|
Bivalvia | Unionoida | Unionoidea | Unionidae | Ambleminae tribe QUADRULINI Ihering, 1901: cladomics
1 subtribe, 5 genera
Cladomics Summary
number of species = 47
number of subtaxa (rank: subtribe) = 6
analyzed in 169 cladograms in 104 publications
maximum number of terminals in an analysis = 22, mean = 2.77
maximum number of subtaxa (subtribe) in an analysis = 3, mean = 1.35
Basic Monophyly
number of cladograms that test basic monophyly (i.e., more than 1 included species) = 73
maximum number of included terminals = 22, mean = 5.1
maximum number of subtaxa (subtribe) = 3, mean = 1.81
proportion of cladograms that support basic monophyly = 0.79
Rigorous Monophyly
number of cladograms that test rigorous monophyly = 39
(Rigorous monophyly refers to cladograms with terminal species representing more than one subtaxon (subtribe) taxon and at least one rigorous outgroup (tribe).
maximum number of included terminals = 22, mean = 6.85
maximum number of subtaxa (subtribe) = 3, mean = 2.44
proportion of cladograms that support rigorous monophyly = 0.69
Pairwise Cladogram Clustering
The graphs below depict in two dimensions all pair-wise ingroup distances within the cladome. See the Cladmoics FARQ for more information.
Neemuchwala et al., 2023 [Publ 4292], fig. 2
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 22 |
22 |
20 |
3 |
5 |
4 |
4 |
0.0 |
*** Top Monophyly Cladogram ***
+-------- Pleurobema_strodeanum
+--------------------+
| | +----- Amblema_plicata
| +--+
| | +-- Lampsilis_cardium
| +--+
--+ +-- Popenaias_popeii
|
| +-------------------------- Megalonaias_nervosa **
| |
| | +-------- Uniomerus_carolinianus **
| | +--------------+
+--+ | | +----- Uniomerus_columbensis **
| | +--+
| | | +-- Uniomerus_tetralasmus **
| | +--+
| | +-- Uniomerus_declivis **
+--+
| +-------- Theliderma_cylindrica **
| +-----------+
| | | +----- Theliderma_metanevra **
| | +--+
| | | +-- Theliderma_sparsa **
| | +--+
+--+ +-- Theliderma_johnsoni **
|
| +----------------- Cyclonaias_tuberculata **
| |
| | +-------- Quadrula_nobilis **
| | +-----+
+--+ | | +----- Quadrula_verrucosa **
| | +--+
| | | +-- Quadrula_fragosa **
| | +--+
| | +-- Quadrula_quadrula **
+--+
| +----- Pustulosa_nodulata **
| +-----+
| | | +-- Pustulosa_necki **
| | +--+
| | +-- Pustulosa_petrina **
+--+
| +-- Pustulosa_kieneriana **
| +-----+
| | +-- Pustulosa_pustulosa **
+--+
| +----- Pustulosa_succissa **
+--+
| +-- Pustulosa_infucata **
+--+
+-- Pustulosa_kleiniana **
Inoue et al., 2018 [Publ 3858], fig. 1, 3a-c
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 8 |
8 |
2 |
2 |
197 |
174 |
4 |
19.0 |
*** Top Sister-Group Cladogram ***
+-- Reginaia_ebenus
+--------------------------------------------+
| +-- Reginaia_rotulata
|
| +-- Amblema_plicata
+--------------------------------------------+
| +-- Amblema_elliottii
|
| +----- Popenaias_popeii
| |
| | +-- Dromus_dromas
| | |
| | +-- Epioblasma_rangiana
| | |
+-----------------------------------------+ +-- Glebula_rotundata
| | |
| | +-- Lampsilis_ovata
| | |
| | +-- Lampsilis_virescens
| | |
| | +-- Lemiox_rimosus
| +--+
| +-- Obliquaria_reflexa
| |
| +-- Obovaria_olivaria
| |
--+ +-- Obovaria_unicolor
| |
| +-- Plectomerus_dombeyanus
| |
| +-- Potamilus_alatus
| |
| +-- Potamilus_capax
|
| +-- Uniomerus_declivis **
| +--+
| | +-- Uniomerus_tetralasmus **
| |
| | +-- Cyclonaias_tuberculata **
| +--------------------------------------+ |
| | | +-- Pustulosa_pustulosa[1] **
| | | |
| | | +-- Theliderma_metanevra **
| | +--+
| | +-- Pustulosa_pustulosa[2] **
| | |
| | +-- Quadrula_quadrula **
| | |
| | +-- Quadrula_verrucosa **
| |
| | +----------------------------------------- Elliptoideus_sloatianus
+--+ |
| | +-- Elliptio_lanceolata
| | +-----------------------------------+
| | | +-- Parvaspina_collina
| | |
| | | +----------------------------------- Plethobasus_cyphyus
| | | |
| | | +----------------------------------- Eurynia_dilatata
| | | |
| | | +----------------------------------- Hemistena_lata
| | | |
| | | | +-- Elliptio_complanata
+--+ | | |
| | | +-- Elliptio_hopetonensis
| | | |
| | | +-- Elliptio_producta[1]
| | | |
| | | +-- Elliptio_shepardiana
| | | |
| | | +-- Elliptio_angustata
| | | |
| | | +-- Elliptio_arca
| | | |
| | | +-- Elliptio_arctata
| | | |
+--+ | +-- Elliptio_jayensis
| | |
| | +-- Elliptio_crassidens
| | |
| +--------------------------------+-- Elliptio_dariensis
| | |
| | +-- Elliptio_fisheriana
| | |
| | +-- Elliptio_folliculata
| | |
| | +-- Elliptio_fumata
| | |
| | +-- Elliptio_icterina
| | |
| | +-- Elliptio_mcmichaeli
| | |
| | +-- Elliptio_producta[2]
| | |
| | +-- Elliptio_pullata
| | |
| | +-- Elliptio_purpurella
| | |
| | +-- Elliptio_congaraea
| |
+--+ +-- Pleuronaia_barnesiana
| |
| +-----------------------------+-- Pleuronaia_dolabelloides
| | |
| | +-- Pleuronaia_gibber
| |
| | +-------------------------- Pleurobema_plenum
| | |
| | | +----------------------- Pleurobema_stabile
| | | |
| | | | +----- Pleurobema_pyriforme
| | +--+ | +--+
| | | | | | | +-- Pleurobema_georgianum[2]
| | | | | | +--+
| | | | | +-----------+ +-- Pleurobema_athearni
| | | | | | |
| | | +--+ | | +----- Pleurobema_decisum
| | | | | | |
| | | | | +--+ +-- Pleurobema_georgianum[1]
| | | | | | |
| | | | | +--+-- Pleurobema_hanleyianum[1]
| | | | | |
| | | | | +-- Pleurobema_hanleyianum[2]
| | | | |
| | | +--+ +-------------- Pleurobema_beadleianum[2]
| | | | |
| | | | +--+ +----------- Pleurobema_strodeanum[2]
| | | | | | |
| | | | | +--+----------- Pleurobema_rubellum
| | | | | |
| | | | | | +-------- Pleurobema_beadleianum[1]
| | | | | +--+
| | | | | | +----- Pleurobema_taitianum
| | | | | +--+
| | | | | | +-- Pleurobema_perovatum
| | | +--+ +--+
| | | | +-- Pleurobema_strodeanum[1]
| | | |
| | | | +----- Pleurobema_oviforme[6]
| | | | |
| | | | +--+----- Pleurobema_clava[3]
| | | | | |
| | | | | | +-- Pleurobema_oviforme[5]
| | | | | +--+
| | | +--------+ +-- Pleurobema_clava[2]
+--+ | |
| | | +----- Pleurobema_oviforme[2]
| | | |
| | | +----- Pleurobema_oviforme[3]
| | +--+
| | +----- Pleurobema_oviforme[4]
| | |
| | | +-- Pleurobema_clava[1]
| | +--+
| | +-- Pleurobema_oviforme[1]
| |
| | +-------------------------- Pleurobema_cordatum
| | |
| | | +----- Fusconaia_subrotunda
| | | +-----------------+
| +--+ | | +-- Fusconaia_cor
| | | | +--+
| | | | +-- Fusconaia_cuneolus
| | | |
| | | | +-------------------- Fusconaia_ozarkensis
| | +--+ |
| | | | +-- Fusconaia_escambia
| | | | +--------------+
| | | | | +-- Fusconaia_burkei
| | | | |
| | | | | +----- Fusconaia_flava[27]
| | | | | |
| | +--+ | | +-- Fusconaia_iheringi
| | | | +--+
| | | | | +-- Fusconaia_mitchelli
| | | +-----------+
| | | | | +-- Fusconaia_askewi[1]
| | | | | |
| | | | | +-- Fusconaia_askewi[2]
| | | | | |
| | +--+ | +-- Fusconaia_askewi[3]
| | | +--+
| | | +-- Fusconaia_askewi[4]
| | | |
| | | +-- Fusconaia_askewi[5]
| | | |
| | | +-- Fusconaia_chunii
| | |
+--+ | +-------------- Fusconaia_masoni
| | |
| | | +----------- Fusconaia_flava[26]
| | | |
| | | | +-- Fusconaia_cerina[8]
| +--+ | +-----+
| | | | +-- Fusconaia_flava[25]
| | | |
| | | | +----- Fusconaia_flava[5]
| | | | |
| | | | +----- Fusconaia_flava[6]
| +--+ | |
| | | +----- Fusconaia_flava[7]
| | | |
| | | +----- Fusconaia_flava[8]
| | | |
| | | +----- Fusconaia_cerina[4]
| | | |
| | | +----- Fusconaia_flava[9]
| | | |
| +--+ +----- Fusconaia_flava[10]
| | |
| | +----- Fusconaia_flava[11]
| | |
| | +----- Fusconaia_flava[12]
| | |
| | +----- Fusconaia_flava[13]
| | |
| | +----- Fusconaia_flava[14]
| | |
| | +----- Fusconaia_flava[15]
| | |
| | +----- Fusconaia_flava[16]
| | |
| | +----- Fusconaia_flava[17]
| +--+
| +----- Fusconaia_flava[18]
| |
| +----- Fusconaia_flava[19]
| |
| +----- Fusconaia_flava[20]
| |
| +----- Fusconaia_flava[21]
| |
| +----- Fusconaia_flava[22]
| |
| +----- Fusconaia_flava[23]
| |
| +----- Fusconaia_flava[24]
| |
| +----- Fusconaia_cerina[5]
| |
| +----- Fusconaia_cerina[6]
| |
| +----- Fusconaia_cerina[7]
| |
| | +-- Fusconaia_flava[1]
| | |
| | +-- Fusconaia_flava[2]
| | |
| | +-- Fusconaia_flava[3]
| | |
| +--+-- Fusconaia_flava[4]
| |
| +-- Fusconaia_cerina[1]
| |
| +-- Fusconaia_cerina[2]
| |
| +-- Fusconaia_cerina[3]
|
| +----------------- Pleurobema_riddellii
| |
| | +-------------- Pleurobema_sintoxia[21]
| | |
| | | +----------- Pleurobema_sintoxia[19]
| | | |
| | | +----------- Pleurobema_sintoxia[20]
| | | |
+-----------+ | | +----- Pleurobema_sintoxia[18]
| | | |
| | | +----- Pleurobema_rubrum[19]
| | | |
| | | +----- Pleurobema_rubrum[20]
| | +-----+
| | | +----- Pleurobema_rubrum[21]
| | | |
| | | | +-- Pleurobema_rubrum[17]
+--+ | | |
| | | +-- Pleurobema_sintoxia[16]
| | +--+
| | +-- Pleurobema_sintoxia[17]
| | |
| | +-- Pleurobema_rubrum[18]
| |
| | +-------- Pleurobema[4]
| | |
| | +-------- Pleurobema[5]
| | |
| | +-------- Pleurobema_sintoxia[1]
| | |
| | +-------- Pleurobema[8]
| | |
| | +-------- Pleurobema[9]
+--+ |
| +-------- Pleurobema[10]
| |
| +-------- Pleurobema_sintoxia[3]
| |
| +-------- Pleurobema_sintoxia[4]
| |
| +-------- Pleurobema_sintoxia[5]
| |
| +-------- Pleurobema_sintoxia[6]
| |
| +-------- Pleurobema_sintoxia[7]
| |
| +-------- Pleurobema_sintoxia[8]
| |
| +-------- Pleurobema_sintoxia[9]
| |
| +-------- Pleurobema_sintoxia[10]
| |
| +-------- Pleurobema_sintoxia[11]
| |
| +-------- Pleurobema_sintoxia[12]
| |
| +-------- Pleurobema_sintoxia[13]
| |
| +-------- Pleurobema_sintoxia[14]
| |
| +-------- Pleurobema_sintoxia[15]
| |
+--+-------- Pleurobema_rubrum[4]
|
+-------- Pleurobema_rubrum[5]
|
+-------- Pleurobema_rubrum[6]
|
+-------- Pleurobema_rubrum[7]
|
+-------- Pleurobema_rubrum[8]
|
+-------- Pleurobema_rubrum[9]
|
+-------- Pleurobema_rubrum[10]
|
+-------- Pleurobema_rubrum[11]
|
+-------- Pleurobema_rubrum[12]
|
+-------- Pleurobema_rubrum[13]
|
+-------- Pleurobema_rubrum[14]
|
+-------- Pleurobema_rubrum[15]
|
+-------- Pleurobema_rubrum[16]
|
| +-- Pleurobema[6]
+-----+
| +-- Pleurobema_rubrum[1]
|
| +-- Pleurobema_sintoxia[2]
+-----+
| +-- Pleurobema_rubrum[2]
|
| +-- Pleurobema[7]
+-----+
| +-- Pleurobema_rubrum[3]
|
| +----- Pleurobema[3]
+--+
| +-- Pleurobema[1]
+--+
+-- Pleurobema[2]
Lopes-Lima et al., 2019 [Publ 3901], fig. 3
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 19 |
25 |
23 |
3 |
10 |
6 |
3 |
27.0 |
+-------------------------------- Megalonaias_nervosa **
|
| +-- Pleurobema_oviforme
| +-----+
| | +-- Pleurobema_riddellii
| |
--+ +--------------------+ +-- Lampsilis_ornata
| | | +--+
| | | | +-- Potamilus_leptodon
| | +--+
| | | +-- Amblema_elliottii
| | +--+
+--+ +-- Amblema_plicata
|
| +-------------------------- Uniomerus_tetralasmus **
| |
| | +----------- Theliderma_cylindrica **
| | +-----------+
| | | | +-------- Theliderma_sparsa **
+--+ | +--+
| | | +----- Theliderma_intermedia **
| | +--+
| | | +-- Theliderma_metanevra **
| | +--+
+--+ +-- Theliderma_johnsoni **
|
| +----- Quadrula_quadrula **
| +--------------+
| | | +-- Quadrula_nobilis **
| | +--+
| | +-- Quadrula_verrucosa **
+--+
| +----------------- Pustulosa_kieneriana **
| |
| | +-- Pustulosa_infucata **
+--+ +-----------+
| | +-- Pustulosa_kleiniana **
| |
+--+ +-- Pustulosa_pustulosa **
| +--------+
| | +-- Pustulosa_succissa **
+--+
| +-------- Cyclonaias_tuberculata **
+--+
| +----- Pustulosa_necki **
+--+
| +-- Pustulosa_petrina **
+--+
+-- Pustulosa_nodulata **
Lopes-Lima et al., 2019 [Publ 3901], fig. 2
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 18 |
18 |
15 |
3 |
10 |
6 |
3 |
23.0 |
+-- Lampsilis_ornata
+-----------------------------+
| +-- Potamilus_leptodon
|
| +-- Pleurobema_riddellii
--+ +--------------------------+
| | +-- Elliptio_crassidens
| |
| | +----- Megalonaias_nervosa **
+--+ |
| +--------------------+ +-- Uniomerus_carolinianus **
| | | |
| | +--+-- Uniomerus_declivis **
| | |
+--+ +-- Uniomerus_tetralasmus **
|
| +----------------------- Quadrula_quadrula **
| |
| | +-- Quadrula_verrucosa **
+--+ +-----------------+
| | +-- Quadrula_nobilis **
| |
+--+ +-- Theliderma_metanevra **
| +--------------+
| | +-- Theliderma_johnsoni **
| |
+--+ +----- Pustulosa_kieneriana **
| +--------+
| | | +-- Pustulosa_infucata **
| | +--+
+--+ +-- Pustulosa_kleiniana **
|
| +----------- Cyclonaias_tuberculata **
| |
+--+ +-- Pustulosa_pustulosa **
| +-----+
| | +-- Pustulosa_succissa **
+--+
| +----- Pustulosa_nodulata **
+--+
| +-- Pustulosa_necki **
+--+
+-- Pustulosa_petrina **
Lopes-Lima et al., 2019 [Publ 3901], fig. 4
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 16 |
16 |
14 |
3 |
10 |
6 |
3 |
22.0 |
+-- Amblema_elliottii
+-----+
| +-- Amblema_plicata
|
+-----------------------+ +-- Pleurobema_riddellii
| | +--+
| | | +-- Pleurobema_oviforme
| +--+
| | +-- Lampsilis_ornata
--+ +--+
| +-- Potamilus_leptodon
|
| +----------------------------- Megalonaias_nervosa **
| |
+--+ +-------------------------- Uniomerus_tetralasmus **
| |
| | +-- Theliderma_metanevra **
+--+ +--------------------+
| | +-- Theliderma_johnsoni **
| |
+--+ +-------------------- Quadrula_quadrula **
| |
| | +-- Quadrula_verrucosa **
+--+ +--------------+
| | +-- Quadrula_nobilis **
| |
| | +----- Pustulosa_kieneriana **
+--+ +--------+
| | | +-- Pustulosa_infucata **
| | +--+
| | +-- Pustulosa_kleiniana **
+--+
| +-- Pustulosa_pustulosa **
| +--------+
| | +-- Pustulosa_succissa **
+--+
| +-------- Cyclonaias_tuberculata **
+--+
| +----- Pustulosa_necki **
+--+
| +-- Pustulosa_petrina **
+--+
+-- Pustulosa_nodulata **
Szumowski et al., 2012 [Publ 3569], fig. 5
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 15 |
15 |
12 |
2 |
0 |
0 |
0 |
20.0 |
+----------------- Megalonaias_nervosa **
|
| +-------- Theliderma_cylindrica **
+--------+
| | +----- Theliderma_intermedia **
| +--+
| | +-- Theliderma_metanevra **
--+ +--+
| +-- Theliderma_sparsa **
|
| +-------- Quadrula_quadrula[1] **
| +-----+
| | | +----- Quadrula_verrucosa **
| | +--+
| | | +-- Quadrula_nobilis **
+--+ +--+
| +-- Quadrula_quadrula[2] **
|
| +----------- Pustulosa_kieneriana **
| |
+--+ +-------- Pustulosa_infucata **
| |
+--+ +-- Pustulosa_pustulosa **
| +--+
| | +-- Pustulosa_succissa **
+--+
| +-- Pustulosa_nodulata **
+--+
+-- Pustulosa_petrina **
Serb et al., 2003 [Publ 2659], fig. 2
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 14 |
14 |
11 |
2 |
15 |
15 |
3 |
17.0 |
+-- Lampsilis_siliquoidea
+--------------------+
| +-- Lampsilis_ornata
|
| +-------------------- Obliquaria_reflexa
| |
| +-------------------- Amblema_plicata
| |
| +-------------------- Reginaia_rotulata
| |
--+ +-------------------- Potamilus_alatus
| |
| | +-- Elliptoideus_sloatianus
| +-----------------+
| | +-- Plectomerus_dombeyanus
| |
| | +----- Medionidus_conradicus
| +--------------+
| | | +-- Dromus_dromas
+--+ +--+
| +-- Cyprogenia_aberti
|
| +-------- Hemistena_lata
+-----------+
| | +----- Elliptio_complanata
| +--+
| | +-- Fusconaia_burkei
| +--+
| +-- Fusconaia_flava
|
| +----------------- Megalonaias_nervosa **
| |
| | +-------- Theliderma_cylindrica **
| | +-----+
+--+ | | +----- Theliderma_intermedia **
| | +--+
| | | +-- Theliderma_sparsa **
| | +--+
| | +-- Theliderma_metanevra **
+--+
| +----- Quadrula_verrucosa **
| +-----+
| | | +-- Quadrula_quadrula **
| | +--+
| | +-- Quadrula_nobilis **
+--+
| +-------- Pustulosa_kieneriana **
| |
| +-------- Pustulosa_infucata **
+--+
| +-- Pustulosa_pustulosa **
| +--+
| | +-- Pustulosa_succissa **
+--+
| +-- Pustulosa_nodulata **
+--+
+-- Pustulosa_petrina **
Serb et al., 2003 [Publ 2659], fig. 3
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 14 |
14 |
12 |
2 |
15 |
15 |
3 |
22.0 |
+----- Reginaia_rotulata
+-----------------------+
| | +-- Elliptoideus_sloatianus
| +--+
| +-- Plectomerus_dombeyanus
|
--+ +-------- Hemistena_lata
| +-----------------+
| | | +----- Elliptio_complanata
| | +--+
| | | +-- Fusconaia_burkei
| | +--+
+--+ +-- Fusconaia_flava
|
| +----------------------- Megalonaias_nervosa **
| |
| | +-- Quadrula_verrucosa **
+--+ +-----------------+
| | +-- Quadrula_nobilis **
| |
+--+ +----------------- Quadrula_quadrula **
| |
| | +-------- Theliderma_cylindrica **
+--+ +-----+
| | | +----- Theliderma_sparsa **
| | +--+
| | | +-- Theliderma_intermedia **
+--+ +--+
| +-- Theliderma_metanevra **
|
| +----------- Pustulosa_kieneriana **
| |
+--+ +-------- Pustulosa_infucata **
| |
+--+ +-- Pustulosa_succissa **
| +--+
| | +-- Pustulosa_pustulosa **
+--+
| +-- Pustulosa_nodulata **
+--+
+-- Pustulosa_petrina **
Johnson et al., 2018 [Publ 3844], fig. 2
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 14 |
14 |
12 |
3 |
2 |
2 |
2 |
20.0 |
+-------------------------------- Amblema_plicata
|
--+ +----------------------------- Elliptio_crassidens
| |
+--+ +-------------------------- Megalonaias_nervosa **
| |
+--+ +----------------------- Uniomerus_tetralasmus **
| |
+--+ +-------------------- Theliderma_metanevra **
| |
| | +-- Quadrula_quadrula **
+--+ +--------------+
| | +-- Quadrula_verrucosa **
| |
| | +----- Pustulosa_kieneriana **
+--+ +--------+
| | | +-- Pustulosa_infucata **
| | +--+
| | +-- Pustulosa_kleiniana **
+--+
| +-- Pustulosa_pustulosa **
| +--------+
| | +-- Pustulosa_succissa **
+--+
| +-------- Cyclonaias_tuberculata **
+--+
| +----- Pustulosa_necki **
+--+
| +-- Pustulosa_nodulata **
+--+
+-- Pustulosa_petrina **
Campbell et al., 2005 [Publ 2660], fig. 1
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 11 |
11 |
8 |
3 |
106 |
94 |
4 |
20.0 |
+-- Megalonaias_nervosa **
+-----------+
| +-- Uniomerus_declivis **
|
+-----------------------------+ +----------- Theliderma_metanevra **
| | |
| | | +----- Quadrula_verrucosa **
| +--+ +--+
| | | | +-- Quadrula_quadrula **
| | | +--+
| | | +-- Quadrula_nobilis **
| +--+
| | +----- Pustulosa_kieneriana **
| | |
| | | +-- Cyclonaias_tuberculata **
| +--+--+
| | +-- Pustulosa_succissa **
| |
| | +-- Pustulosa_infucata **
| +--+
| +-- Pustulosa_kleiniana **
|
| +----------------------------------- Elliptoideus_sloatianus
| |
| | +-------------------------------- Hemistena_lata
| | |
| +-----+ | +----- Plethobasus_cyphyus
| | | | +-----------------------+
| | | | | | +-- Elliptio_arca
| | | | | +--+
| | | | | +-- Elliptio_crassidens
| | +--+ |
| | | | +-- Eurynia_dilatata
| | | | +-----------------------+
| | | | | +-- Parvaspina_collina
--+ | | | |
| | | | | +----- Pleuronaia_gibber
| | | | +--------------------+
| | | | | | +-- Pleuronaia_barnesiana
| | +--+ | +--+
| | | | +-- Pleuronaia_dolabelloides
| | | |
| | | | +----------- Fusconaia_subrotunda
| | | | |
| | | | | +-- Fusconaia_cor
| | | +--------------+ +-----+
| | | | | | +-- Fusconaia_cuneolus
| | | | | |
| | | | | | +-- Fusconaia_cerina[1]
| | | | +--+ |
| | | | | +--+-- Fusconaia_cerina[2]
| | +--+ | | |
| | | | | +-- Fusconaia_flava
| | | +--+
| | | | +-- Fusconaia_burkei[1]
| | | | |
| | | | +-- Fusconaia_burkei[2]
| | | +--+
| | | +-- Fusconaia_burkei[3]
| | | |
| | | +-- Fusconaia_escambia
| | |
| | | +-- Pleurobema_rubrum[2]
| | | +--+
| | | | +-- Pleurobema_cordatum
| | | +-----------------+
| | | | | +-- Pleurobema_rubrum[1]
+--+ | | +--+
| | | +-- Pleurobema_sintoxia
| +--+
| | +-------------------- Pleurobema_pyriforme
| | |
| | | +-- Pleurobema_georgianum[2]
| | | |
| +--+ +--------------+-- Pleurobema_georgianum[3]
| | | |
| | | +-- Pleurobema_athearni
| | |
| | | +----- Pleurobema_decisum
| +--+ |
| | +--------+ +-- Pleurobema_hanleyianum[1]
| | | | |
| | | +--+-- Pleurobema_hanleyianum[2]
| | | |
| | | +-- Pleurobema_georgianum[1]
| +--+
| | +-- Pleurobema_oviforme[1]
| | |
| | +--------+-- Pleurobema_oviforme[2]
| | | |
| +--+ +-- Pleurobema_clava
| |
| | +-------- Pleurobema_strodeanum
| +--+
| | +----- Pleurobema_rubellum
| +--+
| | +-- Pleurobema_perovatum
| +--+
| +-- Pleurobema_taitianum
|
| +----- Plectomerus_dombeyanus
| +--------------------------------+
| | | +-- Reginaia_ebenus
| | +--+
| | +-- Reginaia_rotulata
| |
| | +----- Popenaias_popeii
| | +-----------------------------+
| | | | +-- Amblema_elliottii
+--+ | +--+
| | +-- Amblema_plicata
| |
| | +-- Toxolasma_parvum
| | +-----+
| | | +-- Toxolasma_texasiense
| | +-----------------------+
| | | | +----- Obliquaria_reflexa
+--+ | +--+
| | | +-- Cyrtonaias_tampicoensis
| | +--+
| | +-- Glebula_rotundata
| |
| | +-- Potamilus_fragilis
| | +--+
| | | +-- Potamilus_leptodon
| | +-----------+
| | | | +-- Potamilus_alatus
| | | +--+
+--+ | +-- Potamilus_purpuratus
| |
| +-----------+ +-- Sagittunio_nasutus
| | | +--+
| | | | +-- Medionidus_conradicus
| | | +--------+
| | | | | +-- Ptychobranchus_fasciolaris
| | | | +--+
| | +--+ +-- Venustaconcha_ellipsiformis[2]
| | |
| | | +----------- Medionidus_acutissimus
| | +--+
| | | +-------- Lemiox_rimosus
| | +--+
+--+ | +----- Dromus_dromas
| +--+
| | +-- Cyprogenia_aberti
| +--+
| +-- Cyprogenia_stegaria
|
| +-- Ellipsaria_lineolata
| +-----------------------+
| | +-- Truncilla_truncata
| |
| | +----------------------- Villosa_villosa
| | |
| | | +-- Obovaria_olivaria
+--+ | +--------------+
| | | +-- Venustaconcha_ellipsiformis[1]
| | |
| | +--+ +----- Obovaria_unicolor
| | | | +--------+
| | | | | | +-- Obovaria_arkansasensis
+--+ | | | +--+
| | +--+ +-- Obovaria_subrotunda
| | |
| | | +----------- Epioblasma_brevidens
| | +--+
| | | +-------- Epioblasma_triquetra
| | +--+
| | | +----- Epioblasma_walkeri[2]
| | +--+
| | | +-- Epioblasma_capsaeformis
+--+ +--+
| +-- Epioblasma_walkeri[1]
|
| +-- Ligumia_recta
| +--------+
| | +-- Cambarunio_iris
| +--+
| | | +-------- Lampsilis_teres
| | +--+
| | | +----- Ortmanniana_ligamentina
| | +--+
| | | +-- Lampsilis_siliquoidea
| | +--+
| | +-- Leaunio_vanuxemensis
+-----+
| +-- Lampsilis_cardium
| +--+
| | +-- Lampsilis_ovata
| +-----+
| | | +-- Ortmanniana_pectorosa
| | +--+
| | +-- Lampsilis_ornata
| |
+--+ +----- Hamiota_australis[2]
| +--+
| | | +-- Hamiota_australis[1]
| | +--+
| | +-- Hamiota_subangulata
+--+
| +----- Hamiota_altilis[2]
| |
| | +-- Hamiota_altilis[1]
+--+--+
| +-- Hamiota_perovalis[1]
|
| +-- Hamiota_altilis[3]
+--+
+-- Hamiota_perovalis[2]
Campbell et al., 2005 [Publ 2660], fig. 2
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 11 |
11 |
8 |
3 |
104 |
92 |
4 |
20.0 |
+-------------------- Megalonaias_nervosa **
|
+-----------------+ +----------------- Uniomerus_declivis **
| | |
| +--+ +-------------- Theliderma_metanevra **
| | |
| +--+ +----------- Quadrula_quadrula **
| | |
| | | +-- Quadrula_nobilis **
| +--+ +-----+
| | | +-- Quadrula_verrucosa **
| | |
| +--+ +----- Pustulosa_kieneriana **
| | |
| | | +-- Cyclonaias_tuberculata **
| +--+--+
| | +-- Pustulosa_succissa **
| |
| | +-- Pustulosa_infucata **
| +--+
| +-- Pustulosa_kleiniana **
|
| +-------------------------------- Elliptoideus_sloatianus
| |
| | +----------------------------- Plethobasus_cyphyus
| | |
| | +----------------------------- Parvaspina_collina
| +--+ |
| | | | +-- Elliptio_arca
| | | +--------------------------+
| | | | +-- Elliptio_crassidens
| | | |
| | | | +-------------------------- Eurynia_dilatata
| | +--+ |
--+ | | +-------------------------- Hemistena_lata
| | | |
| | | | +-- Pleurobema_rubrum[1]
| | | | +--+
| | | | | +-- Pleurobema_sintoxia
| | | | +-----------------+
| | | | | | +-- Pleurobema_rubrum[2]
| | | | | +--+
| | +--+ | +-- Pleurobema_cordatum
| | | |
| | | | +-------------- Fusconaia_subrotunda
| | | | |
| | | | | +-- Fusconaia_cor
| | | +--------+ +--------+
| | | | | | +-- Fusconaia_cuneolus
| | | | | |
| | | | +--+ +-- Fusconaia_cerina
| | | | | +-----+
| | | | | | +-- Fusconaia_flava
| | +--+ +--+
| | | | +----- Fusconaia_burkei[2]
| | | +--+
| | | | +-- Fusconaia_escambia
| | | +--+
| | | +-- Fusconaia_burkei[1]
| | |
| | | +----- Pleuronaia_gibber
| | | +--------------+
| | | | | +-- Pleuronaia_barnesiana
| | | | +--+
| | | | +-- Pleuronaia_dolabelloides
| | | |
| | | | +-------- Pleurobema_rubellum
+--+ +--+ +--------+
| | | | +----- Pleurobema_strodeanum
| | | +--+
| | | | +-- Pleurobema_perovatum
| | | +--+
| | | +-- Pleurobema_taitianum
| +--+
| | +-- Pleurobema_oviforme[1]
| | |
| | +-----------+-- Pleurobema_oviforme[2]
| | | |
| | | +-- Pleurobema_clava
| +--+
| | +----------- Pleurobema_pyriforme
| | |
| +--+ +-- Pleurobema_athearni
| | +-----+
| | | +-- Pleurobema_georgianum[2]
| +--+
| | +----- Pleurobema_decisum
| | |
| +--+ +-- Pleurobema_hanleyianum[1]
| | |
| +--+-- Pleurobema_hanleyianum[2]
| |
| +-- Pleurobema_georgianum[1]
|
| +-------- Popenaias_popeii
| |
| +--------------+ +-- Amblema_elliottii
| | | +--+
| | | | +-- Amblema_plicata
| | +--+
| | | +-- Reginaia_ebenus
| | | |
| | +--+-- Reginaia_rotulata[1]
| | |
| | +-- Reginaia_rotulata[2]
| |
| | +----- Plectomerus_dombeyanus
| | +--------------+
+-----------+ | | +-- Toxolasma_parvum
| | +--+
| | +-- Toxolasma_texasiense
| |
| | +----- Obliquaria_reflexa
| | +-----------+
| | | | +-- Cyrtonaias_tampicoensis
| | | +--+
| | | +-- Glebula_rotundata
| | |
| | | +-------------- Sagittunio_nasutus
| | | |
+--+ | +-------------- Leaunio_vanuxemensis
| | |
| | | +-- Ellipsaria_lineolata
| | +-----------+
| | | +-- Truncilla_truncata
| | |
| | | +----- Lampsilis_teres
| | +--------+
| | | | +-- Ortmanniana_ligamentina
| | | +--+
| | | +-- Lampsilis_siliquoidea
| | |
| | | +----- Villosa_villosa
| | +--------+
+--+ | | +-- Ligumia_recta
| | +--+
| | +-- Cambarunio_iris
| |
| | +-------- Ortmanniana_pectorosa
| +-----+
| | | +----- Lampsilis_ornata
| | +--+
| | | +-- Lampsilis_cardium
| | +--+
| | +-- Lampsilis_ovata
| |
| | +-- Potamilus_fragilis
| | +--+
| | | +-- Potamilus_leptodon
| +--------+
| | | +-- Potamilus_alatus
| | +--+
| | +-- Potamilus_purpuratus
| |
| | +-- Ptychobranchus_fasciolaris
+--+ +--------+
| | +-- Venustaconcha_ellipsiformis[1]
| |
+--+ +-------- Medionidus_conradicus
| | |
| | | +-- Lemiox_rimosus
| +--+-----+
| | +-- Medionidus_acutissimus
| |
| | +----- Dromus_dromas
| +--+
| | +-- Cyprogenia_aberti
| +--+
| +-- Cyprogenia_stegaria
|
| +----- Hamiota_australis[2]
| +-----+
| | | +-- Hamiota_australis[1]
| | +--+
| | +-- Hamiota_subangulata
+--+
| | +-- Hamiota_altilis[3]
| | +-----+
| | | +-- Hamiota_perovalis[2]
| +--+
| | +----- Hamiota_altilis[2]
| +--+
| | +-- Hamiota_altilis[1]
| +--+
| +-- Hamiota_perovalis[1]
|
| +----------- Obovaria_olivaria
| |
| +----------- Venustaconcha_ellipsiformis[2]
| |
| | +----- Obovaria_unicolor
| +-----+
+--+ | +-- Obovaria_arkansasensis
| +--+
| +-- Obovaria_subrotunda
|
| +-- Epioblasma_brevidens
| +-----+
| | +-- Epioblasma_triquetra
+--+
| +----- Epioblasma_walkeri[1]
+--+
| +-- Epioblasma_walkeri[2]
+--+
+-- Epioblasma_capsaeformis
Campbell & Lydeard, 2012 [Publ 3309], fig. 1-2
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 11 |
11 |
9 |
2 |
63 |
63 |
4 |
21.0 |
+-- Uniomerus_declivis **
+-----------------+
| +-- Uniomerus_tetralasmus **
+--------------------+
| | +----------------- Theliderma_metanevra **
| | |
| +--+ +-------------- Quadrula_verrucosa **
| | |
| | | +-- Quadrula_quadrula **
| +--+ +--------+
| | | +-- Quadrula_nobilis **
| | |
| +--+ +-- Pustulosa_infucata **
| | +-----+
| | | +-- Pustulosa_kleiniana **
| +--+
| | +----- Cyclonaias_tuberculata **
| +--+
| | +-- Pustulosa_pustulosa **
| +--+
| +-- Pustulosa_succissa **
|
| +----- Plectomerus_dombeyanus
| +-----------------+
| | | +-- Reginaia_ebenus
| | +--+
--+ | +-- Reginaia_rotulata
| |
| | +----- Popenaias_popeii
| +--------------+ +--------------+
| | | | | +-- Amblema_elliottii
| | | | +--+
| | | | +-- Amblema_plicata
| | | |
| | | | +----------------- Potamilus_alatus
| | +--+ |
| | | +----------------- Potamilus_capax
| | | |
| | | +----------------- Glebula_rotundata
| | | |
| | | | +-- Toxolasma_lividum
| | | +--------------+
| | +--+ +-- Paetulunio_fabalis
| | |
| | | +-- Dromus_dromas
| | +--------------+
| | | +-- Lemiox_rimosus
| | |
| | | +-- Lampsilis_ovata
+--+ | +-----------+
| | | +-- Obliquaria_reflexa
| +--+
| | +----------- Lampsilis_virescens
| | |
| +--+ +-- Epioblasma_rangiana
| | +-----+
| | | +-- Obovaria_arkansasensis
| +--+
| | +----- Obovaria_olivaria
| +--+
| | +-- Venustaconcha_ellipsiformis
| +--+
| +-- Venustaconcha_pleasii
|
| +----------------------------------- Elliptoideus_sloatianus
| |
| | +-------------------------------- Pleurobema_stabile
| | |
| | | +-- Pleurobema_clava
| | | +-----------+
+--+ | | +-- Pleurobema_oviforme
| | |
| | +--------------+ +----------- Pleurobema_pyriforme
| | | | |
| | | | | +-- Pleurobema_athearni
| | | | | +--+
+--+ | +--+ | +-- Pleurobema_georgianum
| | | +--+
| | | | | +-- Pleurobema_decisum
| | | | +--+
| | | | +-- Pleurobema_hanleyianum
| | +--+
| | | +----- Pleurobema_beadleianum
| | | |
| | | +----- Pleurobema_strodeanum
| | +--+
| | +----- Pleurobema_rubellum
+--+ |
| | +-- Pleurobema_perovatum
| +--+
| +-- Pleurobema_taitianum
|
| +-------- Pleurobema_riddellii
| +-----------------+
| | | +----- Pleurobema_cordatum
| | +--+
| | | +-- Pleurobema_rubrum
| | +--+
| | +-- Pleurobema_sintoxia
| |
| | +----------------- Fusconaia_subrotunda
| | |
+--+ +-----+ +-- Fusconaia_cor
| | | +-----------+
| | | | +-- Fusconaia_cuneolus
| | +--+
| | | +----------- Fusconaia_ozarkensis
| | | |
| | +--+ +-- Fusconaia_escambia
| | | +-----+
| | | | +-- Fusconaia_burkei
| | +--+
+--+ | +----- Fusconaia_masoni
| +--+
| | +-- Fusconaia_flava
| +--+
| +-- Fusconaia_cerina
|
| +----- Pleuronaia_gibber
| +--------------+
| | | +-- Pleuronaia_barnesiana
| | +--+
| | +-- Pleuronaia_dolabelloides
+--+
| +-- Eurynia_dilatata
| +--------------+
| | +-- Parvaspina_collina
| |
+--+ +-- Hemistena_lata
| +-----------+
| | +-- Plethobasus_cyphyus
| |
+--+ +----------- Elliptio_shepardiana
| |
| | +-------- Elliptio_complanata
+--+ |
| | +----- Elliptio_purpurella
| +--+
| | | +-- Elliptio_arca
+--+ +--+
| +-- Elliptio_arctata
|
| +-- Elliptio_mcmichaeli
| |
+-----+-- Elliptio_pullata
|
+-- Elliptio_crassidens
Campbell & Lydeard, 2012 [Publ 3309], fig. 3
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 11 |
11 |
8 |
2 |
63 |
63 |
4 |
22.0 |
+-- Uniomerus_declivis **
+--------------+
| +-- Uniomerus_tetralasmus **
|
+--------------------+ +-- Quadrula_verrucosa **
| | +-----------+
| | | +-- Quadrula_nobilis **
| | |
| +--+ +----------- Quadrula_quadrula **
| | |
| | +----------- Theliderma_metanevra **
| +--+
| | +-- Pustulosa_infucata **
| | +-----+
| | | +-- Pustulosa_kleiniana **
| +--+
| | +----- Cyclonaias_tuberculata **
| +--+
| | +-- Pustulosa_pustulosa **
| +--+
| +-- Pustulosa_succissa **
|
| +-- Amblema_elliottii
| +--+
| | +-- Amblema_plicata
| +--------------------------+
| | | +-- Reginaia_ebenus
--+ | +--+
| | +-- Reginaia_rotulata
| +--+
| | | +----------------------------- Popenaias_popeii
| | | |
| | | | +-- Glebula_rotundata
| | +--+ +-----------------------+
| | | | +-- Paetulunio_fabalis
| | | |
| | +--+ +-- Toxolasma_lividum
| | | +--------------------+
| | | | +-- Plectomerus_dombeyanus
| | | |
| | +--+ +-------------------- Obliquaria_reflexa
| | | |
| | | | +-- Dromus_dromas
| | | | +--+
| | +--+ | +-- Lemiox_rimosus
| | | +-----------+
| | | | | +-- Potamilus_alatus
| | | | +--+
| | +--+ +-- Potamilus_capax
| | |
| | | +-------------- Lampsilis_ovata
+--+ | |
| +--+ +----------- Lampsilis_virescens
| | |
| +--+ +-- Epioblasma_rangiana
| | +-----+
| | | +-- Obovaria_arkansasensis
| +--+
| | +----- Obovaria_olivaria
| +--+
| | +-- Venustaconcha_ellipsiformis
| +--+
| +-- Venustaconcha_pleasii
|
| +-------------------------------- Elliptoideus_sloatianus
| |
| | +-- Eurynia_dilatata
| | +--------------------------+
| | | +-- Parvaspina_collina
| | |
| | | +-- Hemistena_lata
| | +--------------------------+
| | | +-- Plethobasus_cyphyus
+--+ |
| | +----------- Elliptio_shepardiana
| | |
| | | +----- Elliptio_crassidens
| +-----------------+ +--+
| | | | | +-- Elliptio_mcmichaeli
| | | | +--+
| | +--+ +-- Elliptio_pullata
| | |
| | | +----- Elliptio_complanata
+--+ | |
| +--+ +-- Elliptio_purpurella
| | |
| +--+-- Elliptio_arca
| |
| +-- Elliptio_arctata
|
| +----- Pleuronaia_gibber
| +--------------------+
| | | +-- Pleuronaia_barnesiana
| | +--+
| | +-- Pleuronaia_dolabelloides
| |
| | +-------------- Fusconaia_ozarkensis
| | |
| | | +----- Fusconaia_subrotunda
| | +--------+ +-----+
+--+ | | | | +-- Fusconaia_cor
| | | | +--+
| | | | +-- Fusconaia_cuneolus
| | +--+
| | | +-- Fusconaia_escambia
| | | +-----+
| | | | +-- Fusconaia_burkei
| | +--+
| | | +----- Fusconaia_masoni
+--+ +--+
| | +-- Fusconaia_cerina
| +--+
| +-- Fusconaia_flava
|
| +-------- Pleurobema_riddellii
| +-----------+
| | | +----- Pleurobema_cordatum
| | +--+
| | | +-- Pleurobema_rubrum
| | +--+
+--+ +-- Pleurobema_sintoxia
|
| +----------------- Pleurobema_stabile
| |
| | +----------- Pleurobema_strodeanum
| | +--+
+--+ | | +-------- Pleurobema_rubellum
| | +--+
| | | +----- Pleurobema_beadleianum
| | +--+
| | | +-- Pleurobema_perovatum
+--+ +--+
| +-- Pleurobema_taitianum
|
| +-- Pleurobema_clava
| +--------+
| | +-- Pleurobema_oviforme
+--+
| +-------- Pleurobema_pyriforme
| |
+--+ +-- Pleurobema_athearni
| +--+
| | +-- Pleurobema_georgianum
+--+
| +-- Pleurobema_decisum
+--+
+-- Pleurobema_hanleyianum
Zhou et al., 2007 [Publ 2658], fig. 2
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 7 |
7 |
5 |
1 |
14 |
0 |
0 |
23.0 |
+-- Theliderma_intermedia **
+-----------+
| +-- Theliderma_sparsa **
--+
| +----------- Quadrula_quadrula **
+--+
| +-------- Pustulosa_infucata **
+--+
| +----- Quadrula_verrucosa **
+--+
| +-- Pustulosa_kieneriana **
+--+
+-- Pustulosa_succissa **
Boyer et al., 2011 [Publ 2718], fig. 3
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 7 |
7 |
5 |
2 |
28 |
22 |
3 |
23.0 |
+-- Fusconaia_flava
+--+
| +-- Pleurobema_sintoxia
+--------------+
| | +-- Eurynia_dilatata
| +--+
+--+ +-- Elliptio_crassidens
| |
| | +----------------- Megalonaias_nervosa **
| +--+
| | +-------------- Quadrula_quadrula **
| +--+
| | +----------- Quadrula_verrucosa **
| +--+
| | +-------- Theliderma_metanevra **
--+ +--+
| | +----- Pustulosa_pustulosa **
| +--+
| | +-- Cyclonaias_tuberculata **
| +--+
| +-- Pustulosa_nodulata **
|
| +----------------- Amblema_plicata
| |
| | +-------------- Toxolasma_parvum
+-----+ |
| | +----------- Obliquaria_reflexa
| | |
+--+ | +-------- Obovaria_olivaria
| | |
| | +-------- Lampsilis_cardium
| | |
+--+ | +-- Epioblasma_triquetra
| +-----+
| | +-- Lampsilis_siliquoidea
| |
| | +-- Ortmanniana_ligamentina
| +-----+
| | +-- Lampsilis_higginsii
+--+
| +-- Ligumia_recta
+-----+
| +-- Lampsilis_teres
|
| +----- Potamilus_fragilis
+--+
| | +-- Potamilus_ohiensis
| +--+
| +-- Potamilus_alatus
|
| +----- Ellipsaria_lineolata
+--+
| +-- Truncilla_truncata
+--+
+-- Truncilla_donaciformis
Boyer et al., 2011 [Publ 2718], fig. 4
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 7 |
29 |
25 |
2 |
31 |
22 |
3 |
41.0 |
+----------- Theliderma_metanevra **
|
+--------------------+ +-- Quadrula_verrucosa **
| | +-----+
| | | +-- Quadrula_quadrula **
| +--+
| | +----- Pustulosa_pustulosa **
| +--+
--+ | +-- Cyclonaias_tuberculata **
| +--+
| +-- Pustulosa_nodulata **
|
| +----------------------------- Megalonaias_nervosa **
| |
+--+ +-------------------------- Toxolasma_parvum
| |
| | +----------------------- Amblema_plicata
+--+ |
| | +-- Reginaia_ebenus
| | +-----------------+
| | | +-- Obliquaria_reflexa
+--+ |
| | +-------- Pleurobema_sintoxia
| +-----------+
| | | +----- Elliptio_crassidens
| | +--+
| | | +-- Fusconaia_flava
+--+ +--+
| +-- Eurynia_dilatata
|
| +----- Potamilus_fragilis
| +-----------+
| | | +-- Potamilus_ohiensis
| | +--+
| | +-- Potamilus_alatus
| |
+--+ +----- Ellipsaria_lineolata
| +--------+
| | | +-- Truncilla_donaciformis
| | +--+
| | +-- Truncilla_truncata
+--+
| +----------- Lampsilis_cardium
| |
| | +----- Ligumia_recta
| +-----+
+--+ | +-- Obovaria_olivaria
| +--+
| +-- Epioblasma_triquetra
|
| +-------- Lampsilis_teres
+--+
| +----- Lampsilis_siliquoidea
+--+
| +-- Ortmanniana_ligamentina
+--+
+-- Lampsilis_higginsii
Campbell & Lydeard, 2012 [Publ 3308], fig. 3
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 7 |
7 |
5 |
2 |
44 |
44 |
4 |
21.0 |
+-------------- Uniomerus_declivis **
|
+--------------------------------+ +-- Pustulosa_infucata **
| | +--------+
| | | +-- Pustulosa_kleiniana **
| +--+
| | +-------- Quadrula_quadrula **
| +--+
| | +----- Cyclonaias_tuberculata **
| +--+
| | +-- Pustulosa_succissa **
| +--+
| +-- Pustulosa_pustulosa **
--+
| +-------------- Elliptoideus_sloatianus
| +-----------------------------+
| | | +----------- Plectomerus_dombeyanus
| | +--+
| | | +-------- Obovaria_unicolor
| | +--+
| | | +----- Obovaria_olivaria
| | +--+
| | | +-- Lampsilis_ovata
| | +--+
| | +-- Lemiox_rimosus
+--+
| +----------- Cyrtonaias_tampicoensis
| |
| | +----- Popenaias_popeii
| +-----------------------------+ +--+
| | | | | +-- Amblema_elliottii
| | | | +--+
| | +--+ +-- Amblema_plicata
| | |
| | | +-- Reginaia_ebenus[1]
| | | |
+--+ +-----+-- Reginaia_ebenus[2]
| |
| +-- Reginaia_rotulata
|
| +-- Elliptio_arca
| +-----------------------------------+
| | +-- Elliptio_crassidens
| |
+--+ +----------------------------------- Eurynia_dilatata
| |
| | +-------------------------------- Parvaspina_collina
| | |
+--+ | +-- Hemistena_lata
| +-----------------------------+
| | +-- Plethobasus_cyphyus
| |
+--+ +----- Pleuronaia_barnesiana
| +-----------------------+
| | | +-- Pleuronaia_dolabelloides
| | +--+
| | +-- Pleuronaia_gibber
| |
| | +-- Pleurobema_stabile
+--+ +-----+
| | +-- Pleurobema_sintoxia
| +-----------------+
| | | +----- Pleurobema_decisum
| | +--+
| | | +-- Pleurobema_clava
+--+ +--+
| +-- Pleurobema_pyriforme
|
| +----------------------- Pleurobema_cordatum
| |
+--+ +-------------------- Fusconaia_subrotunda
| |
| | +-- Fusconaia_cor
+--+ +--------------+
| | +-- Fusconaia_cuneolus
| |
+--+ +-------------- Fusconaia_ozarkensis
| |
| | +-- Fusconaia_flava[6]
| | +--------+
+--+ | +-- Fusconaia_masoni
| |
| | +-- Fusconaia_escambia
| | |
+--+ +-- Fusconaia_burkei[1]
| +-----+
| | +-- Fusconaia_burkei[2]
| | |
| | +-- Fusconaia_burkei[3]
+--+
| +----- Fusconaia_flava[5]
| |
| | +-- Fusconaia_cerina
+--+ |
| +-- Fusconaia_flava[1]
| |
+--+-- Fusconaia_flava[2]
|
+-- Fusconaia_flava[3]
|
+-- Fusconaia_flava[4]
Campbell & Lydeard, 2012 [Publ 3308], fig. 1
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 6 |
6 |
4 |
1 |
37 |
37 |
4 |
22.0 |
+----------- Quadrula_quadrula **
|
+-----------------------------------+ +-- Pustulosa_infucata **
| | +-----+
| | | +-- Pustulosa_kleiniana **
| +--+
| | +----- Cyclonaias_tuberculata **
| +--+
| | +-- Pustulosa_succissa **
| +--+
| +-- Pustulosa_pustulosa **
|
| +----- Popenaias_popeii
| +-----------+
--+ | | +-- Amblema_elliottii
| | +--+
| | +-- Amblema_plicata
| +--------------------------+
| | | +----- Cyrtonaias_tampicoensis
| | | +--------+
| | | | | +-- Reginaia_ebenus
| | | | +--+
| | +--+ +-- Reginaia_rotulata
| | |
| | | +----------- Plectomerus_dombeyanus
| | +--+
| | | +-------- Lemiox_rimosus
+--+ +--+
| | +----- Lampsilis_ovata
| +--+
| | +-- Obovaria_unicolor
| +--+
| +-- Obovaria_olivaria
|
| +----------------------------------------- Elliptoideus_sloatianus
| |
| | +-------------------------------------- Pleurobema_stabile
+--+ |
| | +----- Pleurobema_pyriforme
| | +-----------------------------+
+--+ | | +-- Pleurobema_clava
| | +--+
| | +-- Pleurobema_decisum
| |
+--+ +-- Pleurobema_cordatum
| +-----------------------------+
| | +-- Pleurobema_sintoxia
| |
| | +-- Eurynia_dilatata
| | +-----+
+--+ | +-- Hemistena_lata
| +--------------------+
| | | +----- Plethobasus_cyphyus
| | +--+
| | | +-- Elliptio_arca
| | +--+
+--+ +-- Elliptio_crassidens
|
| +----- Pleuronaia_gibber
| +--------------------+
| | | +-- Pleuronaia_barnesiana
| | +--+
+--+ +-- Pleuronaia_dolabelloides
|
| +----------------------- Parvaspina_collina
| |
+--+ +-------------------- Fusconaia_subrotunda
| |
+--+ +-- Fusconaia_cor
| +--------------+
| | +-- Fusconaia_cuneolus
+--+
| +-------------- Fusconaia_ozarkensis
| |
+--+ +-- Fusconaia_burkei
| +--------+
| | +-- Fusconaia_escambia
+--+
| +-------- Fusconaia_masoni
+--+
| +----- Fusconaia_cerina[2]
+--+
| +-- Fusconaia_flava
+--+
+-- Fusconaia_cerina[1]
Campbell & Lydeard, 2012 [Publ 3308], fig. 2
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 6 |
6 |
4 |
1 |
37 |
37 |
4 |
22.0 |
+----------- Quadrula_quadrula **
|
+--------------------+ +-- Pustulosa_infucata **
| | +-----+
| | | +-- Pustulosa_kleiniana **
| +--+
| | +----- Cyclonaias_tuberculata **
| +--+
| | +-- Pustulosa_succissa **
| +--+
| +-- Pustulosa_pustulosa **
|
| +-- Reginaia_ebenus
| +--+
| | +-- Reginaia_rotulata
--+ +-----------+
| | | +-- Amblema_elliottii
| | +--+
| +-----------+ +-- Amblema_plicata
| | |
| | | +-------------- Popenaias_popeii
| | | |
| | +--+ +----------- Cyrtonaias_tampicoensis
| | | |
| | +--+----------- Plectomerus_dombeyanus
| | |
| | | +-------- Lemiox_rimosus
| | +--+
| | | +----- Lampsilis_ovata
+--+ +--+
| | +-- Obovaria_unicolor
| +--+
| +-- Obovaria_olivaria
|
| +-------------------------- Elliptoideus_sloatianus
| |
| | +----------------------- Hemistena_lata
| | |
| | +----------------------- Plethobasus_cyphyus
| | |
+--+ | +-- Elliptio_arca
| +--------------------+
| | +-- Elliptio_crassidens
| |
| | +-- Eurynia_dilatata
| +--------------------+
| | +-- Parvaspina_collina
+--+
| +----- Pleuronaia_gibber
| +--------------+
| | | +-- Pleuronaia_barnesiana
| | +--+
| | +-- Pleuronaia_dolabelloides
| |
| | +-- Pleurobema_cordatum
| | +--------+
| | | +-- Pleurobema_sintoxia
| +--------+
| | | +-------- Pleurobema_stabile
+--+ +--+
| | +----- Pleurobema_pyriforme
| +--+
| | +-- Pleurobema_clava
| +--+
| +-- Pleurobema_decisum
|
| +----- Fusconaia_subrotunda
| +-----------+
| | | +-- Fusconaia_cor
| | +--+
+--+ +-- Fusconaia_cuneolus
|
| +-------------- Fusconaia_ozarkensis
| |
+--+ +-- Fusconaia_burkei
| +--------+
| | +-- Fusconaia_escambia
+--+
| +-------- Fusconaia_masoni
+--+
| +----- Fusconaia_cerina[2]
+--+
| +-- Fusconaia_flava
+--+
+-- Fusconaia_cerina[1]
Campbell & Lydeard, 2012 [Publ 3308], fig. 4
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 6 |
6 |
4 |
1 |
37 |
37 |
4 |
22.0 |
+----------- Quadrula_quadrula **
|
+--------------------------------------+ +-- Pustulosa_infucata **
| | +-----+
| | | +-- Pustulosa_kleiniana **
| +--+
| | +----- Cyclonaias_tuberculata **
| +--+
| | +-- Pustulosa_succissa **
| +--+
| +-- Pustulosa_pustulosa **
|
| +----- Popenaias_popeii
| +--------+
--+ | | +-- Amblema_elliottii
| | +--+
| | +-- Amblema_plicata
| +--------------------------------+
| | | +----------- Plectomerus_dombeyanus
| | | |
| | | | +----- Cyrtonaias_tampicoensis
| | | +-----+
| | +--+ | +-- Reginaia_ebenus
| | | +--+
| | | +-- Reginaia_rotulata
| | |
+--+ | +-------- Lemiox_rimosus
| +--+
| | +----- Lampsilis_ovata
| +--+
| | +-- Obovaria_unicolor
| +--+
| +-- Obovaria_olivaria
|
| +-------------------------------------------- Elliptoideus_sloatianus
| |
+--+ +----------------------------------------- Pleurobema_stabile
| |
+--+ +-------------------------------------- Hemistena_lata
| |
+--+ +----------------------------------- Eurynia_dilatata
| |
| | +-- Pleurobema_cordatum
+--+ +-----------------------------+
| | +-- Pleurobema_sintoxia
| |
| | +----- Pleurobema_clava
+--+ +-----------------------+
| | | +-- Pleurobema_decisum
| | +--+
| | +-- Pleurobema_pyriforme
| |
+--+ +-------- Parvaspina_collina
| +-----------------+
| | | +----- Pleuronaia_gibber
| | +--+
| | | +-- Pleuronaia_barnesiana
| | +--+
+--+ +-- Pleuronaia_dolabelloides
|
| +----- Plethobasus_cyphyus
| +-----------------+
| | | +-- Elliptio_arca
| | +--+
+--+ +-- Elliptio_crassidens
|
| +-------------------- Fusconaia_subrotunda
| |
+--+ +-- Fusconaia_cor
| +--------------+
| | +-- Fusconaia_cuneolus
+--+
| +-------------- Fusconaia_ozarkensis
| |
+--+ +-- Fusconaia_burkei
| +--------+
| | +-- Fusconaia_escambia
+--+
| +-------- Fusconaia_masoni
+--+
| +----- Fusconaia_cerina[2]
+--+
| +-- Fusconaia_flava
+--+
+-- Fusconaia_cerina[1]
Pfeiffer et al., 2016 [Publ 3660], fig. 2
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 6 |
6 |
4 |
2 |
33 |
32 |
3 |
22.0 |
+----------------------------- Elliptoideus_sloatianus
|
| +-------------- Uniomerus_declivis **
| +-----------+
| | | +----------- Quadrula_quadrula **
| | +--+
--+ | | +-------- Quadrula_verrucosa **
| | +--+
| | | +----- Pustulosa_pustulosa **
| | +--+
| | | +-- Pustulosa_petrina **
| | +--+
+--+ +-- Cyclonaias_tuberculata **
|
| +-- Elliptio_fumata
| +--------------------+
| | +-- Elliptio_crassidens
| |
| | +-------------------- Parvaspina_collina
| | |
| | +-------------------- Plethobasus_cyphyus
+--+ |
| | +-- Hemistena_lata
| +-----------------+
| | +-- Eurynia_dilatata
| |
| | +----- Pleurobema_riddellii
+--+ +--+
| | | +-- Pleurobema_sintoxia
| | +--+
| +--------+ +-- Pleurobema_cordatum
| | |
| | | +----- Pleuronaia_dolabelloides
| | | |
| | +--+ +-- Pleurobema_pyriforme
| | | |
+--+ +--+-- Pleurobema_strodeanum
| |
| +-- Pleurobema_clava
|
| +-------------- Fusconaia_askewi
| |
| | +-- Fusconaia_mitchelli
+--+ +--------+
| | +-- Fusconaia_iheringi
| |
+--+ +-- Fusconaia_escambia
| +-----+
| | +-- Fusconaia_burkei
+--+
| +----- Fusconaia_flava[2]
+--+
| +-- Fusconaia_cerina
+--+
+-- Fusconaia_flava[1]
Krebs et al., 2003 [Publ 2691], fig. 3
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 5 |
22 |
20 |
3 |
29 |
17 |
3 |
38.0 |
+-------- Megalonaias_nervosa **
+-----------------+
| | +----- Quadrula_quadrula[2] **
| +--+
| | +-- Quadrula_quadrula[1] **
| +--+
| +-- Theliderma_metanevra **
|
--+ +----------- Elliptoideus_sloatianus
| +-----------+
| | | +-------- Pleurobema_pyriforme
| | +--+
| | | +----- Uniomerus_carolinianus **
| | +--+
| | | +-- Fusconaia_flava
+--+ +--+
| +-- Eurynia_dilatata
|
| +-------------------- Amblema_plicata
| |
| | +----- Toxolasma_parvum
| | +-----------+
+--+ | | +-- Plectomerus_dombeyanus
| | +--+
| | +-- Obliquaria_reflexa
| |
+--+ +----- Ptychobranchus_subtentus
| +--------+
| | | +-- Potamilus_fragilis
| | +--+
| | +-- Medionidus_conradicus
+--+
| +-- Villosa_delumbis
| +--------+
| | +-- Ellipsaria_lineolata
+--+
| +-------- Lampsilis_teres
+--+
| +----- Lampsilis_siliquoidea
+--+
| +-- Potamilus_purpuratus
+--+
+-- Potamilus_alatus
Pfeiffer et al., 2019 [Publ 3866], fig. 1
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 5 |
5 |
3 |
3 |
34 |
33 |
4 |
19.0 |
+----------- Megalonaias_nervosa **
+-----------------------+
| | +-------- Uniomerus_declivis **
| +--+
| | +----- Quadrula_verrucosa **
| +--+
| | +-- Cyclonaias_tuberculata **
| +--+
| +-- Quadrula_quadrula **
|
| +-------------- Elliptoideus_sloatianus
--+ +-----------------+
| | | +----------- Elliptio_fumata
| | +--+
| | | +-------- Pleuronaia_barnesiana
| | +--+
| | | +----- Pleurobema_riddellii
| | +--+
| | | +-- Fusconaia_flava
| | +--+
| | +-- Pleurobema_strodeanum
| |
+--+ +-- Amblema_plicata
| +--+
| | +-- Amblema_neislerii
| +-----------------------+
| | | +-- Reginaia_ebenus
| | +--+
| | +-- Reginaia_rotulata
| |
| | +----------------- Popenaias_popeii
| | |
| | +--------+ +-------------- Psoronaias_guatemalensis[2]
| | | | |
+--+ | +--+ +----------- Psoronaias_guatemalensis[1]
| | | |
| | +--+ +-------- Psoronaias_semigranosa[3]
| | | |
| | | | +----- Psoronaias_semigranosa[2]
| | +--+ |
| | | +----- Psoronaias_usumasintae[2]
| | | |
| | +--+----- Psoronaias_percompressa
+--+ |
| | +-- Psoronaias_semigranosa[1]
| | |
| +--+-- Psoronaias_profunda
| |
| +-- Psoronaias_usumasintae[1]
|
| +-- Cyrtonaias_tampicoensis
| +--------------------+
| | +-- Glebula_rotundata
| |
+--+ +-------------------- Potamilus_alatus
| |
| | +-- Truncilla_truncata
+--+ +--------------+
| | +-- Ptychobranchus_fasciolaris
| |
+--+ +-------------- Pachynaias_spheniopsis
| |
| | +-- Cyrtonaias_explicata
+--+ +--------+
| | +-- Sphenonaias_microdon
| |
+--+ +-- Lampsilis_cardium
| +-----+
| | +-- Ortmanniana_ligamentina
+--+
| +----- Cambarunio_iris
+--+
| +-- Epioblasma_triquetra
+--+
+-- Ligumia_recta
Lydeard et al., 1996 [Publ 2656], fig. 3
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 4 |
18 |
14 |
3 |
23 |
14 |
3 |
34.0 |
+----- Megalonaias_nervosa **
+--------------------+
| | +-- Quadrula_quadrula **
| +--+
| +-- Theliderma_metanevra **
|
--+ +----- Elliptoideus_sloatianus
| |
| +-----------------+ +-- Eurynia_dilatata
| | | |
| | +--+-- Pleurobema_pyriforme
| | |
+--+ +-- Uniomerus_carolinianus **
|
| +-------------------- Amblema_plicata
| |
+--+ +-- Plectomerus_dombeyanus
| +--------------+
| | +-- Obliquaria_reflexa
+--+
| +-------------- Potamilus_fragilis
| |
+--+ +----------- Ptychobranchus_subtentus
| |
+--+ +-------- Medionidus_conradicus
| |
+--+ +----- Potamilus_purpuratus
| |
| | +-- Ellipsaria_lineolata
+--+--+
| +-- Villosa_delumbis
|
| +-- Lampsilis_teres
+--+
+-- Lampsilis_siliquoidea
Lydeard et al., 1996 [Publ 2656], fig. 4
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 4 |
18 |
16 |
3 |
24 |
14 |
3 |
34.0 |
+----- Megalonaias_nervosa **
+-----------------------------+
| | +-- Quadrula_quadrula **
| +--+
| +-- Theliderma_metanevra **
|
--+ +-------- Elliptoideus_sloatianus
| +-----------------------+
| | | +----- Uniomerus_carolinianus **
| | +--+
| | | +-- Eurynia_dilatata
+--+ +--+
| +-- Pleurobema_pyriforme
|
| +----------------------------- Amblema_plicata
+--+
| +-------------------------- Plectomerus_dombeyanus
+--+
| +----------------------- Obliquaria_reflexa
+--+
| +-------------------- Medionidus_conradicus
+--+
| +----------------- Ptychobranchus_subtentus
+--+
| +-------------- Potamilus_fragilis
+--+
| +----------- Potamilus_purpuratus
+--+
| +-------- Ellipsaria_lineolata
+--+
| +----- Villosa_delumbis
+--+
| +-- Lampsilis_teres
+--+
+-- Lampsilis_siliquoidea
Rosenberg et al., 1994 [Publ 2719], fig. 3
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 4 |
10 |
0 |
3 |
15 |
7 |
3 |
26.0 |
+-- Pyganodon_cataracta
|
+-- Pyganodon_grandis
|
+-- Utterbackia_imbecillis
|
+-- Lampsilis_teres
|
+-- Lampsilis_straminea
--+
+-- Megalonaias_nervosa **
|
+-- Obliquaria_reflexa
|
+-- Theliderma_cylindrica **
|
+-- Quadrula_quadrula **
|
+-- Uniomerus_tetralasmus **
Rosenberg et al., 1997 [Publ 2720], fig. 2A
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 4 |
17 |
0 |
3 |
19 |
8 |
3 |
33.0 |
+-- Gonidea_angulata
|
+-- Pyganodon_grandis
|
+-- Utterbackia_imbecillis
|
+-- Pyganodon_cataracta
|
+-- Lampsilis_straminea
|
+-- Lampsilis_teres
|
+-- Megalonaias_nervosa **
|
+-- Obliquaria_reflexa
|
--+-- Theliderma_cylindrica **
|
+-- Quadrula_quadrula **
|
+-- Uniomerus_tetralasmus **
|
+-- Amblema_plicata
|
+-- Elliptio_complanata
|
+-- Fusconaia_cerina
|
+-- Plectomerus_dombeyanus
|
+-- Pleurobema_cordatum
|
+-- Unio_pictorum
Rosenberg et al., 1997 [Publ 2720], fig. 2B
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 4 |
17 |
0 |
3 |
19 |
7 |
3 |
33.0 |
+-- Gonidea_angulata
|
+-- Pyganodon_grandis
|
+-- Utterbackia_imbecillis
|
+-- Pyganodon_cataracta
|
+-- Lampsilis_straminea
|
+-- Lampsilis_teres
|
+-- Megalonaias_nervosa **
|
+-- Obliquaria_reflexa
|
--+-- Theliderma_cylindrica **
|
+-- Quadrula_quadrula **
|
+-- Uniomerus_tetralasmus **
|
+-- Amblema_plicata
|
+-- Elliptio_complanata
|
+-- Fusconaia_cerina
|
+-- Plectomerus_dombeyanus
|
+-- Aculamprotula_tortuosa
|
+-- Unio_pictorum
Breton et al., 2011 [Publ 3376], fig. 3
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 4 |
34 |
31 |
2 |
35 |
15 |
3 |
49.0 |
+----- Quadrula_quadrula **
+--------------------------+
| | +-- Cyclonaias_tuberculata **
| +--+
| +-- Pustulosa_pustulosa **
|
| +-------------------------- Pronodularia_japanensis
| |
| +--+ +----------------------- Sinohyriopsis_cumingii
| | | |
| | +--+ +-------------------- Cristaria_plicata
| | | |
--+ | | | +-- Lasmigona_complanata
| | +--+ +--------------+
| | | | +-- Lasmigona_costata
| | | |
| | +--+ +-------------- Strophitus_undulatus
| | | |
| | | | +-- Platynaias_compressa
| | | | +--+
| | +--+ | +-- Platynaias_subviridis
| | | +-----+
| | | | | +-- Alasmidonta_marginata
+--+ | | +--+
| | | +-- Alasmidonta_undulata
| +--+
| | +-- Pyganodon_lacustris
| | +-----+
| | | +-- Pyganodon_grandis
| +--+
| | +----- Utterbackia_peninsularis[2]
| +--+
| | +-- Utterbackia_peninsularis[1]
| +--+
| +-- Utterbackia_imbecillis
|
| +-------------------------- Megalonaias_nervosa **
| |
+--+ +----------------------- Fusconaia_flava
| |
+--+ +-------------------- Reginaia_ebenus
| |
| | +-------- Toxolasma_paulum[2]
+--+ +--------+
| | | +----- Toxolasma_lividum
| | +--+
| | | +-- Toxolasma_parvum
+--+ +--+
| +-- Toxolasma_paulum[1]
|
| +-------------- Ellipsaria_lineolata
| |
+--+ +----------- Truncilla_macrodon
| |
| | +-- Lemiox_rimosus
+--+ +-----+
| | +-- Potamilus_metnecktayi
| |
+--+ +-- Cambarunio_iris
| +--+
| | +-- Venustaconcha_ellipsiformis
+--+
| +-- Lampsilis_powellii
| |
+--+-- Lampsilis_ornata
|
+-- Lampsilis_siliquoidea
Pfeiffer et al., 2018 [Publ 3835], fig. 2
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 4 |
4 |
2 |
3 |
0 |
0 |
0 |
22.0 |
+-------- Quadrula_quadrula **
--+
| +----- Uniomerus_declivis **
+--+
| +-- Megalonaias_nervosa **
+--+
+-- Megalonaias_nickliniana **
Huang et al., 2019 [Publ 3843], fig. 2
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 4 |
4 |
2 |
3 |
200 |
21 |
3 |
20.0 |
+-- Megalonaias_nervosa **
+--+
| +-- Uniomerus_carolinianus **
+-----------------------+
| | +-- Quadrula_quadrula **
| +--+
| +-- Quadrula_verrucosa **
|
| +-------- Elliptoideus_sloatianus
--+ |
| +-----------------+ +-- Elliptio_complanata
| | | +--+
| | | | +-- Eurynia_dilatata
| | +--+
| | | +-- Fusconaia_flava
+--+ +--+
| +-- Pleurobema_sintoxia
|
| +----------------------- Amblema_plicata
| |
| | +----- Toxolasma_parvum
+--+ +--------------+
| | | +-- Obliquaria_reflexa
| | +--+
| | +-- Plectomerus_dombeyanus
+--+
| +----------------- Potamilus_leptodon
| |
| | +-------------- Potamilus_alatus
+--+ |
| | +----------- Venustaconcha_ellipsiformis
| | |
+--+ +----------- Leaunio_lienosus
| |
| | +-- Ellipsaria_lineolata
| +--------+
+--+ +-- Truncilla_truncata
|
| +-- Medionidus_conradicus
+--------+
| +-- Ptychobranchus_subtentus
|
| +-------- Cambarunio_iris
+--+
| +----- Ortmanniana_ligamentina
+--+
| +-- Lampsilis_cardium
+--+
+-- Lampsilis_ornata
Huang et al., 2019 [Publ 3843], fig. S3
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 4 |
4 |
2 |
3 |
197 |
21 |
3 |
20.0 |
+-- Megalonaias_nervosa **
+--+
| +-- Uniomerus_carolinianus **
+--------------------------------+
| | +-- Quadrula_quadrula **
| +--+
| +-- Quadrula_verrucosa **
|
| +-------- Elliptoideus_sloatianus
--+ |
| +--------------------------+ +-- Elliptio_complanata
| | | +--+
| | | | +-- Eurynia_dilatata
| | +--+
| | | +-- Fusconaia_flava
+--+ +--+
| +-- Pleurobema_sintoxia
|
| +-------------------------------- Amblema_plicata
| |
+--+ +----------------------------- Toxolasma_parvum
| |
+--+ +-- Obliquaria_reflexa
| +-----------------------+
| | +-- Plectomerus_dombeyanus
+--+
| +----------------------- Potamilus_leptodon
| |
+--+ +-------------------- Potamilus_alatus
| |
| | +-- Medionidus_conradicus
+--+ +--------------+
| | +-- Ptychobranchus_subtentus
| |
+--+ +-- Ellipsaria_lineolata
| +-----------+
| | +-- Truncilla_truncata
+--+
| +----------- Leaunio_lienosus
| |
+--+ +-- Venustaconcha_ellipsiformis
| +-----+
| | +-- Cambarunio_iris
+--+
| +----- Ortmanniana_ligamentina
+--+
| +-- Lampsilis_cardium
+--+
+-- Lampsilis_ornata
Huang et al., 2019 [Publ 3843], fig. S4
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 4 |
127 |
70 |
3 |
198 |
21 |
3 |
142.0 |
+-------------------- Chamberlainia_somsakpanhai
|
+-------------------- Megalonaias_nervosa **
|
+-------------------- Uniomerus_carolinianus **
|
| +-- Quadrula_quadrula **
+-----------------+
| +-- Quadrula_verrucosa **
|
| +-- Sinohyriopsis_cumingii
+-----------------+
| +-- Sinohyriopsis_schlegelii
|
| +-- Potomida_littoralis
| +-----+
| | +-- Pronodularia_japanensis
+-----------+
| | +----- Lamprotula_leaii
| +--+
| | +-- Lamprotula_caveata
| +--+
| +-- Lamprotula_cornuumlunae
|
| +-------- Sinosolenaia_carinata
| |
| | +----- Gonidea_angulata
| +--+
| | | +-- Leguminaia_wheatleyi
| | +--+
| | +-- Microcondylaea_bonellii
+-----------+
| | +----- Ptychorhynchus_pfisteri
| +--+
| | | +-- Parvasolenaia_rivularis
| | +--+
| | +-- Koreosolenaia_sitgyensis
| |
| | +----- Sinosolenaia_recognita
| +--+
| | +-- Sinosolenaia_iridinea
| +--+
| +-- Sinosolenaia_oleivora
|
| +----- Indopseudodon_salwenianus
| +-----+
| | | +-- Indopseudodon_indawgyiensis
| | +--+
| | +-- Indopseudodon_bogani
| |
| | +-------- Nyeinchanconcha_nyeinchani
| | |
| | +-------- Thaiconcha_callifera
+--------+ |
| | +-------- Sundadontina_cumingii
| | |
| | +-------- Sundadontina_tumida[1]
| | |
| | +-------- Sundadontina_tumida[2]
| | |
| | | +----- Pilsbryoconcha_exilis[2]
| +--+--+
| | | +-- Pilsbryoconcha_carinifera
| | +--+
| | +-- Pilsbryoconcha_exilis[1]
| |
| | +----- Pseudodon_vagulus
| +--+
| | | +-- Pseudodon_mekongi
| | +--+
| | +-- Pseudodon_vondembuschianus
| |
| | +----- Bineurus_loeiensis
| +--+
| | +-- Bineurus_mouhotii
| +--+
| +-- Bineurus_exilis
|
| +-------------- Amblema_plicata
| |
| +-------------- Toxolasma_parvum
| |
| | +-- Obliquaria_reflexa
| +-----------+
| | +-- Plectomerus_dombeyanus
| |
| | +----------- Elliptoideus_sloatianus
| +--+
| | | +-------- Elliptio_complanata
| | +--+
| | | +----- Eurynia_dilatata
+-----+ +--+
| | | +-- Fusconaia_flava
| | +--+
| | +-- Pleurobema_sintoxia
| |
| | +-------- Ellipsaria_lineolata
| | |
| | +-------- Potamilus_leptodon
| | |
| | +-------- Medionidus_conradicus
| | |
--+ | +-------- Potamilus_alatus
| | |
| | +-------- Ptychobranchus_subtentus
| +-----+
| +-------- Truncilla_truncata
| |
| +-------- Venustaconcha_ellipsiformis
| |
| +-------- Cambarunio_iris
| |
| +-------- Leaunio_lienosus
| |
| | +----- Ortmanniana_ligamentina
| +--+
| | +-- Lampsilis_cardium
| +--+
| +-- Lampsilis_ornata
|
| +-------------- Hyriopsis_desowitzi
| |
| +-------------- Hyriopsis_myersiana
| |
| +-------------- Hyriopsis_panhai
| |
| +--+ +-- Hyriopsis_bialata
| | +-----------+
| | | +-- Hyriopsis_khoratensis
| | |
| | | +----------- Rectidens_sumatrensis
| | | |
| | +--+ +-- Ensidens_telus
| | | +-----+
| | | | +-- Ensidens_ingallsianus
| | +--+
| | | +----- Ensidens_sagittarius
| | +--+
+--+ | +-- Ensidens_spiculus
| | +--+
| | +-- Ensidens_jaculus
| |
| | +----------- Physunio_modelli
| | |
| | +----------- Lens
| | |
| | +----------- Physunio_superbus
| | |
| | | +-------- Yaukthwa_inlenensis
| | +--+
| +-----+ | +----- Yaukthwa_panhai
| | +--+
| | | +-- Yaukthwa_nesemanni
| | +--+
| | +-- Yaukthwa_rectangularis
| |
| | +-- Lens_comptus
| | +-----+
| | | +-- Lens_novoselovi
| +--+
| | +----- Lens_eximius
| +--+
| | +-- Lens_contradens
| +--+
| +-- Lens_micropterus
|
| +----------------- Acuticosta_chinensis
| |
| +----------------- Alasmidonta_marginata
| |
| +----------------- Anodonta_anatina
| |
| +----------------- Anemina_arcaeformis
| |
| +----------------- Anodonta_cygnea
| |
| +----------------- Cristaria_plicata
| |
| +----------------- Anodonta_nuttalliana
| |
| +----------------- Lasmigona_costata
| |
| +----------------- Lepidodesma_languilati
| |
| +----------------- Pseudanodonta_complanata
| |
| +----------------- Pyganodon_grandis
| |
| +----------------- Simpsonaias_ambigua
| |
| +----------------- Strophitus_undulatus
| |
| | +-- Schistodesmus_lampreyanus
| +--------------+
| | +-- Schistodesmus_spinosus
| |
| | +-- Utterbackia_imbecillis
| +--------------+
| | +-- Utterbackia_peninsularis
| |
| | +-------- Lanceolaria_gladiola
| +--------+
| | | +----- Lanceolaria_grayii
| | +--+
+--+ | +-- Lanceolaria_lanceolata
| +--+
| +-- Lanceolaria_triformis
|
| +-------- Unio_tumidus
+--------+
| | +----- Unio_crassus
| +--+
| | +-- Unio_delphinus
| +--+
| +-- Unio_pictorum
|
| +-------- Aculamprotula_tortuosa
| |
+--------+ +-- Aculamprotula_coreana
| | +--+
| | | +-- Aculamprotula_scripta
| +--+
| | +-- Aculamprotula_fibrosa
| +--+
| +-- Aculamprotula_tientsinensis
|
| +-------- Cuneopsis_heudei
| |
| +-------- Cuneopsis_celtiformis
| |
+--------+-------- Tchangsinaia_piscicula
| |
| +-------- Cuneopsis_rufescens
| |
| | +----- Nodularia_douglasiae[2]
| +--+
| | +-- Nodularia_douglasiae[1]
| +--+
| +-- Nodularia_nuxpersicae
|
| +-------------- Pletholophus_tenuis
| |
+--+ +-- Sinanodonta_angula
| +--------+
| | +-- Sinanodonta_lucida
+--+
| +-------- Sinanodonta_woodiana[1]
+--+
| +----- Sinanodonta_jourdyi
+--+
| +-- Sinanodonta_elliptica
+--+
+-- Sinanodonta_woodiana[2]
Lydeard et al., 2000 [Publ 2654], fig. 2a
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 3 |
3 |
1 |
1 |
14 |
12 |
3 |
19.0 |
+----- Quadrula_nobilis **
+-----------+
| | +-- Pustulosa_infucata **
| +--+
--+ +-- Pustulosa_succissa **
|
| +-------------- Pleurobema_decisum
+--+
| +----------- Fusconaia_flava
+--+
| +-------- Fusconaia_burkei[1]
+--+
| +----- Fusconaia_burkei[2]
+--+
| +-- Fusconaia_burkei[3]
+--+
+-- Fusconaia_escambia
Lydeard et al., 2000 [Publ 2654], fig. 2b
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 3 |
3 |
0 |
1 |
14 |
12 |
3 |
20.0 |
+-- Reginaia_rotulata[1]
|
+-----------------+-- Reginaia_rotulata[2]
| |
| +-- Reginaia_ebenus
|
--+ +-- Quadrula_nobilis **
| |
| +--------------+-- Pustulosa_infucata **
| | |
+--+ +-- Pustulosa_succissa **
|
| +-------------- Pleurobema_decisum
+--+
| +----------- Fusconaia_flava
+--+
| +-------- Fusconaia_burkei[1]
+--+
| +----- Fusconaia_burkei[2]
+--+
| +-- Fusconaia_burkei[3]
+--+
+-- Fusconaia_escambia
Lydeard et al., 2000 [Publ 2654], fig. 2c
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 3 |
3 |
1 |
1 |
13 |
11 |
3 |
19.0 |
+----- Quadrula_nobilis **
--+
| +-- Pustulosa_infucata **
+--+
+-- Pustulosa_succissa **
Lydeard et al., 2000 [Publ 2654], fig. 2d
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 3 |
3 |
0 |
1 |
13 |
11 |
3 |
20.0 |
+----- Lampsilis_teres
+--------------+
| | +-- Obovaria_unicolor
| +--+
| +-- Obovaria_olivaria
--+
| +-- Quadrula_nobilis **
| |
| +--------------+-- Pustulosa_infucata **
| | |
+--+ +-- Pustulosa_succissa **
|
| +-------------- Pleurobema_decisum
+--+
| +----------- Fusconaia_flava
+--+
| +-------- Fusconaia_burkei[1]
+--+
| +----- Fusconaia_burkei[2]
+--+
| +-- Fusconaia_burkei[3]
+--+
+-- Fusconaia_escambia
Amaro et al., 2017 [Publ 3791], fig. 2
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 3 |
3 |
1 |
1 |
15 |
4 |
2 |
21.0 |
+----- Toxolasma_parvum
+--+
| | +-- Venustaconcha_ellipsiformis
| +--+
--+ +-- Lampsilis_siliquoidea
|
| +----- Quadrula_quadrula[2] **
+--+
| +-- Quadrula_quadrula[1] **
+--+
+-- Quadrula_nobilis **
Chase et al., 2018 [Publ 3847], fig. 4
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 3 |
3 |
1 |
2 |
36 |
14 |
3 |
19.0 |
+-------- Fusconaia_flava
--+
| +----- Megalonaias_nervosa **
+--+
| +-- Cyclonaias_tuberculata **
+--+
+-- Pustulosa_pustulosa **
Graf & Cummings, 2006 [Publ 1968], fig. 2
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 2 |
2 |
0 |
1 |
45 |
12 |
3 |
-- |
+-- Quadrula_verrucosa **
+--------------------+
| +-- Quadrula_quadrula **
|
--+ +-------------------- Amblema_plicata
| |
| | +----- Eurynia_dilatata
+--+ +-----------+
| | | +-- Fusconaia_flava
| | +--+
| | +-- Pleurobema_sintoxia
+--+
| +-------------- Obliquaria_reflexa
| |
| +-------------- Truncilla_truncata
+--+
+-------------- Ptychobranchus_fasciolaris
|
| +----------- Ligumia_recta
+--+
| +-------- Cambarunio_iris
+--+
| +----- Epioblasma_triquetra
+--+
| +-- Lampsilis_cardium
+--+
+-- Ortmanniana_ligamentina
Huang et al., 2002 [Publ 2657], fig. 1
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 2 |
14 |
0 |
2 |
32 |
12 |
3 |
32.0 |
+----------------------------- Megalonaias_nervosa **
|
| +----- Elliptoideus_sloatianus
--+ +--------------------+
| | | +-- Uniomerus_carolinianus **
| | +--+
+--+ +-- Pleurobema_pyriforme
|
| +----------------------- Amblema_plicata
| |
+--+ +-- Obliquaria_reflexa
| +-----------------+
| | +-- Plectomerus_dombeyanus
+--+
| +----------------- Medionidus_conradicus
+--+
| +-------------- Potamilus_fragilis
+--+
| +----------- Ptychobranchus_subtentus
+--+
| +-------- Potamilus_purpuratus
+--+
| +----- Lampsilis_teres
+--+
| +-- Villosa_delumbis
+--+
+-- Ellipsaria_lineolata
Huang et al., 2002 [Publ 2657], fig. 2
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 2 |
14 |
0 |
2 |
32 |
12 |
3 |
32.0 |
+----------- Megalonaias_nervosa **
|
| +----- Elliptoideus_sloatianus
| +--+
--+ | | +-- Uniomerus_carolinianus **
| | +--+
| | +-- Pleurobema_pyriforme
| |
| | +----- Potamilus_fragilis
+--+ |
| +----- Medionidus_conradicus
| |
| +----- Amblema_plicata
| |
| | +-- Obliquaria_reflexa
+--+--+
| +-- Plectomerus_dombeyanus
|
| +-- Villosa_delumbis
| |
| +-- Ellipsaria_lineolata
| |
+--+-- Lampsilis_teres
|
+-- Potamilus_purpuratus
|
+-- Ptychobranchus_subtentus
Chapman et al., 2008 [Publ 2667], fig. 1
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 2 |
2 |
0 |
1 |
19 |
18 |
4 |
-- |
+-- Quadrula_quadrula **
+--------------------------------------+
| +-- Pustulosa_pustulosa **
|
--+ +-- Pleurobema_sintoxia
| +-----------------------------------+
| | +-- Fusconaia_flava
+--+
| +----------------------------------- Amblema_plicata
| |
+--+ +-------------------------------- Plectomerus_dombeyanus
| |
+--+ +----------------------------- Popenaias_popeii
| |
+--+ +-------------------------- Toxolasma_lividum
| |
+--+ +----------------------- Obliquaria_reflexa
| |
| | +-- Cyrtonaias_tampicoensis
+--+ +-----------------+
| | +-- Glebula_rotundata
| |
+--+ +-- Ptychobranchus_fasciolaris
| +--------------+
| | +-- Lemiox_rimosus
| |
+--+ +-- Venustaconcha_ellipsiformis
| +-----------+
| | +-- Obovaria_olivaria
+--+
| +----------- Hamiota_subangulata
+--+
| +-------- Lampsilis_ovata
+--+
| +----- Ortmanniana_ligamentina
+--+
| +-- Lampsilis_straminea
+--+
+-- Lampsilis_hydiana
Curole & Kocher, 2005 [Publ 2682], fig. 1
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 2 |
2 |
0 |
1 |
9 |
7 |
2 |
-- |
+----- Amblema_plicata
+--------+
| | +-- Quadrula_quadrula **
| +--+
--+ +-- Pustulosa_pustulosa **
|
| +----------- Plectomerus_dombeyanus
+--+
| +-------- Leaunio_lienosus
+--+
| +----- Lampsilis_straminea
+--+
| +-- Lampsilis_teres
+--+
+-- Potamilus_purpuratus
Curole & Kocher, 2005 [Publ 2682], fig. 1-M
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 2 |
2 |
0 |
1 |
10 |
7 |
2 |
-- |
+-- Quadrula_quadrula **
+--------------+
| +-- Pustulosa_pustulosa **
--+
| +-------------- Plectomerus_dombeyanus
| |
+--+ +----------- Amblema_plicata
| |
+--+ +-- Potamilus_purpuratus
| +-----+
| | +-- Glebula_rotundata
+--+
| +----- Leaunio_lienosus
+--+
| +-- Lampsilis_straminea
+--+
+-- Lampsilis_teres
Curole & Kocher, 2002 [Publ 2712], fig. 1-F
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 2 |
2 |
0 |
1 |
2 |
1 |
1 |
-- |
+----- Lampsilis_teres
--+
| +-- Quadrula_quadrula **
+--+
+-- Pustulosa_pustulosa **
Curole & Kocher, 2002 [Publ 2712], fig. 1a-M
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 2 |
2 |
0 |
1 |
2 |
1 |
1 |
-- |
+-- Quadrula_quadrula **
--+
+-- Pustulosa_pustulosa **
Curole & Kocher, 2002 [Publ 2712], fig. 1b-M
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 2 |
2 |
0 |
1 |
2 |
1 |
1 |
-- |
+-- Quadrula_quadrula **
--+
+-- Pustulosa_pustulosa **
Hoeh et al., 2009 [Publ 2716], fig. 3
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 2 |
2 |
0 |
1 |
45 |
12 |
3 |
-- |
+----- Obliquaria_reflexa
+--+
| | +-- Quadrula_verrucosa **
| +--+
--+ +-- Quadrula_quadrula **
|
| +----- Eurynia_dilatata
+--+
| +-- Pleurobema_sintoxia
+--+
+-- Fusconaia_flava
Hoeh et al., 2009 [Publ 2716], fig. 4
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 2 |
2 |
0 |
1 |
44 |
11 |
2 |
-- |
+-- Eurynia_dilatata
|
+-----+-- Pleurobema_sintoxia
| |
--+ +-- Fusconaia_flava
|
| +----- Obliquaria_reflexa
+--+
| +-- Quadrula_verrucosa **
+--+
+-- Quadrula_quadrula **
Hoeh et al., 2009 [Publ 2716], fig. 5
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 2 |
2 |
0 |
1 |
44 |
11 |
3 |
-- |
+----- Obliquaria_reflexa
+-----+
| | +-- Quadrula_verrucosa **
| +--+
| +-- Quadrula_quadrula **
|
--+ +-------- Amblema_plicata
| |
| | +----- Eurynia_dilatata
| +--+
| | | +-- Pleurobema_sintoxia
+--+ +--+
| +-- Fusconaia_flava
|
| +----- Truncilla_truncata
| |
| +----- Ptychobranchus_fasciolaris
+--+
| +-- Ortmanniana_ligamentina
| |
| +-- Cambarunio_iris
+--+
+-- Ligumia_recta
|
+-- Epioblasma_triquetra
Chapman et al., 2008 [Publ 2744], fig. 2
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 2 |
2 |
0 |
1 |
19 |
18 |
4 |
-- |
+-- Quadrula_quadrula **
+--------------------------------------+
| +-- Pustulosa_pustulosa **
|
--+ +-- Pleurobema_sintoxia
| +-----------------------------------+
| | +-- Fusconaia_flava
+--+
| +----------------------------------- Amblema_plicata
| |
+--+ +-------------------------------- Plectomerus_dombeyanus
| |
+--+ +----------------------------- Popenaias_popeii
| |
+--+ +-------------------------- Toxolasma_lividum
| |
+--+ +----------------------- Obliquaria_reflexa
| |
| | +-- Cyrtonaias_tampicoensis
+--+ +-----------------+
| | +-- Glebula_rotundata
| |
+--+ +-- Ptychobranchus_fasciolaris
| +--------------+
| | +-- Lemiox_rimosus
| |
+--+ +-- Venustaconcha_ellipsiformis
| +-----------+
| | +-- Obovaria_olivaria
+--+
| +----------- Hamiota_subangulata
+--+
| +-------- Lampsilis_ovata
+--+
| +----- Ortmanniana_ligamentina
+--+
| +-- Lampsilis_straminea
+--+
+-- Lampsilis_hydiana
Jadhav & Jamkhedkar, 2009 [Publ 3697], fig. 11
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 2 |
2 |
0 |
1 |
24 |
11 |
3 |
-- |
+-- Quadrula_quadrula **
--+
+-- Quadrula_verrucosa **
Lopes-Lima et al., 2017 [Publ 3745], fig. 1-2
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 2 |
2 |
0 |
1 |
79 |
7 |
3 |
-- |
+-------- Amblema_plicata
+--+
| | +----- Cambarunio_iris
| +--+
| | +-- Lampsilis_cardium
| +--+
--+ +-- Ortmanniana_ligamentina
|
| +-- Quadrula_quadrula **
| +-----+
| | +-- Quadrula_verrucosa **
+--+
| +----- Elliptio_complanata
+--+
| +-- Pleurobema_sintoxia
+--+
+-- Eurynia_dilatata
Bolotov et al., 2017 [Publ 3761], fig. S1
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 2 |
2 |
0 |
1 |
139 |
7 |
3 |
-- |
+-------- Amblema_plicata
+--+
| | +----- Ortmanniana_ligamentina
| +--+
| | +-- Lampsilis_cardium
| +--+
--+ +-- Cambarunio_iris
|
| +-- Quadrula_quadrula **
| +-----+
| | +-- Quadrula_verrucosa **
+--+
| +----- Pleurobema_sintoxia
+--+
| +-- Eurynia_dilatata
+--+
+-- Elliptio_complanata
Bolotov et al., 2017 [Publ 3761], fig. S2
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 2 |
2 |
0 |
1 |
139 |
7 |
3 |
-- |
+-------- Amblema_plicata
+--+
| | +----- Cambarunio_iris
| +--+
| | +-- Ortmanniana_ligamentina
| +--+
--+ +-- Lampsilis_cardium
|
| +-- Quadrula_quadrula **
| +-----+
| | +-- Quadrula_verrucosa **
+--+
| +----- Pleurobema_sintoxia
+--+
| +-- Elliptio_complanata
+--+
+-- Eurynia_dilatata
Bolotov et al., 2017 [Publ 3764], fig. S1
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 2 |
9 |
0 |
1 |
129 |
7 |
3 |
27.0 |
+-- Amblema_plicata
|
+-- Ortmanniana_ligamentina
|
+-- Lampsilis_cardium
|
+-- Cambarunio_iris
|
--+-- Elliptio_complanata
|
+-- Eurynia_dilatata
|
+-- Pleurobema_sintoxia
|
+-- Quadrula_quadrula **
|
+-- Quadrula_verrucosa **
Huang et al., 2019 [Publ 3843], fig. 3
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 2 |
2 |
0 |
2 |
55 |
8 |
2 |
-- |
+-- Quadrula_quadrula **
+-----------------+
| +-- Uniomerus_tetralasmus **
--+
| +----------------- Elliptio_complanata
| |
+--+ +-------------- Toxolasma_parvum
| |
+--+ +-- Potamilus_leptodon
| +--------+
| | +-- Potamilus_alatus
+--+
| +-------- Leaunio_lienosus
+--+
| +----- Venustaconcha_ellipsiformis
+--+
| +-- Lampsilis_cardium
+--+
+-- Lampsilis_ornata
Pieri et al., 2018 [Publ 3867], fig. 3
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 2 |
2 |
0 |
1 |
35 |
34 |
1 |
-- |
+-- Quadrula_quadrula **
+-----------------------------------+
| +-- Pustulosa_petrina **
|
| +----------------------------------- Eurynia_dilatata
| |
--+ | +----- Pleurobema_clava
| +-----------------------------+
| | | +-- Pleurobema_riddellii
| | +--+
| | +-- Pleurobema_sintoxia
+--+
| +----- Fusconaia_subrotunda
| +--------------------------+
| | | +-- Fusconaia_cor
| | +--+
| | +-- Fusconaia_cuneolus
| |
+--+ +----------------------------- Fusconaia_ozarkensis
| |
| | +----------- Fusconaia_cerina[3]
| | |
| | +--------------+ +-------- Fusconaia_flava[1]
+--+ | | |
| | +--+ +----- Fusconaia_flava[3]
| | | |
| | +--+ +-- Fusconaia_cerina[1]
| | | |
| | +--+-- Fusconaia_cerina[2]
+--+ |
| +-- Fusconaia_flava[2]
|
| +----------------------- Fusconaia_masoni
| |
| | +-- Fusconaia_burkei
| +--------------------+
+--+ +-- Fusconaia_escambia
|
| +-------------------- Fusconaia_mitchelli
| |
+--+ +----------------- Fusconaia_chunii
| |
| | +-------------- Fusconaia_askewi[9]
+--+ |
| | +----------- Fusconaia_askewi[6]
| | |
+--+ +----------- Fusconaia_askewi[7]
| |
| +----------- Fusconaia_askewi[8]
| |
| +----------- Fusconaia_lananensis[5]
+--+
+----------- Fusconaia_lananensis[6]
|
| +-------- Fusconaia_askewi[5]
| |
| +-------- Fusconaia_lananensis[4]
| |
+--+ +----- Fusconaia_askewi[2]
| |
| +----- Fusconaia_askewi[3]
| |
+--+----- Fusconaia_askewi[4]
|
| +-- Fusconaia_askewi[1]
| |
| +-- Fusconaia_lananensis[1]
+--+
+-- Fusconaia_lananensis[2]
|
+-- Fusconaia_lananensis[3]
Burlakova et al., 2019 [Publ 3899], fig. 1
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 2 |
2 |
0 |
2 |
31 |
31 |
4 |
-- |
+-- Quadrula_quadrula **
+-----------------------+
| +-- Uniomerus_declivis **
|
| +----------------------- Plectomerus_dombeyanus
--+ |
| | +-------------------- Popenaias_popeii
| | |
| | | +-- Elliptio_crassidens
| | +-----------------+
+--+ | +-- Pleurobema_rubellum
| |
| | +-- Cyrtonaias_tampicoensis
| | +--+
| | | +-- Glebula_rotundata
| +--------------+
| | | +-- Reginaia_ebenus
+--+ +--+
| +-- Obliquaria_reflexa
|
| +----------------- Toxolasma_lividum
| |
| | +-- Potamilus_fragilis
| | +-----------+
| | | +-- Potamilus_metnecktayi
| | |
| | | +----- Medionidus_acutissimus
| | +--------+
+--+ | | +-- Lemiox_rimosus
| | +--+
| | +-- Dromus_dromas
| |
| | +----- Ellipsaria_lineolata
| | |
| +--------+ +-- Truncilla_cognata
| | | |
+--+ | +-- Truncilla_donaciformis
| +--+
| +-- Truncilla_macrodon
| |
| +-- Truncilla_truncata
|
| +----------- Cambarunio_simus
| |
| +----------- Leaunio_lienosus
| |
| +----------- Villosa_villosa
| |
| | +-- Sagittunio_nasutus
| +--------+
+--+ +-- Lampsilis_ovata
|
| +----- Venustaconcha_pleasii
+-----+
| | +-- Epioblasma_rangiana
| +--+
| +-- Obovaria_unicolor
|
| +-------- Lampsilis_teres
+--+
| +----- Lampsilis_siliquoidea
+--+
| +-- Lampsilis_hydiana
+--+
+-- Lampsilis_straminea
Pfeiffer et al., 2019 [Publ 3937], fig. 4-5
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 2 |
2 |
0 |
2 |
55 |
13 |
4 |
-- |
+-- Quadrula_quadrula **
+--------------------+
| +-- Uniomerus_tetralasmus **
|
| +----- Elliptoideus_sloatianus
--+ +--------------+
| | | +-- Elliptio_complanata
| | +--+
| | +-- Pleurobema_strodeanum
| |
+--+ +----- Reginaia_ebenus
| +-----------+
| | | +-- Plectomerus_dombeyanus
| | +--+
| | +-- Amblema_plicata
+--+
| +-- Psoronaias_semigranosa
| +-----------+
| | +-- Popenaias_popeii
+--+
| +----------- Obliquaria_reflexa
+--+
| +-------- Cyrtonaias_tampicoensis
+--+
| +----- Leaunio_lienosus
+--+
| +-- Lampsilis_cardium
+--+
+-- Pachynaias_spheniopsis
Pfeiffer et al., 2019 [Publ 3937], fig. 6
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 2 |
2 |
0 |
2 |
55 |
13 |
4 |
-- |
+-- Quadrula_quadrula **
+-----------------+
| +-- Uniomerus_tetralasmus **
|
| +----- Elliptoideus_sloatianus
| +--+
| | | +-- Elliptio_complanata
--+ | +--+
| +--------+ +-- Pleurobema_strodeanum
| | |
| | | +----- Reginaia_ebenus
| | +--+
| | | +-- Plectomerus_dombeyanus
+--+ +--+
| +-- Amblema_plicata
|
| +-- Psoronaias_semigranosa
| +-----------+
| | +-- Popenaias_popeii
+--+
| +----------- Obliquaria_reflexa
+--+
| +-------- Cyrtonaias_tampicoensis
+--+
| +----- Leaunio_lienosus
+--+
| +-- Lampsilis_cardium
+--+
+-- Pachynaias_spheniopsis
Inoue et al., 2020 [Publ 3941], fig. 3
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 2 |
2 |
0 |
1 |
42 |
41 |
3 |
-- |
+-- Pustulosa_petrina **
+-----------------------------------------+
| +-- Quadrula_quadrula **
|
| +-------------- Parvaspina_collina
| |
| | +-- Eurynia_dilatata
| +--------------------------+ +--------+
| | | | +-- Hemistena_lata
--+ | | |
| | +--+ +-- Pleurobema_clava
| | | +-----+
| | | | +-- Pleurobema_sintoxia
| | +--+
| | | +----- Pleuronaia_dolabelloides
| | +--+
| | | +-- Fusconaia_flava
+--+ +--+
| +-- Fusconaia_subrotunda
|
| +-- Amblema_plicata
| +--+
| | +-- Amblema_elliottii
| +--------------------------------+
| | | +-- Reginaia_ebenus
| | +--+
| | +-- Reginaia_rotulata
| |
+--+ +----------------------------------- Toxolasma_lividum
| |
| | +-------- Potamilus_fragilis
| | |
| | +-----------------------+ +-- Potamilus_ohiensis
+--+ | | +--+
| | | | +-- Potamilus_amphichaenus
| | +--+
| | | +-- Potamilus_metnecktayi
| | +--+
| | +-- Potamilus_purpuratus
+--+
| +-------- Ellipsaria_lineolata
| +--------------------+
| | | +----- Truncilla_macrodon
| | +--+
| | | +-- Truncilla_donaciformis
| | +--+
| | +-- Truncilla_cognata
+--+
| +-------------------------- Leaunio_lienosus
| |
| | +-------------------- Lampsilis_bracteata
| | |
| | +--+ +----------------- Lampsilis_fasciola
| | | | |
+--+ | +--+ +-------------- Lampsilis_ornata
| | | |
| | +--+ +-- Lampsilis_ovata
| | | +--------+
| | | | +-- Lampsilis_cariosa
| | +--+
| | | +-------- Lampsilis_cardium[3]
+--+ +--+
| | +----- Lampsilis_cardium[2]
| +--+
| | +-- Lampsilis_satura
| +--+
| +-- Lampsilis_cardium[1]
|
| +-- Lampsilis_higginsii
| +--------------+
| | +-- Ortmanniana_ligamentina
+-----+
| +-------------- Lampsilis_teres
| |
+--+ +-- Lampsilis_radiata
| +--------+
| | +-- Lampsilis_siliquoidea
+--+
| +-------- Lampsilis_bergmanni
+--+
| +----- Lampsilis_hydiana
+--+
| +-- Lampsilis_virescens
+--+
+-- Lampsilis_straminea
Inoue et al., 2020 [Publ 3941], fig. 4
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 2 |
2 |
0 |
1 |
41 |
40 |
3 |
-- |
+-- Pustulosa_petrina **
+--------------------------------------------+
| +-- Quadrula_quadrula **
|
| +-- Reginaia_ebenus
| +-----------------------------------------+
--+ | +-- Reginaia_rotulata
| |
| | +-------------- Parvaspina_collina
| | |
| | | +-- Eurynia_dilatata
| | +--------------------------+ +--------+
+--+ | | | +-- Hemistena_lata
| | | |
| | +--+ +-- Pleurobema_clava
| | | +-----+
| | | | +-- Pleurobema_sintoxia
| | +--+
| | | +----- Pleuronaia_dolabelloides
+--+ +--+
| | +-- Fusconaia_flava
| +--+
| +-- Fusconaia_subrotunda
|
| +-- Amblema_plicata
| +-----------------------------------+
| | +-- Amblema_elliottii
| |
+--+ +----------------------------------- Toxolasma_lividum
| |
| | +-------------------------------- Ellipsaria_lineolata
+--+ |
| | +----- Truncilla_cognata
| | +-----------------------+
| | | | +-- Truncilla_macrodon
+--+ | +--+
| | +-- Truncilla_donaciformis
| |
| | +-------- Potamilus_fragilis
| | |
+--+ +-----------------+ +-- Potamilus_ohiensis
| | | +--+
| | | | +-- Potamilus_amphichaenus
| | +--+
| | | +-- Potamilus_metnecktayi
| | +--+
| | +-- Potamilus_purpuratus
+--+
| +----------------------- Leaunio_lienosus
| |
| | +----------------- Lampsilis_bracteata
| | |
| | +--+ +-- Lampsilis_ornata
| | | | +-----------+
+--+ | | | +-- Lampsilis_fasciola
| | +--+
| | | +----------- Lampsilis_ovata
| | +--+
| | | +-------- Lampsilis_cardium[3]
| | +--+
+--+ | +----- Lampsilis_cardium[2]
| +--+
| | +-- Lampsilis_satura
| +--+
| +-- Lampsilis_cardium[1]
|
| +-------------- Lampsilis_teres
| |
| | +-- Lampsilis_higginsii
+-----+ +--------+
| | +-- Ortmanniana_ligamentina
| |
+--+ +-- Lampsilis_radiata
| +-----+
| | +-- Lampsilis_siliquoidea
| |
+--+ +-- Lampsilis_hydiana
| +--+
| | +-- Lampsilis_bergmanni
+--+
| +-- Lampsilis_virescens
+--+
+-- Lampsilis_straminea
Fukata & Iigo, 2020 [Publ 4106], fig. 1
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 2 |
2 |
0 |
2 |
52 |
9 |
2 |
-- |
+-- Quadrula_quadrula **
+-----------------+
| +-- Uniomerus_tetralasmus **
--+
| +----------------- Elliptio_complanata
| |
+--+ +-- Potamilus_leptodon
| +-----------+
| | +-- Potamilus_alatus
+--+
| +----------- Leaunio_lienosus
| |
+--+ +-- Lampsilis_cardium
| +-----+
| | +-- Lampsilis_ornata
+--+
| +----- Venustaconcha_ellipsiformis
+--+
| +-- Lampsilis_powellii
+--+
+-- Lampsilis_siliquoidea
Teiga-Teixeira et al., 2020 [Publ 4173], fig. 1
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 2 |
2 |
0 |
2 |
17 |
12 |
4 |
-- |
+-- Uniomerus_tetralasmus **
+-----------------------+
| +-- Quadrula_quadrula **
|
--+ +-- Elliptio_complanata
| +--------------------+
| | +-- Pleurobema_oviforme
+--+
| +-------------------- Amblema_plicata
| |
+--+ +----------------- Popenaias_popeii
| |
+--+ +-------------- Toxolasma_parvum
| |
| | +-- Potamilus_alatus
+--+ +--------+
| | +-- Potamilus_leptodon
| |
+--+ +-- Lampsilis_cardium
| +-----+
| | +-- Lampsilis_ornata
+--+
| +----- Venustaconcha_ellipsiformis
+--+
| +-- Lampsilis_powellii
+--+
+-- Lampsilis_siliquoidea
Porto-Hannes et al., 2021 [Publ 4227], fig. 3
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 2 |
16 |
0 |
1 |
14 |
14 |
3 |
33.0 |
+-------------------- Quadrula_quadrula **
|
+-------------------- Quadrula_verrucosa **
|
--+ +-- Eurynia_dilatata
| +--------------+
| | +-- Pleurobema_sintoxia
| |
| | +-- Amblema_plicata
+--+ +-----------+
| | +-- Amblema_elliottii
| |
| | +-- Obovaria_olivaria
+--+ +--------+
| | +-- Ortmanniana_ligamentina
| |
| | +-- Obliquaria_reflexa
| +--------+
+--+ +-- Lampsilis_ornata
|
| +----- Lampsilis_radiata[2]
| +--+
| | | +-- Lampsilis_siliquoidea
| | +--+
+--+ +-- Lampsilis_radiata[1]
|
| +----- Lampsilis_virescens
+--+
| +-- Lampsilis_hydiana
+--+
+-- Lampsilis_straminea
Porto-Hannes et al., 2021 [Publ 4227], fig. 4
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 2 |
12 |
0 |
1 |
10 |
10 |
3 |
29.0 |
+-------------------- Quadrula_quadrula **
|
+-------------------- Pustulosa_pustulosa **
--+
| +----------------- Pleurobema_sintoxia
| |
+--+ +-------------- Amblema_plicata
| |
+--+ +----------- Obovaria_olivaria
| |
+--+ +-------- Ortmanniana_ligamentina
| |
+--+ +----- Lampsilis_hydiana
| |
| +----- Lampsilis_straminea
+--+
| +-- Lampsilis_siliquoidea
| |
| +-- Lampsilis_radiata
+--+
+-- Lampsilis[1]
|
+-- Lampsilis[2]
Wu et al., 2022 [Publ 4238], fig. 3
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 2 |
2 |
0 |
2 |
72 |
13 |
4 |
-- |
+-- Uniomerus_tetralasmus **
+--------------------------+
| +-- Quadrula_quadrula **
|
--+ +-- Pleurobema_oviforme
| +-----------------------+
| | +-- Elliptio_complanata
+--+
| +----------------------- Amblema_plicata
| |
+--+ +-------------------- Popenaias_popeii
| |
+--+ +----------------- Toxolasma_parvum
| |
+--+ +-- Potamilus_alatus
| +-----------+
| | +-- Potamilus_leptodon
+--+
| +----------- Leaunio_lienosus
| |
+--+ +-- Lampsilis_cardium
| +-----+
| | +-- Lampsilis_ornata
+--+
| +----- Venustaconcha_ellipsiformis
+--+
| +-- Lampsilis_siliquoidea
+--+
+-- Lampsilis_powellii
Jeratthitikul et al., 2022 [Publ 4260], fig. S1, 1
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 2 |
2 |
0 |
1 |
74 |
7 |
3 |
-- |
+-------- Amblema_plicata
+--+
| | +----- Ortmanniana_ligamentina
| +--+
| | +-- Cambarunio_iris
| +--+
--+ +-- Lampsilis_cardium
|
| +-- Quadrula_quadrula **
| +-----+
| | +-- Quadrula_verrucosa **
+--+
| +----- Elliptio_complanata
+--+
| +-- Pleurobema_sintoxia
+--+
+-- Eurynia_dilatata
Jeratthitikul et al., 2022 [Publ 4260], fig. S2, 2
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 2 |
2 |
0 |
1 |
74 |
7 |
3 |
-- |
+-------- Amblema_plicata
+--+
| | +----- Ortmanniana_ligamentina
| +--+
| | +-- Lampsilis_cardium
| +--+
--+ +-- Cambarunio_iris
|
| +-- Quadrula_quadrula **
| +-----+
| | +-- Quadrula_verrucosa **
+--+
| +----- Pleurobema_sintoxia
+--+
| +-- Eurynia_dilatata
+--+
+-- Elliptio_complanata
Mejia-Trujillo & Smith, 2022 [Publ 4274], fig. 2-F
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 2 |
2 |
0 |
2 |
13 |
13 |
4 |
-- |
+-------- Amblema_plicata
|
--+ +-- Quadrula_quadrula **
| +--+
| | +-- Uniomerus_tetralasmus **
+--+
| +-- Pleurobema_oviforme
+--+
+-- Elliptio_complanata
Dai et al., 2023 [Publ 4285], fig. 5
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 2 |
2 |
0 |
2 |
51 |
13 |
4 |
-- |
+-- Quadrula_quadrula **
+--------------------------+
| +-- Uniomerus_tetralasmus **
|
--+ +-- Elliptio_complanata
| +-----------------------+
| | +-- Pleurobema_oviforme
+--+
| +----------------------- Amblema_plicata
| |
+--+ +-------------------- Popenaias_popeii
| |
+--+ +----------------- Toxolasma_parvum
| |
+--+ +-- Potamilus_leptodon
| +-----------+
| | +-- Potamilus_alatus
+--+
| +----------- Leaunio_lienosus
| |
+--+ +-- Lampsilis_cardium
| +-----+
| | +-- Lampsilis_ornata
+--+
| +----- Venustaconcha_ellipsiformis
+--+
| +-- Lampsilis_powellii
+--+
+-- Lampsilis_siliquoidea
Graf & Ó Foighil, 2000 [Publ 2648], fig. 2A
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 1 |
1 |
0 |
1 |
29 |
16 |
3 |
-- |
+-------- Quadrula_quadrula **
+--------+
| | +----- Eurynia_dilatata
| +--+
| | +-- Pleurobema_sintoxia
| +--+
--+ +-- Fusconaia_flava
|
| +-------------- Amblema_plicata
| |
| | +----------- Ligumia_recta
| | |
+--+ +----------- Truncilla_truncata
| |
| | +-- Ptychobranchus_fasciolaris
| +--------+
+--+ +-- Sagittunio_nasutus
|
| +----- Lampsilis_siliquoidea
| +--+
| | | +-- Epioblasma_triquetra
| | +--+
| | +-- Epioblasma_brevidens
+--+
| +----- Ortmanniana_ligamentina
| |
| +----- Cambarunio_iris
+--+
+----- Leaunio_vanuxemensis
|
| +-- Lampsilis_cardium
+--+
+-- Lampsilis_fasciola
Graf & Ó Foighil, 2000 [Publ 2648], fig. 2B
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 1 |
1 |
0 |
1 |
29 |
16 |
3 |
-- |
+-------- Quadrula_quadrula **
+-----------------+
| | +----- Eurynia_dilatata
| +--+
| | +-- Pleurobema_sintoxia
--+ +--+
| +-- Fusconaia_flava
|
| +----------------------- Amblema_plicata
| |
+--+ +-------------------- Truncilla_truncata
| |
+--+ +----------------- Ligumia_recta
| |
| | +-- Ptychobranchus_fasciolaris
+--+ +-----------+
| | +-- Sagittunio_nasutus
| |
+--+ +----------- Lampsilis_siliquoidea
| |
| | +----- Ortmanniana_ligamentina
+--+ +--+
| | | +-- Lampsilis_cardium
| | +--+
| | +-- Lampsilis_fasciola
+--+
| +-- Epioblasma_brevidens
| +--+
| | +-- Epioblasma_triquetra
+--+
| +-- Cambarunio_iris
+--+
+-- Leaunio_vanuxemensis
Bogan & Hoeh, 2000 [Publ 2650], fig. 1
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 1 |
1 |
0 |
1 |
33 |
10 |
3 |
-- |
+-------- Toxolasma_lividum
|
+-------- Amblema_plicata
|
| +-- Glebula_rotundata
+-----+
| +-- Cyrtonaias_tampicoensis
|
--+ +----- Potamilus_alatus
+--+
| | +-- Ortmanniana_ligamentina
| +--+
| +-- Ligumia_recta
|
| +----- Quadrula_quadrula **
| |
+--+ +-- Eurynia_dilatata
| |
+--+-- Pleurobema_clava
|
+-- Fusconaia_flava
Hoeh et al., 2001 [Publ 2652], fig. 14.2
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 1 |
1 |
0 |
1 |
30 |
10 |
3 |
-- |
-- Quadrula_quadrula
Hoeh et al., 2001 [Publ 2652], fig. 14.4
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 1 |
1 |
0 |
1 |
30 |
10 |
3 |
-- |
+-------- Quadrula_quadrula **
+-----+
| | +----- Eurynia_dilatata
| +--+
| | +-- Fusconaia_flava
| +--+
--+ +-- Pleurobema_clava
|
| +-- Cyrtonaias_tampicoensis
| +--------+
| | +-- Glebula_rotundata
+--+
| +-------- Toxolasma_lividum
+--+
| +----- Potamilus_alatus
+--+
| +-- Ligumia_recta
+--+
+-- Ortmanniana_ligamentina
Hoeh et al., 2002 [Publ 2653], fig. 2
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 1 |
1 |
0 |
1 |
30 |
10 |
3 |
-- |
+----------- Amblema_plicata
--+
| +-------- Quadrula_quadrula **
+--+
| +----- Eurynia_dilatata
+--+
| +-- Pleurobema_clava
+--+
+-- Fusconaia_flava
Roe & Hoeh, 2003 [Publ 2655], fig. 4.2
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 1 |
1 |
0 |
1 |
30 |
10 |
3 |
-- |
-- Quadrula_quadrula
Roe & Hoeh, 2003 [Publ 2655], fig. 4.3
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 1 |
1 |
0 |
1 |
30 |
10 |
3 |
-- |
+-------- Quadrula_quadrula **
+-----+
| | +----- Eurynia_dilatata
| +--+
| | +-- Fusconaia_flava
| +--+
--+ +-- Pleurobema_clava
|
| +-- Cyrtonaias_tampicoensis
| +--------+
| | +-- Glebula_rotundata
+--+
| +-------- Toxolasma_lividum
+--+
| +----- Ortmanniana_ligamentina
+--+
| +-- Ligumia_recta
+--+
+-- Potamilus_alatus
Campbell et al., 2008 [Publ 2675], fig. 4
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 1 |
1 |
0 |
1 |
28 |
28 |
3 |
-- |
+----------------------- Uniomerus_declivis **
|
+----------------------- Plectomerus_dombeyanus
|
| +-------------------- Cyrtonaias_tampicoensis
| |
| +-------------------- Eurynia_dilatata
| |
| +-------------------- Fusconaia_cerina
| |
--+ +-------------------- Fusconaia_flava
| |
| +-------------------- Hemistena_lata
| |
| +-------------------- Plethobasus_cyphyus
| |
| +-------------------- Parvaspina_collina
| |
| +-------------------- Pleurobema_cordatum
| |
| +-------------------- Pleuronaia_gibber
+--+
+-------------------- Pleurobema_oviforme[2]
|
+-------------------- Pleurobema_stabile
|
| +----- Elliptio_arca
+--------------+
| | +-- Elliptio_crassidens
| +--+
| +-- Elliptio_arctata
|
| +----------------- Pleurobema_decisum
| |
| +----------------- Pleurobema_clava
| |
| +----------------- Pleurobema_oviforme[1]
| |
+--+ +-- Pleurobema_rubrum
+--------------+
| +-- Pleurobema_sintoxia
|
| +-------------- Pleurobema_beadleianum
| |
+--+ +----------- Pleurobema_pyriforme
| |
+--+ +-------- Pleurobema_rubellum
| |
| +-------- Pleurobema_hanleyianum[2]
+--+
+-------- Pleurobema_georgianum[2]
|
| +----- Pleurobema_hanleyianum[1]
+--+
| +-- Pleurobema_georgianum[1]
+--+
+-- Pleurobema_athearni
Campbell et al., 2008 [Publ 2675], fig. 5
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 1 |
1 |
0 |
1 |
29 |
29 |
3 |
-- |
+-------------- Uniomerus_declivis **
|
+-------------- Plectomerus_dombeyanus
|
| +-- Elliptio_arctata
| +--------+
| | +-- Elliptio_crassidens
| |
| | +-------- Pleurobema_decisum
--+ | |
| | +-------- Pleurobema_rubellum
| | |
| | +-------- Pleurobema_georgianum[2]
| | |
| | +-------- Pleurobema_stabile
| | |
| | +-------- Pleurobema_pyriforme
| | |
+--+ +-------- Pleurobema_oviforme[2]
| |
| +-------- Pleurobema_hanleyianum[2]
| |
| +-------- Pleuronaia_gibber
| |
| +-------- Pleurobema_georgianum[3]
| |
| +-------- Pleurobema_cordatum
| |
| +-------- Pleurobema_beadleianum
| |
| +-------- Hemistena_lata
+--+
+-------- Fusconaia_flava
|
+-------- Fusconaia_cerina
|
+-------- Eurynia_dilatata
|
+-------- Elliptio_arca
|
+-------- Cyrtonaias_tampicoensis
|
| +-- Plethobasus_cyphyus
+-----+
| +-- Parvaspina_collina
|
| +-- Pleurobema_athearni
| |
+-----+-- Pleurobema_georgianum[1]
| |
| +-- Pleurobema_hanleyianum[1]
|
| +----- Pleurobema_oviforme[1]
| |
+--+----- Pleurobema_clava
|
| +-- Pleurobema_rubrum
+--+
+-- Pleurobema_sintoxia
Krebs, 2004 [Publ 2692], fig. 2
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 1 |
1 |
0 |
1 |
4 |
2 |
2 |
-- |
-- Quadrula_quadrula
Turner et al., 2000 [Publ 2722], fig. 2
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 1 |
1 |
0 |
1 |
15 |
13 |
3 |
-- |
-- Quadrula_quadrula
Whelan et al., 2011 [Publ 3380], fig. 1 ML
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 1 |
1 |
0 |
1 |
36 |
4 |
3 |
-- |
+----------- Pleurobema_sintoxia
--+
| +-------- Quadrula_quadrula **
+--+
| +----- Amblema_plicata
+--+
| +-- Truncilla_truncata
+--+
+-- Lampsilis_cardium
Whelan et al., 2011 [Publ 3380], fig. 1 MP
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 1 |
1 |
0 |
1 |
36 |
4 |
3 |
-- |
+-------- Quadrula_quadrula **
|
--+-------- Pleurobema_sintoxia
|
| +----- Amblema_plicata
+--+
| +-- Lampsilis_cardium
+--+
+-- Truncilla_truncata
Kandl et al., 2001 [Publ 3384], fig. 4
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 1 |
1 |
0 |
1 |
12 |
12 |
1 |
-- |
+-------------- Elliptio
|
+-------------- Uniomerus_carolinianus **
--+
| +----------- Pleurobema_sintoxia
| |
+--+ +-------- Pleurobema_strodeanum
| |
| | +----- Pleurobema_pyriforme[7]
+--+ |
| | +-- Pleurobema_pyriforme[1]
| | |
+--+ +-- Pleurobema_pyriforme[2]
| |
| +-- Pleurobema_pyriforme[3]
| |
| +-- Pleurobema_pyriforme[4]
+--+
+-- Pleurobema_pyriforme[5]
|
+-- Pleurobema[1]
|
+-- Pleurobema_pyriforme[6]
|
+-- Pleurobema[2]
Kandl et al., 2001 [Publ 3384], fig. 5
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 1 |
1 |
0 |
1 |
13 |
13 |
2 |
-- |
+-------------------- Amblema_plicata
|
+-------------------- Quadrula_quadrula **
--+
| +----------------- Elliptio_crassidens
| |
+--+ +-------------- Fusconaia_flava
| |
+--+ +----------- Pleurobema_sintoxia
| |
| | +-- Pleurobema_clava
+--+ +-----+
| | +-- Pleurobema_strodeanum
| |
+--+ +----- Pleurobema_pyriforme[6]
| |
| | +-- Pleurobema_pyriforme[1]
+--+ |
| +-- Pleurobema
| |
| +-- Pleurobema_pyriforme[2]
+--+
+-- Pleurobema_pyriforme[3]
|
+-- Pleurobema_pyriforme[4]
|
+-- Pleurobema_pyriforme[5]
Mulvey et al., 1997 [Publ 3390], fig. 1, 3
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 1 |
1 |
0 |
1 |
4 |
3 |
1 |
-- |
+-------- Megalonaias_nervosa **
--+
| +----- Amblema_plicata
+--+
| +-- Amblema_neislerii
+--+
+-- Amblema_elliottii
Inoue et al., 2013 [Publ 3528], fig. 3
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 1 |
1 |
0 |
1 |
16 |
14 |
3 |
-- |
+-- Eurynia_dilatata
+--------------------+
| +-- Quadrula_quadrula **
|
| +-------- Paetulunio_fabalis
--+ +-----------+
| | | +----- Amblema_plicata
| | +--+
| | | +-- Reginaia_ebenus
+--+ +--+
| +-- Reginaia_rotulata
|
| +----------------- Lampsilis_ornata
| |
+--+ +-------------- Leaunio_vanuxemensis
| |
| | +-- Villosa_villosa
+--+-----------+
| +-- Cambarunio_iris
|
| +----------- Obovaria_olivaria
+--+
| +-------- Obovaria_arkansasensis
+--+
| +----- Obovaria_retusa
+--+
| +-- Obovaria_unicolor
+--+
+-- Obovaria_subrotunda
Pfeiffer & Graf, 2013 [Publ 3531], fig. 2
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 1 |
1 |
0 |
1 |
34 |
4 |
3 |
-- |
+----------- Quadrula_quadrula **
--+
| +-------- Eurynia_dilatata
+--+
| +----- Amblema_plicata
+--+
| +-- Lampsilis_ornata
+--+
+-- Truncilla_truncata
Reis et al., 2013 [Publ 3580], fig. 4
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 1 |
1 |
0 |
1 |
13 |
1 |
1 |
-- |
+-- Amblema_plicata
--+
+-- Quadrula_nobilis **
Reis et al., 2013 [Publ 3580], fig. 5
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 1 |
1 |
0 |
1 |
15 |
1 |
1 |
-- |
+-- Amblema_plicata
--+
+-- Theliderma_metanevra **
Pfeiffer & Graf, 2015 [Publ 3620], fig. 2
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 1 |
1 |
0 |
1 |
39 |
4 |
2 |
-- |
+----------- Quadrula_quadrula **
--+
| +-------- Amblema_plicata
+--+
| +----- Potamilus_alatus
+--+
| +-- Lampsilis_cardium
+--+
+-- Truncilla_truncata
Chen et al., 2012 [Publ 3624], fig. 4
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 1 |
1 |
0 |
1 |
6 |
2 |
1 |
-- |
+----- Quadrula_quadrula **
--+
| +-- Lampsilis_ornata
+--+
+-- Venustaconcha_ellipsiformis
Huang et al., 2013 [Publ 3629], fig. 3
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 1 |
1 |
0 |
1 |
15 |
3 |
1 |
-- |
+-------- Quadrula_quadrula **
--+
| +----- Toxolasma_parvum
+--+
| +-- Lampsilis_ornata
+--+
+-- Venustaconcha_ellipsiformis
Huang et al., 2013 [Publ 3629], fig. 3-M
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 1 |
1 |
0 |
1 |
5 |
1 |
1 |
-- |
+-- Quadrula_quadrula **
--+
+-- Venustaconcha_ellipsiformis
Inoue et al., 2014 [Publ 3635], fig. 3A
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 1 |
1 |
0 |
1 |
20 |
3 |
3 |
-- |
-- Quadrula_quadrula
Xue et al., 2016 [Publ 3659], fig. 1-M
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 1 |
1 |
0 |
1 |
1 |
1 |
1 |
-- |
+-- Quadrula_quadrula **
--+
+-- Venustaconcha_ellipsiformis
Froufe et al., 2015 [Publ 3664], fig. 1
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 1 |
1 |
0 |
1 |
11 |
1 |
1 |
-- |
+-- Venustaconcha_ellipsiformis
--+
+-- Quadrula_quadrula **
Froufe et al., 2015 [Publ 3664], fig. 1-M
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 1 |
1 |
0 |
1 |
7 |
1 |
1 |
-- |
+-- Venustaconcha_ellipsiformis
--+
+-- Quadrula_quadrula **
Wei et al., 2016 [Publ 3665], fig. 0
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 1 |
1 |
0 |
1 |
5 |
1 |
1 |
-- |
-- Quadrula_quadrula
Plazzi et al., 2011 [Publ 3676], fig. 5
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 1 |
1 |
0 |
1 |
17 |
4 |
2 |
-- |
+-------- Quadrula_quadrula **
|
--+-------- Parvaspina_collina
|
| +----- Lampsilis_ornata
+--+
| +-- Epioblasma_rangiana
+--+
+-- Venustaconcha_ellipsiformis
Sheng et al., 2014 [Publ 3704], fig. 2
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 1 |
1 |
0 |
1 |
7 |
2 |
1 |
-- |
+-- Venustaconcha_ellipsiformis
--+
+-- Quadrula_quadrula **
Zhou et al., 2016 [Publ 3710], fig. 1
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 1 |
1 |
0 |
1 |
19 |
3 |
1 |
-- |
+-------- Quadrula_quadrula **
--+
| +----- Toxolasma_parvum
+--+
| +-- Venustaconcha_ellipsiformis
+--+
+-- Lampsilis_ornata
Wu et al., 2016 [Publ 3711], fig. 1
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 1 |
1 |
0 |
1 |
20 |
2 |
1 |
-- |
+----- Quadrula_quadrula **
--+
| +-- Venustaconcha_ellipsiformis
+--+
+-- Lampsilis_ornata
Soroka & Burzynski, 2018 [Publ 3762], fig. 3
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 1 |
1 |
0 |
1 |
6 |
1 |
1 |
-- |
+-- Venustaconcha_ellipsiformis
--+
+-- Quadrula_quadrula **
Soroka & Burzynski, 2018 [Publ 3762], fig. 3-M
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 1 |
1 |
0 |
1 |
6 |
1 |
1 |
-- |
+-- Venustaconcha_ellipsiformis
--+
+-- Quadrula_quadrula **
Guerra et al., 2017 [Publ 3772], fig. 2a
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 1 |
1 |
0 |
1 |
24 |
3 |
1 |
-- |
+-------- Quadrula_quadrula **
--+
| +----- Toxolasma_parvum
+--+
| +-- Venustaconcha_ellipsiformis
+--+
+-- Lampsilis_ornata
Guerra et al., 2017 [Publ 3772], fig. 2a-M
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 1 |
1 |
0 |
1 |
7 |
1 |
1 |
-- |
+-- Venustaconcha_ellipsiformis
--+
+-- Quadrula_quadrula **
Guerra et al., 2017 [Publ 3772], fig. 2b
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 1 |
1 |
0 |
1 |
24 |
3 |
1 |
-- |
+-------- Quadrula_quadrula **
--+
| +----- Toxolasma_parvum
+--+
| +-- Venustaconcha_ellipsiformis
+--+
+-- Lampsilis_ornata
Guerra et al., 2017 [Publ 3772], fig. 2b-M
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 1 |
1 |
0 |
1 |
7 |
1 |
1 |
-- |
+-- Venustaconcha_ellipsiformis
--+
+-- Quadrula_quadrula **
Burzynski et al., 2017 [Publ 3774], fig. 1 bottom
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 1 |
1 |
0 |
1 |
12 |
1 |
1 |
-- |
+-- Quadrula_quadrula **
--+
+-- Venustaconcha_ellipsiformis
Burzynski et al., 2017 [Publ 3774], fig. 1 top
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 1 |
1 |
0 |
1 |
12 |
1 |
1 |
-- |
+-- Quadrula_quadrula **
--+
+-- Venustaconcha_ellipsiformis
Fonseca et al., 2016 [Publ 3775], fig. 1
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 1 |
1 |
0 |
1 |
31 |
3 |
1 |
-- |
+-------- Quadrula_quadrula **
--+
| +----- Toxolasma_parvum
+--+
| +-- Lampsilis_ornata
+--+
+-- Venustaconcha_ellipsiformis
Fonseca et al., 2016 [Publ 3775], fig. 1-M
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 1 |
1 |
0 |
1 |
8 |
1 |
1 |
-- |
+-- Venustaconcha_ellipsiformis
--+
+-- Quadrula_quadrula **
Perkins et al., 2017 [Publ 3777], fig. 3
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 1 |
1 |
0 |
1 |
27 |
27 |
2 |
-- |
+----------------------- Plectomerus_dombeyanus
|
+----------------------- Uniomerus_declivis **
|
| +-- Parvaspina_steinstansana
| |
| +-- Parvaspina_collina[1]
--+ |
| +-----------------+-- Parvaspina_collina[2]
| | |
| | +-- Parvaspina_collina[3]
| | |
| | +-- Parvaspina_collina[4]
| |
+--+ +----------------- Elliptio_arca
| |
| +----------------- Hemistena_lata
| |
| | +-- Elliptio_arctata
| +--------------+
| | +-- Elliptio_crassidens
+--+
| +-------------- Eurynia_dilatata
| |
| | +----------- Pleurobema_oviforme
| | |
| | +----------- Pleurobema_clava
| | |
+--+ +----------- Pleurobema_pyriforme
| |
| +----------- Pleurobema_georgianum
| |
| +----------- Pleurobema_beadleianum
| |
| +----------- Pleurobema_hanleyianum
+--+
+----------- Pleurobema_decisum
|
| +----- Pleurobema_rubellum
+-----+
| | +-- Pleurobema_sintoxia
| +--+
| +-- Pleurobema_rubrum
|
| +-------- Elliptio_spinosa
| |
+--+ +----- Fusconaia_escambia
| |
| +----- Fusconaia_flava
+--+
+----- Fusconaia_cerina
|
| +-- Pleuronaia_gibber
+--+
+-- Pleuronaia_dolabelloides
Zhou et al., 2017 [Publ 3788], fig. 2
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 1 |
1 |
0 |
1 |
16 |
2 |
1 |
-- |
+----- Quadrula_quadrula **
--+
| +-- Potamilus_leptodon
+--+
+-- Lampsilis_ornata
Amaro et al., 2017 [Publ 3791], fig. 2-M
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 1 |
1 |
0 |
1 |
9 |
3 |
2 |
-- |
+-- Fusconaia_flava
+--+
| +-- Quadrula_quadrula **
--+
| +-- Lampsilis_siliquoidea
+--+
+-- Venustaconcha_ellipsiformis
Amaro et al., 2017 [Publ 3791], fig. 3A
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 1 |
1 |
0 |
1 |
11 |
2 |
1 |
-- |
+----- Quadrula_quadrula **
--+
| +-- Toxolasma_parvum
+--+
+-- Venustaconcha_ellipsiformis
Amaro et al., 2017 [Publ 3791], fig. 3A-M
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 1 |
1 |
0 |
1 |
6 |
1 |
1 |
-- |
-- Quadrula_quadrula
Amaro et al., 2017 [Publ 3791], fig. 3B
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 1 |
1 |
0 |
1 |
11 |
2 |
1 |
-- |
-- Quadrula_quadrula
Amaro et al., 2017 [Publ 3791], fig. 3B-M
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 1 |
1 |
0 |
1 |
6 |
1 |
1 |
-- |
-- Quadrula_quadrula
Lopes-Lima et al., 2017 [Publ 3792], fig. 2
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 1 |
1 |
0 |
1 |
32 |
3 |
1 |
-- |
+-------- Quadrula_quadrula **
--+
| +----- Toxolasma_parvum
+--+
| +-- Lampsilis_ornata
+--+
+-- Venustaconcha_ellipsiformis
Lopes-Lima et al., 2017 [Publ 3792], fig. 2-M
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 1 |
1 |
0 |
1 |
9 |
1 |
1 |
-- |
+-- Venustaconcha_ellipsiformis
--+
+-- Quadrula_quadrula **
Wen et al., 2017 [Publ 3795], fig. 3
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 1 |
1 |
0 |
1 |
15 |
5 |
1 |
-- |
+----------- Quadrula_quadrula **
|
--+ +-------- Toxolasma_parvum
| |
+--+ +-- Lampsilis_ornata
| +--+
| | +-- Venustaconcha_ellipsiformis
+--+
| +-- Potamilus_leptodon
+--+
+-- Potamilus_alatus
Wen et al., 2017 [Publ 3795], fig. 3-M
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 1 |
1 |
0 |
1 |
5 |
2 |
1 |
-- |
+----- Quadrula_quadrula **
--+
| +-- Potamilus_alatus
+--+
+-- Venustaconcha_ellipsiformis
Feng et al., 2016 [Publ 3800], fig. 2
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 1 |
1 |
0 |
1 |
14 |
2 |
1 |
-- |
+----- Quadrula_quadrula **
--+
| +-- Potamilus_leptodon
+--+
+-- Lampsilis_ornata
Bolotov et al., 2018 [Publ 3826], fig. 2
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 1 |
1 |
0 |
1 |
84 |
6 |
3 |
-- |
-- Quadrula_quadrula
Huang et al., 2018 [Publ 3828], fig. 6
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 1 |
1 |
0 |
1 |
42 |
5 |
1 |
-- |
+----------- Quadrula_quadrula **
|
--+ +-------- Toxolasma_parvum
| |
+--+ +-- Lampsilis_ornata
| +--+
| | +-- Venustaconcha_ellipsiformis
+--+
| +-- Potamilus_leptodon
+--+
+-- Potamilus_alatus
Huang et al., 2018 [Publ 3828], fig. 6-M
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 1 |
1 |
0 |
1 |
16 |
2 |
1 |
-- |
+----- Quadrula_quadrula **
--+
| +-- Venustaconcha_ellipsiformis
+--+
+-- Potamilus_alatus
Wu et al., 2018 [Publ 3836], fig. 2
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 1 |
1 |
0 |
1 |
52 |
8 |
3 |
-- |
-- Quadrula_quadrula
Huang et al., 2019 [Publ 3843], fig. 3-M
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 1 |
1 |
0 |
1 |
22 |
2 |
1 |
-- |
+----- Quadrula_quadrula **
--+
| +-- Venustaconcha_ellipsiformis
+--+
+-- Potamilus_alatus
Wang et al., 2018 [Publ 3846], fig. 2
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 1 |
1 |
0 |
1 |
31 |
5 |
1 |
-- |
+----------- Quadrula_quadrula **
|
--+ +-------- Toxolasma_parvum
| |
+--+ +-- Venustaconcha_ellipsiformis
| +--+
| | +-- Lampsilis_ornata
+--+
| +-- Potamilus_leptodon
+--+
+-- Potamilus_alatus
Chase et al., 2018 [Publ 3861], fig. 1
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 1 |
1 |
0 |
1 |
10 |
2 |
1 |
-- |
+----- Quadrula_quadrula **
--+
| +-- Lampsilis_powellii
+--+
+-- Lampsilis_siliquoidea
Chase et al., 2018 [Publ 3861], fig. 1-M
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 1 |
1 |
0 |
1 |
10 |
2 |
1 |
-- |
+----- Quadrula_quadrula **
--+
| +-- Lampsilis_powellii
+--+
+-- Lampsilis_siliquoidea
Burzynski & Soroka, 2018 [Publ 3862], fig. 5a
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 1 |
1 |
0 |
1 |
19 |
2 |
1 |
-- |
+----- Quadrula_quadrula **
--+
| +-- Potamilus_alatus
+--+
+-- Venustaconcha_ellipsiformis
Aunins et al., 2018 [Publ 3863], fig. 1
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 1 |
1 |
0 |
1 |
40 |
3 |
1 |
-- |
+-------- Quadrula_quadrula **
--+
| +----- Toxolasma_parvum
+--+
| +-- Lampsilis_ornata
+--+
+-- Venustaconcha_ellipsiformis
Wu et al., 2019 [Publ 3864], fig. 3
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 1 |
1 |
0 |
1 |
46 |
5 |
1 |
-- |
+----------- Quadrula_quadrula **
|
--+ +-------- Toxolasma_parvum
| |
+--+ +-- Lampsilis_ornata
| +--+
| | +-- Venustaconcha_ellipsiformis
+--+
| +-- Potamilus_leptodon
+--+
+-- Potamilus_alatus
Wang et al., 2018 [Publ 3865], fig. 2
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 1 |
1 |
0 |
1 |
31 |
5 |
1 |
-- |
+----------- Quadrula_quadrula **
|
--+ +-------- Toxolasma_parvum
| |
+--+ +-- Potamilus_leptodon
| +--+
| | +-- Potamilus_alatus
+--+
| +-- Venustaconcha_ellipsiformis
+--+
+-- Lampsilis_ornata
Cha et al., 2018 [Publ 3916], fig. 1
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 1 |
1 |
0 |
1 |
38 |
4 |
1 |
-- |
+----------- Quadrula_quadrula **
--+
| +-------- Toxolasma_parvum
+--+
| +----- Lampsilis_ornata
+--+
| +-- Potamilus_alatus
+--+
+-- Potamilus_leptodon
Froufe et al., 2019 [Publ 3936], fig. 3
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 1 |
1 |
0 |
1 |
44 |
5 |
1 |
-- |
+----------- Quadrula_quadrula **
|
--+ +-------- Toxolasma_parvum
| |
+--+ +-- Lampsilis_ornata
| +--+
| | +-- Venustaconcha_ellipsiformis
+--+
| +-- Potamilus_leptodon
+--+
+-- Potamilus_alatus
Froufe et al., 2019 [Publ 3936], fig. 3-M
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 1 |
1 |
0 |
1 |
18 |
2 |
1 |
-- |
+----- Quadrula_quadrula **
--+
| +-- Potamilus_alatus
+--+
+-- Venustaconcha_ellipsiformis
Froufe et al., 2019 [Publ 3936], fig. 4
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 1 |
1 |
0 |
1 |
18 |
2 |
1 |
-- |
+----- Quadrula_quadrula **
--+
| +-- Potamilus_alatus
+--+
+-- Venustaconcha_ellipsiformis
Froufe et al., 2019 [Publ 3936], fig. 6
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 1 |
1 |
0 |
1 |
44 |
5 |
1 |
-- |
+----------- Quadrula_quadrula **
|
--+ +-------- Toxolasma_parvum
| |
+--+ +-- Potamilus_leptodon
| +--+
| | +-- Potamilus_alatus
+--+
| +-- Lampsilis_ornata
+--+
+-- Venustaconcha_ellipsiformis
Smith et al., 2019 [Publ 3951], fig. 2
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 1 |
1 |
0 |
1 |
41 |
41 |
3 |
-- |
+----------------------------------------------- Quadrula_quadrula **
|
| +-- Amblema_plicata
--+ +-----------------------------------------+
| | +-- Reginaia_ebenus
| |
+--+ +----------------------------------------- Plectomerus_dombeyanus
| |
| | +-------------------------------------- Toxolasma_lividum
+--+ |
| | +----- Popenaias_popeii
| | +-----------------------------+
+--+ | | +-- Glebula_rotundata
| | +--+
| | +-- Cyrtonaias_tampicoensis
| |
| | +-------------------------------- Obliquaria_reflexa
+--+ |
| | +-- Medionidus_conradicus
| | +--+
| | | +-- Lemiox_rimosus
| | +-----------------------+
| | | | +-- Cyprogenia_stegaria
| | | +--+
+--+ | +-- Ptychobranchus_fasciolaris
| |
| | +-- Hamiota_subangulata
| | +--------------------+
| | | +-- Villosa_villosa
| | |
| | +--+ +-------------------- Ligumia_recta
| | | | |
+--+ | | | +----- Venustaconcha_ellipsiformis
| | +--+ +-----------+
| | | | | +-- Obovaria_subrotunda
| | | | +--+
| | +--+ +-- Epioblasma_rangiana
| | |
| | | +-------------- Ortmanniana_pectorosa
| | +--+
| | | +----------- Ortmanniana_ligamentina
| | +--+
| | | +-------- Lampsilis_ovata
| | +--+
+--+ | +----- Lampsilis_dolabraeformis
| +--+
| | +-- Lampsilis_satura
| +--+
| +-- Lampsilis_cardium
|
| +-------------- Atlanticoncha_ochracea
| +-----+
| | | +----------- Ellipsaria_lineolata
| | +--+
| | | +-------- Truncilla_truncata
| | +--+
| | | +----- Truncilla_cognata
| | +--+
+-----+ | +-- Truncilla_macrodon
| +--+
| +-- Truncilla_donaciformis
|
| +----------------- Potamilus_capax
| |
| | +----- Potamilus_inflatus
+--+ +--------+
| | | +-- Potamilus_fragilis
| | +--+
| | +-- Potamilus_leptodon
+--+
| +----- Potamilus_ohiensis
| +-----+
| | | +-- Potamilus_amphichaenus
| | +--+
+--+ +-- Potamilus_streckersoni
|
| +-------- Potamilus_metnecktayi
+--+
| +----- Potamilus_purpuratus[1]
+--+
| +-- Potamilus_purpuratus[2]
+--+
+-- Potamilus_alatus
Lopes-Lima et al., 2020 [Publ 4080], fig. 2-4
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 1 |
1 |
0 |
1 |
121 |
3 |
3 |
-- |
+-- Amblema_plicata
|
+-- Elliptio_complanata
--+
+-- Lampsilis_cardium
|
+-- Quadrula_quadrula **
Bolotov et al., 2020 [Publ 4084], fig. 1
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 1 |
1 |
0 |
1 |
35 |
6 |
3 |
-- |
+-- Amblema_plicata
|
+-- Ortmanniana_ligamentina
|
+-- Lampsilis_cardium
|
--+-- Cambarunio_iris
|
+-- Elliptio_complanata
|
+-- Pleurobema_sintoxia
|
+-- Quadrula_quadrula **
Wu et al., 2020 [Publ 4121], fig. 3
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 1 |
1 |
0 |
1 |
49 |
5 |
1 |
-- |
+----------- Quadrula_quadrula **
|
--+ +-------- Toxolasma_parvum
| |
+--+ +-- Venustaconcha_ellipsiformis
| +--+
| | +-- Lampsilis_ornata
+--+
| +-- Potamilus_alatus
+--+
+-- Potamilus_leptodon
Inoue et al., 2020 [Publ 4128], fig. 2
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 1 |
1 |
0 |
1 |
28 |
28 |
3 |
-- |
+-------------------------- Quadrula_quadrula **
|
| +-- Reginaia_ebenus
| +--------------------+
| | +-- Reginaia_rotulata
--+ |
| | +----- Amblema_plicata
| | +--------------+
| | | | +-- Amblema_elliottii
| | | +--+
+--+ | +-- Amblema_neislerii
| |
| | +----- Popenaias_popeii
| | +--+
| | | | +-- Popenaias_berezai
| | | +--+
| | | +-- Popenaias_semirasa
+--+ |
| +--------+ +-- Psoronaias_profunda
| | | |
| | | +-- Psoronaias_semigranosa[1]
| | | |
| | | +-- Psoronaias_semigranosa[2]
| | | |
| | +-----+-- Psoronaias_guatemalensis[1]
| | |
| | +-- Psoronaias_guatemalensis[2]
+--+ |
| +-- Psoronaias_percompressa
| |
| +-- Psoronaias_semigranosa[3]
|
| +-- Glebula_rotundata
| +-----------+
| | +-- Cyrtonaias_tampicoensis
| |
+--+ +----------- Potamilus_alatus
| |
| | +-- Ptychobranchus_fasciolaris
+--+ +-----+
| | +-- Truncilla_truncata
| |
| | +----- Ortmanniana_ligamentina
+--+ |
| +----- Epioblasma_triquetra
| |
| +----- Lampsilis_cardium
| |
+--+----- Ligumia_recta
|
+----- Cambarunio_iris
|
+----- Pachynaias_spheniopsis
|
| +-- Cyrtonaias_explicata
+--+
+-- Sphenonaias_microdon
Inoue et al., 2020 [Publ 4128], fig. 3
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 1 |
1 |
0 |
1 |
28 |
28 |
3 |
-- |
+----------------------------------------- Quadrula_quadrula **
|
| +-- Reginaia_ebenus
| +-----------------------------------+
--+ | +-- Reginaia_rotulata
| |
| | +----- Amblema_neislerii
| | +-----------------------------+
| | | | +-- Amblema_plicata
+--+ | +--+
| | +-- Amblema_elliottii
| |
| | +----- Popenaias_popeii
| | +--------+
| | | | +-- Popenaias_berezai
| | | +--+
+--+ | +-- Popenaias_semirasa
| +-----------------+
| | | +----------- Psoronaias_profunda
| | | |
| | | | +----- Psoronaias_guatemalensis[1]
| | +--+ +--+
| | | | | +-- Psoronaias_semigranosa[1]
| | | | +--+
| | +--+ +-- Psoronaias_percompressa
+--+ |
| | +----- Psoronaias_semigranosa[3]
| +--+
| | +-- Psoronaias_guatemalensis[2]
| +--+
| +-- Psoronaias_semigranosa[2]
|
| +----------------------------- Cyrtonaias_tampicoensis
| |
+--+ +-------------------------- Ptychobranchus_fasciolaris
| |
+--+ +----------------------- Ligumia_recta
| |
+--+ +-------------------- Potamilus_alatus
| |
+--+ +----------------- Pachynaias_spheniopsis
| |
| | +-- Epioblasma_triquetra
+--+ +-----------+
| | +-- Truncilla_truncata
| |
+--+ +-- Ortmanniana_ligamentina
| +--------+
| | +-- Lampsilis_cardium
+--+
| +-------- Cambarunio_iris
+--+
| +----- Sphenonaias_microdon[2]
+--+
| +-- Cyrtonaias_explicata
+--+
+-- Sphenonaias_microdon[1]
Smith et al., 2020 [Publ 4129], fig. 1
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 1 |
1 |
0 |
1 |
27 |
27 |
4 |
-- |
+-------------------------------------- Quadrula_quadrula **
|
| +-- Pleurobema_strodeanum
--+ +--------------------------------+
| | +-- Elliptoideus_sloatianus
| |
| | +----- Reginaia_ebenus
+--+ +--------------------------+
| | | +-- Amblema_plicata
| | +--+
| | +-- Plectomerus_dombeyanus
+--+
| +-- Popenaias_popeii
| +--------------------------+
| | +-- Psoronaias_semigranosa
| |
+--+ +-------------------------- Obliquaria_reflexa
| |
| | +----------------------- Cyrtonaias_tampicoensis
+--+ |
| | +-- Pachynaias_spheniopsis
| | +-----------------+
+--+ | +-- Lampsilis_cardium
| |
| | +----------- Ellipsaria_lineolata
| | +-----+
+--+ | | +-------- Truncilla_truncata
| | +--+
| | | +----- Truncilla_donaciformis
| | +--+
| | | +-- Truncilla_cognata
| | +--+
+--+ +-- Truncilla_macrodon
|
| +-------------- Atlanticoncha_ochracea
| |
| | +-- Potamilus_fragilis
| | +-----+
| | | +-- Potamilus_leptodon
+--+ +--+
| | | +----- Potamilus_metnecktayi
| | +--+
| | | +-- Potamilus_alatus
| | +--+
+--+ +-- Potamilus_purpuratus
|
| +-- Potamilus_capax
| +-----+
| | +-- Potamilus_inflatus
+--+
| +----- Potamilus_ohiensis
+--+
| +-- Potamilus_streckersoni
+--+
+-- Potamilus_amphichaenus
Smith et al., 2020 [Publ 4129], fig. 2-3
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 1 |
1 |
0 |
1 |
27 |
27 |
4 |
-- |
+----------------------------------------- Quadrula_quadrula **
|
| +-- Pleurobema_strodeanum
--+ +-----------------------------------+
| | +-- Elliptoideus_sloatianus
| |
| | +----- Reginaia_ebenus
+--+ +-----------------------------+
| | | +-- Amblema_plicata
| | +--+
| | +-- Plectomerus_dombeyanus
+--+
| +-- Popenaias_popeii
| +-----------------------------+
| | +-- Psoronaias_semigranosa
| |
+--+ +----------------------------- Obliquaria_reflexa
| |
| | +-------------------------- Cyrtonaias_tampicoensis
+--+ |
| | +-- Pachynaias_spheniopsis
| | +--------------------+
+--+ | +-- Lampsilis_cardium
| |
| | +----------- Ellipsaria_lineolata
| | +--------+
+--+ | | +-------- Truncilla_truncata
| | +--+
| | | +----- Truncilla_donaciformis
| | +--+
| | | +-- Truncilla_cognata
+--+ +--+
| +-- Truncilla_macrodon
|
| +----------------- Atlanticoncha_ochracea
| |
| | +-- Potamilus_capax
+--+ +-----------+
| | +-- Potamilus_inflatus
| |
| | +----- Potamilus_ohiensis
+--+ +-----+
| | | +-- Potamilus_streckersoni
| | +--+
| | +-- Potamilus_amphichaenus
+--+
| +-- Potamilus_fragilis
| +-----+
| | +-- Potamilus_leptodon
+--+
| +----- Potamilus_metnecktayi
+--+
| +-- Potamilus_alatus
+--+
+-- Potamilus_purpuratus
Smith et al., 2020 [Publ 4136], fig. 1
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 1 |
1 |
0 |
1 |
19 |
19 |
1 |
-- |
+-------------------------- Quadrula_quadrula **
|
--+ +----------------------- Elliptoideus_sloatianus
| |
| | +-------------------- Elliptio_crassidens
+--+ |
| +-------------------- Plethobasus_cyphyus
| |
+--+ +----------------- Parvaspina_steinstansana
| |
| | +-- Hemistena_lata
| +--------------+
| | +-- Eurynia_dilatata
+--+
| +-- Pleuronaia_barnesiana
| +-----------+
| | +-- Pleurobema_clava
| |
| | +----------- Fusconaia_subrotunda
+--+ |
| | +-- Fusconaia_cor
| +--------+
| | +-- Fusconaia_cuneolus
+--+
| +-------- Fusconaia_flava
| |
| +-------- Fusconaia_masoni
| |
| | +-- Fusconaia_burkei
+--+-----+
| +-- Fusconaia_escambia
|
| +-- Fusconaia_iheringi
| +--+
| | +-- Fusconaia_mitchelli
+--+
| +-- Fusconaia_askewi
+--+
+-- Fusconaia_chunii
Wu et al., 2018 [Publ 4143], fig. 2
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 1 |
1 |
0 |
1 |
21 |
2 |
1 |
-- |
+----- Quadrula_quadrula **
--+
| +-- Venustaconcha_ellipsiformis
+--+
+-- Lampsilis_ornata
Wu et al., 2020 [Publ 4147], fig. 3
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 1 |
1 |
0 |
1 |
29 |
2 |
1 |
-- |
+----- Quadrula_quadrula **
--+
| +-- Toxolasma_parvum
+--+
+-- Venustaconcha_ellipsiformis
Guerra et al., 2019 [Publ 4168], fig. 1
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 1 |
1 |
0 |
1 |
40 |
3 |
1 |
-- |
+-------- Quadrula_quadrula **
--+
| +----- Toxolasma_parvum
+--+
| +-- Venustaconcha_ellipsiformis
+--+
+-- Potamilus_alatus
Guerra et al., 2019 [Publ 4168], fig. 1-M
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 1 |
1 |
0 |
1 |
21 |
2 |
1 |
-- |
+----- Quadrula_quadrula **
--+
| +-- Venustaconcha_ellipsiformis
+--+
+-- Potamilus_alatus
Guerra et al., 2019 [Publ 4168], fig. 2
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 1 |
1 |
0 |
1 |
40 |
3 |
1 |
-- |
+-------- Quadrula_quadrula **
--+
| +----- Toxolasma_parvum
+--+
| +-- Venustaconcha_ellipsiformis
+--+
+-- Potamilus_alatus
Guerra et al., 2019 [Publ 4168], fig. 2-M
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 1 |
1 |
0 |
1 |
21 |
2 |
1 |
-- |
+----- Quadrula_quadrula **
--+
| +-- Venustaconcha_ellipsiformis
+--+
+-- Potamilus_alatus
Chen et al., 2020 [Publ 4170], fig. 1
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 1 |
1 |
0 |
1 |
33 |
6 |
1 |
-- |
+-------------- Quadrula_quadrula **
--+
| +----------- Toxolasma_parvum
+--+
| +-------- Lampsilis_ornata
+--+
| +----- Potamilus_leptodon
+--+
| +-- Lampsilis_siliquoidea
+--+
+-- Lampsilis_powellii
Zieritz et al., 2021 [Publ 4231], fig. 3-F, 5
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 1 |
1 |
0 |
1 |
48 |
5 |
1 |
-- |
+----------- Quadrula_quadrula **
|
--+ +-------- Toxolasma_parvum
| |
+--+ +-- Potamilus_alatus
| +--+
| | +-- Potamilus_leptodon
+--+
| +-- Venustaconcha_ellipsiformis
+--+
+-- Lampsilis_ornata
Zieritz et al., 2021 [Publ 4231], fig. 3-M, 4
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 1 |
1 |
0 |
1 |
22 |
2 |
1 |
-- |
+----- Quadrula_quadrula **
--+
| +-- Venustaconcha_ellipsiformis
+--+
+-- Potamilus_alatus
Pavan-Kumar et al., 2022 [Publ 4263], fig. 4
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 1 |
1 |
0 |
1 |
35 |
6 |
1 |
-- |
+-------------- Quadrula_quadrula **
|
--+ +----------- Toxolasma_parvum
| |
+--+ +-------- Potamilus_leptodon
| |
+--+ +-- Lampsilis_cardium
| +--+
| | +-- Lampsilis_ornata
+--+
| +-- Lampsilis_siliquoidea
+--+
+-- Venustaconcha_ellipsiformis
Pavan-Kumar et al., 2022 [Publ 4263], fig. S5
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 1 |
1 |
0 |
1 |
38 |
6 |
1 |
-- |
+-------------- Quadrula_quadrula **
|
--+ +----------- Toxolasma_parvum
| |
+--+ +-------- Potamilus_leptodon
| |
+--+ +-- Lampsilis_cardium
| +--+
| | +-- Lampsilis_ornata
+--+
| +-- Lampsilis_siliquoidea
+--+
+-- Venustaconcha_ellipsiformis
Mejia-Trujillo & Smith, 2022 [Publ 4274], fig. 2-M
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 1 |
1 |
0 |
1 |
5 |
5 |
1 |
-- |
+----------- Quadrula_quadrula **
|
--+ +-- Potamilus_streckersoni
| +-----+
| | +-- Potamilus_alatus
+--+
| +----- Venustaconcha_ellipsiformis
+--+
| +-- Lampsilis_powellii
+--+
+-- Lampsilis_siliquoidea
Chen et al., 2023 [Publ 4302], fig. 1
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 1 |
1 |
0 |
1 |
39 |
1 |
1 |
-- |
+-- Lampsilis_siliquoidea
--+
+-- Quadrula_nobilis **
Chen et al., 2023 [Publ 4302], fig. 2
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 1 |
1 |
0 |
1 |
39 |
1 |
1 |
-- |
+-- Lampsilis_siliquoidea
--+
+-- Quadrula_nobilis **
| 
