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Page last updated
Sat 03 May 2025

The Freshwater Mussels (Unionoida) of the World (and other less consequential bivalves)

FM(U)otW(aolcb) is the web version of the MUSSEL Project Database. Follow the links to browse the data or use the custom Google search field. Either way, you win!

Cladomics pages are experimental — even more so than the rest of this site!

The purpose of the cladomics pages for each genus and family-group level taxon is to provide statistics and references indicating the extent to which published phylogenetic work supports the classification used here.

The Cladomics Summary provides an overview of the number of studies that have included terminal species representing this taxon as well as an indication of how frequently it has been recovered as monophyletic.

Following the summary, the available published clades representing the taxon are listed.

For more information, visit the Cladomics FARQ (frequently asked rhetorical questions) page. Abbreviations for the various statistics are explained on the FARQ!

Search the database pages:

Bivalvia | Unionoida | Unionoidea | Unionidae | Gonideinae

tribe PSEUDODONTINI Frierson, 1927: cladomics
     2 subtribes

Cladomics Summary

number of species = 52
number of subtaxa (rank: subtribe) = 2

analyzed in 92 cladograms in 54 publications
maximum number of terminals in an analysis = 43, mean = 8.4
maximum number of subtaxa (subtribe) in an analysis = 2, mean = 1.5

Basic Monophyly

number of cladograms that test basic monophyly (i.e., more than 1 included species) = 82
maximum number of included terminals = 43, mean = 9.3
maximum number of subtaxa (subtribe) = 2, mean = 1.56
proportion of cladograms that support basic monophyly = 0.99

Rigorous Monophyly

number of cladograms that test rigorous monophyly = 37
(Rigorous monophyly refers to cladograms with terminal species representing more than one subtaxon (subtribe) taxon and at least one rigorous outgroup (tribe).
maximum number of included terminals = 43, mean = 13.84
maximum number of subtaxa (subtribe) = 2, mean = 2.0
proportion of cladograms that support rigorous monophyly = 0.97

Pairwise Cladogram Clustering

The graphs below depict in two dimensions all pair-wise ingroup distances within the cladome. See the Cladmoics FARQ for more information.


PCoA Ordination


NJ tree

Zieritz et al., 2024 [Publ 4381], fig. 2

iT tT bi riS oT roT roS iD
10 10 8 2 36 25 8 0.0

*** Top Monophyly Cladogram *** *** Top Sister-Group Cladogram ***

        +----- Ctenodesmini     
  +-----+                       
  |     |  +-- Rectidentini     
  |     +--+                    
  |        +-- Contradentini    
--+                             
  |     +----- Caudiculatus     
  |  +--+                       
  |  |  |  +-- Lamprotulini     
  |  |  +--+                    
  +--+     +-- Gonideini        
     |                          
     |  +----- Schepmaniini     
     +--+                       
        |  +-- Indopseudodontina **
        +--+                    
           +-- Pseudodontina **

Jeratthitikul et al., 2025 [Publ 4423], fig. 1

iT tT bi riS oT roT roS iD
43 43 42 2 7 7 3 40.0 
        +-- Lamprotulini     
  +-----+                    
  |     +-- Gonideini        
--+                          
  |  +----- Schepmaniini     
  +--+                       
     |  +-- Indopseudodontina **
     +--+                    
        +-- Pseudodontina **

Bolotov et al., 2023 [Publ 4312], fig. S2

iT tT bi riS oT roT roS iD
41 41 39 2 0 0 0 36.0 
  +-- Indopseudodontina **
--+                    
  +-- Pseudodontina **

Jeratthitikul et al., 2024 [Publ 4387], fig. 1

iT tT bi riS oT roT roS iD
40 40 41 2 18 18 7 37.0 
        +-------- Chamberlainiini  
  +-----+                          
  |     |  +----- Ctenodesmini     
  |     +--+                       
  |        |  +-- Rectidentini     
--+        +--+                    
  |           +-- Contradentini    
  |                                
  |  +----------- Lamprotulini     
  +--+                             
     |  +-------- Gonideini        
     +--+                          
        |  +----- Schepmaniini     
        +--+                       
           |  +-- Indopseudodontina **
           +--+                    
              +-- Pseudodontina **

Bolotov et al., 2023 [Publ 4312], fig. 2

iT tT bi riS oT roT roS iD
39 39 38 2 0 0 0 34.0 
  +-- Indopseudodontina **
--+                    
  +-- Pseudodontina **

Bolotov et al., 2023 [Publ 4312], fig. S1

iT tT bi riS oT roT roS iD
39 39 37 2 0 0 0 35.0 
  +-- Indopseudodontina **
--+                    
  +-- Pseudodontina **

Konopleva et al., 2023 [Publ 4262], fig. 8

iT tT bi riS oT roT roS iD
30 30 29 2 52 38 4 25.0 
        +-- Indopseudodontina **
  +-----+                    
  |     +-- Pseudodontina **    
  |                          
--+     +-- Gonideini        
  |  +--+                    
  |  |  +-- Lamprotulini     
  +--+                       
     |  +-- Rectidentini     
     +--+                    
        +-- Contradentini

Konopleva et al., 2021 [Publ 4213], fig. 1

iT tT bi riS oT roT roS iD
23 23 21 2 4 4 2 20.0 
  +----- Lamprotula       
  |                       
  +----- Leguminaia       
  |                       
--+----- Potomida         
  |                       
  +----- Gonidea          
  |                       
  |  +-- Indopseudodontina **
  +--+                    
     +-- Pseudodontina **

Jeratthitikul et al., 2021 [Publ 4219], fig. 2

iT tT bi riS oT roT roS iD
22 22 20 2 14 14 5 22.0 
           +-- Contradentini    
  +--------+                    
  |        +-- Rectidentini     
  |                             
--+        +-- Indopseudodontina **
  |  +-----+                    
  |  |     +-- Pseudodontina **    
  +--+                          
     |  +----- Gonideini        
     +--+                       
        |  +-- Chamberlainiini  
        +--+                    
           +-- Lamprotulini

Jeratthitikul et al., 2021 [Publ 4219], fig. 1

iT tT bi riS oT roT roS iD
22 22 19 2 14 14 5 22.0 
           +-- Contradentini    
  +--------+                    
  |        +-- Rectidentini     
  |                             
--+        +-- Indopseudodontina **
  |  +-----+                    
  |  |     +-- Pseudodontina **    
  +--+                          
     |  +----- Gonideini        
     +--+                       
        |  +-- Chamberlainiini  
        +--+                    
           +-- Lamprotulini

Bolotov et al., 2020 [Publ 4100], fig. S1

iT tT bi riS oT roT roS iD
21 21 20 2 91 51 5 22.0 
           +-- Rectidentini     
  +--------+                    
  |        +-- Contradentini    
  |                             
--+        +-- Indopseudodontina **
  |  +-----+                    
  |  |     +-- Pseudodontina **    
  +--+                          
     |  +----- Gonideini        
     +--+                       
        |  +-- Chamberlainiini  
        +--+                    
           +-- Lamprotulini

Bolotov et al., 2020 [Publ 4100], fig. 1

iT tT bi riS oT roT roS iD
21 21 20 2 97 51 5 22.0 
        +-- Rectidentini             
  +-----+                            
  |     +-- Contradentini            
  |                                  
  |     +-- Indopseudodontina **        
  |  +--+                            
  |  |  +-- Pseudodontina **            
--+  |                               
  |  |  +-- Sinosolenaia_iridinea    
  |  |  |                            
  |  |  +-- Sinosolenaia_recognita__1
  |  |  |                            
  |  |  +-- Sinosolenaia_carinata    
  |  |  |                            
  +--+  +-- Leguminaia               
     |  |                            
     |  +-- Microcondylaea           
     |  |                            
     |  +-- Gonidea                  
     |  |                            
     |  +-- Ptychorhynchus           
     |  |                            
     |  +-- Parvasolenaia            
     |  |                            
     +--+-- Lamprotula_leaii         
        |                            
        +-- Lamprotula               
        |                            
        +-- Lamprotula_caveata       
        |                            
        +-- Potomida                 
        |                            
        +-- Pronodularia             
        |                            
        +-- Sinohyriopsis            
        |                            
        +-- Chamberlainia            
        |                            
        +-- Sinosolenaia_recognita__2
        |                            
        +-- Sinosolenaia_oleivora

Jeratthitikul et al., 2022 [Publ 4260], fig. S2_2

iT tT bi riS oT roT roS iD
17 17 15 2 59 14 5 17.0 
  +-- Indopseudodontina **
--+                    
  +-- Pseudodontina **

Jeratthitikul et al., 2022 [Publ 4260], fig. S1_1

iT tT bi riS oT roT roS iD
17 17 15 2 59 14 5 17.0 
  +-- Indopseudodontina **
--+                    
  +-- Pseudodontina **

Dai et al., 2023 [Publ 4285], fig. 3-BI

iT tT bi riS oT roT roS iD
17 17 14 2 28 26 5 18.0 
                       +-- Indopseudodontina **
  +--------------------+                    
  |                    +-- Pseudodontina **    
  |                                         
--+                    +-- Rectidentini     
  |  +-----------------+                    
  |  |                 +-- Contradentini    
  |  |                                      
  +--+  +----------------- Inversidens      
     |  |                                   
     |  |              +-- Chamberlainiini  
     |  +--------------+                    
     +--+              +-- Lamprotulini     
        |                                   
        |  +-------------- Sinosolenaia     
        |  |                                
        |  |        +----- Gonidea          
        +--+  +-----+                       
           |  |     |  +-- Leguminaia       
           |  |     +--+                    
           |  |        +-- Microcondylaea   
           +--+                             
              |        +-- Koreosolenaia    
              |  +-----+                    
              |  |     +-- Parvasolenaia    
              +--+                          
                 |  +----- Postolata        
                 +--+                       
                    |  +-- Ptychorhynchus   
                    +--+                    
                       +-- Obovalis

Dai et al., 2023 [Publ 4285], fig. 3-ML

iT tT bi riS oT roT roS iD
17 17 15 2 28 26 5 19.0 
                       +-- Indopseudodontina **
  +--------------------+                    
  |                    +-- Pseudodontina **    
  |                                         
  |                    +-- Rectidentini     
--+  +-----------------+                    
  |  |                 +-- Contradentini    
  |  |                                      
  |  |              +----- Inversidens      
  +--+  +-----------+                       
     |  |           |  +-- Chamberlainiini  
     |  |           +--+                    
     |  |              +-- Lamprotulini     
     +--+                                   
        |  +-------------- Sinosolenaia     
        |  |                                
        |  |        +----- Gonidea          
        +--+  +-----+                       
           |  |     |  +-- Leguminaia       
           |  |     +--+                    
           |  |        +-- Microcondylaea   
           +--+                             
              |        +-- Koreosolenaia    
              |  +-----+                    
              |  |     +-- Parvasolenaia    
              +--+                          
                 |  +----- Postolata        
                 +--+                       
                    |  +-- Ptychorhynchus   
                    +--+                    
                       +-- Obovalis

Huang et al., 2019 [Publ 3843], fig. S3

iT tT bi riS oT roT roS iD
15 15 16 2 182 40 5 17.0 
  +-- Indopseudodontina **
--+                    
  +-- Pseudodontina **

Huang et al., 2019 [Publ 3843], fig. 2

iT tT bi riS oT roT roS iD
15 15 15 2 184 41 5 19.0 
  +----------- Chamberlainiini  
--+                             
  |  +-------- Lamprotulini     
  +--+                          
     |  +----- Gonideini        
     +--+                       
        |  +-- Indopseudodontina **
        +--+                    
           +-- Pseudodontina **

Huang et al., 2019 [Publ 3843], fig. S4

iT tT bi riS oT roT roS iD
15 15 9 2 184 41 5 17.0 
  +-- Indopseudodontina **
--+                    
  +-- Pseudodontina **

Bolotov et al., 2017 [Publ 3764], fig. S1

iT tT bi riS oT roT roS iD
15 15 13 2 110 36 5 18.0 
  +-- Indopseudodontina **
--+                    
  +-- Pseudodontina **

Bolotov et al., 2017 [Publ 3761], fig. S2

iT tT bi riS oT roT roS iD
14 14 13 2 125 35 5 19.0 
              +----- Gonideini                 
  +-----------+                                
  |           |  +-- Chamberlainiini           
  |           +--+                             
  |              +-- Lamprotulini              
--+                                            
  |  +-------------- Pilsbryoconcha **            
  |  |                                         
  |  |  +----------- Indopseudodontina **         
  +--+  |                                      
     |  |        +-- Thaiconcha **                
     |  +--------+                             
     |  |        +-- Bineurus **                  
     +--+                                      
        |     +----- Pseudodon_vagulus **         
        |  +--+                                
        |  |  |  +-- Pseudodon_mekongi **         
        |  |  +--+                             
        |  |     +-- Pseudodon_vondembuschianus **
        +--+                                   
           |     +-- Pseudodon_inoscularis **     
           |  +--+                             
           |  |  +-- Sundadontina_tumida **       
           +--+                                
              |  +-- Nyeinchanconcha **           
              +--+                             
                 +-- Sundadontina_cumingii **

Bolotov et al., 2017 [Publ 3761], fig. S1

iT tT bi riS oT roT roS iD
13 13 13 2 126 36 5 15.0 
  +-- Indopseudodontina **
--+                    
  +-- Pseudodontina **

Jeratthitikul & Sutcharit, 2023 [Publ 4289], fig. 1

iT tT bi riS oT roT roS iD
13 13 11 2 0 0 0 16.0 
  +-- Indopseudodontina **
--+                    
  +-- Pseudodontina **

Bolotov et al., 2018 [Publ 3826], fig. 2

iT tT bi riS oT roT roS iD
11 11 2 2 72 24 4 13.0 
  +-- Indopseudodontina **
--+                    
  +-- Pseudodontina **

Konopleva et al., 2019 [Publ 3900], fig. 1

iT tT bi riS oT roT roS iD
11 11 2 2 41 27 2 13.0 
  +-- Indopseudodontina **
--+                    
  +-- Pseudodontina **

Dai et al., 2024 [Publ 4389], fig. 3

iT tT bi riS oT roT roS iD
11 11 9 1 36 31 7 9.0 
           +----- Ctenodesmini   
  +--------+                     
  |        |  +-- Rectidentini   
  |        +--+                  
--+           +-- Contradentini  
  |                              
  |  +----------- Chamberlainiini
  +--+                           
     |  +-------- Lamprotulini   
     +--+                        
        |  +----- Gonideini      
        +--+                     
           |  +-- Schepmaniini   
           +--+                  
              +-- Pseudodontina **

Zieritz et al., 2021 [Publ 4218], fig. 5

iT tT bi riS oT roT roS iD
9 9 7 2 23 19 7 3.0 
  +----------------- Chamberlainiini  
  |                                   
--+  +-------------- Lamprotulini     
  |  |                                
  |  |        +----- Ctenodesmini     
  +--+  +-----+                       
     |  |     |  +-- Rectidentini     
     |  |     +--+                    
     +--+        +-- Contradentini    
        |                             
        |  +-------- Gonideini        
        +--+                          
           |  +----- Schepmaniini     
           +--+                       
              |  +-- Indopseudodontina **
              +--+                    
                 +-- Pseudodontina **

Wu et al., 2024 [Publ 4350], fig. 6-7

iT tT bi riS oT roT roS iD
8 8 6 2 29 27 7 4.0 
  +-------------- Chamberlainiini  
  |                                
  |        +----- Ctenodesmini     
--+  +-----+                       
  |  |     |  +-- Rectidentini     
  |  |     +--+                    
  |  |        +-- Contradentini    
  +--+                             
     |        +-- Lamprotulini     
     |  +-----+                    
     |  |     +-- Gonideini        
     +--+                          
        |  +----- Schepmaniini     
        +--+                       
           |  +-- Indopseudodontina **
           +--+                    
              +-- Pseudodontina **

Wu et al., 2024 [Publ 4350], fig. 5b

iT tT bi riS oT roT roS iD
8 8 6 2 29 27 7 6.0 
  +----------------------------- Schepmaniini       
  |                                                 
  |                          +-- Indopseudodontina **  
  |  +-----------------------+                      
--+  |                       +-- Pseudodontina **      
  |  |                                              
  |  |              +----------- Hyriopsis_bialata  
  |  |              |                               
  +--+  +-----------+        +-- Rectidens          
     |  |           |  +-----+                      
     |  |           |  |     +-- Ctenodesmini       
     |  |           +--+                            
     |  |              |  +----- Ensidens           
     |  |              +--+                         
     +--+                 |  +-- Hyriopsis_desowitzi
        |                 +--+                      
        |                    +-- Contradentini      
        |                                           
        |  +-------------------- Inversidens        
        |  |                                        
        |  |              +----- Lamprotula         
        +--+  +-----------+                         
           |  |           |  +-- Chamberlainiini    
           |  |           +--+                      
           +--+              +-- Potomida           
              |                                     
              |  +-------------- Sinosolenaia       
              |  |                                  
              +--+           +-- Gonidea            
                 |  +--------+                      
                 |  |        +-- Microcondylaea     
                 +--+                               
                    |  +-------- Koreosolenaia      
                    +--+                            
                       |  +----- Parvasolenaia      
                       +--+                         
                          |  +-- Postolata          
                          +--+                      
                             +-- Ptychorhynchus

Wu et al., 2024 [Publ 4350], fig. 5a

iT tT bi riS oT roT roS iD
8 8 6 2 29 27 7 4.0 
  +-------------------------- Chamberlainiini  
  |                                            
  |                    +----- Ctenodesmini     
  |  +-----------------+                       
--+  |                 |  +-- Rectidentini     
  |  |                 +--+                    
  |  |                    +-- Contradentini    
  |  |                                         
  +--+                 +----- Schepmaniini     
     |  +--------------+                       
     |  |              |  +-- Indopseudodontina **
     |  |              +--+                    
     |  |                 +-- Pseudodontina **    
     +--+                                      
        |                 +-- Inversidens      
        |  +--------------+                    
        |  |              +-- Lamprotulini     
        +--+                                   
           |  +-------------- Sinosolenaia     
           |  |                                
           +--+           +-- Gonidea          
              |  +--------+                    
              |  |        +-- Microcondylaea   
              +--+                             
                 |  +-------- Parvasolenaia    
                 +--+                          
                    |  +----- Koreosolenaia    
                    +--+                       
                       |  +-- Postolata        
                       +--+                    
                          +-- Ptychorhynchus

Wu et al., 2024 [Publ 4350], fig. 5c

iT tT bi riS oT roT roS iD
8 8 5 2 29 27 7 7.0 
  +----------------------------- Schepmania_nieuwenhuisi__1
  |                                                        
  +----------------------------- Schepmania_nieuwenhuisi__2
  |                                                        
  |                          +-- Indopseudodontina **         
--+  +-----------------------+                             
  |  |                       +-- Pseudodontina **             
  |  |                                                     
  |  |                       +-- Ensidens                  
  |  |                       |                             
  |  |                       +-- Hyriopsis_desowitzi       
  +--+                       |                             
     |                       +-- Rectidens                 
     |  +--------------------+                             
     |  |                    +-- Hyriopsis_bialata         
     |  |                    |                             
     |  |                    +-- Ctenodesmini              
     |  |                    |                             
     +--+                    +-- Contradentini             
        |                                                  
        |  +-------------------- Inversidens               
        |  |                                               
        |  |              +----- Lamprotula                
        +--+  +-----------+                                
           |  |           |  +-- Chamberlainiini           
           |  |           +--+                             
           +--+              +-- Potomida                  
              |                                            
              |  +-------------- Sinosolenaia              
              |  |                                         
              +--+  +----------- Gonidea                   
                 |  |                                      
                 +--+----------- Microcondylaea            
                    |                                      
                    |  +-------- Koreosolenaia             
                    +--+                                   
                       |  +----- Parvasolenaia             
                       +--+                                
                          |  +-- Postolata                 
                          +--+                             
                             +-- Ptychorhynchus

Lopes-Lima et al., 2020 [Publ 4080], fig. 2-4

iT tT bi riS oT roT roS iD
8 8 5 2 113 25 5 13.0 
        +-- Chamberlainiini  
  +-----+                    
  |     +-- Lamprotulini     
--+                          
  |  +----- Gonideini        
  +--+                       
     |  +-- Indopseudodontina **
     +--+                    
        +-- Pseudodontina **

Zieritz et al., 2024 [Publ 4381], fig. 8A

iT tT bi riS oT roT roS iD
8 8 6 1 0 0 0 12.0 
-- (Pseudodontina

Wu et al., 2024 [Publ 4388], fig. 5

iT tT bi riS oT roT roS iD
8 8 6 2 30 28 7 4.0 
  +----------------- Chamberlainiini  
  |                                   
  |           +----- Ctenodesmini     
--+  +--------+                       
  |  |        |  +-- Rectidentini     
  |  |        +--+                    
  +--+           +-- Contradentini    
     |                                
     |  +----------- Lamprotulini     
     +--+                             
        |  +-------- Gonideini        
        +--+                          
           |  +----- Schepmaniini     
           +--+                       
              |  +-- Indopseudodontina **
              +--+                    
                 +-- Pseudodontina **

Liu et al., 2024 [Publ 4380], fig. 5

iT tT bi riS oT roT roS iD
8 8 6 2 29 27 7 4.0 
           +----- Schepmaniini     
  +--------+                       
  |        |  +-- Indopseudodontina **
  |        +--+                    
--+           +-- Pseudodontina **    
  |                                
  |  +----------- Gonideini        
  |  |                             
  +--+        +-- Chamberlainiini  
     |  +-----+                    
     |  |     +-- Lamprotulini     
     +--+                          
        |  +----- Ctenodesmini     
        +--+                       
           |  +-- Rectidentini     
           +--+                    
              +-- Contradentini

Bolotov et al., 2022 [Publ 4250], fig. 8

iT tT bi riS oT roT roS iD
6 6 4 2 43 17 3 9.0 
  +----- Contradentini    
--+                       
  |  +-- Pseudodontina **    
  +--+                    
     +-- Indopseudodontina **

Wu et al., 2021 [Publ 4366], fig. 6

iT tT bi riS oT roT roS iD
6 6 4 2 85 23 5 11.0 
  +----------- Chamberlainiini  
--+                             
  |  +-------- Lamprotulini     
  +--+                          
     |  +----- Gonideini        
     +--+                       
        |  +-- Indopseudodontina **
        +--+                    
           +-- Pseudodontina **

Konopleva et al., 2017 [Publ 3807], fig. 6

iT tT bi riS oT roT roS iD
6 6 4 1 25 4 2 13.0 
  +----- Pseudodontina **
--+                   
  |  +-- Rectidentini 
  +--+                
     +-- Contradentini

Wu et al., 2021 [Publ 4366], fig. 2_I

iT tT bi riS oT roT roS iD
6 6 4 2 85 23 5 11.0 
  +----------- Chamberlainiini  
--+                             
  |  +-------- Lamprotulini     
  +--+                          
     |  +----- Gonideini        
     +--+                       
        |  +-- Indopseudodontina **
        +--+                    
           +-- Pseudodontina **

Pfeiffer & Graf, 2015 [Publ 3620], fig. 2

iT tT bi riS oT roT roS iD
6 6 4 1 34 8 4 13.0 
        +-- Rectidentini 
  +-----+                
  |     +-- Contradentini
--+                      
  |  +----- Pseudodontina **
  +--+                   
     |  +-- Gonideini    
     +--+                
        +-- Lamprotulini

Pfeiffer et al., 2019 [Publ 3937], fig. 4-5

iT tT bi riS oT roT roS iD
5 5 3 2 52 12 5 9.0 
  +-------------- Chamberlainiini  
  |                                
--+           +-- Rectidentini     
  |  +--------+                    
  |  |        +-- Contradentini    
  +--+                             
     |  +-------- Lamprotulini     
     +--+                          
        |  +----- Gonideini        
        +--+                       
           |  +-- Indopseudodontina **
           +--+                    
              +-- Pseudodontina **

Lopes-Lima et al., 2017 [Publ 3745], fig. 1-2

iT tT bi riS oT roT roS iD
5 5 3 1 75 22 5 13.0 
              +-- Chamberlainiini
  +-----------+                  
  |           +-- Lamprotulini   
  |                              
--+           +-- Parvasolenaia  
  |  +--------+                  
  |  |        +-- Pseudodontina **  
  +--+                           
     |  +-------- Sinosolenaia   
     +--+                        
        |  +----- Gonidea        
        +--+                     
           |  +-- Microcondylaea 
           +--+                  
              +-- Leguminaia

Wu et al., 2021 [Publ 4215], fig. 3

iT tT bi riS oT roT roS iD
5 19 17 2 19 14 3 26.0 
                 +----- Chamberlainia    
  +--------------+                       
  |              |  +-- Sundadontina **     
  |              +--+                    
  |                 +-- Sinohyriopsis    
  |                                      
--+              +----- Indopseudodontina **
  |  +-----------+                       
  |  |           |  +-- Pseudodon **        
  |  |           +--+                    
  |  |              +-- Pilsbryoconcha **   
  +--+                                   
     |              +-- Sinosolenaia     
     |  +-----------+                    
     |  |           +-- Lamprotulini     
     |  |                                
     +--+           +-- Ptychorhynchus   
        |  +--------+                    
        |  |        +-- Parvasolenaia    
        +--+                             
           |  +-------- Inversidens      
           +--+                          
              |  +----- Gonidea          
              +--+                       
                 |  +-- Microcondylaea   
                 +--+                    
                    +-- Leguminaia

Pfeiffer et al., 2019 [Publ 3937], fig. 6

iT tT bi riS oT roT roS iD
5 5 3 2 52 12 5 9.0 
  +----------- Chamberlainiini  
  |                             
  |        +-- Rectidentini     
--+  +-----+                    
  |  |     +-- Contradentini    
  |  |                          
  +--+     +-- Gonideini        
     |  +--+                    
     |  |  +-- Lamprotulini     
     +--+                       
        |  +-- Indopseudodontina **
        +--+                    
           +-- Pseudodontina **

Bolotov et al., 2022 [Publ 4239], fig. S2

iT tT bi riS oT roT roS iD
4 4 0 2 54 0 0 10.0 
  +-- Pseudodon **        
  |                    
  +-- Nyeinchanconcha **  
--+                    
  +-- Indopseudodontina **
  |                    
  +-- Sundadontina **

Bolotov et al., 2022 [Publ 4252], fig. S1

iT tT bi riS oT roT roS iD
4 4 0 2 57 0 0 10.0 
  +-- Pseudodon **        
  |                    
  +-- Nyeinchanconcha **  
--+                    
  +-- Indopseudodontina **
  |                    
  +-- Sundadontina **

Bolotov et al., 2022 [Publ 4239], fig. S1

iT tT bi riS oT roT roS iD
4 4 0 2 54 0 0 10.0 
  +-- Pseudodon **        
  |                    
  +-- Nyeinchanconcha **  
--+                    
  +-- Indopseudodontina **
  |                    
  +-- Sundadontina **

Bolotov et al., 2022 [Publ 4252], fig. 2

iT tT bi riS oT roT roS iD
4 4 0 2 57 0 0 10.0 
  +-- Pseudodon **        
  |                    
  +-- Nyeinchanconcha **  
--+                    
  +-- Indopseudodontina **
  |                    
  +-- Sundadontina **

Bolotov et al., 2022 [Publ 4239], fig. 2

iT tT bi riS oT roT roS iD
4 4 0 2 54 0 0 10.0 
  +-- Pseudodon **        
  |                    
  +-- Nyeinchanconcha **  
--+                    
  +-- Indopseudodontina **
  |                    
  +-- Sundadontina **

Konopleva et al., 2019 [Publ 4041], fig. 3

iT tT bi riS oT roT roS iD
3 3 0 1 31 31 2 11.0 
  +----- Indopseudodontina **
--+                       
  |  +-- Rectidentini     
  +--+                    
     +-- Contradentini

Kongim et al., 2023 [Publ 4290], fig. 1

iT tT bi riS oT roT roS iD
3 3 1 2 12 12 3 10.0 
  +----------- Chamberlainiini  
--+                             
  |  +-------- Lamprotulini     
  +--+                          
     |  +----- Gonideini        
     +--+                       
        |  +-- Indopseudodontina **
        +--+                    
           +-- Pseudodontina **

Hoeh et al., 2009 [Publ 2716], fig. 3

iT tT bi riS oT roT roS iD
2 2 0 1 45 2 2 -- 
  +-- Gonideini    
--+                
  +-- Pseudodontina **

Graf & Cummings, 2006 [Publ 1968], fig. 2

iT tT bi riS oT roT roS iD
2 2 0 1 45 2 2 -- 
-- (Pseudodontina

Fukata & Iigo, 2020 [Publ 4106], fig. 1

iT tT bi riS oT roT roS iD
2 2 0 1 52 10 3 -- 
  +-------- Chamberlainiini
--+                        
  |  +----- Pseudodontina **  
  +--+                     
     |  +-- Lamprotulini   
     +--+                  
        +-- Gonideini

Froufe et al., 2019 [Publ 3936], fig. 6

iT tT bi riS oT roT roS iD
2 2 0 1 43 11 3 -- 
  +-------- Chamberlainiini
--+                        
  |  +----- Lamprotulini   
  +--+                     
     |  +-- Pseudodontina **  
     +--+                  
        +-- Gonideini

Graf, 2002 [Publ 2651], fig. 2

iT tT bi riS oT roT roS iD
2 2 0 1 16 2 2 -- 
-- (Pseudodontina

Froufe et al., 2019 [Publ 3936], fig. 3-M

iT tT bi riS oT roT roS iD
2 2 0 1 17 6 3 -- 
  +-------- Chamberlainiini
--+                        
  |  +----- Lamprotulini   
  +--+                     
     |  +-- Gonideini      
     +--+                  
        +-- Pseudodontina **

Wu et al., 2024 [Publ 4350], fig. 8a_S1

iT tT bi riS oT roT roS iD
2 2 0 1 90 19 5 -- 
  +----------- Chamberlainiini
  |                           
--+        +-- Rectidentini   
  |  +-----+                  
  |  |     +-- Contradentini  
  +--+                        
     |  +----- Lamprotulini   
     +--+                     
        |  +-- Pseudodontina **  
        +--+                  
           +-- Gonideini

Matos et al., 2024 [Publ 4384], fig. 3

iT tT bi riS oT roT roS iD
2 2 0 1 18 16 5 -- 
  +----------- Chamberlainiini
  |                           
--+        +-- Contradentini  
  |  +-----+                  
  |  |     +-- Rectidentini   
  +--+                        
     |  +----- Lamprotulini   
     +--+                     
        |  +-- Pseudodontina **  
        +--+                  
           +-- Gonideini

Pavan-Kumar et al., 2022 [Publ 4263], fig. 4

iT tT bi riS oT roT roS iD
2 2 0 1 34 9 3 -- 
  +-------- Chamberlainiini
--+                        
  |  +----- Lamprotulini   
  +--+                     
     |  +-- Pseudodontina **  
     +--+                  
        +-- Gonideini

Zieritz et al., 2020 [Publ 4102], fig. 4

iT tT bi riS oT roT roS iD
2 2 0 2 29 29 6 -- 
  +----------------- Chamberlainiini  
  |                                   
--+  +-------------- Lamprotulini     
  |  |                                
  +--+  +----------- Gonideini        
     |  |                             
     +--+        +-- Pseudodontina **    
        |  +-----+                    
        |  |     +-- Indopseudodontina **
        +--+                          
           |  +----- Contradentini    
           +--+                       
              |  +-- Ctenodesmini     
              +--+                    
                 +-- Rectidentini

Chen et al., 2020 [Publ 4170], fig. 1

iT tT bi riS oT roT roS iD
2 2 0 1 32 9 3 -- 
  +----- Pseudodontina **  
--+                     
  |  +-- Lamprotulini   
  +--+                  
     +-- Chamberlainiini

Wu et al., 2024 [Publ 4350], fig. 8b_S2

iT tT bi riS oT roT roS iD
2 2 0 1 90 19 5 -- 
  +----------- Chamberlainiini
  |                           
--+        +-- Rectidentini   
  |  +-----+                  
  |  |     +-- Contradentini  
  +--+                        
     |  +----- Lamprotulini   
     +--+                     
        |  +-- Pseudodontina **  
        +--+                  
           +-- Gonideini

Zieritz et al., 2021 [Publ 4231], fig. 3-M_4

iT tT bi riS oT roT roS iD
2 2 0 1 21 10 5 -- 
  +----------- Chamberlainiini
  |                           
--+        +-- Contradentini  
  |  +-----+                  
  |  |     +-- Rectidentini   
  +--+                        
     |  +----- Pseudodontina **  
     +--+                     
        |  +-- Lamprotulini   
        +--+                  
           +-- Gonideini

Hoeh et al., 2009 [Publ 2716], fig. 5

iT tT bi riS oT roT roS iD
2 2 0 1 44 2 2 -- 
-- (Pseudodontina

Dai et al., 2023 [Publ 4285], fig. 5

iT tT bi riS oT roT roS iD
2 2 0 1 51 17 5 -- 
  +----------- Chamberlainiini
  |                           
--+        +-- Rectidentini   
  |  +-----+                  
  |  |     +-- Contradentini  
  +--+                        
     |  +----- Lamprotulini   
     +--+                     
        |  +-- Pseudodontina **  
        +--+                  
           +-- Gonideini

Froufe et al., 2019 [Publ 3936], fig. 4

iT tT bi riS oT roT roS iD
2 2 0 1 17 6 3 -- 
  +-------- Chamberlainiini
--+                        
  |  +----- Lamprotulini   
  +--+                     
     |  +-- Gonideini      
     +--+                  
        +-- Pseudodontina **

Wu et al., 2021 [Publ 4366], fig. 4

iT tT bi riS oT roT roS iD
2 2 0 1 67 11 3 -- 
  +-------- Chamberlainiini
--+                        
  |  +----- Lamprotulini   
  +--+                     
     |  +-- Pseudodontina **  
     +--+                  
        +-- Gonideini

Wu et al., 2021 [Publ 4366], fig. 3-BI

iT tT bi riS oT roT roS iD
2 2 0 1 72 12 3 -- 
  +-------- Chamberlainiini
--+                        
  |  +----- Lamprotulini   
  +--+                     
     |  +-- Pseudodontina **  
     +--+                  
        +-- Gonideini

Froufe et al., 2019 [Publ 3936], fig. 3

iT tT bi riS oT roT roS iD
2 2 0 1 43 11 3 -- 
  +-------- Chamberlainiini
--+                        
  |  +----- Lamprotulini   
  +--+                     
     |  +-- Pseudodontina **  
     +--+                  
        +-- Gonideini

Wu et al., 2023 [Publ 4339], fig. 4_S5-nt

iT tT bi riS oT roT roS iD
2 2 0 1 94 18 5 -- 
  +----------- Chamberlainiini
  |                           
--+        +-- Rectidentini   
  |  +-----+                  
  |  |     +-- Contradentini  
  +--+                        
     |  +----- Lamprotulini   
     +--+                     
        |  +-- Pseudodontina **  
        +--+                  
           +-- Gonideini

Pfeiffer & Graf, 2013 [Publ 3531], fig. 2

iT tT bi riS oT roT roS iD
2 2 0 1 29 9 4 -- 
  +-------- Chamberlainiini
  |                        
--+     +-- Gonideini      
  |  +--+                  
  |  |  +-- Pseudodontina **  
  +--+                     
     |  +-- Contradentini  
     +--+                  
        +-- Lamprotulini

Dai et al., 2024 [Publ 4358], fig. 4

iT tT bi riS oT roT roS iD
2 2 0 1 50 9 3 -- 
  +-------- Chamberlainiini
--+                        
  |  +----- Lamprotulini   
  +--+                     
     |  +-- Pseudodontina **  
     +--+                  
        +-- Gonideini

Hoeh et al., 2009 [Publ 2716], fig. 4

iT tT bi riS oT roT roS iD
2 2 0 1 44 2 2 -- 
-- (Pseudodontina

Wu et al., 2024 [Publ 4388], fig. 6

iT tT bi riS oT roT roS iD
2 2 0 1 52 21 5 -- 
  +----------- Chamberlainiini
  |                           
--+        +-- Rectidentini   
  |  +-----+                  
  |  |     +-- Contradentini  
  +--+                        
     |  +----- Lamprotulini   
     +--+                     
        |  +-- Pseudodontina **  
        +--+                  
           +-- Gonideini

Wu et al., 2023 [Publ 4339], fig. 4_S5-aa

iT tT bi riS oT roT roS iD
2 2 0 1 94 18 5 -- 
  +----------- Chamberlainiini
  |                           
--+        +-- Rectidentini   
  |  +-----+                  
  |  |     +-- Contradentini  
  +--+                        
     |  +----- Lamprotulini   
     +--+                     
        |  +-- Pseudodontina **  
        +--+                  
           +-- Gonideini

Wu et al., 2021 [Publ 4366], fig. 3-ML

iT tT bi riS oT roT roS iD
2 2 0 1 72 12 3 -- 
  +-------- Chamberlainiini
--+                        
  |  +----- Lamprotulini   
  +--+                     
     |  +-- Pseudodontina **  
     +--+                  
        +-- Gonideini

Zieritz et al., 2021 [Publ 4231], fig. 3-F_5

iT tT bi riS oT roT roS iD
2 2 0 1 47 15 5 -- 
  +----------- Chamberlainiini
  |                           
--+        +-- Contradentini  
  |  +-----+                  
  |  |     +-- Rectidentini   
  +--+                        
     |  +----- Lamprotulini   
     +--+                     
        |  +-- Pseudodontina **  
        +--+                  
           +-- Gonideini

Wu et al., 2022 [Publ 4238], fig. 3

iT tT bi riS oT roT roS iD
2 2 0 1 72 12 3 -- 
  +-------- Chamberlainiini
--+                        
  |  +----- Lamprotulini   
  +--+                     
     |  +-- Pseudodontina **  
     +--+                  
        +-- Gonideini

Matos et al., 2024 [Publ 4383], fig. 3

iT tT bi riS oT roT roS iD
2 2 0 1 18 16 5 -- 
  +----------- Chamberlainiini
  |                           
--+        +-- Contradentini  
  |  +-----+                  
  |  |     +-- Rectidentini   
  +--+                        
     |  +----- Lamprotulini   
     +--+                     
        |  +-- Pseudodontina **  
        +--+                  
           +-- Gonideini

Pavan-Kumar et al., 2022 [Publ 4263], fig. S5

iT tT bi riS oT roT roS iD
2 2 0 1 37 10 3 -- 
  +-------- Chamberlainiini
--+                        
  |  +----- Gonideini      
  +--+                     
     |  +-- Pseudodontina **  
     +--+                  
        +-- Lamprotulini

Zhang et al., 2024 [Publ 4397], fig. 5

iT tT bi riS oT roT roS iD
2 2 0 1 26 24 5 -- 
  +----------- Chamberlainiini
  |                           
--+        +-- Rectidentini   
  |  +-----+                  
  |  |     +-- Contradentini  
  +--+                        
     |  +----- Lamprotulini   
     +--+                     
        |  +-- Pseudodontina **  
        +--+                  
           +-- Gonideini

Walker et al., 2006 [Publ 2666], fig. 6

iT tT bi riS oT roT roS iD
1 1 0 1 35 1 1 -- 
  +-- Gonideini    
--+                
  +-- Pseudodontina **

Walker et al., 2006 [Publ 2666], fig. 4

iT tT bi riS oT roT roS iD
1 1 0 1 14 2 1 -- 
-- Pseudodon_vondembuschianus

Bolotov et al., 2024 [Publ 4369], fig. S1

iT tT bi riS oT roT roS iD
1 1 0 1 62 0 0 -- 
-- Indopseudodon_kayinensis

Walker et al., 2006 [Publ 2666], fig. 2

iT tT bi riS oT roT roS iD
1 1 0 1 14 2 2 -- 
  +-- Gonideini    
--+                
  +-- Pseudodontina **

Walker et al., 2006 [Publ 2666], fig. 3

iT tT bi riS oT roT roS iD
1 1 0 1 14 2 1 -- 
-- Pseudodon_vondembuschianus

Walker et al., 2006 [Publ 2666], fig. 6-M

iT tT bi riS oT roT roS iD
1 1 0 1 13 1 1 -- 
  +-- Gonideini    
--+                
  +-- Pseudodontina **

Whelan et al., 2011 [Publ 3380], fig. 1-ML

iT tT bi riS oT roT roS iD
1 1 0 1 35 5 3 -- 
  +-- Pseudodontina **
--+                
  +-- Contradentini

Whelan et al., 2011 [Publ 3380], fig. 1-MP

iT tT bi riS oT roT roS iD
1 1 0 1 35 5 3 -- 
-- Pseudodon_vondembuschianus

Jeratthitikul et al., 2019 [Publ 3911], fig. 4

iT tT bi riS oT roT roS iD
1 1 0 1 8 7 2 -- 
-- Pilsbryoconcha_exilis

Dai et al., 2024 [Publ 4386], fig. 6

iT tT bi riS oT roT roS iD
1 1 0 1 39 9 3 -- 
  +-------- Chamberlainiini
--+                        
  |  +----- Lamprotulini   
  +--+                     
     |  +-- Pseudodontina **  
     +--+                  
        +-- Gonideini

Page updated: 10:43:39 Sat 03 May 2025
 
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