_gibber **          
                    |  +--------------+                                 
                    |  |              |  +-- Pleuronaia_barnesiana **      
                    |  |              +--+                              
                    |  |                 +-- Pleuronaia_dolabelloides **   
                    +--+                                                
                       |                 +-- Eurynia_dilatata **           
                       |  +--------------+                              
                       |  |              +-- Parvaspina_collina **         
                       |  |                                             
                       +--+              +-- Hemistena_lata **             
                          |  +-----------+                              
                          |  |           +-- Plethobasus_cyphyus **        
                          |  |                                          
                          +--+  +----------- Elliptio_shepardiana **       
                             |  |                                       
                             |  |  +-------- Elliptio_complanata **        
                             +--+  |                                    
                                |  |  +----- Elliptio_purpurella **        
                                |  +--+                                 
                                |  |  |  +-- Elliptio_arca **              
                                +--+  +--+                              
                                   |     +-- Elliptio_arctata **           
                                   |                                    
                                   |     +-- Elliptio_mcmichaeli **        
                                   |     |                              
                                   +-----+-- Elliptio_pullata **           
                                         |                              
                                         +-- Elliptio_crassidens **        

Campbell & Lydeard, 2012 [Publ 3309], fig. 3

iT tT bi riS oT roT roS iD
73 73 67 5 0 0 0 72.0
                                      +-- Uniomerus_declivis **         
                       +--------------+                              
                       |              +-- Uniomerus_tetralasmus **      
                       |                                             
  +--------------------+              +-- Quadrula_verrucosa **         
  |                    |  +-----------+                              
  |                    |  |           +-- Quadrula_nobilis **           
  |                    |  |                                          
  |                    +--+  +----------- Quadrula_quadrula **          
  |                       |  |                                       
  |                       |  +----------- Theliderma_metanevra **       
  |                       +--+                                       
  |                          |        +-- Pustulosa_infucata **         
  |                          |  +-----+                              
  |                          |  |     +-- Pustulosa_kleiniana **        
  |                          +--+                                    
  |                             |  +----- Cyclonaias_tuberculata **     
  |                             +--+                                 
  |                                |  +-- Pustulosa_pustulosa **        
  |                                +--+                              
  |                                   +-- Pustulosa_succissa **         
  |                                                                  
  |                                   +-- Amblema_elliottii **          
  |                                +--+                              
  |                                |  +-- Amblema_plicata **            
  |     +--------------------------+                                 
  |     |                          |  +-- Reginaia_ebenus **            
--+     |                          +--+                              
  |     |                             +-- Reginaia_rotulata **          
  |  +--+                                                            
  |  |  |  +----------------------------- Popenaias_popeii **           
  |  |  |  |                                                         
  |  |  |  |                          +-- Glebula_rotundata **          
  |  |  +--+  +-----------------------+                              
  |  |     |  |                       +-- Paetulunio_fabalis **         
  |  |     |  |                                                      
  |  |     +--+                       +-- Toxolasma_lividum **          
  |  |        |  +--------------------+                              
  |  |        |  |                    +-- Plectomerus_dombeyanus **     
  |  |        |  |                                                   
  |  |        +--+  +-------------------- Obliquaria_reflexa **         
  |  |           |  |                                                
  |  |           |  |                 +-- Dromus_dromas **              
  |  |           |  |              +--+                              
  |  |           +--+              |  +-- Lemiox_rimosus **             
  |  |              |  +-----------+                                 
  |  |              |  |           |  +-- Potamilus_alatus **           
  |  |              |  |           +--+                              
  |  |              +--+              +-- Potamilus_capax **            
  |  |                 |                                             
  |  |                 |  +-------------- Lampsilis_ovata **            
  +--+                 |  |                                          
     |                 +--+  +----------- Lampsilis_virescens **        
     |                    |  |                                       
     |                    +--+        +-- Epioblasma_rangiana **        
     |                       |  +-----+                              
     |                       |  |     +-- Obovaria_arkansasensis **     
     |                       +--+                                    
     |                          |  +----- Obovaria_olivaria **          
     |                          +--+                                 
     |                             |  +-- Venustaconcha_ellipsiformis **
     |                             +--+                              
     |                                +-- Venustaconcha_pleasii **      
     |                                                               
     |  +-------------------------------- Elliptoideus_sloatianus **    
     |  |                                                            
     |  |                             +-- Eurynia_dilatata **           
     |  |  +--------------------------+                              
     |  |  |                          +-- Parvaspina_collina **         
     |  |  |                                                         
     |  |  |                          +-- Hemistena_lata **             
     |  |  +--------------------------+                              
     |  |  |                          +-- Plethobasus_cyphyus **        
     +--+  |                                                         
        |  |                 +----------- Elliptio_shepardiana **       
        |  |                 |                                       
        |  |                 |     +----- Elliptio_crassidens **        
        |  +-----------------+  +--+                                 
        |  |                 |  |  |  +-- Elliptio_mcmichaeli **        
        |  |                 |  |  +--+                              
        |  |                 +--+     +-- Elliptio_pullata **           
        |  |                    |                                    
        |  |                    |  +----- Elliptio_complanata **        
        +--+                    |  |                                 
           |                    +--+  +-- Elliptio_purpurella **        
           |                       |  |                              
           |                       +--+-- Elliptio_arca **              
           |                          |                              
           |                          +-- Elliptio_arctata **           
           |                                                         
           |                       +----- Pleuronaia_gibber **          
           |  +--------------------+                                 
           |  |                    |  +-- Pleuronaia_barnesiana **      
           |  |                    +--+                              
           |  |                       +-- Pleuronaia_dolabelloides **   
           |  |                                                      
           |  |           +-------------- Fusconaia_ozarkensis **       
           |  |           |                                          
           |  |           |        +----- Fusconaia_subrotunda **       
           |  |  +--------+  +-----+                                 
           +--+  |        |  |     |  +-- Fusconaia_cor **              
              |  |        |  |     +--+                              
              |  |        |  |        +-- Fusconaia_cuneolus **         
              |  |        +--+                                       
              |  |           |        +-- Fusconaia_escambia **         
              |  |           |  +-----+                              
              |  |           |  |     +-- Fusconaia_burkei **           
              |  |           +--+                                    
              |  |              |  +----- Fusconaia_masoni **           
              +--+              +--+                                 
                 |                 |  +-- Fusconaia_cerina **           
                 |                 +--+                              
                 |                    +-- Fusconaia_flava **            
                 |                                                   
                 |                 +----- Pleurobema_riddellii **       
                 |  +--------------+                                 
                 |  |              |  +-- Pleurobema_sintoxia **        
                 |  |              +--+                              
                 |  |                 +-- Pleurobema_cordatum **        
                 +--+                                                
                    |  +----------------- Pleurobema_stabile **         
                    |  |                                             
                    |  |     +----------- Pleurobema_strodeanum **      
                    |  |  +--+                                       
                    +--+  |  |  +-------- Pleurobema_rubellum **        
                       |  |  +--+                                    
                       |  |     |  +----- Pleurobema_beadleianum **     
                       |  |     +--+                                 
                       |  |        |  +-- Pleurobema_perovatum **       
                       +--+        +--+                              
                          |           +-- Pleurobema_taitianum **       
                          |                                          
                          |           +-- Pleurobema_clava **           
                          |  +--------+                              
                          |  |        +-- Pleurobema_oviforme **        
                          +--+                                       
                             |  +-------- Pleurobema_pyriforme **       
                             |  |                                    
                             +--+     +-- Pleurobema_athearni **        
                                |  +--+                              
                                |  |  +-- Pleurobema_georgianum **      
                                +--+                                 
                                   |  +-- Pleurobema_decisum **         
                                   +--+                              
                                      +-- Pleurobema_hanleyianum **     

Zanatta & Murphy, 2006 [Publ 2672], fig. 2

iT tT bi riS oT roT roS iD
55 55 47 4 2 1 1 55.0
                                            +-------- Pleuronaia_dolabelloides **   
                                            |                                    
  +-----------------------------------------+-------- Pleurobema_clava **           
  |                                         |                                    
  |                                         |  +----- Fusconaia_flava **            
  |                                         +--+                                 
  |                                            |  +-- Hemistena_lata **             
  |                                            +--+                              
  |                                               +-- Eurynia_dilatata **           
--+                                                                              
  |  +----------------------------------------------- Amblema_plicata **            
  |  |                                                                           
  |  +----------------------------------------------- Popenaias_popeii **           
  |  |                                                                           
  |  |                                         +----- Paetulunio_fabalis **         
  |  |                                         |                                 
  |  |  +--------------------------------------+  +-- Toxolasma_parvum **           
  +--+  |                                      |  |                              
     |  |                                      +--+-- Toxolasma_lividum **          
     |  |                                         |                              
     |  |                                         +-- Toxolasma_texasiense **       
     |  |                                                                        
     |  |                                      +----- Obliquaria_reflexa **         
     |  |  +-----------------------------------+                                 
     +--+  |                                   |  +-- Glebula_rotundata **          
        |  |                                   +--+                              
        |  |                                      +-- Cyrtonaias_tampicoensis **    
        |  |                                                                     
        |  |                                   +----- Ptychobranchus_fasciolaris ** 
        |  |                                +--+                                 
        |  |                                |  |  +-- Dromus_dromas **              
        |  |                                |  +--+                              
        +--+  +-----------------------------+     +-- Cyprogenia_stegaria **        
           |  |                             |                                    
           |  |                             |  +----- Medionidus_conradicus **      
           |  |                             +--+                                 
           |  |                                |  +-- Medionidus_acutissimus **     
           |  |                                +--+                              
           |  |                                   +-- Lemiox_rimosus **             
           |  |                                                                  
           |  |  +----------------------------------- Sagittunio_nasutus **         
           |  |  |                                                               
           +--+  |                    +-------------- Potamilus_leptodon **         
              |  |                    |                                          
              |  +--------------------+  +----------- Potamilus_fragilis **         
              |  |                    |  |                                       
              |  |                    +--+        +-- Potamilus_alatus **           
              |  |                       |  +-----+                              
              |  |                       |  |     +-- Potamilus_purpuratus **       
              |  |                       +--+                                    
              |  |                          |  +----- Potamilus_inflatus **         
              |  |                          +--+                                 
              +--+                             |  +-- Potamilus_ohiensis **         
                 |                             +--+                              
                 |                                +-- Potamilus_amphichaenus **     
                 |                                                               
                 |                                +-- Truncilla_truncata **         
                 |  +-----------------------------+                              
                 |  |                             +-- Ellipsaria_lineolata **       
                 |  |                                                            
                 |  |                             +-- Ortmanniana_pectorosa **      
                 |  |                       +-----+                              
                 |  |                       |     +-- Lampsilis_fasciola **         
                 |  |  +--------------------+                                    
                 +--+  |                    |  +----- Lampsilis_ornata **           
                    |  |                    +--+                                 
                    |  |                       |  +-- Lampsilis_ovata **            
                    |  |                       +--+                              
                    |  |                          +-- Lampsilis_cardium **          
                    |  |                                                         
                    |  |                          +-- Ligumia_recta **              
                    +--+                 +--------+                              
                       |                 |        +-- Cambarunio_iris **            
                       |  +--------------+                                       
                       |  |              |  +-------- Lampsilis_teres **            
                       |  |              |  |                                    
                       |  |              +--+     +-- Hamiota_australis **          
                       |  |                 |  +--+                              
                       |  |                 |  |  +-- Hamiota_subangulata **        
                       +--+                 +--+                                 
                          |                    |  +-- Hamiota_altilis **            
                          |                    +--+                              
                          |                       +-- Hamiota_perovalis **          
                          |                                                      
                          |  +----------------------- Leaunio_vanuxemensis **       
                          |  |                                                   
                          +--+  +-------------------- Ortmanniana_ligamentina **    
                             |  |                                                
                             +--+  +----------------- Lampsilis_siliquoidea **      
                                |  |                                             
                                |  |  +-------------- Obovaria_olivaria **          
                                +--+  |                                          
                                   |  +-------------- Venustaconcha_ellipsiformis **
                                   |  |                                          
                                   |  |        +----- Obovaria_unicolor **          
                                   +--+--------+                                 
                                      |        |  +-- Obovaria_subrotunda **        
                                      |        +--+                              
                                      |           +-- Obovaria_arkansasensis **     
                                      |                                          
                                      |  +----------- Epioblasma_brevidens **       
                                      +--+                                       
                                         |  +-------- Epioblasma_triquetra **       
                                         +--+                                    
                                            |  +----- Epioblasma_rangiana **        
                                            +--+                                 
                                               |  +-- Epioblasma_walkeri **         
                                               +--+                              
                                                  +-- Epioblasma_capsaeformis **    

Zanatta & Murphy, 2006 [Publ 2672], fig. 1

iT tT bi riS oT roT roS iD
54 54 49 4 2 1 1 55.0
  +-------------------------------------------------- Pyganodon_grandis          
  |                                                                              
  |                                               +-- Amblema_plicata **            
  |                                            +--+                              
  |                                            |  +-- Popenaias_popeii **           
  |                                            |                                 
--+  +-----------------------------------------+  +-- Pleurobema_clava **           
  |  |                                         |  |                              
  |  |                                         |  +-- Eurynia_dilatata **           
  |  |                                         +--+                              
  |  |                                            +-- Pleuronaia_dolabelloides **   
  |  |                                            |                              
  +--+                                            +-- Fusconaia_flava **            
     |                                                                           
     |                                            +-- Cyrtonaias_tampicoensis **    
     |  +-----------------------------------------+                              
     |  |                                         +-- Glebula_rotundata **          
     |  |                                                                        
     |  |  +----------------------------------------- Paetulunio_fabalis **         
     +--+  |                                                                     
        |  |                                +-------- Obliquaria_reflexa **         
        |  |  +-----------------------------+                                    
        |  |  |                             |  +----- Toxolasma_texasiense **       
        |  |  |                             +--+                                 
        +--+  |                                |  +-- Toxolasma_parvum **           
           |  |                                +--+                              
           |  |                                   +-- Toxolasma_lividum **          
           |  |                                                                  
           |  |  +----------------------------------- Sagittunio_nasutus **         
           |  |  |                                                               
           +--+  |                                +-- Medionidus_conradicus **      
              |  |                          +-----+                              
              |  |                          |     +-- Ptychobranchus_fasciolaris ** 
              |  |                 +--------+                                    
              |  |                 |        |  +----- Medionidus_acutissimus **     
              |  |                 |        |  |                                 
              |  |                 |        +--+----- Lemiox_rimosus **             
              |  |                 |           |                                 
              |  |  +--------------+           |  +-- Dromus_dromas **              
              +--+  |              |           +--+                              
                 |  |              |              +-- Cyprogenia_stegaria **        
                 |  |              |                                             
                 |  |              |  +-------------- Potamilus_leptodon **         
                 |  |              |  |                                          
                 |  |              +--+  +----------- Potamilus_fragilis **         
                 |  |                 |  |                                       
                 |  |                 +--+        +-- Potamilus_alatus **           
                 |  |                    |  +-----+                              
                 |  |                    |  |     +-- Potamilus_purpuratus **       
                 |  |                    +--+                                    
                 +--+                       |  +----- Potamilus_inflatus **         
                    |                       +--+                                 
                    |                          |  +-- Potamilus_ohiensis **         
                    |                          +--+                              
                    |                             +-- Potamilus_amphichaenus **     
                    |                                                            
                    |                          +----- Lampsilis_teres **            
                    |  +-----------------------+                                 
                    |  |                       |  +-- Truncilla_truncata **         
                    |  |                       +--+                              
                    |  |                          +-- Ellipsaria_lineolata **       
                    |  |                                                         
                    |  |  +-------------------------- Leaunio_vanuxemensis **       
                    +--+  |                                                      
                       |  |                       +-- Hamiota_australis **          
                       |  |                    +--+                              
                       |  |                    |  +-- Hamiota_subangulata **        
                       |  |              +-----+                                 
                       |  |              |     |  +-- Hamiota_altilis **            
                       |  |              |     +--+                              
                       +--+              |        +-- Hamiota_perovalis **          
                          |  +-----------+                                       
                          |  |           |        +-- Lampsilis_fasciola **         
                          |  |           |  +-----+                              
                          |  |           |  |     +-- Ortmanniana_pectorosa **      
                          |  |           +--+                                    
                          |  |              |  +----- Lampsilis_ornata **           
                          |  |              +--+                                 
                          |  |                 |  +-- Lampsilis_ovata **            
                          +--+                 +--+                              
                             |                    +-- Lampsilis_cardium **          
                             |                                                   
                             |              +-------- Lampsilis_siliquoidea **      
                             |  +-----------+                                    
                             |  |           |  +----- Ortmanniana_ligamentina **    
                             |  |           +--+                                 
                             |  |              |  +-- Cambarunio_iris **            
                             |  |              +--+                              
                             +--+                 +-- Ligumia_recta **              
                                |                                                
                                |  +----------------- Venustaconcha_ellipsiformis **
                                |  |                                             
                                |  |        +-------- Obovaria_olivaria **          
                                +--+  +-----+                                    
                                   |  |     |  +----- Obovaria_unicolor **          
                                   |  |     +--+                                 
                                   |  |        |  +-- Obovaria_subrotunda **        
                                   +--+        +--+                              
                                      |           +-- Obovaria_arkansasensis **     
                                      |                                          
                                      |  +----------- Epioblasma_brevidens **       
                                      +--+                                       
                                         |  +-------- Epioblasma_triquetra **       
                                         +--+                                    
                                            |  +----- Epioblasma_rangiana **        
                                            +--+                                 
                                               |  +-- Epioblasma_walkeri **         
                                               +--+                              
                                                  +-- Epioblasma_capsaeformis **    

Campbell & Lydeard, 2012 [Publ 3308], fig. 3

iT tT bi riS oT roT roS iD
51 51 42 5 0 0 0 59.0
                                   +-------------- Uniomerus_declivis **      
                                   |                                       
  +--------------------------------+           +-- Pustulosa_infucata **      
  |                                |  +--------+                           
  |                                |  |        +-- Pustulosa_kleiniana **     
  |                                +--+                                    
  |                                   |  +-------- Quadrula_quadrula **       
  |                                   +--+                                 
  |                                      |  +----- Cyclonaias_tuberculata **  
  |                                      +--+                              
  |                                         |  +-- Pustulosa_succissa **      
  |                                         +--+                           
  |                                            +-- Pustulosa_pustulosa **     
--+                                                                        
  |                                +-------------- Elliptoideus_sloatianus ** 
  |  +-----------------------------+                                       
  |  |                             |  +----------- Plectomerus_dombeyanus **  
  |  |                             +--+                                    
  |  |                                |  +-------- Obovaria_unicolor **       
  |  |                                +--+                                 
  |  |                                   |  +----- Obovaria_olivaria **       
  |  |                                   +--+                              
  |  |                                      |  +-- Lampsilis_ovata **         
  |  |                                      +--+                           
  |  |                                         +-- Lemiox_rimosus **          
  +--+                                                                     
     |                                +----------- Cyrtonaias_tampicoensis ** 
     |                                |                                    
     |                                |     +----- Popenaias_popeii **        
     |  +-----------------------------+  +--+                              
     |  |                             |  |  |  +-- Amblema_elliottii **       
     |  |                             |  |  +--+                           
     |  |                             +--+     +-- Amblema_plicata **         
     |  |                                |                                 
     |  |                                |     +-- Reginaia_ebenus[1] **      
     |  |                                |     |                           
     +--+                                +-----+-- Reginaia_ebenus[2] **      
        |                                      |                           
        |                                      +-- Reginaia_rotulata **       
        |                                                                  
        |                                      +-- Elliptio_arca **           
        |  +-----------------------------------+                           
        |  |                                   +-- Elliptio_crassidens **     
        |  |                                                               
        +--+  +----------------------------------- Eurynia_dilatata **        
           |  |                                                            
           |  |  +-------------------------------- Parvaspina_collina **      
           |  |  |                                                         
           +--+  |                             +-- Hemistena_lata **          
              |  +-----------------------------+                           
              |  |                             +-- Plethobasus_cyphyus **     
              |  |                                                         
              +--+                          +----- Pleuronaia_barnesiana **   
                 |  +-----------------------+                              
                 |  |                       |  +-- Pleuronaia_dolabelloides **
                 |  |                       +--+                           
                 |  |                          +-- Pleuronaia_gibber **       
                 |  |                                                      
                 |  |                          +-- Pleurobema_stabile **      
                 +--+                    +-----+                           
                    |                    |     +-- Pleurobema_sintoxia **     
                    |  +-----------------+                                 
                    |  |                 |  +----- Pleurobema_decisum **      
                    |  |                 +--+                              
                    |  |                    |  +-- Pleurobema_clava **        
                    +--+                    +--+                           
                       |                       +-- Pleurobema_pyriforme **    
                       |                                                   
                       |  +----------------------- Pleurobema_cordatum **     
                       |  |                                                
                       +--+  +-------------------- Fusconaia_subrotunda **    
                          |  |                                             
                          |  |                 +-- Fusconaia_cor **           
                          +--+  +--------------+                           
                             |  |              +-- Fusconaia_cuneolus **      
                             |  |                                          
                             +--+  +-------------- Fusconaia_ozarkensis **    
                                |  |                                       
                                |  |           +-- Fusconaia_flava[6] **      
                                |  |  +--------+                           
                                +--+  |        +-- Fusconaia_masoni **        
                                   |  |                                    
                                   |  |        +-- Fusconaia_escambia **      
                                   |  |        |                           
                                   +--+        +-- Fusconaia_burkei[1] **     
                                      |  +-----+                           
                                      |  |     +-- Fusconaia_burkei[2] **     
                                      |  |     |                           
                                      |  |     +-- Fusconaia_burkei[3] **     
                                      +--+                                 
                                         |  +----- Fusconaia_flava[5] **      
                                         |  |                              
                                         |  |  +-- Fusconaia_cerina **        
                                         +--+  |                           
                                            |  +-- Fusconaia_flava[1] **      
                                            |  |                           
                                            +--+-- Fusconaia_flava[2] **      
                                               |                           
                                               +-- Fusconaia_flava[3] **      
                                               |                           
                                               +-- Fusconaia_flava[4] **      

Burlakova et al., 2012 [Publ 3486], fig. 1

iT tT bi riS oT roT roS iD
44 44 42 4 0 0 0 57.0
                                               +-- Amblema_elliottii **       
  +--------------------------------------------+                           
  |                                            +-- Amblema_plicata **         
  |                                                                        
--+                                            +-- Lampsilis_ovata **         
  |  +-----------------------------------------+                           
  |  |                                         +-- Plectomerus_dombeyanus **  
  |  |                                                                     
  +--+  +----------------------------------------- Popenaias_popeii **        
     |  |                                                                  
     |  |  +-------------------------------------- Cyrtonaias_tampicoensis ** 
     +--+  |                                                               
        |  |  +----------------------------------- Parvaspina_collina **      
        |  |  |                                                            
        +--+  |                             +----- Plethobasus_cyphyus **     
           |  |  +--------------------------+                              
           |  |  |                          |  +-- Elliptio_arca **           
           |  |  |                          +--+                           
           |  |  |                             +-- Elliptio_crassidens **     
           +--+  |                                                         
              |  |           +-------------------- Fusconaia_subrotunda **    
              |  |           |                                             
              |  |           |                 +-- Fusconaia_cor **           
              |  |  +--------+  +--------------+                           
              |  |  |        |  |              +-- Fusconaia_cuneolus **      
              |  |  |        |  |                                          
              +--+  |        +--+  +-------------- Fusconaia_ozarkensis **    
                 |  |           |  |                                       
                 |  |           |  |     +-------- Fusconaia_askewi[3] **     
                 |  |           |  |  +--+                                 
                 |  |           +--+  |  |  +----- Fusconaia_askewi[2] **     
                 |  |              |  |  +--+                              
                 |  |              |  |     |  +-- Fusconaia_lananensis **    
                 |  |              |  |     +--+                           
                 |  |              +--+        +-- Fusconaia_askewi[1] **     
                 |  |                 |                                    
                 |  |                 |        +-- Fusconaia_burkei **        
                 |  |                 |  +-----+                           
                 +--+                 |  |     +-- Fusconaia_escambia **      
                    |                 +--+                                 
                    |                    |  +----- Fusconaia_masoni **        
                    |                    +--+                              
                    |                       |  +-- Fusconaia_flava **         
                    |                       +--+                           
                    |                          +-- Fusconaia_cerina **        
                    |                                                      
                    |                          +-- Eurynia_dilatata **        
                    |  +-----------------------+                           
                    |  |                       +-- Hemistena_lata **          
                    |  |                                                   
                    |  |                    +----- Pleuronaia_gibber **       
                    |  |              +-----+                              
                    |  |              |     |  +-- Pleuronaia_dolabelloides **
                    +--+              |     +--+                           
                       |  +-----------+        +-- Pleuronaia_barnesiana **   
                       |  |           |                                    
                       |  |           |  +-------- Pleurobema_riddellii **    
                       |  |           +--+                                 
                       |  |              |  +----- Pleurobema_sintoxia[2] **  
                       |  |              +--+                              
                       +--+                 |  +-- Pleurobema_sintoxia[1] **  
                          |                 +--+                           
                          |                    +-- Pleurobema_cordatum **     
                          |                                                
                          |  +-------------------- Pleurobema_stabile **      
                          |  |                                             
                          +--+  +----------------- Pleurobema_pyriforme **    
                             |  |                                          
                             |  |           +----- Pleurobema_georgianum **   
                             +--+  +--------+                              
                                |  |        |  +-- Pleurobema_hanleyianum **  
                                |  |        +--+                           
                                |  |           +-- Pleurobema_decisum **      
                                +--+                                       
                                   |           +-- Pleurobema_clava **        
                                   |  +--------+                           
                                   |  |        +-- Pleurobema_oviforme **     
                                   +--+                                    
                                      |  +-------- Pleurobema_beadleianum **  
                                      |  |                                 
                                      +--+     +-- Pleurobema_rubellum **     
                                         |  +--+                           
                                         |  |  +-- Pleurobema_strodeanum **   
                                         +--+                              
                                            |  +-- Pleurobema_perovatum **    
                                            +--+                           
                                               +-- Pleurobema_taitianum **    

Campbell & Lydeard, 2012 [Publ 3308], fig. 1

iT tT bi riS oT roT roS iD
43 43 41 5 0 0 0 52.0
                                      +----------- Quadrula_quadrula **       
                                      |                                    
  +-----------------------------------+        +-- Pustulosa_infucata **      
  |                                   |  +-----+                           
  |                                   |  |     +-- Pustulosa_kleiniana **     
  |                                   +--+                                 
  |                                      |  +----- Cyclonaias_tuberculata **  
  |                                      +--+                              
  |                                         |  +-- Pustulosa_succissa **      
  |                                         +--+                           
  |                                            +-- Pustulosa_pustulosa **     
  |                                                                        
  |                                         +----- Popenaias_popeii **        
  |                             +-----------+                              
--+                             |           |  +-- Amblema_elliottii **       
  |                             |           +--+                           
  |                             |              +-- Amblema_plicata **         
  |  +--------------------------+                                          
  |  |                          |           +----- Cyrtonaias_tampicoensis ** 
  |  |                          |  +--------+                              
  |  |                          |  |        |  +-- Reginaia_ebenus **         
  |  |                          |  |        +--+                           
  |  |                          +--+           +-- Reginaia_rotulata **       
  |  |                             |                                       
  |  |                             |  +----------- Plectomerus_dombeyanus **  
  |  |                             +--+                                    
  |  |                                |  +-------- Lemiox_rimosus **          
  +--+                                +--+                                 
     |                                   |  +----- Lampsilis_ovata **         
     |                                   +--+                              
     |                                      |  +-- Obovaria_unicolor **       
     |                                      +--+                           
     |                                         +-- Obovaria_olivaria **       
     |                                                                     
     |  +----------------------------------------- Elliptoideus_sloatianus ** 
     |  |                                                                  
     |  |  +-------------------------------------- Pleurobema_stabile **      
     +--+  |                                                               
        |  |                                +----- Pleurobema_pyriforme **    
        |  |  +-----------------------------+                              
        +--+  |                             |  +-- Pleurobema_clava **        
           |  |                             +--+                           
           |  |                                +-- Pleurobema_decisum **      
           |  |                                                            
           +--+                                +-- Pleurobema_cordatum **     
              |  +-----------------------------+                           
              |  |                             +-- Pleurobema_sintoxia **     
              |  |                                                         
              |  |                             +-- Eurynia_dilatata **        
              |  |                       +-----+                           
              +--+                       |     +-- Hemistena_lata **          
                 |  +--------------------+                                 
                 |  |                    |  +----- Plethobasus_cyphyus **     
                 |  |                    +--+                              
                 |  |                       |  +-- Elliptio_arca **           
                 |  |                       +--+                           
                 +--+                          +-- Elliptio_crassidens **     
                    |                                                      
                    |                       +----- Pleuronaia_gibber **       
                    |  +--------------------+                              
                    |  |                    |  +-- Pleuronaia_barnesiana **   
                    |  |                    +--+                           
                    +--+                       +-- Pleuronaia_dolabelloides **
                       |                                                   
                       |  +----------------------- Parvaspina_collina **      
                       |  |                                                
                       +--+  +-------------------- Fusconaia_subrotunda **    
                          |  |                                             
                          +--+                 +-- Fusconaia_cor **           
                             |  +--------------+                           
                             |  |              +-- Fusconaia_cuneolus **      
                             +--+                                          
                                |  +-------------- Fusconaia_ozarkensis **    
                                |  |                                       
                                +--+           +-- Fusconaia_burkei **        
                                   |  +--------+                           
                                   |  |        +-- Fusconaia_escambia **      
                                   +--+                                    
                                      |  +-------- Fusconaia_masoni **        
                                      +--+                                 
                                         |  +----- Fusconaia_cerina[2] **     
                                         +--+                              
                                            |  +-- Fusconaia_flava **         
                                            +--+                           
                                               +-- Fusconaia_cerina[1] **     

Campbell & Lydeard, 2012 [Publ 3308], fig. 2

iT tT bi riS oT roT roS iD
43 43 36 5 0 0 0 50.0
                       +----------- Quadrula_quadrula **       
                       |                                    
  +--------------------+        +-- Pustulosa_infucata **      
  |                    |  +-----+                           
  |                    |  |     +-- Pustulosa_kleiniana **     
  |                    +--+                                 
  |                       |  +----- Cyclonaias_tuberculata **  
  |                       +--+                              
  |                          |  +-- Pustulosa_succissa **      
  |                          +--+                           
  |                             +-- Pustulosa_pustulosa **     
  |                                                         
  |                             +-- Reginaia_ebenus **         
  |                          +--+                           
  |                          |  +-- Reginaia_rotulata **       
--+              +-----------+                              
  |              |           |  +-- Amblema_elliottii **       
  |              |           +--+                           
  |  +-----------+              +-- Amblema_plicata **         
  |  |           |                                          
  |  |           |  +-------------- Popenaias_popeii **        
  |  |           |  |                                       
  |  |           +--+  +----------- Cyrtonaias_tampicoensis ** 
  |  |              |  |                                    
  |  |              +--+----------- Plectomerus_dombeyanus **  
  |  |                 |                                    
  |  |                 |  +-------- Lemiox_rimosus **          
  |  |                 +--+                                 
  |  |                    |  +----- Lampsilis_ovata **         
  +--+                    +--+                              
     |                       |  +-- Obovaria_unicolor **       
     |                       +--+                           
     |                          +-- Obovaria_olivaria **       
     |                                                      
     |  +-------------------------- Elliptoideus_sloatianus ** 
     |  |                                                   
     |  |  +----------------------- Hemistena_lata **          
     |  |  |                                                
     |  |  +----------------------- Plethobasus_cyphyus **     
     |  |  |                                                
     +--+  |                    +-- Elliptio_arca **           
        |  +--------------------+                           
        |  |                    +-- Elliptio_crassidens **     
        |  |                                                
        |  |                    +-- Eurynia_dilatata **        
        |  +--------------------+                           
        |  |                    +-- Parvaspina_collina **      
        +--+                                                
           |                 +----- Pleuronaia_gibber **       
           |  +--------------+                              
           |  |              |  +-- Pleuronaia_barnesiana **   
           |  |              +--+                           
           |  |                 +-- Pleuronaia_dolabelloides **
           |  |                                             
           |  |                 +-- Pleurobema_cordatum **     
           |  |        +--------+                           
           |  |        |        +-- Pleurobema_sintoxia **     
           |  +--------+                                    
           |  |        |  +-------- Pleurobema_stabile **      
           +--+        +--+                                 
              |           |  +----- Pleurobema_pyriforme **    
              |           +--+                              
              |              |  +-- Pleurobema_clava **        
              |              +--+                           
              |                 +-- Pleurobema_decisum **      
              |                                             
              |              +----- Fusconaia_subrotunda **    
              |  +-----------+                              
              |  |           |  +-- Fusconaia_cor **           
              |  |           +--+                           
              +--+              +-- Fusconaia_cuneolus **      
                 |                                          
                 |  +-------------- Fusconaia_ozarkensis **    
                 |  |                                       
                 +--+           +-- Fusconaia_burkei **        
                    |  +--------+                           
                    |  |        +-- Fusconaia_escambia **      
                    +--+                                    
                       |  +-------- Fusconaia_masoni **        
                       +--+                                 
                          |  +----- Fusconaia_cerina[2] **     
                          +--+                              
                             |  +-- Fusconaia_flava **         
                             +--+                           
                                +-- Fusconaia_cerina[1] **     

Campbell & Lydeard, 2012 [Publ 3308], fig. 4

iT tT bi riS oT roT roS iD
43 43 40 5 0 0 0 52.0
                                         +----------- Quadrula_quadrula **       
                                         |                                    
  +--------------------------------------+        +-- Pustulosa_infucata **      
  |                                      |  +-----+                           
  |                                      |  |     +-- Pustulosa_kleiniana **     
  |                                      +--+                                 
  |                                         |  +----- Cyclonaias_tuberculata **  
  |                                         +--+                              
  |                                            |  +-- Pustulosa_succissa **      
  |                                            +--+                           
  |                                               +-- Pustulosa_pustulosa **     
  |                                                                           
  |                                            +----- Popenaias_popeii **        
  |                                   +--------+                              
--+                                   |        |  +-- Amblema_elliottii **       
  |                                   |        +--+                           
  |                                   |           +-- Amblema_plicata **         
  |  +--------------------------------+                                       
  |  |                                |  +----------- Plectomerus_dombeyanus **  
  |  |                                |  |                                    
  |  |                                |  |     +----- Cyrtonaias_tampicoensis ** 
  |  |                                |  +-----+                              
  |  |                                +--+     |  +-- Reginaia_ebenus **         
  |  |                                   |     +--+                           
  |  |                                   |        +-- Reginaia_rotulata **       
  |  |                                   |                                    
  +--+                                   |  +-------- Lemiox_rimosus **          
     |                                   +--+                                 
     |                                      |  +----- Lampsilis_ovata **         
     |                                      +--+                              
     |                                         |  +-- Obovaria_unicolor **       
     |                                         +--+                           
     |                                            +-- Obovaria_olivaria **       
     |                                                                        
     |  +-------------------------------------------- Elliptoideus_sloatianus ** 
     |  |                                                                     
     +--+  +----------------------------------------- Pleurobema_stabile **      
        |  |                                                                  
        +--+  +-------------------------------------- Hemistena_lata **          
           |  |                                                               
           +--+  +----------------------------------- Eurynia_dilatata **        
              |  |                                                            
              |  |                                +-- Pleurobema_cordatum **     
              +--+  +-----------------------------+                           
                 |  |                             +-- Pleurobema_sintoxia **     
                 |  |                                                         
                 |  |                          +----- Pleurobema_clava **        
                 +--+  +-----------------------+                              
                    |  |                       |  +-- Pleurobema_decisum **      
                    |  |                       +--+                           
                    |  |                          +-- Pleurobema_pyriforme **    
                    |  |                                                      
                    +--+                    +-------- Parvaspina_collina **      
                       |  +-----------------+                                 
                       |  |                 |  +----- Pleuronaia_gibber **       
                       |  |                 +--+                              
                       |  |                    |  +-- Pleuronaia_barnesiana **   
                       |  |                    +--+                           
                       +--+                       +-- Pleuronaia_dolabelloides **
                          |                                                   
                          |                    +----- Plethobasus_cyphyus **     
                          |  +-----------------+                              
                          |  |                 |  +-- Elliptio_arca **           
                          |  |                 +--+                           
                          +--+                    +-- Elliptio_crassidens **     
                             |                                                
                             |  +-------------------- Fusconaia_subrotunda **    
                             |  |                                             
                             +--+                 +-- Fusconaia_cor **           
                                |  +--------------+                           
                                |  |              +-- Fusconaia_cuneolus **      
                                +--+                                          
                                   |  +-------------- Fusconaia_ozarkensis **    
                                   |  |                                       
                                   +--+           +-- Fusconaia_burkei **        
                                      |  +--------+                           
                                      |  |        +-- Fusconaia_escambia **      
                                      +--+                                    
                                         |  +-------- Fusconaia_masoni **        
                                         +--+                                 
                                            |  +----- Fusconaia_cerina[2] **     
                                            +--+                              
                                               |  +-- Fusconaia_flava **         
                                               +--+                           
                                                  +-- Fusconaia_cerina[1] **     

Burlakova et al., 2012 [Publ 3486], fig. 3

iT tT bi riS oT roT roS iD
43 43 26 2 0 0 0 51.0
  +----------------------------------- Plectomerus_dombeyanus **  
  |                                                            
  |  +-------------------------------- Hemistena_lata **          
  |  |                                                         
--+  +-------------------------------- Pleurobema_stabile **      
  |  |                                                         
  |  |  +----------------------------- Eurynia_dilatata **        
  |  |  |                                                      
  +--+  |                    +-------- Pleurobema_riddellii **    
     |  |                    |                                 
     |  |  +-----------------+  +----- Pleurobema_sintoxia[2] **  
     |  |  |                 |  |                              
     |  |  |                 +--+----- Pleurobema_sintoxia[3] **  
     |  |  |                    |                              
     +--+  |                    |  +-- Pleurobema_cordatum **     
        |  |                    +--+                           
        |  |                       +-- Pleurobema_sintoxia[1] **  
        |  |                                                   
        |  |  +----------------------- Parvaspina_collina **      
        |  |  |                                                
        |  |  |                 +----- Pleuronaia_gibber **       
        |  |  +-----------------+                              
        |  |  |                 |  +-- Pleuronaia_dolabelloides **
        +--+  |                 +--+                           
           |  |                    +-- Pleuronaia_barnesiana **   
           |  |                                                
           |  |                 +----- Plethobasus_cyphyus **     
           |  +-----------------+                              
           |  |                 |  +-- Elliptio_arca **           
           |  |                 +--+                           
           |  |                    +-- Elliptio_crassidens **     
           |  |                                                
           |  |                    +-- Pleurobema_clava **        
           |  |           +--------+                           
           |  |           |        +-- Pleurobema_oviforme **     
           |  |           |                                    
           |  |           |  +-------- Pleurobema_decisum **      
           +--+-----------+  |                                 
              |           |  +-------- Pleurobema_pyriforme **    
              |           |  |                                 
              |           |  +-------- Pleurobema_hanleyianum **  
              |           +--+                                 
              |              +-------- Pleurobema_georgianum **   
              |              |                                 
              |              |  +----- Pleurobema_rubellum **     
              |              |  |                              
              |              |  +----- Pleurobema_beadleianum **  
              |              +--+                              
              |                 +----- Pleurobema_strodeanum **   
              |                 |                              
              |                 |  +-- Pleurobema_perovatum **    
              |                 +--+                           
              |                    +-- Pleurobema_taitianum **    
              |                                                
              |  +-------------------- Fusconaia_subrotunda **    
              |  |                                             
              |  |                 +-- Fusconaia_cor **           
              +--+  +--------------+                           
                 |  |              +-- Fusconaia_cuneolus **      
                 |  |                                          
                 |  |  +-------------- Fusconaia_ozarkensis **    
                 +--+  |                                       
                    |  |     +-------- Fusconaia_masoni **        
                    |  |     |                                 
                    |  |     |     +-- Fusconaia_burkei **        
                    |  |  +--+-----+                           
                    +--+  |  |     +-- Fusconaia_escambia **      
                       |  |  |                                 
                       |  |  |  +----- Fusconaia_cerina **        
                       |  |  +--+                              
                       |  |     |  +-- Fusconaia_askewi[1] **     
                       |  |     +--+                           
                       +--+        +-- Fusconaia_flava **         
                          |                                    
                          |        +-- Fusconaia_askewi[2] **     
                          |        |                           
                          |        +-- Fusconaia_askewi[3] **     
                          |        |                           
                          |        +-- Fusconaia_askewi[4] **     
                          +--------+                           
                                   +-- Fusconaia_lananensis[1] ** 
                                   |                           
                                   +-- Fusconaia_lananensis[2] ** 
                                   |                           
                                   +-- Fusconaia_askewi[5] **     

Inoue et al., 2020 [Publ 3941], fig. 3

iT tT bi riS oT roT roS iD
43 43 41 4 1 1 1 43.0
  +----------------------------------------------- Arcidens_confragosus    
  |                                                                        
  |                                            +-- Pustulosa_petrina **       
  |  +-----------------------------------------+                           
  |  |                                         +-- Quadrula_quadrula **       
--+  |                                                                     
  |  |                             +-------------- Parvaspina_collina **      
  |  |                             |                                       
  |  |                             |           +-- Eurynia_dilatata **        
  |  |  +--------------------------+  +--------+                           
  |  |  |                          |  |        +-- Hemistena_lata **          
  +--+  |                          |  |                                    
     |  |                          +--+        +-- Pleurobema_clava **        
     |  |                             |  +-----+                           
     |  |                             |  |     +-- Pleurobema_sintoxia **     
     |  |                             +--+                                 
     |  |                                |  +----- Pleuronaia_dolabelloides **
     |  |                                +--+                              
     |  |                                   |  +-- Fusconaia_flava **         
     +--+                                   +--+                           
        |                                      +-- Fusconaia_subrotunda **    
        |                                                                  
        |                                      +-- Amblema_plicata **         
        |                                   +--+                           
        |                                   |  +-- Amblema_elliottii **       
        |  +--------------------------------+                              
        |  |                                |  +-- Reginaia_ebenus **         
        |  |                                +--+                           
        |  |                                   +-- Reginaia_rotulata **       
        |  |                                                               
        +--+  +----------------------------------- Toxolasma_lividum **       
           |  |                                                            
           |  |                          +-------- Potamilus_fragilis **      
           |  |                          |                                 
           |  |  +-----------------------+     +-- Potamilus_ohiensis **      
           +--+  |                       |  +--+                           
              |  |                       |  |  +-- Potamilus_amphichaenus **  
              |  |                       +--+                              
              |  |                          |  +-- Potamilus_metnecktayi **   
              |  |                          +--+                           
              |  |                             +-- Potamilus_purpuratus **    
              +--+                                                         
                 |                       +-------- Ellipsaria_lineolata **    
                 |  +--------------------+                                 
                 |  |                    |  +----- Truncilla_macrodon **      
                 |  |                    +--+                              
                 |  |                       |  +-- Truncilla_donaciformis **  
                 |  |                       +--+                           
                 |  |                          +-- Truncilla_cognata **       
                 +--+                                                      
                    |  +-------------------------- Leaunio_lienosus **        
                    |  |                                                   
                    |  |     +-------------------- Lampsilis_bracteata **     
                    |  |     |                                             
                    |  |  +--+  +----------------- Lampsilis_fasciola **      
                    |  |  |  |  |                                          
                    +--+  |  +--+  +-------------- Lampsilis_ornata **        
                       |  |     |  |                                       
                       |  |     +--+           +-- Lampsilis_ovata **         
                       |  |        |  +--------+                           
                       |  |        |  |        +-- Lampsilis_cariosa **       
                       |  |        +--+                                    
                       |  |           |  +-------- Lampsilis_cardium[3] **    
                       +--+           +--+                                 
                          |              |  +----- Lampsilis_cardium[2] **    
                          |              +--+                              
                          |                 |  +-- Lampsilis_satura **        
                          |                 +--+                           
                          |                    +-- Lampsilis_cardium[1] **    
                          |                                                
                          |                    +-- Lampsilis_higginsii **     
                          |     +--------------+                           
                          |     |              +-- Ortmanniana_ligamentina ** 
                          +-----+                                          
                                |  +-------------- Lampsilis_teres **         
                                |  |                                       
                                +--+           +-- Lampsilis_radiata **       
                                   |  +--------+                           
                                   |  |        +-- Lampsilis_siliquoidea **   
                                   +--+                                    
                                      |  +-------- Lampsilis_bergmanni **     
                                      +--+                                 
                                         |  +----- Lampsilis_hydiana **       
                                         +--+                              
                                            |  +-- Lampsilis_virescens **     
                                            +--+                           
                                               +-- Lampsilis_straminea **     

Inoue et al., 2020 [Publ 3941], fig. 4

iT tT bi riS oT roT roS iD
42 42 40 4 1 1 1 44.0
  +-------------------------------------------------- Arcidens_confragosus    
  |                                                                           
  |                                               +-- Pustulosa_petrina **       
  |  +--------------------------------------------+                           
--+  |                                            +-- Quadrula_quadrula **       
  |  |                                                                        
  |  |                                            +-- Reginaia_ebenus **         
  |  |  +-----------------------------------------+                           
  +--+  |                                         +-- Reginaia_rotulata **       
     |  |                                                                     
     |  |                             +-------------- Parvaspina_collina **      
     |  |                             |                                       
     |  |                             |           +-- Eurynia_dilatata **        
     |  |  +--------------------------+  +--------+                           
     +--+  |                          |  |        +-- Hemistena_lata **          
        |  |                          |  |                                    
        |  |                          +--+        +-- Pleurobema_clava **        
        |  |                             |  +-----+                           
        |  |                             |  |     +-- Pleurobema_sintoxia **     
        |  |                             +--+                                 
        |  |                                |  +----- Pleuronaia_dolabelloides **
        +--+                                +--+                              
           |                                   |  +-- Fusconaia_flava **         
           |                                   +--+                           
           |                                      +-- Fusconaia_subrotunda **    
           |                                                                  
           |                                      +-- Amblema_plicata **         
           |  +-----------------------------------+                           
           |  |                                   +-- Amblema_elliottii **       
           |  |                                                               
           +--+  +----------------------------------- Toxolasma_lividum **       
              |  |                                                            
              |  |  +-------------------------------- Ellipsaria_lineolata **    
              +--+  |                                                         
                 |  |                          +----- Truncilla_cognata **       
                 |  |  +-----------------------+                              
                 |  |  |                       |  +-- Truncilla_macrodon **      
                 +--+  |                       +--+                           
                    |  |                          +-- Truncilla_donaciformis **  
                    |  |                                                      
                    |  |                    +-------- Potamilus_fragilis **      
                    |  |                    |                                 
                    +--+  +-----------------+     +-- Potamilus_ohiensis **      
                       |  |                 |  +--+                           
                       |  |                 |  |  +-- Potamilus_amphichaenus **  
                       |  |                 +--+                              
                       |  |                    |  +-- Potamilus_metnecktayi **   
                       |  |                    +--+                           
                       |  |                       +-- Potamilus_purpuratus **    
                       +--+                                                   
                          |  +----------------------- Leaunio_lienosus **        
                          |  |                                                
                          |  |     +----------------- Lampsilis_bracteata **     
                          |  |     |                                          
                          |  |  +--+              +-- Lampsilis_ornata **        
                          |  |  |  |  +-----------+                           
                          +--+  |  |  |           +-- Lampsilis_fasciola **      
                             |  |  +--+                                       
                             |  |     |  +----------- Lampsilis_ovata **         
                             |  |     +--+                                    
                             |  |        |  +-------- Lampsilis_cardium[3] **    
                             |  |        +--+                                 
                             +--+           |  +----- Lampsilis_cardium[2] **    
                                |           +--+                              
                                |              |  +-- Lampsilis_satura **        
                                |              +--+                           
                                |                 +-- Lampsilis_cardium[1] **    
                                |                                             
                                |     +-------------- Lampsilis_teres **         
                                |     |                                       
                                |     |           +-- Lampsilis_higginsii **     
                                +-----+  +--------+                           
                                      |  |        +-- Ortmanniana_ligamentina ** 
                                      |  |                                    
                                      +--+        +-- Lampsilis_radiata **       
                                         |  +-----+                           
                                         |  |     +-- Lampsilis_siliquoidea **   
                                         |  |                                 
                                         +--+     +-- Lampsilis_hydiana **       
                                            |  +--+                           
                                            |  |  +-- Lampsilis_bergmanni **     
                                            +--+                              
                                               |  +-- Lampsilis_virescens **     
                                               +--+                           
                                                  +-- Lampsilis_straminea **     

Smith et al., 2019 [Publ 3951], fig. 2

iT tT bi riS oT roT roS iD
42 42 40 4 0 0 0 43.0
  +----------------------------------------------- Quadrula_quadrula **          
  |                                                                           
  |                                            +-- Amblema_plicata **            
--+  +-----------------------------------------+                              
  |  |                                         +-- Reginaia_ebenus **            
  |  |                                                                        
  +--+  +----------------------------------------- Plectomerus_dombeyanus **     
     |  |                                                                     
     |  |  +-------------------------------------- Toxolasma_lividum **          
     +--+  |                                                                  
        |  |                                +----- Popenaias_popeii **           
        |  |  +-----------------------------+                                 
        +--+  |                             |  +-- Glebula_rotundata **          
           |  |                             +--+                              
           |  |                                +-- Cyrtonaias_tampicoensis **    
           |  |                                                               
           |  |  +-------------------------------- Obliquaria_reflexa **         
           +--+  |                                                            
              |  |                             +-- Medionidus_conradicus **      
              |  |                          +--+                              
              |  |                          |  +-- Lemiox_rimosus **             
              |  |  +-----------------------+                                 
              |  |  |                       |  +-- Cyprogenia_stegaria **        
              |  |  |                       +--+                              
              +--+  |                          +-- Ptychobranchus_fasciolaris ** 
                 |  |                                                         
                 |  |                          +-- Hamiota_subangulata **        
                 |  |     +--------------------+                              
                 |  |     |                    +-- Villosa_villosa **            
                 |  |     |                                                   
                 |  |  +--+  +-------------------- Ligumia_recta **              
                 |  |  |  |  |                                                
                 +--+  |  |  |              +----- Venustaconcha_ellipsiformis **
                    |  |  +--+  +-----------+                                 
                    |  |     |  |           |  +-- Obovaria_subrotunda **        
                    |  |     |  |           +--+                              
                    |  |     +--+              +-- Epioblasma_rangiana **        
                    |  |        |                                             
                    |  |        |  +-------------- Ortmanniana_pectorosa **      
                    |  |        +--+                                          
                    |  |           |  +----------- Ortmanniana_ligamentina **    
                    |  |           +--+                                       
                    |  |              |  +-------- Lampsilis_ovata **            
                    |  |              +--+                                    
                    +--+                 |  +----- Lampsilis_dolabraeformis **   
                       |                 +--+                                 
                       |                    |  +-- Lampsilis_satura **           
                       |                    +--+                              
                       |                       +-- Lampsilis_cardium **          
                       |                                                      
                       |           +-------------- Atlanticoncha_ochracea **     
                       |     +-----+                                          
                       |     |     |  +----------- Ellipsaria_lineolata **       
                       |     |     +--+                                       
                       |     |        |  +-------- Truncilla_truncata **         
                       |     |        +--+                                    
                       |     |           |  +----- Truncilla_cognata **          
                       |     |           +--+                                 
                       +-----+              |  +-- Truncilla_macrodon **         
                             |              +--+                              
                             |                 +-- Truncilla_donaciformis **     
                             |                                                
                             |  +----------------- Potamilus_capax **            
                             |  |                                             
                             |  |           +----- Potamilus_inflatus **         
                             +--+  +--------+                                 
                                |  |        |  +-- Potamilus_fragilis **         
                                |  |        +--+                              
                                |  |           +-- Potamilus_leptodon **         
                                +--+                                          
                                   |        +----- Potamilus_ohiensis **         
                                   |  +-----+                                 
                                   |  |     |  +-- Potamilus_amphichaenus **     
                                   |  |     +--+                              
                                   +--+        +-- Potamilus_streckersoni **     
                                      |                                       
                                      |  +-------- Potamilus_metnecktayi **      
                                      +--+                                    
                                         |  +----- Potamilus_purpuratus[1] **    
                                         +--+                                 
                                            |  +-- Potamilus_purpuratus[2] **    
                                            +--+                              
                                               +-- Potamilus_alatus **           

Burlakova et al., 2012 [Publ 3486], fig. 2

iT tT bi riS oT roT roS iD
40 40 30 1 0 0 0 53.0
  +----------------------------------- Parvaspina_collina **      
  |                                                            
  |  +-------------------------------- Pleurobema_plenum **       
  |  |                                                         
--+  |                             +-- Plethobasus_cyphyus **     
  |  |                       +-----+                           
  |  |                       |     +-- Hemistena_lata **          
  |  |  +--------------------+                                 
  +--+  |                    |  +----- Eurynia_dilatata **        
     |  |                    +--+                              
     |  |                       |  +-- Elliptio_arca **           
     |  |                       +--+                           
     |  |                          +-- Elliptio_crassidens **     
     |  |                                                      
     |  |  +-------------------------- Pleurobema_cordatum[2] **  
     +--+  |                                                   
        |  |           +-------------- Fusconaia_subrotunda **    
        |  |           |                                       
        |  |           |           +-- Fusconaia_cor **           
        |  |           |  +--------+                           
        |  |  +--------+  |        +-- Fusconaia_cuneolus **      
        |  |  |        |  |                                    
        |  |  |        |  |  +-------- Fusconaia_flava[2] **      
        |  |  |        |  |  |                                 
        +--+  |        +--+  +-------- Fusconaia_ozarkensis **    
           |  |           |  |                                 
           |  |           |  +-------- Fusconaia_masoni **        
           |  |           |  |                                 
           |  |           |  |  +----- Fusconaia_askewi **        
           |  |           +--+--+                              
           |  |              |  |  +-- Fusconaia_burkei **        
           |  |              |  +--+                           
           |  |              |     +-- Fusconaia_escambia **      
           |  |              |                                 
           +--+              |     +-- Fusconaia_flava[1] **      
              |              |     |                           
              |              +-----+-- Fusconaia_cerina[1] **     
              |                    |                           
              |                    +-- Fusconaia_cerina[2] **     
              |                                                
              |              +-------- Pleurobema_stabile **      
              |              |                                 
              |  +-----------+  +----- Pleurobema_riddellii **    
              |  |           |  |                              
              |  |           +--+  +-- Pleurobema_cordatum[1] **  
              |  |              |  |                           
              |  |              |  +-- Pleurobema_sintoxia[1] **  
              |  |              +--+                           
              |  |                 +-- Pleurobema_sintoxia[2] **  
              +--+                 |                           
                 |                 +-- Pleurobema_sintoxia[3] **  
                 |                                             
                 |              +----- Pleuronaia_barnesiana **   
                 |  +-----------+                              
                 |  |           |  +-- Pleuronaia_dolabelloides **
                 |  |           +--+                           
                 |  |              +-- Pleuronaia_gibber **       
                 +--+                                          
                    |              +-- Pleurobema_georgianum **   
                    |  +-----------+                           
                    |  |           +-- Pleurobema_pyriforme **    
                    |  |                                       
                    +--+           +-- Pleurobema_decisum **      
                       |  +--------+                           
                       |  |        +-- Pleurobema_hanleyianum **  
                       |  |                                    
                       +--+        +-- Pleurobema_clava **        
                          |  +-----+                           
                          |  |     +-- Pleurobema_oviforme **     
                          |  |                                 
                          +--+  +----- Pleurobema_rubellum **     
                             |  |                              
                             |  +----- Pleurobema_beadleianum **  
                             +--+                              
                                +----- Pleurobema_strodeanum **   
                                |                              
                                |  +-- Pleurobema_perovatum **    
                                +--+                           
                                   +-- Pleurobema_taitianum **    

Burdick & White, 2007 [Publ 2679], fig. 3

iT tT bi riS oT roT roS iD
38 38 31 3 0 0 0 46.0
  +----------------------------------------- Lampsilis_ornata **      
  |                                                                
--+  +-------------------------------------- Parvaspina_collina **    
  |  |                                                             
  |  |  +----------------------------------- Reginaia_ebenus **       
  +--+  |                                                          
     |  |                             +----- Eurynia_dilatata **      
     |  |  +--------------------------+                            
     |  |  |                          |  +-- Plethobasus_cyphyus **   
     +--+  |                          +--+                         
        |  |                             +-- Elliptio_crassidens **   
        |  |                                                       
        |  |  +----------------------------- Pleurobema_plenum **     
        |  |  |                                                    
        |  |  |                          +-- Pleuronaia_barnesiana ** 
        +--+  +--------------------------+                         
           |  |                          +-- Pleuronaia_gibber **     
           |  |                                                    
           |  |              +-------------- Pleurobema_stabile **    
           |  |              |                                     
           |  |              |           +-- Pleurobema_rubellum **   
           |  |              |     +-----+                         
           |  |              |     |     +-- Pleurobema_strodeanum ** 
           +--+  +-----------+  +--+                               
              |  |           |  |  |  +----- Pleurobema_beadleianum **
              |  |           |  |  +--+                            
              |  |           |  |     |  +-- Pleurobema_perovatum **  
              |  |           |  |     +--+                         
              |  |           +--+        +-- Pleurobema_taitianum **  
              |  |              |                                  
              |  |              |        +-- Pleurobema_clava **      
              |  |              |  +-----+                         
              |  |              |  |     +-- Pleurobema_oviforme **   
              |  |              |  |                               
              |  |              +--+     +-- Pleurobema_decisum **    
              +--+                 |  +--+                         
                 |                 |  |  +-- Pleurobema_hanleyianum **
                 |                 +--+                            
                 |                    |  +-- Pleurobema_pyriforme **  
                 |                    +--+                         
                 |                       +-- Pleurobema_georgianum ** 
                 |                                                 
                 |                    +----- Pleurobema_sintoxia[1] **
                 |                    |                            
                 |  +-----------------+  +-- Pleurobema_cordatum[2] **
                 |  |                 |  |                         
                 |  |                 +--+-- Pleurobema_sintoxia[2] **
                 |  |                    |                         
                 +--+                    +-- Pleurobema_sintoxia[3] **
                    |                                              
                    |  +-------------------- Pleurobema_cordatum[1] **
                    |  |                                           
                    +--+  +----------------- Fusconaia_subrotunda **  
                       |  |                                        
                       |  |              +-- Fusconaia_cor **         
                       +--+  +-----------+                         
                          |  |           +-- Fusconaia_cuneolus **    
                          |  |                                     
                          +--+  +----------- Fusconaia_ozarkensis **  
                             |  |                                  
                             |  |        +-- Fusconaia_masoni **      
                             +--+  +-----+                         
                                |  |     +-- Fusconaia_flava[3] **    
                                |  |                               
                                +--+  +----- Fusconaia_flava[1] **    
                                   |  |                            
                                   |  +----- Fusconaia_cerina[4] **   
                                   +--+                            
                                      |  +-- Fusconaia_cerina[1] **   
                                      |  |                         
                                      |  +-- Fusconaia_cerina[2] **   
                                      +--+                         
                                         +-- Fusconaia_cerina[3] **   
                                         |                         
                                         +-- Fusconaia_flava[2] **    

Pfeiffer et al., 2016 [Publ 3660], fig. 2

iT tT bi riS oT roT roS iD
38 38 33 4 1 1 1 47.0
  +-------------------------------------- Utterbackiana_suborbiculata
  |                                                                  
  |                                   +-- Amblema_plicata **            
  |                                +--+                              
  |                                |  +-- Amblema_neislerii **          
  |  +-----------------------------+                                 
--+  |                             |  +-- Reginaia_rotulata **          
  |  |                             +--+                              
  |  |                                +-- Reginaia_ebenus **            
  |  |                                                               
  |  |                                +-- Cyrtonaias_tampicoensis **    
  |  |                          +-----+                              
  |  |                          |     +-- Glebula_rotundata **          
  +--+                          |                                    
     |  +-----------------------+     +-- Lampsilis_teres **            
     |  |                       |  +--+                              
     |  |                       |  |  +-- Leaunio_lienosus **           
     |  |                       +--+                                 
     |  |                          |  +-- Truncilla_macrodon **         
     |  |                          +--+                              
     |  |                             +-- Potamilus_fragilis **         
     +--+                                                            
        |  +----------------------------- Elliptoideus_sloatianus **    
        |  |                                                         
        |  |              +-------------- Uniomerus_declivis **         
        |  |  +-----------+                                          
        |  |  |           |  +----------- Quadrula_quadrula **          
        |  |  |           +--+                                       
        +--+  |              |  +-------- Quadrula_verrucosa **         
           |  |              +--+                                    
           |  |                 |  +----- Pustulosa_pustulosa **        
           |  |                 +--+                                 
           |  |                    |  +-- Pustulosa_petrina **          
           |  |                    +--+                              
           +--+                       +-- Cyclonaias_tuberculata **     
              |                                                      
              |                       +-- Elliptio_fumata **            
              |  +--------------------+                              
              |  |                    +-- Elliptio_crassidens **        
              |  |                                                   
              |  |  +-------------------- Parvaspina_collina **         
              |  |  |                                                
              |  |  +-------------------- Plethobasus_cyphyus **        
              +--+  |                                                
                 |  |                 +-- Hemistena_lata **             
                 |  +-----------------+                              
                 |  |                 +-- Eurynia_dilatata **           
                 |  |                                                
                 |  |              +----- Pleurobema_riddellii **       
                 +--+           +--+                                 
                    |           |  |  +-- Pleurobema_sintoxia **        
                    |           |  +--+                              
                    |  +--------+     +-- Pleurobema_cordatum **        
                    |  |        |                                    
                    |  |        |  +----- Pleuronaia_dolabelloides **   
                    |  |        |  |                                 
                    |  |        +--+  +-- Pleurobema_pyriforme **       
                    |  |           |  |                              
                    +--+           +--+-- Pleurobema_strodeanum **      
                       |              |                              
                       |              +-- Pleurobema_clava **           
                       |                                             
                       |  +-------------- Fusconaia_askewi **           
                       |  |                                          
                       |  |           +-- Fusconaia_mitchelli **        
                       +--+  +--------+                              
                          |  |        +-- Fusconaia_iheringi **         
                          |  |                                       
                          +--+        +-- Fusconaia_escambia **         
                             |  +-----+                              
                             |  |     +-- Fusconaia_burkei **           
                             +--+                                    
                                |  +----- Fusconaia_flava[2] **         
                                +--+                                 
                                   |  +-- Fusconaia_cerina **           
                                   +--+                              
                                      +-- Fusconaia_flava[1] **         

Pfeiffer et al., 2019 [Publ 3866], fig. 1

iT tT bi riS oT roT roS iD
38 38 33 5 1 1 1 42.0
  +-------------------------------------- Gonidea_angulata           
  |                                                                  
  |                          +----------- Megalonaias_nervosa **        
  |  +-----------------------+                                       
  |  |                       |  +-------- Uniomerus_declivis **         
  |  |                       +--+                                    
--+  |                          |  +----- Quadrula_verrucosa **         
  |  |                          +--+                                 
  |  |                             |  +-- Cyclonaias_tuberculata **     
  |  |                             +--+                              
  |  |                                +-- Quadrula_quadrula **          
  |  |                                                               
  |  |                    +-------------- Elliptoideus_sloatianus **    
  +--+  +-----------------+                                          
     |  |                 |  +----------- Elliptio_fumata **            
     |  |                 +--+                                       
     |  |                    |  +-------- Pleuronaia_barnesiana **      
     |  |                    +--+                                    
     |  |                       |  +----- Pleurobema_riddellii **       
     |  |                       +--+                                 
     |  |                          |  +-- Fusconaia_flava **            
     |  |                          +--+                              
     |  |                             +-- Pleurobema_strodeanum **      
     |  |                                                            
     +--+                             +-- Amblema_plicata **            
        |                          +--+                              
        |                          |  +-- Amblema_neislerii **          
        |  +-----------------------+                                 
        |  |                       |  +-- Reginaia_ebenus **            
        |  |                       +--+                              
        |  |                          +-- Reginaia_rotulata **          
        |  |                                                         
        |  |           +----------------- Popenaias_popeii **           
        |  |           |                                             
        |  |  +--------+  +-------------- Psoronaias_guatemalensis[2] **
        |  |  |        |  |                                          
        +--+  |        +--+  +----------- Psoronaias_guatemalensis[1] **
           |  |           |  |                                       
           |  |           +--+  +-------- Psoronaias_semigranosa[3] **  
           |  |              |  |                                    
           |  |              |  |  +----- Psoronaias_semigranosa[2] **  
           |  |              +--+  |                                 
           |  |                 |  +----- Psoronaias_usumasintae[2] **  
           |  |                 |  |                                 
           |  |                 +--+----- Psoronaias_percompressa **    
           +--+                    |                                 
              |                    |  +-- Psoronaias_semigranosa[1] **  
              |                    |  |                              
              |                    +--+-- Psoronaias_profunda **        
              |                       |                              
              |                       +-- Psoronaias_usumasintae[1] **  
              |                                                      
              |                       +-- Cyrtonaias_tampicoensis **    
              |  +--------------------+                              
              |  |                    +-- Glebula_rotundata **          
              |  |                                                   
              +--+  +-------------------- Potamilus_alatus **           
                 |  |                                                
                 |  |                 +-- Truncilla_truncata **         
                 +--+  +--------------+                              
                    |  |              +-- Ptychobranchus_fasciolaris ** 
                    |  |                                             
                    +--+  +-------------- Pachynaias_spheniopsis **     
                       |  |                                          
                       |  |           +-- Cyrtonaias_explicata **       
                       +--+  +--------+                              
                          |  |        +-- Sphenonaias_microdon **       
                          |  |                                       
                          +--+        +-- Lampsilis_cardium **          
                             |  +-----+                              
                             |  |     +-- Ortmanniana_ligamentina **    
                             +--+                                    
                                |  +----- Cambarunio_iris **            
                                +--+                                 
                                   |  +-- Epioblasma_triquetra **       
                                   +--+                              
                                      +-- Ligumia_recta **              

Perkins et al., 2017 [Publ 3777], fig. 2

iT tT bi riS oT roT roS iD
37 37 34 2 0 0 0 48.0
  +----------------------------------------- Amblema_plicata **         
  |                                                                  
  +----------------------------------------- Plectomerus_dombeyanus **  
  |                                                                  
  |                                      +-- Pleurobema_clava **        
  |                          +-----------+                           
  |                          |           +-- Pleurobema_oviforme **     
  |                          |                                       
  |                          |     +-------- Pleurobema_pyriforme **    
  |  +-----------------------+  +--+                                 
--+  |                       |  |  |  +----- Pleurobema_georgianum **   
  |  |                       |  |  +--+                              
  |  |                       |  |     |  +-- Pleurobema_hanleyianum **  
  |  |                       +--+     +--+                           
  |  |                          |        +-- Pleurobema_decisum **      
  |  |                          |                                    
  |  |                          |  +-------- Pleurobema_rubellum **     
  |  |                          +--+                                 
  |  |                             |  +----- Pleurobema_perovatum **    
  |  |                             +--+                              
  +--+                                |  +-- Pleurobema_beadleianum **  
     |                                +--+                           
     |                                   +-- Pleurobema_strodeanum **   
     |                                                               
     |                                +----- Pleuronaia_gibber **       
     |  +-----------------------------+                              
     |  |                             |  +-- Pleuronaia_barnesiana **   
     |  |                             +--+                           
     |  |                                +-- Pleuronaia_dolabelloides **
     |  |                                                            
     |  |                                +-- Fusconaia_masoni **        
     +--+                             +--+                           
        |                             |  +-- Fusconaia_flava **         
        |  +--------------------------+                              
        |  |                          |  +-- Fusconaia_subrotunda **    
        |  |                          +--+                           
        |  |                             +-- Fusconaia_cor **           
        +--+                                                         
           |                             +-- Parvaspina_collina **      
           |  +--------------------------+                           
           |  |                          +-- Parvaspina_steinstansana **
           |  |                                                      
           +--+                          +-- Eurynia_dilatata **        
              |  +-----------------------+                           
              |  |                       +-- Hemistena_lata **          
              +--+                                                   
                 |  +----------------------- Elliptio_producta[2] **    
                 |  |                                                
                 +--+  +-------------------- Elliptio_producta[1] **    
                    |  |                                             
                    +--+  +----------------- Elliptio_spinosa **        
                       |  |                                          
                       |  |  +-------------- Elliptio_icterina **       
                       +--+  |                                       
                          |  |     +-------- Elliptio_roanokensis **    
                          |  |  +--+                                 
                          +--+  |  |  +----- Elliptio_complanata **     
                             |  |  +--+                              
                             |  |     |  +-- Elliptio_crassidens **     
                             |  |     +--+                           
                             +--+        +-- Elliptio_pullata **        
                                |                                    
                                |        +-- Elliptio_congaraea **      
                                |  +-----+                           
                                |  |     +-- Elliptio_hopetonensis **   
                                |  |                                 
                                +--+     +-- Elliptio_folliculata **    
                                   |  +--+                           
                                   |  |  +-- Elliptio_jayensis **       
                                   +--+                              
                                      |  +-- Elliptio_arca **           
                                      +--+                           
                                         +-- Elliptio_arctata **        

Pieri et al., 2018 [Publ 3867], fig. 3

iT tT bi riS oT roT roS iD
36 36 22 2 1 1 1 38.0
  +----------------------------------------- Arcidens_confragosus   
  |                                                                 
  |                                      +-- Quadrula_quadrula **      
  |  +-----------------------------------+                          
--+  |                                   +-- Pustulosa_petrina **      
  |  |                                                              
  |  |  +----------------------------------- Eurynia_dilatata **       
  |  |  |                                                           
  +--+  |                             +----- Pleurobema_clava **       
     |  +-----------------------------+                             
     |  |                             |  +-- Pleurobema_riddellii **   
     |  |                             +--+                          
     |  |                                +-- Pleurobema_sintoxia **    
     +--+                                                           
        |                             +----- Fusconaia_subrotunda **   
        |  +--------------------------+                             
        |  |                          |  +-- Fusconaia_cor **          
        |  |                          +--+                          
        |  |                             +-- Fusconaia_cuneolus **     
        |  |                                                        
        +--+  +----------------------------- Fusconaia_ozarkensis **   
           |  |                                                     
           |  |                 +----------- Fusconaia_cerina[3] **    
           |  |                 |                                   
           |  |  +--------------+  +-------- Fusconaia_flava[1] **     
           +--+  |              |  |                                
              |  |              +--+  +----- Fusconaia_flava[3] **     
              |  |                 |  |                             
              |  |                 +--+  +-- Fusconaia_cerina[1] **    
              |  |                    |  |                          
              |  |                    +--+-- Fusconaia_cerina[2] **    
              +--+                       |                          
                 |                       +-- Fusconaia_flava[2] **     
                 |                                                  
                 |  +----------------------- Fusconaia_masoni **       
                 |  |                                               
                 |  |                    +-- Fusconaia_burkei **       
                 |  +--------------------+                          
                 +--+                    +-- Fusconaia_escambia **     
                    |                                               
                    |  +-------------------- Fusconaia_mitchelli **    
                    |  |                                            
                    +--+  +----------------- Fusconaia_chunii **       
                       |  |                                         
                       |  |  +-------------- Fusconaia_askewi[9] **    
                       +--+  |                                      
                          |  |  +----------- Fusconaia_askewi[6] **    
                          |  |  |                                   
                          +--+  +----------- Fusconaia_askewi[7] **    
                             |  |                                   
                             |  +----------- Fusconaia_askewi[8] **    
                             |  |                                   
                             |  +----------- Fusconaia_lananensis[5] **
                             +--+                                   
                                +----------- Fusconaia_lananensis[6] **
                                |                                   
                                |  +-------- Fusconaia_askewi[5] **    
                                |  |                                
                                |  +-------- Fusconaia_lananensis[4] **
                                |  |                                
                                +--+  +----- Fusconaia_askewi[2] **    
                                   |  |                             
                                   |  +----- Fusconaia_askewi[3] **    
                                   |  |                             
                                   +--+----- Fusconaia_askewi[4] **    
                                      |                             
                                      |  +-- Fusconaia_askewi[1] **    
                                      |  |                          
                                      |  +-- Fusconaia_lananensis[1] **
                                      +--+                          
                                         +-- Fusconaia_lananensis[2] **
                                         |                          
                                         +-- Fusconaia_lananensis[3] **

Burlakova et al., 2019 [Publ 3899], fig. 1

iT tT bi riS oT roT roS iD
33 33 21 5 0 0 0 38.0
                          +-- Quadrula_quadrula **      
  +-----------------------+                          
  |                       +-- Uniomerus_declivis **     
  |                                                  
  |  +----------------------- Plectomerus_dombeyanus ** 
--+  |                                               
  |  |  +-------------------- Popenaias_popeii **       
  |  |  |                                            
  |  |  |                 +-- Elliptio_crassidens **    
  |  |  +-----------------+                          
  +--+  |                 +-- Pleurobema_rubellum **    
     |  |                                            
     |  |                 +-- Cyrtonaias_tampicoensis **
     |  |              +--+                          
     |  |              |  +-- Glebula_rotundata **      
     |  +--------------+                             
     |  |              |  +-- Reginaia_ebenus **        
     +--+              +--+                          
        |                 +-- Obliquaria_reflexa **     
        |                                            
        |  +----------------- Toxolasma_lividum **      
        |  |                                         
        |  |              +-- Potamilus_fragilis **     
        |  |  +-----------+                          
        |  |  |           +-- Potamilus_metnecktayi **  
        |  |  |                                      
        |  |  |        +----- Medionidus_acutissimus ** 
        |  |  +--------+                             
        +--+  |        |  +-- Lemiox_rimosus **         
           |  |        +--+                          
           |  |           +-- Dromus_dromas **          
           |  |                                      
           |  |        +----- Ellipsaria_lineolata **   
           |  |        |                             
           |  +--------+  +-- Truncilla_cognata **      
           |  |        |  |                          
           +--+        |  +-- Truncilla_donaciformis ** 
              |        +--+                          
              |           +-- Truncilla_macrodon **     
              |           |                          
              |           +-- Truncilla_truncata **     
              |                                      
              |  +----------- Cambarunio_simus **       
              |  |                                   
              |  +----------- Leaunio_lienosus **       
              |  |                                   
              |  +----------- Villosa_villosa **        
              |  |                                   
              |  |        +-- Sagittunio_nasutus **     
              |  +--------+                          
              +--+        +-- Lampsilis_ovata **        
                 |                                   
                 |     +----- Venustaconcha_pleasii **  
                 +-----+                             
                 |     |  +-- Epioblasma_rangiana **    
                 |     +--+                          
                 |        +-- Obovaria_unicolor **      
                 |                                   
                 |  +-------- Lampsilis_teres **        
                 +--+                                
                    |  +----- Lampsilis_siliquoidea **  
                    +--+                             
                       |  +-- Lampsilis_hydiana **      
                       +--+                          
                          +-- Lampsilis_straminea **    

Serb et al., 2003 [Publ 2659], fig. 2

iT tT bi riS oT roT roS iD
29 29 20 4 0 0 0 38.0
                       +-- Lampsilis_siliquoidea **  
  +--------------------+                          
  |                    +-- Lampsilis_ornata **       
  |                                               
  |  +-------------------- Obliquaria_reflexa **     
  |  |                                            
  |  +-------------------- Amblema_plicata **        
  |  |                                            
  |  +-------------------- Reginaia_rotulata **      
  |  |                                            
--+  +-------------------- Potamilus_alatus **       
  |  |                                            
  |  |                 +-- Elliptoideus_sloatianus **
  |  +-----------------+                          
  |  |                 +-- Plectomerus_dombeyanus ** 
  |  |                                            
  |  |              +----- Medionidus_conradicus **  
  |  +--------------+                             
  |  |              |  +-- Dromus_dromas **          
  +--+              +--+                          
     |                 +-- Cyprogenia_aberti **      
     |                                            
     |           +-------- Hemistena_lata **         
     +-----------+                                
     |           |  +----- Elliptio_complanata **    
     |           +--+                             
     |              |  +-- Fusconaia_burkei **       
     |              +--+                          
     |                 +-- Fusconaia_flava **        
     |                                            
     |  +----------------- Megalonaias_nervosa **    
     |  |                                         
     |  |        +-------- Theliderma_cylindrica **  
     |  |  +-----+                                
     +--+  |     |  +----- Theliderma_intermedia **  
        |  |     +--+                             
        |  |        |  +-- Theliderma_sparsa **      
        |  |        +--+                          
        |  |           +-- Theliderma_metanevra **   
        +--+                                      
           |        +----- Quadrula_verrucosa **     
           |  +-----+                             
           |  |     |  +-- Quadrula_quadrula **      
           |  |     +--+                          
           |  |        +-- Quadrula_nobilis **       
           +--+                                   
              |  +-------- Pustulosa_kieneriana **   
              |  |                                
              |  +-------- Pustulosa_infucata **     
              +--+                                
                 |     +-- Pustulosa_pustulosa **    
                 |  +--+                          
                 |  |  +-- Pustulosa_succissa **     
                 +--+                             
                    |  +-- Pustulosa_nodulata **     
                    +--+                          
                       +-- Pustulosa_petrina **      

Serb et al., 2003 [Publ 2659], fig. 3

iT tT bi riS oT roT roS iD
29 29 27 4 0 0 0 40.0
                                            +-- Lampsilis_siliquoidea **  
  +-----------------------------------------+                          
  |                                         +-- Lampsilis_ornata **       
  |                                                                    
--+  +----------------------------------------- Potamilus_alatus **       
  |  |                                                                 
  |  |                                   +----- Medionidus_conradicus **  
  +--+  +--------------------------------+                             
     |  |                                |  +-- Dromus_dromas **          
     |  |                                +--+                          
     |  |                                   +-- Cyprogenia_aberti **      
     +--+                                                              
        |  +----------------------------------- Obliquaria_reflexa **     
        |  |                                                           
        |  |  +-------------------------------- Amblema_plicata **        
        +--+  |                                                        
           |  |                          +----- Reginaia_rotulata **      
           |  |  +-----------------------+                             
           +--+  |                       |  +-- Elliptoideus_sloatianus **
              |  |                       +--+                          
              |  |                          +-- Plectomerus_dombeyanus ** 
              |  |                                                     
              +--+                    +-------- Hemistena_lata **         
                 |  +-----------------+                                
                 |  |                 |  +----- Elliptio_complanata **    
                 |  |                 +--+                             
                 |  |                    |  +-- Fusconaia_burkei **       
                 |  |                    +--+                          
                 +--+                       +-- Fusconaia_flava **        
                    |                                                  
                    |  +----------------------- Megalonaias_nervosa **    
                    |  |                                               
                    |  |                    +-- Quadrula_verrucosa **     
                    +--+  +-----------------+                          
                       |  |                 +-- Quadrula_nobilis **       
                       |  |                                            
                       +--+  +----------------- Quadrula_quadrula **      
                          |  |                                         
                          |  |        +-------- Theliderma_cylindrica **  
                          +--+  +-----+                                
                             |  |     |  +----- Theliderma_sparsa **      
                             |  |     +--+                             
                             |  |        |  +-- Theliderma_intermedia **  
                             +--+        +--+                          
                                |           +-- Theliderma_metanevra **   
                                |                                      
                                |  +----------- Pustulosa_kieneriana **   
                                |  |                                   
                                +--+  +-------- Pustulosa_infucata **     
                                   |  |                                
                                   +--+     +-- Pustulosa_succissa **     
                                      |  +--+                          
                                      |  |  +-- Pustulosa_pustulosa **    
                                      +--+                             
                                         |  +-- Pustulosa_nodulata **     
                                         +--+                          
                                            +-- Pustulosa_petrina **      

Campbell et al., 2008 [Publ 2675], fig. 5

iT tT bi riS oT roT roS iD
29 29 6 4 0 0 0 44.0
  +----------- Uniomerus_declivis **       
  |                                     
  +----------- Plectomerus_dombeyanus **   
  |                                     
  |        +-- Elliptio_arctata **         
  |  +-----+                            
  |  |     +-- Elliptio_crassidens **      
  |  |                                  
  |  |  +----- Pleurobema_decisum **       
--+  |  |                               
  |  |  +----- Pleurobema_rubellum **      
  |  |  |                               
  |  |  +----- Pleurobema_georgianum[2] ** 
  |  |  |                               
  |  |  +----- Pleurobema_stabile **       
  |  |  |                               
  |  |  +----- Pleurobema_pyriforme **     
  |  |  |                               
  +--+  +----- Pleurobema_oviforme[2] **   
     |  |                               
     |  +----- Pleurobema_hanleyianum[2] **
     |  |                               
     |  +----- Pleuronaia_gibber **        
     |  |                               
     |  +----- Pleurobema_georgianum[3] ** 
     |  |                               
     |  +----- Pleurobema_cordatum **      
     |  |                               
     |  +----- Pleurobema_beadleianum **   
     |  |                               
     |  +----- Hemistena_lata **           
     +--+                               
        +----- Fusconaia_flava **          
        |                               
        +----- Fusconaia_cerina **         
        |                               
        +----- Eurynia_dilatata **         
        |                               
        +----- Elliptio_arca **            
        |                               
        +----- Cyrtonaias_tampicoensis **  
        |                               
        |  +-- Plethobasus_cyphyus **      
        +--+                            
        |  +-- Parvaspina_collina **       
        |                               
        |  +-- Pleurobema_sintoxia **      
        |  |                            
        +--+-- Pleurobema_oviforme[1] **   
        |  |                            
        |  +-- Pleurobema_clava **         
        |                               
        |  +-- Pleurobema_athearni **      
        |  |                            
        +--+-- Pleurobema_georgianum[1] ** 
           |                            
           +-- Pleurobema_hanleyianum[1] **

Boyer et al., 2011 [Publ 2718], fig. 3

iT tT bi riS oT roT roS iD
29 29 22 4 6 6 1 35.0
                          +----- Alasmidonta_marginata  
                          |                             
                          +----- Pyganodon_grandis      
                          |                             
  +-----------------------+  +-- Lasmigona_complanata   
  |                       +--+                          
  |                       |  +-- Lasmigona_costata      
  |                       |                             
  |                       |  +-- Platynaias_compressa   
  |                       +--+                          
  |                          +-- Strophitus_undulatus   
  |                                                     
--+  +-------------------------- Reginaia_ebenus **        
  |  |                                                  
  |  |                       +-- Fusconaia_flava **        
  |  |                    +--+                          
  |  |                    |  +-- Pleurobema_sintoxia **    
  |  |     +--------------+                             
  |  |     |              |  +-- Eurynia_dilatata **       
  |  |     |              +--+                          
  +--+  +--+                 +-- Elliptio_crassidens **    
     |  |  |                                            
     |  |  |  +----------------- Megalonaias_nervosa **    
     |  |  +--+                                         
     |  |     |  +-------------- Quadrula_quadrula **      
     |  |     +--+                                      
     |  |        |  +----------- Quadrula_verrucosa **     
     |  |        +--+                                   
     |  |           |  +-------- Theliderma_metanevra **   
     +--+           +--+                                
        |              |  +----- Pustulosa_pustulosa **    
        |              +--+                             
        |                 |  +-- Cyclonaias_tuberculata ** 
        |                 +--+                          
        |                    +-- Pustulosa_nodulata **     
        |                                               
        |     +----------------- Amblema_plicata **        
        |     |                                         
        |     |  +-------------- Toxolasma_parvum **       
        +-----+  |                                      
              |  |  +----------- Obliquaria_reflexa **     
              |  |  |                                   
              +--+  |  +-------- Obovaria_olivaria **      
                 |  |  |                                
                 |  |  +-------- Lampsilis_cardium **      
                 |  |  |                                
                 +--+  |     +-- Epioblasma_triquetra **   
                    |  +-----+                          
                    |  |     +-- Lampsilis_siliquoidea **  
                    |  |                                
                    |  |     +-- Ortmanniana_ligamentina **
                    |  +-----+                          
                    |  |     +-- Lampsilis_higginsii **    
                    +--+                                
                       |     +-- Ligumia_recta **          
                       +-----+                          
                       |     +-- Lampsilis_teres **        
                       |                                
                       |  +----- Potamilus_fragilis **     
                       +--+                             
                       |  |  +-- Potamilus_ohiensis **     
                       |  +--+                          
                       |     +-- Potamilus_alatus **       
                       |                                
                       |  +----- Ellipsaria_lineolata **   
                       +--+                             
                          |  +-- Truncilla_truncata **     
                          +--+                          
                             +-- Truncilla_donaciformis ** 

Boyer et al., 2011 [Publ 2718], fig. 4

iT tT bi riS oT roT roS iD
29 29 25 4 9 9 1 36.0
                       +-------------- Strophitus_undulatus      
                       |                                         
                       |        +----- Lasmigona_complanata      
  +--------------------+  +-----+                                
  |                    |  |     |  +-- Lasmigona_costata         
  |                    |  |     +--+                             
  |                    +--+        +-- Arcidens_confragosus      
  |                       |                                      
  |                       |  +-------- Utterbackia_imbecillis    
  |                       |  |                                   
  |                       +--+-------- Alasmidonta_marginata     
  |                          |                                   
  |                          |  +----- Pyganodon_grandis         
  |                          +--+                                
--+                             |  +-- Anodontoides_ferussacianus
  |                             +--+                             
  |                                +-- Platynaias_compressa      
  |                                                              
  |                       +----------- Theliderma_metanevra **      
  |                       |                                      
  |  +--------------------+        +-- Quadrula_verrucosa **        
  |  |                    |  +-----+                             
  |  |                    |  |     +-- Quadrula_quadrula **         
  |  |                    +--+                                   
  |  |                       |  +----- Pustulosa_pustulosa **       
  |  |                       +--+                                
  +--+                          |  +-- Cyclonaias_tuberculata **    
     |                          +--+                             
     |                             +-- Pustulosa_nodulata **        
     |                                                           
     |  +----------------------------- Megalonaias_nervosa **       
     |  |                                                        
     +--+  +-------------------------- Toxolasma_parvum **          
        |  |                                                     
        |  |  +----------------------- Amblema_plicata **           
        +--+  |                                                  
           |  |                    +-- Reginaia_ebenus **           
           |  |  +-----------------+                             
           |  |  |                 +-- Obliquaria_reflexa **        
           +--+  |                                               
              |  |           +-------- Pleurobema_sintoxia **       
              |  +-----------+                                   
              |  |           |  +----- Elliptio_crassidens **       
              |  |           +--+                                
              |  |              |  +-- Fusconaia_flava **           
              +--+              +--+                             
                 |                 +-- Eurynia_dilatata **          
                 |                                               
                 |              +----- Potamilus_fragilis **        
                 |  +-----------+                                
                 |  |           |  +-- Potamilus_ohiensis **        
                 |  |           +--+                             
                 |  |              +-- Potamilus_alatus **          
                 |  |                                            
                 +--+           +----- Ellipsaria_lineolata **      
                    |  +--------+                                
                    |  |        |  +-- Truncilla_donaciformis **    
                    |  |        +--+                             
                    |  |           +-- Truncilla_truncata **        
                    +--+                                         
                       |  +----------- Lampsilis_cardium **         
                       |  |                                      
                       |  |     +----- Ligumia_recta **             
                       |  +-----+                                
                       +--+     |  +-- Obovaria_olivaria **         
                          |     +--+                             
                          |        +-- Epioblasma_triquetra **      
                          |                                      
                          |  +-------- Lampsilis_teres **           
                          +--+                                   
                             |  +----- Lampsilis_siliquoidea **     
                             +--+                                
                                |  +-- Ortmanniana_ligamentina **   
                                +--+                             
                                   +-- Lampsilis_higginsii **       

Inoue et al., 2020 [Publ 4128], fig. 2

iT tT bi riS oT roT roS iD
29 29 17 4 0 0 0 36.0
  +-------------------------- Quadrula_quadrula **          
  |                                                      
  |                       +-- Reginaia_ebenus **            
  |  +--------------------+                              
  |  |                    +-- Reginaia_rotulata **          
--+  |                                                   
  |  |                 +----- Amblema_plicata **            
  |  |  +--------------+                                 
  |  |  |              |  +-- Amblema_elliottii **          
  |  |  |              +--+                              
  +--+  |                 +-- Amblema_neislerii **          
     |  |                                                
     |  |              +----- Popenaias_popeii **           
     |  |           +--+                                 
     |  |           |  |  +-- Popenaias_berezai **          
     |  |           |  +--+                              
     |  |           |     +-- Popenaias_semirasa **         
     +--+           |                                    
        |  +--------+     +-- Psoronaias_profunda **        
        |  |        |     |                              
        |  |        |     +-- Psoronaias_semigranosa[1] **  
        |  |        |     |                              
        |  |        |     +-- Psoronaias_semigranosa[2] **  
        |  |        |     |                              
        |  |        +-----+-- Psoronaias_guatemalensis[1] **
        |  |              |                              
        |  |              +-- Psoronaias_guatemalensis[2] **
        +--+              |                              
           |              +-- Psoronaias_percompressa **    
           |              |                              
           |              +-- Psoronaias_semigranosa[3] **  
           |                                             
           |              +-- Glebula_rotundata **          
           |  +-----------+                              
           |  |           +-- Cyrtonaias_tampicoensis **    
           |  |                                          
           +--+  +----------- Potamilus_alatus **           
              |  |                                       
              |  |        +-- Ptychobranchus_fasciolaris ** 
              +--+  +-----+                              
                 |  |     +-- Truncilla_truncata **         
                 |  |                                    
                 |  |  +----- Ortmanniana_ligamentina **    
                 +--+  |                                 
                    |  +----- Epioblasma_triquetra **       
                    |  |                                 
                    |  +----- Lampsilis_cardium **          
                    |  |                                 
                    +--+----- Ligumia_recta **              
                       |                                 
                       +----- Cambarunio_iris **            
                       |                                 
                       +----- Pachynaias_spheniopsis **     
                       |                                 
                       |  +-- Cyrtonaias_explicata **       
                       +--+                              
                          +-- Sphenonaias_microdon **       

Inoue et al., 2020 [Publ 4128], fig. 3

iT tT bi riS oT roT roS iD
29 29 27 4 0 0 0 35.0
  +----------------------------------------- Quadrula_quadrula **          
  |                                                                     
  |                                      +-- Reginaia_ebenus **            
  |  +-----------------------------------+                              
--+  |                                   +-- Reginaia_rotulata **          
  |  |                                                                  
  |  |                                +----- Amblema_neislerii **          
  |  |  +-----------------------------+                                 
  |  |  |                             |  +-- Amblema_plicata **            
  +--+  |                             +--+                              
     |  |                                +-- Amblema_elliottii **          
     |  |                                                               
     |  |                             +----- Popenaias_popeii **           
     |  |                    +--------+                                 
     |  |                    |        |  +-- Popenaias_berezai **          
     |  |                    |        +--+                              
     +--+                    |           +-- Popenaias_semirasa **         
        |  +-----------------+                                          
        |  |                 |  +----------- Psoronaias_profunda **        
        |  |                 |  |                                       
        |  |                 |  |     +----- Psoronaias_guatemalensis[1] **
        |  |                 +--+  +--+                                 
        |  |                    |  |  |  +-- Psoronaias_semigranosa[1] **  
        |  |                    |  |  +--+                              
        |  |                    +--+     +-- Psoronaias_percompressa **    
        +--+                       |                                    
           |                       |  +----- Psoronaias_semigranosa[3] **  
           |                       +--+                                 
           |                          |  +-- Psoronaias_guatemalensis[2] **
           |                          +--+                              
           |                             +-- Psoronaias_semigranosa[2] **  
           |                                                            
           |  +----------------------------- Cyrtonaias_tampicoensis **    
           |  |                                                         
           +--+  +-------------------------- Ptychobranchus_fasciolaris ** 
              |  |                                                      
              +--+  +----------------------- Ligumia_recta **              
                 |  |                                                   
                 +--+  +-------------------- Potamilus_alatus **           
                    |  |                                                
                    +--+  +----------------- Pachynaias_spheniopsis **     
                       |  |                                             
                       |  |              +-- Epioblasma_triquetra **       
                       +--+  +-----------+                              
                          |  |           +-- Truncilla_truncata **         
                          |  |                                          
                          +--+           +-- Ortmanniana_ligamentina **    
                             |  +--------+                              
                             |  |        +-- Lampsilis_cardium **          
                             +--+                                       
                                |  +-------- Cambarunio_iris **            
                                +--+                                    
                                   |  +----- Sphenonaias_microdon[2] **    
                                   +--+                                 
                                      |  +-- Cyrtonaias_explicata **       
                                      +--+                              
                                         +-- Sphenonaias_microdon[1] **    

Campbell et al., 2008 [Publ 2675], fig. 2

iT tT bi riS oT roT roS iD
28 28 18 3 0 0 0 41.0
  +----------------------- Cyrtonaias_tampicoensis ** 
  |                                                
  +----------------------- Plectomerus_dombeyanus **  
  |                                                
  |                    +-- Pleurobema_cordatum **     
  |                    |                           
  |        +-----------+-- Pleurobema_sintoxia[1] **  
  |        |           |                           
  |        |           +-- Pleurobema_sintoxia[2] **  
  |        |                                       
--+  +-----+           +-- Fusconaia_cerina **        
  |  |     |  +--------+                           
  |  |     |  |        +-- Fusconaia_flava **         
  |  |     |  |                                    
  |  |     +--+  +-------- Pleuronaia_gibber **       
  |  |        |  |                                 
  |  |        |  |  +----- Eurynia_dilatata **        
  |  |        +--+  |                              
  |  |           |  +----- Hemistena_lata **          
  |  |           |  |                              
  |  |           +--+----- Plethobasus_cyphyus **     
  +--+              |                              
     |              +----- Parvaspina_collina **      
     |              |                              
     |              |  +-- Elliptio_arca **           
     |              +--+                           
     |                 +-- Elliptio_crassidens **     
     |                                             
     |  +----------------- Pleurobema_stabile **      
     |  |                                          
     |  |  +-------------- Pleurobema_pyriforme **    
     |  |  |                                       
     +--+  |           +-- Pleurobema_georgianum[2] **
        |  +-----------+                           
        |  |           +-- Pleurobema_athearni **     
        |  |                                       
        |  |        +----- Pleurobema_decisum **      
        +--+--------+                              
           |        |  +-- Pleurobema_hanleyianum **  
           |        +--+                           
           |           +-- Pleurobema_georgianum[1] **
           |                                       
           |           +-- Pleurobema_clava **        
           |  +--------+                           
           |  |        +-- Pleurobema_oviforme **     
           +--+                                    
              |  +-------- Pleurobema_rubellum **     
              |  |                                 
              +--+-------- Pleurobema_strodeanum **   
                 |                                 
                 |  +----- Pleurobema_beadleianum **  
                 +--+                              
                    |  +-- Pleurobema_taitianum **    
                    +--+                           
                       +-- Pleurobema_perovatum **    

Campbell et al., 2008 [Publ 2675], fig. 4

iT tT bi riS oT roT roS iD
28 28 9 4 0 0 0 41.0
  +----------------------- Uniomerus_declivis **       
  |                                                 
  +----------------------- Plectomerus_dombeyanus **   
  |                                                 
  |  +-------------------- Cyrtonaias_tampicoensis **  
  |  |                                              
  |  +-------------------- Eurynia_dilatata **         
  |  |                                              
  |  +-------------------- Fusconaia_cerina **         
  |  |                                              
--+  +-------------------- Fusconaia_flava **          
  |  |                                              
  |  +-------------------- Hemistena_lata **           
  |  |                                              
  |  +-------------------- Plethobasus_cyphyus **      
  |  |                                              
  |  +-------------------- Parvaspina_collina **       
  |  |                                              
  |  +-------------------- Pleurobema_cordatum **      
  |  |                                              
  +--+-------------------- Pleuronaia_gibber **        
     |                                              
     +-------------------- Pleurobema_oviforme[2] **   
     |                                              
     +-------------------- Pleurobema_stabile **       
     |                                              
     |              +----- Elliptio_arca **            
     +--------------+                               
     |              |  +-- Elliptio_crassidens **      
     |              +--+                            
     |                 +-- Elliptio_arctata **         
     |                                              
     |  +----------------- Pleurobema_decisum **       
     |  |                                           
     |  +----------------- Pleurobema_sintoxia **      
     |  |                                           
     |  +----------------- Pleurobema_clava **         
     +--+                                           
        +----------------- Pleurobema_oviforme[1] **   
        |                                           
        |  +-------------- Pleurobema_beadleianum **   
        |  |                                        
        +--+  +----------- Pleurobema_pyriforme **     
           |  |                                     
           +--+  +-------- Pleurobema_rubellum **      
              |  |                                  
              |  +-------- Pleurobema_hanleyianum[2] **
              +--+                                  
                 +-------- Pleurobema_georgianum[2] ** 
                 |                                  
                 |  +----- Pleurobema_hanleyianum[1] **
                 +--+                               
                    |  +-- Pleurobema_georgianum[1] ** 
                    +--+                            
                       +-- Pleurobema_athearni **      

Campbell et al., 2008 [Publ 2675], fig. 6

iT tT bi riS oT roT roS iD
28 28 23 3 0 0 0 40.0
  +----------------------------------------- Cyrtonaias_tampicoensis ** 
  |                                                                  
  +----------------------------------------- Plectomerus_dombeyanus **  
--+                                                                  
  |  +-------------------------------------- Parvaspina_collina **      
  |  |                                                               
  +--+  +----------------------------------- Hemistena_lata **          
     |  |                                                            
     |  |                             +----- Plethobasus_cyphyus **     
     +--+  +--------------------------+                              
        |  |                          |  +-- Elliptio_arca **           
        |  |                          +--+                           
        |  |                             +-- Elliptio_crassidens **     
        +--+                                                         
           |                             +-- Fusconaia_cerina **        
           |  +--------------------------+                           
           |  |                          +-- Fusconaia_flava **         
           |  |                                                      
           +--+                          +-- Eurynia_dilatata **        
              |  +-----------------------+                           
              |  |                       +-- Pleuronaia_gibber **       
              |  |                                                   
              +--+                    +----- Pleurobema_sintoxia[2] **  
                 |  +-----------------+                              
                 |  |                 |  +-- Pleurobema_cordatum **     
                 |  |                 +--+                           
                 +--+                    +-- Pleurobema_sintoxia[1] **  
                    |                                                
                    |  +-------------------- Pleurobema_stabile **      
                    |  |                                             
                    +--+  +----------------- Pleurobema_pyriforme **    
                       |  |                                          
                       |  |           +----- Pleurobema_decisum **      
                       +--+  +--------+                              
                          |  |        |  +-- Pleurobema_hanleyianum **  
                          |  |        +--+                           
                          |  |           +-- Pleurobema_georgianum[2] **
                          +--+                                       
                             |           +-- Pleurobema_georgianum[1] **
                             |  +--------+                           
                             |  |        +-- Pleurobema_athearni **     
                             |  |                                    
                             +--+        +-- Pleurobema_oviforme **     
                                |  +-----+                           
                                |  |     +-- Pleurobema_clava **        
                                |  |                                 
                                +--+  +----- Pleurobema_beadleianum **  
                                   |  |                              
                                   |  +----- Pleurobema_rubellum **     
                                   +--+                              
                                      +----- Pleurobema_strodeanum **   
                                      |                              
                                      |  +-- Pleurobema_perovatum **    
                                      +--+                           
                                         +-- Pleurobema_taitianum **    

Smith et al., 2020 [Publ 4129], fig. 1

iT tT bi riS oT roT roS iD
28 28 26 5 0 0 0 32.0
  +-------------------------------------- Quadrula_quadrula **      
  |                                                              
  |                                   +-- Pleurobema_strodeanum **  
--+  +--------------------------------+                          
  |  |                                +-- Elliptoideus_sloatianus **
  |  |                                                           
  |  |                             +----- Reginaia_ebenus **        
  +--+  +--------------------------+                             
     |  |                          |  +-- Amblema_plicata **        
     |  |                          +--+                          
     |  |                             +-- Plectomerus_dombeyanus ** 
     +--+                                                        
        |                             +-- Popenaias_popeii **       
        |  +--------------------------+                          
        |  |                          +-- Psoronaias_semigranosa ** 
        |  |                                                     
        +--+  +-------------------------- Obliquaria_reflexa **     
           |  |                                                  
           |  |  +----------------------- Cyrtonaias_tampicoensis **
           +--+  |                                               
              |  |                    +-- Pachynaias_spheniopsis ** 
              |  |  +-----------------+                          
              +--+  |                 +-- Lampsilis_cardium **      
                 |  |                                            
                 |  |        +----------- Ellipsaria_lineolata **   
                 |  |  +-----+                                   
                 +--+  |     |  +-------- Truncilla_truncata **     
                    |  |     +--+                                
                    |  |        |  +----- Truncilla_donaciformis ** 
                    |  |        +--+                             
                    |  |           |  +-- Truncilla_cognata **      
                    |  |           +--+                          
                    +--+              +-- Truncilla_macrodon **     
                       |                                         
                       |  +-------------- Atlanticoncha_ochracea ** 
                       |  |                                      
                       |  |           +-- Potamilus_fragilis **     
                       |  |     +-----+                          
                       |  |     |     +-- Potamilus_leptodon **     
                       +--+  +--+                                
                          |  |  |  +----- Potamilus_metnecktayi **  
                          |  |  +--+                             
                          |  |     |  +-- Potamilus_alatus **       
                          |  |     +--+                          
                          +--+        +-- Potamilus_purpuratus **   
                             |                                   
                             |        +-- Potamilus_capax **        
                             |  +-----+                          
                             |  |     +-- Potamilus_inflatus **     
                             +--+                                
                                |  +----- Potamilus_ohiensis **     
                                +--+                             
                                   |  +-- Potamilus_streckersoni ** 
                                   +--+                          
                                      +-- Potamilus_amphichaenus ** 

Smith et al., 2020 [Publ 4129], fig. 2-3

iT tT bi riS oT roT roS iD
28 28 26 5 0 0 0 32.0
  +----------------------------------------- Quadrula_quadrula **      
  |                                                                 
  |                                      +-- Pleurobema_strodeanum **  
--+  +-----------------------------------+                          
  |  |                                   +-- Elliptoideus_sloatianus **
  |  |                                                              
  |  |                                +----- Reginaia_ebenus **        
  +--+  +-----------------------------+                             
     |  |                             |  +-- Amblema_plicata **        
     |  |                             +--+                          
     |  |                                +-- Plectomerus_dombeyanus ** 
     +--+                                                           
        |                                +-- Popenaias_popeii **       
        |  +-----------------------------+                          
        |  |                             +-- Psoronaias_semigranosa ** 
        |  |                                                        
        +--+  +----------------------------- Obliquaria_reflexa **     
           |  |                                                     
           |  |  +-------------------------- Cyrtonaias_tampicoensis **
           +--+  |                                                  
              |  |                       +-- Pachynaias_spheniopsis ** 
              |  |  +--------------------+                          
              +--+  |                    +-- Lampsilis_cardium **      
                 |  |                                               
                 |  |           +----------- Ellipsaria_lineolata **   
                 |  |  +--------+                                   
                 +--+  |        |  +-------- Truncilla_truncata **     
                    |  |        +--+                                
                    |  |           |  +----- Truncilla_donaciformis ** 
                    |  |           +--+                             
                    |  |              |  +-- Truncilla_cognata **      
                    +--+              +--+                          
                       |                 +-- Truncilla_macrodon **     
                       |                                            
                       |  +----------------- Atlanticoncha_ochracea ** 
                       |  |                                         
                       |  |              +-- Potamilus_capax **        
                       +--+  +-----------+                          
                          |  |           +-- Potamilus_inflatus **     
                          |  |                                      
                          |  |        +----- Potamilus_ohiensis **     
                          +--+  +-----+                             
                             |  |     |  +-- Potamilus_streckersoni ** 
                             |  |     +--+                          
                             |  |        +-- Potamilus_amphichaenus ** 
                             +--+                                   
                                |        +-- Potamilus_fragilis **     
                                |  +-----+                          
                                |  |     +-- Potamilus_leptodon **     
                                +--+                                
                                   |  +----- Potamilus_metnecktayi **  
                                   +--+                             
                                      |  +-- Potamilus_alatus **       
                                      +--+                          
                                         +-- Potamilus_purpuratus **   

Perkins et al., 2017 [Publ 3777], fig. 3

iT tT bi riS oT roT roS iD
27 27 10 3 0 0 0 40.0
  +----------------------- Plectomerus_dombeyanus **  
  |                                                
  +----------------------- Uniomerus_declivis **      
  |                                                
  |                    +-- Parvaspina_steinstansana **
  |                    |                           
  |                    +-- Parvaspina_collina[1] **   
--+                    |                           
  |  +-----------------+-- Parvaspina_collina[2] **   
  |  |                 |                           
  |  |                 +-- Parvaspina_collina[3] **   
  |  |                 |                           
  |  |                 +-- Parvaspina_collina[4] **   
  |  |                                             
  +--+  +----------------- Elliptio_arca **           
     |  |                                          
     |  +----------------- Hemistena_lata **          
     |  |                                          
     |  |              +-- Elliptio_arctata **        
     |  +--------------+                           
     |  |              +-- Elliptio_crassidens **     
     +--+                                          
        |  +-------------- Eurynia_dilatata **        
        |  |                                       
        |  |  +----------- Pleurobema_oviforme **     
        |  |  |                                    
        |  |  +----------- Pleurobema_clava **        
        |  |  |                                    
        +--+  +----------- Pleurobema_pyriforme **    
           |  |                                    
           |  +----------- Pleurobema_georgianum **   
           |  |                                    
           |  +----------- Pleurobema_beadleianum **  
           |  |                                    
           +--+----------- Pleurobema_hanleyianum **  
              |                                    
              +----------- Pleurobema_decisum **      
              |                                    
              |        +-- Pleurobema_sintoxia **     
              +--------+                           
              |        +-- Pleurobema_rubellum **     
              |                                    
              |  +-------- Elliptio_spinosa **        
              |  |                                 
              +--+  +----- Fusconaia_escambia **      
                 |  |                              
                 |  +----- Fusconaia_flava **         
                 +--+                              
                    +----- Fusconaia_cerina **        
                    |                              
                    |  +-- Pleuronaia_gibber **       
                    +--+                           
                       +-- Pleuronaia_dolabelloides **

Neemuchwala et al., 2023 [Publ 4292], fig. 2

iT tT bi riS oT roT roS iD
26 26 24 5 1 1 1 32.0
  +-------------------------------- Gonidea_angulata      
  |                                                       
  |                       +-------- Pleurobema_strodeanum ** 
  |  +--------------------+                               
--+  |                    |  +----- Amblema_plicata **       
  |  |                    +--+                            
  |  |                       |  +-- Lampsilis_cardium **     
  |  |                       +--+                         
  +--+                          +-- Popenaias_popeii **      
     |                                                    
     |  +-------------------------- Megalonaias_nervosa **   
     |  |                                                 
     |  |                 +-------- Uniomerus_carolinianus **
     |  |  +--------------+                               
     +--+  |              |  +----- Uniomerus_columbensis ** 
        |  |              +--+                            
        |  |                 |  +-- Uniomerus_tetralasmus ** 
        |  |                 +--+                         
        |  |                    +-- Uniomerus_declivis **    
        +--+                                              
           |              +-------- Theliderma_cylindrica ** 
           |  +-----------+                               
           |  |           |  +----- Theliderma_metanevra **  
           |  |           +--+                            
           |  |              |  +-- Theliderma_sparsa **     
           |  |              +--+                         
           +--+                 +-- Theliderma_johnsoni **   
              |                                           
              |  +----------------- Cyclonaias_tuberculata **
              |  |                                        
              |  |        +-------- Quadrula_nobilis **      
              |  |  +-----+                               
              +--+  |     |  +----- Quadrula_verrucosa **    
                 |  |     +--+                            
                 |  |        |  +-- Quadrula_fragosa **      
                 |  |        +--+                         
                 |  |           +-- Quadrula_quadrula **     
                 +--+                                     
                    |        +----- Pustulosa_nodulata **    
                    |  +-----+                            
                    |  |     |  +-- Pustulosa_necki **       
                    |  |     +--+                         
                    |  |        +-- Pustulosa_petrina **     
                    +--+                                  
                       |        +-- Pustulosa_kieneriana **  
                       |  +-----+                         
                       |  |     +-- Pustulosa_pustulosa **   
                       +--+                               
                          |  +----- Pustulosa_succissa **    
                          +--+                            
                             |  +-- Pustulosa_infucata **    
                             +--+                         
                                +-- Pustulosa_kleiniana **   

Huang et al., 2019 [Publ 3843], fig. S3

iT tT bi riS oT roT roS iD
25 25 23 4 176 137 3 22.0
                                         +----- Chamberlainia_somsakpanhai 
  +--------------------------------------+                                 
  |                                      |  +-- Sinohyriopsis_cumingii     
  |                                      +--+                              
  |                                         +-- Sinohyriopsis_schlegelii   
  |                                                                        
  |                                         +-- Potomida_littoralis        
  |                                   +-----+                              
  |                                   |     +-- Pronodularia_japanensis    
  |                             +-----+                                    
  |                             |     |  +----- Lamprotula_leaii           
  |                             |     +--+                                 
  |                             |        |  +-- Lamprotula_caveata         
  |                             |        +--+                              
  |                             |           +-- Lamprotula_cornuumlunae    
--+  +--------------------------+                                          
  |  |                          |        +----- Sinosolenaia_recognita     
  |  |                          |  +-----+                                 
  |  |                          |  |     |  +-- Sinosolenaia_oleivora      
  |  |                          |  |     +--+                              
  |  |                          |  |        +-- Sinosolenaia_iridinea      
  |  |                          +--+                                       
  |  |                             |     +----- Gonidea_angulata           
  |  |                             |  +--+                                 
  |  |                             |  |  |  +-- Leguminaia_wheatleyi       
  |  |                             |  |  +--+                              
  |  |                             +--+     +-- Microcondylaea_bonellii    
  |  |                                |                                    
  |  |                                |  +----- Ptychorhynchus_pfisteri    
  +--+                                +--+                                 
     |                                   |  +-- Parvasolenaia_rivularis    
     |                                   +--+                              
     |                                      +-- Koreosolenaia_sitgyensis   
     |                                                                     
     |                                      +-- Megalonaias_nervosa **        
     |                                   +--+                              
     |                                   |  +-- Uniomerus_carolinianus **     
     |  +--------------------------------+                                 
     |  |                                |  +-- Quadrula_quadrula **          
     |  |                                +--+                              
     |  |                                   +-- Quadrula_verrucosa **         
     |  |                                                                  
     |  |                             +-------- Elliptoideus_sloatianus **    
     +--+                             |                                    
        |  +--------------------------+     +-- Elliptio_complanata **        
        |  |                          |  +--+                              
        |  |                          |  |  +-- Eurynia_dilatata **           
        |  |                          +--+                                 
        |  |                             |  +-- Fusconaia_flava **            
        +--+                             +--+                              
           |                                +-- Pleurobema_sintoxia **        
           |                                                               
           |  +-------------------------------- Amblema_plicata **            
           |  |                                                            
           +--+  +----------------------------- Toxolasma_parvum **           
              |  |                                                         
              +--+                          +-- Obliquaria_reflexa **         
                 |  +-----------------------+                              
                 |  |                       +-- Plectomerus_dombeyanus **     
                 +--+                                                      
                    |  +----------------------- Potamilus_leptodon **         
                    |  |                                                   
                    +--+  +-------------------- Potamilus_alatus **           
                       |  |                                                
                       |  |                 +-- Medionidus_conradicus **      
                       +--+  +--------------+                              
                          |  |              +-- Ptychobranchus_subtentus **   
                          |  |                                             
                          +--+              +-- Ellipsaria_lineolata **       
                             |  +-----------+                              
                             |  |           +-- Truncilla_truncata **         
                             +--+                                          
                                |  +----------- Leaunio_lienosus **           
                                |  |                                       
                                +--+        +-- Venustaconcha_ellipsiformis **
                                   |  +-----+                              
                                   |  |     +-- Cambarunio_iris **            
                                   +--+                                    
                                      |  +----- Ortmanniana_ligamentina **    
                                      +--+                                 
                                         |  +-- Lampsilis_cardium **          
                                         +--+                              
                                            +-- Lampsilis_ornata **           

Huang et al., 2019 [Publ 3843], fig. S4

iT tT bi riS oT roT roS iD
25 127 70 4 177 138 3 128.0
                          +-- Parreysia_keralaensis      
                          |                              
     +--------------------+-- Parreysia_nagpoorensis     
     |                    |                              
     |                    +-- Parreysia_plagiosoma       
     |                                                   
     |                 +----- Trapezidens_exolescens     
     |        +--------+                                 
     |        |        |  +-- Trapezidens_angustior      
     |        |        +--+                              
     +--------+           +-- Trapezidens_dolichorhynchus
     |        |                                          
     |        |  +----------- Lamellidens_corrianus      
     |        |  |                                       
     |        +--+  +-------- Lamellidens_savadiensis    
     |           |  |                                    
     |           |  |  +----- Lamellidens_marginalis     
     |           +--+  |                                 
     |              |  +----- Lamellidens_ferrugineus    
     |              |  |                                 
  +--+              +--+  +-- Lamellidens_mainwaringi[1] 
  |  |                 |  |                              
  |  |                 |  +-- Lamellidens_mainwaringi[2] 
  |  |                 |  |                              
  |  |                 +--+-- Lamellidens_lamellatus     
  |  |                    |                              
  |  |                    +-- Lamellidens_generosus      
  |  |                    |                              
  |  |                    +-- Lamellidens_jenkinsianus   
  |  |                                                   
  |  |                 +----- Leoparreysia_tavoyensis    
  |  |  +--------------+                                 
  |  |  |              |  +-- Leoparreysia_canefrii      
  |  |  |              +--+                              
  |  |  |                 +-- Leoparreysia_olivacea      
  |  |  |                                                
  |  |  |           +-------- Coelatura_aegyptiaca[2]    
  |  |  |           |                                    
  |  |  |           +-------- Nitia_teretiuscula         
  |  +--+  +--------+                                    
  |     |  |        |  +----- Coelatura_aegyptiaca[1]    
  |     |  |        |  |                                 
  |     |  |        +--+  +-- Coelatura_gabonensis       
  |     |  |           |  |                              
  |     |  |           +--+-- Coelatura_kunenensis       
  |     |  |              |                              
  |     |  |              +-- Prisodontopsis_mweruensis  
  |     +--+                                             
  |        |           +----- Indonaia_shurtleffiana     
  |        |           |                                 
  |        |           |  +-- Indonaia_bonneaudii        
  |        |  +--------+--+                              
  |        |  |        |  +-- Indonaia_rugosa            
  |        |  |        |                                 
  |        |  |        |  +-- Indonaia_andersoniana      
  |        |  |        +--+                              
  |        +--+           +-- Indonaia_caerulea          
  |           |                                          
  |           |  +----------- Indochinella_pugio         
  |           |  |                                       
  |           |  |     +----- Scabies_humilis            
  |           +--+  +--+                                 
  |              |  |  |  +-- Scabies_phaselus           
  |              |  |  +--+                              
  |              +--+     +-- Scabies_mandarinus         
  |                 |                                    
  |                 |  +----- Radiatula_mouhoti          
  |                 +--+                                 
  |                    |  +-- Radiatula_myitkyinae       
  |                    +--+                              
  |                       +-- Radiatula_chaudhurii       
  |                                                      
  |     +-------------------- Chamberlainia_somsakpanhai 
  |     |                                                
  |     +-------------------- Megalonaias_nervosa **        
  |     |                                                
  |     +-------------------- Uniomerus_carolinianus **     
  |     |                                                
  |     |                 +-- Quadrula_quadrula **          
  |     +-----------------+                              
  |     |                 +-- Quadrula_verrucosa **         
  |     |                                                
  |     |                 +-- Sinohyriopsis_cumingii     
  |     +-----------------+                              
  |     |                 +-- Sinohyriopsis_schlegelii   
  |     |                                                
  |     |                 +-- Potomida_littoralis        
  |     |           +-----+                              
  |     |           |     +-- Pronodularia_japanensis    
  |     +-----------+                                    
  |     |           |  +----- Lamprotula_leaii           
  |     |           +--+                                 
  |     |              |  +-- Lamprotula_caveata         
  |     |              +--+                              
  |     |                 +-- Lamprotula_cornuumlunae    
  |     |                                                
  |     |           +-------- Sinosolenaia_carinata      
--+     |           |                                    
  |     |           |  +----- Gonidea_angulata           
  |     |           +--+                                 
  |     |           |  |  +-- Leguminaia_wheatleyi       
  |     |           |  +--+                              
  |     |           |     +-- Microcondylaea_bonellii    
  |     +-----------+                                    
  |     |           |  +----- Ptychorhynchus_pfisteri    
  |     |           +--+                                 
  |     |           |  |  +-- Parvasolenaia_rivularis    
  |     |           |  +--+                              
  |     |           |     +-- Koreosolenaia_sitgyensis   
  |     |           |                                    
  |     |           |  +----- Sinosolenaia_recognita     
  |     |           +--+                                 
  |     |              |  +-- Sinosolenaia_iridinea      
  |     |              +--+                              
  |     |                 +-- Sinosolenaia_oleivora      
  |     |                                                
  |     |              +----- Indopseudodon_salwenianus  
  |     |        +-----+                                 
  |     |        |     |  +-- Indopseudodon_indawgyiensis
  |     |        |     +--+                              
  |     |        |        +-- Indopseudodon_bogani       
  |     |        |                                       
  |     |        |  +-------- Nyeinchanconcha_nyeinchani 
  |     |        |  |                                    
  |     |        |  +-------- Thaiconcha_callifera       
  |     +--------+  |                                    
  |     |        |  +-------- Sundadontina_cumingii      
  |     |        |  |                                    
  |     |        |  +-------- Sundadontina_tumida[1]     
  |     |        |  |                                    
  |     |        |  +-------- Sundadontina_tumida[2]     
  |     |        |  |                                    
  |     |        |  |  +----- Pilsbryoconcha_exilis[2]   
  |     |        +--+--+                                 
  |     |           |  |  +-- Pilsbryoconcha_carinifera  
  |     |           |  +--+                              
  |     |           |     +-- Pilsbryoconcha_exilis[1]   
  |     |           |                                    
  |     |           |  +----- Pseudodon_vagulus          
  |     |           +--+                                 
  |     |           |  |  +-- Pseudodon_mekongi          
  |     |           |  +--+                              
  |     |           |     +-- Pseudodon_vondembuschianus 
  |     |           |                                    
  |     |           |  +----- Bineurus_loeiensis         
  |     |           +--+                                 
  |     |              |  +-- Bineurus_mouhotii          
  |     |              +--+                              
  |     |                 +-- Bineurus_exilis            
  |     |                                                
  |     |     +-------------- Amblema_plicata **            
  |     |     |                                          
  |     |     +-------------- Toxolasma_parvum **           
  |     |     |                                          
  |     |     |           +-- Obliquaria_reflexa **         
  |     |     +-----------+                              
  |     |     |           +-- Plectomerus_dombeyanus **     
  |     |     |                                          
  |     |     |  +----------- Elliptoideus_sloatianus **    
  |     |     +--+                                       
  |     |     |  |  +-------- Elliptio_complanata **        
  |     |     |  +--+                                    
  |     |     |     |  +----- Eurynia_dilatata **           
  |     +-----+     +--+                                 
  |     |     |        |  +-- Fusconaia_flava **            
  |     |     |        +--+                              
  |     |     |           +-- Pleurobema_sintoxia **        
  |     |     |                                          
  |     |     |     +-------- Ellipsaria_lineolata **       
  |     |     |     |                                    
  |     |     |     +-------- Potamilus_leptodon **         
  |     |     |     |                                    
  |     |     |     +-------- Medionidus_conradicus **      
  |     |     |     |                                    
  +-----+     |     +-------- Potamilus_alatus **           
        |     |     |                                    
        |     |     +-------- Ptychobranchus_subtentus **   
        |     +-----+                                    
        |           +-------- Truncilla_truncata **         
        |           |                                    
        |           +-------- Venustaconcha_ellipsiformis **
        |           |                                    
        |           +-------- Cambarunio_iris **            
        |           |                                    
        |           +-------- Leaunio_lienosus **           
        |           |                                    
        |           |  +----- Ortmanniana_ligamentina **    
        |           +--+                                 
        |              |  +-- Lampsilis_cardium **          
        |              +--+                              
        |                 +-- Lampsilis_ornata **           
        |                                                
        |     +-------------- Hyriopsis_desowitzi        
        |     |                                          
        |     +-------------- Hyriopsis_myersiana        
        |     |                                          
        |     +-------------- Hyriopsis_panhai           
        |     |                                          
        |  +--+           +-- Hyriopsis_bialata          
        |  |  +-----------+                              
        |  |  |           +-- Hyriopsis_khoratensis      
        |  |  |                                          
        |  |  |  +----------- Rectidens_sumatrensis      
        |  |  |  |                                       
        |  |  +--+        +-- Ensidens_telus             
        |  |     |  +-----+                              
        |  |     |  |     +-- Ensidens_ingallsianus      
        |  |     +--+                                    
        |  |        |  +----- Ensidens_sagittarius       
        |  |        +--+                                 
        +--+           |  +-- Ensidens_spiculus          
        |  |           +--+                              
        |  |              +-- Ensidens_jaculus           
        |  |                                             
        |  |     +----------- Physunio_modelli           
        |  |     |                                       
        |  |     +----------- Lens                       
        |  |     |                                       
        |  |     +----------- Physunio_superbus          
        |  |     |                                       
        |  |     |  +-------- Yaukthwa_inlenensis        
        |  |     +--+                                    
        |  +-----+  |  +----- Yaukthwa_panhai            
        |        |  +--+                                 
        |        |     |  +-- Yaukthwa_nesemanni         
        |        |     +--+                              
        |        |        +-- Yaukthwa_rectangularis     
        |        |                                       
        |        |        +-- Lens_comptus               
        |        |  +-----+                              
        |        |  |     +-- Lens_novoselovi            
        |        +--+                                    
        |           |  +----- Lens_eximius               
        |           +--+                                 
        |              |  +-- Lens_contradens            
        |              +--+                              
        |                 +-- Lens_micropterus           
        |                                                
        |  +----------------- Acuticosta_chinensis       
        |  |                                             
        |  +----------------- Alasmidonta_marginata      
        |  |                                             
        |  +----------------- Anodonta_anatina           
        |  |                                             
        |  +----------------- Anemina_arcaeformis        
        |  |                                             
        |  +----------------- Anodonta_cygnea            
        |  |                                             
        |  +----------------- Cristaria_plicata          
        |  |                                             
        |  +----------------- Anodonta_nuttalliana       
        |  |                                             
        |  +----------------- Lasmigona_costata          
        |  |                                             
        |  +----------------- Lepidodesma_languilati     
        |  |                                             
        |  +----------------- Pseudanodonta_complanata   
        |  |                                             
        |  +----------------- Pyganodon_grandis          
        |  |                                             
        |  +----------------- Simpsonaias_ambigua        
        |  |                                             
        |  +----------------- Strophitus_undulatus       
        |  |                                             
        |  |              +-- Schistodesmus_lampreyanus  
        |  +--------------+                              
        |  |              +-- Schistodesmus_spinosus     
        |  |                                             
        |  |              +-- Utterbackia_imbecillis     
        |  +--------------+                              
        |  |              +-- Utterbackia_peninsularis   
        |  |                                             
        |  |        +-------- Lanceolaria_gladiola       
        |  +--------+                                    
        |  |        |  +----- Lanceolaria_grayii         
        |  |        +--+                                 
        +--+           |  +-- Lanceolaria_lanceolata     
           |           +--+                              
           |              +-- Lanceolaria_triformis      
           |                                             
           |        +-------- Unio_tumidus               
           +--------+                                    
           |        |  +----- Unio_crassus               
           |        +--+                                 
           |           |  +-- Unio_delphinus             
           |           +--+                              
           |              +-- Unio_pictorum              
           |                                             
           |        +-------- Aculamprotula_tortuosa     
           |        |                                    
           +--------+     +-- Aculamprotula_coreana      
           |        |  +--+                              
           |        |  |  +-- Aculamprotula_scripta      
           |        +--+                                 
           |           |  +-- Aculamprotula_fibrosa      
           |           +--+                              
           |              +-- Aculamprotula_tientsinensis
           |                                             
           |        +-------- Cuneopsis_heudei           
           |        |                                    
           |        +-------- Cuneopsis_celtiformis      
           |        |                                    
           +--------+-------- Tchangsinaia_piscicula     
           |        |                                    
           |        +-------- Cuneopsis_rufescens        
           |        |                                    
           |        |  +----- Nodularia_douglasiae[2]    
           |        +--+                                 
           |           |  +-- Nodularia_douglasiae[1]    
           |           +--+                              
           |              +-- Nodularia_nuxpersicae      
           |                                             
           |  +-------------- Pletholophus_tenuis        
           |  |                                          
           +--+           +-- Sinanodonta_angula         
              |  +--------+                              
              |  |        +-- Sinanodonta_lucida         
              +--+                                       
                 |  +-------- Sinanodonta_woodiana[1]    
                 +--+                                    
                    |  +----- Sinanodonta_jourdyi        
                    +--+                                 
                       |  +-- Sinanodonta_elliptica      
                       +--+                              
                          +-- Sinanodonta_woodiana[2]    

Lopes-Lima et al., 2019 [Publ 3901], fig. 3

iT tT bi riS oT roT roS iD
25 25 23 4 4 2 1 30.0
                                   +-- Anodonta_nuttalliana  
  +--------------------------------+                         
  |                                +-- Pyganodon_grandis     
  |                                                          
  |  +-------------------------------- Megalonaias_nervosa **   
--+  |                                                       
  |  |                             +-- Pleurobema_oviforme **   
  |  |                       +-----+                         
  |  |                       |     +-- Pleurobema_riddellii **  
  |  |                       |                               
  +--+  +--------------------+     +-- Lampsilis_ornata **      
     |  |                    |  +--+                         
     |  |                    |  |  +-- Potamilus_leptodon **    
     |  |                    +--+                            
     |  |                       |  +-- Amblema_elliottii **     
     |  |                       +--+                         
     +--+                          +-- Amblema_plicata **       
        |                                                    
        |  +-------------------------- Uniomerus_tetralasmus ** 
        |  |                                                 
        |  |              +----------- Theliderma_cylindrica ** 
        |  |  +-----------+                                  
        |  |  |           |  +-------- Theliderma_sparsa **     
        +--+  |           +--+                               
           |  |              |  +----- Theliderma_intermedia ** 
           |  |              +--+                            
           |  |                 |  +-- Theliderma_metanevra **  
           |  |                 +--+                         
           +--+                    +-- Theliderma_johnsoni **   
              |                                              
              |                 +----- Quadrula_quadrula **     
              |  +--------------+                            
              |  |              |  +-- Quadrula_nobilis **      
              |  |              +--+                         
              |  |                 +-- Quadrula_verrucosa **    
              +--+                                           
                 |  +----------------- Pustulosa_kieneriana **  
                 |  |                                        
                 |  |              +-- Pustulosa_infucata **    
                 +--+  +-----------+                         
                    |  |           +-- Pustulosa_kleiniana **   
                    |  |                                     
                    +--+           +-- Pustulosa_pustulosa **   
                       |  +--------+                         
                       |  |        +-- Pustulosa_succissa **    
                       +--+                                  
                          |  +-------- Cyclonaias_tuberculata **
                          +--+                               
                             |  +----- Pustulosa_necki **       
                             +--+                            
                                |  +-- Pustulosa_petrina **     
                                +--+                         
                                   +-- Pustulosa_nodulata **    

Lopes-Lima et al., 2019 [Publ 3901], fig. 2

iT tT bi riS oT roT roS iD
24 24 21 4 4 2 1 33.0
                                      +-- Anodonta_nuttalliana  
  +-----------------------------------+                         
  |                                   +-- Pyganodon_grandis     
  |                                                             
  |                                   +-- Amblema_elliottii **     
--+  +--------------------------------+                         
  |  |                                +-- Amblema_plicata **       
  |  |                                                          
  |  |                                +-- Lampsilis_ornata **      
  +--+  +-----------------------------+                         
     |  |                             +-- Potamilus_leptodon **    
     |  |                                                       
     |  |                             +-- Pleurobema_riddellii **  
     +--+  +--------------------------+                         
        |  |                          +-- Elliptio_crassidens **   
        |  |                                                    
        |  |                       +----- Megalonaias_nervosa **   
        +--+                       |                            
           |  +--------------------+  +-- Uniomerus_carolinianus **
           |  |                    |  |                         
           |  |                    +--+-- Uniomerus_declivis **    
           |  |                       |                         
           +--+                       +-- Uniomerus_tetralasmus ** 
              |                                                 
              |  +----------------------- Quadrula_quadrula **     
              |  |                                              
              |  |                    +-- Quadrula_verrucosa **    
              +--+  +-----------------+                         
                 |  |                 +-- Quadrula_nobilis **      
                 |  |                                           
                 +--+                 +-- Theliderma_metanevra **  
                    |  +--------------+                         
                    |  |              +-- Theliderma_johnsoni **   
                    |  |                                        
                    +--+           +----- Pustulosa_kieneriana **  
                       |  +--------+                            
                       |  |        |  +-- Pustulosa_infucata **    
                       |  |        +--+                         
                       +--+           +-- Pustulosa_kleiniana **   
                          |                                     
                          |  +----------- Cyclonaias_tuberculata **
                          |  |                                  
                          +--+        +-- Pustulosa_pustulosa **   
                             |  +-----+                         
                             |  |     +-- Pustulosa_succissa **    
                             +--+                               
                                |  +----- Pustulosa_nodulata **    
                                +--+                            
                                   |  +-- Pustulosa_necki **       
                                   +--+                         
                                      +-- Pustulosa_petrina **     

Krebs et al., 2003 [Publ 2691], fig. 3

iT tT bi riS oT roT roS iD
22 22 20 4 12 10 1 31.0
                    +----------- Utterbackiana_couperiana
                    |                                    
                    +----------- Platynaias_compressa    
                    |                                    
                    +----------- Alasmidonta_triangulata 
                    |                                    
  +-----------------+        +-- Lasmigona_complanata    
  |                 +--------+                           
  |                 |        +-- Lasmigona_costata       
  |                 |                                    
  |                 |        +-- Strophitus_radiatus     
  |                 |  +-----+                           
  |                 |  |     +-- Strophitus_undulatus    
  |                 +--+                                 
  |                    |  +----- Utterbackia_imbecillis  
  |                    +--+                              
  |                       |  +-- Pyganodon_grandis       
--+                       +--+                           
  |                          +-- Anodonta                
  |                                                      
  |                    +-------- Megalonaias_nervosa **     
  |  +-----------------+                                 
  |  |                 |  +----- Quadrula_quadrula[2] **    
  |  |                 +--+                              
  |  |                    |  +-- Quadrula_quadrula[1] **    
  |  |                    +--+                           
  |  |                       +-- Theliderma_metanevra **    
  |  |                                                   
  +--+              +----------- Elliptoideus_sloatianus ** 
     |  +-----------+                                    
     |  |           |  +-------- Pleurobema_pyriforme **    
     |  |           +--+                                 
     |  |              |  +----- Uniomerus_carolinianus **  
     |  |              +--+                              
     |  |                 |  +-- Fusconaia_flava **         
     +--+                 +--+                           
        |                    +-- Eurynia_dilatata **        
        |                                                
        |  +-------------------- Amblema_plicata **         
        |  |                                             
        |  |              +----- Toxolasma_parvum **        
        |  |  +-----------+                              
        +--+  |           |  +-- Plectomerus_dombeyanus **  
           |  |           +--+                           
           |  |              +-- Obliquaria_reflexa **      
           |  |                                          
           +--+           +----- Ptychobranchus_subtentus **
              |  +--------+                              
              |  |        |  +-- Potamilus_fragilis **      
              |  |        +--+                           
              |  |           +-- Medionidus_conradicus **   
              +--+                                       
                 |           +-- Villosa_delumbis **        
                 |  +--------+                           
                 |  |        +-- Ellipsaria_lineolata **    
                 +--+                                    
                    |  +-------- Lampsilis_teres **         
                    +--+                                 
                       |  +----- Lampsilis_siliquoidea **   
                       +--+                              
                          |  +-- Potamilus_purpuratus **    
                          +--+                           
                             +-- Potamilus_alatus **        

Lopes-Lima et al., 2019 [Publ 3901], fig. 4

iT tT bi riS oT roT roS iD
22 22 20 4 4 2 1 29.0
                                   +-- Anodonta_nuttalliana  
  +--------------------------------+                         
  |                                +-- Pyganodon_grandis     
  |                                                          
  |                                +-- Amblema_elliottii **     
  |                          +-----+                         
  |                          |     +-- Amblema_plicata **       
--+                          |                               
  |  +-----------------------+     +-- Pleurobema_riddellii **  
  |  |                       |  +--+                         
  |  |                       |  |  +-- Pleurobema_oviforme **   
  |  |                       +--+                            
  |  |                          |  +-- Lampsilis_ornata **      
  +--+                          +--+                         
     |                             +-- Potamilus_leptodon **    
     |                                                       
     |  +----------------------------- Megalonaias_nervosa **   
     |  |                                                    
     +--+  +-------------------------- Uniomerus_tetralasmus ** 
        |  |                                                 
        |  |                       +-- Theliderma_metanevra **  
        +--+  +--------------------+                         
           |  |                    +-- Theliderma_johnsoni **   
           |  |                                              
           +--+  +-------------------- Quadrula_quadrula **     
              |  |                                           
              |  |                 +-- Quadrula_verrucosa **    
              +--+  +--------------+                         
                 |  |              +-- Quadrula_nobilis **      
                 |  |                                        
                 |  |           +----- Pustulosa_kieneriana **  
                 +--+  +--------+                            
                    |  |        |  +-- Pustulosa_infucata **    
                    |  |        +--+                         
                    |  |           +-- Pustulosa_kleiniana **   
                    +--+                                     
                       |           +-- Pustulosa_pustulosa **   
                       |  +--------+                         
                       |  |        +-- Pustulosa_succissa **    
                       +--+                                  
                          |  +-------- Cyclonaias_tuberculata **
                          +--+                               
                             |  +----- Pustulosa_necki **       
                             +--+                            
                                |  +-- Pustulosa_petrina **     
                                +--+                         
                                   +-- Pustulosa_nodulata **    

Chapman et al., 2008 [Publ 2667], fig. 1

iT tT bi riS oT roT roS iD
20 20 18 5 1 1 1 26.0
  +-------------------------------------------- Pronodularia_japanensis    
  |                                                                        
  |                                         +-- Quadrula_quadrula **          
--+  +--------------------------------------+                              
  |  |                                      +-- Pustulosa_pustulosa **        
  |  |                                                                     
  +--+                                      +-- Pleurobema_sintoxia **        
     |  +-----------------------------------+                              
     |  |                                   +-- Fusconaia_flava **            
     +--+                                                                  
        |  +----------------------------------- Amblema_plicata **            
        |  |                                                               
        +--+  +-------------------------------- Plectomerus_dombeyanus **     
           |  |                                                            
           +--+  +----------------------------- Popenaias_popeii **           
              |  |                                                         
              +--+  +-------------------------- Toxolasma_lividum **          
                 |  |                                                      
                 +--+  +----------------------- Obliquaria_reflexa **         
                    |  |                                                   
                    |  |                    +-- Cyrtonaias_tampicoensis **    
                    +--+  +-----------------+                              
                       |  |                 +-- Glebula_rotundata **          
                       |  |                                                
                       +--+                 +-- Ptychobranchus_fasciolaris ** 
                          |  +--------------+                              
                          |  |              +-- Lemiox_rimosus **             
                          |  |                                             
                          +--+              +-- Venustaconcha_ellipsiformis **
                             |  +-----------+                              
                             |  |           +-- Obovaria_olivaria **          
                             +--+                                          
                                |  +----------- Hamiota_subangulata **        
                                +--+                                       
                                   |  +-------- Lampsilis_ovata **            
                                   +--+                                    
                                      |  +----- Ortmanniana_ligamentina **    
                                      +--+                                 
                                         |  +-- Lampsilis_straminea **        
                                         +--+                              
                                            +-- Lampsilis_hydiana **          

Chapman et al., 2008 [Publ 2744], fig. 2

iT tT bi riS oT roT roS iD
20 20 18 5 1 1 1 26.0
  +-------------------------------------------- Pronodularia_japanensis    
  |                                                                        
  |                                         +-- Quadrula_quadrula **          
--+  +--------------------------------------+                              
  |  |                                      +-- Pustulosa_pustulosa **        
  |  |                                                                     
  +--+                                      +-- Pleurobema_sintoxia **        
     |  +-----------------------------------+                              
     |  |                                   +-- Fusconaia_flava **            
     +--+                                                                  
        |  +----------------------------------- Amblema_plicata **            
        |  |                                                               
        +--+  +-------------------------------- Plectomerus_dombeyanus **     
           |  |                                                            
           +--+  +----------------------------- Popenaias_popeii **           
              |  |                                                         
              +--+  +-------------------------- Toxolasma_lividum **          
                 |  |                                                      
                 +--+  +----------------------- Obliquaria_reflexa **         
                    |  |                                                   
                    |  |                    +-- Cyrtonaias_tampicoensis **    
                    +--+  +-----------------+                              
                       |  |                 +-- Glebula_rotundata **          
                       |  |                                                
                       +--+                 +-- Ptychobranchus_fasciolaris ** 
                          |  +--------------+                              
                          |  |              +-- Lemiox_rimosus **             
                          |  |                                             
                          +--+              +-- Venustaconcha_ellipsiformis **
                             |  +-----------+                              
                             |  |           +-- Obovaria_olivaria **          
                             +--+                                          
                                |  +----------- Hamiota_subangulata **        
                                +--+                                       
                                   |  +-------- Lampsilis_ovata **            
                                   +--+                                    
                                      |  +----- Ortmanniana_ligamentina **    
                                      +--+                                 
                                         |  +-- Lampsilis_straminea **        
                                         +--+                              
                                            +-- Lampsilis_hydiana **          

Porto-Hannes et al., 2019 [Publ 3969], fig. 2

iT tT bi riS oT roT roS iD
20 20 16 2 0 0 0 27.0
  +-------------------------- Amblema_plicata **        
  |                                                  
  |  +----------------------- Obliquaria_reflexa **     
  |  |                                               
--+  |                    +-- Cambarunio_iris **        
  |  |  +-----------------+                          
  |  |  |                 +-- Leaunio_lienosus **       
  |  |  |                                            
  +--+  |        +----------- Lampsilis_teres **        
     |  |        |                                   
     |  +--------+  +-------- Ortmanniana_ligamentina **
     |  |        |  |                                
     |  |        +--+     +-- Lampsilis_hydiana **      
     |  |           |  +--+                          
     +--+           |  |  +-- Lampsilis_virescens **    
        |           +--+                             
        |              |  +-- Lampsilis_siliquoidea **  
        |              +--+                          
        |                 +-- Lampsilis_radiata **      
        |                                            
        |              +----- Obovaria_arkansasensis ** 
        |  +-----------+                             
        |  |           |  +-- Obovaria_olivaria **      
        |  |           +--+                          
        +--+              +-- Obovaria_subrotunda **    
           |                                         
           |  +-------------- Lampsilis_bracteata **    
           |  |                                      
           +--+  +----------- Lampsilis_ornata **       
              |  |                                   
              +--+  +-------- Lampsilis_ovata **        
                 |  |                                
                 +--+  +----- Lampsilis_cardium[3] **   
                    |  |                             
                    +--+  +-- Lampsilis_satura **       
                       |  |                          
                       +--+-- Lampsilis_cardium[1] **   
                          |                          
                          +-- Lampsilis_cardium[2] **   

Porto-Hannes et al., 2019 [Publ 3969], fig. 3

iT tT bi riS oT roT roS iD
20 20 13 2 0 0 0 27.0
  +-------------------- Obliquaria_reflexa **     
  |                                            
  +-------------------- Amblema_plicata **        
  |                                            
  |  +----------------- Lampsilis_teres **        
  |  |                                         
  |  |              +-- Leaunio_lienosus **       
  |  +--------------+                          
--+  |              +-- Cambarunio_iris **        
  |  |                                         
  |  |           +----- Obovaria_arkansasensis ** 
  |  +-----------+                             
  |  |           |  +-- Obovaria_olivaria **      
  |  |           +--+                          
  |  |              +-- Obovaria_subrotunda **    
  |  |                                         
  +--+        +-------- Ortmanniana_ligamentina **
     |        |                                
     +--------+     +-- Lampsilis_hydiana **      
     |        |  +--+                          
     |        |  |  +-- Lampsilis_virescens **    
     |        +--+                             
     |           |  +-- Lampsilis_siliquoidea **  
     |           +--+                          
     |              +-- Lampsilis_radiata **      
     |                                         
     |  +-------------- Lampsilis_bracteata **    
     |  |                                      
     +--+  +----------- Lampsilis_ornata **       
        |  |                                   
        +--+  +-------- Lampsilis_ovata **        
           |  |                                
           +--+  +----- Lampsilis_cardium[3] **   
              |  |                             
              +--+  +-- Lampsilis_satura **       
                 |  |                          
                 +--+-- Lampsilis_cardium[1] **   
                    |                          
                    +-- Lampsilis_cardium[2] **   

Smith et al., 2020 [Publ 4136], fig. 1

iT tT bi riS oT roT roS iD
20 20 13 2 0 0 0 33.0
  +-------------------------- Quadrula_quadrula **       
  |                                                   
--+  +----------------------- Elliptoideus_sloatianus ** 
  |  |                                                
  |  |  +-------------------- Elliptio_crassidens **     
  +--+  |                                             
     |  +-------------------- Plethobasus_cyphyus **     
     |  |                                             
     +--+  +----------------- Parvaspina_steinstansana **
        |  |                                          
        |  |              +-- Hemistena_lata **          
        |  +--------------+                           
        |  |              +-- Eurynia_dilatata **        
        +--+                                          
           |              +-- Pleuronaia_barnesiana **   
           |  +-----------+                           
           |  |           +-- Pleurobema_clava **        
           |  |                                       
           |  |  +----------- Fusconaia_subrotunda **    
           +--+  |                                    
              |  |        +-- Fusconaia_cor **           
              |  +--------+                           
              |  |        +-- Fusconaia_cuneolus **      
              +--+                                    
                 |  +-------- Fusconaia_flava **         
                 |  |                                 
                 |  +-------- Fusconaia_masoni **        
                 |  |                                 
                 |  |     +-- Fusconaia_burkei **        
                 +--+-----+                           
                    |     +-- Fusconaia_escambia **      
                    |                                 
                    |     +-- Fusconaia_iheringi **      
                    |  +--+                           
                    |  |  +-- Fusconaia_mitchelli **     
                    +--+                              
                       |  +-- Fusconaia_askewi **        
                       +--+                           
                          +-- Fusconaia_chunii **        

Breton et al., 2011 [Publ 3376], fig. 3

iT tT bi riS oT roT roS iD
19 34 31 4 20 15 2 38.0
                             +----- Quadrula_quadrula **          
  +--------------------------+                                 
  |                          |  +-- Cyclonaias_tuberculata **     
  |                          +--+                              
  |                             +-- Pustulosa_pustulosa **        
  |                                                            
  |     +-------------------------- Pronodularia_japanensis    
  |     |                                                      
  |  +--+  +----------------------- Sinohyriopsis_cumingii     
  |  |  |  |                                                   
  |  |  +--+  +-------------------- Cristaria_plicata          
  |  |     |  |                                                
--+  |     |  |                 +-- Lasmigona_complanata       
  |  |     +--+  +--------------+                              
  |  |        |  |              +-- Lasmigona_costata          
  |  |        |  |                                             
  |  |        +--+  +-------------- Strophitus_undulatus       
  |  |           |  |                                          
  |  |           |  |           +-- Platynaias_compressa       
  |  |           |  |        +--+                              
  |  |           +--+        |  +-- Platynaias_subviridis      
  |  |              |  +-----+                                 
  |  |              |  |     |  +-- Alasmidonta_marginata      
  +--+              |  |     +--+                              
     |              |  |        +-- Alasmidonta_undulata       
     |              +--+                                       
     |                 |        +-- Pyganodon_lacustris        
     |                 |  +-----+                              
     |                 |  |     +-- Pyganodon_grandis          
     |                 +--+                                    
     |                    |  +----- Utterbackia_peninsularis[2]
     |                    +--+                                 
     |                       |  +-- Utterbackia_peninsularis[1]
     |                       +--+                              
     |                          +-- Utterbackia_imbecillis     
     |                                                         
     |  +-------------------------- Megalonaias_nervosa **        
     |  |                                                      
     +--+  +----------------------- Fusconaia_flava **            
        |  |                                                   
        +--+  +-------------------- Reginaia_ebenus **            
           |  |                                                
           |  |           +-------- Toxolasma_paulum[2] **        
           +--+  +--------+                                    
              |  |        |  +----- Toxolasma_lividum **          
              |  |        +--+                                 
              |  |           |  +-- Toxolasma_parvum **           
              +--+           +--+                              
                 |              +-- Toxolasma_paulum[1] **        
                 |                                             
                 |  +-------------- Ellipsaria_lineolata **       
                 |  |                                          
                 +--+  +----------- Truncilla_macrodon **         
                    |  |                                       
                    |  |        +-- Lemiox_rimosus **             
                    +--+  +-----+                              
                       |  |     +-- Potamilus_metnecktayi **      
                       |  |                                    
                       +--+     +-- Cambarunio_iris **            
                          |  +--+                              
                          |  |  +-- Venustaconcha_ellipsiformis **
                          +--+                                 
                             |  +-- Lampsilis_powellii **         
                             |  |                              
                             +--+-- Lampsilis_ornata **           
                                |                              
                                +-- Lampsilis_siliquoidea **      

Lydeard et al., 1996 [Publ 2656], fig. 3

iT tT bi riS oT roT roS iD
18 18 14 4 9 6 1 30.0
                          +----- Utterbackia_imbecillis  
                       +--+                              
                       |  |  +-- Utterbackiana_couperiana
                       |  +--+                           
  +--------------------+     +-- Utterbackiana           
  |                    |                                 
  |                    |  +----- Strophitus_radiatus     
  |                    +--+                              
  |                       |  +-- Lasmigona_costata       
  |                       +--+                           
  |                          +-- Alasmidonta_triangulata 
--+                                                      
  |                       +----- Megalonaias_nervosa **     
  |  +--------------------+                              
  |  |                    |  +-- Quadrula_quadrula **       
  |  |                    +--+                           
  |  |                       +-- Theliderma_metanevra **    
  |  |                                                   
  +--+                    +----- Elliptoideus_sloatianus ** 
     |                    |                              
     |  +-----------------+  +-- Eurynia_dilatata **        
     |  |                 |  |                           
     |  |                 +--+-- Pleurobema_pyriforme **    
     |  |                    |                           
     +--+                    +-- Uniomerus_carolinianus **  
        |                                                
        |  +-------------------- Amblema_plicata **         
        |  |                                             
        +--+                 +-- Plectomerus_dombeyanus **  
           |  +--------------+                           
           |  |              +-- Obliquaria_reflexa **      
           +--+                                          
              |  +-------------- Potamilus_fragilis **      
              |  |                                       
              +--+  +----------- Ptychobranchus_subtentus **
                 |  |                                    
                 +--+  +-------- Medionidus_conradicus **   
                    |  |                                 
                    +--+  +----- Potamilus_purpuratus **    
                       |  |                              
                       |  |  +-- Ellipsaria_lineolata **    
                       +--+--+                           
                          |  +-- Villosa_delumbis **        
                          |                              
                          |  +-- Lampsilis_teres **         
                          +--+                           
                             +-- Lampsilis_siliquoidea **   

Lydeard et al., 1996 [Publ 2656], fig. 4

iT tT bi riS oT roT roS iD
18 18 16 4 10 6 1 30.0
                                   +----- Utterbackia_imbecillis  
                                +--+                              
                                |  |  +-- Utterbackiana_couperiana
                                |  +--+                           
  +-----------------------------+     +-- Anodonta                
  |                             |                                 
  |                             |  +----- Strophitus_radiatus     
  |                             +--+                              
  |                                |  +-- Lasmigona_costata       
  |                                +--+                           
  |                                   +-- Alasmidonta_triangulata 
--+                                                               
  |                                +----- Megalonaias_nervosa **     
  |  +-----------------------------+                              
  |  |                             |  +-- Quadrula_quadrula **       
  |  |                             +--+                           
  |  |                                +-- Theliderma_metanevra **    
  |  |                                                            
  +--+                          +-------- Elliptoideus_sloatianus ** 
     |  +-----------------------+                                 
     |  |                       |  +----- Uniomerus_carolinianus **  
     |  |                       +--+                              
     |  |                          |  +-- Eurynia_dilatata **        
     +--+                          +--+                           
        |                             +-- Pleurobema_pyriforme **    
        |                                                         
        |  +----------------------------- Amblema_plicata **         
        +--+                                                      
           |  +-------------------------- Plectomerus_dombeyanus **  
           +--+                                                   
              |  +----------------------- Obliquaria_reflexa **      
              +--+                                                
                 |  +-------------------- Medionidus_conradicus **   
                 +--+                                             
                    |  +----------------- Ptychobranchus_subtentus **
                    +--+                                          
                       |  +-------------- Potamilus_fragilis **      
                       +--+                                       
                          |  +----------- Potamilus_purpuratus **    
                          +--+                                    
                             |  +-------- Ellipsaria_lineolata **    
                             +--+                                 
                                |  +----- Villosa_delumbis **        
                                +--+                              
                                   |  +-- Lampsilis_teres **         
                                   +--+                           
                                      +-- Lampsilis_siliquoidea **   

Graf & Ó Foighil, 2000 [Publ 2648], fig. 2A

iT tT bi riS oT roT roS iD
17 17 11 4 13 8 1 28.0
                       +-- Unio_pictorum             
  +--------------------+                             
  |                    +-- Unio_caffer               
  |                                                  
  |           +----------- Anodonta_cygnea           
  |           |                                      
  |  +--------+        +-- Alasmidonta_marginata     
--+  |        |  +-----+                             
  |  |        |  |     +-- Lasmigona_costata         
  |  |        +--+                                   
  |  |           |  +----- Strophitus_undulatus      
  |  |           +--+                                
  |  |              |  +-- Pyganodon_grandis         
  |  |              +--+                             
  +--+                 +-- Pyganodon_fragilis        
     |                                               
     |           +-------- Quadrula_quadrula **         
     |  +--------+                                   
     |  |        |  +----- Eurynia_dilatata **          
     |  |        +--+                                
     |  |           |  +-- Pleurobema_sintoxia **       
     |  |           +--+                             
     +--+              +-- Fusconaia_flava **           
        |                                            
        |  +-------------- Amblema_plicata **           
        |  |                                         
        |  |  +----------- Ligumia_recta **             
        |  |  |                                      
        +--+  +----------- Truncilla_truncata **        
           |  |                                      
           |  |        +-- Ptychobranchus_fasciolaris **
           |  +--------+                             
           +--+        +-- Sagittunio_nasutus **        
              |                                      
              |     +----- Lampsilis_siliquoidea **     
              |  +--+                                
              |  |  |  +-- Epioblasma_triquetra **      
              |  |  +--+                             
              |  |     +-- Epioblasma_brevidens **      
              +--+                                   
                 |  +----- Ortmanniana_ligamentina **   
                 |  |                                
                 |  +----- Cambarunio_iris **           
                 +--+                                
                    +----- Leaunio_vanuxemensis **      
                    |                                
                    |  +-- Lampsilis_cardium **         
                    +--+                             
                       +-- Lampsilis_fasciola **        

Graf & Ó Foighil, 2000 [Publ 2648], fig. 2B

iT tT bi riS oT roT roS iD
17 17 15 4 13 8 1 28.0
                                +-- Unio_pictorum             
  +-----------------------------+                             
  |                             +-- Unio_caffer               
  |                                                           
  |                    +----------- Anodonta_cygnea           
  |                    |                                      
  |  +-----------------+        +-- Alasmidonta_marginata     
--+  |                 |  +-----+                             
  |  |                 |  |     +-- Lasmigona_costata         
  |  |                 +--+                                   
  |  |                    |  +----- Strophitus_undulatus      
  |  |                    +--+                                
  |  |                       |  +-- Pyganodon_grandis         
  +--+                       +--+                             
     |                          +-- Pyganodon_fragilis        
     |                                                        
     |                    +-------- Quadrula_quadrula **         
     |  +-----------------+                                   
     |  |                 |  +----- Eurynia_dilatata **          
     |  |                 +--+                                
     |  |                    |  +-- Pleurobema_sintoxia **       
     +--+                    +--+                             
        |                       +-- Fusconaia_flava **           
        |                                                     
        |  +----------------------- Amblema_plicata **           
        |  |                                                  
        +--+  +-------------------- Truncilla_truncata **        
           |  |                                               
           +--+  +----------------- Ligumia_recta **             
              |  |                                            
              |  |              +-- Ptychobranchus_fasciolaris **
              +--+  +-----------+                             
                 |  |           +-- Sagittunio_nasutus **        
                 |  |                                         
                 +--+  +----------- Lampsilis_siliquoidea **     
                    |  |                                      
                    |  |     +----- Ortmanniana_ligamentina **   
                    +--+  +--+                                
                       |  |  |  +-- Lampsilis_cardium **         
                       |  |  +--+                             
                       |  |     +-- Lampsilis_fasciola **        
                       +--+                                   
                          |     +-- Epioblasma_brevidens **      
                          |  +--+                             
                          |  |  +-- Epioblasma_triquetra **      
                          +--+                                
                             |  +-- Cambarunio_iris **           
                             +--+                             
                                +-- Leaunio_vanuxemensis **      

Grobler et al., 2011 [Publ 3359], fig. 2

iT tT bi riS oT roT roS iD
17 17 13 2 0 0 0 26.0
  +----------------- Amblema_plicata **        
  |                                         
  |     +----------- Lemiox_rimosus **         
  |  +--+                                   
--+  |  |  +-------- Medionidus_conradicus **  
  |  |  +--+                                
  |  |     |  +----- Dromus_dromas **          
  |  |     +--+                             
  |  |        |  +-- Cyprogenia_stegaria **    
  +--+        +--+                          
     |           +-- Cyprogenia_aberti **      
     |                                      
     |  +----------- Potamilus_alatus **       
     |  |                                   
     |  |     +----- Lampsilis_siliquoidea **  
     |  |  +--+                             
     +--+  |  |  +-- Lampsilis_higginsii **    
        |  |  +--+                          
        |  |     +-- Ortmanniana_ligamentina **
        |  |                                
        |  |  +----- Lampsilis_fasciola **     
        +--+--+                             
           |  |  +-- Lampsilis_cardium **      
           |  +--+                          
           |     +-- Lampsilis_ovata **        
           |                                
           |  +----- Cambarunio_iris **        
           |  |                             
           +--+----- Ligumia_recta **          
              |                             
              |  +-- Epioblasma_brevidens **   
              +--+                          
                 +-- Epioblasma_capsaeformis **

Chase et al., 2018 [Publ 3847], fig. 4

iT tT bi riS oT roT roS iD
17 18 16 4 22 17 1 29.0
                          +-- Unio_delphinus             
     +--------------------+                              
     |                    +-- Unio_tumidus               
     |                                                   
  +--+  +-------------------- Sinanodonta_woodiana       
  |  |  |                                                
  |  |  |                 +-- Anodonta_cygnea            
  |  |  |  +--------------+                              
  |  +--+  |              +-- Anodonta_anatina           
  |     |  |                                             
  |     |  |              +-- Pyganodon_lacustris        
  |     |  |        +-----+                              
  |     +--+        |     +-- Pyganodon_grandis          
  |        |  +-----+                                    
  |        |  |     |  +----- Utterbackia_peninsularis[2]
  |        |  |     +--+                                 
  |        |  |        |  +-- Utterbackia_peninsularis[1]
  |        |  |        +--+                              
  |        +--+           +-- Utterbackia_imbecillis     
  |           |                                          
  |           |           +-- Lasmigona_costata          
  |           |  +--------+                              
  |           |  |        +-- Lasmigona_complanata       
  |           |  |                                       
--+           +--+        +-- Alasmidonta_undulata       
  |              |  +-----+                              
  |              |  |     +-- Alasmidonta_marginata      
  |              +--+                                    
  |                 |  +----- Strophitus_undulatus       
  |                 +--+                                 
  |                    |  +-- Platynaias_compressa       
  |                    +--+                              
  |                       +-- Platynaias_subviridis      
  |                                                      
  |                       +-- Echyridella_menziesii      
  |              +--------+                              
  |              |        +-- Reginaia_ebenus **            
  |     +--------+                                       
  |     |        |  +-------- Fusconaia_flava **            
  |     |        +--+                                    
  |     |           |  +----- Megalonaias_nervosa **        
  |     |           +--+                                 
  |     |              |  +-- Cyclonaias_tuberculata **     
  |     |              +--+                              
  +-----+                 +-- Pustulosa_pustulosa **        
        |                                                
        |           +-------- Toxolasma_parvum **           
        |  +--------+                                    
        |  |        |  +----- Toxolasma_lividum **          
        |  |        +--+                                 
        |  |           |  +-- Toxolasma_paulum **           
        |  |           +--+                              
        +--+              +-- Toxolasma_texasiense **       
           |                                             
           |  +-------------- Venustaconcha_ellipsiformis **
           |  |                                          
           |  |        +----- Lemiox_rimosus **             
           +--+  +-----+                                 
              |  |     |  +-- Lampsilis_powellii **         
              |  |     +--+                              
              +--+        +-- Lampsilis_siliquoidea **      
                 |                                       
                 |  +-------- Potamilus_metnecktayi **      
                 +--+                                    
                    |  +----- Cambarunio_iris **            
                    +--+                                 
                       |  +-- Ellipsaria_lineolata **       
                       +--+                              
                          +-- Truncilla_macrodon **         

Wu et al., 2023 [Publ 4339], fig. 4, S5-aa

iT tT bi riS oT roT roS iD
17 17 15 5 79 68 3 0.0
                                      +-- Quadrula_quadrula **           
           +--------------------------+                               
           |                          +-- Uniomerus_tetralasmus **       
           |                                                          
  +--------+                          +-- Elliptio_complanata **         
  |        |  +-----------------------+                               
  |        |  |                       +-- Pleurobema_oviforme **         
  |        +--+                                                       
  |           |  +----------------------- Amblema_plicata **             
  |           |  |                                                    
  |           +--+  +-------------------- Popenaias_popeii **            
  |              |  |                                                 
  |              +--+  +----------------- Toxolasma_parvum **            
  |                 |  |                                              
  |                 |  |           +----- Potamilus_leptodon **          
  |                 +--+  +--------+                                  
  |                    |  |        |  +-- Potamilus_alatus **            
  |                    |  |        +--+                               
  |                    |  |           +-- Potamilus_streckersoni **      
  |                    +--+                                           
  |                       |  +----------- Leaunio_lienosus **            
  |                       |  |                                        
  |                       |  |     +----- Lampsilis_ornata **            
  |                       +--+  +--+                                  
  |                          |  |  |  +-- Lampsilis_cardium **           
  |                          |  |  +--+                               
  |                          +--+     +-- Lampsilis_cariosa **           
  |                             |                                     
  |                             |  +----- Venustaconcha_ellipsiformis ** 
  |                             +--+                                  
  |                                |  +-- Lampsilis_powellii **          
  |                                +--+                               
  |                                   +-- Lampsilis_siliquoidea **       
--+                                                                   
  |                                +----- Chamberlainia_somsakpanhai  
  |              +-----------------+                                  
  |              |                 |  +-- Sinohyriopsis_cumingii      
  |              |                 +--+                               
  |              |                    +-- Sinohyriopsis_schlegelii    
  |              |                                                    
  |              |                    +-- Hyriopsis_bialata           
  |              |                 +--+                               
  |  +-----------+                 |  +-- Rectidens_sumatrensis       
  |  |           |  +--------------+                                  
  |  |           |  |              |  +-- Lens_contradens             
  |  |           |  |              +--+                               
  |  |           |  |                 +-- Physunio_superbus           
  |  |           |  |                                                 
  |  |           |  |        +----------- Potomida_littoralis         
  |  |           +--+  +-----+                                        
  |  |              |  |     |  +-------- Pronodularia_japanensis     
  |  |              |  |     +--+                                     
  |  |              |  |        |  +----- Lamprotula_leaii            
  |  |              |  |        +--+                                  
  |  |              |  |           |  +-- Lamprotula_caveata          
  |  |              +--+           +--+                               
  |  |                 |              +-- Lamprotula_cornuumlunae     
  |  |                 |                                              
  |  |                 |              +-- Pseudodon_vondembuschianus  
  |  |                 |  +-----------+                               
  |  |                 |  |           +-- Pilsbryoconcha_exilis       
  |  |                 +--+                                           
  |  |                    |  +----------- Microcondylaea_bonellii     
  +--+                    |  |                                        
     |                    +--+        +-- Parvasolenaia_rivularis     
     |                       |  +-----+                               
     |                       |  |     +-- Ptychorhynchus_pfisteri     
     |                       +--+                                     
     |                          |  +----- Sinosolenaia_carinata       
     |                          +--+                                  
     |                             |  +-- Sinosolenaia_iridinea       
     |                             +--+                               
     |                                +-- Sinosolenaia_oleivora       
     |                                                                
     |  +-------------------------------- Lamellidens_marginalis      
     |  |                                                             
     |  |                             +-- Lepidodesma_aligera         
     |  |  +--------------------------+                               
     |  |  |                          +-- Lepidodesma_languilati      
     |  |  |                                                          
     |  |  |                    +-------- Aculamprotula_tortuosa      
     |  |  |                    |                                     
     |  |  |     +--------------+     +-- Aculamprotula_coreana       
     |  |  |     |              |  +--+                               
     +--+  |     |              |  |  +-- Aculamprotula_tientsinensis 
        |  |     |              +--+                                  
        |  |     |                 |  +-- Aculamprotula_polysticta    
        |  |     |                 +--+                               
        |  |     |                    +-- Aculamprotula_scripta       
        |  |     |                                                    
        |  |  +--+        +-------------- Unio_tumidus                
        |  |  |  |  +-----+                                           
        |  |  |  |  |     |  +----------- Unio_crassus                
        |  |  |  |  |     +--+                                        
        +--+  |  |  |        |  +-------- Unio_elongatulus            
           |  |  |  |        +--+                                     
           |  |  |  |           |  +----- Unio_pictorum               
           |  |  |  |           +--+                                  
           |  |  |  |              |  +-- Unio_delphinus              
           |  |  +--+              +--+                               
           |  |     |                 +-- Unio_mancus                 
           |  |     |                                                 
           |  |     |                 +-- Nodularia_breviconcha       
           |  |     |              +--+                               
           |  |     |              |  +-- Nodularia_nipponensis       
           |  |     |  +-----------+                                  
           |  |     |  |           |  +-- Pseudocuneopsis_capitata    
           |  |     |  |           +--+                               
           |  |     +--+              +-- Pseudocuneopsis_sichuanensis
           |  |        |                                              
           |  |        |  +-------------- Tchangsinaia_piscicula      
           |  |        |  |                                           
           +--+        +--+           +-- Schistodesmus_lampreyanus   
              |           |  +--------+                               
              |           |  |        +-- Schistodesmus_spinosus      
              |           +--+                                        
              |              |  +-------- Cuneopsis_rufescens         
              |              +--+                                     
              |                 |  +----- Cuneopsis_celtiformis       
              |                 +--+                                  
              |                    |  +-- Cuneopsis_demangei          
              |                    +--+                               
              |                       +-- Cuneopsis_heudei            
              |                                                       
              |                 +-------- Acuticosta_chinensis        
              |        +--------+                                     
              |        |        |  +----- Lanceolaria_gladiola        
              |        |        +--+                                  
              |        |           |  +-- Lanceolaria_grayii          
              |        |           +--+                               
              |        |              +-- Lanceolaria_lanceolata      
              |        |                                              
              |        |           +----- Cristaria_plicata           
              |        |     +-----+                                  
              |        |     |     |  +-- Anemina_arcaeformis         
              +--------+     |     +--+                               
                       |  +--+        +-- Anemina_euscaphys           
                       |  |  |                                        
                       |  |  |  +-------- Beringiana_fukuharai        
                       |  |  +--+                                     
                       |  |     |  +----- Sinanodonta_lucida          
                       |  |     +--+                                  
                       |  |        |  +-- Sinanodonta_tumens          
                       |  |        +--+                               
                       |  |           +-- Sinanodonta_woodiana        
                       +--+                                           
                          |           +-- Anodonta_nuttalliana        
                          |     +-----+                               
                          |     |     +-- Pseudanodonta_complanata    
                          |  +--+                                     
                          |  |  |  +----- Anodonta_anatina            
                          |  |  +--+                                  
                          |  |     |  +-- Anodonta_cygnea             
                          |  |     +--+                               
                          |  |        +-- Anodonta_exulcerata         
                          +--+                                        
                             |     +----- Pyganodon_grandis           
                             |  +--+                                  
                             |  |  |  +-- Utterbackia_peninsularis    
                             |  |  +--+                               
                             |  |     +-- Utterbackia_imbecillis      
                             +--+                                     
                                |     +-- Alasmidonta_varicosa        
                                |  +--+                               
                                |  |  +-- Platynaias_compressa        
                                +--+                                  
                                   |  +-- Prolasmidonta_heterodon     
                                   +--+                               
                                      +-- Lasmigona_complanata        

Wu et al., 2024 [Publ 4350], fig. 8a, S1

iT tT bi riS oT roT roS iD
17 17 15 5 75 69 3 0.0
                                      +-- Quadrula_quadrula **           
           +--------------------------+                               
           |                          +-- Uniomerus_tetralasmus **       
           |                                                          
  +--------+                          +-- Elliptio_complanata **         
  |        |  +-----------------------+                               
  |        |  |                       +-- Pleurobema_oviforme **         
  |        +--+                                                       
  |           |  +----------------------- Amblema_plicata **             
  |           |  |                                                    
  |           +--+  +-------------------- Popenaias_popeii **            
  |              |  |                                                 
  |              +--+  +----------------- Toxolasma_parvum **            
  |                 |  |                                              
  |                 |  |           +----- Potamilus_leptodon **          
  |                 +--+  +--------+                                  
  |                    |  |        |  +-- Potamilus_alatus **            
  |                    |  |        +--+                               
  |                    |  |           +-- Potamilus_streckersoni **      
  |                    +--+                                           
  |                       |  +----------- Leaunio_lienosus **            
  |                       |  |                                        
  |                       |  |     +----- Lampsilis_ornata **            
  |                       +--+  +--+                                  
  |                          |  |  |  +-- Lampsilis_cardium **           
  |                          |  |  +--+                               
  |                          +--+     +-- Lampsilis_cariosa **           
  |                             |                                     
  |                             |  +----- Venustaconcha_ellipsiformis ** 
  |                             +--+                                  
  |                                |  +-- Lampsilis_powellii **          
  |                                +--+                               
  |                                   +-- Lampsilis_siliquoidea **       
  |                                                                   
--+                                +----- Chamberlainia_somsakpanhai  
  |           +--------------------+                                  
  |           |                    |  +-- Sinohyriopsis_cumingii      
  |           |                    +--+                               
  |           |                       +-- Sinohyriopsis_schlegelii    
  |           |                                                       
  |           |                       +-- Hyriopsis_bialata           
  |           |                    +--+                               
  |  +--------+                    |  +-- Rectidens_sumatrensis       
  |  |        |  +-----------------+                                  
  |  |        |  |                 |  +-- Lens_contradens             
  |  |        |  |                 +--+                               
  |  |        |  |                    +-- Physunio_superbus           
  |  |        |  |                                                    
  |  |        |  |           +----------- Potomida_littoralis         
  |  |        +--+  +--------+                                        
  |  |           |  |        |  +-------- Pronodularia_japanensis     
  |  |           |  |        +--+                                     
  |  |           |  |           |  +----- Lamprotula_leaii            
  |  |           |  |           +--+                                  
  |  |           |  |              |  +-- Lamprotula_caveata          
  |  |           +--+              +--+                               
  |  |              |                 +-- Lamprotula_cornuumlunae     
  |  |              |                                                 
  |  |              |                 +-- Pseudodon_vondembuschianus  
  |  |              |  +--------------+                               
  |  |              |  |              +-- Pilsbryoconcha_exilis       
  |  |              +--+                                              
  |  |                 |  +-------------- Inversidens_rentianensis    
  |  |                 |  |                                           
  +--+                 +--+           +-- Parvasolenaia_rivularis     
     |                    |  +--------+                               
     |                    |  |        +-- Ptychorhynchus_pfisteri     
     |                    +--+                                        
     |                       |  +-------- Microcondylaea_bonellii     
     |                       +--+                                     
     |                          |  +----- Sinosolenaia_carinata       
     |                          +--+                                  
     |                             |  +-- Sinosolenaia_iridinea       
     |                             +--+                               
     |                                +-- Sinosolenaia_oleivora       
     |                                                                
     |  +-------------------------------- Lamellidens_marginalis      
     |  |                                                             
     |  |                             +-- Lepidodesma_aligera         
     |  |  +--------------------------+                               
     |  |  |                          +-- Lepidodesma_languilati      
     |  |  |                                                          
     |  |  |                    +-------- Aculamprotula_tortuosa      
     |  |  |                    |                                     
     |  |  |     +--------------+     +-- Aculamprotula_coreana       
     |  |  |     |              |  +--+                               
     +--+  |     |              |  |  +-- Aculamprotula_tientsinensis 
        |  |     |              +--+                                  
        |  |     |                 |  +-- Aculamprotula_polysticta    
        |  |     |                 +--+                               
        |  |     |                    +-- Aculamprotula_scripta       
        |  |     |                                                    
        |  |  +--+        +-------------- Unio_tumidus                
        |  |  |  |  +-----+                                           
        |  |  |  |  |     |  +----------- Unio_crassus                
        |  |  |  |  |     +--+                                        
        +--+  |  |  |        |  +-------- Unio_elongatulus            
           |  |  |  |        +--+                                     
           |  |  |  |           |  +----- Unio_pictorum               
           |  |  |  |           +--+                                  
           |  |  |  |              |  +-- Unio_delphinus              
           |  |  +--+              +--+                               
           |  |     |                 +-- Unio_mancus                 
           |  |     |                                                 
           |  |     |                 +-- Nodularia_breviconcha       
           |  |     |  +--------------+                               
           |  |     |  |              +-- Nodularia_douglasiae        
           |  |     |  |                                              
           |  |     |  |              +-- Pseudocuneopsis_capitata    
           |  |     |  +--------------+                               
           |  |     +--+              +-- Pseudocuneopsis_sichuanensis
           |  |        |                                              
           |  |        |              +-- Schistodesmus_lampreyanus   
           |  |        |  +-----------+                               
           +--+        |  |           +-- Schistodesmus_spinosus      
              |        +--+                                           
              |           |  +----------- Tchangsinaia_piscicula      
              |           +--+                                        
              |              |  +-------- Cuneopsis_rufescens         
              |              +--+                                     
              |                 |  +----- Cuneopsis_celtiformis       
              |                 +--+                                  
              |                    |  +-- Cuneopsis_demangei          
              |                    +--+                               
              |                       +-- Cuneopsis_heudei            
              |                                                       
              |                 +-------- Acuticosta_chinensis        
              |  +--------------+                                     
              |  |              |  +----- Lanceolaria_gladiola        
              |  |              +--+                                  
              |  |                 |  +-- Lanceolaria_grayii          
              |  |                 +--+                               
              |  |                    +-- Lanceolaria_lanceolata      
              |  |                                                    
              |  |        +-------------- Cristaria_plicata           
              |  |        |                                           
              +--+  +-----+           +-- Anemina_arcaeformis         
                 |  |     |  +--------+                               
                 |  |     |  |        +-- Anemina_euscaphys           
                 |  |     +--+                                        
                 |  |        |  +-------- Beringiana_fukuharai        
                 |  |        +--+                                     
                 |  |           |  +----- Sinanodonta_lucida          
                 |  |           +--+                                  
                 |  |              |  +-- Sinanodonta_tumens          
                 +--+              +--+                               
                    |                 +-- Sinanodonta_woodiana        
                    |                                                 
                    |           +-------- Pseudanodonta_complanata    
                    |  +--------+                                     
                    |  |        |  +----- Anodonta_anatina            
                    |  |        +--+                                  
                    |  |           |  +-- Anodonta_cygnea             
                    |  |           +--+                               
                    +--+              +-- Anodonta_exulcerata         
                       |                                              
                       |  +-------------- Anodonta_nuttalliana        
                       |  |                                           
                       |  |        +----- Pyganodon_grandis           
                       +--+  +-----+                                  
                          |  |     |  +-- Utterbackia_imbecillis      
                          |  |     +--+                               
                          +--+        +-- Utterbackia_peninsularis    
                             |                                        
                             |  +-------- Lasmigona_complanata        
                             +--+                                     
                                |  +----- Prolasmidonta_heterodon     
                                +--+                                  
                                   |  +-- Alasmidonta_varicosa        
                                   +--+                               
                                      +-- Platynaias_compressa        

Wu et al., 2024 [Publ 4350], fig. 8b, S2

iT tT bi riS oT roT roS iD
17 17 14 5 75 69 3 1.0
                                      +-- Quadrula_quadrula **           
           +--------------------------+                               
           |                          +-- Uniomerus_tetralasmus **       
           |                                                          
  +--------+                          +-- Elliptio_complanata **         
  |        |  +-----------------------+                               
  |        |  |                       +-- Pleurobema_oviforme **         
  |        +--+                                                       
  |           |  +----------------------- Amblema_plicata **             
  |           |  |                                                    
  |           +--+  +-------------------- Popenaias_popeii **            
  |              |  |                                                 
  |              +--+  +----------------- Toxolasma_parvum **            
  |                 |  |                                              
  |                 |  |           +----- Potamilus_leptodon **          
  |                 +--+  +--------+                                  
  |                    |  |        |  +-- Potamilus_alatus **            
  |                    |  |        +--+                               
  |                    +--+           +-- Potamilus_streckersoni **      
  |                       |                                           
  |                       |  +----------- Leaunio_lienosus **            
  |                       |  |                                        
  |                       +--+  +-------- Venustaconcha_ellipsiformis ** 
  |                          |  |                                     
  |                          |  |     +-- Lampsilis_powellii **          
  |                          +--+-----+                               
  |                             |     +-- Lampsilis_siliquoidea **       
  |                             |                                     
  |                             |  +----- Lampsilis_ornata **            
  |                             +--+                                  
  |                                |  +-- Lampsilis_cardium **           
  |                                +--+                               
  |                                   +-- Lampsilis_cariosa **           
  |                                                                   
--+                                +----- Chamberlainia_somsakpanhai  
  |           +--------------------+                                  
  |           |                    |  +-- Sinohyriopsis_cumingii      
  |           |                    +--+                               
  |           |                       +-- Sinohyriopsis_schlegelii    
  |           |                                                       
  |           |                       +-- Hyriopsis_bialata           
  |           |                    +--+                               
  |  +--------+                    |  +-- Rectidens_sumatrensis       
  |  |        |  +-----------------+                                  
  |  |        |  |                 |  +-- Lens_contradens             
  |  |        |  |                 +--+                               
  |  |        |  |                    +-- Physunio_superbus           
  |  |        |  |                                                    
  |  |        |  |           +----------- Potomida_littoralis         
  |  |        +--+  +--------+                                        
  |  |           |  |        |  +-------- Pronodularia_japanensis     
  |  |           |  |        +--+                                     
  |  |           |  |           |  +----- Lamprotula_leaii            
  |  |           |  |           +--+                                  
  |  |           |  |              |  +-- Lamprotula_caveata          
  |  |           +--+              +--+                               
  |  |              |                 +-- Lamprotula_cornuumlunae     
  |  |              |                                                 
  |  |              |                 +-- Pseudodon_vondembuschianus  
  |  |              |  +--------------+                               
  |  |              |  |              +-- Pilsbryoconcha_exilis       
  |  |              +--+                                              
  |  |                 |  +-------------- Inversidens_rentianensis    
  |  |                 |  |                                           
  +--+                 +--+           +-- Parvasolenaia_rivularis     
     |                    |  +--------+                               
     |                    |  |        +-- Ptychorhynchus_pfisteri     
     |                    +--+                                        
     |                       |  +-------- Microcondylaea_bonellii     
     |                       +--+                                     
     |                          |  +----- Sinosolenaia_carinata       
     |                          +--+                                  
     |                             |  +-- Sinosolenaia_iridinea       
     |                             +--+                               
     |                                +-- Sinosolenaia_oleivora       
     |                                                                
     |  +-------------------------------- Lamellidens_marginalis      
     |  |                                                             
     |  |                             +-- Lepidodesma_aligera         
     |  |  +--------------------------+                               
     |  |  |                          +-- Lepidodesma_languilati      
     |  |  |                                                          
     |  |  |                    +-------- Aculamprotula_tortuosa      
     |  |  |                    |                                     
     |  |  |     +--------------+     +-- Aculamprotula_coreana       
     |  |  |     |              |  +--+                               
     +--+  |     |              |  |  +-- Aculamprotula_tientsinensis 
        |  |     |              +--+                                  
        |  |     |                 |  +-- Aculamprotula_polysticta    
        |  |     |                 +--+                               
        |  |     |                    +-- Aculamprotula_scripta       
        |  |     |                                                    
        |  |  +--+        +-------------- Unio_tumidus                
        |  |  |  |  +-----+                                           
        |  |  |  |  |     |  +----------- Unio_crassus                
        |  |  |  |  |     +--+                                        
        +--+  |  |  |        |  +-------- Unio_elongatulus            
           |  |  |  |        +--+                                     
           |  |  |  |           |  +----- Unio_pictorum               
           |  |  |  |           +--+                                  
           |  |  |  |              |  +-- Unio_delphinus              
           |  |  +--+              +--+                               
           |  |     |                 +-- Unio_mancus                 
           |  |     |                                                 
           |  |     |                 +-- Nodularia_breviconcha       
           |  |     |  +--------------+                               
           |  |     |  |              +-- Nodularia_douglasiae        
           |  |     |  |                                              
           |  |     |  |              +-- Pseudocuneopsis_capitata    
           |  |     |  +--------------+                               
           |  |     +--+              +-- Pseudocuneopsis_sichuanensis
           |  |        |                                              
           |  |        |              +-- Schistodesmus_lampreyanus   
           +--+        |  +-----------+                               
              |        |  |           +-- Schistodesmus_spinosus      
              |        +--+                                           
              |           |  +----------- Tchangsinaia_piscicula      
              |           +--+                                        
              |              |  +-------- Cuneopsis_rufescens         
              |              +--+                                     
              |                 |  +----- Cuneopsis_celtiformis       
              |                 +--+                                  
              |                    |  +-- Cuneopsis_demangei          
              |                    +--+                               
              |                       +-- Cuneopsis_heudei            
              |                                                       
              |                 +-------- Acuticosta_chinensis        
              |  +--------------+                                     
              |  |              |  +----- Lanceolaria_gladiola        
              |  |              +--+                                  
              |  |                 |  +-- Lanceolaria_grayii          
              |  |                 +--+                               
              |  |                    +-- Lanceolaria_lanceolata      
              |  |                                                    
              |  |        +-------------- Cristaria_plicata           
              +--+        |                                           
                 |  +-----+           +-- Anemina_arcaeformis         
                 |  |     |  +--------+                               
                 |  |     |  |        +-- Anemina_euscaphys           
                 |  |     +--+                                        
                 |  |        |  +-------- Beringiana_fukuharai        
                 |  |        +--+                                     
                 |  |           |  +----- Sinanodonta_lucida          
                 |  |           +--+                                  
                 +--+              |  +-- Sinanodonta_tumens          
                    |              +--+                               
                    |                 +-- Sinanodonta_woodiana        
                    |                                                 
                    |  +----------------- Anodonta_nuttalliana        
                    |  |                                              
                    |  |        +-------- Pseudanodonta_complanata    
                    |  |  +-----+                                     
                    +--+  |     |  +----- Anodonta_anatina            
                       |  |     +--+                                  
                       |  |        |  +-- Anodonta_cygnea             
                       |  |        +--+                               
                       +--+           +-- Anodonta_exulcerata         
                          |                                           
                          |        +----- Pyganodon_grandis           
                          |  +-----+                                  
                          |  |     |  +-- Utterbackia_imbecillis      
                          |  |     +--+                               
                          +--+        +-- Utterbackia_peninsularis    
                             |                                        
                             |  +-------- Lasmigona_complanata        
                             +--+                                     
                                |  +----- Prolasmidonta_heterodon     
                                +--+                                  
                                   |  +-- Alasmidonta_varicosa        
                                   +--+                               
                                      +-- Platynaias_compressa        

Roe et al., 2001 [Publ 2689], fig. 3a

iT tT bi riS oT roT roS iD
16 16 8 1 0 0 0 25.0
  +-------------- Ligumia_recta **       
  |                                   
  +-------------- Obliquaria_reflexa **  
  |                                   
  |        +----- Lampsilis_teres **     
  +--------+                          
--+        |  +-- Lampsilis_ovata **     
  |        +--+                       
  |           +-- Lampsilis_ornata **    
  |                                   
  |  +----------- Villosa_villosa **     
  |  |                                
  |  |     +----- Hamiota_australis[2] **
  +--+  +--+                          
     |  |  |  +-- Hamiota_subangulata ** 
     |  |  +--+                       
     |  |     +-- Hamiota_australis[1] **
     +--+                             
        |  +----- Hamiota_altilis[3] **  
        |  |                          
        |  |  +-- Hamiota_perovalis[1] **
        +--+  |                       
           |  +-- Hamiota_perovalis[2] **
           |  |                       
           |  +-- Hamiota_altilis[1] **  
           +--+                       
              +-- Hamiota_altilis[2] **  
              |                       
              +-- Hamiota_altilis[4] **  
              |                       
              +-- Hamiota_altilis[5] **  

Szumowski et al., 2012 [Publ 3569], fig. 4

iT tT bi riS oT roT roS iD
16 16 12 2 0 0 0 23.0
  +-------------------------- Amblema_plicata **        
  |                                                  
--+  +----------------------- Ellipsaria_lineolata **   
  |  |                                               
  +--+  +-------------------- Truncilla_truncata **     
     |  |                                            
     +--+  +----------------- Obliquaria_reflexa **     
        |  |                                         
        |  |  +-------------- Potamilus_alatus **       
        +--+  |                                      
           |  |        +----- Lampsilis_ornata **       
           |  |  +-----+                             
           |  |  |     |  +-- Lampsilis_cardium **      
           +--+  |     +--+                          
              |  |        +-- Lampsilis_ovata **        
              |  |                                   
              |  |  +-------- Lampsilis_teres **        
              |  +--+                                
              +--+  |  +----- Lampsilis_siliquoidea **  
                 |  +--+                             
                 |     |  +-- Ortmanniana_ligamentina **
                 |     +--+                          
                 |        +-- Lampsilis_higginsii **    
                 |                                   
                 |     +----- Hamiota_altilis **        
                 |     |                             
                 +-----+  +-- Epioblasma_triquetra **   
                       |  |                          
                       +--+-- Cambarunio_iris **        
                          |                          
                          +-- Ligumia_recta **          

Johnson et al., 2018 [Publ 3844], fig. 2

iT tT bi riS oT roT roS iD
16 16 14 3 0 0 0 28.0
  +-------------------------------- Amblema_plicata **       
  |                                                       
--+  +----------------------------- Elliptio_crassidens **   
  |  |                                                    
  +--+  +-------------------------- Megalonaias_nervosa **   
     |  |                                                 
     +--+  +----------------------- Uniomerus_tetralasmus ** 
        |  |                                              
        +--+  +-------------------- Theliderma_metanevra **  
           |  |                                           
           |  |                 +-- Quadrula_quadrula **     
           +--+  +--------------+                         
              |  |              +-- Quadrula_verrucosa **    
              |  |                                        
              |  |           +----- Pustulosa_kieneriana **  
              +--+  +--------+                            
                 |  |        |  +-- Pustulosa_infucata **    
                 |  |        +--+                         
                 |  |           +-- Pustulosa_kleiniana **   
                 +--+                                     
                    |           +-- Pustulosa_pustulosa **   
                    |  +--------+                         
                    |  |        +-- Pustulosa_succissa **    
                    +--+                                  
                       |  +-------- Cyclonaias_tuberculata **
                       +--+                               
                          |  +----- Pustulosa_necki **       
                          +--+                            
                             |  +-- Pustulosa_nodulata **    
                             +--+                         
                                +-- Pustulosa_petrina **     

Inoue et al., 2020 [Publ 3941], fig. 5

iT tT bi riS oT roT roS iD
16 16 14 1 0 0 0 25.0
  +-------------------- Truncilla_macrodon **   
  |                                          
  |     +-------------- Lampsilis_bracteata **  
  |     |                                    
  |  +--+           +-- Lampsilis_ornata **     
--+  |  |  +--------+                        
  |  |  |  |        +-- Lampsilis_fasciola **   
  |  |  +--+                                 
  |  |     |  +-------- Lampsilis_ovata **      
  |  |     +--+                              
  |  |        |  +----- Lampsilis_cariosa **    
  +--+        +--+                           
     |           |  +-- Lampsilis_cardium **    
     |           +--+                        
     |              +-- Lampsilis_satura **     
     |                                       
     |  +-------------- Lampsilis_higginsii **  
     |  |                                    
     +--+  +----------- Lampsilis_teres **      
        |  |                                 
        |  |        +-- Lampsilis_radiata **    
        +--+  +-----+                        
           |  |     +-- Lampsilis_siliquoidea **
           |  |                              
           +--+     +-- Lampsilis_hydiana **    
              |  +--+                        
              |  |  +-- Lampsilis_bergmanni **  
              +--+                           
                 |  +-- Lampsilis_straminea **  
                 +--+                        
                    +-- Lampsilis_virescens **  

Porto-Hannes et al., 2021 [Publ 4227], fig. 3

iT tT bi riS oT roT roS iD
16 16 12 4 0 0 0 23.0
  +-------------------- Quadrula_quadrula **      
  |                                            
  +-------------------- Quadrula_verrucosa **     
  |                                            
--+                 +-- Eurynia_dilatata **       
  |  +--------------+                          
  |  |              +-- Pleurobema_sintoxia **    
  |  |                                         
  |  |              +-- Amblema_plicata **        
  +--+  +-----------+                          
     |  |           +-- Amblema_elliottii **      
     |  |                                      
     |  |           +-- Obovaria_olivaria **      
     +--+  +--------+                          
        |  |        +-- Ortmanniana_ligamentina **
        |  |                                   
        |  |        +-- Obliquaria_reflexa **     
        |  +--------+                          
        +--+        +-- Lampsilis_ornata **       
           |                                   
           |     +----- Lampsilis_radiata[2] **   
           |  +--+                             
           |  |  |  +-- Lampsilis_siliquoidea **  
           |  |  +--+                          
           +--+     +-- Lampsilis_radiata[1] **   
              |                                
              |  +----- Lampsilis_virescens **    
              +--+                             
                 |  +-- Lampsilis_hydiana **      
                 +--+                          
                    +-- Lampsilis_straminea **    

Lydeard et al., 2000 [Publ 2654], fig. 2a

iT tT bi riS oT roT roS iD
15 17 11 4 2 1 1 31.0
  +-------------------- Anodonta_cygnea       
  |                                           
  +-------------------- Cumberlandia_monodonta
  |                                           
  |              +----- Lampsilis_teres **       
  +--------------+                            
  |              |  +-- Obovaria_unicolor **     
  |              +--+                         
  |                 +-- Obovaria_olivaria **     
  |                                           
--+                 +-- Reginaia_rotulata[1] **  
  |                 |                         
  +-----------------+-- Reginaia_rotulata[2] **  
  |                 |                         
  |                 +-- Reginaia_ebenus **       
  |                                           
  |              +----- Quadrula_nobilis **      
  |  +-----------+                            
  |  |           |  +-- Pustulosa_infucata **    
  |  |           +--+                         
  +--+              +-- Pustulosa_succissa **    
     |                                        
     |  +-------------- Pleurobema_decisum **    
     +--+                                     
        |  +----------- Fusconaia_flava **       
        +--+                                  
           |  +-------- Fusconaia_burkei[1] **   
           +--+                               
              |  +----- Fusconaia_burkei[2] **   
              +--+                            
                 |  +-- Fusconaia_burkei[3] **   
                 +--+                         
                    +-- Fusconaia_escambia **    

Lydeard et al., 2000 [Publ 2654], fig. 2b

iT tT bi riS oT roT roS iD
15 15 11 4 2 1 1 30.0
                    +----- Lampsilis_teres **     
  +-----------------+                          
  |                 |  +-- Obovaria_unicolor **   
  |                 +--+                       
  |                    +-- Obovaria_olivaria **   
  |                                            
--+                    +-- Reginaia_rotulata[1] **
  |                    |                       
  |  +-----------------+-- Reginaia_rotulata[2] **
  |  |                 |                       
  |  |                 +-- Reginaia_ebenus **     
  |  |                                         
  +--+                 +-- Quadrula_nobilis **    
     |                 |                       
     |  +--------------+-- Pustulosa_infucata **  
     |  |              |                       
     +--+              +-- Pustulosa_succissa **  
        |                                      
        |  +-------------- Pleurobema_decisum **  
        +--+                                   
           |  +----------- Fusconaia_flava **     
           +--+                                
              |  +-------- Fusconaia_burkei[1] ** 
              +--+                             
                 |  +----- Fusconaia_burkei[2] ** 
                 +--+                          
                    |  +-- Fusconaia_burkei[3] ** 
                    +--+                       
                       +-- Fusconaia_escambia **  

Inoue et al., 2013 [Publ 3528], fig. 3

iT tT bi riS oT roT roS iD
15 15 12 4 2 2 2 26.0
  +-------------------------- Gonidea_angulata      
  |                                                 
  +-------------------------- Pyganodon_grandis     
  |                                                 
--+                       +-- Eurynia_dilatata **      
  |  +--------------------+                         
  |  |                    +-- Quadrula_quadrula **     
  |  |                                              
  |  |              +-------- Paetulunio_fabalis **    
  +--+  +-----------+                               
     |  |           |  +----- Amblema_plicata **       
     |  |           +--+                            
     |  |              |  +-- Reginaia_ebenus **       
     +--+              +--+                         
        |                 +-- Reginaia_rotulata **     
        |                                           
        |  +----------------- Lampsilis_ornata **      
        |  |                                        
        +--+  +-------------- Leaunio_vanuxemensis **  
           |  |                                     
           |  |           +-- Villosa_villosa **       
           +--+-----------+                         
              |           +-- Cambarunio_iris **       
              |                                     
              |  +----------- Obovaria_olivaria **     
              +--+                                  
                 |  +-------- Obovaria_arkansasensis **
                 +--+                               
                    |  +----- Obovaria_retusa **       
                    +--+                            
                       |  +-- Obovaria_unicolor **     
                       +--+                         
                          +-- Obovaria_subrotunda **   

Szumowski et al., 2012 [Publ 3569], fig. 5

iT tT bi riS oT roT roS iD
15 15 12 1 0 0 0 30.0
  +----------------- Megalonaias_nervosa **  
  |                                       
  |        +-------- Theliderma_cylindrica **
  +--------+                              
  |        |  +----- Theliderma_intermedia **
  |        +--+                           
  |           |  +-- Theliderma_metanevra ** 
--+           +--+                        
  |              +-- Theliderma_sparsa **    
  |                                       
  |        +-------- Quadrula_quadrula[1] ** 
  |  +-----+                              
  |  |     |  +----- Quadrula_verrucosa **   
  |  |     +--+                           
  |  |        |  +-- Quadrula_nobilis **     
  +--+        +--+                        
     |           +-- Quadrula_quadrula[2] ** 
     |                                    
     |  +----------- Pustulosa_kieneriana ** 
     |  |                                 
     +--+  +-------- Pustulosa_infucata **   
        |  |                              
        +--+     +-- Pustulosa_pustulosa **  
           |  +--+                        
           |  |  +-- Pustulosa_succissa **   
           +--+                           
              |  +-- Pustulosa_nodulata **   
              +--+                        
                 +-- Pustulosa_petrina **    

Pfeiffer et al., 2019 [Publ 3937], fig. 4-5

iT tT bi riS oT roT roS iD
15 15 13 5 42 35 3 17.0
                             +-- Quadrula_quadrula **            
        +--------------------+                                
        |                    +-- Uniomerus_tetralasmus **        
        |                                                     
        |                 +----- Elliptoideus_sloatianus **      
  +-----+  +--------------+                                   
  |     |  |              |  +-- Elliptio_complanata **          
  |     |  |              +--+                                
  |     |  |                 +-- Pleurobema_strodeanum **        
  |     |  |                                                  
  |     +--+              +----- Reginaia_ebenus **              
  |        |  +-----------+                                   
  |        |  |           |  +-- Plectomerus_dombeyanus **       
  |        |  |           +--+                                
  |        |  |              +-- Amblema_plicata **              
  |        +--+                                               
  |           |              +-- Psoronaias_semigranosa **       
  |           |  +-----------+                                
  |           |  |           +-- Popenaias_popeii **             
  |           +--+                                            
  |              |  +----------- Obliquaria_reflexa **           
  |              +--+                                         
  |                 |  +-------- Cyrtonaias_tampicoensis **      
  |                 +--+                                      
  |                    |  +----- Leaunio_lienosus **             
  |                    +--+                                   
  |                       |  +-- Lampsilis_cardium **            
  |                       +--+                                
  |                          +-- Pachynaias_spheniopsis **       
  |                                                           
--+                          +-- Chamberlainia_hainesiana     
  |     +--------------------+                                
  |     |                    +-- Sinohyriopsis_cumingii       
  |     |                                                     
  |     |     +----------------- Lamprotula_cornuumlunae      
  |     |  +--+                                               
  |     |  |  |  +-------------- Gonidea_angulata             
  |  +--+  |  +--+                                            
  |  |  |  |     |  +----------- Indopseudodon_crebristriatus 
  |  |  |  |     +--+                                         
  |  |  |  |        |  +-------- Bineurus_anodontinum         
  |  |  |  |        +--+                                      
  |  |  |  |           |  +----- Pilsbryoconcha_exilis        
  |  |  |  |           +--+                                   
  |  |  +--+              |  +-- Pseudodon_vondembuschianus   
  |  |     |              +--+                                
  |  |     |                 +-- Pseudodon_inoscularis        
  |  |     |                                                  
  |  |     |              +----- Ensidens_ingallsianus        
  |  |     |        +-----+                                   
  |  |     |        |     |  +-- Hyriopsis_bialata            
  |  |     |        |     +--+                                
  |  |     |        |        +-- Rectidens_sumatrensis        
  |  |     +--------+                                         
  |  |              |        +-- Trapezoideus_foliaceus       
  |  |              |  +-----+                                
  +--+              |  |     +-- Physunio_superbus            
     |              +--+                                      
     |                 |  +----- Yaukthwa_paiensis            
     |                 +--+                                   
     |                    |  +-- Lens_contradens              
     |                    +--+                                
     |                       +-- Solenaia_emarginata          
     |                                                        
     |              +----------- Lanceolaria_lanceolata       
     |              |                                         
     |              |     +----- Unio_gibbus                  
     |     +--------+  +--+                                   
     |     |        |  |  |  +-- Nodularia_douglasiae         
     |     |        |  |  +--+                                
     |     |        |  |     +-- Tchangsinaia_piscicula       
     |     |        +--+                                      
     |     |           |     +-- Cristaria_plicata            
     |     |           |  +--+                                
     |     |           |  |  +-- Sinanodonta_woodiana         
     +-----+           +--+                                   
           |              |  +-- Anodonta_cygnea              
           |              +--+                                
           |                 +-- Pyganodon_grandis            
           |                                                  
           |                 +-- Trapezidens_dolichorhynchus  
           |  +--------------+                                
           |  |              +-- Lamellidens_savadiensis      
           |  |                                               
           +--+              +-- Leoparreysia_tavoyensis      
              |  +-----------+                                
              |  |           +-- Leoparreysia_olivacea        
              |  |                                            
              +--+           +-- Coelatura_choziensis         
                 |  +--------+                                
                 |  |        +-- Prisodontopsis_aviculaeformis
                 +--+                                         
                    |  +-------- Indonaia_caerulea            
                    +--+                                      
                       |  +----- Indochinella_pugio           
                       +--+                                   
                          |  +-- Scabies_phaselus             
                          +--+                                
                             +-- Harmandia_somboriensis       

Pfeiffer et al., 2019 [Publ 3937], fig. 6

iT tT bi riS oT roT roS iD
15 15 13 5 42 35 3 19.0
                       +-- Quadrula_quadrula **            
     +-----------------+                                
     |                 +-- Uniomerus_tetralasmus **        
     |                                                  
     |              +----- Elliptoideus_sloatianus **      
     |           +--+                                   
     |           |  |  +-- Elliptio_complanata **          
  +--+           |  +--+                                
  |  |  +--------+     +-- Pleurobema_strodeanum **        
  |  |  |        |                                      
  |  |  |        |  +----- Reginaia_ebenus **              
  |  |  |        +--+                                   
  |  |  |           |  +-- Plectomerus_dombeyanus **       
  |  +--+           +--+                                
  |     |              +-- Amblema_plicata **              
  |     |                                               
  |     |              +-- Psoronaias_semigranosa **       
  |     |  +-----------+                                
  |     |  |           +-- Popenaias_popeii **             
  |     +--+                                            
  |        |  +----------- Obliquaria_reflexa **           
  |        +--+                                         
  |           |  +-------- Cyrtonaias_tampicoensis **      
  |           +--+                                      
  |              |  +----- Leaunio_lienosus **             
  |              +--+                                   
  |                 |  +-- Lampsilis_cardium **            
  |                 +--+                                
  |                    +-- Pachynaias_spheniopsis **       
  |                                                     
  |                    +-- Chamberlainia_hainesiana     
  |  +-----------------+                                
  |  |                 +-- Sinohyriopsis_cumingii       
  |  |                                                  
  |  |                 +-- Gonidea_angulata             
  |  |     +-----------+                                
  |  |     |           +-- Lamprotula_cornuumlunae      
  |  |  +--+                                            
  +--+  |  |  +----------- Indopseudodon_crebristriatus 
  |  |  |  +--+                                         
  |  |  |     |  +-------- Pilsbryoconcha_exilis        
  |  |  |     +--+                                      
--+  |  |        |  +----- Bineurus_anodontinum         
  |  |  |        +--+                                   
  |  |  |           |  +-- Pseudodon_vondembuschianus   
  |  +--+           +--+                                
  |     |              +-- Pseudodon_inoscularis        
  |     |                                               
  |     |           +----- Ensidens_ingallsianus        
  |     |     +-----+                                   
  |     |     |     |  +-- Hyriopsis_bialata            
  |     |     |     +--+                                
  |     |     |        +-- Rectidens_sumatrensis        
  |     +-----+                                         
  |           |        +-- Trapezoideus_foliaceus       
  |           |  +-----+                                
  |           |  |     +-- Physunio_superbus            
  |           +--+                                      
  |              |  +----- Yaukthwa_paiensis            
  |              +--+                                   
  |                 |  +-- Lens_contradens              
  |                 +--+                                
  |                    +-- Solenaia_emarginata          
  |                                                     
  |              +-------- Lanceolaria_lanceolata       
  |           +--+                                      
  |           |  |  +----- Unio_gibbus                  
  |           |  +--+                                   
  |           |     |  +-- Tchangsinaia_piscicula       
  |           |     +--+                                
  |  +--------+        +-- Nodularia_douglasiae         
  |  |        |                                         
  |  |        |        +-- Cristaria_plicata            
  |  |        |     +--+                                
  |  |        |     |  +-- Sinanodonta_woodiana         
  |  |        +-----+                                   
  |  |              |  +-- Anodonta_cygnea              
  +--+              +--+                                
     |                 +-- Pyganodon_grandis            
     |                                                  
     |                 +-- Trapezidens_dolichorhynchus  
     |  +--------------+                                
     |  |              +-- Lamellidens_savadiensis      
     |  |                                               
     +--+              +-- Leoparreysia_tavoyensis      
        |  +-----------+                                
        |  |           +-- Leoparreysia_olivacea        
        |  |                                            
        +--+           +-- Coelatura_choziensis         
           |  +--------+                                
           |  |        +-- Prisodontopsis_aviculaeformis
           +--+                                         
              |  +-------- Indonaia_caerulea            
              +--+                                      
                 |  +----- Indochinella_pugio           
                 +--+                                   
                    |  +-- Scabies_phaselus             
                    +--+                                
                       +-- Harmandia_somboriensis       

Wu et al., 2022 [Publ 4238], fig. 3

iT tT bi riS oT roT roS iD
15 15 13 5 59 50 2 2.0
                                      +-- Uniomerus_tetralasmus **       
           +--------------------------+                               
           |                          +-- Quadrula_quadrula **           
           |                                                          
  +--------+                          +-- Pleurobema_oviforme **         
  |        |  +-----------------------+                               
  |        |  |                       +-- Elliptio_complanata **         
  |        +--+                                                       
  |           |  +----------------------- Amblema_plicata **             
  |           |  |                                                    
  |           +--+  +-------------------- Popenaias_popeii **            
  |              |  |                                                 
  |              +--+  +----------------- Toxolasma_parvum **            
  |                 |  |                                              
  |                 +--+              +-- Potamilus_alatus **            
  |                    |  +-----------+                               
  |                    |  |           +-- Potamilus_leptodon **          
  |                    +--+                                           
  |                       |  +----------- Leaunio_lienosus **            
  |                       |  |                                        
  |                       +--+        +-- Lampsilis_cardium **           
  |                          |  +-----+                               
  |                          |  |     +-- Lampsilis_ornata **            
  |                          +--+                                     
  |                             |  +----- Venustaconcha_ellipsiformis ** 
  |                             +--+                                  
  |                                |  +-- Lampsilis_siliquoidea **       
  |                                +--+                               
--+                                   +-- Lampsilis_powellii **          
  |                                                                   
  |                                +----- Chamberlainia_somsakpanhai  
  |                    +-----------+                                  
  |                    |           |  +-- Sinohyriopsis_schlegelii    
  |                    |           +--+                               
  |                    |              +-- Sinohyriopsis_cumingii      
  |                    |                                              
  |  +-----------------+        +-------- Potomida_littoralis         
  |  |                 |  +-----+                                     
  |  |                 |  |     |  +----- Pronodularia_japanensis     
  |  |                 |  |     +--+                                  
  |  |                 |  |        |  +-- Lamprotula_caveata          
  |  |                 |  |        +--+                               
  |  |                 +--+           +-- Lamprotula_leaii            
  |  |                    |                                           
  |  |                    |           +-- Pilsbryoconcha_exilis       
  |  |                    |  +--------+                               
  |  |                    |  |        +-- Pseudodon_vondembuschianus  
  |  |                    |  |                                        
  |  |                    +--+        +-- Ptychorhynchus_pfisteri     
  |  |                       |  +-----+                               
  |  |                       |  |     +-- Parvasolenaia_rivularis     
  |  |                       +--+                                     
  |  |                          |  +----- Microcondylaea_bonellii     
  +--+                          +--+                                  
     |                             |  +-- Sinosolenaia_carinata       
     |                             +--+                               
     |                                +-- Sinosolenaia_recognita      
     |                                                                
     |  +-------------------------------- Lepidodesma_languilati      
     |  |                                                             
     |  |                       +-------- Acuticosta_chinensis        
     |  |              +--------+                                     
     |  |              |        |  +----- Lanceolaria_gladiola        
     |  |              |        +--+                                  
     |  |              |           |  +-- Lanceolaria_grayii          
     |  |              |           +--+                               
     |  |              |              +-- Lanceolaria_lanceolata      
     |  |              |                                              
     |  |  +-----------+        +-------- Cristaria_plicata           
     |  |  |           |        |                                     
     +--+  |           |  +-----+     +-- Sinanodonta_woodiana        
        |  |           |  |     |  +--+                               
        |  |           |  |     |  |  +-- Sinanodonta_lucida          
        |  |           |  |     +--+                                  
        |  |           |  |        |  +-- Anemina_arcaeformis         
        |  |           +--+        +--+                               
        |  |              |           +-- Anemina_euscaphys           
        |  |              |                                           
        |  |              |           +-- Anodonta_cygnea             
        |  |              |  +--------+                               
        |  |              |  |        +-- Anodonta_anatina            
        |  |              +--+                                        
        |  |                 |  +-------- Platynaias_compressa        
        +--+                 +--+                                     
           |                    |  +----- Pyganodon_grandis           
           |                    +--+                                  
           |                       |  +-- Utterbackia_imbecillis      
           |                       +--+                               
           |                          +-- Utterbackia_peninsularis    
           |                                                          
           |                    +-------- Aculamprotula_tortuosa      
           |                    |                                     
           |  +-----------------+     +-- Aculamprotula_tientsinensis 
           |  |                 |  +--+                               
           |  |                 |  |  +-- Aculamprotula_coreana       
           |  |                 +--+                                  
           |  |                    |  +-- Aculamprotula_polysticta    
           |  |                    +--+                               
           |  |                       +-- Aculamprotula_scripta       
           +--+                                                       
              |                 +-------- Unio_tumidus                
              |  +--------------+                                     
              |  |              |  +----- Unio_crassus                
              |  |              +--+                                  
              |  |                 |  +-- Unio_pictorum               
              |  |                 +--+                               
              +--+                    +-- Unio_delphinus              
                 |                                                    
                 |                    +-- Pseudocuneopsis_capitata    
                 |  +-----------------+                               
                 |  |                 +-- Pseudocuneopsis_sichuanensis
                 |  |                                                 
                 +--+                 +-- Nodularia_douglasiae        
                    |  +--------------+                               
                    |  |              +-- Nodularia_breviconcha       
                    |  |                                              
                    +--+              +-- Schistodesmus_lampreyanus   
                       |  +-----------+                               
                       |  |           +-- Schistodesmus_spinosus      
                       +--+                                           
                          |  +----------- Tchangsinaia_piscicula      
                          +--+                                        
                             |  +-------- Cuneopsis_rufescens         
                             +--+                                     
                                |  +----- Cuneopsis_celtiformis       
                                +--+                                  
                                   |  +-- Cuneopsis_heudei            
                                   +--+                               
                                      +-- Cuneopsis_demangei          

Mejia-Trujillo & Smith, 2022 [Publ 4274], fig. 2-F

iT tT bi riS oT roT roS iD
15 15 13 5 0 0 0 10.0
     +-------------- Toxolasma_parvum **           
     |                                          
  +--+  +----------- Popenaias_popeii **           
  |  |  |                                       
  |  +--+  +-------- Amblema_plicata **            
  |     |  |                                    
  |     +--+     +-- Quadrula_quadrula **          
  |        |  +--+                              
  |        |  |  +-- Uniomerus_tetralasmus **      
  |        +--+                                 
--+           |  +-- Pleurobema_oviforme **        
  |           +--+                              
  |              +-- Elliptio_complanata **        
  |                                             
  |           +----- Potamilus_leptodon **         
  |     +-----+                                 
  |     |     |  +-- Potamilus_streckersoni **     
  |     |     +--+                              
  |     |        +-- Potamilus_alatus **           
  +-----+                                       
        |        +-- Lampsilis_cardium **          
        |  +-----+                              
        |  |     +-- Lampsilis_ornata **           
        +--+                                    
           |  +----- Venustaconcha_ellipsiformis **
           +--+                                 
              |  +-- Lampsilis_siliquoidea **      
              +--+                              
                 +-- Lampsilis_powellii **         

Dai et al., 2023 [Publ 4285], fig. 5

iT tT bi riS oT roT roS iD
15 15 13 5 38 36 2 2.0
                                +-- Quadrula_quadrula **          
     +--------------------------+                              
     |                          +-- Uniomerus_tetralasmus **      
     |                                                         
  +--+                          +-- Elliptio_complanata **        
  |  |  +-----------------------+                              
  |  |  |                       +-- Pleurobema_oviforme **        
  |  +--+                                                      
  |     |  +----------------------- Amblema_plicata **            
  |     |  |                                                   
  |     +--+  +-------------------- Popenaias_popeii **           
  |        |  |                                                
  |        +--+  +----------------- Toxolasma_parvum **           
  |           |  |                                             
  |           +--+              +-- Potamilus_leptodon **         
  |              |  +-----------+                              
  |              |  |           +-- Potamilus_alatus **           
  |              +--+                                          
  |                 |  +----------- Leaunio_lienosus **           
  |                 |  |                                       
  |                 +--+        +-- Lampsilis_cardium **          
  |                    |  +-----+                              
  |                    |  |     +-- Lampsilis_ornata **           
  |                    +--+                                    
  |                       |  +----- Venustaconcha_ellipsiformis **
  |                       +--+                                 
  |                          |  +-- Lampsilis_powellii **         
  |                          +--+                              
--+                             +-- Lampsilis_siliquoidea **      
  |                                                            
  |           +-------------------- Lepidodesma_languilati     
  |           |                                                
  |           |     +-------------- Aculamprotula_tientsinensis
  |           |     |                                          
  |           |  +--+  +----------- Unio_crassus               
  |           |  |  |  |                                       
  |        +--+  |  +--+        +-- Pseudocuneopsis_capitata   
  |        |  |  |     |  +-----+                              
  |        |  |  |     |  |     +-- Nodularia_douglasiae       
  |        |  |  |     +--+                                    
  |        |  |  |        |  +----- Schistodesmus_lampreyanus  
  |        |  |  |        +--+                                 
  |        |  +--+           |  +-- Cuneopsis_celtiformis      
  |        |     |           +--+                              
  |        |     |              +-- Tchangsinaia_piscicula     
  |        |     |                                             
  |        |     |              +-- Acuticosta_chinensis       
  |        |     |  +-----------+                              
  |        |     |  |           +-- Lanceolaria_grayii         
  |        |     |  |                                          
  |        |     +--+        +----- Cristaria_plicata          
  |        |        |  +-----+                                 
  |        |        |  |     |  +-- Anemina_arcaeformis        
  +--------+        |  |     +--+                              
           |        +--+        +-- Sinanodonta_woodiana       
           |           |                                       
           |           |  +-------- Anodonta_cygnea            
           |           +--+                                    
           |              |  +----- Platynaias_compressa       
           |              +--+                                 
           |                 |  +-- Pyganodon_grandis          
           |                 +--+                              
           |                    +-- Utterbackia_imbecillis     
           |                                                   
           |                 +----- Chamberlainia_somsakpanhai 
           |  +--------------+                                 
           |  |              |  +-- Sinohyriopsis_cumingii     
           |  |              +--+                              
           |  |                 +-- Sinohyriopsis_schlegelii   
           |  |                                                
           |  |                 +-- Hyriopsis_bialata          
           +--+              +--+                              
              |              |  +-- Rectidens_sumatrensis      
              |  +-----------+                                 
              |  |           |  +-- Lens_contradens            
              |  |           +--+                              
              |  |              +-- Physunio_superbus          
              |  |                                             
              +--+        +-------- Potomida_littoralis        
                 |  +-----+                                    
                 |  |     |  +----- Pronodularia_japanensis    
                 |  |     +--+                                 
                 |  |        |  +-- Lamprotula_caveata         
                 |  |        +--+                              
                 +--+           +-- Lamprotula_leaii           
                    |                                          
                    |           +-- Pseudodon_vondembuschianus 
                    |  +--------+                              
                    |  |        +-- Pilsbryoconcha_exilis      
                    |  |                                       
                    +--+     +----- Parvasolenaia_rivularis    
                       |  +--+                                 
                       |  |  |  +-- Postolata_guangxiensis     
                       |  |  +--+                              
                       +--+     +-- Ptychorhynchus_pfisteri    
                          |                                    
                          |  +----- Microcondylaea_bonellii    
                          +--+                                 
                             |  +-- Sinosolenaia_oleivora      
                             +--+                              
                                +-- Sinosolenaia_carinata      

Graf & Cummings, 2006 [Publ 1968], fig. 2

iT tT bi riS oT roT roS iD
14 14 10 4 33 11 3 26.0
  +----------------------------- Coelatura_aegyptiaca      
  |                                                        
  +----------------------------- Lens_contradens           
  |                                                        
  |                          +-- Unio_pictorum             
  |                 +--------+                             
  |                 |        +-- Unio_caffer               
  +-----------------+                                      
  |                 |  +-------- Lasmigona_costata         
--+                 +--+                                   
  |                    |  +----- Alasmidonta_marginata     
  |                    +--+                                
  |                       |  +-- Pyganodon_grandis         
  |                       +--+                             
  |                          +-- Strophitus_undulatus      
  |                                                        
  |  +-------------------------- Gonidea_angulata          
  |  |                                                     
  |  |                       +-- Quadrula_verrucosa **        
  +--+  +--------------------+                             
     |  |                    +-- Quadrula_quadrula **         
     |  |                                                  
     +--+  +-------------------- Amblema_plicata **           
        |  |                                               
        |  |              +----- Eurynia_dilatata **          
        +--+  +-----------+                                
           |  |           |  +-- Fusconaia_flava **           
           |  |           +--+                             
           |  |              +-- Pleurobema_sintoxia **       
           +--+                                            
              |  +-------------- Obliquaria_reflexa **        
              |  |                                         
              |  +-------------- Truncilla_truncata **        
              +--+                                         
                 +-------------- Ptychobranchus_fasciolaris **
                 |                                         
                 |  +----------- Ligumia_recta **             
                 +--+                                      
                    |  +-------- Cambarunio_iris **           
                    +--+                                   
                       |  +----- Epioblasma_triquetra **      
                       +--+                                
                          |  +-- Lampsilis_cardium **         
                          +--+                             
                             +-- Ortmanniana_ligamentina **   

Lydeard et al., 2000 [Publ 2654], fig. 2c

iT tT bi riS oT roT roS iD
14 16 9 4 2 1 1 30.0
  +-------------- Anodonta_cygnea       
  |                                     
  +-------------- Cumberlandia_monodonta
  |                                     
  |        +----- Quadrula_nobilis **      
  +--------+                            
  |        |  +-- Pustulosa_infucata **    
  |        +--+                         
  |           +-- Pustulosa_succissa **    
--+                                     
  |        +----- Obovaria_unicolor **     
  +--------+                            
  |        |  +-- Obovaria_olivaria **     
  |        +--+                         
  |           +-- Lampsilis_teres **       
  |                                     
  |           +-- Reginaia_ebenus **       
  |  +--------+                         
  |  |        +-- Reginaia_rotulata **     
  +--+                                  
     |  +-------- Pleurobema_decisum **    
     |  |                               
     +--+  +----- Fusconaia_flava **       
        |  |                            
        +--+  +-- Fusconaia_burkei[1] **   
           |  |                         
           |  +-- Fusconaia_burkei[2] **   
           +--+                         
              +-- Fusconaia_burkei[3] **   
              |                         
              +-- Fusconaia_escambia **    

Lydeard et al., 2000 [Publ 2654], fig. 2d

iT tT bi riS oT roT roS iD
14 14 11 4 2 1 1 29.0
                       +-- Reginaia_ebenus **    
  +--------------------+                      
  |                    +-- Reginaia_rotulata **  
  |                                           
  |                 +----- Lampsilis_teres **    
--+  +--------------+                         
  |  |              |  +-- Obovaria_unicolor **  
  |  |              +--+                      
  |  |                 +-- Obovaria_olivaria **  
  +--+                                        
     |                 +-- Quadrula_nobilis **   
     |                 |                      
     |  +--------------+-- Pustulosa_infucata ** 
     |  |              |                      
     +--+              +-- Pustulosa_succissa ** 
        |                                     
        |  +-------------- Pleurobema_decisum ** 
        +--+                                  
           |  +----------- Fusconaia_flava **    
           +--+                               
              |  +-------- Fusconaia_burkei[1] **
              +--+                            
                 |  +----- Fusconaia_burkei[2] **
                 +--+                         
                    |  +-- Fusconaia_burkei[3] **
                    +--+                      
                       +-- Fusconaia_escambia ** 

Huang et al., 2002 [Publ 2657], fig. 1

iT tT bi riS oT roT roS iD
14 14 12 4 20 18 2 27.0
  +----------------------------------- Sinohyriopsis_cumingii  
  |                                                            
--+  +-------------------------------- Ptychorhynchus_pfisteri 
  |  |                                                         
  +--+  +----------------------------- Megalonaias_nervosa **     
     |  |                                                      
     |  |                       +----- Elliptoideus_sloatianus ** 
     +--+  +--------------------+                              
        |  |                    |  +-- Uniomerus_carolinianus **  
        |  |                    +--+                           
        +--+                       +-- Pleurobema_pyriforme **    
           |                                                   
           |  +----------------------- Amblema_plicata **         
           |  |                                                
           +--+                    +-- Obliquaria_reflexa **      
              |  +-----------------+                           
              |  |                 +-- Plectomerus_dombeyanus **  
              +--+                                             
                 |  +----------------- Medionidus_conradicus **   
                 +--+                                          
                    |  +-------------- Potamilus_fragilis **      
                    +--+                                       
                       |  +----------- Ptychobranchus_subtentus **
                       +--+                                    
                          |  +-------- Potamilus_purpuratus **    
                          +--+                                 
                             |  +----- Lampsilis_teres **         
                             +--+                              
                                |  +-- Villosa_delumbis **        
                                +--+                           
                                   +-- Ellipsaria_lineolata **    

Huang et al., 2002 [Publ 2657], fig. 2

iT tT bi riS oT roT roS iD
14 14 6 4 20 18 2 27.0
           +-------- Strophitus_radiatus      
           |                                  
           +-------- Schistodesmus_lampreyanus
           |                                  
           +-------- Lepidodesma_languilati   
           |                                  
           +-------- Acuticosta_ovata         
           |                                  
           |     +-- Sinanodonta_woodiana     
           +-----+                            
           |     +-- Cristaria_plicata        
  +--------+                                  
  |        |     +-- Lanceolaria_grayii       
  |        +-----+                            
  |        |     +-- Lanceolaria_lanceolata   
  |        |                                  
  |        |     +-- Nodularia_douglasiae     
  |        +-----+                            
  |        |     +-- Tchangsinaia_piscicula   
  |        |                                  
  |        |  +----- Utterbackia_imbecillis   
  |        |  |                               
--+        +--+----- Utterbackiana_couperiana 
  |           |                               
  |           |  +-- Alasmidonta_triangulata  
  |           +--+                            
  |              +-- Lasmigona_costata        
  |                                           
  |  +-------------- Ptychorhynchus_pfisteri  
  |  |                                        
  |  +-------------- Sinohyriopsis_cumingii   
  |  |                                        
  |  +-------------- Lamprotula_leaii         
  |  |                                        
  +--+-------------- Sinosolenaia_oleivora    
     |                                        
     |  +----------- Megalonaias_nervosa **      
     |  |                                     
     |  |     +----- Elliptoideus_sloatianus **  
     |  |  +--+                               
     +--+  |  |  +-- Uniomerus_carolinianus **   
        |  |  +--+                            
        |  |     +-- Pleurobema_pyriforme **     
        |  |                                  
        |  |  +----- Potamilus_fragilis **       
        +--+  |                               
           |  +----- Medionidus_conradicus **    
           |  |                               
           |  +----- Amblema_plicata **          
           |  |                               
           |  |  +-- Obliquaria_reflexa **       
           +--+--+                            
              |  +-- Plectomerus_dombeyanus **   
              |                               
              |  +-- Villosa_delumbis **         
              |  |                            
              |  +-- Ellipsaria_lineolata **     
              |  |                            
              +--+-- Lampsilis_teres **          
                 |                            
                 +-- Potamilus_purpuratus **     
                 |                            
                 +-- Ptychobranchus_subtentus ** 

Hoeh et al., 2009 [Publ 2716], fig. 3

iT tT bi riS oT roT roS iD
14 14 12 4 33 11 3 28.0
  +-------------------------- Lens_contradens           
  |                                                     
  |                    +----- Gonidea_angulata          
  |  +-----------------+                                
  |  |                 |  +-- Pilsbryoconcha_exilis     
--+  |                 +--+                             
  |  |                    +-- Pseudodon_vondembuschianus
  |  |                                                  
  |  |           +----------- Amblema_plicata **           
  |  |           |                                      
  +--+           |     +----- Obliquaria_reflexa **        
     |  +--------+  +--+                                
     |  |        |  |  |  +-- Quadrula_verrucosa **        
     |  |        |  |  +--+                             
     |  |        +--+     +-- Quadrula_quadrula **         
     |  |           |                                   
     |  |           |  +----- Eurynia_dilatata **          
     +--+           +--+                                
        |              |  +-- Pleurobema_sintoxia **       
        |              +--+                             
        |                 +-- Fusconaia_flava **           
        |                                               
        |  +----------------- Ptychobranchus_fasciolaris **
        +--+                                            
           |  +-------------- Truncilla_truncata **        
           +--+                                         
              |  +----------- Ortmanniana_ligamentina **   
              +--+                                      
                 |  +-------- Lampsilis_cardium **         
                 +--+                                   
                    |  +----- Cambarunio_iris **           
                    +--+                                
                       |  +-- Ligumia_recta **             
                       +--+                             
                          +-- Epioblasma_triquetra **      

Turner et al., 2000 [Publ 2722], fig. 2

iT tT bi riS oT roT roS iD
14 15 11 4 2 1 1 22.0
                          +-- Quadrula_quadrula **       
  +-----------------------+                           
  |                       +-- Utterbackiana_couperiana
  |                                                   
--+                       +-- Amblema_plicata **         
  |  +--------------------+                           
  |  |                    +-- Eurynia_dilatata **        
  +--+                                                
     |  +-------------------- Medionidus_conradicus **   
     |  |                                             
     +--+  +----------------- Lampsilis_teres **         
        |  |                                          
        |  +----------------- Villosa_delumbis **        
        +--+                                          
           |              +-- Lampsilis_satura **        
           |  +-----------+                           
           |  |           +-- Lampsilis_cardium **       
           +--+                                       
              |  +----------- Lampsilis_reeveiana **     
              |  |                                    
              +--+  +-------- Lampsilis_hydiana[1] **    
                 |  |                                 
                 +--+-------- Lampsilis_hydiana[2] **    
                    |                                 
                    |  +----- Lampsilis_siliquoidea[2] **
                    +--+                              
                       |  +-- Lampsilis_siliquoidea[1] **
                       +--+                           
                          +-- Lampsilis_powellii **      

Kandl et al., 2001 [Publ 3384], fig. 5

iT tT bi riS oT roT roS iD
14 14 7 3 0 0 0 21.0
  +-------------------- Amblema_plicata **        
  |                                            
  +-------------------- Quadrula_quadrula **      
--+                                            
  |  +----------------- Elliptio_crassidens **    
  |  |                                         
  +--+  +-------------- Fusconaia_flava **        
     |  |                                      
     +--+  +----------- Pleurobema_sintoxia **    
        |  |                                   
        |  |        +-- Pleurobema_clava **       
        +--+  +-----+                          
           |  |     +-- Pleurobema_strodeanum **  
           |  |                                
           +--+  +----- Pleurobema_pyriforme[6] **
              |  |                             
              |  |  +-- Pleurobema_pyriforme[1] **
              +--+  |                          
                 |  +-- Pleurobema **             
                 |  |                          
                 |  +-- Pleurobema_pyriforme[2] **
                 +--+                          
                    +-- Pleurobema_pyriforme[3] **
                    |                          
                    +-- Pleurobema_pyriforme[4] **
                    |                          
                    +-- Pleurobema_pyriforme[5] **

Hewitt et al., 2019 [Publ 3962], fig. 2

iT tT bi riS oT roT roS iD
14 14 9 2 2 1 1 19.0
  +-------------------------- Pyganodon_grandis    
  |                                                
--+  +----------------------- Fusconaia_flava **      
  |  |                                             
  +--+  +-------------------- Lampsilis_siliquoidea **
     |  |                                          
     |  |  +----------------- Lampsilis_fasciola **   
     +--+  |                                       
        |  |              +-- Epioblasma_rangiana **  
        |  +--------------+                        
        +--+              +-- Lampsilis_radiata **    
           |                                       
           |  +-------------- Lampsilis_ornata **     
           |  |                                    
           +--+           +-- Lampsilis_cardium[1] ** 
              |  +--------+                        
              |  |        +-- Lampsilis_ovata[1] **   
              +--+                                 
                 |  +-------- Lampsilis_cardium[6] ** 
                 |  |                              
                 +--+  +----- Lampsilis_cardium[5] ** 
                    |  |                           
                    +--+  +-- Lampsilis_cardium[2] ** 
                       |  |                        
                       |  +-- Lampsilis_ovata[2] **   
                       +--+                        
                          +-- Lampsilis_cardium[3] ** 
                          |                        
                          +-- Lampsilis_cardium[4] ** 

Teiga-Teixeira et al., 2020 [Publ 4173], fig. 1

iT tT bi riS oT roT roS iD
14 14 12 5 5 3 2 3.0
                          +----- Lanceolaria_grayii         
  +-----------------------+                                 
  |                       |  +-- Chamberlainia_somsakpanhai 
  |                       +--+                              
  |                          +-- Lamprotula_leaii           
--+                                                         
  |                          +-- Uniomerus_tetralasmus **      
  |  +-----------------------+                              
  |  |                       +-- Quadrula_quadrula **          
  |  |                                                      
  +--+                       +-- Elliptio_complanata **        
     |  +--------------------+                              
     |  |                    +-- Pleurobema_oviforme **        
     +--+                                                   
        |  +-------------------- Amblema_plicata **            
        |  |                                                
        +--+  +----------------- Popenaias_popeii **           
           |  |                                             
           +--+  +-------------- Toxolasma_parvum **           
              |  |                                          
              |  |           +-- Potamilus_alatus **           
              +--+  +--------+                              
                 |  |        +-- Potamilus_leptodon **         
                 |  |                                       
                 +--+        +-- Lampsilis_cardium **          
                    |  +-----+                              
                    |  |     +-- Lampsilis_ornata **           
                    +--+                                    
                       |  +----- Venustaconcha_ellipsiformis **
                       +--+                                 
                          |  +-- Lampsilis_powellii **         
                          +--+                              
                             +-- Lampsilis_siliquoidea **      

Serb & Barnhart, 2008 [Publ 2674], fig. 3

iT tT bi riS oT roT roS iD
13 13 10 1 0 0 0 20.0
                          +-- Lampsilis_siliquoidea **     
  +-----------------------+                             
  |                       +-- Lampsilis_ornata **          
--+                                                     
  |  +----------------------- Obliquaria_reflexa **        
  |  |                                                  
  +--+  +-------------------- Ptychobranchus_fasciolaris **
     |  |                                               
     +--+                 +-- Cyprogenia_aberti[6] **      
        |  +--------------+                             
        |  |              +-- Dromus_dromas **             
        +--+                                            
           |  +-------------- Cyprogenia_aberti[4] **      
           |  |                                         
           +--+  +----------- Cyprogenia_aberti[3] **      
              |  |                                      
              +--+  +-------- Cyprogenia_aberti[5] **      
                 |  |                                   
                 +--+  +----- Cyprogenia_stegaria[2] **    
                    |  |                                
                    +--+----- Cyprogenia_aberti[2] **      
                       |                                
                       |  +-- Cyprogenia_stegaria[1] **    
                       +--+                             
                          +-- Cyprogenia_aberti[1] **      

Serb & Barnhart, 2008 [Publ 2674], fig. 4

iT tT bi riS oT roT roS iD
13 13 9 1 0 0 0 20.0
  +----------------------- Obliquaria_reflexa **        
  |                                                  
  |                    +-- Lampsilis_ornata **          
  +--------------------+                             
--+                    +-- Lampsilis_siliquoidea **     
  |                                                  
  |  +-------------------- Ptychobranchus_fasciolaris **
  |  |                                               
  +--+                 +-- Cyprogenia_aberti[6] **      
     |  +--------------+                             
     |  |              +-- Dromus_dromas **             
     +--+                                            
        |  +-------------- Cyprogenia_aberti[5] **      
        |  |                                         
        +--+  +----------- Cyprogenia_aberti[4] **      
           |  |                                      
           +--+  +-------- Cyprogenia_aberti[3] **      
              |  |                                   
              +--+  +----- Cyprogenia_stegaria[2] **    
                 |  |                                
                 +--+----- Cyprogenia_aberti[2] **      
                    |                                
                    |  +-- Cyprogenia_stegaria[1] **    
                    +--+                             
                       +-- Cyprogenia_aberti[1] **      

Serb, 2006 [Publ 2678], fig. 2

iT tT bi riS oT roT roS iD
13 13 7 1 0 0 0 20.0
  +-------------- Obliquaria_reflexa **        
  |                                         
  |        +----- Ptychobranchus_fasciolaris **
  +--------+                                
  |        |  +-- Lampsilis_ornata **          
--+        +--+                             
  |           +-- Lampsilis_siliquoidea **     
  |                                         
  |           +-- Cyprogenia_aberti[6] **      
  |  +--------+                             
  |  |        +-- Dromus_dromas **             
  +--+                                      
     |  +-------- Cyprogenia_aberti[5] **      
     |  |                                   
     +--+  +----- Cyprogenia_aberti[4] **      
        |  |                                
        |  |  +-- Cyprogenia_stegaria[1] **    
        +--+  |                             
           |  +-- Cyprogenia_stegaria[2] **    
           |  |                             
           +--+-- Cyprogenia_aberti[1] **      
              |                             
              +-- Cyprogenia_aberti[2] **      
              |                             
              +-- Cyprogenia_aberti[3] **      

Hoeh et al., 2009 [Publ 2716], fig. 4

iT tT bi riS oT roT roS iD
13 13 10 3 33 11 3 27.0
                          +-- Unio_pictorum             
                 +--------+                             
                 |        +-- Unio_caffer               
  +--------------+                                      
  |              |  +-------- Lasmigona_costata         
  |              +--+                                   
  |                 |  +----- Alasmidonta_marginata     
  |                 +--+                                
  |                    |  +-- Strophitus_undulatus      
  |                    +--+                             
--+                       +-- Pyganodon_grandis         
  |                                                     
  |  +----------------------- Gonidea_angulata          
  |  |                                                  
  |  |                    +-- Eurynia_dilatata **          
  |  |                    |                             
  |  |              +-----+-- Pleurobema_sintoxia **       
  +--+              |     |                             
     |  +-----------+     +-- Fusconaia_flava **           
     |  |           |                                   
     |  |           |  +----- Obliquaria_reflexa **        
     |  |           +--+                                
     +--+              |  +-- Quadrula_verrucosa **        
        |              +--+                             
        |                 +-- Quadrula_quadrula **         
        |                                               
        |  +----------------- Ptychobranchus_fasciolaris **
        +--+                                            
           |  +-------------- Truncilla_truncata **        
           +--+                                         
              |  +----------- Ortmanniana_ligamentina **   
              +--+                                      
                 |  +-------- Lampsilis_cardium **         
                 +--+                                   
                    |  +----- Cambarunio_iris **           
                    +--+                                
                       |  +-- Ligumia_recta **             
                       +--+                             
                          +-- Epioblasma_triquetra **      

Hoeh et al., 2009 [Publ 2716], fig. 5

iT tT bi riS oT roT roS iD
13 13 7 4 33 11 3 25.0
                 +-- Unio_pictorum             
        +--------+                             
        |        +-- Unio_caffer               
  +-----+                                      
  |     |  +-------- Lasmigona_costata         
  |     +--+                                   
  |        |  +----- Alasmidonta_marginata     
  |        +--+                                
  |           |  +-- Strophitus_undulatus      
--+           +--+                             
  |              +-- Pyganodon_grandis         
  |                                            
  |  +-------------- Gonidea_angulata          
  |  |                                         
  |  |        +----- Obliquaria_reflexa **        
  |  |  +-----+                                
  +--+  |     |  +-- Quadrula_verrucosa **        
     |  |     +--+                             
     |  |        +-- Quadrula_quadrula **         
     |  |                                      
     +--+  +-------- Amblema_plicata **           
        |  |                                   
        |  |  +----- Eurynia_dilatata **          
        |  +--+                                
        |  |  |  +-- Pleurobema_sintoxia **       
        +--+  +--+                             
           |     +-- Fusconaia_flava **           
           |                                   
           |  +----- Truncilla_truncata **        
           |  |                                
           |  +----- Ptychobranchus_fasciolaris **
           +--+                                
              |  +-- Ortmanniana_ligamentina **   
              |  |                             
              |  +-- Cambarunio_iris **           
              +--+                             
                 +-- Ligumia_recta **             
                 |                             
                 +-- Epioblasma_triquetra **      

Kandl et al., 2001 [Publ 3384], fig. 4

iT tT bi riS oT roT roS iD
13 13 4 2 0 0 0 22.5
  +-------------- Elliptio **               
  |                                      
  +-------------- Uniomerus_carolinianus ** 
--+                                      
  |  +----------- Pleurobema_sintoxia **    
  |  |                                   
  +--+  +-------- Pleurobema_strodeanum **  
     |  |                                
     |  |  +----- Pleurobema_pyriforme[7] **
     +--+  |                             
        |  |  +-- Pleurobema_pyriforme[1] **
        |  |  |                          
        +--+  +-- Pleurobema_pyriforme[2] **
           |  |                          
           |  +-- Pleurobema_pyriforme[3] **
           |  |                          
           |  +-- Pleurobema_pyriforme[4] **
           +--+                          
              +-- Pleurobema_pyriforme[5] **
              |                          
              +-- Pleurobema[1] **          
              |                          
              +-- Pleurobema_pyriforme[6] **
              |                          
              +-- Pleurobema[2] **          

Jadhav & Jamkhedkar, 2009 [Publ 3697], fig. 11

iT tT bi riS oT roT roS iD
13 14 12 4 13 13 3 26.0
           +-------------------- Nodularia_douglasiae      
           |                                               
  +--------+  +----------------- Anodonta_cygnea           
  |        |  |                                            
  |        +--+  +-------------- Unio_pictorum[2]          
  |           |  |                                         
  |           +--+           +-- Unio_pictorum[1]          
  |              |  +--------+                             
  |              |  |        +-- Unio_caffer               
  |              +--+                                      
  |                 |  +-------- Anodonta                  
  |                 |  |                                   
--+                 +--+  +----- Lasmigona_costata         
  |                    |  |                                
  |                    +--+  +-- Strophitus_undulatus      
  |                       |  |                             
  |                       +--+-- Alasmidonta_marginata     
  |                          |                             
  |                          +-- Pyganodon_grandis         
  |                                                        
  |  +-------------------------- Pronodularia_japanensis   
  |  |                                                     
  |  |                       +-- Quadrula_quadrula **         
  +--+  +--------------------+                             
     |  |                    +-- Quadrula_verrucosa **        
     |  |                                                  
     +--+  +-------------------- Gonidea_angulata          
        |  |                                               
        |  |              +----- Eurynia_dilatata **          
        +--+  +-----------+                                
           |  |           |  +-- Cambarunio_iris **           
           |  |           +--+                             
           |  |              +-- Ptychobranchus_fasciolaris **
           +--+                                            
              |              +-- Amblema_plicata **           
              |  +-----------+                             
              |  |           +-- Pleurobema_sintoxia **       
              +--+                                         
                 |  +----------- Truncilla_truncata **        
                 |  |                                      
                 +--+        +-- Lampsilis_cardium **         
                    |  +-----+                             
                    |  |     +-- Epioblasma_triquetra **      
                    +--+                                   
                       |  +----- Ligumia_recta **             
                       +--+                                
                          |  +-- Obliquaria_reflexa **        
                          +--+                             
                             +-- Ortmanniana_ligamentina **   

Serb, 2006 [Publ 2678], fig. 3

iT tT bi riS oT roT roS iD
12 12 8 1 0 0 0 20.0
  +-------------------- Obliquaria_reflexa **        
  |                                               
  |                 +-- Lampsilis_ornata **          
  +-----------------+                             
--+                 +-- Lampsilis_siliquoidea **     
  |                                               
  |  +----------------- Ptychobranchus_fasciolaris **
  |  |                                            
  +--+              +-- Cyprogenia_aberti[5] **      
     |  +-----------+                             
     |  |           +-- Dromus_dromas **             
     +--+                                         
        |  +----------- Cyprogenia_aberti[4] **      
        |  |                                      
        +--+  +-------- Cyprogenia_aberti[3] **      
           |  |                                   
           +--+  +----- Cyprogenia_stegaria[2] **    
              |  |                                
              +--+----- Cyprogenia_aberti[2] **      
                 |                                
                 |  +-- Cyprogenia_stegaria[1] **    
                 +--+                             
                    +-- Cyprogenia_aberti[1] **      

Serb, 2006 [Publ 2678], fig. 4

iT tT bi riS oT roT roS iD
12 12 8 1 0 0 0 20.0
  +-------------------- Obliquaria_reflexa **        
  |                                               
  |                 +-- Lampsilis_ornata **          
  +-----------------+                             
--+                 +-- Lampsilis_siliquoidea **     
  |                                               
  |  +----------------- Ptychobranchus_fasciolaris **
  |  |                                            
  +--+              +-- Cyprogenia_aberti[5] **      
     |  +-----------+                             
     |  |           +-- Dromus_dromas **             
     +--+                                         
        |  +----------- Cyprogenia_aberti[4] **      
        |  |                                      
        +--+  +-------- Cyprogenia_aberti[3] **      
           |  |                                   
           +--+  +----- Cyprogenia_stegaria[2] **    
              |  |                                
              +--+----- Cyprogenia_aberti[2] **      
                 |                                
                 |  +-- Cyprogenia_stegaria[1] **    
                 +--+                             
                    +-- Cyprogenia_aberti[1] **      

Rosenberg et al., 1997 [Publ 2720], fig. 2A

iT tT bi riS oT roT roS iD
12 17 0 4 11 6 2 28.0
  +----- Anodonta_anatina      
  |                            
  |  +-- Gonidea_angulata      
  |  |                         
  |  +-- Pyganodon_grandis     
  |  |                         
  |  +-- Utterbackia_imbecillis
  |  |                         
  |  +-- Pyganodon_cataracta   
--+  |                         
  |  +-- Lampsilis_straminea **   
  |  |                         
  |  +-- Lampsilis_teres **       
  |  |                         
  |  +-- Megalonaias_nervosa **   
  |  |                         
  |  +-- Obliquaria_reflexa **    
  |  |                         
  +--+-- Theliderma_cylindrica ** 
     |                         
     +-- Quadrula_quadrula **     
     |                         
     +-- Uniomerus_tetralasmus ** 
     |                         
     +-- Amblema_plicata **       
     |                         
     +-- Elliptio_complanata **   
     |                         
     +-- Fusconaia_cerina **      
     |                         
     +-- Plectomerus_dombeyanus **
     |                         
     +-- Pleurobema_cordatum **   
     |                         
     +-- Unio_pictorum         

Roe, 2013 [Publ 3571], fig. 4

iT tT bi riS oT roT roS iD
12 12 9 1 0 0 0 24.0
  +----------------------- Obliquaria_reflexa **            
  |                                                      
  |                    +-- Lampsilis_siliquoidea **         
  +--------------------+                                 
--+                    +-- Lampsilis_ornata **              
  |                                                      
  |  +-------------------- Medionidus_conradicus **         
  |  |                                                   
  +--+                 +-- Cyprogenia_aberti **             
     |  +--------------+                                 
     |  |              +-- Dromus_dromas **                 
     +--+                                                
        |  +-------------- Ptychobranchus_subtentus **      
        +--+                                             
           |  +----------- Ptychobranchus_jonesi **         
           +--+                                          
              |  +-------- Ptychobranchus_greenii **        
              +--+                                       
                 |  +----- Ptychobranchus_occidentalis[2] **
                 +--+                                    
                    |  +-- Ptychobranchus_occidentalis[1] **
                    +--+                                 
                       +-- Ptychobranchus_fasciolaris **    

Smith et al., 2020 [Publ 4136], fig. 3

iT tT bi riS oT roT roS iD
12 12 10 1 0 0 0 28.0
  +-------------- Pleurobema_clava **    
  |                                   
  |        +----- Fusconaia_subrotunda **
  |  +-----+                          
--+  |     |  +-- Fusconaia_cor **       
  |  |     +--+                       
  |  |        +-- Fusconaia_cuneolus **  
  |  |                                
  |  |        +-- Fusconaia_iheringi **  
  +--+     +--+                       
     |     |  +-- Fusconaia_mitchelli ** 
     |  +--+                          
     |  |  |  +-- Fusconaia_askewi **    
     |  |  +--+                       
     |  |     +-- Fusconaia_chunii **    
     +--+                             
        |     +-- Fusconaia_flava **     
        |  +--+                       
        |  |  +-- Fusconaia_masoni **    
        +--+                          
           |  +-- Fusconaia_burkei **    
           +--+                       
              +-- Fusconaia_escambia **  

Porto-Hannes et al., 2021 [Publ 4227], fig. 4

iT tT bi riS oT roT roS iD
12 12 6 4 0 0 0 21.0
  +-------------------- Quadrula_quadrula **      
  |                                            
  +-------------------- Pustulosa_pustulosa **    
--+                                            
  |  +----------------- Pleurobema_sintoxia **    
  |  |                                         
  +--+  +-------------- Amblema_plicata **        
     |  |                                      
     +--+  +----------- Obovaria_olivaria **      
        |  |                                   
        +--+  +-------- Ortmanniana_ligamentina **
           |  |                                
           +--+  +----- Lampsilis_hydiana **      
              |  |                             
              |  +----- Lampsilis_straminea **    
              +--+                             
                 |  +-- Lampsilis_siliquoidea **  
                 |  |                          
                 |  +-- Lampsilis_radiata **      
                 +--+                          
                    +-- Lampsilis[1] **           
                    |                          
                    +-- Lampsilis[2] **           

Stodola et al., 2021 [Publ 4266], fig. 4

iT tT bi riS oT roT roS iD
12 12 7 1 0 0 0 25.0
              +-- Ortmanniana_ligamentina **
              |                          
  +-----------+-- Lampsilis_abrupta **      
  |           |                          
  |           +-- Lampsilis_higginsii **    
--+                                      
  |  +----------- Lampsilis_sietmani **     
  |  |                                   
  |  |        +-- Lampsilis_siliquoidea **  
  +--+  +-----+                          
     |  |     +-- Lampsilis_radiata **      
     |  |                                
     +--+  +----- Lampsilis_hydiana[2] **   
        |  |                             
        |  +----- Lampsilis_straminea[2] ** 
        |  |                             
        +--+  +-- Lampsilis_hydiana[1] **   
           +--+                          
           |  +-- Lampsilis_bergmanni **    
           |                             
           |  +-- Lampsilis_straminea[1] ** 
           +--+                          
              +-- Lampsilis_virescens **    

Dai et al., 2024 [Publ 4358], fig. 4

iT tT bi riS oT roT roS iD
12 12 10 5 40 38 2 5.0
                                      +-- Quadrula_quadrula **            
                 +--------------------+                                
                 |                    +-- Uniomerus_tetralasmus **        
                 |                                                     
  +--------------+                    +-- Elliptio_complanata **          
  |              |  +-----------------+                                
  |              |  |                 +-- Pleurobema_oviforme **          
  |              +--+                                                  
  |                 |  +----------------- Amblema_plicata **              
  |                 |  |                                               
  |                 +--+  +-------------- Popenaias_popeii **             
  |                    |  |                                            
  |                    +--+  +----------- Toxolasma_parvum **             
  |                       |  |                                         
  |                       +--+        +-- Potamilus_leptodon **           
  |                          |  +-----+                                
  |                          |  |     +-- Potamilus_alatus **             
  |                          +--+                                      
  |                             |  +----- Leaunio_lienosus **             
  |                             +--+                                   
  |                                |  +-- Lampsilis_cardium **            
  |                                +--+                                
--+                                   +-- Venustaconcha_ellipsiformis **  
  |                                                                    
  |                                   +-- Chamberlainia_somsakpanhai   
  |                    +--------------+                                
  |                    |              +-- Sinohyriopsis_cumingii       
  |                    |                                               
  |                    |           +----- Potomida_littoralis          
  |  +-----------------+  +--------+                                   
  |  |                 |  |        |  +-- Lamprotula_leaii             
  |  |                 |  |        +--+                                
  |  |                 |  |           +-- Pronodularia_japanensis      
  |  |                 +--+                                            
  |  |                    |           +-- Pseudodon_vondembuschianus   
  |  |                    |  +--------+                                
  |  |                    |  |        +-- Pilsbryoconcha_exilis        
  |  |                    +--+                                         
  |  |                       |  +-------- Microcondylaea_bonellii      
  |  |                       +--+                                      
  |  |                          |  +----- Sinosolenaia_oleivora        
  +--+                          +--+                                   
     |                             |  +-- Ptychorhynchus_pfisteri      
     |                             +--+                                
     |                                +-- Parvasolenaia_rivularis      
     |                                                                 
     |  +-------------------------------- Lepidodesma_languilati       
     |  |                                                              
     |  |              +----------------- Lanceolaria_gladiola         
     |  |              |                                               
     |  |  +-----------+  +-------------- Acuticosta_chinensis         
     |  |  |           |  |                                            
     |  |  |           |  |        +----- Cristaria_plicata            
     +--+  |           +--+  +-----+                                   
        |  |              |  |     |  +-- Anemina_arcaeformis          
        |  |              |  |     +--+                                
        |  |              +--+        +-- Sinanodonta_woodiana         
        |  |                 |                                         
        |  |                 |  +-------- Anodonta_anatina             
        |  |                 +--+                                      
        +--+                    |  +----- Platynaias_compressa         
           |                    +--+                                   
           |                       |  +-- Pyganodon_grandis            
           |                       +--+                                
           |                          +-- Utterbackia_imbecillis       
           |                                                           
           |                          +-- Aculamprotula_scripta        
           |  +-----------------------+                                
           |  |                       +-- Aculamprotula_tientsinensis  
           |  |                                                        
           |  |                       +-- Unio_crassus                 
           +--+  +--------------------+                                
              |  |                    +-- Unio_delphinus               
              |  |                                                     
              |  |              +-------- Pseudocuneopsis_capitata     
              +--+  +-----------+                                      
                 |  |           |  +----- Pseudocuneopsis_yangshuoensis
                 |  |           +--+                                   
                 |  |              |  +-- Pseudocuneopsis_yemaoi       
                 |  |              +--+                                
                 +--+                 +-- Pseudocuneopsis_sichuanensis 
                    |                                                  
                    |                 +-- Nodularia_breviconcha        
                    |  +--------------+                                
                    |  |              +-- Nodularia_douglasiae         
                    |  |                                               
                    +--+              +-- Schistodesmus_lampreyanus    
                       |  +-----------+                                
                       |  |           +-- Schistodesmus_spinosus       
                       +--+                                            
                          |  +----------- Tchangsinaia_piscicula       
                          +--+                                         
                             |  +-------- Cuneopsis_rufescens          
                             +--+                                      
                                |  +----- Cuneopsis_celtiformis        
                                +--+                                   
                                   |  +-- Cuneopsis_heudei             
                                   +--+                                
                                      +-- Cuneopsis_demangei           

Bogan & Hoeh, 2000 [Publ 2650], fig. 1

iT tT bi riS oT roT roS iD
11 11 5 4 23 7 3 24.0
  +-------- Toxolasma_lividum **      
  |                                
  +-------- Amblema_plicata **        
  |                                
  |     +-- Glebula_rotundata **      
  +-----+                          
  |     +-- Cyrtonaias_tampicoensis **
  |                                
--+  +----- Potamilus_alatus **       
  +--+                             
  |  |  +-- Ortmanniana_ligamentina **
  |  +--+                          
  |     +-- Ligumia_recta **          
  |                                
  |  +----- Quadrula_quadrula **      
  |  |                             
  +--+  +-- Eurynia_dilatata **       
     |  |                          
     +--+-- Pleurobema_clava **       
        |                          
        +-- Fusconaia_flava **        

Hoeh et al., 2001 [Publ 2652], fig. 14.2

iT tT bi riS oT roT roS iD
11 28 16 4 20 7 3 39.0
  +-------------------- Amblema_plicata **        
  |                                            
  +-------------------- Coelatura_aegyptiaca   
  |                                            
  +-------------------- Eurynia_dilatata **       
  |                                            
  +-------------------- Fusconaia_flava **        
  |                                            
  +-------------------- Gonidea_angulata       
  |                                            
  +-------------------- Pleurobema_clava **       
  |                                            
  +-------------------- Quadrula_quadrula **      
  |                                            
--+        +----------- Ortmanniana_ligamentina **
  +--------+                                   
  |        |  +-------- Glebula_rotundata **      
  |        +--+                                
  |           |  +----- Potamilus_alatus **       
  |           +--+                             
  |              |  +-- Ligumia_recta **          
  |              +--+                          
  |                 +-- Toxolasma_lividum **      
  |                                            
  |  +----------------- Cyrtonaias_tampicoensis **
  |  |                                         
  |  |              +-- Unio_pictorum          
  |  |  +-----------+                          
  +--+  |           +-- Unio_tumidus           
     |  |                                      
     |  |        +----- Pyganodon_grandis      
     |  |  +-----+                             
     +--+  |     |  +-- Anodonta_cygnea        
        |  |     +--+                          
        |  |        +-- Strophitus_undulatus   
        |  |                                   
        |  |     +----- Diplodon_gratus        
        +--+     |                             
           |     +----- Echyridella_menziesii  
           |  +--+                             
           |  |  +----- Lortiella_rugata       
           |  |  |                             
           |  |  |  +-- Castalia_stevensi      
           |  |  +--+                          
           +--+     +-- Velesunio_angasi       
              |                                
              |     +-- Mutela_dubia           
              |  +--+                          
              |  |  +-- Mutela_rostrata        
              +--+                             
                 |  +-- Anodontites_guanarensis
                 |  |                          
                 +--+-- Anodontites_trigona    
                    |                          
                    +-- Monocondylaea_minuana  

Hoeh et al., 2001 [Publ 2652], fig. 14.4

iT tT bi riS oT roT roS iD
11 11 9 4 20 7 3 24.0
  +----------------------- Gonidea_angulata       
  |                                               
  |              +-------- Unio_pictorum          
  |              |                                
  |  +-----------+-------- Unio_tumidus           
--+  |           |                                
  |  |           |  +----- Anodonta_cygnea        
  |  |           +--+                             
  |  |              |  +-- Strophitus_undulatus   
  |  |              +--+                          
  +--+                 +-- Pyganodon_grandis      
     |                                            
     |  +----------------- Amblema_plicata **        
     |  |                                         
     |  |        +-------- Quadrula_quadrula **      
     |  |  +-----+                                
     +--+  |     |  +----- Eurynia_dilatata **       
        |  |     +--+                             
        |  |        |  +-- Fusconaia_flava **        
        |  |        +--+                          
        +--+           +-- Pleurobema_clava **       
           |                                      
           |           +-- Cyrtonaias_tampicoensis **
           |  +--------+                          
           |  |        +-- Glebula_rotundata **      
           +--+                                   
              |  +-------- Toxolasma_lividum **      
              +--+                                
                 |  +----- Potamilus_alatus **       
                 +--+                             
                    |  +-- Ligumia_recta **          
                    +--+                          
                       +-- Ortmanniana_ligamentina **

Hoeh et al., 2002 [Publ 2653], fig. 2

iT tT bi riS oT roT roS iD
11 11 8 4 20 7 3 25.0
        +-------------- Gonidea_angulata       
  +-----+                                      
  |     |  +----------- Unio_pictorum          
  |     +--+                                   
  |        |  +-------- Unio_tumidus           
  |        +--+                                
  |           |  +----- Anodonta_cygnea        
  |           +--+                             
--+              |  +-- Pyganodon_grandis      
  |              +--+                          
  |                 +-- Strophitus_undulatus   
  |                                            
  |                 +-- Glebula_rotundata **      
  |  +--------------+                          
  |  |              +-- Cyrtonaias_tampicoensis **
  |  |                                         
  +--+  +-------------- Toxolasma_lividum **      
     |  |                                      
     |  |        +----- Potamilus_alatus **       
     |  +--------+                             
     +--+        |  +-- Ligumia_recta **          
        |        +--+                          
        |           +-- Ortmanniana_ligamentina **
        |                                      
        |  +----------- Amblema_plicata **        
        +--+                                   
           |  +-------- Quadrula_quadrula **      
           +--+                                
              |  +----- Eurynia_dilatata **       
              +--+                             
                 |  +-- Pleurobema_clava **       
                 +--+                          
                    +-- Fusconaia_flava **        

Roe & Hoeh, 2003 [Publ 2655], fig. 4.2

iT tT bi riS oT roT roS iD
11 16 12 4 20 7 3 29.0
  +----------------------------- Gonidea_angulata       
  |                                                     
--+  +-------------------------- Coelatura_aegyptiaca   
  |  |                                                  
  +--+  +----------------------- Fusconaia_flava **        
     |  |                                               
     |  +----------------------- Amblema_plicata **        
     +--+                                               
        |  +-------------------- Quadrula_quadrula **      
        |  |                                            
        |  |  +----------------- Pleurobema_clava **       
        +--+  |                                         
           |  |  +-------------- Eurynia_dilatata **       
           |  |  |                                      
           +--+  |  +----------- Ortmanniana_ligamentina **
              |  +--+                                   
              |  |  |  +-------- Glebula_rotundata **      
              |  |  +--+                                
              |  |     |  +----- Potamilus_alatus **       
              +--+     +--+                             
                 |        |  +-- Ligumia_recta **          
                 |        +--+                          
                 |           +-- Toxolasma_lividum **      
                 |                                      
                 |  +----------- Cyrtonaias_tampicoensis **
                 |  |                                   
                 +--+        +-- Unio_pictorum          
                    |  +-----+                          
                    |  |     +-- Unio_tumidus           
                    +--+                                
                       |  +----- Pyganodon_grandis      
                       +--+                             
                          |  +-- Anodonta_cygnea        
                          +--+                          
                             +-- Strophitus_undulatus   

Roe & Hoeh, 2003 [Publ 2655], fig. 4.3

iT tT bi riS oT roT roS iD
11 11 9 4 20 7 3 24.0
                       +-- Unio_pictorum          
                 +-----+                          
                 |     +-- Unio_tumidus           
  +--------------+                                
  |              |  +----- Anodonta_cygnea        
  |              +--+                             
  |                 |  +-- Pyganodon_grandis      
  |                 +--+                          
--+                    +-- Strophitus_undulatus   
  |                                               
  |  +-------------------- Gonidea_angulata       
  |  |                                            
  |  |  +----------------- Amblema_plicata **        
  +--+  |                                         
     |  |        +-------- Quadrula_quadrula **      
     |  |  +-----+                                
     +--+  |     |  +----- Eurynia_dilatata **       
        |  |     +--+                             
        |  |        |  +-- Fusconaia_flava **        
        |  |        +--+                          
        +--+           +-- Pleurobema_clava **       
           |                                      
           |           +-- Cyrtonaias_tampicoensis **
           |  +--------+                          
           |  |        +-- Glebula_rotundata **      
           +--+                                   
              |  +-------- Toxolasma_lividum **      
              +--+                                
                 |  +----- Ortmanniana_ligamentina **
                 +--+                             
                    |  +-- Ligumia_recta **          
                    +--+                          
                       +-- Potamilus_alatus **       

Roe & Lydeard, 1998 [Publ 2693], fig. 3

iT tT bi riS oT roT roS iD
11 11 5 2 0 0 0 22.0
  +-------------- Fusconaia_cerina **       
  |                                      
  +-------------- Obliquaria_reflexa **     
--+                                      
  |  +----------- Potamilus_capax **        
  |  |                                   
  |  +----------- Lampsilis_ornata **       
  +--+                                   
     |  +-------- Potamilus_fragilis **     
     |  |                                
     |  |  +----- Potamilus_inflatus **     
     +--+  |                             
        |  |  +-- Potamilus_ohiensis **     
        |  +--+                          
        +--+  +-- Potamilus_amphichaenus ** 
           |                             
           |  +-- Potamilus_purpuratus[1] **
           |  |                          
           +--+-- Potamilus_purpuratus[2] **
              |                          
              +-- Potamilus_alatus **       

Rosenberg et al., 1994 [Publ 2719], fig. 3

iT tT bi riS oT roT roS iD
11 15 2 4 8 5 2 29.0
  +-------- Gonidea_angulata      
  |                               
  |     +-- Elliptio_complanata **   
  |     |                         
  |     +-- Fusconaia_cerina **      
  |     |                         
--+  +--+-- Plectomerus_dombeyanus **
  |  |  |                         
  |  |  +-- Pleurobema_cordatum **   
  |  |  |                         
  |  |  +-- Unio_pictorum         
  |  |                            
  |  |  +-- Pyganodon_cataracta   
  +--+  |                         
     |  +-- Pyganodon_grandis     
     |  |                         
     |  +-- Utterbackia_imbecillis
     |  |                         
     |  +-- Lampsilis_teres **       
     |  |                         
     |  +-- Lampsilis_straminea **   
     +--+                         
        +-- Megalonaias_nervosa **   
        |                         
        +-- Obliquaria_reflexa **    
        |                         
        +-- Theliderma_cylindrica ** 
        |                         
        +-- Quadrula_quadrula **     
        |                         
        +-- Uniomerus_tetralasmus ** 

Rosenberg et al., 1997 [Publ 2720], fig. 2B

iT tT bi riS oT roT roS iD
11 17 0 4 12 7 2 28.0
  +----- Anodonta              
  |                            
  |  +-- Gonidea_angulata      
  |  |                         
  |  +-- Pyganodon_grandis     
  |  |                         
  |  +-- Utterbackia_imbecillis
  |  |                         
  |  +-- Pyganodon_cataracta   
--+  |                         
  |  +-- Lampsilis_straminea **   
  |  |                         
  |  +-- Lampsilis_teres **       
  |  |                         
  |  +-- Megalonaias_nervosa **   
  |  |                         
  |  +-- Obliquaria_reflexa **    
  |  |                         
  +--+-- Theliderma_cylindrica ** 
     |                         
     +-- Quadrula_quadrula **     
     |                         
     +-- Uniomerus_tetralasmus ** 
     |                         
     +-- Amblema_plicata **       
     |                         
     +-- Elliptio_complanata **   
     |                         
     +-- Fusconaia_cerina **      
     |                         
     +-- Plectomerus_dombeyanus **
     |                         
     +-- Aculamprotula_tortuosa
     |                         
     +-- Unio_pictorum         

Fukata & Iigo, 2020 [Publ 4106], fig. 1

iT tT bi riS oT roT roS iD
11 11 9 3 43 40 2 6.0
                          +-- Quadrula_quadrula **          
        +-----------------+                              
        |                 +-- Uniomerus_tetralasmus **      
     +--+                                                
     |  |  +----------------- Elliptio_complanata **        
     |  |  |                                             
     |  +--+              +-- Potamilus_leptodon **         
     |     |  +-----------+                              
     |     |  |           +-- Potamilus_alatus **           
     |     +--+                                          
     |        |  +----------- Leaunio_lienosus **           
     |        |  |                                       
     |        +--+        +-- Lampsilis_cardium **          
     |           |  +-----+                              
     |           |  |     +-- Lampsilis_ornata **           
  +--+           +--+                                    
  |  |              |  +----- Venustaconcha_ellipsiformis **
  |  |              +--+                                 
  |  |                 |  +-- Lampsilis_powellii **         
  |  |                 +--+                              
  |  |                    +-- Lampsilis_siliquoidea **      
  |  |                                                   
  |  |                    +-- Sinohyriopsis_cumingii     
  |  |        +-----------+                              
  |  |        |           +-- Sinohyriopsis_schlegelii   
  |  |        |                                          
  |  |        |           +-- Pilsbryoconcha_exilis      
  |  +--------+  +--------+                              
  |           |  |        +-- Pseudodon_vondembuschianus 
  |           |  |                                       
  |           |  |        +-- Lamprotula_caveata         
  |           |  |     +--+                              
  |           +--+     |  +-- Lamprotula_leaii           
  |              |  +--+                                 
--+              |  |  |  +-- Pronodularia_japanensis    
  |              |  |  +--+                              
  |              |  |     +-- Potomida_littoralis        
  |              +--+                                    
  |                 |     +-- Sinosolenaia_carinata      
  |                 |  +--+                              
  |                 |  |  +-- Sinosolenaia_oleivora      
  |                 +--+                                 
  |                    |  +-- Parvasolenaia_rivularis    
  |                    +--+                              
  |                       +-- Ptychorhynchus_pfisteri    
  |                                                      
  |  +----------------------- Lepidodesma_languilati     
  |  |                                                   
  |  |                 +----- Lanceolaria_gladiola       
  |  |  +--------------+                                 
  |  |  |              |  +-- Lanceolaria_grayii         
  |  |  |              +--+                              
  |  |  |                 +-- Lanceolaria_lanceolata     
  +--+  |                                                
     |  |     +-------------- Acuticosta_chinensis       
     |  |     |                                          
     |  |     |     +-------- Cristaria_plicata          
     |  |     |     |                                    
     |  |     |  +--+     +-- Sinanodonta_woodiana       
     |  |  +--+  |  |  +--+                              
     +--+  |  |  |  |  |  +-- Sinanodonta_lucida         
        |  |  |  |  +--+                                 
        |  |  |  |     |  +-- Anemina_arcaeformis        
        |  |  |  |     +--+                              
        |  |  +--+        +-- Anemina_euscaphys          
        |  |     |                                       
        |  |     |        +-- Anodonta_anatina           
        |  |     |  +-----+                              
        |  |     |  |     +-- Anodonta_cygnea            
        |  |     |  |                                    
        |  |     +--+     +-- Pyganodon_grandis          
        +--+        |  +--+                              
           |        |  |  +-- Utterbackia_peninsularis   
           |        +--+                                 
           |           |  +-- Alasmidonta_varicosa       
           |           +--+                              
           |              +-- Prolasmidonta_heterodon    
           |                                             
           |           +----- Aculamprotula_tortuosa     
           |  +--------+                                 
           |  |        |  +-- Aculamprotula_tientsinensis
           |  |        +--+                              
           |  |           +-- Aculamprotula_coreana      
           |  |                                          
           +--+     +-------- Unio_tumidus               
              |  +--+                                    
              |  |  |  +----- Unio_crassus               
              |  |  +--+                                 
              |  |     |  +-- Unio_pictorum              
              |  |     +--+                              
              +--+        +-- Unio_delphinus             
                 |                                       
                 |        +-- Schistodesmus_lampreyanus  
                 |  +-----+                              
                 |  |     +-- Cuneopsis_heudei           
                 +--+                                    
                    |  +----- Pseudocuneopsis_capitata   
                    +--+                                 
                       |  +-- Nodularia_douglasiae       
                       +--+                              
                          +-- Tchangsinaia_piscicula     

Buhay et al., 2002 [Publ 2661], fig. 2A

iT tT bi riS oT roT roS iD
10 10 3 1 0 0 0 22.0
  +-------- Lampsilis_siliquoidea **     
  |                                   
  |     +-- Cambarunio_taeniatus **      
  +-----+                             
  |     +-- Villosa_villosa **           
--+                                   
  |  +----- Epioblasma_triquetra **      
  |  |                                
  |  +----- Epioblasma_brevidens **      
  |  |                                
  +--+  +-- Epioblasma_capsaeformis[1] **
     |  |                             
     |  +-- Epioblasma_capsaeformis[2] **
     |  |                             
     +--+-- Epioblasma_capsaeformis[3] **
        |                             
        +-- Epioblasma_walkeri[1] **     
        |                             
        +-- Epioblasma_walkeri[2] **     

Buhay et al., 2002 [Publ 2661], fig. 2B

iT tT bi riS oT roT roS iD
10 10 5 1 0 0 0 22.0
  +-------------- Lampsilis_siliquoidea **     
  |                                         
  |           +-- Cambarunio_taeniatus **      
  +-----------+                             
--+           +-- Villosa_villosa **           
  |                                         
  |  +----------- Epioblasma_brevidens **      
  |  |                                      
  +--+  +-------- Epioblasma_triquetra **      
     |  |                                   
     +--+  +----- Epioblasma_capsaeformis[3] **
        |  |                                
        +--+  +-- Epioblasma_capsaeformis[1] **
           |  |                             
           |  +-- Epioblasma_capsaeformis[2] **
           +--+                             
              +-- Epioblasma_walkeri[1] **     
              |                             
              +-- Epioblasma_walkeri[2] **     

Huang et al., 2019 [Publ 3843], fig. 3

iT tT bi riS oT roT roS iD
10 10 8 3 47 37 2 7.0
                                +-- Sinohyriopsis_cumingii     
                    +-----------+                              
                    |           +-- Sinohyriopsis_schlegelii   
                    |                                          
                    |     +-------- Potomida_littoralis        
           +--------+  +--+                                    
           |        |  |  |  +----- Pronodularia_japanensis    
           |        |  |  +--+                                 
           |        |  |     |  +-- Lamprotula_caveata         
           |        |  |     +--+                              
           |        +--+        +-- Lamprotula_leaii           
           |           |                                       
           |           |        +-- Ptychorhynchus_pfisteri    
           |           |     +--+                              
  +--------+           |     |  +-- Parvasolenaia_rivularis    
  |        |           +-----+                                 
  |        |                 |  +-- Sinosolenaia_oleivora      
  |        |                 +--+                              
  |        |                    +-- Sinosolenaia_carinata      
  |        |                                                   
  |        |                    +-- Quadrula_quadrula **          
  |        |  +-----------------+                              
  |        |  |                 +-- Uniomerus_tetralasmus **      
  |        +--+                                                
  |           |  +----------------- Elliptio_complanata **        
  |           |  |                                             
  |           +--+  +-------------- Toxolasma_parvum **           
  |              |  |                                          
  |              +--+           +-- Potamilus_leptodon **         
  |                 |  +--------+                              
  |                 |  |        +-- Potamilus_alatus **           
--+                 +--+                                       
  |                    |  +-------- Leaunio_lienosus **           
  |                    +--+                                    
  |                       |  +----- Venustaconcha_ellipsiformis **
  |                       +--+                                 
  |                          |  +-- Lampsilis_cardium **          
  |                          +--+                              
  |                             +-- Lampsilis_ornata **           
  |                                                            
  |  +----------------------------- Lepidodesma_languilati     
  |  |                                                         
  |  |                    +-------- Aculamprotula_tortuosa     
  |  |              +-----+                                    
  |  |              |     |  +----- Aculamprotula_scripta      
  |  |              |     +--+                                 
  |  |              |        |  +-- Aculamprotula_tientsinensis
  |  |              |        +--+                              
  |  |  +-----------+           +-- Aculamprotula_coreana      
  +--+  |           |                                          
     |  |           |        +----- Nodularia_douglasiae       
     |  |           |  +-----+                                 
     |  |           |  |     |  +-- Tchangsinaia_piscicula     
     |  |           |  |     +--+                              
     |  |           +--+        +-- Schistodesmus_lampreyanus  
     |  |              |                                       
     |  |              |  +-------- Unio_tumidus               
     |  |              +--+                                    
     +--+                 |  +----- Unio_crassus               
        |                 +--+                                 
        |                    |  +-- Unio_delphinus             
        |                    +--+                              
        |                       +-- Unio_pictorum              
        |                                                      
        |                    +----- Lanceolaria_gladiola       
        |  +-----------------+                                 
        |  |                 |  +-- Lanceolaria_lanceolata     
        |  |                 +--+                              
        |  |                    +-- Lanceolaria_grayii         
        +--+                                                   
           |  +-------------------- Acuticosta_chinensis       
           |  |                                                
           |  |           +-------- Cristaria_plicata          
           |  |           |                                    
           +--+  +--------+     +-- Sinanodonta_lucida         
              |  |        |  +--+                              
              |  |        |  |  +-- Sinanodonta_woodiana       
              |  |        +--+                                 
              +--+           |  +-- Anemina_arcaeformis        
                 |           +--+                              
                 |              +-- Anemina_euscaphys          
                 |                                             
                 |  +-------------- Anodonta_cygnea            
                 +--+                                          
                    |  +----------- Anodonta_anatina           
                    +--+                                       
                       |  +-------- Lasmigona_costata          
                       +--+                                    
                          |  +----- Pyganodon_grandis          
                          +--+                                 
                             |  +-- Utterbackia_imbecillis     
                             +--+                              
                                +-- Utterbackia_peninsularis   

Keogh & Simons, 2019 [Publ 3935], fig. S1

iT tT bi riS oT roT roS iD
10 10 8 3 0 0 0 21.0
                    +-- Amblema_plicata **      
  +-----------------+                        
  |                 +-- Fusconaia_mitchelli **  
--+                                          
  |  +----------------- Potamilus_fragilis **   
  |  |                                       
  +--+  +-------------- Leaunio_lienosus **     
     |  |                                    
     +--+           +-- Lampsilis_floridensis **
        |  +--------+                        
        |  |        +-- Lampsilis_teres **      
        +--+                                 
           |  +-------- Lampsilis_sietmani **   
           +--+                              
              |  +----- Lampsilis_siliquoidea **
              +--+                           
                 |  +-- Lampsilis_hydiana **    
                 +--+                        
                    +-- Lampsilis_straminea **  

Curole & Kocher, 2005 [Publ 2682], fig. 1

iT tT bi riS oT roT roS iD
9 9 7 3 2 1 1 22.0
  +----------------- Glebula_rotundata **     
  |                                        
  |           +----- Amblema_plicata **       
--+  +--------+                            
  |  |        |  +-- Quadrula_quadrula **     
  |  |        +--+                         
  +--+           +-- Pustulosa_pustulosa **   
     |                                     
     |  +----------- Plectomerus_dombeyanus **
     +--+                                  
        |  +-------- Leaunio_lienosus **      
        +--+                               
           |  +----- Lampsilis_straminea **   
           +--+                            
              |  +-- Lampsilis_teres **       
              +--+                         
                 +-- Potamilus_purpuratus **  

Curole & Kocher, 2005 [Publ 2682], fig. 1-M

iT tT bi riS oT roT roS iD
9 9 7 3 3 2 1 18.0
                    +-- Utterbackiana_implicata
  +-----------------+                          
  |                 +-- Pyganodon_grandis      
  |                                            
--+                 +-- Quadrula_quadrula **      
  |  +--------------+                          
  |  |              +-- Pustulosa_pustulosa **    
  +--+                                         
     |  +-------------- Plectomerus_dombeyanus ** 
     |  |                                      
     +--+  +----------- Amblema_plicata **        
        |  |                                   
        +--+        +-- Potamilus_purpuratus **   
           |  +-----+                          
           |  |     +-- Glebula_rotundata **      
           +--+                                
              |  +----- Leaunio_lienosus **       
              +--+                             
                 |  +-- Lampsilis_straminea **    
                 +--+                          
                    +-- Lampsilis_teres **        

Lopes-Lima et al., 2017 [Publ 3745], fig. 1-2

iT tT bi riS oT roT roS iD
9 9 7 4 72 60 3 19.0
              +-------- Amblema_plicata **           
           +--+                                   
           |  |  +----- Cambarunio_iris **           
           |  +--+                                
           |     |  +-- Lampsilis_cardium **         
           |     +--+                             
  +--------+        +-- Ortmanniana_ligamentina **   
  |        |                                      
  |        |        +-- Quadrula_quadrula **         
  |        |  +-----+                             
  |        |  |     +-- Quadrula_verrucosa **        
  |        +--+                                   
  |           |  +----- Elliptio_complanata **       
  |           +--+                                
  |              |  +-- Pleurobema_sintoxia **       
  |              +--+                             
  |                 +-- Eurynia_dilatata **          
--+                                               
  |                 +-- Chamberlainia_somsakpanhai
  |           +-----+                             
  |           |     +-- Sinohyriopsis_cumingii    
  |           |                                   
  |  +--------+     +-- Lamprotula_caveata        
  |  |        |  +--+                             
  |  |        |  |  +-- Lamprotula_leaii          
  |  |        +--+                                
  |  |           |  +-- Potomida_littoralis       
  |  |           +--+                             
  |  |              +-- Pronodularia_japanensis   
  +--+                                            
     |              +-- Sinosolenaia_carinata     
     |        +-----+                             
     |        |     +-- Sinosolenaia_oleivora     
     |  +-----+                                   
     |  |     |  +----- Gonidea_angulata          
     |  |     +--+                                
     |  |        |  +-- Microcondylaea_bonellii   
     +--+        +--+                             
        |           +-- Leguminaia_wheatleyi      
        |                                         
        |  +----------- Parvasolenaia_rivularis   
        |  |                                      
        +--+        +-- Pilsbryoconcha_exilis     
           |  +-----+                             
           |  |     +-- Pilsbryoconcha_acuta      
           +--+                                   
              |  +----- Pseudodon_vagulus         
              +--+                                
                 |  +-- Bineurus_exilis           
                 +--+                             
                    +-- Nyeinchanconcha_nyeinchani

Bolotov et al., 2017 [Publ 3761], fig. S1

iT tT bi riS oT roT roS iD
9 9 7 4 132 107 3 19.0
                    +-------- Amblema_plicata **           
                 +--+                                   
                 |  |  +----- Ortmanniana_ligamentina **   
                 |  +--+                                
                 |     |  +-- Lampsilis_cardium **         
                 |     +--+                             
           +-----+        +-- Cambarunio_iris **           
           |     |                                      
           |     |        +-- Quadrula_quadrula **         
           |     |  +-----+                             
           |     |  |     +-- Quadrula_verrucosa **        
           |     +--+                                   
           |        |  +----- Pleurobema_sintoxia **       
           |        +--+                                
           |           |  +-- Eurynia_dilatata **          
           |           +--+                             
           |              +-- Elliptio_complanata **       
  +--------+                                            
  |        |              +-- Sinohyriopsis_cumingii    
  |        |        +-----+                             
  |        |        |     +-- Chamberlainia_somsakpanhai
  |        |        |                                   
  |        |  +-----+     +-- Potomida_littoralis       
  |        |  |     |  +--+                             
  |        |  |     |  |  +-- Pronodularia_japanensis   
  |        |  |     +--+                                
  |        |  |        |  +-- Lamprotula_caveata        
  |        |  |        +--+                             
  |        +--+           +-- Lamprotula_leaii          
  |           |                                         
  |           |        +----- Sinosolenaia_carinata     
  |           |  +-----+                                
  |           |  |     |  +-- Sinosolenaia_oleivora     
  |           |  |     +--+                             
  |           +--+        +-- Sinosolenaia_recognita    
  |              |                                      
  |              |  +-------- Gonidea_angulata          
  |              +--+                                   
--+                 |  +----- Microcondylaea_bonellii   
  |                 +--+                                
  |                    |  +-- Leguminaia_wheatleyi      
  |                    +--+                             
  |                       +-- Leguminaia                
  |                                                     
  |                    +----- Unio_tumidus              
  |           +--------+                                
  |           |        |  +-- Unio_pictorum             
  |           |        +--+                             
  |           |           +-- Unio_crassus              
  |  +--------+                                         
  |  |        |  +----------- Aculamprotula_tortuosa    
  |  |        |  |                                      
  |  |        |  |        +-- Nodularia_douglasiae      
  |  |        +--+  +-----+                             
  |  |           |  |     +-- Nodularia_nuxpersicae     
  |  |           |  |                                   
  |  |           +--+     +-- Cuneopsis_rufescens       
  |  |              |  +--+                             
  |  |              |  |  +-- Cuneopsis_heudei          
  |  |              +--+                                
  +--+                 |  +-- Tchangsinaia_piscicula    
     |                 +--+                             
     |                    +-- Schistodesmus_lampreyanus 
     |                                                  
     |                    +-- Lanceolaria_grayii        
     |  +-----------------+                             
     |  |                 +-- Lanceolaria_gladiola      
     |  |                                               
     |  |                 +-- Cristaria_plicata         
     |  |        +--------+                             
     |  |        |        +-- Cristaria_radiata         
     +--+  +-----+                                      
        |  |     |  +-------- Pletholophus_tenuis       
        |  |     +--+                                   
        |  |        |  +----- Sinanodonta_lucida        
        |  |        +--+                                
        |  |           |  +-- Sinanodonta_jourdyi       
        +--+           +--+                             
           |              +-- Sinanodonta_lauta         
           |                                            
           |  +-------------- Anodonta_anatina          
           |  |                                         
           |  |        +----- Pseudanodonta_complanata  
           +--+  +-----+                                
              |  |     |  +-- Anodonta_cygnea           
              |  |     +--+                             
              |  |        +-- Anodonta_nuttalliana      
              +--+                                      
                 |        +-- Lasmigona_costata         
                 |  +-----+                             
                 |  |     +-- Alasmidonta_marginata     
                 +--+                                   
                    |  +----- Simpsonaias_ambigua       
                    +--+                                
                       |  +-- Pyganodon_grandis         
                       +--+                             
                          +-- Strophitus_undulatus      

Bolotov et al., 2017 [Publ 3761], fig. S2

iT tT bi riS oT roT roS iD
9 9 7 4 132 107 3 19.0
                 +-------- Lens_contradens            
           +-----+                                    
           |     |  +----- Lens_eximius               
           |     +--+                                 
           |        |  +-- Lens_comptus               
           |        +--+                              
        +--+           +-- Lens_novoselovi            
        |  |                                          
        |  |        +----- Physunio                   
        |  |  +-----+                                 
        |  |  |     |  +-- Physunio_modelli           
        |  |  |     +--+                              
        |  +--+        +-- Physunio_superbus          
        |     |                                       
        |     |  +-------- Yaukthwa_inlenensis        
        |     +--+                                    
        |        |  +----- Yaukthwa_panhai            
  +-----+        +--+                                 
  |     |           |  +-- Yaukthwa_rectangularis     
  |     |           +--+                              
  |     |              +-- Yaukthwa_nesemanni         
  |     |                                             
  |     |           +----- Hyriopsis_panhai           
  |     |           |                                 
  |     |  +--------+----- Hyriopsis_khoratensis      
  |     |  |        |                                 
  |     |  |        |  +-- Hyriopsis_myersiana        
  |     |  |        +--+                              
  |     +--+           +-- Hyriopsis_bialata          
  |        |                                          
  |        |  +----------- Hyriopsis_desowitzi        
  |        |  |                                       
  |        |  +----------- Rectidens_sumatrensis      
  |        +--+                                       
  |           |        +-- Ensidens_telus             
  |           |  +-----+                              
  |           |  |     +-- Ensidens_ingallsianus      
  |           +--+                                    
  |              |  +----- Ensidens_sagittarius       
  |              +--+                                 
  |                 |  +-- Ensidens_spiculus          
--+                 +--+                              
  |                    +-- Ensidens_jaculus           
  |                                                   
  |              +-------- Amblema_plicata **            
  |           +--+                                    
  |           |  |  +----- Cambarunio_iris **            
  |           |  +--+                                 
  |           |     |  +-- Ortmanniana_ligamentina **    
  |           |     +--+                              
  |  +--------+        +-- Lampsilis_cardium **          
  |  |        |                                       
  |  |        |        +-- Quadrula_quadrula **          
  |  |        |  +-----+                              
  |  |        |  |     +-- Quadrula_verrucosa **         
  |  |        +--+                                    
  |  |           |  +----- Pleurobema_sintoxia **        
  |  |           +--+                                 
  |  |              |  +-- Elliptio_complanata **        
  |  |              +--+                              
  |  |                 +-- Eurynia_dilatata **           
  |  |                                                
  |  |           +-------- Sinohyriopsis_cumingii     
  |  |           |                                    
  |  |     +-----+     +-- Lamprotula_caveata         
  +--+     |     |  +--+                              
     |     |     |  |  +-- Lamprotula_leaii           
     |     |     +--+                                 
     |     |        |  +-- Potomida_littoralis        
     |  +--+        +--+                              
     |  |  |           +-- Pronodularia_japanensis    
     |  |  |                                          
     |  |  |  +----------- Sinosolenaia_carinata      
     |  |  |  |                                       
     |  |  |  |        +-- Sinosolenaia_oleivora      
     |  |  +--+--------+                              
     |  |     |        +-- Sinosolenaia_recognita     
     |  |     |                                       
     |  |     |  +-------- Gonidea_angulata           
     |  |     +--+                                    
     |  |        |  +----- Microcondylaea_bonellii    
     +--+        +--+                                 
        |           |  +-- Leguminaia_wheatleyi       
        |           +--+                              
        |              +-- Leguminaia                 
        |                                             
        |        +-------- Pilsbryoconcha_exilis[3]   
        |  +-----+                                    
        |  |     |  +----- Pilsbryoconcha_exilis[2]   
        |  |     +--+                                 
        |  |        |  +-- Pilsbryoconcha_exilis[1]   
        |  |        +--+                              
        |  |           +-- Pilsbryoconcha_carinifera  
        |  |                                          
        +--+  +----------- Indopseudodon_indawgyiensis
           |  |                                       
           |  |  +-------- Thaiconcha_callifera       
           |  +--+                                    
           |  |  |  +----- Bineurus_exilis            
           |  |  +--+                                 
           |  |     |  +-- Bineurus_mouhotii          
           +--+     +--+                              
              |        +-- Bineurus_loeiensis         
              |                                       
              |     +----- Pseudodon_vagulus          
              |  +--+                                 
              |  |  |  +-- Pseudodon_mekongi          
              |  |  +--+                              
              |  |     +-- Pseudodon_vondembuschianus 
              +--+                                    
                 |     +-- Pseudodon_inoscularis      
                 |  +--+                              
                 |  |  +-- Sundadontina_tumida        
                 +--+                                 
                    |  +-- Nyeinchanconcha_nyeinchani 
                    +--+                              
                       +-- Sundadontina_cumingii      

Bolotov et al., 2017 [Publ 3764], fig. S1

iT tT bi riS oT roT roS iD
9 9 0 4 122 122 3 20.0
        +-- Amblema_plicata **           
        |                             
        +-- Ortmanniana_ligamentina **   
        |                             
        +-- Lampsilis_cardium **         
        |                             
        +-- Cambarunio_iris **           
        |                             
     +--+-- Elliptio_complanata **       
     |  |                             
     |  +-- Eurynia_dilatata **          
     |  |                             
     |  +-- Pleurobema_sintoxia **       
     |  |                             
     |  +-- Quadrula_quadrula **         
     |  |                             
     |  +-- Quadrula_verrucosa **        
     |                                
     |  +-- Chamberlainia_somsakpanhai
  +--+  |                             
  |  |  +-- Sinohyriopsis_cumingii    
  |  |  |                             
  |  |  +-- Lamprotula_caveata        
  |  |  |                             
  |  |  +-- Lamprotula_leaii          
  |  |  |                             
  |  |  +-- Potomida_littoralis       
  |  |  |                             
  |  |  +-- Pronodularia_japanensis   
  |  |  |                             
  |  +--+-- Gonidea_angulata          
  |     |                             
  |     +-- Leguminaia_wheatleyi      
  |     |                             
  |     +-- Leguminaia                
  |     |                             
  |     +-- Microcondylaea_bonellii   
  |     |                             
  |     +-- Sinosolenaia_carinata     
  |     |                             
  |     +-- Sinosolenaia_oleivora     
  |     |                             
  |     +-- Sinosolenaia_recognita    
--+                                   
  |     +-- Unio_crassus              
  |     |                             
  |     +-- Unio_pictorum             
  |     |                             
  |     +-- Unio_tumidus              
  |     |                             
  |     +-- Aculamprotula_tortuosa    
  |     |                             
  |     +-- Cuneopsis_heudei          
  |  +--+                             
  |  |  +-- Tchangsinaia_piscicula    
  |  |  |                             
  |  |  +-- Cuneopsis_rufescens       
  |  |  |                             
  |  |  +-- Nodularia_douglasiae      
  |  |  |                             
  |  |  +-- Nodularia_nuxpersicae     
  |  |  |                             
  |  |  +-- Schistodesmus_lampreyanus 
  |  |                                
  |  |  +-- Alasmidonta_marginata     
  |  |  |                             
  |  |  +-- Anodonta_anatina          
  +--+  |                             
     |  +-- Anodonta_cygnea           
     |  |                             
     |  +-- Anodonta_nuttalliana      
     |  |                             
     |  +-- Lasmigona_costata         
     |  |                             
     |  +-- Pseudanodonta_complanata  
     |  |                             
     |  +-- Pyganodon_grandis         
     |  |                             
     |  +-- Simpsonaias_ambigua       
     |  |                             
     |  +-- Strophitus_undulatus      
     +--+                             
        +-- Sinanodonta_lauta         
        |                             
        +-- Cristaria_plicata         
        |                             
        +-- Cristaria_radiata         
        |                             
        +-- Pletholophus_tenuis       
        |                             
        +-- Sinanodonta_lucida        
        |                             
        +-- Sinanodonta_jourdyi       
        |                             
        +-- Lanceolaria_gladiola      
        |                             
        +-- Lanceolaria_grayii[1]     
        |                             
        +-- Lanceolaria_grayii[2]     

Wu et al., 2018 [Publ 3836], fig. 2

iT tT bi riS oT roT roS iD
9 51 43 4 44 42 2 61.0
  +-------------------------------- Quadrula_quadrula **          
  |                                                            
  |                             +-- Amblema_plicata **            
  +-----------------------------+                              
  |                             +-- Obovaria_arkansasensis **     
  |                                                            
  |                    +----------- Pleurobema_sintoxia **        
  |                    |                                       
  +--------------------+        +-- Elliptio_complanata **        
  |                    |  +-----+                              
  |                    |  |     +-- Eurynia_dilatata **           
  |                    +--+                                    
  |                       |  +----- Fusconaia_askewi **           
  |                       +--+                                 
  |                          |  +-- Fusconaia_cerina **           
  |                          +--+                              
--+                             +-- Fusconaia_burkei **           
  |                                                            
  |                          +----- Potomida_littoralis        
  |                    +-----+                                 
  |                    |     |  +-- Lamprotula_caveata         
  |                    |     +--+                              
  |  +-----------------+        +-- Lamprotula_leaii           
  |  |                 |                                       
  |  |                 |  +-------- Sinosolenaia_carinata      
  |  |                 |  |                                    
  |  |                 +--+  +----- Gonidea_angulata           
  |  |                    |  |                                 
  |  |                    +--+----- Sinosolenaia_oleivora      
  |  |                       |                                 
  |  |                       |  +-- Parvasolenaia_rivularis    
  |  |                       +--+                              
  |  |                          +-- Ptychorhynchus_pfisteri    
  +--+                                                         
     |  +-------------------------- Sinohyriopsis_cumingii     
     |  |                                                      
     |  |  +----------------------- Lepidodesma_languilati     
     |  |  |                                                   
     |  |  |           +----------- Aculamprotula_tortuosa     
     |  |  |     +-----+                                       
     |  |  |     |     |  +-------- Aculamprotula_scripta      
     |  |  |     |     +--+                                    
     |  |  |     |        |  +----- Aculamprotula_coreana      
     |  |  |     |        +--+                                 
     +--+  |     |           |  +-- Aculamprotula_fibrosa      
        |  +-----+           +--+                              
        |  |     |              +-- Aculamprotula_tientsinensis
        |  |     |                                             
        |  |     |              +-- Unio_crassus               
        |  |     |  +-----------+                              
        |  |     |  |           +-- Unio_pictorum              
        |  |     |  |                                          
        |  |     +--+        +----- Nodularia_douglasiae       
        |  |        |  +-----+                                 
        |  |        |  |     |  +-- Schistodesmus_lampreyanus  
        +--+        |  |     +--+                              
           |        +--+        +-- Schistodesmus_spinosus     
           |           |                                       
           |           |  +-------- Tchangsinaia_piscicula     
           |           +--+                                    
           |              |  +----- Cuneopsis_rufescens        
           |              +--+                                 
           |                 |  +-- Cuneopsis_celtiformis      
           |                 +--+                              
           |                    +-- Cuneopsis_heudei           
           |                                                   
           |           +----------- Acuticosta_chinensis       
           |  +--------+                                       
           |  |        |  +-------- Lanceolaria_gladiola       
           |  |        +--+                                    
           |  |           |  +----- Lanceolaria_grayii         
           |  |           +--+                                 
           |  |              |  +-- Lanceolaria_lanceolata     
           |  |              +--+                              
           |  |                 +-- Lanceolaria_triformis      
           +--+                                                
              |           +-------- Anemina_arcaeformis        
              |           |                                    
              |           +-------- Cristaria_plicata          
              |  +--------+                                    
              |  |        |     +-- Sinanodonta_woodiana       
              |  |        |  +--+                              
              |  |        |  |  +-- Sinanodonta_elliptica      
              |  |        +--+                                 
              +--+           |  +-- Sinanodonta_lucida         
                 |           +--+                              
                 |              +-- Sinanodonta_angula         
                 |                                             
                 |  +-------------- Anodonta_cygnea            
                 |  |                                          
                 +--+  +----------- Anodonta_anatina           
                    |  |                                       
                    +--+  +-------- Strophitus_undulatus       
                       |  |                                    
                       +--+  +----- Platynaias_compressa       
                          |  |                                 
                          +--+----- Alasmidonta_marginata      
                             |                                 
                             |  +-- Pyganodon_cataracta        
                             +--+                              
                                +-- Pyganodon_grandis          

Jeratthitikul et al., 2022 [Publ 4260], fig. S1, 1

iT tT bi riS oT roT roS iD
9 9 7 4 67 57 2 19.0
                          +-- Chamberlainia_somsakpanhai
                    +-----+                             
                    |     +-- Sinohyriopsis_cumingii    
  +-----------------+                                   
  |                 |  +----- Lamprotula_leaii          
  |                 +--+                                
  |                    |  +-- Potomida_littoralis       
  |                    +--+                             
  |                       +-- Pronodularia_japanensis   
  |                                                     
  |                 +-------- Amblema_plicata **           
  |              +--+                                   
  |              |  |  +----- Ortmanniana_ligamentina **   
  |              |  +--+                                
  |              |     |  +-- Cambarunio_iris **           
  |              |     +--+                             
--+  +-----------+        +-- Lampsilis_cardium **         
  |  |           |                                      
  |  |           |        +-- Quadrula_quadrula **         
  |  |           |  +-----+                             
  |  |           |  |     +-- Quadrula_verrucosa **        
  |  |           +--+                                   
  |  |              |  +----- Elliptio_complanata **       
  |  |              +--+                                
  |  |                 |  +-- Pleurobema_sintoxia **       
  |  |                 +--+                             
  |  |                    +-- Eurynia_dilatata **          
  |  |                                                  
  |  |                 +----- Unio_tumidus              
  +--+     +-----------+                                
     |     |           |  +-- Unio_pictorum             
     |     |           +--+                             
     |  +--+              +-- Unio_crassus              
     |  |  |                                            
     |  |  |  +-------------- Aculamprotula_tortuosa    
     |  |  |  |                                         
     |  |  +--+           +-- Nodularia_douglasiae      
     |  |     |  +--------+                             
     |  |     |  |        +-- Nodularia_nuxpersicae     
     |  |     +--+                                      
     |  |        |  +-------- Schistodesmus_lampreyanus 
     |  |        +--+                                   
     |  |           |  +----- Cuneopsis_heudei          
     +--+           +--+                                
        |              |  +-- Tchangsinaia_piscicula    
        |              +--+                             
        |                 +-- Cuneopsis_rufescens       
        |                                               
        |                 +-- Lanceolaria_grayii        
        |  +--------------+                             
        |  |              +-- Lanceolaria_gladiola      
        |  |                                            
        |  |     +----------- Cristaria_plicata         
        |  |  +--+                                      
        +--+  |  |  +-------- Pletholophus_tenuis       
           |  |  +--+                                   
           |  |     |  +----- Sinanodonta_lucida        
           |  |     +--+                                
           |  |        |  +-- Sinanodonta_lauta         
           |  |        +--+                             
           +--+           +-- Sinanodonta_jourdyi       
              |                                         
              |     +-------- Pseudanodonta_complanata  
              |  +--+                                   
              |  |  |  +----- Anodonta_nuttalliana      
              |  |  +--+                                
              |  |     |  +-- Anodonta_anatina          
              |  |     +--+                             
              +--+        +-- Anodonta_cygnea           
                 |                                      
                 |        +-- Pyganodon_grandis         
                 |  +-----+                             
                 |  |     +-- Simpsonaias_ambigua       
                 +--+                                   
                    |  +----- Strophitus_undulatus      
                    +--+                                
                       |  +-- Alasmidonta_marginata     
                       +--+                             
                          +-- Lasmigona_costata         

Jeratthitikul et al., 2022 [Publ 4260], fig. S2, 2

iT tT bi riS oT roT roS iD
9 9 7 4 67 57 2 19.0
                          +-- Chamberlainia_somsakpanhai
                    +-----+                             
                    |     +-- Sinohyriopsis_cumingii    
  +-----------------+                                   
  |                 |  +----- Potomida_littoralis       
  |                 +--+                                
  |                    |  +-- Lamprotula_leaii          
  |                    +--+                             
  |                       +-- Pronodularia_japanensis   
  |                                                     
  |                 +-------- Amblema_plicata **           
  |              +--+                                   
  |              |  |  +----- Ortmanniana_ligamentina **   
  |              |  +--+                                
  |              |     |  +-- Lampsilis_cardium **         
  |              |     +--+                             
--+  +-----------+        +-- Cambarunio_iris **           
  |  |           |                                      
  |  |           |        +-- Quadrula_quadrula **         
  |  |           |  +-----+                             
  |  |           |  |     +-- Quadrula_verrucosa **        
  |  |           +--+                                   
  |  |              |  +----- Pleurobema_sintoxia **       
  |  |              +--+                                
  |  |                 |  +-- Eurynia_dilatata **          
  |  |                 +--+                             
  |  |                    +-- Elliptio_complanata **       
  |  |                                                  
  |  |                 +----- Unio_tumidus              
  +--+     +-----------+                                
     |     |           |  +-- Unio_pictorum             
     |     |           +--+                             
     |  +--+              +-- Unio_crassus              
     |  |  |                                            
     |  |  |  +-------------- Aculamprotula_tortuosa    
     |  |  |  |                                         
     |  |  +--+           +-- Nodularia_douglasiae      
     |  |     |  +--------+                             
     |  |     |  |        +-- Nodularia_nuxpersicae     
     |  |     +--+                                      
     |  |        |  +-------- Schistodesmus_lampreyanus 
     |  |        +--+                                   
     |  |           |  +----- Cuneopsis_heudei          
     +--+           +--+                                
        |              |  +-- Tchangsinaia_piscicula    
        |              +--+                             
        |                 +-- Cuneopsis_rufescens       
        |                                               
        |                 +-- Lanceolaria_grayii        
        |  +--------------+                             
        |  |              +-- Lanceolaria_gladiola      
        |  |                                            
        |  |     +----------- Cristaria_plicata         
        |  |  +--+                                      
        +--+  |  |  +-------- Pletholophus_tenuis       
           |  |  +--+                                   
           |  |     |  +----- Sinanodonta_lucida        
           |  |     +--+                                
           |  |        |  +-- Sinanodonta_jourdyi       
           |  |        +--+                             
           +--+           +-- Sinanodonta_lauta         
              |                                         
              |     +-------- Pseudanodonta_complanata  
              |  +--+                                   
              |  |  |  +----- Anodonta_anatina          
              |  |  +--+                                
              |  |     |  +-- Anodonta_nuttalliana      
              |  |     +--+                             
              +--+        +-- Anodonta_cygnea           
                 |                                      
                 |        +-- Alasmidonta_marginata     
                 |  +-----+                             
                 |  |     +-- Lasmigona_costata         
                 +--+                                   
                    |  +----- Pyganodon_grandis         
                    +--+                                
                       |  +-- Strophitus_undulatus      
                       +--+                             
                          +-- Simpsonaias_ambigua       

Smith, 2021 [Publ 4331], fig. 1

iT tT bi riS oT roT roS iD
9 9 7 4 11 7 3 14.0
                 +----- Sinohyriopsis_cumingii     
           +-----+                                 
           |     |  +-- Lens_contradens            
           |     +--+                              
  +--------+        +-- Lamprotula_leaii           
  |        |                                       
  |        |  +-------- Scabies_phaselus           
  |        +--+                                    
  |           |  +----- Cristaria_plicata          
  |           +--+                                 
  |              |  +-- Pyganodon_grandis          
--+              +--+                              
  |                 +-- Utterbackia_peninsularis   
  |                                                
  |                 +-- Uniomerus_tetralasmus **      
  |  +--------------+                              
  |  |              +-- Megalonaias_nervosa **        
  |  |                                             
  +--+              +-- Elliptio_crassidens **        
     |  +-----------+                              
     |  |           +-- Elliptio_complanata **        
     +--+                                          
        |  +----------- Amblema_plicata **            
        +--+                                       
           |  +-------- Potamilus_streckersoni **     
           +--+                                    
              |  +----- Venustaconcha_ellipsiformis **
              +--+                                 
                 |  +-- Lampsilis_siliquoidea **      
                 +--+                              
                    +-- Lampsilis_cardium **          

Keogh & Simons, 2019 [Publ 3935], fig. 4

iT tT bi riS oT roT roS iD
8 8 6 1 0 0 0 21.0
  +----------------- Lampsilis_sietmani **    
  |                                        
--+              +-- Lampsilis_floridensis ** 
  |  +-----------+                         
  |  |           +-- Lampsilis_teres **       
  +--+                                     
     |  +----------- Lampsilis_siliquoidea ** 
     +--+                                  
        |  +-------- Lampsilis_hydiana[2] **  
        +--+                               
           |  +----- Lampsilis_straminea[2] **
           +--+                            
              |  +-- Lampsilis_hydiana[1] **  
              +--+                         
                 +-- Lampsilis_straminea[1] **

Jones & Neves, 2010 [Publ 2646], fig. 16

iT tT bi riS oT roT roS iD
7 7 5 1 0 0 0 23.0
              +-- Epioblasma_triquetra **   
  +-----------+                          
  |           +-- Epioblasma_brevidens **   
--+                                      
  |  +----------- Epioblasma_rangiana **    
  +--+                                   
     |  +-------- Epioblasma_capsaeformis **
     +--+                                
        |  +----- Epioblasma_ahlstedti **   
        +--+                             
           |  +-- Epioblasma_aureola **     
           +--+                          
              +-- Epioblasma_walkeri **     

Jones & Neves, 2010 [Publ 2646], fig. 17

iT tT bi riS oT roT roS iD
7 7 1 1 0 0 0 23.0
  +----- Epioblasma_triquetra **   
  |                             
  +----- Epioblasma_brevidens **   
  |                             
--+  +-- Epioblasma_capsaeformis **
  |  |                          
  |  +-- Epioblasma_aureola **     
  |  |                          
  +--+-- Epioblasma_walkeri **     
     |                          
     +-- Epioblasma_rangiana **    
     |                          
     +-- Epioblasma_ahlstedti **   

Zhou et al., 2007 [Publ 2658], fig. 2

iT tT bi riS oT roT roS iD
7 7 5 1 14 13 2 22.0
              +-- Theliderma_intermedia **
  +-----------+                        
  |           +-- Theliderma_sparsa **    
--+                                    
  |  +----------- Quadrula_quadrula **    
  +--+                                 
     |  +-------- Pustulosa_infucata **   
     +--+                              
        |  +----- Quadrula_verrucosa **   
        +--+                           
           |  +-- Pustulosa_kieneriana ** 
           +--+                        
              +-- Pustulosa_succissa **   

Buhay et al., 2002 [Publ 2661], fig. 1

iT tT bi riS oT roT roS iD
7 7 3 1 0 0 0 21.0
  +----------- Lampsilis_siliquoidea[1] **
  |                                    
  +----------- Lampsilis_siliquoidea[2] **
--+                                    
  |  +-------- Epioblasma_brevidens **    
  |  |                                 
  +--+  +----- Epioblasma_triquetra **    
     |  |                              
     +--+  +-- Epioblasma_capsaeformis ** 
        |  |                           
        +--+-- Epioblasma_walkeri[1] **   
           |                           
           +-- Epioblasma_walkeri[2] **   

Amaro et al., 2017 [Publ 3791], fig. 2

iT tT bi riS oT roT roS iD
7 9 7 3 11 7 2 19.0
              +-- Lasmigona_costata          
        +-----+                              
        |     +-- Platynaias_compressa       
  +-----+                                    
  |     |  +----- Pyganodon_grandis          
  |     +--+                                 
  |        |  +-- Utterbackia_peninsularis   
  |        +--+                              
  |           +-- Utterbackia_imbecillis     
--+                                          
  |        +----- Fusconaia_flava **            
  |  +-----+                                 
  |  |     |  +-- Pronodularia_japanensis    
  |  |     +--+                              
  |  |        +-- Sinosolenaia_carinata      
  +--+                                       
     |     +----- Toxolasma_parvum **           
     |  +--+                                 
     |  |  |  +-- Venustaconcha_ellipsiformis **
     |  |  +--+                              
     +--+     +-- Lampsilis_siliquoidea **      
        |                                    
        |  +----- Quadrula_quadrula[2] **       
        +--+                                 
           |  +-- Quadrula_quadrula[1] **       
           +--+                              
              +-- Quadrula_nobilis **           

Bolotov et al., 2018 [Publ 3826], fig. 2

iT tT bi riS oT roT roS iD
7 13 0 4 78 78 3 24.0
                             +-- Ensidens_telus         
                             |                          
                             +-- Ensidens_ingallsianus  
                             |                          
                             +-- Ensidens_jaculus       
                             |                          
                             +-- Hyriopsis_desowitzi    
                          +--+                          
                          |  +-- Hyriopsis_myersiana    
                          |  |                          
                          |  +-- Hyriopsis_khoratensis  
                          |  |                          
                          |  +-- Ensidens_spiculus      
                          |  |                          
                          |  +-- Ensidens_sagittarius   
                          |                             
  +-----------------------+  +-- Lens_eximius           
  |                       |  |                          
  |                       |  +-- Physunio_modelli[1]    
  |                       |  |                          
  |                       |  +-- Yaukthwa_nesemanni     
  |                       |  |                          
  |                       |  +-- Yaukthwa_panhai        
  |                       |  |                          
  |                       |  +-- Lens_comptus           
  |                       |  |                          
  |                       +--+-- Lens_novoselovi        
  |                          |                          
  |                          +-- Physunio_modelli[2]    
  |                          |                          
  |                          +-- Physunio               
  |                          |                          
  |                          +-- Yaukthwa_inlenensis    
  |                          |                          
  |                          +-- Lens_contradens        
  |                          |                          
  |                          +-- Lens_micropterus       
  |                                                     
  |                          +-- Lamprotula_leaii       
--+                          |                          
  |                          +-- Potomida_littoralis    
  |                          |                          
  |                          +-- Pronodularia_japanensis
  |                          |                          
  |                          +-- Gonidea_angulata       
  |                          |                          
  |                          +-- Leguminaia_wheatleyi   
  |                          |                          
  |                          +-- Microcondylaea_bonellii
  |                          |                          
  |  +-----------------------+-- Amblema_plicata **        
  |  |                       |                          
  |  |                       +-- Ortmanniana_ligamentina **
  |  |                       |                          
  |  |                       +-- Lampsilis_cardium **      
  |  |                       |                          
  |  |                       +-- Cambarunio_iris **        
  |  |                       |                          
  |  |                       +-- Elliptio_complanata **    
  |  |                       |                          
  |  |                       +-- Pleurobema_sintoxia **    
  |  |                       |                          
  +--+                       +-- Quadrula_quadrula **      
     |                                                  
     |                    +----- Unio_tumidus           
     |                 +--+                             
     |                 |  |  +-- Unio_crassus           
     |                 |  +--+                          
     |  +--------------+     +-- Unio_pictorum          
     |  |              |                                
     |  |              |  +----- Nodularia_douglasiae[1]
     |  |              +--+                             
     |  |                 |  +-- Nodularia_nuxpersicae  
     |  |                 +--+                          
     +--+                    +-- Nodularia_douglasiae[2]
        |                                               
        |                    +-- Lanceolaria_gladiola   
        |  +-----------------+                          
        |  |                 +-- Lanceolaria_grayii     
        |  |                                            
        |  |           +-------- Cristaria_plicata      
        +--+  +--------+                                
           |  |        |  +----- Pletholophus_tenuis    
           |  |        +--+                             
           |  |           |  +-- Sinanodonta_jourdyi    
           +--+           +--+                          
              |              +-- Sinanodonta_lauta      
              |                                         
              |  +-------------- Anodonta_anatina       
              +--+                                      
                 |  +----------- Pyganodon_grandis      
                 +--+                                   
                    |  +-------- Simpsonaias_ambigua    
                    +--+                                
                       |  +----- Strophitus_undulatus   
                       +--+                             
                          |  +-- Alasmidonta_marginata  
                          +--+                          
                             +-- Lasmigona_costata      

Bolotov et al., 2020 [Publ 4084], fig. 1

iT tT bi riS oT roT roS iD
7 7 0 4 29 25 2 18.0
  +-- Amblema_plicata **        
  |                          
  +-- Ortmanniana_ligamentina **
  |                          
  +-- Lampsilis_cardium **      
  |                          
--+-- Cambarunio_iris **        
  |                          
  +-- Elliptio_complanata **    
  |                          
  +-- Pleurobema_sintoxia **    
  |                          
  +-- Quadrula_quadrula **      

Chen et al., 2020 [Publ 4170], fig. 1

iT tT bi riS oT roT roS iD
7 16 14 2 27 26 2 27.0
                       +----- Prolasmidonta_heterodon   
                    +--+                                
                    |  |  +-- Alasmidonta_varicosa      
                    |  +--+                             
                    |     +-- Anemina_euscaphys         
  +-----------------+                                   
  |                 |     +-- Lamprotula_caveata        
  |                 |  +--+                             
  |                 |  |  +-- Chamberlainia_somsakpanhai
  |                 +--+                                
  |                    |  +-- Pseudodon_vondembuschianus
  |                    +--+                             
  |                       +-- Pilsbryoconcha_exilis     
  |                                                     
  |                 +-------- Schistodesmus_lampreyanus 
  |           +-----+                                   
  |           |     |  +----- Lepidodesma_languilati    
  |           |     +--+                                
--+           |        |  +-- Aculamprotula_coreana     
  |           |        +--+                             
  |  +--------+           +-- Aculamprotula_tortuosa    
  |  |        |                                         
  |  |        |        +----- Platynaias_compressa      
  |  |        |  +-----+                                
  |  |        |  |     |  +-- Utterbackia_imbecillis    
  |  |        |  |     +--+                             
  |  |        +--+        +-- Pyganodon_grandis         
  |  |           |                                      
  |  |           |  +-------- Sinanodonta_lucida        
  |  |           +--+                                   
  |  |              |  +----- Cristaria_plicata         
  |  |              +--+                                
  +--+                 |  +-- Anodonta_cygnea           
     |                 +--+                             
     |                    +-- Anodonta_anatina          
     |                                                  
     |     +----------------- Gibbosula_crassa          
     |  +--+                                            
     |  |  |  +-------------- Quadrula_quadrula **         
     |  |  +--+                                         
     |  |     |  +----------- Toxolasma_parvum **          
     |  |     +--+                                      
     |  |        |  +-------- Lampsilis_ornata **          
     |  |        +--+                                   
     |  |           |  +----- Potamilus_leptodon **        
     +--+           +--+                                
        |              |  +-- Lampsilis_siliquoidea **     
        |              +--+                             
        |                 +-- Lampsilis_powellii **        
        |                                               
        |                 +-- Utterbackia_peninsularis  
        |  +--------------+                             
        |  |              +-- Potamilus_alatus **          
        +--+                                            
           |  +-------------- Lamprotula_leaii          
           |  |                                         
           +--+           +-- Sinohyriopsis_cumingii    
              |  +--------+                             
              |  |        +-- Sinohyriopsis_schlegelii  
              +--+                                      
                 |  +-------- Parvasolenaia_rivularis   
                 +--+                                   
                    |  +----- Sinosolenaia_carinata     
                    +--+                                
                       |  +-- Sinosolenaia_iridinea     
                       +--+                             
                          +-- Sinosolenaia_oleivora     

Pavan-Kumar et al., 2022 [Publ 4263], fig. 4

iT tT bi riS oT roT roS iD
7 7 5 2 29 27 3 10.0
                 +-------------- Quadrula_quadrula **          
                 |                                          
  +--------------+  +----------- Toxolasma_parvum **           
  |              |  |                                       
  |              +--+  +-------- Potamilus_leptodon **         
  |                 |  |                                    
  |                 +--+     +-- Lampsilis_cardium **          
  |                    |  +--+                              
  |                    |  |  +-- Lampsilis_ornata **           
  |                    +--+                                 
  |                       |  +-- Lampsilis_siliquoidea **      
  |                       +--+                              
  |                          +-- Venustaconcha_ellipsiformis **
  |                                                         
  |                       +----- Chamberlainia_somsakpanhai 
--+           +-----------+                                 
  |           |           |  +-- Sinohyriopsis_cumingii     
  |           |           +--+                              
  |           |              +-- Sinohyriopsis_schlegelii   
  |  +--------+                                             
  |  |        |              +-- Lamprotula_leaii           
  |  |        |  +-----------+                              
  |  |        |  |           +-- Potomida_littoralis        
  |  |        |  |                                          
  |  |        +--+           +-- Pseudodon_vondembuschianus 
  |  |           |  +--------+                              
  |  |           |  |        +-- Pilsbryoconcha_exilis      
  |  |           +--+                                       
  |  |              |  +-------- Parvasolenaia_rivularis    
  +--+              +--+                                    
     |                 |  +----- Microcondylaea_bonellii    
     |                 +--+                                 
     |                    |  +-- Sinosolenaia_carinata      
     |                    +--+                              
     |                       +-- Sinosolenaia_oleivora      
     |                                                      
     |  +----------------------- Lamellidens_marginalis     
     |  |                                                   
     |  |  +-------------------- Lepidodesma_languilati     
     +--+  |                                                
        |  |                 +-- Aculamprotula_tientsinensis
        |  |        +--------+                              
        |  |        |        +-- Aculamprotula_tortuosa     
        +--+  +-----+                                       
           |  |     |  +-------- Unio_pictorum              
           |  |     +--+                                    
           |  |        |  +----- Schistodesmus_lampreyanus  
           |  |        +--+                                 
           |  |           |  +-- Nodularia_douglasiae       
           +--+           +--+                              
              |              +-- Tchangsinaia_piscicula     
              |                                             
              |              +-- Lanceolaria_lanceolata     
              |  +-----------+                              
              |  |           +-- Lanceolaria_gladiola       
              +--+                                          
                 |  +----------- Acuticosta_chinensis       
                 |  |                                       
                 +--+        +-- Anodonta_anatina           
                    |  +-----+                              
                    |  |     +-- Pyganodon_grandis          
                    +--+                                    
                       |  +----- Cristaria_plicata          
                       +--+                                 
                          |  +-- Sinanodonta_lucida         
                          +--+                              
                             +-- Anemina_euscaphys          

Pavan-Kumar et al., 2022 [Publ 4263], fig. S5

iT tT bi riS oT roT roS iD
7 7 5 2 32 30 3 10.0
     +-------------------- Lamellidens_marginalis     
     |                                                
     |                 +-- Anodonta_anatina           
     |        +--------+                              
     |        |        +-- Pyganodon_grandis          
     |  +-----+                                       
  +--+  |     |  +-------- Cristaria_plicata          
  |  |  |     |  |                                    
  |  |  |     +--+     +-- Anemina_arcaeformis        
  |  |  |        |  +--+                              
  |  |  |        |  |  +-- Anemina_euscaphys          
  |  |  |        +--+                                 
  |  |  |           |  +-- Sinanodonta_lucida         
  |  +--+           +--+                              
  |     |              +-- Sinanodonta_woodiana       
  |     |                                             
  |     |           +----- Acuticosta_chinensis       
  |     |  +--------+                                 
  |     |  |        |  +-- Lanceolaria_lanceolata     
  |     |  |        +--+                              
  |     |  |           +-- Lanceolaria_gladiola       
  |     +--+                                          
  |        |        +----- Lepidodesma_languilati     
  |        |  +-----+                                 
  |        |  |     |  +-- Aculamprotula_tientsinensis
  |        |  |     +--+                              
--+        +--+        +-- Aculamprotula_tortuosa     
  |           |                                       
  |           |  +-------- Unio_pictorum              
  |           +--+                                    
  |              |  +----- Schistodesmus_lampreyanus  
  |              +--+                                 
  |                 |  +-- Nodularia_douglasiae       
  |                 +--+                              
  |                    +-- Tchangsinaia_piscicula     
  |                                                   
  |        +-------------- Quadrula_quadrula **          
  |        |                                          
  |     +--+  +----------- Toxolasma_parvum **           
  |     |  |  |                                       
  |     |  +--+  +-------- Potamilus_leptodon **         
  |     |     |  |                                    
  |     |     +--+     +-- Lampsilis_cardium **          
  |     |        |  +--+                              
  |     |        |  |  +-- Lampsilis_ornata **           
  |     |        +--+                                 
  |     |           |  +-- Lampsilis_siliquoidea **      
  +-----+           +--+                              
        |              +-- Venustaconcha_ellipsiformis **
        |                                             
        |           +----- Chamberlainia_somsakpanhai 
        |  +--------+                                 
        |  |        |  +-- Sinohyriopsis_schlegelii   
        |  |        +--+                              
        |  |           +-- Sinohyriopsis_cumingii     
        |  |                                          
        |  |           +-- Sinosolenaia_carinata      
        +--+        +--+                              
           |        |  +-- Sinosolenaia_oleivora      
           |  +-----+                                 
           |  |     |  +-- Parvasolenaia_rivularis    
           |  |     +--+                              
           |  |        +-- Microcondylaea_bonellii    
           +--+                                       
              |        +-- Pseudodon_vondembuschianus 
              |  +-----+                              
              |  |     +-- Pilsbryoconcha_exilis      
              +--+                                    
                 |  +----- Potomida_littoralis        
                 +--+                                 
                    |  +-- Lamprotula_leaii           
                    +--+                              
                       +-- Pronodularia_japanensis    

Walker et al., 2006 [Publ 2666], fig. 2

iT tT bi riS oT roT roS iD
6 7 5 2 9 6 2 21.0
  +-------------------- Pronodularia_japanensis   
  |                                               
  |                 +-- Gonidea_angulata          
--+  +--------------+                             
  |  |              +-- Pseudodon_vondembuschianus
  |  |                                            
  +--+              +-- Fusconaia_flava **           
     |  +-----------+                             
     |  |           +-- Cumberlandia_monodonta    
     +--+                                         
        |  +----------- Cyrtonaias_tampicoensis **   
        +--+                                      
           |  +-------- Potamilus_purpuratus **      
           +--+                                   
              |  +----- Ligumia_recta **             
              +--+                                
                 |  +-- Ortmanniana_ligamentina **   
                 +--+                             
                    +-- Lampsilis_teres **           

Walker et al., 2006 [Publ 2666], fig. 3

iT tT bi riS oT roT roS iD
6 6 4 2 9 6 2 20.0
  +-------------------- Inversidens            
  |                                            
  |              +----- Anodonta               
--+  +-----------+                             
  |  |           |  +-- Pyganodon_fragilis     
  |  |           +--+                          
  +--+              +-- Pyganodon_grandis      
     |                                         
     |  +-------------- Fusconaia_flava **        
     +--+                                      
        |  +----------- Potamilus_purpuratus **   
        +--+                                   
           |  +-------- Cyrtonaias_tampicoensis **
           +--+                                
              |  +----- Ligumia_recta **          
              +--+                             
                 |  +-- Ortmanniana_ligamentina **
                 +--+                          
                    +-- Lampsilis_teres **        

Walker et al., 2006 [Publ 2666], fig. 4

iT tT bi riS oT roT roS iD
6 6 4 2 9 6 2 20.0
  +-------------------- Inversidens            
  |                                            
  |              +----- Anodonta               
--+  +-----------+                             
  |  |           |  +-- Pyganodon_fragilis     
  |  |           +--+                          
  +--+              +-- Pyganodon_grandis      
     |                                         
     |  +-------------- Fusconaia_flava **        
     +--+                                      
        |  +----------- Cyrtonaias_tampicoensis **
        +--+                                   
           |  +-------- Potamilus_purpuratus **   
           +--+                                
              |  +----- Ligumia_recta **          
              +--+                             
                 |  +-- Ortmanniana_ligamentina **
                 +--+                          
                    +-- Lampsilis_teres **        

Walker et al., 2006 [Publ 2666], fig. 6

iT tT bi riS oT roT roS iD
6 6 3 2 32 4 2 20.0
              +-- Gonidea_angulata       
  +-----------+                          
  |           +-- Sundadontina_tumida    
--+                                      
  |  +----------- Cyrtonaias_tampicoensis **
  |  |                                   
  +--+----------- Fusconaia_flava **        
     |                                   
     |  +-------- Potamilus_purpuratus **   
     +--+                                
        |  +----- Lampsilis_teres **        
        +--+                             
           |  +-- Ligumia_recta **          
           +--+                          
              +-- Ortmanniana_ligamentina **

Walker et al., 2006 [Publ 2666], fig. 6-M

iT tT bi riS oT roT roS iD
6 6 4 2 9 4 2 20.0
                    +-- Gonidea_angulata       
  +-----------------+                          
  |                 +-- Sundadontina_tumida    
  |                                            
--+                 +-- Pyganodon_fragilis     
  |  +--------------+                          
  |  |              +-- Pyganodon_grandis      
  +--+                                         
     |  +-------------- Fusconaia_flava **        
     +--+                                      
        |  +----------- Potamilus_purpuratus **   
        +--+                                   
           |  +-------- Cyrtonaias_tampicoensis **
           +--+                                
              |  +----- Ligumia_recta **          
              +--+                             
                 |  +-- Lampsilis_teres **        
                 +--+                          
                    +-- Ortmanniana_ligamentina **

Hoeh et al., 2002 [Publ 2673], fig. 1-F

iT tT bi riS oT roT roS iD
6 6 4 2 3 2 1 20.0
                 +-- Pyganodon_fragilis     
  +--------------+                          
  |              +-- Pyganodon_grandis      
--+                                         
  |  +-------------- Fusconaia_flava **        
  +--+                                      
     |  +----------- Cyrtonaias_tampicoensis **
     +--+                                   
        |  +-------- Ortmanniana_ligamentina **
        +--+                                
           |  +----- Lampsilis_teres **        
           +--+                             
              |  +-- Potamilus_purpuratus **   
              +--+                          
                 +-- Ligumia_recta **          

Hoeh et al., 2002 [Publ 2673], fig. 1-M

iT tT bi riS oT roT roS iD
6 6 4 2 3 2 1 20.0
                 +-- Pyganodon_fragilis     
  +--------------+                          
  |              +-- Pyganodon_grandis      
--+                                         
  |  +-------------- Fusconaia_flava **        
  +--+                                      
     |  +----------- Potamilus_purpuratus **   
     +--+                                   
        |  +-------- Cyrtonaias_tampicoensis **
        +--+                                
           |  +----- Ligumia_recta **          
           +--+                             
              |  +-- Lampsilis_teres **        
              +--+                          
                 +-- Ortmanniana_ligamentina **

Jones et al., 2006 [Publ 2681], fig. 2

iT tT bi riS oT roT roS iD
6 6 4 1 0 0 0 21.0
           +-- Epioblasma_triquetra **      
  +--------+                             
  |        +-- Epioblasma_brevidens **      
--+                                      
  |  +-------- Epioblasma_capsaeformis[2] **
  +--+                                   
     |  +----- Epioblasma_capsaeformis[1] **
     +--+                                
        |  +-- Epioblasma_walkeri **        
        +--+                             
           +-- Epioblasma_rangiana **       

Lane et al., 2016 [Publ 3735], fig. 3

iT tT bi riS oT roT roS iD
6 6 1 1 0 0 0 20.0
  +----- Paetulunio_fabalis **         
  |                                 
  +----- Epioblasma_capsaeformis **    
--+                                 
  |  +-- Venustaconcha_trabalis[1] **  
  |  |                              
  |  +-- Venustaconcha_trabalis[2] **  
  +--+                              
     +-- Venustaconcha_ellipsiformis **
     |                              
     +-- Venustaconcha_pleasii **      

Wen et al., 2017 [Publ 3795], fig. 3

iT tT bi riS oT roT roS iD
6 6 4 2 10 8 2 11.0
     +----------- Quadrula_quadrula **          
     |                                       
  +--+  +-------- Toxolasma_parvum **           
  |  |  |                                    
  |  +--+     +-- Lampsilis_ornata **           
  |     |  +--+                              
  |     |  |  +-- Venustaconcha_ellipsiformis **
  |     +--+                                 
--+        |  +-- Potamilus_leptodon **         
  |        +--+                              
  |           +-- Potamilus_alatus **           
  |                                          
  |  +----------- Lamprotula_leaii           
  |  |                                       
  |  |     +----- Anodonta_anatina           
  +--+  +--+                                 
     |  |  |  +-- Sinanodonta_woodiana       
     |  |  +--+                              
     |  |     +-- Cristaria_plicata          
     +--+                                    
        |     +-- Aculamprotula_coreana      
        |  +--+                              
        |  |  +-- Aculamprotula_tortuosa     
        +--+                                 
           |  +-- Lepidodesma_languilati     
           +--+                              
              +-- Unio_pictorum              

Huang et al., 2018 [Publ 3828], fig. 6

iT tT bi riS oT roT roS iD
6 6 4 2 37 27 2 11.0
           +----------- Quadrula_quadrula **          
           |                                       
     +-----+  +-------- Toxolasma_parvum **           
     |     |  |                                    
     |     +--+     +-- Lampsilis_ornata **           
     |        |  +--+                              
     |        |  |  +-- Venustaconcha_ellipsiformis **
     |        +--+                                 
     |           |  +-- Potamilus_leptodon **         
  +--+           +--+                              
  |  |              +-- Potamilus_alatus **           
  |  |                                             
  |  |              +-- Sinohyriopsis_cumingii     
  |  |  +-----------+                              
  |  |  |           +-- Sinohyriopsis_schlegelii   
  |  |  |                                          
  |  +--+           +-- Sinosolenaia_oleivora      
  |     |  +--------+                              
  |     |  |        +-- Sinosolenaia_carinata      
  |     +--+                                       
  |        |  +-------- Potomida_littoralis        
  |        +--+                                    
  |           |  +----- Pronodularia_japanensis    
  |           +--+                                 
  |              |  +-- Lamprotula_caveata         
--+              +--+                              
  |                 +-- Lamprotula_leaii           
  |                                                
  |              +----- Aculamprotula_tortuosa     
  |     +--------+                                 
  |     |        |  +-- Aculamprotula_coreana      
  |     |        +--+                              
  |     |           +-- Aculamprotula_tientsinensis
  |  +--+                                          
  |  |  |           +-- Tchangsinaia_piscicula     
  |  |  |  +--------+                              
  |  |  |  |        +-- Nodularia_douglasiae       
  |  |  +--+                                       
  |  |     |  +-------- Unio_tumidus               
  |  |     +--+                                    
  |  |        |  +----- Unio_crassus               
  |  |        +--+                                 
  +--+           |  +-- Unio_delphinus             
     |           +--+                              
     |              +-- Unio_pictorum              
     |                                             
     |     +----------- Anodonta_anatina           
     |  +--+                                       
     |  |  |  +-------- Platynaias_compressa       
     |  |  +--+                                    
     |  |     |  +----- Pyganodon_grandis          
     |  |     +--+                                 
     +--+        |  +-- Utterbackia_imbecillis     
        |        +--+                              
        |           +-- Utterbackia_peninsularis   
        |                                          
        |     +-------- Cristaria_plicata          
        |     |                                    
        +-----+     +-- Anemina_arcaeformis        
              |  +--+                              
              |  |  +-- Anemina_euscaphys          
              +--+                                 
                 |  +-- Sinanodonta_woodiana       
                 +--+                              
                    +-- Sinanodonta_lucida         

Wang et al., 2018 [Publ 3846], fig. 2

iT tT bi riS oT roT roS iD
6 6 4 2 26 26 2 11.0
              +----------- Quadrula_quadrula **          
              |                                       
           +--+  +-------- Toxolasma_parvum **           
           |  |  |                                    
           |  +--+     +-- Venustaconcha_ellipsiformis **
           |     |  +--+                              
           |     |  |  +-- Lampsilis_ornata **           
           |     +--+                                 
           |        |  +-- Potamilus_leptodon **         
  +--------+        +--+                              
  |        |           +-- Potamilus_alatus **           
  |        |                                          
  |        |           +-- Sinohyriopsis_cumingii     
  |        |  +--------+                              
  |        |  |        +-- Sinohyriopsis_schlegelii   
  |        |  |                                       
  |        +--+     +----- Potomida_littoralis        
  |           |  +--+                                 
  |           |  |  |  +-- Lamprotula_caveata         
  |           |  |  +--+                              
  |           +--+     +-- Lamprotula_leaii           
  |              |                                    
  |              |  +----- Ptychorhynchus_pfisteri    
  |              +--+                                 
--+                 |  +-- Sinosolenaia_carinata      
  |                 +--+                              
  |                    +-- Sinosolenaia_oleivora      
  |                                                   
  |                 +----- Anodonta_anatina           
  |           +-----+                                 
  |           |     |  +-- Pyganodon_grandis          
  |           |     +--+                              
  |  +--------+        +-- Platynaias_compressa       
  |  |        |                                       
  |  |        |  +-------- Cristaria_plicata          
  |  |        |  |                                    
  |  |        +--+     +-- Anemina_euscaphys          
  |  |           |  +--+                              
  |  |           |  |  +-- Anemina_arcaeformis        
  |  |           +--+                                 
  +--+              |  +-- Sinanodonta_woodiana       
     |              +--+                              
     |                 +-- Sinanodonta_lucida         
     |                                                
     |                 +-- Lanceolaria_grayii         
     |  +--------------+                              
     |  |              +-- Lanceolaria_lanceolata     
     |  |                                             
     +--+  +-------------- Aculamprotula_tientsinensis
        |  |                                          
        |  |        +----- Nodularia_douglasiae       
        +--+  +-----+                                 
           |  |     |  +-- Tchangsinaia_piscicula     
           |  |     +--+                              
           +--+        +-- Cuneopsis_heudei           
              |                                       
              |  +-------- Unio_tumidus               
              +--+                                    
                 |  +----- Unio_crassus               
                 +--+                                 
                    |  +-- Unio_pictorum              
                    +--+                              
                       +-- Unio_delphinus             

Wu et al., 2019 [Publ 3864], fig. 3

iT tT bi riS oT roT roS iD
6 6 4 2 41 35 2 11.0
                    +----------- Quadrula_quadrula **          
                    |                                       
  +-----------------+  +-------- Toxolasma_parvum **           
  |                 |  |                                    
  |                 +--+     +-- Lampsilis_ornata **           
  |                    |  +--+                              
  |                    |  |  +-- Venustaconcha_ellipsiformis **
  |                    +--+                                 
  |                       |  +-- Potamilus_leptodon **         
  |                       +--+                              
  |                          +-- Potamilus_alatus **           
  |                                                         
  |                          +-- Sinohyriopsis_cumingii     
  |                 +--------+                              
--+                 |        +-- Sinohyriopsis_schlegelii   
  |                 |                                       
  |  +--------------+     +----- Potomida_littoralis        
  |  |              |  +--+                                 
  |  |              |  |  |  +-- Lamprotula_leaii           
  |  |              |  |  +--+                              
  |  |              +--+     +-- Pronodularia_japanensis    
  |  |                 |                                    
  |  |                 |  +----- Ptychorhynchus_pfisteri    
  |  |                 +--+                                 
  |  |                    |  +-- Sinosolenaia_oleivora      
  |  |                    +--+                              
  |  |                       +-- Sinosolenaia_carinata      
  +--+                                                      
     |  +----------------------- Lepidodesma_languilati     
     |  |                                                   
     |  |              +-------- Aculamprotula_tortuosa     
     |  |     +--------+                                    
     |  |     |        |  +----- Aculamprotula_scripta      
     |  |     |        +--+                                 
     |  |     |           |  +-- Aculamprotula_tientsinensis
     |  |     |           +--+                              
     |  |     |              +-- Aculamprotula_coreana      
     |  |  +--+                                             
     +--+  |  |        +-------- Unio_tumidus               
        |  |  |  +-----+                                    
        |  |  |  |     |  +----- Unio_crassus               
        |  |  |  |     +--+                                 
        |  |  |  |        |  +-- Unio_delphinus             
        |  |  +--+        +--+                              
        |  |     |           +-- Unio_pictorum              
        |  |     |                                          
        |  |     |  +----------- Pseudocuneopsis_capitata   
        |  |     +--+                                       
        |  |        |  +-------- Nodularia_douglasiae       
        +--+        +--+                                    
           |           |  +----- Tchangsinaia_piscicula     
           |           +--+                                 
           |              |  +-- Cuneopsis_heudei           
           |              +--+                              
           |                 +-- Schistodesmus_lampreyanus  
           |                                                
           |              +----- Acuticosta_chinensis       
           |  +-----------+                                 
           |  |           |  +-- Lanceolaria_grayii         
           |  |           +--+                              
           |  |              +-- Lanceolaria_lanceolata     
           |  |                                             
           +--+     +----------- Anodonta_anatina           
              |  +--+                                       
              |  |  |  +-------- Platynaias_compressa       
              |  |  +--+                                    
              |  |     |  +----- Pyganodon_grandis          
              |  |     +--+                                 
              +--+        |  +-- Utterbackia_peninsularis   
                 |        +--+                              
                 |           +-- Utterbackia_imbecillis     
                 |                                          
                 |     +-------- Cristaria_plicata          
                 |     |                                    
                 +-----+     +-- Anemina_arcaeformis        
                       |  +--+                              
                       |  |  +-- Anemina_euscaphys          
                       +--+                                 
                          |  +-- Sinanodonta_lucida         
                          +--+                              
                             +-- Sinanodonta_woodiana       

Wang et al., 2018 [Publ 3865], fig. 2

iT tT bi riS oT roT roS iD
6 6 4 2 26 26 2 11.0
           +----------- Quadrula_quadrula **          
           |                                       
        +--+  +-------- Toxolasma_parvum **           
        |  |  |                                    
        |  +--+     +-- Potamilus_leptodon **         
        |     |  +--+                              
        |     |  |  +-- Potamilus_alatus **           
        |     +--+                                 
        |        |  +-- Venustaconcha_ellipsiformis **
  +-----+        +--+                              
  |     |           +-- Lampsilis_ornata **           
  |     |                                          
  |     |           +-- Sinohyriopsis_schlegelii   
  |     |  +--------+                              
  |     |  |        +-- Sinohyriopsis_cumingii     
  |     |  |                                       
  |     +--+     +----- Ptychorhynchus_pfisteri    
  |        |  +--+                                 
  |        |  |  |  +-- Sinosolenaia_oleivora      
  |        |  |  +--+                              
  |        +--+     +-- Sinosolenaia_carinata      
  |           |                                    
  |           |  +----- Potomida_littoralis        
--+           +--+                                 
  |              |  +-- Lamprotula_caveata         
  |              +--+                              
  |                 +-- Lamprotula_leaii           
  |                                                
  |     +-------------- Aculamprotula_tientsinensis
  |     |                                          
  |  +--+           +-- Tchangsinaia_piscicula     
  |  |  |  +--------+                              
  |  |  |  |        +-- Nodularia_douglasiae       
  |  |  +--+                                       
  |  |     |  +-------- Unio_tumidus               
  |  |     +--+                                    
  |  |        |  +----- Unio_crassus               
  |  |        +--+                                 
  +--+           |  +-- Unio_pictorum              
     |           +--+                              
     |              +-- Unio_delphinus             
     |                                             
     |           +----- Acuticosta_chinensis       
     |  +--------+                                 
     |  |        |  +-- Lanceolaria_grayii         
     |  |        +--+                              
     |  |           +-- Lanceolaria_lanceolata     
     +--+                                          
        |        +----- Anodonta_anatina           
        |  +-----+                                 
        |  |     |  +-- Pyganodon_grandis          
        |  |     +--+                              
        +--+        +-- Platynaias_compressa       
           |                                       
           |  +-------- Cristaria_plicata          
           |  |                                    
           +--+     +-- Anemina_euscaphys          
              |  +--+                              
              |  |  +-- Anemina_arcaeformis        
              +--+                                 
                 |  +-- Sinanodonta_woodiana       
                 +--+                              
                    +-- Sinanodonta_lucida         

Froufe et al., 2019 [Publ 3936], fig. 3

iT tT bi riS oT roT roS iD
6 6 4 2 39 36 2 11.0
                    +----------- Quadrula_quadrula **          
                    |                                       
  +-----------------+  +-------- Toxolasma_parvum **           
  |                 |  |                                    
  |                 +--+     +-- Lampsilis_ornata **           
  |                    |  +--+                              
  |                    |  |  +-- Venustaconcha_ellipsiformis **
  |                    +--+                                 
  |                       |  +-- Potamilus_leptodon **         
  |                       +--+                              
  |                          +-- Potamilus_alatus **           
  |                                                         
  |                       +----- Chamberlainia_somsakpanhai 
  |           +-----------+                                 
  |           |           |  +-- Sinohyriopsis_cumingii     
  |           |           +--+                              
  |           |              +-- Sinohyriopsis_schlegelii   
--+           |                                             
  |  +--------+        +-------- Potomida_littoralis        
  |  |        |  +-----+                                    
  |  |        |  |     |  +----- Pronodularia_japanensis    
  |  |        |  |     +--+                                 
  |  |        |  |        |  +-- Lamprotula_leaii           
  |  |        |  |        +--+                              
  |  |        +--+           +-- Lamprotula_caveata         
  |  |           |                                          
  |  |           |           +-- Pilsbryoconcha_exilis      
  |  |           |  +--------+                              
  |  |           |  |        +-- Pseudodon_vondembuschianus 
  |  |           +--+                                       
  |  |              |  +-------- Ptychorhynchus_pfisteri    
  |  |              +--+                                    
  +--+                 |  +----- Microcondylaea_bonellii    
     |                 +--+                                 
     |                    |  +-- Sinosolenaia_carinata      
     |                    +--+                              
     |                       +-- Sinosolenaia_oleivora      
     |                                                      
     |  +----------------------- Lepidodesma_languilati     
     |  |                                                   
     |  |                 +----- Aculamprotula_tortuosa     
     |  |        +--------+                                 
     |  |        |        |  +-- Aculamprotula_tientsinensis
     |  |        |        +--+                              
     |  |        |           +-- Aculamprotula_coreana      
     |  |  +-----+                                          
     +--+  |     |        +----- Nodularia_douglasiae       
        |  |     |  +-----+                                 
        |  |     |  |     |  +-- Tchangsinaia_piscicula     
        |  |     |  |     +--+                              
        |  |     +--+        +-- Schistodesmus_lampreyanus  
        |  |        |                                       
        |  |        |  +-------- Unio_tumidus               
        |  |        +--+                                    
        |  |           |  +----- Unio_crassus               
        +--+           +--+                                 
           |              |  +-- Unio_pictorum              
           |              +--+                              
           |                 +-- Unio_delphinus             
           |                                                
           |                 +-- Lanceolaria_lanceolata     
           |  +--------------+                              
           |  |              +-- Lanceolaria_grayii         
           |  |                                             
           |  |        +-------- Cristaria_plicata          
           +--+        |                                    
              |  +-----+     +-- Anemina_arcaeformis        
              |  |     |  +--+                              
              |  |     |  |  +-- Anemina_euscaphys          
              |  |     +--+                                 
              +--+        |  +-- Sinanodonta_lucida         
                 |        +--+                              
                 |           +-- Sinanodonta_woodiana       
                 |                                          
                 |  +----------- Anodonta_anatina           
                 +--+                                       
                    |  +-------- Lasmigona_complanata       
                    +--+                                    
                       |  +----- Pyganodon_grandis          
                       +--+                                 
                          |  +-- Utterbackia_peninsularis   
                          +--+                              
                             +-- Utterbackia_imbecillis     

Froufe et al., 2019 [Publ 3936], fig. 6

iT tT bi riS oT roT roS iD
6 6 4 2 39 36 2 11.0
                    +----------- Quadrula_quadrula **          
                    |                                       
  +-----------------+  +-------- Toxolasma_parvum **           
  |                 |  |                                    
  |                 +--+     +-- Potamilus_leptodon **         
  |                    |  +--+                              
  |                    |  |  +-- Potamilus_alatus **           
  |                    +--+                                 
  |                       |  +-- Lampsilis_ornata **           
  |                       +--+                              
  |                          +-- Venustaconcha_ellipsiformis **
  |                                                         
  |                       +----- Chamberlainia_somsakpanhai 
  |           +-----------+                                 
  |           |           |  +-- Sinohyriopsis_cumingii     
  |           |           +--+                              
  |           |              +-- Sinohyriopsis_schlegelii   
--+           |                                             
  |  +--------+        +-------- Potomida_littoralis        
  |  |        |  +-----+                                    
  |  |        |  |     |  +----- Pronodularia_japanensis    
  |  |        |  |     +--+                                 
  |  |        |  |        |  +-- Lamprotula_leaii           
  |  |        |  |        +--+                              
  |  |        +--+           +-- Lamprotula_caveata         
  |  |           |                                          
  |  |           |           +-- Pilsbryoconcha_exilis      
  |  |           |  +--------+                              
  |  |           |  |        +-- Pseudodon_vondembuschianus 
  |  |           +--+                                       
  |  |              |  +-------- Ptychorhynchus_pfisteri    
  |  |              +--+                                    
  +--+                 |  +----- Microcondylaea_bonellii    
     |                 +--+                                 
     |                    |  +-- Sinosolenaia_carinata      
     |                    +--+                              
     |                       +-- Sinosolenaia_oleivora      
     |                                                      
     |  +----------------------- Lepidodesma_languilati     
     |  |                                                   
     |  |                 +----- Aculamprotula_tortuosa     
     |  |        +--------+                                 
     |  |        |        |  +-- Aculamprotula_tientsinensis
     |  |        |        +--+                              
     |  |        |           +-- Aculamprotula_coreana      
     |  |  +-----+                                          
     +--+  |     |        +----- Nodularia_douglasiae       
        |  |     |  +-----+                                 
        |  |     |  |     |  +-- Tchangsinaia_piscicula     
        |  |     |  |     +--+                              
        |  |     +--+        +-- Schistodesmus_lampreyanus  
        |  |        |                                       
        |  |        |  +-------- Unio_tumidus               
        |  |        +--+                                    
        |  |           |  +----- Unio_crassus               
        +--+           +--+                                 
           |              |  +-- Unio_pictorum              
           |              +--+                              
           |                 +-- Unio_delphinus             
           |                                                
           |                 +-- Lanceolaria_lanceolata     
           |  +--------------+                              
           |  |              +-- Lanceolaria_grayii         
           |  |                                             
           |  |     +----------- Anodonta_anatina           
           +--+  +--+                                       
              |  |  |  +-------- Lasmigona_complanata       
              |  |  +--+                                    
              |  |     |  +----- Pyganodon_grandis          
              |  |     +--+                                 
              +--+        |  +-- Utterbackia_peninsularis   
                 |        +--+                              
                 |           +-- Utterbackia_imbecillis     
                 |                                          
                 |     +-------- Cristaria_plicata          
                 |     |                                    
                 +-----+     +-- Sinanodonta_lucida         
                       |  +--+                              
                       |  |  +-- Sinanodonta_woodiana       
                       +--+                                 
                          |  +-- Anemina_arcaeformis        
                          +--+                              
                             +-- Anemina_euscaphys          

Wu et al., 2020 [Publ 4121], fig. 3

iT tT bi riS oT roT roS iD
6 6 4 2 44 36 2 11.0
                    +----------- Quadrula_quadrula **          
                    |                                       
  +-----------------+  +-------- Toxolasma_parvum **           
  |                 |  |                                    
  |                 +--+     +-- Venustaconcha_ellipsiformis **
  |                    |  +--+                              
  |                    |  |  +-- Lampsilis_ornata **           
  |                    +--+                                 
  |                       |  +-- Potamilus_alatus **           
  |                       +--+                              
  |                          +-- Potamilus_leptodon **         
  |                                                         
  |                          +-- Sinohyriopsis_schlegelii   
  |                 +--------+                              
--+                 |        +-- Sinohyriopsis_cumingii     
  |                 |                                       
  |  +--------------+     +----- Potomida_littoralis        
  |  |              |  +--+                                 
  |  |              |  |  |  +-- Pronodularia_japanensis    
  |  |              |  |  +--+                              
  |  |              +--+     +-- Lamprotula_leaii           
  |  |                 |                                    
  |  |                 |  +----- Ptychorhynchus_pfisteri    
  |  |                 +--+                                 
  |  |                    |  +-- Sinosolenaia_oleivora      
  |  |                    +--+                              
  |  |                       +-- Sinosolenaia_carinata      
  +--+                                                      
     |  +----------------------- Lepidodesma_languilati     
     |  |                                                   
     |  |                 +----- Acuticosta_chinensis       
     |  |     +-----------+                                 
     |  |     |           |  +-- Lanceolaria_lanceolata     
     |  |     |           +--+                              
     |  |     |              +-- Lanceolaria_grayii         
     |  |     |                                             
     |  |  +--+     +----------- Anodonta_anatina           
     |  |  |  |  +--+                                       
     |  |  |  |  |  |  +-------- Platynaias_compressa       
     +--+  |  |  |  +--+                                    
        |  |  |  |     |  +----- Pyganodon_grandis          
        |  |  |  |     +--+                                 
        |  |  +--+        |  +-- Utterbackia_peninsularis   
        |  |     |        +--+                              
        |  |     |           +-- Utterbackia_imbecillis     
        |  |     |                                          
        |  |     |     +-------- Cristaria_plicata          
        |  |     |     |                                    
        |  |     +-----+     +-- Sinanodonta_woodiana       
        |  |           |  +--+                              
        +--+           |  |  +-- Sinanodonta_lauta          
           |           +--+                                 
           |              |  +-- Anemina_arcaeformis        
           |              +--+                              
           |                 +-- Anemina_euscaphys          
           |                                                
           |           +-------- Aculamprotula_tortuosa     
           |           |                                    
           |  +--------+     +-- Aculamprotula_coreana      
           |  |        |  +--+                              
           |  |        |  |  +-- Aculamprotula_tientsinensis
           |  |        +--+                                 
           |  |           |  +-- Aculamprotula_polysticta   
           |  |           +--+                              
           +--+              +-- Aculamprotula_scripta      
              |                                             
              |        +-------- Unio_tumidus               
              |  +-----+                                    
              |  |     |  +----- Unio_crassus               
              |  |     +--+                                 
              |  |        |  +-- Unio_delphinus             
              +--+        +--+                              
                 |           +-- Unio_pictorum              
                 |                                          
                 |  +----------- Pseudocuneopsis_capitata   
                 +--+                                       
                    |  +-------- Nodularia_douglasiae       
                    +--+                                    
                       |  +----- Tchangsinaia_piscicula     
                       +--+                                 
                          |  +-- Cuneopsis_heudei           
                          +--+                              
                             +-- Schistodesmus_lampreyanus  

Zieritz et al., 2021 [Publ 4231], fig. 3-F, 5

iT tT bi riS oT roT roS iD
6 6 4 2 43 40 2 11.0
                    +----------- Quadrula_quadrula **          
                    |                                       
  +-----------------+  +-------- Toxolasma_parvum **           
  |                 |  |                                    
  |                 +--+     +-- Potamilus_alatus **           
  |                    |  +--+                              
  |                    |  |  +-- Potamilus_leptodon **         
  |                    +--+                                 
  |                       |  +-- Venustaconcha_ellipsiformis **
  |                       +--+                              
  |                          +-- Lampsilis_ornata **           
  |                                                         
  |                       +----- Chamberlainia_somsakpanhai 
  |        +--------------+                                 
  |        |              |  +-- Sinohyriopsis_cumingii     
  |        |              +--+                              
  |        |                 +-- Sinohyriopsis_schlegelii   
  |        |                                                
  |        |                 +-- Lens_contradens            
--+  +-----+              +--+                              
  |  |     |              |  +-- Physunio_superbus          
  |  |     |  +-----------+                                 
  |  |     |  |           |  +-- Hyriopsis_bialata          
  |  |     |  |           +--+                              
  |  |     |  |              +-- Rectidens_sumatrensis      
  |  |     |  |                                             
  |  |     +--+        +-------- Potomida_littoralis        
  |  |        |  +-----+                                    
  |  |        |  |     |  +----- Pronodularia_japanensis    
  |  |        |  |     +--+                                 
  |  |        |  |        |  +-- Lamprotula_caveata         
  |  |        |  |        +--+                              
  |  |        +--+           +-- Lamprotula_leaii           
  |  |           |                                          
  |  |           |           +-- Pilsbryoconcha_exilis      
  |  |           |  +--------+                              
  +--+           |  |        +-- Pseudodon_vondembuschianus 
     |           +--+                                       
     |              |  +-------- Ptychorhynchus_pfisteri    
     |              +--+                                    
     |                 |  +----- Microcondylaea_bonellii    
     |                 +--+                                 
     |                    |  +-- Sinosolenaia_recognita     
     |                    +--+                              
     |                       +-- Sinosolenaia_carinata      
     |                                                      
     |  +----------------------- Lepidodesma_languilati     
     |  |                                                   
     |  |                 +----- Aculamprotula_tortuosa     
     |  |        +--------+                                 
     |  |        |        |  +-- Aculamprotula_tientsinensis
     |  |        |        +--+                              
     |  |        |           +-- Aculamprotula_coreana      
     |  |  +-----+                                          
     +--+  |     |        +----- Nodularia_douglasiae       
        |  |     |  +-----+                                 
        |  |     |  |     |  +-- Schistodesmus_lampreyanus  
        |  |     |  |     +--+                              
        |  |     +--+        +-- Tchangsinaia_piscicula     
        |  |        |                                       
        |  |        |  +-------- Unio_tumidus               
        |  |        +--+                                    
        |  |           |  +----- Unio_crassus               
        +--+           +--+                                 
           |              |  +-- Unio_pictorum              
           |              +--+                              
           |                 +-- Unio_delphinus             
           |                                                
           |                 +-- Lanceolaria_grayii         
           |  +--------------+                              
           |  |              +-- Lanceolaria_lanceolata     
           |  |                                             
           |  |     +----------- Anodonta_anatina           
           +--+  +--+                                       
              |  |  |  +-------- Platynaias_compressa       
              |  |  +--+                                    
              |  |     |  +----- Pyganodon_grandis          
              |  |     +--+                                 
              +--+        |  +-- Utterbackia_imbecillis     
                 |        +--+                              
                 |           +-- Utterbackia_peninsularis   
                 |                                          
                 |     +-------- Cristaria_plicata          
                 |     |                                    
                 +-----+     +-- Sinanodonta_lucida         
                       |  +--+                              
                       |  |  +-- Sinanodonta_woodiana       
                       +--+                                 
                          |  +-- Anemina_arcaeformis        
                          +--+                              
                             +-- Anemina_euscaphys          

Mejia-Trujillo & Smith, 2022 [Publ 4274], fig. 2-M

iT tT bi riS oT roT roS iD
6 6 4 2 0 0 0 11.0
  +----------- Quadrula_quadrula **          
  |                                       
--+        +-- Potamilus_streckersoni **     
  |  +-----+                              
  |  |     +-- Potamilus_alatus **           
  +--+                                    
     |  +----- Venustaconcha_ellipsiformis **
     +--+                                 
        |  +-- Lampsilis_powellii **         
        +--+                              
           +-- Lampsilis_siliquoidea **      

Wu et al., 2023 [Publ 4339], fig. 5

iT tT bi riS oT roT roS iD
6 6 4 3 13 11 1 17.0
                 +-- Elliptio_complanata **     
        +--------+                           
        |        +-- Quadrula_quadrula **       
  +-----+                                    
  |     |  +-------- Obliquaria_reflexa **      
  |     +--+                                 
  |        |  +----- Truncilla_truncata **      
  |        +--+                              
  |           |  +-- Lampsilis_cardium **       
  |           +--+                           
--+              +-- Potamilus_alatus **        
  |                                          
  |              +-- Lepidodesma_languilati  
  |  +-----------+                           
  |  |           +-- Lepidodesma_aligera     
  |  |                                       
  |  |     +-------- Anemina_arcaeformis     
  +--+  +--+                                 
     |  |  |  +----- Cristaria_plicata       
     |  |  +--+                              
     |  |     |  +-- Sinanodonta_tumens      
     |  |     +--+                           
     +--+        +-- Sinanodonta_woodiana    
        |                                    
        |        +-- Platynaias              
        |  +-----+                           
        |  |     +-- Platynaias_compressa    
        +--+                                 
           |  +----- Anodonta_anatina        
           +--+                              
              |  +-- Pseudanodonta_complanata
              +--+                           
                 +-- Anodonta_cygnea         

Manendo et al., 2008 [Publ 2669], fig. 1

iT tT bi riS oT roT roS iD
5 5 3 3 8 8 1 19.0
              +-- Lampsilis_siliquoidea **   
        +-----+                           
        |     +-- Sagittunio_nasutus **      
  +-----+                                 
  |     |  +----- Amblema_plicata **         
  |     +--+                              
  |        |  +-- Pleuronaia_dolabelloides **
  |        +--+                           
  |           +-- Fusconaia_flava **         
--+                                       
  |        +----- Unio_crassus            
  |  +-----+                              
  |  |     |  +-- Unio_pictorum           
  |  |     +--+                           
  |  |        +-- Unio_tumidus            
  +--+                                    
     |        +-- Anodonta_cygnea         
     |  +-----+                           
     |  |     +-- Anodonta_anatina        
     +--+                                 
        |  +----- Pseudanodonta_complanata
        +--+                              
           |  +-- Platynaias_subviridis   
           +--+                           
              +-- Pyganodon_grandis       

Whelan et al., 2011 [Publ 3380], fig. 1 ML

iT tT bi riS oT roT roS iD
5 5 3 4 32 20 3 17.0
  +----------------- Gonidea_angulata       
  |                                         
  |           +----- Pronodularia_japanensis
--+  +--------+                             
  |  |        |  +-- Potomida_littoralis    
  |  |        +--+                          
  +--+           +-- Potomida_semirugata    
     |                                      
     |  +----------- Pleurobema_sintoxia **    
     +--+                                   
        |  +-------- Quadrula_quadrula **      
        +--+                                
           |  +----- Amblema_plicata **        
           +--+                             
              |  +-- Truncilla_truncata **     
              +--+                          
                 +-- Lampsilis_cardium **      

Whelan et al., 2011 [Publ 3380], fig. 1 MP

iT tT bi riS oT roT roS iD
5 5 2 4 32 20 3 16.0
  +-------------- Gonidea_angulata       
  |                                      
  |        +----- Pronodularia_japanensis
--+  +-----+                             
  |  |     |  +-- Potomida_littoralis    
  |  |     +--+                          
  +--+        +-- Potomida_semirugata    
     |                                   
     |  +-------- Quadrula_quadrula **      
     |  |                                
     +--+-------- Pleurobema_sintoxia **    
        |                                
        |  +----- Amblema_plicata **        
        +--+                             
           |  +-- Lampsilis_cardium **      
           +--+                          
              +-- Truncilla_truncata **     

Krebs et al., 2013 [Publ 3515], fig. 2A

iT tT bi riS oT roT roS iD
5 5 3 1 0 0 0 21.0
  +-------- Lampsilis_radiata **      
  |                                
--+     +-- Lampsilis_ovata **        
  |  +--+                          
  |  |  +-- Ortmanniana_ligamentina **
  +--+                             
     |  +-- Lampsilis_hydiana **      
     +--+                          
        +-- Lampsilis_straminea **    

Pfeiffer & Graf, 2013 [Publ 3531], fig. 2

iT tT bi riS oT roT roS iD
5 5 3 4 30 27 3 15.0
  +----------- Quadrula_quadrula ** 
--+                              
  |  +-------- Eurynia_dilatata **  
  +--+                           
     |  +----- Amblema_plicata **   
     +--+                        
        |  +-- Lampsilis_ornata **  
        +--+                     
           +-- Truncilla_truncata **

Pfeiffer & Graf, 2015 [Publ 3620], fig. 2

iT tT bi riS oT roT roS iD
5 5 3 3 35 25 3 14.0
                    +----- Unio_pictorum             
  +-----------------+                                
  |                 |  +-- Pyganodon_grandis         
  |                 +--+                             
  |                    +-- Lasmigona_costata         
  |                                                  
  |           +----------- Quadrula_quadrula **         
  |  +--------+                                      
--+  |        |  +-------- Amblema_plicata **           
  |  |        +--+                                   
  |  |           |  +----- Potamilus_alatus **          
  |  |           +--+                                
  |  |              |  +-- Lampsilis_cardium **         
  |  |              +--+                             
  |  |                 +-- Truncilla_truncata **        
  +--+                                               
     |           +-------- Ensidens_telus            
     |  +--------+                                   
     |  |        |  +----- Lens_micropterus          
     |  |        +--+                                
     |  |           |  +-- Physunio                  
     |  |           +--+                             
     |  |              +-- Physunio_modelli          
     +--+                                            
        |        +-------- Gonidea_angulata          
        |  +-----+                                   
        |  |     |  +----- Potomida_littoralis       
        |  |     +--+                                
        |  |        |  +-- Pronodularia_japanensis   
        +--+        +--+                             
           |           +-- Lamprotula_caveata        
           |                                         
           |  +----------- Pilsbryoconcha_carinifera 
           |  |                                      
           +--+        +-- Pseudodon_vondembuschianus
              |  +-----+                             
              |  |     +-- Pseudodon_vagulus         
              +--+                                   
                 |  +----- Sundadontina_tumida       
                 +--+                                
                    |  +-- Nyeinchanconcha_nyeinchani
                    +--+                             
                       +-- Bineurus_exilis           

Plazzi et al., 2011 [Publ 3676], fig. 5

iT tT bi riS oT roT roS iD
5 5 2 3 13 12 2 16.0
  +-------- Quadrula_quadrula **          
  |                                    
--+-------- Parvaspina_collina **         
  |                                    
  |  +----- Lampsilis_ornata **           
  +--+                                 
     |  +-- Epioblasma_rangiana **        
     +--+                              
        +-- Venustaconcha_ellipsiformis **

Cha et al., 2018 [Publ 3916], fig. 1

iT tT bi riS oT roT roS iD
5 5 3 2 34 31 2 12.0
                    +----------- Quadrula_quadrula **          
  +-----------------+                                       
  |                 |  +-------- Toxolasma_parvum **           
  |                 +--+                                    
  |                    |  +----- Lampsilis_ornata **           
  |                    +--+                                 
  |                       |  +-- Potamilus_alatus **           
  |                       +--+                              
  |                          +-- Potamilus_leptodon **         
  |                                                         
  |                          +-- Sinohyriopsis_cumingii     
  |              +-----------+                              
  |              |           +-- Sinohyriopsis_schlegelii   
--+              |                                          
  |  +-----------+        +----- Ptychorhynchus_pfisteri    
  |  |           |  +-----+                                 
  |  |           |  |     |  +-- Sinosolenaia_carinata      
  |  |           |  |     +--+                              
  |  |           +--+        +-- Sinosolenaia_oleivora      
  |  |              |                                       
  |  |              |  +-------- Potomida_littoralis        
  |  |              +--+                                    
  |  |                 |  +----- Pronodularia_japanensis    
  |  |                 +--+                                 
  |  |                    |  +-- Lamprotula_caveata         
  +--+                    +--+                              
     |                       +-- Lamprotula_leaii           
     |                                                      
     |  +----------------------- Lepidodesma_languilati     
     |  |                                                   
     |  |                 +----- Aculamprotula_tortuosa     
     |  |        +--------+                                 
     |  |        |        |  +-- Aculamprotula_coreana      
     |  |        |        +--+                              
     |  |        |           +-- Aculamprotula_tientsinensis
     |  |  +-----+                                          
     +--+  |     |        +----- Schistodesmus_lampreyanus  
        |  |     |  +-----+                                 
        |  |     |  |     |  +-- Nodularia_douglasiae       
        |  |     |  |     +--+                              
        |  |     +--+        +-- Tchangsinaia_piscicula     
        |  |        |                                       
        |  |        |  +-------- Unio_tumidus               
        |  |        +--+                                    
        +--+           |  +----- Unio_crassus               
           |           +--+                                 
           |              |  +-- Unio_pictorum              
           |              +--+                              
           |                 +-- Unio_delphinus             
           |                                                
           |  +----------------- Lanceolaria_grayii         
           |  |                                             
           |  |     +----------- Anodonta_anatina           
           |  |  +--+                                       
           +--+  |  |  +-------- Platynaias_compressa       
              |  |  +--+                                    
              |  |     |  +----- Pyganodon_grandis          
              |  |     +--+                                 
              +--+        |  +-- Utterbackia_peninsularis   
                 |        +--+                              
                 |           +-- Utterbackia_imbecillis     
                 |                                          
                 |     +-------- Cristaria_plicata          
                 |     |                                    
                 +-----+     +-- Anemina_arcaeformis        
                       |  +--+                              
                       |  |  +-- Anemina_euscaphys          
                       +--+                                 
                          |  +-- Sinanodonta_lucida         
                          +--+                              
                             +-- Sinanodonta_woodiana       

Saxon et al., 2023 [Publ 4338], fig. 4

iT tT bi riS oT roT roS iD
5 5 3 1 0 0 0 21.0
  +----------- Ellipsaria_lineolata **  
--+                                  
  |  +-------- Truncilla_truncata **    
  +--+                               
     |  +----- Truncilla_macrodon **    
     +--+                            
        |  +-- Truncilla_cognata **     
        +--+                         
           +-- Truncilla_donaciformis **

Graf, 2002 [Publ 2651], fig. 2

iT tT bi riS oT roT roS iD
4 4 2 3 14 8 2 18.0
           +-- Unio_pictorum        
        +--+                        
        |  +-- Unio_caffer          
  +-----+                           
  |     |  +-- Alasmidonta_marginata
  |     +--+                        
--+        +-- Pyganodon_grandis    
  |                                 
  |  +-------- Gonidea_angulata     
  |  |                              
  +--+     +-- Lampsilis_cardium **    
     |  +--+                        
     |  |  +-- Obliquaria_reflexa **   
     +--+                           
        |  +-- Eurynia_dilatata **     
        +--+                        
           +-- Amblema_plicata **      

Moyer & Díaz-Ferguson, 2012 [Publ 3375], fig. 3

iT tT bi riS oT roT roS iD
4 4 2 1 3 3 1 19.0
        +----- Lasmigona_costata    
  +-----+                           
  |     |  +-- Platynaias_compressa 
  |     +--+                        
--+        +-- Platynaias_subviridis
  |                                 
  |  +-------- Lampsilis_ovata[2] **   
  +--+                              
     |  +----- Lampsilis_ovata[1] **   
     +--+                           
        |  +-- Lampsilis_virescens **  
        +--+                        
           +-- Lampsilis_fasciola **   

Mulvey et al., 1997 [Publ 3390], fig. 1, 3

iT tT bi riS oT roT roS iD
4 4 2 2 1 1 1 18.0
  +----------- Utterbackia_imbecillis
--+                                  
  |  +-------- Megalonaias_nervosa **   
  +--+                               
     |  +----- Amblema_plicata **       
     +--+                            
        |  +-- Amblema_neislerii **     
        +--+                         
           +-- Amblema_elliottii **     

Zanatta & Harris, 2013 [Publ 3530], fig. 2

iT tT bi riS oT roT roS iD
4 4 2 1 1 1 1 17.0
  +----------- Pyganodon_grandis          
--+                                       
  |  +-------- Lampsilis_cardium **          
  +--+                                    
     |  +----- Epioblasma_rangiana **        
     +--+                                 
        |  +-- Venustaconcha_pleasii **      
        +--+                              
           +-- Venustaconcha_ellipsiformis **

Gangloff et al., 2013 [Publ 3551], fig. 2

iT tT bi riS oT roT roS iD
4 4 2 1 0 0 0 19.0
     +-- Ligumia_recta **              
  +--+                              
  |  +-- Sagittunio_nasutus **         
--+                                 
  |  +-- Venustaconcha_ellipsiformis **
  +--+                              
     +-- Venustaconcha_pleasii **      

Huang et al., 2013 [Publ 3629], fig. 3

iT tT bi riS oT roT roS iD
4 4 2 2 12 11 2 13.0
     +-------------- Unio_pictorum              
     |                                          
  +--+           +-- Sinanodonta_woodiana       
  |  |  +--------+                              
  |  |  |        +-- Cristaria_plicata          
  |  +--+                                       
  |     |  +-------- Platynaias_compressa       
  |     +--+                                    
  |        |  +----- Pyganodon_grandis          
  |        +--+                                 
  |           |  +-- Utterbackia_imbecillis     
--+           +--+                              
  |              +-- Utterbackia_peninsularis   
  |                                             
  |        +-------- Quadrula_quadrula **          
  |     +--+                                    
  |     |  |  +----- Toxolasma_parvum **           
  |     |  +--+                                 
  |     |     |  +-- Lampsilis_ornata **           
  |     |     +--+                              
  +-----+        +-- Venustaconcha_ellipsiformis **
        |                                       
        |        +-- Sinosolenaia_carinata      
        |     +--+                              
        |     |  +-- Pronodularia_japanensis    
        +-----+                                 
              |  +-- Sinohyriopsis_cumingii     
              +--+                              
                 +-- Sinohyriopsis_schlegelii   

Inoue et al., 2014 [Publ 3635], fig. 3A

iT tT bi riS oT roT roS iD
4 20 16 4 17 16 2 31.0
  +-------------------------- Amblema_plicata **            
  |                                                      
  +-------------------------- Eurynia_dilatata **           
  |                                                      
--+-------------------------- Lampsilis_ornata **           
  |                                                      
  |  +----------------------- Quadrula_quadrula **          
  |  |                                                   
  |  |  +-------------------- Gonidea_angulata           
  +--+  |                                                
     |  |        +----------- Pseudodontoideus_subvexus  
     |  |  +-----+                                       
     +--+  |     |  +-------- Lasmigona_costata          
        |  |     +--+                                    
        |  |        |  +----- Lasmigona_complanata       
        |  |        +--+                                 
        |  |           |  +-- Arcidens_confragosus       
        +--+           +--+                              
           |              +-- Arcidens_wheeleri          
           |                                             
           |  +-------------- Strophitus_undulatus       
           |  |                                          
           |  |     +-------- Pyganodon_grandis          
           |  |  +--+                                    
           +--+  |  |  +----- Utterbackia_peninsularis[2]
              |  |  +--+                                 
              |  |     |  +-- Utterbackia_imbecillis     
              |  |     +--+                              
              +--+        +-- Utterbackia_peninsularis[1]
                 |                                       
                 |        +-- Platynaias_compressa       
                 |  +-----+                              
                 |  |     +-- Platynaias_subviridis      
                 +--+                                    
                    |  +----- Alasmidonta_marginata      
                    +--+                                 
                       |  +-- Alasmidonta_undulata       
                       +--+                              
                          +-- Alasmidonta_varicosa       

Zhou et al., 2016 [Publ 3710], fig. 1

iT tT bi riS oT roT roS iD
4 4 2 2 16 15 2 13.0
     +-------- Quadrula_quadrula **          
  +--+                                    
  |  |  +----- Toxolasma_parvum **           
  |  +--+                                 
  |     |  +-- Venustaconcha_ellipsiformis **
  |     +--+                              
--+        +-- Lampsilis_ornata **           
  |                                       
  |        +-- Sinohyriopsis_cumingii     
  |  +-----+                              
  |  |     +-- Sinohyriopsis_schlegelii   
  |  |                                    
  +--+     +-- Lamprotula_leaii           
     |  +--+                              
     |  |  +-- Pronodularia_japanensis    
     +--+                                 
        |  +-- Sinosolenaia_carinata      
        +--+                              
           +-- Sinosolenaia_oleivora      

Guerra et al., 2017 [Publ 3772], fig. 2a

iT tT bi riS oT roT roS iD
4 4 2 2 21 15 2 13.0
                 +-------- Quadrula_quadrula **          
              +--+                                    
              |  |  +----- Toxolasma_parvum **           
              |  +--+                                 
              |     |  +-- Venustaconcha_ellipsiformis **
              |     +--+                              
  +-----------+        +-- Lampsilis_ornata **           
  |           |                                       
  |           |        +-- Sinohyriopsis_schlegelii   
  |           |  +-----+                              
  |           |  |     +-- Sinohyriopsis_cumingii     
  |           +--+                                    
  |              |  +----- Pronodularia_japanensis    
--+              +--+                                 
  |                 |  +-- Sinosolenaia_oleivora      
  |                 +--+                              
  |                    +-- Sinosolenaia_carinata      
  |                                                   
  |                    +-- Aculamprotula_tortuosa     
  |  +-----------------+                              
  |  |                 +-- Unio_pictorum              
  +--+                                                
     |  +----------------- Sinanodonta_woodiana       
     |  |                                             
     +--+  +-------------- Cristaria_plicata          
        |  |                                          
        +--+  +----------- Anodonta_anatina           
           |  |                                       
           +--+        +-- Platynaias_subviridis      
              |  +-----+                              
              |  |     +-- Platynaias_compressa       
              +--+                                    
                 |  +----- Pyganodon_grandis          
                 +--+                                 
                    |  +-- Utterbackia_peninsularis   
                    +--+                              
                       +-- Utterbackia_imbecillis     

Guerra et al., 2017 [Publ 3772], fig. 2b

iT tT bi riS oT roT roS iD
4 4 2 2 21 15 2 13.0
                    +-------- Quadrula_quadrula **          
  +-----------------+                                    
  |                 |  +----- Toxolasma_parvum **           
  |                 +--+                                 
  |                    |  +-- Venustaconcha_ellipsiformis **
  |                    +--+                              
  |                       +-- Lampsilis_ornata **           
  |                                                      
--+                       +-- Sinohyriopsis_schlegelii   
  |                 +-----+                              
  |                 |     +-- Sinohyriopsis_cumingii     
  |  +--------------+                                    
  |  |              |  +----- Pronodularia_japanensis    
  |  |              +--+                                 
  |  |                 |  +-- Sinosolenaia_oleivora      
  +--+                 +--+                              
     |                    +-- Sinosolenaia_carinata      
     |                                                   
     |  +-------------------- Unio_pictorum              
     |  |                                                
     +--+  +----------------- Aculamprotula_tortuosa     
        |  |                                             
        +--+              +-- Cristaria_plicata          
           |  +-----------+                              
           |  |           +-- Sinanodonta_woodiana       
           +--+                                          
              |  +----------- Anodonta_anatina           
              |  |                                       
              +--+        +-- Platynaias_subviridis      
                 |  +-----+                              
                 |  |     +-- Platynaias_compressa       
                 +--+                                    
                    |  +----- Pyganodon_grandis          
                    +--+                                 
                       |  +-- Utterbackia_peninsularis   
                       +--+                              
                          +-- Utterbackia_imbecillis     

Fonseca et al., 2016 [Publ 3775], fig. 1

iT tT bi riS oT roT roS iD
4 4 2 2 28 26 2 13.0
                 +-------- Quadrula_quadrula **          
  +--------------+                                    
  |              |  +----- Toxolasma_parvum **           
  |              +--+                                 
  |                 |  +-- Lampsilis_ornata **           
  |                 +--+                              
  |                    +-- Venustaconcha_ellipsiformis **
  |                                                   
  |                    +-- Sinohyriopsis_cumingii     
  |           +--------+                              
  |           |        +-- Sinohyriopsis_schlegelii   
  |           |                                       
--+  +--------+        +-- Sinosolenaia_oleivora      
  |  |        |  +-----+                              
  |  |        |  |     +-- Sinosolenaia_carinata      
  |  |        +--+                                    
  |  |           |  +----- Potomida_littoralis        
  |  |           +--+                                 
  |  |              |  +-- Lamprotula_leaii           
  |  |              +--+                              
  |  |                 +-- Pronodularia_japanensis    
  |  |                                                
  |  |                 +-- Lanceolaria_lanceolata     
  |  |     +-----------+                              
  +--+     |           +-- Lanceolaria_grayii         
     |     |                                          
     |  +--+           +-- Aculamprotula_tortuosa     
     |  |  |  +--------+                              
     |  |  |  |        +-- Aculamprotula_coreana      
     |  |  |  |                                       
     |  |  +--+        +-- Unio_pictorum              
     |  |     |  +-----+                              
     |  |     |  |     +-- Unio_delphinus             
     |  |     +--+                                    
     |  |        |  +----- Schistodesmus_lampreyanus  
     |  |        +--+                                 
     +--+           |  +-- Nodularia_douglasiae       
        |           +--+                              
        |              +-- Tchangsinaia_piscicula     
        |                                             
        |        +-------- Cristaria_plicata          
        |        |                                    
        |  +-----+     +-- Anemina_euscaphys          
        |  |     |  +--+                              
        |  |     |  |  +-- Anemina_arcaeformis        
        |  |     +--+                                 
        +--+        |  +-- Sinanodonta_woodiana       
           |        +--+                              
           |           +-- Sinanodonta_lucida         
           |                                          
           |  +----------- Anodonta_anatina           
           +--+                                       
              |  +-------- Platynaias_compressa       
              +--+                                    
                 |  +----- Pyganodon_grandis          
                 +--+                                 
                    |  +-- Utterbackia_peninsularis   
                    +--+                              
                       +-- Utterbackia_imbecillis     

Amaro et al., 2017 [Publ 3791], fig. 2-M

iT tT bi riS oT roT roS iD
4 4 2 3 6 6 2 16.0
           +-- Fusconaia_flava **            
        +--+                              
        |  +-- Quadrula_quadrula **          
  +-----+                                 
  |     |  +-- Lampsilis_siliquoidea **      
  |     +--+                              
  |        +-- Venustaconcha_ellipsiformis **
--+                                       
  |        +-- Pronodularia_japanensis    
  |  +-----+                              
  |  |     +-- Sinosolenaia_carinata      
  +--+                                    
     |  +----- Pyganodon_grandis          
     |  |                                 
     +--+  +-- Lasmigona_costata          
        |  |                              
        +--+-- Sinanodonta_woodiana       
           |                              
           +-- Utterbackia_peninsularis   

Lopes-Lima et al., 2017 [Publ 3792], fig. 2

iT tT bi riS oT roT roS iD
4 4 2 2 29 26 2 13.0
                 +-------- Quadrula_quadrula **          
  +--------------+                                    
  |              |  +----- Toxolasma_parvum **           
  |              +--+                                 
  |                 |  +-- Lampsilis_ornata **           
  |                 +--+                              
  |                    +-- Venustaconcha_ellipsiformis **
  |                                                   
  |                    +-- Sinohyriopsis_cumingii     
  |           +--------+                              
  |           |        +-- Sinohyriopsis_schlegelii   
  |           |                                       
--+  +--------+        +-- Sinosolenaia_oleivora      
  |  |        |  +-----+                              
  |  |        |  |     +-- Sinosolenaia_carinata      
  |  |        +--+                                    
  |  |           |  +----- Potomida_littoralis        
  |  |           +--+                                 
  |  |              |  +-- Pronodularia_japanensis    
  |  |              +--+                              
  |  |                 +-- Lamprotula_leaii           
  |  |                                                
  |  |                 +-- Lanceolaria_lanceolata     
  |  |     +-----------+                              
  +--+     |           +-- Lanceolaria_grayii         
     |     |                                          
     |  +--+           +-- Aculamprotula_tortuosa     
     |  |  |  +--------+                              
     |  |  |  |        +-- Aculamprotula_coreana      
     |  |  |  |                                       
     |  |  +--+        +-- Unio_delphinus             
     |  |     |  +-----+                              
     |  |     |  |     +-- Unio_pictorum              
     |  |     +--+                                    
     |  |        |  +----- Schistodesmus_lampreyanus  
     |  |        +--+                                 
     +--+           |  +-- Tchangsinaia_piscicula     
        |           +--+                              
        |              +-- Nodularia_douglasiae       
        |                                             
        |     +----------- Anodonta_anatina           
        |  +--+                                       
        |  |  |  +-------- Platynaias_compressa       
        |  |  +--+                                    
        |  |     |  +----- Pyganodon_grandis          
        |  |     +--+                                 
        +--+        |  +-- Utterbackia_peninsularis   
           |        +--+                              
           |           +-- Utterbackia_imbecillis     
           |                                          
           |     +-------- Cristaria_plicata          
           |     |                                    
           +-----+     +-- Sinanodonta_lucida         
                 |  +--+                              
                 |  |  +-- Sinanodonta_woodiana       
                 +--+                                 
                    |  +-- Anemina_arcaeformis        
                    +--+                              
                       +-- Anemina_euscaphys          

Pfeiffer et al., 2018 [Publ 3835], fig. 2

iT tT bi riS oT roT roS iD
4 4 2 1 0 0 0 18.0
  +-------- Quadrula_quadrula **      
--+                                
  |  +----- Uniomerus_declivis **     
  +--+                             
     |  +-- Megalonaias_nervosa **    
     +--+                          
        +-- Megalonaias_nickliniana **

Aunins et al., 2018 [Publ 3863], fig. 1

iT tT bi riS oT roT roS iD
4 4 2 2 37 34 2 13.0
                       +-------- Quadrula_quadrula **          
  +--------------------+                                    
  |                    |  +----- Toxolasma_parvum **           
  |                    +--+                                 
  |                       |  +-- Lampsilis_ornata **           
  |                       +--+                              
  |                          +-- Venustaconcha_ellipsiformis **
  |                                                         
  |                          +-- Sinohyriopsis_cumingii     
  |                 +--------+                              
  |                 |        +-- Sinohyriopsis_schlegelii   
--+                 |                                       
  |  +--------------+        +-- Lamprotula_caveata         
  |  |              |  +-----+                              
  |  |              |  |     +-- Potomida_littoralis        
  |  |              +--+                                    
  |  |                 |  +----- Sinosolenaia_oleivora      
  |  |                 +--+                                 
  |  |                    |  +-- Ptychorhynchus_pfisteri    
  |  |                    +--+                              
  |  |                       +-- Sinosolenaia_carinata      
  +--+                                                      
     |  +----------------------- Lepidodesma_languilati     
     |  |                                                   
     |  |                 +----- Aculamprotula_tortuosa     
     |  |        +--------+                                 
     |  |        |        |  +-- Aculamprotula_coreana      
     |  |        |        +--+                              
     |  |        |           +-- Aculamprotula_tientsinensis
     |  |        |                                          
     |  |  +-----+     +-------- Nodularia_douglasiae       
     +--+  |     |  +--+                                    
        |  |     |  |  |  +----- Tchangsinaia_piscicula     
        |  |     |  |  +--+                                 
        |  |     |  |     |  +-- Cuneopsis_heudei           
        |  |     +--+     +--+                              
        |  |        |        +-- Schistodesmus_lampreyanus  
        |  |        |                                       
        |  |        |  +-------- Unio_tumidus               
        |  |        +--+                                    
        |  |           |  +----- Unio_crassus               
        +--+           +--+                                 
           |              |  +-- Unio_pictorum              
           |              +--+                              
           |                 +-- Unio_delphinus             
           |                                                
           |              +----- Acuticosta_chinensis       
           |  +-----------+                                 
           |  |           |  +-- Lanceolaria_grayii         
           |  |           +--+                              
           |  |              +-- Lanceolaria_lanceolata     
           |  |                                             
           |  |        +-------- Cristaria_plicata          
           +--+        |                                    
              |  +-----+     +-- Anemina_arcaeformis        
              |  |     |  +--+                              
              |  |     |  |  +-- Anemina_euscaphys          
              |  |     +--+                                 
              |  |        |  +-- Sinanodonta_woodiana       
              |  |        +--+                              
              +--+           +-- Sinanodonta_lucida         
                 |                                          
                 |           +-- Anodonta_anatina           
                 |  +--------+                              
                 |  |        +-- Anodonta_cygnea            
                 |  |                                       
                 +--+        +-- Prolasmidonta_heterodon    
                    |  +-----+                              
                    |  |     +-- Alasmidonta_varicosa       
                    +--+                                    
                       |  +----- Pyganodon_grandis          
                       +--+                                 
                          |  +-- Utterbackia_imbecillis     
                          +--+                              
                             +-- Utterbackia_peninsularis   

Lopes-Lima et al., 2020 [Publ 4080], fig. 2-4

iT tT bi riS oT roT roS iD
4 4 0 4 118 107 3 15.0
                          +-- Lens_contradens            
                          |                              
                          +-- Ensidens_sagittarius       
                          |                              
  +-----------------------+-- Physunio_modelli           
  |                       |                              
  |                       +-- Rectidens_sumatrensis      
  |                       |                              
  |                       +-- Yaukthwa_nesemanni         
  |                                                      
  |                       +-- Amblema_plicata **            
  |                       |                              
  |                       +-- Elliptio_complanata **        
  |  +--------------------+                              
--+  |                    +-- Lampsilis_cardium **          
  |  |                    |                              
  |  |                    +-- Quadrula_quadrula **          
  |  |                                                   
  |  |                 +----- Chamberlainia_somsakpanhai 
  |  |     +-----------+                                 
  |  |     |           |  +-- Sinohyriopsis_cumingii     
  |  |     |           +--+                              
  |  |  +--+              +-- Sinohyriopsis_schlegelii   
  |  |  |  |                                             
  |  |  |  |  +-------------- Pronodularia_japanensis    
  +--+  |  |  |                                          
     |  |  +--+  +----------- Potomida_littoralis        
     |  |     |  |                                       
     |  |     +--+  +-------- Lamprotula_caveata         
     |  |        |  |                                    
     |  |        +--+  +----- Lamprotula_leaii[2]        
     |  |           |  |                                 
     |  |           +--+----- Lamprotula_leaii[3]        
     |  |              |                                 
     |  |              |  +-- Lamprotula_seomjinensis    
     |  |              +--+                              
     |  |                 +-- Lamprotula_leaii[1]        
     +--+                                                
        |                 +-- Indopseudodon_indawgyiensis
        |        +--------+                              
        |        |        +-- Indopseudodon_salwenianus  
        |        |                                       
        |  +-----+        +-- Pilsbryoconcha_exilis      
        |  |     |  +-----+                              
        |  |     |  |     +-- Pilsbryoconcha_carinifera  
        |  |     +--+                                    
        |  |        |  +----- Pseudodon_mekongi          
        |  |        |  |                                 
        |  |        +--+  +-- Nyeinchanconcha_nyeinchani 
        |  |           |  |                              
        |  |           +--+-- Bineurus_mouhotii          
        |  |              |                              
        +--+              +-- Sundadontina_tumida        
           |                                             
           |        +-------- Inversidens_brandtii       
           |        |                                    
           |  +-----+     +-- Obovalis_omiensis          
           |  |     |  +--+                              
           |  |     |  |  +-- Ptychorhynchus_pfisteri    
           |  |     +--+                                 
           |  |        |  +-- Parvasolenaia_rivularis    
           |  |        +--+                              
           +--+           +-- Koreosolenaia_sitgyensis   
              |                                          
              |        +----- Gonidea_angulata           
              |  +-----+                                 
              |  |     |  +-- Leguminaia_wheatleyi       
              |  |     +--+                              
              +--+        +-- Microcondylaea_bonellii    
                 |                                       
                 |  +-------- Sinosolenaia_carinata      
                 +--+                                    
                    |  +----- Sinosolenaia_recognita     
                    +--+                                 
                       |  +-- Sinosolenaia_iridinea      
                       +--+                              
                          +-- Sinosolenaia_oleivora      

Guerra et al., 2019 [Publ 4168], fig. 1

iT tT bi riS oT roT roS iD
4 4 2 2 37 17 2 13.0
                 +-------- Quadrula_quadrula **          
  +--------------+                                    
  |              |  +----- Toxolasma_parvum **           
  |              +--+                                 
  |                 |  +-- Venustaconcha_ellipsiformis **
  |                 +--+                              
  |                    +-- Potamilus_alatus **           
  |                                                   
--+              +-------- Sinosolenaia_carinata      
  |  +-----------+                                    
  |  |           |  +----- Pronodularia_japanensis    
  |  |           +--+                                 
  |  |              |  +-- Lamprotula_leaii           
  |  |              +--+                              
  |  |                 +-- Potomida_littoralis        
  |  |                                                
  +--+     +-------------- Lanceolaria_lanceolata     
     |  +--+                                          
     |  |  |  +----------- Aculamprotula_tortuosa     
     |  |  +--+                                       
     |  |     |  +-------- Schistodesmus_lampreyanus  
     |  |     +--+                                    
     |  |        |  +----- Unio_tumidus               
     +--+        +--+                                 
        |           |  +-- Unio_crassus               
        |           +--+                              
        |              +-- Unio_delphinus             
        |                                             
        |  +-------------- Sinanodonta_woodiana       
        |  |                                          
        +--+  +----------- Anodonta_anatina           
           |  |                                       
           +--+        +-- Platynaias_compressa       
              |  +-----+                              
              |  |     +-- Platynaias_subviridis      
              +--+                                    
                 |  +----- Pyganodon_grandis          
                 +--+                                 
                    |  +-- Utterbackia_peninsularis   
                    +--+                              
                       +-- Utterbackia_imbecillis     

Guerra et al., 2019 [Publ 4168], fig. 2

iT tT bi riS oT roT roS iD
4 4 2 2 37 17 2 13.0
                       +-------- Quadrula_quadrula **          
  +--------------------+                                    
  |                    |  +----- Toxolasma_parvum **           
  |                    +--+                                 
  |                       |  +-- Venustaconcha_ellipsiformis **
  |                       +--+                              
  |                          +-- Potamilus_alatus **           
--+                                                         
  |                    +-------- Sinosolenaia_carinata      
  |  +-----------------+                                    
  |  |                 |  +----- Pronodularia_japanensis    
  |  |                 +--+                                 
  |  |                    |  +-- Lamprotula_leaii           
  |  |                    +--+                              
  +--+                       +-- Potomida_littoralis        
     |                                                      
     |  +----------------------- Aculamprotula_tortuosa     
     |  |                                                   
     |  |              +-------- Schistodesmus_lampreyanus  
     +--+  +-----------+                                    
        |  |           |  +----- Unio_tumidus               
        |  |           +--+                                 
        |  |              |  +-- Unio_crassus               
        +--+              +--+                              
           |                 +-- Unio_delphinus             
           |                                                
           |  +----------------- Lanceolaria_lanceolata     
           |  |                                             
           +--+  +-------------- Sinanodonta_woodiana       
              |  |                                          
              +--+  +----------- Anodonta_anatina           
                 |  |                                       
                 +--+        +-- Platynaias_compressa       
                    |  +-----+                              
                    |  |     +-- Platynaias_subviridis      
                    +--+                                    
                       |  +----- Pyganodon_grandis          
                       +--+                                 
                          |  +-- Utterbackia_peninsularis   
                          +--+                              
                             +-- Utterbackia_imbecillis     

Patterson et al., 2021 [Publ 4222], fig. 3

iT tT bi riS oT roT roS iD
4 4 2 2 0 0 0 18.0
  +-------- Villosa_villosa **  
--+                          
  |  +----- Amblema_neislerii **
  +--+                       
     |  +-- Amblema_plicata **  
     +--+                    
        +-- Amblema_elliottii **

Johnson et al., 2023 [Publ 4323], fig. 3

iT tT bi riS oT roT roS iD
4 4 2 1 0 0 0 20.0
  +-------- Fusconaia_cerina **      
--+                               
  |  +----- Pleurobema_riddellii **  
  +--+                            
     |  +-- Pleurobema_beadleianum **
     +--+                         
        +-- Pleurobema_clava **      

Krebs, 2004 [Publ 2692], fig. 2

iT tT bi riS oT roT roS iD
3 5 3 3 2 2 1 17.0
  +-------- Quadrula_quadrula **    
  |                              
--+     +-- Lasmigona_costata    
  |  +--+                        
  |  |  +-- Pyganodon_grandis    
  +--+                           
     |  +-- Lampsilis_siliquoidea **
     +--+                        
        +-- Fusconaia_flava **      

Curole & Kocher, 2002 [Publ 2712], fig. 1-F

iT tT bi riS oT roT roS iD
3 3 1 2 1 0 0 17.0
  +----- Lampsilis_teres **    
--+                         
  |  +-- Quadrula_quadrula **  
  +--+                      
     +-- Pustulosa_pustulosa **

Curole & Kocher, 2002 [Publ 2712], fig. 1a-M

iT tT bi riS oT roT roS iD
3 4 2 2 1 1 1 18.0
     +-- Utterbackiana_implicata
  +--+                          
  |  +-- Lampsilis_teres **        
--+                             
  |  +-- Quadrula_quadrula **      
  +--+                          
     +-- Pustulosa_pustulosa **    

Curole & Kocher, 2002 [Publ 2712], fig. 1b-M

iT tT bi riS oT roT roS iD
3 4 2 2 1 1 1 18.0
  +-------- Lampsilis_teres **        
--+                                
  |  +----- Utterbackiana_implicata
  +--+                             
     |  +-- Quadrula_quadrula **      
     +--+                          
        +-- Pustulosa_pustulosa **    

Chen et al., 2012 [Publ 3624], fig. 4

iT tT bi riS oT roT roS iD
3 3 1 2 4 4 2 14.0
     +----- Quadrula_quadrula **          
  +--+                                 
  |  |  +-- Lampsilis_ornata **           
  |  +--+                              
  |     +-- Venustaconcha_ellipsiformis **
--+                                    
  |     +-- Pyganodon_grandis          
  |  +--+                              
  |  |  +-- Cristaria_plicata          
  +--+                                 
     |  +-- Sinohyriopsis_cumingii     
     +--+                              
        +-- Lamprotula_leaii           

Sheng et al., 2014 [Publ 3704], fig. 2

iT tT bi riS oT roT roS iD
3 6 4 2 5 5 2 19.0
  +----------------- Cristaria_plicata          
  |                                             
--+  +-------------- Pyganodon_grandis          
  |  |                                          
  +--+  +----------- Lampsilis_ornata **           
     |  |                                       
     +--+        +-- Venustaconcha_ellipsiformis **
        |  +-----+                              
        |  |     +-- Quadrula_quadrula **          
        +--+                                    
           |  +----- Lamprotula_leaii           
           +--+                                 
              |  +-- Sinohyriopsis_cumingii     
              +--+                              
                 +-- Sinohyriopsis_schlegelii   

Wu et al., 2016 [Publ 3711], fig. 1

iT tT