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Page last updated
Wed 27 Sep 2023

The Freshwater Mussels (Unionoida) of the World (and other less consequential bivalves)

FM(U)otW(aolcb) is the web version of the MUSSEL Project Database. Follow the links to browse the data or use the custom Google search field. Either way, you win!

Cladomics pages are experimental — even more so than the rest of this site!

The purpose of the cladomics pages for each genus and family-group level taxon is to provide statistics and references indicating the extent to which published phylogenetic work supports the classification used here.

The Cladomics Summary provides an overview of the number of studies that have included terminal species representing this taxon as well as an indication of how frequently it has been recovered as monophyletic.

Following the summary, the available published clades representing the taxon are listed.

For more information, visit the Cladomics FARQ (frequently asked rhetorical questions) page. Abbreviations for the various statistics are explained on the FARQ!

Search the database pages:

Bivalvia | Unionoida | Unionoidea | Unionidae | Gonideinae | Pseudodontini

subtribe PILSBRYOCONCHINA Bolotov, Vikhrev & Tumpeesuwan, 2017: cladomics
     8 genera

Cladomics Summary

number of species = 39
number of subtaxa (rank: genus) = 8

analyzed in 61 cladograms in 37 publications
maximum number of terminals in an analysis = 26, mean = 6.1
maximum number of subtaxa (genus) in an analysis = 8, mean = 3.49

Basic Monophyly

number of cladograms that test basic monophyly (i.e., more than 1 included species) = 52
maximum number of included terminals = 26, mean = 6.98
maximum number of subtaxa (genus) = 8, mean = 3.92
proportion of cladograms that support basic monophyly = 0.83

Rigorous Monophyly

number of cladograms that test rigorous monophyly = 31
(Rigorous monophyly refers to cladograms with terminal species representing more than one subtaxon (genus) taxon and at least one rigorous outgroup (subtribe).
maximum number of included terminals = 26, mean = 10.0
maximum number of subtaxa (genus) = 8, mean = 4.97
proportion of cladograms that support rigorous monophyly = 0.71


Pairwise Cladogram Clustering

The graphs below depict in two dimensions all pair-wise ingroup distances within the cladome. See the Cladmoics FARQ for more information.


PCoA Ordination


NJ tree


Konopleva et al., 2023 [Publ 4262], fig. 8

iT tT bi riS oT roT roS iD
26 26 24 8 58 5 1 0.0

*** Top Monophyly Cladogram *** *** Top Sister-Group Cladogram ***

                       +----------- Pseudodon_kayinensis         
  +--------------------+                                         
  |                    |  +-------- Pseudodon_inoscularis        
  |                    +--+                                      
  |                       |  +----- Pseudodon_bogani             
  |                       +--+                                   
  |                          |  +-- Pseudodon_salwenianus        
  |                          +--+                                
  |                             +-- Pseudodon_manueli            
--+                                                              
  |                             +-- Pilsbryoconcha_schomburgki **   
  |                       +-----+                                
  |                       |     +-- Pilsbryoconcha_exilis[1] **     
  |  +--------------------+                                      
  |  |                    |  +----- Pilsbryoconcha **               
  |  |                    +--+                                   
  |  |                       |  +-- Pilsbryoconcha_exilis[2] **     
  |  |                       +--+                                
  +--+                          +-- Pilsbryoconcha_linguaeformis ** 
     |                                                           
     |  +-------------------------- Songkhlanaia_tamodienica **     
     |  |                                                        
     |  |                       +-- Namkongnaia_inkhavilayi **      
     |  |  +--------------------+                                
     +--+  |                    +-- Namkongnaia_lemeslei **         
        |  |                                                     
        |  |              +-------- Monodontina_lenyanensis **      
        |  |  +-----------+                                      
        |  |  |           |  +----- Monodontina_cambodjensis **     
        +--+  |           +--+                                   
           |  |              |  +-- Monodontina_vondembuschiana **  
           |  |              +--+                                
           |  |                 +-- Monodontina_mekongi **          
           |  |                                                  
           |  |                 +-- Sundadontina_taskaevi **        
           +--+        +--------+                                
              |        |        +-- Sundadontina_brandti **         
              |  +-----+                                         
              |  |     |  +-------- Sundadontina_tumida **          
              |  |     +--+                                      
              |  |        |  +----- Sundadontina_tanintharyiensis **
              |  |        +--+                                   
              |  |           |  +-- Sundadontina_plugpomenica **    
              +--+           +--+                                
                 |              +-- Sundadontina_cumingii **        
                 |                                               
                 |  +-------------- Nyeinchanconcha_nyeinchani **   
                 |  |                                            
                 |  |        +----- Thaiconcha_thaiensis **         
                 +--+  +-----+                                   
                    |  |     |  +-- Thaiconcha_munelliptica **      
                    |  |     +--+                                
                    +--+        +-- Thaiconcha_callifera **         
                       |                                         
                       |  +-------- Bineurus_loeiensis **           
                       +--+                                      
                          |  +----- Bineurus_mouhotii **            
                          +--+                                   
                             |  +-- Bineurus_anodontinum **         
                             +--+                                
                                +-- Bineurus_exilis **              

Konopleva et al., 2021 [Publ 4213], fig. 1

iT tT bi riS oT roT roS iD
20 20 17 7 9 5 1 12.0
              +----------- Pseudodon_kayinensis         
  +-----------+                                         
  |           |  +-------- Pseudodon_inoscularis        
  |           +--+                                      
  |              |  +----- Pseudodon_bogani             
  |              +--+                                   
  |                 |  +-- Pseudodon_salwenianus        
  |                 +--+                                
  |                    +-- Pseudodon_manueli            
  |                                                     
--+                 +----- Pilsbryoconcha_exilis[2] **     
  |  +--------------+                                   
  |  |              |  +-- Pilsbryoconcha_exilis[1] **     
  |  |              +--+                                
  |  |                 +-- Namkongnaia_lemeslei **         
  |  |                                                  
  |  |     +-------------- Nyeinchanconcha_nyeinchani **   
  |  |     |                                            
  |  |     |           +-- Thaiconcha_munelliptica **      
  +--+  +--+           |                                
     |  |  |  +--------+-- Thaiconcha_thaiensis **         
     |  |  |  |        |                                
     |  |  +--+        +-- Thaiconcha_callifera **         
     |  |     |                                         
     |  |     |  +-------- Bineurus_loeiensis **           
     |  |     +--+                                      
     |  |        |  +----- Bineurus_mouhotii **            
     |  |        +--+                                   
     +--+           |  +-- Bineurus_anodontinum **         
        |           +--+                                
        |              +-- Bineurus_exilis **              
        |                                               
        |        +-------- Monodontina_lenyanensis **      
        |     +--+                                      
        |     |  |  +----- Monodontina_cambodjensis **     
        |     |  +--+                                   
        |     |     |  +-- Monodontina_vondembuschiana **  
        |     |     +--+                                
        +-----+        +-- Monodontina_mekongi **          
              |                                         
              |        +-- Sundadontina_taskaevi **        
              |  +-----+                                
              |  |     +-- Sundadontina_brandti **         
              +--+                                      
                 |  +----- Sundadontina_tumida **          
                 +--+                                   
                    |  +-- Sundadontina_cumingii **        
                    +--+                                
                       +-- Sundadontina_tanintharyiensis **

Bolotov et al., 2020 [Publ 4100], fig. 1

iT tT bi riS oT roT roS iD
18 18 16 6 103 5 1 19.0
              +----------- Pseudodon_kayinensis         
  +-----------+                                         
  |           |  +-------- Pseudodon_inoscularis        
  |           +--+                                      
  |              |  +----- Pseudodon_bogani             
  |              +--+                                   
  |                 |  +-- Pseudodon_salwenianus        
  |                 +--+                                
--+                    +-- Pseudodon_manueli            
  |                                                     
  |                 +----- Bineurus_loeiensis **           
  |  +--------------+                                   
  |  |              |  +-- Bineurus_exilis **              
  |  |              +--+                                
  |  |                 +-- Bineurus_mouhotii **            
  |  |                                                  
  +--+  +----------------- Nyeinchanconcha_nyeinchani **   
     |  |                                               
     |  |              +-- Thaiconcha_callifera **         
     |  |        +-----+                                
     |  |        |     +-- Thaiconcha_munelliptica **      
     +--+  +-----+                                      
        |  |     |  +----- Pilsbryoconcha_exilis[2] **     
        |  |     +--+                                   
        |  |        |  +-- Pilsbryoconcha_exilis[1] **     
        |  |        +--+                                
        |  |           +-- Pilsbryoconcha **               
        +--+                                            
           |     +-------- Monodontina_lenyanensis **      
           |  +--+                                      
           |  |  |  +----- Monodontina_cambodjensis **     
           |  |  +--+                                   
           |  |     |  +-- Monodontina_vondembuschiana **  
           |  |     +--+                                
           +--+        +-- Monodontina_mekongi **          
              |                                         
              |        +-- Sundadontina_taskaevi **        
              |  +-----+                                
              |  |     +-- Sundadontina_brandti **         
              +--+                                      
                 |  +----- Sundadontina_tumida **          
                 +--+                                   
                    |  +-- Sundadontina_cumingii **        
                    +--+                                
                       +-- Sundadontina_tanintharyiensis **

Bolotov et al., 2020 [Publ 4100], fig. S1

iT tT bi riS oT roT roS iD
18 18 16 6 97 5 1 21.0
              +----------- Pseudodon_kayinensis         
  +-----------+                                         
  |           |  +-------- Pseudodon_inoscularis        
  |           +--+                                      
  |              |  +----- Pseudodon_bogani             
  |              +--+                                   
  |                 |  +-- Pseudodon_salwenianus        
  |                 +--+                                
--+                    +-- Pseudodon_manueli            
  |                                                     
  |                 +----- Bineurus_loeiensis **           
  |  +--------------+                                   
  |  |              |  +-- Bineurus_exilis **              
  |  |              +--+                                
  |  |                 +-- Bineurus_mouhotii **            
  |  |                                                  
  +--+  +----------------- Nyeinchanconcha_nyeinchani **   
     |  |                                               
     |  |              +-- Thaiconcha_callifera **         
     |  |        +-----+                                
     |  |        |     +-- Thaiconcha_munelliptica **      
     +--+  +-----+                                      
        |  |     |  +----- Pilsbryoconcha_exilis[2] **     
        |  |     +--+                                   
        |  |        |  +-- Pilsbryoconcha_exilis[1] **     
        |  |        +--+                                
        |  |           +-- Pilsbryoconcha **               
        +--+                                            
           |     +-------- Monodontina_lenyanensis **      
           |  +--+                                      
           |  |  |  +----- Monodontina_cambodjensis **     
           |  |  +--+                                   
           |  |     |  +-- Monodontina_vondembuschiana **  
           |  |     +--+                                
           +--+        +-- Monodontina_mekongi **          
              |                                         
              |        +-- Sundadontina_tanintharyiensis **
              |  +-----+                                
              |  |     +-- Sundadontina_cumingii **        
              +--+                                      
                 |  +----- Sundadontina_tumida **          
                 +--+                                   
                    |  +-- Sundadontina_taskaevi **        
                    +--+                                
                       +-- Sundadontina_brandti **         

Jeratthitikul et al., 2021 [Publ 4219], fig. 1

iT tT bi riS oT roT roS iD
18 18 15 7 19 5 1 16.0
              +----------- Pseudodon_kayinensis        
  +-----------+                                        
  |           |  +-------- Pseudodon_inoscularis       
  |           +--+                                     
  |              |  +----- Pseudodon_bogani            
  |              +--+                                  
  |                 |  +-- Pseudodon_manueli           
  |                 +--+                               
  |                    +-- Pseudodon_salwenianus       
--+                                                    
  |                 +----- Pilsbryoconcha_exilis **       
  |  +--------------+                                  
  |  |              |  +-- Pilsbryoconcha_linguaeformis **
  |  |              +--+                               
  |  |                 +-- Pilsbryoconcha **              
  |  |                                                 
  |  |           +-------- Nyeinchanconcha_nyeinchani **  
  |  |           |                                     
  +--+  +--------+  +----- Thaiconcha_callifera **        
     |  |        |  |                                  
     |  |        +--+  +-- Bineurus_panhai **             
     |  |           |  |                               
     |  |           +--+-- Bineurus_exilis **             
     |  |              |                               
     |  |              +-- Bineurus_mouhotii **           
     +--+                                              
        |              +-- Sundadontina_brandti **        
        |           +--+                               
        |           |  +-- Sundadontina_taskaevi **       
        |  +--------+                                  
        |  |        |  +-- Sundadontina_tumida **         
        |  |        +--+                               
        |  |           +-- Sundadontina_cumingii **       
        +--+                                           
           |           +-- Namkongnaia_lemeslei **        
           |  +--------+                               
           |  |        +-- Namkongnaia_inkhavilayi **     
           +--+                                        
              |  +-------- Monodontina_lenyanensis **     
              +--+                                     
                 |  +----- Monodontina_cambodjensis **    
                 +--+                                  
                    |  +-- Monodontina_vondembuschiana ** 
                    +--+                               
                       +-- Monodontina_mekongi **         

Jeratthitikul et al., 2021 [Publ 4219], fig. 2

iT tT bi riS oT roT roS iD
18 18 16 7 19 5 1 22.0
              +----------- Pseudodon_kayinensis        
  +-----------+                                        
  |           |  +-------- Pseudodon_inoscularis       
  |           +--+                                     
  |              |  +----- Pseudodon_bogani            
  |              +--+                                  
  |                 |  +-- Pseudodon_manueli           
  |                 +--+                               
  |                    +-- Pseudodon_salwenianus       
--+                                                    
  |                 +----- Pilsbryoconcha_exilis **       
  |  +--------------+                                  
  |  |              |  +-- Pilsbryoconcha_linguaeformis **
  |  |              +--+                               
  |  |                 +-- Pilsbryoconcha **              
  |  |                                                 
  |  |                 +-- Nyeinchanconcha_nyeinchani **  
  |  |        +--------+                               
  +--+        |        +-- Sundadontina_cumingii **       
     |  +-----+                                        
     |  |     |  +-------- Thaiconcha_callifera **        
     |  |     +--+                                     
     |  |        |  +----- Bineurus_mouhotii **           
     |  |        +--+                                  
     |  |           |  +-- Bineurus_panhai **             
     |  |           +--+                               
     +--+              +-- Bineurus_exilis **             
        |                                              
        |           +----- Sundadontina_tumida **         
        |  +--------+                                  
        |  |        |  +-- Sundadontina_brandti **        
        |  |        +--+                               
        |  |           +-- Sundadontina_taskaevi **       
        +--+                                           
           |           +-- Namkongnaia_lemeslei **        
           |  +--------+                               
           |  |        +-- Namkongnaia_inkhavilayi **     
           +--+                                        
              |  +-------- Monodontina_lenyanensis **     
              +--+                                     
                 |  +----- Monodontina_cambodjensis **    
                 +--+                                  
                    |  +-- Monodontina_vondembuschiana ** 
                    +--+                               
                       +-- Monodontina_mekongi **         

Jeratthitikul et al., 2022 [Publ 4260], fig. S1, 1

iT tT bi riS oT roT roS iD
15 15 13 7 61 2 1 31.0
                       +-- Pseudodon_inoscularis       
  +--------------------+                               
  |                    +-- Pseudodon_kayinensis        
  |                                                    
  |                    +-- Nyeinchanconcha_nyeinchani **  
  |              +-----+                               
--+              |     +-- Sundadontina_cumingii **       
  |  +-----------+                                     
  |  |           |  +----- Thaiconcha_callifera **        
  |  |           +--+                                  
  |  |              |  +-- Bineurus_panhai **             
  |  |              +--+                               
  +--+                 +-- Bineurus_mouhotii **           
     |                                                 
     |                 +-- Namkongnaia_inkhavilayi **     
     |  +--------------+                               
     |  |              +-- Monodontina_vondembuschiana ** 
     |  |                                              
     +--+              +-- Pilsbryoconcha_exilis **       
        |  +-----------+                               
        |  |           +-- Pilsbryoconcha_schomburgki **  
        |  |                                           
        +--+           +-- Pilsbryoconcha_carinifera **   
           |  +--------+                               
           |  |        +-- Pilsbryoconcha_linguaeformis **
           +--+                                        
              |  +-------- Pilsbryoconcha_hoikaab **      
              +--+                                     
                 |  +----- Pilsbryoconcha_mekongiana **   
                 +--+                                  
                    |  +-- Pilsbryoconcha_acuta **        
                    +--+                               
                       +-- Pilsbryoconcha_kittitati **    

Jeratthitikul et al., 2022 [Publ 4260], fig. S2, 2

iT tT bi riS oT roT roS iD
15 15 13 7 61 2 1 31.0
                       +-- Pseudodon_inoscularis       
  +--------------------+                               
  |                    +-- Pseudodon_kayinensis        
  |                                                    
  |                    +-- Nyeinchanconcha_nyeinchani **  
  |              +-----+                               
--+              |     +-- Sundadontina_cumingii **       
  |  +-----------+                                     
  |  |           |  +----- Thaiconcha_callifera **        
  |  |           +--+                                  
  |  |              |  +-- Bineurus_panhai **             
  |  |              +--+                               
  +--+                 +-- Bineurus_mouhotii **           
     |                                                 
     |                 +-- Namkongnaia_inkhavilayi **     
     |  +--------------+                               
     |  |              +-- Monodontina_vondembuschiana ** 
     |  |                                              
     +--+              +-- Pilsbryoconcha_exilis **       
        |  +-----------+                               
        |  |           +-- Pilsbryoconcha_schomburgki **  
        |  |                                           
        +--+           +-- Pilsbryoconcha_carinifera **   
           |  +--------+                               
           |  |        +-- Pilsbryoconcha_linguaeformis **
           +--+                                        
              |  +-------- Pilsbryoconcha_hoikaab **      
              +--+                                     
                 |  +----- Pilsbryoconcha_mekongiana **   
                 +--+                                  
                    |  +-- Pilsbryoconcha_acuta **        
                    +--+                               
                       +-- Pilsbryoconcha_kittitati **    

Dai et al., 2023 [Publ 4285], fig. 3-BI

iT tT bi riS oT roT roS iD
13 13 10 6 31 3 1 20.0
           +----- Pseudodon_inoscularis        
  +--------+                                   
  |        |  +-- Pseudodon_bogani             
  |        +--+                                
  |           +-- Pseudodon_manueli            
  |                                            
  |     +-------- Pilsbryoconcha_exilis **        
  |  +--+                                      
--+  |  |  +----- Monodontina_lenyanensis **      
  |  |  +--+                                   
  |  |     |  +-- Monodontina_mekongi **          
  |  |     +--+                                
  |  |        +-- Monodontina_cambodjensis **     
  |  |                                         
  |  |  +-------- Nyeinchanconcha_nyeinchani **   
  |  +--+                                      
  +--+  |  +----- Bineurus_loeiensis **           
     |  +--+                                   
     |     |  +-- Bineurus_mouhotii **            
     |     +--+                                
     |        +-- Bineurus_exilis **              
     |                                         
     |        +-- Thaiconcha_callifera **         
     |  +-----+                                
     |  |     +-- Thaiconcha_munelliptica **      
     +--+                                      
        |  +----- Sundadontina_tanintharyiensis **
        +--+                                   
           |  +-- Sundadontina_brandti **         
           +--+                                
              +-- Sundadontina_taskaevi **        

Dai et al., 2023 [Publ 4285], fig. 3-ML

iT tT bi riS oT roT roS iD
13 13 11 6 31 3 1 21.0
              +----- Pseudodon_inoscularis        
  +-----------+                                   
  |           |  +-- Pseudodon_bogani             
  |           +--+                                
  |              +-- Pseudodon_manueli            
  |                                               
  |        +-------- Nyeinchanconcha_nyeinchani **   
--+  +-----+                                      
  |  |     |  +----- Bineurus_loeiensis **           
  |  |     +--+                                   
  |  |        |  +-- Bineurus_mouhotii **            
  |  |        +--+                                
  |  |           +-- Bineurus_exilis **              
  +--+                                            
     |     +-------- Pilsbryoconcha_exilis **        
     |  +--+                                      
     |  |  |  +----- Monodontina_lenyanensis **      
     |  |  +--+                                   
     |  |     |  +-- Monodontina_mekongi **          
     |  |     +--+                                
     +--+        +-- Monodontina_cambodjensis **     
        |                                         
        |        +-- Thaiconcha_callifera **         
        |  +-----+                                
        |  |     +-- Thaiconcha_munelliptica **      
        +--+                                      
           |  +----- Sundadontina_tanintharyiensis **
           +--+                                   
              |  +-- Sundadontina_brandti **         
              +--+                                
                 +-- Sundadontina_taskaevi **        

Huang et al., 2019 [Publ 3843], fig. 2

iT tT bi riS oT roT roS iD
12 12 9 6 189 3 1 18.0
                 +----- Pseudodon_manueli          
  +--------------+                                 
  |              |  +-- Pseudodon_inoscularis      
  |              +--+                              
  |                 +-- Pseudodon_bogani           
--+                                                
  |  +----------------- Pilsbryoconcha_exilis **      
  |  |                                             
  |  |     +----------- Nyeinchanconcha_nyeinchani ** 
  |  |  +--+                                       
  +--+  |  |  +-------- Thaiconcha_callifera **       
     |  |  +--+                                    
     |  |     |  +----- Bineurus_loeiensis **         
     |  |     +--+                                 
     |  |        |  +-- Bineurus_mouhotii **          
     +--+        +--+                              
        |           +-- Bineurus_exilis **            
        |                                          
        |        +----- Monodontina_cambodjensis **   
        |     +--+                                 
        |     |  |  +-- Monodontina_mekongi **        
        |     |  +--+                              
        +-----+     +-- Monodontina_vondembuschiana **
              |                                    
              |     +-- Sundadontina_cumingii **      
              |     |                              
              +-----+-- Sundadontina_tumida[1] **     
                    |                              
                    +-- Sundadontina_tumida[2] **     

Huang et al., 2019 [Publ 3843], fig. S3

iT tT bi riS oT roT roS iD
12 12 10 6 186 3 1 22.0
                 +----- Pseudodon_inoscularis         
  +--------------+                                    
  |              |  +-- Pseudodon_bogani              
  |              +--+                                 
  |                 +-- Pseudodon_manueli             
--+                                                   
  |  +----------------- Pilsbryoconcha_exilis **         
  |  |                                                
  |  |     +----------- Sundadontina_tumida **           
  |  |  +--+                                          
  +--+  |  |  +-------- Monodontina_cambodjensis **      
     |  |  +--+                                       
     |  |     |  +----- Monodontina_vondembuschiana[1] **
     |  |     +--+                                    
     |  |        |  +-- Monodontina_mekongi **           
     +--+        +--+                                 
        |           +-- Monodontina_vondembuschiana[2] **
        |                                             
        |           +-- Nyeinchanconcha_nyeinchani **    
        |  +--------+                                 
        |  |        +-- Sundadontina_cumingii **         
        +--+                                          
           |  +-------- Thaiconcha_callifera **          
           +--+                                       
              |  +----- Bineurus_loeiensis **            
              +--+                                    
                 |  +-- Bineurus_mouhotii **             
                 +--+                                 
                    +-- Bineurus_exilis **               

Huang et al., 2019 [Publ 3843], fig. S4

iT tT bi riS oT roT roS iD
12 12 4 6 188 3 1 21.0
        +----- Pseudodon_manueli          
  +-----+                                 
  |     |  +-- Pseudodon_inoscularis      
  |     +--+                              
  |        +-- Pseudodon_bogani           
  |                                       
  |  +-------- Pilsbryoconcha_exilis **      
  |  |                                    
--+  +-------- Nyeinchanconcha_nyeinchani ** 
  |  |                                    
  |  +-------- Thaiconcha_callifera **       
  |  |                                    
  |  +-------- Sundadontina_cumingii **      
  |  |                                    
  |  +-------- Sundadontina_tumida[1] **     
  +--+                                    
     +-------- Sundadontina_tumida[2] **     
     |                                    
     |  +----- Monodontina_cambodjensis **   
     +--+                                 
     |  |  +-- Monodontina_mekongi **        
     |  +--+                              
     |     +-- Monodontina_vondembuschiana **
     |                                    
     |  +----- Bineurus_loeiensis **         
     +--+                                 
        |  +-- Bineurus_mouhotii **          
        +--+                              
           +-- Bineurus_exilis **            

Bolotov et al., 2017 [Publ 3761], fig. S1

iT tT bi riS oT roT roS iD
11 11 9 6 127 1 1 23.0
  +-------------------- Pseudodon_inoscularis      
  |                                                
  |  +----------------- Pilsbryoconcha_exilis **      
--+  |                                             
  |  |        +-------- Sundadontina_tumida **        
  |  |  +-----+                                    
  +--+  |     |  +----- Monodontina_cambodjensis **   
     |  |     +--+                                 
     |  |        |  +-- Monodontina_mekongi **        
     |  |        +--+                              
     +--+           +-- Monodontina_vondembuschiana **
        |                                          
        |           +-- Nyeinchanconcha_nyeinchani ** 
        |  +--------+                              
        |  |        +-- Sundadontina_cumingii **      
        +--+                                       
           |  +-------- Thaiconcha_callifera **       
           +--+                                    
              |  +----- Bineurus_exilis **            
              +--+                                 
                 |  +-- Bineurus_mouhotii **          
                 +--+                              
                    +-- Bineurus_loeiensis **         

Bolotov et al., 2017 [Publ 3761], fig. S2

iT tT bi riS oT roT roS iD
11 13 10 6 127 2 1 24.0
  +-------------- Pilsbryoconcha_exilis **      
  |                                          
  |  +----------- Pseudodon_inoscularis[2]   
  |  |                                       
--+  |  +-------- Thaiconcha_callifera **       
  |  +--+                                    
  |  |  |  +----- Bineurus_exilis **            
  |  |  +--+                                 
  |  |     |  +-- Bineurus_mouhotii **          
  +--+     +--+                              
     |        +-- Bineurus_loeiensis **         
     |                                       
     |     +----- Monodontina_cambodjensis **   
     |  +--+                                 
     |  |  |  +-- Monodontina_mekongi **        
     |  |  +--+                              
     |  |     +-- Monodontina_vondembuschiana **
     +--+                                    
        |     +-- Pseudodon_inoscularis[1]   
        |  +--+                              
        |  |  +-- Sundadontina_tumida **        
        +--+                                 
           |  +-- Nyeinchanconcha_nyeinchani ** 
           +--+                              
              +-- Sundadontina_cumingii **      

Bolotov et al., 2017 [Publ 3764], fig. S1

iT tT bi riS oT roT roS iD
11 11 7 6 117 3 1 25.0
           +----- Pseudodon_inoscularis      
  +--------+                                 
  |        |  +-- Pseudodon_manueli          
  |        +--+                              
  |           +-- Pseudodon_bogani           
  |                                          
  |           +-- Thaiconcha_callifera **       
--+  +--------+                              
  |  |        +-- Pilsbryoconcha_exilis **      
  |  |                                       
  |  |        +-- Nyeinchanconcha_nyeinchani ** 
  |  +--------+                              
  |  |        +-- Sundadontina_cumingii **      
  |  |                                       
  +--+     +----- Bineurus_exilis **            
     +-----+                                 
     |     |  +-- Bineurus_mouhotii **          
     |     +--+                              
     |        +-- Bineurus_loeiensis **         
     |                                       
     |  +-------- Sundadontina_tumida **        
     +--+                                    
        |  +----- Monodontina_cambodjensis **   
        +--+                                 
           |  +-- Monodontina_mekongi **        
           +--+                              
              +-- Monodontina_vondembuschiana **

Jeratthitikul & Sutcharit, 2023 [Publ 4289], fig. 1

iT tT bi riS oT roT roS iD
11 11 9 7 2 2 1 29.0
                 +-- Pseudodon_inoscularis      
  +--------------+                              
  |              +-- Pseudodon_kayinensis       
  |                                             
  |     +----------- Bineurus_mouhotii **          
  |  +--+                                       
--+  |  |  +-------- Bineurus_loeiensis **         
  |  |  +--+                                    
  |  |     |  +----- Bineurus_panhai **            
  |  |     +--+                                 
  |  |        |  +-- Bineurus_anodontinum **       
  +--+        +--+                              
     |           +-- Bineurus_exilis **            
     |                                          
     |        +----- Namkongnaia_inkhavilayi **    
     |     +--+                                 
     |     |  |  +-- Sundadontina_cumingii **      
     |     |  +--+                              
     +-----+     +-- Nyeinchanconcha_nyeinchani ** 
           |                                    
           |  +----- Monodontina_vondembuschiana **
           +--+                                 
              |  +-- Pilsbryoconcha_exilis **      
              +--+                              
                 +-- Thaiconcha_callifera **       

Bolotov et al., 2018 [Publ 3826], fig. 2

iT tT bi riS oT roT roS iD
8 8 0 3 77 3 1 24.0
     +-- Pseudodon_inoscularis      
     |                              
  +--+-- Pseudodon_bogani           
  |  |                              
  |  +-- Pseudodon_manueli          
  |                                 
  |  +-- Bineurus_mouhotii **          
--+  |                              
  |  +-- Monodontina_cambodjensis **   
  |  |                              
  |  +-- Monodontina_mekongi **        
  |  |                              
  |  +-- Pilsbryoconcha_exilis[1] **   
  +--+                              
     +-- Pilsbryoconcha_exilis[2] **   
     |                              
     +-- Bineurus_loeiensis **         
     |                              
     +-- Bineurus_exilis **            
     |                              
     +-- Monodontina_vondembuschiana **

Konopleva et al., 2019 [Publ 3900], fig. 1

iT tT bi riS oT roT roS iD
8 8 0 3 44 3 1 24.0
     +-- Pseudodon_inoscularis      
     |                              
  +--+-- Pseudodon_bogani           
  |  |                              
  |  +-- Pseudodon_manueli          
  |                                 
  |  +-- Bineurus_mouhotii **          
--+  |                              
  |  +-- Monodontina_cambodjensis **   
  |  |                              
  |  +-- Monodontina_mekongi **        
  |  |                              
  |  +-- Pilsbryoconcha_exilis[1] **   
  +--+                              
     +-- Pilsbryoconcha_exilis[2] **   
     |                              
     +-- Bineurus_loeiensis **         
     |                              
     +-- Bineurus_exilis **            
     |                              
     +-- Monodontina_vondembuschiana **

Zieritz et al., 2021 [Publ 4218], fig. 5

iT tT bi riS oT roT roS iD
7 7 5 5 25 2 1 19.5
                 +-- Pseudodon_bogani           
  +--------------+                              
  |              +-- Pseudodon_crebristriatus   
--+                                             
  |  +-------------- Pilsbryoconcha_exilis **      
  |  |                                          
  +--+           +-- Monodontina_mekongi **        
     |  +--------+                              
     |  |        +-- Monodontina_vondembuschiana **
     +--+                                       
        |  +-------- Sundadontina **               
        +--+                                    
           |  +----- Thaiconcha_callifera **       
           +--+                                 
              |  +-- Bineurus_loeiensis **         
              +--+                              
                 +-- Bineurus_anodontinum **       

Pfeiffer & Graf, 2015 [Publ 3620], fig. 2

iT tT bi riS oT roT roS iD
6 6 4 5 34 0 0 20.0
  +----------- Pilsbryoconcha_exilis **      
  |                                       
--+        +-- Monodontina_vondembuschiana **
  |  +-----+                              
  |  |     +-- Monodontina_cambodjensis **   
  +--+                                    
     |  +----- Sundadontina_tumida **        
     +--+                                 
        |  +-- Nyeinchanconcha_nyeinchani ** 
        +--+                              
           +-- Bineurus_exilis **            

Konopleva et al., 2017 [Publ 3807], fig. 6

iT tT bi riS oT roT roS iD
6 6 4 5 25 0 0 24.0
  +----------- Sundadontina_tumida **        
  |                                       
--+        +-- Nyeinchanconcha_nyeinchani ** 
  |  +-----+                              
  |  |     +-- Bineurus_exilis **            
  +--+                                    
     |  +----- Pilsbryoconcha_exilis **      
     +--+                                 
        |  +-- Monodontina_cambodjensis **   
        +--+                              
           +-- Monodontina_vondembuschiana **

Lopes-Lima et al., 2017 [Publ 3745], fig. 1-2

iT tT bi riS oT roT roS iD
5 5 3 4 76 0 0 23.0
        +-- Pilsbryoconcha_exilis **     
  +-----+                             
  |     +-- Pilsbryoconcha_acuta **      
--+                                   
  |  +----- Monodontina_cambodjensis **  
  +--+                                
     |  +-- Bineurus_exilis **           
     +--+                             
        +-- Nyeinchanconcha_nyeinchani **

Lopes-Lima et al., 2020 [Publ 4080], fig. 2-4

iT tT bi riS oT roT roS iD
5 5 2 5 116 2 1 22.0
           +-- Pseudodon_inoscularis     
  +--------+                             
  |        +-- Pseudodon_manueli         
--+                                      
  |  +-------- Pilsbryoconcha_exilis **     
  |  |                                   
  +--+  +----- Monodontina_mekongi **       
     |  |                                
     +--+  +-- Nyeinchanconcha_nyeinchani **
        |  |                             
        +--+-- Bineurus_mouhotii **         
           |                             
           +-- Sundadontina_tumida **       

Pfeiffer et al., 2019 [Publ 3937], fig. 4-5

iT tT bi riS oT roT roS iD
3 4 2 3 54 2 1 26.0
  +----------- Pseudodon_crebristriatus   
--+                                       
  |  +-------- Bineurus_anodontinum **       
  +--+                                    
     |  +----- Pilsbryoconcha_exilis **      
     +--+                                 
        |  +-- Monodontina_vondembuschiana **
        +--+                              
           +-- Pseudodon_inoscularis      

Pfeiffer et al., 2019 [Publ 3937], fig. 6

iT tT bi riS oT roT roS iD
3 4 2 3 54 2 1 24.0
  +----------- Pseudodon_crebristriatus   
--+                                       
  |  +-------- Pilsbryoconcha_exilis **      
  +--+                                    
     |  +----- Bineurus_anodontinum **       
     +--+                                 
        |  +-- Monodontina_vondembuschiana **
        +--+                              
           +-- Pseudodon_inoscularis      

Wu et al., 2021 [Publ 4215], fig. 3

iT tT bi riS oT roT roS iD
3 19 17 3 21 2 1 41.0
                 +----- Chamberlainia_somsakpanhai
  +--------------+                                
  |              |  +-- Sundadontina_cumingii **     
  |              +--+                             
  |                 +-- Sinohyriopsis_schlegelii  
  |                                               
  |                 +-- Pseudodon_bogani          
  |              +--+                             
--+              |  +-- Pseudodon_manueli         
  |  +-----------+                                
  |  |           |  +-- Monodontina_cambodjensis **  
  |  |           +--+                             
  |  |              +-- Pilsbryoconcha_exilis **     
  |  |                                            
  |  |        +-------- Sinosolenaia_carinata     
  +--+        |                                   
     |  +-----+     +-- Lamprotula_caveata        
     |  |     |  +--+                             
     |  |     |  |  +-- Lamprotula_leaii          
     |  |     +--+                                
     |  |        |  +-- Potomida_littoralis       
     |  |        +--+                             
     +--+           +-- Pronodularia_japanensis   
        |                                         
        |           +-- Ptychorhynchus_pfisteri   
        |  +--------+                             
        |  |        +-- Parvasolenaia_rivularis   
        |  |                                      
        +--+        +-- Inversidens_brandtii      
           |  +-----+                             
           |  |     +-- Inversidens_rentianensis  
           +--+                                   
              |  +----- Gonidea_angulata          
              +--+                                
                 |  +-- Microcondylaea_bonellii   
                 +--+                             
                    +-- Leguminaia_wheatleyi      

Bolotov et al., 2022 [Publ 4239], fig. 2

iT tT bi riS oT roT roS iD
3 4 0 3 55 1 1 25.0
  +-- Monodontina_lenyanensis **      
  |                                
  +-- Nyeinchanconcha_nyeinchani **   
--+                                
  +-- Pseudodon_kayinensis         
  |                                
  +-- Sundadontina_tanintharyiensis **

Bolotov et al., 2022 [Publ 4239], fig. S1

iT tT bi riS oT roT roS iD
3 4 0 3 55 1 1 25.0
  +-- Monodontina_lenyanensis **      
  |                                
  +-- Nyeinchanconcha_nyeinchani **   
--+                                
  +-- Pseudodon_kayinensis         
  |                                
  +-- Sundadontina_tanintharyiensis **

Bolotov et al., 2022 [Publ 4239], fig. S2

iT tT bi riS oT roT roS iD
3 4 0 3 55 1 1 25.0
  +-- Monodontina_lenyanensis **      
  |                                
  +-- Nyeinchanconcha_nyeinchani **   
--+                                
  +-- Pseudodon_kayinensis         
  |                                
  +-- Sundadontina_tanintharyiensis **

Bolotov et al., 2022 [Publ 4252], fig. 2

iT tT bi riS oT roT roS iD
3 4 0 3 58 1 1 25.0
  +-- Monodontina_lenyanensis **      
  |                                
  +-- Nyeinchanconcha_nyeinchani **   
--+                                
  +-- Pseudodon_kayinensis         
  |                                
  +-- Sundadontina_tanintharyiensis **

Bolotov et al., 2022 [Publ 4252], fig. S1

iT tT bi riS oT roT roS iD
3 4 0 3 58 1 1 25.0
  +-- Monodontina_lenyanensis **      
  |                                
  +-- Nyeinchanconcha_nyeinchani **   
--+                                
  +-- Pseudodon_kayinensis         
  |                                
  +-- Sundadontina_tanintharyiensis **

Graf & Cummings, 2006 [Publ 1968], fig. 2

iT tT bi riS oT roT roS iD
2 2 0 2 45 0 0 --
  +-- Pilsbryoconcha_exilis **      
--+                              
  +-- Monodontina_vondembuschiana **

Graf, 2002 [Publ 2651], fig. 2

iT tT bi riS oT roT roS iD
2 2 0 2 16 0 0 --
  +-- Pilsbryoconcha_exilis **      
--+                              
  +-- Monodontina_vondembuschiana **

Hoeh et al., 2009 [Publ 2716], fig. 3

iT tT bi riS oT roT roS iD
2 2 0 2 45 0 0 --
  +-- Pilsbryoconcha_exilis **      
--+                              
  +-- Monodontina_vondembuschiana **

Hoeh et al., 2009 [Publ 2716], fig. 4

iT tT bi riS oT roT roS iD
2 2 0 2 44 0 0 --
  +-- Pilsbryoconcha_exilis **      
--+                              
  +-- Monodontina_vondembuschiana **

Hoeh et al., 2009 [Publ 2716], fig. 5

iT tT bi riS oT roT roS iD
2 2 0 2 44 0 0 --
  +-- Pilsbryoconcha_exilis **      
--+                              
  +-- Monodontina_vondembuschiana **

Pfeiffer & Graf, 2013 [Publ 3531], fig. 2

iT tT bi riS oT roT roS iD
2 2 0 2 33 0 0 --
  +-- Monodontina_cambodjensis **
--+                           
  +-- Sundadontina_tumida **     

Froufe et al., 2019 [Publ 3936], fig. 3

iT tT bi riS oT roT roS iD
2 2 0 2 43 0 0 --
  +-- Pilsbryoconcha_exilis **      
--+                              
  +-- Monodontina_vondembuschiana **

Froufe et al., 2019 [Publ 3936], fig. 3-M

iT tT bi riS oT roT roS iD
2 2 0 2 17 0 0 --
  +-- Pilsbryoconcha_exilis **      
--+                              
  +-- Monodontina_vondembuschiana **

Froufe et al., 2019 [Publ 3936], fig. 4

iT tT bi riS oT roT roS iD
2 2 0 2 17 0 0 --
  +-- Pilsbryoconcha_exilis **      
--+                              
  +-- Monodontina_vondembuschiana **

Froufe et al., 2019 [Publ 3936], fig. 6

iT tT bi riS oT roT roS iD
2 2 0 2 43 0 0 --
  +-- Pilsbryoconcha_exilis **      
--+                              
  +-- Monodontina_vondembuschiana **

Fukata & Iigo, 2020 [Publ 4106], fig. 1

iT tT bi riS oT roT roS iD
2 2 0 2 52 0 0 --
  +-- Pilsbryoconcha_exilis **      
--+                              
  +-- Monodontina_vondembuschiana **

Chen et al., 2020 [Publ 4170], fig. 1

iT tT bi riS oT roT roS iD
2 2 0 2 32 0 0 --
  +-- Monodontina_vondembuschiana **
--+                              
  +-- Pilsbryoconcha_exilis **      

Zieritz et al., 2021 [Publ 4231], fig. 3-F, 5

iT tT bi riS oT roT roS iD
2 2 0 2 47 0 0 --
  +-- Pilsbryoconcha_exilis **      
--+                              
  +-- Monodontina_vondembuschiana **

Zieritz et al., 2021 [Publ 4231], fig. 3-M, 4

iT tT bi riS oT roT roS iD
2 2 0 2 21 0 0 --
  +-- Pilsbryoconcha_exilis **      
--+                              
  +-- Monodontina_vondembuschiana **

Wu et al., 2022 [Publ 4238], fig. 3

iT tT bi riS oT roT roS iD
2 2 0 2 72 0 0 --
  +-- Pilsbryoconcha_exilis **      
--+                              
  +-- Monodontina_vondembuschiana **

Bolotov et al., 2022 [Publ 4250], fig. 8

iT tT bi riS oT roT roS iD
2 2 0 2 48 5 1 --
              +-- Monodontina_lenyanensis **      
  +-----------+                                
  |           +-- Sundadontina_tanintharyiensis **
--+                                            
  |  +----------- Pseudodon_kayinensis         
  +--+                                         
     |  +-------- Pseudodon_inoscularis        
     +--+                                      
        |  +----- Pseudodon_bogani             
        +--+                                   
           |  +-- Pseudodon_salwenianus        
           +--+                                
              +-- Pseudodon_manueli            

Pavan-Kumar et al., 2022 [Publ 4263], fig. 4

iT tT bi riS oT roT roS iD
2 2 0 2 34 0 0 --
  +-- Monodontina_vondembuschiana **
--+                              
  +-- Pilsbryoconcha_exilis **      

Pavan-Kumar et al., 2022 [Publ 4263], fig. S5

iT tT bi riS oT roT roS iD
2 2 0 2 37 0 0 --
  +-- Monodontina_vondembuschiana **
--+                              
  +-- Pilsbryoconcha_exilis **      

Dai et al., 2023 [Publ 4285], fig. 5

iT tT bi riS oT roT roS iD
2 2 0 2 51 0 0 --
  +-- Monodontina_vondembuschiana **
--+                              
  +-- Pilsbryoconcha_exilis **      

Kongim et al., 2023 [Publ 4290], fig. 1

iT tT bi riS oT roT roS iD
2 2 0 2 13 1 1 --
  +----- Pseudodon_inoscularis  
--+                             
  |  +-- Namkongnaia_inkhavilayi **
  +--+                          
     +-- Pilsbryoconcha_exilis **  

Walker et al., 2006 [Publ 2666], fig. 2

iT tT bi riS oT roT roS iD
1 1 0 1 14 0 0 --
-- Monodontina_vondembuschiana

Walker et al., 2006 [Publ 2666], fig. 3

iT tT bi riS oT roT roS iD
1 1 0 1 14 0 0 --
-- Monodontina_vondembuschiana

Walker et al., 2006 [Publ 2666], fig. 4

iT tT bi riS oT roT roS iD
1 1 0 1 14 0 0 --
-- Monodontina_vondembuschiana

Walker et al., 2006 [Publ 2666], fig. 6

iT tT bi riS oT roT roS iD
1 1 0 1 37 0 0 --
-- Sundadontina_tumida

Walker et al., 2006 [Publ 2666], fig. 6-M

iT tT bi riS oT roT roS iD
1 1 0 1 14 0 0 --
-- Sundadontina_tumida

Whelan et al., 2011 [Publ 3380], fig. 1 ML

iT tT bi riS oT roT roS iD
1 1 0 1 36 0 0 --
-- Monodontina_vondembuschiana

Whelan et al., 2011 [Publ 3380], fig. 1 MP

iT tT bi riS oT roT roS iD
1 1 0 1 36 0 0 --
-- Monodontina_vondembuschiana

Jeratthitikul et al., 2019 [Publ 3911], fig. 4

iT tT bi riS oT roT roS iD
1 1 0 1 21 0 0 --
-- Pilsbryoconcha_exilis

Zieritz et al., 2020 [Publ 4102], fig. 4

iT tT bi riS oT roT roS iD
1 1 0 1 30 1 1 --
  +-- Pilsbryoconcha_exilis **
--+                        
  +-- Pseudodon_inoscularis

Page updated: 10:53:20 Wed 27 Sep 2023

 
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