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Page last updated
Wed 18 Jan 2023

The Freshwater Mussels (Unionoida) of the World (and other less consequential bivalves)

FM(U)otW(aolcb) is the web version of the MUSSEL Project Database. Follow the links to browse the data or use the custom Google search field. Either way, you win!

Cladomics pages are experimental — even more so than the rest of this site!

The purpose of the cladomics pages for each genus and family-group level taxon is to provide statistics and references indicating the extent to which published phylogenetic work supports the classification used here.

The Cladomics Summary provides an overview of the number of studies that have included terminal species representing this taxon as well as an indication of how frequently it has been recovered as monophyletic.

Following the summary, the available published clades representing the taxon are listed.

For more information, visit the Cladomics FARQ (frequently asked rhetorical questions) page. Abbreviations for the various statistics are explained on the FARQ!

Search the database pages:

Bivalvia | Unionoida | Unionoidea | Unionidae | Parreysiinae

tribe LAMELLIDENTINI Modell, 1942: cladomics
     3 genera

Cladomics Summary

number of species = 20
number of subtaxa (rank: genus) = 3

analyzed in 36 cladograms in 23 publications
maximum number of terminal species in an analysis = 16, mean = 5.97
maximum number of subtaxa (genus) in an analysis = 2, mean = 1.72

Basic Monophyly

number of cladograms that test basic monophyly (i.e., more than 1 included species) = 27
maximum number of included species = 16, mean = 7.63
maximum number of subtaxa (genus) = 2, mean = 1.96
proportion of cladograms that support basic monophyly = 0.93

Rigorous Monophyly

number of cladograms that test rigorous monophyly = 25
(Rigorous monophyly refers to cladograms with terminal species representing more than one subtaxon (genus) taxon and at least one rigorous outgroup (tribe).
maximum number of included species = 16, mean = 7.96
maximum number of subtaxa (genus) = 2, mean = 2.0
proportion of cladograms that support rigorous monophyly = 0.96


Bolotov et al., 2017, fig. S1

iT tT bi riS oT roT roS iD
12 38 27 2 113 26 4 0.0

*** Top Monophyly Cladogram ***

                       +----- Trapezidens_exolescens **     
              +--------+                                 
              |        |  +-- Trapezidens_dolichorhynchus **
              |        +--+                              
  +-----------+           +-- Trapezidens_angustior **      
  |           |                                          
  |           |  +----------- Lamellidens_lamellatus **     
  |           |  |                                       
  |           +--+  +-------- Lamellidens_jenkinsianus **   
  |              |  |                                    
  |              |  |  +----- Lamellidens_generosus **      
  |              +--+  |                                 
  |                 |  +----- Lamellidens_mainwaringi **    
  |                 |  |                                 
  |                 +--+  +-- Lamellidens_ferrugineus **    
  |                    +--+                              
  |                    |  +-- Lamellidens_marginalis **     
--+                    |                                 
  |                    |  +-- Lamellidens_savadiensis **    
  |                    +--+                              
  |                       +-- Lamellidens_corrianus[1] **   
  |                                                      
  |                 +-------- Parreysia_keralaensis      
  |  +--------------+                                    
  |  |              |  +----- Parreysia_plagiosoma       
  |  |              +--+                                 
  |  |                 |  +-- Parreysia_corrugata        
  |  |                 +--+                              
  |  |                    +-- Parreysia_nagpoorensis     
  |  |                                                   
  |  |              +-------- Leoparreysia_olivacea      
  +--+  +-----------+                                    
     |  |           |  +----- Leoparreysia_tavoyensis[2] 
     |  |           +--+                                 
     |  |              |  +-- Leoparreysia_canefrii      
     |  |              +--+                              
     |  |                 +-- Leoparreysia_tavoyensis[1] 
     |  |                                                
     |  |                 +-- Nitia_teretiuscula         
     |  |                 |                              
     +--+                 +-- Coelatura_gabonensis       
        |                 |                              
        |                 +-- Prisodontopsis_mweruensis  
        |  +--------------+                              
        |  |              +-- Coelatura_kunenensis       
        |  |              |                              
        |  |              +-- Coelatura_aegyptiaca[1]    
        |  |              |                              
        |  |              +-- Coelatura_aegyptiaca[2]    
        |  |                                             
        +--+        +-------- Indonaia_shurtleffiana     
           |        |                                    
           |  +-----+  +----- Indonaia_rugosa            
           |  |     |  |                                 
           |  |     |  |  +-- Indonaia_andersoniana      
           |  |     +--+--+                              
           |  |        |  +-- Indonaia_caerulea          
           |  |        |                                 
           +--+        |  +-- Indonaia_bonneaudii        
              |        +--+                              
              |           +-- Lamellidens_corrianus[2] **   
              |                                          
              |  +----------- Indochinella_pugio         
              |  |                                       
              |  |        +-- Scabies_humilis            
              +--+        |                              
                 |  +-----+-- Scabies_phaselus           
                 |  |     |                              
                 +--+     +-- Scabies_mandarinus         
                    |                                    
                    |  +----- Radiatula_mouhoti          
                    +--+                                 
                       |  +-- Radiatula_myitkyinae       
                       +--+                              
                          +-- Radiatula_chaudhurii       

Bolotov et al., 2022, fig. S1

iT tT bi riS oT roT roS iD
16 16 13 2 45 41 4 41.0

*** Top Sister-Group Cladogram ***

                                               +-- Parreysia_keralaensis      
                                               |                              
                                               +-- Parreysia_corrugata        
                                               |                              
  +--------------------------------------------+-- Parreysia_plagiosoma       
  |                                            |                              
  |                                            +-- Parreysia_rakhinensis      
  |                                            |                              
  |                                            +-- Parreysia_nagpoorensis     
  |                                                                           
  |                                +-------------- Trapezidens_mogaungensis **   
  |                             +--+                                          
  |                             |  |  +----------- Trapezidens_scutum **         
  |                             |  +--+                                       
  |                             |     |  +-------- Trapezidens_exolescens **     
  |                             |     +--+                                    
  |                             |        |  +----- Trapezidens_yeti **           
  |                             |        +--+                                 
  |                             |           |  +-- Trapezidens_dolichorhynchus **
  |  +--------------------------+           +--+                              
--+  |                          |              +-- Trapezidens_angustior **      
  |  |                          |                                             
  |  |                          |        +-------- Lamellidens_savadiensis **    
  |  |                          |  +-----+                                    
  |  |                          |  |     |  +----- Lamellidens_chindwinensis **  
  |  |                          |  |     +--+                                 
  |  |                          |  |        |  +-- Lamellidens_corrianus **      
  |  |                          |  |        +--+                              
  |  |                          +--+           +-- Lamellidens_nongyangensis **  
  |  |                             |                                          
  |  |                             |           +-- Lamellidens_marginalis **     
  |  |                             |  +--------+                              
  |  |                             |  |        +-- Lamellidens_ferrugineus **    
  |  |                             +--+                                       
  |  |                                |  +-------- Lamellidens_generosus **      
  |  |                                +--+                                    
  |  |                                   |  +----- Lamellidens_mainwaringi **    
  +--+                                   +--+                                 
     |                                      |  +-- Lamellidens_jenkinsianus **   
     |                                      +--+                              
     |                                         +-- Lamellidens_lamellatus **     
     |                                                                        
     |                                         +-- Leoparreysia_burmana       
     |                                         |                              
     |                                         +-- Leoparreysia_canefrii      
     |                                         |                              
     |  +--------------------------------------+-- Leoparreysia_olivacea      
     |  |                                      |                              
     |  |                                      +-- Leoparreysia_tavoyensis    
     |  |                                      |                              
     |  |                                      +-- Leoparreysia_whitteni      
     |  |                                                                     
     |  |                                      +-- Coelatura_aegyptiaca       
     |  |                                      |                              
     |  |                                      +-- Coelatura_kunenensis       
     +--+  +-----------------------------------+                              
        |  |                                   +-- Nitia_teretiuscula         
        |  |                                   |                              
        |  |                                   +-- Nyassunio_nyassaensis      
        |  |                                                                  
        |  |                                   +-- Indonaia_cylindrica        
        |  |                          +--------+                              
        |  |                          |        +-- Indonaia_gratiosa          
        |  |                          |                                       
        +--+  +-----------------------+     +----- Indonaia_shurtleffiana     
           |  |                       |  +--+                                 
           |  |                       |  |  |  +-- Indonaia_bonneaudii        
           |  |                       |  |  +--+                              
           |  |                       +--+     +-- Indonaia_rugosa            
           |  |                          |                                    
           |  |                          |  +----- Indonaia_andersoniana      
           |  |                          +--+                                 
           |  |                             |  +-- Indonaia_caerulea          
           +--+                             +--+                              
              |                                +-- Indonaia_subclathrata      
              |                                                               
              |  +-------------------------------- Indochinella_pugio         
              |  |                                                            
              |  |                          +----- Radiatula_chaudhurii       
              |  |                    +-----+                                 
              |  |                    |     |  +-- Radiatula_kachinensis      
              |  |                    |     +--+                              
              +--+  +-----------------+        +-- Radiatula_myitkyinae       
                 |  |                 |                                       
                 |  |                 |  +-------- Radiatula_crispisulcata    
                 |  |                 +--+                                    
                 |  |                    |  +----- Radiatula_mouhoti          
                 |  |                    +--+                                 
                 +--+                       |  +-- Radiatula_myitthanensis    
                    |                       +--+                              
                    |                          +-- Radiatula_chindwinensis    
                    |                                                         
                    |  +-------------------------- Scabiellus_songkramensis   
                    |  |                                                      
                    +--+  +----------------------- Harmandia_somboriensis     
                       |  |                                                   
                       +--+  +-------------------- Unionetta_fabagina         
                          |  |                                                
                          +--+  +----------------- Scabies_crispata           
                             |  |                                             
                             +--+              +-- Scabies_humilis            
                                |  +-----------+                              
                                |  |           +-- Scabies_mandarinus         
                                +--+                                          
                                   |  +----------- Scabies_nucleus            
                                   +--+                                       
                                      |  +-------- Scabies_pilata             
                                      +--+                                    
                                         |  +----- Scabies_anceps             
                                         +--+                                 
                                            |  +-- Scabies_phaselus           
                                            +--+                              
                                               +-- Scabies_scobinatus         

Bolotov et al., 2022, fig. 2

iT tT bi riS oT roT roS iD
16 16 11 2 45 41 4 42.0
                                   +----------- Trapezidens_mogaungensis **   
                                   |                                       
                             +-----+  +-------- Trapezidens_exolescens **     
                             |     |  |                                    
                             |     +--+  +----- Trapezidens_scutum **         
                             |        |  |                                 
                             |        +--+  +-- Trapezidens_angustior **      
                             |           |  |                              
  +--------------------------+           +--+-- Trapezidens_dolichorhynchus **
  |                          |              |                              
  |                          |              +-- Trapezidens_yeti **           
  |                          |                                             
  |                          |  +-------------- Lamellidens_marginalis **     
  |                          |  |                                          
  |                          |  |  +----------- Lamellidens_ferrugineus **    
  |                          +--+  |                                       
  |                             |  |  +-------- Lamellidens_generosus **      
  |                             |  +--+                                    
  |                             |  |  |  +----- Lamellidens_mainwaringi **    
  |                             +--+  +--+                                 
  |                                |     |  +-- Lamellidens_lamellatus **     
  |                                |     +--+                              
  |                                |        +-- Lamellidens_jenkinsianus **   
  |                                |                                       
  |                                |  +-------- Lamellidens_nongyangensis **  
--+                                +--+                                    
  |                                   |  +----- Lamellidens_savadiensis **    
  |                                   +--+                                 
  |                                      |  +-- Lamellidens_corrianus **      
  |                                      +--+                              
  |                                         +-- Lamellidens_chindwinensis **  
  |                                                                        
  |                                +----------- Parreysia_keralaensis      
  |  +-----------------------------+                                       
  |  |                             |  +-------- Parreysia_corrugata        
  |  |                             +--+                                    
  |  |                                |  +----- Parreysia_rakhinensis      
  |  |                                +--+                                 
  |  |                                   |  +-- Parreysia_nagpoorensis     
  |  |                                   +--+                              
  |  |                                      +-- Parreysia_plagiosoma       
  |  |                                                                     
  |  |                                      +-- Leoparreysia_burmana       
  +--+                                +-----+                              
     |                                |     +-- Leoparreysia_whitteni      
     |  +-----------------------------+                                    
     |  |                             |  +----- Leoparreysia_olivacea      
     |  |                             +--+                                 
     |  |                                |  +-- Leoparreysia_canefrii      
     |  |                                +--+                              
     |  |                                   +-- Leoparreysia_tavoyensis    
     |  |                                                                  
     |  |                             +-------- Nitia_teretiuscula         
     +--+  +--------------------------+                                    
        |  |                          |  +----- Coelatura_aegyptiaca       
        |  |                          +--+                                 
        |  |                             |  +-- Nyassunio_nyassaensis      
        |  |                             +--+                              
        |  |                                +-- Coelatura_kunenensis       
        |  |                                                               
        |  |                 +----------------- Indonaia_cylindrica        
        |  |                 |                                             
        +--+  +--------------+  +-------------- Indonaia_gratiosa          
           |  |              |  |                                          
           |  |              +--+  +----------- Indonaia_andersoniana      
           |  |                 |  |                                       
           |  |                 +--+        +-- Indonaia_caerulea          
           |  |                    |  +-----+                              
           |  |                    |  |     +-- Indonaia_subclathrata      
           |  |                    +--+                                    
           |  |                       |  +----- Indonaia_rugosa            
           +--+                       +--+                                 
              |                          |  +-- Indonaia_bonneaudii        
              |                          +--+                              
              |                             +-- Indonaia_shurtleffiana     
              |                                                            
              |  +----------------------------- Indochinella_pugio         
              |  |                                                         
              |  |                 +----------- Radiatula_crispisulcata    
              |  |                 |                                       
              |  |                 |     +----- Radiatula_chaudhurii       
              +--+  +--------------+  +--+                                 
                 |  |              |  |  |  +-- Radiatula_myitkyinae       
                 |  |              |  |  +--+                              
                 |  |              +--+     +-- Radiatula_kachinensis      
                 |  |                 |                                    
                 |  |                 |  +----- Radiatula_chindwinensis    
                 +--+                 +--+                                 
                    |                    |  +-- Radiatula_myitthanensis    
                    |                    +--+                              
                    |                       +-- Radiatula_mouhoti          
                    |                                                      
                    |  +----------------------- Scabiellus_songkramensis   
                    |  |                                                   
                    +--+  +-------------------- Harmandia_somboriensis     
                       |  |                                                
                       +--+  +----------------- Unionetta_fabagina         
                          |  |                                             
                          +--+              +-- Scabies_crispata           
                             |  +-----------+                              
                             |  |           +-- Scabies_mandarinus         
                             +--+                                          
                                |  +----------- Scabies_nucleus            
                                |  |                                       
                                +--+        +-- Scabies_humilis            
                                   |  +-----+                              
                                   |  |     +-- Scabies_pilata             
                                   +--+                                    
                                      |  +----- Scabies_phaselus           
                                      +--+                                 
                                         |  +-- Scabies_anceps             
                                         +--+                              
                                            +-- Scabies_scobinatus         

Bolotov et al., 2022, fig. 2

iT tT bi riS oT roT roS iD
14 14 11 2 44 40 4 37.0
                                               +-- Trapezidens_exolescens **     
                                         +-----+                              
                                         |     +-- Trapezidens_scutum **         
                             +-----------+                                    
                             |           |  +----- Trapezidens_angustior **      
                             |           +--+                                 
                             |              |  +-- Trapezidens_dolichorhynchus **
  +--------------------------+              +--+                              
  |                          |                 +-- Trapezidens_yeti **           
  |                          |                                                
  |                          |  +----------------- Lamellidens_corrianus **      
  |                          |  |                                             
  |                          +--+  +-------------- Lamellidens_nongyangensis **  
  |                             |  |                                          
  |                             +--+  +----------- Lamellidens_savadiensis **    
  |                                |  |                                       
  |                                |  |        +-- Lamellidens_ferrugineus **    
  |                                +--+  +-----+                              
  |                                   |  |     +-- Lamellidens_marginalis **     
  |                                   |  |                                    
  |                                   +--+     +-- Lamellidens_lamellatus **     
  |                                      |  +--+                              
  |                                      |  |  +-- Lamellidens_jenkinsianus **   
  |                                      +--+                                 
  |                                         |  +-- Lamellidens_generosus **      
  |                                         +--+                              
--+                                            +-- Lamellidens_mainwaringi **    
  |                                                                           
  |                                   +----------- Parreysia_keralaensis      
  |                                +--+                                       
  |                                |  |  +-------- Parreysia_plagiosoma       
  |                                |  +--+                                    
  |                                |     |  +----- Parreysia_nagpoorensis     
  |                                |     +--+                                 
  |                                |        |  +-- Parreysia_corrugata        
  |  +-----------------------------+        +--+                              
  |  |                             |           +-- Parreysia_rakhinensis      
  |  |                             |                                          
  |  |                             |           +-- Leoparreysia_burmana       
  |  |                             |     +-----+                              
  |  |                             |     |     +-- Leoparreysia_whitteni      
  |  |                             +-----+                                    
  |  |                                   |  +----- Leoparreysia_olivacea      
  |  |                                   +--+                                 
  |  |                                      |  +-- Leoparreysia_canefrii      
  |  |                                      +--+                              
  +--+                                         +-- Leoparreysia_tavoyensis    
     |                                                                        
     |                                   +-------- Nitia_teretiuscula         
     |  +--------------------------------+                                    
     |  |                                |  +----- Coelatura_aegyptiaca       
     |  |                                +--+                                 
     |  |                                   |  +-- Coelatura_kunenensis       
     |  |                                   +--+                              
     |  |                                      +-- Nyassunio_nyassaensis      
     |  |                                                                     
     |  |                                      +-- Indonaia_cylindrica        
     |  |                             +--------+                              
     +--+                             |        +-- Indonaia_gratiosa          
        |                             |                                       
        |  +--------------------------+     +----- Indonaia_andersoniana      
        |  |                          |  +--+                                 
        |  |                          |  |  |  +-- Indonaia_caerulea          
        |  |                          |  |  +--+                              
        |  |                          +--+     +-- Indonaia_subclathrata      
        |  |                             |                                    
        |  |                             |  +----- Indonaia_shurtleffiana     
        |  |                             +--+                                 
        +--+                                |  +-- Indonaia_bonneaudii        
           |                                +--+                              
           |                                   +-- Indonaia_rugosa            
           |                                                                  
           |  +----------------------------------- Indochinella_pugio         
           |  |                                                               
           |  |                       +----------- Radiatula_crispisulcata    
           |  |                       |                                       
           +--+  +--------------------+        +-- Radiatula_chaudhurii       
              |  |                    |  +-----+                              
              |  |                    |  |     +-- Radiatula_myitkyinae       
              |  |                    +--+                                    
              |  |                       |  +----- Radiatula_mouhoti          
              +--+                       +--+                                 
                 |                          |  +-- Radiatula_chindwinensis    
                 |                          +--+                              
                 |                             +-- Radiatula_myitthanensis    
                 |                                                            
                 |  +----------------------------- Scabiellus_songkramensis   
                 +--+                                                         
                    |  +-------------------------- Harmandia_somboriensis     
                    +--+                                                      
                       |  +----------------------- Unionetta_fabagina         
                       +--+                                                   
                          |  +-------------------- Scabies_crispata           
                          +--+                                                
                             |  +----------------- Scabies_mandarinus         
                             +--+                                             
                                |  +-------------- Scabies_nucleus            
                                +--+                                          
                                   |  +----------- Scabies_pilata             
                                   +--+                                       
                                      |  +-------- Scabies_humilis            
                                      +--+                                    
                                         |  +----- Scabies_anceps             
                                         +--+                                 
                                            |  +-- Scabies_phaselus           
                                            +--+                              
                                               +-- Scabies_scobinatus         

Bolotov et al., 2022, fig. S1

iT tT bi riS oT roT roS iD
14 14 10 2 44 40 4 39.0
                                      +-------- Trapezidens_exolescens **     
                                      |                                    
                             +--------+  +----- Trapezidens_scutum **         
                             |        |  |                                 
                             |        +--+  +-- Trapezidens_angustior **      
                             |           |  |                              
                             |           +--+-- Trapezidens_dolichorhynchus **
                             |              |                              
  +--------------------------+              +-- Trapezidens_yeti **           
  |                          |                                             
  |                          |           +----- Lamellidens_savadiensis **    
  |                          |  +--------+                                 
  |                          |  |        |  +-- Lamellidens_corrianus **      
  |                          |  |        +--+                              
  |                          |  |           +-- Lamellidens_nongyangensis **  
  |                          +--+                                          
  |                             |           +-- Lamellidens_ferrugineus **    
  |                             |  +--------+                              
  |                             |  |        +-- Lamellidens_marginalis **     
  |                             +--+                                       
  |                                |  +-------- Lamellidens_lamellatus **     
  |                                +--+                                    
  |                                   |  +----- Lamellidens_jenkinsianus **   
--+                                   +--+                                 
  |                                      |  +-- Lamellidens_generosus **      
  |                                      +--+                              
  |                                         +-- Lamellidens_mainwaringi **    
  |                                                                        
  |                                +----------- Parreysia_keralaensis      
  |  +-----------------------------+                                       
  |  |                             |  +-------- Parreysia_corrugata        
  |  |                             +--+                                    
  |  |                                |  +----- Parreysia_rakhinensis      
  |  |                                +--+                                 
  |  |                                   |  +-- Parreysia_plagiosoma       
  |  |                                   +--+                              
  |  |                                      +-- Parreysia_nagpoorensis     
  |  |                                                                     
  |  |                                      +-- Leoparreysia_burmana       
  +--+                                +-----+                              
     |                                |     +-- Leoparreysia_whitteni      
     |  +-----------------------------+                                    
     |  |                             |  +----- Leoparreysia_olivacea      
     |  |                             +--+                                 
     |  |                                |  +-- Leoparreysia_canefrii      
     |  |                                +--+                              
     |  |                                   +-- Leoparreysia_tavoyensis    
     |  |                                                                  
     |  |                             +-------- Nitia_teretiuscula         
     +--+  +--------------------------+                                    
        |  |                          |  +----- Coelatura_aegyptiaca       
        |  |                          +--+                                 
        |  |                             |  +-- Coelatura_kunenensis       
        |  |                             +--+                              
        |  |                                +-- Nyassunio_nyassaensis      
        |  |                                                               
        |  |                 +----------------- Indonaia_cylindrica        
        |  |                 |                                             
        +--+  +--------------+  +-------------- Indonaia_gratiosa          
           |  |              |  |                                          
           |  |              +--+  +----------- Indonaia_andersoniana      
           |  |                 |  |                                       
           |  |                 +--+        +-- Indonaia_caerulea          
           |  |                    |  +-----+                              
           |  |                    |  |     +-- Indonaia_subclathrata      
           |  |                    +--+                                    
           |  |                       |  +----- Indonaia_rugosa            
           +--+                       +--+                                 
              |                          |  +-- Indonaia_bonneaudii        
              |                          +--+                              
              |                             +-- Indonaia_shurtleffiana     
              |                                                            
              |  +----------------------------- Indochinella_pugio         
              |  |                                                         
              |  |                    +-------- Radiatula_crispisulcata    
              |  |                    |                                    
              |  |  +-----------------+     +-- Radiatula_chaudhurii       
              +--+  |                 |  +--+                              
                 |  |                 |  |  +-- Radiatula_myitkyinae       
                 |  |                 +--+                                 
                 |  |                    |  +-- Radiatula_myitthanensis    
                 |  |                    |  |                              
                 +--+                    +--+-- Radiatula_mouhoti          
                    |                       |                              
                    |                       +-- Radiatula_chindwinensis    
                    |                                                      
                    |  +----------------------- Scabiellus_songkramensis   
                    |  |                                                   
                    +--+  +-------------------- Harmandia_somboriensis     
                       |  |                                                
                       +--+  +----------------- Unionetta_fabagina         
                          |  |                                             
                          +--+              +-- Scabies_crispata           
                             |  +-----------+                              
                             |  |           +-- Scabies_mandarinus         
                             +--+                                          
                                |  +----------- Scabies_nucleus            
                                |  |                                       
                                +--+        +-- Scabies_humilis            
                                   |  +-----+                              
                                   |  |     +-- Scabies_pilata             
                                   +--+                                    
                                      |  +----- Scabies_phaselus           
                                      +--+                                 
                                         |  +-- Scabies_anceps             
                                         +--+                              
                                            +-- Scabies_scobinatus         

Bolotov et al., 2022, fig. S2

iT tT bi riS oT roT roS iD
14 14 11 2 44 40 4 39.0
                                      +----------- Parreysia_keralaensis      
  +-----------------------------------+                                       
  |                                   |  +-------- Parreysia_corrugata        
  |                                   +--+                                    
  |                                      |  +----- Parreysia_nagpoorensis     
  |                                      +--+                                 
  |                                         |  +-- Parreysia_plagiosoma       
  |                                         +--+                              
  |                                            +-- Parreysia_rakhinensis      
  |                                                                           
  |                                   +----------- Trapezidens_scutum **         
  |                             +-----+                                       
  |                             |     |  +-------- Trapezidens_exolescens **     
  |                             |     +--+                                    
  |                             |        |  +----- Trapezidens_yeti **           
  |                             |        +--+                                 
  |                             |           |  +-- Trapezidens_dolichorhynchus **
--+                             |           +--+                              
  |  +--------------------------+              +-- Trapezidens_angustior **      
  |  |                          |                                             
  |  |                          |           +----- Lamellidens_savadiensis **    
  |  |                          |  +--------+                                 
  |  |                          |  |        |  +-- Lamellidens_corrianus **      
  |  |                          |  |        +--+                              
  |  |                          |  |           +-- Lamellidens_nongyangensis **  
  |  |                          +--+                                          
  |  |                             |           +-- Lamellidens_marginalis **     
  |  |                             |  +--------+                              
  |  |                             |  |        +-- Lamellidens_ferrugineus **    
  |  |                             +--+                                       
  |  |                                |  +-------- Lamellidens_lamellatus **     
  |  |                                +--+                                    
  |  |                                   |  +----- Lamellidens_jenkinsianus **   
  +--+                                   +--+                                 
     |                                      |  +-- Lamellidens_mainwaringi **    
     |                                      +--+                              
     |                                         +-- Lamellidens_generosus **      
     |                                                                        
     |                                +----------- Leoparreysia_burmana       
     |  +-----------------------------+                                       
     |  |                             |  +-------- Leoparreysia_whitteni      
     |  |                             +--+                                    
     |  |                                |  +----- Leoparreysia_olivacea      
     |  |                                +--+                                 
     |  |                                   |  +-- Leoparreysia_canefrii      
     |  |                                   +--+                              
     |  |                                      +-- Leoparreysia_tavoyensis    
     |  |                                                                     
     +--+                                +-------- Nitia_teretiuscula         
        |  +-----------------------------+                                    
        |  |                             |  +----- Coelatura_aegyptiaca       
        |  |                             +--+                                 
        |  |                                |  +-- Coelatura_kunenensis       
        |  |                                +--+                              
        |  |                                   +-- Nyassunio_nyassaensis      
        |  |                                                                  
        |  |                                   +-- Indonaia_cylindrica        
        |  |                          +--------+                              
        +--+                          |        +-- Indonaia_gratiosa          
           |                          |                                       
           |  +-----------------------+     +----- Indonaia_shurtleffiana     
           |  |                       |  +--+                                 
           |  |                       |  |  |  +-- Indonaia_bonneaudii        
           |  |                       |  |  +--+                              
           |  |                       +--+     +-- Indonaia_rugosa            
           |  |                          |                                    
           |  |                          |  +----- Indonaia_andersoniana      
           |  |                          +--+                                 
           +--+                             |  +-- Indonaia_caerulea          
              |                             +--+                              
              |                                +-- Indonaia_subclathrata      
              |                                                               
              |  +-------------------------------- Indochinella_pugio         
              |  |                                                            
              |  |                    +----------- Radiatula_crispisulcata    
              |  |                    |                                       
              |  |  +-----------------+        +-- Radiatula_chaudhurii       
              +--+  |                 |  +-----+                              
                 |  |                 |  |     +-- Radiatula_myitkyinae       
                 |  |                 +--+                                    
                 |  |                    |  +----- Radiatula_mouhoti          
                 |  |                    +--+                                 
                 +--+                       |  +-- Radiatula_myitthanensis    
                    |                       +--+                              
                    |                          +-- Radiatula_chindwinensis    
                    |                                                         
                    |  +-------------------------- Scabiellus_songkramensis   
                    |  |                                                      
                    +--+  +----------------------- Harmandia_somboriensis     
                       |  |                                                   
                       +--+  +-------------------- Unionetta_fabagina         
                          |  |                                                
                          +--+  +----------------- Scabies_crispata           
                             |  |                                             
                             +--+              +-- Scabies_humilis            
                                |  +-----------+                              
                                |  |           +-- Scabies_mandarinus         
                                +--+                                          
                                   |  +----------- Scabies_nucleus            
                                   +--+                                       
                                      |  +-------- Scabies_pilata             
                                      +--+                                    
                                         |  +----- Scabies_anceps             
                                         +--+                                 
                                            |  +-- Scabies_phaselus           
                                            +--+                              
                                               +-- Scabies_scobinatus         

Huang et al., 2019, fig. 2

iT tT bi riS oT roT roS iD
12 12 9 2 187 24 4 35.0
                       +----- Trapezidens_exolescens **     
     +-----------------+                                 
     |                 |  +-- Trapezidens_dolichorhynchus **
     |                 +--+                              
  +--+                    +-- Trapezidens_angustior **      
  |  |                                                   
  |  |  +-------------------- Lamellidens_corrianus **      
  |  |  |                                                
  |  +--+  +----------------- Lamellidens_savadiensis **    
  |     |  |                                             
  |     +--+              +-- Lamellidens_lamellatus **     
  |        |  +-----------+                              
  |        |  |           +-- Lamellidens_jenkinsianus **   
  |        +--+                                          
  |           |  +----------- Lamellidens_generosus **      
  |           +--+                                       
  |              |  +-------- Lamellidens_mainwaringi[2] ** 
  |              +--+                                    
--+                 |  +----- Lamellidens_mainwaringi[1] ** 
  |                 +--+                                 
  |                    |  +-- Lamellidens_ferrugineus **    
  |                    +--+                              
  |                       +-- Lamellidens_marginalis **     
  |                                                      
  |                    +----- Parreysia_keralaensis      
  |  +-----------------+                                 
  |  |                 |  +-- Parreysia_nagpoorensis     
  |  |                 +--+                              
  |  |                    +-- Parreysia_plagiosoma       
  |  |                                                   
  |  |                    +-- Leoparreysia_tavoyensis    
  |  |                    |                              
  +--+  +-----------------+-- Leoparreysia_canefrii      
     |  |                 |                              
     |  |                 +-- Leoparreysia_olivacea      
     |  |                                                
     |  |           +-------- Coelatura_aegyptiaca[2]    
     |  |           |                                    
     |  |  +--------+  +----- Coelatura_aegyptiaca[1]    
     +--+  |        |  |                                 
        |  |        |  +----- Nitia_teretiuscula         
        |  |        +--+                                 
        |  |           |  +-- Coelatura_gabonensis       
        |  |           |  |                              
        |  |           +--+-- Coelatura_kunenensis       
        |  |              |                              
        +--+              +-- Prisodontopsis_mweruensis  
           |                                             
           |        +-------- Indonaia_shurtleffiana     
           |        |                                    
           |  +-----+  +----- Indonaia_bonneaudii        
           |  |     |  |                                 
           |  |     +--+----- Indonaia_rugosa            
           |  |        |                                 
           |  |        |  +-- Indonaia_andersoniana      
           +--+        +--+                              
              |           +-- Indonaia_caerulea          
              |                                          
              |  +----------- Indochinella_pugio         
              |  |                                       
              |  |     +----- Scabies_humilis            
              +--+  +--+                                 
                 |  |  |  +-- Scabies_phaselus           
                 |  |  +--+                              
                 +--+     +-- Scabies_mandarinus         
                    |                                    
                    |  +----- Radiatula_myitkyinae       
                    +--+                                 
                       |  +-- Radiatula_chaudhurii       
                       +--+                              
                          +-- Radiatula_mouhoti          

Huang et al., 2019, fig. S3

iT tT bi riS oT roT roS iD
12 12 6 2 185 24 4 35.0
                       +----- Trapezidens_exolescens **     
                 +-----+                                 
                 |     |  +-- Trapezidens_angustior **      
                 |     +--+                              
                 |        +-- Trapezidens_dolichorhynchus **
                 |                                       
  +--------------+        +-- Lamellidens_corrianus **      
  |              |  +-----+                              
  |              |  |     +-- Lamellidens_savadiensis **    
  |              |  |                                    
  |              |  |  +----- Lamellidens_lamellatus **     
  |              +--+  |                                 
  |                 |  +----- Lamellidens_jenkinsianus **   
  |                 |  |                                 
  |                 |  |  +-- Lamellidens_marginalis **     
  |                 +--+--+                              
  |                    |  +-- Lamellidens_ferrugineus **    
  |                    |                                 
--+                    |  +-- Lamellidens_mainwaringi[1] ** 
  |                    |  |                              
  |                    +--+-- Lamellidens_mainwaringi[2] ** 
  |                       |                              
  |                       +-- Lamellidens_generosus **      
  |                                                      
  |                    +----- Parreysia_keralaensis      
  |  +-----------------+                                 
  |  |                 |  +-- Parreysia_nagpoorensis     
  |  |                 +--+                              
  |  |                    +-- Parreysia_plagiosoma       
  |  |                                                   
  |  |                 +----- Leoparreysia_canefrii      
  +--+  +--------------+                                 
     |  |              |  +-- Leoparreysia_tavoyensis    
     |  |              +--+                              
     |  |                 +-- Leoparreysia_olivacea      
     |  |                                                
     |  |        +----------- Coelatura_aegyptiaca[2]    
     |  |        |                                       
     +--+  +-----+        +-- Coelatura_aegyptiaca[1]    
        |  |     |  +-----+                              
        |  |     |  |     +-- Nitia_teretiuscula         
        |  |     +--+                                    
        |  |        |  +----- Coelatura_kunenensis       
        |  |        +--+                                 
        |  |           |  +-- Coelatura_gabonensis       
        |  |           +--+                              
        +--+              +-- Prisodontopsis_mweruensis  
           |                                             
           |        +-------- Indonaia_shurtleffiana     
           |        |                                    
           |  +-----+  +----- Indonaia_bonneaudii        
           |  |     |  |                                 
           |  |     +--+----- Indonaia_rugosa            
           |  |        |                                 
           |  |        |  +-- Indonaia_andersoniana      
           +--+        +--+                              
              |           +-- Indonaia_caerulea          
              |                                          
              |  +----------- Indochinella_pugio         
              |  |                                       
              |  |     +----- Scabies_phaselus           
              +--+  +--+                                 
                 |  |  |  +-- Scabies_humilis            
                 |  |  +--+                              
                 +--+     +-- Scabies_mandarinus         
                    |                                    
                    |  +----- Radiatula_mouhoti          
                    +--+                                 
                       |  +-- Radiatula_myitkyinae       
                       +--+                              
                          +-- Radiatula_chaudhurii       

Huang et al., 2019, fig. S4

iT tT bi riS oT roT roS iD
12 12 5 2 187 24 4 34.0
                       +-- Parreysia_keralaensis      
                       |                              
  +--------------------+-- Parreysia_nagpoorensis     
  |                    |                              
  |                    +-- Parreysia_plagiosoma       
  |                                                   
  |                 +----- Trapezidens_exolescens **     
  |        +--------+                                 
  |        |        |  +-- Trapezidens_angustior **      
  |        |        +--+                              
  +--------+           +-- Trapezidens_dolichorhynchus **
  |        |                                          
  |        |  +----------- Lamellidens_corrianus **      
  |        |  |                                       
  |        +--+  +-------- Lamellidens_savadiensis **    
  |           |  |                                    
  |           |  |  +----- Lamellidens_marginalis **     
  |           +--+  |                                 
  |              |  +----- Lamellidens_ferrugineus **    
  |              |  |                                 
--+              +--+  +-- Lamellidens_mainwaringi[1] ** 
  |                 |  |                              
  |                 |  +-- Lamellidens_mainwaringi[2] ** 
  |                 |  |                              
  |                 +--+-- Lamellidens_lamellatus **     
  |                    |                              
  |                    +-- Lamellidens_generosus **      
  |                    |                              
  |                    +-- Lamellidens_jenkinsianus **   
  |                                                   
  |                 +----- Leoparreysia_tavoyensis    
  |  +--------------+                                 
  |  |              |  +-- Leoparreysia_canefrii      
  |  |              +--+                              
  |  |                 +-- Leoparreysia_olivacea      
  |  |                                                
  |  |           +-------- Coelatura_aegyptiaca[2]    
  |  |           |                                    
  |  |           +-------- Nitia_teretiuscula         
  +--+  +--------+                                    
     |  |        |  +----- Coelatura_aegyptiaca[1]    
     |  |        |  |                                 
     |  |        +--+  +-- Coelatura_gabonensis       
     |  |           |  |                              
     |  |           +--+-- Coelatura_kunenensis       
     |  |              |                              
     |  |              +-- Prisodontopsis_mweruensis  
     +--+                                             
        |           +----- Indonaia_shurtleffiana     
        |           |                                 
        |           |  +-- Indonaia_bonneaudii        
        |  +--------+--+                              
        |  |        |  +-- Indonaia_rugosa            
        |  |        |                                 
        |  |        |  +-- Indonaia_andersoniana      
        |  |        +--+                              
        +--+           +-- Indonaia_caerulea          
           |                                          
           |  +----------- Indochinella_pugio         
           |  |                                       
           |  |     +----- Scabies_humilis            
           +--+  +--+                                 
              |  |  |  +-- Scabies_phaselus           
              |  |  +--+                              
              +--+     +-- Scabies_mandarinus         
                 |                                    
                 |  +----- Radiatula_mouhoti          
                 +--+                                 
                    |  +-- Radiatula_myitkyinae       
                    +--+                              
                       +-- Radiatula_chaudhurii       

Konopleva et al., 2020, fig. 2

iT tT bi riS oT roT roS iD
9 9 6 2 3 3 1 35.0
                 +-- Leoparreysia_olivacea      
                 |                              
  +--------------+-- Leoparreysia_canefrii      
  |              |                              
  |              +-- Leoparreysia_tavoyensis    
  |                                             
--+        +-------- Lamellidens_savadiensis **    
  |  +-----+                                    
  |  |     |  +----- Lamellidens_generosus **      
  |  |     +--+                                 
  |  |        |  +-- Lamellidens_mainwaringi **    
  +--+        +--+                              
     |           +-- Lamellidens_ferrugineus **    
     |                                          
     |  +----------- Trapezidens_scutum **         
     +--+                                       
        |  +-------- Trapezidens_exolescens **     
        +--+                                    
           |  +----- Trapezidens_yeti **           
           +--+                                 
              |  +-- Trapezidens_angustior **      
              +--+                              
                 +-- Trapezidens_dolichorhynchus **

Bolotov et al., 2022, fig. 8

iT tT bi riS oT roT roS iD
9 9 6 2 41 15 3 35.0
                    +-------- Lamellidens_savadiensis **    
              +-----+                                    
              |     |  +----- Lamellidens_ferrugineus **    
              |     +--+                                 
              |        |  +-- Lamellidens_generosus **      
  +-----------+        +--+                              
  |           |           +-- Lamellidens_mainwaringi **    
  |           |                                          
  |           |  +----------- Trapezidens_exolescens **     
  |           +--+                                       
  |              |  +-------- Trapezidens_scutum **         
  |              +--+                                    
  |                 |  +----- Trapezidens_yeti **           
  |                 +--+                                 
--+                    |  +-- Trapezidens_angustior **      
  |                    +--+                              
  |                       +-- Trapezidens_dolichorhynchus **
  |                                                      
  |  +----------------------- Parreysia_rakhinensis      
  |  |                                                   
  |  |                    +-- Leoparreysia_whitteni      
  |  |              +-----+                              
  |  |              |     +-- Leoparreysia_burmana       
  +--+  +-----------+                                    
     |  |           |  +----- Leoparreysia_olivacea      
     |  |           +--+                                 
     |  |              |  +-- Leoparreysia_canefrii      
     |  |              +--+                              
     +--+                 +-- Leoparreysia_tavoyensis    
        |                                                
        |                 +-- Indonaia_andersoniana      
        |  +--------------+                              
        |  |              +-- Indonaia_subclathrata      
        +--+                                             
           |  +-------------- Indochinella_pugio         
           |  |                                          
           +--+  +----------- Radiatula_crispisulcata    
              |  |                                       
              +--+        +-- Radiatula_chaudhurii       
                 |  +-----+                              
                 |  |     +-- Radiatula_myitkyinae       
                 +--+                                    
                    |  +----- Radiatula_mouhoti          
                    +--+                                 
                       |  +-- Radiatula_chindwinensis    
                       +--+                              
                          +-- Radiatula_myitthanensis    

Bolotov et al., 2017, fig. S1

iT tT bi riS oT roT roS iD
8 8 5 2 130 21 4 34.0
                          +-- Trapezidens_dolichorhynchus **
              +-----------+                              
              |           +-- Trapezidens_exolescens **     
  +-----------+                                          
  |           |  +----------- Lamellidens_corrianus **      
  |           |  |                                       
  |           +--+  +-------- Lamellidens_savadiensis **    
  |              |  |                                    
  |              +--+     +-- Lamellidens_jenkinsianus **   
  |                 |  +--+                              
  |                 |  |  +-- Lamellidens_mainwaringi **    
  |                 +--+                                 
--+                    |  +-- Lamellidens_ferrugineus **    
  |                    +--+                              
  |                       +-- Lamellidens_marginalis **     
  |                                                      
  |                       +-- Parreysia_nagpoorensis     
  |  +--------------------+                              
  |  |                    +-- Parreysia_plagiosoma       
  |  |                                                   
  |  |                    +-- Leoparreysia_tavoyensis    
  |  |  +-----------------+                              
  +--+  |                 +-- Leoparreysia_olivacea      
     |  |                                                
     |  |     +-------------- Nitia_teretiuscula         
     |  |  +--+                                          
     |  |  |  |  +----------- Coelatura_aegyptiaca[2]    
     +--+  |  +--+                                       
        |  |     |  +-------- Coelatura_aegyptiaca[1]    
        |  |     +--+                                    
        |  |        |  +----- Prisodontopsis_mweruensis  
        |  |        +--+                                 
        |  |           |  +-- Coelatura_gabonensis       
        +--+           +--+                              
           |              +-- Coelatura_kunenensis       
           |                                             
           |        +-------- Indonaia_shurtleffiana     
           |  +-----+                                    
           |  |     |  +----- Indonaia_andersoniana      
           |  |     +--+                                 
           |  |        |  +-- Indonaia_rugosa            
           |  |        +--+                              
           +--+           +-- Indonaia_bonneaudii        
              |                                          
              |           +-- Indochinella_pugio         
              |  +--------+                              
              |  |        +-- Scabies_crispata           
              |  |                                       
              +--+        +-- Radiatula_myitkyinae       
                 |  +-----+                              
                 |  |     +-- Radiatula_chaudhurii       
                 +--+                                    
                    |  +----- Scabies_mandarinus         
                    +--+                                 
                       |  +-- Scabies_humilis            
                       +--+                              
                          +-- Scabies_phaselus           

Bolotov et al., 2017, fig. S2

iT tT bi riS oT roT roS iD
8 8 4 2 130 21 4 35.0
                       +-- Parreysia_nagpoorensis     
           +-----------+                              
           |           +-- Parreysia_plagiosoma       
           |                                          
  +--------+           +-- Trapezidens_dolichorhynchus **
  |        |  +--------+                              
  |        |  |        +-- Trapezidens_exolescens **     
  |        |  |                                       
  |        +--+  +-------- Lamellidens_savadiensis **    
  |           |  |                                    
  |           |  +-------- Lamellidens_corrianus **      
  |           +--+                                    
  |              |     +-- Lamellidens_marginalis **     
  |              |  +--+                              
  |              |  |  +-- Lamellidens_ferrugineus **    
--+              +--+                                 
  |                 |  +-- Lamellidens_jenkinsianus **   
  |                 +--+                              
  |                    +-- Lamellidens_mainwaringi **    
  |                                                   
  |                    +-- Leoparreysia_olivacea      
  |  +-----------------+                              
  |  |                 +-- Leoparreysia_tavoyensis    
  |  |                                                
  |  |     +-------------- Nitia_teretiuscula         
  |  |  +--+                                          
  |  |  |  |  +----------- Coelatura_aegyptiaca[1]    
  +--+  |  +--+                                       
     |  |     |  +-------- Coelatura_aegyptiaca[2]    
     |  |     +--+                                    
     |  |        |  +----- Coelatura_gabonensis       
     |  |        +--+                                 
     |  |           |  +-- Coelatura_kunenensis       
     +--+           +--+                              
        |              +-- Prisodontopsis_mweruensis  
        |                                             
        |        +-------- Indonaia_shurtleffiana     
        |  +-----+                                    
        |  |     |  +----- Indonaia_andersoniana      
        |  |     +--+                                 
        |  |        |  +-- Indonaia_rugosa            
        |  |        +--+                              
        +--+           +-- Indonaia_bonneaudii        
           |                                          
           |           +-- Indochinella_pugio         
           |  +--------+                              
           |  |        +-- Scabies_crispata           
           |  |                                       
           +--+        +-- Radiatula_myitkyinae       
              |  +-----+                              
              |  |     +-- Radiatula_chaudhurii       
              +--+                                    
                 |  +----- Scabies_mandarinus         
                 +--+                                 
                    |  +-- Scabies_humilis            
                    +--+                              
                       +-- Scabies_phaselus           

Bolotov et al., 2020, fig. 1

iT tT bi riS oT roT roS iD
8 8 6 2 113 33 4 34.0
                                +-------- Trapezidens_exolescens **     
                             +--+                                    
                             |  |  +----- Trapezidens_scutum **         
                             |  +--+                                 
                             |     |  +-- Trapezidens_dolichorhynchus **
  +--------------------------+     +--+                              
  |                          |        +-- Trapezidens_angustior **      
  |                          |                                       
  |                          |  +-------- Lamellidens_savadiensis **    
  |                          +--+                                    
  |                             |  +----- Lamellidens_generosus **      
  |                             +--+                                 
  |                                |  +-- Lamellidens_ferrugineus **    
  |                                +--+                              
  |                                   +-- Lamellidens_mainwaringi **    
  |                                                                  
  |                          +----------- Parreysia_keralaensis      
  |                       +--+                                       
--+                       |  |  +-------- Parreysia_plagiosoma       
  |                       |  +--+                                    
  |                       |     |  +----- Parreysia_nagpoorensis[2]  
  |                       |     +--+                                 
  |                       |        |  +-- Parreysia_nagpoorensis[1]  
  |  +--------------------+        +--+                              
  |  |                    |           +-- Parreysia_rakhinensis      
  |  |                    |                                          
  |  |                    |           +-- Leoparreysia_burmana       
  |  |                    |     +-----+                              
  |  |                    |     |     +-- Leoparreysia_whitteni      
  |  |                    +-----+                                    
  |  |                          |  +----- Leoparreysia_tavoyensis    
  |  |                          +--+                                 
  +--+                             |  +-- Leoparreysia_olivacea      
     |                             +--+                              
     |                                +-- Leoparreysia_canefrii      
     |                                                               
     |                                +-- Coelatura_aegyptiaca       
     |  +-----------------------------+                              
     |  |                             +-- Nitia_teretiuscula         
     |  |                                                            
     |  |                    +----------- Indonaia_shurtleffiana     
     |  |  +-----------------+                                       
     +--+  |                 |  +-------- Indonaia_rugosa            
        |  |                 +--+                                    
        |  |                    |  +----- Indonaia_andersoniana      
        |  |                    +--+                                 
        |  |                       |  +-- Indonaia_subclathrata      
        +--+                       +--+                              
           |                          +-- Indonaia_caerulea          
           |                                                         
           |  +-------------------------- Indochinella_pugio         
           |  |                                                      
           |  |                 +-------- Radiatula_crispisulcata    
           |  |  +--------------+                                    
           +--+  |              |  +----- Radiatula_mouhoti          
              |  |              +--+                                 
              |  |                 |  +-- Radiatula_chaudhurii       
              |  |                 +--+                              
              +--+                    +-- Radiatula_myitkyinae       
                 |                                                   
                 |  +-------------------- Scabiellus_songkramensis   
                 |  |                                                
                 |  |                 +-- Unionetta_fabagina         
                 +--+  +--------------+                              
                    |  |              +-- Harmandia_somboriensis     
                    |  |                                             
                    +--+  +-------------- Scabies_nucleus            
                       |  |                                          
                       |  |           +-- Scabies_crispata           
                       +--+  +--------+                              
                          |  |        +-- Scabies_mandarinus         
                          |  |                                       
                          +--+        +-- Scabies_humilis            
                             |  +-----+                              
                             |  |     +-- Scabies_pilata             
                             +--+                                    
                                |  +----- Scabies_anceps             
                                +--+                                 
                                   |  +-- Scabies_phaselus           
                                   +--+                              
                                      +-- Scabies_scobinatus         

Bolotov et al., 2020, fig. S1

iT tT bi riS oT roT roS iD
8 8 6 2 107 32 4 34.0
                                      +-------- Trapezidens_exolescens **     
                                   +--+                                    
                                   |  |  +----- Trapezidens_scutum **         
                                   |  +--+                                 
                                   |     |  +-- Trapezidens_dolichorhynchus **
  +--------------------------------+     +--+                              
  |                                |        +-- Trapezidens_angustior **      
  |                                |                                       
  |                                |  +-------- Lamellidens_savadiensis **    
  |                                +--+                                    
  |                                   |  +----- Lamellidens_generosus **      
  |                                   +--+                                 
  |                                      |  +-- Lamellidens_ferrugineus **    
  |                                      +--+                              
  |                                         +-- Lamellidens_mainwaringi **    
  |                                                                        
  |                                +----------- Parreysia_keralaensis      
  |                             +--+                                       
--+                             |  |  +-------- Parreysia_nagpoorensis[2]  
  |                             |  +--+                                    
  |                             |     |  +----- Parreysia_nagpoorensis[1]  
  |                             |     +--+                                 
  |                             |        |  +-- Parreysia_rakhinensis      
  |  +--------------------------+        +--+                              
  |  |                          |           +-- Parreysia_plagiosoma       
  |  |                          |                                          
  |  |                          |           +-- Leoparreysia_burmana       
  |  |                          |     +-----+                              
  |  |                          |     |     +-- Leoparreysia_whitteni      
  |  |                          +-----+                                    
  |  |                                |  +----- Leoparreysia_olivacea      
  |  |                                +--+                                 
  +--+                                   |  +-- Leoparreysia_tavoyensis    
     |                                   +--+                              
     |                                      +-- Leoparreysia_canefrii      
     |                                                                     
     |                                      +-- Coelatura_aegyptiaca       
     |  +-----------------------------------+                              
     |  |                                   +-- Nitia_teretiuscula         
     |  |                                                                  
     |  |                             +-------- Indonaia_shurtleffiana     
     +--+  +--------------------------+                                    
        |  |                          |  +----- Indonaia_andersoniana      
        |  |                          +--+                                 
        |  |                             |  +-- Indonaia_caerulea          
        |  |                             +--+                              
        +--+                                +-- Indonaia_subclathrata      
           |                                                               
           |  +-------------------------------- Indochinella_pugio         
           |  |                                                            
           |  |                          +----- Radiatula_mouhoti          
           +--+  +-----------------------+                                 
              |  |                       |  +-- Radiatula_myitkyinae       
              |  |                       +--+                              
              +--+                          +-- Radiatula_chaudhurii       
                 |                                                         
                 |  +-------------------------- Radiatula_crispisulcata    
                 |  |                                                      
                 +--+  +----------------------- Scabiellus_songkramensis   
                    |  |                                                   
                    +--+  +-------------------- Harmandia_somboriensis     
                       |  |                                                
                       +--+  +----------------- Unionetta_fabagina         
                          |  |                                             
                          +--+  +-------------- Scabies_mandarinus         
                             |  |                                          
                             +--+  +----------- Scabies_crispata           
                                |  |                                       
                                +--+  +-------- Scabies_nucleus            
                                   |  |                                    
                                   +--+     +-- Scabies_humilis            
                                      |  +--+                              
                                      |  |  +-- Scabies_pilata             
                                      +--+                                 
                                         |  +-- Scabies_phaselus           
                                         |  |                              
                                         +--+-- Scabies_scobinatus         
                                            |                              
                                            +-- Scabies_anceps             

Bolotov et al., 2018, fig. 2

iT tT bi riS oT roT roS iD
6 6 3 2 79 18 4 34.0
                          +-- Trapezidens_exolescens **     
                 +--------+                              
                 |        +-- Trapezidens_dolichorhynchus **
  +--------------+                                       
  |              |  +-------- Lamellidens_savadiensis **    
  |              +--+                                    
  |                 |  +----- Lamellidens_ferrugineus **    
  |                 +--+                                 
  |                    |  +-- Lamellidens_generosus **      
  |                    +--+                              
--+                       +-- Lamellidens_mainwaringi **    
  |                                                      
  |                       +-- Parreysia_keralaensis      
  |  +--------------------+                              
  |  |                    +-- Parreysia_nagpoorensis     
  |  |                                                   
  |  |                 +----- Leoparreysia_olivacea      
  +--+  +--------------+                                 
     |  |              |  +-- Leoparreysia_canefrii      
     |  |              +--+                              
     |  |                 +-- Leoparreysia_tavoyensis    
     |  |                                                
     +--+                 +-- Coelatura_aegyptiaca       
        |  +--------------+                              
        |  |              +-- Nitia_teretiuscula         
        |  |                                             
        |  |        +-------- Indonaia_shurtleffiana     
        +--+  +-----+                                    
           |  |     |  +----- Indonaia_rugosa            
           |  |     +--+                                 
           |  |        |  +-- Indonaia_caerulea          
           |  |        +--+                              
           +--+           +-- Indonaia_andersoniana      
              |                                          
              |  +----------- Indochinella_pugio         
              |  |                                       
              |  |     +----- Radiatula_myitkyinae       
              +--+  +--+                                 
                 |  |  |  +-- Radiatula_mouhoti          
                 |  |  +--+                              
                 +--+     +-- Radiatula_chaudhurii       
                    |                                    
                    |  +----- Scabies_mandarinus         
                    +--+                                 
                       |  +-- Scabies_humilis            
                       +--+                              
                          +-- Scabies_phaselus           

Konopleva et al., 2017, fig. 5

iT tT bi riS oT roT roS iD
5 5 2 2 0 0 0 35.0
        +-- Trapezidens_exolescens **     
  +-----+                              
  |     +-- Trapezidens_dolichorhynchus **
--+                                    
  |  +----- Lamellidens_savadiensis **    
  +--+                                 
     |  +-- Lamellidens_mainwaringi **    
     +--+                              
        +-- Lamellidens_jenkinsianus **   

Konopleva et al., 2017, fig. 6

iT tT bi riS oT roT roS iD
5 5 2 2 26 6 3 35.0
              +-- Trapezidens_exolescens **     
        +-----+                              
        |     +-- Trapezidens_dolichorhynchus **
  +-----+                                    
  |     |  +----- Lamellidens_savadiensis **    
  |     +--+                                 
  |        |  +-- Lamellidens_jenkinsianus **   
  |        +--+                              
--+           +-- Lamellidens_mainwaringi **    
  |                                          
  |           +-- Leoparreysia_tavoyensis    
  |  +--------+                              
  |  |        +-- Leoparreysia_olivacea      
  +--+                                       
     |  +-------- Coelatura_aegyptiaca       
     +--+                                    
        |  +----- Indochinella_pugio         
        +--+                                 
           |  +-- Radiatula_chaudhurii       
           +--+                              
              +-- Scabies_mandarinus         

Whelan et al., 2011, fig. 1 ML

iT tT bi riS oT roT roS iD
2 2 0 2 34 8 3 --
                    +-- Lamellidens_savadiensis **      
  +-----------------+                                
  |                 +-- Trapezidens_dolichorhynchus **  
--+                                                  
  |  +----------------- Leoparreysia_tavoyensis      
  |  |                                               
  +--+              +-- Radiatula_chaudhurii         
     |  +-----------+                                
     |  |           +-- Indochinella_pugio           
     +--+                                            
        |  +----------- Nitia_teretiuscula           
        +--+                                         
           |  +-------- Coelatura_aegyptiaca[2]      
           +--+                                      
              |  +----- Coelatura_aegyptiaca[1]      
              +--+                                   
                 |  +-- Coelatura_gabonensis         
                 +--+                                
                    +-- Prisodontopsis_aviculaeformis

Whelan et al., 2011, fig. 1 MP

iT tT bi riS oT roT roS iD
2 2 0 2 34 8 3 --
                 +-- Lamellidens_savadiensis **      
  +--------------+                                
  |              +-- Trapezidens_dolichorhynchus **  
--+                                               
  |  +-------------- Leoparreysia_tavoyensis      
  |  |                                            
  +--+           +-- Radiatula_chaudhurii         
     |  +--------+                                
     |  |        +-- Indochinella_pugio           
     +--+                                         
        |  +-------- Nitia_teretiuscula           
        |  |                                      
        +--+  +----- Coelatura_aegyptiaca[1]      
           |  |                                   
           +--+----- Coelatura_aegyptiaca[2]      
              |                                   
              |  +-- Coelatura_gabonensis         
              +--+                                
                 +-- Prisodontopsis_aviculaeformis

Pfeiffer & Graf, 2015, fig. 2

iT tT bi riS oT roT roS iD
2 2 0 2 38 6 3 --
              +-- Trapezidens_dolichorhynchus **
  +-----------+                              
  |           +-- Lamellidens_savadiensis **    
  |                                          
--+           +-- Leoparreysia_olivacea      
  |  +--------+                              
  |  |        +-- Leoparreysia_tavoyensis    
  +--+                                       
     |  +-------- Coelatura_aegyptiaca       
     +--+                                    
        |  +----- Indochinella_pugio         
        +--+                                 
           |  +-- Radiatula_chaudhurii       
           +--+                              
              +-- Scabies_mandarinus         

Pfeiffer et al., 2018, fig. 2

iT tT bi riS oT roT roS iD
2 2 0 2 27 26 4 --
                                            +-- Trapezidens_dolichorhynchus **  
  +-----------------------------------------+                                
  |                                         +-- Lamellidens_savadiensis **      
  |                                                                          
  |                                      +----- Parreysia_keralaensis        
--+  +-----------------------------------+                                   
  |  |                                   |  +-- Parreysia_nagpoorensis       
  |  |                                   +--+                                
  |  |                                      +-- Parreysia_plagiosoma         
  |  |                                                                       
  +--+                                      +-- Leoparreysia_tavoyensis      
     |  +-----------------------------------+                                
     |  |                                   +-- Leoparreysia_olivacea        
     |  |                                                                    
     |  |                                +----- Nitia_teretiuscula           
     +--+  +-----------------------------+                                   
        |  |                             |  +-- Coelatura_aegyptiaca         
        |  |                             +--+                                
        |  |                                +-- Prisodontopsis_aviculaeformis
        +--+                                                                 
           |                             +----- Indonaia_bonneaudii          
           |  +--------------------------+                                   
           |  |                          |  +-- Indonaia_andersoniana        
           |  |                          +--+                                
           +--+                             +-- Indonaia_caerulea            
              |                                                              
              |  +----------------------------- Indochinella_pugio           
              |  |                                                           
              +--+                          +-- Radiatula_chaudhurii         
                 |  +-----------------------+                                
                 |  |                       +-- Radiatula_myitkyinae         
                 +--+                                                        
                    |  +----------------------- Scabiellus_songkramensis     
                    |  |                                                     
                    +--+  +-------------------- Unionetta_fabagina[2]        
                       |  |                                                  
                       +--+  +----------------- Harmandia_somboriensis       
                          |  |                                               
                          +--+  +-------------- Unionetta_fabagina[1]        
                             |  |                                            
                             +--+           +-- Scabies_crispata             
                                |  +--------+                                
                                |  |        +-- Scabies_mandarinus           
                                +--+                                         
                                   |  +-------- Scabies_nucleus              
                                   |  |                                      
                                   +--+     +-- Scabies_pilata               
                                      |  +--+                                
                                      |  |  +-- Scabies_humilis              
                                      +--+                                   
                                         |  +-- Scabies_phaselus             
                                         |  |                                
                                         +--+-- Scabies_anceps               
                                            |                                
                                            +-- Scabies_scobinatus           

Jeratthitikul et al., 2019, fig. 1

iT tT bi riS oT roT roS iD
2 2 0 2 21 21 4 --
                                                  +-- Lamellidens_mainwaringi ** 
  +-----------------------------------------------+                           
  |                                               +-- Trapezidens_exolescens **  
--+                                                                           
  |  +----------------------------------------------- Leoparreysia_canefrii   
  |  |                                                                        
  +--+  +-------------------------------------------- Parreysia_keralaensis   
     |  |                                                                     
     +--+  +----------------------------------------- Coelatura_aegyptiaca    
        |  |                                                                  
        |  |                                   +----- Indonaia_rugosa         
        +--+  +--------------------------------+                              
           |  |                                |  +-- Indonaia_caerulea       
           |  |                                +--+                           
           |  |                                   +-- Indonaia_andersoniana   
           +--+                                                               
              |  +----------------------------------- Indochinella_pugio      
              |  |                                                            
              |  |                             +----- Radiatula_mouhoti       
              +--+  +--------------------------+                              
                 |  |                          |  +-- Radiatula_myitkyinae    
                 |  |                          +--+                           
                 +--+                             +-- Radiatula_chaudhurii    
                    |                                                         
                    |  +----------------------------- Scabiellus_songkramensis
                    +--+                                                      
                       |  +-------------------------- Harmandia_somboriensis  
                       +--+                                                   
                          |  +----------------------- Unionetta_fabagina      
                          +--+                                                
                             |  +-------------------- Scabies_crispata        
                             +--+                                             
                                |  +----------------- Scabies_mandarinus      
                                +--+                                          
                                   |  +-------------- Scabies_pilata          
                                   +--+                                       
                                      |  +----------- Scabies_humilis[2]      
                                      +--+                                    
                                         |  +-------- Scabies_humilis[1]      
                                         +--+                                 
                                            |  +----- Scabies_scobinatus      
                                            +--+                              
                                               |  +-- Scabies_phaselus        
                                               +--+                           
                                                  +-- Scabies_anceps          

Pfeiffer et al., 2019, fig. 4-5

iT tT bi riS oT roT roS iD
2 2 0 2 55 8 3 --
                 +-- Trapezidens_dolichorhynchus **  
  +--------------+                                
  |              +-- Lamellidens_savadiensis **      
  |                                               
--+              +-- Leoparreysia_tavoyensis      
  |  +-----------+                                
  |  |           +-- Leoparreysia_olivacea        
  |  |                                            
  +--+           +-- Coelatura_choziensis         
     |  +--------+                                
     |  |        +-- Prisodontopsis_aviculaeformis
     +--+                                         
        |  +-------- Indonaia_caerulea            
        +--+                                      
           |  +----- Indochinella_pugio           
           +--+                                   
              |  +-- Scabies_phaselus             
              +--+                                
                 +-- Harmandia_somboriensis       

Pfeiffer et al., 2019, fig. 6

iT tT bi riS oT roT roS iD
2 2 0 2 55 8 3 --
                 +-- Trapezidens_dolichorhynchus **  
  +--------------+                                
  |              +-- Lamellidens_savadiensis **      
  |                                               
--+              +-- Leoparreysia_tavoyensis      
  |  +-----------+                                
  |  |           +-- Leoparreysia_olivacea        
  |  |                                            
  +--+           +-- Coelatura_choziensis         
     |  +--------+                                
     |  |        +-- Prisodontopsis_aviculaeformis
     +--+                                         
        |  +-------- Indonaia_caerulea            
        +--+                                      
           |  +----- Indochinella_pugio           
           +--+                                   
              |  +-- Scabies_phaselus             
              +--+                                
                 +-- Harmandia_somboriensis       

Sonowal et al., 2021, fig. 4

iT tT bi riS oT roT roS iD
2 10 0 1 14 0 0 44.0
  +-------------------------- Lamellidens_mainwaringi **    
--+                                                      
  |  +----------------------- Lamprotula_leaii           
  +--+                                                   
     |  +-------------------- Potomida_littoralis        
     +--+                                                
        |  +----------------- Lampsilis_cardium          
        +--+                                             
           |  +-------------- Elliptio_complanata        
           +--+                                          
              |  +----------- Aculamprotula_tientsinensis
              +--+                                       
                 |  +-------- Unio_pictorum              
                 +--+                                    
                    |  +----- Anodonta_cygnea            
                    +--+                                 
                       |  +-- Unio_ravoisieri            
                       +--+                              
                          +-- Lamellidens_corrianus **      

Konopleva et al., 2023, fig. S1, 4

iT tT bi riS oT roT roS iD
2 2 0 2 54 5 3 --
              +-- Trapezidens_exolescens ** 
  +-----------+                          
  |           +-- Lamellidens_savadiensis **
--+                                      
  |  +----------- Parreysia_rakhinensis  
  +--+                                   
     |  +-------- Leoparreysia_tavoyensis
     +--+                                
        |  +----- Indonaia_andersoniana  
        +--+                             
           |  +-- Indochinella_pugio     
           +--+                          
              +-- Radiatula_mouhoti      

Pfeiffer & Graf, 2013, fig. 2

iT tT bi riS oT roT roS iD
1 1 0 1 34 1 1 --
  +-- Lamellidens_savadiensis **
--+                          
  +-- Coelatura_aegyptiaca   

Jadhav & Jamkhedkar, 2009, fig. 11

iT tT bi riS oT roT roS iD
1 1 0 1 25 0 0 --
-- Lamellidens_corrianus

Lopes-Lima et al., 2017, fig. 1-2

iT tT bi riS oT roT roS iD
1 1 0 1 80 3 3 --
  +-------- Trapezidens_dolichorhynchus **
--+                                    
  |  +----- Leoparreysia_tavoyensis    
  +--+                                 
     |  +-- Coelatura_aegyptiaca       
     +--+                              
        +-- Indochinella_pugio         

Araujo et al., 2017, fig. 2

iT tT bi riS oT roT roS iD
1 1 0 1 13 1 1 --
  +-- Lamellidens_savadiensis **
--+                          
  +-- Coelatura_aegyptiaca   

Choi et al., 2020, fig. 5

iT tT bi riS oT roT roS iD
1 1 0 1 8 2 2 --
  +----- Radiatula_mouhoti    
--+                           
  |  +-- Lamellidens_generosus **
  +--+                        
     +-- Leoparreysia_canefrii

Sonowal et al., 2021, fig. 3a

iT tT bi riS oT roT roS iD
1 1 0 1 14 0 0 --
-- Lamellidens_corrianus

Sonowal et al., 2021, fig. 3b

iT tT bi riS oT roT roS iD
1 1 0 1 14 0 0 --
-- Lamellidens_mainwaringi

Pavan-Kumar et al., 2022, fig. 4

iT tT bi riS oT roT roS iD
1 1 0 1 35 0 0 --
-- Lamellidens_marginalis

Pavan-Kumar et al., 2022, fig. S5

iT tT bi riS oT roT roS iD
1 1 0 1 38 0 0 --
-- Lamellidens_marginalis

Page updated: 08:55:56 Wed 18 Jan 2023

 
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