|
Bivalvia | Unionoida | Unionoidea | Unionidae subfamily PARREYSIINAE Henderson, 1935: cladomics
5 tribes
Cladomics Summary
number of species = 110
number of subtaxa (rank: tribe) = 5
analyzed in 76 cladograms in 47 publications
maximum number of terminals in an analysis = 57, mean = 12.75
maximum number of subtaxa (tribe) in an analysis = 5, mean = 2.47
Basic Monophyly
number of cladograms that test basic monophyly (i.e., more than 1 included species) = 46
maximum number of included terminals = 57, mean = 20.41
maximum number of subtaxa (tribe) = 5, mean = 3.43
proportion of cladograms that support basic monophyly = 0.96
Rigorous Monophyly
number of cladograms that test rigorous monophyly = 32
(Rigorous monophyly refers to cladograms with terminal species representing more than one subtaxon (tribe) taxon and at least one rigorous outgroup (subfamily).
maximum number of included terminals = 57, mean = 23.06
maximum number of subtaxa (tribe) = 5, mean = 4.03
proportion of cladograms that support rigorous monophyly = 0.97
Pairwise Cladogram Clustering
The graphs below depict in two dimensions all pair-wise ingroup distances within the cladome. See the Cladmoics FARQ for more information.
Bolotov et al., 2017 [Publ 3761], fig. S1
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 29 |
29 |
27 |
5 |
112 |
87 |
3 |
0.0 |
*** Top Monophyly Cladogram ***
+-- Trapezidens_dolichorhynchus **
+-----------+
| +-- Trapezidens_exolescens **
+-----------+
| | +----------- Lamellidens_corrianus **
| | |
| +--+ +-------- Lamellidens_savadiensis **
| | |
| +--+ +-- Lamellidens_jenkinsianus **
| | +--+
| | | +-- Lamellidens_mainwaringi **
| +--+
+--------+ | +-- Lamellidens_ferrugineus **
| | +--+
| | +-- Lamellidens_marginalis **
| |
| | +-- Parreysia_nagpoorensis **
| | +--------------------+
| | | +-- Parreysia_plagiosoma **
| | |
| | | +-- Leoparreysia_tavoyensis **
| | | +-----------------+
| +--+ | +-- Leoparreysia_olivacea **
| | |
| | | +-------------- Nitia_teretiuscula **
| | | +--+
| | | | | +----------- Coelatura_aegyptiaca[2] **
| +--+ | +--+
| | | | +-------- Coelatura_aegyptiaca[1] **
| | | +--+
| | | | +----- Prisodontopsis_mweruensis **
| | | +--+
| | | | +-- Coelatura_gabonensis **
| +--+ +--+
| | +-- Coelatura_kunenensis **
| |
| | +-------- Indonaia_shurtleffiana **
| | +-----+
| | | | +----- Indonaia_andersoniana **
| | | +--+
| | | | +-- Indonaia_rugosa **
| | | +--+
| +--+ +-- Indonaia_bonneaudii **
| |
| | +-- Indochinella_pugio **
| | +--------+
| | | +-- Scabies_crispata **
| | |
| +--+ +-- Radiatula_myitkyinae **
| | +-----+
| | | +-- Radiatula_chaudhurii **
| +--+
| | +----- Scabies_mandarinus **
| +--+
| | +-- Scabies_humilis **
| +--+
| +-- Scabies_phaselus **
|
| +-- Ensidens_telus
| +-----+
--+ | +-- Ensidens_ingallsianus
| +-----+
| | | +----- Ensidens_sagittarius
| | +--+
| | | +-- Ensidens_spiculus
| | +--+
| +--+ +-- Ensidens_jaculus
| | |
| | | +-- Hyriopsis_bialata
| | | +--------+
| | | | +-- Hyriopsis_khoratensis
| | +--+
| | | +-------- Hyriopsis_panhai
| | +--+
| | | +----- Hyriopsis_myersiana
| | +--+
| +--------------+ | +-- Rectidens_sumatrensis
| | | +--+
| | | +-- Hyriopsis_desowitzi
| | |
| | | +----- Physunio
| | | +--------+
| | | | | +-- Physunio_modelli
| | | | +--+
| | | | +-- Physunio_superbus
| | | |
| | +--+ +-------- Yaukthwa_inlenensis
| | | +--+
| | | | | +----- Yaukthwa_panhai
| | | | +--+
| | | | | +-- Yaukthwa_rectangularis
| | | | +--+
| | +--+ +-- Yaukthwa_nesemanni
| | |
| | | +-- Lens_contradens
| | | +-----+
| | | | +-- Lens_micropterus
| | +--+
| | | +----- Lens_eximius
| | +--+
| | | +-- Lens_comptus
| | +--+
| | +-- Lens_novoselovi
| |
| | +-------------------- Indopseudodon_indawgyiensis
| | |
| | | +-------- Pilsbryoconcha_exilis[1]
| | | +--------+
+--+ +--------+ | | +----- Pilsbryoconcha_exilis[3]
| | | | +--+
| | | | | +-- Pilsbryoconcha_exilis[2]
| | | | +--+
| | | | +-- Pilsbryoconcha_carinifera
| | +--+
| | | +-------- Sundadontina_tumida
| | | +-----+
| | | | | +----- Pseudodon_vagulus
| | | | +--+
| | | | | +-- Pseudodon_mekongi
| | | | +--+
| | +--+ +-- Pseudodon_vondembuschianus
| | |
| | | +-- Nyeinchanconcha_nyeinchani
| | | +--------+
| | | | +-- Sundadontina_cumingii
| | +--+
| | | +-------- Thaiconcha_callifera
| | +--+
| | | +----- Bineurus_exilis
| | +--+
| | | +-- Bineurus_mouhotii
| | +--+
| | +-- Bineurus_loeiensis
| |
| | +-------- Amblema_plicata
| | +--+
| | | | +----- Ortmanniana_ligamentina
| | | +--+
| | | | +-- Lampsilis_cardium
| | | +--+
+--+ +-----+ +-- Cambarunio_iris
| | |
| | | +-- Quadrula_quadrula
| | | +-----+
| | | | +-- Quadrula_verrucosa
| | +--+
| | | +----- Pleurobema_sintoxia
| | +--+
| | | +-- Eurynia_dilatata
| | +--+
| | +-- Elliptio_complanata
| +--------+
| | | +-- Sinohyriopsis_cumingii
| | | +-----+
| | | | +-- Chamberlainia_somsakpanhai
| | | |
| | | +-----+ +-- Potomida_littoralis
| | | | | +--+
| | | | | | +-- Pronodularia_japanensis
| | | | +--+
| | | | | +-- Lamprotula_caveata
| | | | +--+
| | +--+ +-- Lamprotula_leaii
| | |
| | | +----- Sinosolenaia_carinata
| | | +-----+
| | | | | +-- Sinosolenaia_oleivora
| | | | +--+
| | +--+ +-- Sinosolenaia_recognita
| | |
| | | +-------- Gonidea_angulata
| | +--+
+--+ | +----- Microcondylaea_bonellii
| +--+
| | +-- Leguminaia_wheatleyi
| +--+
| +-- Leguminaia
|
| +----- Unio_tumidus
| +--------+
| | | +-- Unio_pictorum
| | +--+
| | +-- Unio_crassus
| +--------+
| | | +----------- Aculamprotula_tortuosa
| | | |
| | | | +-- Nodularia_douglasiae
| | +--+ +-----+
| | | | +-- Nodularia_nuxpersicae
| | | |
| | +--+ +-- Cuneopsis_rufescens
| | | +--+
| | | | +-- Cuneopsis_heudei
| | +--+
+--+ | +-- Tchangsinaia_piscicula
| +--+
| +-- Schistodesmus_lampreyanus
|
| +-- Lanceolaria_grayii
| +-----------------+
| | +-- Lanceolaria_gladiola
| |
| | +-- Cristaria_plicata
| | +--------+
| | | +-- Cristaria_radiata
+--+ +-----+
| | | +-------- Pletholophus_tenuis
| | +--+
| | | +----- Sinanodonta_lucida
| | +--+
| | | +-- Sinanodonta_jourdyi
+--+ +--+
| +-- Sinanodonta_lauta
|
| +-------------- Anodonta_anatina
| |
| | +----- Pseudanodonta_complanata
+--+ +-----+
| | | +-- Anodonta_cygnea
| | +--+
| | +-- Anodonta_nuttalliana
+--+
| +-- Lasmigona_costata
| +-----+
| | +-- Alasmidonta_marginata
+--+
| +----- Simpsonaias_ambigua
+--+
| +-- Pyganodon_grandis
+--+
+-- Strophitus_undulatus
Huang et al., 2019 [Publ 3843], fig. 2
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 36 |
36 |
30 |
5 |
168 |
129 |
3 |
17.0 |
*** Top Sister-Group Cladogram ***
+----- Trapezidens_exolescens **
+-----------------+
| | +-- Trapezidens_dolichorhynchus **
| +--+
+--+ +-- Trapezidens_angustior **
| |
| | +-------------------- Lamellidens_corrianus **
| | |
| +--+ +----------------- Lamellidens_savadiensis **
| | |
| +--+ +-- Lamellidens_lamellatus **
| | +-----------+
| | | +-- Lamellidens_jenkinsianus **
| +--+
| | +----------- Lamellidens_generosus **
| +--+
| | +-------- Lamellidens_mainwaringi[2] **
| +--+
+--------------+ | +----- Lamellidens_mainwaringi[1] **
| | +--+
| | | +-- Lamellidens_ferrugineus **
| | +--+
| | +-- Lamellidens_marginalis **
| |
| | +----- Parreysia_keralaensis **
| | +-----------------+
| | | | +-- Parreysia_nagpoorensis **
| | | +--+
| | | +-- Parreysia_plagiosoma **
| | |
| | | +-- Leoparreysia_tavoyensis **
| | | |
| +--+ +-----------------+-- Leoparreysia_canefrii **
| | | |
| | | +-- Leoparreysia_olivacea **
| | |
| | | +-------- Coelatura_aegyptiaca[2] **
| | | |
| | | +--------+ +----- Coelatura_aegyptiaca[1] **
| +--+ | | |
| | | | +----- Nitia_teretiuscula **
| | | +--+
| | | | +-- Coelatura_gabonensis **
| | | | |
| | | +--+-- Coelatura_kunenensis **
| | | |
| +--+ +-- Prisodontopsis_mweruensis **
| |
| | +-------- Indonaia_shurtleffiana **
| | |
| | +-----+ +----- Indonaia_bonneaudii **
| | | | |
| | | +--+----- Indonaia_rugosa **
| | | |
| | | | +-- Indonaia_andersoniana **
| +--+ +--+
| | +-- Indonaia_caerulea **
| |
| | +----------- Indochinella_pugio **
| | |
| | | +----- Scabies_humilis **
| +--+ +--+
| | | | +-- Scabies_phaselus **
| | | +--+
| +--+ +-- Scabies_mandarinus **
| |
| | +----- Radiatula_myitkyinae **
| +--+
| | +-- Radiatula_chaudhurii **
| +--+
| +-- Radiatula_mouhoti **
|
| +-------------------------------- Acuticosta_chinensis
| |
| | +----------------------------- Lepidodesma_languilati
| | |
| | | +-------- Aculamprotula_tortuosa
| | | |
| | | +-----------+ +----- Aculamprotula_coreana
| | | | | |
| | | | +--+----- Aculamprotula_scripta
| | | | |
--+ | | | | +-- Aculamprotula_fibrosa
| | | | +--+
| | | | +-- Aculamprotula_tientsinensis
| +-----+ +--------+
| | | | | +-------- Unio_tumidus
| | | | | +--------+
| | | | | | | +----- Unio_crassus
| | | | | | +--+
| | | | | | | +-- Unio_delphinus
| | | | | | +--+
| | | | +--+ +-- Unio_pictorum
| | | | |
| | | | | +-- Schistodesmus_lampreyanus
| | | | | +-----------+
| | | | | | +-- Schistodesmus_spinosus
| | | | | |
| | | | +--+ +----- Nodularia_douglasiae[1]
| | +--+ | +-----+
| | | | | | +-- Nodularia_douglasiae[2]
| | | | | +--+
| | | +--+ +-- Nodularia_nuxpersicae
| | | |
| | | | +-------- Tchangsinaia_piscicula
| | | +--+
| | | | +----- Cuneopsis_rufescens
| | | +--+
| | | | +-- Cuneopsis_heudei
| | | +--+
| | | +-- Cuneopsis_celtiformis
| | |
| | | +-------- Lanceolaria_gladiola
| | | |
| | | +-----------------+ +----- Lanceolaria_grayii[1]
| | | | | |
| | | | +--+----- Lanceolaria_grayii[2]
| | | | |
| | | | | +-- Lanceolaria_lanceolata
| | | | +--+
| | | | +-- Lanceolaria_triformis
| | | |
| | | | +-------------------- Anemina_arcaeformis
| | | | |
| | | | +--+ +----------------- Cristaria_plicata
| | +--+ | | |
| | | | +--+ +-------------- Pletholophus_tenuis
| | | | | |
| | | | +--+ +-- Sinanodonta_angula
| | | | | +--------+
| | | | | | +-- Sinanodonta_lucida
| | | | +--+
| | | | | +-------- Sinanodonta_woodiana[2]
| | | | +--+
| | | | | +----- Sinanodonta_jourdyi
| | | | +--+
| | +--+ | +-- Sinanodonta_elliptica
| | | +--+
| | | +-- Sinanodonta_woodiana[1]
| | |
| | | +-------- Alasmidonta_marginata
| | | |
| | | +-------- Lasmigona_costata
| | | |
| | | +-------- Pseudanodonta_complanata
| | | |
| | | +-------- Simpsonaias_ambigua
+--+ | |
| +--------------+-------- Strophitus_undulatus
| |
| | +----- Anodonta_anatina
| +--+
| | | +-- Anodonta_cygnea
| | +--+
| | +-- Anodonta_nuttalliana
| |
| | +----- Pyganodon_grandis
| +--+
| | +-- Utterbackia_imbecillis
| +--+
| +-- Utterbackia_peninsularis
|
| +----------- Hyriopsis_desowitzi
| |
| +----------- Rectidens_sumatrensis
| |
| | +-- Hyriopsis_bialata
| | |
| | +-- Hyriopsis_myersiana
| +-----+--------+
| | | +-- Hyriopsis_panhai
| | | |
| | | +-- Hyriopsis_khoratensis
| | |
| | | +-- Ensidens_ingallsianus
| | | +-----+
| | | | +-- Ensidens_telus
| | +--+
| | | +----- Ensidens_sagittarius
| +-----------------+ +--+
| | | | +-- Ensidens_spiculus
| | | +--+
| | | +-- Ensidens_jaculus
| | |
| | | +----- Physunio
| | | +--------+
| | | | | +-- Physunio_modelli
| | | | +--+
| | | | +-- Physunio_superbus
| | | |
| | +--+ +-------- Yaukthwa_inlenensis
| | | +--+
| | | | | +----- Yaukthwa_panhai
| | | | +--+
| | | | | +-- Yaukthwa_nesemanni
| | | | +--+
| | +--+ +-- Yaukthwa_rectangularis
| | |
| | | +-- Lens_contradens
| | | +-----+
| | | | +-- Lens_micropterus
| | +--+
| | | +----- Lens_eximius
| | +--+
| | | +-- Lens_comptus
| | +--+
| | +-- Lens_novoselovi
| |
| | +-- Megalonaias_nervosa
| | +--+
+--+ | +-- Uniomerus_carolinianus
| +-----------------------+
| | | +-- Quadrula_quadrula
| | +--+
| | +-- Quadrula_verrucosa
| |
| | +-------- Elliptoideus_sloatianus
| +--+ |
| | | +-----------------+ +-- Elliptio_complanata
| | | | | +--+
| | | | | | +-- Eurynia_dilatata
| | | | +--+
| | | | | +-- Fusconaia_flava
| | +--+ +--+
| | | +-- Pleurobema_sintoxia
| | |
| | | +----------------------- Amblema_plicata
| | | |
| | | | +----- Toxolasma_parvum
| | +--+ +--------------+
| | | | | +-- Obliquaria_reflexa
| | | | +--+
| | | | +-- Plectomerus_dombeyanus
| | +--+
| | | +----------------- Potamilus_leptodon
| | | |
| | | | +-------------- Potamilus_alatus
| | +--+ |
| | | | +----------- Venustaconcha_ellipsiformis
| | | | |
| | +--+ +----------- Leaunio_lienosus
| | | |
+--+ | | +-- Ellipsaria_lineolata
| | +--------+
| +--+ +-- Truncilla_truncata
| |
| | +-- Medionidus_conradicus
| +--------+
| | +-- Ptychobranchus_subtentus
| |
| | +-------- Cambarunio_iris
| +--+
| | +----- Ortmanniana_ligamentina
| +--+
| | +-- Lampsilis_cardium
| +--+
| +-- Lampsilis_ornata
|
| +----- Chamberlainia_somsakpanhai
| +-----------------------+
| | | +-- Sinohyriopsis_cumingii
| | +--+
| | +-- Sinohyriopsis_schlegelii
| |
| | +-- Potomida_littoralis
| | +-----+
| | | +-- Pronodularia_japanensis
| | +-----------------+
+--+ | | +----- Lamprotula_leaii
| | +--+
| | | +-- Lamprotula_caveata
| | +--+
| | +-- Lamprotula_cornuumlunae
| |
| | +-------- Sinosolenaia_carinata
| | +--+
| | | | +----- Sinosolenaia_recognita
| | | +--+
+--+ | | +-- Sinosolenaia_oleivora
| | +--+
| +-----------+ +-- Sinosolenaia_iridinea
| | |
| | | +----- Gonidea_angulata
| | | +--+
| | | | | +-- Leguminaia_wheatleyi
| | | | +--+
| | +--+ +-- Microcondylaea_bonellii
| | |
| | | +-- Ptychorhynchus_pfisteri
| | | |
+--+ +-----+-- Parvasolenaia_rivularis
| |
| +-- Koreosolenaia_sitgyensis
|
| +----- Indopseudodon_salwenianus
| +--------------+
| | | +-- Indopseudodon_indawgyiensis
| | +--+
| | +-- Indopseudodon_bogani
| |
| | +-------- Pilsbryoconcha_exilis[3]
+--+ +--------+
| | | +----- Pilsbryoconcha_exilis[2]
| | +--+
| | | +-- Pilsbryoconcha_exilis[1]
| | +--+
| | +-- Pilsbryoconcha_carinifera
| |
+--+ +----------- Nyeinchanconcha_nyeinchani
| +--+
| | | +-------- Thaiconcha_callifera
| | +--+
| | | +----- Bineurus_loeiensis
| | +--+
| | | +-- Bineurus_mouhotii
+--+ +--+
| +-- Bineurus_exilis
|
| +----- Pseudodon_vagulus
| +--+
| | | +-- Pseudodon_mekongi
| | +--+
+-----+ +-- Pseudodon_vondembuschianus
|
| +-- Sundadontina_cumingii
| |
+-----+-- Sundadontina_tumida[1]
|
+-- Sundadontina_tumida[2]
Bolotov et al., 2022 [Publ 4252], fig. 2
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 57 |
57 |
53 |
5 |
4 |
4 |
1 |
49.0 |
+-- Pseudodon_lenyanensis
|
+-- Nyeinchanconcha_nyeinchani
+--------------------------------------------+
| +-- Indopseudodon_kayinensis
| |
| +-- Sundadontina_tanintharyiensis
|
| +----------- Trapezidens_mogaungensis **
| |
| +-----+ +-------- Trapezidens_exolescens **
| | | |
| | +--+ +----- Trapezidens_scutum **
| | | |
| | +--+ +-- Trapezidens_angustior **
| | | |
| +--------------------------+ +--+-- Trapezidens_dolichorhynchus **
| | | |
--+ | | +-- Trapezidens_yeti **
| | |
| | | +-------------- Lamellidens_marginalis **
| | | |
| | | | +----------- Lamellidens_ferrugineus **
| | +--+ |
| | | | +-------- Lamellidens_generosus **
| | | +--+
| | | | | +----- Lamellidens_mainwaringi **
| | +--+ +--+
| | | | +-- Lamellidens_lamellatus **
| | | +--+
| | | +-- Lamellidens_jenkinsianus **
| | |
| | | +-------- Lamellidens_nongyangensis **
+--+ +--+
| | +----- Lamellidens_savadiensis **
| +--+
| | +-- Lamellidens_corrianus **
| +--+
| +-- Lamellidens_chindwinensis **
|
| +----------- Parreysia_keralaensis **
| +-----------------------------+
| | | +-------- Parreysia_corrugata **
| | +--+
| | | +----- Parreysia_rakhinensis **
| | +--+
| | | +-- Parreysia_nagpoorensis **
| | +--+
| | +-- Parreysia_plagiosoma **
| |
| | +-- Leoparreysia_burmana **
+--+ +-----+
| | +-- Leoparreysia_whitteni **
| +-----------------------------+
| | | +----- Leoparreysia_olivacea **
| | +--+
| | | +-- Leoparreysia_canefrii **
| | +--+
| | +-- Leoparreysia_tavoyensis **
| |
| | +-------- Nitia_teretiuscula **
+--+ +--------------------------+
| | | +----- Coelatura_aegyptiaca **
| | +--+
| | | +-- Nyassunio_nyassaensis **
| | +--+
| | +-- Coelatura_kunenensis **
| |
| | +----------------- Indonaia_cylindrica **
| | |
+--+ +--------------+ +-------------- Indonaia_gratiosa **
| | | |
| | +--+ +----------- Indonaia_andersoniana **
| | | |
| | +--+ +-- Indonaia_caerulea **
| | | +-----+
| | | | +-- Indonaia_subclathrata **
| | +--+
| | | +----- Indonaia_rugosa **
+--+ +--+
| | +-- Indonaia_bonneaudii **
| +--+
| +-- Indonaia_shurtleffiana **
|
| +----------------------------- Indochinella_pugio **
| |
| | +----------- Radiatula_crispisulcata **
| | |
| | | +----- Radiatula_chaudhurii **
+--+ +--------------+ +--+
| | | | | +-- Radiatula_myitkyinae **
| | | | +--+
| | +--+ +-- Radiatula_kachinensis **
| | |
| | | +----- Radiatula_chindwinensis **
+--+ +--+
| | +-- Radiatula_myitthanensis **
| +--+
| +-- Radiatula_mouhoti **
|
| +----------------------- Scabiellus_songkramensis **
| |
+--+ +-------------------- Harmandia_somboriensis **
| |
+--+ +----------------- Unionetta_fabagina **
| |
+--+ +-- Scabies_crispata **
| +-----------+
| | +-- Scabies_mandarinus **
+--+
| +----------- Scabies_nucleus **
| |
+--+ +-- Scabies_humilis **
| +-----+
| | +-- Scabies_pilata **
+--+
| +----- Scabies_phaselus **
+--+
| +-- Scabies_anceps **
+--+
+-- Scabies_scobinatus **
Bolotov et al., 2022 [Publ 4252], fig. S1
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 57 |
57 |
47 |
5 |
4 |
4 |
1 |
47.0 |
+-- Pseudodon_lenyanensis
|
+-- Nyeinchanconcha_nyeinchani
+-----------------------------------------------+
| +-- Indopseudodon_kayinensis
| |
| +-- Sundadontina_tanintharyiensis
|
| +-- Parreysia_keralaensis **
| |
| +-- Parreysia_corrugata **
| |
| +--------------------------------------------+-- Parreysia_plagiosoma **
| | |
| | +-- Parreysia_rakhinensis **
--+ | |
| | +-- Parreysia_nagpoorensis **
| |
| | +-------------- Trapezidens_mogaungensis **
| | +--+
| | | | +----------- Trapezidens_scutum **
| | | +--+
| | | | +-------- Trapezidens_exolescens **
| | | +--+
| | | | +----- Trapezidens_yeti **
| | | +--+
| | | | +-- Trapezidens_dolichorhynchus **
| | +--------------------------+ +--+
+--+ | | +-- Trapezidens_angustior **
| | |
| | | +-------- Lamellidens_savadiensis **
| | | +-----+
| | | | | +----- Lamellidens_chindwinensis **
| | | | +--+
| | | | | +-- Lamellidens_corrianus **
| | | | +--+
| | +--+ +-- Lamellidens_nongyangensis **
| | |
| | | +-- Lamellidens_marginalis **
| | | +--------+
| | | | +-- Lamellidens_ferrugineus **
| | +--+
| | | +-------- Lamellidens_generosus **
| | +--+
| | | +----- Lamellidens_mainwaringi **
+--+ +--+
| | +-- Lamellidens_jenkinsianus **
| +--+
| +-- Lamellidens_lamellatus **
|
| +-- Leoparreysia_burmana **
| |
| +-- Leoparreysia_canefrii **
| |
| +--------------------------------------+-- Leoparreysia_olivacea **
| | |
| | +-- Leoparreysia_tavoyensis **
| | |
| | +-- Leoparreysia_whitteni **
| |
| | +-- Coelatura_aegyptiaca **
| | |
| | +-- Coelatura_kunenensis **
+--+ +-----------------------------------+
| | +-- Nitia_teretiuscula **
| | |
| | +-- Nyassunio_nyassaensis **
| |
| | +-- Indonaia_cylindrica **
| | +--------+
| | | +-- Indonaia_gratiosa **
| | |
+--+ +-----------------------+ +----- Indonaia_shurtleffiana **
| | | +--+
| | | | | +-- Indonaia_bonneaudii **
| | | | +--+
| | +--+ +-- Indonaia_rugosa **
| | |
| | | +----- Indonaia_andersoniana **
| | +--+
| | | +-- Indonaia_caerulea **
+--+ +--+
| +-- Indonaia_subclathrata **
|
| +-------------------------------- Indochinella_pugio **
| |
| | +----- Radiatula_chaudhurii **
| | +-----+
| | | | +-- Radiatula_kachinensis **
| | | +--+
+--+ +-----------------+ +-- Radiatula_myitkyinae **
| | |
| | | +-------- Radiatula_crispisulcata **
| | +--+
| | | +----- Radiatula_mouhoti **
| | +--+
+--+ | +-- Radiatula_myitthanensis **
| +--+
| +-- Radiatula_chindwinensis **
|
| +-------------------------- Scabiellus_songkramensis **
| |
+--+ +----------------------- Harmandia_somboriensis **
| |
+--+ +-------------------- Unionetta_fabagina **
| |
+--+ +----------------- Scabies_crispata **
| |
+--+ +-- Scabies_humilis **
| +-----------+
| | +-- Scabies_mandarinus **
+--+
| +----------- Scabies_nucleus **
+--+
| +-------- Scabies_pilata **
+--+
| +----- Scabies_anceps **
+--+
| +-- Scabies_phaselus **
+--+
+-- Scabies_scobinatus **
Bolotov et al., 2022 [Publ 4239], fig. 2
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 54 |
54 |
52 |
5 |
4 |
4 |
1 |
39.0 |
+-- Pseudodon_lenyanensis
|
+-- Nyeinchanconcha_nyeinchani
+-----------------------------------------------+
| +-- Indopseudodon_kayinensis
| |
| +-- Sundadontina_tanintharyiensis
|
| +-- Trapezidens_exolescens **
| +-----+
| | +-- Trapezidens_scutum **
| +-----------+
| | | +----- Trapezidens_angustior **
| | +--+
| | | +-- Trapezidens_dolichorhynchus **
| +--------------------------+ +--+
| | | +-- Trapezidens_yeti **
| | |
--+ | | +----------------- Lamellidens_corrianus **
| | | |
| | +--+ +-------------- Lamellidens_nongyangensis **
| | | |
| | +--+ +----------- Lamellidens_savadiensis **
| | | |
| | | | +-- Lamellidens_ferrugineus **
| | +--+ +-----+
| | | | +-- Lamellidens_marginalis **
| | | |
| | +--+ +-- Lamellidens_lamellatus **
| | | +--+
| | | | +-- Lamellidens_jenkinsianus **
| | +--+
| | | +-- Lamellidens_generosus **
| | +--+
+--+ +-- Lamellidens_mainwaringi **
|
| +----------- Parreysia_keralaensis **
| +--+
| | | +-------- Parreysia_plagiosoma **
| | +--+
| | | +----- Parreysia_nagpoorensis **
| | +--+
| | | +-- Parreysia_corrugata **
| +-----------------------------+ +--+
| | | +-- Parreysia_rakhinensis **
| | |
| | | +-- Leoparreysia_burmana **
| | | +-----+
| | | | +-- Leoparreysia_whitteni **
| | +-----+
| | | +----- Leoparreysia_olivacea **
| | +--+
| | | +-- Leoparreysia_canefrii **
| | +--+
+--+ +-- Leoparreysia_tavoyensis **
|
| +-------- Nitia_teretiuscula **
| +--------------------------------+
| | | +----- Coelatura_aegyptiaca **
| | +--+
| | | +-- Coelatura_kunenensis **
| | +--+
| | +-- Nyassunio_nyassaensis **
| |
| | +-- Indonaia_cylindrica **
| | +--------+
+--+ | +-- Indonaia_gratiosa **
| |
| +--------------------------+ +----- Indonaia_andersoniana **
| | | +--+
| | | | | +-- Indonaia_caerulea **
| | | | +--+
| | +--+ +-- Indonaia_subclathrata **
| | |
| | | +----- Indonaia_shurtleffiana **
| | +--+
+--+ | +-- Indonaia_bonneaudii **
| +--+
| +-- Indonaia_rugosa **
|
| +----------------------------------- Indochinella_pugio **
| |
| | +----------- Radiatula_crispisulcata **
| | |
+--+ +--------------------+ +-- Radiatula_chaudhurii **
| | | +-----+
| | | | +-- Radiatula_myitkyinae **
| | +--+
| | | +----- Radiatula_mouhoti **
+--+ +--+
| | +-- Radiatula_chindwinensis **
| +--+
| +-- Radiatula_myitthanensis **
|
| +----------------------------- Scabiellus_songkramensis **
+--+
| +-------------------------- Harmandia_somboriensis **
+--+
| +----------------------- Unionetta_fabagina **
+--+
| +-------------------- Scabies_crispata **
+--+
| +----------------- Scabies_mandarinus **
+--+
| +-------------- Scabies_nucleus **
+--+
| +----------- Scabies_pilata **
+--+
| +-------- Scabies_humilis **
+--+
| +----- Scabies_anceps **
+--+
| +-- Scabies_phaselus **
+--+
+-- Scabies_scobinatus **
Bolotov et al., 2022 [Publ 4239], fig. S1
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 54 |
54 |
50 |
5 |
4 |
4 |
1 |
43.0 |
+-- Pseudodon_lenyanensis
|
+-- Nyeinchanconcha_nyeinchani
+--------------------------------------------+
| +-- Indopseudodon_kayinensis
| |
| +-- Sundadontina_tanintharyiensis
|
| +-------- Trapezidens_exolescens **
| |
| +--------+ +----- Trapezidens_scutum **
| | | |
| | +--+ +-- Trapezidens_angustior **
| | | |
| | +--+-- Trapezidens_dolichorhynchus **
| | |
| +--------------------------+ +-- Trapezidens_yeti **
--+ | |
| | | +----- Lamellidens_savadiensis **
| | | +--------+
| | | | | +-- Lamellidens_corrianus **
| | | | +--+
| | | | +-- Lamellidens_nongyangensis **
| | +--+
| | | +-- Lamellidens_ferrugineus **
| | | +--------+
| | | | +-- Lamellidens_marginalis **
| | +--+
| | | +-------- Lamellidens_lamellatus **
| | +--+
| | | +----- Lamellidens_jenkinsianus **
+--+ +--+
| | +-- Lamellidens_generosus **
| +--+
| +-- Lamellidens_mainwaringi **
|
| +----------- Parreysia_keralaensis **
| +-----------------------------+
| | | +-------- Parreysia_corrugata **
| | +--+
| | | +----- Parreysia_rakhinensis **
| | +--+
| | | +-- Parreysia_plagiosoma **
| | +--+
| | +-- Parreysia_nagpoorensis **
| |
| | +-- Leoparreysia_burmana **
+--+ +-----+
| | +-- Leoparreysia_whitteni **
| +-----------------------------+
| | | +----- Leoparreysia_olivacea **
| | +--+
| | | +-- Leoparreysia_canefrii **
| | +--+
| | +-- Leoparreysia_tavoyensis **
| |
| | +-------- Nitia_teretiuscula **
+--+ +--------------------------+
| | | +----- Coelatura_aegyptiaca **
| | +--+
| | | +-- Coelatura_kunenensis **
| | +--+
| | +-- Nyassunio_nyassaensis **
| |
| | +----------------- Indonaia_cylindrica **
| | |
+--+ +--------------+ +-------------- Indonaia_gratiosa **
| | | |
| | +--+ +----------- Indonaia_andersoniana **
| | | |
| | +--+ +-- Indonaia_caerulea **
| | | +-----+
| | | | +-- Indonaia_subclathrata **
| | +--+
| | | +----- Indonaia_rugosa **
+--+ +--+
| | +-- Indonaia_bonneaudii **
| +--+
| +-- Indonaia_shurtleffiana **
|
| +----------------------------- Indochinella_pugio **
| |
| | +-------- Radiatula_crispisulcata **
| | |
| | +-----------------+ +-- Radiatula_chaudhurii **
+--+ | | +--+
| | | | +-- Radiatula_myitkyinae **
| | +--+
| | | +-- Radiatula_myitthanensis **
| | | |
+--+ +--+-- Radiatula_mouhoti **
| |
| +-- Radiatula_chindwinensis **
|
| +----------------------- Scabiellus_songkramensis **
| |
+--+ +-------------------- Harmandia_somboriensis **
| |
+--+ +----------------- Unionetta_fabagina **
| |
+--+ +-- Scabies_crispata **
| +-----------+
| | +-- Scabies_mandarinus **
+--+
| +----------- Scabies_nucleus **
| |
+--+ +-- Scabies_humilis **
| +-----+
| | +-- Scabies_pilata **
+--+
| +----- Scabies_phaselus **
+--+
| +-- Scabies_anceps **
+--+
+-- Scabies_scobinatus **
Bolotov et al., 2022 [Publ 4239], fig. S2
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 54 |
54 |
52 |
5 |
4 |
4 |
1 |
43.0 |
+-- Pseudodon_lenyanensis
|
+-- Nyeinchanconcha_nyeinchani
+-----------------------------------------------+
| +-- Indopseudodon_kayinensis
| |
| +-- Sundadontina_tanintharyiensis
|
| +----------- Parreysia_keralaensis **
| +-----------------------------------+
| | | +-------- Parreysia_corrugata **
| | +--+
| | | +----- Parreysia_nagpoorensis **
| | +--+
--+ | | +-- Parreysia_plagiosoma **
| | +--+
| | +-- Parreysia_rakhinensis **
| |
| | +----------- Trapezidens_scutum **
| | +-----+
| | | | +-------- Trapezidens_exolescens **
| | | +--+
| | | | +----- Trapezidens_yeti **
| | | +--+
| | | | +-- Trapezidens_dolichorhynchus **
+--+ | +--+
| +--------------------------+ +-- Trapezidens_angustior **
| | |
| | | +----- Lamellidens_savadiensis **
| | | +--------+
| | | | | +-- Lamellidens_corrianus **
| | | | +--+
| | | | +-- Lamellidens_nongyangensis **
| | +--+
| | | +-- Lamellidens_marginalis **
| | | +--------+
| | | | +-- Lamellidens_ferrugineus **
| | +--+
| | | +-------- Lamellidens_lamellatus **
| | +--+
| | | +----- Lamellidens_jenkinsianus **
+--+ +--+
| | +-- Lamellidens_mainwaringi **
| +--+
| +-- Lamellidens_generosus **
|
| +----------- Leoparreysia_burmana **
| +-----------------------------+
| | | +-------- Leoparreysia_whitteni **
| | +--+
| | | +----- Leoparreysia_olivacea **
| | +--+
| | | +-- Leoparreysia_canefrii **
| | +--+
| | +-- Leoparreysia_tavoyensis **
| |
+--+ +-------- Nitia_teretiuscula **
| +-----------------------------+
| | | +----- Coelatura_aegyptiaca **
| | +--+
| | | +-- Coelatura_kunenensis **
| | +--+
| | +-- Nyassunio_nyassaensis **
| |
| | +-- Indonaia_cylindrica **
| | +--------+
+--+ | +-- Indonaia_gratiosa **
| |
| +-----------------------+ +----- Indonaia_shurtleffiana **
| | | +--+
| | | | | +-- Indonaia_bonneaudii **
| | | | +--+
| | +--+ +-- Indonaia_rugosa **
| | |
| | | +----- Indonaia_andersoniana **
| | +--+
+--+ | +-- Indonaia_caerulea **
| +--+
| +-- Indonaia_subclathrata **
|
| +-------------------------------- Indochinella_pugio **
| |
| | +----------- Radiatula_crispisulcata **
| | |
| | +-----------------+ +-- Radiatula_chaudhurii **
+--+ | | +-----+
| | | | +-- Radiatula_myitkyinae **
| | +--+
| | | +----- Radiatula_mouhoti **
| | +--+
+--+ | +-- Radiatula_myitthanensis **
| +--+
| +-- Radiatula_chindwinensis **
|
| +-------------------------- Scabiellus_songkramensis **
| |
+--+ +----------------------- Harmandia_somboriensis **
| |
+--+ +-------------------- Unionetta_fabagina **
| |
+--+ +----------------- Scabies_crispata **
| |
+--+ +-- Scabies_humilis **
| +-----------+
| | +-- Scabies_mandarinus **
+--+
| +----------- Scabies_nucleus **
+--+
| +-------- Scabies_pilata **
+--+
| +----- Scabies_anceps **
+--+
| +-- Scabies_phaselus **
+--+
+-- Scabies_scobinatus **
Bolotov et al., 2017 [Publ 3764], fig. S1
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 41 |
41 |
28 |
5 |
90 |
90 |
3 |
39.0 |
+----- Trapezidens_exolescens **
+--------+
| | +-- Trapezidens_dolichorhynchus **
| +--+
+-----------+ +-- Trapezidens_angustior **
| |
| | +----------- Lamellidens_lamellatus **
| | |
| +--+ +-------- Lamellidens_jenkinsianus **
| | |
| | | +----- Lamellidens_generosus **
| +--+ |
| | +----- Lamellidens_mainwaringi **
| | |
| +--+ +-- Lamellidens_ferrugineus **
| +--+
| | +-- Lamellidens_marginalis **
| |
+--+ | +-- Lamellidens_savadiensis **
| | +--+
| | +-- Lamellidens_corrianus[1] **
| |
| | +-------- Parreysia_keralaensis **
| | +--------------+
| | | | +----- Parreysia_plagiosoma **
| | | +--+
| | | | +-- Parreysia_corrugata **
| | | +--+
| | | +-- Parreysia_nagpoorensis **
| | |
| | | +-------- Leoparreysia_olivacea **
| | | +-----------+
| | | | | +----- Leoparreysia_tavoyensis[2] **
| +--+ | +--+
| | | | +-- Leoparreysia_canefrii **
| | | +--+
| | | +-- Leoparreysia_tavoyensis[1] **
| | |
| | | +-- Nitia_teretiuscula **
| | | |
| | | +-- Coelatura_gabonensis **
| | | |
| | | +-- Prisodontopsis_mweruensis **
| +--+ |
| | +-- Coelatura_kunenensis **
| | |
| | +--------------+-- Coelatura_aegyptiaca[1] **
| | | |
| | | +-- Coelatura_aegyptiaca[2] **
| | | |
| | | +-- Prisodontopsis_aviculaeformis **
| | | |
| | | +-- Coelatura_luapulaensis **
| | | |
| | | +-- Coelatura_choziensis **
| +--+
| | +-------- Indonaia_shurtleffiana **
| | |
| | +-----+ +----- Indonaia_rugosa **
| | | | |
| | | | | +-- Indonaia_andersoniana **
| | | +--+--+
| | | | +-- Indonaia_caerulea **
| | | |
| +--+ | +-- Indonaia_bonneaudii **
| | +--+
| | +-- Lamellidens_corrianus[2] **
| |
| | +----------- Indochinella_pugio **
| | |
| | | +-- Scabies_humilis **
| +--+ |
| | +-----+-- Scabies_phaselus **
--+ | | |
| +--+ +-- Scabies_mandarinus **
| |
| | +----- Radiatula_mouhoti **
| +--+
| | +-- Radiatula_myitkyinae **
| +--+
| +-- Radiatula_chaudhurii **
|
| +----- Indopseudodon_indawgyiensis
| +-----------+
| | | +-- Indopseudodon_salwenianus
| | +--+
| | +-- Indopseudodon_bogani
| |
| | +-- Nyeinchanconcha_nyeinchani
| | +-----------+
| +-----+ | +-- Sundadontina_cumingii
| | | |
| | | | +----- Bineurus_exilis
| | | +--------+
| | | | | +-- Bineurus_mouhotii
| | | | +--+
| | | | +-- Bineurus_loeiensis
| | | |
| | +--+ +-------- Sundadontina_tumida
| | +-----+
| | | | +----- Pseudodon_vagulus
| | | +--+
| | | | +-- Pseudodon_mekongi
| | | +--+
| | | +-- Pseudodon_vondembuschianus
| | |
| | | +----------- Thaiconcha_callifera
| | +--+
| | | +-------- Pilsbryoconcha_exilis[3]
| | +--+
| | | +----- Pilsbryoconcha_exilis[2]
| | +--+
| | | +-- Pilsbryoconcha_exilis[1]
| | +--+
| | +-- Pilsbryoconcha_carinifera
| |
| | +-- Ensidens_telus
| | +-----+
| | | +-- Ensidens_ingallsianus
| | +--+
| | | | +----- Ensidens_jaculus
| | | +--+
| | | | +-- Ensidens_spiculus
| | | +--+
| | +-----+ +-- Ensidens_sagittarius
| | | |
| | | | +-------- Hyriopsis_desowitzi
| | | | |
| | | | +-------- Rectidens_sumatrensis
+-----+ | +--+
| | | +----- Hyriopsis_panhai
| | | |
| | +--+ +-- Hyriopsis_myersiana
| | | |
| +--+ +--+-- Hyriopsis_bialata
| | | |
| | | +-- Hyriopsis_khoratensis
| | |
| | | +----- Physunio
| | | +--------+
| | | | | +-- Physunio_superbus
| | | | +--+
| | | | +-- Physunio_modelli
| | | |
| | +--+ +-------- Yaukthwa_inlenensis
| | | +--+
| | | | | +----- Yaukthwa_panhai
| | | | +--+
| | | | | +-- Yaukthwa_nesemanni
| | | | +--+
| | +--+ +-- Yaukthwa_rectangularis
| | |
| | | +-- Lens_contradens
| | | +-----+
| | | | +-- Lens_micropterus
| | +--+
| | | +----- Lens_eximius
| | +--+
| | | +-- Lens_comptus
| | +--+
| | +-- Lens_novoselovi
| |
| | +-- Amblema_plicata
| | |
| | +-- Ortmanniana_ligamentina
| | |
| | +-- Lampsilis_cardium
| | |
| | +-- Cambarunio_iris
+--+ |
| +--+-- Elliptio_complanata
| | |
| | +-- Eurynia_dilatata
| | |
| | +-- Pleurobema_sintoxia
| | |
| | +-- Quadrula_quadrula
| | |
| | +-- Quadrula_verrucosa
| |
| | +-- Chamberlainia_somsakpanhai
| +--+ |
| | | +-- Sinohyriopsis_cumingii
| | | |
| | | +-- Lamprotula_caveata
| | | |
| | | +-- Lamprotula_leaii
| | | |
| | | +-- Potomida_littoralis
| | | |
| | | +-- Pronodularia_japanensis
| | | |
| | +--+-- Gonidea_angulata
| | |
| | +-- Leguminaia_wheatleyi
| | |
| | +-- Leguminaia
| | |
| | +-- Microcondylaea_bonellii
| | |
| | +-- Sinosolenaia_carinata
| | |
| | +-- Sinosolenaia_oleivora
| | |
| | +-- Sinosolenaia_recognita
+-----------+
| +-- Unio_crassus
| |
| +-- Unio_pictorum
| |
| +-- Unio_tumidus
| |
| +-- Aculamprotula_tortuosa
| |
| +-- Cuneopsis_heudei
| +--+
| | +-- Tchangsinaia_piscicula
| | |
| | +-- Cuneopsis_rufescens
| | |
| | +-- Nodularia_douglasiae
| | |
| | +-- Nodularia_nuxpersicae
| | |
| | +-- Schistodesmus_lampreyanus
| |
| | +-- Alasmidonta_marginata
| | |
| | +-- Anodonta_anatina
+--+ |
| +-- Anodonta_cygnea
| |
| +-- Anodonta_nuttalliana
| |
| +-- Lasmigona_costata
| |
| +-- Pseudanodonta_complanata
| |
| +-- Pyganodon_grandis
| |
| +-- Simpsonaias_ambigua
| |
| +-- Strophitus_undulatus
+--+
+-- Sinanodonta_lauta
|
+-- Cristaria_plicata
|
+-- Cristaria_radiata
|
+-- Pletholophus_tenuis
|
+-- Sinanodonta_lucida
|
+-- Sinanodonta_jourdyi
|
+-- Lanceolaria_gladiola
|
+-- Lanceolaria_grayii[1]
|
+-- Lanceolaria_grayii[2]
Bolotov et al., 2020 [Publ 4100], fig. 1
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 41 |
41 |
39 |
5 |
79 |
74 |
1 |
36.0 |
+-------- Trapezidens_exolescens **
+--+
| | +----- Trapezidens_scutum **
| +--+
| | +-- Trapezidens_dolichorhynchus **
+--------------------------+ +--+
| | +-- Trapezidens_angustior **
| |
| | +-------- Lamellidens_savadiensis **
| +--+
| | +----- Lamellidens_generosus **
| +--+
| | +-- Lamellidens_ferrugineus **
| +--+
| +-- Lamellidens_mainwaringi **
|
| +----------- Parreysia_keralaensis **
| +--+
+--+ | | +-------- Parreysia_plagiosoma **
| | | +--+
| | | | +----- Parreysia_nagpoorensis[2] **
| | | +--+
| | | | +-- Parreysia_nagpoorensis[1] **
| | +--------------------+ +--+
| | | | +-- Parreysia_rakhinensis **
| | | |
| | | | +-- Leoparreysia_burmana **
| | | | +-----+
| | | | | +-- Leoparreysia_whitteni **
| | | +-----+
| | | | +----- Leoparreysia_tavoyensis **
| | | +--+
| +--+ | +-- Leoparreysia_olivacea **
| | +--+
| | +-- Leoparreysia_canefrii **
| |
| | +-- Coelatura_aegyptiaca **
| | +-----------------------------+
| | | +-- Nitia_teretiuscula **
| | |
| | | +----------- Indonaia_shurtleffiana **
| | | +-----------------+
| +--+ | | +-------- Indonaia_rugosa **
| | | +--+
| | | | +----- Indonaia_andersoniana **
| | | +--+
| | | | +-- Indonaia_subclathrata **
| +--+ +--+
| | +-- Indonaia_caerulea **
| |
| | +-------------------------- Indochinella_pugio **
| | |
| | | +-------- Radiatula_crispisulcata **
| | | +--------------+
| +--+ | | +----- Radiatula_mouhoti **
| | | +--+
| | | | +-- Radiatula_chaudhurii **
| | | +--+
| +--+ +-- Radiatula_myitkyinae **
| |
| | +-------------------- Scabiellus_songkramensis **
| | |
| | | +-- Unionetta_fabagina **
| +--+ +--------------+
| | | +-- Harmandia_somboriensis **
| | |
| +--+ +-------------- Scabies_nucleus **
| | |
| | | +-- Scabies_crispata **
| +--+ +--------+
| | | +-- Scabies_mandarinus **
| | |
| +--+ +-- Scabies_humilis **
| | +-----+
| | | +-- Scabies_pilata **
| +--+
| | +----- Scabies_anceps **
| +--+
| | +-- Scabies_phaselus **
| +--+
--+ +-- Scabies_scobinatus **
|
| +----------- Hyriopsis_desowitzi
| |
| +--+ +-------- Rectidens_sumatrensis
| | | |
| | +--+ +-- Hyriopsis_khoratensis
| | | +--+
| | | | +-- Hyriopsis_bialata
| | +--+
| | | +-- Hyriopsis_myersiana
| +--------+ +--+
| | | +-- Hyriopsis_panhai
| | |
| | | +-- Ensidens_telus
| | | +-----+
| | | | +-- Ensidens_ingallsianus
| | | |
| | +-----+ +-- Ensidens_jaculus
| | | +--+
| | | | +-- Ensidens_spiculus
| | +--+
| | | +-- Ensidens_dugasti
| | +--+
| | +-- Ensidens_sagittarius
| +-----+
| | | +-- Trapezoideus_lenya
| | | +-----+
| | | | +-- Trapezoideus_foliaceus
| | | +-----------+
| | | | | +----- Physunio_superbus
| | | | +--+
| | | | | +-- Physunio_modelli
| | | | +--+
| | | | +-- Physunio
| | | |
| | | | +-- Yaukthwa_elongatula
| | | | +-----------+
| | | | | +-- Yaukthwa_baniensis
| | +--+ |
| | | +--+ +----- Yaukthwa_inlenensis
| | | | | +-----+
| | | | | | | +-- Yaukthwa_avaensis
| | | | | | +--+
| | | | +--+ +-- Yaukthwa_paiensis
| | | | |
| | | | | +-------- Yaukthwa_panhai
| | | | +--+
| | | | | +----- Yaukthwa_rectangularis
| | +--+ +--+
| | | | +-- Yaukthwa_zayleymanensis
| | | +--+
| | | +-- Yaukthwa_nesemanni
| | |
| | | +----- Lens_misellus
| | | +-----+
| | | | | +-- Lens_contradens
| | | | +--+
| | | | +-- Lens_micropterus
| | +-----+
| | | +----- Lens_pallegoixi
| | | +--+
+-----------+ | | | +-- Lens_comptus
| | | +--+
| +--+ +-- Lens_novoselovi
| |
| | +----- Lens_maenamensis
| +--+
| | +-- Lens_eximius
| +--+
| +-- Lens_rolfbrandti
|
| +-- Sinosolenaia_iridinea
| |
| +-- Sinosolenaia_recognita[1]
| |
| +-- Sinosolenaia_carinata
| |
| +-- Leguminaia_wheatleyi
| |
| +-- Microcondylaea_bonellii
| |
| +-- Gonidea_angulata
| |
| +-- Ptychorhynchus_pfisteri
| |
| +-- Parvasolenaia_rivularis
| |
| +-----------------------+-- Lamprotula_leaii
| | |
| | +-- Lamprotula
| | |
| | +-- Lamprotula_caveata
| | |
| | +-- Potomida_littoralis
| | |
| | +-- Pronodularia_japanensis
| | |
| | +-- Sinohyriopsis_cumingii
| | |
+--+ +-- Chamberlainia_somsakpanhai
| |
| +-- Sinosolenaia_recognita[2]
| |
| +-- Sinosolenaia_oleivora
|
| +-------- Indopseudodon_kayinensis
| +--------------+
| | | +----- Indopseudodon_indawgyiensis
| | +--+
| | | +-- Indopseudodon_salwenianus
| | +--+
| | +-- Indopseudodon_bogani
+--+
| +----- Bineurus_loeiensis
| +--------------+
| | | +-- Bineurus_exilis
| | +--+
| | +-- Bineurus_mouhotii
| |
+--+ +----------------- Nyeinchanconcha_nyeinchani
| |
| | +-- Thaiconcha_callifera
| | +-----+
| | | +-- Thaiconcha_munelliptica
+--+ +-----+
| | | +----- Pilsbryoconcha_exilis[2]
| | +--+
| | | +-- Pilsbryoconcha_exilis[1]
| | +--+
| | +-- Pilsbryoconcha
+--+
| +-------- Pseudodon_lenyanensis
| +--+
| | | +----- Pseudodon_vagulus
| | +--+
| | | +-- Pseudodon_vondembuschianus
| | +--+
+--+ +-- Pseudodon_mekongi
|
| +-- Sundadontina_taskaevi
| +-----+
| | +-- Sundadontina_brandti
+--+
| +----- Sundadontina_tumida
+--+
| +-- Sundadontina_cumingii
+--+
+-- Sundadontina_tanintharyiensis
Bolotov et al., 2020 [Publ 4100], fig. S1
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 40 |
40 |
37 |
5 |
74 |
74 |
1 |
37.0 |
+-------- Trapezidens_exolescens **
+--+
| | +----- Trapezidens_scutum **
| +--+
| | +-- Trapezidens_dolichorhynchus **
+--------------------------------+ +--+
| | +-- Trapezidens_angustior **
| |
| | +-------- Lamellidens_savadiensis **
| +--+
| | +----- Lamellidens_generosus **
| +--+
| | +-- Lamellidens_ferrugineus **
| +--+
| +-- Lamellidens_mainwaringi **
|
| +----------- Parreysia_keralaensis **
| +--+
+--+ | | +-------- Parreysia_nagpoorensis[2] **
| | | +--+
| | | | +----- Parreysia_nagpoorensis[1] **
| | | +--+
| | | | +-- Parreysia_rakhinensis **
| | +--------------------------+ +--+
| | | | +-- Parreysia_plagiosoma **
| | | |
| | | | +-- Leoparreysia_burmana **
| | | | +-----+
| | | | | +-- Leoparreysia_whitteni **
| | | +-----+
| | | | +----- Leoparreysia_olivacea **
| | | +--+
| +--+ | +-- Leoparreysia_tavoyensis **
| | +--+
| | +-- Leoparreysia_canefrii **
| |
| | +-- Coelatura_aegyptiaca **
| | +-----------------------------------+
| | | +-- Nitia_teretiuscula **
| | |
| | | +-------- Indonaia_shurtleffiana **
| +--+ +--------------------------+
| | | | +----- Indonaia_andersoniana **
| | | +--+
| | | | +-- Indonaia_caerulea **
| | | +--+
| +--+ +-- Indonaia_subclathrata **
| |
| | +-------------------------------- Indochinella_pugio **
| | |
| | | +----- Radiatula_mouhoti **
| +--+ +-----------------------+
| | | | +-- Radiatula_myitkyinae **
| | | +--+
| +--+ +-- Radiatula_chaudhurii **
| |
| | +-------------------------- Radiatula_crispisulcata **
| | |
| +--+ +----------------------- Scabiellus_songkramensis **
| | |
| +--+ +-------------------- Harmandia_somboriensis **
| | |
| +--+ +----------------- Unionetta_fabagina **
| | |
| +--+ +-------------- Scabies_mandarinus **
| | |
| +--+ +----------- Scabies_crispata **
| | |
| +--+ +-------- Scabies_nucleus **
| | |
| +--+ +-- Scabies_humilis **
| | +--+
| | | +-- Scabies_pilata **
| +--+
| | +-- Scabies_phaselus **
| | |
| +--+-- Scabies_scobinatus **
| |
--+ +-- Scabies_anceps **
|
| +-------------- Hyriopsis_desowitzi
| +--+
| | | +----------- Rectidens_sumatrensis
| | +--+
| | | +-------- Hyriopsis_panhai
| | +--+
| | | +----- Hyriopsis_myersiana
| | +--+
| +--------+ | +-- Hyriopsis_khoratensis
| | | +--+
| | | +-- Hyriopsis_bialata
| | |
| | | +-- Ensidens_telus
| | | +-----+
| | | | +-- Ensidens_ingallsianus
| | | |
| | +--------+ +-- Ensidens_dugasti
| | | +--+
| | | | +-- Ensidens_sagittarius
| | +--+
| +--+ | +-- Ensidens_spiculus
| | | +--+
| | | +-- Ensidens_jaculus
| | |
| | | +----- Physunio
| | | +-----------------+
| | | | | +-- Physunio_modelli
| | | | +--+
| | | | +-- Physunio_superbus
| | | |
| | | | +-- Trapezoideus_lenya
| | | | +-----------------+
| | +--+ | +-- Trapezoideus_foliaceus
| | | |
| | | | +----- Lens_misellus
| | | | +-----+
| | | | | | +-- Lens_contradens
| | | | | +--+
| | | | | +-- Lens_micropterus
| | +--+ +-----+
| | | | | +----- Lens_maenamensis
| | | | | +--+
| | | | | | | +-- Lens_eximius
| | | | | | +--+
| | | | +--+ +-- Lens_rolfbrandti
| | | | |
| | | | | +----- Lens_pallegoixi
| | +--+ +--+
| | | | +-- Lens_novoselovi
| | | +--+
| | | +-- Lens_comptus
| | |
| | | +-- Yaukthwa_elongatula
| | | +-----------+
| | | | +-- Yaukthwa_baniensis
| | | |
| | +--+ +----- Yaukthwa_inlenensis
| | | +-----+
+-----------------+ | | | +-- Yaukthwa_avaensis
| | | +--+
| +--+ +-- Yaukthwa_paiensis
| |
| | +-------- Yaukthwa_panhai
| +--+
| | +----- Yaukthwa_rectangularis
| +--+
| | +-- Yaukthwa_zayleymanensis
| +--+
| +-- Yaukthwa_nesemanni
|
| +-- Sinohyriopsis_cumingii
| +--+
| | +-- Chamberlainia_somsakpanhai
| |
| +--+ +-- Lamprotula_leaii
| | | |
| | | +-- Lamprotula
| | | |
| | +--+-- Lamprotula_caveata
| | |
| | +-- Potomida_littoralis
| | |
| | +-- Pronodularia_japanensis
| +-----------------+
| | | +-- Sinosolenaia_iridinea
| | | |
| | | +-- Sinosolenaia_recognita[1]
| | | |
| | | +-- Sinosolenaia_carinata
| | | |
| | | +-- Leguminaia_wheatleyi
| | | |
| | | +-- Microcondylaea_bonellii
| | +-----+
| | +-- Gonidea_angulata
| | |
| | +-- Ptychorhynchus_pfisteri
+--+ |
| +-- Parvasolenaia_rivularis
| |
| +-- Sinosolenaia_recognita[2]
| |
| +-- Sinosolenaia_oleivora
|
| +-------- Indopseudodon_kayinensis
| +--------------+
| | | +----- Indopseudodon_indawgyiensis
| | +--+
| | | +-- Indopseudodon_salwenianus
| | +--+
| | +-- Indopseudodon_bogani
+--+
| +----- Bineurus_loeiensis
| +--------------+
| | | +-- Bineurus_exilis
| | +--+
| | +-- Bineurus_mouhotii
| |
+--+ +----------------- Nyeinchanconcha_nyeinchani
| |
| | +-- Thaiconcha_callifera
| | +-----+
| | | +-- Thaiconcha_munelliptica
+--+ +-----+
| | | +----- Pilsbryoconcha_exilis[2]
| | +--+
| | | +-- Pilsbryoconcha_exilis[1]
| | +--+
| | +-- Pilsbryoconcha
+--+
| +-------- Pseudodon_lenyanensis
| +--+
| | | +----- Pseudodon_vagulus
| | +--+
| | | +-- Pseudodon_vondembuschianus
| | +--+
+--+ +-- Pseudodon_mekongi
|
| +-- Sundadontina_tanintharyiensis
| +-----+
| | +-- Sundadontina_cumingii
+--+
| +----- Sundadontina_tumida
+--+
| +-- Sundadontina_taskaevi
+--+
+-- Sundadontina_brandti
Huang et al., 2019 [Publ 3843], fig. S3
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 36 |
36 |
30 |
5 |
165 |
126 |
3 |
20.0 |
+----- Trapezidens_exolescens **
+-----+
| | +-- Trapezidens_angustior **
| +--+
| +-- Trapezidens_dolichorhynchus **
|
+--------------+ +-- Lamellidens_corrianus **
| | +-----+
| | | +-- Lamellidens_savadiensis **
| | |
| | | +----- Lamellidens_lamellatus **
| +--+ |
| | +----- Lamellidens_jenkinsianus **
| | |
| | | +-- Lamellidens_marginalis **
| +--+--+
| | +-- Lamellidens_ferrugineus **
| |
+-----------------------------+ | +-- Lamellidens_mainwaringi[1] **
| | | |
| | +--+-- Lamellidens_mainwaringi[2] **
| | |
| | +-- Lamellidens_generosus **
| |
| | +----- Parreysia_keralaensis **
| | +-----------------+
| | | | +-- Parreysia_nagpoorensis **
| | | +--+
| | | +-- Parreysia_plagiosoma **
| | |
| | | +----- Leoparreysia_canefrii **
| +--+ +--------------+
| | | | +-- Leoparreysia_tavoyensis **
| | | +--+
| | | +-- Leoparreysia_olivacea **
| | |
| | | +----------- Coelatura_aegyptiaca[2] **
| | | |
| +--+ +-----+ +-- Coelatura_aegyptiaca[1] **
| | | | +-----+
| | | | | +-- Nitia_teretiuscula **
| | | +--+
| | | | +----- Coelatura_kunenensis **
| | | +--+
| | | | +-- Coelatura_gabonensis **
| | | +--+
| +--+ +-- Prisodontopsis_mweruensis **
| |
| | +-------- Indonaia_shurtleffiana **
| | |
| | +-----+ +----- Indonaia_bonneaudii **
| | | | |
| | | +--+----- Indonaia_rugosa **
| | | |
| | | | +-- Indonaia_andersoniana **
| +--+ +--+
| | +-- Indonaia_caerulea **
| |
| | +----------- Indochinella_pugio **
| | |
| | | +----- Scabies_phaselus **
| +--+ +--+
| | | | +-- Scabies_humilis **
| | | +--+
| +--+ +-- Scabies_mandarinus **
| |
| | +----- Radiatula_mouhoti **
| +--+
| | +-- Radiatula_myitkyinae **
| +--+
--+ +-- Radiatula_chaudhurii **
|
| +----- Indopseudodon_indawgyiensis
| +--------------+
| | | +-- Indopseudodon_bogani
| | +--+
| | +-- Indopseudodon_salwenianus
| |
| | +-------- Pilsbryoconcha_exilis[3]
| +--------------------------------+ +--------+
| | | | | +----- Pilsbryoconcha_exilis[2]
| | | | +--+
| | | | | +-- Pilsbryoconcha_carinifera
| | | | +--+
| | | | +-- Pilsbryoconcha_exilis[1]
| | | |
| | +--+ +----------- Sundadontina_tumida
| | | +--+
| | | | | +-------- Pseudodon_vagulus
| | | | +--+
| | | | | +----- Pseudodon_vondembuschianus[1]
| | | | +--+
| | | | | +-- Pseudodon_mekongi
| | +--+ +--+
| | | +-- Pseudodon_vondembuschianus[2]
| | |
| | | +-- Nyeinchanconcha_nyeinchani
| | | +--------+
| | | | +-- Sundadontina_cumingii
| | +--+
| | | +-------- Thaiconcha_callifera
| | +--+
| | | +----- Bineurus_loeiensis
| | +--+
| | | +-- Bineurus_mouhotii
| | +--+
| | +-- Bineurus_exilis
| |
| | +-- Ensidens_telus
| | +-----+
| | | +-- Ensidens_ingallsianus
| | +-----+
| | | | +----- Ensidens_sagittarius
| | | +--+
| | | | +-- Ensidens_spiculus
| | +--+ +--+
| | | | +-- Ensidens_jaculus
| | | |
| | | | +----------- Hyriopsis_desowitzi
| | | | |
| | | +--+ +-------- Rectidens_sumatrensis
| | | | |
| | | +--+ +-- Hyriopsis_panhai
+--+ | | +--+
| | | | +-- Hyriopsis_khoratensis
| +--------------------------------+ +--+
| | | | +-- Hyriopsis_bialata
| | | +--+
| | | +-- Hyriopsis_myersiana
| | |
| | | +----- Physunio
| | | +--------+
| | | | | +-- Physunio_superbus
| | | | +--+
| | | | +-- Physunio_modelli
| | | |
| | +--+ +-------- Yaukthwa_inlenensis
| | | +--+
| | | | | +----- Yaukthwa_panhai
| | | | +--+
| | | | | +-- Yaukthwa_nesemanni
| | | | +--+
| | +--+ +-- Yaukthwa_rectangularis
| | |
| | | +-- Lens_contradens
| | | +-----+
| | | | +-- Lens_micropterus
| | +--+
| | | +----- Lens_eximius
| | +--+
| | | +-- Lens_novoselovi
| | +--+
| | +-- Lens_comptus
| |
| | +----- Chamberlainia_somsakpanhai
| | +--------------------------------------+
| | | | +-- Sinohyriopsis_cumingii
| | | +--+
| | | +-- Sinohyriopsis_schlegelii
| | |
| | | +-- Potomida_littoralis
| | | +-----+
| | | | +-- Pronodularia_japanensis
| | | +-----+
| | | | | +----- Lamprotula_leaii
| | | | +--+
| | | | | +-- Lamprotula_caveata
+--+ | | +--+
| | | +-- Lamprotula_cornuumlunae
| +--+ +--------------------------+
| | | | | +----- Sinosolenaia_recognita
| | | | | +-----+
| | | | | | | +-- Sinosolenaia_oleivora
| | | | | | +--+
| | | | | | +-- Sinosolenaia_iridinea
| | | | +--+
| | | | | +----- Gonidea_angulata
| | | | | +--+
| | | | | | | +-- Leguminaia_wheatleyi
| | | | | | +--+
| | | | +--+ +-- Microcondylaea_bonellii
| | | | |
| | | | | +----- Ptychorhynchus_pfisteri
| | +--+ +--+
| | | | +-- Parvasolenaia_rivularis
| | | +--+
| | | +-- Koreosolenaia_sitgyensis
| | |
| | | +-- Megalonaias_nervosa
| | | +--+
| | | | +-- Uniomerus_carolinianus
| | | +--------------------------------+
| | | | | +-- Quadrula_quadrula
| | | | +--+
| | | | +-- Quadrula_verrucosa
| | | |
| | | | +-------- Elliptoideus_sloatianus
| | +--+ |
| | | +--------------------------+ +-- Elliptio_complanata
| | | | | +--+
| | | | | | +-- Eurynia_dilatata
| | | | +--+
| | | | | +-- Fusconaia_flava
| | +--+ +--+
| | | +-- Pleurobema_sintoxia
| | |
| | | +-------------------------------- Amblema_plicata
| | | |
| | +--+ +----------------------------- Toxolasma_parvum
| | | |
+--+ +--+ +-- Obliquaria_reflexa
| | +-----------------------+
| | | +-- Plectomerus_dombeyanus
| +--+
| | +----------------------- Potamilus_leptodon
| | |
| +--+ +-------------------- Potamilus_alatus
| | |
| | | +-- Medionidus_conradicus
| +--+ +--------------+
| | | +-- Ptychobranchus_subtentus
| | |
| +--+ +-- Ellipsaria_lineolata
| | +-----------+
| | | +-- Truncilla_truncata
| +--+
| | +----------- Leaunio_lienosus
| | |
| +--+ +-- Venustaconcha_ellipsiformis
| | +-----+
| | | +-- Cambarunio_iris
| +--+
| | +----- Ortmanniana_ligamentina
| +--+
| | +-- Lampsilis_cardium
| +--+
| +-- Lampsilis_ornata
|
| +----------------------------------- Acuticosta_chinensis
| |
| | +-------- Aculamprotula_tortuosa
| | |
| | +-----------+ +-- Aculamprotula_coreana
| | | | +--+
| | | | | +-- Aculamprotula_scripta
| | | +--+
| | | | +-- Aculamprotula_fibrosa
| | | +--+
| | | +-- Aculamprotula_tientsinensis
| | +-----------+
| | | | +-------- Unio_tumidus
+-----------+ | | +--------+
| | | | | +----- Unio_crassus
| | | | +--+
| | | | | +-- Unio_delphinus
| | | | +--+
| | +--+ +-- Unio_pictorum
| | |
| | | +----- Nodularia_douglasiae[1]
| | | +--------+
| | | | | +-- Nodularia_douglasiae[2]
| | | | +--+
| | +--+ +-- Nodularia_nuxpersicae
| | |
+--+ | +----------- Schistodesmus_lampreyanus
| +--+
| | +-------- Tchangsinaia_piscicula
| +--+
| | +----- Cuneopsis_rufescens
| +--+
| | +-- Cuneopsis_heudei
| +--+
| +-- Cuneopsis_celtiformis
|
| +----------------------------- Lepidodesma_languilati
| |
| | +-------- Lanceolaria_gladiola
| | +-----------------+
| | | | +----- Lanceolaria_grayii
| | | +--+
+--+ | | +-- Lanceolaria_lanceolata
| | +--+
| | +-- Lanceolaria_triformis
| |
| | +-------------------- Anemina_arcaeformis
| | |
| | +--+ +----------------- Cristaria_plicata
+--+ | | |
| | +--+ +-------------- Pletholophus_tenuis
| | | |
| | +--+ +-- Sinanodonta_angula
| | | +--------+
| | | | +-- Sinanodonta_lucida
| | +--+
| | | +-------- Sinanodonta_woodiana[2]
| | +--+
| | | +----- Sinanodonta_jourdyi
+--+ +--+
| | +-- Sinanodonta_elliptica
| +--+
| +-- Sinanodonta_woodiana[1]
|
| +-- Alasmidonta_marginata
| +--------+
| | +-- Lasmigona_costata
| |
| | +----- Anodonta_anatina
| +-----+
| | | +-- Anodonta_cygnea
+-----------+ +--+
| +-- Anodonta_nuttalliana
|
| +----- Pseudanodonta_complanata
| +--+
| | | +-- Simpsonaias_ambigua
| | +--+
+--+ +-- Strophitus_undulatus
|
| +----- Pyganodon_grandis
+--+
| +-- Utterbackia_imbecillis
+--+
+-- Utterbackia_peninsularis
Huang et al., 2019 [Publ 3843], fig. S4
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 36 |
36 |
25 |
5 |
166 |
127 |
3 |
16.0 |
+-- Parreysia_keralaensis **
|
+--------------------+-- Parreysia_nagpoorensis **
| |
| +-- Parreysia_plagiosoma **
|
| +----- Trapezidens_exolescens **
| +--------+
| | | +-- Trapezidens_angustior **
| | +--+
+--------+ +-- Trapezidens_dolichorhynchus **
| |
| | +----------- Lamellidens_corrianus **
| | |
| +--+ +-------- Lamellidens_savadiensis **
| | |
| | | +----- Lamellidens_marginalis **
| +--+ |
| | +----- Lamellidens_ferrugineus **
| | |
+--+ +--+ +-- Lamellidens_mainwaringi[1] **
| | | |
| | | +-- Lamellidens_mainwaringi[2] **
| | | |
| | +--+-- Lamellidens_lamellatus **
| | |
| | +-- Lamellidens_generosus **
| | |
| | +-- Lamellidens_jenkinsianus **
| |
| | +----- Leoparreysia_tavoyensis **
| | +--------------+
| | | | +-- Leoparreysia_canefrii **
| | | +--+
| | | +-- Leoparreysia_olivacea **
| | |
| | | +-------- Coelatura_aegyptiaca[2] **
| | | |
| | | +-------- Nitia_teretiuscula **
| +--+ +--------+
| | | | +----- Coelatura_aegyptiaca[1] **
| | | | |
| | | +--+ +-- Coelatura_gabonensis **
| | | | |
| | | +--+-- Coelatura_kunenensis **
| | | |
| | | +-- Prisodontopsis_mweruensis **
| +--+
| | +----- Indonaia_shurtleffiana **
| | |
| | | +-- Indonaia_bonneaudii **
| | +--------+--+
| | | | +-- Indonaia_rugosa **
| | | |
| | | | +-- Indonaia_andersoniana **
| | | +--+
| +--+ +-- Indonaia_caerulea **
| |
| | +----------- Indochinella_pugio **
| | |
| | | +----- Scabies_humilis **
| +--+ +--+
| | | | +-- Scabies_phaselus **
| | | +--+
| +--+ +-- Scabies_mandarinus **
| |
| | +----- Radiatula_mouhoti **
| +--+
| | +-- Radiatula_myitkyinae **
| +--+
| +-- Radiatula_chaudhurii **
|
| +-------------------- Chamberlainia_somsakpanhai
| |
| +-------------------- Megalonaias_nervosa
| |
| +-------------------- Uniomerus_carolinianus
| |
| | +-- Quadrula_quadrula
| +-----------------+
| | +-- Quadrula_verrucosa
| |
| | +-- Sinohyriopsis_cumingii
| +-----------------+
| | +-- Sinohyriopsis_schlegelii
| |
| | +-- Potomida_littoralis
| | +-----+
| | | +-- Pronodularia_japanensis
| +-----------+
| | | +----- Lamprotula_leaii
| | +--+
| | | +-- Lamprotula_caveata
| | +--+
| | +-- Lamprotula_cornuumlunae
| |
| | +-------- Sinosolenaia_carinata
--+ | |
| | | +----- Gonidea_angulata
| | +--+
| | | | +-- Leguminaia_wheatleyi
| | | +--+
| | | +-- Microcondylaea_bonellii
| +-----------+
| | | +----- Ptychorhynchus_pfisteri
| | +--+
| | | | +-- Parvasolenaia_rivularis
| | | +--+
| | | +-- Koreosolenaia_sitgyensis
| | |
| | | +----- Sinosolenaia_recognita
| | +--+
| | | +-- Sinosolenaia_iridinea
| | +--+
| | +-- Sinosolenaia_oleivora
| |
| | +----- Indopseudodon_salwenianus
| | +-----+
| | | | +-- Indopseudodon_indawgyiensis
| | | +--+
| | | +-- Indopseudodon_bogani
| | |
| | | +-------- Nyeinchanconcha_nyeinchani
| | | |
| | | +-------- Thaiconcha_callifera
| +--------+ |
| | | +-------- Sundadontina_cumingii
| | | |
| | | +-------- Sundadontina_tumida[1]
| | | |
| | | +-------- Sundadontina_tumida[2]
| | | |
| | | | +----- Pilsbryoconcha_exilis[2]
| | +--+--+
| | | | +-- Pilsbryoconcha_carinifera
| | | +--+
| | | +-- Pilsbryoconcha_exilis[1]
| | |
| | | +----- Pseudodon_vagulus
| | +--+
| | | | +-- Pseudodon_mekongi
| | | +--+
| | | +-- Pseudodon_vondembuschianus
| | |
| | | +----- Bineurus_loeiensis
| | +--+
| | | +-- Bineurus_mouhotii
| | +--+
| | +-- Bineurus_exilis
| |
| | +-------------- Amblema_plicata
| | |
| | +-------------- Toxolasma_parvum
| | |
| | | +-- Obliquaria_reflexa
| | +-----------+
| | | +-- Plectomerus_dombeyanus
| | |
| | | +----------- Elliptoideus_sloatianus
| | +--+
| | | | +-------- Elliptio_complanata
| | | +--+
| | | | +----- Eurynia_dilatata
| +-----+ +--+
| | | | +-- Fusconaia_flava
| | | +--+
| | | +-- Pleurobema_sintoxia
| | |
| | | +-------- Ellipsaria_lineolata
| | | |
| | | +-------- Potamilus_leptodon
| | | |
| | | +-------- Medionidus_conradicus
| | | |
+-----+ | +-------- Potamilus_alatus
| | |
| | +-------- Ptychobranchus_subtentus
| +-----+
| +-------- Truncilla_truncata
| |
| +-------- Venustaconcha_ellipsiformis
| |
| +-------- Cambarunio_iris
| |
| +-------- Leaunio_lienosus
| |
| | +----- Ortmanniana_ligamentina
| +--+
| | +-- Lampsilis_cardium
| +--+
| +-- Lampsilis_ornata
|
| +-------------- Hyriopsis_desowitzi
| |
| +-------------- Hyriopsis_myersiana
| |
| +-------------- Hyriopsis_panhai
| |
| +--+ +-- Hyriopsis_bialata
| | +-----------+
| | | +-- Hyriopsis_khoratensis
| | |
| | | +----------- Rectidens_sumatrensis
| | | |
| | +--+ +-- Ensidens_telus
| | | +-----+
| | | | +-- Ensidens_ingallsianus
| | +--+
| | | +----- Ensidens_sagittarius
| | +--+
+--+ | +-- Ensidens_spiculus
| | +--+
| | +-- Ensidens_jaculus
| |
| | +----------- Physunio_modelli
| | |
| | +----------- Lens
| | |
| | +----------- Physunio_superbus
| | |
| | | +-------- Yaukthwa_inlenensis
| | +--+
| +-----+ | +----- Yaukthwa_panhai
| | +--+
| | | +-- Yaukthwa_nesemanni
| | +--+
| | +-- Yaukthwa_rectangularis
| |
| | +-- Lens_comptus
| | +-----+
| | | +-- Lens_novoselovi
| +--+
| | +----- Lens_eximius
| +--+
| | +-- Lens_contradens
| +--+
| +-- Lens_micropterus
|
| +----------------- Acuticosta_chinensis
| |
| +----------------- Alasmidonta_marginata
| |
| +----------------- Anodonta_anatina
| |
| +----------------- Anemina_arcaeformis
| |
| +----------------- Anodonta_cygnea
| |
| +----------------- Cristaria_plicata
| |
| +----------------- Anodonta_nuttalliana
| |
| +----------------- Lasmigona_costata
| |
| +----------------- Lepidodesma_languilati
| |
| +----------------- Pseudanodonta_complanata
| |
| +----------------- Pyganodon_grandis
| |
| +----------------- Simpsonaias_ambigua
| |
| +----------------- Strophitus_undulatus
| |
| | +-- Schistodesmus_lampreyanus
| +--------------+
| | +-- Schistodesmus_spinosus
| |
| | +-- Utterbackia_imbecillis
| +--------------+
| | +-- Utterbackia_peninsularis
| |
| | +-------- Lanceolaria_gladiola
| +--------+
| | | +----- Lanceolaria_grayii
| | +--+
+--+ | +-- Lanceolaria_lanceolata
| +--+
| +-- Lanceolaria_triformis
|
| +-------- Unio_tumidus
+--------+
| | +----- Unio_crassus
| +--+
| | +-- Unio_delphinus
| +--+
| +-- Unio_pictorum
|
| +-------- Aculamprotula_tortuosa
| |
+--------+ +-- Aculamprotula_coreana
| | +--+
| | | +-- Aculamprotula_scripta
| +--+
| | +-- Aculamprotula_fibrosa
| +--+
| +-- Aculamprotula_tientsinensis
|
| +-------- Cuneopsis_heudei
| |
| +-------- Cuneopsis_celtiformis
| |
+--------+-------- Tchangsinaia_piscicula
| |
| +-------- Cuneopsis_rufescens
| |
| | +----- Nodularia_douglasiae[2]
| +--+
| | +-- Nodularia_douglasiae[1]
| +--+
| +-- Nodularia_nuxpersicae
|
| +-------------- Pletholophus_tenuis
| |
+--+ +-- Sinanodonta_angula
| +--------+
| | +-- Sinanodonta_lucida
+--+
| +-------- Sinanodonta_woodiana[1]
+--+
| +----- Sinanodonta_jourdyi
+--+
| +-- Sinanodonta_elliptica
+--+
+-- Sinanodonta_woodiana[2]
Ortiz-Sepulveda et al., 2020 [Publ 4101], fig. 1
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 36 |
36 |
22 |
3 |
0 |
0 |
0 |
43.0 |
+-------------------------------------- Leoparreysia_tavoyensis **
|
| +----------- Indonaia_andersoniana **
| |
| +-----------------------+ +-------- Indochinella_pugio **
| | | |
--+ | | | +----- Radiatula_mouhoti **
| | +--+ |
| | | +----- Radiatula_myitkyinae **
| | | |
| | +--+----- Radiatula_chaudhurii **
| | |
| | | +-- Scabies_humilis **
+--+ | |
| +--+-- Scabies_phaselus **
| |
| +-- Scabies_mandarinus **
|
| +-------------------------------- Grandidieria_burtoni **
| |
| | +-- Pseudospatha_tanganyicensis **
| | +--------------------------+
+--+ | +-- Moncetia_anceyi **
| |
| | +-- Nitia_acuminata **
| | |
+--+ +-----------------------+-- Nitia_monceti **
| | |
| | +-- Nitia_teretiuscula **
| |
| | +-- Coelatura_aegyptiaca **
| | |
+--+ +-- Coelatura_alluaudi **
| |
| +--------------------+-- Coelatura_bakeri **
| | |
| | +-- Coelatura_hauttecoeuri **
| | |
| | +-- Coelatura_stuhlmanni **
+--+
| +-- Coelatura_leopoldvillensis **
| +-----------------+
| | +-- Coelatura_mesafricana **
| |
| | +-- Coelatura_horei **
+--+ |
| +--------------+-- Coelatura_ratidota **
| | |
| | +-- Coelatura **
+--+
| +-------------- Coelatura_gabonensis **
| |
| +-------------- Coelatura_stanleyvillensis **
+--+
| +----------- Coelatura_kunenensis **
| |
| | +----- Prisodontopsis_aviculaeformis **
+--+ +--+
| | | +-- Coelatura_briarti **
| | +--+
| | +-- Prisodontopsis_mweruensis **
+--+
| +-- Coelatura_choziensis **
| |
| +-- Coelatura_hypsiprymna **
| |
+-----+-- Coelatura_luapulaensis **
|
+-- Coelatura_mossambicensis **
|
+-- Nyassunio_nyassaensis **
Ortiz-Sepulveda et al., 2020 [Publ 4101], fig. 2
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 36 |
36 |
20 |
3 |
0 |
0 |
0 |
45.0 |
+-------------------------------- Leoparreysia_tavoyensis **
|
| +----- Radiatula_mouhoti **
| |
| +----- Radiatula_myitkyinae **
| |
| +----- Radiatula_chaudhurii **
| |
--+ +-----------------------+----- Indochinella_pugio **
| | |
| | +----- Indonaia_andersoniana **
| | |
| | | +-- Scabies_humilis **
| | | |
| | +--+-- Scabies_phaselus **
| | |
+--+ +-- Scabies_mandarinus **
|
| +----- Grandidieria_burtoni **
| +--------------------+
| | | +-- Pseudospatha_tanganyicensis **
| | +--+
| | +-- Moncetia_anceyi **
| |
| | +-- Nitia_acuminata **
+--+ |
| +--------------------+-- Nitia_monceti **
| | |
| | +-- Nitia_teretiuscula **
| |
| | +-- Coelatura_aegyptiaca **
| | |
+--+ +-- Coelatura_alluaudi **
| |
| +-----------------+-- Coelatura_bakeri **
| | |
| | +-- Coelatura_hauttecoeuri **
| | |
| | +-- Coelatura_stuhlmanni **
+--+
| +-- Coelatura_leopoldvillensis **
| +--------------+
| | +-- Coelatura_mesafricana **
| |
| | +-- Coelatura_horei **
+--+ |
| +-----------+-- Coelatura_ratidota **
| | |
| | +-- Coelatura **
| |
+--+ +-- Coelatura_gabonensis **
| +--------+
| | +-- Coelatura_stanleyvillensis **
| |
+--+ +-------- Coelatura_kunenensis **
| |
| | +-- Coelatura_briarti **
| | |
+--+ +--+-- Prisodontopsis_mweruensis **
| | |
| | +-- Prisodontopsis_aviculaeformis **
| |
+--+ +-- Coelatura_choziensis **
| |
| +-- Coelatura_hypsiprymna **
| |
+--+-- Coelatura_luapulaensis **
|
+-- Coelatura_mossambicensis **
|
+-- Nyassunio_nyassaensis **
Bolotov et al., 2017 [Publ 3761], fig. S2
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 29 |
29 |
26 |
5 |
112 |
87 |
3 |
6.0 |
+-- Parreysia_nagpoorensis **
+-----------+
| +-- Parreysia_plagiosoma **
|
+--------+ +-- Trapezidens_dolichorhynchus **
| | +--------+
| | | +-- Trapezidens_exolescens **
| | |
| +--+ +-------- Lamellidens_savadiensis **
| | |
| | +-------- Lamellidens_corrianus **
| +--+
| | +-- Lamellidens_marginalis **
| | +--+
| | | +-- Lamellidens_ferrugineus **
+-----+ +--+
| | | +-- Lamellidens_jenkinsianus **
| | +--+
| | +-- Lamellidens_mainwaringi **
| |
| | +-- Leoparreysia_olivacea **
| | +-----------------+
| | | +-- Leoparreysia_tavoyensis **
| | |
| | | +-------------- Nitia_teretiuscula **
| | | +--+
| | | | | +----------- Coelatura_aegyptiaca[1] **
| +--+ | +--+
| | | | +-------- Coelatura_aegyptiaca[2] **
| | | +--+
| | | | +----- Coelatura_gabonensis **
| | | +--+
| | | | +-- Coelatura_kunenensis **
| +--+ +--+
| | +-- Prisodontopsis_mweruensis **
| |
| | +-------- Indonaia_shurtleffiana **
| | +-----+
| | | | +----- Indonaia_andersoniana **
| | | +--+
| | | | +-- Indonaia_rugosa **
| | | +--+
| +--+ +-- Indonaia_bonneaudii **
| |
| | +-- Indochinella_pugio **
| | +--------+
| | | +-- Scabies_crispata **
| | |
| +--+ +-- Radiatula_myitkyinae **
| | +-----+
| | | +-- Radiatula_chaudhurii **
| +--+
| | +----- Scabies_mandarinus **
| +--+
| | +-- Scabies_humilis **
| +--+
| +-- Scabies_phaselus **
|
| +----- Unio_tumidus
| +-----+
| | | +-- Unio_crassus
| | +--+
| | +-- Unio_pictorum
| +-----+
--+ | | +-------- Aculamprotula_tortuosa
| | | |
| | | +-------- Schistodesmus_lampreyanus
| | | |
| | +--+ +-- Nodularia_douglasiae
| | +-----+
| | | +-- Nodularia_nuxpersicae
| | |
| | | +----- Cuneopsis_heudei
| | +--+
| | | +-- Tchangsinaia_piscicula
| | +--+
| +--------+ +-- Cuneopsis_rufescens
| | |
| | | +-- Lanceolaria_grayii
| | | +-----------+
| | | | +-- Lanceolaria_gladiola
| | | |
| | | | +-- Cristaria_plicata
| | | | +--------+
| | | | | +-- Cristaria_radiata
| | | +--+
| | | | | +-------- Pletholophus_tenuis
| | | | +--+
| | | | | +----- Sinanodonta_lucida
| | +--+ +--+
| | | | +-- Sinanodonta_lauta
| | | +--+
| | | +-- Sinanodonta_jourdyi
| | |
| | | +-------- Anodonta_anatina
| | | |
| | | +-------- Pseudanodonta_complanata
| | | |
| | | | +-- Anodonta_cygnea
| | +-----+-----+
| | | +-- Anodonta_nuttalliana
| | |
| | | +----- Pyganodon_grandis
| | | |
| | | +----- Simpsonaias_ambigua
| | +--+
| | +----- Strophitus_undulatus
| | |
| | | +-- Alasmidonta_marginata
| | +--+
| | +-- Lasmigona_costata
| |
| | +-------- Lens_contradens
| | +-----+
+--+ | | +----- Lens_eximius
| | +--+
| | | +-- Lens_comptus
| | +--+
| +--+ +-- Lens_novoselovi
| | |
| | | +----- Physunio
| | | +-----+
| | | | | +-- Physunio_modelli
| | | | +--+
| | +--+ +-- Physunio_superbus
| | |
| | | +-------- Yaukthwa_inlenensis
| | +--+
| | | +----- Yaukthwa_panhai
| +-----+ +--+
| | | | +-- Yaukthwa_rectangularis
| | | +--+
| | | +-- Yaukthwa_nesemanni
| | |
| | | +----- Hyriopsis_panhai
| | | |
| | | +--------+----- Hyriopsis_khoratensis
| | | | |
| | | | | +-- Hyriopsis_myersiana
| | | | +--+
| | +--+ +-- Hyriopsis_bialata
| | |
| | | +----------- Hyriopsis_desowitzi
| | | |
| | | +----------- Rectidens_sumatrensis
| | +--+
| | | +-- Ensidens_telus
| | | +-----+
| | | | +-- Ensidens_ingallsianus
| | +--+
| | | +----- Ensidens_sagittarius
| | +--+
| | | +-- Ensidens_spiculus
+--+ +--+
| +-- Ensidens_jaculus
|
| +-------- Amblema_plicata
| +--+
| | | +----- Cambarunio_iris
| | +--+
| | | +-- Ortmanniana_ligamentina
| | +--+
| +--------+ +-- Lampsilis_cardium
| | |
| | | +-- Quadrula_quadrula
| | | +-----+
| | | | +-- Quadrula_verrucosa
| | +--+
| | | +----- Pleurobema_sintoxia
| | +--+
| | | +-- Elliptio_complanata
| | +--+
| | +-- Eurynia_dilatata
| |
| | +-------- Sinohyriopsis_cumingii
| | |
| | +-----+ +-- Lamprotula_caveata
+--+ | | +--+
| | | | +-- Lamprotula_leaii
| | +--+
| | | +-- Potomida_littoralis
| +--+ +--+
| | | +-- Pronodularia_japanensis
| | |
| | | +----------- Sinosolenaia_carinata
| | | |
| | | | +-- Sinosolenaia_oleivora
| | +--+--------+
| | | +-- Sinosolenaia_recognita
| | |
| | | +-------- Gonidea_angulata
| | +--+
| | | +----- Microcondylaea_bonellii
+--+ +--+
| | +-- Leguminaia_wheatleyi
| +--+
| +-- Leguminaia
|
| +-------- Pilsbryoconcha_exilis[3]
| +-----+
| | | +----- Pilsbryoconcha_exilis[2]
| | +--+
| | | +-- Pilsbryoconcha_exilis[1]
| | +--+
| | +-- Pilsbryoconcha_carinifera
| |
+--+ +----------- Indopseudodon_indawgyiensis
| |
| | +-------- Thaiconcha_callifera
| +--+
| | | +----- Bineurus_exilis
| | +--+
| | | +-- Bineurus_mouhotii
+--+ +--+
| +-- Bineurus_loeiensis
|
| +----- Pseudodon_vagulus
| +--+
| | | +-- Pseudodon_mekongi
| | +--+
| | +-- Pseudodon_vondembuschianus
+--+
| +-- Pseudodon_inoscularis
| +--+
| | +-- Sundadontina_tumida
+--+
| +-- Nyeinchanconcha_nyeinchani
+--+
+-- Sundadontina_cumingii
Pfeiffer et al., 2018 [Publ 3827], fig. 2
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 28 |
28 |
25 |
5 |
1 |
0 |
0 |
27.0 |
+-- Trapezidens_dolichorhynchus **
+-----------------------------------------+
| +-- Lamellidens_savadiensis **
|
| +----- Parreysia_keralaensis **
--+ +-----------------------------------+
| | | +-- Parreysia_nagpoorensis **
| | +--+
| | +-- Parreysia_plagiosoma **
| |
+--+ +-- Leoparreysia_tavoyensis **
| +-----------------------------------+
| | +-- Leoparreysia_olivacea **
| |
| | +----- Nitia_teretiuscula **
+--+ +-----------------------------+
| | | +-- Coelatura_aegyptiaca **
| | +--+
| | +-- Prisodontopsis_aviculaeformis **
+--+
| +----- Indonaia_bonneaudii **
| +--------------------------+
| | | +-- Indonaia_andersoniana **
| | +--+
+--+ +-- Indonaia_caerulea **
|
| +----------------------------- Indochinella_pugio **
| |
+--+ +-- Radiatula_chaudhurii **
| +-----------------------+
| | +-- Radiatula_myitkyinae **
+--+
| +----------------------- Scabiellus_songkramensis **
| |
+--+ +-------------------- Unionetta_fabagina[2] **
| |
+--+ +----------------- Harmandia_somboriensis **
| |
+--+ +-------------- Unionetta_fabagina[1] **
| |
+--+ +-- Scabies_crispata **
| +--------+
| | +-- Scabies_mandarinus **
+--+
| +-------- Scabies_nucleus **
| |
+--+ +-- Scabies_pilata **
| +--+
| | +-- Scabies_humilis **
+--+
| +-- Scabies_phaselus **
| |
+--+-- Scabies_anceps **
|
+-- Scabies_scobinatus **
Graf et al., 2014 [Publ 3591], fig. 3
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 25 |
25 |
12 |
2 |
2 |
1 |
1 |
29.0 |
+----------------------------- Leoparreysia_tavoyensis **
|
--+ +-------------------------- Nitia_teretiuscula **
| |
+--+ +----------------------- Coelatura_aegyptiaca[2] **
| |
+--+ +-------------------- Coelatura_aegyptiaca[1] **
| |
| | +----------------- Coelatura_gabonensis **
+--+ |
| | +-------------- Coelatura_kunenensis **
| | |
| | | +-- Coelatura_choziensis[1] **
+--+ | |
| | +-- Coelatura_choziensis[2] **
| | |
| | +-- Coelatura_luapulaensis[1] **
| | |
+--+ +--------+-- Coelatura_luapulaensis[2] **
| | |
| | +-- Coelatura_luapulaensis[3] **
| | |
| | +-- Coelatura_luapulaensis[4] **
| | |
| | +-- Coelatura_luapulaensis[5] **
| |
+--+ +----- Prisodontopsis_mweruensis[2] **
| |
| +----- Prisodontopsis_mweruensis[3] **
| +--+
| | | +-- Prisodontopsis_mweruensis[4] **
| | | |
| | +--+-- Prisodontopsis_mweruensis[5] **
| | |
+--+ +-- Prisodontopsis_aviculaeformis[7] **
|
| +----- Prisodontopsis_aviculaeformis[1] **
| |
| | +-- Prisodontopsis_aviculaeformis[2] **
+--+ |
| +-- Prisodontopsis_aviculaeformis[3] **
| |
| +-- Prisodontopsis_aviculaeformis[4] **
+--+
+-- Prisodontopsis_aviculaeformis[5] **
|
+-- Prisodontopsis_aviculaeformis[6] **
|
+-- Prisodontopsis_mweruensis[1] **
Bolotov et al., 2018 [Publ 3826], fig. 2
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 24 |
24 |
22 |
5 |
61 |
61 |
3 |
15.0 |
+-- Trapezidens_exolescens **
+--------+
| +-- Trapezidens_dolichorhynchus **
+--------------+
| | +-------- Lamellidens_savadiensis **
| +--+
| | +----- Lamellidens_ferrugineus **
| +--+
| | +-- Lamellidens_generosus **
| +--+
+--------+ +-- Lamellidens_mainwaringi **
| |
| | +-- Parreysia_keralaensis **
| | +--------------------+
| | | +-- Parreysia_nagpoorensis **
| | |
| | | +----- Leoparreysia_olivacea **
| +--+ +--------------+
| | | | +-- Leoparreysia_canefrii **
| | | +--+
| | | +-- Leoparreysia_tavoyensis **
| | |
| +--+ +-- Coelatura_aegyptiaca **
| | +--------------+
| | | +-- Nitia_teretiuscula **
| | |
| | | +-------- Indonaia_shurtleffiana **
| +--+ +-----+
| | | | +----- Indonaia_rugosa **
| | | +--+
| | | | +-- Indonaia_caerulea **
| | | +--+
| +--+ +-- Indonaia_andersoniana **
| |
| | +----------- Indochinella_pugio **
| | |
| | | +----- Radiatula_myitkyinae **
| +--+ +--+
| | | | +-- Radiatula_mouhoti **
| | | +--+
| +--+ +-- Radiatula_chaudhurii **
| |
| | +----- Scabies_mandarinus **
| +--+
| | +-- Scabies_humilis **
| +--+
--+ +-- Scabies_phaselus **
|
| +-- Indopseudodon_indawgyiensis
| |
| +--+-- Indopseudodon_bogani
| | |
| | +-- Indopseudodon_salwenianus
| |
| | +-- Bineurus_mouhotii
| +--------------------------+ |
| | | +-- Pseudodon_vagulus
| | | |
| | | +-- Pseudodon_mekongi
| | | |
| | | +-- Pilsbryoconcha_exilis
| | +--+
| | +-- Pilsbryoconcha_carinifera
| | |
| | +-- Bineurus_loeiensis
| | |
| | +-- Bineurus_exilis
| | |
| | +-- Pseudodon_vondembuschianus
| |
| | +-- Ensidens_telus
| | |
| | +-- Ensidens_ingallsianus
| | |
| | +-- Ensidens_jaculus
| | |
| | +-- Hyriopsis_desowitzi
| | +--+
| | | +-- Hyriopsis_myersiana
| | | |
| | | +-- Hyriopsis_khoratensis
| | | |
+--+ | +-- Ensidens_spiculus
| | |
| | +-- Ensidens_sagittarius
| |
| +-----------------------+ +-- Lens_eximius
| | | |
| | | +-- Physunio_modelli[1]
| | | |
| | | +-- Yaukthwa_nesemanni
| | | |
| | | +-- Yaukthwa_panhai
| | | |
| | | +-- Lens_comptus
| | | |
| | +--+-- Lens_novoselovi
| | |
| | +-- Physunio_modelli[2]
| | |
| | +-- Physunio
| | |
| | +-- Yaukthwa_inlenensis
| | |
| | +-- Lens_contradens
| | |
| | +-- Lens_micropterus
| |
| | +-- Lamprotula_leaii
+--+ |
| +-- Potomida_littoralis
| |
| +-- Pronodularia_japanensis
| |
| +-- Gonidea_angulata
| |
| +-- Leguminaia_wheatleyi
| |
| +-- Microcondylaea_bonellii
| |
| +-----------------------+-- Amblema_plicata
| | |
| | +-- Ortmanniana_ligamentina
| | |
| | +-- Lampsilis_cardium
| | |
| | +-- Cambarunio_iris
| | |
| | +-- Elliptio_complanata
| | |
| | +-- Pleurobema_sintoxia
| | |
+--+ +-- Quadrula_quadrula
|
| +----- Unio_tumidus
| +--+
| | | +-- Unio_crassus
| | +--+
| +--------------+ +-- Unio_pictorum
| | |
| | | +----- Nodularia_douglasiae[1]
| | +--+
| | | +-- Nodularia_nuxpersicae
| | +--+
+--+ +-- Nodularia_douglasiae[2]
|
| +-- Lanceolaria_gladiola
| +-----------------+
| | +-- Lanceolaria_grayii
| |
| | +-------- Cristaria_plicata
+--+ +--------+
| | | +----- Pletholophus_tenuis
| | +--+
| | | +-- Sinanodonta_jourdyi
+--+ +--+
| +-- Sinanodonta_lauta
|
| +-------------- Anodonta_anatina
+--+
| +----------- Pyganodon_grandis
+--+
| +-------- Simpsonaias_ambigua
+--+
| +----- Strophitus_undulatus
+--+
| +-- Alasmidonta_marginata
+--+
+-- Lasmigona_costata
Bolotov et al., 2022 [Publ 4250], fig. 8
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 24 |
24 |
22 |
4 |
25 |
23 |
1 |
30.0 |
+-- Pseudodon_lenyanensis
+--------+
| +-- Sundadontina_tanintharyiensis
+--------+
| | +-------- Indopseudodon_kayinensis
| +--+
| | +----- Indopseudodon_indawgyiensis
| +--+
| | +-- Indopseudodon_salwenianus
| +--+
+--------+ +-- Indopseudodon_bogani
| |
| | +----- Trapezoideus_mitanensis
| | +-----------+
| | | | +-- Trapezoideus_foliaceus
| | | +--+
| | | +-- Trapezoideus_lenya
| +--+
| | +-- Yaukthwa_elongatula
| | +-----------+
| | | +-- Yaukthwa_baniensis
| | |
| +--+ +----- Yaukthwa_inlenensis
| | +-----+
| | | | +-- Yaukthwa_paiensis
| | | +--+
| +--+ +-- Yaukthwa_avaensis
| |
| | +-------- Yaukthwa_panhai
| +--+
--+ | +----- Yaukthwa_rectangularis
| +--+
| | +-- Yaukthwa_nesemanni
| +--+
| +-- Yaukthwa_zayleymanensis
|
| +-------- Lamellidens_savadiensis **
| +-----+
| | | +----- Lamellidens_ferrugineus **
| | +--+
| | | +-- Lamellidens_generosus **
| +-----------+ +--+
| | | +-- Lamellidens_mainwaringi **
| | |
| | | +----------- Trapezidens_exolescens **
| | +--+
| | | +-------- Trapezidens_scutum **
| | +--+
| | | +----- Trapezidens_yeti **
| | +--+
+--+ | +-- Trapezidens_angustior **
| +--+
| +-- Trapezidens_dolichorhynchus **
|
| +----------------------- Parreysia_rakhinensis **
| |
| | +-- Leoparreysia_whitteni **
| | +-----+
| | | +-- Leoparreysia_burmana **
+--+ +-----------+
| | | +----- Leoparreysia_olivacea **
| | +--+
| | | +-- Leoparreysia_canefrii **
| | +--+
+--+ +-- Leoparreysia_tavoyensis **
|
| +-- Indonaia_andersoniana **
| +--------------+
| | +-- Indonaia_subclathrata **
+--+
| +-------------- Indochinella_pugio **
| |
+--+ +----------- Radiatula_crispisulcata **
| |
+--+ +-- Radiatula_chaudhurii **
| +-----+
| | +-- Radiatula_myitkyinae **
+--+
| +----- Radiatula_mouhoti **
+--+
| +-- Radiatula_chindwinensis **
+--+
+-- Radiatula_myitthanensis **
Jeratthitikul et al., 2019 [Publ 3922], fig. 1
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 23 |
23 |
21 |
5 |
0 |
0 |
0 |
32.0 |
+-- Lamellidens_mainwaringi **
+-----------------------------------------------+
| +-- Trapezidens_exolescens **
--+
| +----------------------------------------------- Leoparreysia_canefrii **
| |
+--+ +-------------------------------------------- Parreysia_keralaensis **
| |
+--+ +----------------------------------------- Coelatura_aegyptiaca **
| |
| | +----- Indonaia_rugosa **
+--+ +--------------------------------+
| | | +-- Indonaia_caerulea **
| | +--+
| | +-- Indonaia_andersoniana **
+--+
| +----------------------------------- Indochinella_pugio **
| |
| | +----- Radiatula_mouhoti **
+--+ +--------------------------+
| | | +-- Radiatula_myitkyinae **
| | +--+
+--+ +-- Radiatula_chaudhurii **
|
| +----------------------------- Scabiellus_songkramensis **
+--+
| +-------------------------- Harmandia_somboriensis **
+--+
| +----------------------- Unionetta_fabagina **
+--+
| +-------------------- Scabies_crispata **
+--+
| +----------------- Scabies_mandarinus **
+--+
| +-------------- Scabies_pilata **
+--+
| +----------- Scabies_nucleus **
+--+
| +-------- Scabies_humilis **
+--+
| +----- Scabies_scobinatus **
+--+
| +-- Scabies_phaselus **
+--+
+-- Scabies_anceps **
Bolotov et al., 2022 [Publ 4250], fig. S2
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 15 |
15 |
13 |
3 |
0 |
0 |
0 |
31.0 |
+----- Parreysia_rakhinensis **
+--------------+
| | +-- Leoparreysia_tavoyensis **
| +--+
| +-- Leoparreysia_olivacea **
|
--+ +----------- Indonaia_rugosa **
| +-----+
| | | +-------- Indonaia_shurtleffiana **
| | +--+
| | | +----- Indonaia_andersoniana **
| | +--+
+--+ | +-- Indonaia_subclathrata **
| +--+
| +-- Indonaia_caerulea **
|
| +-------------- Indochinella_pugio **
| |
+--+ +----------- Radiatula_crispisulcata **
| |
+--+ +-- Radiatula_chaudhurii **
| +-----+
| | +-- Radiatula_myitkyinae **
+--+
| +----- Radiatula_chindwinensis **
+--+
| +-- Radiatula_mouhoti **
+--+
+-- Radiatula_myitthanensis **
Konopleva et al., 2023 [Publ 4262], fig. 8
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 14 |
14 |
10 |
1 |
70 |
70 |
1 |
37.0 |
+----------------------- Indonaia_andersoniana **
|
+--------------+ +-------------------- Indochinella_pugio **
| | |
| +--+ +----------------- Radiatula_mouhoti **
| | |
| +--+ +-------------- Scabiellus_songkramensis **
| | |
| | | +-- Unionetta_fabagina **
| +--+ +--------+
| | | +-- Harmandia_somboriensis **
| | |
| | | +-- Scabies_humilis **
| +--+ +-----+
| | | +-- Scabies_pilata **
| | |
| | | +----- Scabies_crispata **
| | +--+
| +--+ | +-- Scabies_nucleus **
| | +--+
| | +-- Scabies_mandarinus **
| |
| | +-- Scabies_phaselus **
| | |
| +-----+-- Scabies_anceps **
| |
| +-- Scabies_scobinatus **
|
| +-------- Indopseudodon_kayinensis
| +-----------------------+
| | | +----- Indopseudodon_indawgyiensis
| | +--+
| | | +-- Indopseudodon_salwenianus
| | +--+
| | +-- Indopseudodon_bogani
| |
| | +-- Pilsbryoconcha_schomburgki
--+ +--+ +--------+
| | | | +-- Pilsbryoconcha_exilis[1]
| | | +-----------------+
| | | | | +-------- Pilsbryoconcha
| | | | +--+
| | | | | +----- Pilsbryoconcha_linguaeformis
| | | | +--+
| | | | | +-- Pilsbryoconcha_exilis[2]
| | +--+ +--+
| | | +-- Pilsbryoconcha_carinifera
| | |
| | | +-------------------------- Songkhlanaia_tamodienica
| | | |
| | | | +-- Namkongnaia_inkhavilayi
| | | | +--------------------+
| | +--+ | +-- Namkongnaia_lemeslei
| | | |
| | | | +-------- Pseudodon_lenyanensis
| | | | +-----------+
| | | | | | +----- Pseudodon_vagulus
| | +--+ | +--+
| | | | | +-- Pseudodon_vondembuschianus
| | | | +--+
| | | | +-- Pseudodon_mekongi
| | | |
| | | | +-- Sundadontina_taskaevi
| | +--+ +--------+
| | | | +-- Sundadontina_brandti
| | | +-----+
| | | | | +-------- Sundadontina_tumida
| | | | +--+
| | | | | +----- Sundadontina_tanintharyiensis
| | | | +--+
| | | | | +-- Sundadontina_plugpomenica
| | +--+ +--+
| | | +-- Sundadontina_cumingii
+--+ |
| | +-------------- Nyeinchanconcha_nyeinchani
| | |
| | | +----- Thaiconcha_thaiensis
| +--+ +-----+
| | | | +-- Thaiconcha_munelliptica
| | | +--+
| +--+ +-- Thaiconcha_callifera
| |
| | +-------- Bineurus_loeiensis
| +--+
| | +----- Bineurus_mouhotii
| +--+
| | +-- Bineurus_anodontinum
| +--+
| +-- Bineurus_exilis
|
| +-- Gonidea_angulata
| +--+
| | +-- Leguminaia_wheatleyi
| +-----------------------+
| | | +-- Lamprotula_leaii
| | +--+
| | +-- Potomida_littoralis
| |
| | +-- Ensidens_ingallsianus
| | +-----+
| | | +-- Ensidens_telus
| | |
| | +--------------+ +-- Ensidens_jaculus
| | | | +--+
| | | | | +-- Ensidens_spiculus[1]
| | | +--+
| | | | +-- Ensidens_spiculus[2]
| | +--+ +--+
| | | | +-- Ensidens_dugasti
+-----+ | |
| | | +-- Hyriopsis_desowitzi
| | | +-----------------+
| | | | +-- Hyriopsis_delaportei
| | +--+
| | | +----------------- Rectidens_sumatrensis
| | | |
| | +--+ +-------------- Hyriopsis_panhai
| | | |
| | +--+ +----------- Hyriopsis_myersiana
| | | |
| | +--+ +-- Hyriopsis_khoratensis
| | | +-----+
| | | | +-- Hyriopsis_bialata
| | +--+
| | | +----- Hyriopsis_sakhonensis
+--+ +--+
| | +-- Hyriopsis_phuphaniensis
| +--+
| +-- Hyriopsis_kratiensis
|
| +----- Physunio_modelli
| +-----+
| | | +-- Physunio_superbus
| | +--+
| | +-- Physunio_massini
| +-----+
| | | +-- Trapezoideus_thachiadensis
| | | +-----+
| | | | +-- Trapezoideus_subviridis
| | +--+
| | | +----- Trapezoideus_mitanensis
| | +--+
| | | +-- Trapezoideus_foliaceus
+--------+ +--+
| +-- Trapezoideus_lenya
|
| +-- Yaukthwa_nesemanni
| +-----------+
| | +-- Yaukthwa_elongatula
| |
| | +----- Lens_misellus
+--+ +-----+
| | | +-- Lens_contradens
| | +--+
| | +-- Lens_micropterus
+--+
| +----- Lens_pallegoixi
| +--+
| | | +-- Lens_novoselovi
| | +--+
+--+ +-- Lens_comptus
|
| +----- Lens_maenamensis
+--+
| +-- Lens_eximius
+--+
+-- Lens_rolfbrandti
Bolotov et al., 2019 [Publ 3974], fig. 1
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 13 |
13 |
11 |
3 |
5 |
5 |
1 |
28.0 |
+----------- Yaukthwa_inlenensis
+--------+
| | +-------- Yaukthwa_elongatula
| +--+
| | +----- Yaukthwa_panhai
| +--+
| | +-- Yaukthwa_nesemanni
| +--+
| +-- Yaukthwa_rectangularis
--+
| +-- Leoparreysia_whitteni **
| +-----+
| | +-- Leoparreysia_burmana **
| +--------+
| | | +----- Leoparreysia_olivacea **
| | +--+
| | | +-- Leoparreysia_tavoyensis **
| | +--+
+--+ +-- Leoparreysia_canefrii **
|
| +-- Nitia_teretiuscula **
| +-----------+
| | +-- Coelatura_aegyptiaca **
+--+
| +----------- Indochinella_pugio **
| |
+--+ +-- Radiatula_myitkyinae **
| +-----+
| | +-- Radiatula_chaudhurii **
+--+
| +----- Radiatula_chindwinensis **
+--+
| +-- Radiatula_mouhoti **
+--+
+-- Radiatula_myitthanensis **
Konopleva et al., 2019 [Publ 3900], fig. 1
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 12 |
12 |
9 |
1 |
40 |
38 |
1 |
24.0 |
+-- Indopseudodon_indawgyiensis
|
+--+-- Indopseudodon_bogani
| |
| +-- Indopseudodon_salwenianus
|
| +-- Bineurus_mouhotii
+--------------------------+ |
| | +-- Pseudodon_vagulus
| | |
| | +-- Pseudodon_mekongi
| | |
| | +-- Pilsbryoconcha_exilis
| +--+
| +-- Pilsbryoconcha_carinifera
| |
| +-- Bineurus_loeiensis
| |
| +-- Bineurus_exilis
| |
| +-- Pseudodon_vondembuschianus
|
| +----------- Indonaia_shurtleffiana **
| +--+
| | | +-------- Indonaia_rugosa **
| | +--+
--+ | | +----- Indonaia_andersoniana **
| | +--+
| | | +-- Indonaia_caerulea **
| +--------------+ +--+
| | | +-- Indonaia_subclathrata **
| | |
| | | +----------- Indochinella_pugio **
| | | |
| | | | +-- Scabies_humilis **
| | +--+ |
| | | +-----+-- Scabies_mandarinus **
| | | | |
| | +--+ +-- Scabies_phaselus **
| | |
| | | +----- Radiatula_mouhoti **
| | +--+
| | | +-- Radiatula_myitkyinae **
| | +--+
| | +-- Radiatula_chaudhurii **
| |
+--+ +-- Ensidens_ingallsianus
| +-----+
| | +-- Ensidens_telus
| +-----+
| | | +----- Ensidens_sagittarius
| | +--+
| | | +-- Ensidens_jaculus
| +-----------+ +--+
| | | +-- Ensidens_spiculus
| | |
| | | +----------- Hyriopsis_desowitzi
| | | |
| | +--+----------- Rectidens_sumatrensis
| | |
| | | +-------- Hyriopsis_panhai
| | +--+
| | | +----- Hyriopsis_myersiana
| | +--+
+--+ | +-- Hyriopsis_khoratensis
| +--+
| +-- Hyriopsis_bialata
|
| +-- Yaukthwa_paiensis
| +-----+
| | +-- Yaukthwa_inlenensis
| +--------------+
| | | +----- Yaukthwa_panhai
| | +--+
| | | +-- Yaukthwa_nesemanni
| | +--+
+--+ +-- Yaukthwa_rectangularis
|
| +----- Physunio_superbus
| +--------------+
| | | +-- Physunio_modelli
| | +--+
+--+ +-- Physunio
|
| +----------------- Trapezoideus_foliaceus
| |
+--+ +-- Lens_contradens
| +-----------+
| | +-- Lens_micropterus
+--+
| +----------- Lens_misellus
+--+
| +-------- Lens_eximius
+--+
| +----- Lens_pallegoixi
+--+
| +-- Lens_comptus
+--+
+-- Lens_novoselovi
Konopleva et al., 2020 [Publ 4072], fig. 2
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 12 |
12 |
9 |
2 |
0 |
0 |
0 |
27.0 |
+-- Leoparreysia_olivacea **
|
+--------------+-- Leoparreysia_canefrii **
| |
| +-- Leoparreysia_tavoyensis **
|
--+ +-------- Lamellidens_savadiensis **
| +-----+
| | | +----- Lamellidens_generosus **
| | +--+
| | | +-- Lamellidens_mainwaringi **
+--+ +--+
| +-- Lamellidens_ferrugineus **
|
| +----------- Trapezidens_scutum **
+--+
| +-------- Trapezidens_exolescens **
+--+
| +----- Trapezidens_yeti **
+--+
| +-- Trapezidens_angustior **
+--+
+-- Trapezidens_dolichorhynchus **
Konopleva et al., 2017 [Publ 3807], fig. 6
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 11 |
11 |
9 |
4 |
20 |
10 |
1 |
18.0 |
+-------- Ensidens_sagittarius
+-----+
| | +----- Lens_micropterus
| +--+
| | +-- Physunio
+--+ +--+
| | +-- Physunio_modelli
| |
| | +----------- Sundadontina_tumida
| | |
| +--+ +-- Nyeinchanconcha_nyeinchani
| | +-----+
| | | +-- Bineurus_exilis
| +--+
| | +----- Pilsbryoconcha_carinifera
| +--+
--+ | +-- Pseudodon_vagulus
| +--+
| +-- Pseudodon_vondembuschianus
|
| +-- Trapezidens_exolescens **
| +-----+
| | +-- Trapezidens_dolichorhynchus **
| +-----+
| | | +----- Lamellidens_savadiensis **
| | +--+
| | | +-- Lamellidens_jenkinsianus **
| | +--+
+--+ +-- Lamellidens_mainwaringi **
|
| +-- Leoparreysia_tavoyensis **
| +--------+
| | +-- Leoparreysia_olivacea **
+--+
| +-------- Coelatura_aegyptiaca **
+--+
| +----- Indochinella_pugio **
+--+
| +-- Radiatula_chaudhurii **
+--+
+-- Scabies_mandarinus **
Whelan et al., 2011 [Publ 3380], fig. 1 ML
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 10 |
10 |
8 |
4 |
27 |
15 |
3 |
21.0 |
+-- Lamellidens_savadiensis **
+-----------------+
| +-- Trapezidens_dolichorhynchus **
+-----+
| | +----------------- Leoparreysia_tavoyensis **
| | |
| +--+ +-- Radiatula_chaudhurii **
| | +-----------+
| | | +-- Indochinella_pugio **
| +--+
| | +----------- Nitia_teretiuscula **
| +--+
| | +-------- Coelatura_aegyptiaca[2] **
| +--+
--+ | +----- Coelatura_aegyptiaca[1] **
| +--+
| | +-- Coelatura_gabonensis **
| +--+
| +-- Prisodontopsis_aviculaeformis **
|
| +-- Pseudodon_vondembuschianus
| +--------------------+
| | +-- Lens_contradens
| |
| | +-- Pyganodon_grandis
| | +--+
+--+ | +-- Lasmigona_costata
| +--------------+
| | | +-- Unio_pictorum
| | +--+
| | +-- Unio_caffer
+--+
| +----------------- Gonidea_angulata
| |
| | +----- Pronodularia_japanensis
+--+ +--------+
| | | +-- Potomida_littoralis
| | +--+
+--+ +-- Potomida_semirugata
|
| +----------- Pleurobema_sintoxia
+--+
| +-------- Quadrula_quadrula
+--+
| +----- Amblema_plicata
+--+
| +-- Truncilla_truncata
+--+
+-- Lampsilis_cardium
Whelan et al., 2011 [Publ 3380], fig. 1 MP
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 10 |
10 |
7 |
4 |
27 |
15 |
3 |
21.0 |
+-- Lamellidens_savadiensis **
+--------------+
| +-- Trapezidens_dolichorhynchus **
+--------+
| | +-------------- Leoparreysia_tavoyensis **
| | |
| +--+ +-- Radiatula_chaudhurii **
| | +--------+
| | | +-- Indochinella_pugio **
| +--+
| | +-------- Nitia_teretiuscula **
| | |
| +--+ +----- Coelatura_aegyptiaca[1] **
--+ | |
| +--+----- Coelatura_aegyptiaca[2] **
| |
| | +-- Coelatura_gabonensis **
| +--+
| +-- Prisodontopsis_aviculaeformis **
|
| +----------------------- Lens_contradens
| |
| | +-------------------- Pseudodon_vondembuschianus
+--+ |
| | +-- Pyganodon_grandis
| | +--+
+--+ | +-- Lasmigona_costata
| +-----------+
| | | +-- Unio_pictorum
| | +--+
| | +-- Unio_caffer
+--+
| +-------------- Gonidea_angulata
| |
| | +----- Pronodularia_japanensis
+--+ +-----+
| | | +-- Potomida_littoralis
| | +--+
+--+ +-- Potomida_semirugata
|
| +-------- Quadrula_quadrula
| |
+--+-------- Pleurobema_sintoxia
|
| +----- Amblema_plicata
+--+
| +-- Lampsilis_cardium
+--+
+-- Truncilla_truncata
Pfeiffer et al., 2019 [Publ 3937], fig. 4-5
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 10 |
10 |
8 |
4 |
47 |
40 |
3 |
24.0 |
+-- Chamberlainia_hainesiana
+--------------------+
| +-- Sinohyriopsis_cumingii
|
| +----------------- Lamprotula_cornuumlunae
| +--+
| | | +-------------- Gonidea_angulata
+--+ | +--+
| | | | +----------- Indopseudodon_crebristriatus
| | | +--+
| | | | +-------- Bineurus_anodontinum
| | | +--+
| | | | +----- Pilsbryoconcha_exilis
| | | +--+
| +--+ | +-- Pseudodon_vondembuschianus
| | +--+
| | +-- Pseudodon_inoscularis
| |
| | +----- Ensidens_ingallsianus
| | +-----+
| | | | +-- Hyriopsis_bialata
| | | +--+
| | | +-- Rectidens_sumatrensis
| +--------+
| | +-- Trapezoideus_foliaceus
| | +-----+
--+ | | +-- Physunio_superbus
| +--+
| | +----- Yaukthwa_paiensis
| +--+
| | +-- Lens_contradens
| +--+
| +-- Solenaia_emarginata
|
| +----------- Lanceolaria_lanceolata
| |
| | +----- Unio_gibbus
| +--------+ +--+
| | | | | +-- Nodularia_douglasiae
| | | | +--+
| | | | +-- Tchangsinaia_piscicula
| | +--+
| | | +-- Cristaria_plicata
| | | +--+
| | | | +-- Sinanodonta_woodiana
+-----+ +--+
| | +-- Anodonta_cygnea
| +--+
| +-- Pyganodon_grandis
|
| +-- Trapezidens_dolichorhynchus **
| +--------------+
| | +-- Lamellidens_savadiensis **
| |
+--+ +-- Leoparreysia_tavoyensis **
| +-----------+
| | +-- Leoparreysia_olivacea **
| |
+--+ +-- Coelatura_choziensis **
| +--------+
| | +-- Prisodontopsis_aviculaeformis **
+--+
| +-------- Indonaia_caerulea **
+--+
| +----- Indochinella_pugio **
+--+
| +-- Scabies_phaselus **
+--+
+-- Harmandia_somboriensis **
Pfeiffer et al., 2019 [Publ 3937], fig. 6
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 10 |
10 |
8 |
4 |
47 |
40 |
3 |
24.0 |
+-- Quadrula_quadrula
+-----------------+
| +-- Uniomerus_tetralasmus
|
| +----- Elliptoideus_sloatianus
| +--+
| | | +-- Elliptio_complanata
+--+ | +--+
| | +--------+ +-- Pleurobema_strodeanum
| | | |
| | | | +----- Reginaia_ebenus
| | | +--+
| | | | +-- Plectomerus_dombeyanus
| +--+ +--+
| | +-- Amblema_plicata
| |
| | +-- Psoronaias_semigranosa
| | +-----------+
| | | +-- Popenaias_popeii
| +--+
| | +----------- Obliquaria_reflexa
| +--+
| | +-------- Cyrtonaias_tampicoensis
| +--+
| | +----- Leaunio_lienosus
| +--+
| | +-- Lampsilis_cardium
| +--+
| +-- Pachynaias_spheniopsis
|
| +-- Chamberlainia_hainesiana
| +-----------------+
| | +-- Sinohyriopsis_cumingii
| |
| | +-- Gonidea_angulata
| | +-----------+
| | | +-- Lamprotula_cornuumlunae
| | +--+
+--+ | | +----------- Indopseudodon_crebristriatus
| | | +--+
| | | | +-------- Pilsbryoconcha_exilis
| | | +--+
--+ | | | +----- Bineurus_anodontinum
| | | +--+
| | | | +-- Pseudodon_vondembuschianus
| +--+ +--+
| | +-- Pseudodon_inoscularis
| |
| | +----- Ensidens_ingallsianus
| | +-----+
| | | | +-- Hyriopsis_bialata
| | | +--+
| | | +-- Rectidens_sumatrensis
| +-----+
| | +-- Trapezoideus_foliaceus
| | +-----+
| | | +-- Physunio_superbus
| +--+
| | +----- Yaukthwa_paiensis
| +--+
| | +-- Lens_contradens
| +--+
| +-- Solenaia_emarginata
|
| +-------- Lanceolaria_lanceolata
| +--+
| | | +----- Unio_gibbus
| | +--+
| | | +-- Tchangsinaia_piscicula
| | +--+
| +--------+ +-- Nodularia_douglasiae
| | |
| | | +-- Cristaria_plicata
| | | +--+
| | | | +-- Sinanodonta_woodiana
| | +-----+
| | | +-- Anodonta_cygnea
+--+ +--+
| +-- Pyganodon_grandis
|
| +-- Trapezidens_dolichorhynchus **
| +--------------+
| | +-- Lamellidens_savadiensis **
| |
+--+ +-- Leoparreysia_tavoyensis **
| +-----------+
| | +-- Leoparreysia_olivacea **
| |
+--+ +-- Coelatura_choziensis **
| +--------+
| | +-- Prisodontopsis_aviculaeformis **
+--+
| +-------- Indonaia_caerulea **
+--+
| +----- Indochinella_pugio **
+--+
| +-- Scabies_phaselus **
+--+
+-- Harmandia_somboriensis **
Bolotov et al., 2022 [Publ 4250], fig. S4
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 9 |
9 |
6 |
1 |
0 |
0 |
0 |
28.0 |
+-------- Lamellidens_savadiensis **
+--+
| | +----- Lamellidens_ferrugineus **
| +--+
| | +-- Lamellidens_mainwaringi **
--+ +--+
| +-- Lamellidens_generosus **
|
| +-------- Trapezidens_scutum **
| |
+--+ +----- Trapezidens_exolescens **
| |
+--+ +-- Trapezidens_dolichorhynchus **
| |
+--+-- Trapezidens_angustior **
|
+-- Trapezidens_yeti **
Pfeiffer & Graf, 2015 [Publ 3620], fig. 2
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 8 |
8 |
6 |
4 |
32 |
22 |
3 |
21.0 |
+-- Trapezidens_dolichorhynchus **
+-----------+
| +-- Lamellidens_savadiensis **
|
+-----------+ +-- Leoparreysia_olivacea **
| | +--------+
| | | +-- Leoparreysia_tavoyensis **
| +--+
| | +-------- Coelatura_aegyptiaca **
| +--+
| | +----- Indochinella_pugio **
| +--+
| | +-- Radiatula_chaudhurii **
| +--+
--+ +-- Scabies_mandarinus **
|
| +----- Unio_pictorum
| +-----------------+
| | | +-- Pyganodon_grandis
| | +--+
| | +-- Lasmigona_costata
| |
| | +----------- Quadrula_quadrula
| | +--------+
+--+ | | +-------- Amblema_plicata
| | +--+
| | | +----- Potamilus_alatus
| | +--+
| | | +-- Lampsilis_cardium
| | +--+
| | +-- Truncilla_truncata
+--+
| +-------- Ensidens_telus
| +--------+
| | | +----- Lens_micropterus
| | +--+
| | | +-- Physunio
| | +--+
| | +-- Physunio_modelli
+--+
| +-------- Gonidea_angulata
| +-----+
| | | +----- Potomida_littoralis
| | +--+
| | | +-- Pronodularia_japanensis
+--+ +--+
| +-- Lamprotula_caveata
|
| +----------- Pilsbryoconcha_carinifera
| |
+--+ +-- Pseudodon_vondembuschianus
| +-----+
| | +-- Pseudodon_vagulus
+--+
| +----- Sundadontina_tumida
+--+
| +-- Nyeinchanconcha_nyeinchani
+--+
+-- Bineurus_exilis
Konopleva et al., 2023 [Publ 4269], fig. S1, 4
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 7 |
7 |
5 |
4 |
49 |
48 |
3 |
24.0 |
+-- Trapezidens_exolescens **
+-----------+
| +-- Lamellidens_savadiensis **
+-----------------+
| | +----------- Parreysia_rakhinensis **
| +--+
| | +-------- Leoparreysia_tavoyensis **
| +--+
| | +----- Indonaia_andersoniana **
| +--+
| | +-- Indochinella_pugio **
| +--+
| +-- Radiatula_mouhoti **
|
| +-- Lampsilis_ovata
| +--------------+
| | +-- Amblema_plicata
| |
| | +----------- Sinosolenaia_recognita
| | |
| | +--+ +-- Ptychorhynchus_pfisteri
| +-----------+ | | +-----+
--+ | | | | | +-- Parvasolenaia_rivularis
| | | | +--+
| | | | | +----- Gonidea_angulata
| | | | +--+
| | | | | +-- Microcondylaea_bonellii
| | +--+ +--+
| | | +-- Leguminaia_wheatleyi
| | |
| | | +----- Chamberlainia_somsakpanhai
| | | +-----+
| | | | | +-- Sinohyriopsis_schlegelii
| | | | +--+
| | +--+ +-- Sinohyriopsis_cumingii
| | |
| | | +-------- Potomida_littoralis
| | +--+
| | | +----- Pronodularia_japanensis
| | +--+
| | | +-- Lamprotula_caveata
+--+ +--+
| +-- Lamprotula_leaii
|
| +----------------- Middendorffinaia_mongolica
| |
| | +-------- Aculamprotula_tortuosa
| | +-----+
| +--------+ | | +----- Aculamprotula_scripta
| | | | +--+
| | | | | +-- Aculamprotula_tientsinensis
| | | | +--+
| | +--+ +-- Aculamprotula_coreana
| | |
| | | +----- Unio_tumidus
| | | +-----+
| | | | | +-- Unio_pictorum
| | | | +--+
| | +--+ +-- Unio_crassus
| | |
| | | +-------- Nodularia_douglasiae
+--+ +--+
| | +----- Tchangsinaia_piscicula
| +--+
| | +-- Cuneopsis_rufescens
| +--+
| +-- Cuneopsis_heudei
|
| +-------- Lepidodesma_languilati
| +--------------+
| | | +----- Lanceolaria_gladiola
| | +--+
| | | +-- Lanceolaria_grayii
| | +--+
+--+ +-- Lanceolaria_triformis
|
| +-------------------- Acuticosta_chinensis
| |
| | +----- Anodonta_anatina
| | +-----------+
+--+ | | +-- Pseudanodonta_complanata
| | +--+
| | +-- Anodonta_cygnea
| |
| | +-------- Anemina_arcaeformis
+--+ +-----+
| | | +----- Amuranodonta_kijaensis
| | +--+
| | | +-- Buldowskia_suifunica
| | +--+
| | +-- Buldowskia_shadini
+--+
| +-------- Beringiana_beringiana
| +--+
| | | +----- Sinanodonta_schrenkii
| | +--+
| | | +-- Sinanodonta_woodiana
+--+ +--+
| +-- Sinanodonta_lauta
|
| +-------- Cristaria_radiata
| |
+--+ +-- Cristaria_truncata
| +--+
| | +-- Cristaria_plicata
+--+
| +-- Cristaria_bellua
+--+
+-- Cristaria_clessini
Konopleva et al., 2017 [Publ 3807], fig. 5
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 5 |
5 |
3 |
1 |
0 |
0 |
0 |
24.0 |
+-- Trapezidens_exolescens **
+-----+
| +-- Trapezidens_dolichorhynchus **
--+
| +----- Lamellidens_savadiensis **
+--+
| +-- Lamellidens_mainwaringi **
+--+
+-- Lamellidens_jenkinsianus **
Bolotov et al., 2022 [Publ 4250], fig. S3
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 5 |
5 |
3 |
1 |
0 |
0 |
0 |
30.0 |
+----------- Leoparreysia_burmana **
--+
| +-------- Leoparreysia_whitteni **
+--+
| +----- Leoparreysia_olivacea **
+--+
| +-- Leoparreysia_tavoyensis **
+--+
+-- Leoparreysia_canefrii **
Lopes-Lima et al., 2017 [Publ 3745], fig. 1-2
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 4 |
4 |
2 |
4 |
77 |
65 |
3 |
25.0 |
+-------- Trapezidens_dolichorhynchus **
--+
| +----- Leoparreysia_tavoyensis **
+--+
| +-- Coelatura_aegyptiaca **
+--+
+-- Indochinella_pugio **
Lopes-Lima et al., 2020 [Publ 4080], fig. 2-4
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 3 |
3 |
0 |
3 |
119 |
108 |
3 |
28.0 |
+-- Coelatura_aegyptiaca **
|
+-----------------------------------+-- Leoparreysia_tavoyensis **
| |
| +-- Radiatula_mouhoti **
|
| +-- Lens_contradens
| |
| +-- Ensidens_sagittarius
| |
| +-----------------------+-- Physunio_modelli
| | |
| | +-- Rectidens_sumatrensis
| | |
| | +-- Yaukthwa_nesemanni
| |
| | +-- Amblema_plicata
| | |
| | +-- Elliptio_complanata
| | +--------------------+
| +--------+ | +-- Lampsilis_cardium
| | | | |
| | | | +-- Quadrula_quadrula
| | | |
| | | | +----- Chamberlainia_somsakpanhai
| | | | +-----------+
| | | | | | +-- Sinohyriopsis_cumingii
| | | | | +--+
| | | | +--+ +-- Sinohyriopsis_schlegelii
| | | | | |
| | | | | | +-------------- Pronodularia_japanensis
| | +--+ | | |
| | | | +--+ +----------- Potomida_littoralis
| | | | | |
| | | | +--+ +-------- Lamprotula_caveata
| | | | | |
--+ | | | +--+ +----- Lamprotula_leaii[2]
| | | | | |
| | | | +--+----- Lamprotula_leaii[3]
| | | | |
| | | | | +-- Lamprotula_seomjinensis
| | | | +--+
| | | | +-- Lamprotula_leaii[1]
| | +--+
| | | +-- Indopseudodon_indawgyiensis
| | | +--------+
| | | | +-- Indopseudodon_salwenianus
| | | |
| | | +-----+ +-- Pilsbryoconcha_exilis
| | | | | +-----+
| | | | | | +-- Pilsbryoconcha_carinifera
| | | | +--+
| | | | | +----- Pseudodon_mekongi
| | | | | |
| | | | +--+ +-- Nyeinchanconcha_nyeinchani
| | | | | |
| | | | +--+-- Bineurus_mouhotii
| | | | |
| | +--+ +-- Sundadontina_tumida
| | |
| | | +-------- Inversidens_brandtii
| | | |
| | | +-----+ +-- Obovalis_omiensis
| | | | | +--+
| | | | | | +-- Ptychorhynchus_pfisteri
| | | | +--+
| | | | | +-- Parvasolenaia_rivularis
| | | | +--+
| | +--+ +-- Koreosolenaia_sitgyensis
| | |
+--+ | +----- Gonidea_angulata
| | +-----+
| | | | +-- Leguminaia_wheatleyi
| | | +--+
| +--+ +-- Microcondylaea_bonellii
| |
| | +-------- Sinosolenaia_carinata
| +--+
| | +----- Sinosolenaia_recognita
| +--+
| | +-- Sinosolenaia_iridinea
| +--+
| +-- Sinosolenaia_oleivora
|
| +-------------------------------- Acuticosta_chinensis
| |
| +-------------------------------- Lepidodesma_languilati
| |
| | +----------- Aculamprotula_tortuosa
| | +--------------+
| | | | +-------- Aculamprotula_scripta
| | | +--+
| | | | +----- Aculamprotula_coreana
| | | +--+
| | | | +-- Aculamprotula_fibrosa
| | | +--+
| +-----+ +-- Aculamprotula_tientsinensis
| | |
| | | +----------------------- Middendorffinaia_mongolica
| | | |
| | | | +----- Unio_gibbus
| | | | |
| | | | +--------------+----- Unio_tumidus
| | +--+ | |
| | | | | +-- Unio_crassus
| | | | +--+
| | | | +-- Unio_pictorum
| | | |
| | | | +-------------- Nodularia_breviconcha
| | +--+ +--+
| | | | | +----------- Nodularia[2]
| | | | +--+
| | | | | +-------- Nodularia_nuxpersicae
| | | | +--+
| | | | | +----- Nodularia_nipponensis
| | | | +--+
| | | | | +-- Nodularia_douglasiae
| | +--+ +--+
| | | +-- Nodularia[1]
| | |
+--+ | +-- Schistodesmus_lampreyanus
| | +-----------+
| | | +-- Schistodesmus_spinosus
| | |
| | | +----- Cuneopsis_rufescens
| | +--------+
| +--+ | +-- Cuneopsis_heudei
| | +--+
| | +-- Cuneopsis_celtiformis
| |
| | +----------- Tchangsinaia_piscicula
| +--+
| | +-------- Inversiunio_jokohamensis
| +--+
| | +----- Inversiunio_verrucosus
| +--+
| | +-- Inversiunio_reinianus
| +--+
| +-- Inversiunio_yanagawensis
|
| +----------------- Lanceolaria_oxyrhyncha
| |
| +-----------+ +-------------- Lanceolaria_gladiola
| | | |
| | +--+ +-- Lanceolaria_bilirata
| | | +--------+
| | | | +-- Lanceolaria_grayii
| | +--+
| | | +-------- Lanceolaria_acrorrhyncha
| | +--+
| | | +----- Lanceolaria_kihirai
| | +--+
| | | +-- Lanceolaria_lanceolata
| | +--+
| | +-- Lanceolaria_triformis
| |
| | +-------- Pseudanodonta_complanata
+--+ |
| | +----- Anodonta_cygnea
| +--+
| | | +-- Anodonta_anatina
| +-----------------+ +--+
| | | +-- Anodonta_nuttalliana
| | |
| | | +----- Pyganodon_grandis
| | | |
| | | +----- Simpsonaias_ambigua
| | +--+
| | +----- Strophitus_undulatus
| | |
| | | +-- Alasmidonta_marginata
| | +--+
| | +-- Lasmigona_costata
+--+
| +-- Cristaria_plicata
| +--------------------+
| | +-- Cristaria_clessini
| |
| | +-------------- Anemina_arcaeformis
| | |
| +--------+ +----------- Amuranodonta_kijaensis
| | | |
| | +--+ +-------- Buldowskia_suifunica
| | | |
| | +--+ +-- Buldowskia_flavotincta
+--+ | +--+
| | | +-- Buldowskia_shadini
| +--+
| | +-- Buldowskia_iwakawai
| +--+
| +-- Buldowskia_kamiyai
|
| +-------------------- Pletholophus_tenuis
| |
| | +-------- Beringiana_japonica
| | +--------+
+--+ | | +----- Beringiana_beringiana
| | +--+
| | | +-- Beringiana_fukuharai
| | +--+
+--+ +-- Beringiana_gosannensis
|
| +-- Sinanodonta_angula
| +-----------+
| | +-- Sinanodonta_lucida
| |
+--+ +----------- Sinanodonta_schrenkii
| |
| | +-- Sinanodonta_calipygos
| +--------+
+--+ +-- Sinanodonta_tumens
|
| +-------- Sinanodonta_woodiana[2]
| |
+--+ +----- Sinanodonta_elliptica
| |
+--+----- Sinanodonta_jourdyi
|
| +-- Sinanodonta_lauta
+--+
+-- Sinanodonta_woodiana[1]
Choi et al., 2020 [Publ 4169], fig. 5
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 3 |
3 |
1 |
3 |
6 |
6 |
1 |
31.0 |
+----- Radiatula_mouhoti **
+--------+
| | +-- Lamellidens_generosus **
| +--+
--+ +-- Leoparreysia_canefrii **
|
| +----------- Nodularia_breviconcha
| |
+--+ +-- Nodularia[1]
| +-----+
| | +-- Nodularia[2]
+--+
| +----- Nodularia_douglasiae
+--+
| +-- Nodularia_nipponensis
+--+
+-- Nodularia_nuxpersicae
Chen et al., 2023 [Publ 4302], fig. 1
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 3 |
3 |
0 |
3 |
37 |
35 |
3 |
27.0 |
+-- Pronodularia_japanensis
+--+
| +-- Lamprotula_leaii
+-----------------------------+
| | +-- Lampsilis_siliquoidea
| +--+
| +-- Quadrula_quadrula
|
| +-- Coelatura_aegyptiaca **
--+ |
| +-----------------------------+-- Indonaia_andersoniana **
| | |
| | +-- Parreysia_nagpoorensis **
| |
| | +-- Anemina_arcaeformis
+--+ +--------------------------+
| | +-- Amuranodonta_kijaensis
| |
| | +----- Pletholophus_tenuis
| | +--------------------+
| | | | +-- Sinanodonta_jourdyi
+--+ | +--+
| | +-- Beringiana_beringiana
| |
| | +----------------------- Lanceolaria_gladiola
| | |
| | +----------------------- Acuticosta_chinensis
+--+ |
| +----------------------- Lepidodesma_languilati
| |
| +----------------------- Lanceolaria_oxyrhyncha
| |
| +----------------------- Cristaria_plicata
| |
| | +-- Alasmidonta_marginata
+--+ +--+
| | +-- Lasmigona_costata
+-----------------+
| | +-- Anodonta_anatina
| +--+
| +-- Pseudanodonta_complanata
|
| +-------------------- Aculamprotula_fibrosa
| |
| | +----- Middendorffinaia_mongolica
| | +-----------+
+--+ | | +-- Nodularia_douglasiae
| | +--+
| | +-- Nodularia_breviconcha
| |
| | +-- Cuneopsis_celtiformis
+--+ +-----------+
| | +-- Cuneopsis_heudei
| |
| | +-- Diaurora_aurorea
| | +--+
| | | +-- Diaurora_laeve
| +--------+
+--+ | +-- Schistodesmus_lampreyanus
| +--+
| +-- Schistodesmus_spinosus
|
| +-- Pseudocuneopsis_sichuanensis
| +--------+
| | +-- Pseudocuneopsis_capitata
| |
+--+ +-- Inversiunio_yanagawensis
| +-----+
| | +-- Inversiunio_reinianus
+--+
| +----- Tchangsinaia_piscicula
+--+
| +-- Unio_pictorum
+--+
+-- Unio_crassus
Chen et al., 2023 [Publ 4302], fig. 2
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 3 |
3 |
1 |
3 |
37 |
35 |
3 |
26.0 |
+-- Pronodularia_japanensis
+--+
| +-- Lamprotula_leaii
+--------------------------+
| | +-- Lampsilis_siliquoidea
| +--+
| +-- Quadrula_quadrula
|
| +----- Parreysia_nagpoorensis **
--+ +-----------------------+
| | | +-- Coelatura_aegyptiaca **
| | +--+
| | +-- Indonaia_andersoniana **
| |
| | +-------------------------- Acuticosta_chinensis
+--+ |
| | +----------------------- Lepidodesma_languilati
| | |
| | | +-- Lanceolaria_gladiola
| | | +--------------+
| | | | +-- Lanceolaria_oxyrhyncha
| | | |
+--+ | | +-- Alasmidonta_marginata
| +-----+ +--+
| | | | +-- Lasmigona_costata
| | | +--------+
| | | | | +-- Anodonta_anatina
| | | | +--+
| | +--+ +-- Pseudanodonta_complanata
| | |
| | | +----------- Cristaria_plicata
+--+ | |
| +--+ +-- Anemina_arcaeformis
| | +-----+
| | | +-- Amuranodonta_kijaensis
| +--+
| | +----- Pletholophus_tenuis
| +--+
| | +-- Sinanodonta_jourdyi
| +--+
| +-- Beringiana_beringiana
|
| +-------------------- Aculamprotula_fibrosa
| |
| | +----- Middendorffinaia_mongolica
| | +-----------+
+--+ | | +-- Nodularia_douglasiae
| | +--+
| | +-- Nodularia_breviconcha
| |
| | +-- Cuneopsis_celtiformis
+--+ +-----------+
| | +-- Cuneopsis_heudei
| |
| | +-- Diaurora_aurorea
| | +--+
| | | +-- Diaurora_laeve
| +--------+
+--+ | +-- Schistodesmus_lampreyanus
| +--+
| +-- Schistodesmus_spinosus
|
| +-- Pseudocuneopsis_sichuanensis
| +--------+
| | +-- Pseudocuneopsis_capitata
| |
+--+ +-- Inversiunio_yanagawensis
| +-----+
| | +-- Inversiunio_reinianus
+--+
| +----- Tchangsinaia_piscicula
+--+
| +-- Unio_pictorum
+--+
+-- Unio_crassus
Graf, 2000 [Publ 2647], fig. 1
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 2 |
12 |
0 |
2 |
16 |
4 |
2 |
37.5 |
+-------- Unio
+-----------+
| | +----- Elliptio
| +--+
| | +-- Pyganodon
| +--+
--+ +-- Lampsilis
|
| +----------------- Parreysia **
| |
| | +-------------- Grandidieria **
+--+ |
| | +-------- Castalia
| | +--+
+--+ | | +----- Diplodon
| | +--+
| | | +-- Hyridella
| | +--+
+--+ +-- Velesunio
|
| +-- Mutela[1]
| +-----+
| | +-- Mutela[2]
+--+
| +----- Anodontites
| |
+--+----- Mycetopoda
|
| +-- Etheria
+--+
+-- Mulleria
Pfeiffer & Graf, 2013 [Publ 3531], fig. 2
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 2 |
2 |
0 |
2 |
33 |
30 |
3 |
-- |
+-- Lamellidens_savadiensis **
--+
+-- Coelatura_aegyptiaca **
Araujo et al., 2017 [Publ 3801], fig. 2
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 2 |
2 |
0 |
2 |
12 |
11 |
3 |
-- |
+-- Lamellidens_savadiensis **
+--------------+
| +-- Coelatura_aegyptiaca **
|
| +-- Amblema_plicata
--+ +-----------+
| | +-- Lampsilis_teres
| |
| | +-- Schistodesmus_lampreyanus
| | +--+
+--+ | +-- Aculamprotula_tortuosa
| +-----+
| | | +-- Anodonta_anatina
| | +--+
| | +-- Unio_delphinus
+--+
| +-- Gonidea_angulata
| +-----+
| | +-- Microcondylaea_bonellii
+--+
| +----- Potomida_littoralis
+--+
| +-- Lamprotula_leaii
+--+
+-- Pronodularia_japanensis
Sonowal et al., 2021 [Publ 4221], fig. 4
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 2 |
10 |
0 |
1 |
14 |
8 |
3 |
35.0 |
+-------------------------- Lamellidens_mainwaringi **
--+
| +----------------------- Lamprotula_leaii
+--+
| +-------------------- Potomida_littoralis
+--+
| +----------------- Lampsilis_cardium
+--+
| +-------------- Elliptio_complanata
+--+
| +----------- Aculamprotula_tientsinensis
+--+
| +-------- Unio_pictorum
+--+
| +----- Anodonta_cygnea
+--+
| +-- Unio_ravoisieri
+--+
+-- Lamellidens_corrianus **
Lyubas et al., 2022 [Publ 4255], fig. 7
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 2 |
2 |
0 |
1 |
29 |
29 |
2 |
-- |
+-- Parreysia_corrugata **
+-----------------------------------+
| +-- Parreysia_keralaensis **
|
| +----------------------------------- Microcondylaea_bonellii
--+ |
| | +----------- Pseudanodonta_complanata
| | +--------------------+
| | | | +-------- Anodonta_anatina
+--+ | +--+
| | | +----- Anodonta_cygnea
| | +--+
| | | +-- Anodonta_exulcerata
| | +--+
+--+ +-- Anodonta_seddoni
|
| +----------------------------- Unio_tumidus
| |
| | +-- Unio_durieui
| | +-----------------------+
| | | +-- Unio_gibbus
+--+ |
| | +-------------- Unio_damascensis
| | |
| | +-------------- Unio_sesirmensis
| | +--------+
+--+ | | +----------- Unio_ionicus
| | | |
| | +--+ +-- Unio_bruguierianus
| | | +-----+
| | | | +-- Unio_tumidiformis
| | +--+
| | | +----- Unio_crassus[2]
+--+ +--+
| | +-- Unio_carneus
| +--+
| +-- Unio_crassus[1]
|
| +----- Unio_caffer
| +--------------+
| | | +-- Unio_terminalis
| | +--+
+--+ +-- Unio_delicatus
|
| +----------------- Unio_pictorum
| |
+--+ +-------------- Unio_eucirrus
| |
+--+ +-- Unio_elongatulus
| +--------+
| | +-- Unio_mancus
+--+
| +-------- Unio_ravoisieri
| |
+--+ +-- Unio_delphinus
| +--+
| | +-- Unio_foucauldianus
+--+
| +-- Unio_hueti
+--+
+-- Unio_tigridis
Lyubas et al., 2022 [Publ 4255], fig. S1
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 2 |
2 |
0 |
1 |
29 |
29 |
2 |
-- |
+-- Parreysia_corrugata **
+-----------------------+
| +-- Parreysia_keralaensis **
|
| +----------------------- Microcondylaea_bonellii
--+ |
| | +----------- Anodonta_anatina
| | +--------+
| | | | +-------- Pseudanodonta_complanata
| | | +--+
+--+ | | +----- Anodonta_exulcerata
| | +--+
| | | +-- Anodonta_seddoni
| | +--+
| | +-- Anodonta_cygnea
| |
+--+ +----------------- Unio_tumidus
| |
| | +-- Unio_durieui
| +--------------+
| | +-- Unio_gibbus
| |
| | +----------- Unio_sesirmensis
| | |
| | +--+ +-------- Unio_damascensis
+--+ | | |
| | | | +----- Unio_ionicus
| | +--+ |
| | | | +-- Unio_bruguierianus
| | | +--+
| | +--+ +-- Unio_tumidiformis
| | |
| | | +-- Unio_carneus
| | | |
| | +--+-- Unio_crassus[1]
+--+ |
| +-- Unio_crassus[2]
|
| +-------- Unio_elongatulus
| |
| +-------- Unio_mancus
| |
| | +-- Unio_pictorum
| +-----+
| | +-- Unio_eucirrus
| |
+-----+ +----- Unio_caffer
+--+
| | +-- Unio_terminalis
| +--+
| +-- Unio_delicatus
|
| +----- Unio_ravoisieri
| |
| | +-- Unio_delphinus
+--+--+
| +-- Unio_foucauldianus
|
| +-- Unio_hueti
+--+
+-- Unio_tigridis
Graf & Cummings, 2006 [Publ 1968], fig. 2
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 1 |
1 |
0 |
1 |
46 |
24 |
3 |
-- |
+-- Pilsbryoconcha_exilis
+-----------------------------+
| +-- Pseudodon_vondembuschianus
|
| +----------------------------- Coelatura_aegyptiaca **
| |
| +----------------------------- Lens_contradens
--+ |
| | +-- Unio_pictorum
| | +--------+
| | | +-- Unio_caffer
| +-----------------+
| | | +-------- Lasmigona_costata
+--+ +--+
| | +----- Alasmidonta_marginata
| +--+
| | +-- Pyganodon_grandis
| +--+
| +-- Strophitus_undulatus
|
| +-------------------------- Gonidea_angulata
| |
| | +-- Quadrula_verrucosa
+--+ +--------------------+
| | +-- Quadrula_quadrula
| |
+--+ +-------------------- Amblema_plicata
| |
| | +----- Eurynia_dilatata
+--+ +-----------+
| | | +-- Fusconaia_flava
| | +--+
| | +-- Pleurobema_sintoxia
+--+
| +-------------- Obliquaria_reflexa
| |
| +-------------- Truncilla_truncata
+--+
+-------------- Ptychobranchus_fasciolaris
|
| +----------- Ligumia_recta
+--+
| +-------- Cambarunio_iris
+--+
| +----- Epioblasma_triquetra
+--+
| +-- Lampsilis_cardium
+--+
+-- Ortmanniana_ligamentina
Bogan & Hoeh, 2000 [Publ 2650], fig. 1
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 1 |
1 |
0 |
1 |
33 |
17 |
3 |
-- |
-- Coelatura_aegyptiaca
Hoeh et al., 2001 [Publ 2652], fig. 14.2
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 1 |
1 |
0 |
1 |
30 |
17 |
3 |
-- |
-- Coelatura_aegyptiaca
Hoeh et al., 2001 [Publ 2652], fig. 14.4
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 1 |
1 |
0 |
1 |
30 |
17 |
3 |
-- |
-- Coelatura_aegyptiaca
Hoeh et al., 2002 [Publ 2653], fig. 2
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 1 |
1 |
0 |
1 |
30 |
17 |
3 |
-- |
-- Coelatura_aegyptiaca
Roe & Hoeh, 2003 [Publ 2655], fig. 4.2
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 1 |
1 |
0 |
1 |
30 |
17 |
3 |
-- |
+----------------------------- Gonidea_angulata
|
--+ +-------------------------- Coelatura_aegyptiaca **
| |
+--+ +----------------------- Fusconaia_flava
| |
| +----------------------- Amblema_plicata
+--+
| +-------------------- Quadrula_quadrula
| |
| | +----------------- Pleurobema_clava
+--+ |
| | +-------------- Eurynia_dilatata
| | |
+--+ | +----------- Ortmanniana_ligamentina
| +--+
| | | +-------- Glebula_rotundata
| | +--+
| | | +----- Potamilus_alatus
+--+ +--+
| | +-- Ligumia_recta
| +--+
| +-- Toxolasma_lividum
|
| +----------- Cyrtonaias_tampicoensis
| |
+--+ +-- Unio_pictorum
| +-----+
| | +-- Unio_tumidus
+--+
| +----- Pyganodon_grandis
+--+
| +-- Anodonta_cygnea
+--+
+-- Strophitus_undulatus
Roe & Hoeh, 2003 [Publ 2655], fig. 4.3
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 1 |
1 |
0 |
1 |
30 |
17 |
3 |
-- |
+-------------------------- Coelatura_aegyptiaca **
|
| +-- Unio_pictorum
| +-----+
| | +-- Unio_tumidus
--+ +--------------+
| | | +----- Anodonta_cygnea
| | +--+
| | | +-- Pyganodon_grandis
| | +--+
+--+ +-- Strophitus_undulatus
|
| +-------------------- Gonidea_angulata
| |
| | +----------------- Amblema_plicata
+--+ |
| | +-------- Quadrula_quadrula
| | +-----+
+--+ | | +----- Eurynia_dilatata
| | +--+
| | | +-- Fusconaia_flava
| | +--+
+--+ +-- Pleurobema_clava
|
| +-- Cyrtonaias_tampicoensis
| +--------+
| | +-- Glebula_rotundata
+--+
| +-------- Toxolasma_lividum
+--+
| +----- Ortmanniana_ligamentina
+--+
| +-- Ligumia_recta
+--+
+-- Potamilus_alatus
Hoeh et al., 2009 [Publ 2716], fig. 3
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 1 |
1 |
0 |
1 |
46 |
24 |
3 |
-- |
-- Coelatura_aegyptiaca
Hoeh et al., 2009 [Publ 2716], fig. 4
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 1 |
1 |
0 |
1 |
45 |
23 |
3 |
-- |
-- Coelatura_aegyptiaca
Hoeh et al., 2009 [Publ 2716], fig. 5
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 1 |
1 |
0 |
1 |
45 |
23 |
3 |
-- |
-- Coelatura_aegyptiaca
Jadhav & Jamkhedkar, 2009 [Publ 3697], fig. 11
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 1 |
1 |
0 |
1 |
25 |
25 |
3 |
-- |
+-------------------------------- Lamellidens_corrianus **
|
| +-------------------- Nodularia_douglasiae
| |
| +--------+ +----------------- Anodonta_cygnea
| | | |
| | +--+ +-------------- Unio_pictorum[2]
--+ | | |
| | +--+ +-- Unio_pictorum[1]
| | | +--------+
| | | | +-- Unio_caffer
| | +--+
| | | +-------- Anodonta
| | | |
+--+ +--+ +----- Lasmigona_costata
| | |
| +--+ +-- Strophitus_undulatus
| | |
| +--+-- Alasmidonta_marginata
| |
| +-- Pyganodon_grandis
|
| +-------------------------- Pronodularia_japanensis
| |
| | +-- Quadrula_quadrula
+--+ +--------------------+
| | +-- Quadrula_verrucosa
| |
+--+ +-------------------- Gonidea_angulata
| |
| | +----- Eurynia_dilatata
+--+ +-----------+
| | | +-- Cambarunio_iris
| | +--+
| | +-- Ptychobranchus_fasciolaris
+--+
| +-- Amblema_plicata
| +-----------+
| | +-- Pleurobema_sintoxia
+--+
| +----------- Truncilla_truncata
| |
+--+ +-- Lampsilis_cardium
| +-----+
| | +-- Epioblasma_triquetra
+--+
| +----- Ligumia_recta
+--+
| +-- Obliquaria_reflexa
+--+
+-- Ortmanniana_ligamentina
Combosch et al., 2017 [Publ 3806], fig. 1
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 1 |
1 |
0 |
1 |
14 |
2 |
2 |
-- |
+----- Unio_pictorum
--+
| +-- Lampsilis_cardium
+--+
+-- Scabies_crispata **
Combosch et al., 2017 [Publ 3806], fig. 2
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 1 |
1 |
0 |
1 |
14 |
2 |
2 |
-- |
+----- Unio_pictorum
--+
| +-- Scabies_crispata **
+--+
+-- Lampsilis_cardium
Huang et al., 2018 [Publ 3828], fig. 3, 5
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 1 |
1 |
0 |
1 |
28 |
5 |
3 |
-- |
+-- Lampsilis_cardium
+--+
| +-- Scabies_crispata **
--+
| +-- Unio_pictorum
+--+
+-- Aculamprotula_tientsinensis
Jeratthitikul et al., 2019 [Publ 3911], fig. 4
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 1 |
1 |
0 |
1 |
21 |
21 |
1 |
-- |
+-- Pilsbryoconcha_exilis
+--------------------------+
| +-- Scabies_crispata **
|
--+ +-------------------------- Ensidens_ingallsianus
| |
| | +-- Physunio_modelli
+--+ +--------------------+
| | +-- Physunio_superbus
| |
| | +-- Lens_maenamensis
+--+ +-----------------+
| | +-- Lens_rolfbrandti
| |
| | +----------------- Lens_contradens[12]
+--+ |
| | +----- Lens_micropterus[3]
| | |
| +-----------+ +-- Lens_contradens[10]
| | | |
+--+ +--+-- Lens_micropterus[2]
| |
| +-- Lens_contradens[11]
|
| +-------------- Lens_contradens[9]
| |
+--+ +----------- Lens_contradens[8]
| |
+--+ +-------- Lens_contradens[7]
| |
+--+ +----- Lens_contradens[6]
| |
| | +-- Lens_contradens[1]
+--+ |
| +-- Lens_contradens[2]
| |
| +-- Lens_contradens[3]
+--+
+-- Lens_contradens[4]
|
+-- Lens_micropterus[1]
|
+-- Lens_contradens[5]
Wu et al., 2020 [Publ 4121], fig. 2
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 1 |
1 |
0 |
1 |
33 |
10 |
3 |
-- |
+-- Lampsilis_cardium
+--------------+
| +-- Scabies_crispata **
|
| +-------- Sinosolenaia_carinata
--+ +-----+
| | | +----- Potomida_littoralis
| | +--+
| | | +-- Lamprotula_caveata
+--+ +--+
| +-- Lamprotula_leaii
|
| +----------- Unio_pictorum
+--+
| +-------- Aculamprotula_tortuosa
+--+
| +----- Aculamprotula_polysticta
+--+
| +-- Aculamprotula_fibrosa
+--+
+-- Aculamprotula_tientsinensis
Wu et al., 2020 [Publ 4121], fig. S1
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 1 |
1 |
0 |
1 |
33 |
10 |
3 |
-- |
+-- Lampsilis_cardium
+--------------+
| +-- Scabies_crispata **
|
| +-------- Sinosolenaia_carinata
--+ +-----+
| | | +----- Potomida_littoralis
| | +--+
| | | +-- Lamprotula_caveata
+--+ +--+
| +-- Lamprotula_leaii
|
| +----------- Unio_pictorum
+--+
| +-------- Aculamprotula_tortuosa
+--+
| +----- Aculamprotula_polysticta
+--+
| +-- Aculamprotula_tientsinensis
+--+
+-- Aculamprotula_fibrosa
Sano et al., 2020 [Publ 4138], fig. 2
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 1 |
1 |
0 |
1 |
72 |
49 |
3 |
-- |
+----------------------------------------- Scabies_crispata **
|
| +----------------- Lampsilis_cardium
| |
| | +----- Sinohyriopsis_cumingii[2]
| +--------------------+ +--------+
| | | | | +-- Sinohyriopsis_schlegelii
| | | | +--+
| | +--+ +-- Sinohyriopsis_cumingii[1]
--+ | |
| | | +----------- Inversidens_brandtii
| | | |
| | +--+ +-- Ptychorhynchus_pfisteri
| | | +-----+
| | | | +-- Obovalis_omiensis
| | +--+
| | | +----- Lamprotula_leaii
| | +--+
+--+ | +-- Pronodularia_japanensis
| +--+
| +-- Potomida_littoralis
|
| +----- Buldowskia_iwakawai
| +-----------------------------+
| | | +-- Anemina_arcaeformis
| | +--+
| | +-- Anemina_euscaphys
| |
| | +-------------- Pletholophus_tenuis
| | |
| | | +----- Beringiana_fukuharai
| | +-----------------+ +-----+
+--+ | | | | +-- Beringiana_japonica
| | | | +--+
| | +--+ +-- Beringiana_beringiana
| | |
| | | +-------- Sinanodonta_lucida
| | +--+
| | | +----- Sinanodonta_tumens
| | +--+
| | | +-- Sinanodonta_lauta
+--+ +--+
| +-- Sinanodonta_woodiana
|
| +----------------------------- Cristaria_plicata
| |
| | +-- Anodonta_cygnea
| | +--------+
| | | +-- Anodonta_anatina
| | |
| | +--------------+ +----- Alasmidonta_varicosa
+--+ | | +--+
| | | | | +-- Prolasmidonta_heterodon
| | | | +--+
| | +--+ +-- Platynaias_compressa
| | |
| | | +----- Pyganodon_grandis
| | +--+
+--+ | +-- Utterbackia_imbecillis
| +--+
| +-- Utterbackia_peninsularis
|
| +----- Lanceolaria_grayii[1]
| +-----------------+
| | | +-- Lanceolaria_lanceolata
| | +--+
| | +-- Lanceolaria_grayii[2]
+--+
| +-------------------- Lepidodesma_languilati
| |
| | +----- Aculamprotula_tortuosa
| | +-----------+
+--+ | | +-- Aculamprotula_tientsinensis
| | +--+
| | +-- Aculamprotula_coreana
| |
+--+ +----- Unio_crassus
| +--------+
| | | +-- Unio_delphinus
| | +--+
| | +-- Unio_pictorum
| |
+--+ +-------- Inversiunio_jokohamensis
| +--+
| | | +----- Inversiunio_yanagawensis[2]
| | +--+
| | | +-- Inversiunio_reinianus
| | +--+
+--+ +-- Inversiunio_yanagawensis[1]
|
| +-- Tchangsinaia_piscicula
| +-----+
| | +-- Schistodesmus_lampreyanus
+--+
| +----- Nodularia_douglasiae[2]
+--+
| +-- Nodularia_nipponensis
+--+
+-- Nodularia_douglasiae[1]
Sano et al., 2020 [Publ 4138], fig. 3
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 1 |
1 |
0 |
1 |
71 |
48 |
3 |
-- |
+----------------------------------------- Scabies_crispata **
|
| +----- Sinohyriopsis_cumingii[2]
| +--------+
| | | +-- Sinohyriopsis_schlegelii
| | +--+
| | +-- Sinohyriopsis_cumingii[1]
| +-----------------------+
| | | +----------- Lampsilis_cardium
--+ | | |
| | | | +----- Lamprotula_leaii
| | +--+ +--+
| | | | | +-- Potomida_littoralis
| | | | +--+
| | +--+ +-- Pronodularia_japanensis
| | |
| | | +----- Inversidens_brandtii
| | +--+
| | | +-- Obovalis_omiensis
+--+ +--+
| +-- Ptychorhynchus_pfisteri
|
| +----- Buldowskia_iwakawai
| +-----------------------------+
| | | +-- Anemina_arcaeformis
| | +--+
| | +-- Anemina_euscaphys
| |
| | +-------- Pletholophus_tenuis
| | +--+
| | | | +----- Beringiana_fukuharai
| | | +--+
+--+ | | +-- Beringiana_japonica
| +--------------------+ +--+
| | | +-- Beringiana_beringiana
| | |
| | | +-------- Sinanodonta_lucida
| | +--+
| | | +----- Sinanodonta_tumens
| | +--+
| | | +-- Sinanodonta_woodiana
| | +--+
+--+ +-- Sinanodonta_lauta
|
| +----------------------------- Cristaria_plicata
| |
| | +-- Anodonta_anatina
| | +--------+
| | | +-- Anodonta_cygnea
| | |
| | +--------------+ +----- Pyganodon_grandis
+--+ | | +--+
| | | | | +-- Utterbackia_imbecillis
| | | | +--+
| | +--+ +-- Utterbackia_peninsularis
| | |
| | | +----- Alasmidonta_varicosa
| | +--+
+--+ | +-- Platynaias_compressa
| +--+
| +-- Prolasmidonta_heterodon
|
| +----- Lanceolaria_grayii[1]
| +-----------------+
| | | +-- Lanceolaria_lanceolata
| | +--+
| | +-- Lanceolaria_grayii[2]
+--+
| +-------------------- Lepidodesma_languilati
| |
| | +----- Aculamprotula_tortuosa
| | +-----------+
+--+ | | +-- Aculamprotula_tientsinensis
| | +--+
| | +-- Aculamprotula_coreana
| |
+--+ +----- Unio_crassus
| +--------+
| | | +-- Unio_delphinus
| | +--+
| | +-- Unio_pictorum
| |
+--+ +-------- Inversiunio_jokohamensis
| +--+
| | | +----- Inversiunio_yanagawensis[2]
| | +--+
| | | +-- Inversiunio_reinianus
| | +--+
+--+ +-- Inversiunio_yanagawensis[1]
|
| +-- Nodularia_nipponensis
| +--+
| | +-- Nodularia_douglasiae
+-----+
| +-- Schistodesmus_lampreyanus
+--+
+-- Tchangsinaia_piscicula
Sano et al., 2020 [Publ 4138], fig. 4
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 1 |
1 |
0 |
1 |
71 |
48 |
3 |
-- |
+-------------------------------------------- Scabies_crispata **
|
| +----------------- Lampsilis_cardium
| |
| | +----- Sinohyriopsis_cumingii[2]
| +-----------------------+ +--------+
| | | | | +-- Sinohyriopsis_schlegelii
| | | | +--+
| | +--+ +-- Sinohyriopsis_cumingii[1]
--+ | |
| | | +----------- Inversidens_brandtii
| | | |
| | +--+ +-- Obovalis_omiensis
| | | +-----+
| | | | +-- Ptychorhynchus_pfisteri
| | +--+
| | | +----- Lamprotula_leaii
| | +--+
+--+ | +-- Pronodularia_japanensis
| +--+
| +-- Potomida_littoralis
|
| +----- Buldowskia_iwakawai
| +--------------------------------+
| | | +-- Anemina_arcaeformis
| | +--+
| | +-- Anemina_euscaphys
| |
| | +-------- Sinanodonta_lucida
| | +--+
| | | | +----- Sinanodonta_tumens
| | | +--+
+--+ | | +-- Sinanodonta_woodiana
| +-----------------------+ +--+
| | | +-- Sinanodonta_lauta
| | |
| | | +-------- Pletholophus_tenuis
| | +--+
| | | +----- Beringiana_fukuharai
| | +--+
| | | +-- Beringiana_japonica
| | +--+
+--+ +-- Beringiana_beringiana
|
| +-------------------------------- Cristaria_plicata
| |
| | +-- Anodonta_cygnea
| | +--------+
| | | +-- Anodonta_anatina
| | |
| | +-----------------+ +----- Alasmidonta_varicosa
+--+ | | +--+
| | | | | +-- Prolasmidonta_heterodon
| | | | +--+
| | +--+ +-- Platynaias_compressa
| | |
| | | +----- Pyganodon_grandis
| | +--+
+--+ | +-- Utterbackia_imbecillis
| +--+
| +-- Utterbackia_peninsularis
|
| +-------------------------- Lepidodesma_languilati
| |
| | +----- Lanceolaria_grayii[1]
| | +-----------------+
+--+ | | +-- Lanceolaria_lanceolata
| | +--+
| | +-- Lanceolaria_grayii[2]
| |
+--+ +----- Aculamprotula_tortuosa
| +--------------+
| | | +-- Aculamprotula_tientsinensis
| | +--+
| | +-- Aculamprotula_coreana
| |
+--+ +----- Unio_crassus
| +-----------+
| | | +-- Unio_delphinus
| | +--+
| | +-- Unio_pictorum
+--+
| +-- Nodularia_douglasiae
| +-----------+
| | +-- Nodularia_nipponensis
| |
+--+ +-- Schistodesmus_lampreyanus
| +--------+
| | +-- Tchangsinaia_piscicula
+--+
| +-------- Inversiunio_jokohamensis
+--+
| +----- Inversiunio_yanagawensis[2]
+--+
| +-- Inversiunio_reinianus
+--+
+-- Inversiunio_yanagawensis[1]
Wu et al., 2021 [Publ 4215], fig. 3
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 1 |
1 |
0 |
1 |
23 |
23 |
2 |
-- |
+-------------------------- Indonaia_andersoniana **
|
| +-------- Acuticosta_chinensis
| +--------------+
| | | +----- Nodularia_douglasiae
--+ | +--+
| | | +-- Inversiunio_jokohamensis
| | +--+
| | +-- Inversiunio_reinianus
| |
+--+ +----- Chamberlainia_somsakpanhai
| +--------------+
| | | +-- Sundadontina_cumingii
| | +--+
| | +-- Sinohyriopsis_schlegelii
| |
| | +-- Indopseudodon_bogani
| | +--+
+--+ | +-- Indopseudodon_salwenianus
| +-----------+
| | | +-- Pseudodon_vagulus
| | +--+
| | +-- Pilsbryoconcha_carinifera
| |
| | +-------- Sinosolenaia_carinata
+--+ |
| +-----+ +-- Lamprotula_caveata
| | | +--+
| | | | +-- Lamprotula_leaii
| | +--+
| | | +-- Potomida_littoralis
| | +--+
+--+ +-- Pronodularia_japanensis
|
| +-- Ptychorhynchus_pfisteri
| +--------+
| | +-- Parvasolenaia_rivularis
| |
+--+ +-- Inversidens_brandtii
| +-----+
| | +-- Inversidens_rentianensis
+--+
| +----- Gonidea_angulata
+--+
| +-- Microcondylaea_bonellii
+--+
+-- Leguminaia_wheatleyi
Sonowal et al., 2021 [Publ 4221], fig. 3a
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 1 |
1 |
0 |
1 |
14 |
8 |
3 |
-- |
-- Lamellidens_corrianus
Sonowal et al., 2021 [Publ 4221], fig. 3b
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 1 |
1 |
0 |
1 |
14 |
8 |
3 |
-- |
+-- Aculamprotula_tientsinensis
|
+-- Lamprotula_leaii
|
+-- Lampsilis_cardium
|
+-- Potomida_littoralis
|
--+-- Unio_pictorum
|
+-- Anodonta_cygnea
|
+-- Unio_ravoisieri
|
+-- Lamellidens_mainwaringi **
|
+-- Elliptio_complanata
Pavan-Kumar et al., 2022 [Publ 4263], fig. 4
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 1 |
1 |
0 |
1 |
35 |
33 |
3 |
-- |
+----- Chamberlainia_somsakpanhai
+-----------+
| | +-- Sinohyriopsis_cumingii
| +--+
| +-- Sinohyriopsis_schlegelii
+--------+
| | +-- Lamprotula_leaii
| | +-----------+
| | | +-- Potomida_littoralis
| | |
| +--+ +-- Pseudodon_vondembuschianus
| | +--------+
| | | +-- Pilsbryoconcha_exilis
| +--+
| | +-------- Parvasolenaia_rivularis
--+ +--+
| | +----- Microcondylaea_bonellii
| +--+
| | +-- Sinosolenaia_carinata
| +--+
| +-- Sinosolenaia_oleivora
|
| +----------------------- Lamellidens_marginalis **
| |
| | +-------------------- Lepidodesma_languilati
+--+ |
| | +-- Aculamprotula_tientsinensis
| | +--------+
| | | +-- Aculamprotula_tortuosa
+--+ +-----+
| | | +-------- Unio_pictorum
| | +--+
| | | +----- Schistodesmus_lampreyanus
| | +--+
| | | +-- Nodularia_douglasiae
+--+ +--+
| +-- Tchangsinaia_piscicula
|
| +-- Lanceolaria_lanceolata
| +-----------+
| | +-- Lanceolaria_gladiola
+--+
| +----------- Acuticosta_chinensis
| |
+--+ +-- Anodonta_anatina
| +-----+
| | +-- Pyganodon_grandis
+--+
| +----- Cristaria_plicata
+--+
| +-- Sinanodonta_lucida
+--+
+-- Anemina_euscaphys
Pavan-Kumar et al., 2022 [Publ 4263], fig. S5
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 1 |
1 |
0 |
1 |
38 |
36 |
3 |
-- |
+-------------------- Lamellidens_marginalis **
|
| +-- Anodonta_anatina
| +--------+
| | +-- Pyganodon_grandis
| +-----+
+--+ | | +-------- Cristaria_plicata
| | | | |
| | | +--+ +-- Anemina_arcaeformis
| | | | +--+
| | | | | +-- Anemina_euscaphys
| | | +--+
| | | | +-- Sinanodonta_lucida
| +--+ +--+
| | +-- Sinanodonta_woodiana
| |
| | +----- Acuticosta_chinensis
| | +--------+
| | | | +-- Lanceolaria_lanceolata
| | | +--+
| | | +-- Lanceolaria_gladiola
| +--+
| | +----- Lepidodesma_languilati
| | +-----+
| | | | +-- Aculamprotula_tientsinensis
| | | +--+
--+ +--+ +-- Aculamprotula_tortuosa
| |
| | +-------- Unio_pictorum
| +--+
| | +----- Schistodesmus_lampreyanus
| +--+
| | +-- Nodularia_douglasiae
| +--+
| +-- Tchangsinaia_piscicula
|
| +-------------- Quadrula_quadrula
| |
| +--+ +----------- Toxolasma_parvum
| | | |
| | +--+ +-------- Potamilus_leptodon
| | | |
| | +--+ +-- Lampsilis_cardium
| | | +--+
| | | | +-- Lampsilis_ornata
| | +--+
| | | +-- Lampsilis_siliquoidea
+-----+ +--+
| +-- Venustaconcha_ellipsiformis
|
| +----- Chamberlainia_somsakpanhai
| +--------+
| | | +-- Sinohyriopsis_schlegelii
| | +--+
| | +-- Sinohyriopsis_cumingii
| |
| | +-- Sinosolenaia_carinata
+--+ +--+
| | +-- Sinosolenaia_oleivora
| +-----+
| | | +-- Parvasolenaia_rivularis
| | +--+
| | +-- Microcondylaea_bonellii
+--+
| +-- Pseudodon_vondembuschianus
| +-----+
| | +-- Pilsbryoconcha_exilis
+--+
| +----- Potomida_littoralis
+--+
| +-- Lamprotula_leaii
+--+
+-- Pronodularia_japanensis
Smith, 2021 [Publ 4331], fig. 1
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 1 |
1 |
0 |
1 |
19 |
15 |
3 |
-- |
+----- Sinohyriopsis_cumingii
+-----+
| | +-- Lens_contradens
| +--+
--+ +-- Lamprotula_leaii
|
| +-------- Scabies_phaselus **
+--+
| +----- Cristaria_plicata
+--+
| +-- Pyganodon_grandis
+--+
+-- Utterbackia_peninsularis
Wu et al., 2023 [Publ 4339], fig. 4, S5-aa
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 1 |
1 |
0 |
1 |
95 |
84 |
3 |
-- |
+----- Chamberlainia_somsakpanhai
+-----------------+
| | +-- Sinohyriopsis_cumingii
| +--+
| +-- Sinohyriopsis_schlegelii
|
| +-- Hyriopsis_bialata
| +--+
+-----------+ | +-- Rectidens_sumatrensis
| | +--------------+
| | | | +-- Lens_contradens
| | | +--+
| | | +-- Physunio_superbus
| | |
| | | +----------- Potomida_littoralis
| +--+ +-----+
| | | | +-------- Pronodularia_japanensis
| | | +--+
| | | | +----- Lamprotula_leaii
| | | +--+
| | | | +-- Lamprotula_caveata
| +--+ +--+
| | +-- Lamprotula_cornuumlunae
| |
| | +-- Pseudodon_vondembuschianus
| | +-----------+
| | | +-- Pilsbryoconcha_exilis
| +--+
| | +----------- Microcondylaea_bonellii
--+ | |
| +--+ +-- Parvasolenaia_rivularis
| | +-----+
| | | +-- Ptychorhynchus_pfisteri
| +--+
| | +----- Sinosolenaia_carinata
| +--+
| | +-- Sinosolenaia_iridinea
| +--+
| +-- Sinosolenaia_oleivora
|
| +-------------------------------- Lamellidens_marginalis **
| |
| | +-- Lepidodesma_aligera
| | +--------------------------+
| | | +-- Lepidodesma_languilati
| | |
| | | +-------- Aculamprotula_tortuosa
| | | |
| | | +--------------+ +-- Aculamprotula_coreana
| | | | | +--+
+--+ | | | | +-- Aculamprotula_tientsinensis
| | | +--+
| | | | +-- Aculamprotula_polysticta
| | | +--+
| | | +-- Aculamprotula_scripta
| | |
| | +--+ +-------------- Unio_tumidus
| | | | +-----+
| | | | | | +----------- Unio_crassus
| | | | | +--+
+--+ | | | | +-------- Unio_elongatulus
| | | | +--+
| | | | | +----- Unio_pictorum
| | | | +--+
| | | | | +-- Unio_delphinus
| | +--+ +--+
| | | +-- Unio_mancus
| | |
| | | +-- Nodularia_breviconcha
| | | +--+
| | | | +-- Nodularia_nipponensis
| | | +-----------+
| | | | | +-- Pseudocuneopsis_capitata
| | | | +--+
| | +--+ +-- Pseudocuneopsis_sichuanensis
| | |
| | | +-------------- Tchangsinaia_piscicula
| | | |
+--+ +--+ +-- Schistodesmus_lampreyanus
| | +--------+
| | | +-- Schistodesmus_spinosus
| +--+
| | +-------- Cuneopsis_rufescens
| +--+
| | +----- Cuneopsis_celtiformis
| +--+
| | +-- Cuneopsis_demangei
| +--+
| +-- Cuneopsis_heudei
|
| +-------- Acuticosta_chinensis
| +--------+
| | | +----- Lanceolaria_gladiola
| | +--+
| | | +-- Lanceolaria_grayii
| | +--+
| | +-- Lanceolaria_lanceolata
| |
| | +----- Cristaria_plicata
| | +-----+
| | | | +-- Anemina_arcaeformis
+--------+ | +--+
| +--+ +-- Anemina_euscaphys
| | |
| | | +-------- Beringiana_fukuharai
| | +--+
| | | +----- Sinanodonta_lucida
| | +--+
| | | +-- Sinanodonta_tumens
| | +--+
| | +-- Sinanodonta_woodiana
+--+
| +-- Anodonta_nuttalliana
| +-----+
| | +-- Pseudanodonta_complanata
| +--+
| | | +----- Anodonta_anatina
| | +--+
| | | +-- Anodonta_cygnea
| | +--+
| | +-- Anodonta_exulcerata
+--+
| +----- Pyganodon_grandis
| +--+
| | | +-- Utterbackia_peninsularis
| | +--+
| | +-- Utterbackia_imbecillis
+--+
| +-- Alasmidonta_varicosa
| +--+
| | +-- Platynaias_compressa
+--+
| +-- Prolasmidonta_heterodon
+--+
+-- Lasmigona_complanata
Wu et al., 2023 [Publ 4339], fig. 4, S5-nt
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 1 |
1 |
0 |
1 |
96 |
85 |
3 |
-- |
+----- Chamberlainia_somsakpanhai
+-----------------+
| | +-- Sinohyriopsis_cumingii
| +--+
| +-- Sinohyriopsis_schlegelii
|
| +-- Hyriopsis_bialata
| +--+
+--------------+ | +-- Rectidens_sumatrensis
| | +--------------+
| | | | +-- Lens_contradens
| | | +--+
| | | +-- Physunio_superbus
| | |
| | | +----------- Potomida_littoralis
| +--+ +-----+
| | | | +-------- Pronodularia_japanensis
| | | +--+
| | | | +----- Lamprotula_leaii
| | | +--+
| | | | +-- Lamprotula_caveata
| +--+ +--+
| | +-- Lamprotula_cornuumlunae
| |
| | +-- Pseudodon_vondembuschianus
| | +-----------+
| | | +-- Pilsbryoconcha_exilis
| | |
| +--+ +-- Ptychorhynchus_pfisteri
--+ | +--------+
| | | +-- Microcondylaea_bonellii
| +--+
| | +-------- Parvasolenaia_rivularis
| +--+
| | +----- Sinosolenaia_carinata
| +--+
| | +-- Sinosolenaia_iridinea
| +--+
| +-- Sinosolenaia_oleivora
|
| +----------------------------------- Lamellidens_marginalis **
| |
| | +-- Lepidodesma_aligera
| | +-----------------------------+
| | | +-- Lepidodesma_languilati
| | |
| | | +-------- Acuticosta_chinensis
| | | +--------------+
| | | | | +----- Lanceolaria_gladiola
| | | | +--+
+--+ | | | +-- Lanceolaria_grayii
| | | +--+
| | | +-- Lanceolaria_lanceolata
| | |
| | | +-------------- Cristaria_plicata
| | +-----+ |
| | | | +-----+ +-- Anemina_arcaeformis
| | | | | | +--------+
| | | | | | | +-- Anemina_euscaphys
| | | | | +--+
+--+ | | | | +-------- Beringiana_fukuharai
| | | | +--+
| | | | | +----- Sinanodonta_lucida
| | | | +--+
| | +--+ | +-- Sinanodonta_tumens
| | | +--+
| | | +-- Sinanodonta_woodiana
| | |
| | | +----------------- Anodonta_nuttalliana
| | | |
| | | +----------------- Pseudanodonta_complanata
| | | |
| | +--+ +-------------- Anodonta_cygnea
| | | |
| | | | +-- Anodonta_exulcerata
| | | +-----------+
| | | | +-- Anodonta_anatina
| | +--+
+--+ | +----- Pyganodon_grandis
| | +-----+
| | | | +-- Utterbackia_imbecillis
| | | +--+
| +--+ +-- Utterbackia_peninsularis
| |
| | +-------- Lasmigona_complanata
| +--+
| | +----- Prolasmidonta_heterodon
| +--+
| | +-- Alasmidonta_varicosa
| +--+
| +-- Platynaias_compressa
|
| +-------- Aculamprotula_tortuosa
| |
| +-----------------+ +-- Aculamprotula_coreana
| | | +--+
| | | | +-- Aculamprotula_tientsinensis
| | +--+
| | | +-- Aculamprotula_polysticta
| | +--+
| | +-- Aculamprotula_scripta
| |
+--+ +----------- Unio_tumidus
| |
| +-----------+ +-------- Unio_crassus
| | | |
| | +--+ +----- Unio_elongatulus
| | | |
| | +--+ +-- Unio_delphinus
| | | |
| | +--+-- Unio_mancus
+--+ |
| +-- Unio_pictorum
|
| +-- Pseudocuneopsis_capitata
| +-----------------+
| | +-- Pseudocuneopsis_sichuanensis
| |
+--+ +-- Nodularia_breviconcha
| +--------------+
| | +-- Nodularia_nipponensis
| |
+--+ +----- Tchangsinaia_piscicula
| +--------+
| | | +-- Schistodesmus_lampreyanus[2]
| | +--+
+--+ +-- Schistodesmus_spinosus
|
| +----------- Schistodesmus_lampreyanus[1]
+--+
| +-------- Cuneopsis_rufescens
+--+
| +----- Cuneopsis_celtiformis
+--+
| +-- Cuneopsis_demangei
+--+
+-- Cuneopsis_heudei
Wu et al., 2024 [Publ 4350], fig. 8a, S1
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 1 |
1 |
0 |
1 |
91 |
85 |
3 |
-- |
+----- Chamberlainia_somsakpanhai
+--------------------+
| | +-- Sinohyriopsis_cumingii
| +--+
| +-- Sinohyriopsis_schlegelii
|
| +-- Hyriopsis_bialata
| +--+
+--------+ | +-- Rectidens_sumatrensis
| | +-----------------+
| | | | +-- Lens_contradens
| | | +--+
| | | +-- Physunio_superbus
| | |
| | | +----------- Potomida_littoralis
| +--+ +--------+
| | | | +-------- Pronodularia_japanensis
| | | +--+
| | | | +----- Lamprotula_leaii
| | | +--+
| | | | +-- Lamprotula_caveata
| +--+ +--+
| | +-- Lamprotula_cornuumlunae
| |
| | +-- Pseudodon_vondembuschianus
| | +--------------+
| | | +-- Pilsbryoconcha_exilis
| +--+
| | +-------------- Inversidens_rentianensis
| | |
--+ +--+ +-- Parvasolenaia_rivularis
| | +--------+
| | | +-- Ptychorhynchus_pfisteri
| +--+
| | +-------- Microcondylaea_bonellii
| +--+
| | +----- Sinosolenaia_carinata
| +--+
| | +-- Sinosolenaia_iridinea
| +--+
| +-- Sinosolenaia_oleivora
|
| +-------------------------------- Lamellidens_marginalis **
| |
| | +-- Lepidodesma_aligera
| | +--------------------------+
| | | +-- Lepidodesma_languilati
| | |
| | | +-------- Aculamprotula_tortuosa
| | | |
| | | +--------------+ +-- Aculamprotula_coreana
| | | | | +--+
+--+ | | | | +-- Aculamprotula_tientsinensis
| | | +--+
| | | | +-- Aculamprotula_polysticta
| | | +--+
| | | +-- Aculamprotula_scripta
| | |
| | +--+ +-------------- Unio_tumidus
| | | | +-----+
| | | | | | +----------- Unio_crassus
| | | | | +--+
+--+ | | | | +-------- Unio_elongatulus
| | | | +--+
| | | | | +----- Unio_pictorum
| | | | +--+
| | | | | +-- Unio_delphinus
| | +--+ +--+
| | | +-- Unio_mancus
| | |
| | | +-- Nodularia_breviconcha
| | | +--------------+
| | | | +-- Nodularia_douglasiae
| | | |
| | | | +-- Pseudocuneopsis_capitata
| | | +--------------+
| | +--+ +-- Pseudocuneopsis_sichuanensis
| | |
| | | +-- Schistodesmus_lampreyanus
| | | +-----------+
+--+ | | +-- Schistodesmus_spinosus
| +--+
| | +----------- Tchangsinaia_piscicula
| +--+
| | +-------- Cuneopsis_rufescens
| +--+
| | +----- Cuneopsis_celtiformis
| +--+
| | +-- Cuneopsis_demangei
| +--+
| +-- Cuneopsis_heudei
|
| +-------- Acuticosta_chinensis
| +--------------+
| | | +----- Lanceolaria_gladiola
| | +--+
| | | +-- Lanceolaria_grayii
| | +--+
| | +-- Lanceolaria_lanceolata
| |
| | +-------------- Cristaria_plicata
| | |
+--+ +-----+ +-- Anemina_arcaeformis
| | | +--------+
| | | | +-- Anemina_euscaphys
| | +--+
| | | +-------- Beringiana_fukuharai
| | +--+
| | | +----- Sinanodonta_lucida
| | +--+
| | | +-- Sinanodonta_tumens
+--+ +--+
| +-- Sinanodonta_woodiana
|
| +-------- Pseudanodonta_complanata
| +--------+
| | | +----- Anodonta_anatina
| | +--+
| | | +-- Anodonta_cygnea
| | +--+
+--+ +-- Anodonta_exulcerata
|
| +-------------- Anodonta_nuttalliana
| |
| | +----- Pyganodon_grandis
+--+ +-----+
| | | +-- Utterbackia_imbecillis
| | +--+
+--+ +-- Utterbackia_peninsularis
|
| +-------- Lasmigona_complanata
+--+
| +----- Prolasmidonta_heterodon
+--+
| +-- Alasmidonta_varicosa
+--+
+-- Platynaias_compressa
Wu et al., 2024 [Publ 4350], fig. 8b, S2
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 1 |
1 |
0 |
1 |
91 |
85 |
3 |
-- |
+----- Chamberlainia_somsakpanhai
+--------------------+
| | +-- Sinohyriopsis_cumingii
| +--+
| +-- Sinohyriopsis_schlegelii
|
| +-- Hyriopsis_bialata
| +--+
+--------+ | +-- Rectidens_sumatrensis
| | +-----------------+
| | | | +-- Lens_contradens
| | | +--+
| | | +-- Physunio_superbus
| | |
| | | +----------- Potomida_littoralis
| +--+ +--------+
| | | | +-------- Pronodularia_japanensis
| | | +--+
| | | | +----- Lamprotula_leaii
| | | +--+
| | | | +-- Lamprotula_caveata
| +--+ +--+
| | +-- Lamprotula_cornuumlunae
| |
| | +-- Pseudodon_vondembuschianus
| | +--------------+
| | | +-- Pilsbryoconcha_exilis
| +--+
| | +-------------- Inversidens_rentianensis
| | |
--+ +--+ +-- Parvasolenaia_rivularis
| | +--------+
| | | +-- Ptychorhynchus_pfisteri
| +--+
| | +-------- Microcondylaea_bonellii
| +--+
| | +----- Sinosolenaia_carinata
| +--+
| | +-- Sinosolenaia_iridinea
| +--+
| +-- Sinosolenaia_oleivora
|
| +-------------------------------- Lamellidens_marginalis **
| |
| | +-- Lepidodesma_aligera
| | +--------------------------+
| | | +-- Lepidodesma_languilati
| | |
| | | +-------- Aculamprotula_tortuosa
| | | |
| | | +--------------+ +-- Aculamprotula_coreana
| | | | | +--+
+--+ | | | | +-- Aculamprotula_tientsinensis
| | | +--+
| | | | +-- Aculamprotula_polysticta
| | | +--+
| | | +-- Aculamprotula_scripta
| | |
| | +--+ +-------------- Unio_tumidus
| | | | +-----+
| | | | | | +----------- Unio_crassus
| | | | | +--+
+--+ | | | | +-------- Unio_elongatulus
| | | | +--+
| | | | | +----- Unio_pictorum
| | | | +--+
| | | | | +-- Unio_delphinus
| | +--+ +--+
| | | +-- Unio_mancus
| | |
| | | +-- Nodularia_breviconcha
| | | +--------------+
| | | | +-- Nodularia_douglasiae
| | | |
| | | | +-- Pseudocuneopsis_capitata
| | | +--------------+
| | +--+ +-- Pseudocuneopsis_sichuanensis
| | |
| | | +-- Schistodesmus_lampreyanus
+--+ | +-----------+
| | | +-- Schistodesmus_spinosus
| +--+
| | +----------- Tchangsinaia_piscicula
| +--+
| | +-------- Cuneopsis_rufescens
| +--+
| | +----- Cuneopsis_celtiformis
| +--+
| | +-- Cuneopsis_demangei
| +--+
| +-- Cuneopsis_heudei
|
| +-------- Acuticosta_chinensis
| +--------------+
| | | +----- Lanceolaria_gladiola
| | +--+
| | | +-- Lanceolaria_grayii
| | +--+
| | +-- Lanceolaria_lanceolata
| |
| | +-------------- Cristaria_plicata
+--+ |
| +-----+ +-- Anemina_arcaeformis
| | | +--------+
| | | | +-- Anemina_euscaphys
| | +--+
| | | +-------- Beringiana_fukuharai
| | +--+
| | | +----- Sinanodonta_lucida
| | +--+
+--+ | +-- Sinanodonta_tumens
| +--+
| +-- Sinanodonta_woodiana
|
| +----------------- Anodonta_nuttalliana
| |
| | +-------- Pseudanodonta_complanata
| | +-----+
+--+ | | +----- Anodonta_anatina
| | +--+
| | | +-- Anodonta_cygnea
| | +--+
+--+ +-- Anodonta_exulcerata
|
| +----- Pyganodon_grandis
| +-----+
| | | +-- Utterbackia_imbecillis
| | +--+
+--+ +-- Utterbackia_peninsularis
|
| +-------- Lasmigona_complanata
+--+
| +----- Prolasmidonta_heterodon
+--+
| +-- Alasmidonta_varicosa
+--+
+-- Platynaias_compressa
| 
