|
Bivalvia | Unionoida superfamily ETHERIOIDEA Deshayes, 1832: cladomics
3 families
Cladomics Summary
number of species = 112
number of subtaxa (rank: family) = 3
analyzed in 64 cladograms in 36 publications
maximum number of terminals in an analysis = 10, mean = 4.39
maximum number of subtaxa (family) in an analysis = 3, mean = 2.31
Basic Monophyly
number of cladograms that test basic monophyly (i.e., more than 1 included species) = 53
maximum number of included terminals = 10, mean = 5.09
maximum number of subtaxa (family) = 3, mean = 2.58
proportion of cladograms that support basic monophyly = 0.92
Rigorous Monophyly
number of cladograms that test rigorous monophyly = 53
(Rigorous monophyly refers to cladograms with terminal species representing more than one subtaxon (family) taxon and at least one rigorous outgroup (superfamily).
maximum number of included terminals = 10, mean = 5.09
maximum number of subtaxa (family) = 3, mean = 2.58
proportion of cladograms that support rigorous monophyly = 0.92
Pairwise Cladogram Clustering
The graphs below depict in two dimensions all pair-wise ingroup distances within the cladome. See the Cladmoics FARQ for more information.
Lopes-Lima et al., 2017 [Publ 3745], fig. 1-2
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 6 |
6 |
4 |
3 |
75 |
74 |
3 |
0.0 |
*** Top Monophyly Cladogram *** *** Top Sister-Group Cladogram ***
+-- Cumberlandia_monodonta
+--------------------------------+
| +-- Margaritifera_margaritifera
|
| +----------- Mutela_rostrata **
| |
| +--------------------+ +-- Aspatharia_pfeifferiana **
| | | +-----+
| | | | +-- Chambardia_wahlbergi **
| | +--+
| | | +----- Etheria_elliptica **
| | +--+
| | | +-- Tamsiella_schroeteriana **
--+ | +--+
| | +-- Mycetopoda_siliquosa **
| |
| | +-------- Trapezidens_dolichorhynchus
| | +--------------------+
| | | | +----- Leoparreysia_tavoyensis
| | | +--+
| | | | +-- Coelatura_aegyptiaca
| | | +--+
| | | +-- Indochinella_pugio
| | |
| | | +----- Unio_gibbus
+--+ | |
| | +-----------+----- Unio_tumidus
| | | |
| | | | +-- Unio_crassus
| | | +--+
| | +--+ +-- Unio_pictorum
| | | |
| | | | +-------------- Aculamprotula_tortuosa
| | | | |
| | | +--+ +-- Nodularia_nuxpersicae
| | | | +--------+
| | | | | +-- Nodularia_douglasiae
| | | +--+
| | | | +-------- Schistodesmus_lampreyanus
| | | +--+
| | | | +----- Tchangsinaia_piscicula
| | | +--+
| | +-----+ | +-- Cuneopsis_heudei
| | | | +--+
| | | | +-- Cuneopsis_rufescens
+--+ | |
| | | +-------- Lanceolaria_gladiola
| | | +--------+
| | | | | +----- Lanceolaria_grayii[2]
| | | | +--+
| | | | | +-- Lanceolaria_lanceolata
| | | | +--+
| | | | +-- Lanceolaria_grayii[1]
| | | |
| | +--+ +----------- Cristaria_plicata
| | | +--+
| | | | | +-------- Anemina_arcaeformis
| | | | +--+
| | | | | +----- Pletholophus_tenuis
| | | | +--+
| | | | | +-- Sinanodonta_jourdyi
| | | | +--+
| | +--+ +-- Sinanodonta_lucida
| | |
| | | +----- Pseudanodonta_complanata
| | | |
| | | +-----+ +-- Anodonta_anatina
| | | | | |
| | | | +--+-- Anodonta_cygnea
| | | | |
| | +--+ +-- Anodonta_nuttalliana
| | |
| | | +-- Pyganodon_grandis
+--+ | +-----+
| | | +-- Simpsonaias_ambigua
| +--+
| | +----- Strophitus_undulatus
| +--+
| | +-- Alasmidonta_marginata
| +--+
| +-- Lasmigona_costata
|
| +-- Lens_contradens
| +--+
| | +-- Lens_comptus
| +--+
| | | +-- Physunio_modelli
| | +--+
| | +-- unionoid_species
| |
| +--------------+ +----- Hyriopsis_desowitzi
| | | |
| | | +----- Rectidens_sumatrensis
| | | |
| | | | +-- Hyriopsis_khoratensis
| | +--+--+
| | | +-- Hyriopsis_myersiana
| | |
| | | +-- Ensidens_ingallsianus
| | | |
| | +--+-- Ensidens_telus
| | |
| | +-- Ensidens_jaculus
| |
| | +-------- Amblema_plicata
+--+ +--+
| | | +----- Cambarunio_iris
| | +--+
| | | +-- Lampsilis_cardium
| | +--+
| +--------+ +-- Ortmanniana_ligamentina
| | |
| | | +-- Quadrula_quadrula
| | | +-----+
| | | | +-- Quadrula_verrucosa
| | +--+
| | | +----- Elliptio_complanata
| | +--+
| | | +-- Pleurobema_sintoxia
| | +--+
| | +-- Eurynia_dilatata
+--+
| +-- Chamberlainia_somsakpanhai
| +-----+
| | +-- Sinohyriopsis_cumingii
| |
| +--------+ +-- Lamprotula_caveata
| | | +--+
| | | | +-- Lamprotula_leaii
| | +--+
| | | +-- Potomida_littoralis
| | +--+
| | +-- Pronodularia_japanensis
+--+
| +-- Sinosolenaia_carinata
| +-----+
| | +-- Sinosolenaia_oleivora
| +-----+
| | | +----- Gonidea_angulata
| | +--+
| | | +-- Microcondylaea_bonellii
+--+ +--+
| +-- Leguminaia_wheatleyi
|
| +----------- Parvasolenaia_rivularis
| |
+--+ +-- Pilsbryoconcha_exilis
| +-----+
| | +-- Pilsbryoconcha_acuta
+--+
| +----- Pseudodon_vagulus
+--+
| +-- Bineurus_exilis
+--+
+-- Nyeinchanconcha_nyeinchani
Graf & Cummings, 2006 [Publ 1968], fig. 2
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 10 |
10 |
8 |
3 |
37 |
36 |
2 |
12.0 |
+----- Velesunio_ambiguus
+--------+
| | +-- Velesunio_angasi
| +--+
+--+ +-- Lortiella_rugata
| |
| | +----------- Echyridella_menziesii
| | |
| +--+ +-- Hyridella_depressa
| | +-----+
| | | +-- Hyridella_australis
| +--+
| | +----- Castalia
| +--+
+--------------------+ | +-- Diplodon_chilensis
| | +--+
| | +-- Diplodon_gratus
| |
| | +----- Etheria_elliptica **
| | +-----+
| | | | +-- Mulleria_rivoli **
| | | +--+
| | | +-- Pseudomulleria_dalyi **
| | |
| +-----+ +----- Monocondylaea_minuana **
| | +--+
| | | | +-- Anodontites_trigona **
| | | +--+
| | | +-- Anodontites_guanarensis **
--+ +--+
| | +-- Chambardia_wahlbergi **
| | +--+
| | | +-- Aspatharia_rugifera **
| +--+
| | +-- Mutela_dubia **
| +--+
| +-- Mutela_rostrata **
|
| +-- Margaritifera_margaritifera
| +--------------------------------+
| | +-- Cumberlandia_monodonta
| |
| | +-- Pilsbryoconcha_exilis
| | +-----------------------------+
+--+ | +-- Pseudodon_vondembuschianus
| |
| | +----------------------------- Coelatura_aegyptiaca
| | |
| | +----------------------------- Lens_contradens
+--+ |
| | +-- Unio_pictorum
| | +--------+
| | | +-- Unio_caffer
| +-----------------+
| | | +-------- Lasmigona_costata
+--+ +--+
| | +----- Alasmidonta_marginata
| +--+
| | +-- Pyganodon_grandis
| +--+
| +-- Strophitus_undulatus
|
| +-------------------------- Gonidea_angulata
| |
| | +-- Quadrula_verrucosa
+--+ +--------------------+
| | +-- Quadrula_quadrula
| |
+--+ +-------------------- Amblema_plicata
| |
| | +----- Eurynia_dilatata
+--+ +-----------+
| | | +-- Fusconaia_flava
| | +--+
| | +-- Pleurobema_sintoxia
+--+
| +-------------- Obliquaria_reflexa
| |
| +-------------- Truncilla_truncata
+--+
+-------------- Ptychobranchus_fasciolaris
|
| +----------- Ligumia_recta
+--+
| +-------- Cambarunio_iris
+--+
| +----- Epioblasma_triquetra
+--+
| +-- Lampsilis_cardium
+--+
+-- Ortmanniana_ligamentina
Hoeh et al., 2009 [Publ 2716], fig. 3
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 10 |
37 |
34 |
3 |
37 |
36 |
2 |
38.0 |
+----- Velesunio_ambiguus
+--------+
| | +-- Velesunio_angasi
| +--+
+-----------------------------+ +-- Lortiella_rugata
| |
| | +----------- Echyridella_menziesii
| | |
| +--+ +-- Hyridella_depressa
| | +-----+
| | | +-- Hyridella_australis
| +--+
| | +----- Castalia
| +--+
| | +-- Diplodon_chilensis
| +--+
| +-- Diplodon_gratus
--+
| +----- Chambardia_wahlbergi **
| |
| +--------+----- Aspatharia_rugifera **
| | |
| | | +-- Mutela_rostrata **
| +--------------------------+ +--+
| | | +-- Mutela_dubia **
| | |
| | | +----------- Etheria_elliptica **
| | +--+
| | | +-------- Mulleria_rivoli **
| | +--+
| | | +----- Monocondylaea_minuana **
+--+ +--+
| | +-- Anodontites_trigona **
| +--+
| +-- Anodontites_guanarensis **
|
| +-- Margaritifera_margaritifera
| +-----------------------------------+
| | +-- Cumberlandia_monodonta
| |
+--+ +----------------------------------- Coelatura_aegyptiaca
| |
| | +-------------------------------- Pseudomulleria_dalyi **
| | |
+--+ | +-- Unio_pictorum
| | +--------+
| | | +-- Unio_caffer
| | +-----------------+
+--+ | | +-------- Lasmigona_costata
| | +--+
| | | +----- Alasmidonta_marginata
| | +--+
| | | +-- Strophitus_undulatus
| | +--+
+--+ +-- Pyganodon_grandis
|
| +-------------------------- Lens_contradens
| |
| | +----- Gonidea_angulata
| | +-----------------+
| | | | +-- Pilsbryoconcha_exilis
+--+ | +--+
| | +-- Pseudodon_vondembuschianus
| |
| | +----------- Amblema_plicata
| | |
+--+ | +----- Obliquaria_reflexa
| +--------+ +--+
| | | | | +-- Quadrula_verrucosa
| | | | +--+
| | +--+ +-- Quadrula_quadrula
| | |
| | | +----- Eurynia_dilatata
+--+ +--+
| | +-- Pleurobema_sintoxia
| +--+
| +-- Fusconaia_flava
|
| +----------------- Ptychobranchus_fasciolaris
+--+
| +-------------- Truncilla_truncata
+--+
| +----------- Ortmanniana_ligamentina
+--+
| +-------- Lampsilis_cardium
+--+
| +----- Cambarunio_iris
+--+
| +-- Ligumia_recta
+--+
+-- Epioblasma_triquetra
Hoeh et al., 2009 [Publ 2716], fig. 4
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 10 |
36 |
33 |
3 |
36 |
35 |
2 |
38.0 |
+----- Velesunio_ambiguus
+--------+
| | +-- Velesunio_angasi
| +--+
+--------------------------------+ +-- Lortiella_rugata
| |
| | +----------- Echyridella_menziesii
| | |
| +--+ +-- Hyridella_depressa
| | +-----+
| | | +-- Hyridella_australis
| +--+
| | +----- Castalia
| +--+
| | +-- Diplodon_chilensis
| +--+
| +-- Diplodon_gratus
--+
| +-- Etheria_elliptica **
| +--------+
| | +-- Mulleria_rivoli **
| |
| | +----- Monocondylaea_minuana **
| +--------------------------------+ +--+
| | | | | +-- Anodontites_trigona **
| | | | +--+
| | | | +-- Anodontites_guanarensis **
| | +--+
| | | +-- Mutela_rostrata **
| | | +--+
+--+ | | +-- Mutela_dubia **
| +--+
| | +-- Chambardia_wahlbergi **
| +--+
| +-- Aspatharia_rugifera **
|
| +----------------------------------------- Coelatura_aegyptiaca
| |
+--+ +-------------------------------------- Pseudomulleria_dalyi **
| |
| | +-- Margaritifera_margaritifera
+--+ +--------------------------------+
| | +-- Cumberlandia_monodonta
| |
| | +-- Pilsbryoconcha_exilis
+--+ +-----------------------------+
| | +-- Pseudodon_vondembuschianus
| |
| | +----------------------------- Lens_contradens
+--+ |
| | +-- Unio_pictorum
| | +--------+
| | | +-- Unio_caffer
| | +--------------+
+--+ | | +-------- Lasmigona_costata
| | +--+
| | | +----- Alasmidonta_marginata
| | +--+
| | | +-- Strophitus_undulatus
| | +--+
+--+ +-- Pyganodon_grandis
|
| +----------------------- Gonidea_angulata
| |
| | +-- Eurynia_dilatata
| | |
| | +-----+-- Pleurobema_sintoxia
+--+ | |
| +-----------+ +-- Fusconaia_flava
| | |
| | | +----- Obliquaria_reflexa
| | +--+
+--+ | +-- Quadrula_verrucosa
| +--+
| +-- Quadrula_quadrula
|
| +----------------- Ptychobranchus_fasciolaris
+--+
| +-------------- Truncilla_truncata
+--+
| +----------- Ortmanniana_ligamentina
+--+
| +-------- Lampsilis_cardium
+--+
| +----- Cambarunio_iris
+--+
| +-- Ligumia_recta
+--+
+-- Epioblasma_triquetra
Hoeh et al., 2009 [Publ 2716], fig. 5
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 10 |
10 |
8 |
3 |
36 |
35 |
2 |
12.0 |
+-- Margaritifera_margaritifera
+--------------------+
| +-- Cumberlandia_monodonta
|
| +----- Velesunio_ambiguus
| +--------+
| | | +-- Velesunio_angasi
| | +--+
--+ +-----+ +-- Lortiella_rugata
| | |
| | | +----------- Echyridella_menziesii
| | | |
| | +--+ +-- Hyridella_depressa
| | | +-----+
| | | | +-- Hyridella_australis
| | +--+
+--+ | +----- Castalia
| +--+
| | +-- Diplodon_chilensis
| +--+
| +-- Diplodon_gratus
|
| +----------------- Pseudomulleria_dalyi **
| |
+--+ +-------------- Mulleria_rivoli **
| |
+--+ +----------- Etheria_elliptica **
| |
| | +----- Monocondylaea_minuana **
+--+ +--+
| | | +-- Anodontites_trigona **
| | +--+
| | +-- Anodontites_guanarensis **
+--+
| +-- Mutela_rostrata **
| +--+
| | +-- Mutela_dubia **
+--+
| +-- Chambardia_wahlbergi **
+--+
+-- Aspatharia_rugifera **
Huang et al., 2019 [Publ 3843], fig. 2
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 10 |
10 |
8 |
3 |
194 |
192 |
2 |
6.0 |
+----- Alathyria_jacksoni
+--+
| | +-- Alathyria_profuga
| +--+
+--------+ +-- Velesunio_ambiguus
| |
| | +----- Alathyria_pertexta
| +--+
| | +-- Lortiella_froggatti
+--------------------------------+ +--+
| | +-- Microdontia_anodontaeformis
| |
| | +-------------- Westralunio_carteri
| | |
| +--+ +----------- Echyridella_menziesii
| | |
| | | +----- Diplodon_suavidicus
| +--+ +--+
| | | | +-- Castalia_ambigua
| | | +--+
| +--+ +-- Triplodon_corrugatus
| |
| | +----- Hyridella_australis
| +--+
| | +-- Hyridella_depressa
| +--+
| +-- Hyridella_drapeta
|
| +-------- Anodontites_trapesialis **
| +--+
| | | +----- Mutela_rostrata **
| | +--+
| | | +-- Mutela_dubia **
| | +--+
| +-----------------------------------+ +-- Mutela_hargeri **
| | |
| | | +----- Mycetopoda_siliquosa **
| | | +--+
--+ | | | | +-- Tamsiella_schroeteriana **
| | | | +--+
| | +--+ +-- Lamproscapha_ensiformis **
| | |
| | | +----- Etheria_elliptica **
| | +--+
| | | +-- Aspatharia_pfeifferiana **
| | +--+
| | +-- Chambardia_wahlbergi **
| |
| | +----------------- Gibbosula_rochechouartii
| | |
| | | +-------------- Gibbosula_laosensis
| | +--------------------------+ |
| | | | | +----------- Cumberlandia_monodonta
| | | | | |
| | | +--+ | +----- Pseudunio_marocanus
| | | | +-----+
| | | | | | +-- Pseudunio_auricularius
| | | | | +--+
| | | +--+ +-- Pseudunio_homsensis
| | | |
| | | | +-- Margaritifera_dahurica
| | | | +-----+
| | | | | +-- Margaritifera_margaritifera
| | | | |
| | | +--+ +----- Margaritifera_laevis
| | | | |
| | | | | +-- Margaritifera_falcata
| | | +--+--+
+--+ | | +-- Margaritifera_middendorffi
| | |
| | | +-- Margaritifera_hembeli
| | +--+
| | +-- Margaritifera_marrianae
| |
| | +----- Trapezidens_exolescens
| | +-----------------+
| | | | +-- Trapezidens_dolichorhynchus
| | | +--+
| | +--+ +-- Trapezidens_angustior
| | | |
| | | | +-------------------- Lamellidens_corrianus
| | | | |
| | | +--+ +----------------- Lamellidens_savadiensis
| | | | |
| | | +--+ +-- Lamellidens_lamellatus
| | | | +-----------+
| | | | | +-- Lamellidens_jenkinsianus
| | | +--+
| | | | +----------- Lamellidens_generosus
| | | +--+
| | | | +-------- Lamellidens_mainwaringi[2]
| | | +--+
| | +--------------+ | +----- Lamellidens_mainwaringi[1]
| | | | +--+
| | | | | +-- Lamellidens_ferrugineus
| | | | +--+
| | | | +-- Lamellidens_marginalis
| | | |
| | | | +----- Parreysia_keralaensis
| | | | +-----------------+
| | | | | | +-- Parreysia_nagpoorensis
| | | | | +--+
| | | | | +-- Parreysia_plagiosoma
+--+ | | |
| | | | +-- Leoparreysia_tavoyensis
| | | | |
| | +--+ +-----------------+-- Leoparreysia_canefrii
| | | | |
| | | | +-- Leoparreysia_olivacea
| | | |
| | | | +-------- Coelatura_aegyptiaca[2]
| | | | |
| | | | +--------+ +----- Coelatura_aegyptiaca[1]
| | +--+ | | |
| | | | | +----- Nitia_teretiuscula
| | | | +--+
| | | | | +-- Coelatura_gabonensis
| | | | | |
| | | | +--+-- Coelatura_kunenensis
| | | | |
| | +--+ +-- Prisodontopsis_mweruensis
| | |
| | | +-------- Indonaia_shurtleffiana
| | | |
| | | +-----+ +----- Indonaia_bonneaudii
| | | | | |
| | | | +--+----- Indonaia_rugosa
| | | | |
| | | | | +-- Indonaia_andersoniana
| | +--+ +--+
| | | +-- Indonaia_caerulea
| | |
| | | +----------- Indochinella_pugio
| | | |
| | | | +----- Scabies_humilis
| | +--+ +--+
| | | | | +-- Scabies_phaselus
| | | | +--+
| | +--+ +-- Scabies_mandarinus
| | |
| | | +----- Radiatula_myitkyinae
| | +--+
| | | +-- Radiatula_chaudhurii
| | +--+
| | +-- Radiatula_mouhoti
| |
| | +-------------------------------- Acuticosta_chinensis
| | |
| | | +----------------------------- Lepidodesma_languilati
| | | |
| | | | +-------- Aculamprotula_tortuosa
| | | | |
| | | | +-----------+ +----- Aculamprotula_coreana
| | | | | | |
| | | | | +--+----- Aculamprotula_scripta
| | | | | |
+--+ | | | | +-- Aculamprotula_fibrosa
| | | | +--+
| | | | +-- Aculamprotula_tientsinensis
| +-----+ +--------+
| | | | | +-------- Unio_tumidus
| | | | | +--------+
| | | | | | | +----- Unio_crassus
| | | | | | +--+
| | | | | | | +-- Unio_delphinus
| | | | | | +--+
| | | | +--+ +-- Unio_pictorum
| | | | |
| | | | | +-- Schistodesmus_lampreyanus
| | | | | +-----------+
| | | | | | +-- Schistodesmus_spinosus
| | | | | |
| | | | +--+ +----- Nodularia_douglasiae[1]
| | +--+ | +-----+
| | | | | | +-- Nodularia_douglasiae[2]
| | | | | +--+
| | | +--+ +-- Nodularia_nuxpersicae
| | | |
| | | | +-------- Tchangsinaia_piscicula
| | | +--+
| | | | +----- Cuneopsis_rufescens
| | | +--+
| | | | +-- Cuneopsis_heudei
| | | +--+
| | | +-- Cuneopsis_celtiformis
| | |
| | | +-------- Lanceolaria_gladiola
| | | |
| | | +-----------------+ +----- Lanceolaria_grayii[1]
| | | | | |
| | | | +--+----- Lanceolaria_grayii[2]
| | | | |
| | | | | +-- Lanceolaria_lanceolata
| | | | +--+
| | | | +-- Lanceolaria_triformis
| | | |
| | | | +-------------------- Anemina_arcaeformis
| | | | |
| | | | +--+ +----------------- Cristaria_plicata
| | +--+ | | |
| | | | +--+ +-------------- Pletholophus_tenuis
| | | | | |
| | | | +--+ +-- Sinanodonta_angula
| | | | | +--------+
| | | | | | +-- Sinanodonta_lucida
| | | | +--+
| | | | | +-------- Sinanodonta_woodiana[2]
| | | | +--+
| | | | | +----- Sinanodonta_jourdyi
| | | | +--+
| | +--+ | +-- Sinanodonta_elliptica
| | | +--+
| | | +-- Sinanodonta_woodiana[1]
| | |
| | | +-------- Alasmidonta_marginata
| | | |
| | | +-------- Lasmigona_costata
| | | |
| | | +-------- Pseudanodonta_complanata
| | | |
| | | +-------- Simpsonaias_ambigua
+--+ | |
| +--------------+-------- Strophitus_undulatus
| |
| | +----- Anodonta_anatina
| +--+
| | | +-- Anodonta_cygnea
| | +--+
| | +-- Anodonta_nuttalliana
| |
| | +----- Pyganodon_grandis
| +--+
| | +-- Utterbackia_imbecillis
| +--+
| +-- Utterbackia_peninsularis
|
| +----------- Hyriopsis_desowitzi
| |
| +----------- Rectidens_sumatrensis
| |
| | +-- Hyriopsis_bialata
| | |
| | +-- Hyriopsis_myersiana
| +-----+--------+
| | | +-- Hyriopsis_panhai
| | | |
| | | +-- Hyriopsis_khoratensis
| | |
| | | +-- Ensidens_ingallsianus
| | | +-----+
| | | | +-- Ensidens_telus
| | +--+
| | | +----- Ensidens_sagittarius
| +-----------------+ +--+
| | | | +-- Ensidens_spiculus
| | | +--+
| | | +-- Ensidens_jaculus
| | |
| | | +----- Physunio
| | | +--------+
| | | | | +-- Physunio_modelli
| | | | +--+
| | | | +-- Physunio_superbus
| | | |
| | +--+ +-------- Yaukthwa_inlenensis
| | | +--+
| | | | | +----- Yaukthwa_panhai
| | | | +--+
| | | | | +-- Yaukthwa_nesemanni
| | | | +--+
| | +--+ +-- Yaukthwa_rectangularis
| | |
| | | +-- Lens_contradens
| | | +-----+
| | | | +-- Lens_micropterus
| | +--+
| | | +----- Lens_eximius
| | +--+
| | | +-- Lens_comptus
| | +--+
| | +-- Lens_novoselovi
| |
| | +-- Megalonaias_nervosa
| | +--+
+--+ | +-- Uniomerus_carolinianus
| +-----------------------+
| | | +-- Quadrula_quadrula
| | +--+
| | +-- Quadrula_verrucosa
| |
| | +-------- Elliptoideus_sloatianus
| +--+ |
| | | +-----------------+ +-- Elliptio_complanata
| | | | | +--+
| | | | | | +-- Eurynia_dilatata
| | | | +--+
| | | | | +-- Fusconaia_flava
| | +--+ +--+
| | | +-- Pleurobema_sintoxia
| | |
| | | +----------------------- Amblema_plicata
| | | |
| | | | +----- Toxolasma_parvum
| | +--+ +--------------+
| | | | | +-- Obliquaria_reflexa
| | | | +--+
| | | | +-- Plectomerus_dombeyanus
| | +--+
| | | +----------------- Potamilus_leptodon
| | | |
| | | | +-------------- Potamilus_alatus
| | +--+ |
| | | | +----------- Venustaconcha_ellipsiformis
| | | | |
| | +--+ +----------- Leaunio_lienosus
| | | |
+--+ | | +-- Ellipsaria_lineolata
| | +--------+
| +--+ +-- Truncilla_truncata
| |
| | +-- Medionidus_conradicus
| +--------+
| | +-- Ptychobranchus_subtentus
| |
| | +-------- Cambarunio_iris
| +--+
| | +----- Ortmanniana_ligamentina
| +--+
| | +-- Lampsilis_cardium
| +--+
| +-- Lampsilis_ornata
|
| +----- Chamberlainia_somsakpanhai
| +-----------------------+
| | | +-- Sinohyriopsis_cumingii
| | +--+
| | +-- Sinohyriopsis_schlegelii
| |
| | +-- Potomida_littoralis
| | +-----+
| | | +-- Pronodularia_japanensis
| | +-----------------+
+--+ | | +----- Lamprotula_leaii
| | +--+
| | | +-- Lamprotula_caveata
| | +--+
| | +-- Lamprotula_cornuumlunae
| |
| | +-------- Sinosolenaia_carinata
| | +--+
| | | | +----- Sinosolenaia_recognita
| | | +--+
+--+ | | +-- Sinosolenaia_oleivora
| | +--+
| +-----------+ +-- Sinosolenaia_iridinea
| | |
| | | +----- Gonidea_angulata
| | | +--+
| | | | | +-- Leguminaia_wheatleyi
| | | | +--+
| | +--+ +-- Microcondylaea_bonellii
| | |
| | | +-- Ptychorhynchus_pfisteri
| | | |
+--+ +-----+-- Parvasolenaia_rivularis
| |
| +-- Koreosolenaia_sitgyensis
|
| +----- Indopseudodon_salwenianus
| +--------------+
| | | +-- Indopseudodon_indawgyiensis
| | +--+
| | +-- Indopseudodon_bogani
| |
| | +-------- Pilsbryoconcha_exilis[3]
+--+ +--------+
| | | +----- Pilsbryoconcha_exilis[2]
| | +--+
| | | +-- Pilsbryoconcha_exilis[1]
| | +--+
| | +-- Pilsbryoconcha_carinifera
| |
+--+ +----------- Nyeinchanconcha_nyeinchani
| +--+
| | | +-------- Thaiconcha_callifera
| | +--+
| | | +----- Bineurus_loeiensis
| | +--+
| | | +-- Bineurus_mouhotii
+--+ +--+
| +-- Bineurus_exilis
|
| +----- Pseudodon_vagulus
| +--+
| | | +-- Pseudodon_mekongi
| | +--+
+-----+ +-- Pseudodon_vondembuschianus
|
| +-- Sundadontina_cumingii
| |
+-----+-- Sundadontina_tumida[1]
|
+-- Sundadontina_tumida[2]
Huang et al., 2019 [Publ 3843], fig. S3
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 10 |
10 |
8 |
3 |
191 |
189 |
2 |
6.0 |
+----- Velesunio_ambiguus
+--+
| | +-- Alathyria_jacksoni
| +--+
+--------+ +-- Alathyria_profuga
| |
| | +----- Alathyria_pertexta
| +--+
| | +-- Lortiella_froggatti
+-----------------------------------------------+ +--+
| | +-- Microdontia_anodontaeformis
| |
| | +-------------- Westralunio_carteri
| | |
| +--+ +----------- Echyridella_menziesii
| | |
| | | +----- Diplodon_suavidicus
| +--+ +--+
| | | | +-- Castalia_ambigua
| | | +--+
| +--+ +-- Triplodon_corrugatus
| |
| | +----- Hyridella_australis
| +--+
| | +-- Hyridella_depressa
| +--+
| +-- Hyridella_drapeta
|
| +-------- Anodontites_trapesialis **
| +--+
| | | +----- Mutela_rostrata **
| | +--+
| | | +-- Mutela_dubia **
| | +--+
| +--------------------------------------------------+ +-- Mutela_hargeri **
| | |
| | | +----- Mycetopoda_siliquosa **
--+ | | +--+
| | | | | +-- Tamsiella_schroeteriana **
| | | | +--+
| | +--+ +-- Lamproscapha_ensiformis **
| | |
| | | +----- Etheria_elliptica **
| | +--+
| | | +-- Aspatharia_pfeifferiana **
| | +--+
| | +-- Chambardia_wahlbergi **
| |
| | +-------------------- Gibbosula_rochechouartii
| | |
| | | +----------------- Gibbosula_laosensis
| | +--------------------------------------+ |
| | | | | +-------- Cumberlandia_monodonta
| | | | | +-----+
| | | +--+ | | +----- Pseudunio_marocanus
| | | | | +--+
| | | | | | +-- Pseudunio_auricularius
| | | | | +--+
| | | +--+ +-- Pseudunio_homsensis
| | | |
| | | | +-- Margaritifera_dahurica
| | | | +--------+
| | | | | +-- Margaritifera_margaritifera
| | | +--+
| | | | +-------- Margaritifera_falcata
+--+ | | |
| | +--+ +-- Margaritifera_hembeli
| | | +--+
| | | | +-- Margaritifera_marrianae
| | +--+
| | | +-- Margaritifera_laevis
| | +--+
| | +-- Margaritifera_middendorffi
| |
| | +----- Trapezidens_exolescens
| | +-----+
| | | | +-- Trapezidens_angustior
| | | +--+
| | | +-- Trapezidens_dolichorhynchus
| | |
| | +--------------+ +-- Lamellidens_corrianus
| | | | +-----+
| | | | | +-- Lamellidens_savadiensis
| | | | |
| | | | | +----- Lamellidens_lamellatus
| | | +--+ |
| | | | +----- Lamellidens_jenkinsianus
| | | | |
| | | | | +-- Lamellidens_marginalis
| | | +--+--+
| | | | +-- Lamellidens_ferrugineus
| | | |
| | +-----------------------------+ | +-- Lamellidens_mainwaringi[1]
| | | | | |
| | | | +--+-- Lamellidens_mainwaringi[2]
| | | | |
| | | | +-- Lamellidens_generosus
+--+ | |
| | | +----- Parreysia_keralaensis
| | | +-----------------+
| | | | | +-- Parreysia_nagpoorensis
| | | | +--+
| | | | +-- Parreysia_plagiosoma
| | | |
| | | | +----- Leoparreysia_canefrii
| | +--+ +--------------+
| | | | | +-- Leoparreysia_tavoyensis
| | | | +--+
| | | | +-- Leoparreysia_olivacea
| | | |
| | | | +----------- Coelatura_aegyptiaca[2]
| | | | |
| | +--+ +-----+ +-- Coelatura_aegyptiaca[1]
| | | | | +-----+
| | | | | | +-- Nitia_teretiuscula
| | | | +--+
| | | | | +----- Coelatura_kunenensis
| | | | +--+
| | | | | +-- Coelatura_gabonensis
| | | | +--+
| | +--+ +-- Prisodontopsis_mweruensis
| | |
| | | +-------- Indonaia_shurtleffiana
| | | |
| | | +-----+ +----- Indonaia_bonneaudii
| | | | | |
| | | | +--+----- Indonaia_rugosa
| | | | |
| | | | | +-- Indonaia_andersoniana
| | +--+ +--+
| | | +-- Indonaia_caerulea
| | |
| | | +----------- Indochinella_pugio
| | | |
| | | | +----- Scabies_phaselus
| | +--+ +--+
| | | | | +-- Scabies_humilis
| | | | +--+
| | +--+ +-- Scabies_mandarinus
| | |
| | | +----- Radiatula_mouhoti
| | +--+
| | | +-- Radiatula_myitkyinae
| | +--+
+--+ +-- Radiatula_chaudhurii
|
| +----- Indopseudodon_indawgyiensis
| +--------------+
| | | +-- Indopseudodon_bogani
| | +--+
| | +-- Indopseudodon_salwenianus
| |
| | +-------- Pilsbryoconcha_exilis[3]
| +--------------------------------+ +--------+
| | | | | +----- Pilsbryoconcha_exilis[2]
| | | | +--+
| | | | | +-- Pilsbryoconcha_carinifera
| | | | +--+
| | | | +-- Pilsbryoconcha_exilis[1]
| | | |
| | +--+ +----------- Sundadontina_tumida
| | | +--+
| | | | | +-------- Pseudodon_vagulus
| | | | +--+
| | | | | +----- Pseudodon_vondembuschianus[1]
| | | | +--+
| | | | | +-- Pseudodon_mekongi
| | +--+ +--+
| | | +-- Pseudodon_vondembuschianus[2]
| | |
| | | +-- Nyeinchanconcha_nyeinchani
| | | +--------+
| | | | +-- Sundadontina_cumingii
| | +--+
| | | +-------- Thaiconcha_callifera
| | +--+
| | | +----- Bineurus_loeiensis
| | +--+
| | | +-- Bineurus_mouhotii
| | +--+
| | +-- Bineurus_exilis
| |
| | +-- Ensidens_telus
| | +-----+
| | | +-- Ensidens_ingallsianus
| | +-----+
| | | | +----- Ensidens_sagittarius
| | | +--+
| | | | +-- Ensidens_spiculus
| | +--+ +--+
| | | | +-- Ensidens_jaculus
| | | |
| | | | +----------- Hyriopsis_desowitzi
| | | | |
| | | +--+ +-------- Rectidens_sumatrensis
| | | | |
| | | +--+ +-- Hyriopsis_panhai
+--+ | | +--+
| | | | +-- Hyriopsis_khoratensis
| +--------------------------------+ +--+
| | | | +-- Hyriopsis_bialata
| | | +--+
| | | +-- Hyriopsis_myersiana
| | |
| | | +----- Physunio
| | | +--------+
| | | | | +-- Physunio_superbus
| | | | +--+
| | | | +-- Physunio_modelli
| | | |
| | +--+ +-------- Yaukthwa_inlenensis
| | | +--+
| | | | | +----- Yaukthwa_panhai
| | | | +--+
| | | | | +-- Yaukthwa_nesemanni
| | | | +--+
| | +--+ +-- Yaukthwa_rectangularis
| | |
| | | +-- Lens_contradens
| | | +-----+
| | | | +-- Lens_micropterus
| | +--+
| | | +----- Lens_eximius
| | +--+
| | | +-- Lens_novoselovi
| | +--+
| | +-- Lens_comptus
| |
| | +----- Chamberlainia_somsakpanhai
| | +--------------------------------------+
| | | | +-- Sinohyriopsis_cumingii
| | | +--+
| | | +-- Sinohyriopsis_schlegelii
| | |
| | | +-- Potomida_littoralis
| | | +-----+
| | | | +-- Pronodularia_japanensis
| | | +-----+
| | | | | +----- Lamprotula_leaii
| | | | +--+
| | | | | +-- Lamprotula_caveata
+--+ | | +--+
| | | +-- Lamprotula_cornuumlunae
| +--+ +--------------------------+
| | | | | +----- Sinosolenaia_recognita
| | | | | +-----+
| | | | | | | +-- Sinosolenaia_oleivora
| | | | | | +--+
| | | | | | +-- Sinosolenaia_iridinea
| | | | +--+
| | | | | +----- Gonidea_angulata
| | | | | +--+
| | | | | | | +-- Leguminaia_wheatleyi
| | | | | | +--+
| | | | +--+ +-- Microcondylaea_bonellii
| | | | |
| | | | | +----- Ptychorhynchus_pfisteri
| | +--+ +--+
| | | | +-- Parvasolenaia_rivularis
| | | +--+
| | | +-- Koreosolenaia_sitgyensis
| | |
| | | +-- Megalonaias_nervosa
| | | +--+
| | | | +-- Uniomerus_carolinianus
| | | +--------------------------------+
| | | | | +-- Quadrula_quadrula
| | | | +--+
| | | | +-- Quadrula_verrucosa
| | | |
| | | | +-------- Elliptoideus_sloatianus
| | +--+ |
| | | +--------------------------+ +-- Elliptio_complanata
| | | | | +--+
| | | | | | +-- Eurynia_dilatata
| | | | +--+
| | | | | +-- Fusconaia_flava
| | +--+ +--+
| | | +-- Pleurobema_sintoxia
| | |
| | | +-------------------------------- Amblema_plicata
| | | |
| | +--+ +----------------------------- Toxolasma_parvum
| | | |
+--+ +--+ +-- Obliquaria_reflexa
| | +-----------------------+
| | | +-- Plectomerus_dombeyanus
| +--+
| | +----------------------- Potamilus_leptodon
| | |
| +--+ +-------------------- Potamilus_alatus
| | |
| | | +-- Medionidus_conradicus
| +--+ +--------------+
| | | +-- Ptychobranchus_subtentus
| | |
| +--+ +-- Ellipsaria_lineolata
| | +-----------+
| | | +-- Truncilla_truncata
| +--+
| | +----------- Leaunio_lienosus
| | |
| +--+ +-- Venustaconcha_ellipsiformis
| | +-----+
| | | +-- Cambarunio_iris
| +--+
| | +----- Ortmanniana_ligamentina
| +--+
| | +-- Lampsilis_cardium
| +--+
| +-- Lampsilis_ornata
|
| +----------------------------------- Acuticosta_chinensis
| |
| | +-------- Aculamprotula_tortuosa
| | |
| | +-----------+ +-- Aculamprotula_coreana
| | | | +--+
| | | | | +-- Aculamprotula_scripta
| | | +--+
| | | | +-- Aculamprotula_fibrosa
| | | +--+
| | | +-- Aculamprotula_tientsinensis
| | +-----------+
| | | | +-------- Unio_tumidus
+-----------+ | | +--------+
| | | | | +----- Unio_crassus
| | | | +--+
| | | | | +-- Unio_delphinus
| | | | +--+
| | +--+ +-- Unio_pictorum
| | |
| | | +----- Nodularia_douglasiae[1]
| | | +--------+
| | | | | +-- Nodularia_douglasiae[2]
| | | | +--+
| | +--+ +-- Nodularia_nuxpersicae
| | |
+--+ | +----------- Schistodesmus_lampreyanus
| +--+
| | +-------- Tchangsinaia_piscicula
| +--+
| | +----- Cuneopsis_rufescens
| +--+
| | +-- Cuneopsis_heudei
| +--+
| +-- Cuneopsis_celtiformis
|
| +----------------------------- Lepidodesma_languilati
| |
| | +-------- Lanceolaria_gladiola
| | +-----------------+
| | | | +----- Lanceolaria_grayii
| | | +--+
+--+ | | +-- Lanceolaria_lanceolata
| | +--+
| | +-- Lanceolaria_triformis
| |
| | +-------------------- Anemina_arcaeformis
| | |
| | +--+ +----------------- Cristaria_plicata
+--+ | | |
| | +--+ +-------------- Pletholophus_tenuis
| | | |
| | +--+ +-- Sinanodonta_angula
| | | +--------+
| | | | +-- Sinanodonta_lucida
| | +--+
| | | +-------- Sinanodonta_woodiana[2]
| | +--+
| | | +----- Sinanodonta_jourdyi
+--+ +--+
| | +-- Sinanodonta_elliptica
| +--+
| +-- Sinanodonta_woodiana[1]
|
| +-- Alasmidonta_marginata
| +--------+
| | +-- Lasmigona_costata
| |
| | +----- Anodonta_anatina
| +-----+
| | | +-- Anodonta_cygnea
+-----------+ +--+
| +-- Anodonta_nuttalliana
|
| +----- Pseudanodonta_complanata
| +--+
| | | +-- Simpsonaias_ambigua
| | +--+
+--+ +-- Strophitus_undulatus
|
| +----- Pyganodon_grandis
+--+
| +-- Utterbackia_imbecillis
+--+
+-- Utterbackia_peninsularis
Huang et al., 2019 [Publ 3843], fig. S4
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 10 |
10 |
6 |
3 |
192 |
190 |
2 |
6.0 |
+----------- Etheria_elliptica **
|
| +-- Aspatharia_pfeifferiana **
+--------+
| +-- Chambardia_wahlbergi **
|
+-----------------------+ +----- Mycetopoda_siliquosa **
| +-----+
| | | +-- Tamsiella_schroeteriana **
| | +--+
| | +-- Lamproscapha_ensiformis **
| |
| | +-------- Anodontites_trapesialis **
| +--+
| | +----- Mutela_hargeri **
| +--+
| | +-- Mutela_dubia **
| +--+
| +-- Mutela_rostrata **
|
| +----------- Echyridella_menziesii
| |
| +----------- Westralunio_carteri
| |
| | +----- Diplodon_suavidicus
| +-----+
| | | +-- Castalia_ambigua
| | +--+
| | +-- Triplodon_corrugatus
| |
| +--------------------+ +----- Hyridella_australis
| | +-----+
| | | | +-- Hyridella_depressa
| | | +--+
| | | +-- Hyridella_drapeta
| | |
| | | +----- Velesunio_ambiguus
| | | +--+
--+ | | | | +-- Alathyria_jacksoni
| | | | +--+
| | +--+ +-- Alathyria_profuga
| | |
| | | +----- Alathyria_pertexta
| | +--+
| | | +-- Lortiella_froggatti
| | +--+
| | +-- Microdontia_anodontaeformis
| |
| | +-------------------- Gibbosula_rochechouartii
| | |
| | +--------+ +----------------- Gibbosula_laosensis
| | | | |
| | | +--+ +-------------- Cumberlandia_monodonta
| | | | |
| | | | | +----- Pseudunio_marocanus
| | | +--+ +-----+
| | | | | | +-- Pseudunio_auricularius
| | | | | +--+
| | | | | +-- Pseudunio_homsensis
| | | +--+
| | | | +-------- Margaritifera_falcata
| | | | |
| | | | | +-- Margaritifera_dahurica
| | | | +-----+
| | | +--+ +-- Margaritifera_margaritifera
| | | |
| | | | +----- Margaritifera_laevis
| | | | |
| | | +--+----- Margaritifera_middendorffi
| | | |
+--+ | | +-- Margaritifera_hembeli
| | +--+
| | +-- Margaritifera_marrianae
| |
| | +-- Parreysia_keralaensis
| | |
| | +--------------------+-- Parreysia_nagpoorensis
| | | |
| | | +-- Parreysia_plagiosoma
| | |
| | | +----- Trapezidens_exolescens
| | | +--------+
| | | | | +-- Trapezidens_angustior
| | | | +--+
| | +--------+ +-- Trapezidens_dolichorhynchus
| | | |
| | | | +----------- Lamellidens_corrianus
| | | | |
| | | +--+ +-------- Lamellidens_savadiensis
| | | | |
| | | | | +----- Lamellidens_marginalis
| | | +--+ |
| | | | +----- Lamellidens_ferrugineus
| | | | |
| | +--+ +--+ +-- Lamellidens_mainwaringi[1]
| | | | | |
| | | | | +-- Lamellidens_mainwaringi[2]
| | | | | |
| | | | +--+-- Lamellidens_lamellatus
| | | | |
| | | | +-- Lamellidens_generosus
| | | | |
| | | | +-- Lamellidens_jenkinsianus
| | | |
| | | | +----- Leoparreysia_tavoyensis
| | | | +--------------+
| | | | | | +-- Leoparreysia_canefrii
| | | | | +--+
| | | | | +-- Leoparreysia_olivacea
| | | | |
+--+ | | | +-------- Coelatura_aegyptiaca[2]
| | | | |
| | | | +-------- Nitia_teretiuscula
| | +--+ +--------+
| | | | | +----- Coelatura_aegyptiaca[1]
| | | | | |
| | | | +--+ +-- Coelatura_gabonensis
| | | | | |
| | | | +--+-- Coelatura_kunenensis
| | | | |
| | | | +-- Prisodontopsis_mweruensis
| | +--+
| | | +----- Indonaia_shurtleffiana
| | | |
| | | | +-- Indonaia_bonneaudii
| | | +--------+--+
| | | | | +-- Indonaia_rugosa
| | | | |
| | | | | +-- Indonaia_andersoniana
| | | | +--+
| | +--+ +-- Indonaia_caerulea
| | |
| | | +----------- Indochinella_pugio
| | | |
| | | | +----- Scabies_humilis
| | +--+ +--+
| | | | | +-- Scabies_phaselus
| | | | +--+
| | +--+ +-- Scabies_mandarinus
| | |
| | | +----- Radiatula_mouhoti
| | +--+
| | | +-- Radiatula_myitkyinae
| | +--+
| | +-- Radiatula_chaudhurii
| |
| | +-------------------- Chamberlainia_somsakpanhai
| | |
| | +-------------------- Megalonaias_nervosa
| | |
| | +-------------------- Uniomerus_carolinianus
| | |
| | | +-- Quadrula_quadrula
| | +-----------------+
| | | +-- Quadrula_verrucosa
| | |
| | | +-- Sinohyriopsis_cumingii
| | +-----------------+
| | | +-- Sinohyriopsis_schlegelii
| | |
| | | +-- Potomida_littoralis
| | | +-----+
| | | | +-- Pronodularia_japanensis
| | +-----------+
| | | | +----- Lamprotula_leaii
| | | +--+
| | | | +-- Lamprotula_caveata
| | | +--+
| | | +-- Lamprotula_cornuumlunae
| | |
| | | +-------- Sinosolenaia_carinata
+--+ | |
| | | +----- Gonidea_angulata
| | +--+
| | | | +-- Leguminaia_wheatleyi
| | | +--+
| | | +-- Microcondylaea_bonellii
| +-----------+
| | | +----- Ptychorhynchus_pfisteri
| | +--+
| | | | +-- Parvasolenaia_rivularis
| | | +--+
| | | +-- Koreosolenaia_sitgyensis
| | |
| | | +----- Sinosolenaia_recognita
| | +--+
| | | +-- Sinosolenaia_iridinea
| | +--+
| | +-- Sinosolenaia_oleivora
| |
| | +----- Indopseudodon_salwenianus
| | +-----+
| | | | +-- Indopseudodon_indawgyiensis
| | | +--+
| | | +-- Indopseudodon_bogani
| | |
| | | +-------- Nyeinchanconcha_nyeinchani
| | | |
| | | +-------- Thaiconcha_callifera
| +--------+ |
| | | +-------- Sundadontina_cumingii
| | | |
| | | +-------- Sundadontina_tumida[1]
| | | |
| | | +-------- Sundadontina_tumida[2]
| | | |
| | | | +----- Pilsbryoconcha_exilis[2]
| | +--+--+
| | | | +-- Pilsbryoconcha_carinifera
| | | +--+
| | | +-- Pilsbryoconcha_exilis[1]
| | |
| | | +----- Pseudodon_vagulus
| | +--+
| | | | +-- Pseudodon_mekongi
| | | +--+
| | | +-- Pseudodon_vondembuschianus
| | |
| | | +----- Bineurus_loeiensis
| | +--+
| | | +-- Bineurus_mouhotii
| | +--+
| | +-- Bineurus_exilis
| |
| | +-------------- Amblema_plicata
| | |
| | +-------------- Toxolasma_parvum
| | |
| | | +-- Obliquaria_reflexa
| | +-----------+
| | | +-- Plectomerus_dombeyanus
| | |
| | | +----------- Elliptoideus_sloatianus
| | +--+
| | | | +-------- Elliptio_complanata
| | | +--+
| | | | +----- Eurynia_dilatata
| +-----+ +--+
| | | | +-- Fusconaia_flava
| | | +--+
| | | +-- Pleurobema_sintoxia
| | |
| | | +-------- Ellipsaria_lineolata
| | | |
| | | +-------- Potamilus_leptodon
| | | |
| | | +-------- Medionidus_conradicus
| | | |
+-----+ | +-------- Potamilus_alatus
| | |
| | +-------- Ptychobranchus_subtentus
| +-----+
| +-------- Truncilla_truncata
| |
| +-------- Venustaconcha_ellipsiformis
| |
| +-------- Cambarunio_iris
| |
| +-------- Leaunio_lienosus
| |
| | +----- Ortmanniana_ligamentina
| +--+
| | +-- Lampsilis_cardium
| +--+
| +-- Lampsilis_ornata
|
| +-------------- Hyriopsis_desowitzi
| |
| +-------------- Hyriopsis_myersiana
| |
| +-------------- Hyriopsis_panhai
| |
| +--+ +-- Hyriopsis_bialata
| | +-----------+
| | | +-- Hyriopsis_khoratensis
| | |
| | | +----------- Rectidens_sumatrensis
| | | |
| | +--+ +-- Ensidens_telus
| | | +-----+
| | | | +-- Ensidens_ingallsianus
| | +--+
| | | +----- Ensidens_sagittarius
| | +--+
+--+ | +-- Ensidens_spiculus
| | +--+
| | +-- Ensidens_jaculus
| |
| | +----------- Physunio_modelli
| | |
| | +----------- Lens
| | |
| | +----------- Physunio_superbus
| | |
| | | +-------- Yaukthwa_inlenensis
| | +--+
| +-----+ | +----- Yaukthwa_panhai
| | +--+
| | | +-- Yaukthwa_nesemanni
| | +--+
| | +-- Yaukthwa_rectangularis
| |
| | +-- Lens_comptus
| | +-----+
| | | +-- Lens_novoselovi
| +--+
| | +----- Lens_eximius
| +--+
| | +-- Lens_contradens
| +--+
| +-- Lens_micropterus
|
| +----------------- Acuticosta_chinensis
| |
| +----------------- Alasmidonta_marginata
| |
| +----------------- Anodonta_anatina
| |
| +----------------- Anemina_arcaeformis
| |
| +----------------- Anodonta_cygnea
| |
| +----------------- Cristaria_plicata
| |
| +----------------- Anodonta_nuttalliana
| |
| +----------------- Lasmigona_costata
| |
| +----------------- Lepidodesma_languilati
| |
| +----------------- Pseudanodonta_complanata
| |
| +----------------- Pyganodon_grandis
| |
| +----------------- Simpsonaias_ambigua
| |
| +----------------- Strophitus_undulatus
| |
| | +-- Schistodesmus_lampreyanus
| +--------------+
| | +-- Schistodesmus_spinosus
| |
| | +-- Utterbackia_imbecillis
| +--------------+
| | +-- Utterbackia_peninsularis
| |
| | +-------- Lanceolaria_gladiola
| +--------+
| | | +----- Lanceolaria_grayii
| | +--+
+--+ | +-- Lanceolaria_lanceolata
| +--+
| +-- Lanceolaria_triformis
|
| +-------- Unio_tumidus
+--------+
| | +----- Unio_crassus
| +--+
| | +-- Unio_delphinus
| +--+
| +-- Unio_pictorum
|
| +-------- Aculamprotula_tortuosa
| |
+--------+ +-- Aculamprotula_coreana
| | +--+
| | | +-- Aculamprotula_scripta
| +--+
| | +-- Aculamprotula_fibrosa
| +--+
| +-- Aculamprotula_tientsinensis
|
| +-------- Cuneopsis_heudei
| |
| +-------- Cuneopsis_celtiformis
| |
+--------+-------- Tchangsinaia_piscicula
| |
| +-------- Cuneopsis_rufescens
| |
| | +----- Nodularia_douglasiae[2]
| +--+
| | +-- Nodularia_douglasiae[1]
| +--+
| +-- Nodularia_nuxpersicae
|
| +-------------- Pletholophus_tenuis
| |
+--+ +-- Sinanodonta_angula
| +--------+
| | +-- Sinanodonta_lucida
+--+
| +-------- Sinanodonta_woodiana[1]
+--+
| +----- Sinanodonta_jourdyi
+--+
| +-- Sinanodonta_elliptica
+--+
+-- Sinanodonta_woodiana[2]
Walker et al., 2006 [Publ 2666], fig. 6
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 9 |
9 |
7 |
3 |
29 |
28 |
2 |
11.0 |
+----------------- Castalia_stevensi
|
+----------------- Diplodon_gratus
+--------+
| | +-------------- Hyridella_depressa
| | |
| | +-------------- Hyridella_australis
| +--+
| | +----------- Echyridella_menziesii
| | |
| | | +-- Velesunio_ambiguus
| +--+ +-----+
| | | +-- Velesunio[3]
| | |
| +--+ +----- Velesunio[2]
| | |
| | | +-- Alathyria_jacksoni
| +--+--+
| | +-- Velesunio[1]
| |
--+ | +-- Velesunio_angasi
| +--+
| +-- Lortiella_rugata
|
| +-- Mutela_dubia **
| +--------------+
| | +-- Mutela_rostrata **
| +-----+
| | | +-------------- Etheria_elliptica **
| | | |
| | +--+ +----------- Chambardia_rubens **
| | | |
| | +--+ +-------- Mulleria_rivoli **
| | | |
| | +--+ +-- Anodontites_guanarensis **
| | | +--+
| | | | +-- Anodontites_trigona **
+--+ +--+
| | +-- Tamsiella_tamsiana **
| +--+
| +-- Monocondylaea_minuana **
|
| +-- Pyganodon_fragilis
| +-----------------+
| | +-- Pyganodon_grandis
| |
| | +----------- Cumberlandia_monodonta
| | |
+--+ +-----+ +-------- Pseudunio_auricularius
| | | |
| | +--+ +-- Margaritifera_margaritifera
| | | +--+
| | | | +-- Margaritifera_dahurica
| | +--+
+--+ | +-- Margaritifera_falcata
| +--+
| +-- Margaritifera_middendorffi
|
| +-- Gonidea_angulata
| +-----------+
| | +-- Sundadontina_tumida
+--+
| +----------- Cyrtonaias_tampicoensis
| |
+--+----------- Fusconaia_flava
|
| +-------- Potamilus_purpuratus
+--+
| +----- Lampsilis_teres
+--+
| +-- Ligumia_recta
+--+
+-- Ortmanniana_ligamentina
Bogan & Hoeh, 2000 [Publ 2650], fig. 1
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 8 |
28 |
21 |
3 |
26 |
25 |
2 |
29.0 |
+-- Castalia_stevensi
+-----+
| +-- Diplodon_gratus
+--------------------+
| | +----- Echyridella_menziesii
| +--+
| | +-- Velesunio_angasi
| +--+
| +-- Lortiella_rugata
|
| +-- Mutela_dubia **
--+ +-----------+
| | +-- Mutela_rostrata **
| +-----------+
| | | +----------- Etheria_elliptica **
| | +--+
| | | +-------- Mulleria_rivoli **
| | +--+
| | | +----- Monocondylaea_minuana **
| | +--+
+--+ | +-- Anodontites_trigona **
| +--+
| +-- Anodontites_guanarensis **
|
| +----------------------- Coelatura_aegyptiaca
| |
| | +-------------------- Pseudomulleria_dalyi **
+--+ |
| | +----- Anodonta_cygnea
| | +-----------+
+--+ | | +-- Strophitus_undulatus
| | +--+
| | +-- Pyganodon_grandis
| |
+--+ +-------------- Gonidea_angulata
| |
| | +-- Unio_pictorum
| +-----------+
| | +-- Unio_tumidus
| |
+--+ +-- Cumberlandia_monodonta
| +--------+
| | +-- Margaritifera_margaritifera
| |
| | +-------- Toxolasma_lividum
| | |
+--+ +-------- Amblema_plicata
| |
| | +-- Glebula_rotundata
| +-----+
| | +-- Cyrtonaias_tampicoensis
| |
+--+ +----- Potamilus_alatus
+--+
| | +-- Ortmanniana_ligamentina
| +--+
| +-- Ligumia_recta
|
| +----- Quadrula_quadrula
| |
+--+ +-- Eurynia_dilatata
| |
+--+-- Pleurobema_clava
|
+-- Fusconaia_flava
Guerra et al., 2019 [Publ 4168], fig. 1
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 7 |
7 |
5 |
2 |
34 |
33 |
2 |
9.0 |
+----------------- Mutela_dubia **
+--------------+
| | +-------------- Chambardia_rubens **
| +--+
| | +----------- Anodontites_trapesialis **
| +--+
| | +-------- Monocondylaea_parchappii **
| +--+
| | +----- Lamproscapha_ensiformis **
| +--+
| | +-- Tamsiella_schroeteriana **
| +--+
| +-- Fossula_fossiculifera **
--+
| +-- Echyridella_menziesii
| +-----+
| | +-- Westralunio_carteri
| +--------------------+
| | | +----- Diplodon_suavidicus
| | +--+
| | | +-- Castalia_ambigua
| | +--+
| | +-- Prisodon_obliquus
| |
| | +-------------- Gibbosula_crassa
| | |
+--+ +-----------+ +----------- Cumberlandia_monodonta
| | | |
| | +--+ +-- Pseudunio_marocanus
| | | +-----+
| | | | +-- Pseudunio_auricularius
| | +--+
| | | +----- Margaritifera_falcata
| | +--+
| | | +-- Margaritifera_dahurica
| | +--+
+--+ +-- Margaritifera_margaritifera
|
| +-------- Quadrula_quadrula
| +--------------+
| | | +----- Toxolasma_parvum
| | +--+
| | | +-- Venustaconcha_ellipsiformis
| | +--+
| | +-- Potamilus_alatus
| |
+--+ +-------- Sinosolenaia_carinata
| +-----------+
| | | +----- Pronodularia_japanensis
| | +--+
| | | +-- Lamprotula_leaii
| | +--+
| | +-- Potomida_littoralis
| |
+--+ +-------------- Lanceolaria_lanceolata
| +--+
| | | +----------- Aculamprotula_tortuosa
| | +--+
| | | +-------- Schistodesmus_lampreyanus
| | +--+
| | | +----- Unio_tumidus
+--+ +--+
| | +-- Unio_crassus
| +--+
| +-- Unio_delphinus
|
| +-------------- Sinanodonta_woodiana
| |
+--+ +----------- Anodonta_anatina
| |
+--+ +-- Platynaias_compressa
| +-----+
| | +-- Platynaias_subviridis
+--+
| +----- Pyganodon_grandis
+--+
| +-- Utterbackia_peninsularis
+--+
+-- Utterbackia_imbecillis
Guerra et al., 2019 [Publ 4168], fig. 2
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 7 |
7 |
5 |
2 |
34 |
33 |
2 |
11.0 |
+----------------- Chambardia_rubens **
+--------------------+
| | +-------------- Mutela_dubia **
| +--+
| | +----------- Anodontites_trapesialis **
| +--+
| | +-------- Monocondylaea_parchappii **
| +--+
| | +----- Lamproscapha_ensiformis **
| +--+
| | +-- Tamsiella_schroeteriana **
| +--+
| +-- Fossula_fossiculifera **
--+
| +----------- Westralunio_carteri
| +-----------------------+
| | | +-------- Echyridella_menziesii
| | +--+
| | | +----- Diplodon_suavidicus
| | +--+
| | | +-- Castalia_ambigua
| | +--+
| | +-- Prisodon_obliquus
| |
| | +-------------- Gibbosula_crassa
+--+ |
| +-----------------+ +----------- Cumberlandia_monodonta
| | | |
| | +--+ +-- Pseudunio_marocanus
| | | +-----+
| | | | +-- Pseudunio_auricularius
| | +--+
| | | +----- Margaritifera_falcata
| | +--+
| | | +-- Margaritifera_dahurica
+--+ +--+
| +-- Margaritifera_margaritifera
|
| +-------- Quadrula_quadrula
| +--------------------+
| | | +----- Toxolasma_parvum
| | +--+
| | | +-- Venustaconcha_ellipsiformis
| | +--+
| | +-- Potamilus_alatus
+--+
| +-------- Sinosolenaia_carinata
| +-----------------+
| | | +----- Pronodularia_japanensis
| | +--+
| | | +-- Lamprotula_leaii
| | +--+
+--+ +-- Potomida_littoralis
|
| +----------------------- Aculamprotula_tortuosa
| |
| | +-------- Schistodesmus_lampreyanus
+--+ +-----------+
| | | +----- Unio_tumidus
| | +--+
| | | +-- Unio_crassus
+--+ +--+
| +-- Unio_delphinus
|
| +----------------- Lanceolaria_lanceolata
| |
+--+ +-------------- Sinanodonta_woodiana
| |
+--+ +----------- Anodonta_anatina
| |
+--+ +-- Platynaias_compressa
| +-----+
| | +-- Platynaias_subviridis
+--+
| +----- Pyganodon_grandis
+--+
| +-- Utterbackia_peninsularis
+--+
+-- Utterbackia_imbecillis
Graf, 2000 [Publ 2647], fig. 1
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 6 |
6 |
3 |
3 |
12 |
11 |
2 |
7.5 |
+-------------- Grandidieria
|
| +-------- Castalia
| +--+
--+ | | +----- Diplodon
| | +--+
| | | +-- Hyridella
| | +--+
+--+ +-- Velesunio
|
| +-- Mutela[1] **
| +-----+
| | +-- Mutela[2] **
+--+
| +----- Anodontites **
| |
+--+----- Mycetopoda **
|
| +-- Etheria **
+--+
+-- Mulleria **
Combosch et al., 2017 [Publ 3806], fig. 1
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 6 |
6 |
4 |
3 |
9 |
7 |
2 |
7.0 |
+----- Unio_pictorum
+-----------+
| | +-- Lampsilis_cardium
| +--+
--+ +-- Scabies_crispata
|
| +-------------- Margaritifera_margaritifera
| |
+--+ +-- Chambardia_wahlbergi **
| +--------+
| | +-- Aspatharia_pfeifferiana **
+--+
| +-------- Etheria_elliptica **
+--+
| +----- Mutela_hargeri **
+--+
| +-- Tamsiella_schroeteriana **
+--+
+-- Lamproscapha_ensiformis **
Combosch et al., 2017 [Publ 3806], fig. 2
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 6 |
6 |
4 |
3 |
9 |
7 |
2 |
7.0 |
+-- Chambardia_wahlbergi **
+--------+
| +-- Aspatharia_pfeifferiana **
+--+
| | +-------- Etheria_elliptica **
| +--+
| | +----- Mutela_hargeri **
| +--+
| | +-- Tamsiella_schroeteriana **
--+ +--+
| +-- Lamproscapha_ensiformis **
|
| +----- Velesunio_ambiguus
| +-----+
| | | +-- Triplodon_corrugatus
| | +--+
+--+ +-- Hyridella_australis
|
| +-------- Margaritifera_margaritifera
+--+
| +----- Unio_pictorum
+--+
| +-- Scabies_crispata
+--+
+-- Lampsilis_cardium
Huang et al., 2018 [Publ 3828], fig. 3, 5
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 6 |
6 |
4 |
3 |
23 |
21 |
2 |
3.0 |
+----- Triplodon_corrugatus
+-----------------------+
| | +-- Velesunio_ambiguus
| +--+
| +-- Hyridella_australis
|
| +----------- Mutela_hargeri **
| |
| +--------------+ +-- Aspatharia_pfeifferiana **
--+ | | +-----+
| | | | +-- Chambardia_wahlbergi **
| | +--+
| | | +----- Etheria_elliptica **
| | +--+
| | | +-- Tamsiella_schroeteriana **
| | +--+
| | +-- Lamproscapha_ensiformis **
| |
+--+ +-- Potomida_littoralis
| +-----+
| | +-- Lamprotula_leaii
| |
| +--------------+ +-- Lampsilis_cardium
| | | +--+
| | | | +-- Scabies_crispata
| | +--+
| | | +-- Unio_pictorum
| | +--+
+--+ +-- Aculamprotula_tientsinensis
|
| +-- Gibbosula_rochechouartii
| +-----------------+
| | +-- Gibbosula_laosensis
| |
| | +----- Cumberlandia_monodonta
+--+ +-----------+
| | | +-- Pseudunio_auricularius
| | +--+
| | +-- Pseudunio_marocanus
+--+
| +-- Margaritifera_margaritifera
| +-----------+
| | +-- Margaritifera_dahurica
+--+
| +----------- Margaritifera_falcata
+--+
| +-------- Margaritifera_laevis
+--+
| +----- Margaritifera_hembeli
+--+
| +-- Margaritifera_middendorffi
+--+
+-- Margaritifera_marrianae
Lopes-Lima et al., 2018 [Publ 3829], fig. 1
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 6 |
6 |
4 |
3 |
22 |
20 |
2 |
3.0 |
+----- Triplodon_corrugatus
+--------------------+
| | +-- Velesunio_ambiguus
| +--+
| +-- Hyridella_australis
|
| +----------- Mutela_hargeri **
| |
--+ +-----------+ +-- Aspatharia_pfeifferiana **
| | | +-----+
| | | | +-- Chambardia_wahlbergi **
| | +--+
| | | +----- Etheria_elliptica **
| | +--+
| | | +-- Lamproscapha_ensiformis **
| | +--+
+--+ +-- Tamsiella_schroeteriana **
|
| +----- Potomida_littoralis
| +--------------+
| | | +-- Lampsilis_cardium
| | +--+
| | +-- Unio_pictorum
| |
+--+ +----- Gibbosula_rochechouartii
| +-----------+
| | | +-- Gibbosula_crassa
| | +--+
| | +-- Gibbosula_laosensis
| |
+--+ +-------- Cumberlandia_monodonta
| +-----+
| | | +----- Pseudunio_marocanus
| | +--+
| | | +-- Pseudunio_auricularius
| | +--+
+--+ +-- Pseudunio_homsensis
|
| +-- Margaritifera_dahurica
| +--------+
| | +-- Margaritifera_margaritifera
+--+
| +-------- Margaritifera_falcata
| |
+--+ +-- Margaritifera_middendorffi
| +--+
| | +-- Margaritifera_laevis
+--+
| +-- Margaritifera_hembeli
+--+
+-- Margaritifera_marrianae
Lopes-Lima et al., 2018 [Publ 3829], fig. 3
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 6 |
6 |
4 |
3 |
22 |
20 |
2 |
3.0 |
+----- Triplodon_corrugatus
+-----------------------+
| | +-- Velesunio_ambiguus
| +--+
| +-- Hyridella_australis
|
| +----------- Mutela_hargeri **
| |
--+ +--------------+ +-- Chambardia_wahlbergi **
| | | +-----+
| | | | +-- Aspatharia_pfeifferiana **
| | +--+
| | | +----- Etheria_elliptica **
| | +--+
| | | +-- Tamsiella_schroeteriana **
| | +--+
+--+ +-- Lamproscapha_ensiformis **
|
| +----- Potomida_littoralis
| +-----------------+
| | | +-- Lampsilis_cardium
| | +--+
| | +-- Unio_pictorum
| |
+--+ +----- Gibbosula_rochechouartii
| +--------------+
| | | +-- Gibbosula_laosensis
| | +--+
| | +-- Gibbosula_crassa
| |
+--+ +-------- Cumberlandia_monodonta
| +--------+
| | | +----- Pseudunio_marocanus
| | +--+
| | | +-- Pseudunio_auricularius
| | +--+
+--+ +-- Pseudunio_homsensis
|
| +-- Margaritifera_margaritifera
| +-----------+
| | +-- Margaritifera_dahurica
+--+
| +----------- Margaritifera_falcata
+--+
| +-------- Margaritifera_laevis
+--+
| +----- Margaritifera_middendorffi
+--+
| +-- Margaritifera_hembeli
+--+
+-- Margaritifera_marrianae
Wu et al., 2020 [Publ 4121], fig. 2
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 6 |
6 |
4 |
3 |
28 |
26 |
2 |
3.0 |
+----- Triplodon_corrugatus
+--------------------+
| | +-- Hyridella_australis
| +--+
| +-- Velesunio_ambiguus
|
| +----------- Mutela_hargeri **
| |
| +-----------+ +-- Aspatharia_pfeifferiana **
| | | +-----+
--+ | | | +-- Chambardia_wahlbergi **
| | +--+
| | | +----- Etheria_elliptica **
| | +--+
| | | +-- Tamsiella_schroeteriana **
| | +--+
| | +-- Lamproscapha_ensiformis **
| |
| | +-- Lampsilis_cardium
| | +--------------+
+--+ | +-- Scabies_crispata
| |
| | +-------- Sinosolenaia_carinata
| +--+ +-----+
| | | | | +----- Potomida_littoralis
| | | | +--+
| | | | | +-- Lamprotula_caveata
| | +--+ +--+
| | | +-- Lamprotula_leaii
| | |
| | | +----------- Unio_pictorum
| | +--+
| | | +-------- Aculamprotula_tortuosa
+--+ +--+
| | +----- Aculamprotula_polysticta
| +--+
| | +-- Aculamprotula_fibrosa
| +--+
| +-- Aculamprotula_tientsinensis
|
| +----------------- Cumberlandia_monodonta
| |
| | +-- Gibbosula_laosensis
| | +--+
+--+ | +-- Gibbosula_rochechouartii
| +--------+
| | | +-- Pseudunio_auricularius
| | +--+
| | +-- Pseudunio_marocanus
+--+
| +-- Margaritifera_dahurica
| +--------+
| | +-- Margaritifera_margaritifera
+--+
| +-------- Margaritifera_falcata
| |
+--+ +-- Margaritifera_hembeli
| +--+
| | +-- Margaritifera_marrianae
+--+
| +-- Margaritifera_laevis
+--+
+-- Margaritifera_middendorffi
Wu et al., 2020 [Publ 4121], fig. S1
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 6 |
6 |
4 |
3 |
28 |
26 |
2 |
3.0 |
+----- Triplodon_corrugatus
+-----------------------+
| | +-- Hyridella_australis
| +--+
| +-- Velesunio_ambiguus
|
| +----------- Mutela_hargeri **
| |
| +--------------+ +-- Aspatharia_pfeifferiana **
| | | +-----+
--+ | | | +-- Chambardia_wahlbergi **
| | +--+
| | | +----- Etheria_elliptica **
| | +--+
| | | +-- Tamsiella_schroeteriana **
| | +--+
| | +-- Lamproscapha_ensiformis **
| |
| | +-- Lampsilis_cardium
| | +--------------+
+--+ | +-- Scabies_crispata
| |
| | +-------- Sinosolenaia_carinata
| +-----+ +-----+
| | | | | +----- Potomida_littoralis
| | | | +--+
| | | | | +-- Lamprotula_caveata
| | +--+ +--+
| | | +-- Lamprotula_leaii
| | |
| | | +----------- Unio_pictorum
| | +--+
+--+ | +-------- Aculamprotula_tortuosa
| +--+
| | +----- Aculamprotula_polysticta
| +--+
| | +-- Aculamprotula_tientsinensis
| +--+
| +-- Aculamprotula_fibrosa
|
| +-------------------- Gibbosula_rochechouartii
| |
+--+ +----------------- Gibbosula_laosensis
| |
| | +----- Cumberlandia_monodonta
+--+ +--------+
| | | +-- Pseudunio_auricularius
| | +--+
| | +-- Pseudunio_marocanus
+--+
| +-- Margaritifera_dahurica
| +--------+
| | +-- Margaritifera_margaritifera
+--+
| +-------- Margaritifera_falcata
| |
+--+ +-- Margaritifera_hembeli
| +--+
| | +-- Margaritifera_marrianae
+--+
| +-- Margaritifera_laevis
+--+
+-- Margaritifera_middendorffi
Sano et al., 2020 [Publ 4138], fig. 2
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 6 |
6 |
4 |
3 |
67 |
65 |
2 |
3.0 |
+----- Triplodon_corrugatus
+--------------------------------------------+
| | +-- Hyridella_australis
| +--+
| +-- Velesunio_ambiguus
|
| +----------- Mutela_hargeri **
| |
| +-----------------------------------+ +-- Aspatharia_pfeifferiana **
| | | +-----+
| | | | +-- Chambardia_wahlbergi **
--+ | +--+
| | | +----- Etheria_elliptica **
| | +--+
| | | +-- Tamsiella_schroeteriana **
| | +--+
| | +-- Lamproscapha_ensiformis **
| |
| | +-------------------- Cumberlandia_monodonta
| | |
| | | +-- Gibbosula_rochechouartii
| | | +--+
+--+ +-----------------------+ | +-- Gibbosula_laosensis
| | | +-----------+
| | | | | +-- Pseudunio_auricularius
| | | | +--+
| | | | +-- Pseudunio_marocanus
| | +--+
| | | +-- Margaritifera_dahurica
| | | +-----------+
| | | | +-- Margaritifera_margaritifera
| | +--+
| | | +----------- Margaritifera_falcata
| | +--+
| | | +-------- Margaritifera_middendorffi
| | +--+
+--+ | +----- Margaritifera_laevis
| +--+
| | +-- Margaritifera_hembeli
| +--+
| +-- Margaritifera_marrianae
|
| +----------------------------------------- Scabies_crispata
| |
| | +----------------- Lampsilis_cardium
| | |
| | | +----- Sinohyriopsis_cumingii[2]
| | +--------------------+ +--------+
| | | | | | +-- Sinohyriopsis_schlegelii
| | | | | +--+
| | | +--+ +-- Sinohyriopsis_cumingii[1]
+--+ | |
| | | +----------- Inversidens_brandtii
| | | |
| | +--+ +-- Ptychorhynchus_pfisteri
| | | +-----+
| | | | +-- Obovalis_omiensis
| | +--+
| | | +----- Lamprotula_leaii
| | +--+
+--+ | +-- Pronodularia_japanensis
| +--+
| +-- Potomida_littoralis
|
| +----- Buldowskia_iwakawai
| +-----------------------------+
| | | +-- Anemina_arcaeformis
| | +--+
| | +-- Anemina_euscaphys
| |
| | +-------------- Pletholophus_tenuis
| | |
| | | +----- Beringiana_fukuharai
| | +-----------------+ +-----+
+--+ | | | | +-- Beringiana_japonica
| | | | +--+
| | +--+ +-- Beringiana_beringiana
| | |
| | | +-------- Sinanodonta_lucida
| | +--+
| | | +----- Sinanodonta_tumens
| | +--+
| | | +-- Sinanodonta_lauta
+--+ +--+
| +-- Sinanodonta_woodiana
|
| +----------------------------- Cristaria_plicata
| |
| | +-- Anodonta_cygnea
| | +--------+
| | | +-- Anodonta_anatina
| | |
| | +--------------+ +----- Alasmidonta_varicosa
+--+ | | +--+
| | | | | +-- Prolasmidonta_heterodon
| | | | +--+
| | +--+ +-- Platynaias_compressa
| | |
| | | +----- Pyganodon_grandis
| | +--+
+--+ | +-- Utterbackia_imbecillis
| +--+
| +-- Utterbackia_peninsularis
|
| +----- Lanceolaria_grayii[1]
| +-----------------+
| | | +-- Lanceolaria_lanceolata
| | +--+
| | +-- Lanceolaria_grayii[2]
+--+
| +-------------------- Lepidodesma_languilati
| |
| | +----- Aculamprotula_tortuosa
| | +-----------+
+--+ | | +-- Aculamprotula_tientsinensis
| | +--+
| | +-- Aculamprotula_coreana
| |
+--+ +----- Unio_crassus
| +--------+
| | | +-- Unio_delphinus
| | +--+
| | +-- Unio_pictorum
| |
+--+ +-------- Inversiunio_jokohamensis
| +--+
| | | +----- Inversiunio_yanagawensis[2]
| | +--+
| | | +-- Inversiunio_reinianus
| | +--+
+--+ +-- Inversiunio_yanagawensis[1]
|
| +-- Tchangsinaia_piscicula
| +-----+
| | +-- Schistodesmus_lampreyanus
+--+
| +----- Nodularia_douglasiae[2]
+--+
| +-- Nodularia_nipponensis
+--+
+-- Nodularia_douglasiae[1]
Sano et al., 2020 [Publ 4138], fig. 3
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 6 |
6 |
4 |
3 |
66 |
64 |
2 |
3.0 |
+----- Triplodon_corrugatus
+--------------------------------------------+
| | +-- Hyridella_australis
| +--+
| +-- Velesunio_ambiguus
|
| +----------- Mutela_hargeri **
| |
| +-----------------------------------+ +-- Aspatharia_pfeifferiana **
| | | +-----+
| | | | +-- Chambardia_wahlbergi **
--+ | +--+
| | | +----- Etheria_elliptica **
| | +--+
| | | +-- Tamsiella_schroeteriana **
| | +--+
| | +-- Lamproscapha_ensiformis **
| |
| | +----------------------- Cumberlandia_monodonta
| | |
| | | +-- Pseudunio_auricularius
| | +--------------------+ +-----------------+
+--+ | | | +-- Pseudunio_marocanus
| | | |
| | +--+ +-- Gibbosula_laosensis
| | | +--------------+
| | | | +-- Gibbosula_rochechouartii
| | | |
| | +--+ +-- Margaritifera_margaritifera
| | | +-----------+
| | | | +-- Margaritifera_dahurica
| | +--+
| | | +----------- Margaritifera_falcata
| | +--+
| | | +-------- Margaritifera_middendorffi
| | +--+
+--+ | +----- Margaritifera_laevis
| +--+
| | +-- Margaritifera_hembeli
| +--+
| +-- Margaritifera_marrianae
|
| +----------------------------------------- Scabies_crispata
| |
| | +----- Sinohyriopsis_cumingii[2]
| | +--------+
| | | | +-- Sinohyriopsis_schlegelii
| | | +--+
| | | +-- Sinohyriopsis_cumingii[1]
| | +-----------------------+
| | | | +----------- Lampsilis_cardium
+--+ | | |
| | | | +----- Lamprotula_leaii
| | +--+ +--+
| | | | | +-- Potomida_littoralis
| | | | +--+
| | +--+ +-- Pronodularia_japanensis
| | |
| | | +----- Inversidens_brandtii
| | +--+
| | | +-- Obovalis_omiensis
+--+ +--+
| +-- Ptychorhynchus_pfisteri
|
| +----- Buldowskia_iwakawai
| +-----------------------------+
| | | +-- Anemina_arcaeformis
| | +--+
| | +-- Anemina_euscaphys
| |
| | +-------- Pletholophus_tenuis
| | +--+
| | | | +----- Beringiana_fukuharai
| | | +--+
+--+ | | +-- Beringiana_japonica
| +--------------------+ +--+
| | | +-- Beringiana_beringiana
| | |
| | | +-------- Sinanodonta_lucida
| | +--+
| | | +----- Sinanodonta_tumens
| | +--+
| | | +-- Sinanodonta_woodiana
| | +--+
+--+ +-- Sinanodonta_lauta
|
| +----------------------------- Cristaria_plicata
| |
| | +-- Anodonta_anatina
| | +--------+
| | | +-- Anodonta_cygnea
| | |
| | +--------------+ +----- Pyganodon_grandis
+--+ | | +--+
| | | | | +-- Utterbackia_imbecillis
| | | | +--+
| | +--+ +-- Utterbackia_peninsularis
| | |
| | | +----- Alasmidonta_varicosa
| | +--+
+--+ | +-- Platynaias_compressa
| +--+
| +-- Prolasmidonta_heterodon
|
| +----- Lanceolaria_grayii[1]
| +-----------------+
| | | +-- Lanceolaria_lanceolata
| | +--+
| | +-- Lanceolaria_grayii[2]
+--+
| +-------------------- Lepidodesma_languilati
| |
| | +----- Aculamprotula_tortuosa
| | +-----------+
+--+ | | +-- Aculamprotula_tientsinensis
| | +--+
| | +-- Aculamprotula_coreana
| |
+--+ +----- Unio_crassus
| +--------+
| | | +-- Unio_delphinus
| | +--+
| | +-- Unio_pictorum
| |
+--+ +-------- Inversiunio_jokohamensis
| +--+
| | | +----- Inversiunio_yanagawensis[2]
| | +--+
| | | +-- Inversiunio_reinianus
| | +--+
+--+ +-- Inversiunio_yanagawensis[1]
|
| +-- Nodularia_nipponensis
| +--+
| | +-- Nodularia_douglasiae
+-----+
| +-- Schistodesmus_lampreyanus
+--+
+-- Tchangsinaia_piscicula
Sano et al., 2020 [Publ 4138], fig. 4
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 6 |
6 |
4 |
3 |
66 |
64 |
2 |
3.0 |
+----- Triplodon_corrugatus
+-----------------------------------------------+
| | +-- Hyridella_australis
| +--+
| +-- Velesunio_ambiguus
|
| +----------- Mutela_hargeri **
| |
| +--------------------------------------+ +-- Aspatharia_pfeifferiana **
| | | +-----+
| | | | +-- Chambardia_wahlbergi **
--+ | +--+
| | | +----- Etheria_elliptica **
| | +--+
| | | +-- Tamsiella_schroeteriana **
| | +--+
| | +-- Lamproscapha_ensiformis **
| |
| | +-------------------- Cumberlandia_monodonta
| | |
| | | +-- Pseudunio_auricularius
| | +--------------------------+ +--------------+
+--+ | | | +-- Pseudunio_marocanus
| | | |
| | +--+ +-- Gibbosula_rochechouartii
| | | +-----------+
| | | | +-- Gibbosula_laosensis
| | | |
| | +--+ +-- Margaritifera_margaritifera
| | | +--------+
| | | | +-- Margaritifera_dahurica
| | +--+
| | | +-------- Margaritifera_falcata
| | | |
| | +--+ +-- Margaritifera_laevis
| | | +--+
+--+ | | +-- Margaritifera_middendorffi
| +--+
| | +-- Margaritifera_hembeli
| +--+
| +-- Margaritifera_marrianae
|
| +-------------------------------------------- Scabies_crispata
| |
| | +----------------- Lampsilis_cardium
| | |
| | | +----- Sinohyriopsis_cumingii[2]
| | +-----------------------+ +--------+
| | | | | | +-- Sinohyriopsis_schlegelii
| | | | | +--+
| | | +--+ +-- Sinohyriopsis_cumingii[1]
+--+ | |
| | | +----------- Inversidens_brandtii
| | | |
| | +--+ +-- Obovalis_omiensis
| | | +-----+
| | | | +-- Ptychorhynchus_pfisteri
| | +--+
| | | +----- Lamprotula_leaii
| | +--+
+--+ | +-- Pronodularia_japanensis
| +--+
| +-- Potomida_littoralis
|
| +----- Buldowskia_iwakawai
| +--------------------------------+
| | | +-- Anemina_arcaeformis
| | +--+
| | +-- Anemina_euscaphys
| |
| | +-------- Sinanodonta_lucida
| | +--+
| | | | +----- Sinanodonta_tumens
| | | +--+
+--+ | | +-- Sinanodonta_woodiana
| +-----------------------+ +--+
| | | +-- Sinanodonta_lauta
| | |
| | | +-------- Pletholophus_tenuis
| | +--+
| | | +----- Beringiana_fukuharai
| | +--+
| | | +-- Beringiana_japonica
| | +--+
+--+ +-- Beringiana_beringiana
|
| +-------------------------------- Cristaria_plicata
| |
| | +-- Anodonta_cygnea
| | +--------+
| | | +-- Anodonta_anatina
| | |
| | +-----------------+ +----- Alasmidonta_varicosa
+--+ | | +--+
| | | | | +-- Prolasmidonta_heterodon
| | | | +--+
| | +--+ +-- Platynaias_compressa
| | |
| | | +----- Pyganodon_grandis
| | +--+
+--+ | +-- Utterbackia_imbecillis
| +--+
| +-- Utterbackia_peninsularis
|
| +-------------------------- Lepidodesma_languilati
| |
| | +----- Lanceolaria_grayii[1]
| | +-----------------+
+--+ | | +-- Lanceolaria_lanceolata
| | +--+
| | +-- Lanceolaria_grayii[2]
| |
+--+ +----- Aculamprotula_tortuosa
| +--------------+
| | | +-- Aculamprotula_tientsinensis
| | +--+
| | +-- Aculamprotula_coreana
| |
+--+ +----- Unio_crassus
| +-----------+
| | | +-- Unio_delphinus
| | +--+
| | +-- Unio_pictorum
+--+
| +-- Nodularia_douglasiae
| +-----------+
| | +-- Nodularia_nipponensis
| |
+--+ +-- Schistodesmus_lampreyanus
| +--------+
| | +-- Tchangsinaia_piscicula
+--+
| +-------- Inversiunio_jokohamensis
+--+
| +----- Inversiunio_yanagawensis[2]
+--+
| +-- Inversiunio_reinianus
+--+
+-- Inversiunio_yanagawensis[1]
Hoeh et al., 2001 [Publ 2652], fig. 14.2
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 5 |
5 |
2 |
2 |
26 |
25 |
2 |
8.0 |
+----- Pyganodon_grandis
+-----+
| | +-- Anodonta_cygnea
| +--+
| +-- Strophitus_undulatus
|
| +----- Diplodon_gratus
--+ |
| +----- Echyridella_menziesii
| +--+
| | +----- Lortiella_rugata
| | |
| | | +-- Castalia_stevensi
| | +--+
+--+ +-- Velesunio_angasi
|
| +-- Mutela_dubia **
| +--+
| | +-- Mutela_rostrata **
+--+
| +-- Anodontites_guanarensis **
| |
+--+-- Anodontites_trigona **
|
+-- Monocondylaea_minuana **
Hoeh et al., 2001 [Publ 2652], fig. 14.4
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 5 |
5 |
3 |
2 |
26 |
25 |
2 |
8.0 |
+-- Castalia_stevensi
+-----+
| +-- Diplodon_gratus
+--------------------------+
| | +----- Echyridella_menziesii
| +--+
| | +-- Lortiella_rugata
| +--+
| +-- Velesunio_angasi
|
--+ +-- Mutela_dubia **
| +-----+
| | +-- Mutela_rostrata **
| +-----------------------+
| | | +----- Monocondylaea_minuana **
| | +--+
| | | +-- Anodontites_guanarensis **
| | +--+
+--+ +-- Anodontites_trigona **
|
| +----------------------------- Coelatura_aegyptiaca
| |
| | +-- Cumberlandia_monodonta
+--+ +-----------------------+
| | +-- Margaritifera_margaritifera
| |
| | +----------------------- Gonidea_angulata
+--+ |
| | +-------- Unio_pictorum
| | |
| | +-----------+-------- Unio_tumidus
+--+ | |
| | | +----- Anodonta_cygnea
| | +--+
| | | +-- Strophitus_undulatus
| | +--+
+--+ +-- Pyganodon_grandis
|
| +----------------- Amblema_plicata
| |
| | +-------- Quadrula_quadrula
| | +-----+
+--+ | | +----- Eurynia_dilatata
| | +--+
| | | +-- Fusconaia_flava
| | +--+
+--+ +-- Pleurobema_clava
|
| +-- Cyrtonaias_tampicoensis
| +--------+
| | +-- Glebula_rotundata
+--+
| +-------- Toxolasma_lividum
+--+
| +----- Potamilus_alatus
+--+
| +-- Ligumia_recta
+--+
+-- Ortmanniana_ligamentina
Hoeh et al., 2002 [Publ 2653], fig. 2
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 5 |
5 |
3 |
2 |
26 |
25 |
2 |
8.0 |
+-- Castalia_stevensi
+-----+
| +-- Diplodon_gratus
+-----------------------+
| | +----- Echyridella_menziesii
| +--+
| | +-- Velesunio_angasi
| +--+
| +-- Lortiella
|
--+ +-- Mutela_dubia **
| +-----+
| | +-- Mutela_rostrata **
| +--------------------+
| | | +----- Monocondylaea_minuana **
| | +--+
| | | +-- Anodontites_guanarensis **
| | +--+
+--+ +-- Anodontites_trigona **
|
| +-------------------------- Coelatura_aegyptiaca
| |
| | +-- Margaritifera_margaritifera
| | +--------------------+
+--+ | +-- Cumberlandia_monodonta
| |
| | +-------------- Gonidea_angulata
| | +-----+
+--+ | | +----------- Unio_pictorum
| | +--+
| | | +-------- Unio_tumidus
| | +--+
| | | +----- Anodonta_cygnea
| | +--+
+--+ | +-- Pyganodon_grandis
| +--+
| +-- Strophitus_undulatus
|
| +-- Glebula_rotundata
| +--------------+
| | +-- Cyrtonaias_tampicoensis
| |
+--+ +-------------- Toxolasma_lividum
| |
| | +----- Potamilus_alatus
| +--------+
+--+ | +-- Ligumia_recta
| +--+
| +-- Ortmanniana_ligamentina
|
| +----------- Amblema_plicata
+--+
| +-------- Quadrula_quadrula
+--+
| +----- Eurynia_dilatata
+--+
| +-- Pleurobema_clava
+--+
+-- Fusconaia_flava
Roe & Hoeh, 2003 [Publ 2655], fig. 4.2
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 5 |
5 |
3 |
2 |
26 |
25 |
2 |
8.0 |
+-- Mutela_dubia **
+-----+
| +-- Mutela_rostrata **
+--+
| | +----- Monocondylaea_minuana **
| +--+
| | +-- Anodontites_guanarensis **
| +--+
+-----------------------+ +-- Anodontites_trigona **
| |
| | +-- Castalia_stevensi
| | +--+
| | | +-- Velesunio_angasi
| +-----+
| | +-- Echyridella_menziesii
--+ | |
| +--+-- Lortiella_rugata
| |
| +-- Diplodon_gratus
|
| +-- Cumberlandia_monodonta
| +-----------------------------+
| | +-- Margaritifera_margaritifera
+--+
| +----------------------------- Gonidea_angulata
| |
+--+ +-------------------------- Coelatura_aegyptiaca
| |
+--+ +----------------------- Fusconaia_flava
| |
| +----------------------- Amblema_plicata
+--+
| +-------------------- Quadrula_quadrula
| |
| | +----------------- Pleurobema_clava
+--+ |
| | +-------------- Eurynia_dilatata
| | |
+--+ | +----------- Ortmanniana_ligamentina
| +--+
| | | +-------- Glebula_rotundata
| | +--+
| | | +----- Potamilus_alatus
+--+ +--+
| | +-- Ligumia_recta
| +--+
| +-- Toxolasma_lividum
|
| +----------- Cyrtonaias_tampicoensis
| |
+--+ +-- Unio_pictorum
| +-----+
| | +-- Unio_tumidus
+--+
| +----- Pyganodon_grandis
+--+
| +-- Anodonta_cygnea
+--+
+-- Strophitus_undulatus
Roe & Hoeh, 2003 [Publ 2655], fig. 4.3
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 5 |
5 |
3 |
2 |
26 |
25 |
2 |
8.0 |
+-- Mutela_dubia **
+-----+
| +-- Mutela_rostrata **
+--+
| | +----- Monocondylaea_minuana **
| +--+
| | +-- Anodontites_guanarensis **
| +--+
+-----------------+ +-- Anodontites_trigona **
| |
| | +-- Castalia_stevensi
| | +-----+
| | | +-- Diplodon_gratus
| +--+
| | +----- Echyridella_menziesii
| +--+
--+ | +-- Velesunio_angasi
| +--+
| +-- Lortiella_rugata
|
| +-------------------------- Coelatura_aegyptiaca
| |
| | +-- Unio_pictorum
| | +-----+
| | | +-- Unio_tumidus
+--+ +--------------+
| | | +----- Anodonta_cygnea
| | +--+
| | | +-- Pyganodon_grandis
| | +--+
+--+ +-- Strophitus_undulatus
|
| +-------------------- Gonidea_angulata
| |
| | +----------------- Amblema_plicata
+--+ |
| | +-------- Quadrula_quadrula
| | +-----+
+--+ | | +----- Eurynia_dilatata
| | +--+
| | | +-- Fusconaia_flava
| | +--+
+--+ +-- Pleurobema_clava
|
| +-- Cyrtonaias_tampicoensis
| +--------+
| | +-- Glebula_rotundata
+--+
| +-------- Toxolasma_lividum
+--+
| +----- Ortmanniana_ligamentina
+--+
| +-- Ligumia_recta
+--+
+-- Potamilus_alatus
Whelan et al., 2011 [Publ 3380], fig. 1 ML
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 5 |
5 |
3 |
2 |
32 |
31 |
2 |
3.0 |
+-- Margaritifera_margaritifera
+-----------------------------+
| +-- Cumberlandia_monodonta
|
| +-- Aspatharia_pfeifferiana **
| +-----+
| | +-- Chambardia_wahlbergi **
| +--------------------+
| | | +----- Mutela_rostrata **
--+ | +--+
| | | +-- Mycetopoda_siliquosa **
| | +--+
| | +-- Tamsiella_schroeteriana **
| |
| | +-- Lamellidens_savadiensis
| | +-----------------+
| | | +-- Trapezidens_dolichorhynchus
+--+ +-----+
| | | +----------------- Leoparreysia_tavoyensis
| | | |
| | +--+ +-- Radiatula_chaudhurii
| | | +-----------+
| | | | +-- Indochinella_pugio
| | +--+
| | | +----------- Nitia_teretiuscula
| | +--+
| | | +-------- Coelatura_aegyptiaca[2]
| | +--+
+--+ | +----- Coelatura_aegyptiaca[1]
| +--+
| | +-- Coelatura_gabonensis
| +--+
| +-- Prisodontopsis_aviculaeformis
|
| +-- Pseudodon_vondembuschianus
| +--------------------+
| | +-- Lens_contradens
| |
| | +-- Pyganodon_grandis
| | +--+
+--+ | +-- Lasmigona_costata
| +--------------+
| | | +-- Unio_pictorum
| | +--+
| | +-- Unio_caffer
+--+
| +----------------- Gonidea_angulata
| |
| | +----- Pronodularia_japanensis
+--+ +--------+
| | | +-- Potomida_littoralis
| | +--+
+--+ +-- Potomida_semirugata
|
| +----------- Pleurobema_sintoxia
+--+
| +-------- Quadrula_quadrula
+--+
| +----- Amblema_plicata
+--+
| +-- Truncilla_truncata
+--+
+-- Lampsilis_cardium
Whelan et al., 2011 [Publ 3380], fig. 1 MP
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 5 |
5 |
3 |
2 |
32 |
31 |
2 |
3.0 |
+-- Aspatharia_pfeifferiana **
+-----+
| +-- Chambardia_wahlbergi **
+-----------------------+
| | +----- Mutela_rostrata **
| +--+
| | +-- Mycetopoda_siliquosa **
| +--+
| +-- Tamsiella_schroeteriana **
|
| +-- Margaritifera_margaritifera
| +--------+
| | +-- Cumberlandia_monodonta
--+ +-----------------+
| | | +-------- Velesunio_ambiguus
| | +--+
| | | +----- Hyridella_australis
| | +--+
| | | +-- Castalia_ambigua
| | +--+
| | +-- Triplodon_corrugatus
| |
| | +-- Lamellidens_savadiensis
| | +--------------+
+--+ | +-- Trapezidens_dolichorhynchus
| +--------+
| | | +-------------- Leoparreysia_tavoyensis
| | | |
| | +--+ +-- Radiatula_chaudhurii
| | | +--------+
| | | | +-- Indochinella_pugio
| | +--+
| | | +-------- Nitia_teretiuscula
| | | |
| | +--+ +----- Coelatura_aegyptiaca[1]
+--+ | |
| +--+----- Coelatura_aegyptiaca[2]
| |
| | +-- Coelatura_gabonensis
| +--+
| +-- Prisodontopsis_aviculaeformis
|
| +----------------------- Lens_contradens
| |
| | +-------------------- Pseudodon_vondembuschianus
+--+ |
| | +-- Pyganodon_grandis
| | +--+
+--+ | +-- Lasmigona_costata
| +-----------+
| | | +-- Unio_pictorum
| | +--+
| | +-- Unio_caffer
+--+
| +-------------- Gonidea_angulata
| |
| | +----- Pronodularia_japanensis
+--+ +-----+
| | | +-- Potomida_littoralis
| | +--+
+--+ +-- Potomida_semirugata
|
| +-------- Quadrula_quadrula
| |
+--+-------- Pleurobema_sintoxia
|
| +----- Amblema_plicata
+--+
| +-- Lampsilis_cardium
+--+
+-- Truncilla_truncata
Lopes-Lima et al., 2020 [Publ 4080], fig. 2-4
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 5 |
5 |
3 |
3 |
117 |
115 |
2 |
5.0 |
+-- Margaritifera_laevis
+-----------------------------------------+
| +-- Margaritifera_middendorffi
|
| +-- Lamproscapha_ensiformis **
| +-----+
| | +-- Mycetopoda_siliquosa **
| +--------------------------------+
| | | +----- Etheria_elliptica **
| | +--+
| | | +-- Aspatharia_pfeifferiana **
| | +--+
| | +-- Chambardia_wahlbergi **
| |
--+ | +-- Coelatura_aegyptiaca
| | |
| | +-----------------------------------+-- Leoparreysia_tavoyensis
| | | |
| | | +-- Radiatula_mouhoti
| | |
| | | +-- Lens_contradens
| | | |
| | | +-- Ensidens_sagittarius
| | | |
| | | +-----------------------+-- Physunio_modelli
| | | | |
| | | | +-- Rectidens_sumatrensis
| | | | |
+--+ | | +-- Yaukthwa_nesemanni
| | |
| | | +-- Amblema_plicata
| | | |
| | | +-- Elliptio_complanata
| | | +--------------------+
| | +--------+ | +-- Lampsilis_cardium
| | | | | |
| | | | | +-- Quadrula_quadrula
| | | | |
| | | | | +----- Chamberlainia_somsakpanhai
| | | | | +-----------+
| | | | | | | +-- Sinohyriopsis_cumingii
| | | | | | +--+
| | | | | +--+ +-- Sinohyriopsis_schlegelii
| | | | | | |
| | | | | | | +-------------- Pronodularia_japanensis
| | | +--+ | | |
| | | | | +--+ +----------- Potomida_littoralis
| | | | | | |
| | | | | +--+ +-------- Lamprotula_caveata
| | | | | | |
+--+ | | | +--+ +----- Lamprotula_leaii[2]
| | | | | |
| | | | +--+----- Lamprotula_leaii[3]
| | | | |
| | | | | +-- Lamprotula_seomjinensis
| | | | +--+
| | | | +-- Lamprotula_leaii[1]
| | +--+
| | | +-- Indopseudodon_indawgyiensis
| | | +--------+
| | | | +-- Indopseudodon_salwenianus
| | | |
| | | +-----+ +-- Pilsbryoconcha_exilis
| | | | | +-----+
| | | | | | +-- Pilsbryoconcha_carinifera
| | | | +--+
| | | | | +----- Pseudodon_mekongi
| | | | | |
| | | | +--+ +-- Nyeinchanconcha_nyeinchani
| | | | | |
| | | | +--+-- Bineurus_mouhotii
| | | | |
| | +--+ +-- Sundadontina_tumida
| | |
| | | +-------- Inversidens_brandtii
| | | |
| | | +-----+ +-- Obovalis_omiensis
| | | | | +--+
| | | | | | +-- Ptychorhynchus_pfisteri
| | | | +--+
| | | | | +-- Parvasolenaia_rivularis
| | | | +--+
| | +--+ +-- Koreosolenaia_sitgyensis
| | |
+--+ | +----- Gonidea_angulata
| | +-----+
| | | | +-- Leguminaia_wheatleyi
| | | +--+
| +--+ +-- Microcondylaea_bonellii
| |
| | +-------- Sinosolenaia_carinata
| +--+
| | +----- Sinosolenaia_recognita
| +--+
| | +-- Sinosolenaia_iridinea
| +--+
| +-- Sinosolenaia_oleivora
|
| +-------------------------------- Acuticosta_chinensis
| |
| +-------------------------------- Lepidodesma_languilati
| |
| | +----------- Aculamprotula_tortuosa
| | +--------------+
| | | | +-------- Aculamprotula_scripta
| | | +--+
| | | | +----- Aculamprotula_coreana
| | | +--+
| | | | +-- Aculamprotula_fibrosa
| | | +--+
| +-----+ +-- Aculamprotula_tientsinensis
| | |
| | | +----------------------- Middendorffinaia_mongolica
| | | |
| | | | +----- Unio_gibbus
| | | | |
| | | | +--------------+----- Unio_tumidus
| | +--+ | |
| | | | | +-- Unio_crassus
| | | | +--+
| | | | +-- Unio_pictorum
| | | |
| | | | +-------------- Nodularia_breviconcha
| | +--+ +--+
| | | | | +----------- Nodularia[2]
| | | | +--+
| | | | | +-------- Nodularia_nuxpersicae
| | | | +--+
| | | | | +----- Nodularia_nipponensis
| | | | +--+
| | | | | +-- Nodularia_douglasiae
| | +--+ +--+
| | | +-- Nodularia[1]
| | |
+--+ | +-- Schistodesmus_lampreyanus
| | +-----------+
| | | +-- Schistodesmus_spinosus
| | |
| | | +----- Cuneopsis_rufescens
| | +--------+
| +--+ | +-- Cuneopsis_heudei
| | +--+
| | +-- Cuneopsis_celtiformis
| |
| | +----------- Tchangsinaia_piscicula
| +--+
| | +-------- Inversiunio_jokohamensis
| +--+
| | +----- Inversiunio_verrucosus
| +--+
| | +-- Inversiunio_reinianus
| +--+
| +-- Inversiunio_yanagawensis
|
| +----------------- Lanceolaria_oxyrhyncha
| |
| +-----------+ +-------------- Lanceolaria_gladiola
| | | |
| | +--+ +-- Lanceolaria_bilirata
| | | +--------+
| | | | +-- Lanceolaria_grayii
| | +--+
| | | +-------- Lanceolaria_acrorrhyncha
| | +--+
| | | +----- Lanceolaria_kihirai
| | +--+
| | | +-- Lanceolaria_lanceolata
| | +--+
| | +-- Lanceolaria_triformis
| |
| | +-------- Pseudanodonta_complanata
+--+ |
| | +----- Anodonta_cygnea
| +--+
| | | +-- Anodonta_anatina
| +-----------------+ +--+
| | | +-- Anodonta_nuttalliana
| | |
| | | +----- Pyganodon_grandis
| | | |
| | | +----- Simpsonaias_ambigua
| | +--+
| | +----- Strophitus_undulatus
| | |
| | | +-- Alasmidonta_marginata
| | +--+
| | +-- Lasmigona_costata
+--+
| +-- Cristaria_plicata
| +--------------------+
| | +-- Cristaria_clessini
| |
| | +-------------- Anemina_arcaeformis
| | |
| +--------+ +----------- Amuranodonta_kijaensis
| | | |
| | +--+ +-------- Buldowskia_suifunica
| | | |
| | +--+ +-- Buldowskia_flavotincta
+--+ | +--+
| | | +-- Buldowskia_shadini
| +--+
| | +-- Buldowskia_iwakawai
| +--+
| +-- Buldowskia_kamiyai
|
| +-------------------- Pletholophus_tenuis
| |
| | +-------- Beringiana_japonica
| | +--------+
+--+ | | +----- Beringiana_beringiana
| | +--+
| | | +-- Beringiana_fukuharai
| | +--+
+--+ +-- Beringiana_gosannensis
|
| +-- Sinanodonta_angula
| +-----------+
| | +-- Sinanodonta_lucida
| |
+--+ +----------- Sinanodonta_schrenkii
| |
| | +-- Sinanodonta_calipygos
| +--------+
+--+ +-- Sinanodonta_tumens
|
| +-------- Sinanodonta_woodiana[2]
| |
+--+ +----- Sinanodonta_elliptica
| |
+--+----- Sinanodonta_jourdyi
|
| +-- Sinanodonta_lauta
+--+
+-- Sinanodonta_woodiana[1]
Bolotov et al., 2016 [Publ 3743], fig. 2
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 4 |
4 |
2 |
3 |
26 |
23 |
2 |
4.0 |
+----- Diplodon_suavidicus
+-----+
| | +-- Triplodon_corrugatus
| +--+
| +-- Castalia_ambigua
+--------------+
| | +----- Microdontia_anodontaeformis
| | +--+
| | | | +-- Lortiella_froggatti
| | | +--+
| +--+ +-- Alathyria_pertexta
| |
| | +----- Velesunio_ambiguus
| +--+
| | +-- Alathyria_jacksoni
--+ +--+
| +-- Alathyria_profuga
|
| +-------- Tamsiella_schroeteriana **
| +--------------+
| | | +----- Etheria_elliptica **
| | +--+
| | | +-- Aspatharia_pfeifferiana **
| | +--+
| | +-- Chambardia_wahlbergi **
+--+
| +-------- Lampsilis_cardium
| +-----------+
| | | +----- Anodonta_anatina
| | +--+
| | | +-- Unio_pictorum
| | +--+
+--+ +-- Unio_ravoisieri
|
| +-- Pseudunio_auricularius
| +--------------+
| | +-- Pseudunio_marocanus
| |
+--+ +-- Gibbosula_laosensis
| +-----------+
| | +-- Cumberlandia_monodonta
| |
+--+ +-- Margaritifera_dahurica
| +--------+
| | +-- Margaritifera_margaritifera
+--+
| +-------- Margaritifera_falcata
+--+
| +----- Margaritifera_laevis
+--+
| +-- Margaritifera_middendorffi
+--+
+-- Margaritifera_marrianae
Bolotov et al., 2016 [Publ 3743], fig. 4
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 4 |
4 |
2 |
3 |
26 |
23 |
2 |
4.0 |
+----- Diplodon_suavidicus
+-----+
| | +-- Triplodon_corrugatus
| +--+
| +-- Castalia_ambigua
+--------------+
| | +----- Microdontia_anodontaeformis
| | +--+
| | | | +-- Alathyria_pertexta
| | | +--+
| +--+ +-- Lortiella_froggatti
| |
| | +----- Velesunio_ambiguus
| +--+
| | +-- Alathyria_jacksoni
--+ +--+
| +-- Alathyria_profuga
|
| +-------- Tamsiella_schroeteriana **
| +--------------+
| | | +----- Etheria_elliptica **
| | +--+
| | | +-- Chambardia_wahlbergi **
| | +--+
| | +-- Aspatharia_pfeifferiana **
+--+
| +-------- Lampsilis_cardium
| +-----------+
| | | +----- Anodonta_anatina
| | +--+
| | | +-- Unio_pictorum
| | +--+
+--+ +-- Unio_ravoisieri
|
| +-- Pseudunio_auricularius
| +--------------+
| | +-- Pseudunio_marocanus
| |
+--+ +-- Gibbosula_laosensis
| +-----------+
| | +-- Cumberlandia_monodonta
| |
+--+ +-- Margaritifera_margaritifera
| +--------+
| | +-- Margaritifera_dahurica
+--+
| +-------- Margaritifera_falcata
+--+
| +----- Margaritifera_laevis
+--+
| +-- Margaritifera_marrianae
+--+
+-- Margaritifera_middendorffi
Bolotov et al., 2017 [Publ 3761], fig. S1
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 4 |
4 |
2 |
3 |
137 |
135 |
2 |
4.0 |
+-------- Tamsiella_schroeteriana **
+-----+
| | +----- Etheria_elliptica **
| +--+
| | +-- Chambardia_wahlbergi **
| +--+
| +-- Aspatharia_pfeifferiana **
+-----------------------+
| | +----- Diplodon_suavidicus
| | +-----+
| | | | +-- Castalia_ambigua
| | | +--+
| | | +-- Triplodon_corrugatus
| +--+
| | +----- Alathyria_pertexta
| | +--+
| | | | +-- Microdontia_anodontaeformis
| | | +--+
| +--+ +-- Lortiella_froggatti
| |
| | +----- Velesunio_ambiguus
| +--+
| | +-- Alathyria_jacksoni
| +--+
| +-- Alathyria_profuga
|
| +-- Trapezidens_dolichorhynchus
| +-----------+
| | +-- Trapezidens_exolescens
| +-----------+
| | | +----------- Lamellidens_corrianus
| | | |
| | +--+ +-------- Lamellidens_savadiensis
| | | |
| | +--+ +-- Lamellidens_jenkinsianus
| | | +--+
| | | | +-- Lamellidens_mainwaringi
| | +--+
| +--------+ | +-- Lamellidens_ferrugineus
| | | +--+
| | | +-- Lamellidens_marginalis
| | |
| | | +-- Parreysia_nagpoorensis
| | | +--------------------+
| | | | +-- Parreysia_plagiosoma
| | | |
--+ | | | +-- Leoparreysia_tavoyensis
| | | | +-----------------+
| | +--+ | +-- Leoparreysia_olivacea
| | | |
| | | | +-------------- Nitia_teretiuscula
| | | | +--+
| | | | | | +----------- Coelatura_aegyptiaca[2]
| | +--+ | +--+
| | | | | +-------- Coelatura_aegyptiaca[1]
| | | | +--+
| | | | | +----- Prisodontopsis_mweruensis
| | | | +--+
| | | | | +-- Coelatura_gabonensis
| | +--+ +--+
| | | +-- Coelatura_kunenensis
| | |
| | | +-------- Indonaia_shurtleffiana
| | | +-----+
| | | | | +----- Indonaia_andersoniana
| | | | +--+
| | | | | +-- Indonaia_rugosa
| | | | +--+
| | +--+ +-- Indonaia_bonneaudii
| | |
| | | +-- Indochinella_pugio
| | | +--------+
| | | | +-- Scabies_crispata
| | | |
| | +--+ +-- Radiatula_myitkyinae
| | | +-----+
| | | | +-- Radiatula_chaudhurii
| | +--+
| | | +----- Scabies_mandarinus
| | +--+
| | | +-- Scabies_humilis
| | +--+
| | +-- Scabies_phaselus
| |
| | +-- Ensidens_telus
| | +-----+
+--+ | +-- Ensidens_ingallsianus
| +-----+
| | | +----- Ensidens_sagittarius
| | +--+
| | | +-- Ensidens_spiculus
| | +--+
| +--+ +-- Ensidens_jaculus
| | |
| | | +-- Hyriopsis_bialata
| | | +--------+
| | | | +-- Hyriopsis_khoratensis
| | +--+
| | | +-------- Hyriopsis_panhai
| | +--+
| | | +----- Hyriopsis_myersiana
| | +--+
| +--------------+ | +-- Rectidens_sumatrensis
| | | +--+
| | | +-- Hyriopsis_desowitzi
| | |
| | | +----- Physunio
| | | +--------+
| | | | | +-- Physunio_modelli
| | | | +--+
| | | | +-- Physunio_superbus
| | | |
| | +--+ +-------- Yaukthwa_inlenensis
| | | +--+
| | | | | +----- Yaukthwa_panhai
| | | | +--+
| | | | | +-- Yaukthwa_rectangularis
| | | | +--+
| | +--+ +-- Yaukthwa_nesemanni
| | |
| | | +-- Lens_contradens
| | | +-----+
| | | | +-- Lens_micropterus
| | +--+
| | | +----- Lens_eximius
| | +--+
| | | +-- Lens_comptus
| | +--+
| | +-- Lens_novoselovi
| |
| | +-------------------- Indopseudodon_indawgyiensis
| | |
| | | +-------- Pilsbryoconcha_exilis[1]
| | | +--------+
+--+ +--------+ | | +----- Pilsbryoconcha_exilis[3]
| | | | +--+
| | | | | +-- Pilsbryoconcha_exilis[2]
| | | | +--+
| | | | +-- Pilsbryoconcha_carinifera
| | +--+
| | | +-------- Sundadontina_tumida
| | | +-----+
| | | | | +----- Pseudodon_vagulus
| | | | +--+
| | | | | +-- Pseudodon_mekongi
| | | | +--+
| | +--+ +-- Pseudodon_vondembuschianus
| | |
| | | +-- Nyeinchanconcha_nyeinchani
| | | +--------+
| | | | +-- Sundadontina_cumingii
| | +--+
| | | +-------- Thaiconcha_callifera
| | +--+
| | | +----- Bineurus_exilis
| | +--+
| | | +-- Bineurus_mouhotii
| | +--+
| | +-- Bineurus_loeiensis
| |
| | +-------- Amblema_plicata
| | +--+
| | | | +----- Ortmanniana_ligamentina
| | | +--+
| | | | +-- Lampsilis_cardium
| | | +--+
+--+ +-----+ +-- Cambarunio_iris
| | |
| | | +-- Quadrula_quadrula
| | | +-----+
| | | | +-- Quadrula_verrucosa
| | +--+
| | | +----- Pleurobema_sintoxia
| | +--+
| | | +-- Eurynia_dilatata
| | +--+
| | +-- Elliptio_complanata
| +--------+
| | | +-- Sinohyriopsis_cumingii
| | | +-----+
| | | | +-- Chamberlainia_somsakpanhai
| | | |
| | | +-----+ +-- Potomida_littoralis
| | | | | +--+
| | | | | | +-- Pronodularia_japanensis
| | | | +--+
| | | | | +-- Lamprotula_caveata
| | | | +--+
| | +--+ +-- Lamprotula_leaii
| | |
| | | +----- Sinosolenaia_carinata
| | | +-----+
| | | | | +-- Sinosolenaia_oleivora
| | | | +--+
| | +--+ +-- Sinosolenaia_recognita
| | |
| | | +-------- Gonidea_angulata
| | +--+
+--+ | +----- Microcondylaea_bonellii
| +--+
| | +-- Leguminaia_wheatleyi
| +--+
| +-- Leguminaia
|
| +----- Unio_tumidus
| +--------+
| | | +-- Unio_pictorum
| | +--+
| | +-- Unio_crassus
| +--------+
| | | +----------- Aculamprotula_tortuosa
| | | |
| | | | +-- Nodularia_douglasiae
| | +--+ +-----+
| | | | +-- Nodularia_nuxpersicae
| | | |
| | +--+ +-- Cuneopsis_rufescens
| | | +--+
| | | | +-- Cuneopsis_heudei
| | +--+
+--+ | +-- Tchangsinaia_piscicula
| +--+
| +-- Schistodesmus_lampreyanus
|
| +-- Lanceolaria_grayii
| +-----------------+
| | +-- Lanceolaria_gladiola
| |
| | +-- Cristaria_plicata
| | +--------+
| | | +-- Cristaria_radiata
+--+ +-----+
| | | +-------- Pletholophus_tenuis
| | +--+
| | | +----- Sinanodonta_lucida
| | +--+
| | | +-- Sinanodonta_jourdyi
+--+ +--+
| +-- Sinanodonta_lauta
|
| +-------------- Anodonta_anatina
| |
| | +----- Pseudanodonta_complanata
+--+ +-----+
| | | +-- Anodonta_cygnea
| | +--+
| | +-- Anodonta_nuttalliana
+--+
| +-- Lasmigona_costata
| +-----+
| | +-- Alasmidonta_marginata
+--+
| +----- Simpsonaias_ambigua
+--+
| +-- Pyganodon_grandis
+--+
+-- Strophitus_undulatus
Bolotov et al., 2017 [Publ 3761], fig. S2
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 4 |
4 |
2 |
3 |
137 |
135 |
2 |
2.0 |
+-------------------- Chamberlainia_somsakpanhai
|
+-----------+ +----------------- Cumberlandia_monodonta
| | |
| | | +-------------- Gibbosula_laosensis
| +--+ |
| | | +-- Pseudunio_auricularius
| | +-----------+
| +--+ +-- Pseudunio_marocanus
| |
| | +-- Margaritifera_dahurica
| | +--------+
| | | +-- Margaritifera_margaritifera
| +--+
| | +-------- Margaritifera_falcata
| +--+
| | +----- Margaritifera_middendorffi
| +--+
| | +-- Margaritifera_laevis
| +--+
| +-- Margaritifera_marrianae
|
| +-- Tamsiella_schroeteriana **
| +--+
| | +-- Etheria_elliptica **
| +--------+
| | | +-- Aspatharia_pfeifferiana **
| | +--+
| | +-- Chambardia_wahlbergi **
| |
+-----------------+ +----- Diplodon_suavidicus
| | +-----+
| | | | +-- Castalia_ambigua
| | | +--+
| | | +-- Triplodon_corrugatus
| +--+
| | +----- Velesunio_ambiguus
| | +--+
| | | | +-- Alathyria_jacksoni
| | | +--+
| +--+ +-- Alathyria_profuga
| |
| | +----- Microdontia_anodontaeformis
| +--+
| | +-- Alathyria_pertexta
| +--+
| +-- Lortiella_froggatti
|
| +-- Parreysia_nagpoorensis
| +-----------+
| | +-- Parreysia_plagiosoma
| |
| +--------+ +-- Trapezidens_dolichorhynchus
| | | +--------+
| | | | +-- Trapezidens_exolescens
| | | |
| | +--+ +-------- Lamellidens_savadiensis
--+ | | |
| | | +-------- Lamellidens_corrianus
| | +--+
| | | +-- Lamellidens_marginalis
| | | +--+
| | | | +-- Lamellidens_ferrugineus
| +-----+ +--+
| | | | +-- Lamellidens_jenkinsianus
| | | +--+
| | | +-- Lamellidens_mainwaringi
| | |
| | | +-- Leoparreysia_olivacea
| | | +-----------------+
| | | | +-- Leoparreysia_tavoyensis
| | | |
| | | | +-------------- Nitia_teretiuscula
| | | | +--+
| | | | | | +----------- Coelatura_aegyptiaca[1]
| | +--+ | +--+
| | | | | +-------- Coelatura_aegyptiaca[2]
| | | | +--+
| | | | | +----- Coelatura_gabonensis
| | | | +--+
| | | | | +-- Coelatura_kunenensis
| | +--+ +--+
| | | +-- Prisodontopsis_mweruensis
| | |
| | | +-------- Indonaia_shurtleffiana
| | | +-----+
| | | | | +----- Indonaia_andersoniana
| | | | +--+
| | | | | +-- Indonaia_rugosa
| | | | +--+
| | +--+ +-- Indonaia_bonneaudii
| | |
| | | +-- Indochinella_pugio
| | | +--------+
| | | | +-- Scabies_crispata
| | | |
| | +--+ +-- Radiatula_myitkyinae
| | | +-----+
| | | | +-- Radiatula_chaudhurii
| | +--+
| | | +----- Scabies_mandarinus
| | +--+
| | | +-- Scabies_humilis
| | +--+
| | +-- Scabies_phaselus
| |
| | +----- Unio_tumidus
| | +-----+
| | | | +-- Unio_crassus
| | | +--+
| | | +-- Unio_pictorum
| | +-----+
+--+ | | +-------- Aculamprotula_tortuosa
| | | |
| | | +-------- Schistodesmus_lampreyanus
| | | |
| | +--+ +-- Nodularia_douglasiae
| | +-----+
| | | +-- Nodularia_nuxpersicae
| | |
| | | +----- Cuneopsis_heudei
| | +--+
| | | +-- Tchangsinaia_piscicula
| | +--+
| +--------+ +-- Cuneopsis_rufescens
| | |
| | | +-- Lanceolaria_grayii
| | | +-----------+
| | | | +-- Lanceolaria_gladiola
| | | |
| | | | +-- Cristaria_plicata
| | | | +--------+
| | | | | +-- Cristaria_radiata
| | | +--+
| | | | | +-------- Pletholophus_tenuis
| | | | +--+
| | | | | +----- Sinanodonta_lucida
| | +--+ +--+
| | | | +-- Sinanodonta_lauta
| | | +--+
| | | +-- Sinanodonta_jourdyi
| | |
| | | +-------- Anodonta_anatina
| | | |
| | | +-------- Pseudanodonta_complanata
| | | |
| | | | +-- Anodonta_cygnea
| | +-----+-----+
| | | +-- Anodonta_nuttalliana
| | |
| | | +----- Pyganodon_grandis
| | | |
| | | +----- Simpsonaias_ambigua
| | +--+
| | +----- Strophitus_undulatus
| | |
| | | +-- Alasmidonta_marginata
| | +--+
| | +-- Lasmigona_costata
| |
| | +-------- Lens_contradens
| | +-----+
+--+ | | +----- Lens_eximius
| | +--+
| | | +-- Lens_comptus
| | +--+
| +--+ +-- Lens_novoselovi
| | |
| | | +----- Physunio
| | | +-----+
| | | | | +-- Physunio_modelli
| | | | +--+
| | +--+ +-- Physunio_superbus
| | |
| | | +-------- Yaukthwa_inlenensis
| | +--+
| | | +----- Yaukthwa_panhai
| +-----+ +--+
| | | | +-- Yaukthwa_rectangularis
| | | +--+
| | | +-- Yaukthwa_nesemanni
| | |
| | | +----- Hyriopsis_panhai
| | | |
| | | +--------+----- Hyriopsis_khoratensis
| | | | |
| | | | | +-- Hyriopsis_myersiana
| | | | +--+
| | +--+ +-- Hyriopsis_bialata
| | |
| | | +----------- Hyriopsis_desowitzi
| | | |
| | | +----------- Rectidens_sumatrensis
| | +--+
| | | +-- Ensidens_telus
| | | +-----+
| | | | +-- Ensidens_ingallsianus
| | +--+
| | | +----- Ensidens_sagittarius
| | +--+
| | | +-- Ensidens_spiculus
+--+ +--+
| +-- Ensidens_jaculus
|
| +-------- Amblema_plicata
| +--+
| | | +----- Cambarunio_iris
| | +--+
| | | +-- Ortmanniana_ligamentina
| | +--+
| +--------+ +-- Lampsilis_cardium
| | |
| | | +-- Quadrula_quadrula
| | | +-----+
| | | | +-- Quadrula_verrucosa
| | +--+
| | | +----- Pleurobema_sintoxia
| | +--+
| | | +-- Elliptio_complanata
| | +--+
| | +-- Eurynia_dilatata
| |
| | +-------- Sinohyriopsis_cumingii
| | |
| | +-----+ +-- Lamprotula_caveata
+--+ | | +--+
| | | | +-- Lamprotula_leaii
| | +--+
| | | +-- Potomida_littoralis
| +--+ +--+
| | | +-- Pronodularia_japanensis
| | |
| | | +----------- Sinosolenaia_carinata
| | | |
| | | | +-- Sinosolenaia_oleivora
| | +--+--------+
| | | +-- Sinosolenaia_recognita
| | |
| | | +-------- Gonidea_angulata
| | +--+
| | | +----- Microcondylaea_bonellii
+--+ +--+
| | +-- Leguminaia_wheatleyi
| +--+
| +-- Leguminaia
|
| +-------- Pilsbryoconcha_exilis[3]
| +-----+
| | | +----- Pilsbryoconcha_exilis[2]
| | +--+
| | | +-- Pilsbryoconcha_exilis[1]
| | +--+
| | +-- Pilsbryoconcha_carinifera
| |
+--+ +----------- Indopseudodon_indawgyiensis
| |
| | +-------- Thaiconcha_callifera
| +--+
| | | +----- Bineurus_exilis
| | +--+
| | | +-- Bineurus_mouhotii
+--+ +--+
| +-- Bineurus_loeiensis
|
| +----- Pseudodon_vagulus
| +--+
| | | +-- Pseudodon_mekongi
| | +--+
| | +-- Pseudodon_vondembuschianus
+--+
| +-- Pseudodon_inoscularis
| +--+
| | +-- Sundadontina_tumida
+--+
| +-- Nyeinchanconcha_nyeinchani
+--+
+-- Sundadontina_cumingii
Graf et al., 2015 [Publ 3619], fig. 1
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 3 |
3 |
1 |
3 |
17 |
16 |
2 |
3.0 |
+-- Unio_pictorum
+-----+
| +-- Margaritifera_margaritifera
+--------+
| | +----- Chambardia_wahlbergi **
| +--+
| | +-- Tamsiella_schroeteriana **
| +--+
| +-- Etheria_elliptica **
|
| +----------- Westralunio_carteri
| |
--+ | +----- Alathyria_jacksoni
| +--+ +--+
| | | | | +-- Velesunio_ambiguus
| | | | +--+
| | +--+ +-- Alathyria_profuga
| | |
| | | +----- Alathyria_pertexta
| | +--+
| | | +-- Microdontia_anodontaeformis
+--+ +--+
| +-- Lortiella_froggatti
|
| +----- Diplodon_suavidicus
| +--+
| | | +-- Triplodon_corrugatus
| | +--+
| | +-- Castalia_ambigua
+-----+
| +-- Hyridella_depressa
| +--+
| | +-- Hyridella_drapeta
+--+
| +-- Hyridella_australis
+--+
+-- Cucumerunio_novaehollandiae
Pfeiffer & Graf, 2015 [Publ 3620], fig. 2
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 3 |
3 |
1 |
3 |
37 |
36 |
2 |
6.0 |
+-------- Velesunio_ambiguus
+--------------------------+
| | +----- Hyridella_australis
| +--+
| | +-- Triplodon_corrugatus
| +--+
| +-- Castalia_ambigua
|
--+ +----- Etheria_elliptica **
| +--------------------------+
| | | +-- Lamproscapha_ensiformis **
| | +--+
| | +-- Chambardia_wahlbergi **
| |
| | +-- Margaritifera_margaritifera
| | +--------------------------+
+--+ | +-- Cumberlandia_monodonta
| |
| | +-- Trapezidens_dolichorhynchus
| | +-----------+
| | | +-- Lamellidens_savadiensis
| | |
| | +-----------+ +-- Leoparreysia_olivacea
+--+ | | +--------+
| | | | +-- Leoparreysia_tavoyensis
| | +--+
| | | +-------- Coelatura_aegyptiaca
| | +--+
| | | +----- Indochinella_pugio
| | +--+
| | | +-- Radiatula_chaudhurii
| | +--+
+--+ +-- Scabies_mandarinus
|
| +----- Unio_pictorum
| +-----------------+
| | | +-- Pyganodon_grandis
| | +--+
| | +-- Lasmigona_costata
| |
| | +----------- Quadrula_quadrula
| | +--------+
+--+ | | +-------- Amblema_plicata
| | +--+
| | | +----- Potamilus_alatus
| | +--+
| | | +-- Lampsilis_cardium
| | +--+
| | +-- Truncilla_truncata
+--+
| +-------- Ensidens_telus
| +--------+
| | | +----- Lens_micropterus
| | +--+
| | | +-- Physunio
| | +--+
| | +-- Physunio_modelli
+--+
| +-------- Gonidea_angulata
| +-----+
| | | +----- Potomida_littoralis
| | +--+
| | | +-- Pronodularia_japanensis
+--+ +--+
| +-- Lamprotula_caveata
|
| +----------- Pilsbryoconcha_carinifera
| |
+--+ +-- Pseudodon_vondembuschianus
| +-----+
| | +-- Pseudodon_vagulus
+--+
| +----- Sundadontina_tumida
+--+
| +-- Nyeinchanconcha_nyeinchani
+--+
+-- Bineurus_exilis
Konopleva et al., 2017 [Publ 3807], fig. 6
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 3 |
3 |
1 |
3 |
28 |
26 |
2 |
5.0 |
+----- Diplodon_suavidicus
+--------------------+
| | +-- Triplodon_corrugatus
| +--+
| +-- Castalia_ambigua
|
| +----- Tamsiella_schroeteriana **
--+ +-----------------+
| | | +-- Etheria_elliptica **
| | +--+
| | +-- Chambardia_wahlbergi **
| |
| | +-- Margaritifera_dahurica
| | +-----------------+
+--+ | +-- Margaritifera_laevis
| |
| | +-------- Ensidens_sagittarius
| | +-----+
| | | | +----- Lens_micropterus
| | | +--+
| | | | +-- Physunio
| | +--+ +--+
+--+ | | +-- Physunio_modelli
| | |
| | | +----------- Sundadontina_tumida
| | | |
| | +--+ +-- Nyeinchanconcha_nyeinchani
| | | +-----+
| | | | +-- Bineurus_exilis
| | +--+
| | | +----- Pilsbryoconcha_carinifera
| | +--+
+--+ | +-- Pseudodon_vagulus
| +--+
| +-- Pseudodon_vondembuschianus
|
| +-- Trapezidens_exolescens
| +-----+
| | +-- Trapezidens_dolichorhynchus
| +-----+
| | | +----- Lamellidens_savadiensis
| | +--+
| | | +-- Lamellidens_jenkinsianus
| | +--+
+--+ +-- Lamellidens_mainwaringi
|
| +-- Leoparreysia_tavoyensis
| +--------+
| | +-- Leoparreysia_olivacea
+--+
| +-------- Coelatura_aegyptiaca
+--+
| +----- Indochinella_pugio
+--+
| +-- Radiatula_chaudhurii
+--+
+-- Scabies_mandarinus
Pfeiffer et al., 2019 [Publ 3937], fig. 4-5
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 3 |
3 |
1 |
3 |
54 |
53 |
2 |
5.0 |
+----- Tamsiella_schroeteriana **
+--------------------------------+
| | +-- Etheria_elliptica **
| +--+
| +-- Aspatharia_pfeifferiana **
|
| +----------------------------------- Hyridella_australis
| |
--+ | +-- Margaritifera_margaritifera
| | +-----------------------------+
| | | +-- Margaritifera_hembeli
| | |
| | | +-- Quadrula_quadrula
| | | +--------------------+
| | | | +-- Uniomerus_tetralasmus
+--+ | |
| | | +----- Elliptoideus_sloatianus
| | +-----+ +--------------+
| | | | | | +-- Elliptio_complanata
| | | | | +--+
| | | | | +-- Pleurobema_strodeanum
| | | | |
| | | +--+ +----- Reginaia_ebenus
| | | | +-----------+
| | | | | | +-- Plectomerus_dombeyanus
+--+ | | | +--+
| | | | +-- Amblema_plicata
| | +--+
| | | +-- Psoronaias_semigranosa
| | | +-----------+
| | | | +-- Popenaias_popeii
| | +--+
| | | +----------- Obliquaria_reflexa
| | +--+
| | | +-------- Cyrtonaias_tampicoensis
| | +--+
| | | +----- Leaunio_lienosus
| | +--+
| | | +-- Lampsilis_cardium
| | +--+
| | +-- Pachynaias_spheniopsis
| |
+--+ +-- Chamberlainia_hainesiana
| +--------------------+
| | +-- Sinohyriopsis_cumingii
| |
| | +----------------- Lamprotula_cornuumlunae
| | +--+
| | | | +-------------- Gonidea_angulata
| +--+ | +--+
| | | | | +----------- Indopseudodon_crebristriatus
| | | | +--+
| | | | | +-------- Bineurus_anodontinum
| | | | +--+
| | | | | +----- Pilsbryoconcha_exilis
| | | | +--+
| | +--+ | +-- Pseudodon_vondembuschianus
| | | +--+
| | | +-- Pseudodon_inoscularis
| | |
| | | +----- Ensidens_ingallsianus
| | | +-----+
| | | | | +-- Hyriopsis_bialata
| | | | +--+
| | | | +-- Rectidens_sumatrensis
| | +--------+
| | | +-- Trapezoideus_foliaceus
| | | +-----+
+--+ | | +-- Physunio_superbus
| +--+
| | +----- Yaukthwa_paiensis
| +--+
| | +-- Lens_contradens
| +--+
| +-- Solenaia_emarginata
|
| +----------- Lanceolaria_lanceolata
| |
| | +----- Unio_gibbus
| +--------+ +--+
| | | | | +-- Nodularia_douglasiae
| | | | +--+
| | | | +-- Tchangsinaia_piscicula
| | +--+
| | | +-- Cristaria_plicata
| | | +--+
| | | | +-- Sinanodonta_woodiana
+-----+ +--+
| | +-- Anodonta_cygnea
| +--+
| +-- Pyganodon_grandis
|
| +-- Trapezidens_dolichorhynchus
| +--------------+
| | +-- Lamellidens_savadiensis
| |
+--+ +-- Leoparreysia_tavoyensis
| +-----------+
| | +-- Leoparreysia_olivacea
| |
+--+ +-- Coelatura_choziensis
| +--------+
| | +-- Prisodontopsis_aviculaeformis
+--+
| +-------- Indonaia_caerulea
+--+
| +----- Indochinella_pugio
+--+
| +-- Scabies_phaselus
+--+
+-- Harmandia_somboriensis
Pfeiffer et al., 2019 [Publ 3937], fig. 6
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 3 |
3 |
1 |
3 |
54 |
53 |
2 |
5.0 |
+----- Tamsiella_schroeteriana **
+-----------------------+
| | +-- Etheria_elliptica **
| +--+
| +-- Aspatharia_pfeifferiana **
|
| +----- Hyridella_australis
| +--------------------+
| | | +-- Margaritifera_margaritifera
| | +--+
| | +-- Margaritifera_hembeli
| |
| | +-- Quadrula_quadrula
| | +-----------------+
| | | +-- Uniomerus_tetralasmus
--+ | |
| | | +----- Elliptoideus_sloatianus
| | | +--+
| | | | | +-- Elliptio_complanata
| | +--+ | +--+
| | | | +--------+ +-- Pleurobema_strodeanum
| | | | | |
| | | | | | +----- Reginaia_ebenus
| | | | | +--+
| | | | | | +-- Plectomerus_dombeyanus
| | | +--+ +--+
| | | | +-- Amblema_plicata
| | | |
| | | | +-- Psoronaias_semigranosa
| | | | +-----------+
+--+ | | | +-- Popenaias_popeii
| | +--+
| | | +----------- Obliquaria_reflexa
| | +--+
| | | +-------- Cyrtonaias_tampicoensis
| | +--+
| | | +----- Leaunio_lienosus
| | +--+
| | | +-- Lampsilis_cardium
| | +--+
| | +-- Pachynaias_spheniopsis
| |
| | +-- Chamberlainia_hainesiana
| | +-----------------+
| | | +-- Sinohyriopsis_cumingii
| | |
| | | +-- Gonidea_angulata
| | | +-----------+
| | | | +-- Lamprotula_cornuumlunae
| | | +--+
| +--+ | | +----------- Indopseudodon_crebristriatus
| | | | +--+
| | | | | +-------- Pilsbryoconcha_exilis
| | | | +--+
+--+ | | | +----- Bineurus_anodontinum
| | | +--+
| | | | +-- Pseudodon_vondembuschianus
| +--+ +--+
| | +-- Pseudodon_inoscularis
| |
| | +----- Ensidens_ingallsianus
| | +-----+
| | | | +-- Hyriopsis_bialata
| | | +--+
| | | +-- Rectidens_sumatrensis
| +-----+
| | +-- Trapezoideus_foliaceus
| | +-----+
| | | +-- Physunio_superbus
| +--+
| | +----- Yaukthwa_paiensis
| +--+
| | +-- Lens_contradens
| +--+
| +-- Solenaia_emarginata
|
| +-------- Lanceolaria_lanceolata
| +--+
| | | +----- Unio_gibbus
| | +--+
| | | +-- Tchangsinaia_piscicula
| | +--+
| +--------+ +-- Nodularia_douglasiae
| | |
| | | +-- Cristaria_plicata
| | | +--+
| | | | +-- Sinanodonta_woodiana
| | +-----+
| | | +-- Anodonta_cygnea
+--+ +--+
| +-- Pyganodon_grandis
|
| +-- Trapezidens_dolichorhynchus
| +--------------+
| | +-- Lamellidens_savadiensis
| |
+--+ +-- Leoparreysia_tavoyensis
| +-----------+
| | +-- Leoparreysia_olivacea
| |
+--+ +-- Coelatura_choziensis
| +--------+
| | +-- Prisodontopsis_aviculaeformis
+--+
| +-------- Indonaia_caerulea
+--+
| +----- Indochinella_pugio
+--+
| +-- Scabies_phaselus
+--+
+-- Harmandia_somboriensis
Elderkin et al., 2016 [Publ 3693], fig. 2
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 2 |
3 |
0 |
2 |
1 |
1 |
1 |
5.0 |
+-- Etheria_elliptica **
|
--+-- Aspatharia_pfeifferiana **
|
+-- Velesunio_ambiguus
Guerra et al., 2017 [Publ 3772], fig. 2a
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 2 |
2 |
0 |
2 |
23 |
22 |
2 |
-- |
+-- Mutela_dubia **
+-----------------------------+
| +-- Anodontites_trapesialis **
|
| +----------------------------- Echyridella_menziesii
--+ |
| | +-- Margaritifera_falcata
| | +-----------------------+
| | | +-- Cumberlandia_monodonta
+--+ |
| | +-------- Quadrula_quadrula
| | +--+
| | | | +----- Toxolasma_parvum
| | | +--+
| | | | +-- Venustaconcha_ellipsiformis
+--+ | +--+
| +-----------+ +-- Lampsilis_ornata
| | |
| | | +-- Sinohyriopsis_schlegelii
| | | +-----+
| | | | +-- Sinohyriopsis_cumingii
| | +--+
| | | +----- Pronodularia_japanensis
+--+ +--+
| | +-- Sinosolenaia_oleivora
| +--+
| +-- Sinosolenaia_carinata
|
| +-- Aculamprotula_tortuosa
| +-----------------+
| | +-- Unio_pictorum
+--+
| +----------------- Sinanodonta_woodiana
| |
+--+ +-------------- Cristaria_plicata
| |
+--+ +----------- Anodonta_anatina
| |
+--+ +-- Platynaias_subviridis
| +-----+
| | +-- Platynaias_compressa
+--+
| +----- Pyganodon_grandis
+--+
| +-- Utterbackia_peninsularis
+--+
+-- Utterbackia_imbecillis
Guerra et al., 2017 [Publ 3772], fig. 2b
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 2 |
2 |
0 |
2 |
23 |
22 |
2 |
-- |
+-- Mutela_dubia **
+--------------------------------+
| +-- Anodontites_trapesialis **
|
--+ +-------------------------------- Echyridella_menziesii
| |
| | +-- Margaritifera_falcata
| | +--------------------------+
+--+ | +-- Cumberlandia_monodonta
| |
| | +-------- Quadrula_quadrula
| | +-----------------+
+--+ | | +----- Toxolasma_parvum
| | +--+
| | | +-- Venustaconcha_ellipsiformis
| | +--+
| | +-- Lampsilis_ornata
| |
+--+ +-- Sinohyriopsis_schlegelii
| +-----+
| | +-- Sinohyriopsis_cumingii
| +--------------+
| | | +----- Pronodularia_japanensis
| | +--+
| | | +-- Sinosolenaia_oleivora
+--+ +--+
| +-- Sinosolenaia_carinata
|
| +-------------------- Unio_pictorum
| |
+--+ +----------------- Aculamprotula_tortuosa
| |
+--+ +-- Cristaria_plicata
| +-----------+
| | +-- Sinanodonta_woodiana
+--+
| +----------- Anodonta_anatina
| |
+--+ +-- Platynaias_subviridis
| +-----+
| | +-- Platynaias_compressa
+--+
| +----- Pyganodon_grandis
+--+
| +-- Utterbackia_peninsularis
+--+
+-- Utterbackia_imbecillis
Huang et al., 2018 [Publ 3828], fig. 6
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 2 |
2 |
0 |
2 |
41 |
40 |
2 |
-- |
+-- Anodontites_trapesialis **
+--------------------------+
| +-- Mutela_dubia **
|
| +-------------------------- Echyridella_menziesii
| |
| | +-------------- Gibbosula_rochechouartii
--+ | +--------+
| | | | +----------- Cumberlandia_monodonta
| | | +--+
| | | | +-------- Pseudunio_marocanus
| | | +--+
| | | | +----- Margaritifera_falcata
| | | +--+
+--+ | | +-- Margaritifera_margaritifera
| | +--+
| | +-- Margaritifera_dahurica
| |
| | +----------- Quadrula_quadrula
| | |
| | +-----+ +-------- Toxolasma_parvum
| | | | |
| | | +--+ +-- Lampsilis_ornata
| | | | +--+
| | | | | +-- Venustaconcha_ellipsiformis
+--+ | +--+
| | | +-- Potamilus_leptodon
| +--+ +--+
| | | +-- Potamilus_alatus
| | |
| | | +-- Sinohyriopsis_cumingii
| | | +-----------+
| | | | +-- Sinohyriopsis_schlegelii
| | | |
| | +--+ +-- Sinosolenaia_oleivora
| | | +--------+
| | | | +-- Sinosolenaia_carinata
| | +--+
| | | +-------- Potomida_littoralis
| | +--+
| | | +----- Pronodularia_japanensis
| | +--+
| | | +-- Lamprotula_caveata
+--+ +--+
| +-- Lamprotula_leaii
|
| +----- Aculamprotula_tortuosa
| +--------+
| | | +-- Aculamprotula_coreana
| | +--+
| | +-- Aculamprotula_tientsinensis
| +--+
| | | +-- Tchangsinaia_piscicula
| | | +--------+
| | | | +-- Nodularia_douglasiae
| | +--+
| | | +-------- Unio_tumidus
| | +--+
| | | +----- Unio_crassus
| | +--+
+--+ | +-- Unio_delphinus
| +--+
| +-- Unio_pictorum
|
| +----------- Anodonta_anatina
| +--+
| | | +-------- Platynaias_compressa
| | +--+
| | | +----- Pyganodon_grandis
| | +--+
+--+ | +-- Utterbackia_imbecillis
| +--+
| +-- Utterbackia_peninsularis
|
| +-------- Cristaria_plicata
| |
+-----+ +-- Anemina_arcaeformis
| +--+
| | +-- Anemina_euscaphys
+--+
| +-- Sinanodonta_woodiana
+--+
+-- Sinanodonta_lucida
Huang et al., 2019 [Publ 3843], fig. 3
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 2 |
2 |
0 |
2 |
55 |
54 |
2 |
-- |
+-- Anodontites_trapesialis **
+--------------------------------------+
| +-- Mutela_dubia **
|
| +-------------------------------------- Echyridella_menziesii
| |
| | +-------------- Gibbosula_rochechouartii
| | +--------------------+
--+ | | | +----------- Cumberlandia_monodonta
| | | +--+
| | | | +-------- Pseudunio_marocanus
| | | +--+
| | | | +----- Margaritifera_falcata
| | | +--+
| | | | +-- Margaritifera_margaritifera
| | | +--+
+--+ | +-- Margaritifera_dahurica
| |
| | +-- Sinohyriopsis_cumingii
| | +-----------+
| | | +-- Sinohyriopsis_schlegelii
| | |
| | | +-------- Potomida_littoralis
| | +--------+ +--+
| | | | | | +----- Pronodularia_japanensis
| | | | | +--+
| | | | | | +-- Lamprotula_caveata
| | | | | +--+
+--+ | +--+ +-- Lamprotula_leaii
| | |
| | | +-- Ptychorhynchus_pfisteri
| | | +--+
| +--------+ | | +-- Parvasolenaia_rivularis
| | | +-----+
| | | | +-- Sinosolenaia_oleivora
| | | +--+
| | | +-- Sinosolenaia_carinata
| | |
| | | +-- Quadrula_quadrula
| | | +-----------------+
| | | | +-- Uniomerus_tetralasmus
| | +--+
| | | +----------------- Elliptio_complanata
| | | |
| | +--+ +-------------- Toxolasma_parvum
| | | |
| | +--+ +-- Potamilus_leptodon
| | | +--------+
| | | | +-- Potamilus_alatus
+--+ +--+
| | +-------- Leaunio_lienosus
| +--+
| | +----- Venustaconcha_ellipsiformis
| +--+
| | +-- Lampsilis_cardium
| +--+
| +-- Lampsilis_ornata
|
| +----------------------------- Lepidodesma_languilati
| |
| | +-------- Aculamprotula_tortuosa
| | +-----+
| | | | +----- Aculamprotula_scripta
| | | +--+
| | | | +-- Aculamprotula_tientsinensis
| | | +--+
| | +-----------+ +-- Aculamprotula_coreana
+--+ | |
| | | +----- Nodularia_douglasiae
| | | +-----+
| | | | | +-- Tchangsinaia_piscicula
| | | | +--+
| | +--+ +-- Schistodesmus_lampreyanus
| | |
| | | +-------- Unio_tumidus
| | +--+
+--+ | +----- Unio_crassus
| +--+
| | +-- Unio_delphinus
| +--+
| +-- Unio_pictorum
|
| +----- Lanceolaria_gladiola
| +-----------------+
| | | +-- Lanceolaria_lanceolata
| | +--+
| | +-- Lanceolaria_grayii
+--+
| +-------------------- Acuticosta_chinensis
| |
| | +-------- Cristaria_plicata
| | |
+--+ +--------+ +-- Sinanodonta_lucida
| | | +--+
| | | | +-- Sinanodonta_woodiana
| | +--+
+--+ | +-- Anemina_arcaeformis
| +--+
| +-- Anemina_euscaphys
|
| +-------------- Anodonta_cygnea
+--+
| +----------- Anodonta_anatina
+--+
| +-------- Lasmigona_costata
+--+
| +----- Pyganodon_grandis
+--+
| +-- Utterbackia_imbecillis
+--+
+-- Utterbackia_peninsularis
Lemer et al., 2019 [Publ 4093], fig. 1
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 2 |
2 |
0 |
2 |
4 |
2 |
1 |
-- |
+-- Etheria_elliptica **
+--+
| +-- Aspatharia_pfeifferiana **
--+
| +-- Margaritifera_margaritifera
+--+
+-- Lampsilis_cardium
Wu et al., 2020 [Publ 4121], fig. 3
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 2 |
2 |
0 |
2 |
48 |
47 |
2 |
-- |
+-------------------------------------- Echyridella_menziesii
|
| +-- Anodontites_trapesialis **
| +--------------------------------+
| | +-- Mutela_dubia **
--+ |
| | +-------- Gibbosula_rochechouartii
| | +-----------------------+
| | | | +----- Cumberlandia_monodonta
| | | +--+
+--+ | | +-- Margaritifera_falcata
| | +--+
| | +-- Margaritifera_dahurica
| |
| | +----------- Quadrula_quadrula
| | |
| | +-----------------+ +-------- Toxolasma_parvum
+--+ | | |
| | +--+ +-- Venustaconcha_ellipsiformis
| | | +--+
| | | | +-- Lampsilis_ornata
| | +--+
| | | +-- Potamilus_alatus
| | +--+
| | +-- Potamilus_leptodon
| |
| | +-- Sinohyriopsis_schlegelii
| | +--------+
+--+ | +-- Sinohyriopsis_cumingii
| |
| +--------------+ +----- Potomida_littoralis
| | | +--+
| | | | | +-- Pronodularia_japanensis
| | | | +--+
| | +--+ +-- Lamprotula_leaii
| | |
| | | +----- Ptychorhynchus_pfisteri
| | +--+
| | | +-- Sinosolenaia_oleivora
| | +--+
| | +-- Sinosolenaia_carinata
+--+
| +----------------------- Lepidodesma_languilati
| |
| | +----- Acuticosta_chinensis
| | +-----------+
| | | | +-- Lanceolaria_lanceolata
| | | +--+
| | | +-- Lanceolaria_grayii
| | |
| | +--+ +----------- Anodonta_anatina
| | | | +--+
| | | | | | +-------- Platynaias_compressa
+--+ | | | +--+
| | | | | +----- Pyganodon_grandis
| | | | +--+
| | +--+ | +-- Utterbackia_peninsularis
| | | +--+
| | | +-- Utterbackia_imbecillis
| | |
| | | +-------- Cristaria_plicata
| | | |
| | +-----+ +-- Sinanodonta_woodiana
| | | +--+
+--+ | | +-- Sinanodonta_lauta
| +--+
| | +-- Anemina_arcaeformis
| +--+
| +-- Anemina_euscaphys
|
| +-------- Aculamprotula_tortuosa
| |
| +--------+ +-- Aculamprotula_coreana
| | | +--+
| | | | +-- Aculamprotula_tientsinensis
| | +--+
| | | +-- Aculamprotula_polysticta
| | +--+
+--+ +-- Aculamprotula_scripta
|
| +-------- Unio_tumidus
| +-----+
| | | +----- Unio_crassus
| | +--+
| | | +-- Unio_delphinus
+--+ +--+
| +-- Unio_pictorum
|
| +----------- Pseudocuneopsis_capitata
+--+
| +-------- Nodularia_douglasiae
+--+
| +----- Tchangsinaia_piscicula
+--+
| +-- Cuneopsis_heudei
+--+
+-- Schistodesmus_lampreyanus
Wu et al., 2022 [Publ 4238], fig. 3
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 2 |
2 |
0 |
2 |
72 |
72 |
2 |
-- |
+-- Mutela_dubia **
+--------------------------------------------+
| +-- Anodontites_trapesialis **
|
| +-------------------------------------------- Echyridella_menziesii
| |
| | +-------------- Gibbosula_rochechouartii
| | +--------------------------+
--+ | | | +----------- Cumberlandia_monodonta
| | | +--+
| | | | +-------- Pseudunio_marocanus
| | | +--+
| | | | +----- Margaritifera_falcata
| | | +--+
| | | | +-- Margaritifera_dahurica
+--+ | +--+
| | +-- Margaritifera_margaritifera
| |
| | +-- Uniomerus_tetralasmus
| | +--------------------------+
| | | +-- Quadrula_quadrula
| | |
| | +--------+ +-- Pleurobema_oviforme
| | | | +-----------------------+
| | | | | +-- Elliptio_complanata
| | | +--+
+--+ | | +----------------------- Amblema_plicata
| | | |
| | +--+ +-------------------- Popenaias_popeii
| | | |
| | +--+ +----------------- Toxolasma_parvum
| | | |
| | +--+ +-- Potamilus_alatus
| | | +-----------+
| | | | +-- Potamilus_leptodon
| | +--+
| | | +----------- Leaunio_lienosus
| | | |
| | +--+ +-- Lampsilis_cardium
| | | +-----+
| | | | +-- Lampsilis_ornata
| | +--+
| | | +----- Venustaconcha_ellipsiformis
| | +--+
| | | +-- Lampsilis_siliquoidea
| | +--+
+--+ +-- Lampsilis_powellii
|
| +----- Chamberlainia_somsakpanhai
| +-----------+
| | | +-- Sinohyriopsis_schlegelii
| | +--+
| | +-- Sinohyriopsis_cumingii
| |
| +-----------------+ +-------- Potomida_littoralis
| | | +-----+
| | | | | +----- Pronodularia_japanensis
| | | | +--+
| | | | | +-- Lamprotula_caveata
| | | | +--+
| | +--+ +-- Lamprotula_leaii
| | |
| | | +-- Pilsbryoconcha_exilis
| | | +--------+
| | | | +-- Pseudodon_vondembuschianus
| | | |
| | +--+ +-- Ptychorhynchus_pfisteri
| | | +-----+
| | | | +-- Parvasolenaia_rivularis
| | +--+
| | | +----- Microcondylaea_bonellii
+--+ +--+
| | +-- Sinosolenaia_carinata
| +--+
| +-- Sinosolenaia_recognita
|
| +-------------------------------- Lepidodesma_languilati
| |
| | +-------- Acuticosta_chinensis
| | +--------+
| | | | +----- Lanceolaria_gladiola
| | | +--+
| | | | +-- Lanceolaria_grayii
| | | +--+
| | | +-- Lanceolaria_lanceolata
| | |
| | +-----------+ +-------- Cristaria_plicata
| | | | |
+--+ | | +-----+ +-- Sinanodonta_woodiana
| | | | | +--+
| | | | | | +-- Sinanodonta_lucida
| | | | +--+
| | | | | +-- Anemina_arcaeformis
| | +--+ +--+
| | | +-- Anemina_euscaphys
| | |
| | | +-- Anodonta_cygnea
| | | +--------+
| | | | +-- Anodonta_anatina
| | +--+
| | | +-------- Platynaias_compressa
+--+ +--+
| | +----- Pyganodon_grandis
| +--+
| | +-- Utterbackia_imbecillis
| +--+
| +-- Utterbackia_peninsularis
|
| +-------- Aculamprotula_tortuosa
| |
| +-----------------+ +-- Aculamprotula_tientsinensis
| | | +--+
| | | | +-- Aculamprotula_coreana
| | +--+
| | | +-- Aculamprotula_polysticta
| | +--+
| | +-- Aculamprotula_scripta
+--+
| +-------- Unio_tumidus
| +--------------+
| | | +----- Unio_crassus
| | +--+
| | | +-- Unio_pictorum
| | +--+
+--+ +-- Unio_delphinus
|
| +-- Pseudocuneopsis_capitata
| +-----------------+
| | +-- Pseudocuneopsis_sichuanensis
| |
+--+ +-- Nodularia_douglasiae
| +--------------+
| | +-- Nodularia_breviconcha
| |
+--+ +-- Schistodesmus_lampreyanus
| +-----------+
| | +-- Schistodesmus_spinosus
+--+
| +----------- Tchangsinaia_piscicula
+--+
| +-------- Cuneopsis_rufescens
+--+
| +----- Cuneopsis_celtiformis
+--+
| +-- Cuneopsis_heudei
+--+
+-- Cuneopsis_demangei
Smith, 2021 [Publ 4331], fig. 1
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 2 |
2 |
0 |
2 |
18 |
17 |
1 |
-- |
+-- Etheria_elliptica **
+-----------------------+
| +-- Aspatharia_pfeifferiana **
|
| +----------------------- Margaritifera_margaritifera
--+ |
| | +----- Sinohyriopsis_cumingii
| | +-----+
| | | | +-- Lens_contradens
| | | +--+
+--+ +--------+ +-- Lamprotula_leaii
| | |
| | | +-------- Scabies_phaselus
| | +--+
| | | +----- Cristaria_plicata
| | +--+
| | | +-- Pyganodon_grandis
+--+ +--+
| +-- Utterbackia_peninsularis
|
| +-- Uniomerus_tetralasmus
| +--------------+
| | +-- Megalonaias_nervosa
| |
+--+ +-- Elliptio_crassidens
| +-----------+
| | +-- Elliptio_complanata
+--+
| +----------- Amblema_plicata
+--+
| +-------- Potamilus_streckersoni
+--+
| +----- Venustaconcha_ellipsiformis
+--+
| +-- Lampsilis_siliquoidea
+--+
+-- Lampsilis_cardium
Wu et al., 2023 [Publ 4339], fig. 4, S5-aa
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 2 |
2 |
0 |
2 |
94 |
93 |
2 |
-- |
+----------------------------------------------- Echyridella_menziesii
|
| +-- Anodontites_trapesialis **
| +-----------------------------------------+
| | +-- Mutela_dubia **
| |
| | +-- Gibbosula_crassa
| | +-----------+
--+ | | +-- Gibbosula_rochechouartii
| | +--------------------------+
| | | | +----------- Cumberlandia_monodonta
| | | +--+
| | | | +-------- Pseudunio_marocanus
| | | +--+
| | | | +----- Margaritifera_falcata
| | | +--+
| | | | +-- Margaritifera_dahurica
+--+ | +--+
| | +-- Margaritifera_margaritifera
| |
| | +-- Quadrula_quadrula
| | +--------------------------+
| | | +-- Uniomerus_tetralasmus
| | |
| | +--------+ +-- Elliptio_complanata
| | | | +-----------------------+
| | | | | +-- Pleurobema_oviforme
| | | +--+
| | | | +----------------------- Amblema_plicata
| | | | |
| | | +--+ +-------------------- Popenaias_popeii
+--+ | | |
| | +--+ +----------------- Toxolasma_parvum
| | | |
| | | | +----- Potamilus_leptodon
| | +--+ +--------+
| | | | | +-- Potamilus_alatus
| | | | +--+
| | | | +-- Potamilus_streckersoni
| | +--+
| | | +----------- Leaunio_lienosus
| | | |
| | | | +----- Lampsilis_ornata
| | +--+ +--+
| | | | | +-- Lampsilis_cardium
| | | | +--+
| | +--+ +-- Lampsilis_cariosa
| | |
| | | +----- Venustaconcha_ellipsiformis
| | +--+
| | | +-- Lampsilis_powellii
| | +--+
| | +-- Lampsilis_siliquoidea
+--+
| +----- Chamberlainia_somsakpanhai
| +-----------------+
| | | +-- Sinohyriopsis_cumingii
| | +--+
| | +-- Sinohyriopsis_schlegelii
| |
| | +-- Hyriopsis_bialata
| | +--+
| +-----------+ | +-- Rectidens_sumatrensis
| | | +--------------+
| | | | | +-- Lens_contradens
| | | | +--+
| | | | +-- Physunio_superbus
| | | |
| | | | +----------- Potomida_littoralis
| | +--+ +-----+
| | | | | +-------- Pronodularia_japanensis
| | | | +--+
| | | | | +----- Lamprotula_leaii
| | | | +--+
| | | | | +-- Lamprotula_caveata
| | +--+ +--+
| | | +-- Lamprotula_cornuumlunae
| | |
| | | +-- Pseudodon_vondembuschianus
| | | +-----------+
| | | | +-- Pilsbryoconcha_exilis
| | +--+
| | | +----------- Microcondylaea_bonellii
+--+ | |
| +--+ +-- Parvasolenaia_rivularis
| | +-----+
| | | +-- Ptychorhynchus_pfisteri
| +--+
| | +----- Sinosolenaia_carinata
| +--+
| | +-- Sinosolenaia_iridinea
| +--+
| +-- Sinosolenaia_oleivora
|
| +-------------------------------- Lamellidens_marginalis
| |
| | +-- Lepidodesma_aligera
| | +--------------------------+
| | | +-- Lepidodesma_languilati
| | |
| | | +-------- Aculamprotula_tortuosa
| | | |
| | | +--------------+ +-- Aculamprotula_coreana
| | | | | +--+
+--+ | | | | +-- Aculamprotula_tientsinensis
| | | +--+
| | | | +-- Aculamprotula_polysticta
| | | +--+
| | | +-- Aculamprotula_scripta
| | |
| | +--+ +-------------- Unio_tumidus
| | | | +-----+
| | | | | | +----------- Unio_crassus
| | | | | +--+
+--+ | | | | +-------- Unio_elongatulus
| | | | +--+
| | | | | +----- Unio_pictorum
| | | | +--+
| | | | | +-- Unio_delphinus
| | +--+ +--+
| | | +-- Unio_mancus
| | |
| | | +-- Nodularia_breviconcha
| | | +--+
| | | | +-- Nodularia_nipponensis
| | | +-----------+
| | | | | +-- Pseudocuneopsis_capitata
| | | | +--+
| | +--+ +-- Pseudocuneopsis_sichuanensis
| | |
| | | +-------------- Tchangsinaia_piscicula
| | | |
+--+ +--+ +-- Schistodesmus_lampreyanus
| | +--------+
| | | +-- Schistodesmus_spinosus
| +--+
| | +-------- Cuneopsis_rufescens
| +--+
| | +----- Cuneopsis_celtiformis
| +--+
| | +-- Cuneopsis_demangei
| +--+
| +-- Cuneopsis_heudei
|
| +-------- Acuticosta_chinensis
| +--------+
| | | +----- Lanceolaria_gladiola
| | +--+
| | | +-- Lanceolaria_grayii
| | +--+
| | +-- Lanceolaria_lanceolata
| |
| | +----- Cristaria_plicata
| | +-----+
| | | | +-- Anemina_arcaeformis
+--------+ | +--+
| +--+ +-- Anemina_euscaphys
| | |
| | | +-------- Beringiana_fukuharai
| | +--+
| | | +----- Sinanodonta_lucida
| | +--+
| | | +-- Sinanodonta_tumens
| | +--+
| | +-- Sinanodonta_woodiana
+--+
| +-- Anodonta_nuttalliana
| +-----+
| | +-- Pseudanodonta_complanata
| +--+
| | | +----- Anodonta_anatina
| | +--+
| | | +-- Anodonta_cygnea
| | +--+
| | +-- Anodonta_exulcerata
+--+
| +----- Pyganodon_grandis
| +--+
| | | +-- Utterbackia_peninsularis
| | +--+
| | +-- Utterbackia_imbecillis
+--+
| +-- Alasmidonta_varicosa
| +--+
| | +-- Platynaias_compressa
+--+
| +-- Prolasmidonta_heterodon
+--+
+-- Lasmigona_complanata
Wu et al., 2023 [Publ 4339], fig. 4, S5-nt
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 2 |
2 |
0 |
2 |
95 |
94 |
2 |
-- |
+-------------------------------------------------- Echyridella_menziesii
|
| +-- Anodontites_trapesialis **
| +--------------------------------------------+
| | +-- Mutela_dubia **
| |
| | +-- Gibbosula_crassa
| | +-----------+
--+ | | +-- Gibbosula_rochechouartii
| | +-----------------------------+
| | | | +----------- Cumberlandia_monodonta
| | | +--+
| | | | +-------- Pseudunio_marocanus
| | | +--+
| | | | +----- Margaritifera_falcata
| | | +--+
| | | | +-- Margaritifera_dahurica
+--+ | +--+
| | +-- Margaritifera_margaritifera
| |
| | +-- Quadrula_quadrula
| | +--------------------------+
| | | +-- Uniomerus_tetralasmus
| | |
| | +-----------+ +-- Elliptio_complanata
| | | | +-----------------------+
| | | | | +-- Pleurobema_oviforme
| | | +--+
| | | | +----------------------- Amblema_plicata
| | | | |
| | | +--+ +-------------------- Popenaias_popeii
+--+ | | |
| | +--+ +----------------- Toxolasma_parvum
| | | |
| | | | +----- Potamilus_leptodon
| | +--+ +--------+
| | | | | +-- Potamilus_alatus
| | | | +--+
| | | | +-- Potamilus_streckersoni
| | +--+
| | | +----------- Leaunio_lienosus
| | | |
| | | | +----- Lampsilis_ornata
| | +--+ +--+
| | | | | +-- Lampsilis_cardium
| | | | +--+
| | +--+ +-- Lampsilis_cariosa
| | |
| | | +----- Venustaconcha_ellipsiformis
| | +--+
| | | +-- Lampsilis_powellii
| | +--+
| | +-- Lampsilis_siliquoidea
+--+
| +----- Chamberlainia_somsakpanhai
| +-----------------+
| | | +-- Sinohyriopsis_cumingii
| | +--+
| | +-- Sinohyriopsis_schlegelii
| |
| | +-- Hyriopsis_bialata
| | +--+
| +--------------+ | +-- Rectidens_sumatrensis
| | | +--------------+
| | | | | +-- Lens_contradens
| | | | +--+
| | | | +-- Physunio_superbus
| | | |
| | | | +----------- Potomida_littoralis
| | +--+ +-----+
| | | | | +-------- Pronodularia_japanensis
| | | | +--+
| | | | | +----- Lamprotula_leaii
| | | | +--+
| | | | | +-- Lamprotula_caveata
| | +--+ +--+
| | | +-- Lamprotula_cornuumlunae
| | |
| | | +-- Pseudodon_vondembuschianus
| | | +-----------+
| | | | +-- Pilsbryoconcha_exilis
| | | |
| | +--+ +-- Ptychorhynchus_pfisteri
+--+ | +--------+
| | | +-- Microcondylaea_bonellii
| +--+
| | +-------- Parvasolenaia_rivularis
| +--+
| | +----- Sinosolenaia_carinata
| +--+
| | +-- Sinosolenaia_iridinea
| +--+
| +-- Sinosolenaia_oleivora
|
| +----------------------------------- Lamellidens_marginalis
| |
| | +-- Lepidodesma_aligera
| | +-----------------------------+
| | | +-- Lepidodesma_languilati
| | |
| | | +-------- Acuticosta_chinensis
| | | +--------------+
| | | | | +----- Lanceolaria_gladiola
| | | | +--+
+--+ | | | +-- Lanceolaria_grayii
| | | +--+
| | | +-- Lanceolaria_lanceolata
| | |
| | | +-------------- Cristaria_plicata
| | +-----+ |
| | | | +-----+ +-- Anemina_arcaeformis
| | | | | | +--------+
| | | | | | | +-- Anemina_euscaphys
| | | | | +--+
+--+ | | | | +-------- Beringiana_fukuharai
| | | | +--+
| | | | | +----- Sinanodonta_lucida
| | | | +--+
| | +--+ | +-- Sinanodonta_tumens
| | | +--+
| | | +-- Sinanodonta_woodiana
| | |
| | | +----------------- Anodonta_nuttalliana
| | | |
| | | +----------------- Pseudanodonta_complanata
| | | |
| | +--+ +-------------- Anodonta_cygnea
| | | |
| | | | +-- Anodonta_exulcerata
| | | +-----------+
| | | | +-- Anodonta_anatina
| | +--+
+--+ | +----- Pyganodon_grandis
| | +-----+
| | | | +-- Utterbackia_imbecillis
| | | +--+
| +--+ +-- Utterbackia_peninsularis
| |
| | +-------- Lasmigona_complanata
| +--+
| | +----- Prolasmidonta_heterodon
| +--+
| | +-- Alasmidonta_varicosa
| +--+
| +-- Platynaias_compressa
|
| +-------- Aculamprotula_tortuosa
| |
| +-----------------+ +-- Aculamprotula_coreana
| | | +--+
| | | | +-- Aculamprotula_tientsinensis
| | +--+
| | | +-- Aculamprotula_polysticta
| | +--+
| | +-- Aculamprotula_scripta
| |
+--+ +----------- Unio_tumidus
| |
| +-----------+ +-------- Unio_crassus
| | | |
| | +--+ +----- Unio_elongatulus
| | | |
| | +--+ +-- Unio_delphinus
| | | |
| | +--+-- Unio_mancus
+--+ |
| +-- Unio_pictorum
|
| +-- Pseudocuneopsis_capitata
| +-----------------+
| | +-- Pseudocuneopsis_sichuanensis
| |
+--+ +-- Nodularia_breviconcha
| +--------------+
| | +-- Nodularia_nipponensis
| |
+--+ +----- Tchangsinaia_piscicula
| +--------+
| | | +-- Schistodesmus_lampreyanus[2]
| | +--+
+--+ +-- Schistodesmus_spinosus
|
| +----------- Schistodesmus_lampreyanus[1]
+--+
| +-------- Cuneopsis_rufescens
+--+
| +----- Cuneopsis_celtiformis
+--+
| +-- Cuneopsis_demangei
+--+
+-- Cuneopsis_heudei
Wu et al., 2024 [Publ 4350], fig. 8a, S1
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 2 |
2 |
0 |
2 |
90 |
89 |
2 |
-- |
+----------------------------------------------- Echyridella_menziesii
|
| +-- Anodontites_trapesialis **
| +-----------------------------------------+
| | +-- Mutela_dubia **
| |
| | +-- Margaritifera_dahurica
--+ | +--------------------------------------+
| | | +-- Margaritifera_margaritifera
| | |
| | | +-- Quadrula_quadrula
| | | +--------------------------+
| | | | +-- Uniomerus_tetralasmus
| | | |
+--+ | +--------+ +-- Elliptio_complanata
| | | | +-----------------------+
| | | | | +-- Pleurobema_oviforme
| | | +--+
| | | | +----------------------- Amblema_plicata
| | | | |
| | | +--+ +-------------------- Popenaias_popeii
| | | | |
| | | +--+ +----------------- Toxolasma_parvum
| | | | |
| | | | | +----- Potamilus_leptodon
+--+ | +--+ +--------+
| | | | | +-- Potamilus_alatus
| | | | +--+
| | | | +-- Potamilus_streckersoni
| | +--+
| | | +----------- Leaunio_lienosus
| | | |
| | | | +----- Lampsilis_ornata
| | +--+ +--+
| | | | | +-- Lampsilis_cardium
| | | | +--+
| | +--+ +-- Lampsilis_cariosa
| | |
| | | +----- Venustaconcha_ellipsiformis
| | +--+
| | | +-- Lampsilis_powellii
| | +--+
| | +-- Lampsilis_siliquoidea
| |
+--+ +----- Chamberlainia_somsakpanhai
| +--------------------+
| | | +-- Sinohyriopsis_cumingii
| | +--+
| | +-- Sinohyriopsis_schlegelii
| |
| | +-- Hyriopsis_bialata
| | +--+
| +--------+ | +-- Rectidens_sumatrensis
| | | +-----------------+
| | | | | +-- Lens_contradens
| | | | +--+
| | | | +-- Physunio_superbus
| | | |
| | | | +----------- Potomida_littoralis
| | +--+ +--------+
| | | | | +-------- Pronodularia_japanensis
| | | | +--+
| | | | | +----- Lamprotula_leaii
| | | | +--+
| | | | | +-- Lamprotula_caveata
| | +--+ +--+
| | | +-- Lamprotula_cornuumlunae
| | |
| | | +-- Pseudodon_vondembuschianus
| | | +--------------+
| | | | +-- Pilsbryoconcha_exilis
| | +--+
| | | +-------------- Inversidens_rentianensis
| | | |
+--+ +--+ +-- Parvasolenaia_rivularis
| | +--------+
| | | +-- Ptychorhynchus_pfisteri
| +--+
| | +-------- Microcondylaea_bonellii
| +--+
| | +----- Sinosolenaia_carinata
| +--+
| | +-- Sinosolenaia_iridinea
| +--+
| +-- Sinosolenaia_oleivora
|
| +-------------------------------- Lamellidens_marginalis
| |
| | +-- Lepidodesma_aligera
| | +--------------------------+
| | | +-- Lepidodesma_languilati
| | |
| | | +-------- Aculamprotula_tortuosa
| | | |
| | | +--------------+ +-- Aculamprotula_coreana
| | | | | +--+
+--+ | | | | +-- Aculamprotula_tientsinensis
| | | +--+
| | | | +-- Aculamprotula_polysticta
| | | +--+
| | | +-- Aculamprotula_scripta
| | |
| | +--+ +-------------- Unio_tumidus
| | | | +-----+
| | | | | | +----------- Unio_crassus
| | | | | +--+
+--+ | | | | +-------- Unio_elongatulus
| | | | +--+
| | | | | +----- Unio_pictorum
| | | | +--+
| | | | | +-- Unio_delphinus
| | +--+ +--+
| | | +-- Unio_mancus
| | |
| | | +-- Nodularia_breviconcha
| | | +--------------+
| | | | +-- Nodularia_douglasiae
| | | |
| | | | +-- Pseudocuneopsis_capitata
| | | +--------------+
| | +--+ +-- Pseudocuneopsis_sichuanensis
| | |
| | | +-- Schistodesmus_lampreyanus
| | | +-----------+
+--+ | | +-- Schistodesmus_spinosus
| +--+
| | +----------- Tchangsinaia_piscicula
| +--+
| | +-------- Cuneopsis_rufescens
| +--+
| | +----- Cuneopsis_celtiformis
| +--+
| | +-- Cuneopsis_demangei
| +--+
| +-- Cuneopsis_heudei
|
| +-------- Acuticosta_chinensis
| +--------------+
| | | +----- Lanceolaria_gladiola
| | +--+
| | | +-- Lanceolaria_grayii
| | +--+
| | +-- Lanceolaria_lanceolata
| |
| | +-------------- Cristaria_plicata
| | |
+--+ +-----+ +-- Anemina_arcaeformis
| | | +--------+
| | | | +-- Anemina_euscaphys
| | +--+
| | | +-------- Beringiana_fukuharai
| | +--+
| | | +----- Sinanodonta_lucida
| | +--+
| | | +-- Sinanodonta_tumens
+--+ +--+
| +-- Sinanodonta_woodiana
|
| +-------- Pseudanodonta_complanata
| +--------+
| | | +----- Anodonta_anatina
| | +--+
| | | +-- Anodonta_cygnea
| | +--+
+--+ +-- Anodonta_exulcerata
|
| +-------------- Anodonta_nuttalliana
| |
| | +----- Pyganodon_grandis
+--+ +-----+
| | | +-- Utterbackia_imbecillis
| | +--+
+--+ +-- Utterbackia_peninsularis
|
| +-------- Lasmigona_complanata
+--+
| +----- Prolasmidonta_heterodon
+--+
| +-- Alasmidonta_varicosa
+--+
+-- Platynaias_compressa
Wu et al., 2024 [Publ 4350], fig. 8b, S2
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 2 |
2 |
0 |
2 |
90 |
89 |
2 |
-- |
+----------------------------------------------- Echyridella_menziesii
|
| +-- Anodontites_trapesialis **
| +-----------------------------------------+
| | +-- Mutela_dubia **
| |
| | +-- Margaritifera_dahurica
--+ | +--------------------------------------+
| | | +-- Margaritifera_margaritifera
| | |
| | | +-- Quadrula_quadrula
| | | +--------------------------+
| | | | +-- Uniomerus_tetralasmus
| | | |
+--+ | +--------+ +-- Elliptio_complanata
| | | | +-----------------------+
| | | | | +-- Pleurobema_oviforme
| | | +--+
| | | | +----------------------- Amblema_plicata
| | | | |
| | | +--+ +-------------------- Popenaias_popeii
| | | | |
| | | +--+ +----------------- Toxolasma_parvum
| | | | |
| | | | | +----- Potamilus_leptodon
+--+ | +--+ +--------+
| | | | | +-- Potamilus_alatus
| | | | +--+
| | +--+ +-- Potamilus_streckersoni
| | |
| | | +----------- Leaunio_lienosus
| | | |
| | +--+ +-------- Venustaconcha_ellipsiformis
| | | |
| | | | +-- Lampsilis_powellii
| | +--+-----+
| | | +-- Lampsilis_siliquoidea
| | |
| | | +----- Lampsilis_ornata
| | +--+
| | | +-- Lampsilis_cardium
| | +--+
| | +-- Lampsilis_cariosa
| |
+--+ +----- Chamberlainia_somsakpanhai
| +--------------------+
| | | +-- Sinohyriopsis_cumingii
| | +--+
| | +-- Sinohyriopsis_schlegelii
| |
| | +-- Hyriopsis_bialata
| | +--+
| +--------+ | +-- Rectidens_sumatrensis
| | | +-----------------+
| | | | | +-- Lens_contradens
| | | | +--+
| | | | +-- Physunio_superbus
| | | |
| | | | +----------- Potomida_littoralis
| | +--+ +--------+
| | | | | +-------- Pronodularia_japanensis
| | | | +--+
| | | | | +----- Lamprotula_leaii
| | | | +--+
| | | | | +-- Lamprotula_caveata
| | +--+ +--+
| | | +-- Lamprotula_cornuumlunae
| | |
| | | +-- Pseudodon_vondembuschianus
| | | +--------------+
| | | | +-- Pilsbryoconcha_exilis
| | +--+
| | | +-------------- Inversidens_rentianensis
| | | |
+--+ +--+ +-- Parvasolenaia_rivularis
| | +--------+
| | | +-- Ptychorhynchus_pfisteri
| +--+
| | +-------- Microcondylaea_bonellii
| +--+
| | +----- Sinosolenaia_carinata
| +--+
| | +-- Sinosolenaia_iridinea
| +--+
| +-- Sinosolenaia_oleivora
|
| +-------------------------------- Lamellidens_marginalis
| |
| | +-- Lepidodesma_aligera
| | +--------------------------+
| | | +-- Lepidodesma_languilati
| | |
| | | +-------- Aculamprotula_tortuosa
| | | |
| | | +--------------+ +-- Aculamprotula_coreana
| | | | | +--+
+--+ | | | | +-- Aculamprotula_tientsinensis
| | | +--+
| | | | +-- Aculamprotula_polysticta
| | | +--+
| | | +-- Aculamprotula_scripta
| | |
| | +--+ +-------------- Unio_tumidus
| | | | +-----+
| | | | | | +----------- Unio_crassus
| | | | | +--+
+--+ | | | | +-------- Unio_elongatulus
| | | | +--+
| | | | | +----- Unio_pictorum
| | | | +--+
| | | | | +-- Unio_delphinus
| | +--+ +--+
| | | +-- Unio_mancus
| | |
| | | +-- Nodularia_breviconcha
| | | +--------------+
| | | | +-- Nodularia_douglasiae
| | | |
| | | | +-- Pseudocuneopsis_capitata
| | | +--------------+
| | +--+ +-- Pseudocuneopsis_sichuanensis
| | |
| | | +-- Schistodesmus_lampreyanus
+--+ | +-----------+
| | | +-- Schistodesmus_spinosus
| +--+
| | +----------- Tchangsinaia_piscicula
| +--+
| | +-------- Cuneopsis_rufescens
| +--+
| | +----- Cuneopsis_celtiformis
| +--+
| | +-- Cuneopsis_demangei
| +--+
| +-- Cuneopsis_heudei
|
| +-------- Acuticosta_chinensis
| +--------------+
| | | +----- Lanceolaria_gladiola
| | +--+
| | | +-- Lanceolaria_grayii
| | +--+
| | +-- Lanceolaria_lanceolata
| |
| | +-------------- Cristaria_plicata
+--+ |
| +-----+ +-- Anemina_arcaeformis
| | | +--------+
| | | | +-- Anemina_euscaphys
| | +--+
| | | +-------- Beringiana_fukuharai
| | +--+
| | | +----- Sinanodonta_lucida
| | +--+
+--+ | +-- Sinanodonta_tumens
| +--+
| +-- Sinanodonta_woodiana
|
| +----------------- Anodonta_nuttalliana
| |
| | +-------- Pseudanodonta_complanata
| | +-----+
+--+ | | +----- Anodonta_anatina
| | +--+
| | | +-- Anodonta_cygnea
| | +--+
+--+ +-- Anodonta_exulcerata
|
| +----- Pyganodon_grandis
| +-----+
| | | +-- Utterbackia_imbecillis
| | +--+
+--+ +-- Utterbackia_peninsularis
|
| +-------- Lasmigona_complanata
+--+
| +----- Prolasmidonta_heterodon
+--+
| +-- Alasmidonta_varicosa
+--+
+-- Platynaias_compressa
Graf & Ó Foighil, 2000 [Publ 2648], fig. 2A
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 1 |
1 |
0 |
1 |
29 |
28 |
2 |
-- |
+-------------------------------- Mutela_rostrata **
|
| +----------------------------- Hyridella_depressa
--+ |
| | +-- Margaritifera_margaritifera
| | +-----------------------+
+--+ | +-- Cumberlandia_monodonta
| |
| | +-- Unio_pictorum
| | +--------------------+
+--+ | +-- Unio_caffer
| |
| | +----------- Anodonta_cygnea
| | |
| | +--------+ +-- Alasmidonta_marginata
+--+ | | +-----+
| | | | +-- Lasmigona_costata
| | +--+
| | | +----- Strophitus_undulatus
| | +--+
| | | +-- Pyganodon_grandis
| | +--+
+--+ +-- Pyganodon_fragilis
|
| +-------- Quadrula_quadrula
| +--------+
| | | +----- Eurynia_dilatata
| | +--+
| | | +-- Pleurobema_sintoxia
| | +--+
+--+ +-- Fusconaia_flava
|
| +-------------- Amblema_plicata
| |
| | +----------- Ligumia_recta
| | |
+--+ +----------- Truncilla_truncata
| |
| | +-- Ptychobranchus_fasciolaris
| +--------+
+--+ +-- Sagittunio_nasutus
|
| +----- Lampsilis_siliquoidea
| +--+
| | | +-- Epioblasma_triquetra
| | +--+
| | +-- Epioblasma_brevidens
+--+
| +----- Ortmanniana_ligamentina
| |
| +----- Cambarunio_iris
+--+
+----- Leaunio_vanuxemensis
|
| +-- Lampsilis_cardium
+--+
+-- Lampsilis_fasciola
Graf & Ó Foighil, 2000 [Publ 2648], fig. 2B
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 1 |
1 |
0 |
1 |
29 |
28 |
2 |
-- |
+----------------------------------------- Mutela_rostrata **
|
| +-------------------------------------- Hyridella_depressa
--+ |
| | +-- Margaritifera_margaritifera
| | +--------------------------------+
+--+ | +-- Cumberlandia_monodonta
| |
| | +-- Unio_pictorum
| | +-----------------------------+
+--+ | +-- Unio_caffer
| |
| | +----------- Anodonta_cygnea
| | |
| | +-----------------+ +-- Alasmidonta_marginata
+--+ | | +-----+
| | | | +-- Lasmigona_costata
| | +--+
| | | +----- Strophitus_undulatus
| | +--+
| | | +-- Pyganodon_grandis
+--+ +--+
| +-- Pyganodon_fragilis
|
| +-------- Quadrula_quadrula
| +-----------------+
| | | +----- Eurynia_dilatata
| | +--+
| | | +-- Pleurobema_sintoxia
+--+ +--+
| +-- Fusconaia_flava
|
| +----------------------- Amblema_plicata
| |
+--+ +-------------------- Truncilla_truncata
| |
+--+ +----------------- Ligumia_recta
| |
| | +-- Ptychobranchus_fasciolaris
+--+ +-----------+
| | +-- Sagittunio_nasutus
| |
+--+ +----------- Lampsilis_siliquoidea
| |
| | +----- Ortmanniana_ligamentina
+--+ +--+
| | | +-- Lampsilis_cardium
| | +--+
| | +-- Lampsilis_fasciola
+--+
| +-- Epioblasma_brevidens
| +--+
| | +-- Epioblasma_triquetra
+--+
| +-- Cambarunio_iris
+--+
+-- Leaunio_vanuxemensis
Hoeh et al., 1998 [Publ 2695], fig. 3
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 1 |
1 |
0 |
1 |
4 |
3 |
1 |
-- |
+-------- Mutela_rostrata **
--+
| +----- Anodonta_cygnea
+--+
| +-- Margaritifera_margaritifera
+--+
+-- Amblema_plicata
Hoeh et al., 1998 [Publ 2695], fig. 4
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 1 |
1 |
0 |
1 |
4 |
3 |
1 |
-- |
+-------- Mutela_rostrata **
--+
| +----- Amblema_plicata
+--+
| +-- Margaritifera_margaritifera
+--+
+-- Anodonta_cygnea
Bieler et al., 2014 [Publ 3622], fig. 26, 27, 29, 30
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 1 |
1 |
0 |
1 |
4 |
3 |
2 |
-- |
+----- Unio_pictorum
--+
| +-- Aspatharia_pfeifferiana **
+--+
+-- Margaritifera_margaritifera
Bieler et al., 2014 [Publ 3622], fig. 28
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 1 |
1 |
0 |
1 |
4 |
3 |
2 |
-- |
+-- Aspatharia_pfeifferiana **
+--+
| +-- Margaritifera_margaritifera
--+
| +-- Unio_pictorum
+--+
+-- Velesunio_ambiguus
González & Giribet, 2015 [Publ 3632], fig. 2
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 1 |
1 |
0 |
1 |
5 |
3 |
2 |
-- |
+-- Aspatharia_pfeifferiana **
+--+
| +-- Margaritifera_margaritifera
--+
| +-- Unio_pictorum
+--+
+-- Velesunio_ambiguus
González & Giribet, 2015 [Publ 3632], fig. 3
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 1 |
1 |
0 |
1 |
5 |
3 |
2 |
-- |
+----- Unio_pictorum
|
--+----- Velesunio_ambiguus
|
| +-- Aspatharia_pfeifferiana **
+--+
+-- Margaritifera_margaritifera
González & Giribet, 2015 [Publ 3632], fig. 4
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 1 |
1 |
0 |
1 |
5 |
3 |
2 |
-- |
+-------- Aspatharia_pfeifferiana **
--+
| +----- Velesunio_ambiguus
+--+
| +-- Unio_pictorum
+--+
+-- Margaritifera_margaritifera
González & Giribet, 2015 [Publ 3632], fig. 6
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 1 |
1 |
0 |
1 |
5 |
3 |
2 |
-- |
+----- Unio_pictorum
--+
| +-- Aspatharia_pfeifferiana **
+--+
+-- Margaritifera_margaritifera
Wu et al., 2019 [Publ 3864], fig. 3
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 1 |
1 |
0 |
1 |
46 |
46 |
2 |
-- |
+-- Echyridella_menziesii
+--------+
| +-- Mutela_dubia **
--+
| +-------- Gibbosula_rochechouartii
+--+
| +----- Cumberlandia_monodonta
+--+
| +-- Margaritifera_dahurica
+--+
+-- Margaritifera_falcata
| 
