Bivalvia | Unionoida | Unionoidea | Unionidae subfamily PARREYSIINAE Henderson, 1935: cladomics 5 tribes
Cladomics Summary
number of species = 114
number of subtaxa (rank: tribe) = 5
analyzed in 87 cladograms in 53 publications
maximum number of terminals in an analysis = 62, mean = 12.76
maximum number of subtaxa (tribe) in an analysis = 5, mean = 2.55
Basic Monophyly
number of cladograms that test basic monophyly (i.e., more than 1 included species) = 56
maximum number of included terminals = 62, mean = 19.27
maximum number of subtaxa (tribe) = 5, mean = 3.41
proportion of cladograms that support basic monophyly = 0.96
Rigorous Monophyly
number of cladograms that test rigorous monophyly = 40
(Rigorous monophyly refers to cladograms with terminal species representing more than one subtaxon (tribe) taxon and at least one rigorous outgroup (subfamily).
maximum number of included terminals = 62, mean = 20.35
maximum number of subtaxa (tribe) = 5, mean = 3.85
proportion of cladograms that support rigorous monophyly = 0.97
Pairwise Cladogram Clustering
The graphs below depict in two dimensions all pair-wise ingroup distances within the cladome. See the Cladmoics FARQ for more information.
Bolotov et al., 2017 [Publ 3761], fig. S1
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
29 |
29 |
27 |
5 |
110 |
85 |
3 |
0.0 |
*** Top Monophyly Cladogram ***
+----------- Lamellidentini **
+--------+
| | +-------- Parreysiini **
| +--+
| | +----- Leoparreysiini **
| +--+
| | +-- Coelaturini **
--+ +--+
| +-- Indochinellini **
|
| +-- Rectidentini
| +--------------+
| | +-- Contradentini
+--+
| +-------------- Pseudodontini
+--+
| +----------- Unioninae
+--+
| +-------- Ambleminae
+--+
| +----- Gonideini
+--+
| +-- Chamberlainiini
+--+
+-- Lamprotulini
Huang et al., 2019 [Publ 3843], fig. 2
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
34 |
34 |
30 |
5 |
164 |
125 |
3 |
18.0 |
*** Top Sister-Group Cladogram ***
+----------- Lamellidentini **
+--------+
| | +-------- Parreysiini **
| +--+
| | +----- Leoparreysiini **
| +--+
--+ | +-- Coelaturini **
| +--+
| +-- Indochinellini **
|
| +----------------- Unioninae
| |
+--+ +-- Rectidentini
| +-----------+
| | +-- Contradentini
+--+
| +----------- Ambleminae
+--+
| +-------- Chamberlainiini
+--+
| +----- Lamprotulini
+--+
| +-- Gonideini
+--+
+-- Pseudodontini
Bolotov et al., 2024 [Publ 4369], fig. 2
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
62 |
62 |
60 |
5 |
0 |
0 |
0 |
53.0 |
+-------- Lamellidentini **
|
--+ +-- Leoparreysiini **
| +--+
| | +-- Parreysiini **
+--+
| +-- Coelaturini **
+--+
+-- Indochinellini **
Bolotov et al., 2024 [Publ 4369], fig. S1
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
62 |
62 |
60 |
5 |
1 |
1 |
1 |
51.0 |
+-------------- Gonideinae
|
--+ +----------- Parreysia **
| |
+--+ +-- Leoparreysiini **
| +-----+
| | +-- Lamellidentini **
+--+
| +----- Assamnaia **
+--+
| +-- Coelaturini **
+--+
+-- Indochinellini **
Bolotov et al., 2022 [Publ 4252], fig. 2
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
57 |
57 |
53 |
5 |
4 |
4 |
1 |
49.0 |
+-------------- Gonideinae
--+
| +----------- Lamellidentini **
+--+
| +-------- Parreysiini **
+--+
| +----- Leoparreysiini **
+--+
| +-- Coelaturini **
+--+
+-- Indochinellini **
Bolotov et al., 2022 [Publ 4252], fig. S1
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
57 |
57 |
47 |
5 |
4 |
4 |
1 |
46.0 |
+-------------- Gonideinae
--+
| +----------- Parreysiini **
+--+
| +-------- Lamellidentini **
+--+
| +----- Leoparreysiini **
+--+
| +-- Coelaturini **
+--+
+-- Indochinellini **
Bolotov et al., 2022 [Publ 4239], fig. S1
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
54 |
54 |
50 |
5 |
4 |
4 |
1 |
43.0 |
+-------------- Gonideinae
--+
| +----------- Lamellidentini **
+--+
| +-------- Parreysiini **
+--+
| +----- Leoparreysiini **
+--+
| +-- Coelaturini **
+--+
+-- Indochinellini **
Bolotov et al., 2022 [Publ 4239], fig. S2
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
54 |
54 |
52 |
5 |
4 |
4 |
1 |
43.0 |
+-------------- Gonideinae
--+
| +----------- Parreysiini **
+--+
| +-------- Lamellidentini **
+--+
| +----- Leoparreysiini **
+--+
| +-- Coelaturini **
+--+
+-- Indochinellini **
Bolotov et al., 2022 [Publ 4239], fig. 2
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
54 |
54 |
52 |
5 |
4 |
4 |
1 |
39.0 |
+----------- Gonideinae
|
--+ +-------- Lamellidentini **
| |
+--+ +-- Parreysiini **
| +--+
| | +-- Leoparreysiini **
+--+
| +-- Coelaturini **
+--+
+-- Indochinellini **
Bolotov et al., 2020 [Publ 4100], fig. 1
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
41 |
41 |
39 |
5 |
77 |
72 |
1 |
36.0 |
+----------- Gonideinae
|
--+ +-------- Lamellidentini **
| |
+--+ +-- Parreysiini **
| +--+
| | +-- Leoparreysiini **
+--+
| +-- Coelaturini **
+--+
+-- Indochinellini **
Bolotov et al., 2020 [Publ 4100], fig. S1
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
40 |
40 |
37 |
5 |
72 |
72 |
1 |
37.0 |
+----------- Gonideinae
|
--+ +-------- Lamellidentini **
| |
+--+ +-- Parreysiini **
| +--+
| | +-- Leoparreysiini **
+--+
| +-- Coelaturini **
+--+
+-- Indochinellini **
Bolotov et al., 2017 [Publ 3764], fig. S1
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
38 |
38 |
28 |
5 |
87 |
87 |
3 |
39.0 |
+-------------- Pseudodontini
|
| +-- Rectidentini
+-----------+ +--------+
| | | +-- Contradentini
| | |
| +--+ +-------- Unioninae
| | |
| | | +----- Ambleminae
| | | |
| +--+ | +-- Chamberlainia
| | | |
| | | +-- Sinohyriopsis
| | | |
| | | +-- Lamprotula_caveata
| +--+ |
| | +-- Lamprotula_leaii
| | |
| | +-- Potomida
| | |
| | +-- Pronodularia
| | |
| +--+-- Gonidea
| |
| +-- Leguminaia_wheatleyi
--+ |
| +-- Leguminaia
| |
| +-- Microcondylaea
| |
| +-- Sinosolenaia_carinata
| |
| +-- Sinosolenaia_oleivora
| |
| +-- Sinosolenaia_recognita
|
| +-------------- Trapezidens **
| |
| +--------+ +----------- Lamellidens_lamellatus **
| | | |
| | +--+ +-------- Lamellidens_jenkinsianus **
| | | |
| | | | +----- Lamellidens_generosus **
| | +--+ |
| | | +----- Lamellidens_mainwaringi **
| | | |
| | +--+ +-- Lamellidens_ferrugineus **
+--+ +--+
| | +-- Lamellidens_marginalis **
| |
| | +-- Lamellidens_savadiensis **
| +--+
| +-- Lamellidens_corrianus__1 **
|
| +-------------------- Parreysiini **
| |
+--+ +----------------- Leoparreysiini **
| |
+--+ +-------------- Coelaturini **
| |
| | +----- Indochinella **
+--+ +-----+
| | | +-- Scabies **
| | +--+
+--+ +-- Radiatula **
|
| +-------- Indonaia_shurtleffiana **
| |
+--+ +----- Indonaia_rugosa **
| |
| | +-- Indonaia_andersoniana **
+--+--+
| +-- Indonaia_caerulea **
|
| +-- Indonaia_bonneaudii **
+--+
+-- Lamellidens_corrianus__2 **
Ortiz-Sepulveda et al., 2020 [Publ 4101], fig. 1
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
36 |
36 |
22 |
3 |
0 |
0 |
0 |
44.0 |
+----- Leoparreysiini **
--+
| +-- Indochinellini **
+--+
+-- Coelaturini **
Ortiz-Sepulveda et al., 2020 [Publ 4101], fig. 2
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
36 |
36 |
20 |
3 |
0 |
0 |
0 |
46.0 |
+----- Leoparreysiini **
--+
| +-- Indochinellini **
+--+
+-- Coelaturini **
Huang et al., 2019 [Publ 3843], fig. S3
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
34 |
34 |
30 |
5 |
162 |
123 |
3 |
21.0 |
+----------- Lamellidentini **
+--------+
| | +-------- Parreysiini **
| +--+
| | +----- Leoparreysiini **
| +--+
--+ | +-- Coelaturini **
| +--+
| +-- Indochinellini **
|
| +----------------- Pseudodontini
| |
+--+ +-- Rectidentini
| +-----------+
| | +-- Contradentini
+--+
| +----------- Unioninae
+--+
| +-------- Chamberlainiini
+--+
| +----- Ambleminae
+--+
| +-- Lamprotulini
+--+
+-- Gonideini
Huang et al., 2019 [Publ 3843], fig. S4
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
34 |
34 |
25 |
5 |
164 |
125 |
3 |
17.0 |
+-------- Parreysiini **
|
+-----+-------- Lamellidentini **
| |
| | +----- Leoparreysiini **
| +--+
| | +-- Coelaturini **
| +--+
| +-- Indochinellini **
|
| +----------- Chamberlainia
| |
| +----------- Megalonaias
--+ |
| +----------- Uniomerus
| |
| +----------- Quadrulina
| |
| +----------- Sinohyriopsis
| |
| +----------- Lamprotulini
| |
| +----------- Gonideini
| |
| +----------- Pseudodontini
+--+
+----------- Unioninae
|
| +-- Rectidentini
+--------+
| +-- Contradentini
|
| +-------- Amblema
| |
| +-------- Toxolasma
| |
| +-------- Pleurobemini
| |
| | +-- Obliquaria
| +-----+
| | +-- Plectomerus
+--+
| +----- Ellipsaria
| |
| +----- Potamilus_leptodon
| |
| +----- Medionidus
| |
| +----- Potamilus_alatus
| |
| +----- Ptychobranchus
+--+
+----- Truncilla
|
+----- Venustaconcha
|
+----- Cambarunio
|
+----- Leaunio
|
| +-- Ortmanniana
+--+
+-- Lampsilis
Bolotov et al., 2017 [Publ 3761], fig. S2
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
29 |
29 |
26 |
5 |
110 |
85 |
3 |
6.0 |
+-- Parreysiini **
+-----+
| +-- Lamellidentini **
+-----------+
| | +----- Leoparreysiini **
| +--+
| | +-- Coelaturini **
--+ +--+
| +-- Indochinellini **
|
| +----------------- Unioninae
| |
+--+ +-- Contradentini
| +-----------+
| | +-- Rectidentini
+--+
| +----------- Ambleminae
+--+
| +-------- Pseudodontini
+--+
| +----- Gonideini
+--+
| +-- Chamberlainiini
+--+
+-- Lamprotulini
Pfeiffer et al., 2018 [Publ 3827], fig. 2
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
27 |
27 |
25 |
5 |
1 |
0 |
0 |
28.0 |
+----------- Lamellidentini **
--+
| +-------- Parreysiini **
+--+
| +----- Leoparreysiini **
+--+
| +-- Coelaturini **
+--+
+-- Indochinellini **
Bolotov et al., 2022 [Publ 4250], fig. 8
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
24 |
24 |
22 |
4 |
25 |
23 |
1 |
31.0 |
+----------- Gonideinae
--+
| +-------- Lamellidentini **
+--+
| +----- Parreysiini **
+--+
| +-- Leoparreysiini **
+--+
+-- Indochinellini **
Bolotov et al., 2018 [Publ 3826], fig. 2
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
24 |
24 |
22 |
5 |
59 |
59 |
3 |
16.0 |
+----------- Lamellidentini **
+--+
| | +-------- Parreysiini **
| +--+
| | +----- Leoparreysiini **
| +--+
| | +-- Coelaturini **
--+ +--+
| +-- Indochinellini **
|
| +----------- Pseudodontini
| |
| | +-- Rectidentini
| | +-----+
+--+ | +-- Contradentini
| |
| | +----- Unioninae
| | |
+--+ | +-- Lamprotula
| | |
| | +-- Potomida
| | |
| | +-- Pronodularia
+--+ |
| +-- Gonidea
| |
| +-- Leguminaia
| |
| +-- Microcondylaea
| |
+--+-- Amblemini
|
+-- Ortmanniana
|
+-- Lampsilis
|
+-- Cambarunio
|
+-- Elliptio
|
+-- Pleurobema
|
+-- Quadrulini
Jeratthitikul et al., 2019 [Publ 3922], fig. 1
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
23 |
23 |
21 |
5 |
0 |
0 |
0 |
33.0 |
+----------- Lamellidentini **
--+
| +-------- Leoparreysiini **
+--+
| +----- Parreysiini **
+--+
| +-- Coelaturini **
+--+
+-- Indochinellini **
Bolotov et al., 2022 [Publ 4250], fig. S2
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
15 |
15 |
13 |
3 |
0 |
0 |
0 |
32.0 |
+----- Indochinellini **
--+
| +-- Parreysiini **
+--+
+-- Leoparreysiini **
Konopleva et al., 2023 [Publ 4262], fig. 8
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
14 |
14 |
10 |
1 |
68 |
68 |
1 |
38.0 |
+-- Indochinellini **
--+
+-- Gonideinae
Wu et al., 2021 [Publ 4366], fig. 2_I
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
13 |
13 |
11 |
4 |
78 |
61 |
3 |
23.0 |
+-------- Lamellidentini **
+-----------+
| | +----- Leoparreysiini **
| +--+
| | +-- Coelaturini **
--+ +--+
| +-- Indochinellini **
|
| +----------------- Unioninae
| |
+--+ +-- Rectidentini
| +-----------+
| | +-- Contradentini
+--+
| +----------- Ambleminae
+--+
| +-------- Chamberlainiini
+--+
| +----- Lamprotulini
+--+
| +-- Pseudodontini
+--+
+-- Gonideini
Wu et al., 2021 [Publ 4366], fig. 6
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
13 |
13 |
11 |
4 |
78 |
61 |
3 |
25.0 |
+-------- Lamellidentini **
+-----------+
| | +----- Leoparreysiini **
| +--+
| | +-- Coelaturini **
--+ +--+
| +-- Indochinellini **
|
| +----------------- Unioninae
| |
+--+ +-- Contradentini
| +-----------+
| | +-- Rectidentini
+--+
| +----------- Ambleminae
+--+
| +-------- Chamberlainiini
+--+
| +----- Lamprotulini
+--+
| +-- Pseudodontini
+--+
+-- Gonideini
Bolotov et al., 2019 [Publ 3974], fig. 1
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
13 |
13 |
11 |
3 |
5 |
5 |
1 |
29.0 |
+-------- Gonideinae
--+
| +----- Leoparreysiini **
+--+
| +-- Coelaturini **
+--+
+-- Indochinellini **
Konopleva et al., 2020 [Publ 4072], fig. 2
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
12 |
12 |
9 |
2 |
0 |
0 |
0 |
28.0 |
+-- Leoparreysiini **
--+
+-- Lamellidentini **
Konopleva et al., 2019 [Publ 3900], fig. 1
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
12 |
12 |
9 |
1 |
40 |
38 |
1 |
25.0 |
+-------- Pseudodontini
--+
| +----- Indochinellini **
+--+
| +-- Rectidentini
+--+
+-- Contradentini
Konopleva et al., 2017 [Publ 3807], fig. 6
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
11 |
11 |
9 |
4 |
20 |
10 |
1 |
19.0 |
+----------- Gonideinae
--+
| +-------- Lamellidentini **
+--+
| +----- Leoparreysiini **
+--+
| +-- Coelaturini **
+--+
+-- Indochinellini **
Pfeiffer et al., 2019 [Publ 3937], fig. 4-5
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
10 |
10 |
8 |
4 |
47 |
40 |
3 |
25.0 |
+----------------- Ambleminae
--+
| +-------------- Gonideinae
+--+
| +----------- Unioninae
+--+
| +-------- Lamellidentini **
+--+
| +----- Leoparreysiini **
+--+
| +-- Coelaturini **
+--+
+-- Indochinellini **
Pfeiffer et al., 2019 [Publ 3937], fig. 6
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
10 |
10 |
8 |
4 |
47 |
40 |
3 |
25.0 |
+-------------- Ambleminae
|
--+-------------- Gonideinae
|
| +----------- Unioninae
+--+
| +-------- Lamellidentini **
+--+
| +----- Leoparreysiini **
+--+
| +-- Coelaturini **
+--+
+-- Indochinellini **
Graf et al., 2014 [Publ 3591], fig. 3
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
9 |
9 |
12 |
2 |
2 |
1 |
1 |
30.0 |
+-- Leoparreysiini **
--+
+-- Coelaturini **
Whelan et al., 2011 [Publ 3380], fig. 1-ML
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
9 |
9 |
8 |
4 |
27 |
15 |
3 |
22.0 |
+-------- Lamellidentini **
+-----+
| | +----- Leoparreysiini **
| +--+
| | +-- Indochinellini **
| +--+
--+ +-- Coelaturini **
|
| +-- Pseudodontini
| +--------+
| | +-- Contradentini
+--+
| +-------- Unioninae
+--+
| +----- Gonideini
+--+
| +-- Lamprotulini
+--+
+-- Ambleminae
Bolotov et al., 2022 [Publ 4250], fig. S4
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
9 |
9 |
6 |
1 |
0 |
0 |
0 |
29.0 |
-- (Lamellidentini
Whelan et al., 2011 [Publ 3380], fig. 1-MP
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
9 |
9 |
7 |
4 |
27 |
15 |
3 |
22.0 |
+-------- Lamellidentini **
+--------+
| | +----- Leoparreysiini **
| +--+
| | +-- Indochinellini **
--+ +--+
| +-- Coelaturini **
|
| +-------------- Contradentini
+--+
| +----------- Pseudodontini
+--+
| +-------- Unioninae
+--+
| +----- Gonideini
+--+
| +-- Lamprotulini
+--+
+-- Ambleminae
Pfeiffer & Graf, 2015 [Publ 3620], fig. 2
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
8 |
8 |
6 |
4 |
32 |
22 |
3 |
22.0 |
+----- Unioninae
+-----+
| | +-- Ambleminae
| +--+
--+ +-- Gonideinae
|
| +-------- Lamellidentini **
+--+
| +----- Leoparreysiini **
+--+
| +-- Coelaturini **
+--+
+-- Indochinellini **
Konopleva et al., 2023 [Publ 4269], fig. S1_4
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
7 |
7 |
5 |
4 |
49 |
48 |
3 |
25.0 |
+----- Unioninae
+-----+
| | +-- Ambleminae
| +--+
--+ +-- Gonideinae
|
| +-------- Lamellidentini **
+--+
| +----- Parreysiini **
+--+
| +-- Leoparreysiini **
+--+
+-- Indochinellini **
Zieritz et al., 2024 [Publ 4381], fig. 2
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
5 |
5 |
3 |
4 |
41 |
39 |
3 |
27.0 |
+----------------- Ambleminae
|
| +----- Unioninae
| +-----+
| | | +-- Chamberlainia
| | +--+
--+ +--+ +-- Discomya
| | |
| | | +-------- Lamellidentini **
| | +--+
| | | +----- Leoparreysiini **
| | +--+
| | | +-- Indochinellini **
+--+ +--+
| +-- Coelaturini **
|
| +----- Ctenodesmini
| +-----+
| | | +-- Rectidentini
| | +--+
| | +-- Contradentini
+--+
| +-- Schepmaniini
| +-----+
| | +-- Pseudodontini
+--+
| +----- Caudiculatus
+--+
| +-- Potomida
+--+
+-- Gonideini
Bolotov et al., 2022 [Publ 4250], fig. S3
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
5 |
5 |
3 |
1 |
0 |
0 |
0 |
31.0 |
-- (Leoparreysia
Konopleva et al., 2017 [Publ 3807], fig. 5
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
5 |
5 |
3 |
1 |
0 |
0 |
0 |
25.0 |
-- (Lamellidentini
Lopes-Lima et al., 2017 [Publ 3745], fig. 1-2
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
4 |
4 |
2 |
4 |
76 |
64 |
3 |
26.0 |
+-------- Lamellidentini **
--+
| +----- Leoparreysiini **
+--+
| +-- Coelaturini **
+--+
+-- Indochinellini **
Imsong & Murali, 2023 [Publ 4370], fig. 2
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
3 |
3 |
1 |
3 |
0 |
0 |
0 |
30.0 |
+----- Lamellidentini **
--+
| +-- Indochinellini **
+--+
+-- Parreysiini **
Lopes-Lima et al., 2020 [Publ 4080], fig. 2-4
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
3 |
3 |
0 |
3 |
117 |
106 |
3 |
29.0 |
+-- Coelaturini **
|
+--------------+-- Leoparreysiini **
| |
| +-- Indochinellini **
|
--+ +-------------- Unioninae
| |
| | +-- Lens
| | |
| | +-- Ensidens
+--+ |
| +--------+-- Physunio
| | |
| | +-- Rectidens
| | |
+--+ +-- Yaukthwa
|
| +-------- Ambleminae
| |
+--+ +-- Chamberlainiini
| +--+
| | +-- Lamprotulini
+--+
| +-- Pseudodontini
+--+
+-- Gonideini
Dai et al., 2024 [Publ 4386], fig. 5-BI
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
3 |
3 |
1 |
2 |
18 |
16 |
1 |
29.0 |
+----- Unioninae
--+
| +-- Lamellidentini **
+--+
+-- Indochinellini **
Chen et al., 2023 [Publ 4302], fig. 1
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
3 |
3 |
0 |
3 |
37 |
35 |
3 |
28.0 |
+-- Gonideinae
+-----+
| +-- Ambleminae
--+
| +----- Unioninae
| |
+--+ +-- Coelaturini **
| |
+--+-- Indochinellini **
|
+-- Parreysiini **
Choi et al., 2020 [Publ 4169], fig. 5
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
3 |
3 |
1 |
3 |
7 |
7 |
1 |
32.0 |
+-------- Unioninae
--+
| +----- Indochinellini **
+--+
| +-- Lamellidentini **
+--+
+-- Leoparreysiini **
Chen et al., 2023 [Publ 4302], fig. 2
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
3 |
3 |
1 |
3 |
37 |
35 |
3 |
27.0 |
+-- Gonideinae
+--------+
| +-- Ambleminae
--+
| +-------- Unioninae
+--+
| +----- Parreysiini **
+--+
| +-- Coelaturini **
+--+
+-- Indochinellini **
Dai et al., 2024 [Publ 4386], fig. 5-ML
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
3 |
3 |
1 |
2 |
18 |
16 |
1 |
29.0 |
+----- Unioninae
--+
| +-- Lamellidentini **
+--+
+-- Indochinellini **
Lyubas et al., 2022 [Publ 4255], fig. 7
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
2 |
2 |
0 |
1 |
29 |
29 |
2 |
-- |
+----- Parreysiini **
--+
| +-- Gonideinae
+--+
+-- Unioninae
Dai et al., 2024 [Publ 4385], fig. 3-BI
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
2 |
2 |
0 |
2 |
29 |
27 |
1 |
-- |
+----- Unioninae
--+
| +-- Lamellidentini **
+--+
+-- Indochinellini **
Araujo et al., 2017 [Publ 3801], fig. 2
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
2 |
2 |
0 |
2 |
12 |
11 |
3 |
-- |
+-- Lamellidentini **
+-----+
| +-- Coelaturini **
--+
| +----- Ambleminae
+--+
| +-- Unioninae
+--+
+-- Gonideinae
Lyubas et al., 2022 [Publ 4255], fig. S1
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
2 |
2 |
0 |
1 |
29 |
29 |
2 |
-- |
+----- Parreysiini **
--+
| +-- Gonideinae
+--+
+-- Unioninae
Pfeiffer & Graf, 2013 [Publ 3531], fig. 2
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
2 |
2 |
0 |
2 |
32 |
29 |
3 |
-- |
+-- Lamellidentini **
--+
+-- Coelaturini **
Dai et al., 2024 [Publ 4385], fig. 3-ML
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
2 |
2 |
0 |
2 |
29 |
27 |
1 |
-- |
+----- Unioninae
--+
| +-- Lamellidentini **
+--+
+-- Indochinellini **
Sonowal et al., 2021 [Publ 4221], fig. 4
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
2 |
10 |
0 |
1 |
14 |
8 |
3 |
36.0 |
+-------------------------- Lamellidens_mainwaringi **
--+
| +----------------------- Lamprotula
+--+
| +-------------------- Potomida
+--+
| +----------------- Lampsilini
+--+
| +-------------- Pleurobemini
+--+
| +----------- Aculamprotula
+--+
| +-------- Unio_pictorum
+--+
| +----- Anodontini
+--+
| +-- Unio_ravoisieri
+--+
+-- Lamellidens_corrianus **
Graf, 2000 [Publ 2647], fig. 1
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
2 |
11 |
0 |
2 |
15 |
4 |
2 |
38.5 |
+-------- Unionini
+-----------+
| | +----- Pleurobemini
| +--+
| | +-- Anodontini
--+ +--+
| +-- Lampsilini
|
| +----------------- Parreysiini **
| |
+--+ +-------------- Coelaturini **
| |
| | +-------- Castaliini
+--+ +--+
| | | +----- Rhipidodontini
| | +--+
| | | +-- Hyridellini
+--+ +--+
| +-- Velesunioninae
|
| +-------- Iridininae
| |
+--+ +----- Anodontitinae
| |
+--+----- Mycetopodinae
|
| +-- Etheria
+--+
+-- Mulleria
Huang et al., 2018 [Publ 3828], fig. 3_5
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
1 |
1 |
0 |
1 |
28 |
5 |
3 |
-- |
+-------- Gonideinae
--+
| +----- Unioninae
+--+
| +-- Ambleminae
+--+
+-- Indochinellini **
Wu et al., 2020 [Publ 4121], fig. 2
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
1 |
1 |
0 |
1 |
33 |
10 |
3 |
-- |
+-- Ambleminae
+--+
| +-- Indochinellini **
--+
| +-- Gonideinae
+--+
+-- Unioninae
Wu et al., 2024 [Publ 4350], fig. 8a_S1
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
1 |
1 |
0 |
1 |
91 |
85 |
3 |
-- |
+-------- Ambleminae
--+
| +----- Gonideinae
+--+
| +-- Lamellidentini **
+--+
+-- Unioninae
Pavan-Kumar et al., 2022 [Publ 4263], fig. S5
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
1 |
1 |
0 |
1 |
38 |
36 |
3 |
-- |
+-- Lamellidentini **
+--+
| +-- Unioninae
--+
| +-- Ambleminae
+--+
+-- Gonideinae
Sonowal et al., 2021 [Publ 4221], fig. 3a
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
1 |
1 |
0 |
1 |
14 |
8 |
3 |
-- |
-- Lamellidens_corrianus
Sano et al., 2020 [Publ 4138], fig. 2
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
1 |
1 |
0 |
1 |
68 |
45 |
3 |
-- |
+-------- Indochinellini **
--+
| +----- Unioninae
+--+
| +-- Ambleminae
+--+
+-- Gonideinae
Roe & Hoeh, 2003 [Publ 2655], fig. 4f2
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
1 |
1 |
0 |
1 |
30 |
17 |
3 |
-- |
+----------------------------- Gonideinae
|
--+ +-------------------------- Coelaturini **
| |
+--+ +----------------------- Fusconaia
| |
| +----------------------- Amblemini
+--+
| +-------------------- Quadrulini
| |
+--+ +----------------- Pleurobema
| |
+--+ +-------------- Eurynia
| |
| | +-- Cyrtonaias
+--+-----------+
| +-- Unioninae
|
| +----------- Ortmanniana
+--+
| +-------- Glebula
+--+
| +----- Potamilus
+--+
| +-- Ligumia
+--+
+-- Toxolasma
Roe & Hoeh, 2003 [Publ 2655], fig. 4f3
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
1 |
1 |
0 |
1 |
30 |
17 |
3 |
-- |
+-------- Coelaturini **
--+
| +----- Unioninae
+--+
| +-- Gonideinae
+--+
+-- Ambleminae
Wu et al., 2024 [Publ 4350], fig. 8b_S2
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
1 |
1 |
0 |
1 |
91 |
85 |
3 |
-- |
+-------- Ambleminae
--+
| +----- Gonideinae
+--+
| +-- Lamellidentini **
+--+
+-- Unioninae
Sonowal et al., 2021 [Publ 4221], fig. 3b
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
1 |
1 |
0 |
1 |
14 |
8 |
3 |
-- |
+-- Aculamprotula
|
+-- Lamprotula
|
+-- Lampsilini
|
+-- Potomida
|
--+-- Unio_pictorum
|
+-- Anodontini
|
+-- Unio_ravoisieri
|
+-- Lamellidentini **
|
+-- Pleurobemini
Combosch et al., 2017 [Publ 3806], fig. 1
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
1 |
1 |
0 |
1 |
14 |
2 |
2 |
-- |
+----- Unioninae
--+
| +-- Ambleminae
+--+
+-- Indochinellini **
Jadhav & Jamkhedkar, 2009 [Publ 3697], fig. 11
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
1 |
1 |
0 |
1 |
24 |
24 |
3 |
-- |
+-------------------------------- Lamellidentini **
|
--+ +----------------------------- Unioninae
| |
+--+ +-------------------------- Lamprotulini
| |
+--+ +----------------------- Quadrulini
| |
+--+ +-------------------- Gonideini
| |
| | +----- Eurynia
+--+ +-----------+
| | | +-- Cambarunio
| | +--+
| | +-- Ptychobranchus
+--+
| +-- Amblemini
| +-----------+
| | +-- Pleurobema
+--+
| +----------- Truncilla
| |
+--+ +-- Lampsilis
| +-----+
| | +-- Epioblasma
+--+
| +----- Ligumia
+--+
| +-- Obliquaria
+--+
+-- Ortmanniana
Sano et al., 2020 [Publ 4138], fig. 3
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
1 |
1 |
0 |
1 |
68 |
45 |
3 |
-- |
+-------------- Indochinellini **
--+
| +----------- Unioninae
+--+
| +-------- Chamberlainiini
+--+
| +----- Ambleminae
+--+
| +-- Lamprotulini
+--+
+-- Gonideini
Wu et al., 2021 [Publ 4215], fig. 3
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
1 |
1 |
0 |
1 |
23 |
23 |
2 |
-- |
+----- Indochinellini **
--+
| +-- Unioninae
+--+
+-- Gonideinae
Pavan-Kumar et al., 2022 [Publ 4263], fig. 4
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
1 |
1 |
0 |
1 |
35 |
33 |
3 |
-- |
+-------- Ambleminae
--+
| +----- Gonideinae
+--+
| +-- Lamellidentini **
+--+
+-- Unioninae
Wu et al., 2023 [Publ 4339], fig. 4_S5-nt
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
1 |
1 |
0 |
1 |
95 |
84 |
3 |
-- |
+-------- Ambleminae
--+
| +----- Gonideinae
+--+
| +-- Lamellidentini **
+--+
+-- Unioninae
Smith, 2021 [Publ 4331], fig. 1
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
1 |
1 |
0 |
1 |
19 |
15 |
3 |
-- |
+-------- Ambleminae
--+
| +----- Gonideinae
+--+
| +-- Indochinellini **
+--+
+-- Unioninae
Hoeh et al., 2001 [Publ 2652], fig. 14f2
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
1 |
1 |
0 |
1 |
30 |
17 |
3 |
-- |
-- Coelatura_aegyptiaca
Hoeh et al., 2009 [Publ 2716], fig. 5
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
1 |
1 |
0 |
1 |
45 |
23 |
3 |
-- |
-- Coelatura_aegyptiaca
Wu et al., 2020 [Publ 4121], fig. S1
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
1 |
1 |
0 |
1 |
33 |
10 |
3 |
-- |
+-- Ambleminae
+--+
| +-- Indochinellini **
--+
| +-- Gonideinae
+--+
+-- Unioninae
Graf & Cummings, 2006 [Publ 1968], fig. 2
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
1 |
1 |
0 |
1 |
46 |
24 |
3 |
-- |
+-------- Pseudodontini
|
--+ +----- Coelaturini **
| |
| +----- Contradentini
+--+
+----- Unioninae
|
| +-- Gonideini
+--+
+-- Ambleminae
Sano et al., 2020 [Publ 4138], fig. 4
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
1 |
1 |
0 |
1 |
68 |
45 |
3 |
-- |
+-------- Indochinellini **
--+
| +----- Unioninae
+--+
| +-- Ambleminae
+--+
+-- Gonideinae
Hoeh et al., 2009 [Publ 2716], fig. 3
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
1 |
1 |
0 |
1 |
46 |
24 |
3 |
-- |
-- Coelatura_aegyptiaca
Dai et al., 2024 [Publ 4386], fig. 3
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
1 |
1 |
0 |
1 |
12 |
12 |
1 |
-- |
+-- Indochinellini **
--+
+-- Unioninae
Hoeh et al., 2001 [Publ 2652], fig. 14f4
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
1 |
1 |
0 |
1 |
30 |
17 |
3 |
-- |
-- Coelatura_aegyptiaca
Hoeh et al., 2002 [Publ 2653], fig. 2
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
1 |
1 |
0 |
1 |
30 |
17 |
3 |
-- |
-- Coelatura_aegyptiaca
Combosch et al., 2017 [Publ 3806], fig. 2
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
1 |
1 |
0 |
1 |
14 |
2 |
2 |
-- |
+----- Unioninae
--+
| +-- Indochinellini **
+--+
+-- Ambleminae
Jeratthitikul et al., 2019 [Publ 3911], fig. 4
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
1 |
1 |
0 |
1 |
8 |
8 |
1 |
-- |
+-- Pseudodontini
+--+
| +-- Indochinellini **
--+
| +-- Rectidentini
+--+
+-- Contradentini
Hoeh et al., 2009 [Publ 2716], fig. 4
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
1 |
1 |
0 |
1 |
45 |
23 |
3 |
-- |
-- Coelatura_aegyptiaca
Bogan & Hoeh, 2000 [Publ 2650], fig. 1
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
1 |
1 |
0 |
1 |
33 |
17 |
3 |
-- |
-- Coelatura_aegyptiaca
Wu et al., 2023 [Publ 4339], fig. 4_S5-aa
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
1 |
1 |
0 |
1 |
95 |
84 |
3 |
-- |
+-------- Ambleminae
--+
| +----- Gonideinae
+--+
| +-- Lamellidentini **
+--+
+-- Unioninae
|