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Page last updated
Wed 04 Dec 2024

The Freshwater Mussels (Unionoida) of the World (and other less consequential bivalves)

FM(U)otW(aolcb) is the web version of the MUSSEL Project Database. Follow the links to browse the data or use the custom Google search field. Either way, you win!

Cladomics pages are experimental — even more so than the rest of this site!

The purpose of the cladomics pages for each genus and family-group level taxon is to provide statistics and references indicating the extent to which published phylogenetic work supports the classification used here.

The Cladomics Summary provides an overview of the number of studies that have included terminal species representing this taxon as well as an indication of how frequently it has been recovered as monophyletic.

Following the summary, the available published clades representing the taxon are listed.

For more information, visit the Cladomics FARQ (frequently asked rhetorical questions) page. Abbreviations for the various statistics are explained on the FARQ!

Search the database pages:

Bivalvia | Unionoida | Unionoidea | Margaritiferidae

subfamily GIBBOSULINAE Bogan, Bolotov, Froufe & Lopes-Lima in Lopes-Lima et al., 2018: cladomics
     1 genus

Cladomics Summary

number of species = 4
number of subtaxa (rank: genus) = 1

analyzed in 68 cladograms in 37 publications
maximum number of terminals in an analysis = 3, mean = 1.31
maximum number of subtaxa (genus) in an analysis = 1, mean = 1.0

Basic Monophyly

number of cladograms that test basic monophyly (i.e., more than 1 included species) = 18
maximum number of included terminals = 3, mean = 2.17
maximum number of subtaxa (genus) = 1, mean = 1.0
proportion of cladograms that support basic monophyly = 0.78


Pairwise Cladogram Clustering

The graphs below depict in two dimensions all pair-wise ingroup distances within the cladome. See the Cladmoics FARQ for more information.


PCoA Ordination


NJ tree


Lyubas et al., 2019 [Publ 3963], fig. 10

iT tT bi riS oT roT roS iD
3 3 1 1 13 13 1 0.0

*** Top Monophyly Cladogram *** *** Top Sister-Group Cladogram ***

  +-- Gibbosula **       
--+                   
  +-- Margaritiferinae

Lopes-Lima et al., 2018 [Publ 3829], fig. 3

iT tT bi riS oT roT roS iD
3 3 1 1 25 11 1 0.0
  +-- Gibbosula **       
--+                   
  +-- Margaritiferinae

Lopes-Lima et al., 2018 [Publ 3829], fig. 1

iT tT bi riS oT roT roS iD
3 3 1 1 25 11 1 0.0
  +-- Gibbosula **       
--+                   
  +-- Margaritiferinae

Wu et al., 2021 [Publ 4366], fig. 3-ML

iT tT bi riS oT roT roS iD
2 2 0 1 72 5 1 --
  +-- Gibbosula **       
--+                   
  +-- Margaritiferinae

Sano et al., 2020 [Publ 4138], fig. 2

iT tT bi riS oT roT roS iD
2 2 0 1 67 10 1 --
  +-------- Cumberlandia 
--+                      
  |  +----- Margaritifera
  +--+                   
     |  +-- Gibbosula **    
     +--+                
        +-- Pseudunio    

Sano et al., 2020 [Publ 4138], fig. 4

iT tT bi riS oT roT roS iD
2 2 0 1 67 10 1 --
  +-------- Cumberlandia 
--+                      
  |  +----- Pseudunio    
  +--+                   
     |  +-- Gibbosula **    
     +--+                
        +-- Margaritifera

Huang et al., 2019 [Publ 3843], fig. S4

iT tT bi riS oT roT roS iD
2 13 0 1 196 11 1 12.0
  +----- Gibbosula_rochechouartii **
--+                              
  |  +-- Gibbosula_laosensis **     
  +--+                           
     +-- Margaritiferinae        

Sano et al., 2020 [Publ 4138], fig. 3

iT tT bi riS oT roT roS iD
2 2 0 1 67 10 1 --
  +-------- Cumberlandia 
--+                      
  |  +----- Pseudunio    
  +--+                   
     |  +-- Gibbosula **    
     +--+                
        +-- Margaritifera

Huang et al., 2019 [Publ 3843], fig. S3

iT tT bi riS oT roT roS iD
2 13 0 1 194 11 1 12.0
  +----- Gibbosula_rochechouartii **
--+                              
  |  +-- Gibbosula_laosensis **     
  +--+                           
     +-- Margaritiferinae        

Wu et al., 2021 [Publ 4366], fig. 3-BI

iT tT bi riS oT roT roS iD
2 2 0 1 72 5 1 --
  +-- Gibbosula **       
--+                   
  +-- Margaritiferinae

Wu et al., 2021 [Publ 4366], fig. 4

iT tT bi riS oT roT roS iD
2 2 0 1 67 5 1 --
  +-- Gibbosula **       
--+                   
  +-- Margaritiferinae

Wu et al., 2023 [Publ 4339], fig. 4_S5-aa

iT tT bi riS oT roT roS iD
2 2 0 1 94 5 1 --
  +-- Gibbosula **       
--+                   
  +-- Margaritiferinae

Wu et al., 2020 [Publ 4121], fig. S1

iT tT bi riS oT roT roS iD
2 12 0 1 32 10 1 11.0
  +----- Gibbosula_rochechouartii **
--+                              
  |  +-- Gibbosula_laosensis **     
  +--+                           
     +-- Margaritiferinae        

Huang et al., 2018 [Publ 3828], fig. 3_5

iT tT bi riS oT roT roS iD
2 2 0 1 27 10 1 --
  +-- Gibbosula **       
--+                   
  +-- Margaritiferinae

Wu et al., 2023 [Publ 4339], fig. 4_S5-nt

iT tT bi riS oT roT roS iD
2 2 0 1 94 5 1 --
  +-- Gibbosula **       
--+                   
  +-- Margaritiferinae

Wu et al., 2020 [Publ 4121], fig. 2

iT tT bi riS oT roT roS iD
2 2 0 1 32 10 1 --
  +-------- Cumberlandia 
--+                      
  |  +----- Margaritifera
  +--+                   
     |  +-- Gibbosula **    
     +--+                
        +-- Pseudunio    

Gomes-dos-Santos et al., 2019 [Publ 3943], fig. 1

iT tT bi riS oT roT roS iD
2 2 0 1 8 5 1 --
  +-- Gibbosula **       
--+                   
  +-- Margaritiferinae

Huang et al., 2019 [Publ 3843], fig. 2

iT tT bi riS oT roT roS iD
2 13 0 1 196 11 1 12.0
  +----- Gibbosula_rochechouartii **
--+                              
  |  +-- Gibbosula_laosensis **     
  +--+                           
     +-- Margaritiferinae        

Bolotov et al., 2017 [Publ 3761], fig. S1

iT tT bi riS oT roT roS iD
1 1 0 1 138 9 1 --
  +-------- Cumberlandia 
--+                      
  |  +----- Margaritifera
  +--+                   
     |  +-- Gibbosula **    
     +--+                
        +-- Pseudunio    

Dai et al., 2024 [Publ 4358], fig. 4

iT tT bi riS oT roT roS iD
1 1 0 1 51 1 1 --
  +-- Gibbosula **       
--+                   
  +-- Margaritiferinae

Vikhrev et al., 2017 [Publ 3810], fig. 4

iT tT bi riS oT roT roS iD
1 1 0 1 9 1 1 --
  +-- Gibbosula **       
--+                   
  +-- Margaritiferinae

Bolotov et al., 2022 [Publ 4250], fig. 8

iT tT bi riS oT roT roS iD
1 1 0 1 48 1 1 --
-- Gibbosula_laosensis

Bogan et al., 2023 [Publ 4295], fig. 6

iT tT bi riS oT roT roS iD
1 1 0 1 12 0 0 --
-- Gibbosula_crassa

Dai et al., 2024 [Publ 4389], fig. 3

iT tT bi riS oT roT roS iD
1 1 0 1 46 1 1 --
  +-- Margaritiferinae
--+                   
  +-- Gibbosula **       

Konopleva et al., 2023 [Publ 4269], fig. S1_4

iT tT bi riS oT roT roS iD
1 1 0 1 55 0 0 --
-- Gibbosula_laosensis

Vikhrev et al., 2018 [Publ 3812], fig. 3

iT tT bi riS oT roT roS iD
1 1 0 1 12 10 1 --
  +-- Gibbosula **       
--+                   
  +-- Margaritiferinae

Dai et al., 2024 [Publ 4386], fig. 5-BI

iT tT bi riS oT roT roS iD
1 1 0 1 20 1 1 --
  +-- Margaritiferinae
--+                   
  +-- Gibbosula **       

Araujo et al., 2016 [Publ 3746], fig. 2a

iT tT bi riS oT roT roS iD
1 1 0 1 10 10 1 --
  +-------- Cumberlandia 
--+                      
  |  +----- Pseudunio    
  +--+                   
     |  +-- Gibbosula **    
     +--+                
        +-- Margaritifera

Dai et al., 2023 [Publ 4285], fig. 3-BI

iT tT bi riS oT roT roS iD
1 1 0 1 44 1 1 --
  +-- Gibbosula **       
--+                   
  +-- Margaritiferinae

Huang et al., 2019 [Publ 3843], fig. 1

iT tT bi riS oT roT roS iD
1 1 0 1 41 1 1 --
  +-- Margaritiferinae
--+                   
  +-- Gibbosula **       

Huang et al., 2018 [Publ 3828], fig. 6

iT tT bi riS oT roT roS iD
1 1 0 1 42 5 1 --
  +-- Gibbosula **       
--+                   
  +-- Margaritiferinae

Wu et al., 2021 [Publ 4366], fig. 2_I

iT tT bi riS oT roT roS iD
1 1 0 1 90 6 1 --
  +-- Gibbosula **       
--+                   
  +-- Margaritiferinae

Sano et al., 2020 [Publ 4138], fig. 1

iT tT bi riS oT roT roS iD
1 1 0 1 33 2 1 --
  +-- Gibbosula **       
--+                   
  +-- Margaritiferinae

Chen et al., 2023 [Publ 4302], fig. 2

iT tT bi riS oT roT roS iD
1 1 0 1 39 1 1 --
  +-- Gibbosula **       
--+                   
  +-- Margaritiferinae

Bolotov et al., 2016 [Publ 3743], fig. 4

iT tT bi riS oT roT roS iD
1 1 0 1 29 9 1 --
  +-------- Pseudunio    
--+                      
  |  +----- Margaritifera
  +--+                   
     |  +-- Gibbosula **    
     +--+                
        +-- Cumberlandia 

Bolotov et al., 2016 [Publ 3743], fig. 2

iT tT bi riS oT roT roS iD
1 1 0 1 29 9 1 --
  +-------- Pseudunio    
--+                      
  |  +----- Margaritifera
  +--+                   
     |  +-- Gibbosula **    
     +--+                
        +-- Cumberlandia 

Zhou et al., 2007 [Publ 2658], fig. 2

iT tT bi riS oT roT roS iD
1 1 0 1 20 0 0 --
-- Gibbosula_rochechouartii

Dai et al., 2024 [Publ 4385], fig. 3-ML

iT tT bi riS oT roT roS iD
1 1 0 1 30 1 1 --
  +-- Gibbosula **       
--+                   
  +-- Margaritiferinae

Wu et al., 2019 [Publ 3864], fig. 3

iT tT bi riS oT roT roS iD
1 1 0 1 46 3 1 --
  +-- Gibbosula **       
--+                   
  +-- Margaritiferinae

Dai et al., 2023 [Publ 4285], fig. 5

iT tT bi riS oT roT roS iD
1 1 0 1 52 1 1 --
  +-- Gibbosula **       
--+                   
  +-- Margaritiferinae

Guerra et al., 2019 [Publ 4168], fig. 2

iT tT bi riS oT roT roS iD
1 1 0 1 40 6 1 --
  +-- Gibbosula **       
--+                   
  +-- Margaritiferinae

Bolotov et al., 2016 [Publ 3732], fig. 2

iT tT bi riS oT roT roS iD
1 1 0 1 7 1 1 --
  +-- Gibbosula **       
--+                   
  +-- Margaritiferinae

Konopleva et al., 2019 [Publ 3900], fig. 1

iT tT bi riS oT roT roS iD
1 1 0 1 51 1 1 --
  +-- Gibbosula **       
--+                   
  +-- Margaritiferinae

Bogan et al., 2023 [Publ 4295], fig. 4

iT tT bi riS oT roT roS iD
1 1 0 1 6 0 0 --
-- Gibbosula_crassa

Dai et al., 2024 [Publ 4386], fig. 5-ML

iT tT bi riS oT roT roS iD
1 1 0 1 20 1 1 --
  +-- Margaritiferinae
--+                   
  +-- Gibbosula **       

Bolotov et al., 2020 [Publ 4100], fig. 1

iT tT bi riS oT roT roS iD
1 1 0 1 117 4 1 --
  +-- Gibbosula **                  
  |                              
  +-- Margaritifera_dahurica     
  |                              
--+-- Margaritifera_margaritifera
  |                              
  +-- Margaritifera_laevis       
  |                              
  +-- Margaritifera_middendorffi 

Bespalaya et al., 2018 [Publ 3811], fig. 3

iT tT bi riS oT roT roS iD
1 1 0 1 9 1 1 --
  +-- Gibbosula **       
--+                   
  +-- Margaritiferinae

Jeratthitikul et al., 2022 [Publ 4260], fig. S1_1

iT tT bi riS oT roT roS iD
1 1 0 1 75 9 1 --
  +-------- Cumberlandia 
--+                      
  |  +----- Gibbosula **    
  +--+                   
     |  +-- Pseudunio    
     +--+                
        +-- Margaritifera

Huang et al., 2019 [Publ 3843], fig. S1

iT tT bi riS oT roT roS iD
1 1 0 1 44 1 1 --
  +-- Gibbosula **       
--+                   
  +-- Margaritiferinae

Wu et al., 2022 [Publ 4238], fig. 3

iT tT bi riS oT roT roS iD
1 1 0 1 73 5 1 --
  +-- Gibbosula **       
--+                   
  +-- Margaritiferinae

Chen et al., 2023 [Publ 4302], fig. 1

iT tT bi riS oT roT roS iD
1 1 0 1 39 1 1 --
  +-- Gibbosula **       
--+                   
  +-- Margaritiferinae

Jeratthitikul et al., 2022 [Publ 4260], fig. S2_2

iT tT bi riS oT roT roS iD
1 1 0 1 75 9 1 --
  +-------- Cumberlandia 
--+                      
  |  +----- Margaritifera
  +--+                   
     |  +-- Gibbosula **    
     +--+                
        +-- Pseudunio    

Bolotov et al., 2017 [Publ 3761], fig. S2

iT tT bi riS oT roT roS iD
1 1 0 1 138 9 1 --
  +----- Cumberlandia 
  |                   
--+  +-- Gibbosula **    
  |  |                
  +--+-- Pseudunio    
     |                
     +-- Margaritifera

Zhou et al., 2007 [Publ 2658], fig. 1

iT tT bi riS oT roT roS iD
1 1 0 1 22 2 1 --
-- Gibbosula_rochechouartii

Bogan et al., 2023 [Publ 4295], fig. 3

iT tT bi riS oT roT roS iD
1 1 0 1 30 1 1 --
  +-- Gibbosula **       
--+                   
  +-- Margaritiferinae

Araujo et al., 2016 [Publ 3746], fig. 1_2b

iT tT bi riS oT roT roS iD
1 1 0 1 12 10 1 --
  +-- Gibbosula **       
--+                   
  +-- Margaritiferinae

Dai et al., 2024 [Publ 4385], fig. 3-BI

iT tT bi riS oT roT roS iD
1 1 0 1 30 1 1 --
  +-- Gibbosula **       
--+                   
  +-- Margaritiferinae

Wu et al., 2024 [Publ 4379], fig. 5

iT tT bi riS oT roT roS iD
1 1 0 1 53 1 1 --
  +-- Gibbosula **       
--+                   
  +-- Margaritiferinae

Guerra et al., 2019 [Publ 4168], fig. 1

iT tT bi riS oT roT roS iD
1 1 0 1 40 6 1 --
  +-- Gibbosula **       
--+                   
  +-- Margaritiferinae

Pfeiffer & Graf, 2013 [Publ 3531], fig. 2

iT tT bi riS oT roT roS iD
1 1 0 1 33 1 1 --
-- Gibbosula_rochechouartii

Wu et al., 2021 [Publ 4366], fig. 6

iT tT bi riS oT roT roS iD
1 1 0 1 90 6 1 --
  +-- Gibbosula **       
--+                   
  +-- Margaritiferinae

Chen et al., 2020 [Publ 4170], fig. 1

iT tT bi riS oT roT roS iD
1 1 0 1 33 0 0 --
-- Gibbosula_crassa

Bogan et al., 2023 [Publ 4295], fig. 5

iT tT bi riS oT roT roS iD
1 1 0 1 5 0 0 --
-- Gibbosula_crassa

Wu et al., 2020 [Publ 4121], fig. 3

iT tT bi riS oT roT roS iD
1 1 0 1 49 3 1 --
  +-- Gibbosula **       
--+                   
  +-- Margaritiferinae

Fukata & Iigo, 2020 [Publ 4106], fig. 1

iT tT bi riS oT roT roS iD
1 1 0 1 53 2 1 --
  +-- Gibbosula **       
--+                   
  +-- Margaritiferinae

Dai et al., 2023 [Publ 4285], fig. 3-ML

iT tT bi riS oT roT roS iD
1 1 0 1 44 1 1 --
  +-- Gibbosula **       
--+                   
  +-- Margaritiferinae

Wu et al., 2020 [Publ 4147], fig. 4

iT tT bi riS oT roT roS iD
1 1 0 1 13 2 1 --
  +-- Gibbosula **       
--+                   
  +-- Margaritiferinae

Huang et al., 2019 [Publ 3843], fig. 3

iT tT bi riS oT roT roS iD
1 1 0 1 56 5 1 --
  +-- Gibbosula **       
--+                   
  +-- Margaritiferinae

Page updated: 14:54:15 Wed 04 Dec 2024

 
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