MUSSELp
 
MUSSELpdb
 
 
Taxonomy
 
 
Geography
 
 
 

Page last updated
Tue 09 Jul 2024

The Freshwater Mussels (Unionoida) of the World (and other less consequential bivalves)

FM(U)otW(aolcb) is the web version of the MUSSEL Project Database. Follow the links to browse the data or use the custom Google search field. Either way, you win!

Cladomics pages are experimental — even more so than the rest of this site!

The purpose of the cladomics pages for each genus and family-group level taxon is to provide statistics and references indicating the extent to which published phylogenetic work supports the classification used here.

The Cladomics Summary provides an overview of the number of studies that have included terminal species representing this taxon as well as an indication of how frequently it has been recovered as monophyletic.

Following the summary, the available published clades representing the taxon are listed.

For more information, visit the Cladomics FARQ (frequently asked rhetorical questions) page. Abbreviations for the various statistics are explained on the FARQ!

Search the database pages:

Bivalvia | Unionoida

superfamily ETHERIOIDEA Deshayes, 1832: cladomics
     3 families

Cladomics Summary

number of species = 112
number of subtaxa (rank: family) = 3

analyzed in 69 cladograms in 37 publications
maximum number of terminals in an analysis = 10, mean = 4.3
maximum number of subtaxa (family) in an analysis = 3, mean = 2.32

Basic Monophyly

number of cladograms that test basic monophyly (i.e., more than 1 included species) = 58
maximum number of included terminals = 10, mean = 4.93
maximum number of subtaxa (family) = 3, mean = 2.57
proportion of cladograms that support basic monophyly = 0.93

Rigorous Monophyly

number of cladograms that test rigorous monophyly = 58
(Rigorous monophyly refers to cladograms with terminal species representing more than one subtaxon (family) taxon and at least one rigorous outgroup (superfamily).
maximum number of included terminals = 10, mean = 4.93
maximum number of subtaxa (family) = 3, mean = 2.57
proportion of cladograms that support rigorous monophyly = 0.93


Pairwise Cladogram Clustering

The graphs below depict in two dimensions all pair-wise ingroup distances within the cladome. See the Cladmoics FARQ for more information.


PCoA Ordination


NJ tree


Lopes-Lima et al., 2017 [Publ 3745], fig. 1-2

iT tT bi riS oT roT roS iD
6 6 4 3 75 74 3 0.0

*** Top Monophyly Cladogram *** *** Top Sister-Group Cladogram ***

                                   +-- Cumberlandia_monodonta     
  +--------------------------------+                              
  |                                +-- Margaritifera_margaritifera
  |                                                               
  |                       +----------- Mutela_rostrata **            
  |                       |                                       
  |  +--------------------+        +-- Aspatharia_pfeifferiana **    
  |  |                    |  +-----+                              
  |  |                    |  |     +-- Chambardia_wahlbergi **       
  |  |                    +--+                                    
  |  |                       |  +----- Etheria_elliptica **          
  |  |                       +--+                                 
  |  |                          |  +-- Tamsiella_schroeteriana **    
--+  |                          +--+                              
  |  |                             +-- Mycetopoda_siliquosa **       
  |  |                                                            
  |  |                       +-------- Trapezidens_dolichorhynchus
  |  |  +--------------------+                                    
  |  |  |                    |  +----- Leoparreysia_tavoyensis    
  |  |  |                    +--+                                 
  |  |  |                       |  +-- Coelatura_aegyptiaca       
  |  |  |                       +--+                              
  |  |  |                          +-- Indochinella_pugio         
  |  |  |                                                         
  |  |  |                       +----- Unio_gibbus                
  +--+  |                       |                                 
     |  |           +-----------+----- Unio_tumidus               
     |  |           |           |                                 
     |  |           |           |  +-- Unio_crassus               
     |  |           |           +--+                              
     |  |        +--+              +-- Unio_pictorum              
     |  |        |  |                                             
     |  |        |  |  +-------------- Aculamprotula_tortuosa     
     |  |        |  |  |                                          
     |  |        |  +--+           +-- Nodularia_nuxpersicae      
     |  |        |     |  +--------+                              
     |  |        |     |  |        +-- Nodularia_douglasiae       
     |  |        |     +--+                                       
     |  |        |        |  +-------- Schistodesmus_lampreyanus  
     |  |        |        +--+                                    
     |  |        |           |  +----- Tchangsinaia_piscicula     
     |  |        |           +--+                                 
     |  |  +-----+              |  +-- Cuneopsis_heudei           
     |  |  |     |              +--+                              
     |  |  |     |                 +-- Cuneopsis_rufescens        
     +--+  |     |                                                
        |  |     |           +-------- Lanceolaria_gladiola       
        |  |     |  +--------+                                    
        |  |     |  |        |  +----- Lanceolaria_grayii[2]      
        |  |     |  |        +--+                                 
        |  |     |  |           |  +-- Lanceolaria_lanceolata     
        |  |     |  |           +--+                              
        |  |     |  |              +-- Lanceolaria_grayii[1]      
        |  |     |  |                                             
        |  |     +--+     +----------- Cristaria_plicata          
        |  |        |  +--+                                       
        |  |        |  |  |  +-------- Anemina_arcaeformis        
        |  |        |  |  +--+                                    
        |  |        |  |     |  +----- Pletholophus_tenuis        
        |  |        |  |     +--+                                 
        |  |        |  |        |  +-- Sinanodonta_jourdyi        
        |  |        |  |        +--+                              
        |  |        +--+           +-- Sinanodonta_lucida         
        |  |           |                                          
        |  |           |        +----- Pseudanodonta_complanata   
        |  |           |        |                                 
        |  |           |  +-----+  +-- Anodonta_anatina           
        |  |           |  |     |  |                              
        |  |           |  |     +--+-- Anodonta_cygnea            
        |  |           |  |        |                              
        |  |           +--+        +-- Anodonta_nuttalliana       
        |  |              |                                       
        |  |              |        +-- Pyganodon_grandis          
        +--+              |  +-----+                              
           |              |  |     +-- Simpsonaias_ambigua        
           |              +--+                                    
           |                 |  +----- Strophitus_undulatus       
           |                 +--+                                 
           |                    |  +-- Alasmidonta_marginata      
           |                    +--+                              
           |                       +-- Lasmigona_costata          
           |                                                      
           |                       +-- Lens_contradens            
           |                    +--+                              
           |                    |  +-- Lens_comptus               
           |                 +--+                                 
           |                 |  |  +-- Physunio_modelli           
           |                 |  +--+                              
           |                 |     +-- unionoid_species           
           |                 |                                    
           |  +--------------+  +----- Hyriopsis_desowitzi        
           |  |              |  |                                 
           |  |              |  +----- Rectidens_sumatrensis      
           |  |              |  |                                 
           |  |              |  |  +-- Hyriopsis_khoratensis      
           |  |              +--+--+                              
           |  |                 |  +-- Hyriopsis_myersiana        
           |  |                 |                                 
           |  |                 |  +-- Ensidens_ingallsianus      
           |  |                 |  |                              
           |  |                 +--+-- Ensidens_telus             
           |  |                    |                              
           |  |                    +-- Ensidens_jaculus           
           |  |                                                   
           |  |              +-------- Amblema_plicata            
           +--+           +--+                                    
              |           |  |  +----- Cambarunio_iris            
              |           |  +--+                                 
              |           |     |  +-- Lampsilis_cardium          
              |           |     +--+                              
              |  +--------+        +-- Ortmanniana_ligamentina    
              |  |        |                                       
              |  |        |        +-- Quadrula_quadrula          
              |  |        |  +-----+                              
              |  |        |  |     +-- Quadrula_verrucosa         
              |  |        +--+                                    
              |  |           |  +----- Elliptio_complanata        
              |  |           +--+                                 
              |  |              |  +-- Pleurobema_sintoxia        
              |  |              +--+                              
              |  |                 +-- Eurynia_dilatata           
              +--+                                                
                 |                 +-- Chamberlainia_somsakpanhai 
                 |           +-----+                              
                 |           |     +-- Sinohyriopsis_cumingii     
                 |           |                                    
                 |  +--------+     +-- Lamprotula_caveata         
                 |  |        |  +--+                              
                 |  |        |  |  +-- Lamprotula_leaii           
                 |  |        +--+                                 
                 |  |           |  +-- Potomida_littoralis        
                 |  |           +--+                              
                 |  |              +-- Pronodularia_japanensis    
                 +--+                                             
                    |              +-- Sinosolenaia_carinata      
                    |        +-----+                              
                    |        |     +-- Sinosolenaia_oleivora      
                    |  +-----+                                    
                    |  |     |  +----- Gonidea_angulata           
                    |  |     +--+                                 
                    |  |        |  +-- Microcondylaea_bonellii    
                    +--+        +--+                              
                       |           +-- Leguminaia_wheatleyi       
                       |                                          
                       |  +----------- Parvasolenaia_rivularis    
                       |  |                                       
                       +--+        +-- Pilsbryoconcha_exilis      
                          |  +-----+                              
                          |  |     +-- Pilsbryoconcha_acuta       
                          +--+                                    
                             |  +----- Pseudodon_vagulus          
                             +--+                                 
                                |  +-- Bineurus_exilis            
                                +--+                              
                                   +-- Nyeinchanconcha_nyeinchani 

Graf & Cummings, 2006 [Publ 1968], fig. 2

iT tT bi riS oT roT roS iD
10 10 8 3 37 36 2 12.0
                                   +----- Velesunio_ambiguus         
                          +--------+                                 
                          |        |  +-- Velesunio_angasi           
                          |        +--+                              
                       +--+           +-- Lortiella_rugata           
                       |  |                                          
                       |  |  +----------- Echyridella_menziesii      
                       |  |  |                                       
                       |  +--+        +-- Hyridella_depressa         
                       |     |  +-----+                              
                       |     |  |     +-- Hyridella_australis        
                       |     +--+                                    
                       |        |  +----- Castalia                   
                       |        +--+                                 
  +--------------------+           |  +-- Diplodon_chilensis         
  |                    |           +--+                              
  |                    |              +-- Diplodon_gratus            
  |                    |                                             
  |                    |           +----- Etheria_elliptica **          
  |                    |     +-----+                                 
  |                    |     |     |  +-- Mulleria_rivoli **            
  |                    |     |     +--+                              
  |                    |     |        +-- Pseudomulleria_dalyi **       
  |                    |     |                                       
  |                    +-----+     +----- Monocondylaea_minuana **      
  |                          |  +--+                                 
  |                          |  |  |  +-- Anodontites_trigona **        
  |                          |  |  +--+                              
  |                          |  |     +-- Anodontites_guanarensis **    
--+                          +--+                                    
  |                             |     +-- Chambardia_wahlbergi **       
  |                             |  +--+                              
  |                             |  |  +-- Aspatharia_rugifera **        
  |                             +--+                                 
  |                                |  +-- Mutela_dubia **               
  |                                +--+                              
  |                                   +-- Mutela_rostrata **            
  |                                                                  
  |                                   +-- Margaritifera_margaritifera
  |  +--------------------------------+                              
  |  |                                +-- Cumberlandia_monodonta     
  |  |                                                               
  |  |                                +-- Pilsbryoconcha_exilis      
  |  |  +-----------------------------+                              
  +--+  |                             +-- Pseudodon_vondembuschianus 
     |  |                                                            
     |  |  +----------------------------- Coelatura_aegyptiaca       
     |  |  |                                                         
     |  |  +----------------------------- Lens_contradens            
     +--+  |                                                         
        |  |                          +-- Unio_pictorum              
        |  |                 +--------+                              
        |  |                 |        +-- Unio_caffer                
        |  +-----------------+                                       
        |  |                 |  +-------- Lasmigona_costata          
        +--+                 +--+                                    
           |                    |  +----- Alasmidonta_marginata      
           |                    +--+                                 
           |                       |  +-- Pyganodon_grandis          
           |                       +--+                              
           |                          +-- Strophitus_undulatus       
           |                                                         
           |  +-------------------------- Gonidea_angulata           
           |  |                                                      
           |  |                       +-- Quadrula_verrucosa         
           +--+  +--------------------+                              
              |  |                    +-- Quadrula_quadrula          
              |  |                                                   
              +--+  +-------------------- Amblema_plicata            
                 |  |                                                
                 |  |              +----- Eurynia_dilatata           
                 +--+  +-----------+                                 
                    |  |           |  +-- Fusconaia_flava            
                    |  |           +--+                              
                    |  |              +-- Pleurobema_sintoxia        
                    +--+                                             
                       |  +-------------- Obliquaria_reflexa         
                       |  |                                          
                       |  +-------------- Truncilla_truncata         
                       +--+                                          
                          +-------------- Ptychobranchus_fasciolaris 
                          |                                          
                          |  +----------- Ligumia_recta              
                          +--+                                       
                             |  +-------- Cambarunio_iris            
                             +--+                                    
                                |  +----- Epioblasma_triquetra       
                                +--+                                 
                                   |  +-- Lampsilis_cardium          
                                   +--+                              
                                      +-- Ortmanniana_ligamentina    

Hoeh et al., 2009 [Publ 2716], fig. 3

iT tT bi riS oT roT roS iD
10 37 34 3 37 36 2 38.0
                                         +----- Velesunio_ambiguus         
                                +--------+                                 
                                |        |  +-- Velesunio_angasi           
                                |        +--+                              
  +-----------------------------+           +-- Lortiella_rugata           
  |                             |                                          
  |                             |  +----------- Echyridella_menziesii      
  |                             |  |                                       
  |                             +--+        +-- Hyridella_depressa         
  |                                |  +-----+                              
  |                                |  |     +-- Hyridella_australis        
  |                                +--+                                    
  |                                   |  +----- Castalia                   
  |                                   +--+                                 
  |                                      |  +-- Diplodon_chilensis         
  |                                      +--+                              
  |                                         +-- Diplodon_gratus            
--+                                                                        
  |                                      +----- Chambardia_wahlbergi **       
  |                                      |                                 
  |                             +--------+----- Aspatharia_rugifera **        
  |                             |        |                                 
  |                             |        |  +-- Mutela_rostrata **            
  |  +--------------------------+        +--+                              
  |  |                          |           +-- Mutela_dubia **               
  |  |                          |                                          
  |  |                          |  +----------- Etheria_elliptica **          
  |  |                          +--+                                       
  |  |                             |  +-------- Mulleria_rivoli **            
  |  |                             +--+                                    
  |  |                                |  +----- Monocondylaea_minuana **      
  +--+                                +--+                                 
     |                                   |  +-- Anodontites_trigona **        
     |                                   +--+                              
     |                                      +-- Anodontites_guanarensis **    
     |                                                                     
     |                                      +-- Margaritifera_margaritifera
     |  +-----------------------------------+                              
     |  |                                   +-- Cumberlandia_monodonta     
     |  |                                                                  
     +--+  +----------------------------------- Coelatura_aegyptiaca       
        |  |                                                               
        |  |  +-------------------------------- Pseudomulleria_dalyi **       
        |  |  |                                                            
        +--+  |                             +-- Unio_pictorum              
           |  |                    +--------+                              
           |  |                    |        +-- Unio_caffer                
           |  |  +-----------------+                                       
           +--+  |                 |  +-------- Lasmigona_costata          
              |  |                 +--+                                    
              |  |                    |  +----- Alasmidonta_marginata      
              |  |                    +--+                                 
              |  |                       |  +-- Strophitus_undulatus       
              |  |                       +--+                              
              +--+                          +-- Pyganodon_grandis          
                 |                                                         
                 |  +-------------------------- Lens_contradens            
                 |  |                                                      
                 |  |                    +----- Gonidea_angulata           
                 |  |  +-----------------+                                 
                 |  |  |                 |  +-- Pilsbryoconcha_exilis      
                 +--+  |                 +--+                              
                    |  |                    +-- Pseudodon_vondembuschianus 
                    |  |                                                   
                    |  |           +----------- Amblema_plicata            
                    |  |           |                                       
                    +--+           |     +----- Obliquaria_reflexa         
                       |  +--------+  +--+                                 
                       |  |        |  |  |  +-- Quadrula_verrucosa         
                       |  |        |  |  +--+                              
                       |  |        +--+     +-- Quadrula_quadrula          
                       |  |           |                                    
                       |  |           |  +----- Eurynia_dilatata           
                       +--+           +--+                                 
                          |              |  +-- Pleurobema_sintoxia        
                          |              +--+                              
                          |                 +-- Fusconaia_flava            
                          |                                                
                          |  +----------------- Ptychobranchus_fasciolaris 
                          +--+                                             
                             |  +-------------- Truncilla_truncata         
                             +--+                                          
                                |  +----------- Ortmanniana_ligamentina    
                                +--+                                       
                                   |  +-------- Lampsilis_cardium          
                                   +--+                                    
                                      |  +----- Cambarunio_iris            
                                      +--+                                 
                                         |  +-- Ligumia_recta              
                                         +--+                              
                                            +-- Epioblasma_triquetra       

Hoeh et al., 2009 [Publ 2716], fig. 4

iT tT bi riS oT roT roS iD
10 36 33 3 36 35 2 38.0
                                            +----- Velesunio_ambiguus         
                                   +--------+                                 
                                   |        |  +-- Velesunio_angasi           
                                   |        +--+                              
  +--------------------------------+           +-- Lortiella_rugata           
  |                                |                                          
  |                                |  +----------- Echyridella_menziesii      
  |                                |  |                                       
  |                                +--+        +-- Hyridella_depressa         
  |                                   |  +-----+                              
  |                                   |  |     +-- Hyridella_australis        
  |                                   +--+                                    
  |                                      |  +----- Castalia                   
  |                                      +--+                                 
  |                                         |  +-- Diplodon_chilensis         
  |                                         +--+                              
  |                                            +-- Diplodon_gratus            
--+                                                                           
  |                                            +-- Etheria_elliptica **          
  |                                   +--------+                              
  |                                   |        +-- Mulleria_rivoli **            
  |                                   |                                       
  |                                   |     +----- Monocondylaea_minuana **      
  |  +--------------------------------+  +--+                                 
  |  |                                |  |  |  +-- Anodontites_trigona **        
  |  |                                |  |  +--+                              
  |  |                                |  |     +-- Anodontites_guanarensis **    
  |  |                                +--+                                    
  |  |                                   |     +-- Mutela_rostrata **            
  |  |                                   |  +--+                              
  +--+                                   |  |  +-- Mutela_dubia **               
     |                                   +--+                                 
     |                                      |  +-- Chambardia_wahlbergi **       
     |                                      +--+                              
     |                                         +-- Aspatharia_rugifera **        
     |                                                                        
     |  +----------------------------------------- Coelatura_aegyptiaca       
     |  |                                                                     
     +--+  +-------------------------------------- Pseudomulleria_dalyi **       
        |  |                                                                  
        |  |                                   +-- Margaritifera_margaritifera
        +--+  +--------------------------------+                              
           |  |                                +-- Cumberlandia_monodonta     
           |  |                                                               
           |  |                                +-- Pilsbryoconcha_exilis      
           +--+  +-----------------------------+                              
              |  |                             +-- Pseudodon_vondembuschianus 
              |  |                                                            
              |  |  +----------------------------- Lens_contradens            
              +--+  |                                                         
                 |  |                          +-- Unio_pictorum              
                 |  |                 +--------+                              
                 |  |                 |        +-- Unio_caffer                
                 |  |  +--------------+                                       
                 +--+  |              |  +-------- Lasmigona_costata          
                    |  |              +--+                                    
                    |  |                 |  +----- Alasmidonta_marginata      
                    |  |                 +--+                                 
                    |  |                    |  +-- Strophitus_undulatus       
                    |  |                    +--+                              
                    +--+                       +-- Pyganodon_grandis          
                       |                                                      
                       |  +----------------------- Gonidea_angulata           
                       |  |                                                   
                       |  |                    +-- Eurynia_dilatata           
                       |  |                    |                              
                       |  |              +-----+-- Pleurobema_sintoxia        
                       +--+              |     |                              
                          |  +-----------+     +-- Fusconaia_flava            
                          |  |           |                                    
                          |  |           |  +----- Obliquaria_reflexa         
                          |  |           +--+                                 
                          +--+              |  +-- Quadrula_verrucosa         
                             |              +--+                              
                             |                 +-- Quadrula_quadrula          
                             |                                                
                             |  +----------------- Ptychobranchus_fasciolaris 
                             +--+                                             
                                |  +-------------- Truncilla_truncata         
                                +--+                                          
                                   |  +----------- Ortmanniana_ligamentina    
                                   +--+                                       
                                      |  +-------- Lampsilis_cardium          
                                      +--+                                    
                                         |  +----- Cambarunio_iris            
                                         +--+                                 
                                            |  +-- Ligumia_recta              
                                            +--+                              
                                               +-- Epioblasma_triquetra       

Hoeh et al., 2009 [Publ 2716], fig. 5

iT tT bi riS oT roT roS iD
10 10 8 3 36 35 2 12.0
                       +-- Margaritifera_margaritifera
  +--------------------+                              
  |                    +-- Cumberlandia_monodonta     
  |                                                   
  |                 +----- Velesunio_ambiguus         
  |        +--------+                                 
  |        |        |  +-- Velesunio_angasi           
  |        |        +--+                              
--+  +-----+           +-- Lortiella_rugata           
  |  |     |                                          
  |  |     |  +----------- Echyridella_menziesii      
  |  |     |  |                                       
  |  |     +--+        +-- Hyridella_depressa         
  |  |        |  +-----+                              
  |  |        |  |     +-- Hyridella_australis        
  |  |        +--+                                    
  +--+           |  +----- Castalia                   
     |           +--+                                 
     |              |  +-- Diplodon_chilensis         
     |              +--+                              
     |                 +-- Diplodon_gratus            
     |                                                
     |  +----------------- Pseudomulleria_dalyi **       
     |  |                                             
     +--+  +-------------- Mulleria_rivoli **            
        |  |                                          
        +--+  +----------- Etheria_elliptica **          
           |  |                                       
           |  |     +----- Monocondylaea_minuana **      
           +--+  +--+                                 
              |  |  |  +-- Anodontites_trigona **        
              |  |  +--+                              
              |  |     +-- Anodontites_guanarensis **    
              +--+                                    
                 |     +-- Mutela_rostrata **            
                 |  +--+                              
                 |  |  +-- Mutela_dubia **               
                 +--+                                 
                    |  +-- Chambardia_wahlbergi **       
                    +--+                              
                       +-- Aspatharia_rugifera **        

Huang et al., 2019 [Publ 3843], fig. 2

iT tT bi riS oT roT roS iD
10 10 8 3 194 192 2 6.0
                                               +----- Alathyria_jacksoni         
                                            +--+                                 
                                            |  |  +-- Alathyria_profuga          
                                            |  +--+                              
                                   +--------+     +-- Velesunio_ambiguus         
                                   |        |                                    
                                   |        |  +----- Alathyria_pertexta         
                                   |        +--+                                 
                                   |           |  +-- Lortiella_froggatti        
  +--------------------------------+           +--+                              
  |                                |              +-- Microdontia_anodontaeformis
  |                                |                                             
  |                                |  +-------------- Westralunio_carteri        
  |                                |  |                                          
  |                                +--+  +----------- Echyridella_menziesii      
  |                                   |  |                                       
  |                                   |  |     +----- Diplodon_suavidicus        
  |                                   +--+  +--+                                 
  |                                      |  |  |  +-- Castalia_ambigua           
  |                                      |  |  +--+                              
  |                                      +--+     +-- Triplodon_corrugatus       
  |                                         |                                    
  |                                         |  +----- Hyridella_australis        
  |                                         +--+                                 
  |                                            |  +-- Hyridella_depressa         
  |                                            +--+                              
  |                                               +-- Hyridella_drapeta          
  |                                                                              
  |                                         +-------- Anodontites_trapesialis **    
  |                                      +--+                                    
  |                                      |  |  +----- Mutela_rostrata **            
  |                                      |  +--+                                 
  |                                      |     |  +-- Mutela_dubia **               
  |                                      |     +--+                              
  |  +-----------------------------------+        +-- Mutela_hargeri **             
  |  |                                   |                                       
  |  |                                   |     +----- Mycetopoda_siliquosa **       
  |  |                                   |  +--+                                 
--+  |                                   |  |  |  +-- Tamsiella_schroeteriana **    
  |  |                                   |  |  +--+                              
  |  |                                   +--+     +-- Lamproscapha_ensiformis **    
  |  |                                      |                                    
  |  |                                      |  +----- Etheria_elliptica **          
  |  |                                      +--+                                 
  |  |                                         |  +-- Aspatharia_pfeifferiana **    
  |  |                                         +--+                              
  |  |                                            +-- Chambardia_wahlbergi **       
  |  |                                                                           
  |  |                             +----------------- Gibbosula_rochechouartii   
  |  |                             |                                             
  |  |                             |  +-------------- Gibbosula_laosensis        
  |  |  +--------------------------+  |                                          
  |  |  |                          |  |  +----------- Cumberlandia_monodonta     
  |  |  |                          |  |  |                                       
  |  |  |                          +--+  |     +----- Pseudunio_marocanus        
  |  |  |                             |  +-----+                                 
  |  |  |                             |  |     |  +-- Pseudunio_auricularius     
  |  |  |                             |  |     +--+                              
  |  |  |                             +--+        +-- Pseudunio_homsensis        
  |  |  |                                |                                       
  |  |  |                                |        +-- Margaritifera_dahurica     
  |  |  |                                |  +-----+                              
  |  |  |                                |  |     +-- Margaritifera_margaritifera
  |  |  |                                |  |                                    
  |  |  |                                +--+  +----- Margaritifera_laevis       
  |  |  |                                   |  |                                 
  |  |  |                                   |  |  +-- Margaritifera_falcata      
  |  |  |                                   +--+--+                              
  +--+  |                                      |  +-- Margaritifera_middendorffi 
     |  |                                      |                                 
     |  |                                      |  +-- Margaritifera_hembeli      
     |  |                                      +--+                              
     |  |                                         +-- Margaritifera_marrianae    
     |  |                                                                        
     |  |                                      +----- Trapezidens_exolescens     
     |  |                    +-----------------+                                 
     |  |                    |                 |  +-- Trapezidens_dolichorhynchus
     |  |                    |                 +--+                              
     |  |                 +--+                    +-- Trapezidens_angustior      
     |  |                 |  |                                                   
     |  |                 |  |  +-------------------- Lamellidens_corrianus      
     |  |                 |  |  |                                                
     |  |                 |  +--+  +----------------- Lamellidens_savadiensis    
     |  |                 |     |  |                                             
     |  |                 |     +--+              +-- Lamellidens_lamellatus     
     |  |                 |        |  +-----------+                              
     |  |                 |        |  |           +-- Lamellidens_jenkinsianus   
     |  |                 |        +--+                                          
     |  |                 |           |  +----------- Lamellidens_generosus      
     |  |                 |           +--+                                       
     |  |                 |              |  +-------- Lamellidens_mainwaringi[2] 
     |  |                 |              +--+                                    
     |  |  +--------------+                 |  +----- Lamellidens_mainwaringi[1] 
     |  |  |              |                 +--+                                 
     |  |  |              |                    |  +-- Lamellidens_ferrugineus    
     |  |  |              |                    +--+                              
     |  |  |              |                       +-- Lamellidens_marginalis     
     |  |  |              |                                                      
     |  |  |              |                    +----- Parreysia_keralaensis      
     |  |  |              |  +-----------------+                                 
     |  |  |              |  |                 |  +-- Parreysia_nagpoorensis     
     |  |  |              |  |                 +--+                              
     |  |  |              |  |                    +-- Parreysia_plagiosoma       
     +--+  |              |  |                                                   
        |  |              |  |                    +-- Leoparreysia_tavoyensis    
        |  |              |  |                    |                              
        |  |              +--+  +-----------------+-- Leoparreysia_canefrii      
        |  |                 |  |                 |                              
        |  |                 |  |                 +-- Leoparreysia_olivacea      
        |  |                 |  |                                                
        |  |                 |  |           +-------- Coelatura_aegyptiaca[2]    
        |  |                 |  |           |                                    
        |  |                 |  |  +--------+  +----- Coelatura_aegyptiaca[1]    
        |  |                 +--+  |        |  |                                 
        |  |                    |  |        |  +----- Nitia_teretiuscula         
        |  |                    |  |        +--+                                 
        |  |                    |  |           |  +-- Coelatura_gabonensis       
        |  |                    |  |           |  |                              
        |  |                    |  |           +--+-- Coelatura_kunenensis       
        |  |                    |  |              |                              
        |  |                    +--+              +-- Prisodontopsis_mweruensis  
        |  |                       |                                             
        |  |                       |        +-------- Indonaia_shurtleffiana     
        |  |                       |        |                                    
        |  |                       |  +-----+  +----- Indonaia_bonneaudii        
        |  |                       |  |     |  |                                 
        |  |                       |  |     +--+----- Indonaia_rugosa            
        |  |                       |  |        |                                 
        |  |                       |  |        |  +-- Indonaia_andersoniana      
        |  |                       +--+        +--+                              
        |  |                          |           +-- Indonaia_caerulea          
        |  |                          |                                          
        |  |                          |  +----------- Indochinella_pugio         
        |  |                          |  |                                       
        |  |                          |  |     +----- Scabies_humilis            
        |  |                          +--+  +--+                                 
        |  |                             |  |  |  +-- Scabies_phaselus           
        |  |                             |  |  +--+                              
        |  |                             +--+     +-- Scabies_mandarinus         
        |  |                                |                                    
        |  |                                |  +----- Radiatula_myitkyinae       
        |  |                                +--+                                 
        |  |                                   |  +-- Radiatula_chaudhurii       
        |  |                                   +--+                              
        |  |                                      +-- Radiatula_mouhoti          
        |  |                                                                     
        |  |        +-------------------------------- Acuticosta_chinensis       
        |  |        |                                                            
        |  |        |  +----------------------------- Lepidodesma_languilati     
        |  |        |  |                                                         
        |  |        |  |                    +-------- Aculamprotula_tortuosa     
        |  |        |  |                    |                                    
        |  |        |  |        +-----------+  +----- Aculamprotula_coreana      
        |  |        |  |        |           |  |                                 
        |  |        |  |        |           +--+----- Aculamprotula_scripta      
        |  |        |  |        |              |                                 
        +--+        |  |        |              |  +-- Aculamprotula_fibrosa      
           |        |  |        |              +--+                              
           |        |  |        |                 +-- Aculamprotula_tientsinensis
           |  +-----+  +--------+                                                
           |  |     |  |        |           +-------- Unio_tumidus               
           |  |     |  |        |  +--------+                                    
           |  |     |  |        |  |        |  +----- Unio_crassus               
           |  |     |  |        |  |        +--+                                 
           |  |     |  |        |  |           |  +-- Unio_delphinus             
           |  |     |  |        |  |           +--+                              
           |  |     |  |        +--+              +-- Unio_pictorum              
           |  |     |  |           |                                             
           |  |     |  |           |              +-- Schistodesmus_lampreyanus  
           |  |     |  |           |  +-----------+                              
           |  |     |  |           |  |           +-- Schistodesmus_spinosus     
           |  |     |  |           |  |                                          
           |  |     |  |           +--+        +----- Nodularia_douglasiae[1]    
           |  |     +--+              |  +-----+                                 
           |  |        |              |  |     |  +-- Nodularia_douglasiae[2]    
           |  |        |              |  |     +--+                              
           |  |        |              +--+        +-- Nodularia_nuxpersicae      
           |  |        |                 |                                       
           |  |        |                 |  +-------- Tchangsinaia_piscicula     
           |  |        |                 +--+                                    
           |  |        |                    |  +----- Cuneopsis_rufescens        
           |  |        |                    +--+                                 
           |  |        |                       |  +-- Cuneopsis_heudei           
           |  |        |                       +--+                              
           |  |        |                          +-- Cuneopsis_celtiformis      
           |  |        |                                                         
           |  |        |                    +-------- Lanceolaria_gladiola       
           |  |        |                    |                                    
           |  |        |  +-----------------+  +----- Lanceolaria_grayii[1]      
           |  |        |  |                 |  |                                 
           |  |        |  |                 +--+----- Lanceolaria_grayii[2]      
           |  |        |  |                    |                                 
           |  |        |  |                    |  +-- Lanceolaria_lanceolata     
           |  |        |  |                    +--+                              
           |  |        |  |                       +-- Lanceolaria_triformis      
           |  |        |  |                                                      
           |  |        |  |     +-------------------- Anemina_arcaeformis        
           |  |        |  |     |                                                
           |  |        |  |  +--+  +----------------- Cristaria_plicata          
           |  |        +--+  |  |  |                                             
           |  |           |  |  +--+  +-------------- Pletholophus_tenuis        
           |  |           |  |     |  |                                          
           |  |           |  |     +--+           +-- Sinanodonta_angula         
           |  |           |  |        |  +--------+                              
           |  |           |  |        |  |        +-- Sinanodonta_lucida         
           |  |           |  |        +--+                                       
           |  |           |  |           |  +-------- Sinanodonta_woodiana[2]    
           |  |           |  |           +--+                                    
           |  |           |  |              |  +----- Sinanodonta_jourdyi        
           |  |           |  |              +--+                                 
           |  |           +--+                 |  +-- Sinanodonta_elliptica      
           |  |              |                 +--+                              
           |  |              |                    +-- Sinanodonta_woodiana[1]    
           |  |              |                                                   
           |  |              |              +-------- Alasmidonta_marginata      
           |  |              |              |                                    
           |  |              |              +-------- Lasmigona_costata          
           |  |              |              |                                    
           |  |              |              +-------- Pseudanodonta_complanata   
           |  |              |              |                                    
           |  |              |              +-------- Simpsonaias_ambigua        
           +--+              |              |                                    
              |              +--------------+-------- Strophitus_undulatus       
              |                             |                                    
              |                             |  +----- Anodonta_anatina           
              |                             +--+                                 
              |                             |  |  +-- Anodonta_cygnea            
              |                             |  +--+                              
              |                             |     +-- Anodonta_nuttalliana       
              |                             |                                    
              |                             |  +----- Pyganodon_grandis          
              |                             +--+                                 
              |                                |  +-- Utterbackia_imbecillis     
              |                                +--+                              
              |                                   +-- Utterbackia_peninsularis   
              |                                                                  
              |                          +----------- Hyriopsis_desowitzi        
              |                          |                                       
              |                          +----------- Rectidens_sumatrensis      
              |                          |                                       
              |                          |        +-- Hyriopsis_bialata          
              |                          |        |                              
              |                          |        +-- Hyriopsis_myersiana        
              |                    +-----+--------+                              
              |                    |     |        +-- Hyriopsis_panhai           
              |                    |     |        |                              
              |                    |     |        +-- Hyriopsis_khoratensis      
              |                    |     |                                       
              |                    |     |        +-- Ensidens_ingallsianus      
              |                    |     |  +-----+                              
              |                    |     |  |     +-- Ensidens_telus             
              |                    |     +--+                                    
              |                    |        |  +----- Ensidens_sagittarius       
              |  +-----------------+        +--+                                 
              |  |                 |           |  +-- Ensidens_spiculus          
              |  |                 |           +--+                              
              |  |                 |              +-- Ensidens_jaculus           
              |  |                 |                                             
              |  |                 |           +----- Physunio                   
              |  |                 |  +--------+                                 
              |  |                 |  |        |  +-- Physunio_modelli           
              |  |                 |  |        +--+                              
              |  |                 |  |           +-- Physunio_superbus          
              |  |                 |  |                                          
              |  |                 +--+     +-------- Yaukthwa_inlenensis        
              |  |                    |  +--+                                    
              |  |                    |  |  |  +----- Yaukthwa_panhai            
              |  |                    |  |  +--+                                 
              |  |                    |  |     |  +-- Yaukthwa_nesemanni         
              |  |                    |  |     +--+                              
              |  |                    +--+        +-- Yaukthwa_rectangularis     
              |  |                       |                                       
              |  |                       |        +-- Lens_contradens            
              |  |                       |  +-----+                              
              |  |                       |  |     +-- Lens_micropterus           
              |  |                       +--+                                    
              |  |                          |  +----- Lens_eximius               
              |  |                          +--+                                 
              |  |                             |  +-- Lens_comptus               
              |  |                             +--+                              
              |  |                                +-- Lens_novoselovi            
              |  |                                                               
              |  |                                +-- Megalonaias_nervosa        
              |  |                             +--+                              
              +--+                             |  +-- Uniomerus_carolinianus     
                 |     +-----------------------+                                 
                 |     |                       |  +-- Quadrula_quadrula          
                 |     |                       +--+                              
                 |     |                          +-- Quadrula_verrucosa         
                 |     |                                                         
                 |     |                    +-------- Elliptoideus_sloatianus    
                 |  +--+                    |                                    
                 |  |  |  +-----------------+     +-- Elliptio_complanata        
                 |  |  |  |                 |  +--+                              
                 |  |  |  |                 |  |  +-- Eurynia_dilatata           
                 |  |  |  |                 +--+                                 
                 |  |  |  |                    |  +-- Fusconaia_flava            
                 |  |  +--+                    +--+                              
                 |  |     |                       +-- Pleurobema_sintoxia        
                 |  |     |                                                      
                 |  |     |  +----------------------- Amblema_plicata            
                 |  |     |  |                                                   
                 |  |     |  |                 +----- Toxolasma_parvum           
                 |  |     +--+  +--------------+                                 
                 |  |        |  |              |  +-- Obliquaria_reflexa         
                 |  |        |  |              +--+                              
                 |  |        |  |                 +-- Plectomerus_dombeyanus     
                 |  |        +--+                                                
                 |  |           |  +----------------- Potamilus_leptodon         
                 |  |           |  |                                             
                 |  |           |  |  +-------------- Potamilus_alatus           
                 |  |           +--+  |                                          
                 |  |              |  |  +----------- Venustaconcha_ellipsiformis
                 |  |              |  |  |                                       
                 |  |              +--+  +----------- Leaunio_lienosus           
                 |  |                 |  |                                       
                 +--+                 |  |        +-- Ellipsaria_lineolata       
                    |                 |  +--------+                              
                    |                 +--+        +-- Truncilla_truncata         
                    |                    |                                       
                    |                    |        +-- Medionidus_conradicus      
                    |                    +--------+                              
                    |                    |        +-- Ptychobranchus_subtentus   
                    |                    |                                       
                    |                    |  +-------- Cambarunio_iris            
                    |                    +--+                                    
                    |                       |  +----- Ortmanniana_ligamentina    
                    |                       +--+                                 
                    |                          |  +-- Lampsilis_cardium          
                    |                          +--+                              
                    |                             +-- Lampsilis_ornata           
                    |                                                            
                    |                          +----- Chamberlainia_somsakpanhai 
                    |  +-----------------------+                                 
                    |  |                       |  +-- Sinohyriopsis_cumingii     
                    |  |                       +--+                              
                    |  |                          +-- Sinohyriopsis_schlegelii   
                    |  |                                                         
                    |  |                          +-- Potomida_littoralis        
                    |  |                    +-----+                              
                    |  |                    |     +-- Pronodularia_japanensis    
                    |  |  +-----------------+                                    
                    +--+  |                 |  +----- Lamprotula_leaii           
                       |  |                 +--+                                 
                       |  |                    |  +-- Lamprotula_caveata         
                       |  |                    +--+                              
                       |  |                       +-- Lamprotula_cornuumlunae    
                       |  |                                                      
                       |  |                 +-------- Sinosolenaia_carinata      
                       |  |              +--+                                    
                       |  |              |  |  +----- Sinosolenaia_recognita     
                       |  |              |  +--+                                 
                       +--+              |     |  +-- Sinosolenaia_oleivora      
                          |              |     +--+                              
                          |  +-----------+        +-- Sinosolenaia_iridinea      
                          |  |           |                                       
                          |  |           |     +----- Gonidea_angulata           
                          |  |           |  +--+                                 
                          |  |           |  |  |  +-- Leguminaia_wheatleyi       
                          |  |           |  |  +--+                              
                          |  |           +--+     +-- Microcondylaea_bonellii    
                          |  |              |                                    
                          |  |              |     +-- Ptychorhynchus_pfisteri    
                          |  |              |     |                              
                          +--+              +-----+-- Parvasolenaia_rivularis    
                             |                    |                              
                             |                    +-- Koreosolenaia_sitgyensis   
                             |                                                   
                             |                 +----- Indopseudodon_salwenianus  
                             |  +--------------+                                 
                             |  |              |  +-- Indopseudodon_indawgyiensis
                             |  |              +--+                              
                             |  |                 +-- Indopseudodon_bogani       
                             |  |                                                
                             |  |           +-------- Pilsbryoconcha_exilis[3]   
                             +--+  +--------+                                    
                                |  |        |  +----- Pilsbryoconcha_exilis[2]   
                                |  |        +--+                                 
                                |  |           |  +-- Pilsbryoconcha_exilis[1]   
                                |  |           +--+                              
                                |  |              +-- Pilsbryoconcha_carinifera  
                                |  |                                             
                                +--+     +----------- Nyeinchanconcha_nyeinchani 
                                   |  +--+                                       
                                   |  |  |  +-------- Thaiconcha_callifera       
                                   |  |  +--+                                    
                                   |  |     |  +----- Bineurus_loeiensis         
                                   |  |     +--+                                 
                                   |  |        |  +-- Bineurus_mouhotii          
                                   +--+        +--+                              
                                      |           +-- Bineurus_exilis            
                                      |                                          
                                      |        +----- Pseudodon_vagulus          
                                      |     +--+                                 
                                      |     |  |  +-- Pseudodon_mekongi          
                                      |     |  +--+                              
                                      +-----+     +-- Pseudodon_vondembuschianus 
                                            |                                    
                                            |     +-- Sundadontina_cumingii      
                                            |     |                              
                                            +-----+-- Sundadontina_tumida[1]     
                                                  |                              
                                                  +-- Sundadontina_tumida[2]     

Huang et al., 2019 [Publ 3843], fig. S3

iT tT bi riS oT roT roS iD
10 10 8 3 191 189 2 6.0
                                                              +----- Velesunio_ambiguus           
                                                           +--+                                   
                                                           |  |  +-- Alathyria_jacksoni           
                                                           |  +--+                                
                                                  +--------+     +-- Alathyria_profuga            
                                                  |        |                                      
                                                  |        |  +----- Alathyria_pertexta           
                                                  |        +--+                                   
                                                  |           |  +-- Lortiella_froggatti          
  +-----------------------------------------------+           +--+                                
  |                                               |              +-- Microdontia_anodontaeformis  
  |                                               |                                               
  |                                               |  +-------------- Westralunio_carteri          
  |                                               |  |                                            
  |                                               +--+  +----------- Echyridella_menziesii        
  |                                                  |  |                                         
  |                                                  |  |     +----- Diplodon_suavidicus          
  |                                                  +--+  +--+                                   
  |                                                     |  |  |  +-- Castalia_ambigua             
  |                                                     |  |  +--+                                
  |                                                     +--+     +-- Triplodon_corrugatus         
  |                                                        |                                      
  |                                                        |  +----- Hyridella_australis          
  |                                                        +--+                                   
  |                                                           |  +-- Hyridella_depressa           
  |                                                           +--+                                
  |                                                              +-- Hyridella_drapeta            
  |                                                                                               
  |                                                        +-------- Anodontites_trapesialis **      
  |                                                     +--+                                      
  |                                                     |  |  +----- Mutela_rostrata **              
  |                                                     |  +--+                                   
  |                                                     |     |  +-- Mutela_dubia **                 
  |                                                     |     +--+                                
  |  +--------------------------------------------------+        +-- Mutela_hargeri **               
  |  |                                                  |                                         
  |  |                                                  |     +----- Mycetopoda_siliquosa **         
--+  |                                                  |  +--+                                   
  |  |                                                  |  |  |  +-- Tamsiella_schroeteriana **      
  |  |                                                  |  |  +--+                                
  |  |                                                  +--+     +-- Lamproscapha_ensiformis **      
  |  |                                                     |                                      
  |  |                                                     |  +----- Etheria_elliptica **            
  |  |                                                     +--+                                   
  |  |                                                        |  +-- Aspatharia_pfeifferiana **      
  |  |                                                        +--+                                
  |  |                                                           +-- Chambardia_wahlbergi **         
  |  |                                                                                            
  |  |                                         +-------------------- Gibbosula_rochechouartii     
  |  |                                         |                                                  
  |  |                                         |  +----------------- Gibbosula_laosensis          
  |  |  +--------------------------------------+  |                                               
  |  |  |                                      |  |        +-------- Cumberlandia_monodonta       
  |  |  |                                      |  |  +-----+                                      
  |  |  |                                      +--+  |     |  +----- Pseudunio_marocanus          
  |  |  |                                         |  |     +--+                                   
  |  |  |                                         |  |        |  +-- Pseudunio_auricularius       
  |  |  |                                         |  |        +--+                                
  |  |  |                                         +--+           +-- Pseudunio_homsensis          
  |  |  |                                            |                                            
  |  |  |                                            |           +-- Margaritifera_dahurica       
  |  |  |                                            |  +--------+                                
  |  |  |                                            |  |        +-- Margaritifera_margaritifera  
  |  |  |                                            +--+                                         
  |  |  |                                               |  +-------- Margaritifera_falcata        
  +--+  |                                               |  |                                      
     |  |                                               +--+     +-- Margaritifera_hembeli        
     |  |                                                  |  +--+                                
     |  |                                                  |  |  +-- Margaritifera_marrianae      
     |  |                                                  +--+                                   
     |  |                                                     |  +-- Margaritifera_laevis         
     |  |                                                     +--+                                
     |  |                                                        +-- Margaritifera_middendorffi   
     |  |                                                                                         
     |  |                                                     +----- Trapezidens_exolescens       
     |  |                                               +-----+                                   
     |  |                                               |     |  +-- Trapezidens_angustior        
     |  |                                               |     +--+                                
     |  |                                               |        +-- Trapezidens_dolichorhynchus  
     |  |                                               |                                         
     |  |                                +--------------+        +-- Lamellidens_corrianus        
     |  |                                |              |  +-----+                                
     |  |                                |              |  |     +-- Lamellidens_savadiensis      
     |  |                                |              |  |                                      
     |  |                                |              |  |  +----- Lamellidens_lamellatus       
     |  |                                |              +--+  |                                   
     |  |                                |                 |  +----- Lamellidens_jenkinsianus     
     |  |                                |                 |  |                                   
     |  |                                |                 |  |  +-- Lamellidens_marginalis       
     |  |                                |                 +--+--+                                
     |  |                                |                    |  +-- Lamellidens_ferrugineus      
     |  |                                |                    |                                   
     |  |  +-----------------------------+                    |  +-- Lamellidens_mainwaringi[1]   
     |  |  |                             |                    |  |                                
     |  |  |                             |                    +--+-- Lamellidens_mainwaringi[2]   
     |  |  |                             |                       |                                
     |  |  |                             |                       +-- Lamellidens_generosus        
     +--+  |                             |                                                        
        |  |                             |                    +----- Parreysia_keralaensis        
        |  |                             |  +-----------------+                                   
        |  |                             |  |                 |  +-- Parreysia_nagpoorensis       
        |  |                             |  |                 +--+                                
        |  |                             |  |                    +-- Parreysia_plagiosoma         
        |  |                             |  |                                                     
        |  |                             |  |                 +----- Leoparreysia_canefrii        
        |  |                             +--+  +--------------+                                   
        |  |                                |  |              |  +-- Leoparreysia_tavoyensis      
        |  |                                |  |              +--+                                
        |  |                                |  |                 +-- Leoparreysia_olivacea        
        |  |                                |  |                                                  
        |  |                                |  |        +----------- Coelatura_aegyptiaca[2]      
        |  |                                |  |        |                                         
        |  |                                +--+  +-----+        +-- Coelatura_aegyptiaca[1]      
        |  |                                   |  |     |  +-----+                                
        |  |                                   |  |     |  |     +-- Nitia_teretiuscula           
        |  |                                   |  |     +--+                                      
        |  |                                   |  |        |  +----- Coelatura_kunenensis         
        |  |                                   |  |        +--+                                   
        |  |                                   |  |           |  +-- Coelatura_gabonensis         
        |  |                                   |  |           +--+                                
        |  |                                   +--+              +-- Prisodontopsis_mweruensis    
        |  |                                      |                                               
        |  |                                      |        +-------- Indonaia_shurtleffiana       
        |  |                                      |        |                                      
        |  |                                      |  +-----+  +----- Indonaia_bonneaudii          
        |  |                                      |  |     |  |                                   
        |  |                                      |  |     +--+----- Indonaia_rugosa              
        |  |                                      |  |        |                                   
        |  |                                      |  |        |  +-- Indonaia_andersoniana        
        |  |                                      +--+        +--+                                
        |  |                                         |           +-- Indonaia_caerulea            
        |  |                                         |                                            
        |  |                                         |  +----------- Indochinella_pugio           
        |  |                                         |  |                                         
        |  |                                         |  |     +----- Scabies_phaselus             
        |  |                                         +--+  +--+                                   
        |  |                                            |  |  |  +-- Scabies_humilis              
        |  |                                            |  |  +--+                                
        |  |                                            +--+     +-- Scabies_mandarinus           
        |  |                                               |                                      
        |  |                                               |  +----- Radiatula_mouhoti            
        |  |                                               +--+                                   
        |  |                                                  |  +-- Radiatula_myitkyinae         
        |  |                                                  +--+                                
        +--+                                                     +-- Radiatula_chaudhurii         
           |                                                                                      
           |                                                  +----- Indopseudodon_indawgyiensis  
           |                                   +--------------+                                   
           |                                   |              |  +-- Indopseudodon_bogani         
           |                                   |              +--+                                
           |                                   |                 +-- Indopseudodon_salwenianus    
           |                                   |                                                  
           |                                   |           +-------- Pilsbryoconcha_exilis[3]     
           |  +--------------------------------+  +--------+                                      
           |  |                                |  |        |  +----- Pilsbryoconcha_exilis[2]     
           |  |                                |  |        +--+                                   
           |  |                                |  |           |  +-- Pilsbryoconcha_carinifera    
           |  |                                |  |           +--+                                
           |  |                                |  |              +-- Pilsbryoconcha_exilis[1]     
           |  |                                |  |                                               
           |  |                                +--+     +----------- Sundadontina_tumida          
           |  |                                   |  +--+                                         
           |  |                                   |  |  |  +-------- Pseudodon_vagulus            
           |  |                                   |  |  +--+                                      
           |  |                                   |  |     |  +----- Pseudodon_vondembuschianus[1]
           |  |                                   |  |     +--+                                   
           |  |                                   |  |        |  +-- Pseudodon_mekongi            
           |  |                                   +--+        +--+                                
           |  |                                      |           +-- Pseudodon_vondembuschianus[2]
           |  |                                      |                                            
           |  |                                      |           +-- Nyeinchanconcha_nyeinchani   
           |  |                                      |  +--------+                                
           |  |                                      |  |        +-- Sundadontina_cumingii        
           |  |                                      +--+                                         
           |  |                                         |  +-------- Thaiconcha_callifera         
           |  |                                         +--+                                      
           |  |                                            |  +----- Bineurus_loeiensis           
           |  |                                            +--+                                   
           |  |                                               |  +-- Bineurus_mouhotii            
           |  |                                               +--+                                
           |  |                                                  +-- Bineurus_exilis              
           |  |                                                                                   
           |  |                                                  +-- Ensidens_telus               
           |  |                                            +-----+                                
           |  |                                            |     +-- Ensidens_ingallsianus        
           |  |                                      +-----+                                      
           |  |                                      |     |  +----- Ensidens_sagittarius         
           |  |                                      |     +--+                                   
           |  |                                      |        |  +-- Ensidens_spiculus            
           |  |                                   +--+        +--+                                
           |  |                                   |  |           +-- Ensidens_jaculus             
           |  |                                   |  |                                            
           |  |                                   |  |  +----------- Hyriopsis_desowitzi          
           |  |                                   |  |  |                                         
           |  |                                   |  +--+  +-------- Rectidens_sumatrensis        
           |  |                                   |     |  |                                      
           |  |                                   |     +--+     +-- Hyriopsis_panhai             
           +--+                                   |        |  +--+                                
              |                                   |        |  |  +-- Hyriopsis_khoratensis        
              |  +--------------------------------+        +--+                                   
              |  |                                |           |  +-- Hyriopsis_bialata            
              |  |                                |           +--+                                
              |  |                                |              +-- Hyriopsis_myersiana          
              |  |                                |                                               
              |  |                                |           +----- Physunio                     
              |  |                                |  +--------+                                   
              |  |                                |  |        |  +-- Physunio_superbus            
              |  |                                |  |        +--+                                
              |  |                                |  |           +-- Physunio_modelli             
              |  |                                |  |                                            
              |  |                                +--+     +-------- Yaukthwa_inlenensis          
              |  |                                   |  +--+                                      
              |  |                                   |  |  |  +----- Yaukthwa_panhai              
              |  |                                   |  |  +--+                                   
              |  |                                   |  |     |  +-- Yaukthwa_nesemanni           
              |  |                                   |  |     +--+                                
              |  |                                   +--+        +-- Yaukthwa_rectangularis       
              |  |                                      |                                         
              |  |                                      |        +-- Lens_contradens              
              |  |                                      |  +-----+                                
              |  |                                      |  |     +-- Lens_micropterus             
              |  |                                      +--+                                      
              |  |                                         |  +----- Lens_eximius                 
              |  |                                         +--+                                   
              |  |                                            |  +-- Lens_novoselovi              
              |  |                                            +--+                                
              |  |                                               +-- Lens_comptus                 
              |  |                                                                                
              |  |                                            +----- Chamberlainia_somsakpanhai   
              |  |     +--------------------------------------+                                   
              |  |     |                                      |  +-- Sinohyriopsis_cumingii       
              |  |     |                                      +--+                                
              |  |     |                                         +-- Sinohyriopsis_schlegelii     
              |  |     |                                                                          
              |  |     |                                         +-- Potomida_littoralis          
              |  |     |                                   +-----+                                
              |  |     |                                   |     +-- Pronodularia_japanensis      
              |  |     |                             +-----+                                      
              |  |     |                             |     |  +----- Lamprotula_leaii             
              |  |     |                             |     +--+                                   
              |  |     |                             |        |  +-- Lamprotula_caveata           
              +--+     |                             |        +--+                                
                 |     |                             |           +-- Lamprotula_cornuumlunae      
                 |  +--+  +--------------------------+                                            
                 |  |  |  |                          |        +----- Sinosolenaia_recognita       
                 |  |  |  |                          |  +-----+                                   
                 |  |  |  |                          |  |     |  +-- Sinosolenaia_oleivora        
                 |  |  |  |                          |  |     +--+                                
                 |  |  |  |                          |  |        +-- Sinosolenaia_iridinea        
                 |  |  |  |                          +--+                                         
                 |  |  |  |                             |     +----- Gonidea_angulata             
                 |  |  |  |                             |  +--+                                   
                 |  |  |  |                             |  |  |  +-- Leguminaia_wheatleyi         
                 |  |  |  |                             |  |  +--+                                
                 |  |  |  |                             +--+     +-- Microcondylaea_bonellii      
                 |  |  |  |                                |                                      
                 |  |  |  |                                |  +----- Ptychorhynchus_pfisteri      
                 |  |  +--+                                +--+                                   
                 |  |     |                                   |  +-- Parvasolenaia_rivularis      
                 |  |     |                                   +--+                                
                 |  |     |                                      +-- Koreosolenaia_sitgyensis     
                 |  |     |                                                                       
                 |  |     |                                      +-- Megalonaias_nervosa          
                 |  |     |                                   +--+                                
                 |  |     |                                   |  +-- Uniomerus_carolinianus       
                 |  |     |  +--------------------------------+                                   
                 |  |     |  |                                |  +-- Quadrula_quadrula            
                 |  |     |  |                                +--+                                
                 |  |     |  |                                   +-- Quadrula_verrucosa           
                 |  |     |  |                                                                    
                 |  |     |  |                             +-------- Elliptoideus_sloatianus      
                 |  |     +--+                             |                                      
                 |  |        |  +--------------------------+     +-- Elliptio_complanata          
                 |  |        |  |                          |  +--+                                
                 |  |        |  |                          |  |  +-- Eurynia_dilatata             
                 |  |        |  |                          +--+                                   
                 |  |        |  |                             |  +-- Fusconaia_flava              
                 |  |        +--+                             +--+                                
                 |  |           |                                +-- Pleurobema_sintoxia          
                 |  |           |                                                                 
                 |  |           |  +-------------------------------- Amblema_plicata              
                 |  |           |  |                                                              
                 |  |           +--+  +----------------------------- Toxolasma_parvum             
                 |  |              |  |                                                           
                 +--+              +--+                          +-- Obliquaria_reflexa           
                    |                 |  +-----------------------+                                
                    |                 |  |                       +-- Plectomerus_dombeyanus       
                    |                 +--+                                                        
                    |                    |  +----------------------- Potamilus_leptodon           
                    |                    |  |                                                     
                    |                    +--+  +-------------------- Potamilus_alatus             
                    |                       |  |                                                  
                    |                       |  |                 +-- Medionidus_conradicus        
                    |                       +--+  +--------------+                                
                    |                          |  |              +-- Ptychobranchus_subtentus     
                    |                          |  |                                               
                    |                          +--+              +-- Ellipsaria_lineolata         
                    |                             |  +-----------+                                
                    |                             |  |           +-- Truncilla_truncata           
                    |                             +--+                                            
                    |                                |  +----------- Leaunio_lienosus             
                    |                                |  |                                         
                    |                                +--+        +-- Venustaconcha_ellipsiformis  
                    |                                   |  +-----+                                
                    |                                   |  |     +-- Cambarunio_iris              
                    |                                   +--+                                      
                    |                                      |  +----- Ortmanniana_ligamentina      
                    |                                      +--+                                   
                    |                                         |  +-- Lampsilis_cardium            
                    |                                         +--+                                
                    |                                            +-- Lampsilis_ornata             
                    |                                                                             
                    |           +----------------------------------- Acuticosta_chinensis         
                    |           |                                                                 
                    |           |                          +-------- Aculamprotula_tortuosa       
                    |           |                          |                                      
                    |           |              +-----------+     +-- Aculamprotula_coreana        
                    |           |              |           |  +--+                                
                    |           |              |           |  |  +-- Aculamprotula_scripta        
                    |           |              |           +--+                                   
                    |           |              |              |  +-- Aculamprotula_fibrosa        
                    |           |              |              +--+                                
                    |           |              |                 +-- Aculamprotula_tientsinensis  
                    |           |  +-----------+                                                  
                    |           |  |           |           +-------- Unio_tumidus                 
                    +-----------+  |           |  +--------+                                      
                                |  |           |  |        |  +----- Unio_crassus                 
                                |  |           |  |        +--+                                   
                                |  |           |  |           |  +-- Unio_delphinus               
                                |  |           |  |           +--+                                
                                |  |           +--+              +-- Unio_pictorum                
                                |  |              |                                               
                                |  |              |           +----- Nodularia_douglasiae[1]      
                                |  |              |  +--------+                                   
                                |  |              |  |        |  +-- Nodularia_douglasiae[2]      
                                |  |              |  |        +--+                                
                                |  |              +--+           +-- Nodularia_nuxpersicae        
                                |  |                 |                                            
                                +--+                 |  +----------- Schistodesmus_lampreyanus    
                                   |                 +--+                                         
                                   |                    |  +-------- Tchangsinaia_piscicula       
                                   |                    +--+                                      
                                   |                       |  +----- Cuneopsis_rufescens          
                                   |                       +--+                                   
                                   |                          |  +-- Cuneopsis_heudei             
                                   |                          +--+                                
                                   |                             +-- Cuneopsis_celtiformis        
                                   |                                                              
                                   |  +----------------------------- Lepidodesma_languilati       
                                   |  |                                                           
                                   |  |                    +-------- Lanceolaria_gladiola         
                                   |  |  +-----------------+                                      
                                   |  |  |                 |  +----- Lanceolaria_grayii           
                                   |  |  |                 +--+                                   
                                   +--+  |                    |  +-- Lanceolaria_lanceolata       
                                      |  |                    +--+                                
                                      |  |                       +-- Lanceolaria_triformis        
                                      |  |                                                        
                                      |  |     +-------------------- Anemina_arcaeformis          
                                      |  |     |                                                  
                                      |  |  +--+  +----------------- Cristaria_plicata            
                                      +--+  |  |  |                                               
                                         |  |  +--+  +-------------- Pletholophus_tenuis          
                                         |  |     |  |                                            
                                         |  |     +--+           +-- Sinanodonta_angula           
                                         |  |        |  +--------+                                
                                         |  |        |  |        +-- Sinanodonta_lucida           
                                         |  |        +--+                                         
                                         |  |           |  +-------- Sinanodonta_woodiana[2]      
                                         |  |           +--+                                      
                                         |  |              |  +----- Sinanodonta_jourdyi          
                                         +--+              +--+                                   
                                            |                 |  +-- Sinanodonta_elliptica        
                                            |                 +--+                                
                                            |                    +-- Sinanodonta_woodiana[1]      
                                            |                                                     
                                            |                    +-- Alasmidonta_marginata        
                                            |           +--------+                                
                                            |           |        +-- Lasmigona_costata            
                                            |           |                                         
                                            |           |     +----- Anodonta_anatina             
                                            |           +-----+                                   
                                            |           |     |  +-- Anodonta_cygnea              
                                            +-----------+     +--+                                
                                                        |        +-- Anodonta_nuttalliana         
                                                        |                                         
                                                        |     +----- Pseudanodonta_complanata     
                                                        |  +--+                                   
                                                        |  |  |  +-- Simpsonaias_ambigua          
                                                        |  |  +--+                                
                                                        +--+     +-- Strophitus_undulatus         
                                                           |                                      
                                                           |  +----- Pyganodon_grandis            
                                                           +--+                                   
                                                              |  +-- Utterbackia_imbecillis       
                                                              +--+                                
                                                                 +-- Utterbackia_peninsularis     

Huang et al., 2019 [Publ 3843], fig. S4

iT tT bi riS oT roT roS iD
10 10 6 3 192 190 2 6.0
                          +----------- Etheria_elliptica **          
                          |                                       
                          |        +-- Aspatharia_pfeifferiana **    
                          +--------+                              
                          |        +-- Chambardia_wahlbergi **       
                          |                                       
  +-----------------------+     +----- Mycetopoda_siliquosa **       
  |                       +-----+                                 
  |                       |     |  +-- Tamsiella_schroeteriana **    
  |                       |     +--+                              
  |                       |        +-- Lamproscapha_ensiformis **    
  |                       |                                       
  |                       |  +-------- Anodontites_trapesialis **    
  |                       +--+                                    
  |                          |  +----- Mutela_hargeri **             
  |                          +--+                                 
  |                             |  +-- Mutela_dubia **               
  |                             +--+                              
  |                                +-- Mutela_rostrata **            
  |                                                               
  |                       +----------- Echyridella_menziesii      
  |                       |                                       
  |                       +----------- Westralunio_carteri        
  |                       |                                       
  |                       |     +----- Diplodon_suavidicus        
  |                       +-----+                                 
  |                       |     |  +-- Castalia_ambigua           
  |                       |     +--+                              
  |                       |        +-- Triplodon_corrugatus       
  |                       |                                       
  |  +--------------------+     +----- Hyridella_australis        
  |  |                    +-----+                                 
  |  |                    |     |  +-- Hyridella_depressa         
  |  |                    |     +--+                              
  |  |                    |        +-- Hyridella_drapeta          
  |  |                    |                                       
  |  |                    |     +----- Velesunio_ambiguus         
  |  |                    |  +--+                                 
--+  |                    |  |  |  +-- Alathyria_jacksoni         
  |  |                    |  |  +--+                              
  |  |                    +--+     +-- Alathyria_profuga          
  |  |                       |                                    
  |  |                       |  +----- Alathyria_pertexta         
  |  |                       +--+                                 
  |  |                          |  +-- Lortiella_froggatti        
  |  |                          +--+                              
  |  |                             +-- Microdontia_anodontaeformis
  |  |                                                            
  |  |           +-------------------- Gibbosula_rochechouartii   
  |  |           |                                                
  |  |  +--------+  +----------------- Gibbosula_laosensis        
  |  |  |        |  |                                             
  |  |  |        +--+  +-------------- Cumberlandia_monodonta     
  |  |  |           |  |                                          
  |  |  |           |  |        +----- Pseudunio_marocanus        
  |  |  |           +--+  +-----+                                 
  |  |  |              |  |     |  +-- Pseudunio_auricularius     
  |  |  |              |  |     +--+                              
  |  |  |              |  |        +-- Pseudunio_homsensis        
  |  |  |              +--+                                       
  |  |  |                 |  +-------- Margaritifera_falcata      
  |  |  |                 |  |                                    
  |  |  |                 |  |     +-- Margaritifera_dahurica     
  |  |  |                 |  +-----+                              
  |  |  |                 +--+     +-- Margaritifera_margaritifera
  |  |  |                    |                                    
  |  |  |                    |  +----- Margaritifera_laevis       
  |  |  |                    |  |                                 
  |  |  |                    +--+----- Margaritifera_middendorffi 
  |  |  |                       |                                 
  +--+  |                       |  +-- Margaritifera_hembeli      
     |  |                       +--+                              
     |  |                          +-- Margaritifera_marrianae    
     |  |                                                         
     |  |                          +-- Parreysia_keralaensis      
     |  |                          |                              
     |  |     +--------------------+-- Parreysia_nagpoorensis     
     |  |     |                    |                              
     |  |     |                    +-- Parreysia_plagiosoma       
     |  |     |                                                   
     |  |     |                 +----- Trapezidens_exolescens     
     |  |     |        +--------+                                 
     |  |     |        |        |  +-- Trapezidens_angustior      
     |  |     |        |        +--+                              
     |  |     +--------+           +-- Trapezidens_dolichorhynchus
     |  |     |        |                                          
     |  |     |        |  +----------- Lamellidens_corrianus      
     |  |     |        |  |                                       
     |  |     |        +--+  +-------- Lamellidens_savadiensis    
     |  |     |           |  |                                    
     |  |     |           |  |  +----- Lamellidens_marginalis     
     |  |     |           +--+  |                                 
     |  |     |              |  +----- Lamellidens_ferrugineus    
     |  |     |              |  |                                 
     |  |  +--+              +--+  +-- Lamellidens_mainwaringi[1] 
     |  |  |  |                 |  |                              
     |  |  |  |                 |  +-- Lamellidens_mainwaringi[2] 
     |  |  |  |                 |  |                              
     |  |  |  |                 +--+-- Lamellidens_lamellatus     
     |  |  |  |                    |                              
     |  |  |  |                    +-- Lamellidens_generosus      
     |  |  |  |                    |                              
     |  |  |  |                    +-- Lamellidens_jenkinsianus   
     |  |  |  |                                                   
     |  |  |  |                 +----- Leoparreysia_tavoyensis    
     |  |  |  |  +--------------+                                 
     |  |  |  |  |              |  +-- Leoparreysia_canefrii      
     |  |  |  |  |              +--+                              
     |  |  |  |  |                 +-- Leoparreysia_olivacea      
     |  |  |  |  |                                                
     +--+  |  |  |           +-------- Coelatura_aegyptiaca[2]    
        |  |  |  |           |                                    
        |  |  |  |           +-------- Nitia_teretiuscula         
        |  |  +--+  +--------+                                    
        |  |     |  |        |  +----- Coelatura_aegyptiaca[1]    
        |  |     |  |        |  |                                 
        |  |     |  |        +--+  +-- Coelatura_gabonensis       
        |  |     |  |           |  |                              
        |  |     |  |           +--+-- Coelatura_kunenensis       
        |  |     |  |              |                              
        |  |     |  |              +-- Prisodontopsis_mweruensis  
        |  |     +--+                                             
        |  |        |           +----- Indonaia_shurtleffiana     
        |  |        |           |                                 
        |  |        |           |  +-- Indonaia_bonneaudii        
        |  |        |  +--------+--+                              
        |  |        |  |        |  +-- Indonaia_rugosa            
        |  |        |  |        |                                 
        |  |        |  |        |  +-- Indonaia_andersoniana      
        |  |        |  |        +--+                              
        |  |        +--+           +-- Indonaia_caerulea          
        |  |           |                                          
        |  |           |  +----------- Indochinella_pugio         
        |  |           |  |                                       
        |  |           |  |     +----- Scabies_humilis            
        |  |           +--+  +--+                                 
        |  |              |  |  |  +-- Scabies_phaselus           
        |  |              |  |  +--+                              
        |  |              +--+     +-- Scabies_mandarinus         
        |  |                 |                                    
        |  |                 |  +----- Radiatula_mouhoti          
        |  |                 +--+                                 
        |  |                    |  +-- Radiatula_myitkyinae       
        |  |                    +--+                              
        |  |                       +-- Radiatula_chaudhurii       
        |  |                                                      
        |  |     +-------------------- Chamberlainia_somsakpanhai 
        |  |     |                                                
        |  |     +-------------------- Megalonaias_nervosa        
        |  |     |                                                
        |  |     +-------------------- Uniomerus_carolinianus     
        |  |     |                                                
        |  |     |                 +-- Quadrula_quadrula          
        |  |     +-----------------+                              
        |  |     |                 +-- Quadrula_verrucosa         
        |  |     |                                                
        |  |     |                 +-- Sinohyriopsis_cumingii     
        |  |     +-----------------+                              
        |  |     |                 +-- Sinohyriopsis_schlegelii   
        |  |     |                                                
        |  |     |                 +-- Potomida_littoralis        
        |  |     |           +-----+                              
        |  |     |           |     +-- Pronodularia_japanensis    
        |  |     +-----------+                                    
        |  |     |           |  +----- Lamprotula_leaii           
        |  |     |           +--+                                 
        |  |     |              |  +-- Lamprotula_caveata         
        |  |     |              +--+                              
        |  |     |                 +-- Lamprotula_cornuumlunae    
        |  |     |                                                
        |  |     |           +-------- Sinosolenaia_carinata      
        +--+     |           |                                    
           |     |           |  +----- Gonidea_angulata           
           |     |           +--+                                 
           |     |           |  |  +-- Leguminaia_wheatleyi       
           |     |           |  +--+                              
           |     |           |     +-- Microcondylaea_bonellii    
           |     +-----------+                                    
           |     |           |  +----- Ptychorhynchus_pfisteri    
           |     |           +--+                                 
           |     |           |  |  +-- Parvasolenaia_rivularis    
           |     |           |  +--+                              
           |     |           |     +-- Koreosolenaia_sitgyensis   
           |     |           |                                    
           |     |           |  +----- Sinosolenaia_recognita     
           |     |           +--+                                 
           |     |              |  +-- Sinosolenaia_iridinea      
           |     |              +--+                              
           |     |                 +-- Sinosolenaia_oleivora      
           |     |                                                
           |     |              +----- Indopseudodon_salwenianus  
           |     |        +-----+                                 
           |     |        |     |  +-- Indopseudodon_indawgyiensis
           |     |        |     +--+                              
           |     |        |        +-- Indopseudodon_bogani       
           |     |        |                                       
           |     |        |  +-------- Nyeinchanconcha_nyeinchani 
           |     |        |  |                                    
           |     |        |  +-------- Thaiconcha_callifera       
           |     +--------+  |                                    
           |     |        |  +-------- Sundadontina_cumingii      
           |     |        |  |                                    
           |     |        |  +-------- Sundadontina_tumida[1]     
           |     |        |  |                                    
           |     |        |  +-------- Sundadontina_tumida[2]     
           |     |        |  |                                    
           |     |        |  |  +----- Pilsbryoconcha_exilis[2]   
           |     |        +--+--+                                 
           |     |           |  |  +-- Pilsbryoconcha_carinifera  
           |     |           |  +--+                              
           |     |           |     +-- Pilsbryoconcha_exilis[1]   
           |     |           |                                    
           |     |           |  +----- Pseudodon_vagulus          
           |     |           +--+                                 
           |     |           |  |  +-- Pseudodon_mekongi          
           |     |           |  +--+                              
           |     |           |     +-- Pseudodon_vondembuschianus 
           |     |           |                                    
           |     |           |  +----- Bineurus_loeiensis         
           |     |           +--+                                 
           |     |              |  +-- Bineurus_mouhotii          
           |     |              +--+                              
           |     |                 +-- Bineurus_exilis            
           |     |                                                
           |     |     +-------------- Amblema_plicata            
           |     |     |                                          
           |     |     +-------------- Toxolasma_parvum           
           |     |     |                                          
           |     |     |           +-- Obliquaria_reflexa         
           |     |     +-----------+                              
           |     |     |           +-- Plectomerus_dombeyanus     
           |     |     |                                          
           |     |     |  +----------- Elliptoideus_sloatianus    
           |     |     +--+                                       
           |     |     |  |  +-------- Elliptio_complanata        
           |     |     |  +--+                                    
           |     |     |     |  +----- Eurynia_dilatata           
           |     +-----+     +--+                                 
           |     |     |        |  +-- Fusconaia_flava            
           |     |     |        +--+                              
           |     |     |           +-- Pleurobema_sintoxia        
           |     |     |                                          
           |     |     |     +-------- Ellipsaria_lineolata       
           |     |     |     |                                    
           |     |     |     +-------- Potamilus_leptodon         
           |     |     |     |                                    
           |     |     |     +-------- Medionidus_conradicus      
           |     |     |     |                                    
           +-----+     |     +-------- Potamilus_alatus           
                 |     |     |                                    
                 |     |     +-------- Ptychobranchus_subtentus   
                 |     +-----+                                    
                 |           +-------- Truncilla_truncata         
                 |           |                                    
                 |           +-------- Venustaconcha_ellipsiformis
                 |           |                                    
                 |           +-------- Cambarunio_iris            
                 |           |                                    
                 |           +-------- Leaunio_lienosus           
                 |           |                                    
                 |           |  +----- Ortmanniana_ligamentina    
                 |           +--+                                 
                 |              |  +-- Lampsilis_cardium          
                 |              +--+                              
                 |                 +-- Lampsilis_ornata           
                 |                                                
                 |     +-------------- Hyriopsis_desowitzi        
                 |     |                                          
                 |     +-------------- Hyriopsis_myersiana        
                 |     |                                          
                 |     +-------------- Hyriopsis_panhai           
                 |     |                                          
                 |  +--+           +-- Hyriopsis_bialata          
                 |  |  +-----------+                              
                 |  |  |           +-- Hyriopsis_khoratensis      
                 |  |  |                                          
                 |  |  |  +----------- Rectidens_sumatrensis      
                 |  |  |  |                                       
                 |  |  +--+        +-- Ensidens_telus             
                 |  |     |  +-----+                              
                 |  |     |  |     +-- Ensidens_ingallsianus      
                 |  |     +--+                                    
                 |  |        |  +----- Ensidens_sagittarius       
                 |  |        +--+                                 
                 +--+           |  +-- Ensidens_spiculus          
                 |  |           +--+                              
                 |  |              +-- Ensidens_jaculus           
                 |  |                                             
                 |  |     +----------- Physunio_modelli           
                 |  |     |                                       
                 |  |     +----------- Lens                       
                 |  |     |                                       
                 |  |     +----------- Physunio_superbus          
                 |  |     |                                       
                 |  |     |  +-------- Yaukthwa_inlenensis        
                 |  |     +--+                                    
                 |  +-----+  |  +----- Yaukthwa_panhai            
                 |        |  +--+                                 
                 |        |     |  +-- Yaukthwa_nesemanni         
                 |        |     +--+                              
                 |        |        +-- Yaukthwa_rectangularis     
                 |        |                                       
                 |        |        +-- Lens_comptus               
                 |        |  +-----+                              
                 |        |  |     +-- Lens_novoselovi            
                 |        +--+                                    
                 |           |  +----- Lens_eximius               
                 |           +--+                                 
                 |              |  +-- Lens_contradens            
                 |              +--+                              
                 |                 +-- Lens_micropterus           
                 |                                                
                 |  +----------------- Acuticosta_chinensis       
                 |  |                                             
                 |  +----------------- Alasmidonta_marginata      
                 |  |                                             
                 |  +----------------- Anodonta_anatina           
                 |  |                                             
                 |  +----------------- Anemina_arcaeformis        
                 |  |                                             
                 |  +----------------- Anodonta_cygnea            
                 |  |                                             
                 |  +----------------- Cristaria_plicata          
                 |  |                                             
                 |  +----------------- Anodonta_nuttalliana       
                 |  |                                             
                 |  +----------------- Lasmigona_costata          
                 |  |                                             
                 |  +----------------- Lepidodesma_languilati     
                 |  |                                             
                 |  +----------------- Pseudanodonta_complanata   
                 |  |                                             
                 |  +----------------- Pyganodon_grandis          
                 |  |                                             
                 |  +----------------- Simpsonaias_ambigua        
                 |  |                                             
                 |  +----------------- Strophitus_undulatus       
                 |  |                                             
                 |  |              +-- Schistodesmus_lampreyanus  
                 |  +--------------+                              
                 |  |              +-- Schistodesmus_spinosus     
                 |  |                                             
                 |  |              +-- Utterbackia_imbecillis     
                 |  +--------------+                              
                 |  |              +-- Utterbackia_peninsularis   
                 |  |                                             
                 |  |        +-------- Lanceolaria_gladiola       
                 |  +--------+                                    
                 |  |        |  +----- Lanceolaria_grayii         
                 |  |        +--+                                 
                 +--+           |  +-- Lanceolaria_lanceolata     
                    |           +--+                              
                    |              +-- Lanceolaria_triformis      
                    |                                             
                    |        +-------- Unio_tumidus               
                    +--------+                                    
                    |        |  +----- Unio_crassus               
                    |        +--+                                 
                    |           |  +-- Unio_delphinus             
                    |           +--+                              
                    |              +-- Unio_pictorum              
                    |                                             
                    |        +-------- Aculamprotula_tortuosa     
                    |        |                                    
                    +--------+     +-- Aculamprotula_coreana      
                    |        |  +--+                              
                    |        |  |  +-- Aculamprotula_scripta      
                    |        +--+                                 
                    |           |  +-- Aculamprotula_fibrosa      
                    |           +--+                              
                    |              +-- Aculamprotula_tientsinensis
                    |                                             
                    |        +-------- Cuneopsis_heudei           
                    |        |                                    
                    |        +-------- Cuneopsis_celtiformis      
                    |        |                                    
                    +--------+-------- Tchangsinaia_piscicula     
                    |        |                                    
                    |        +-------- Cuneopsis_rufescens        
                    |        |                                    
                    |        |  +----- Nodularia_douglasiae[2]    
                    |        +--+                                 
                    |           |  +-- Nodularia_douglasiae[1]    
                    |           +--+                              
                    |              +-- Nodularia_nuxpersicae      
                    |                                             
                    |  +-------------- Pletholophus_tenuis        
                    |  |                                          
                    +--+           +-- Sinanodonta_angula         
                       |  +--------+                              
                       |  |        +-- Sinanodonta_lucida         
                       +--+                                       
                          |  +-------- Sinanodonta_woodiana[1]    
                          +--+                                    
                             |  +----- Sinanodonta_jourdyi        
                             +--+                                 
                                |  +-- Sinanodonta_elliptica      
                                +--+                              
                                   +-- Sinanodonta_woodiana[2]    

Walker et al., 2006 [Publ 2666], fig. 6

iT tT bi riS oT roT roS iD
9 9 7 3 29 28 2 11.0
           +----------------- Castalia_stevensi          
           |                                             
           +----------------- Diplodon_gratus            
  +--------+                                             
  |        |  +-------------- Hyridella_depressa         
  |        |  |                                          
  |        |  +-------------- Hyridella_australis        
  |        +--+                                          
  |           |  +----------- Echyridella_menziesii      
  |           |  |                                       
  |           |  |        +-- Velesunio_ambiguus         
  |           +--+  +-----+                              
  |              |  |     +-- Velesunio[3]               
  |              |  |                                    
  |              +--+  +----- Velesunio[2]               
  |                 |  |                                 
  |                 |  |  +-- Alathyria_jacksoni         
  |                 +--+--+                              
  |                    |  +-- Velesunio[1]               
  |                    |                                 
--+                    |  +-- Velesunio_angasi           
  |                    +--+                              
  |                       +-- Lortiella_rugata           
  |                                                      
  |                       +-- Mutela_dubia **               
  |        +--------------+                              
  |        |              +-- Mutela_rostrata **            
  |  +-----+                                             
  |  |     |  +-------------- Etheria_elliptica **          
  |  |     |  |                                          
  |  |     +--+  +----------- Chambardia_rubens **          
  |  |        |  |                                       
  |  |        +--+  +-------- Mulleria_rivoli **            
  |  |           |  |                                    
  |  |           +--+     +-- Anodontites_guanarensis **    
  |  |              |  +--+                              
  |  |              |  |  +-- Anodontites_trigona **        
  +--+              +--+                                 
     |                 |  +-- Tamsiella_tamsiana **         
     |                 +--+                              
     |                    +-- Monocondylaea_minuana **      
     |                                                   
     |                    +-- Pyganodon_fragilis         
     |  +-----------------+                              
     |  |                 +-- Pyganodon_grandis          
     |  |                                                
     |  |        +----------- Cumberlandia_monodonta     
     |  |        |                                       
     +--+  +-----+  +-------- Pseudunio_auricularius     
        |  |     |  |                                    
        |  |     +--+     +-- Margaritifera_margaritifera
        |  |        |  +--+                              
        |  |        |  |  +-- Margaritifera_dahurica     
        |  |        +--+                                 
        +--+           |  +-- Margaritifera_falcata      
           |           +--+                              
           |              +-- Margaritifera_middendorffi 
           |                                             
           |              +-- Gonidea_angulata           
           |  +-----------+                              
           |  |           +-- Sundadontina_tumida        
           +--+                                          
              |  +----------- Cyrtonaias_tampicoensis    
              |  |                                       
              +--+----------- Fusconaia_flava            
                 |                                       
                 |  +-------- Potamilus_purpuratus       
                 +--+                                    
                    |  +----- Lampsilis_teres            
                    +--+                                 
                       |  +-- Ligumia_recta              
                       +--+                              
                          +-- Ortmanniana_ligamentina    

Bogan & Hoeh, 2000 [Publ 2650], fig. 1

iT tT bi riS oT roT roS iD
8 28 21 3 26 25 2 29.0
                             +-- Castalia_stevensi          
                       +-----+                              
                       |     +-- Diplodon_gratus            
  +--------------------+                                    
  |                    |  +----- Echyridella_menziesii      
  |                    +--+                                 
  |                       |  +-- Velesunio_angasi           
  |                       +--+                              
  |                          +-- Lortiella_rugata           
  |                                                         
  |                          +-- Mutela_dubia **               
--+              +-----------+                              
  |              |           +-- Mutela_rostrata **            
  |  +-----------+                                          
  |  |           |  +----------- Etheria_elliptica **          
  |  |           +--+                                       
  |  |              |  +-------- Mulleria_rivoli **            
  |  |              +--+                                    
  |  |                 |  +----- Monocondylaea_minuana **      
  |  |                 +--+                                 
  +--+                    |  +-- Anodontites_trigona **        
     |                    +--+                              
     |                       +-- Anodontites_guanarensis **    
     |                                                      
     |  +----------------------- Coelatura_aegyptiaca       
     |  |                                                   
     |  |  +-------------------- Pseudomulleria_dalyi **       
     +--+  |                                                
        |  |              +----- Anodonta_cygnea            
        |  |  +-----------+                                 
        +--+  |           |  +-- Strophitus_undulatus       
           |  |           +--+                              
           |  |              +-- Pyganodon_grandis          
           |  |                                             
           +--+  +-------------- Gonidea_angulata           
              |  |                                          
              |  |           +-- Unio_pictorum              
              |  +-----------+                              
              |  |           +-- Unio_tumidus               
              |  |                                          
              +--+           +-- Cumberlandia_monodonta     
                 |  +--------+                              
                 |  |        +-- Margaritifera_margaritifera
                 |  |                                       
                 |  |  +-------- Toxolasma_lividum          
                 |  |  |                                    
                 +--+  +-------- Amblema_plicata            
                    |  |                                    
                    |  |     +-- Glebula_rotundata          
                    |  +-----+                              
                    |  |     +-- Cyrtonaias_tampicoensis    
                    |  |                                    
                    +--+  +----- Potamilus_alatus           
                       +--+                                 
                       |  |  +-- Ortmanniana_ligamentina    
                       |  +--+                              
                       |     +-- Ligumia_recta              
                       |                                    
                       |  +----- Quadrula_quadrula          
                       |  |                                 
                       +--+  +-- Eurynia_dilatata           
                          |  |                              
                          +--+-- Pleurobema_clava           
                             |                              
                             +-- Fusconaia_flava            

Guerra et al., 2019 [Publ 4168], fig. 1

iT tT bi riS oT roT roS iD
7 7 5 2 34 33 2 9.0
                 +----------------- Mutela_dubia **               
  +--------------+                                             
  |              |  +-------------- Chambardia_rubens **          
  |              +--+                                          
  |                 |  +----------- Anodontites_trapesialis **    
  |                 +--+                                       
  |                    |  +-------- Monocondylaea_parchappii **   
  |                    +--+                                    
  |                       |  +----- Lamproscapha_ensiformis **    
  |                       +--+                                 
  |                          |  +-- Tamsiella_schroeteriana **    
  |                          +--+                              
  |                             +-- Fossula_fossiculifera **      
--+                                                            
  |                             +-- Echyridella_menziesii      
  |                       +-----+                              
  |                       |     +-- Westralunio_carteri        
  |  +--------------------+                                    
  |  |                    |  +----- Diplodon_suavidicus        
  |  |                    +--+                                 
  |  |                       |  +-- Castalia_ambigua           
  |  |                       +--+                              
  |  |                          +-- Prisodon_obliquus          
  |  |                                                         
  |  |              +-------------- Gibbosula_crassa           
  |  |              |                                          
  +--+  +-----------+  +----------- Cumberlandia_monodonta     
     |  |           |  |                                       
     |  |           +--+        +-- Pseudunio_marocanus        
     |  |              |  +-----+                              
     |  |              |  |     +-- Pseudunio_auricularius     
     |  |              +--+                                    
     |  |                 |  +----- Margaritifera_falcata      
     |  |                 +--+                                 
     |  |                    |  +-- Margaritifera_dahurica     
     |  |                    +--+                              
     +--+                       +-- Margaritifera_margaritifera
        |                                                      
        |                 +-------- Quadrula_quadrula          
        |  +--------------+                                    
        |  |              |  +----- Toxolasma_parvum           
        |  |              +--+                                 
        |  |                 |  +-- Venustaconcha_ellipsiformis
        |  |                 +--+                              
        |  |                    +-- Potamilus_alatus           
        |  |                                                   
        +--+              +-------- Sinosolenaia_carinata      
           |  +-----------+                                    
           |  |           |  +----- Pronodularia_japanensis    
           |  |           +--+                                 
           |  |              |  +-- Lamprotula_leaii           
           |  |              +--+                              
           |  |                 +-- Potomida_littoralis        
           |  |                                                
           +--+     +-------------- Lanceolaria_lanceolata     
              |  +--+                                          
              |  |  |  +----------- Aculamprotula_tortuosa     
              |  |  +--+                                       
              |  |     |  +-------- Schistodesmus_lampreyanus  
              |  |     +--+                                    
              |  |        |  +----- Unio_tumidus               
              +--+        +--+                                 
                 |           |  +-- Unio_crassus               
                 |           +--+                              
                 |              +-- Unio_delphinus             
                 |                                             
                 |  +-------------- Sinanodonta_woodiana       
                 |  |                                          
                 +--+  +----------- Anodonta_anatina           
                    |  |                                       
                    +--+        +-- Platynaias_compressa       
                       |  +-----+                              
                       |  |     +-- Platynaias_subviridis      
                       +--+                                    
                          |  +----- Pyganodon_grandis          
                          +--+                                 
                             |  +-- Utterbackia_peninsularis   
                             +--+                              
                                +-- Utterbackia_imbecillis     

Guerra et al., 2019 [Publ 4168], fig. 2

iT tT bi riS oT roT roS iD
7 7 5 2 34 33 2 11.0
                       +----------------- Chambardia_rubens **          
  +--------------------+                                             
  |                    |  +-------------- Mutela_dubia **               
  |                    +--+                                          
  |                       |  +----------- Anodontites_trapesialis **    
  |                       +--+                                       
  |                          |  +-------- Monocondylaea_parchappii **   
  |                          +--+                                    
  |                             |  +----- Lamproscapha_ensiformis **    
  |                             +--+                                 
  |                                |  +-- Tamsiella_schroeteriana **    
  |                                +--+                              
  |                                   +-- Fossula_fossiculifera **      
--+                                                                  
  |                          +----------- Westralunio_carteri        
  |  +-----------------------+                                       
  |  |                       |  +-------- Echyridella_menziesii      
  |  |                       +--+                                    
  |  |                          |  +----- Diplodon_suavidicus        
  |  |                          +--+                                 
  |  |                             |  +-- Castalia_ambigua           
  |  |                             +--+                              
  |  |                                +-- Prisodon_obliquus          
  |  |                                                               
  |  |                    +-------------- Gibbosula_crassa           
  +--+                    |                                          
     |  +-----------------+  +----------- Cumberlandia_monodonta     
     |  |                 |  |                                       
     |  |                 +--+        +-- Pseudunio_marocanus        
     |  |                    |  +-----+                              
     |  |                    |  |     +-- Pseudunio_auricularius     
     |  |                    +--+                                    
     |  |                       |  +----- Margaritifera_falcata      
     |  |                       +--+                                 
     |  |                          |  +-- Margaritifera_dahurica     
     +--+                          +--+                              
        |                             +-- Margaritifera_margaritifera
        |                                                            
        |                       +-------- Quadrula_quadrula          
        |  +--------------------+                                    
        |  |                    |  +----- Toxolasma_parvum           
        |  |                    +--+                                 
        |  |                       |  +-- Venustaconcha_ellipsiformis
        |  |                       +--+                              
        |  |                          +-- Potamilus_alatus           
        +--+                                                         
           |                    +-------- Sinosolenaia_carinata      
           |  +-----------------+                                    
           |  |                 |  +----- Pronodularia_japanensis    
           |  |                 +--+                                 
           |  |                    |  +-- Lamprotula_leaii           
           |  |                    +--+                              
           +--+                       +-- Potomida_littoralis        
              |                                                      
              |  +----------------------- Aculamprotula_tortuosa     
              |  |                                                   
              |  |              +-------- Schistodesmus_lampreyanus  
              +--+  +-----------+                                    
                 |  |           |  +----- Unio_tumidus               
                 |  |           +--+                                 
                 |  |              |  +-- Unio_crassus               
                 +--+              +--+                              
                    |                 +-- Unio_delphinus             
                    |                                                
                    |  +----------------- Lanceolaria_lanceolata     
                    |  |                                             
                    +--+  +-------------- Sinanodonta_woodiana       
                       |  |                                          
                       +--+  +----------- Anodonta_anatina           
                          |  |                                       
                          +--+        +-- Platynaias_compressa       
                             |  +-----+                              
                             |  |     +-- Platynaias_subviridis      
                             +--+                                    
                                |  +----- Pyganodon_grandis          
                                +--+                                 
                                   |  +-- Utterbackia_peninsularis   
                                   +--+                              
                                      +-- Utterbackia_imbecillis     

Graf, 2000 [Publ 2647], fig. 1

iT tT bi riS oT roT roS iD
6 6 3 3 12 11 2 7.5
  +-------------- Grandidieria
  |                           
  |     +-------- Castalia    
  |  +--+                     
--+  |  |  +----- Diplodon    
  |  |  +--+                  
  |  |     |  +-- Hyridella   
  |  |     +--+               
  +--+        +-- Velesunio   
     |                        
     |        +-- Mutela[1] **   
     |  +-----+               
     |  |     +-- Mutela[2] **   
     +--+                     
        |  +----- Anodontites ** 
        |  |                  
        +--+----- Mycetopoda **  
           |                  
           |  +-- Etheria **     
           +--+               
              +-- Mulleria **    

Combosch et al., 2017 [Publ 3806], fig. 1

iT tT bi riS oT roT roS iD
6 6 4 3 9 7 2 7.0
              +----- Unio_pictorum              
  +-----------+                                 
  |           |  +-- Lampsilis_cardium          
  |           +--+                              
--+              +-- Scabies_crispata           
  |                                             
  |  +-------------- Margaritifera_margaritifera
  |  |                                          
  +--+           +-- Chambardia_wahlbergi **       
     |  +--------+                              
     |  |        +-- Aspatharia_pfeifferiana **    
     +--+                                       
        |  +-------- Etheria_elliptica **          
        +--+                                    
           |  +----- Mutela_hargeri **             
           +--+                                 
              |  +-- Tamsiella_schroeteriana **    
              +--+                              
                 +-- Lamproscapha_ensiformis **    

Combosch et al., 2017 [Publ 3806], fig. 2

iT tT bi riS oT roT roS iD
6 6 4 3 9 7 2 7.0
              +-- Chambardia_wahlbergi **       
     +--------+                              
     |        +-- Aspatharia_pfeifferiana **    
  +--+                                       
  |  |  +-------- Etheria_elliptica **          
  |  +--+                                    
  |     |  +----- Mutela_hargeri **             
  |     +--+                                 
  |        |  +-- Tamsiella_schroeteriana **    
--+        +--+                              
  |           +-- Lamproscapha_ensiformis **    
  |                                          
  |        +----- Velesunio_ambiguus         
  |  +-----+                                 
  |  |     |  +-- Triplodon_corrugatus       
  |  |     +--+                              
  +--+        +-- Hyridella_australis        
     |                                       
     |  +-------- Margaritifera_margaritifera
     +--+                                    
        |  +----- Unio_pictorum              
        +--+                                 
           |  +-- Scabies_crispata           
           +--+                              
              +-- Lampsilis_cardium          

Huang et al., 2018 [Publ 3828], fig. 3, 5

iT tT bi riS oT roT roS iD
6 6 4 3 23 21 2 3.0
                          +----- Triplodon_corrugatus       
  +-----------------------+                                 
  |                       |  +-- Velesunio_ambiguus         
  |                       +--+                              
  |                          +-- Hyridella_australis        
  |                                                         
  |                 +----------- Mutela_hargeri **             
  |                 |                                       
  |  +--------------+        +-- Aspatharia_pfeifferiana **    
--+  |              |  +-----+                              
  |  |              |  |     +-- Chambardia_wahlbergi **       
  |  |              +--+                                    
  |  |                 |  +----- Etheria_elliptica **          
  |  |                 +--+                                 
  |  |                    |  +-- Tamsiella_schroeteriana **    
  |  |                    +--+                              
  |  |                       +-- Lamproscapha_ensiformis **    
  |  |                                                      
  +--+                       +-- Potomida_littoralis        
     |                 +-----+                              
     |                 |     +-- Lamprotula_leaii           
     |                 |                                    
     |  +--------------+     +-- Lampsilis_cardium          
     |  |              |  +--+                              
     |  |              |  |  +-- Scabies_crispata           
     |  |              +--+                                 
     |  |                 |  +-- Unio_pictorum              
     |  |                 +--+                              
     +--+                    +-- Aculamprotula_tientsinensis
        |                                                   
        |                    +-- Gibbosula_rochechouartii   
        |  +-----------------+                              
        |  |                 +-- Gibbosula_laosensis        
        |  |                                                
        |  |              +----- Cumberlandia_monodonta     
        +--+  +-----------+                                 
           |  |           |  +-- Pseudunio_auricularius     
           |  |           +--+                              
           |  |              +-- Pseudunio_marocanus        
           +--+                                             
              |              +-- Margaritifera_margaritifera
              |  +-----------+                              
              |  |           +-- Margaritifera_dahurica     
              +--+                                          
                 |  +----------- Margaritifera_falcata      
                 +--+                                       
                    |  +-------- Margaritifera_laevis       
                    +--+                                    
                       |  +----- Margaritifera_hembeli      
                       +--+                                 
                          |  +-- Margaritifera_middendorffi 
                          +--+                              
                             +-- Margaritifera_marrianae    

Lopes-Lima et al., 2018 [Publ 3829], fig. 1

iT tT bi riS oT roT roS iD
6 6 4 3 22 20 2 3.0
                       +----- Triplodon_corrugatus       
  +--------------------+                                 
  |                    |  +-- Velesunio_ambiguus         
  |                    +--+                              
  |                       +-- Hyridella_australis        
  |                                                      
  |              +----------- Mutela_hargeri **             
  |              |                                       
--+  +-----------+        +-- Aspatharia_pfeifferiana **    
  |  |           |  +-----+                              
  |  |           |  |     +-- Chambardia_wahlbergi **       
  |  |           +--+                                    
  |  |              |  +----- Etheria_elliptica **          
  |  |              +--+                                 
  |  |                 |  +-- Lamproscapha_ensiformis **    
  |  |                 +--+                              
  +--+                    +-- Tamsiella_schroeteriana **    
     |                                                   
     |                 +----- Potomida_littoralis        
     |  +--------------+                                 
     |  |              |  +-- Lampsilis_cardium          
     |  |              +--+                              
     |  |                 +-- Unio_pictorum              
     |  |                                                
     +--+              +----- Gibbosula_rochechouartii   
        |  +-----------+                                 
        |  |           |  +-- Gibbosula_crassa           
        |  |           +--+                              
        |  |              +-- Gibbosula_laosensis        
        |  |                                             
        +--+        +-------- Cumberlandia_monodonta     
           |  +-----+                                    
           |  |     |  +----- Pseudunio_marocanus        
           |  |     +--+                                 
           |  |        |  +-- Pseudunio_auricularius     
           |  |        +--+                              
           +--+           +-- Pseudunio_homsensis        
              |                                          
              |           +-- Margaritifera_dahurica     
              |  +--------+                              
              |  |        +-- Margaritifera_margaritifera
              +--+                                       
                 |  +-------- Margaritifera_falcata      
                 |  |                                    
                 +--+     +-- Margaritifera_middendorffi 
                    |  +--+                              
                    |  |  +-- Margaritifera_laevis       
                    +--+                                 
                       |  +-- Margaritifera_hembeli      
                       +--+                              
                          +-- Margaritifera_marrianae    

Lopes-Lima et al., 2018 [Publ 3829], fig. 3

iT tT bi riS oT roT roS iD
6 6 4 3 22 20 2 3.0
                          +----- Triplodon_corrugatus       
  +-----------------------+                                 
  |                       |  +-- Velesunio_ambiguus         
  |                       +--+                              
  |                          +-- Hyridella_australis        
  |                                                         
  |                 +----------- Mutela_hargeri **             
  |                 |                                       
--+  +--------------+        +-- Chambardia_wahlbergi **       
  |  |              |  +-----+                              
  |  |              |  |     +-- Aspatharia_pfeifferiana **    
  |  |              +--+                                    
  |  |                 |  +----- Etheria_elliptica **          
  |  |                 +--+                                 
  |  |                    |  +-- Tamsiella_schroeteriana **    
  |  |                    +--+                              
  +--+                       +-- Lamproscapha_ensiformis **    
     |                                                      
     |                    +----- Potomida_littoralis        
     |  +-----------------+                                 
     |  |                 |  +-- Lampsilis_cardium          
     |  |                 +--+                              
     |  |                    +-- Unio_pictorum              
     |  |                                                   
     +--+                 +----- Gibbosula_rochechouartii   
        |  +--------------+                                 
        |  |              |  +-- Gibbosula_laosensis        
        |  |              +--+                              
        |  |                 +-- Gibbosula_crassa           
        |  |                                                
        +--+           +-------- Cumberlandia_monodonta     
           |  +--------+                                    
           |  |        |  +----- Pseudunio_marocanus        
           |  |        +--+                                 
           |  |           |  +-- Pseudunio_auricularius     
           |  |           +--+                              
           +--+              +-- Pseudunio_homsensis        
              |                                             
              |              +-- Margaritifera_margaritifera
              |  +-----------+                              
              |  |           +-- Margaritifera_dahurica     
              +--+                                          
                 |  +----------- Margaritifera_falcata      
                 +--+                                       
                    |  +-------- Margaritifera_laevis       
                    +--+                                    
                       |  +----- Margaritifera_middendorffi 
                       +--+                                 
                          |  +-- Margaritifera_hembeli      
                          +--+                              
                             +-- Margaritifera_marrianae    

Wu et al., 2020 [Publ 4121], fig. 2

iT tT bi riS oT roT roS iD
6 6 4 3 28 26 2 3.0
                       +----- Triplodon_corrugatus       
  +--------------------+                                 
  |                    |  +-- Hyridella_australis        
  |                    +--+                              
  |                       +-- Velesunio_ambiguus         
  |                                                      
  |              +----------- Mutela_hargeri **             
  |              |                                       
  |  +-----------+        +-- Aspatharia_pfeifferiana **    
  |  |           |  +-----+                              
--+  |           |  |     +-- Chambardia_wahlbergi **       
  |  |           +--+                                    
  |  |              |  +----- Etheria_elliptica **          
  |  |              +--+                                 
  |  |                 |  +-- Tamsiella_schroeteriana **    
  |  |                 +--+                              
  |  |                    +-- Lamproscapha_ensiformis **    
  |  |                                                   
  |  |                    +-- Lampsilis_cardium          
  |  |     +--------------+                              
  +--+     |              +-- Scabies_crispata           
     |     |                                             
     |     |        +-------- Sinosolenaia_carinata      
     |  +--+  +-----+                                    
     |  |  |  |     |  +----- Potomida_littoralis        
     |  |  |  |     +--+                                 
     |  |  |  |        |  +-- Lamprotula_caveata         
     |  |  +--+        +--+                              
     |  |     |           +-- Lamprotula_leaii           
     |  |     |                                          
     |  |     |  +----------- Unio_pictorum              
     |  |     +--+                                       
     |  |        |  +-------- Aculamprotula_tortuosa     
     +--+        +--+                                    
        |           |  +----- Aculamprotula_polysticta   
        |           +--+                                 
        |              |  +-- Aculamprotula_fibrosa      
        |              +--+                              
        |                 +-- Aculamprotula_tientsinensis
        |                                                
        |  +----------------- Cumberlandia_monodonta     
        |  |                                             
        |  |              +-- Gibbosula_laosensis        
        |  |           +--+                              
        +--+           |  +-- Gibbosula_rochechouartii   
           |  +--------+                                 
           |  |        |  +-- Pseudunio_auricularius     
           |  |        +--+                              
           |  |           +-- Pseudunio_marocanus        
           +--+                                          
              |           +-- Margaritifera_dahurica     
              |  +--------+                              
              |  |        +-- Margaritifera_margaritifera
              +--+                                       
                 |  +-------- Margaritifera_falcata      
                 |  |                                    
                 +--+     +-- Margaritifera_hembeli      
                    |  +--+                              
                    |  |  +-- Margaritifera_marrianae    
                    +--+                                 
                       |  +-- Margaritifera_laevis       
                       +--+                              
                          +-- Margaritifera_middendorffi 

Wu et al., 2020 [Publ 4121], fig. S1

iT tT bi riS oT roT roS iD
6 6 4 3 28 26 2 3.0
                          +----- Triplodon_corrugatus       
  +-----------------------+                                 
  |                       |  +-- Hyridella_australis        
  |                       +--+                              
  |                          +-- Velesunio_ambiguus         
  |                                                         
  |                 +----------- Mutela_hargeri **             
  |                 |                                       
  |  +--------------+        +-- Aspatharia_pfeifferiana **    
  |  |              |  +-----+                              
--+  |              |  |     +-- Chambardia_wahlbergi **       
  |  |              +--+                                    
  |  |                 |  +----- Etheria_elliptica **          
  |  |                 +--+                                 
  |  |                    |  +-- Tamsiella_schroeteriana **    
  |  |                    +--+                              
  |  |                       +-- Lamproscapha_ensiformis **    
  |  |                                                      
  |  |                       +-- Lampsilis_cardium          
  |  |        +--------------+                              
  +--+        |              +-- Scabies_crispata           
     |        |                                             
     |        |        +-------- Sinosolenaia_carinata      
     |  +-----+  +-----+                                    
     |  |     |  |     |  +----- Potomida_littoralis        
     |  |     |  |     +--+                                 
     |  |     |  |        |  +-- Lamprotula_caveata         
     |  |     +--+        +--+                              
     |  |        |           +-- Lamprotula_leaii           
     |  |        |                                          
     |  |        |  +----------- Unio_pictorum              
     |  |        +--+                                       
     +--+           |  +-------- Aculamprotula_tortuosa     
        |           +--+                                    
        |              |  +----- Aculamprotula_polysticta   
        |              +--+                                 
        |                 |  +-- Aculamprotula_tientsinensis
        |                 +--+                              
        |                    +-- Aculamprotula_fibrosa      
        |                                                   
        |  +-------------------- Gibbosula_rochechouartii   
        |  |                                                
        +--+  +----------------- Gibbosula_laosensis        
           |  |                                             
           |  |           +----- Cumberlandia_monodonta     
           +--+  +--------+                                 
              |  |        |  +-- Pseudunio_auricularius     
              |  |        +--+                              
              |  |           +-- Pseudunio_marocanus        
              +--+                                          
                 |           +-- Margaritifera_dahurica     
                 |  +--------+                              
                 |  |        +-- Margaritifera_margaritifera
                 +--+                                       
                    |  +-------- Margaritifera_falcata      
                    |  |                                    
                    +--+     +-- Margaritifera_hembeli      
                       |  +--+                              
                       |  |  +-- Margaritifera_marrianae    
                       +--+                                 
                          |  +-- Margaritifera_laevis       
                          +--+                              
                             +-- Margaritifera_middendorffi 

Sano et al., 2020 [Publ 4138], fig. 2

iT tT bi riS oT roT roS iD
6 6 4 3 67 65 2 3.0
                                               +----- Triplodon_corrugatus       
  +--------------------------------------------+                                 
  |                                            |  +-- Hyridella_australis        
  |                                            +--+                              
  |                                               +-- Velesunio_ambiguus         
  |                                                                              
  |                                      +----------- Mutela_hargeri **             
  |                                      |                                       
  |  +-----------------------------------+        +-- Aspatharia_pfeifferiana **    
  |  |                                   |  +-----+                              
  |  |                                   |  |     +-- Chambardia_wahlbergi **       
--+  |                                   +--+                                    
  |  |                                      |  +----- Etheria_elliptica **          
  |  |                                      +--+                                 
  |  |                                         |  +-- Tamsiella_schroeteriana **    
  |  |                                         +--+                              
  |  |                                            +-- Lamproscapha_ensiformis **    
  |  |                                                                           
  |  |                          +-------------------- Cumberlandia_monodonta     
  |  |                          |                                                
  |  |                          |                 +-- Gibbosula_rochechouartii   
  |  |                          |              +--+                              
  +--+  +-----------------------+              |  +-- Gibbosula_laosensis        
     |  |                       |  +-----------+                                 
     |  |                       |  |           |  +-- Pseudunio_auricularius     
     |  |                       |  |           +--+                              
     |  |                       |  |              +-- Pseudunio_marocanus        
     |  |                       +--+                                             
     |  |                          |              +-- Margaritifera_dahurica     
     |  |                          |  +-----------+                              
     |  |                          |  |           +-- Margaritifera_margaritifera
     |  |                          +--+                                          
     |  |                             |  +----------- Margaritifera_falcata      
     |  |                             +--+                                       
     |  |                                |  +-------- Margaritifera_middendorffi 
     |  |                                +--+                                    
     +--+                                   |  +----- Margaritifera_laevis       
        |                                   +--+                                 
        |                                      |  +-- Margaritifera_hembeli      
        |                                      +--+                              
        |                                         +-- Margaritifera_marrianae    
        |                                                                        
        |  +----------------------------------------- Scabies_crispata           
        |  |                                                                     
        |  |                       +----------------- Lampsilis_cardium          
        |  |                       |                                             
        |  |                       |           +----- Sinohyriopsis_cumingii[2]  
        |  |  +--------------------+  +--------+                                 
        |  |  |                    |  |        |  +-- Sinohyriopsis_schlegelii   
        |  |  |                    |  |        +--+                              
        |  |  |                    +--+           +-- Sinohyriopsis_cumingii[1]  
        +--+  |                       |                                          
           |  |                       |  +----------- Inversidens_brandtii       
           |  |                       |  |                                       
           |  |                       +--+        +-- Ptychorhynchus_pfisteri    
           |  |                          |  +-----+                              
           |  |                          |  |     +-- Obovalis_omiensis          
           |  |                          +--+                                    
           |  |                             |  +----- Lamprotula_leaii           
           |  |                             +--+                                 
           +--+                                |  +-- Pronodularia_japanensis    
              |                                +--+                              
              |                                   +-- Potomida_littoralis        
              |                                                                  
              |                                +----- Buldowskia_iwakawai        
              |  +-----------------------------+                                 
              |  |                             |  +-- Anemina_arcaeformis        
              |  |                             +--+                              
              |  |                                +-- Anemina_euscaphys          
              |  |                                                               
              |  |                    +-------------- Pletholophus_tenuis        
              |  |                    |                                          
              |  |                    |        +----- Beringiana_fukuharai       
              |  |  +-----------------+  +-----+                                 
              +--+  |                 |  |     |  +-- Beringiana_japonica        
                 |  |                 |  |     +--+                              
                 |  |                 +--+        +-- Beringiana_beringiana      
                 |  |                    |                                       
                 |  |                    |  +-------- Sinanodonta_lucida         
                 |  |                    +--+                                    
                 |  |                       |  +----- Sinanodonta_tumens         
                 |  |                       +--+                                 
                 |  |                          |  +-- Sinanodonta_lauta          
                 +--+                          +--+                              
                    |                             +-- Sinanodonta_woodiana       
                    |                                                            
                    |  +----------------------------- Cristaria_plicata          
                    |  |                                                         
                    |  |                          +-- Anodonta_cygnea            
                    |  |                 +--------+                              
                    |  |                 |        +-- Anodonta_anatina           
                    |  |                 |                                       
                    |  |  +--------------+     +----- Alasmidonta_varicosa       
                    +--+  |              |  +--+                                 
                       |  |              |  |  |  +-- Prolasmidonta_heterodon    
                       |  |              |  |  +--+                              
                       |  |              +--+     +-- Platynaias_compressa       
                       |  |                 |                                    
                       |  |                 |  +----- Pyganodon_grandis          
                       |  |                 +--+                                 
                       +--+                    |  +-- Utterbackia_imbecillis     
                          |                    +--+                              
                          |                       +-- Utterbackia_peninsularis   
                          |                                                      
                          |                    +----- Lanceolaria_grayii[1]      
                          |  +-----------------+                                 
                          |  |                 |  +-- Lanceolaria_lanceolata     
                          |  |                 +--+                              
                          |  |                    +-- Lanceolaria_grayii[2]      
                          +--+                                                   
                             |  +-------------------- Lepidodesma_languilati     
                             |  |                                                
                             |  |              +----- Aculamprotula_tortuosa     
                             |  |  +-----------+                                 
                             +--+  |           |  +-- Aculamprotula_tientsinensis
                                |  |           +--+                              
                                |  |              +-- Aculamprotula_coreana      
                                |  |                                             
                                +--+           +----- Unio_crassus               
                                   |  +--------+                                 
                                   |  |        |  +-- Unio_delphinus             
                                   |  |        +--+                              
                                   |  |           +-- Unio_pictorum              
                                   |  |                                          
                                   +--+     +-------- Inversiunio_jokohamensis   
                                      |  +--+                                    
                                      |  |  |  +----- Inversiunio_yanagawensis[2]
                                      |  |  +--+                                 
                                      |  |     |  +-- Inversiunio_reinianus      
                                      |  |     +--+                              
                                      +--+        +-- Inversiunio_yanagawensis[1]
                                         |                                       
                                         |        +-- Tchangsinaia_piscicula     
                                         |  +-----+                              
                                         |  |     +-- Schistodesmus_lampreyanus  
                                         +--+                                    
                                            |  +----- Nodularia_douglasiae[2]    
                                            +--+                                 
                                               |  +-- Nodularia_nipponensis      
                                               +--+                              
                                                  +-- Nodularia_douglasiae[1]    

Sano et al., 2020 [Publ 4138], fig. 3

iT tT bi riS oT roT roS iD
6 6 4 3 66 64 2 3.0
                                               +----- Triplodon_corrugatus       
  +--------------------------------------------+                                 
  |                                            |  +-- Hyridella_australis        
  |                                            +--+                              
  |                                               +-- Velesunio_ambiguus         
  |                                                                              
  |                                      +----------- Mutela_hargeri **             
  |                                      |                                       
  |  +-----------------------------------+        +-- Aspatharia_pfeifferiana **    
  |  |                                   |  +-----+                              
  |  |                                   |  |     +-- Chambardia_wahlbergi **       
--+  |                                   +--+                                    
  |  |                                      |  +----- Etheria_elliptica **          
  |  |                                      +--+                                 
  |  |                                         |  +-- Tamsiella_schroeteriana **    
  |  |                                         +--+                              
  |  |                                            +-- Lamproscapha_ensiformis **    
  |  |                                                                           
  |  |                       +----------------------- Cumberlandia_monodonta     
  |  |                       |                                                   
  |  |                       |                    +-- Pseudunio_auricularius     
  |  |  +--------------------+  +-----------------+                              
  +--+  |                    |  |                 +-- Pseudunio_marocanus        
     |  |                    |  |                                                
     |  |                    +--+                 +-- Gibbosula_laosensis        
     |  |                       |  +--------------+                              
     |  |                       |  |              +-- Gibbosula_rochechouartii   
     |  |                       |  |                                             
     |  |                       +--+              +-- Margaritifera_margaritifera
     |  |                          |  +-----------+                              
     |  |                          |  |           +-- Margaritifera_dahurica     
     |  |                          +--+                                          
     |  |                             |  +----------- Margaritifera_falcata      
     |  |                             +--+                                       
     |  |                                |  +-------- Margaritifera_middendorffi 
     |  |                                +--+                                    
     +--+                                   |  +----- Margaritifera_laevis       
        |                                   +--+                                 
        |                                      |  +-- Margaritifera_hembeli      
        |                                      +--+                              
        |                                         +-- Margaritifera_marrianae    
        |                                                                        
        |  +----------------------------------------- Scabies_crispata           
        |  |                                                                     
        |  |                                   +----- Sinohyriopsis_cumingii[2]  
        |  |                          +--------+                                 
        |  |                          |        |  +-- Sinohyriopsis_schlegelii   
        |  |                          |        +--+                              
        |  |                          |           +-- Sinohyriopsis_cumingii[1]  
        |  |  +-----------------------+                                          
        |  |  |                       |  +----------- Lampsilis_cardium          
        +--+  |                       |  |                                       
           |  |                       |  |     +----- Lamprotula_leaii           
           |  |                       +--+  +--+                                 
           |  |                          |  |  |  +-- Potomida_littoralis        
           |  |                          |  |  +--+                              
           |  |                          +--+     +-- Pronodularia_japanensis    
           |  |                             |                                    
           |  |                             |  +----- Inversidens_brandtii       
           |  |                             +--+                                 
           |  |                                |  +-- Obovalis_omiensis          
           +--+                                +--+                              
              |                                   +-- Ptychorhynchus_pfisteri    
              |                                                                  
              |                                +----- Buldowskia_iwakawai        
              |  +-----------------------------+                                 
              |  |                             |  +-- Anemina_arcaeformis        
              |  |                             +--+                              
              |  |                                +-- Anemina_euscaphys          
              |  |                                                               
              |  |                          +-------- Pletholophus_tenuis        
              |  |                       +--+                                    
              |  |                       |  |  +----- Beringiana_fukuharai       
              |  |                       |  +--+                                 
              +--+                       |     |  +-- Beringiana_japonica        
                 |  +--------------------+     +--+                              
                 |  |                    |        +-- Beringiana_beringiana      
                 |  |                    |                                       
                 |  |                    |  +-------- Sinanodonta_lucida         
                 |  |                    +--+                                    
                 |  |                       |  +----- Sinanodonta_tumens         
                 |  |                       +--+                                 
                 |  |                          |  +-- Sinanodonta_woodiana       
                 |  |                          +--+                              
                 +--+                             +-- Sinanodonta_lauta          
                    |                                                            
                    |  +----------------------------- Cristaria_plicata          
                    |  |                                                         
                    |  |                          +-- Anodonta_anatina           
                    |  |                 +--------+                              
                    |  |                 |        +-- Anodonta_cygnea            
                    |  |                 |                                       
                    |  |  +--------------+     +----- Pyganodon_grandis          
                    +--+  |              |  +--+                                 
                       |  |              |  |  |  +-- Utterbackia_imbecillis     
                       |  |              |  |  +--+                              
                       |  |              +--+     +-- Utterbackia_peninsularis   
                       |  |                 |                                    
                       |  |                 |  +----- Alasmidonta_varicosa       
                       |  |                 +--+                                 
                       +--+                    |  +-- Platynaias_compressa       
                          |                    +--+                              
                          |                       +-- Prolasmidonta_heterodon    
                          |                                                      
                          |                    +----- Lanceolaria_grayii[1]      
                          |  +-----------------+                                 
                          |  |                 |  +-- Lanceolaria_lanceolata     
                          |  |                 +--+                              
                          |  |                    +-- Lanceolaria_grayii[2]      
                          +--+                                                   
                             |  +-------------------- Lepidodesma_languilati     
                             |  |                                                
                             |  |              +----- Aculamprotula_tortuosa     
                             |  |  +-----------+                                 
                             +--+  |           |  +-- Aculamprotula_tientsinensis
                                |  |           +--+                              
                                |  |              +-- Aculamprotula_coreana      
                                |  |                                             
                                +--+           +----- Unio_crassus               
                                   |  +--------+                                 
                                   |  |        |  +-- Unio_delphinus             
                                   |  |        +--+                              
                                   |  |           +-- Unio_pictorum              
                                   |  |                                          
                                   +--+     +-------- Inversiunio_jokohamensis   
                                      |  +--+                                    
                                      |  |  |  +----- Inversiunio_yanagawensis[2]
                                      |  |  +--+                                 
                                      |  |     |  +-- Inversiunio_reinianus      
                                      |  |     +--+                              
                                      +--+        +-- Inversiunio_yanagawensis[1]
                                         |                                       
                                         |        +-- Nodularia_nipponensis      
                                         |     +--+                              
                                         |     |  +-- Nodularia_douglasiae       
                                         +-----+                                 
                                               |  +-- Schistodesmus_lampreyanus  
                                               +--+                              
                                                  +-- Tchangsinaia_piscicula     

Sano et al., 2020 [Publ 4138], fig. 4

iT tT bi riS oT roT roS iD
6 6 4 3 66 64 2 3.0
                                                  +----- Triplodon_corrugatus       
  +-----------------------------------------------+                                 
  |                                               |  +-- Hyridella_australis        
  |                                               +--+                              
  |                                                  +-- Velesunio_ambiguus         
  |                                                                                 
  |                                         +----------- Mutela_hargeri **             
  |                                         |                                       
  |  +--------------------------------------+        +-- Aspatharia_pfeifferiana **    
  |  |                                      |  +-----+                              
  |  |                                      |  |     +-- Chambardia_wahlbergi **       
--+  |                                      +--+                                    
  |  |                                         |  +----- Etheria_elliptica **          
  |  |                                         +--+                                 
  |  |                                            |  +-- Tamsiella_schroeteriana **    
  |  |                                            +--+                              
  |  |                                               +-- Lamproscapha_ensiformis **    
  |  |                                                                              
  |  |                             +-------------------- Cumberlandia_monodonta     
  |  |                             |                                                
  |  |                             |                 +-- Pseudunio_auricularius     
  |  |  +--------------------------+  +--------------+                              
  +--+  |                          |  |              +-- Pseudunio_marocanus        
     |  |                          |  |                                             
     |  |                          +--+              +-- Gibbosula_rochechouartii   
     |  |                             |  +-----------+                              
     |  |                             |  |           +-- Gibbosula_laosensis        
     |  |                             |  |                                          
     |  |                             +--+           +-- Margaritifera_margaritifera
     |  |                                |  +--------+                              
     |  |                                |  |        +-- Margaritifera_dahurica     
     |  |                                +--+                                       
     |  |                                   |  +-------- Margaritifera_falcata      
     |  |                                   |  |                                    
     |  |                                   +--+     +-- Margaritifera_laevis       
     |  |                                      |  +--+                              
     +--+                                      |  |  +-- Margaritifera_middendorffi 
        |                                      +--+                                 
        |                                         |  +-- Margaritifera_hembeli      
        |                                         +--+                              
        |                                            +-- Margaritifera_marrianae    
        |                                                                           
        |  +-------------------------------------------- Scabies_crispata           
        |  |                                                                        
        |  |                          +----------------- Lampsilis_cardium          
        |  |                          |                                             
        |  |                          |           +----- Sinohyriopsis_cumingii[2]  
        |  |  +-----------------------+  +--------+                                 
        |  |  |                       |  |        |  +-- Sinohyriopsis_schlegelii   
        |  |  |                       |  |        +--+                              
        |  |  |                       +--+           +-- Sinohyriopsis_cumingii[1]  
        +--+  |                          |                                          
           |  |                          |  +----------- Inversidens_brandtii       
           |  |                          |  |                                       
           |  |                          +--+        +-- Obovalis_omiensis          
           |  |                             |  +-----+                              
           |  |                             |  |     +-- Ptychorhynchus_pfisteri    
           |  |                             +--+                                    
           |  |                                |  +----- Lamprotula_leaii           
           |  |                                +--+                                 
           +--+                                   |  +-- Pronodularia_japanensis    
              |                                   +--+                              
              |                                      +-- Potomida_littoralis        
              |                                                                     
              |                                   +----- Buldowskia_iwakawai        
              |  +--------------------------------+                                 
              |  |                                |  +-- Anemina_arcaeformis        
              |  |                                +--+                              
              |  |                                   +-- Anemina_euscaphys          
              |  |                                                                  
              |  |                             +-------- Sinanodonta_lucida         
              |  |                          +--+                                    
              |  |                          |  |  +----- Sinanodonta_tumens         
              |  |                          |  +--+                                 
              +--+                          |     |  +-- Sinanodonta_woodiana       
                 |  +-----------------------+     +--+                              
                 |  |                       |        +-- Sinanodonta_lauta          
                 |  |                       |                                       
                 |  |                       |  +-------- Pletholophus_tenuis        
                 |  |                       +--+                                    
                 |  |                          |  +----- Beringiana_fukuharai       
                 |  |                          +--+                                 
                 |  |                             |  +-- Beringiana_japonica        
                 |  |                             +--+                              
                 +--+                                +-- Beringiana_beringiana      
                    |                                                               
                    |  +-------------------------------- Cristaria_plicata          
                    |  |                                                            
                    |  |                             +-- Anodonta_cygnea            
                    |  |                    +--------+                              
                    |  |                    |        +-- Anodonta_anatina           
                    |  |                    |                                       
                    |  |  +-----------------+     +----- Alasmidonta_varicosa       
                    +--+  |                 |  +--+                                 
                       |  |                 |  |  |  +-- Prolasmidonta_heterodon    
                       |  |                 |  |  +--+                              
                       |  |                 +--+     +-- Platynaias_compressa       
                       |  |                    |                                    
                       |  |                    |  +----- Pyganodon_grandis          
                       |  |                    +--+                                 
                       +--+                       |  +-- Utterbackia_imbecillis     
                          |                       +--+                              
                          |                          +-- Utterbackia_peninsularis   
                          |                                                         
                          |  +-------------------------- Lepidodesma_languilati     
                          |  |                                                      
                          |  |                    +----- Lanceolaria_grayii[1]      
                          |  |  +-----------------+                                 
                          +--+  |                 |  +-- Lanceolaria_lanceolata     
                             |  |                 +--+                              
                             |  |                    +-- Lanceolaria_grayii[2]      
                             |  |                                                   
                             +--+                 +----- Aculamprotula_tortuosa     
                                |  +--------------+                                 
                                |  |              |  +-- Aculamprotula_tientsinensis
                                |  |              +--+                              
                                |  |                 +-- Aculamprotula_coreana      
                                |  |                                                
                                +--+              +----- Unio_crassus               
                                   |  +-----------+                                 
                                   |  |           |  +-- Unio_delphinus             
                                   |  |           +--+                              
                                   |  |              +-- Unio_pictorum              
                                   +--+                                             
                                      |              +-- Nodularia_douglasiae       
                                      |  +-----------+                              
                                      |  |           +-- Nodularia_nipponensis      
                                      |  |                                          
                                      +--+           +-- Schistodesmus_lampreyanus  
                                         |  +--------+                              
                                         |  |        +-- Tchangsinaia_piscicula     
                                         +--+                                       
                                            |  +-------- Inversiunio_jokohamensis   
                                            +--+                                    
                                               |  +----- Inversiunio_yanagawensis[2]
                                               +--+                                 
                                                  |  +-- Inversiunio_reinianus      
                                                  +--+                              
                                                     +-- Inversiunio_yanagawensis[1]

Hoeh et al., 2001 [Publ 2652], fig. 14.2

iT tT bi riS oT roT roS iD
5 5 2 2 26 25 2 8.0
        +----- Pyganodon_grandis      
  +-----+                             
  |     |  +-- Anodonta_cygnea        
  |     +--+                          
  |        +-- Strophitus_undulatus   
  |                                   
  |     +----- Diplodon_gratus        
--+     |                             
  |     +----- Echyridella_menziesii  
  |  +--+                             
  |  |  +----- Lortiella_rugata       
  |  |  |                             
  |  |  |  +-- Castalia_stevensi      
  |  |  +--+                          
  +--+     +-- Velesunio_angasi       
     |                                
     |     +-- Mutela_dubia **           
     |  +--+                          
     |  |  +-- Mutela_rostrata **        
     +--+                             
        |  +-- Anodontites_guanarensis **
        |  |                          
        +--+-- Anodontites_trigona **    
           |                          
           +-- Monocondylaea_minuana **  

Hoeh et al., 2001 [Publ 2652], fig. 14.4

iT tT bi riS oT roT roS iD
5 5 3 2 26 25 2 8.0
                                   +-- Castalia_stevensi          
                             +-----+                              
                             |     +-- Diplodon_gratus            
  +--------------------------+                                    
  |                          |  +----- Echyridella_menziesii      
  |                          +--+                                 
  |                             |  +-- Lortiella_rugata           
  |                             +--+                              
  |                                +-- Velesunio_angasi           
  |                                                               
--+                                +-- Mutela_dubia **               
  |                          +-----+                              
  |                          |     +-- Mutela_rostrata **            
  |  +-----------------------+                                    
  |  |                       |  +----- Monocondylaea_minuana **      
  |  |                       +--+                                 
  |  |                          |  +-- Anodontites_guanarensis **    
  |  |                          +--+                              
  +--+                             +-- Anodontites_trigona **        
     |                                                            
     |  +----------------------------- Coelatura_aegyptiaca       
     |  |                                                         
     |  |                          +-- Cumberlandia_monodonta     
     +--+  +-----------------------+                              
        |  |                       +-- Margaritifera_margaritifera
        |  |                                                      
        |  |  +----------------------- Gonidea_angulata           
        +--+  |                                                   
           |  |              +-------- Unio_pictorum              
           |  |              |                                    
           |  |  +-----------+-------- Unio_tumidus               
           +--+  |           |                                    
              |  |           |  +----- Anodonta_cygnea            
              |  |           +--+                                 
              |  |              |  +-- Strophitus_undulatus       
              |  |              +--+                              
              +--+                 +-- Pyganodon_grandis          
                 |                                                
                 |  +----------------- Amblema_plicata            
                 |  |                                             
                 |  |        +-------- Quadrula_quadrula          
                 |  |  +-----+                                    
                 +--+  |     |  +----- Eurynia_dilatata           
                    |  |     +--+                                 
                    |  |        |  +-- Fusconaia_flava            
                    |  |        +--+                              
                    +--+           +-- Pleurobema_clava           
                       |                                          
                       |           +-- Cyrtonaias_tampicoensis    
                       |  +--------+                              
                       |  |        +-- Glebula_rotundata          
                       +--+                                       
                          |  +-------- Toxolasma_lividum          
                          +--+                                    
                             |  +----- Potamilus_alatus           
                             +--+                                 
                                |  +-- Ligumia_recta              
                                +--+                              
                                   +-- Ortmanniana_ligamentina    

Hoeh et al., 2002 [Publ 2653], fig. 2

iT tT bi riS oT roT roS iD
5 5 3 2 26 25 2 8.0
                                +-- Castalia_stevensi          
                          +-----+                              
                          |     +-- Diplodon_gratus            
  +-----------------------+                                    
  |                       |  +----- Echyridella_menziesii      
  |                       +--+                                 
  |                          |  +-- Velesunio_angasi           
  |                          +--+                              
  |                             +-- Lortiella                  
  |                                                            
--+                             +-- Mutela_dubia **               
  |                       +-----+                              
  |                       |     +-- Mutela_rostrata **            
  |  +--------------------+                                    
  |  |                    |  +----- Monocondylaea_minuana **      
  |  |                    +--+                                 
  |  |                       |  +-- Anodontites_guanarensis **    
  |  |                       +--+                              
  +--+                          +-- Anodontites_trigona **        
     |                                                         
     |  +-------------------------- Coelatura_aegyptiaca       
     |  |                                                      
     |  |                       +-- Margaritifera_margaritifera
     |  |  +--------------------+                              
     +--+  |                    +-- Cumberlandia_monodonta     
        |  |                                                   
        |  |        +-------------- Gonidea_angulata           
        |  |  +-----+                                          
        +--+  |     |  +----------- Unio_pictorum              
           |  |     +--+                                       
           |  |        |  +-------- Unio_tumidus               
           |  |        +--+                                    
           |  |           |  +----- Anodonta_cygnea            
           |  |           +--+                                 
           +--+              |  +-- Pyganodon_grandis          
              |              +--+                              
              |                 +-- Strophitus_undulatus       
              |                                                
              |                 +-- Glebula_rotundata          
              |  +--------------+                              
              |  |              +-- Cyrtonaias_tampicoensis    
              |  |                                             
              +--+  +-------------- Toxolasma_lividum          
                 |  |                                          
                 |  |        +----- Potamilus_alatus           
                 |  +--------+                                 
                 +--+        |  +-- Ligumia_recta              
                    |        +--+                              
                    |           +-- Ortmanniana_ligamentina    
                    |                                          
                    |  +----------- Amblema_plicata            
                    +--+                                       
                       |  +-------- Quadrula_quadrula          
                       +--+                                    
                          |  +----- Eurynia_dilatata           
                          +--+                                 
                             |  +-- Pleurobema_clava           
                             +--+                              
                                +-- Fusconaia_flava            

Roe & Hoeh, 2003 [Publ 2655], fig. 4.2

iT tT bi riS oT roT roS iD
5 5 3 2 26 25 2 8.0
                                   +-- Mutela_dubia **               
                             +-----+                              
                             |     +-- Mutela_rostrata **            
                          +--+                                    
                          |  |  +----- Monocondylaea_minuana **      
                          |  +--+                                 
                          |     |  +-- Anodontites_guanarensis **    
                          |     +--+                              
  +-----------------------+        +-- Anodontites_trigona **        
  |                       |                                       
  |                       |        +-- Castalia_stevensi          
  |                       |     +--+                              
  |                       |     |  +-- Velesunio_angasi           
  |                       +-----+                                 
  |                             |  +-- Echyridella_menziesii      
--+                             |  |                              
  |                             +--+-- Lortiella_rugata           
  |                                |                              
  |                                +-- Diplodon_gratus            
  |                                                               
  |                                +-- Cumberlandia_monodonta     
  |  +-----------------------------+                              
  |  |                             +-- Margaritifera_margaritifera
  +--+                                                            
     |  +----------------------------- Gonidea_angulata           
     |  |                                                         
     +--+  +-------------------------- Coelatura_aegyptiaca       
        |  |                                                      
        +--+  +----------------------- Fusconaia_flava            
           |  |                                                   
           |  +----------------------- Amblema_plicata            
           +--+                                                   
              |  +-------------------- Quadrula_quadrula          
              |  |                                                
              |  |  +----------------- Pleurobema_clava           
              +--+  |                                             
                 |  |  +-------------- Eurynia_dilatata           
                 |  |  |                                          
                 +--+  |  +----------- Ortmanniana_ligamentina    
                    |  +--+                                       
                    |  |  |  +-------- Glebula_rotundata          
                    |  |  +--+                                    
                    |  |     |  +----- Potamilus_alatus           
                    +--+     +--+                                 
                       |        |  +-- Ligumia_recta              
                       |        +--+                              
                       |           +-- Toxolasma_lividum          
                       |                                          
                       |  +----------- Cyrtonaias_tampicoensis    
                       |  |                                       
                       +--+        +-- Unio_pictorum              
                          |  +-----+                              
                          |  |     +-- Unio_tumidus               
                          +--+                                    
                             |  +----- Pyganodon_grandis          
                             +--+                                 
                                |  +-- Anodonta_cygnea            
                                +--+                              
                                   +-- Strophitus_undulatus       

Roe & Hoeh, 2003 [Publ 2655], fig. 4.3

iT tT bi riS oT roT roS iD
5 5 3 2 26 25 2 8.0
                             +-- Mutela_dubia **           
                       +-----+                          
                       |     +-- Mutela_rostrata **        
                    +--+                                
                    |  |  +----- Monocondylaea_minuana **  
                    |  +--+                             
                    |     |  +-- Anodontites_guanarensis **
                    |     +--+                          
  +-----------------+        +-- Anodontites_trigona **    
  |                 |                                   
  |                 |        +-- Castalia_stevensi      
  |                 |  +-----+                          
  |                 |  |     +-- Diplodon_gratus        
  |                 +--+                                
  |                    |  +----- Echyridella_menziesii  
  |                    +--+                             
--+                       |  +-- Velesunio_angasi       
  |                       +--+                          
  |                          +-- Lortiella_rugata       
  |                                                     
  |  +-------------------------- Coelatura_aegyptiaca   
  |  |                                                  
  |  |                       +-- Unio_pictorum          
  |  |                 +-----+                          
  |  |                 |     +-- Unio_tumidus           
  +--+  +--------------+                                
     |  |              |  +----- Anodonta_cygnea        
     |  |              +--+                             
     |  |                 |  +-- Pyganodon_grandis      
     |  |                 +--+                          
     +--+                    +-- Strophitus_undulatus   
        |                                               
        |  +-------------------- Gonidea_angulata       
        |  |                                            
        |  |  +----------------- Amblema_plicata        
        +--+  |                                         
           |  |        +-------- Quadrula_quadrula      
           |  |  +-----+                                
           +--+  |     |  +----- Eurynia_dilatata       
              |  |     +--+                             
              |  |        |  +-- Fusconaia_flava        
              |  |        +--+                          
              +--+           +-- Pleurobema_clava       
                 |                                      
                 |           +-- Cyrtonaias_tampicoensis
                 |  +--------+                          
                 |  |        +-- Glebula_rotundata      
                 +--+                                   
                    |  +-------- Toxolasma_lividum      
                    +--+                                
                       |  +----- Ortmanniana_ligamentina
                       +--+                             
                          |  +-- Ligumia_recta          
                          +--+                          
                             +-- Potamilus_alatus       

Whelan et al., 2011 [Publ 3380], fig. 1 ML

iT tT bi riS oT roT roS iD
5 5 3 2 32 31 2 3.0
                                +-- Margaritifera_margaritifera  
  +-----------------------------+                                
  |                             +-- Cumberlandia_monodonta       
  |                                                              
  |                             +-- Aspatharia_pfeifferiana **      
  |                       +-----+                                
  |                       |     +-- Chambardia_wahlbergi **         
  |  +--------------------+                                      
  |  |                    |  +----- Mutela_rostrata **              
--+  |                    +--+                                   
  |  |                       |  +-- Mycetopoda_siliquosa **         
  |  |                       +--+                                
  |  |                          +-- Tamsiella_schroeteriana **      
  |  |                                                           
  |  |                          +-- Lamellidens_savadiensis      
  |  |        +-----------------+                                
  |  |        |                 +-- Trapezidens_dolichorhynchus  
  +--+  +-----+                                                  
     |  |     |  +----------------- Leoparreysia_tavoyensis      
     |  |     |  |                                               
     |  |     +--+              +-- Radiatula_chaudhurii         
     |  |        |  +-----------+                                
     |  |        |  |           +-- Indochinella_pugio           
     |  |        +--+                                            
     |  |           |  +----------- Nitia_teretiuscula           
     |  |           +--+                                         
     |  |              |  +-------- Coelatura_aegyptiaca[2]      
     |  |              +--+                                      
     +--+                 |  +----- Coelatura_aegyptiaca[1]      
        |                 +--+                                   
        |                    |  +-- Coelatura_gabonensis         
        |                    +--+                                
        |                       +-- Prisodontopsis_aviculaeformis
        |                                                        
        |                       +-- Pseudodon_vondembuschianus   
        |  +--------------------+                                
        |  |                    +-- Lens_contradens              
        |  |                                                     
        |  |                    +-- Pyganodon_grandis            
        |  |                 +--+                                
        +--+                 |  +-- Lasmigona_costata            
           |  +--------------+                                   
           |  |              |  +-- Unio_pictorum                
           |  |              +--+                                
           |  |                 +-- Unio_caffer                  
           +--+                                                  
              |  +----------------- Gonidea_angulata             
              |  |                                               
              |  |           +----- Pronodularia_japanensis      
              +--+  +--------+                                   
                 |  |        |  +-- Potomida_littoralis          
                 |  |        +--+                                
                 +--+           +-- Potomida_semirugata          
                    |                                            
                    |  +----------- Pleurobema_sintoxia          
                    +--+                                         
                       |  +-------- Quadrula_quadrula            
                       +--+                                      
                          |  +----- Amblema_plicata              
                          +--+                                   
                             |  +-- Truncilla_truncata           
                             +--+                                
                                +-- Lampsilis_cardium            

Whelan et al., 2011 [Publ 3380], fig. 1 MP

iT tT bi riS oT roT roS iD
5 5 3 2 32 31 2 3.0
                                +-- Aspatharia_pfeifferiana **      
                          +-----+                                
                          |     +-- Chambardia_wahlbergi **         
  +-----------------------+                                      
  |                       |  +----- Mutela_rostrata **              
  |                       +--+                                   
  |                          |  +-- Mycetopoda_siliquosa **         
  |                          +--+                                
  |                             +-- Tamsiella_schroeteriana **      
  |                                                              
  |                             +-- Margaritifera_margaritifera  
  |                    +--------+                                
  |                    |        +-- Cumberlandia_monodonta       
--+  +-----------------+                                         
  |  |                 |  +-------- Velesunio_ambiguus           
  |  |                 +--+                                      
  |  |                    |  +----- Hyridella_australis          
  |  |                    +--+                                   
  |  |                       |  +-- Castalia_ambigua             
  |  |                       +--+                                
  |  |                          +-- Triplodon_corrugatus         
  |  |                                                           
  |  |                          +-- Lamellidens_savadiensis      
  |  |           +--------------+                                
  +--+           |              +-- Trapezidens_dolichorhynchus  
     |  +--------+                                               
     |  |        |  +-------------- Leoparreysia_tavoyensis      
     |  |        |  |                                            
     |  |        +--+           +-- Radiatula_chaudhurii         
     |  |           |  +--------+                                
     |  |           |  |        +-- Indochinella_pugio           
     |  |           +--+                                         
     |  |              |  +-------- Nitia_teretiuscula           
     |  |              |  |                                      
     |  |              +--+  +----- Coelatura_aegyptiaca[1]      
     +--+                 |  |                                   
        |                 +--+----- Coelatura_aegyptiaca[2]      
        |                    |                                   
        |                    |  +-- Coelatura_gabonensis         
        |                    +--+                                
        |                       +-- Prisodontopsis_aviculaeformis
        |                                                        
        |  +----------------------- Lens_contradens              
        |  |                                                     
        |  |  +-------------------- Pseudodon_vondembuschianus   
        +--+  |                                                  
           |  |                 +-- Pyganodon_grandis            
           |  |              +--+                                
           +--+              |  +-- Lasmigona_costata            
              |  +-----------+                                   
              |  |           |  +-- Unio_pictorum                
              |  |           +--+                                
              |  |              +-- Unio_caffer                  
              +--+                                               
                 |  +-------------- Gonidea_angulata             
                 |  |                                            
                 |  |        +----- Pronodularia_japanensis      
                 +--+  +-----+                                   
                    |  |     |  +-- Potomida_littoralis          
                    |  |     +--+                                
                    +--+        +-- Potomida_semirugata          
                       |                                         
                       |  +-------- Quadrula_quadrula            
                       |  |                                      
                       +--+-------- Pleurobema_sintoxia          
                          |                                      
                          |  +----- Amblema_plicata              
                          +--+                                   
                             |  +-- Lampsilis_cardium            
                             +--+                                
                                +-- Truncilla_truncata           

Lopes-Lima et al., 2020 [Publ 4080], fig. 2-4

iT tT bi riS oT roT roS iD
5 5 3 3 117 115 2 5.0
                                            +-- Margaritifera_laevis       
  +-----------------------------------------+                              
  |                                         +-- Margaritifera_middendorffi 
  |                                                                        
  |                                         +-- Lamproscapha_ensiformis **    
  |                                   +-----+                              
  |                                   |     +-- Mycetopoda_siliquosa **       
  |  +--------------------------------+                                    
  |  |                                |  +----- Etheria_elliptica **          
  |  |                                +--+                                 
  |  |                                   |  +-- Aspatharia_pfeifferiana **    
  |  |                                   +--+                              
  |  |                                      +-- Chambardia_wahlbergi **       
  |  |                                                                     
--+  |                                      +-- Coelatura_aegyptiaca       
  |  |                                      |                              
  |  |  +-----------------------------------+-- Leoparreysia_tavoyensis    
  |  |  |                                   |                              
  |  |  |                                   +-- Radiatula_mouhoti          
  |  |  |                                                                  
  |  |  |                                   +-- Lens_contradens            
  |  |  |                                   |                              
  |  |  |                                   +-- Ensidens_sagittarius       
  |  |  |                                   |                              
  |  |  |           +-----------------------+-- Physunio_modelli           
  |  |  |           |                       |                              
  |  |  |           |                       +-- Rectidens_sumatrensis      
  |  |  |           |                       |                              
  +--+  |           |                       +-- Yaukthwa_nesemanni         
     |  |           |                                                      
     |  |           |                       +-- Amblema_plicata            
     |  |           |                       |                              
     |  |           |                       +-- Elliptio_complanata        
     |  |           |  +--------------------+                              
     |  |  +--------+  |                    +-- Lampsilis_cardium          
     |  |  |        |  |                    |                              
     |  |  |        |  |                    +-- Quadrula_quadrula          
     |  |  |        |  |                                                   
     |  |  |        |  |                 +----- Chamberlainia_somsakpanhai 
     |  |  |        |  |     +-----------+                                 
     |  |  |        |  |     |           |  +-- Sinohyriopsis_cumingii     
     |  |  |        |  |     |           +--+                              
     |  |  |        |  |  +--+              +-- Sinohyriopsis_schlegelii   
     |  |  |        |  |  |  |                                             
     |  |  |        |  |  |  |  +-------------- Pronodularia_japanensis    
     |  |  |        +--+  |  |  |                                          
     |  |  |           |  |  +--+  +----------- Potomida_littoralis        
     |  |  |           |  |     |  |                                       
     |  |  |           |  |     +--+  +-------- Lamprotula_caveata         
     |  |  |           |  |        |  |                                    
     +--+  |           |  |        +--+  +----- Lamprotula_leaii[2]        
        |  |           |  |           |  |                                 
        |  |           |  |           +--+----- Lamprotula_leaii[3]        
        |  |           |  |              |                                 
        |  |           |  |              |  +-- Lamprotula_seomjinensis    
        |  |           |  |              +--+                              
        |  |           |  |                 +-- Lamprotula_leaii[1]        
        |  |           +--+                                                
        |  |              |                 +-- Indopseudodon_indawgyiensis
        |  |              |        +--------+                              
        |  |              |        |        +-- Indopseudodon_salwenianus  
        |  |              |        |                                       
        |  |              |  +-----+        +-- Pilsbryoconcha_exilis      
        |  |              |  |     |  +-----+                              
        |  |              |  |     |  |     +-- Pilsbryoconcha_carinifera  
        |  |              |  |     +--+                                    
        |  |              |  |        |  +----- Pseudodon_mekongi          
        |  |              |  |        |  |                                 
        |  |              |  |        +--+  +-- Nyeinchanconcha_nyeinchani 
        |  |              |  |           |  |                              
        |  |              |  |           +--+-- Bineurus_mouhotii          
        |  |              |  |              |                              
        |  |              +--+              +-- Sundadontina_tumida        
        |  |                 |                                             
        |  |                 |        +-------- Inversidens_brandtii       
        |  |                 |        |                                    
        |  |                 |  +-----+     +-- Obovalis_omiensis          
        |  |                 |  |     |  +--+                              
        |  |                 |  |     |  |  +-- Ptychorhynchus_pfisteri    
        |  |                 |  |     +--+                                 
        |  |                 |  |        |  +-- Parvasolenaia_rivularis    
        |  |                 |  |        +--+                              
        |  |                 +--+           +-- Koreosolenaia_sitgyensis   
        |  |                    |                                          
        +--+                    |        +----- Gonidea_angulata           
           |                    |  +-----+                                 
           |                    |  |     |  +-- Leguminaia_wheatleyi       
           |                    |  |     +--+                              
           |                    +--+        +-- Microcondylaea_bonellii    
           |                       |                                       
           |                       |  +-------- Sinosolenaia_carinata      
           |                       +--+                                    
           |                          |  +----- Sinosolenaia_recognita     
           |                          +--+                                 
           |                             |  +-- Sinosolenaia_iridinea      
           |                             +--+                              
           |                                +-- Sinosolenaia_oleivora      
           |                                                               
           |  +-------------------------------- Acuticosta_chinensis       
           |  |                                                            
           |  +-------------------------------- Lepidodesma_languilati     
           |  |                                                            
           |  |                    +----------- Aculamprotula_tortuosa     
           |  |     +--------------+                                       
           |  |     |              |  +-------- Aculamprotula_scripta      
           |  |     |              +--+                                    
           |  |     |                 |  +----- Aculamprotula_coreana      
           |  |     |                 +--+                                 
           |  |     |                    |  +-- Aculamprotula_fibrosa      
           |  |     |                    +--+                              
           |  +-----+                       +-- Aculamprotula_tientsinensis
           |  |     |                                                      
           |  |     |  +----------------------- Middendorffinaia_mongolica 
           |  |     |  |                                                   
           |  |     |  |                 +----- Unio_gibbus                
           |  |     |  |                 |                                 
           |  |     |  |  +--------------+----- Unio_tumidus               
           |  |     +--+  |              |                                 
           |  |        |  |              |  +-- Unio_crassus               
           |  |        |  |              +--+                              
           |  |        |  |                 +-- Unio_pictorum              
           |  |        |  |                                                
           |  |        |  |     +-------------- Nodularia_breviconcha      
           |  |        +--+  +--+                                          
           |  |           |  |  |  +----------- Nodularia                  
           |  |           |  |  +--+                                       
           |  |           |  |     |  +-------- Nodularia_nuxpersicae      
           |  |           |  |     +--+                                    
           |  |           |  |        |  +----- Nodularia_nipponensis      
           |  |           |  |        +--+                                 
           |  |           |  |           |  +-- Nodularia_douglasiae       
           |  |           +--+           +--+                              
           |  |              |              +-- Nodularia_dualobtusa       
           |  |              |                                             
           +--+              |              +-- Schistodesmus_lampreyanus  
              |              |  +-----------+                              
              |              |  |           +-- Schistodesmus_spinosus     
              |              |  |                                          
              |              |  |        +----- Cuneopsis_rufescens        
              |              |  +--------+                                 
              |              +--+        |  +-- Cuneopsis_heudei           
              |                 |        +--+                              
              |                 |           +-- Cuneopsis_celtiformis      
              |                 |                                          
              |                 |  +----------- Tchangsinaia_piscicula     
              |                 +--+                                       
              |                    |  +-------- Inversiunio_jokohamensis   
              |                    +--+                                    
              |                       |  +----- Inversiunio_verrucosus     
              |                       +--+                                 
              |                          |  +-- Inversiunio_reinianus      
              |                          +--+                              
              |                             +-- Inversiunio_yanagawensis   
              |                                                            
              |              +----------------- Lanceolaria_oxyrhyncha     
              |              |                                             
              |  +-----------+  +-------------- Lanceolaria_gladiola       
              |  |           |  |                                          
              |  |           +--+           +-- Lanceolaria_bilirata       
              |  |              |  +--------+                              
              |  |              |  |        +-- Lanceolaria_grayii         
              |  |              +--+                                       
              |  |                 |  +-------- Lanceolaria_acrorrhyncha   
              |  |                 +--+                                    
              |  |                    |  +----- Lanceolaria_kihirai        
              |  |                    +--+                                 
              |  |                       |  +-- Lanceolaria_lanceolata     
              |  |                       +--+                              
              |  |                          +-- Lanceolaria_triformis      
              |  |                                                         
              |  |                    +-------- Pseudanodonta_complanata   
              +--+                    |                                    
                 |                    |  +----- Anodonta_cygnea            
                 |                    +--+                                 
                 |                    |  |  +-- Anodonta_anatina           
                 |  +-----------------+  +--+                              
                 |  |                 |     +-- Anodonta_nuttalliana       
                 |  |                 |                                    
                 |  |                 |  +----- Pyganodon_grandis          
                 |  |                 |  |                                 
                 |  |                 |  +----- Simpsonaias_ambigua        
                 |  |                 +--+                                 
                 |  |                    +----- Strophitus_undulatus       
                 |  |                    |                                 
                 |  |                    |  +-- Alasmidonta_marginata      
                 |  |                    +--+                              
                 |  |                       +-- Lasmigona_costata          
                 +--+                                                      
                    |                       +-- Cristaria_plicata          
                    |  +--------------------+                              
                    |  |                    +-- Cristaria_clessini         
                    |  |                                                   
                    |  |        +-------------- Anemina_arcaeformis        
                    |  |        |                                          
                    |  +--------+  +----------- Amuranodonta_kijaensis     
                    |  |        |  |                                       
                    |  |        +--+  +-------- Buldowskia_suifunica       
                    |  |           |  |                                    
                    |  |           +--+     +-- Buldowskia_flavotincta     
                    +--+              |  +--+                              
                       |              |  |  +-- Buldowskia_shadini         
                       |              +--+                                 
                       |                 |  +-- Buldowskia_iwakawai        
                       |                 +--+                              
                       |                    +-- Buldowskia_kamiyai         
                       |                                                   
                       |  +-------------------- Pletholophus_tenuis        
                       |  |                                                
                       |  |           +-------- Beringiana_japonica        
                       |  |  +--------+                                    
                       +--+  |        |  +----- Beringiana_beringiana      
                          |  |        +--+                                 
                          |  |           |  +-- Beringiana_fukuharai       
                          |  |           +--+                              
                          +--+              +-- Beringiana_gosannensis     
                             |                                             
                             |              +-- Sinanodonta_angula         
                             |  +-----------+                              
                             |  |           +-- Sinanodonta_lucida         
                             |  |                                          
                             +--+  +----------- Sinanodonta_schrenkii      
                                |  |                                       
                                |  |        +-- Sinanodonta_calipygos      
                                |  +--------+                              
                                +--+        +-- Sinanodonta_tumens         
                                   |                                       
                                   |  +-------- Sinanodonta_woodiana[2]    
                                   |  |                                    
                                   +--+  +----- Sinanodonta_elliptica      
                                      |  |                                 
                                      +--+----- Sinanodonta_jourdyi        
                                         |                                 
                                         |  +-- Sinanodonta_lauta          
                                         +--+                              
                                            +-- Sinanodonta_woodiana[1]    

Wu et al., 2021 [Publ 4366], fig. 2, I

iT tT bi riS oT roT roS iD
5 5 3 3 86 84 2 3.0
                                      +----- Alathyria_jacksoni         
  +-----------------------------------+                                 
  |                                   |  +-- Castalia_ambigua           
  |                                   +--+                              
  |                                      +-- Diplodon_suavidicus        
  |                                                                     
  |                                      +-- Aspatharia_pfeifferiana **    
  |                                +-----+                              
  |                                |     +-- Chambardia_wahlbergi **       
  |  +-----------------------------+                                    
  |  |                             |  +----- Etheria_elliptica **          
  |  |                             +--+                                 
  |  |                                |  +-- Tamsiella_schroeteriana **    
--+  |                                +--+                              
  |  |                                   +-- Lamproscapha_ensiformis **    
  |  |                                                                  
  |  |                          +----------- Gibbosula_laosensis        
  |  |                          |                                       
  |  |                          |        +-- Pseudunio_homsensis        
  |  |  +-----------------------+  +-----+                              
  |  |  |                       |  |     +-- Cumberlandia_monodonta     
  |  |  |                       |  |                                    
  |  |  |                       +--+     +-- Margaritifera_middendorffi 
  |  |  |                          |  +--+                              
  |  |  |                          |  |  +-- Margaritifera_laevis       
  +--+  |                          +--+                                 
     |  |                             |  +-- Margaritifera_dahurica     
     |  |                             +--+                              
     |  |                                +-- Margaritifera_margaritifera
     |  |                                                               
     |  |                          +-------- Trapezidens_exolescens     
     |  |              +-----------+                                    
     |  |              |           |  +----- Lamellidens_savadiensis    
     |  |              |           +--+                                 
     |  |              |              |  +-- Lamellidens_generosus      
     |  |              |              +--+                              
     |  |  +-----------+                 +-- Lamellidens_ferrugineus    
     |  |  |           |                                                
     |  |  |           |              +----- Leoparreysia_olivacea      
     |  |  |           |  +-----------+                                 
     |  |  |           |  |           |  +-- Leoparreysia_canefrii      
     |  |  |           |  |           +--+                              
     +--+  |           +--+              +-- Leoparreysia_tavoyensis    
        |  |              |                                             
        |  |              |  +-------------- Coelatura_aegyptiaca       
        |  |              +--+                                          
        |  |                 |  +----------- Indonaia_andersoniana      
        |  |                 +--+                                       
        |  |                    |  +-------- Scabies_crispata           
        |  |                    +--+                                    
        |  |                       |  +----- Radiatula_myitkyinae       
        |  |                       +--+                                 
        |  |                          |  +-- Radiatula_mouhoti          
        |  |                          +--+                              
        |  |                             +-- Radiatula_chaudhurii       
        |  |                                                            
        |  |     +-------------------------- Acuticosta_chinensis       
        |  |     |                                                      
        |  |     |     +-------------------- Lepidodesma_languilati     
        |  |     |     |                                                
        |  |     |     |              +----- Aculamprotula_tortuosa     
        |  |     |  +--+  +-----------+                                 
        |  |     |  |  |  |           |  +-- Aculamprotula_scripta      
        |  |     |  |  |  |           +--+                              
        |  |     |  |  |  |              +-- Aculamprotula_tientsinensis
        +--+  +--+  |  +--+                                             
           |  |  |  |     |              +-- Unio_pictorum              
           |  |  |  |     |  +-----------+                              
           |  |  |  |     |  |           +-- Unio_tumidus               
           |  |  |  |     +--+                                          
           |  |  |  |        |  +----------- Nodularia_douglasiae       
           |  |  |  |        |  |                                       
           |  |  |  |        +--+        +-- Cuneopsis_heudei           
           |  |  |  |           |  +-----+                              
           |  |  +--+           |  |     +-- Cuneopsis_rufescens        
           |  |     |           +--+                                    
           |  |     |              |  +----- Tchangsinaia_piscicula     
           |  |     |              +--+                                 
           |  |     |                 |  +-- Schistodesmus_lampreyanus  
           |  |     |                 +--+                              
           |  |     |                    +-- Schistodesmus_spinosus     
           |  |     |                                                   
           |  |     |                 +----- Lanceolaria_gladiola       
           |  |     |     +-----------+                                 
           |  |     |     |           |  +-- Lanceolaria_grayii         
           |  |     |     |           +--+                              
           |  |     |     |              +-- Lanceolaria_lanceolata     
           |  |     |     |                                             
           |  |     +-----+        +-------- Anemina_arcaeformis        
           |  |           |  +-----+                                    
           |  |           |  |     |  +----- Cristaria_plicata          
           |  |           |  |     +--+                                 
           |  |           |  |        |  +-- Sinanodonta_lucida         
           |  |           +--+        +--+                              
           +--+              |           +-- Sinanodonta_woodiana       
              |              |                                          
              |              |  +----------- Lasmigona_costata          
              |              |  |                                       
              |              +--+        +-- Anodonta_anatina           
              |                 |  +-----+                              
              |                 |  |     +-- Anodonta_cygnea            
              |                 +--+                                    
              |                    |  +----- Pseudanodonta_complanata   
              |                    +--+                                 
              |                       |  +-- Pyganodon_grandis          
              |                       +--+                              
              |                          +-- Strophitus_undulatus       
              |                                                         
              |                    +-------- Hyriopsis_desowitzi        
              |                 +--+                                    
              |                 |  |  +----- Ensidens_telus             
              |                 |  +--+                                 
              |                 |     |  +-- Ensidens_spiculus          
              |  +--------------+     +--+                              
              |  |              |        +-- Ensidens_sagittarius       
              |  |              |                                       
              |  |              |  +-------- Physunio_modelli           
              |  |              +--+                                    
              |  |                 |  +----- Lens_eximius               
              |  |                 +--+                                 
              |  |                    |  +-- Yaukthwa_nesemanni         
              |  |                    +--+                              
              |  |                       +-- Yaukthwa_panhai            
              +--+                                                      
                 |           +-------------- Quadrula_quadrula          
                 |  +--------+                                          
                 |  |        |  +----------- Elliptio_complanata        
                 |  |        +--+                                       
                 |  |           |  +-------- Obliquaria_reflexa         
                 |  |           +--+                                    
                 |  |              |  +----- Potamilus_alatus           
                 |  |              +--+                                 
                 |  |                 |  +-- Lampsilis_cardium          
                 |  |                 +--+                              
                 +--+                    +-- Truncilla_truncata         
                    |                                                   
                    |                 +----- Chamberlainia_somsakpanhai 
                    |  +--------------+                                 
                    |  |              |  +-- Sinohyriopsis_cumingii     
                    |  |              +--+                              
                    |  |                 +-- Sinohyriopsis_schlegelii   
                    |  |                                                
                    |  |           +-------- Potomida_littoralis        
                    |  |  +--------+                                    
                    +--+  |        |  +----- Pronodularia_japanensis    
                       |  |        +--+                                 
                       |  |           |  +-- Lamprotula_caveata         
                       |  |           +--+                              
                       |  |              +-- Lamprotula_leaii           
                       |  |                                             
                       |  |           +----- Indopseudodon_indawgyiensis
                       +--+        +--+                                 
                          |        |  |  +-- Indopseudodon_bogani       
                          |        |  +--+                              
                          |  +-----+     +-- Indopseudodon_salwenianus  
                          |  |     |                                    
                          |  |     |  +----- Pilsbryoconcha_exilis      
                          |  |     +--+                                 
                          |  |        |  +-- Pseudodon_vagulus          
                          |  |        +--+                              
                          +--+           +-- Pilsbryoconcha_carinifera  
                             |                                          
                             |        +----- Gonidea_angulata           
                             |  +-----+                                 
                             |  |     |  +-- Leguminaia_wheatleyi       
                             |  |     +--+                              
                             +--+        +-- Microcondylaea_bonellii    
                                |                                       
                                |  +-------- Inversidens_brandtii       
                                |  |                                    
                                +--+     +-- Sinosolenaia_carinata      
                                   |  +--+                              
                                   |  |  +-- Sinosolenaia_oleivora      
                                   +--+                                 
                                      |  +-- Parvasolenaia_rivularis    
                                      +--+                              
                                         +-- Ptychorhynchus_pfisteri    

Wu et al., 2021 [Publ 4366], fig. 6

iT tT bi riS oT roT roS iD
5 5 3 3 86 84 2 3.0
                                         +-- Chambardia_wahlbergi **       
                                   +-----+                              
                                   |     +-- Aspatharia_pfeifferiana **    
  +--------------------------------+                                    
  |                                |  +----- Etheria_elliptica **          
  |                                +--+                                 
  |                                   |  +-- Tamsiella_schroeteriana **    
  |                                   +--+                              
  |                                      +-- Lamproscapha_ensiformis **    
  |                                                                     
  |                                   +----- Alathyria_jacksoni         
  |  +--------------------------------+                                 
  |  |                                |  +-- Diplodon_suavidicus        
  |  |                                +--+                              
--+  |                                   +-- Castalia_ambigua           
  |  |                                                                  
  |  |                          +----------- Gibbosula_laosensis        
  |  |                          |                                       
  |  |                          |        +-- Cumberlandia_monodonta     
  |  |  +-----------------------+  +-----+                              
  |  |  |                       |  |     +-- Pseudunio_homsensis        
  |  |  |                       |  |                                    
  |  |  |                       +--+     +-- Margaritifera_middendorffi 
  |  |  |                          |  +--+                              
  |  |  |                          |  |  +-- Margaritifera_laevis       
  |  |  |                          +--+                                 
  +--+  |                             |  +-- Margaritifera_dahurica     
     |  |                             +--+                              
     |  |                                +-- Margaritifera_margaritifera
     |  |                                                               
     |  |                          +-------- Trapezidens_exolescens     
     |  |              +-----------+                                    
     |  |              |           |  +----- Lamellidens_savadiensis    
     |  |              |           +--+                                 
     |  |              |              |  +-- Lamellidens_ferrugineus    
     |  |              |              +--+                              
     |  |  +-----------+                 +-- Lamellidens_generosus      
     |  |  |           |                                                
     |  |  |           |              +----- Leoparreysia_olivacea      
     |  |  |           |  +-----------+                                 
     |  |  |           |  |           |  +-- Leoparreysia_canefrii      
     |  |  |           |  |           +--+                              
     +--+  |           +--+              +-- Leoparreysia_tavoyensis    
        |  |              |                                             
        |  |              |  +-------------- Coelatura_aegyptiaca       
        |  |              +--+                                          
        |  |                 |  +----------- Indonaia_andersoniana      
        |  |                 +--+                                       
        |  |                    |  +-------- Radiatula_myitkyinae       
        |  |                    +--+                                    
        |  |                       |  +----- Scabies_crispata           
        |  |                       +--+                                 
        |  |                          |  +-- Radiatula_mouhoti          
        |  |                          +--+                              
        |  |                             +-- Radiatula_chaudhurii       
        |  |                                                            
        |  |     +-------------------------- Acuticosta_chinensis       
        |  |     |                                                      
        |  |     |                    +----- Aculamprotula_tortuosa     
        |  |     |        +-----------+                                 
        |  |     |        |           |  +-- Aculamprotula_tientsinensis
        |  |     |        |           +--+                              
        |  |     |        |              +-- Aculamprotula_scripta      
        |  |     |  +-----+                                             
        |  |     |  |     |              +-- Unio_pictorum              
        +--+  +--+  |     |  +-----------+                              
           |  |  |  |     |  |           +-- Unio_tumidus               
           |  |  |  |     +--+                                          
           |  |  |  |        |  +----------- Nodularia_douglasiae       
           |  |  |  |        |  |                                       
           |  |  |  |        +--+        +-- Cuneopsis_heudei           
           |  |  |  |           |  +-----+                              
           |  |  |  |           |  |     +-- Cuneopsis_rufescens        
           |  |  |  |           +--+                                    
           |  |  +--+              |  +----- Tchangsinaia_piscicula     
           |  |     |              +--+                                 
           |  |     |                 |  +-- Schistodesmus_spinosus     
           |  |     |                 +--+                              
           |  |     |                    +-- Schistodesmus_lampreyanus  
           |  |     |                                                   
           |  |     |                 +----- Lanceolaria_gladiola       
           |  |     |  +--------------+                                 
           |  |     |  |              |  +-- Lanceolaria_grayii         
           |  |     |  |              +--+                              
           |  |     |  |                 +-- Lanceolaria_lanceolata     
           |  |     +--+                                                
           |  |        |  +----------------- Lepidodesma_languilati     
           |  |        |  |                                             
           |  |        |  |        +-------- Anemina_arcaeformis        
           |  |        +--+  +-----+                                    
           |  |           |  |     |  +----- Cristaria_plicata          
           |  |           |  |     +--+                                 
           |  |           |  |        |  +-- Sinanodonta_woodiana       
           |  |           +--+        +--+                              
           +--+              |           +-- Sinanodonta_lucida         
              |              |                                          
              |              |  +----------- Anodonta_cygnea            
              |              |  |                                       
              |              +--+        +-- Lasmigona_costata          
              |                 |  +-----+                              
              |                 |  |     +-- Anodonta_anatina           
              |                 +--+                                    
              |                    |  +----- Pseudanodonta_complanata   
              |                    +--+                                 
              |                       |  +-- Pyganodon_grandis          
              |                       +--+                              
              |                          +-- Strophitus_undulatus       
              |                                                         
              |                    +-------- Physunio_modelli           
              |                 +--+                                    
              |                 |  |  +----- Lens_eximius               
              |                 |  +--+                                 
              |                 |     |  +-- Yaukthwa_nesemanni         
              |  +--------------+     +--+                              
              |  |              |        +-- Yaukthwa_panhai            
              |  |              |                                       
              |  |              |  +-------- Hyriopsis_desowitzi        
              |  |              +--+                                    
              |  |                 |  +----- Ensidens_telus             
              |  |                 +--+                                 
              |  |                    |  +-- Ensidens_spiculus          
              |  |                    +--+                              
              |  |                       +-- Ensidens_sagittarius       
              |  |                                                      
              +--+                       +-- Quadrula_quadrula          
                 |              +--------+                              
                 |              |        +-- Elliptio_complanata        
                 |  +-----------+                                       
                 |  |           |  +-------- Obliquaria_reflexa         
                 |  |           +--+                                    
                 |  |              |  +----- Potamilus_alatus           
                 |  |              +--+                                 
                 |  |                 |  +-- Lampsilis_cardium          
                 |  |                 +--+                              
                 |  |                    +-- Truncilla_truncata         
                 +--+                                                   
                    |                 +----- Chamberlainia_somsakpanhai 
                    |  +--------------+                                 
                    |  |              |  +-- Sinohyriopsis_cumingii     
                    |  |              +--+                              
                    |  |                 +-- Sinohyriopsis_schlegelii   
                    |  |                                                
                    |  |           +-------- Pronodularia_japanensis    
                    |  |  +--------+                                    
                    +--+  |        |  +----- Potomida_littoralis        
                       |  |        +--+                                 
                       |  |           |  +-- Lamprotula_leaii           
                       |  |           +--+                              
                       |  |              +-- Lamprotula_caveata         
                       |  |                                             
                       |  |           +----- Indopseudodon_indawgyiensis
                       |  |        +--+                                 
                       +--+        |  |  +-- Indopseudodon_salwenianus  
                          |        |  +--+                              
                          |  +-----+     +-- Indopseudodon_bogani       
                          |  |     |                                    
                          |  |     |  +----- Pilsbryoconcha_exilis      
                          |  |     +--+                                 
                          |  |        |  +-- Pseudodon_vagulus          
                          |  |        +--+                              
                          |  |           +-- Pilsbryoconcha_carinifera  
                          +--+                                          
                             |     +-------- Inversidens_brandtii       
                             |  +--+                                    
                             |  |  |  +----- Gonidea_angulata           
                             |  |  +--+                                 
                             |  |     |  +-- Microcondylaea_bonellii    
                             |  |     +--+                              
                             +--+        +-- Leguminaia_wheatleyi       
                                |                                       
                                |        +-- Sinosolenaia_oleivora      
                                |     +--+                              
                                |     |  +-- Sinosolenaia_carinata      
                                +-----+                                 
                                      |  +-- Ptychorhynchus_pfisteri    
                                      +--+                              
                                         +-- Parvasolenaia_rivularis    

Bolotov et al., 2016 [Publ 3743], fig. 2

iT tT bi riS oT roT roS iD
4 4 2 3 26 23 2 4.0
                       +----- Diplodon_suavidicus        
                 +-----+                                 
                 |     |  +-- Triplodon_corrugatus       
                 |     +--+                              
                 |        +-- Castalia_ambigua           
  +--------------+                                       
  |              |     +----- Microdontia_anodontaeformis
  |              |  +--+                                 
  |              |  |  |  +-- Lortiella_froggatti        
  |              |  |  +--+                              
  |              +--+     +-- Alathyria_pertexta         
  |                 |                                    
  |                 |  +----- Velesunio_ambiguus         
  |                 +--+                                 
  |                    |  +-- Alathyria_jacksoni         
--+                    +--+                              
  |                       +-- Alathyria_profuga          
  |                                                      
  |                 +-------- Tamsiella_schroeteriana **    
  |  +--------------+                                    
  |  |              |  +----- Etheria_elliptica **          
  |  |              +--+                                 
  |  |                 |  +-- Aspatharia_pfeifferiana **    
  |  |                 +--+                              
  |  |                    +-- Chambardia_wahlbergi **       
  +--+                                                   
     |              +-------- Lampsilis_cardium          
     |  +-----------+                                    
     |  |           |  +----- Anodonta_anatina           
     |  |           +--+                                 
     |  |              |  +-- Unio_pictorum              
     |  |              +--+                              
     +--+                 +-- Unio_ravoisieri            
        |                                                
        |                 +-- Pseudunio_auricularius     
        |  +--------------+                              
        |  |              +-- Pseudunio_marocanus        
        |  |                                             
        +--+              +-- Gibbosula_laosensis        
           |  +-----------+                              
           |  |           +-- Cumberlandia_monodonta     
           |  |                                          
           +--+           +-- Margaritifera_dahurica     
              |  +--------+                              
              |  |        +-- Margaritifera_margaritifera
              +--+                                       
                 |  +-------- Margaritifera_falcata      
                 +--+                                    
                    |  +----- Margaritifera_laevis       
                    +--+                                 
                       |  +-- Margaritifera_middendorffi 
                       +--+                              
                          +-- Margaritifera_marrianae    

Bolotov et al., 2016 [Publ 3743], fig. 4

iT tT bi riS oT roT roS iD
4 4 2 3 26 23 2 4.0
                       +----- Diplodon_suavidicus        
                 +-----+                                 
                 |     |  +-- Triplodon_corrugatus       
                 |     +--+                              
                 |        +-- Castalia_ambigua           
  +--------------+                                       
  |              |     +----- Microdontia_anodontaeformis
  |              |  +--+                                 
  |              |  |  |  +-- Alathyria_pertexta         
  |              |  |  +--+                              
  |              +--+     +-- Lortiella_froggatti        
  |                 |                                    
  |                 |  +----- Velesunio_ambiguus         
  |                 +--+                                 
  |                    |  +-- Alathyria_jacksoni         
--+                    +--+                              
  |                       +-- Alathyria_profuga          
  |                                                      
  |                 +-------- Tamsiella_schroeteriana **    
  |  +--------------+                                    
  |  |              |  +----- Etheria_elliptica **          
  |  |              +--+                                 
  |  |                 |  +-- Chambardia_wahlbergi **       
  |  |                 +--+                              
  |  |                    +-- Aspatharia_pfeifferiana **    
  +--+                                                   
     |              +-------- Lampsilis_cardium          
     |  +-----------+                                    
     |  |           |  +----- Anodonta_anatina           
     |  |           +--+                                 
     |  |              |  +-- Unio_pictorum              
     |  |              +--+                              
     +--+                 +-- Unio_ravoisieri            
        |                                                
        |                 +-- Pseudunio_auricularius     
        |  +--------------+                              
        |  |              +-- Pseudunio_marocanus        
        |  |                                             
        +--+              +-- Gibbosula_laosensis        
           |  +-----------+                              
           |  |           +-- Cumberlandia_monodonta     
           |  |                                          
           +--+           +-- Margaritifera_margaritifera
              |  +--------+                              
              |  |        +-- Margaritifera_dahurica     
              +--+                                       
                 |  +-------- Margaritifera_falcata      
                 +--+                                    
                    |  +----- Margaritifera_laevis       
                    +--+                                 
                       |  +-- Margaritifera_marrianae    
                       +--+                              
                          +-- Margaritifera_middendorffi 

Bolotov et al., 2017 [Publ 3761], fig. S1

iT tT bi riS oT roT roS iD
4 4 2 3 138 136 2 4.0
                                +-------- Tamsiella_schroeteriana **    
                          +-----+                                    
                          |     |  +----- Etheria_elliptica **          
                          |     +--+                                 
                          |        |  +-- Chambardia_wahlbergi **       
                          |        +--+                              
                          |           +-- Aspatharia_pfeifferiana **    
  +-----------------------+                                          
  |                       |        +----- Diplodon_suavidicus        
  |                       |  +-----+                                 
  |                       |  |     |  +-- Castalia_ambigua           
  |                       |  |     +--+                              
  |                       |  |        +-- Triplodon_corrugatus       
  |                       +--+                                       
  |                          |     +----- Alathyria_pertexta         
  |                          |  +--+                                 
  |                          |  |  |  +-- Microdontia_anodontaeformis
  |                          |  |  +--+                              
  |                          +--+     +-- Lortiella_froggatti        
  |                             |                                    
  |                             |  +----- Velesunio_ambiguus         
  |                             +--+                                 
  |                                |  +-- Alathyria_jacksoni         
  |                                +--+                              
  |                                   +-- Alathyria_profuga          
  |                                                                  
  |                                   +-- Trapezidens_dolichorhynchus
  |                       +-----------+                              
  |                       |           +-- Trapezidens_exolescens     
  |           +-----------+                                          
  |           |           |  +----------- Lamellidens_corrianus      
  |           |           |  |                                       
  |           |           +--+  +-------- Lamellidens_savadiensis    
  |           |              |  |                                    
  |           |              +--+     +-- Lamellidens_jenkinsianus   
  |           |                 |  +--+                              
  |           |                 |  |  +-- Lamellidens_mainwaringi    
  |           |                 +--+                                 
  |           |                    |  +-- Lamellidens_ferrugineus    
  |  +--------+                    +--+                              
  |  |        |                       +-- Lamellidens_marginalis     
  |  |        |                                                      
  |  |        |                       +-- Parreysia_nagpoorensis     
  |  |        |                       |                              
  |  |        |  +--------------------+-- Parreysia_plagiosoma       
  |  |        |  |                    |                              
  |  |        |  |                    +-- Parreysia_corrugata        
--+  |        |  |                                                   
  |  |        |  |                    +-- Leoparreysia_tavoyensis    
  |  |        +--+  +-----------------+                              
  |  |           |  |                 +-- Leoparreysia_olivacea      
  |  |           |  |                                                
  |  |           |  |     +-------------- Nitia_teretiuscula         
  |  |           |  |  +--+                                          
  |  |           |  |  |  |  +----------- Coelatura_aegyptiaca[2]    
  |  |           +--+  |  +--+                                       
  |  |              |  |     |  +-------- Coelatura_aegyptiaca[1]    
  |  |              |  |     +--+                                    
  |  |              |  |        |  +----- Prisodontopsis_mweruensis  
  |  |              |  |        +--+                                 
  |  |              |  |           |  +-- Coelatura_gabonensis       
  |  |              +--+           +--+                              
  |  |                 |              +-- Coelatura_kunenensis       
  |  |                 |                                             
  |  |                 |        +-------- Indonaia_shurtleffiana     
  |  |                 |  +-----+                                    
  |  |                 |  |     |  +----- Indonaia_andersoniana      
  |  |                 |  |     +--+                                 
  |  |                 |  |        |  +-- Indonaia_rugosa            
  |  |                 |  |        +--+                              
  |  |                 +--+           +-- Indonaia_bonneaudii        
  |  |                    |                                          
  |  |                    |           +-- Indochinella_pugio         
  |  |                    |  +--------+                              
  |  |                    |  |        +-- Scabies_crispata           
  |  |                    |  |                                       
  |  |                    +--+        +-- Radiatula_myitkyinae       
  |  |                       |  +-----+                              
  |  |                       |  |     +-- Radiatula_chaudhurii       
  |  |                       +--+                                    
  |  |                          |  +----- Scabies_mandarinus         
  |  |                          +--+                                 
  |  |                             |  +-- Scabies_humilis            
  |  |                             +--+                              
  |  |                                +-- Scabies_phaselus           
  |  |                                                               
  |  |                                +-- Ensidens_telus             
  +--+                          +-----+                              
     |                          |     +-- Ensidens_ingallsianus      
     |                    +-----+                                    
     |                    |     |  +----- Ensidens_sagittarius       
     |                    |     +--+                                 
     |                    |        |  +-- Ensidens_spiculus          
     |                    |        +--+                              
     |                 +--+           +-- Ensidens_jaculus           
     |                 |  |                                          
     |                 |  |           +-- Hyriopsis_bialata          
     |                 |  |  +--------+                              
     |                 |  |  |        +-- Hyriopsis_khoratensis      
     |                 |  +--+                                       
     |                 |     |  +-------- Hyriopsis_panhai           
     |                 |     +--+                                    
     |                 |        |  +----- Hyriopsis_myersiana        
     |                 |        +--+                                 
     |  +--------------+           |  +-- Rectidens_sumatrensis      
     |  |              |           +--+                              
     |  |              |              +-- Hyriopsis_desowitzi        
     |  |              |                                             
     |  |              |           +----- Physunio                   
     |  |              |  +--------+                                 
     |  |              |  |        |  +-- Physunio_modelli           
     |  |              |  |        +--+                              
     |  |              |  |           +-- Physunio_superbus          
     |  |              |  |                                          
     |  |              +--+     +-------- Yaukthwa_inlenensis        
     |  |                 |  +--+                                    
     |  |                 |  |  |  +----- Yaukthwa_panhai            
     |  |                 |  |  +--+                                 
     |  |                 |  |     |  +-- Yaukthwa_rectangularis     
     |  |                 |  |     +--+                              
     |  |                 +--+        +-- Yaukthwa_nesemanni         
     |  |                    |                                       
     |  |                    |        +-- Lens_contradens            
     |  |                    |  +-----+                              
     |  |                    |  |     +-- Lens_micropterus           
     |  |                    +--+                                    
     |  |                       |  +----- Lens_eximius               
     |  |                       +--+                                 
     |  |                          |  +-- Lens_comptus               
     |  |                          +--+                              
     |  |                             +-- Lens_novoselovi            
     |  |                                                            
     |  |           +-------------------- Indopseudodon_indawgyiensis
     |  |           |                                                
     |  |           |           +-------- Pilsbryoconcha_exilis[1]   
     |  |           |  +--------+                                    
     +--+  +--------+  |        |  +----- Pilsbryoconcha_exilis[3]   
        |  |        |  |        +--+                                 
        |  |        |  |           |  +-- Pilsbryoconcha_exilis[2]   
        |  |        |  |           +--+                              
        |  |        |  |              +-- Pilsbryoconcha_carinifera  
        |  |        +--+                                             
        |  |           |        +-------- Sundadontina_tumida        
        |  |           |  +-----+                                    
        |  |           |  |     |  +----- Pseudodon_vagulus          
        |  |           |  |     +--+                                 
        |  |           |  |        |  +-- Pseudodon_mekongi          
        |  |           |  |        +--+                              
        |  |           +--+           +-- Pseudodon_vondembuschianus 
        |  |              |                                          
        |  |              |           +-- Nyeinchanconcha_nyeinchani 
        |  |              |  +--------+                              
        |  |              |  |        +-- Sundadontina_cumingii      
        |  |              +--+                                       
        |  |                 |  +-------- Thaiconcha_callifera       
        |  |                 +--+                                    
        |  |                    |  +----- Bineurus_exilis            
        |  |                    +--+                                 
        |  |                       |  +-- Bineurus_mouhotii          
        |  |                       +--+                              
        |  |                          +-- Bineurus_loeiensis         
        |  |                                                         
        |  |                    +-------- Amblema_plicata            
        |  |                 +--+                                    
        |  |                 |  |  +----- Ortmanniana_ligamentina    
        |  |                 |  +--+                                 
        |  |                 |     |  +-- Lampsilis_cardium          
        |  |                 |     +--+                              
        +--+           +-----+        +-- Cambarunio_iris            
           |           |     |                                       
           |           |     |        +-- Quadrula_quadrula          
           |           |     |  +-----+                              
           |           |     |  |     +-- Quadrula_verrucosa         
           |           |     +--+                                    
           |           |        |  +----- Pleurobema_sintoxia        
           |           |        +--+                                 
           |           |           |  +-- Eurynia_dilatata           
           |           |           +--+                              
           |           |              +-- Elliptio_complanata        
           |  +--------+                                             
           |  |        |              +-- Sinohyriopsis_cumingii     
           |  |        |        +-----+                              
           |  |        |        |     +-- Chamberlainia_somsakpanhai 
           |  |        |        |                                    
           |  |        |  +-----+     +-- Potomida_littoralis        
           |  |        |  |     |  +--+                              
           |  |        |  |     |  |  +-- Pronodularia_japanensis    
           |  |        |  |     +--+                                 
           |  |        |  |        |  +-- Lamprotula_caveata         
           |  |        |  |        +--+                              
           |  |        +--+           +-- Lamprotula_leaii           
           |  |           |                                          
           |  |           |        +----- Sinosolenaia_carinata      
           |  |           |  +-----+                                 
           |  |           |  |     |  +-- Sinosolenaia_oleivora      
           |  |           |  |     +--+                              
           |  |           +--+        +-- Sinosolenaia_recognita     
           |  |              |                                       
           |  |              |  +-------- Gonidea_angulata           
           |  |              +--+                                    
           +--+                 |  +----- Microcondylaea_bonellii    
              |                 +--+                                 
              |                    |  +-- Leguminaia_wheatleyi       
              |                    +--+                              
              |                       +-- Leguminaia                 
              |                                                      
              |                    +----- Unio_tumidus               
              |           +--------+                                 
              |           |        |  +-- Unio_pictorum              
              |           |        +--+                              
              |           |           +-- Unio_crassus               
              |  +--------+                                          
              |  |        |  +----------- Aculamprotula_tortuosa     
              |  |        |  |                                       
              |  |        |  |        +-- Nodularia_douglasiae       
              |  |        +--+  +-----+                              
              |  |           |  |     +-- Nodularia_nuxpersicae      
              |  |           |  |                                    
              |  |           +--+     +-- Cuneopsis_rufescens        
              |  |              |  +--+                              
              |  |              |  |  +-- Cuneopsis_heudei           
              |  |              +--+                                 
              +--+                 |  +-- Tchangsinaia_piscicula     
                 |                 +--+                              
                 |                    +-- Schistodesmus_lampreyanus  
                 |                                                   
                 |                    +-- Lanceolaria_grayii         
                 |  +-----------------+                              
                 |  |                 +-- Lanceolaria_gladiola       
                 |  |                                                
                 |  |                 +-- Cristaria_plicata          
                 |  |        +--------+                              
                 |  |        |        +-- Cristaria_radiata          
                 +--+  +-----+                                       
                    |  |     |  +-------- Pletholophus_tenuis        
                    |  |     +--+                                    
                    |  |        |  +----- Sinanodonta_lucida         
                    |  |        +--+                                 
                    |  |           |  +-- Sinanodonta_jourdyi        
                    +--+           +--+                              
                       |              +-- Sinanodonta_lauta          
                       |                                             
                       |  +-------------- Anodonta_anatina           
                       |  |                                          
                       |  |        +----- Pseudanodonta_complanata   
                       +--+  +-----+                                 
                          |  |     |  +-- Anodonta_cygnea            
                          |  |     +--+                              
                          |  |        +-- Anodonta_nuttalliana       
                          +--+                                       
                             |        +-- Lasmigona_costata          
                             |  +-----+                              
                             |  |     +-- Alasmidonta_marginata      
                             +--+                                    
                                |  +----- Simpsonaias_ambigua        
                                +--+                                 
                                   |  +-- Pyganodon_grandis          
                                   +--+                              
                                      +-- Strophitus_undulatus       

Bolotov et al., 2017 [Publ 3761], fig. S2

iT tT bi riS oT roT roS iD
4 4 2 3 138 136 2 2.0
              +-------------------- Chamberlainia_somsakpanhai 
              |                                                
  +-----------+  +----------------- Cumberlandia_monodonta     
  |           |  |                                             
  |           |  |  +-------------- Gibbosula_laosensis        
  |           +--+  |                                          
  |              |  |           +-- Pseudunio_auricularius     
  |              |  +-----------+                              
  |              +--+           +-- Pseudunio_marocanus        
  |                 |                                          
  |                 |           +-- Margaritifera_dahurica     
  |                 |  +--------+                              
  |                 |  |        +-- Margaritifera_margaritifera
  |                 +--+                                       
  |                    |  +-------- Margaritifera_falcata      
  |                    +--+                                    
  |                       |  +----- Margaritifera_middendorffi 
  |                       +--+                                 
  |                          |  +-- Margaritifera_laevis       
  |                          +--+                              
  |                             +-- Margaritifera_marrianae    
  |                                                            
  |                             +-- Tamsiella_schroeteriana **    
  |                          +--+                              
  |                          |  +-- Etheria_elliptica **          
  |                 +--------+                                 
  |                 |        |  +-- Aspatharia_pfeifferiana **    
  |                 |        +--+                              
  |                 |           +-- Chambardia_wahlbergi **       
  |                 |                                          
  +-----------------+        +----- Diplodon_suavidicus        
  |                 |  +-----+                                 
  |                 |  |     |  +-- Castalia_ambigua           
  |                 |  |     +--+                              
  |                 |  |        +-- Triplodon_corrugatus       
  |                 +--+                                       
  |                    |     +----- Velesunio_ambiguus         
  |                    |  +--+                                 
  |                    |  |  |  +-- Alathyria_jacksoni         
  |                    |  |  +--+                              
  |                    +--+     +-- Alathyria_profuga          
  |                       |                                    
  |                       |  +----- Microdontia_anodontaeformis
  |                       +--+                                 
  |                          |  +-- Alathyria_pertexta         
  |                          +--+                              
  |                             +-- Lortiella_froggatti        
  |                                                            
  |                             +-- Parreysia_nagpoorensis     
  |                             |                              
  |                 +-----------+-- Parreysia_plagiosoma       
  |                 |           |                              
  |                 |           +-- Parreysia_corrugata        
  |                 |                                          
  |        +--------+           +-- Trapezidens_dolichorhynchus
  |        |        |  +--------+                              
  |        |        |  |        +-- Trapezidens_exolescens     
  |        |        |  |                                       
--+        |        +--+  +-------- Lamellidens_savadiensis    
  |        |           |  |                                    
  |        |           |  +-------- Lamellidens_corrianus      
  |        |           +--+                                    
  |        |              |     +-- Lamellidens_marginalis     
  |        |              |  +--+                              
  |        |              |  |  +-- Lamellidens_ferrugineus    
  |  +-----+              +--+                                 
  |  |     |                 |  +-- Lamellidens_jenkinsianus   
  |  |     |                 +--+                              
  |  |     |                    +-- Lamellidens_mainwaringi    
  |  |     |                                                   
  |  |     |                    +-- Leoparreysia_olivacea      
  |  |     |  +-----------------+                              
  |  |     |  |                 +-- Leoparreysia_tavoyensis    
  |  |     |  |                                                
  |  |     |  |     +-------------- Nitia_teretiuscula         
  |  |     |  |  +--+                                          
  |  |     |  |  |  |  +----------- Coelatura_aegyptiaca[1]    
  |  |     +--+  |  +--+                                       
  |  |        |  |     |  +-------- Coelatura_aegyptiaca[2]    
  |  |        |  |     +--+                                    
  |  |        |  |        |  +----- Coelatura_gabonensis       
  |  |        |  |        +--+                                 
  |  |        |  |           |  +-- Coelatura_kunenensis       
  |  |        +--+           +--+                              
  |  |           |              +-- Prisodontopsis_mweruensis  
  |  |           |                                             
  |  |           |        +-------- Indonaia_shurtleffiana     
  |  |           |  +-----+                                    
  |  |           |  |     |  +----- Indonaia_andersoniana      
  |  |           |  |     +--+                                 
  |  |           |  |        |  +-- Indonaia_rugosa            
  |  |           |  |        +--+                              
  |  |           +--+           +-- Indonaia_bonneaudii        
  |  |              |                                          
  |  |              |           +-- Indochinella_pugio         
  |  |              |  +--------+                              
  |  |              |  |        +-- Scabies_crispata           
  |  |              |  |                                       
  |  |              +--+        +-- Radiatula_myitkyinae       
  |  |                 |  +-----+                              
  |  |                 |  |     +-- Radiatula_chaudhurii       
  |  |                 +--+                                    
  |  |                    |  +----- Scabies_mandarinus         
  |  |                    +--+                                 
  |  |                       |  +-- Scabies_humilis            
  |  |                       +--+                              
  |  |                          +-- Scabies_phaselus           
  |  |                                                         
  |  |                       +----- Unio_tumidus               
  |  |                 +-----+                                 
  |  |                 |     |  +-- Unio_crassus               
  |  |                 |     +--+                              
  |  |                 |        +-- Unio_pictorum              
  |  |           +-----+                                       
  +--+           |     |  +-------- Aculamprotula_tortuosa     
     |           |     |  |                                    
     |           |     |  +-------- Schistodesmus_lampreyanus  
     |           |     |  |                                    
     |           |     +--+     +-- Nodularia_douglasiae       
     |           |        +-----+                              
     |           |        |     +-- Nodularia_nuxpersicae      
     |           |        |                                    
     |           |        |  +----- Cuneopsis_heudei           
     |           |        +--+                                 
     |           |           |  +-- Tchangsinaia_piscicula     
     |           |           +--+                              
     |  +--------+              +-- Cuneopsis_rufescens        
     |  |        |                                             
     |  |        |              +-- Lanceolaria_grayii         
     |  |        |  +-----------+                              
     |  |        |  |           +-- Lanceolaria_gladiola       
     |  |        |  |                                          
     |  |        |  |           +-- Cristaria_plicata          
     |  |        |  |  +--------+                              
     |  |        |  |  |        +-- Cristaria_radiata          
     |  |        |  +--+                                       
     |  |        |  |  |  +-------- Pletholophus_tenuis        
     |  |        |  |  +--+                                    
     |  |        |  |     |  +----- Sinanodonta_lucida         
     |  |        +--+     +--+                                 
     |  |           |        |  +-- Sinanodonta_lauta          
     |  |           |        +--+                              
     |  |           |           +-- Sinanodonta_jourdyi        
     |  |           |                                          
     |  |           |     +-------- Anodonta_anatina           
     |  |           |     |                                    
     |  |           |     +-------- Pseudanodonta_complanata   
     |  |           |     |                                    
     |  |           |     |     +-- Anodonta_cygnea            
     |  |           +-----+-----+                              
     |  |                 |     +-- Anodonta_nuttalliana       
     |  |                 |                                    
     |  |                 |  +----- Pyganodon_grandis          
     |  |                 |  |                                 
     |  |                 |  +----- Simpsonaias_ambigua        
     |  |                 +--+                                 
     |  |                    +----- Strophitus_undulatus       
     |  |                    |                                 
     |  |                    |  +-- Alasmidonta_marginata      
     |  |                    +--+                              
     |  |                       +-- Lasmigona_costata          
     |  |                                                      
     |  |                 +-------- Lens_contradens            
     |  |           +-----+                                    
     +--+           |     |  +----- Lens_eximius               
        |           |     +--+                                 
        |           |        |  +-- Lens_comptus               
        |           |        +--+                              
        |        +--+           +-- Lens_novoselovi            
        |        |  |                                          
        |        |  |        +----- Physunio                   
        |        |  |  +-----+                                 
        |        |  |  |     |  +-- Physunio_modelli           
        |        |  |  |     +--+                              
        |        |  +--+        +-- Physunio_superbus          
        |        |     |                                       
        |        |     |  +-------- Yaukthwa_inlenensis        
        |        |     +--+                                    
        |        |        |  +----- Yaukthwa_panhai            
        |  +-----+        +--+                                 
        |  |     |           |  +-- Yaukthwa_rectangularis     
        |  |     |           +--+                              
        |  |     |              +-- Yaukthwa_nesemanni         
        |  |     |                                             
        |  |     |           +----- Hyriopsis_panhai           
        |  |     |           |                                 
        |  |     |  +--------+----- Hyriopsis_khoratensis      
        |  |     |  |        |                                 
        |  |     |  |        |  +-- Hyriopsis_myersiana        
        |  |     |  |        +--+                              
        |  |     +--+           +-- Hyriopsis_bialata          
        |  |        |                                          
        |  |        |  +----------- Hyriopsis_desowitzi        
        |  |        |  |                                       
        |  |        |  +----------- Rectidens_sumatrensis      
        |  |        +--+                                       
        |  |           |        +-- Ensidens_telus             
        |  |           |  +-----+                              
        |  |           |  |     +-- Ensidens_ingallsianus      
        |  |           +--+                                    
        |  |              |  +----- Ensidens_sagittarius       
        |  |              +--+                                 
        |  |                 |  +-- Ensidens_spiculus          
        +--+                 +--+                              
           |                    +-- Ensidens_jaculus           
           |                                                   
           |              +-------- Amblema_plicata            
           |           +--+                                    
           |           |  |  +----- Cambarunio_iris            
           |           |  +--+                                 
           |           |     |  +-- Ortmanniana_ligamentina    
           |           |     +--+                              
           |  +--------+        +-- Lampsilis_cardium          
           |  |        |                                       
           |  |        |        +-- Quadrula_quadrula          
           |  |        |  +-----+                              
           |  |        |  |     +-- Quadrula_verrucosa         
           |  |        +--+                                    
           |  |           |  +----- Pleurobema_sintoxia        
           |  |           +--+                                 
           |  |              |  +-- Elliptio_complanata        
           |  |              +--+                              
           |  |                 +-- Eurynia_dilatata           
           |  |                                                
           |  |           +-------- Sinohyriopsis_cumingii     
           |  |           |                                    
           |  |     +-----+     +-- Lamprotula_caveata         
           +--+     |     |  +--+                              
              |     |     |  |  +-- Lamprotula_leaii           
              |     |     +--+                                 
              |     |        |  +-- Potomida_littoralis        
              |  +--+        +--+                              
              |  |  |           +-- Pronodularia_japanensis    
              |  |  |                                          
              |  |  |  +----------- Sinosolenaia_carinata      
              |  |  |  |                                       
              |  |  |  |        +-- Sinosolenaia_oleivora      
              |  |  +--+--------+                              
              |  |     |        +-- Sinosolenaia_recognita     
              |  |     |                                       
              |  |     |  +-------- Gonidea_angulata           
              |  |     +--+                                    
              |  |        |  +----- Microcondylaea_bonellii    
              +--+        +--+                                 
                 |           |  +-- Leguminaia_wheatleyi       
                 |           +--+                              
                 |              +-- Leguminaia                 
                 |                                             
                 |        +-------- Pilsbryoconcha_exilis[3]   
                 |  +-----+                                    
                 |  |     |  +----- Pilsbryoconcha_exilis[2]   
                 |  |     +--+                                 
                 |  |        |  +-- Pilsbryoconcha_exilis[1]   
                 |  |        +--+                              
                 |  |           +-- Pilsbryoconcha_carinifera  
                 |  |                                          
                 +--+  +----------- Indopseudodon_indawgyiensis
                    |  |                                       
                    |  |  +-------- Thaiconcha_callifera       
                    |  +--+                                    
                    |  |  |  +----- Bineurus_exilis            
                    |  |  +--+                                 
                    |  |     |  +-- Bineurus_mouhotii          
                    +--+     +--+                              
                       |        +-- Bineurus_loeiensis         
                       |                                       
                       |     +----- Pseudodon_vagulus          
                       |  +--+                                 
                       |  |  |  +-- Pseudodon_mekongi          
                       |  |  +--+                              
                       |  |     +-- Pseudodon_vondembuschianus 
                       +--+                                    
                          |     +-- Pseudodon_inoscularis      
                          |  +--+                              
                          |  |  +-- Sundadontina_tumida        
                          +--+                                 
                             |  +-- Nyeinchanconcha_nyeinchani 
                             +--+                              
                                +-- Sundadontina_cumingii      

Graf et al., 2015 [Publ 3619], fig. 1

iT tT bi riS oT roT roS iD
3 3 1 3 17 16 2 3.0
                 +-- Unio_pictorum              
           +-----+                              
           |     +-- Margaritifera_margaritifera
  +--------+                                    
  |        |  +----- Chambardia_wahlbergi **       
  |        +--+                                 
  |           |  +-- Tamsiella_schroeteriana **    
  |           +--+                              
  |              +-- Etheria_elliptica **          
  |                                             
  |     +----------- Westralunio_carteri        
  |     |                                       
--+     |     +----- Alathyria_jacksoni         
  |  +--+  +--+                                 
  |  |  |  |  |  +-- Velesunio_ambiguus         
  |  |  |  |  +--+                              
  |  |  +--+     +-- Alathyria_profuga          
  |  |     |                                    
  |  |     |  +----- Alathyria_pertexta         
  |  |     +--+                                 
  |  |        |  +-- Microdontia_anodontaeformis
  +--+        +--+                              
     |           +-- Lortiella_froggatti        
     |                                          
     |        +----- Diplodon_suavidicus        
     |     +--+                                 
     |     |  |  +-- Triplodon_corrugatus       
     |     |  +--+                              
     |     |     +-- Castalia_ambigua           
     +-----+                                    
           |     +-- Hyridella_depressa         
           |  +--+                              
           |  |  +-- Hyridella_drapeta          
           +--+                                 
              |  +-- Hyridella_australis        
              +--+                              
                 +-- Cucumerunio_novaehollandiae

Pfeiffer & Graf, 2015 [Publ 3620], fig. 2

iT tT bi riS oT roT roS iD
3 3 1 3 37 36 2 6.0
                             +-------- Velesunio_ambiguus         
  +--------------------------+                                    
  |                          |  +----- Hyridella_australis        
  |                          +--+                                 
  |                             |  +-- Triplodon_corrugatus       
  |                             +--+                              
  |                                +-- Castalia_ambigua           
  |                                                               
--+                             +----- Etheria_elliptica **          
  |  +--------------------------+                                 
  |  |                          |  +-- Lamproscapha_ensiformis **    
  |  |                          +--+                              
  |  |                             +-- Chambardia_wahlbergi **       
  |  |                                                            
  |  |                             +-- Margaritifera_margaritifera
  |  |  +--------------------------+                              
  +--+  |                          +-- Cumberlandia_monodonta     
     |  |                                                         
     |  |                          +-- Trapezidens_dolichorhynchus
     |  |              +-----------+                              
     |  |              |           +-- Lamellidens_savadiensis    
     |  |              |                                          
     |  |  +-----------+           +-- Leoparreysia_olivacea      
     +--+  |           |  +--------+                              
        |  |           |  |        +-- Leoparreysia_tavoyensis    
        |  |           +--+                                       
        |  |              |  +-------- Coelatura_aegyptiaca       
        |  |              +--+                                    
        |  |                 |  +----- Indochinella_pugio         
        |  |                 +--+                                 
        |  |                    |  +-- Radiatula_chaudhurii       
        |  |                    +--+                              
        +--+                       +-- Scabies_mandarinus         
           |                                                      
           |                    +----- Unio_pictorum              
           |  +-----------------+                                 
           |  |                 |  +-- Pyganodon_grandis          
           |  |                 +--+                              
           |  |                    +-- Lasmigona_costata          
           |  |                                                   
           |  |           +----------- Quadrula_quadrula          
           |  |  +--------+                                       
           +--+  |        |  +-------- Amblema_plicata            
              |  |        +--+                                    
              |  |           |  +----- Potamilus_alatus           
              |  |           +--+                                 
              |  |              |  +-- Lampsilis_cardium          
              |  |              +--+                              
              |  |                 +-- Truncilla_truncata         
              +--+                                                
                 |           +-------- Ensidens_telus             
                 |  +--------+                                    
                 |  |        |  +----- Lens_micropterus           
                 |  |        +--+                                 
                 |  |           |  +-- Physunio                   
                 |  |           +--+                              
                 |  |              +-- Physunio_modelli           
                 +--+                                             
                    |        +-------- Gonidea_angulata           
                    |  +-----+                                    
                    |  |     |  +----- Potomida_littoralis        
                    |  |     +--+                                 
                    |  |        |  +-- Pronodularia_japanensis    
                    +--+        +--+                              
                       |           +-- Lamprotula_caveata         
                       |                                          
                       |  +----------- Pilsbryoconcha_carinifera  
                       |  |                                       
                       +--+        +-- Pseudodon_vondembuschianus 
                          |  +-----+                              
                          |  |     +-- Pseudodon_vagulus          
                          +--+                                    
                             |  +----- Sundadontina_tumida        
                             +--+                                 
                                |  +-- Nyeinchanconcha_nyeinchani 
                                +--+                              
                                   +-- Bineurus_exilis            

Konopleva et al., 2017 [Publ 3807], fig. 6

iT tT bi riS oT roT roS iD
3 3 1 3 28 26 2 5.0
                       +----- Diplodon_suavidicus        
  +--------------------+                                 
  |                    |  +-- Triplodon_corrugatus       
  |                    +--+                              
  |                       +-- Castalia_ambigua           
  |                                                      
  |                    +----- Tamsiella_schroeteriana **    
--+  +-----------------+                                 
  |  |                 |  +-- Etheria_elliptica **          
  |  |                 +--+                              
  |  |                    +-- Chambardia_wahlbergi **       
  |  |                                                   
  |  |                    +-- Margaritifera_dahurica     
  |  |  +-----------------+                              
  +--+  |                 +-- Margaritifera_laevis       
     |  |                                                
     |  |           +-------- Ensidens_sagittarius       
     |  |     +-----+                                    
     |  |     |     |  +----- Lens_micropterus           
     |  |     |     +--+                                 
     |  |     |        |  +-- Physunio                   
     |  |  +--+        +--+                              
     +--+  |  |           +-- Physunio_modelli           
        |  |  |                                          
        |  |  |  +----------- Sundadontina_tumida        
        |  |  |  |                                       
        |  |  +--+        +-- Nyeinchanconcha_nyeinchani 
        |  |     |  +-----+                              
        |  |     |  |     +-- Bineurus_exilis            
        |  |     +--+                                    
        |  |        |  +----- Pilsbryoconcha_carinifera  
        |  |        +--+                                 
        +--+           |  +-- Pseudodon_vagulus          
           |           +--+                              
           |              +-- Pseudodon_vondembuschianus 
           |                                             
           |              +-- Trapezidens_exolescens     
           |        +-----+                              
           |        |     +-- Trapezidens_dolichorhynchus
           |  +-----+                                    
           |  |     |  +----- Lamellidens_savadiensis    
           |  |     +--+                                 
           |  |        |  +-- Lamellidens_jenkinsianus   
           |  |        +--+                              
           +--+           +-- Lamellidens_mainwaringi    
              |                                          
              |           +-- Leoparreysia_tavoyensis    
              |  +--------+                              
              |  |        +-- Leoparreysia_olivacea      
              +--+                                       
                 |  +-------- Coelatura_aegyptiaca       
                 +--+                                    
                    |  +----- Indochinella_pugio         
                    +--+                                 
                       |  +-- Radiatula_chaudhurii       
                       +--+                              
                          +-- Scabies_mandarinus         

Pfeiffer et al., 2019 [Publ 3937], fig. 4-5

iT tT bi riS oT roT roS iD
3 3 1 3 54 53 2 5.0
                                   +----- Tamsiella_schroeteriana **      
  +--------------------------------+                                   
  |                                |  +-- Etheria_elliptica **            
  |                                +--+                                
  |                                   +-- Aspatharia_pfeifferiana **      
  |                                                                    
  |  +----------------------------------- Hyridella_australis          
  |  |                                                                 
--+  |                                +-- Margaritifera_margaritifera  
  |  |  +-----------------------------+                                
  |  |  |                             +-- Margaritifera_hembeli        
  |  |  |                                                              
  |  |  |                             +-- Quadrula_quadrula            
  |  |  |        +--------------------+                                
  |  |  |        |                    +-- Uniomerus_tetralasmus        
  +--+  |        |                                                     
     |  |        |                 +----- Elliptoideus_sloatianus      
     |  |  +-----+  +--------------+                                   
     |  |  |     |  |              |  +-- Elliptio_complanata          
     |  |  |     |  |              +--+                                
     |  |  |     |  |                 +-- Pleurobema_strodeanum        
     |  |  |     |  |                                                  
     |  |  |     +--+              +----- Reginaia_ebenus              
     |  |  |        |  +-----------+                                   
     |  |  |        |  |           |  +-- Plectomerus_dombeyanus       
     +--+  |        |  |           +--+                                
        |  |        |  |              +-- Amblema_plicata              
        |  |        +--+                                               
        |  |           |              +-- Psoronaias_semigranosa       
        |  |           |  +-----------+                                
        |  |           |  |           +-- Popenaias_popeii             
        |  |           +--+                                            
        |  |              |  +----------- Obliquaria_reflexa           
        |  |              +--+                                         
        |  |                 |  +-------- Cyrtonaias_tampicoensis      
        |  |                 +--+                                      
        |  |                    |  +----- Leaunio_lienosus             
        |  |                    +--+                                   
        |  |                       |  +-- Lampsilis_cardium            
        |  |                       +--+                                
        |  |                          +-- Pachynaias_spheniopsis       
        |  |                                                           
        +--+                          +-- Chamberlainia_hainesiana     
           |     +--------------------+                                
           |     |                    +-- Sinohyriopsis_cumingii       
           |     |                                                     
           |     |     +----------------- Lamprotula_cornuumlunae      
           |     |  +--+                                               
           |     |  |  |  +-------------- Gonidea_angulata             
           |  +--+  |  +--+                                            
           |  |  |  |     |  +----------- Indopseudodon_crebristriatus 
           |  |  |  |     +--+                                         
           |  |  |  |        |  +-------- Bineurus_anodontinum         
           |  |  |  |        +--+                                      
           |  |  |  |           |  +----- Pilsbryoconcha_exilis        
           |  |  |  |           +--+                                   
           |  |  +--+              |  +-- Pseudodon_vondembuschianus   
           |  |     |              +--+                                
           |  |     |                 +-- Pseudodon_inoscularis        
           |  |     |                                                  
           |  |     |              +----- Ensidens_ingallsianus        
           |  |     |        +-----+                                   
           |  |     |        |     |  +-- Hyriopsis_bialata            
           |  |     |        |     +--+                                
           |  |     |        |        +-- Rectidens_sumatrensis        
           |  |     +--------+                                         
           |  |              |        +-- Trapezoideus_foliaceus       
           |  |              |  +-----+                                
           +--+              |  |     +-- Physunio_superbus            
              |              +--+                                      
              |                 |  +----- Yaukthwa_paiensis            
              |                 +--+                                   
              |                    |  +-- Lens_contradens              
              |                    +--+                                
              |                       +-- Solenaia_emarginata          
              |                                                        
              |              +----------- Lanceolaria_lanceolata       
              |              |                                         
              |              |     +----- Unio_gibbus                  
              |     +--------+  +--+                                   
              |     |        |  |  |  +-- Nodularia_douglasiae         
              |     |        |  |  +--+                                
              |     |        |  |     +-- Tchangsinaia_piscicula       
              |     |        +--+                                      
              |     |           |     +-- Cristaria_plicata            
              |     |           |  +--+                                
              |     |           |  |  +-- Sinanodonta_woodiana         
              +-----+           +--+                                   
                    |              |  +-- Anodonta_cygnea              
                    |              +--+                                
                    |                 +-- Pyganodon_grandis            
                    |                                                  
                    |                 +-- Trapezidens_dolichorhynchus  
                    |  +--------------+                                
                    |  |              +-- Lamellidens_savadiensis      
                    |  |                                               
                    +--+              +-- Leoparreysia_tavoyensis      
                       |  +-----------+                                
                       |  |           +-- Leoparreysia_olivacea        
                       |  |                                            
                       +--+           +-- Coelatura_choziensis         
                          |  +--------+                                
                          |  |        +-- Prisodontopsis_aviculaeformis
                          +--+                                         
                             |  +-------- Indonaia_caerulea            
                             +--+                                      
                                |  +----- Indochinella_pugio           
                                +--+                                   
                                   |  +-- Scabies_phaselus             
                                   +--+                                
                                      +-- Harmandia_somboriensis       

Pfeiffer et al., 2019 [Publ 3937], fig. 6

iT tT bi riS oT roT roS iD
3 3 1 3 54 53 2 5.0
                          +----- Tamsiella_schroeteriana **      
  +-----------------------+                                   
  |                       |  +-- Etheria_elliptica **            
  |                       +--+                                
  |                          +-- Aspatharia_pfeifferiana **      
  |                                                           
  |                       +----- Hyridella_australis          
  |  +--------------------+                                   
  |  |                    |  +-- Margaritifera_margaritifera  
  |  |                    +--+                                
  |  |                       +-- Margaritifera_hembeli        
  |  |                                                        
  |  |                       +-- Quadrula_quadrula            
  |  |     +-----------------+                                
  |  |     |                 +-- Uniomerus_tetralasmus        
--+  |     |                                                  
  |  |     |              +----- Elliptoideus_sloatianus      
  |  |     |           +--+                                   
  |  |     |           |  |  +-- Elliptio_complanata          
  |  |  +--+           |  +--+                                
  |  |  |  |  +--------+     +-- Pleurobema_strodeanum        
  |  |  |  |  |        |                                      
  |  |  |  |  |        |  +----- Reginaia_ebenus              
  |  |  |  |  |        +--+                                   
  |  |  |  |  |           |  +-- Plectomerus_dombeyanus       
  |  |  |  +--+           +--+                                
  |  |  |     |              +-- Amblema_plicata              
  |  |  |     |                                               
  |  |  |     |              +-- Psoronaias_semigranosa       
  |  |  |     |  +-----------+                                
  +--+  |     |  |           +-- Popenaias_popeii             
     |  |     +--+                                            
     |  |        |  +----------- Obliquaria_reflexa           
     |  |        +--+                                         
     |  |           |  +-------- Cyrtonaias_tampicoensis      
     |  |           +--+                                      
     |  |              |  +----- Leaunio_lienosus             
     |  |              +--+                                   
     |  |                 |  +-- Lampsilis_cardium            
     |  |                 +--+                                
     |  |                    +-- Pachynaias_spheniopsis       
     |  |                                                     
     |  |                    +-- Chamberlainia_hainesiana     
     |  |  +-----------------+                                
     |  |  |                 +-- Sinohyriopsis_cumingii       
     |  |  |                                                  
     |  |  |                 +-- Gonidea_angulata             
     |  |  |     +-----------+                                
     |  |  |     |           +-- Lamprotula_cornuumlunae      
     |  |  |  +--+                                            
     |  +--+  |  |  +----------- Indopseudodon_crebristriatus 
     |  |  |  |  +--+                                         
     |  |  |  |     |  +-------- Pilsbryoconcha_exilis        
     |  |  |  |     +--+                                      
     +--+  |  |        |  +----- Bineurus_anodontinum         
        |  |  |        +--+                                   
        |  |  |           |  +-- Pseudodon_vondembuschianus   
        |  +--+           +--+                                
        |     |              +-- Pseudodon_inoscularis        
        |     |                                               
        |     |           +----- Ensidens_ingallsianus        
        |     |     +-----+                                   
        |     |     |     |  +-- Hyriopsis_bialata            
        |     |     |     +--+                                
        |     |     |        +-- Rectidens_sumatrensis        
        |     +-----+                                         
        |           |        +-- Trapezoideus_foliaceus       
        |           |  +-----+                                
        |           |  |     +-- Physunio_superbus            
        |           +--+                                      
        |              |  +----- Yaukthwa_paiensis            
        |              +--+                                   
        |                 |  +-- Lens_contradens              
        |                 +--+                                
        |                    +-- Solenaia_emarginata          
        |                                                     
        |              +-------- Lanceolaria_lanceolata       
        |           +--+                                      
        |           |  |  +----- Unio_gibbus                  
        |           |  +--+                                   
        |           |     |  +-- Tchangsinaia_piscicula       
        |           |     +--+                                
        |  +--------+        +-- Nodularia_douglasiae         
        |  |        |                                         
        |  |        |        +-- Cristaria_plicata            
        |  |        |     +--+                                
        |  |        |     |  +-- Sinanodonta_woodiana         
        |  |        +-----+                                   
        |  |              |  +-- Anodonta_cygnea              
        +--+              +--+                                
           |                 +-- Pyganodon_grandis            
           |                                                  
           |                 +-- Trapezidens_dolichorhynchus  
           |  +--------------+                                
           |  |              +-- Lamellidens_savadiensis      
           |  |                                               
           +--+              +-- Leoparreysia_tavoyensis      
              |  +-----------+                                
              |  |           +-- Leoparreysia_olivacea        
              |  |                                            
              +--+           +-- Coelatura_choziensis         
                 |  +--------+                                
                 |  |        +-- Prisodontopsis_aviculaeformis
                 +--+                                         
                    |  +-------- Indonaia_caerulea            
                    +--+                                      
                       |  +----- Indochinella_pugio           
                       +--+                                   
                          |  +-- Scabies_phaselus             
                          +--+                                
                             +-- Harmandia_somboriensis       

Elderkin et al., 2016 [Publ 3693], fig. 2

iT tT bi riS oT roT roS iD
2 3 0 2 1 1 1 5.0
  +-- Etheria_elliptica **      
  |                          
--+-- Aspatharia_pfeifferiana **
  |                          
  +-- Velesunio_ambiguus     

Guerra et al., 2017 [Publ 3772], fig. 2a

iT tT bi riS oT roT roS iD
2 2 0 2 23 22 2 --
                                +-- Mutela_dubia **               
  +-----------------------------+                              
  |                             +-- Anodontites_trapesialis **    
  |                                                            
  |  +----------------------------- Echyridella_menziesii      
--+  |                                                         
  |  |                          +-- Margaritifera_falcata      
  |  |  +-----------------------+                              
  |  |  |                       +-- Cumberlandia_monodonta     
  +--+  |                                                      
     |  |                 +-------- Quadrula_quadrula          
     |  |              +--+                                    
     |  |              |  |  +----- Toxolasma_parvum           
     |  |              |  +--+                                 
     |  |              |     |  +-- Venustaconcha_ellipsiformis
     +--+              |     +--+                              
        |  +-----------+        +-- Lampsilis_ornata           
        |  |           |                                       
        |  |           |        +-- Sinohyriopsis_schlegelii   
        |  |           |  +-----+                              
        |  |           |  |     +-- Sinohyriopsis_cumingii     
        |  |           +--+                                    
        |  |              |  +----- Pronodularia_japanensis    
        +--+              +--+                                 
           |                 |  +-- Sinosolenaia_oleivora      
           |                 +--+                              
           |                    +-- Sinosolenaia_carinata      
           |                                                   
           |                    +-- Aculamprotula_tortuosa     
           |  +-----------------+                              
           |  |                 +-- Unio_pictorum              
           +--+                                                
              |  +----------------- Sinanodonta_woodiana       
              |  |                                             
              +--+  +-------------- Cristaria_plicata          
                 |  |                                          
                 +--+  +----------- Anodonta_anatina           
                    |  |                                       
                    +--+        +-- Platynaias_subviridis      
                       |  +-----+                              
                       |  |     +-- Platynaias_compressa       
                       +--+                                    
                          |  +----- Pyganodon_grandis          
                          +--+                                 
                             |  +-- Utterbackia_peninsularis   
                             +--+                              
                                +-- Utterbackia_imbecillis     

Guerra et al., 2017 [Publ 3772], fig. 2b

iT tT bi riS oT roT roS iD
2 2 0 2 23 22 2 --
                                   +-- Mutela_dubia **               
  +--------------------------------+                              
  |                                +-- Anodontites_trapesialis **    
  |                                                               
--+  +-------------------------------- Echyridella_menziesii      
  |  |                                                            
  |  |                             +-- Margaritifera_falcata      
  |  |  +--------------------------+                              
  +--+  |                          +-- Cumberlandia_monodonta     
     |  |                                                         
     |  |                    +-------- Quadrula_quadrula          
     |  |  +-----------------+                                    
     +--+  |                 |  +----- Toxolasma_parvum           
        |  |                 +--+                                 
        |  |                    |  +-- Venustaconcha_ellipsiformis
        |  |                    +--+                              
        |  |                       +-- Lampsilis_ornata           
        |  |                                                      
        +--+                       +-- Sinohyriopsis_schlegelii   
           |                 +-----+                              
           |                 |     +-- Sinohyriopsis_cumingii     
           |  +--------------+                                    
           |  |              |  +----- Pronodularia_japanensis    
           |  |              +--+                                 
           |  |                 |  +-- Sinosolenaia_oleivora      
           +--+                 +--+                              
              |                    +-- Sinosolenaia_carinata      
              |                                                   
              |  +-------------------- Unio_pictorum              
              |  |                                                
              +--+  +----------------- Aculamprotula_tortuosa     
                 |  |                                             
                 +--+              +-- Cristaria_plicata          
                    |  +-----------+                              
                    |  |           +-- Sinanodonta_woodiana       
                    +--+                                          
                       |  +----------- Anodonta_anatina           
                       |  |                                       
                       +--+        +-- Platynaias_subviridis      
                          |  +-----+                              
                          |  |     +-- Platynaias_compressa       
                          +--+                                    
                             |  +----- Pyganodon_grandis          
                             +--+                                 
                                |  +-- Utterbackia_peninsularis   
                                +--+                              
                                   +-- Utterbackia_imbecillis     

Huang et al., 2018 [Publ 3828], fig. 6

iT tT bi riS oT roT roS iD
2 2 0 2 41 40 2 --
                             +-- Anodontites_trapesialis **    
  +--------------------------+                              
  |                          +-- Mutela_dubia **               
  |                                                         
  |  +-------------------------- Echyridella_menziesii      
  |  |                                                      
  |  |           +-------------- Gibbosula_rochechouartii   
--+  |  +--------+                                          
  |  |  |        |  +----------- Cumberlandia_monodonta     
  |  |  |        +--+                                       
  |  |  |           |  +-------- Pseudunio_marocanus        
  |  |  |           +--+                                    
  |  |  |              |  +----- Margaritifera_falcata      
  |  |  |              +--+                                 
  +--+  |                 |  +-- Margaritifera_margaritifera
     |  |                 +--+                              
     |  |                    +-- Margaritifera_dahurica     
     |  |                                                   
     |  |           +----------- Quadrula_quadrula          
     |  |           |                                       
     |  |     +-----+  +-------- Toxolasma_parvum           
     |  |     |     |  |                                    
     |  |     |     +--+     +-- Lampsilis_ornata           
     |  |     |        |  +--+                              
     |  |     |        |  |  +-- Venustaconcha_ellipsiformis
     +--+     |        +--+                                 
        |     |           |  +-- Potamilus_leptodon         
        |  +--+           +--+                              
        |  |  |              +-- Potamilus_alatus           
        |  |  |                                             
        |  |  |              +-- Sinohyriopsis_cumingii     
        |  |  |  +-----------+                              
        |  |  |  |           +-- Sinohyriopsis_schlegelii   
        |  |  |  |                                          
        |  |  +--+           +-- Sinosolenaia_oleivora      
        |  |     |  +--------+                              
        |  |     |  |        +-- Sinosolenaia_carinata      
        |  |     +--+                                       
        |  |        |  +-------- Potomida_littoralis        
        |  |        +--+                                    
        |  |           |  +----- Pronodularia_japanensis    
        |  |           +--+                                 
        |  |              |  +-- Lamprotula_caveata         
        +--+              +--+                              
           |                 +-- Lamprotula_leaii           
           |                                                
           |              +----- Aculamprotula_tortuosa     
           |     +--------+                                 
           |     |        |  +-- Aculamprotula_coreana      
           |     |        +--+                              
           |     |           +-- Aculamprotula_tientsinensis
           |  +--+                                          
           |  |  |           +-- Tchangsinaia_piscicula     
           |  |  |  +--------+                              
           |  |  |  |        +-- Nodularia_douglasiae       
           |  |  +--+                                       
           |  |     |  +-------- Unio_tumidus               
           |  |     +--+                                    
           |  |        |  +----- Unio_crassus               
           |  |        +--+                                 
           +--+           |  +-- Unio_delphinus             
              |           +--+                              
              |              +-- Unio_pictorum              
              |                                             
              |     +----------- Anodonta_anatina           
              |  +--+                                       
              |  |  |  +-------- Platynaias_compressa       
              |  |  +--+                                    
              |  |     |  +----- Pyganodon_grandis          
              |  |     +--+                                 
              +--+        |  +-- Utterbackia_imbecillis     
                 |        +--+                              
                 |           +-- Utterbackia_peninsularis   
                 |                                          
                 |     +-------- Cristaria_plicata          
                 |     |                                    
                 +-----+     +-- Anemina_arcaeformis        
                       |  +--+                              
                       |  |  +-- Anemina_euscaphys          
                       +--+                                 
                          |  +-- Sinanodonta_woodiana       
                          +--+                              
                             +-- Sinanodonta_lucida         

Huang et al., 2019 [Publ 3843], fig. 3

iT tT bi riS oT roT roS iD
2 2 0 2 55 54 2 --
                                         +-- Anodontites_trapesialis **    
  +--------------------------------------+                              
  |                                      +-- Mutela_dubia **               
  |                                                                     
  |  +-------------------------------------- Echyridella_menziesii      
  |  |                                                                  
  |  |                       +-------------- Gibbosula_rochechouartii   
  |  |  +--------------------+                                          
--+  |  |                    |  +----------- Cumberlandia_monodonta     
  |  |  |                    +--+                                       
  |  |  |                       |  +-------- Pseudunio_marocanus        
  |  |  |                       +--+                                    
  |  |  |                          |  +----- Margaritifera_falcata      
  |  |  |                          +--+                                 
  |  |  |                             |  +-- Margaritifera_margaritifera
  |  |  |                             +--+                              
  +--+  |                                +-- Margaritifera_dahurica     
     |  |                                                               
     |  |                                +-- Sinohyriopsis_cumingii     
     |  |                    +-----------+                              
     |  |                    |           +-- Sinohyriopsis_schlegelii   
     |  |                    |                                          
     |  |                    |     +-------- Potomida_littoralis        
     |  |           +--------+  +--+                                    
     |  |           |        |  |  |  +----- Pronodularia_japanensis    
     |  |           |        |  |  +--+                                 
     |  |           |        |  |     |  +-- Lamprotula_caveata         
     |  |           |        |  |     +--+                              
     +--+           |        +--+        +-- Lamprotula_leaii           
        |           |           |                                       
        |           |           |        +-- Ptychorhynchus_pfisteri    
        |           |           |     +--+                              
        |  +--------+           |     |  +-- Parvasolenaia_rivularis    
        |  |        |           +-----+                                 
        |  |        |                 |  +-- Sinosolenaia_oleivora      
        |  |        |                 +--+                              
        |  |        |                    +-- Sinosolenaia_carinata      
        |  |        |                                                   
        |  |        |                    +-- Quadrula_quadrula          
        |  |        |  +-----------------+                              
        |  |        |  |                 +-- Uniomerus_tetralasmus      
        |  |        +--+                                                
        |  |           |  +----------------- Elliptio_complanata        
        |  |           |  |                                             
        |  |           +--+  +-------------- Toxolasma_parvum           
        |  |              |  |                                          
        |  |              +--+           +-- Potamilus_leptodon         
        |  |                 |  +--------+                              
        |  |                 |  |        +-- Potamilus_alatus           
        +--+                 +--+                                       
           |                    |  +-------- Leaunio_lienosus           
           |                    +--+                                    
           |                       |  +----- Venustaconcha_ellipsiformis
           |                       +--+                                 
           |                          |  +-- Lampsilis_cardium          
           |                          +--+                              
           |                             +-- Lampsilis_ornata           
           |                                                            
           |  +----------------------------- Lepidodesma_languilati     
           |  |                                                         
           |  |                    +-------- Aculamprotula_tortuosa     
           |  |              +-----+                                    
           |  |              |     |  +----- Aculamprotula_scripta      
           |  |              |     +--+                                 
           |  |              |        |  +-- Aculamprotula_tientsinensis
           |  |              |        +--+                              
           |  |  +-----------+           +-- Aculamprotula_coreana      
           +--+  |           |                                          
              |  |           |        +----- Nodularia_douglasiae       
              |  |           |  +-----+                                 
              |  |           |  |     |  +-- Tchangsinaia_piscicula     
              |  |           |  |     +--+                              
              |  |           +--+        +-- Schistodesmus_lampreyanus  
              |  |              |                                       
              |  |              |  +-------- Unio_tumidus               
              |  |              +--+                                    
              +--+                 |  +----- Unio_crassus               
                 |                 +--+                                 
                 |                    |  +-- Unio_delphinus             
                 |                    +--+                              
                 |                       +-- Unio_pictorum              
                 |                                                      
                 |                    +----- Lanceolaria_gladiola       
                 |  +-----------------+                                 
                 |  |                 |  +-- Lanceolaria_lanceolata     
                 |  |                 +--+                              
                 |  |                    +-- Lanceolaria_grayii         
                 +--+                                                   
                    |  +-------------------- Acuticosta_chinensis       
                    |  |                                                
                    |  |           +-------- Cristaria_plicata          
                    |  |           |                                    
                    +--+  +--------+     +-- Sinanodonta_lucida         
                       |  |        |  +--+                              
                       |  |        |  |  +-- Sinanodonta_woodiana       
                       |  |        +--+                                 
                       +--+           |  +-- Anemina_arcaeformis        
                          |           +--+                              
                          |              +-- Anemina_euscaphys          
                          |                                             
                          |  +-------------- Anodonta_cygnea            
                          +--+                                          
                             |  +----------- Anodonta_anatina           
                             +--+                                       
                                |  +-------- Lasmigona_costata          
                                +--+                                    
                                   |  +----- Pyganodon_grandis          
                                   +--+                                 
                                      |  +-- Utterbackia_imbecillis     
                                      +--+                              
                                         +-- Utterbackia_peninsularis   

Lemer et al., 2019 [Publ 4093], fig. 1

iT tT bi riS oT roT roS iD
2 2 0 2 4 2 1 --
     +-- Etheria_elliptica **          
  +--+                              
  |  +-- Aspatharia_pfeifferiana **    
--+                                 
  |  +-- Margaritifera_margaritifera
  +--+                              
     +-- Lampsilis_cardium          

Wu et al., 2020 [Publ 4121], fig. 3

iT tT bi riS oT roT roS iD
2 2 0 2 48 47 2 --
  +-------------------------------------- Echyridella_menziesii      
  |                                                                  
  |                                   +-- Anodontites_trapesialis **    
  |  +--------------------------------+                              
  |  |                                +-- Mutela_dubia **               
--+  |                                                               
  |  |                          +-------- Gibbosula_rochechouartii   
  |  |  +-----------------------+                                    
  |  |  |                       |  +----- Cumberlandia_monodonta     
  |  |  |                       +--+                                 
  +--+  |                          |  +-- Margaritifera_falcata      
     |  |                          +--+                              
     |  |                             +-- Margaritifera_dahurica     
     |  |                                                            
     |  |                    +----------- Quadrula_quadrula          
     |  |                    |                                       
     |  |  +-----------------+  +-------- Toxolasma_parvum           
     +--+  |                 |  |                                    
        |  |                 +--+     +-- Venustaconcha_ellipsiformis
        |  |                    |  +--+                              
        |  |                    |  |  +-- Lampsilis_ornata           
        |  |                    +--+                                 
        |  |                       |  +-- Potamilus_alatus           
        |  |                       +--+                              
        |  |                          +-- Potamilus_leptodon         
        |  |                                                         
        |  |                          +-- Sinohyriopsis_schlegelii   
        |  |                 +--------+                              
        +--+                 |        +-- Sinohyriopsis_cumingii     
           |                 |                                       
           |  +--------------+     +----- Potomida_littoralis        
           |  |              |  +--+                                 
           |  |              |  |  |  +-- Pronodularia_japanensis    
           |  |              |  |  +--+                              
           |  |              +--+     +-- Lamprotula_leaii           
           |  |                 |                                    
           |  |                 |  +----- Ptychorhynchus_pfisteri    
           |  |                 +--+                                 
           |  |                    |  +-- Sinosolenaia_oleivora      
           |  |                    +--+                              
           |  |                       +-- Sinosolenaia_carinata      
           +--+                                                      
              |  +----------------------- Lepidodesma_languilati     
              |  |                                                   
              |  |                 +----- Acuticosta_chinensis       
              |  |     +-----------+                                 
              |  |     |           |  +-- Lanceolaria_lanceolata     
              |  |     |           +--+                              
              |  |     |              +-- Lanceolaria_grayii         
              |  |     |                                             
              |  |  +--+     +----------- Anodonta_anatina           
              |  |  |  |  +--+                                       
              |  |  |  |  |  |  +-------- Platynaias_compressa       
              +--+  |  |  |  +--+                                    
                 |  |  |  |     |  +----- Pyganodon_grandis          
                 |  |  |  |     +--+                                 
                 |  |  +--+        |  +-- Utterbackia_peninsularis   
                 |  |     |        +--+                              
                 |  |     |           +-- Utterbackia_imbecillis     
                 |  |     |                                          
                 |  |     |     +-------- Cristaria_plicata          
                 |  |     |     |                                    
                 |  |     +-----+     +-- Sinanodonta_woodiana       
                 |  |           |  +--+                              
                 +--+           |  |  +-- Sinanodonta_lauta          
                    |           +--+                                 
                    |              |  +-- Anemina_arcaeformis        
                    |              +--+                              
                    |                 +-- Anemina_euscaphys          
                    |                                                
                    |           +-------- Aculamprotula_tortuosa     
                    |           |                                    
                    |  +--------+     +-- Aculamprotula_coreana      
                    |  |        |  +--+                              
                    |  |        |  |  +-- Aculamprotula_tientsinensis
                    |  |        +--+                                 
                    |  |           |  +-- Aculamprotula_polysticta   
                    |  |           +--+                              
                    +--+              +-- Aculamprotula_scripta      
                       |                                             
                       |        +-------- Unio_tumidus               
                       |  +-----+                                    
                       |  |     |  +----- Unio_crassus               
                       |  |     +--+                                 
                       |  |        |  +-- Unio_delphinus             
                       +--+        +--+                              
                          |           +-- Unio_pictorum              
                          |                                          
                          |  +----------- Pseudocuneopsis_capitata   
                          +--+                                       
                             |  +-------- Nodularia_douglasiae       
                             +--+                                    
                                |  +----- Tchangsinaia_piscicula     
                                +--+                                 
                                   |  +-- Cuneopsis_heudei           
                                   +--+                              
                                      +-- Schistodesmus_lampreyanus  

Wu et al., 2022 [Publ 4238], fig. 3

iT tT bi riS oT roT roS iD
2 2 0 2 72 72 2 --
                                               +-- Mutela_dubia **                
  +--------------------------------------------+                               
  |                                            +-- Anodontites_trapesialis **     
  |                                                                            
  |  +-------------------------------------------- Echyridella_menziesii       
  |  |                                                                         
  |  |                             +-------------- Gibbosula_rochechouartii    
  |  |  +--------------------------+                                           
--+  |  |                          |  +----------- Cumberlandia_monodonta      
  |  |  |                          +--+                                        
  |  |  |                             |  +-------- Pseudunio_marocanus         
  |  |  |                             +--+                                     
  |  |  |                                |  +----- Margaritifera_falcata       
  |  |  |                                +--+                                  
  |  |  |                                   |  +-- Margaritifera_dahurica      
  +--+  |                                   +--+                               
     |  |                                      +-- Margaritifera_margaritifera 
     |  |                                                                      
     |  |                                      +-- Uniomerus_tetralasmus       
     |  |           +--------------------------+                               
     |  |           |                          +-- Quadrula_quadrula           
     |  |           |                                                          
     |  |  +--------+                          +-- Pleurobema_oviforme         
     |  |  |        |  +-----------------------+                               
     |  |  |        |  |                       +-- Elliptio_complanata         
     |  |  |        +--+                                                       
     +--+  |           |  +----------------------- Amblema_plicata             
        |  |           |  |                                                    
        |  |           +--+  +-------------------- Popenaias_popeii            
        |  |              |  |                                                 
        |  |              +--+  +----------------- Toxolasma_parvum            
        |  |                 |  |                                              
        |  |                 +--+              +-- Potamilus_alatus            
        |  |                    |  +-----------+                               
        |  |                    |  |           +-- Potamilus_leptodon          
        |  |                    +--+                                           
        |  |                       |  +----------- Leaunio_lienosus            
        |  |                       |  |                                        
        |  |                       +--+        +-- Lampsilis_cardium           
        |  |                          |  +-----+                               
        |  |                          |  |     +-- Lampsilis_ornata            
        |  |                          +--+                                     
        |  |                             |  +----- Venustaconcha_ellipsiformis 
        |  |                             +--+                                  
        |  |                                |  +-- Lampsilis_siliquoidea       
        |  |                                +--+                               
        +--+                                   +-- Lampsilis_powellii          
           |                                                                   
           |                                +----- Chamberlainia_somsakpanhai  
           |                    +-----------+                                  
           |                    |           |  +-- Sinohyriopsis_schlegelii    
           |                    |           +--+                               
           |                    |              +-- Sinohyriopsis_cumingii      
           |                    |                                              
           |  +-----------------+        +-------- Potomida_littoralis         
           |  |                 |  +-----+                                     
           |  |                 |  |     |  +----- Pronodularia_japanensis     
           |  |                 |  |     +--+                                  
           |  |                 |  |        |  +-- Lamprotula_caveata          
           |  |                 |  |        +--+                               
           |  |                 +--+           +-- Lamprotula_leaii            
           |  |                    |                                           
           |  |                    |           +-- Pilsbryoconcha_exilis       
           |  |                    |  +--------+                               
           |  |                    |  |        +-- Pseudodon_vondembuschianus  
           |  |                    |  |                                        
           |  |                    +--+        +-- Ptychorhynchus_pfisteri     
           |  |                       |  +-----+                               
           |  |                       |  |     +-- Parvasolenaia_rivularis     
           |  |                       +--+                                     
           |  |                          |  +----- Microcondylaea_bonellii     
           +--+                          +--+                                  
              |                             |  +-- Sinosolenaia_carinata       
              |                             +--+                               
              |                                +-- Sinosolenaia_recognita      
              |                                                                
              |  +-------------------------------- Lepidodesma_languilati      
              |  |                                                             
              |  |                       +-------- Acuticosta_chinensis        
              |  |              +--------+                                     
              |  |              |        |  +----- Lanceolaria_gladiola        
              |  |              |        +--+                                  
              |  |              |           |  +-- Lanceolaria_grayii          
              |  |              |           +--+                               
              |  |              |              +-- Lanceolaria_lanceolata      
              |  |              |                                              
              |  |  +-----------+        +-------- Cristaria_plicata           
              |  |  |           |        |                                     
              +--+  |           |  +-----+     +-- Sinanodonta_woodiana        
                 |  |           |  |     |  +--+                               
                 |  |           |  |     |  |  +-- Sinanodonta_lucida          
                 |  |           |  |     +--+                                  
                 |  |           |  |        |  +-- Anemina_arcaeformis         
                 |  |           +--+        +--+                               
                 |  |              |           +-- Anemina_euscaphys           
                 |  |              |                                           
                 |  |              |           +-- Anodonta_cygnea             
                 |  |              |  +--------+                               
                 |  |              |  |        +-- Anodonta_anatina            
                 |  |              +--+                                        
                 |  |                 |  +-------- Platynaias_compressa        
                 +--+                 +--+                                     
                    |                    |  +----- Pyganodon_grandis           
                    |                    +--+                                  
                    |                       |  +-- Utterbackia_imbecillis      
                    |                       +--+                               
                    |                          +-- Utterbackia_peninsularis    
                    |                                                          
                    |                    +-------- Aculamprotula_tortuosa      
                    |                    |                                     
                    |  +-----------------+     +-- Aculamprotula_tientsinensis 
                    |  |                 |  +--+                               
                    |  |                 |  |  +-- Aculamprotula_coreana       
                    |  |                 +--+                                  
                    |  |                    |  +-- Aculamprotula_polysticta    
                    |  |                    +--+                               
                    |  |                       +-- Aculamprotula_scripta       
                    +--+                                                       
                       |                 +-------- Unio_tumidus                
                       |  +--------------+                                     
                       |  |              |  +----- Unio_crassus                
                       |  |              +--+                                  
                       |  |                 |  +-- Unio_pictorum               
                       |  |                 +--+                               
                       +--+                    +-- Unio_delphinus              
                          |                                                    
                          |                    +-- Pseudocuneopsis_capitata    
                          |  +-----------------+                               
                          |  |                 +-- Pseudocuneopsis_sichuanensis
                          |  |                                                 
                          +--+                 +-- Nodularia_douglasiae        
                             |  +--------------+                               
                             |  |              +-- Nodularia_breviconcha       
                             |  |                                              
                             +--+              +-- Schistodesmus_lampreyanus   
                                |  +-----------+                               
                                |  |           +-- Schistodesmus_spinosus      
                                +--+                                           
                                   |  +----------- Tchangsinaia_piscicula      
                                   +--+                                        
                                      |  +-------- Cuneopsis_rufescens         
                                      +--+                                     
                                         |  +----- Cuneopsis_celtiformis       
                                         +--+                                  
                                            |  +-- Cuneopsis_heudei            
                                            +--+                               
                                               +-- Cuneopsis_demangei          

Smith, 2021 [Publ 4331], fig. 1

iT tT bi riS oT roT roS iD
2 2 0 2 18 17 1 --
                          +-- Etheria_elliptica **          
  +-----------------------+                              
  |                       +-- Aspatharia_pfeifferiana **    
  |                                                      
  |  +----------------------- Margaritifera_margaritifera
--+  |                                                   
  |  |                 +----- Sinohyriopsis_cumingii     
  |  |           +-----+                                 
  |  |           |     |  +-- Lens_contradens            
  |  |           |     +--+                              
  +--+  +--------+        +-- Lamprotula_leaii           
     |  |        |                                       
     |  |        |  +-------- Scabies_phaselus           
     |  |        +--+                                    
     |  |           |  +----- Cristaria_plicata          
     |  |           +--+                                 
     |  |              |  +-- Pyganodon_grandis          
     +--+              +--+                              
        |                 +-- Utterbackia_peninsularis   
        |                                                
        |                 +-- Uniomerus_tetralasmus      
        |  +--------------+                              
        |  |              +-- Megalonaias_nervosa        
        |  |                                             
        +--+              +-- Elliptio_crassidens        
           |  +-----------+                              
           |  |           +-- Elliptio_complanata        
           +--+                                          
              |  +----------- Amblema_plicata            
              +--+                                       
                 |  +-------- Potamilus_streckersoni     
                 +--+                                    
                    |  +----- Venustaconcha_ellipsiformis
                    +--+                                 
                       |  +-- Lampsilis_siliquoidea      
                       +--+                              
                          +-- Lampsilis_cardium          

Wu et al., 2023 [Publ 4339], fig. 4, S5-aa

iT tT bi riS oT roT roS iD
2 2 0 2 94 93 2 --
  +----------------------------------------------- Echyridella_menziesii       
  |                                                                            
  |                                            +-- Anodontites_trapesialis **     
  |  +-----------------------------------------+                               
  |  |                                         +-- Mutela_dubia **                
  |  |                                                                         
  |  |                                         +-- Gibbosula_crassa            
  |  |                             +-----------+                               
--+  |                             |           +-- Gibbosula_rochechouartii    
  |  |  +--------------------------+                                           
  |  |  |                          |  +----------- Cumberlandia_monodonta      
  |  |  |                          +--+                                        
  |  |  |                             |  +-------- Pseudunio_marocanus         
  |  |  |                             +--+                                     
  |  |  |                                |  +----- Margaritifera_falcata       
  |  |  |                                +--+                                  
  |  |  |                                   |  +-- Margaritifera_dahurica      
  +--+  |                                   +--+                               
     |  |                                      +-- Margaritifera_margaritifera 
     |  |                                                                      
     |  |                                      +-- Quadrula_quadrula           
     |  |           +--------------------------+                               
     |  |           |                          +-- Uniomerus_tetralasmus       
     |  |           |                                                          
     |  |  +--------+                          +-- Elliptio_complanata         
     |  |  |        |  +-----------------------+                               
     |  |  |        |  |                       +-- Pleurobema_oviforme         
     |  |  |        +--+                                                       
     |  |  |           |  +----------------------- Amblema_plicata             
     |  |  |           |  |                                                    
     |  |  |           +--+  +-------------------- Popenaias_popeii            
     +--+  |              |  |                                                 
        |  |              +--+  +----------------- Toxolasma_parvum            
        |  |                 |  |                                              
        |  |                 |  |           +----- Potamilus_leptodon          
        |  |                 +--+  +--------+                                  
        |  |                    |  |        |  +-- Potamilus_alatus            
        |  |                    |  |        +--+                               
        |  |                    |  |           +-- Potamilus_streckersoni      
        |  |                    +--+                                           
        |  |                       |  +----------- Leaunio_lienosus            
        |  |                       |  |                                        
        |  |                       |  |     +----- Lampsilis_ornata            
        |  |                       +--+  +--+                                  
        |  |                          |  |  |  +-- Lampsilis_cardium           
        |  |                          |  |  +--+                               
        |  |                          +--+     +-- Lampsilis_cariosa           
        |  |                             |                                     
        |  |                             |  +----- Venustaconcha_ellipsiformis 
        |  |                             +--+                                  
        |  |                                |  +-- Lampsilis_powellii          
        |  |                                +--+                               
        |  |                                   +-- Lampsilis_siliquoidea       
        +--+                                                                   
           |                                +----- Chamberlainia_somsakpanhai  
           |              +-----------------+                                  
           |              |                 |  +-- Sinohyriopsis_cumingii      
           |              |                 +--+                               
           |              |                    +-- Sinohyriopsis_schlegelii    
           |              |                                                    
           |              |                    +-- Hyriopsis_bialata           
           |              |                 +--+                               
           |  +-----------+                 |  +-- Rectidens_sumatrensis       
           |  |           |  +--------------+                                  
           |  |           |  |              |  +-- Lens_contradens             
           |  |           |  |              +--+                               
           |  |           |  |                 +-- Physunio_superbus           
           |  |           |  |                                                 
           |  |           |  |        +----------- Potomida_littoralis         
           |  |           +--+  +-----+                                        
           |  |              |  |     |  +-------- Pronodularia_japanensis     
           |  |              |  |     +--+                                     
           |  |              |  |        |  +----- Lamprotula_leaii            
           |  |              |  |        +--+                                  
           |  |              |  |           |  +-- Lamprotula_caveata          
           |  |              +--+           +--+                               
           |  |                 |              +-- Lamprotula_cornuumlunae     
           |  |                 |                                              
           |  |                 |              +-- Pseudodon_vondembuschianus  
           |  |                 |  +-----------+                               
           |  |                 |  |           +-- Pilsbryoconcha_exilis       
           |  |                 +--+                                           
           |  |                    |  +----------- Microcondylaea_bonellii     
           +--+                    |  |                                        
              |                    +--+        +-- Parvasolenaia_rivularis     
              |                       |  +-----+                               
              |                       |  |     +-- Ptychorhynchus_pfisteri     
              |                       +--+                                     
              |                          |  +----- Sinosolenaia_carinata       
              |                          +--+                                  
              |                             |  +-- Sinosolenaia_iridinea       
              |                             +--+                               
              |                                +-- Sinosolenaia_oleivora       
              |                                                                
              |  +-------------------------------- Lamellidens_jenkinsianus    
              |  |                                                             
              |  |                             +-- Lepidodesma_aligera         
              |  |  +--------------------------+                               
              |  |  |                          +-- Lepidodesma_languilati      
              |  |  |                                                          
              |  |  |                    +-------- Aculamprotula_tortuosa      
              |  |  |                    |                                     
              |  |  |     +--------------+     +-- Aculamprotula_coreana       
              |  |  |     |              |  +--+                               
              +--+  |     |              |  |  +-- Aculamprotula_tientsinensis 
                 |  |     |              +--+                                  
                 |  |     |                 |  +-- Aculamprotula_polysticta    
                 |  |     |                 +--+                               
                 |  |     |                    +-- Aculamprotula_scripta       
                 |  |     |                                                    
                 |  |  +--+        +-------------- Unio_tumidus                
                 |  |  |  |  +-----+                                           
                 |  |  |  |  |     |  +----------- Unio_crassus                
                 |  |  |  |  |     +--+                                        
                 +--+  |  |  |        |  +-------- Unio_elongatulus            
                    |  |  |  |        +--+                                     
                    |  |  |  |           |  +----- Unio_pictorum               
                    |  |  |  |           +--+                                  
                    |  |  |  |              |  +-- Unio_delphinus              
                    |  |  +--+              +--+                               
                    |  |     |                 +-- Unio_mancus                 
                    |  |     |                                                 
                    |  |     |                 +-- Nodularia_breviconcha       
                    |  |     |              +--+                               
                    |  |     |              |  +-- Nodularia_nipponensis       
                    |  |     |  +-----------+                                  
                    |  |     |  |           |  +-- Pseudocuneopsis_capitata    
                    |  |     |  |           +--+                               
                    |  |     +--+              +-- Pseudocuneopsis_sichuanensis
                    |  |        |                                              
                    |  |        |  +-------------- Tchangsinaia_piscicula      
                    |  |        |  |                                           
                    +--+        +--+           +-- Schistodesmus_lampreyanus   
                       |           |  +--------+                               
                       |           |  |        +-- Schistodesmus_spinosus      
                       |           +--+                                        
                       |              |  +-------- Cuneopsis_rufescens         
                       |              +--+                                     
                       |                 |  +----- Cuneopsis_celtiformis       
                       |                 +--+                                  
                       |                    |  +-- Cuneopsis_demangei          
                       |                    +--+                               
                       |                       +-- Cuneopsis_heudei            
                       |                                                       
                       |                 +-------- Acuticosta_chinensis        
                       |        +--------+                                     
                       |        |        |  +----- Lanceolaria_gladiola        
                       |        |        +--+                                  
                       |        |           |  +-- Lanceolaria_grayii          
                       |        |           +--+                               
                       |        |              +-- Lanceolaria_lanceolata      
                       |        |                                              
                       |        |           +----- Cristaria_plicata           
                       |        |     +-----+                                  
                       |        |     |     |  +-- Anemina_arcaeformis         
                       +--------+     |     +--+                               
                                |  +--+        +-- Anemina_euscaphys           
                                |  |  |                                        
                                |  |  |  +-------- Beringiana_fukuharai        
                                |  |  +--+                                     
                                |  |     |  +----- Sinanodonta_lucida          
                                |  |     +--+                                  
                                |  |        |  +-- Sinanodonta_tumens          
                                |  |        +--+                               
                                |  |           +-- Sinanodonta_woodiana        
                                +--+                                           
                                   |           +-- Anodonta_nuttalliana        
                                   |     +-----+                               
                                   |     |     +-- Pseudanodonta_complanata    
                                   |  +--+                                     
                                   |  |  |  +----- Anodonta_anatina            
                                   |  |  +--+                                  
                                   |  |     |  +-- Anodonta_cygnea             
                                   |  |     +--+                               
                                   |  |        +-- Anodonta_exulcerata         
                                   +--+                                        
                                      |     +----- Pyganodon_grandis           
                                      |  +--+                                  
                                      |  |  |  +-- Utterbackia_peninsularis    
                                      |  |  +--+                               
                                      |  |     +-- Utterbackia_imbecillis      
                                      +--+                                     
                                         |     +-- Alasmidonta_varicosa        
                                         |  +--+                               
                                         |  |  +-- Platynaias_compressa        
                                         +--+                                  
                                            |  +-- Prolasmidonta_heterodon     
                                            +--+                               
                                               +-- Lasmigona_complanata        

Wu et al., 2023 [Publ 4339], fig. 4, S5-nt

iT tT bi riS oT roT roS iD
2 2 0 2 95 94 2 --
  +-------------------------------------------------- Echyridella_menziesii       
  |                                                                               
  |                                               +-- Anodontites_trapesialis **     
  |  +--------------------------------------------+                               
  |  |                                            +-- Mutela_dubia **                
  |  |                                                                            
  |  |                                            +-- Gibbosula_crassa            
  |  |                                +-----------+                               
--+  |                                |           +-- Gibbosula_rochechouartii    
  |  |  +-----------------------------+                                           
  |  |  |                             |  +----------- Cumberlandia_monodonta      
  |  |  |                             +--+                                        
  |  |  |                                |  +-------- Pseudunio_marocanus         
  |  |  |                                +--+                                     
  |  |  |                                   |  +----- Margaritifera_falcata       
  |  |  |                                   +--+                                  
  |  |  |                                      |  +-- Margaritifera_dahurica      
  +--+  |                                      +--+                               
     |  |                                         +-- Margaritifera_margaritifera 
     |  |                                                                         
     |  |                                         +-- Quadrula_quadrula           
     |  |              +--------------------------+                               
     |  |              |                          +-- Uniomerus_tetralasmus       
     |  |              |                                                          
     |  |  +-----------+                          +-- Elliptio_complanata         
     |  |  |           |  +-----------------------+                               
     |  |  |           |  |                       +-- Pleurobema_oviforme         
     |  |  |           +--+                                                       
     |  |  |              |  +----------------------- Amblema_plicata             
     |  |  |              |  |                                                    
     |  |  |              +--+  +-------------------- Popenaias_popeii            
     +--+  |                 |  |                                                 
        |  |                 +--+  +----------------- Toxolasma_parvum            
        |  |                    |  |                                              
        |  |                    |  |           +----- Potamilus_leptodon          
        |  |                    +--+  +--------+                                  
        |  |                       |  |        |  +-- Potamilus_alatus            
        |  |                       |  |        +--+                               
        |  |                       |  |           +-- Potamilus_streckersoni      
        |  |                       +--+                                           
        |  |                          |  +----------- Leaunio_lienosus            
        |  |                          |  |                                        
        |  |                          |  |     +----- Lampsilis_ornata            
        |  |                          +--+  +--+                                  
        |  |                             |  |  |  +-- Lampsilis_cardium           
        |  |                             |  |  +--+                               
        |  |                             +--+     +-- Lampsilis_cariosa           
        |  |                                |                                     
        |  |                                |  +----- Venustaconcha_ellipsiformis 
        |  |                                +--+                                  
        |  |                                   |  +-- Lampsilis_powellii          
        |  |                                   +--+                               
        |  |                                      +-- Lampsilis_siliquoidea       
        +--+                                                                      
           |                                   +----- Chamberlainia_somsakpanhai  
           |                 +-----------------+                                  
           |                 |                 |  +-- Sinohyriopsis_cumingii      
           |                 |                 +--+                               
           |                 |                    +-- Sinohyriopsis_schlegelii    
           |                 |                                                    
           |                 |                    +-- Hyriopsis_bialata           
           |                 |                 +--+                               
           |  +--------------+                 |  +-- Rectidens_sumatrensis       
           |  |              |  +--------------+                                  
           |  |              |  |              |  +-- Lens_contradens             
           |  |              |  |              +--+                               
           |  |              |  |                 +-- Physunio_superbus           
           |  |              |  |                                                 
           |  |              |  |        +----------- Potomida_littoralis         
           |  |              +--+  +-----+                                        
           |  |                 |  |     |  +-------- Pronodularia_japanensis     
           |  |                 |  |     +--+                                     
           |  |                 |  |        |  +----- Lamprotula_leaii            
           |  |                 |  |        +--+                                  
           |  |                 |  |           |  +-- Lamprotula_caveata          
           |  |                 +--+           +--+                               
           |  |                    |              +-- Lamprotula_cornuumlunae     
           |  |                    |                                              
           |  |                    |              +-- Pseudodon_vondembuschianus  
           |  |                    |  +-----------+                               
           |  |                    |  |           +-- Pilsbryoconcha_exilis       
           |  |                    |  |                                           
           |  |                    +--+           +-- Ptychorhynchus_pfisteri     
           +--+                       |  +--------+                               
              |                       |  |        +-- Microcondylaea_bonellii     
              |                       +--+                                        
              |                          |  +-------- Parvasolenaia_rivularis     
              |                          +--+                                     
              |                             |  +----- Sinosolenaia_carinata       
              |                             +--+                                  
              |                                |  +-- Sinosolenaia_iridinea       
              |                                +--+                               
              |                                   +-- Sinosolenaia_oleivora       
              |                                                                   
              |  +----------------------------------- Lamellidens_jenkinsianus    
              |  |                                                                
              |  |                                +-- Lepidodesma_aligera         
              |  |  +-----------------------------+                               
              |  |  |                             +-- Lepidodesma_languilati      
              |  |  |                                                             
              |  |  |                       +-------- Acuticosta_chinensis        
              |  |  |        +--------------+                                     
              |  |  |        |              |  +----- Lanceolaria_gladiola        
              |  |  |        |              +--+                                  
              +--+  |        |                 |  +-- Lanceolaria_grayii          
                 |  |        |                 +--+                               
                 |  |        |                    +-- Lanceolaria_lanceolata      
                 |  |        |                                                    
                 |  |        |        +-------------- Cristaria_plicata           
                 |  |  +-----+        |                                           
                 |  |  |     |  +-----+           +-- Anemina_arcaeformis         
                 |  |  |     |  |     |  +--------+                               
                 |  |  |     |  |     |  |        +-- Anemina_euscaphys           
                 |  |  |     |  |     +--+                                        
                 +--+  |     |  |        |  +-------- Beringiana_fukuharai        
                    |  |     |  |        +--+                                     
                    |  |     |  |           |  +----- Sinanodonta_lucida          
                    |  |     |  |           +--+                                  
                    |  |     +--+              |  +-- Sinanodonta_tumens          
                    |  |        |              +--+                               
                    |  |        |                 +-- Sinanodonta_woodiana        
                    |  |        |                                                 
                    |  |        |  +----------------- Anodonta_nuttalliana        
                    |  |        |  |                                              
                    |  |        |  +----------------- Pseudanodonta_complanata    
                    |  |        |  |                                              
                    |  |        +--+  +-------------- Anodonta_cygnea             
                    |  |           |  |                                           
                    |  |           |  |           +-- Anodonta_exulcerata         
                    |  |           |  +-----------+                               
                    |  |           |  |           +-- Anodonta_anatina            
                    |  |           +--+                                           
                    +--+              |        +----- Pyganodon_grandis           
                       |              |  +-----+                                  
                       |              |  |     |  +-- Utterbackia_imbecillis      
                       |              |  |     +--+                               
                       |              +--+        +-- Utterbackia_peninsularis    
                       |                 |                                        
                       |                 |  +-------- Lasmigona_complanata        
                       |                 +--+                                     
                       |                    |  +----- Prolasmidonta_heterodon     
                       |                    +--+                                  
                       |                       |  +-- Alasmidonta_varicosa        
                       |                       +--+                               
                       |                          +-- Platynaias_compressa        
                       |                                                          
                       |                    +-------- Aculamprotula_tortuosa      
                       |                    |                                     
                       |  +-----------------+     +-- Aculamprotula_coreana       
                       |  |                 |  +--+                               
                       |  |                 |  |  +-- Aculamprotula_tientsinensis 
                       |  |                 +--+                                  
                       |  |                    |  +-- Aculamprotula_polysticta    
                       |  |                    +--+                               
                       |  |                       +-- Aculamprotula_scripta       
                       |  |                                                       
                       +--+              +----------- Unio_tumidus                
                          |              |                                        
                          |  +-----------+  +-------- Unio_crassus                
                          |  |           |  |                                     
                          |  |           +--+  +----- Unio_elongatulus            
                          |  |              |  |                                  
                          |  |              +--+  +-- Unio_delphinus              
                          |  |                 |  |                               
                          |  |                 +--+-- Unio_mancus                 
                          +--+                    |                               
                             |                    +-- Unio_pictorum               
                             |                                                    
                             |                    +-- Pseudocuneopsis_capitata    
                             |  +-----------------+                               
                             |  |                 +-- Pseudocuneopsis_sichuanensis
                             |  |                                                 
                             +--+                 +-- Nodularia_breviconcha       
                                |  +--------------+                               
                                |  |              +-- Nodularia_nipponensis       
                                |  |                                              
                                +--+           +----- Tchangsinaia_piscicula      
                                   |  +--------+                                  
                                   |  |        |  +-- Schistodesmus_lampreyanus[2]
                                   |  |        +--+                               
                                   +--+           +-- Schistodesmus_spinosus      
                                      |                                           
                                      |  +----------- Schistodesmus_lampreyanus[1]
                                      +--+                                        
                                         |  +-------- Cuneopsis_rufescens         
                                         +--+                                     
                                            |  +----- Cuneopsis_celtiformis       
                                            +--+                                  
                                               |  +-- Cuneopsis_demangei          
                                               +--+                               
                                                  +-- Cuneopsis_heudei            

Wu et al., 2024 [Publ 4350], fig. 8a, S1

iT tT bi riS oT roT roS iD
2 2 0 2 90 89 2 --
  +----------------------------------------------- Echyridella_menziesii       
  |                                                                            
  |                                            +-- Anodontites_trapesialis **     
  |  +-----------------------------------------+                               
  |  |                                         +-- Mutela_dubia **                
  |  |                                                                         
  |  |                                         +-- Margaritifera_dahurica      
--+  |  +--------------------------------------+                               
  |  |  |                                      +-- Margaritifera_margaritifera 
  |  |  |                                                                      
  |  |  |                                      +-- Quadrula_quadrula           
  |  |  |           +--------------------------+                               
  |  |  |           |                          +-- Uniomerus_tetralasmus       
  |  |  |           |                                                          
  +--+  |  +--------+                          +-- Elliptio_complanata         
     |  |  |        |  +-----------------------+                               
     |  |  |        |  |                       +-- Pleurobema_oviforme         
     |  |  |        +--+                                                       
     |  |  |           |  +----------------------- Amblema_plicata             
     |  |  |           |  |                                                    
     |  |  |           +--+  +-------------------- Popenaias_popeii            
     |  |  |              |  |                                                 
     |  |  |              +--+  +----------------- Toxolasma_parvum            
     |  |  |                 |  |                                              
     |  |  |                 |  |           +----- Potamilus_leptodon          
     +--+  |                 +--+  +--------+                                  
        |  |                    |  |        |  +-- Potamilus_alatus            
        |  |                    |  |        +--+                               
        |  |                    |  |           +-- Potamilus_streckersoni      
        |  |                    +--+                                           
        |  |                       |  +----------- Leaunio_lienosus            
        |  |                       |  |                                        
        |  |                       |  |     +----- Lampsilis_ornata            
        |  |                       +--+  +--+                                  
        |  |                          |  |  |  +-- Lampsilis_cardium           
        |  |                          |  |  +--+                               
        |  |                          +--+     +-- Lampsilis_cariosa           
        |  |                             |                                     
        |  |                             |  +----- Venustaconcha_ellipsiformis 
        |  |                             +--+                                  
        |  |                                |  +-- Lampsilis_powellii          
        |  |                                +--+                               
        |  |                                   +-- Lampsilis_siliquoidea       
        |  |                                                                   
        +--+                                +----- Chamberlainia_somsakpanhai  
           |           +--------------------+                                  
           |           |                    |  +-- Sinohyriopsis_cumingii      
           |           |                    +--+                               
           |           |                       +-- Sinohyriopsis_schlegelii    
           |           |                                                       
           |           |                       +-- Hyriopsis_bialata           
           |           |                    +--+                               
           |  +--------+                    |  +-- Rectidens_sumatrensis       
           |  |        |  +-----------------+                                  
           |  |        |  |                 |  +-- Lens_contradens             
           |  |        |  |                 +--+                               
           |  |        |  |                    +-- Physunio_superbus           
           |  |        |  |                                                    
           |  |        |  |           +----------- Potomida_littoralis         
           |  |        +--+  +--------+                                        
           |  |           |  |        |  +-------- Pronodularia_japanensis     
           |  |           |  |        +--+                                     
           |  |           |  |           |  +----- Lamprotula_leaii            
           |  |           |  |           +--+                                  
           |  |           |  |              |  +-- Lamprotula_caveata          
           |  |           +--+              +--+                               
           |  |              |                 +-- Lamprotula_cornuumlunae     
           |  |              |                                                 
           |  |              |                 +-- Pseudodon_vondembuschianus  
           |  |              |  +--------------+                               
           |  |              |  |              +-- Pilsbryoconcha_exilis       
           |  |              +--+                                              
           |  |                 |  +-------------- Inversidens_rentianensis    
           |  |                 |  |                                           
           +--+                 +--+           +-- Parvasolenaia_rivularis     
              |                    |  +--------+                               
              |                    |  |        +-- Ptychorhynchus_pfisteri     
              |                    +--+                                        
              |                       |  +-------- Microcondylaea_bonellii     
              |                       +--+                                     
              |                          |  +----- Sinosolenaia_carinata       
              |                          +--+                                  
              |                             |  +-- Sinosolenaia_iridinea       
              |                             +--+                               
              |                                +-- Sinosolenaia_oleivora       
              |                                                                
              |  +-------------------------------- Lamellidens_jenkinsianus    
              |  |                                                             
              |  |                             +-- Lepidodesma_aligera         
              |  |  +--------------------------+                               
              |  |  |                          +-- Lepidodesma_languilati      
              |  |  |                                                          
              |  |  |                    +-------- Aculamprotula_tortuosa      
              |  |  |                    |                                     
              |  |  |     +--------------+     +-- Aculamprotula_coreana       
              |  |  |     |              |  +--+                               
              +--+  |     |              |  |  +-- Aculamprotula_tientsinensis 
                 |  |     |              +--+                                  
                 |  |     |                 |  +-- Aculamprotula_polysticta    
                 |  |     |                 +--+                               
                 |  |     |                    +-- Aculamprotula_scripta       
                 |  |     |                                                    
                 |  |  +--+        +-------------- Unio_tumidus                
                 |  |  |  |  +-----+                                           
                 |  |  |  |  |     |  +----------- Unio_crassus                
                 |  |  |  |  |     +--+                                        
                 +--+  |  |  |        |  +-------- Unio_elongatulus            
                    |  |  |  |        +--+                                     
                    |  |  |  |           |  +----- Unio_pictorum               
                    |  |  |  |           +--+                                  
                    |  |  |  |              |  +-- Unio_delphinus              
                    |  |  +--+              +--+                               
                    |  |     |                 +-- Unio_mancus                 
                    |  |     |                                                 
                    |  |     |                 +-- Nodularia_breviconcha       
                    |  |     |  +--------------+                               
                    |  |     |  |              +-- Nodularia_douglasiae        
                    |  |     |  |                                              
                    |  |     |  |              +-- Pseudocuneopsis_capitata    
                    |  |     |  +--------------+                               
                    |  |     +--+              +-- Pseudocuneopsis_sichuanensis
                    |  |        |                                              
                    |  |        |              +-- Schistodesmus_lampreyanus   
                    |  |        |  +-----------+                               
                    +--+        |  |           +-- Schistodesmus_spinosus      
                       |        +--+                                           
                       |           |  +----------- Tchangsinaia_piscicula      
                       |           +--+                                        
                       |              |  +-------- Cuneopsis_rufescens         
                       |              +--+                                     
                       |                 |  +----- Cuneopsis_celtiformis       
                       |                 +--+                                  
                       |                    |  +-- Cuneopsis_demangei          
                       |                    +--+                               
                       |                       +-- Cuneopsis_heudei            
                       |                                                       
                       |                 +-------- Acuticosta_chinensis        
                       |  +--------------+                                     
                       |  |              |  +----- Lanceolaria_gladiola        
                       |  |              +--+                                  
                       |  |                 |  +-- Lanceolaria_grayii          
                       |  |                 +--+                               
                       |  |                    +-- Lanceolaria_lanceolata      
                       |  |                                                    
                       |  |        +-------------- Cristaria_plicata           
                       |  |        |                                           
                       +--+  +-----+           +-- Anemina_arcaeformis         
                          |  |     |  +--------+                               
                          |  |     |  |        +-- Anemina_euscaphys           
                          |  |     +--+                                        
                          |  |        |  +-------- Beringiana_fukuharai        
                          |  |        +--+                                     
                          |  |           |  +----- Sinanodonta_lucida          
                          |  |           +--+                                  
                          |  |              |  +-- Sinanodonta_tumens          
                          +--+              +--+                               
                             |                 +-- Sinanodonta_woodiana        
                             |                                                 
                             |           +-------- Pseudanodonta_complanata    
                             |  +--------+                                     
                             |  |        |  +----- Anodonta_anatina            
                             |  |        +--+                                  
                             |  |           |  +-- Anodonta_cygnea             
                             |  |           +--+                               
                             +--+              +-- Anodonta_exulcerata         
                                |                                              
                                |  +-------------- Anodonta_nuttalliana        
                                |  |                                           
                                |  |        +----- Pyganodon_grandis           
                                +--+  +-----+                                  
                                   |  |     |  +-- Utterbackia_imbecillis      
                                   |  |     +--+                               
                                   +--+        +-- Utterbackia_peninsularis    
                                      |                                        
                                      |  +-------- Lasmigona_complanata        
                                      +--+                                     
                                         |  +----- Prolasmidonta_heterodon     
                                         +--+                                  
                                            |  +-- Alasmidonta_varicosa        
                                            +--+                               
                                               +-- Platynaias_compressa        

Wu et al., 2024 [Publ 4350], fig. 8b, S2

iT tT bi riS oT roT roS iD
2 2 0 2 90 89 2 --
  +----------------------------------------------- Echyridella_menziesii       
  |                                                                            
  |                                            +-- Anodontites_trapesialis **     
  |  +-----------------------------------------+                               
  |  |                                         +-- Mutela_dubia **                
  |  |                                                                         
  |  |                                         +-- Margaritifera_dahurica      
--+  |  +--------------------------------------+                               
  |  |  |                                      +-- Margaritifera_margaritifera 
  |  |  |                                                                      
  |  |  |                                      +-- Quadrula_quadrula           
  |  |  |           +--------------------------+                               
  |  |  |           |                          +-- Uniomerus_tetralasmus       
  |  |  |           |                                                          
  +--+  |  +--------+                          +-- Elliptio_complanata         
     |  |  |        |  +-----------------------+                               
     |  |  |        |  |                       +-- Pleurobema_oviforme         
     |  |  |        +--+                                                       
     |  |  |           |  +----------------------- Amblema_plicata             
     |  |  |           |  |                                                    
     |  |  |           +--+  +-------------------- Popenaias_popeii            
     |  |  |              |  |                                                 
     |  |  |              +--+  +----------------- Toxolasma_parvum            
     |  |  |                 |  |                                              
     |  |  |                 |  |           +----- Potamilus_leptodon          
     +--+  |                 +--+  +--------+                                  
        |  |                    |  |        |  +-- Potamilus_alatus            
        |  |                    |  |        +--+                               
        |  |                    +--+           +-- Potamilus_streckersoni      
        |  |                       |                                           
        |  |                       |  +----------- Leaunio_lienosus            
        |  |                       |  |                                        
        |  |                       +--+  +-------- Venustaconcha_ellipsiformis 
        |  |                          |  |                                     
        |  |                          |  |     +-- Lampsilis_powellii          
        |  |                          +--+-----+                               
        |  |                             |     +-- Lampsilis_siliquoidea       
        |  |                             |                                     
        |  |                             |  +----- Lampsilis_ornata            
        |  |                             +--+                                  
        |  |                                |  +-- Lampsilis_cardium           
        |  |                                +--+                               
        |  |                                   +-- Lampsilis_cariosa           
        |  |                                                                   
        +--+                                +----- Chamberlainia_somsakpanhai  
           |           +--------------------+                                  
           |           |                    |  +-- Sinohyriopsis_cumingii      
           |           |                    +--+                               
           |           |                       +-- Sinohyriopsis_schlegelii    
           |           |                                                       
           |           |                       +-- Hyriopsis_bialata           
           |           |                    +--+                               
           |  +--------+                    |  +-- Rectidens_sumatrensis       
           |  |        |  +-----------------+                                  
           |  |        |  |                 |  +-- Lens_contradens             
           |  |        |  |                 +--+                               
           |  |        |  |                    +-- Physunio_superbus           
           |  |        |  |                                                    
           |  |        |  |           +----------- Potomida_littoralis         
           |  |        +--+  +--------+                                        
           |  |           |  |        |  +-------- Pronodularia_japanensis     
           |  |           |  |        +--+                                     
           |  |           |  |           |  +----- Lamprotula_leaii            
           |  |           |  |           +--+                                  
           |  |           |  |              |  +-- Lamprotula_caveata          
           |  |           +--+              +--+                               
           |  |              |                 +-- Lamprotula_cornuumlunae     
           |  |              |                                                 
           |  |              |                 +-- Pseudodon_vondembuschianus  
           |  |              |  +--------------+                               
           |  |              |  |              +-- Pilsbryoconcha_exilis       
           |  |              +--+                                              
           |  |                 |  +-------------- Inversidens_rentianensis    
           |  |                 |  |                                           
           +--+                 +--+           +-- Parvasolenaia_rivularis     
              |                    |  +--------+                               
              |                    |  |        +-- Ptychorhynchus_pfisteri     
              |                    +--+                                        
              |                       |  +-------- Microcondylaea_bonellii     
              |                       +--+                                     
              |                          |  +----- Sinosolenaia_carinata       
              |                          +--+                                  
              |                             |  +-- Sinosolenaia_iridinea       
              |                             +--+                               
              |                                +-- Sinosolenaia_oleivora       
              |                                                                
              |  +-------------------------------- Lamellidens_jenkinsianus    
              |  |                                                             
              |  |                             +-- Lepidodesma_aligera         
              |  |  +--------------------------+                               
              |  |  |                          +-- Lepidodesma_languilati      
              |  |  |                                                          
              |  |  |                    +-------- Aculamprotula_tortuosa      
              |  |  |                    |                                     
              |  |  |     +--------------+     +-- Aculamprotula_coreana       
              |  |  |     |              |  +--+                               
              +--+  |     |              |  |  +-- Aculamprotula_tientsinensis 
                 |  |     |              +--+                                  
                 |  |     |                 |  +-- Aculamprotula_polysticta    
                 |  |     |                 +--+                               
                 |  |     |                    +-- Aculamprotula_scripta       
                 |  |     |                                                    
                 |  |  +--+        +-------------- Unio_tumidus                
                 |  |  |  |  +-----+                                           
                 |  |  |  |  |     |  +----------- Unio_crassus                
                 |  |  |  |  |     +--+                                        
                 +--+  |  |  |        |  +-------- Unio_elongatulus            
                    |  |  |  |        +--+                                     
                    |  |  |  |           |  +----- Unio_pictorum               
                    |  |  |  |           +--+                                  
                    |  |  |  |              |  +-- Unio_delphinus              
                    |  |  +--+              +--+                               
                    |  |     |                 +-- Unio_mancus                 
                    |  |     |                                                 
                    |  |     |                 +-- Nodularia_breviconcha       
                    |  |     |  +--------------+                               
                    |  |     |  |              +-- Nodularia_douglasiae        
                    |  |     |  |                                              
                    |  |     |  |              +-- Pseudocuneopsis_capitata    
                    |  |     |  +--------------+                               
                    |  |     +--+              +-- Pseudocuneopsis_sichuanensis
                    |  |        |                                              
                    |  |        |              +-- Schistodesmus_lampreyanus   
                    +--+        |  +-----------+                               
                       |        |  |           +-- Schistodesmus_spinosus      
                       |        +--+                                           
                       |           |  +----------- Tchangsinaia_piscicula      
                       |           +--+                                        
                       |              |  +-------- Cuneopsis_rufescens         
                       |              +--+                                     
                       |                 |  +----- Cuneopsis_celtiformis       
                       |                 +--+                                  
                       |                    |  +-- Cuneopsis_demangei          
                       |                    +--+                               
                       |                       +-- Cuneopsis_heudei            
                       |                                                       
                       |                 +-------- Acuticosta_chinensis        
                       |  +--------------+                                     
                       |  |              |  +----- Lanceolaria_gladiola        
                       |  |              +--+                                  
                       |  |                 |  +-- Lanceolaria_grayii          
                       |  |                 +--+                               
                       |  |                    +-- Lanceolaria_lanceolata      
                       |  |                                                    
                       |  |        +-------------- Cristaria_plicata           
                       +--+        |                                           
                          |  +-----+           +-- Anemina_arcaeformis         
                          |  |     |  +--------+                               
                          |  |     |  |        +-- Anemina_euscaphys           
                          |  |     +--+                                        
                          |  |        |  +-------- Beringiana_fukuharai        
                          |  |        +--+                                     
                          |  |           |  +----- Sinanodonta_lucida          
                          |  |           +--+                                  
                          +--+              |  +-- Sinanodonta_tumens          
                             |              +--+                               
                             |                 +-- Sinanodonta_woodiana        
                             |                                                 
                             |  +----------------- Anodonta_nuttalliana        
                             |  |                                              
                             |  |        +-------- Pseudanodonta_complanata    
                             |  |  +-----+                                     
                             +--+  |     |  +----- Anodonta_anatina            
                                |  |     +--+                                  
                                |  |        |  +-- Anodonta_cygnea             
                                |  |        +--+                               
                                +--+           +-- Anodonta_exulcerata         
                                   |                                           
                                   |        +----- Pyganodon_grandis           
                                   |  +-----+                                  
                                   |  |     |  +-- Utterbackia_imbecillis      
                                   |  |     +--+                               
                                   +--+        +-- Utterbackia_peninsularis    
                                      |                                        
                                      |  +-------- Lasmigona_complanata        
                                      +--+                                     
                                         |  +----- Prolasmidonta_heterodon     
                                         +--+                                  
                                            |  +-- Alasmidonta_varicosa        
                                            +--+                               
                                               +-- Platynaias_compressa        

Wu et al., 2021 [Publ 4366], fig. 3-BI

iT tT bi riS oT roT roS iD
2 2 0 2 72 71 2 --
  +-------------------------------------------- Echyridella_menziesii      
  |                                                                        
  |                                         +-- Anodontites_trapesialis **    
  |  +--------------------------------------+                              
  |  |                                      +-- Mutela_dubia **               
  |  |                                                                     
  |  |                                      +-- Gibbosula_rochechouartii   
--+  |                          +-----------+                              
  |  |                          |           +-- Gibbosula_crassa           
  |  |  +-----------------------+                                          
  |  |  |                       |  +----------- Cumberlandia_monodonta     
  |  |  |                       +--+                                       
  |  |  |                          |  +-------- Pseudunio_marocanus        
  |  |  |                          +--+                                    
  |  |  |                             |  +----- Margaritifera_falcata      
  +--+  |                             +--+                                 
     |  |                                |  +-- Margaritifera_margaritifera
     |  |                                +--+                              
     |  |                                   +-- Margaritifera_dahurica     
     |  |                                                                  
     |  |                                   +-- Uniomerus_tetralasmus      
     |  |        +--------------------------+                              
     |  |        |                          +-- Quadrula_quadrula          
     |  |        |                                                         
     |  |  +-----+                          +-- Pleurobema_oviforme        
     |  |  |     |  +-----------------------+                              
     |  |  |     |  |                       +-- Elliptio_complanata        
     |  |  |     +--+                                                      
     +--+  |        |  +----------------------- Amblema_plicata            
        |  |        |  |                                                   
        |  |        +--+  +-------------------- Popenaias_popeii           
        |  |           |  |                                                
        |  |           +--+  +----------------- Toxolasma_parvum           
        |  |              |  |                                             
        |  |              +--+              +-- Potamilus_alatus           
        |  |                 |  +-----------+                              
        |  |                 |  |           +-- Potamilus_leptodon         
        |  |                 +--+                                          
        |  |                    |  +----------- Leaunio_lienosus           
        |  |                    |  |                                       
        |  |                    +--+        +-- Lampsilis_ornata           
        |  |                       |  +-----+                              
        |  |                       |  |     +-- Lampsilis_cardium          
        |  |                       +--+                                    
        |  |                          |  +----- Venustaconcha_ellipsiformis
        |  |                          +--+                                 
        |  |                             |  +-- Lampsilis_siliquoidea      
        |  |                             +--+                              
        +--+                                +-- Lampsilis_powellii         
           |                                                               
           |                             +----- Chamberlainia_somsakpanhai 
           |                 +-----------+                                 
           |                 |           |  +-- Sinohyriopsis_schlegelii   
           |                 |           +--+                              
           |                 |              +-- Sinohyriopsis_cumingii     
           |                 |                                             
           |  +--------------+        +-------- Potomida_littoralis        
           |  |              |  +-----+                                    
           |  |              |  |     |  +----- Pronodularia_japanensis    
           |  |              |  |     +--+                                 
           |  |              |  |        |  +-- Lamprotula_caveata         
           |  |              |  |        +--+                              
           |  |              +--+           +-- Lamprotula_leaii           
           |  |                 |                                          
           |  |                 |           +-- Pilsbryoconcha_exilis      
           |  |                 |  +--------+                              
           |  |                 |  |        +-- Pseudodon_vondembuschianus 
           |  |                 |  |                                       
           |  |                 +--+        +-- Parvasolenaia_rivularis    
           |  |                    |  +-----+                              
           |  |                    |  |     +-- Ptychorhynchus_pfisteri    
           |  |                    +--+                                    
           |  |                       |  +----- Microcondylaea_bonellii    
           +--+                       +--+                                 
              |                          |  +-- Sinosolenaia_oleivora      
              |                          +--+                              
              |                             +-- Sinosolenaia_carinata      
              |                                                            
              |  +----------------------------- Lepidodesma_languilati     
              |  |                                                         
              |  |                    +-------- Aculamprotula_tortuosa     
              |  |                    |                                    
              |  |           +--------+     +-- Aculamprotula_scripta      
              |  |           |        |  +--+                              
              |  |           |        |  |  +-- Aculamprotula_polysticta   
              |  |           |        +--+                                 
              |  |           |           |  +-- Aculamprotula_coreana      
              |  |           |           +--+                              
              |  |  +--------+              +-- Aculamprotula_tientsinensis
              |  |  |        |                                             
              +--+  |        |        +-------- Unio_tumidus               
                 |  |        |  +-----+                                    
                 |  |        |  |     |  +----- Unio_crassus               
                 |  |        |  |     +--+                                 
                 |  |        |  |        |  +-- Unio_pictorum              
                 |  |        +--+        +--+                              
                 |  |           |           +-- Unio_delphinus             
                 |  |           |                                          
                 |  |           |  +----------- Pseudocuneopsis_capitata   
                 |  |           |  |                                       
                 |  |           +--+        +-- Nodularia_douglasiae       
                 |  |              |  +-----+                              
                 |  |              |  |     +-- Nodularia_breviconcha      
                 +--+              +--+                                    
                    |                 |  +----- Schistodesmus_lampreyanus  
                    |                 +--+                                 
                    |                    |  +-- Cuneopsis_heudei           
                    |                    +--+                              
                    |                       +-- Tchangsinaia_piscicula     
                    |                                                      
                    |                 +-------- Acuticosta_chinensis       
                    |  +--------------+                                    
                    |  |              |  +----- Lanceolaria_gladiola       
                    |  |              +--+                                 
                    |  |                 |  +-- Lanceolaria_grayii         
                    |  |                 +--+                              
                    |  |                    +-- Lanceolaria_lanceolata     
                    |  |                                                   
                    +--+              +-------- Cristaria_plicata          
                       |              |                                    
                       |  +-----------+     +-- Sinanodonta_woodiana       
                       |  |           |  +--+                              
                       |  |           |  |  +-- Sinanodonta_lucida         
                       |  |           +--+                                 
                       |  |              |  +-- Anemina_euscaphys          
                       +--+              +--+                              
                          |                 +-- Anemina_arcaeformis        
                          |                                                
                          |  +----------------- Anodonta_cygnea            
                          |  |                                             
                          +--+  +-------------- Anodonta_anatina           
                             |  |                                          
                             |  |        +----- Pyganodon_grandis          
                             +--+  +-----+                                 
                                |  |     |  +-- Utterbackia_peninsularis   
                                |  |     +--+                              
                                +--+        +-- Utterbackia_imbecillis     
                                   |                                       
                                   |  +-------- Prolasmidonta_heterodon    
                                   +--+                                    
                                      |  +----- Alasmidonta_varicosa       
                                      +--+                                 
                                         |  +-- Platynaias_subviridis      
                                         +--+                              
                                            +-- Platynaias_compressa       

Wu et al., 2021 [Publ 4366], fig. 3-ML

iT tT bi riS oT roT roS iD
2 2 0 2 72 71 2 --
  +-------------------------------------------- Echyridella_menziesii      
  |                                                                        
  |                                         +-- Anodontites_trapesialis **    
  |  +--------------------------------------+                              
  |  |                                      +-- Mutela_dubia **               
  |  |                                                                     
  |  |                                      +-- Gibbosula_rochechouartii   
--+  |                          +-----------+                              
  |  |                          |           +-- Gibbosula_crassa           
  |  |  +-----------------------+                                          
  |  |  |                       |  +----------- Cumberlandia_monodonta     
  |  |  |                       +--+                                       
  |  |  |                          |  +-------- Pseudunio_marocanus        
  |  |  |                          +--+                                    
  |  |  |                             |  +----- Margaritifera_falcata      
  +--+  |                             +--+                                 
     |  |                                |  +-- Margaritifera_margaritifera
     |  |                                +--+                              
     |  |                                   +-- Margaritifera_dahurica     
     |  |                                                                  
     |  |                                   +-- Uniomerus_tetralasmus      
     |  |        +--------------------------+                              
     |  |        |                          +-- Quadrula_quadrula          
     |  |        |                                                         
     |  |  +-----+                          +-- Pleurobema_oviforme        
     |  |  |     |  +-----------------------+                              
     |  |  |     |  |                       +-- Elliptio_complanata        
     |  |  |     +--+                                                      
     +--+  |        |  +----------------------- Amblema_plicata            
        |  |        |  |                                                   
        |  |        +--+  +-------------------- Popenaias_popeii           
        |  |           |  |                                                
        |  |           +--+  +----------------- Toxolasma_parvum           
        |  |              |  |                                             
        |  |              +--+              +-- Potamilus_alatus           
        |  |                 |  +-----------+                              
        |  |                 |  |           +-- Potamilus_leptodon         
        |  |                 +--+                                          
        |  |                    |  +----------- Leaunio_lienosus           
        |  |                    |  |                                       
        |  |                    +--+        +-- Lampsilis_ornata           
        |  |                       |  +-----+                              
        |  |                       |  |     +-- Lampsilis_cardium          
        |  |                       +--+                                    
        |  |                          |  +----- Venustaconcha_ellipsiformis
        |  |                          +--+                                 
        |  |                             |  +-- Lampsilis_siliquoidea      
        |  |                             +--+                              
        +--+                                +-- Lampsilis_powellii         
           |                                                               
           |                             +----- Chamberlainia_somsakpanhai 
           |                 +-----------+                                 
           |                 |           |  +-- Sinohyriopsis_schlegelii   
           |                 |           +--+                              
           |                 |              +-- Sinohyriopsis_cumingii     
           |                 |                                             
           |  +--------------+        +-------- Potomida_littoralis        
           |  |              |  +-----+                                    
           |  |              |  |     |  +----- Pronodularia_japanensis    
           |  |              |  |     +--+                                 
           |  |              |  |        |  +-- Lamprotula_caveata         
           |  |              |  |        +--+                              
           |  |              +--+           +-- Lamprotula_leaii           
           |  |                 |                                          
           |  |                 |           +-- Pilsbryoconcha_exilis      
           |  |                 |  +--------+                              
           |  |                 |  |        +-- Pseudodon_vondembuschianus 
           |  |                 |  |                                       
           |  |                 +--+        +-- Parvasolenaia_rivularis    
           |  |                    |  +-----+                              
           |  |                    |  |     +-- Ptychorhynchus_pfisteri    
           |  |                    +--+                                    
           |  |                       |  +----- Microcondylaea_bonellii    
           +--+                       +--+                                 
              |                          |  +-- Sinosolenaia_oleivora      
              |                          +--+                              
              |                             +-- Sinosolenaia_carinata      
              |                                                            
              |  +----------------------------- Lepidodesma_languilati     
              |  |                                                         
              |  |                    +-------- Aculamprotula_tortuosa     
              |  |                    |                                    
              |  |           +--------+     +-- Aculamprotula_scripta      
              |  |           |        |  +--+                              
              |  |           |        |  |  +-- Aculamprotula_polysticta   
              |  |           |        +--+                                 
              |  |           |           |  +-- Aculamprotula_coreana      
              |  |           |           +--+                              
              |  |  +--------+              +-- Aculamprotula_tientsinensis
              |  |  |        |                                             
              +--+  |        |        +-------- Unio_tumidus               
                 |  |        |  +-----+                                    
                 |  |        |  |     |  +----- Unio_crassus               
                 |  |        |  |     +--+                                 
                 |  |        |  |        |  +-- Unio_pictorum              
                 |  |        +--+        +--+                              
                 |  |           |           +-- Unio_delphinus             
                 |  |           |                                          
                 |  |           |  +----------- Pseudocuneopsis_capitata   
                 |  |           |  |                                       
                 |  |           +--+        +-- Nodularia_douglasiae       
                 |  |              |  +-----+                              
                 |  |              |  |     +-- Nodularia_breviconcha      
                 +--+              +--+                                    
                    |                 |  +----- Tchangsinaia_piscicula     
                    |                 +--+                                 
                    |                    |  +-- Schistodesmus_lampreyanus  
                    |                    +--+                              
                    |                       +-- Cuneopsis_heudei           
                    |                                                      
                    |                 +-------- Acuticosta_chinensis       
                    |  +--------------+                                    
                    |  |              |  +----- Lanceolaria_gladiola       
                    |  |              +--+                                 
                    |  |                 |  +-- Lanceolaria_grayii         
                    |  |                 +--+                              
                    |  |                    +-- Lanceolaria_lanceolata     
                    |  |                                                   
                    +--+              +-------- Cristaria_plicata          
                       |              |                                    
                       |  +-----------+     +-- Sinanodonta_woodiana       
                       |  |           |  +--+                              
                       |  |           |  |  +-- Sinanodonta_lucida         
                       |  |           +--+                                 
                       |  |              |  +-- Anemina_euscaphys          
                       +--+              +--+                              
                          |                 +-- Anemina_arcaeformis        
                          |                                                
                          |  +----------------- Anodonta_cygnea            
                          |  |                                             
                          +--+  +-------------- Anodonta_anatina           
                             |  |                                          
                             |  |        +----- Pyganodon_grandis          
                             +--+  +-----+                                 
                                |  |     |  +-- Utterbackia_peninsularis   
                                |  |     +--+                              
                                +--+        +-- Utterbackia_imbecillis     
                                   |                                       
                                   |  +-------- Prolasmidonta_heterodon    
                                   +--+                                    
                                      |  +----- Alasmidonta_varicosa       
                                      +--+                                 
                                         |  +-- Platynaias_subviridis      
                                         +--+                              
                                            +-- Platynaias_compressa       

Wu et al., 2021 [Publ 4366], fig. 4

iT tT bi riS oT roT roS iD
2 2 0 2 67 66 2 --
  +----------------------------------------- Echyridella_menziesii      
  |                                                                     
  |                                      +-- Mutela_dubia **               
  |  +-----------------------------------+                              
  |  |                                   +-- Anodontites_trapesialis **    
  |  |                                                                  
  |  |                                   +-- Gibbosula_rochechouartii   
--+  |                       +-----------+                              
  |  |                       |           +-- Gibbosula_crassa           
  |  |  +--------------------+                                          
  |  |  |                    |  +----------- Cumberlandia_monodonta     
  |  |  |                    +--+                                       
  |  |  |                       |  +-------- Pseudunio_marocanus        
  |  |  |                       +--+                                    
  +--+  |                          |  +----- Margaritifera_falcata      
     |  |                          +--+                                 
     |  |                             |  +-- Margaritifera_margaritifera
     |  |                             +--+                              
     |  |                                +-- Margaritifera_dahurica     
     |  |                                                               
     |  |        +-------------------------- Quadrula_quadrula          
     |  |        |                                                      
     |  |  +-----+  +----------------------- Pleurobema_oviforme        
     |  |  |     |  |                                                   
     |  |  |     +--+  +-------------------- Amblema_plicata            
     |  |  |        |  |                                                
     +--+  |        +--+  +----------------- Popenaias_popeii           
        |  |           |  |                                             
        |  |           +--+  +-------------- Toxolasma_parvum           
        |  |              |  |                                          
        |  |              |  |           +-- Potamilus_alatus           
        |  |              +--+  +--------+                              
        |  |                 |  |        +-- Potamilus_leptodon         
        |  |                 |  |                                       
        |  |                 +--+        +-- Lampsilis_ornata           
        |  |                    |  +-----+                              
        |  |                    |  |     +-- Lampsilis_cardium          
        |  |                    +--+                                    
        |  |                       |  +----- Venustaconcha_ellipsiformis
        |  |                       +--+                                 
        |  |                          |  +-- Lampsilis_siliquoidea      
        |  |                          +--+                              
        |  |                             +-- Lampsilis_powellii         
        |  |                                                            
        +--+                          +----- Chamberlainia_somsakpanhai 
           |              +-----------+                                 
           |              |           |  +-- Sinohyriopsis_schlegelii   
           |              |           +--+                              
           |              |              +-- Sinohyriopsis_cumingii     
           |              |                                             
           |  +-----------+           +----- Potomida_littoralis        
           |  |           |  +--------+                                 
           |  |           |  |        |  +-- Lamprotula_caveata         
           |  |           |  |        +--+                              
           |  |           |  |           +-- Lamprotula_leaii           
           |  |           +--+                                          
           |  |              |           +-- Pilsbryoconcha_exilis      
           |  |              |  +--------+                              
           |  |              |  |        +-- Pseudodon_vondembuschianus 
           |  |              |  |                                       
           |  |              +--+        +-- Parvasolenaia_rivularis    
           |  |                 |  +-----+                              
           |  |                 |  |     +-- Ptychorhynchus_pfisteri    
           |  |                 +--+                                    
           |  |                    |  +----- Microcondylaea_bonellii    
           |  |                    +--+                                 
           +--+                       |  +-- Sinosolenaia_oleivora      
              |                       +--+                              
              |                          +-- Sinosolenaia_carinata      
              |                                                         
              |  +-------------------------- Lepidodesma_languilati     
              |  |                                                      
              |  |                 +-------- Aculamprotula_tortuosa     
              |  |                 |                                    
              |  |        +--------+     +-- Aculamprotula_scripta      
              |  |        |        |  +--+                              
              |  |        |        |  |  +-- Aculamprotula_polysticta   
              |  |        |        +--+                                 
              |  |        |           |  +-- Aculamprotula_coreana      
              |  |        |           +--+                              
              |  |  +-----+              +-- Aculamprotula_tientsinensis
              |  |  |     |                                             
              +--+  |     |        +-------- Unio_tumidus               
                 |  |     |  +-----+                                    
                 |  |     |  |     |  +----- Unio_crassus               
                 |  |     |  |     +--+                                 
                 |  |     |  |        |  +-- Unio_pictorum              
                 |  |     +--+        +--+                              
                 |  |        |           +-- Unio_delphinus             
                 |  |        |                                          
                 |  |        |  +----------- Pseudocuneopsis_capitata   
                 |  |        |  |                                       
                 |  |        +--+        +-- Nodularia_douglasiae       
                 |  |           |  +-----+                              
                 |  |           |  |     +-- Nodularia_breviconcha      
                 +--+           +--+                                    
                    |              |  +----- Schistodesmus_lampreyanus  
                    |              +--+                                 
                    |                 |  +-- Tchangsinaia_piscicula     
                    |                 +--+                              
                    |                    +-- Cuneopsis_heudei           
                    |                                                   
                    |              +-------- Acuticosta_chinensis       
                    |  +-----------+                                    
                    |  |           |  +----- Lanceolaria_gladiola       
                    |  |           +--+                                 
                    |  |              |  +-- Lanceolaria_grayii         
                    |  |              +--+                              
                    |  |                 +-- Lanceolaria_lanceolata     
                    |  |                                                
                    +--+           +-------- Cristaria_plicata          
                       |           |                                    
                       |  +--------+     +-- Sinanodonta_woodiana       
                       |  |        |  +--+                              
                       |  |        |  |  +-- Sinanodonta_lucida         
                       |  |        +--+                                 
                       |  |           |  +-- Anemina_euscaphys          
                       +--+           +--+                              
                          |              +-- Anemina_arcaeformis        
                          |                                             
                          |  +-------------- Anodonta_cygnea            
                          |  |                                          
                          +--+  +----------- Anodonta_anatina           
                             |  |                                       
                             |  |     +----- Pyganodon_grandis          
                             +--+  +--+                                 
                                |  |  |  +-- Utterbackia_imbecillis     
                                |  |  +--+                              
                                +--+     +-- Utterbackia_peninsularis   
                                   |                                    
                                   |  +----- Prolasmidonta_heterodon    
                                   +--+                                 
                                      |  +-- Platynaias_compressa       
                                      +--+                              
                                         +-- Alasmidonta_varicosa       

Graf & Ó Foighil, 2000 [Publ 2648], fig. 2A

iT tT bi riS oT roT roS iD
1 1 0 1 29 28 2 --
  +-------------------------------- Mutela_rostrata **            
  |                                                            
  |  +----------------------------- Hyridella_depressa         
--+  |                                                         
  |  |                          +-- Margaritifera_margaritifera
  |  |  +-----------------------+                              
  +--+  |                       +-- Cumberlandia_monodonta     
     |  |                                                      
     |  |                       +-- Unio_pictorum              
     |  |  +--------------------+                              
     +--+  |                    +-- Unio_caffer                
        |  |                                                   
        |  |           +----------- Anodonta_cygnea            
        |  |           |                                       
        |  |  +--------+        +-- Alasmidonta_marginata      
        +--+  |        |  +-----+                              
           |  |        |  |     +-- Lasmigona_costata          
           |  |        +--+                                    
           |  |           |  +----- Strophitus_undulatus       
           |  |           +--+                                 
           |  |              |  +-- Pyganodon_grandis          
           |  |              +--+                              
           +--+                 +-- Pyganodon_fragilis         
              |                                                
              |           +-------- Quadrula_quadrula          
              |  +--------+                                    
              |  |        |  +----- Eurynia_dilatata           
              |  |        +--+                                 
              |  |           |  +-- Pleurobema_sintoxia        
              |  |           +--+                              
              +--+              +-- Fusconaia_flava            
                 |                                             
                 |  +-------------- Amblema_plicata            
                 |  |                                          
                 |  |  +----------- Ligumia_recta              
                 |  |  |                                       
                 +--+  +----------- Truncilla_truncata         
                    |  |                                       
                    |  |        +-- Ptychobranchus_fasciolaris 
                    |  +--------+                              
                    +--+        +-- Sagittunio_nasutus         
                       |                                       
                       |     +----- Lampsilis_siliquoidea      
                       |  +--+                                 
                       |  |  |  +-- Epioblasma_triquetra       
                       |  |  +--+                              
                       |  |     +-- Epioblasma_brevidens       
                       +--+                                    
                          |  +----- Ortmanniana_ligamentina    
                          |  |                                 
                          |  +----- Cambarunio_iris            
                          +--+                                 
                             +----- Leaunio_vanuxemensis       
                             |                                 
                             |  +-- Lampsilis_cardium          
                             +--+                              
                                +-- Lampsilis_fasciola         

Graf & Ó Foighil, 2000 [Publ 2648], fig. 2B

iT tT bi riS oT roT roS iD
1 1 0 1 29 28 2 --
  +----------------------------------------- Mutela_rostrata **            
  |                                                                     
  |  +-------------------------------------- Hyridella_depressa         
--+  |                                                                  
  |  |                                   +-- Margaritifera_margaritifera
  |  |  +--------------------------------+                              
  +--+  |                                +-- Cumberlandia_monodonta     
     |  |                                                               
     |  |                                +-- Unio_pictorum              
     |  |  +-----------------------------+                              
     +--+  |                             +-- Unio_caffer                
        |  |                                                            
        |  |                    +----------- Anodonta_cygnea            
        |  |                    |                                       
        |  |  +-----------------+        +-- Alasmidonta_marginata      
        +--+  |                 |  +-----+                              
           |  |                 |  |     +-- Lasmigona_costata          
           |  |                 +--+                                    
           |  |                    |  +----- Strophitus_undulatus       
           |  |                    +--+                                 
           |  |                       |  +-- Pyganodon_grandis          
           +--+                       +--+                              
              |                          +-- Pyganodon_fragilis         
              |                                                         
              |                    +-------- Quadrula_quadrula          
              |  +-----------------+                                    
              |  |                 |  +----- Eurynia_dilatata           
              |  |                 +--+                                 
              |  |                    |  +-- Pleurobema_sintoxia        
              +--+                    +--+                              
                 |                       +-- Fusconaia_flava            
                 |                                                      
                 |  +----------------------- Amblema_plicata            
                 |  |                                                   
                 +--+  +-------------------- Truncilla_truncata         
                    |  |                                                
                    +--+  +----------------- Ligumia_recta              
                       |  |                                             
                       |  |              +-- Ptychobranchus_fasciolaris 
                       +--+  +-----------+                              
                          |  |           +-- Sagittunio_nasutus         
                          |  |                                          
                          +--+  +----------- Lampsilis_siliquoidea      
                             |  |                                       
                             |  |     +----- Ortmanniana_ligamentina    
                             +--+  +--+                                 
                                |  |  |  +-- Lampsilis_cardium          
                                |  |  +--+                              
                                |  |     +-- Lampsilis_fasciola         
                                +--+                                    
                                   |     +-- Epioblasma_brevidens       
                                   |  +--+                              
                                   |  |  +-- Epioblasma_triquetra       
                                   +--+                                 
                                      |  +-- Cambarunio_iris            
                                      +--+                              
                                         +-- Leaunio_vanuxemensis       

Hoeh et al., 1998 [Publ 2695], fig. 3

iT tT bi riS oT roT roS iD
1 1 0 1 4 3 1 --
  +-------- Mutela_rostrata **            
--+                                    
  |  +----- Anodonta_cygnea            
  +--+                                 
     |  +-- Margaritifera_margaritifera
     +--+                              
        +-- Amblema_plicata            

Hoeh et al., 1998 [Publ 2695], fig. 4

iT tT bi riS oT roT roS iD
1 1 0 1 4 3 1 --
  +-------- Mutela_rostrata **            
--+                                    
  |  +----- Amblema_plicata            
  +--+                                 
     |  +-- Margaritifera_margaritifera
     +--+                              
        +-- Anodonta_cygnea            

Bieler et al., 2014 [Publ 3622], fig. 26, 27, 29, 30

iT tT bi riS oT roT roS iD
1 1 0 1 4 3 2 --
  +----- Unio_pictorum              
--+                                 
  |  +-- Aspatharia_pfeifferiana **    
  +--+                              
     +-- Margaritifera_margaritifera

Bieler et al., 2014 [Publ 3622], fig. 28

iT tT bi riS oT roT roS iD
1 1 0 1 4 3 2 --
     +-- Aspatharia_pfeifferiana **    
  +--+                              
  |  +-- Margaritifera_margaritifera
--+                                 
  |  +-- Unio_pictorum              
  +--+                              
     +-- Velesunio_ambiguus         

González & Giribet, 2015 [Publ 3632], fig. 2

iT tT bi riS oT roT roS iD
1 1 0 1 5 3 2 --
     +-- Aspatharia_pfeifferiana **    
  +--+                              
  |  +-- Margaritifera_margaritifera
--+                                 
  |  +-- Unio_pictorum              
  +--+                              
     +-- Velesunio_ambiguus         

González & Giribet, 2015 [Publ 3632], fig. 3

iT tT bi riS oT roT roS iD
1 1 0 1 5 3 2 --
  +----- Unio_pictorum              
  |                                 
--+----- Velesunio_ambiguus         
  |                                 
  |  +-- Aspatharia_pfeifferiana **    
  +--+                              
     +-- Margaritifera_margaritifera

González & Giribet, 2015 [Publ 3632], fig. 4

iT tT bi riS oT roT roS iD
1 1 0 1 5 3 2 --
  +-------- Aspatharia_pfeifferiana **    
--+                                    
  |  +----- Velesunio_ambiguus         
  +--+                                 
     |  +-- Unio_pictorum              
     +--+                              
        +-- Margaritifera_margaritifera

González & Giribet, 2015 [Publ 3632], fig. 6

iT tT bi riS oT roT roS iD
1 1 0 1 5 3 2 --
  +----- Unio_pictorum              
--+                                 
  |  +-- Aspatharia_pfeifferiana **    
  +--+                              
     +-- Margaritifera_margaritifera

Wu et al., 2019 [Publ 3864], fig. 3

iT tT bi riS oT roT roS iD
1 1 0 1 46 46 2 --
           +-- Echyridella_menziesii   
  +--------+                           
  |        +-- Mutela_dubia **            
--+                                    
  |  +-------- Gibbosula_rochechouartii
  +--+                                 
     |  +----- Cumberlandia_monodonta  
     +--+                              
        |  +-- Margaritifera_dahurica  
        +--+                           
           +-- Margaritifera_falcata   

Page updated: 15:06:31 Tue 09 Jul 2024

 
NSF icon MUSSEL icon
"Making the world a better place, one mollusk at a time."