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Page last updated
Mon 06 May 2024

The Freshwater Mussels (Unionoida) of the World (and other less consequential bivalves)

FM(U)otW(aolcb) is the web version of the MUSSEL Project Database. Follow the links to browse the data or use the custom Google search field. Either way, you win!

Cladomics pages are experimental — even more so than the rest of this site!

The purpose of the cladomics pages for each genus and family-group level taxon is to provide statistics and references indicating the extent to which published phylogenetic work supports the classification used here.

The Cladomics Summary provides an overview of the number of studies that have included terminal species representing this taxon as well as an indication of how frequently it has been recovered as monophyletic.

Following the summary, the available published clades representing the taxon are listed.

For more information, visit the Cladomics FARQ (frequently asked rhetorical questions) page. Abbreviations for the various statistics are explained on the FARQ!

Search the database pages:

Bivalvia | Unionoida | Unionoidea | Unionidae | Unioninae

tribe UNIONINI Rafinesque, 1820: cladomics
     13 genera

Cladomics Summary

number of species = 92
number of subtaxa (rank: genus) = 13

analyzed in 221 cladograms in 130 publications
maximum number of terminals in an analysis = 26, mean = 6.12
maximum number of subtaxa (genus) in an analysis = 10, mean = 2.67

Basic Monophyly

number of cladograms that test basic monophyly (i.e., more than 1 included species) = 183
maximum number of included terminals = 26, mean = 7.18
maximum number of subtaxa (genus) = 10, mean = 3.02
proportion of cladograms that support basic monophyly = 0.85

Rigorous Monophyly

number of cladograms that test rigorous monophyly = 99
(Rigorous monophyly refers to cladograms with terminal species representing more than one subtaxon (genus) taxon and at least one rigorous outgroup (tribe).
maximum number of included terminals = 26, mean = 9.14
maximum number of subtaxa (genus) = 10, mean = 4.52
proportion of cladograms that support rigorous monophyly = 0.76


Pairwise Cladogram Clustering

The graphs below depict in two dimensions all pair-wise ingroup distances within the cladome. See the Cladmoics FARQ for more information.


PCoA Ordination


NJ tree


Chen et al., 2023 [Publ 4302], fig. 1

iT tT bi riS oT roT roS iD
17 17 14 10 23 14 4 0.0

*** Top Monophyly Cladogram ***

                       +----- Pletholophus_tenuis         
  +--------------------+                                  
  |                    |  +-- Sinanodonta_jourdyi         
  |                    +--+                               
  |                       +-- Beringiana_beringiana       
  |                                                       
  |  +----------------------- Lanceolaria_gladiola        
  |  |                                                    
  |  +----------------------- Acuticosta_chinensis        
--+  |                                                    
  |  +----------------------- Lepidodesma_languilati      
  |  |                                                    
  |  +----------------------- Lanceolaria_oxyrhyncha      
  |  |                                                    
  |  +----------------------- Cristaria_plicata           
  |  |                                                    
  |  |                    +-- Alasmidonta_marginata       
  +--+                 +--+                               
     |                 |  +-- Lasmigona_costata           
     +-----------------+                                  
     |                 |  +-- Anodonta_anatina            
     |                 +--+                               
     |                    +-- Pseudanodonta_complanata    
     |                                                    
     |  +-------------------- Aculamprotula_fibrosa **       
     |  |                                                 
     |  |              +----- Middendorffinaia_mongolica **  
     |  |  +-----------+                                  
     +--+  |           |  +-- Nodularia_douglasiae **        
        |  |           +--+                               
        |  |              +-- Nodularia_breviconcha **       
        |  |                                              
        |  |              +-- Cuneopsis_celtiformis **       
        +--+  +-----------+                               
           |  |           +-- Cuneopsis_heudei **            
           |  |                                           
           |  |           +-- Diaurora_aurorea **            
           |  |        +--+                               
           |  |        |  +-- Diaurora_laeve **              
           |  +--------+                                  
           +--+        |  +-- Schistodesmus_lampreyanus **   
              |        +--+                               
              |           +-- Schistodesmus_spinosus **      
              |                                           
              |           +-- Pseudocuneopsis_sichuanensis **
              |  +--------+                               
              |  |        +-- Pseudocuneopsis_capitata **    
              |  |                                        
              +--+        +-- Inversiunio_yanagawensis **    
                 |  +-----+                               
                 |  |     +-- Inversiunio_reinianus **       
                 +--+                                     
                    |  +----- Tchangsinaia_piscicula **      
                    +--+                                  
                       |  +-- Unio_pictorum **               
                       +--+                               
                          +-- Unio_crassus **                

Huang et al., 2019 [Publ 3843], fig. S1

iT tT bi riS oT roT roS iD
14 14 12 5 68 48 4 18.0

*** Top Sister-Group Cladogram ***

                       +-- Lepidodesma_languilati     
  +--------------------+                              
  |                    +-- Cristaria_plicata          
  |                                                   
  |           +----------- Lanceolaria_gladiola       
  |           |                                       
  |           |  +-------- Lanceolaria_grayii[5]      
  |  +--------+  |                                    
  |  |        |  |     +-- Lanceolaria_lanceolata     
  |  |        |  +-----+                              
  |  |        |  |     +-- Lanceolaria_triformis      
--+  |        +--+                                    
  |  |           |  +----- Lanceolaria_grayii[4]      
  |  |           |  |                                 
  |  |           |  |  +-- Lanceolaria_eucylindrica[1]
  |  |           |  |  |                              
  |  |           +--+  +-- Lanceolaria_eucylindrica[2]
  |  |              |  |                              
  |  |              |  +-- Lanceolaria_eucylindrica[3]
  |  |              |  |                              
  |  |              +--+-- Lanceolaria_eucylindrica[4]
  +--+                 |                              
     |                 +-- Lanceolaria_grayii[1]      
     |                 |                              
     |                 +-- Lanceolaria_grayii[2]      
     |                 |                              
     |                 +-- Lanceolaria_grayii[3]      
     |                                                
     |        +----------- Aculamprotula_tortuosa **     
     |  +-----+                                       
     |  |     |  +-------- Aculamprotula_tientsinensis **
     |  |     +--+                                    
     |  |        |  +----- Aculamprotula_fibrosa **      
     |  |        +--+                                 
     |  |           |  +-- Aculamprotula_coreana **      
     +--+           +--+                              
        |              +-- Aculamprotula_scripta **      
        |                                             
        |           +----- Nodularia_douglasiae[1] **    
        |  +--------+                                 
        |  |        |  +-- Nodularia_nuxpersicae **      
        |  |        +--+                              
        +--+           +-- Nodularia_douglasiae[2] **    
           |                                          
           |  +----------- Tchangsinaia_piscicula **     
           |  |                                       
           +--+        +-- Schistodesmus_spinosus **     
              |  +-----+                              
              |  |     +-- Schistodesmus_lampreyanus **  
              +--+                                    
                 |  +----- Cuneopsis_rufescens **        
                 +--+                                 
                    |  +-- Cuneopsis_heudei **           
                    +--+                              
                       +-- Cuneopsis_celtiformis **      

Lopes-Lima et al., 2020 [Publ 4080], fig. 2-4

iT tT bi riS oT roT roS iD
26 26 22 8 96 43 4 25.0
  +-------------------------------- Acuticosta_chinensis       
  |                                                            
  +-------------------------------- Lepidodesma_languilati     
  |                                                            
  |                    +----------- Aculamprotula_tortuosa **     
  |     +--------------+                                       
  |     |              |  +-------- Aculamprotula_scripta **      
  |     |              +--+                                    
  |     |                 |  +----- Aculamprotula_coreana **      
  |     |                 +--+                                 
  |     |                    |  +-- Aculamprotula_fibrosa **      
  |     |                    +--+                              
  +-----+                       +-- Aculamprotula_tientsinensis **
  |     |                                                      
  |     |  +----------------------- Middendorffinaia_mongolica ** 
  |     |  |                                                   
  |     |  |                 +----- Unio_gibbus **                
  |     |  |                 |                                 
  |     |  |  +--------------+----- Unio_tumidus **               
  |     +--+  |              |                                 
  |        |  |              |  +-- Unio_crassus **               
  |        |  |              +--+                              
  |        |  |                 +-- Unio_pictorum **              
  |        |  |                                                
  |        |  |     +-------------- Nodularia_breviconcha **      
  |        +--+  +--+                                          
  |           |  |  |  +----------- Nodularia[2] **               
  |           |  |  +--+                                       
  |           |  |     |  +-------- Nodularia_nuxpersicae **      
  |           |  |     +--+                                    
  |           |  |        |  +----- Nodularia_nipponensis **      
  |           |  |        +--+                                 
  |           |  |           |  +-- Nodularia_douglasiae **       
  |           +--+           +--+                              
  |              |              +-- Nodularia[1] **               
  |              |                                             
--+              |              +-- Schistodesmus_lampreyanus **  
  |              |  +-----------+                              
  |              |  |           +-- Schistodesmus_spinosus **     
  |              |  |                                          
  |              |  |        +----- Cuneopsis_rufescens **        
  |              |  +--------+                                 
  |              +--+        |  +-- Cuneopsis_heudei **           
  |                 |        +--+                              
  |                 |           +-- Cuneopsis_celtiformis **      
  |                 |                                          
  |                 |  +----------- Tchangsinaia_piscicula **     
  |                 +--+                                       
  |                    |  +-------- Inversiunio_jokohamensis **   
  |                    +--+                                    
  |                       |  +----- Inversiunio_verrucosus **     
  |                       +--+                                 
  |                          |  +-- Inversiunio_reinianus **      
  |                          +--+                              
  |                             +-- Inversiunio_yanagawensis **   
  |                                                            
  |              +----------------- Lanceolaria_oxyrhyncha     
  |              |                                             
  |  +-----------+  +-------------- Lanceolaria_gladiola       
  |  |           |  |                                          
  |  |           +--+           +-- Lanceolaria_bilirata       
  |  |              |  +--------+                              
  |  |              |  |        +-- Lanceolaria_grayii         
  |  |              +--+                                       
  |  |                 |  +-------- Lanceolaria_acrorrhyncha   
  |  |                 +--+                                    
  |  |                    |  +----- Lanceolaria_kihirai        
  |  |                    +--+                                 
  |  |                       |  +-- Lanceolaria_lanceolata     
  |  |                       +--+                              
  |  |                          +-- Lanceolaria_triformis      
  |  |                                                         
  |  |                    +-------- Pseudanodonta_complanata   
  +--+                    |                                    
     |                    |  +----- Anodonta_cygnea            
     |                    +--+                                 
     |                    |  |  +-- Anodonta_anatina           
     |  +-----------------+  +--+                              
     |  |                 |     +-- Anodonta_nuttalliana       
     |  |                 |                                    
     |  |                 |  +----- Pyganodon_grandis          
     |  |                 |  |                                 
     |  |                 |  +----- Simpsonaias_ambigua        
     |  |                 +--+                                 
     |  |                    +----- Strophitus_undulatus       
     |  |                    |                                 
     |  |                    |  +-- Alasmidonta_marginata      
     |  |                    +--+                              
     |  |                       +-- Lasmigona_costata          
     +--+                                                      
        |                       +-- Cristaria_plicata          
        |  +--------------------+                              
        |  |                    +-- Cristaria_clessini         
        |  |                                                   
        |  |        +-------------- Anemina_arcaeformis        
        |  |        |                                          
        |  +--------+  +----------- Amuranodonta_kijaensis     
        |  |        |  |                                       
        |  |        +--+  +-------- Buldowskia_suifunica       
        |  |           |  |                                    
        |  |           +--+     +-- Buldowskia_flavotincta     
        +--+              |  +--+                              
           |              |  |  +-- Buldowskia_shadini         
           |              +--+                                 
           |                 |  +-- Buldowskia_iwakawai        
           |                 +--+                              
           |                    +-- Buldowskia_kamiyai         
           |                                                   
           |  +-------------------- Pletholophus_tenuis        
           |  |                                                
           |  |           +-------- Beringiana_japonica        
           |  |  +--------+                                    
           +--+  |        |  +----- Beringiana_beringiana      
              |  |        +--+                                 
              |  |           |  +-- Beringiana_fukuharai       
              |  |           +--+                              
              +--+              +-- Beringiana_gosannensis     
                 |                                             
                 |              +-- Sinanodonta_angula         
                 |  +-----------+                              
                 |  |           +-- Sinanodonta_lucida         
                 |  |                                          
                 +--+  +----------- Sinanodonta_schrenkii      
                    |  |                                       
                    |  |        +-- Sinanodonta_calipygos      
                    |  +--------+                              
                    +--+        +-- Sinanodonta_tumens         
                       |                                       
                       |  +-------- Sinanodonta_woodiana[2]    
                       |  |                                    
                       +--+  +----- Sinanodonta_elliptica      
                          |  |                                 
                          +--+----- Sinanodonta_jourdyi        
                             |                                 
                             |  +-- Sinanodonta_lauta          
                             +--+                              
                                +-- Sinanodonta_woodiana[1]    

Wu et al., 2022 [Publ 4238], fig. 2d

iT tT bi riS oT roT roS iD
23 23 21 8 16 16 4 22.0
  +-------------------------------- Lepidodesma_languilati      
  |                                                             
  |                          +----- Lanceolaria_gladiola        
  |           +--------------+                                  
  |           |              |  +-- Lanceolaria_grayii          
  |           |              +--+                               
  |           |                 +-- Lanceolaria_lanceolata      
  |  +--------+                                                 
  |  |        |  +----------------- Acuticosta_chinensis        
  |  |        |  |                                              
  |  |        |  |        +-------- Cristaria_plicata           
  |  |        |  |        |                                     
--+  |        +--+  +-----+     +-- Sinanodonta_woodiana        
  |  |           |  |     |  +--+                               
  |  |           |  |     |  |  +-- Sinanodonta_lucida          
  |  |           |  |     +--+                                  
  |  |           |  |        |  +-- Anemina_arcaeformis         
  |  |           +--+        +--+                               
  |  |              |           +-- Anemina_euscaphys           
  |  |              |                                           
  |  |              |           +-- Anodonta_cygnea             
  |  |              |  +--------+                               
  |  |              |  |        +-- Anodonta_anatina            
  |  |              +--+                                        
  +--+                 |  +-------- Platynaias_compressa        
     |                 +--+                                     
     |                    |  +----- Pyganodon_grandis           
     |                    +--+                                  
     |                       |  +-- Utterbackia_imbecillis      
     |                       +--+                               
     |                          +-- Utterbackia_peninsularis    
     |                                                          
     |                    +-------- Aculamprotula_tortuosa **      
     |                    |                                     
     |  +-----------------+     +-- Aculamprotula_tientsinensis ** 
     |  |                 |  +--+                               
     |  |                 |  |  +-- Aculamprotula_coreana **       
     |  |                 +--+                                  
     |  |                    |  +-- Aculamprotula_polysticta **    
     |  |                    +--+                               
     |  |                       +-- Aculamprotula_scripta **       
     +--+                                                       
        |                 +-------- Unio_tumidus **                
        |  +--------------+                                     
        |  |              |  +----- Unio_crassus **                
        |  |              +--+                                  
        |  |                 |  +-- Unio_pictorum **               
        |  |                 +--+                               
        |  |                    +-- Unio_delphinus **              
        +--+                                                    
           |                    +-- Pseudocuneopsis_capitata **    
           |  +-----------------+                               
           |  |                 +-- Pseudocuneopsis_sichuanensis **
           |  |                                                 
           |  |                 +-- Nodularia_breviconcha **       
           +--+  +--------------+                               
              |  |              +-- Nodularia_douglasiae **        
              |  |                                              
              |  |              +-- Schistodesmus_lampreyanus **   
              +--+  +-----------+                               
                 |  |           +-- Schistodesmus_spinosus **      
                 |  |                                           
                 |  |     +-------- Cuneopsis_rufescens **         
                 +--+  +--+                                     
                    |  |  |  +----- Cuneopsis_celtiformis **       
                    |  |  +--+                                  
                    |  |     |  +-- Cuneopsis_heudei **            
                    +--+     +--+                               
                       |        +-- Cuneopsis_demangei **          
                       |                                        
                       |  +-------- Tchangsinaia_piscicula **      
                       +--+                                     
                          |  +----- Inversiunio_jokohamensis **    
                          +--+                                  
                             |  +-- Inversiunio_reinianus **       
                             +--+                               
                                +-- Inversiunio_yanagawensis **    

Lyubas et al., 2022 [Publ 4255], fig. 7

iT tT bi riS oT roT roS iD
23 23 20 1 8 5 1 36.0
                       +----------- Pseudanodonta_complanata
  +--------------------+                                    
  |                    |  +-------- Anodonta_anatina        
  |                    +--+                                 
  |                       |  +----- Anodonta_cygnea         
  |                       +--+                              
  |                          |  +-- Anodonta_exulcerata     
  |                          +--+                           
--+                             +-- Anodonta_seddoni        
  |                                                         
  |  +----------------------------- Unio_tumidus **            
  |  |                                                      
  |  |                          +-- Unio_durieui **            
  |  |  +-----------------------+                           
  |  |  |                       +-- Unio_gibbus **             
  +--+  |                                                   
     |  |           +-------------- Unio_damascensis **        
     |  |           |                                       
     |  |           +-------------- Unio_sesirmensis **        
     |  |  +--------+                                       
     +--+  |        |  +----------- Unio_ionicus **            
        |  |        |  |                                    
        |  |        +--+        +-- Unio_bruguierianus **      
        |  |           |  +-----+                           
        |  |           |  |     +-- Unio_tumidiformis **       
        |  |           +--+                                 
        |  |              |  +----- Unio_crassus[2] **         
        +--+              +--+                              
           |                 |  +-- Unio_carneus **            
           |                 +--+                           
           |                    +-- Unio_crassus[1] **         
           |                                                
           |                 +----- Unio_caffer **             
           |  +--------------+                              
           |  |              |  +-- Unio_terminalis **         
           |  |              +--+                           
           +--+                 +-- Unio_delicatus **          
              |                                             
              |  +----------------- Unio_pictorum **           
              |  |                                          
              +--+  +-------------- Unio_eucirrus **           
                 |  |                                       
                 +--+           +-- Unio_elongatulus **        
                    |  +--------+                           
                    |  |        +-- Unio_mancus **             
                    +--+                                    
                       |  +-------- Unio_ravoisieri **         
                       |  |                                 
                       +--+     +-- Unio_delphinus **          
                          |  +--+                           
                          |  |  +-- Unio_foucauldianus **      
                          +--+                              
                             |  +-- Unio_hueti **              
                             +--+                           
                                +-- Unio_tigridis **           

Lyubas et al., 2022 [Publ 4255], fig. S1

iT tT bi riS oT roT roS iD
23 23 14 1 8 5 1 36.0
           +----------- Anodonta_anatina        
  +--------+                                    
  |        |  +-------- Pseudanodonta_complanata
  |        +--+                                 
  |           |  +----- Anodonta_exulcerata     
  |           +--+                              
  |              |  +-- Anodonta_seddoni        
  |              +--+                           
  |                 +-- Anodonta_cygnea         
  |                                             
--+  +----------------- Unio_tumidus **            
  |  |                                          
  |  |              +-- Unio_durieui **            
  |  +--------------+                           
  |  |              +-- Unio_gibbus **             
  |  |                                          
  |  |     +----------- Unio_sesirmensis **        
  |  |     |                                    
  |  |  +--+  +-------- Unio_damascensis **        
  +--+  |  |  |                                 
     |  |  |  |  +----- Unio_ionicus **            
     |  |  +--+  |                              
     |  |     |  |  +-- Unio_bruguierianus **      
     |  |     |  +--+                           
     |  |     +--+  +-- Unio_tumidiformis **       
     |  |        |                              
     |  |        |  +-- Unio_carneus **            
     |  |        |  |                           
     |  |        +--+-- Unio_crassus[1] **         
     +--+           |                           
        |           +-- Unio_crassus[2] **         
        |                                       
        |     +-------- Unio_elongatulus **        
        |     |                                 
        |     +-------- Unio_mancus **             
        |     |                                 
        |     |     +-- Unio_pictorum **           
        |     +-----+                           
        |     |     +-- Unio_eucirrus **           
        |     |                                 
        +-----+  +----- Unio_caffer **             
              +--+                              
              |  |  +-- Unio_terminalis **         
              |  +--+                           
              |     +-- Unio_delicatus **          
              |                                 
              |  +----- Unio_ravoisieri **         
              |  |                              
              |  |  +-- Unio_delphinus **          
              +--+--+                           
                 |  +-- Unio_foucauldianus **      
                 |                              
                 |  +-- Unio_hueti **              
                 +--+                           
                    +-- Unio_tigridis **           

Wu et al., 2023 [Publ 4339], fig. 4, S5-nt

iT tT bi riS oT roT roS iD
23 23 20 7 74 25 4 26.0
                                +-- Lepidodesma_aligera         
  +-----------------------------+                               
  |                             +-- Lepidodesma_languilati      
  |                                                             
  |                       +-------- Acuticosta_chinensis        
  |        +--------------+                                     
  |        |              |  +----- Lanceolaria_gladiola        
  |        |              +--+                                  
  |        |                 |  +-- Lanceolaria_grayii          
  |        |                 +--+                               
  |        |                    +-- Lanceolaria_lanceolata      
  |        |                                                    
  |        |        +-------------- Cristaria_plicata           
  |  +-----+        |                                           
  |  |     |  +-----+           +-- Anemina_arcaeformis         
  |  |     |  |     |  +--------+                               
  |  |     |  |     |  |        +-- Anemina_euscaphys           
  |  |     |  |     +--+                                        
--+  |     |  |        |  +-------- Beringiana_fukuharai        
  |  |     |  |        +--+                                     
  |  |     |  |           |  +----- Sinanodonta_lucida          
  |  |     |  |           +--+                                  
  |  |     +--+              |  +-- Sinanodonta_tumens          
  |  |        |              +--+                               
  |  |        |                 +-- Sinanodonta_woodiana        
  |  |        |                                                 
  |  |        |  +----------------- Anodonta_nuttalliana        
  |  |        |  |                                              
  |  |        |  +----------------- Pseudanodonta_complanata    
  |  |        |  |                                              
  |  |        +--+  +-------------- Anodonta_cygnea             
  |  |           |  |                                           
  |  |           |  |           +-- Anodonta_exulcerata         
  |  |           |  +-----------+                               
  |  |           |  |           +-- Anodonta_anatina            
  |  |           +--+                                           
  +--+              |        +----- Pyganodon_grandis           
     |              |  +-----+                                  
     |              |  |     |  +-- Utterbackia_imbecillis      
     |              |  |     +--+                               
     |              +--+        +-- Utterbackia_peninsularis    
     |                 |                                        
     |                 |  +-------- Lasmigona_complanata        
     |                 +--+                                     
     |                    |  +----- Prolasmidonta_heterodon     
     |                    +--+                                  
     |                       |  +-- Alasmidonta_varicosa        
     |                       +--+                               
     |                          +-- Platynaias_compressa        
     |                                                          
     |                    +-------- Aculamprotula_tortuosa **      
     |                    |                                     
     |  +-----------------+     +-- Aculamprotula_coreana **       
     |  |                 |  +--+                               
     |  |                 |  |  +-- Aculamprotula_tientsinensis ** 
     |  |                 +--+                                  
     |  |                    |  +-- Aculamprotula_polysticta **    
     |  |                    +--+                               
     |  |                       +-- Aculamprotula_scripta **       
     |  |                                                       
     +--+              +----------- Unio_tumidus **                
        |              |                                        
        |  +-----------+  +-------- Unio_crassus **                
        |  |           |  |                                     
        |  |           +--+  +----- Unio_elongatulus **            
        |  |              |  |                                  
        |  |              +--+  +-- Unio_delphinus **              
        |  |                 |  |                               
        |  |                 +--+-- Unio_mancus **                 
        +--+                    |                               
           |                    +-- Unio_pictorum **               
           |                                                    
           |                    +-- Pseudocuneopsis_capitata **    
           |  +-----------------+                               
           |  |                 +-- Pseudocuneopsis_sichuanensis **
           |  |                                                 
           +--+                 +-- Nodularia_breviconcha **       
              |  +--------------+                               
              |  |              +-- Nodularia_nipponensis **       
              |  |                                              
              +--+           +----- Tchangsinaia_piscicula **      
                 |  +--------+                                  
                 |  |        |  +-- Schistodesmus_lampreyanus[2] **
                 |  |        +--+                               
                 +--+           +-- Schistodesmus_spinosus **      
                    |                                           
                    |  +----------- Schistodesmus_lampreyanus[1] **
                    +--+                                        
                       |  +-------- Cuneopsis_rufescens **         
                       +--+                                     
                          |  +----- Cuneopsis_celtiformis **       
                          +--+                                  
                             |  +-- Cuneopsis_demangei **          
                             +--+                               
                                +-- Cuneopsis_heudei **            

Wu et al., 2023 [Publ 4339], fig. 4, S5-aa

iT tT bi riS oT roT roS iD
22 22 20 7 74 25 4 25.0
                             +-- Lepidodesma_aligera         
  +--------------------------+                               
  |                          +-- Lepidodesma_languilati      
  |                                                          
  |                    +-------- Aculamprotula_tortuosa **      
  |                    |                                     
  |     +--------------+     +-- Aculamprotula_coreana **       
  |     |              |  +--+                               
  |     |              |  |  +-- Aculamprotula_tientsinensis ** 
  |     |              +--+                                  
  |     |                 |  +-- Aculamprotula_polysticta **    
  |     |                 +--+                               
  |     |                    +-- Aculamprotula_scripta **       
  |     |                                                    
  |  +--+        +-------------- Unio_tumidus **                
  |  |  |  +-----+                                           
  |  |  |  |     |  +----------- Unio_crassus **                
  |  |  |  |     +--+                                        
--+  |  |  |        |  +-------- Unio_elongatulus **            
  |  |  |  |        +--+                                     
  |  |  |  |           |  +----- Unio_pictorum **               
  |  |  |  |           +--+                                  
  |  |  |  |              |  +-- Unio_delphinus **              
  |  |  +--+              +--+                               
  |  |     |                 +-- Unio_mancus **                 
  |  |     |                                                 
  |  |     |                 +-- Nodularia_breviconcha **       
  |  |     |              +--+                               
  |  |     |              |  +-- Nodularia_nipponensis **       
  |  |     |  +-----------+                                  
  |  |     |  |           |  +-- Pseudocuneopsis_capitata **    
  |  |     |  |           +--+                               
  |  |     +--+              +-- Pseudocuneopsis_sichuanensis **
  |  |        |                                              
  |  |        |  +-------------- Tchangsinaia_piscicula **      
  |  |        |  |                                           
  +--+        +--+           +-- Schistodesmus_lampreyanus **   
     |           |  +--------+                               
     |           |  |        +-- Schistodesmus_spinosus **      
     |           +--+                                        
     |              |  +-------- Cuneopsis_rufescens **         
     |              +--+                                     
     |                 |  +----- Cuneopsis_celtiformis **       
     |                 +--+                                  
     |                    |  +-- Cuneopsis_demangei **          
     |                    +--+                               
     |                       +-- Cuneopsis_heudei **            
     |                                                       
     |                 +-------- Acuticosta_chinensis        
     |        +--------+                                     
     |        |        |  +----- Lanceolaria_gladiola        
     |        |        +--+                                  
     |        |           |  +-- Lanceolaria_grayii          
     |        |           +--+                               
     |        |              +-- Lanceolaria_lanceolata      
     |        |                                              
     |        |           +----- Cristaria_plicata           
     |        |     +-----+                                  
     |        |     |     |  +-- Anemina_arcaeformis         
     +--------+     |     +--+                               
              |  +--+        +-- Anemina_euscaphys           
              |  |  |                                        
              |  |  |  +-------- Beringiana_fukuharai        
              |  |  +--+                                     
              |  |     |  +----- Sinanodonta_lucida          
              |  |     +--+                                  
              |  |        |  +-- Sinanodonta_tumens          
              |  |        +--+                               
              |  |           +-- Sinanodonta_woodiana        
              +--+                                           
                 |           +-- Anodonta_nuttalliana        
                 |     +-----+                               
                 |     |     +-- Pseudanodonta_complanata    
                 |  +--+                                     
                 |  |  |  +----- Anodonta_anatina            
                 |  |  +--+                                  
                 |  |     |  +-- Anodonta_cygnea             
                 |  |     +--+                               
                 |  |        +-- Anodonta_exulcerata         
                 +--+                                        
                    |     +----- Pyganodon_grandis           
                    |  +--+                                  
                    |  |  |  +-- Utterbackia_peninsularis    
                    |  |  +--+                               
                    |  |     +-- Utterbackia_imbecillis      
                    +--+                                     
                       |     +-- Alasmidonta_varicosa        
                       |  +--+                               
                       |  |  +-- Platynaias_compressa        
                       +--+                                  
                          |  +-- Prolasmidonta_heterodon     
                          +--+                               
                             +-- Lasmigona_complanata        

Wu et al., 2024 [Publ 4350], fig. 8a, S1

iT tT bi riS oT roT roS iD
22 22 19 7 70 25 4 22.0
                             +-- Lepidodesma_aligera         
  +--------------------------+                               
  |                          +-- Lepidodesma_languilati      
  |                                                          
  |                    +-------- Aculamprotula_tortuosa **      
  |                    |                                     
  |     +--------------+     +-- Aculamprotula_coreana **       
  |     |              |  +--+                               
  |     |              |  |  +-- Aculamprotula_tientsinensis ** 
  |     |              +--+                                  
  |     |                 |  +-- Aculamprotula_polysticta **    
  |     |                 +--+                               
  |     |                    +-- Aculamprotula_scripta **       
  |     |                                                    
  |  +--+        +-------------- Unio_tumidus **                
  |  |  |  +-----+                                           
  |  |  |  |     |  +----------- Unio_crassus **                
  |  |  |  |     +--+                                        
--+  |  |  |        |  +-------- Unio_elongatulus **            
  |  |  |  |        +--+                                     
  |  |  |  |           |  +----- Unio_pictorum **               
  |  |  |  |           +--+                                  
  |  |  |  |              |  +-- Unio_delphinus **              
  |  |  +--+              +--+                               
  |  |     |                 +-- Unio_mancus **                 
  |  |     |                                                 
  |  |     |                 +-- Nodularia_breviconcha **       
  |  |     |  +--------------+                               
  |  |     |  |              +-- Nodularia_douglasiae **        
  |  |     |  |                                              
  |  |     |  |              +-- Pseudocuneopsis_capitata **    
  |  |     |  +--------------+                               
  |  |     +--+              +-- Pseudocuneopsis_sichuanensis **
  |  |        |                                              
  |  |        |              +-- Schistodesmus_lampreyanus **   
  |  |        |  +-----------+                               
  +--+        |  |           +-- Schistodesmus_spinosus **      
     |        +--+                                           
     |           |  +----------- Tchangsinaia_piscicula **      
     |           +--+                                        
     |              |  +-------- Cuneopsis_rufescens **         
     |              +--+                                     
     |                 |  +----- Cuneopsis_celtiformis **       
     |                 +--+                                  
     |                    |  +-- Cuneopsis_demangei **          
     |                    +--+                               
     |                       +-- Cuneopsis_heudei **            
     |                                                       
     |                 +-------- Acuticosta_chinensis        
     |  +--------------+                                     
     |  |              |  +----- Lanceolaria_gladiola        
     |  |              +--+                                  
     |  |                 |  +-- Lanceolaria_grayii          
     |  |                 +--+                               
     |  |                    +-- Lanceolaria_lanceolata      
     |  |                                                    
     |  |        +-------------- Cristaria_plicata           
     |  |        |                                           
     +--+  +-----+           +-- Anemina_arcaeformis         
        |  |     |  +--------+                               
        |  |     |  |        +-- Anemina_euscaphys           
        |  |     +--+                                        
        |  |        |  +-------- Beringiana_fukuharai        
        |  |        +--+                                     
        |  |           |  +----- Sinanodonta_lucida          
        |  |           +--+                                  
        |  |              |  +-- Sinanodonta_tumens          
        +--+              +--+                               
           |                 +-- Sinanodonta_woodiana        
           |                                                 
           |           +-------- Pseudanodonta_complanata    
           |  +--------+                                     
           |  |        |  +----- Anodonta_anatina            
           |  |        +--+                                  
           |  |           |  +-- Anodonta_cygnea             
           |  |           +--+                               
           +--+              +-- Anodonta_exulcerata         
              |                                              
              |  +-------------- Anodonta_nuttalliana        
              |  |                                           
              |  |        +----- Pyganodon_grandis           
              +--+  +-----+                                  
                 |  |     |  +-- Utterbackia_imbecillis      
                 |  |     +--+                               
                 +--+        +-- Utterbackia_peninsularis    
                    |                                        
                    |  +-------- Lasmigona_complanata        
                    +--+                                     
                       |  +----- Prolasmidonta_heterodon     
                       +--+                                  
                          |  +-- Alasmidonta_varicosa        
                          +--+                               
                             +-- Platynaias_compressa        

Wu et al., 2024 [Publ 4350], fig. 8b, S2

iT tT bi riS oT roT roS iD
22 22 19 7 70 25 4 22.0
                             +-- Lepidodesma_aligera         
  +--------------------------+                               
  |                          +-- Lepidodesma_languilati      
  |                                                          
  |                    +-------- Aculamprotula_tortuosa **      
  |                    |                                     
  |     +--------------+     +-- Aculamprotula_coreana **       
  |     |              |  +--+                               
  |     |              |  |  +-- Aculamprotula_tientsinensis ** 
  |     |              +--+                                  
  |     |                 |  +-- Aculamprotula_polysticta **    
  |     |                 +--+                               
  |     |                    +-- Aculamprotula_scripta **       
  |     |                                                    
  |  +--+        +-------------- Unio_tumidus **                
  |  |  |  +-----+                                           
  |  |  |  |     |  +----------- Unio_crassus **                
  |  |  |  |     +--+                                        
--+  |  |  |        |  +-------- Unio_elongatulus **            
  |  |  |  |        +--+                                     
  |  |  |  |           |  +----- Unio_pictorum **               
  |  |  |  |           +--+                                  
  |  |  |  |              |  +-- Unio_delphinus **              
  |  |  +--+              +--+                               
  |  |     |                 +-- Unio_mancus **                 
  |  |     |                                                 
  |  |     |                 +-- Nodularia_breviconcha **       
  |  |     |  +--------------+                               
  |  |     |  |              +-- Nodularia_douglasiae **        
  |  |     |  |                                              
  |  |     |  |              +-- Pseudocuneopsis_capitata **    
  |  |     |  +--------------+                               
  |  |     +--+              +-- Pseudocuneopsis_sichuanensis **
  |  |        |                                              
  |  |        |              +-- Schistodesmus_lampreyanus **   
  +--+        |  +-----------+                               
     |        |  |           +-- Schistodesmus_spinosus **      
     |        +--+                                           
     |           |  +----------- Tchangsinaia_piscicula **      
     |           +--+                                        
     |              |  +-------- Cuneopsis_rufescens **         
     |              +--+                                     
     |                 |  +----- Cuneopsis_celtiformis **       
     |                 +--+                                  
     |                    |  +-- Cuneopsis_demangei **          
     |                    +--+                               
     |                       +-- Cuneopsis_heudei **            
     |                                                       
     |                 +-------- Acuticosta_chinensis        
     |  +--------------+                                     
     |  |              |  +----- Lanceolaria_gladiola        
     |  |              +--+                                  
     |  |                 |  +-- Lanceolaria_grayii          
     |  |                 +--+                               
     |  |                    +-- Lanceolaria_lanceolata      
     |  |                                                    
     |  |        +-------------- Cristaria_plicata           
     +--+        |                                           
        |  +-----+           +-- Anemina_arcaeformis         
        |  |     |  +--------+                               
        |  |     |  |        +-- Anemina_euscaphys           
        |  |     +--+                                        
        |  |        |  +-------- Beringiana_fukuharai        
        |  |        +--+                                     
        |  |           |  +----- Sinanodonta_lucida          
        |  |           +--+                                  
        +--+              |  +-- Sinanodonta_tumens          
           |              +--+                               
           |                 +-- Sinanodonta_woodiana        
           |                                                 
           |  +----------------- Anodonta_nuttalliana        
           |  |                                              
           |  |        +-------- Pseudanodonta_complanata    
           |  |  +-----+                                     
           +--+  |     |  +----- Anodonta_anatina            
              |  |     +--+                                  
              |  |        |  +-- Anodonta_cygnea             
              |  |        +--+                               
              +--+           +-- Anodonta_exulcerata         
                 |                                           
                 |        +----- Pyganodon_grandis           
                 |  +-----+                                  
                 |  |     |  +-- Utterbackia_imbecillis      
                 |  |     +--+                               
                 +--+        +-- Utterbackia_peninsularis    
                    |                                        
                    |  +-------- Lasmigona_complanata        
                    +--+                                     
                       |  +----- Prolasmidonta_heterodon     
                       +--+                                  
                          |  +-- Alasmidonta_varicosa        
                          +--+                               
                             +-- Platynaias_compressa        

Wu et al., 2022 [Publ 4238], fig. 3

iT tT bi riS oT roT roS iD
20 20 18 7 54 16 4 21.0
  +-------------------------------- Lepidodesma_languilati      
  |                                                             
  |                       +-------- Acuticosta_chinensis        
  |              +--------+                                     
  |              |        |  +----- Lanceolaria_gladiola        
  |              |        +--+                                  
  |              |           |  +-- Lanceolaria_grayii          
  |              |           +--+                               
  |              |              +-- Lanceolaria_lanceolata      
  |              |                                              
  |  +-----------+        +-------- Cristaria_plicata           
  |  |           |        |                                     
--+  |           |  +-----+     +-- Sinanodonta_woodiana        
  |  |           |  |     |  +--+                               
  |  |           |  |     |  |  +-- Sinanodonta_lucida          
  |  |           |  |     +--+                                  
  |  |           |  |        |  +-- Anemina_arcaeformis         
  |  |           +--+        +--+                               
  |  |              |           +-- Anemina_euscaphys           
  |  |              |                                           
  |  |              |           +-- Anodonta_cygnea             
  |  |              |  +--------+                               
  |  |              |  |        +-- Anodonta_anatina            
  |  |              +--+                                        
  |  |                 |  +-------- Platynaias_compressa        
  +--+                 +--+                                     
     |                    |  +----- Pyganodon_grandis           
     |                    +--+                                  
     |                       |  +-- Utterbackia_imbecillis      
     |                       +--+                               
     |                          +-- Utterbackia_peninsularis    
     |                                                          
     |                    +-------- Aculamprotula_tortuosa **      
     |                    |                                     
     |  +-----------------+     +-- Aculamprotula_tientsinensis ** 
     |  |                 |  +--+                               
     |  |                 |  |  +-- Aculamprotula_coreana **       
     |  |                 +--+                                  
     |  |                    |  +-- Aculamprotula_polysticta **    
     |  |                    +--+                               
     |  |                       +-- Aculamprotula_scripta **       
     +--+                                                       
        |                 +-------- Unio_tumidus **                
        |  +--------------+                                     
        |  |              |  +----- Unio_crassus **                
        |  |              +--+                                  
        |  |                 |  +-- Unio_pictorum **               
        |  |                 +--+                               
        +--+                    +-- Unio_delphinus **              
           |                                                    
           |                    +-- Pseudocuneopsis_capitata **    
           |  +-----------------+                               
           |  |                 +-- Pseudocuneopsis_sichuanensis **
           |  |                                                 
           +--+                 +-- Nodularia_douglasiae **        
              |  +--------------+                               
              |  |              +-- Nodularia_breviconcha **       
              |  |                                              
              +--+              +-- Schistodesmus_lampreyanus **   
                 |  +-----------+                               
                 |  |           +-- Schistodesmus_spinosus **      
                 +--+                                           
                    |  +----------- Tchangsinaia_piscicula **      
                    +--+                                        
                       |  +-------- Cuneopsis_rufescens **         
                       +--+                                     
                          |  +----- Cuneopsis_celtiformis **       
                          +--+                                  
                             |  +-- Cuneopsis_heudei **            
                             +--+                               
                                +-- Cuneopsis_demangei **          

Huang et al., 2019 [Publ 3843], fig. 2

iT tT bi riS oT roT roS iD
18 18 15 6 186 27 4 22.0
  +-------------------------------- Acuticosta_chinensis       
  |                                                            
  |  +----------------------------- Lepidodesma_languilati     
  |  |                                                         
  |  |                    +-------- Aculamprotula_tortuosa **     
  |  |                    |                                    
  |  |        +-----------+  +----- Aculamprotula_coreana **      
  |  |        |           |  |                                 
  |  |        |           +--+----- Aculamprotula_scripta **      
  |  |        |              |                                 
  |  |        |              |  +-- Aculamprotula_fibrosa **      
  |  |        |              +--+                              
  |  |        |                 +-- Aculamprotula_tientsinensis **
--+  +--------+                                                
  |  |        |           +-------- Unio_tumidus **               
  |  |        |  +--------+                                    
  |  |        |  |        |  +----- Unio_crassus **               
  |  |        |  |        +--+                                 
  |  |        |  |           |  +-- Unio_delphinus **             
  |  |        |  |           +--+                              
  |  |        +--+              +-- Unio_pictorum **              
  |  |           |                                             
  |  |           |              +-- Schistodesmus_lampreyanus **  
  |  |           |  +-----------+                              
  |  |           |  |           +-- Schistodesmus_spinosus **     
  |  |           |  |                                          
  |  |           +--+        +----- Nodularia_douglasiae[1] **    
  +--+              |  +-----+                                 
     |              |  |     |  +-- Nodularia_douglasiae[2] **    
     |              |  |     +--+                              
     |              +--+        +-- Nodularia_nuxpersicae **      
     |                 |                                       
     |                 |  +-------- Tchangsinaia_piscicula **     
     |                 +--+                                    
     |                    |  +----- Cuneopsis_rufescens **        
     |                    +--+                                 
     |                       |  +-- Cuneopsis_heudei **           
     |                       +--+                              
     |                          +-- Cuneopsis_celtiformis **      
     |                                                         
     |                    +-------- Lanceolaria_gladiola       
     |                    |                                    
     |  +-----------------+  +----- Lanceolaria_grayii[1]      
     |  |                 |  |                                 
     |  |                 +--+----- Lanceolaria_grayii[2]      
     |  |                    |                                 
     |  |                    |  +-- Lanceolaria_lanceolata     
     |  |                    +--+                              
     |  |                       +-- Lanceolaria_triformis      
     |  |                                                      
     |  |     +-------------------- Anemina_arcaeformis        
     |  |     |                                                
     |  |  +--+  +----------------- Cristaria_plicata          
     +--+  |  |  |                                             
        |  |  +--+  +-------------- Pletholophus_tenuis        
        |  |     |  |                                          
        |  |     +--+           +-- Sinanodonta_angula         
        |  |        |  +--------+                              
        |  |        |  |        +-- Sinanodonta_lucida         
        |  |        +--+                                       
        |  |           |  +-------- Sinanodonta_woodiana[2]    
        |  |           +--+                                    
        |  |              |  +----- Sinanodonta_jourdyi        
        |  |              +--+                                 
        +--+                 |  +-- Sinanodonta_elliptica      
           |                 +--+                              
           |                    +-- Sinanodonta_woodiana[1]    
           |                                                   
           |              +-------- Alasmidonta_marginata      
           |              |                                    
           |              +-------- Lasmigona_costata          
           |              |                                    
           |              +-------- Pseudanodonta_complanata   
           |              |                                    
           |              +-------- Simpsonaias_ambigua        
           |              |                                    
           +--------------+-------- Strophitus_undulatus       
                          |                                    
                          |  +----- Anodonta_anatina           
                          +--+                                 
                          |  |  +-- Anodonta_cygnea            
                          |  +--+                              
                          |     +-- Anodonta_nuttalliana       
                          |                                    
                          |  +----- Pyganodon_grandis          
                          +--+                                 
                             |  +-- Utterbackia_imbecillis     
                             +--+                              
                                +-- Utterbackia_peninsularis   

Huang et al., 2019 [Publ 3843], fig. S4

iT tT bi riS oT roT roS iD
18 44 21 6 184 26 4 48.0
  +----------------- Acuticosta_chinensis       
  |                                             
  +----------------- Alasmidonta_marginata      
  |                                             
  +----------------- Anodonta_anatina           
  |                                             
  +----------------- Anemina_arcaeformis        
  |                                             
  +----------------- Anodonta_cygnea            
  |                                             
  +----------------- Cristaria_plicata          
  |                                             
  +----------------- Anodonta_nuttalliana       
  |                                             
  +----------------- Lasmigona_costata          
  |                                             
  +----------------- Lepidodesma_languilati     
  |                                             
  +----------------- Pseudanodonta_complanata   
  |                                             
  +----------------- Pyganodon_grandis          
  |                                             
  +----------------- Simpsonaias_ambigua        
  |                                             
  +----------------- Strophitus_undulatus       
  |                                             
  |              +-- Schistodesmus_lampreyanus **  
  +--------------+                              
  |              +-- Schistodesmus_spinosus **     
  |                                             
  |              +-- Utterbackia_imbecillis     
  +--------------+                              
  |              +-- Utterbackia_peninsularis   
  |                                             
  |        +-------- Lanceolaria_gladiola       
  +--------+                                    
  |        |  +----- Lanceolaria_grayii         
  |        +--+                                 
--+           |  +-- Lanceolaria_lanceolata     
  |           +--+                              
  |              +-- Lanceolaria_triformis      
  |                                             
  |        +-------- Unio_tumidus **               
  +--------+                                    
  |        |  +----- Unio_crassus **               
  |        +--+                                 
  |           |  +-- Unio_delphinus **             
  |           +--+                              
  |              +-- Unio_pictorum **              
  |                                             
  |        +-------- Aculamprotula_tortuosa **     
  |        |                                    
  +--------+     +-- Aculamprotula_coreana **      
  |        |  +--+                              
  |        |  |  +-- Aculamprotula_scripta **      
  |        +--+                                 
  |           |  +-- Aculamprotula_fibrosa **      
  |           +--+                              
  |              +-- Aculamprotula_tientsinensis **
  |                                             
  |        +-------- Cuneopsis_heudei **           
  |        |                                    
  |        +-------- Cuneopsis_celtiformis **      
  |        |                                    
  +--------+-------- Tchangsinaia_piscicula **     
  |        |                                    
  |        +-------- Cuneopsis_rufescens **        
  |        |                                    
  |        |  +----- Nodularia_douglasiae[2] **    
  |        +--+                                 
  |           |  +-- Nodularia_douglasiae[1] **    
  |           +--+                              
  |              +-- Nodularia_nuxpersicae **      
  |                                             
  |  +-------------- Pletholophus_tenuis        
  |  |                                          
  +--+           +-- Sinanodonta_angula         
     |  +--------+                              
     |  |        +-- Sinanodonta_lucida         
     +--+                                       
        |  +-------- Sinanodonta_woodiana[1]    
        +--+                                    
           |  +----- Sinanodonta_jourdyi        
           +--+                                 
              |  +-- Sinanodonta_elliptica      
              +--+                              
                 +-- Sinanodonta_woodiana[2]    

Huang et al., 2019 [Publ 3843], fig. S3

iT tT bi riS oT roT roS iD
17 17 15 6 184 26 4 23.0
  +----------------------------------- Acuticosta_chinensis       
  |                                                               
  |                          +-------- Aculamprotula_tortuosa **     
  |                          |                                    
  |              +-----------+     +-- Aculamprotula_coreana **      
  |              |           |  +--+                              
  |              |           |  |  +-- Aculamprotula_scripta **      
  |              |           +--+                                 
  |              |              |  +-- Aculamprotula_fibrosa **      
  |              |              +--+                              
  |              |                 +-- Aculamprotula_tientsinensis **
  |  +-----------+                                                
  |  |           |           +-------- Unio_tumidus **               
--+  |           |  +--------+                                    
  |  |           |  |        |  +----- Unio_crassus **               
  |  |           |  |        +--+                                 
  |  |           |  |           |  +-- Unio_delphinus **             
  |  |           |  |           +--+                              
  |  |           +--+              +-- Unio_pictorum **              
  |  |              |                                             
  |  |              |           +----- Nodularia_douglasiae[1] **    
  |  |              |  +--------+                                 
  |  |              |  |        |  +-- Nodularia_douglasiae[2] **    
  |  |              |  |        +--+                              
  |  |              +--+           +-- Nodularia_nuxpersicae **      
  |  |                 |                                          
  +--+                 |  +----------- Schistodesmus_lampreyanus **  
     |                 +--+                                       
     |                    |  +-------- Tchangsinaia_piscicula **     
     |                    +--+                                    
     |                       |  +----- Cuneopsis_rufescens **        
     |                       +--+                                 
     |                          |  +-- Cuneopsis_heudei **           
     |                          +--+                              
     |                             +-- Cuneopsis_celtiformis **      
     |                                                            
     |  +----------------------------- Lepidodesma_languilati     
     |  |                                                         
     |  |                    +-------- Lanceolaria_gladiola       
     |  |  +-----------------+                                    
     |  |  |                 |  +----- Lanceolaria_grayii         
     |  |  |                 +--+                                 
     +--+  |                    |  +-- Lanceolaria_lanceolata     
        |  |                    +--+                              
        |  |                       +-- Lanceolaria_triformis      
        |  |                                                      
        |  |     +-------------------- Anemina_arcaeformis        
        |  |     |                                                
        |  |  +--+  +----------------- Cristaria_plicata          
        +--+  |  |  |                                             
           |  |  +--+  +-------------- Pletholophus_tenuis        
           |  |     |  |                                          
           |  |     +--+           +-- Sinanodonta_angula         
           |  |        |  +--------+                              
           |  |        |  |        +-- Sinanodonta_lucida         
           |  |        +--+                                       
           |  |           |  +-------- Sinanodonta_woodiana[2]    
           |  |           +--+                                    
           |  |              |  +----- Sinanodonta_jourdyi        
           +--+              +--+                                 
              |                 |  +-- Sinanodonta_elliptica      
              |                 +--+                              
              |                    +-- Sinanodonta_woodiana[1]    
              |                                                   
              |                    +-- Alasmidonta_marginata      
              |           +--------+                              
              |           |        +-- Lasmigona_costata          
              |           |                                       
              |           |     +----- Anodonta_anatina           
              |           +-----+                                 
              |           |     |  +-- Anodonta_cygnea            
              +-----------+     +--+                              
                          |        +-- Anodonta_nuttalliana       
                          |                                       
                          |     +----- Pseudanodonta_complanata   
                          |  +--+                                 
                          |  |  |  +-- Simpsonaias_ambigua        
                          |  |  +--+                              
                          +--+     +-- Strophitus_undulatus       
                             |                                    
                             |  +----- Pyganodon_grandis          
                             +--+                                 
                                |  +-- Utterbackia_imbecillis     
                                +--+                              
                                   +-- Utterbackia_peninsularis   

Chen et al., 2023 [Publ 4302], fig. 2

iT tT bi riS oT roT roS iD
17 17 14 10 23 14 4 0.0
  +-------------------------- Acuticosta_chinensis        
  |                                                       
  |  +----------------------- Lepidodesma_languilati      
  |  |                                                    
  |  |                    +-- Lanceolaria_gladiola        
  |  |     +--------------+                               
  |  |     |              +-- Lanceolaria_oxyrhyncha      
  |  |     |                                              
--+  |     |              +-- Alasmidonta_marginata       
  |  +-----+           +--+                               
  |  |     |           |  +-- Lasmigona_costata           
  |  |     |  +--------+                                  
  |  |     |  |        |  +-- Anodonta_anatina            
  |  |     |  |        +--+                               
  |  |     +--+           +-- Pseudanodonta_complanata    
  |  |        |                                           
  |  |        |  +----------- Cristaria_plicata           
  +--+        |  |                                        
     |        +--+        +-- Anemina_arcaeformis         
     |           |  +-----+                               
     |           |  |     +-- Amuranodonta_kijaensis      
     |           +--+                                     
     |              |  +----- Pletholophus_tenuis         
     |              +--+                                  
     |                 |  +-- Sinanodonta_jourdyi         
     |                 +--+                               
     |                    +-- Beringiana_beringiana       
     |                                                    
     |  +-------------------- Aculamprotula_fibrosa **       
     |  |                                                 
     |  |              +----- Middendorffinaia_mongolica **  
     |  |  +-----------+                                  
     +--+  |           |  +-- Nodularia_douglasiae **        
        |  |           +--+                               
        |  |              +-- Nodularia_breviconcha **       
        |  |                                              
        |  |              +-- Cuneopsis_celtiformis **       
        +--+  +-----------+                               
           |  |           +-- Cuneopsis_heudei **            
           |  |                                           
           |  |           +-- Diaurora_aurorea **            
           |  |        +--+                               
           |  |        |  +-- Diaurora_laeve **              
           |  +--------+                                  
           +--+        |  +-- Schistodesmus_lampreyanus **   
              |        +--+                               
              |           +-- Schistodesmus_spinosus **      
              |                                           
              |           +-- Pseudocuneopsis_sichuanensis **
              |  +--------+                               
              |  |        +-- Pseudocuneopsis_capitata **    
              |  |                                        
              +--+        +-- Inversiunio_yanagawensis **    
                 |  +-----+                               
                 |  |     +-- Inversiunio_reinianus **       
                 +--+                                     
                    |  +----- Tchangsinaia_piscicula **      
                    +--+                                  
                       |  +-- Unio_pictorum **               
                       +--+                               
                          +-- Unio_crassus **                

Dai et al., 2024 [Publ 4358], fig. 4

iT tT bi riS oT roT roS iD
17 17 15 7 35 10 4 20.0
  +-------------------------------- Lepidodesma_languilati       
  |                                                              
  |              +----------------- Lanceolaria_gladiola         
  |              |                                               
  |  +-----------+  +-------------- Acuticosta_chinensis         
  |  |           |  |                                            
  |  |           |  |        +----- Cristaria_plicata            
--+  |           +--+  +-----+                                   
  |  |              |  |     |  +-- Anemina_arcaeformis          
  |  |              |  |     +--+                                
  |  |              +--+        +-- Sinanodonta_woodiana         
  |  |                 |                                         
  |  |                 |  +-------- Anodonta_anatina             
  |  |                 +--+                                      
  +--+                    |  +----- Platynaias_compressa         
     |                    +--+                                   
     |                       |  +-- Pyganodon_grandis            
     |                       +--+                                
     |                          +-- Utterbackia_imbecillis       
     |                                                           
     |                          +-- Aculamprotula_scripta **        
     |  +-----------------------+                                
     |  |                       +-- Aculamprotula_tientsinensis **  
     |  |                                                        
     |  |                       +-- Unio_crassus **                 
     +--+  +--------------------+                                
        |  |                    +-- Unio_delphinus **               
        |  |                                                     
        |  |              +-------- Pseudocuneopsis_capitata **     
        +--+  +-----------+                                      
           |  |           |  +----- Pseudocuneopsis_yangshuoensis **
           |  |           +--+                                   
           |  |              |  +-- Pseudocuneopsis_yemaoi **       
           |  |              +--+                                
           +--+                 +-- Pseudocuneopsis_sichuanensis ** 
              |                                                  
              |                 +-- Nodularia_breviconcha **        
              |  +--------------+                                
              |  |              +-- Nodularia_douglasiae **         
              |  |                                               
              +--+              +-- Schistodesmus_lampreyanus **    
                 |  +-----------+                                
                 |  |           +-- Schistodesmus_spinosus **       
                 +--+                                            
                    |  +----------- Tchangsinaia_piscicula **       
                    +--+                                         
                       |  +-------- Cuneopsis_rufescens **          
                       +--+                                      
                          |  +----- Cuneopsis_celtiformis **        
                          +--+                                   
                             |  +-- Cuneopsis_heudei **             
                             +--+                                
                                +-- Cuneopsis_demangei **           

Lopes-Lima et al., 2021 [Publ 4217], fig. 1

iT tT bi riS oT roT roS iD
16 16 14 1 9 3 1 29.0
                       +----- Anodonta_anatina  
  +--------------------+                        
  |                    |  +-- Anodonta_cygnea   
  |                    +--+                     
  |                       +-- Anodonta_seddoni  
--+                                             
  |  +----------------------- Unio_tumidus **      
  |  |                                          
  |  |              +-------- Unio_damascensis **  
  |  |  +-----------+                           
  +--+  |           |  +----- Unio_sesirmensis **  
     |  |           +--+                        
     |  |              |  +-- Unio_bruguierianus **
     |  |              +--+                     
     +--+                 +-- Unio_crassus **      
        |                                       
        |                 +-- Unio_delicatus **    
        |  +--------------+                     
        |  |              +-- Unio_terminalis **   
        |  |                                    
        +--+              +-- Unio_tigridis **     
           |  +-----------+                     
           |  |           +-- Unio_hueti **        
           |  |                                 
           +--+  +----------- Unio_ravoisieri **   
              |  |                              
              |  |        +-- Unio_pictorum **     
              +--+  +-----+                     
                 |  |     +-- Unio_eucirrus **     
                 |  |                           
                 +--+     +-- Unio_elongatulus **  
                    |  +--+                     
                    |  |  +-- Unio_mancus **       
                    +--+                        
                       |  +-- Unio_foucauldianus **
                       +--+                     
                          +-- Unio_delphinus **    

Araujo et al., 2018 [Publ 3779], fig. 1

iT tT bi riS oT roT roS iD
15 15 10 2 4 3 1 26.0
                    +----- Anodonta_anatina     
  +-----------------+                           
  |                 |  +-- Pyganodon_grandis    
  |                 +--+                        
  |                    +-- Alasmidonta_marginata
--+                                             
  |  +-------------------- Nodularia_douglasiae ** 
  |  |                                          
  |  |                 +-- Unio_durieui **         
  +--+  +--------------+                        
     |  |              +-- Unio_gibbus **          
     |  |                                       
     +--+  +-------------- Unio_tumidus **         
        |  |                                    
        |  |     +-------- Unio_tumidiformis **    
        |  |  +--+                              
        +--+  |  |  +----- Unio_bruguierianus **   
           |  |  +--+                           
           |  |     |  +-- Unio_crassus **         
           |  |     +--+                        
           +--+        +-- Unio_ionicus **         
              |                                 
              |        +-- Unio_delphinus **       
              |     +--+                        
              |     |  +-- Unio_foucauldianus **   
              |     |                           
              +-----+  +-- Unio_pictorum **        
                    |  |                        
                    |  +-- Unio_elongatulus **     
                    |  |                        
                    +--+-- Unio_mancus **          
                       |                        
                       +-- Unio_ravoisieri **      
                       |                        
                       +-- Unio_tigridis **        

Sano et al., 2020 [Publ 4138], fig. 2

iT tT bi riS oT roT roS iD
15 15 13 6 58 24 3 23.0
                    +----- Lanceolaria_grayii[1]      
  +-----------------+                                 
  |                 |  +-- Lanceolaria_lanceolata     
  |                 +--+                              
  |                    +-- Lanceolaria_grayii[2]      
--+                                                   
  |  +-------------------- Lepidodesma_languilati     
  |  |                                                
  |  |              +----- Aculamprotula_tortuosa **     
  |  |  +-----------+                                 
  +--+  |           |  +-- Aculamprotula_tientsinensis **
     |  |           +--+                              
     |  |              +-- Aculamprotula_coreana **      
     |  |                                             
     +--+           +----- Unio_crassus **               
        |  +--------+                                 
        |  |        |  +-- Unio_delphinus **             
        |  |        +--+                              
        |  |           +-- Unio_pictorum **              
        |  |                                          
        +--+     +-------- Inversiunio_jokohamensis **   
           |  +--+                                    
           |  |  |  +----- Inversiunio_yanagawensis[2] **
           |  |  +--+                                 
           |  |     |  +-- Inversiunio_reinianus **      
           |  |     +--+                              
           +--+        +-- Inversiunio_yanagawensis[1] **
              |                                       
              |        +-- Tchangsinaia_piscicula **     
              |  +-----+                              
              |  |     +-- Schistodesmus_lampreyanus **  
              +--+                                    
                 |  +----- Nodularia_douglasiae[2] **    
                 +--+                                 
                    |  +-- Nodularia_nipponensis **      
                    +--+                              
                       +-- Nodularia_douglasiae[1] **    

Froufe et al., 2016 [Publ 3738], fig. 3

iT tT bi riS oT roT roS iD
14 14 12 2 0 0 0 23.0
                             +-- Nodularia_douglasiae ** 
  +--------------------------+                        
  |                          +-- Nodularia_breviconcha **
  |                                                   
--+                          +-- Unio_gibbus **          
  |  +-----------------------+                        
  |  |                       +-- Unio_durieui **         
  +--+                                                
     |  +----------------------- Unio_tumidus **         
     |  |                                             
     +--+                    +-- Unio_crassus **         
        |  +-----------------+                        
        |  |                 +-- Unio_tumidiformis **    
        +--+                                          
           |  +----------------- Unio_delphinus **       
           +--+                                       
              |  +-------------- Unio_foucauldianus **   
              +--+                                    
                 |  +----------- Unio_ravoisieri **      
                 +--+                                 
                    |  +-------- Unio_mancus **          
                    +--+                              
                       |  +----- Unio_elongatulus **     
                       +--+                           
                          |  +-- Unio_pictorum **        
                          +--+                        
                             +-- Unio_caffer **          

Wu et al., 2018 [Publ 3836], fig. 2

iT tT bi riS oT roT roS iD
14 14 12 6 39 19 4 18.0
  +----------------------- Lepidodesma_languilati     
  |                                                   
  |           +----------- Aculamprotula_tortuosa **     
  |     +-----+                                       
  |     |     |  +-------- Aculamprotula_scripta **      
  |     |     +--+                                    
  |     |        |  +----- Aculamprotula_coreana **      
  |     |        +--+                                 
  |     |           |  +-- Aculamprotula_fibrosa **      
  +-----+           +--+                              
  |     |              +-- Aculamprotula_tientsinensis **
  |     |                                             
  |     |              +-- Unio_crassus **               
  |     |  +-----------+                              
  |     |  |           +-- Unio_pictorum **              
  |     |  |                                          
  |     +--+        +----- Nodularia_douglasiae **       
  |        |  +-----+                                 
  |        |  |     |  +-- Schistodesmus_lampreyanus **  
--+        |  |     +--+                              
  |        +--+        +-- Schistodesmus_spinosus **     
  |           |                                       
  |           |  +-------- Tchangsinaia_piscicula **     
  |           +--+                                    
  |              |  +----- Cuneopsis_rufescens **        
  |              +--+                                 
  |                 |  +-- Cuneopsis_celtiformis **      
  |                 +--+                              
  |                    +-- Cuneopsis_heudei **           
  |                                                   
  |           +----------- Acuticosta_chinensis       
  |  +--------+                                       
  |  |        |  +-------- Lanceolaria_gladiola       
  |  |        +--+                                    
  |  |           |  +----- Lanceolaria_grayii         
  |  |           +--+                                 
  |  |              |  +-- Lanceolaria_lanceolata     
  |  |              +--+                              
  |  |                 +-- Lanceolaria_triformis      
  +--+                                                
     |           +-------- Anemina_arcaeformis        
     |           |                                    
     |           +-------- Cristaria_plicata          
     |  +--------+                                    
     |  |        |     +-- Sinanodonta_woodiana       
     |  |        |  +--+                              
     |  |        |  |  +-- Sinanodonta_elliptica      
     |  |        +--+                                 
     +--+           |  +-- Sinanodonta_lucida         
        |           +--+                              
        |              +-- Sinanodonta_angula         
        |                                             
        |  +-------------- Anodonta_cygnea            
        |  |                                          
        +--+  +----------- Anodonta_anatina           
           |  |                                       
           +--+  +-------- Strophitus_undulatus       
              |  |                                    
              +--+  +----- Platynaias_compressa       
                 |  |                                 
                 +--+----- Alasmidonta_marginata      
                    |                                 
                    |  +-- Pyganodon_cataracta        
                    +--+                              
                       +-- Pyganodon_grandis          

Huang et al., 2019 [Publ 3843], fig. 1

iT tT bi riS oT roT roS iD
14 20 18 5 30 14 4 26.0
  +----------------------------- Acuticosta_chinensis       
  |                                                         
  |                          +-- Sinanodonta_angula         
  |                 +--------+                              
  |                 |        +-- Sinanodonta_lucida         
  |  +--------------+                                       
  |  |              |  +-------- Sinanodonta_jourdyi        
--+  |              +--+                                    
  |  |                 |  +----- Sinanodonta_woodiana[2]    
  |  |                 +--+                                 
  |  |                    |  +-- Sinanodonta_elliptica      
  |  |                    +--+                              
  |  |                       +-- Sinanodonta_woodiana[1]    
  |  |                                                      
  +--+              +----------- Aculamprotula_tortuosa **     
     |  +-----------+                                       
     |  |           |  +-------- Aculamprotula_scripta **      
     |  |           +--+                                    
     |  |              |  +----- Aculamprotula_coreana **      
     |  |              +--+                                 
     |  |                 |  +-- Aculamprotula_fibrosa **      
     |  |                 +--+                              
     |  |                    +-- Aculamprotula_tientsinensis **
     +--+                                                   
        |           +----------- Cristaria_plicata          
        |  +--------+                                       
        |  |        |  +-------- Lanceolaria_gladiola       
        |  |        +--+                                    
        |  |           |  +----- Lanceolaria_grayii         
        |  |           +--+                                 
        |  |              |  +-- Lanceolaria_lanceolata     
        +--+              +--+                              
           |                 +-- Lanceolaria_triformis      
           |                                                
           |  +----------------- Lepidodesma_languilati     
           |  |                                             
           |  |              +-- Schistodesmus_lampreyanus **  
           +--+  +-----------+                              
              |  |           +-- Schistodesmus_spinosus **     
              |  |                                          
              +--+        +----- Nodularia_douglasiae[1] **    
                 |  +-----+                                 
                 |  |     |  +-- Nodularia_douglasiae[2] **    
                 |  |     +--+                              
                 +--+        +-- Nodularia_nuxpersicae **      
                    |                                       
                    |  +-------- Cuneopsis_rufescens **        
                    +--+                                    
                       |  +----- Tchangsinaia_piscicula **     
                       +--+                                 
                          |  +-- Cuneopsis_heudei **           
                          +--+                              
                             +-- Cuneopsis_celtiformis **      

Wu et al., 2020 [Publ 4121], fig. 3

iT tT bi riS oT roT roS iD
14 14 12 7 36 14 4 23.0
  +----------------------- Lepidodesma_languilati     
  |                                                   
  |                 +----- Acuticosta_chinensis       
  |     +-----------+                                 
  |     |           |  +-- Lanceolaria_lanceolata     
  |     |           +--+                              
  |     |              +-- Lanceolaria_grayii         
  |     |                                             
  |  +--+     +----------- Anodonta_anatina           
  |  |  |  +--+                                       
  |  |  |  |  |  +-------- Platynaias_compressa       
--+  |  |  |  +--+                                    
  |  |  |  |     |  +----- Pyganodon_grandis          
  |  |  |  |     +--+                                 
  |  |  +--+        |  +-- Utterbackia_peninsularis   
  |  |     |        +--+                              
  |  |     |           +-- Utterbackia_imbecillis     
  |  |     |                                          
  |  |     |     +-------- Cristaria_plicata          
  |  |     |     |                                    
  |  |     +-----+     +-- Sinanodonta_woodiana       
  |  |           |  +--+                              
  +--+           |  |  +-- Sinanodonta_lauta          
     |           +--+                                 
     |              |  +-- Anemina_arcaeformis        
     |              +--+                              
     |                 +-- Anemina_euscaphys          
     |                                                
     |           +-------- Aculamprotula_tortuosa **     
     |           |                                    
     |  +--------+     +-- Aculamprotula_coreana **      
     |  |        |  +--+                              
     |  |        |  |  +-- Aculamprotula_tientsinensis **
     |  |        +--+                                 
     |  |           |  +-- Aculamprotula_polysticta **   
     |  |           +--+                              
     +--+              +-- Aculamprotula_scripta **      
        |                                             
        |        +-------- Unio_tumidus **               
        |  +-----+                                    
        |  |     |  +----- Unio_crassus **               
        |  |     +--+                                 
        |  |        |  +-- Unio_delphinus **             
        +--+        +--+                              
           |           +-- Unio_pictorum **              
           |                                          
           |  +----------- Pseudocuneopsis_capitata **   
           +--+                                       
              |  +-------- Nodularia_douglasiae **       
              +--+                                    
                 |  +----- Tchangsinaia_piscicula **     
                 +--+                                 
                    |  +-- Cuneopsis_heudei **           
                    +--+                              
                       +-- Schistodesmus_lampreyanus **  

Sano et al., 2020 [Publ 4138], fig. 3

iT tT bi riS oT roT roS iD
14 14 12 6 58 24 3 22.0
                    +----- Lanceolaria_grayii[1]      
  +-----------------+                                 
  |                 |  +-- Lanceolaria_lanceolata     
  |                 +--+                              
  |                    +-- Lanceolaria_grayii[2]      
--+                                                   
  |  +-------------------- Lepidodesma_languilati     
  |  |                                                
  |  |              +----- Aculamprotula_tortuosa **     
  |  |  +-----------+                                 
  +--+  |           |  +-- Aculamprotula_tientsinensis **
     |  |           +--+                              
     |  |              +-- Aculamprotula_coreana **      
     |  |                                             
     +--+           +----- Unio_crassus **               
        |  +--------+                                 
        |  |        |  +-- Unio_delphinus **             
        |  |        +--+                              
        |  |           +-- Unio_pictorum **              
        |  |                                          
        +--+     +-------- Inversiunio_jokohamensis **   
           |  +--+                                    
           |  |  |  +----- Inversiunio_yanagawensis[2] **
           |  |  +--+                                 
           |  |     |  +-- Inversiunio_reinianus **      
           |  |     +--+                              
           +--+        +-- Inversiunio_yanagawensis[1] **
              |                                       
              |        +-- Nodularia_nipponensis **      
              |     +--+                              
              |     |  +-- Nodularia_douglasiae **       
              +-----+                                 
                    |  +-- Schistodesmus_lampreyanus **  
                    +--+                              
                       +-- Tchangsinaia_piscicula **     

Sano et al., 2020 [Publ 4138], fig. 4

iT tT bi riS oT roT roS iD
14 14 12 6 58 24 3 22.0
  +-------------------------- Lepidodesma_languilati     
  |                                                      
  |                    +----- Lanceolaria_grayii[1]      
  |  +-----------------+                                 
--+  |                 |  +-- Lanceolaria_lanceolata     
  |  |                 +--+                              
  |  |                    +-- Lanceolaria_grayii[2]      
  |  |                                                   
  +--+                 +----- Aculamprotula_tortuosa **     
     |  +--------------+                                 
     |  |              |  +-- Aculamprotula_tientsinensis **
     |  |              +--+                              
     |  |                 +-- Aculamprotula_coreana **      
     |  |                                                
     +--+              +----- Unio_crassus **               
        |  +-----------+                                 
        |  |           |  +-- Unio_delphinus **             
        |  |           +--+                              
        |  |              +-- Unio_pictorum **              
        +--+                                             
           |              +-- Nodularia_douglasiae **       
           |  +-----------+                              
           |  |           +-- Nodularia_nipponensis **      
           |  |                                          
           +--+           +-- Schistodesmus_lampreyanus **  
              |  +--------+                              
              |  |        +-- Tchangsinaia_piscicula **     
              +--+                                       
                 |  +-------- Inversiunio_jokohamensis **   
                 +--+                                    
                    |  +----- Inversiunio_yanagawensis[2] **
                    +--+                                 
                       |  +-- Inversiunio_reinianus **      
                       +--+                              
                          +-- Inversiunio_yanagawensis[1] **

Liu et al., 2023 [Publ 4310], fig. 4

iT tT bi riS oT roT roS iD
14 14 12 8 10 8 3 24.0
  +-------------------------------- Lanceolaria_gladiola         
  |                                                              
  |                       +-------- Pseudanodonta_complanata     
  |  +--------------------+                                      
--+  |                    |  +----- Lasmigona_costata            
  |  |                    +--+                                   
  |  |                       |  +-- Strophitus_undulatus         
  |  |                       +--+                                
  +--+                          +-- Pyganodon_grandis            
     |                                                           
     |  +-------------------------- Unio_tumidus **                 
     |  |                                                        
     |  |                       +-- Aculamprotula_scripta **        
     +--+  +--------------------+                                
        |  |                    +-- Aculamprotula_tientsinensis **  
        |  |                                                     
        +--+                    +-- Cuneopsis_heudei **             
           |  +-----------------+                                
           |  |                 +-- Cuneopsis_rufescens **          
           |  |                                                  
           +--+              +----- Nodularia_douglasiae **         
              |  +-----------+                                   
              |  |           |  +-- Schistodesmus_lampreyanus **    
              |  |           +--+                                
              +--+              +-- Schistodesmus_spinosus **       
                 |                                               
                 |  +-------------- Tchangsinaia_piscicula **       
                 +--+                                            
                    |  +----------- Inversiunio_yanagawensis **     
                    +--+                                         
                       |  +-------- Pseudocuneopsis_capitata **     
                       +--+                                      
                          |  +----- Pseudocuneopsis_sichuanensis ** 
                          +--+                                   
                             |  +-- Pseudocuneopsis_wuana **        
                             +--+                                
                                +-- Pseudocuneopsis_yangshuoensis **

Liu et al., 2023 [Publ 4310], fig. 5

iT tT bi riS oT roT roS iD
14 14 9 8 10 8 3 21.0
  +----------------------- Lanceolaria_grayii           
  |                                                     
  +----------------------- Lanceolaria_gladiola         
  |                                                     
  +----------------------- Lanceolaria_lanceolata       
  |                                                     
--+              +-------- Lasmigona_costata            
  |  +-----------+                                      
  |  |           |  +----- Pseudanodonta_complanata     
  |  |           +--+                                   
  |  |              |  +-- Pyganodon_grandis            
  |  |              +--+                                
  |  |                 +-- Strophitus_undulatus         
  +--+                                                  
     |  +----------------- Unio_tumidus **                 
     |  |                                               
     |  |              +-- Aculamprotula_tientsinensis **  
     |  +--------------+                                
     |  |              +-- Aculamprotula_scripta **        
     |  |                                               
     +--+              +-- Cuneopsis_rufescens **          
        |  +-----------+                                
        |  |           +-- Cuneopsis_heudei **             
        |  |                                            
        |  |        +----- Nodularia_douglasiae **         
        |  +--------+                                   
        +--+        |  +-- Schistodesmus_lampreyanus **    
           |        +--+                                
           |           +-- Schistodesmus_spinosus **       
           |                                            
           |  +----------- Tchangsinaia_piscicula **       
           |  |                                         
           +--+----------- Inversiunio_yanagawensis **     
              |                                         
              |  +-------- Pseudocuneopsis_capitata **     
              +--+                                      
                 |  +----- Pseudocuneopsis_sichuanensis ** 
                 +--+                                   
                    |  +-- Pseudocuneopsis_wuana **        
                    +--+                                
                       +-- Pseudocuneopsis_yangshuoensis **

Wu et al., 2019 [Publ 3864], fig. 3

iT tT bi riS oT roT roS iD
13 13 11 7 34 14 4 22.0
  +----------------------- Lepidodesma_languilati     
  |                                                   
  |              +-------- Aculamprotula_tortuosa **     
  |     +--------+                                    
  |     |        |  +----- Aculamprotula_scripta **      
  |     |        +--+                                 
  |     |           |  +-- Aculamprotula_tientsinensis **
  |     |           +--+                              
  |     |              +-- Aculamprotula_coreana **      
  |  +--+                                             
--+  |  |        +-------- Unio_tumidus **               
  |  |  |  +-----+                                    
  |  |  |  |     |  +----- Unio_crassus **               
  |  |  |  |     +--+                                 
  |  |  |  |        |  +-- Unio_delphinus **             
  |  |  +--+        +--+                              
  |  |     |           +-- Unio_pictorum **              
  |  |     |                                          
  |  |     |  +----------- Pseudocuneopsis_capitata **   
  |  |     +--+                                       
  |  |        |  +-------- Nodularia_douglasiae **       
  +--+        +--+                                    
     |           |  +----- Tchangsinaia_piscicula **     
     |           +--+                                 
     |              |  +-- Cuneopsis_heudei **           
     |              +--+                              
     |                 +-- Schistodesmus_lampreyanus **  
     |                                                
     |              +----- Acuticosta_chinensis       
     |  +-----------+                                 
     |  |           |  +-- Lanceolaria_grayii         
     |  |           +--+                              
     |  |              +-- Lanceolaria_lanceolata     
     |  |                                             
     +--+     +----------- Anodonta_anatina           
        |  +--+                                       
        |  |  |  +-------- Platynaias_compressa       
        |  |  +--+                                    
        |  |     |  +----- Pyganodon_grandis          
        |  |     +--+                                 
        +--+        |  +-- Utterbackia_peninsularis   
           |        +--+                              
           |           +-- Utterbackia_imbecillis     
           |                                          
           |     +-------- Cristaria_plicata          
           |     |                                    
           +-----+     +-- Anemina_arcaeformis        
                 |  +--+                              
                 |  |  +-- Anemina_euscaphys          
                 +--+                                 
                    |  +-- Sinanodonta_lucida         
                    +--+                              
                       +-- Sinanodonta_woodiana       

Liu et al., 2017 [Publ 3939], fig. 2

iT tT bi riS oT roT roS iD
13 13 8 4 4 3 3 19.0
  +-------------------- Lanceolaria_grayii       
  |                                              
  |              +----- Unio_tumidus **             
  |              |                               
--+  +-----------+----- Unio_gibbus **              
  |  |           |                               
  |  |           |  +-- Unio_crassus **             
  |  |           +--+                            
  |  |              +-- Unio_pictorum **            
  +--+                                           
     |  +-------------- Schistodesmus_lampreyanus **
     |  |                                        
     |  |           +-- Cuneopsis_heudei **         
     |  +-----------+                            
     +--+           +-- Cuneopsis_rufescens **      
        |                                        
        |  +----------- Nodularia_breviconcha **    
        |  |                                     
        +--+  +-------- Nodularia_douglasiae[3] **  
           |  |                                  
           +--+  +----- Nodularia_douglasiae[2] **  
              |  |                               
              +--+  +-- Nodularia_douglasiae[1] **  
                 |  |                            
                 +--+-- Nodularia_nuxpersicae **    
                    |                            
                    +-- Nodularia_nipponensis **    

Sayenko et al., 2021 [Publ 4189], fig. 6

iT tT bi riS oT roT roS iD
13 13 10 4 0 0 0 18.0
                    +-- Unio_pictorum **              
  +-----------------+                              
  |                 +-- Unio_crassus **               
  |                                                
  |           +-------- Inversiunio_jokohamensis **   
--+           |                                    
  |  +--------+  +----- Inversiunio_yanagawensis[2] **
  |  |        |  |                                 
  |  |        +--+----- Inversiunio_verrucosus **     
  |  |           |                                 
  +--+           |  +-- Inversiunio_reinianus **      
     |           +--+                              
     |              +-- Inversiunio_yanagawensis[1] **
     |                                             
     |  +-------------- Middendorffinaia_mongolica ** 
     +--+                                          
        |  +----------- Nodularia_breviconcha **      
        +--+                                       
           |  +-------- Nodularia_douglasiae[2] **    
           +--+                                    
              |  +----- Nodularia_nuxpersicae **      
              +--+                                 
                 |  +-- Nodularia_douglasiae[1] **    
                 +--+                              
                    +-- Nodularia_nipponensis **      

Wu et al., 2023 [Publ 4296], fig. 4

iT tT bi riS oT roT roS iD
13 13 11 8 10 8 3 23.0
                          +----- Lanceolaria_gladiola         
  +-----------------------+                                   
  |                       |  +-- Lanceolaria_lanceolata       
  |                       +--+                                
  |                          +-- Lanceolaria_grayii           
  |                                                           
--+                    +-------- Pseudanodonta_complanata     
  |  +-----------------+                                      
  |  |                 |  +----- Lasmigona_costata            
  |  |                 +--+                                   
  |  |                    |  +-- Strophitus_undulatus         
  |  |                    +--+                                
  +--+                       +-- Pyganodon_grandis            
     |                                                        
     |  +----------------------- Unio_tumidus **                 
     |  |                                                     
     |  |                    +-- Aculamprotula_scripta **        
     +--+  +-----------------+                                
        |  |                 +-- Aculamprotula_tientsinensis **  
        |  |                                                  
        +--+                 +-- Cuneopsis_heudei **             
           |  +--------------+                                
           |  |              +-- Cuneopsis_rufescens **          
           |  |                                               
           +--+           +----- Nodularia_douglasiae **         
              |  +--------+                                   
              |  |        |  +-- Schistodesmus_lampreyanus **    
              |  |        +--+                                
              +--+           +-- Schistodesmus_spinosus **       
                 |                                            
                 |  +----------- Tchangsinaia_piscicula **       
                 +--+                                         
                    |  +-------- Inversiunio_yanagawensis **     
                    +--+                                      
                       |  +----- Pseudocuneopsis_capitata **     
                       +--+                                   
                          |  +-- Pseudocuneopsis_yangshuoensis **
                          +--+                                
                             +-- Pseudocuneopsis_sichuanensis ** 

Fukata & Iigo, 2020 [Publ 4106], fig. 1

iT tT bi riS oT roT roS iD
12 12 10 7 42 16 4 21.0
                 +----- Lanceolaria_gladiola       
  +--------------+                                 
  |              |  +-- Lanceolaria_grayii         
  |              +--+                              
  |                 +-- Lanceolaria_lanceolata     
  |                                                
  |     +-------------- Acuticosta_chinensis       
  |     |                                          
  |     |     +-------- Cristaria_plicata          
  |     |     |                                    
  |     |  +--+     +-- Sinanodonta_woodiana       
  |  +--+  |  |  +--+                              
--+  |  |  |  |  |  +-- Sinanodonta_lucida         
  |  |  |  |  +--+                                 
  |  |  |  |     |  +-- Anemina_arcaeformis        
  |  |  |  |     +--+                              
  |  |  +--+        +-- Anemina_euscaphys          
  |  |     |                                       
  |  |     |        +-- Anodonta_anatina           
  |  |     |  +-----+                              
  |  |     |  |     +-- Anodonta_cygnea            
  |  |     |  |                                    
  |  |     +--+     +-- Pyganodon_grandis          
  +--+        |  +--+                              
     |        |  |  +-- Utterbackia_peninsularis   
     |        +--+                                 
     |           |  +-- Alasmidonta_varicosa       
     |           +--+                              
     |              +-- Prolasmidonta_heterodon    
     |                                             
     |           +----- Aculamprotula_tortuosa **     
     |  +--------+                                 
     |  |        |  +-- Aculamprotula_tientsinensis **
     |  |        +--+                              
     |  |           +-- Aculamprotula_coreana **      
     |  |                                          
     +--+     +-------- Unio_tumidus **               
        |  +--+                                    
        |  |  |  +----- Unio_crassus **               
        |  |  +--+                                 
        |  |     |  +-- Unio_pictorum **              
        |  |     +--+                              
        +--+        +-- Unio_delphinus **             
           |                                       
           |        +-- Schistodesmus_lampreyanus **  
           |  +-----+                              
           |  |     +-- Cuneopsis_heudei **           
           +--+                                    
              |  +----- Pseudocuneopsis_capitata **   
              +--+                                 
                 |  +-- Nodularia_douglasiae **       
                 +--+                              
                    +-- Tchangsinaia_piscicula **     

Sano et al., 2020 [Publ 4138], fig. 1

iT tT bi riS oT roT roS iD
12 34 5 5 36 22 4 41.0
  +-------- Lanceolaria_grayii[1]      
  |                                    
  +-------- Lanceolaria_grayii[2]      
  |                                    
  +-------- Cristaria_plicata[1]       
  |                                    
  +-------- Cristaria_plicata[2]       
  |                                    
  +-------- Sinanodonta_lauta          
  |                                    
  +-------- Beringiana_japonica        
  |                                    
  +-------- Beringiana_fukuharai[1]    
  |                                    
  +-------- Beringiana_beringiana      
  |                                    
  +-------- Beringiana_fukuharai[2]    
  |                                    
  +-------- Beringiana_fukuharai[3]    
  |                                    
  +-------- Buldowskia_iwakawai[1]     
  |                                    
  +-------- Buldowskia_iwakawai[2]     
  |                                    
  +-------- Sinanodonta_tumens[1]      
  |                                    
  +-------- Sinanodonta_tumens[2]      
  |                                    
--+-------- Pletholophus_tenuis        
  |                                    
  +-------- Anemina_arcaeformis        
  |                                    
  +-------- Lanceolaria_lanceolata     
  |                                    
  +-------- Cristaria_plicata[3]       
  |                                    
  +-------- Lanceolaria_gladiola       
  |                                    
  +-------- Sinanodonta_angula         
  |                                    
  +-------- Lepidodesma_languilati     
  |                                    
  |     +-- Aculamprotula_tortuosa **     
  +-----+                              
  |     +-- Acuticosta_chinensis       
  |                                    
  |  +----- Cuneopsis_celtiformis **      
  |  |                                 
  +--+  +-- Nodularia_douglasiae[1] **    
  |  |  |                              
  |  +--+-- Nodularia_douglasiae[2] **    
  |     |                              
  |     +-- Nodularia_nipponensis **      
  |                                    
  |  +----- Schistodesmus_lampreyanus **  
  |  |                                 
  |  |  +-- Inversiunio_reinianus[1] **   
  +--+  |                              
     |  +-- Inversiunio_reinianus[2] **   
     |  |                              
     |  +-- Inversiunio_jokohamensis[1] **
     +--+                              
        +-- Inversiunio_jokohamensis[2] **
        |                              
        +-- Inversiunio_jokohamensis[3] **
        |                              
        +-- Inversiunio_jokohamensis[4] **

Wu et al., 2022 [Publ 4238], fig. 2e

iT tT bi riS oT roT roS iD
12 12 10 7 5 5 2 17.0
                    +-------- Sinanodonta_woodiana        
  +-----------------+                                     
  |                 |  +----- Anodonta_anatina            
  |                 +--+                                  
  |                    |  +-- Utterbackia_peninsularis    
--+                    +--+                               
  |                       +-- Pyganodon_grandis           
  |                                                       
  |  +----------------------- Lanceolaria_lanceolata      
  |  |                                                    
  +--+  +-------------------- Aculamprotula_tortuosa **      
     |  |                                                 
     |  |              +----- Unio_tumidus **                
     +--+  +-----------+                                  
        |  |           |  +-- Unio_crassus **                
        |  |           +--+                               
        |  |              +-- Unio_delphinus **              
        +--+                                              
           |              +-- Pseudocuneopsis_sichuanensis **
           |  +-----------+                               
           |  |           +-- Pseudocuneopsis_capitata **    
           +--+                                           
              |  +----------- Nodularia_douglasiae **        
              |  |                                        
              +--+  +-------- Tchangsinaia_piscicula **      
                 |  |                                     
                 +--+     +-- Cuneopsis_celtiformis **       
                    |  +--+                               
                    |  |  +-- Cuneopsis_heudei **            
                    +--+                                  
                       |  +-- Schistodesmus_lampreyanus **   
                       +--+                               
                          +-- Schistodesmus_spinosus **      

Konopleva et al., 2023 [Publ 4269], fig. S1, 4

iT tT bi riS oT roT roS iD
12 12 10 6 44 21 4 24.0
           +----------------- Middendorffinaia_mongolica ** 
           |                                             
           |        +-------- Aculamprotula_tortuosa **     
           |  +-----+                                    
  +--------+  |     |  +----- Aculamprotula_scripta **      
  |        |  |     +--+                                 
  |        |  |        |  +-- Aculamprotula_tientsinensis **
  |        |  |        +--+                              
  |        +--+           +-- Aculamprotula_coreana **      
  |           |                                          
  |           |        +----- Unio_tumidus **               
  |           |  +-----+                                 
  |           |  |     |  +-- Unio_pictorum **              
  |           |  |     +--+                              
  |           +--+        +-- Unio_crassus **               
  |              |                                       
  |              |  +-------- Nodularia_douglasiae **       
--+              +--+                                    
  |                 |  +----- Tchangsinaia_piscicula **     
  |                 +--+                                 
  |                    |  +-- Cuneopsis_rufescens **        
  |                    +--+                              
  |                       +-- Cuneopsis_heudei **           
  |                                                      
  |                 +-------- Lepidodesma_languilati     
  |  +--------------+                                    
  |  |              |  +----- Lanceolaria_gladiola       
  |  |              +--+                                 
  |  |                 |  +-- Lanceolaria_grayii         
  |  |                 +--+                              
  +--+                    +-- Lanceolaria_triformis      
     |                                                   
     |  +-------------------- Acuticosta_chinensis       
     |  |                                                
     |  |              +----- Anodonta_anatina           
     |  |  +-----------+                                 
     +--+  |           |  +-- Pseudanodonta_complanata   
        |  |           +--+                              
        |  |              +-- Anodonta_cygnea            
        |  |                                             
        |  |        +-------- Anemina_arcaeformis        
        +--+  +-----+                                    
           |  |     |  +----- Amuranodonta_kijaensis     
           |  |     +--+                                 
           |  |        |  +-- Buldowskia_suifunica       
           |  |        +--+                              
           |  |           +-- Buldowskia_shadini         
           +--+                                          
              |     +-------- Beringiana_beringiana      
              |  +--+                                    
              |  |  |  +----- Sinanodonta_schrenkii      
              |  |  +--+                                 
              |  |     |  +-- Sinanodonta_woodiana       
              +--+     +--+                              
                 |        +-- Sinanodonta_lauta          
                 |                                       
                 |  +-------- Cristaria_radiata          
                 |  |                                    
                 +--+     +-- Cristaria_truncata         
                    |  +--+                              
                    |  |  +-- Cristaria_plicata          
                    +--+                                 
                       |  +-- Cristaria_bellua           
                       +--+                              
                          +-- Cristaria_clessini         

Prié & Puillandre, 2014 [Publ 3529], fig. 2

iT tT bi riS oT roT roS iD
11 11 9 1 6 3 1 24.0
                 +----- Anodonta_anatina        
  +--------------+                              
  |              |  +-- Pseudanodonta_complanata
  |              +--+                           
  |                 +-- Anodonta_cygnea         
  |                                             
--+              +----- Unio_tumidus **            
  |  +-----------+                              
  |  |           |  +-- Unio_gibbus **             
  |  |           +--+                           
  |  |              +-- Unio_durieui **            
  +--+                                          
     |           +----- Unio_tumidiformis **       
     |  +--------+                              
     |  |        |  +-- Unio_crassus **            
     |  |        +--+                           
     +--+           +-- Unio_courtillieri **       
        |                                       
        |  +----------- Unio_delphinus **          
        +--+                                    
           |  +-------- Unio_elongatulus **        
           +--+                                 
              |  +----- Unio_mancus **             
              +--+                              
                 |  +-- Unio_pictorum **           
                 +--+                           
                    +-- Unio_ravoisieri **         

Lopes-Lima et al., 2017 [Publ 3745], fig. 1-2

iT tT bi riS oT roT roS iD
11 11 8 6 70 18 2 22.0
                 +----- Unio_gibbus **              
                 |                               
     +-----------+----- Unio_tumidus **             
     |           |                               
     |           |  +-- Unio_crassus **             
     |           +--+                            
  +--+              +-- Unio_pictorum **            
  |  |                                           
  |  |  +-------------- Aculamprotula_tortuosa **   
  |  |  |                                        
  |  +--+           +-- Nodularia_nuxpersicae **    
  |     |  +--------+                            
  |     |  |        +-- Nodularia_douglasiae **     
  |     +--+                                     
  |        |  +-------- Schistodesmus_lampreyanus **
  |        +--+                                  
  |           |  +----- Tchangsinaia_piscicula **   
  |           +--+                               
--+              |  +-- Cuneopsis_heudei **         
  |              +--+                            
  |                 +-- Cuneopsis_rufescens **      
  |                                              
  |           +-------- Lanceolaria_gladiola     
  |  +--------+                                  
  |  |        |  +----- Lanceolaria_grayii[2]    
  |  |        +--+                               
  |  |           |  +-- Lanceolaria_lanceolata   
  |  |           +--+                            
  |  |              +-- Lanceolaria_grayii[1]    
  |  |                                           
  +--+     +----------- Cristaria_plicata        
     |  +--+                                     
     |  |  |  +-------- Anemina_arcaeformis      
     |  |  +--+                                  
     |  |     |  +----- Pletholophus_tenuis      
     |  |     +--+                               
     |  |        |  +-- Sinanodonta_jourdyi      
     |  |        +--+                            
     +--+           +-- Sinanodonta_lucida       
        |                                        
        |        +----- Pseudanodonta_complanata 
        |        |                               
        |  +-----+  +-- Anodonta_anatina         
        |  |     |  |                            
        |  |     +--+-- Anodonta_cygnea          
        |  |        |                            
        +--+        +-- Anodonta_nuttalliana     
           |                                     
           |        +-- Pyganodon_grandis        
           |  +-----+                            
           |  |     +-- Simpsonaias_ambigua      
           +--+                                  
              |  +----- Strophitus_undulatus     
              +--+                               
                 |  +-- Alasmidonta_marginata    
                 +--+                            
                    +-- Lasmigona_costata        

Huang et al., 2019 [Publ 3843], fig. 3

iT tT bi riS oT roT roS iD
11 11 9 5 46 16 4 21.0
  +----------------------------- Lepidodesma_languilati     
  |                                                         
  |                    +-------- Aculamprotula_tortuosa **     
  |              +-----+                                    
  |              |     |  +----- Aculamprotula_scripta **      
  |              |     +--+                                 
  |              |        |  +-- Aculamprotula_tientsinensis **
  |              |        +--+                              
  |  +-----------+           +-- Aculamprotula_coreana **      
--+  |           |                                          
  |  |           |        +----- Nodularia_douglasiae **       
  |  |           |  +-----+                                 
  |  |           |  |     |  +-- Tchangsinaia_piscicula **     
  |  |           |  |     +--+                              
  |  |           +--+        +-- Schistodesmus_lampreyanus **  
  |  |              |                                       
  |  |              |  +-------- Unio_tumidus **               
  |  |              +--+                                    
  +--+                 |  +----- Unio_crassus **               
     |                 +--+                                 
     |                    |  +-- Unio_delphinus **             
     |                    +--+                              
     |                       +-- Unio_pictorum **              
     |                                                      
     |                    +----- Lanceolaria_gladiola       
     |  +-----------------+                                 
     |  |                 |  +-- Lanceolaria_lanceolata     
     |  |                 +--+                              
     |  |                    +-- Lanceolaria_grayii         
     +--+                                                   
        |  +-------------------- Acuticosta_chinensis       
        |  |                                                
        |  |           +-------- Cristaria_plicata          
        |  |           |                                    
        +--+  +--------+     +-- Sinanodonta_lucida         
           |  |        |  +--+                              
           |  |        |  |  +-- Sinanodonta_woodiana       
           |  |        +--+                                 
           +--+           |  +-- Anemina_arcaeformis        
              |           +--+                              
              |              +-- Anemina_euscaphys          
              |                                             
              |  +-------------- Anodonta_cygnea            
              +--+                                          
                 |  +----------- Anodonta_anatina           
                 +--+                                       
                    |  +-------- Lasmigona_costata          
                    +--+                                    
                       |  +----- Pyganodon_grandis          
                       +--+                                 
                          |  +-- Utterbackia_imbecillis     
                          +--+                              
                             +-- Utterbackia_peninsularis   

Aunins et al., 2018 [Publ 3863], fig. 1

iT tT bi riS oT roT roS iD
11 11 9 6 30 16 4 20.0
  +----------------------- Lepidodesma_languilati     
  |                                                   
  |                 +----- Aculamprotula_tortuosa **     
  |        +--------+                                 
  |        |        |  +-- Aculamprotula_coreana **      
  |        |        +--+                              
  |        |           +-- Aculamprotula_tientsinensis **
  |        |                                          
  |  +-----+     +-------- Nodularia_douglasiae **       
--+  |     |  +--+                                    
  |  |     |  |  |  +----- Tchangsinaia_piscicula **     
  |  |     |  |  +--+                                 
  |  |     |  |     |  +-- Cuneopsis_heudei **           
  |  |     +--+     +--+                              
  |  |        |        +-- Schistodesmus_lampreyanus **  
  |  |        |                                       
  |  |        |  +-------- Unio_tumidus **               
  |  |        +--+                                    
  |  |           |  +----- Unio_crassus **               
  +--+           +--+                                 
     |              |  +-- Unio_pictorum **              
     |              +--+                              
     |                 +-- Unio_delphinus **             
     |                                                
     |              +----- Acuticosta_chinensis       
     |  +-----------+                                 
     |  |           |  +-- Lanceolaria_grayii         
     |  |           +--+                              
     |  |              +-- Lanceolaria_lanceolata     
     |  |                                             
     |  |        +-------- Cristaria_plicata          
     +--+        |                                    
        |  +-----+     +-- Anemina_arcaeformis        
        |  |     |  +--+                              
        |  |     |  |  +-- Anemina_euscaphys          
        |  |     +--+                                 
        |  |        |  +-- Sinanodonta_woodiana       
        |  |        +--+                              
        +--+           +-- Sinanodonta_lucida         
           |                                          
           |           +-- Anodonta_anatina           
           |  +--------+                              
           |  |        +-- Anodonta_cygnea            
           |  |                                       
           +--+        +-- Prolasmidonta_heterodon    
              |  +-----+                              
              |  |     +-- Alasmidonta_varicosa       
              +--+                                    
                 |  +----- Pyganodon_grandis          
                 +--+                                 
                    |  +-- Utterbackia_imbecillis     
                    +--+                              
                       +-- Utterbackia_peninsularis   

Choi et al., 2020 [Publ 4169], fig. 2a

iT tT bi riS oT roT roS iD
11 11 9 5 0 0 0 16.0
           +-------- Schistodesmus_lampreyanus **
           |                                  
  +--------+     +-- Unio_pictorum **            
  |        |  +--+                            
  |        |  |  +-- Tchangsinaia_piscicula **   
  |        +--+                               
--+           |  +-- Cuneopsis_heudei **         
  |           +--+                            
  |              +-- Cuneopsis_rufescens **      
  |                                           
  |  +-------------- Nodularia_breviconcha **    
  +--+                                        
     |  +----------- Nodularia[2] **             
     +--+                                     
        |  +-------- Nodularia[1] **             
        +--+                                  
           |  +----- Nodularia_douglasiae **     
           +--+                               
              |  +-- Nodularia_nipponensis **    
              +--+                            
                 +-- Nodularia_nuxpersicae **    

Bolotov et al., 2017 [Publ 3761], fig. S1

iT tT bi riS oT roT roS iD
10 10 8 6 131 17 2 21.0
                    +----- Unio_tumidus **             
           +--------+                               
           |        |  +-- Unio_pictorum **            
           |        +--+                            
           |           +-- Unio_crassus **             
  +--------+                                        
  |        |  +----------- Aculamprotula_tortuosa **   
  |        |  |                                     
  |        |  |        +-- Nodularia_douglasiae **     
  |        +--+  +-----+                            
  |           |  |     +-- Nodularia_nuxpersicae **    
  |           |  |                                  
  |           +--+     +-- Cuneopsis_rufescens **      
  |              |  +--+                            
  |              |  |  +-- Cuneopsis_heudei **         
  |              +--+                               
--+                 |  +-- Tchangsinaia_piscicula **   
  |                 +--+                            
  |                    +-- Schistodesmus_lampreyanus **
  |                                                 
  |                    +-- Lanceolaria_grayii       
  |  +-----------------+                            
  |  |                 +-- Lanceolaria_gladiola     
  |  |                                              
  |  |                 +-- Cristaria_plicata        
  |  |        +--------+                            
  |  |        |        +-- Cristaria_radiata        
  +--+  +-----+                                     
     |  |     |  +-------- Pletholophus_tenuis      
     |  |     +--+                                  
     |  |        |  +----- Sinanodonta_lucida       
     |  |        +--+                               
     |  |           |  +-- Sinanodonta_jourdyi      
     +--+           +--+                            
        |              +-- Sinanodonta_lauta        
        |                                           
        |  +-------------- Anodonta_anatina         
        |  |                                        
        |  |        +----- Pseudanodonta_complanata 
        +--+  +-----+                               
           |  |     |  +-- Anodonta_cygnea          
           |  |     +--+                            
           |  |        +-- Anodonta_nuttalliana     
           +--+                                     
              |        +-- Lasmigona_costata        
              |  +-----+                            
              |  |     +-- Alasmidonta_marginata    
              +--+                                  
                 |  +----- Simpsonaias_ambigua      
                 +--+                               
                    |  +-- Pyganodon_grandis        
                    +--+                            
                       +-- Strophitus_undulatus     

Bolotov et al., 2017 [Publ 3761], fig. S2

iT tT bi riS oT roT roS iD
10 10 6 6 131 17 2 19.0
              +----- Unio_tumidus **             
        +-----+                               
        |     |  +-- Unio_crassus **             
        |     +--+                            
        |        +-- Unio_pictorum **            
  +-----+                                     
  |     |  +-------- Aculamprotula_tortuosa **   
  |     |  |                                  
  |     |  +-------- Schistodesmus_lampreyanus **
  |     |  |                                  
  |     +--+     +-- Nodularia_douglasiae **     
  |        +-----+                            
  |        |     +-- Nodularia_nuxpersicae **    
  |        |                                  
  |        |  +----- Cuneopsis_heudei **         
  |        +--+                               
  |           |  +-- Tchangsinaia_piscicula **   
  |           +--+                            
--+              +-- Cuneopsis_rufescens **      
  |                                           
  |              +-- Lanceolaria_grayii       
  |  +-----------+                            
  |  |           +-- Lanceolaria_gladiola     
  |  |                                        
  |  |           +-- Cristaria_plicata        
  |  |  +--------+                            
  |  |  |        +-- Cristaria_radiata        
  |  +--+                                     
  |  |  |  +-------- Pletholophus_tenuis      
  |  |  +--+                                  
  |  |     |  +----- Sinanodonta_lucida       
  +--+     +--+                               
     |        |  +-- Sinanodonta_lauta        
     |        +--+                            
     |           +-- Sinanodonta_jourdyi      
     |                                        
     |     +-------- Anodonta_anatina         
     |     |                                  
     |     +-------- Pseudanodonta_complanata 
     |     |                                  
     |     |     +-- Anodonta_cygnea          
     +-----+-----+                            
           |     +-- Anodonta_nuttalliana     
           |                                  
           |  +----- Pyganodon_grandis        
           |  |                               
           |  +----- Simpsonaias_ambigua      
           +--+                               
              +----- Strophitus_undulatus     
              |                               
              |  +-- Alasmidonta_marginata    
              +--+                            
                 +-- Lasmigona_costata        

Bolotov et al., 2017 [Publ 3764], fig. S1

iT tT bi riS oT roT roS iD
10 10 0 6 121 18 2 18.0
     +-- Unio_crassus **             
     |                            
     +-- Unio_pictorum **            
     |                            
     +-- Unio_tumidus **             
     |                            
     +-- Aculamprotula_tortuosa **   
     |                            
     +-- Cuneopsis_heudei **         
  +--+                            
  |  +-- Tchangsinaia_piscicula **   
  |  |                            
  |  +-- Cuneopsis_rufescens **      
  |  |                            
  |  +-- Nodularia_douglasiae **     
  |  |                            
  |  +-- Nodularia_nuxpersicae **    
  |  |                            
  |  +-- Schistodesmus_lampreyanus **
  |                               
  |  +-- Alasmidonta_marginata    
  |  |                            
  |  +-- Anodonta_anatina         
--+  |                            
  |  +-- Anodonta_cygnea          
  |  |                            
  |  +-- Anodonta_nuttalliana     
  |  |                            
  |  +-- Lasmigona_costata        
  |  |                            
  |  +-- Pseudanodonta_complanata 
  |  |                            
  |  +-- Pyganodon_grandis        
  |  |                            
  |  +-- Simpsonaias_ambigua      
  |  |                            
  |  +-- Strophitus_undulatus     
  +--+                            
     +-- Sinanodonta_lauta        
     |                            
     +-- Cristaria_plicata        
     |                            
     +-- Cristaria_radiata        
     |                            
     +-- Pletholophus_tenuis      
     |                            
     +-- Sinanodonta_lucida       
     |                            
     +-- Sinanodonta_jourdyi      
     |                            
     +-- Lanceolaria_gladiola     
     |                            
     +-- Lanceolaria_grayii[1]    
     |                            
     +-- Lanceolaria_grayii[2]    

Sano et al., 2017 [Publ 3799], fig. 2

iT tT bi riS oT roT roS iD
10 15 12 4 26 14 4 22.0
                    +-- Cristaria_plicata          
        +-----------+                              
        |           +-- Buldowskia_iwakawai        
        |                                          
  +-----+        +----- Beringiana_fukuharai       
  |     |  +-----+                                 
  |     |  |     |  +-- Beringiana_japonica        
  |     |  |     +--+                              
  |     +--+        +-- Beringiana_beringiana      
  |        |                                       
  |        |  +-------- Pletholophus_tenuis        
  |        +--+                                    
  |           |  +----- Sinanodonta_woodiana       
  |           +--+                                 
--+              |  +-- Sinanodonta_lauta          
  |              +--+                              
  |                 +-- Sinanodonta_tumens         
  |                                                
  |           +-------- Tchangsinaia_piscicula **     
  |  +--------+                                    
  |  |        |  +----- Nodularia_nipponensis **      
  |  |        +--+                                 
  |  |           |  +-- Nodularia_douglasiae **       
  |  |           +--+                              
  |  |              +-- Nodularia_breviconcha **      
  +--+                                             
     |              +-- Schistodesmus_lampreyanus **  
     |  +-----------+                              
     |  |           +-- Acuticosta_ovata           
     |  |                                          
     |  |     +-------- Lepidodesma_languilati     
     +--+  +--+                                    
        |  |  |  +----- Lanceolaria_grayii[1]      
        |  |  +--+                                 
        |  |     |  +-- Lanceolaria_lanceolata     
        +--+     +--+                              
           |        +-- Lanceolaria_grayii[2]      
           |                                       
           |  +-------- Inversiunio_jokohamensis **   
           |  |                                    
           +--+  +----- Inversiunio_yanagawensis[3] **
              |  |                                 
              +--+  +-- Inversiunio_yanagawensis[1] **
                 |  |                              
                 +--+-- Inversiunio_yanagawensis[2] **
                    |                              
                    +-- Inversiunio_reinianus **      

Soroka & Burzynski, 2017 [Publ 3837], fig. 1

iT tT bi riS oT roT roS iD
10 10 8 5 14 14 3 20.0
                    +-- Lanceolaria_lanceolata     
  +-----------------+                              
  |                 +-- Lanceolaria_grayii         
  |                                                
--+  +----------------- Lepidodesma_languilati     
  |  |                                             
  |  |           +----- Aculamprotula_tortuosa **     
  +--+  +--------+                                 
     |  |        |  +-- Aculamprotula_tientsinensis **
     |  |        +--+                              
     |  |           +-- Aculamprotula_coreana **      
     +--+                                          
        |        +----- Schistodesmus_lampreyanus **  
        |  +-----+                                 
        |  |     |  +-- Tchangsinaia_piscicula **     
        |  |     +--+                              
        +--+        +-- Nodularia_douglasiae **       
           |                                       
           |  +-------- Unio_tumidus **               
           +--+                                    
              |  +----- Unio_crassus **               
              +--+                                 
                 |  +-- Unio_pictorum **              
                 +--+                              
                    +-- Unio_delphinus **             

Cha et al., 2018 [Publ 3916], fig. 1

iT tT bi riS oT roT roS iD
10 10 8 5 29 12 3 20.0
  +----------------------- Lepidodesma_languilati     
  |                                                   
  |                 +----- Aculamprotula_tortuosa **     
  |        +--------+                                 
  |        |        |  +-- Aculamprotula_coreana **      
  |        |        +--+                              
  |        |           +-- Aculamprotula_tientsinensis **
  |  +-----+                                          
--+  |     |        +----- Schistodesmus_lampreyanus **  
  |  |     |  +-----+                                 
  |  |     |  |     |  +-- Nodularia_douglasiae **       
  |  |     |  |     +--+                              
  |  |     +--+        +-- Tchangsinaia_piscicula **     
  |  |        |                                       
  |  |        |  +-------- Unio_tumidus **               
  |  |        +--+                                    
  +--+           |  +----- Unio_crassus **               
     |           +--+                                 
     |              |  +-- Unio_pictorum **              
     |              +--+                              
     |                 +-- Unio_delphinus **             
     |                                                
     |  +----------------- Lanceolaria_grayii         
     |  |                                             
     |  |     +----------- Anodonta_anatina           
     |  |  +--+                                       
     +--+  |  |  +-------- Platynaias_compressa       
        |  |  +--+                                    
        |  |     |  +----- Pyganodon_grandis          
        |  |     +--+                                 
        +--+        |  +-- Utterbackia_peninsularis   
           |        +--+                              
           |           +-- Utterbackia_imbecillis     
           |                                          
           |     +-------- Cristaria_plicata          
           |     |                                    
           +-----+     +-- Anemina_arcaeformis        
                 |  +--+                              
                 |  |  +-- Anemina_euscaphys          
                 +--+                                 
                    |  +-- Sinanodonta_lucida         
                    +--+                              
                       +-- Sinanodonta_woodiana       

Froufe et al., 2019 [Publ 3936], fig. 3

iT tT bi riS oT roT roS iD
10 10 8 5 35 13 3 20.0
  +----------------------- Lepidodesma_languilati     
  |                                                   
  |                 +----- Aculamprotula_tortuosa **     
  |        +--------+                                 
  |        |        |  +-- Aculamprotula_tientsinensis **
  |        |        +--+                              
  |        |           +-- Aculamprotula_coreana **      
  |  +-----+                                          
--+  |     |        +----- Nodularia_douglasiae **       
  |  |     |  +-----+                                 
  |  |     |  |     |  +-- Tchangsinaia_piscicula **     
  |  |     |  |     +--+                              
  |  |     +--+        +-- Schistodesmus_lampreyanus **  
  |  |        |                                       
  |  |        |  +-------- Unio_tumidus **               
  |  |        +--+                                    
  |  |           |  +----- Unio_crassus **               
  +--+           +--+                                 
     |              |  +-- Unio_pictorum **              
     |              +--+                              
     |                 +-- Unio_delphinus **             
     |                                                
     |                 +-- Lanceolaria_lanceolata     
     |  +--------------+                              
     |  |              +-- Lanceolaria_grayii         
     |  |                                             
     |  |        +-------- Cristaria_plicata          
     +--+        |                                    
        |  +-----+     +-- Anemina_arcaeformis        
        |  |     |  +--+                              
        |  |     |  |  +-- Anemina_euscaphys          
        |  |     +--+                                 
        +--+        |  +-- Sinanodonta_lucida         
           |        +--+                              
           |           +-- Sinanodonta_woodiana       
           |                                          
           |  +----------- Anodonta_anatina           
           +--+                                       
              |  +-------- Lasmigona_complanata       
              +--+                                    
                 |  +----- Pyganodon_grandis          
                 +--+                                 
                    |  +-- Utterbackia_peninsularis   
                    +--+                              
                       +-- Utterbackia_imbecillis     

Froufe et al., 2019 [Publ 3936], fig. 6

iT tT bi riS oT roT roS iD
10 10 8 5 35 13 3 20.0
  +----------------------- Lepidodesma_languilati     
  |                                                   
  |                 +----- Aculamprotula_tortuosa **     
  |        +--------+                                 
  |        |        |  +-- Aculamprotula_tientsinensis **
  |        |        +--+                              
  |        |           +-- Aculamprotula_coreana **      
  |  +-----+                                          
--+  |     |        +----- Nodularia_douglasiae **       
  |  |     |  +-----+                                 
  |  |     |  |     |  +-- Tchangsinaia_piscicula **     
  |  |     |  |     +--+                              
  |  |     +--+        +-- Schistodesmus_lampreyanus **  
  |  |        |                                       
  |  |        |  +-------- Unio_tumidus **               
  |  |        +--+                                    
  |  |           |  +----- Unio_crassus **               
  +--+           +--+                                 
     |              |  +-- Unio_pictorum **              
     |              +--+                              
     |                 +-- Unio_delphinus **             
     |                                                
     |                 +-- Lanceolaria_lanceolata     
     |  +--------------+                              
     |  |              +-- Lanceolaria_grayii         
     |  |                                             
     |  |     +----------- Anodonta_anatina           
     +--+  +--+                                       
        |  |  |  +-------- Lasmigona_complanata       
        |  |  +--+                                    
        |  |     |  +----- Pyganodon_grandis          
        |  |     +--+                                 
        +--+        |  +-- Utterbackia_peninsularis   
           |        +--+                              
           |           +-- Utterbackia_imbecillis     
           |                                          
           |     +-------- Cristaria_plicata          
           |     |                                    
           +-----+     +-- Sinanodonta_lucida         
                 |  +--+                              
                 |  |  +-- Sinanodonta_woodiana       
                 +--+                                 
                    |  +-- Anemina_arcaeformis        
                    +--+                              
                       +-- Anemina_euscaphys          

Fukata & Iigo, 2019 [Publ 4108], fig. 1

iT tT bi riS oT roT roS iD
10 14 12 7 4 4 2 23.0
     +----------- Aculamprotula_tientsinensis **
  +--+                                       
  |  |  +-------- Acuticosta_chinensis       
  |  +--+                                    
  |     |  +----- Lanceolaria_gladiola       
  |     +--+                                 
  |        |  +-- Lanceolaria_grayii         
  |        +--+                              
--+           +-- Lanceolaria_lanceolata     
  |                                          
  |     +-------- Unio_tumidus **               
  |  +--+                                    
  |  |  |  +----- Unio_crassus **               
  |  |  +--+                                 
  |  |     |  +-- Unio_delphinus **             
  +--+     +--+                              
     |        +-- Unio_pictorum **              
     |                                       
     |  +-------- Pseudocuneopsis_capitata **   
     |  |                                    
     +--+     +-- Nodularia_douglasiae **       
        |  +--+                              
        |  |  +-- Tchangsinaia_piscicula **     
        +--+                                 
           |  +-- Schistodesmus_lampreyanus **  
           +--+                              
              +-- Cuneopsis_heudei **           

Wu et al., 2020 [Publ 4147], fig. 3

iT tT bi riS oT roT roS iD
10 10 8 7 20 7 1 21.0
                 +----- Cristaria_plicata          
           +-----+                                 
           |     |  +-- Sinanodonta_lucida         
           |     +--+                              
  +--------+        +-- Anemina_arcaeformis        
  |        |                                       
  |        |  +-------- Anodonta_anatina           
  |        +--+                                    
  |           |  +----- Pyganodon_grandis          
  |           +--+                                 
  |              |  +-- Utterbackia_peninsularis   
--+              +--+                              
  |                 +-- Utterbackia_imbecillis     
  |                                                
  |           +-------- Aculamprotula_tortuosa **     
  |  +--------+                                    
  |  |        |  +----- Aculamprotula_scripta **      
  |  |        +--+                                 
  |  |           |  +-- Aculamprotula_tientsinensis **
  +--+           +--+                              
     |              +-- Aculamprotula_coreana **      
     |                                             
     |  +-------------- Unio_pictorum **              
     +--+                                          
        |  +----------- Pseudocuneopsis_capitata **   
        +--+                                       
           |  +-------- Nodularia_douglasiae **       
           +--+                                    
              |  +----- Schistodesmus_lampreyanus **  
              +--+                                 
                 |  +-- Cuneopsis_heudei **           
                 +--+                              
                    +-- Tchangsinaia_piscicula **     

Lopes-Lima et al., 2021 [Publ 4217], fig. 2

iT tT bi riS oT roT roS iD
10 10 8 1 0 0 0 25.0
  +----------------- Unio_tumidus **      
  |                                    
--+  +-------------- Unio_bruguierianus **
  |  |                                 
  |  |           +-- Unio_delicatus **    
  +--+  +--------+                     
     |  |        +-- Unio_terminalis **   
     |  |                              
     +--+        +-- Unio_tigridis **     
        |  +-----+                     
        |  |     +-- Unio_hueti **        
        |  |                           
        +--+     +-- Unio_pictorum **     
           |  +--+                     
           |  |  +-- Unio_eucirrus **     
           +--+                        
              |  +-- Unio_elongatulus **  
              +--+                     
                 +-- Unio_mancus **       

Zieritz et al., 2021 [Publ 4231], fig. 3-F, 5

iT tT bi riS oT roT roS iD
10 10 8 5 39 13 3 20.0
  +----------------------- Lepidodesma_languilati     
  |                                                   
  |                 +----- Aculamprotula_tortuosa **     
  |        +--------+                                 
  |        |        |  +-- Aculamprotula_tientsinensis **
  |        |        +--+                              
  |        |           +-- Aculamprotula_coreana **      
  |  +-----+                                          
--+  |     |        +----- Nodularia_douglasiae **       
  |  |     |  +-----+                                 
  |  |     |  |     |  +-- Schistodesmus_lampreyanus **  
  |  |     |  |     +--+                              
  |  |     +--+        +-- Tchangsinaia_piscicula **     
  |  |        |                                       
  |  |        |  +-------- Unio_tumidus **               
  |  |        +--+                                    
  |  |           |  +----- Unio_crassus **               
  +--+           +--+                                 
     |              |  +-- Unio_pictorum **              
     |              +--+                              
     |                 +-- Unio_delphinus **             
     |                                                
     |                 +-- Lanceolaria_grayii         
     |  +--------------+                              
     |  |              +-- Lanceolaria_lanceolata     
     |  |                                             
     |  |     +----------- Anodonta_anatina           
     +--+  +--+                                       
        |  |  |  +-------- Platynaias_compressa       
        |  |  +--+                                    
        |  |     |  +----- Pyganodon_grandis          
        |  |     +--+                                 
        +--+        |  +-- Utterbackia_imbecillis     
           |        +--+                              
           |           +-- Utterbackia_peninsularis   
           |                                          
           |     +-------- Cristaria_plicata          
           |     |                                    
           +-----+     +-- Sinanodonta_lucida         
                 |  +--+                              
                 |  |  +-- Sinanodonta_woodiana       
                 +--+                                 
                    |  +-- Anemina_arcaeformis        
                    +--+                              
                       +-- Anemina_euscaphys          

Jeratthitikul et al., 2022 [Publ 4260], fig. S1, 1

iT tT bi riS oT roT roS iD
10 10 8 6 66 16 2 21.0
                 +----- Unio_tumidus **             
     +-----------+                               
     |           |  +-- Unio_pictorum **            
     |           +--+                            
  +--+              +-- Unio_crassus **             
  |  |                                           
  |  |  +-------------- Aculamprotula_tortuosa **   
  |  |  |                                        
  |  +--+           +-- Nodularia_douglasiae **     
  |     |  +--------+                            
  |     |  |        +-- Nodularia_nuxpersicae **    
  |     +--+                                     
  |        |  +-------- Schistodesmus_lampreyanus **
  |        +--+                                  
  |           |  +----- Cuneopsis_heudei **         
--+           +--+                               
  |              |  +-- Tchangsinaia_piscicula **   
  |              +--+                            
  |                 +-- Cuneopsis_rufescens **      
  |                                              
  |                 +-- Lanceolaria_grayii       
  |  +--------------+                            
  |  |              +-- Lanceolaria_gladiola     
  |  |                                           
  |  |     +----------- Cristaria_plicata        
  |  |  +--+                                     
  +--+  |  |  +-------- Pletholophus_tenuis      
     |  |  +--+                                  
     |  |     |  +----- Sinanodonta_lucida       
     |  |     +--+                               
     |  |        |  +-- Sinanodonta_lauta        
     |  |        +--+                            
     +--+           +-- Sinanodonta_jourdyi      
        |                                        
        |     +-------- Pseudanodonta_complanata 
        |  +--+                                  
        |  |  |  +----- Anodonta_nuttalliana     
        |  |  +--+                               
        |  |     |  +-- Anodonta_anatina         
        |  |     +--+                            
        +--+        +-- Anodonta_cygnea          
           |                                     
           |        +-- Pyganodon_grandis        
           |  +-----+                            
           |  |     +-- Simpsonaias_ambigua      
           +--+                                  
              |  +----- Strophitus_undulatus     
              +--+                               
                 |  +-- Alasmidonta_marginata    
                 +--+                            
                    +-- Lasmigona_costata        

Jeratthitikul et al., 2022 [Publ 4260], fig. S2, 2

iT tT bi riS oT roT roS iD
10 10 8 6 66 16 2 21.0
                 +----- Unio_tumidus **             
     +-----------+                               
     |           |  +-- Unio_pictorum **            
     |           +--+                            
  +--+              +-- Unio_crassus **             
  |  |                                           
  |  |  +-------------- Aculamprotula_tortuosa **   
  |  |  |                                        
  |  +--+           +-- Nodularia_douglasiae **     
  |     |  +--------+                            
  |     |  |        +-- Nodularia_nuxpersicae **    
  |     +--+                                     
  |        |  +-------- Schistodesmus_lampreyanus **
  |        +--+                                  
  |           |  +----- Cuneopsis_heudei **         
--+           +--+                               
  |              |  +-- Tchangsinaia_piscicula **   
  |              +--+                            
  |                 +-- Cuneopsis_rufescens **      
  |                                              
  |                 +-- Lanceolaria_grayii       
  |  +--------------+                            
  |  |              +-- Lanceolaria_gladiola     
  |  |                                           
  |  |     +----------- Cristaria_plicata        
  |  |  +--+                                     
  +--+  |  |  +-------- Pletholophus_tenuis      
     |  |  +--+                                  
     |  |     |  +----- Sinanodonta_lucida       
     |  |     +--+                               
     |  |        |  +-- Sinanodonta_jourdyi      
     |  |        +--+                            
     +--+           +-- Sinanodonta_lauta        
        |                                        
        |     +-------- Pseudanodonta_complanata 
        |  +--+                                  
        |  |  |  +----- Anodonta_anatina         
        |  |  +--+                               
        |  |     |  +-- Anodonta_nuttalliana     
        |  |     +--+                            
        +--+        +-- Anodonta_cygnea          
           |                                     
           |        +-- Alasmidonta_marginata    
           |  +-----+                            
           |  |     +-- Lasmigona_costata        
           +--+                                  
              |  +----- Pyganodon_grandis        
              +--+                               
                 |  +-- Strophitus_undulatus     
                 +--+                            
                    +-- Simpsonaias_ambigua      

Zhou et al., 2007 [Publ 2658], fig. 1

iT tT bi riS oT roT roS iD
9 13 11 4 14 6 3 24.0
                 +-- Schistodesmus_lampreyanus **  
           +-----+                              
           |     +-- Schistodesmus_spinosus **     
           |                                    
  +--------+     +-- Lanceolaria_triformis      
  |        |  +--+                              
  |        |  |  +-- Lanceolaria_lanceolata     
  |        +--+                                 
  |           |  +-- Acuticosta_ovata           
--+           +--+                              
  |              +-- Acuticosta_chinensis       
  |                                             
  |              +-- Tchangsinaia_piscicula **     
  |  +-----------+                              
  |  |           +-- Nodularia_douglasiae **       
  +--+                                          
     |  +----------- Aculamprotula_tortuosa **     
     +--+                                       
        |  +-------- Aculamprotula_scripta **      
        +--+                                    
           |  +----- Aculamprotula_tientsinensis **
           +--+                                 
              |  +-- Aculamprotula_zonata **       
              +--+                              
                 +-- Aculamprotula_fibrosa **      

Khalloufi et al., 2011 [Publ 2748], fig. 2

iT tT bi riS oT roT roS iD
9 9 6 1 2 0 0 22.0
        +----- Unio_tumidus **     
  +-----+                       
  |     |  +-- Unio_gibbus **      
  |     +--+                    
  |        +-- Unio_durieui **     
--+                             
  |        +-- Unio_tumidiformis **
  |  +-----+                    
  |  |     +-- Unio_crassus **     
  +--+                          
     |  +----- Unio **             
     |  |                       
     +--+----- Unio_pictorum **    
        |                       
        |  +-- Unio_delphinus **   
        +--+                    
           +-- Unio_mancus **      

Lopes-Lima et al., 2017 [Publ 3734], fig. 1

iT tT bi riS oT roT roS iD
9 9 5 1 8 3 1 22.0
           +----- Anodonta_anatina        
  +--------+                              
  |        |  +-- Pseudanodonta_complanata
  |        +--+                           
  |           +-- Anodonta_cygnea         
--+                                       
  |           +-- Unio_tumidus **            
  |  +--------+                           
  |  |        +-- Unio_gibbus **             
  |  |                                    
  +--+        +-- Unio_crassus **            
     |  +-----+                           
     |  |     +-- Unio_tumidiformis **       
     +--+                                 
        |  +----- Unio_ravoisieri **         
        |  |                              
        +--+  +-- Unio_mancus **             
           |  |                           
           |  +-- Unio_pictorum **           
           +--+                           
              +-- Unio_delphinus **          
              |                           
              +-- Unio_elongatulus **        

Huang et al., 2018 [Publ 3828], fig. 6

iT tT bi riS oT roT roS iD
9 9 7 4 34 10 1 19.0
              +----- Aculamprotula_tortuosa **     
     +--------+                                 
     |        |  +-- Aculamprotula_coreana **      
     |        +--+                              
     |           +-- Aculamprotula_tientsinensis **
  +--+                                          
  |  |           +-- Tchangsinaia_piscicula **     
  |  |  +--------+                              
  |  |  |        +-- Nodularia_douglasiae **       
  |  +--+                                       
  |     |  +-------- Unio_tumidus **               
  |     +--+                                    
  |        |  +----- Unio_crassus **               
  |        +--+                                 
--+           |  +-- Unio_delphinus **             
  |           +--+                              
  |              +-- Unio_pictorum **              
  |                                             
  |     +----------- Anodonta_anatina           
  |  +--+                                       
  |  |  |  +-------- Platynaias_compressa       
  |  |  +--+                                    
  |  |     |  +----- Pyganodon_grandis          
  |  |     +--+                                 
  +--+        |  +-- Utterbackia_imbecillis     
     |        +--+                              
     |           +-- Utterbackia_peninsularis   
     |                                          
     |     +-------- Cristaria_plicata          
     |     |                                    
     +-----+     +-- Anemina_arcaeformis        
           |  +--+                              
           |  |  +-- Anemina_euscaphys          
           +--+                                 
              |  +-- Sinanodonta_woodiana       
              +--+                              
                 +-- Sinanodonta_lucida         

Kim et al., 2021 [Publ 4197], fig. 1

iT tT bi riS oT roT roS iD
9 13 11 5 6 5 3 20.0
  +----------------- Aculamprotula_tientsinensis **
  |                                             
  |        +-------- Acuticosta_chinensis       
  |  +-----+                                    
--+  |     |  +----- Lanceolaria_gladiola       
  |  |     +--+                                 
  |  |        |  +-- Lanceolaria_grayii         
  |  |        +--+                              
  |  |           +-- Lanceolaria_lanceolata     
  +--+                                          
     |     +-------- Unio_tumidus **               
     |  +--+                                    
     |  |  |  +----- Unio_crassus **               
     |  |  +--+                                 
     |  |     |  +-- Unio_delphinus **             
     |  |     +--+                              
     +--+        +-- Unio_pictorum **              
        |                                       
        |        +-- Nodularia_douglasiae **       
        |     +--+                              
        |     |  +-- Nodularia_breviconcha **      
        +-----+                                 
              |  +-- Schistodesmus_lampreyanus **  
              +--+                              
                 +-- Cuneopsis_heudei **           

Araujo et al., 2009 [Publ 2589], fig. 7A

iT tT bi riS oT roT roS iD
8 9 4 2 4 1 1 20.0
  +----------- Nodularia_douglasiae **
  |                                
  |        +-- Unio_gibbus **         
--+        |                       
  |  +-----+-- Unio_tumidus **        
  |  |     |                       
  |  |     +-- Anodonta_cygnea     
  +--+                             
     |  +----- Unio_pictorum[1] **    
     |  |                          
     |  +----- Unio_pictorum[2] **    
     +--+                          
        +----- Unio_mancus **         
        |                          
        |  +-- Unio_tumidiformis **   
        +--+                       
           +-- Unio_crassus **        

Araujo et al., 2009 [Publ 2589], fig. 7B

iT tT bi riS oT roT roS iD
8 8 6 1 4 1 1 21.0
  +----------------- Anodonta_cygnea  
  |                                   
--+  +-------------- Unio_gibbus **      
  |  |                                
  +--+  +----------- Unio_tumidus **     
     |  |                             
     |  |        +-- Unio_tumidiformis **
     +--+  +-----+                    
        |  |     +-- Unio_crassus **     
        |  |                          
        +--+     +-- Unio_pictorum[1] ** 
           |  +--+                    
           |  |  +-- Unio_mancus **      
           +--+                       
              |  +-- Unio_pictorum[2] ** 
              +--+                    
                 +-- Unio_caffer **      

Reis et al., 2013 [Publ 3580], fig. 5

iT tT bi riS oT roT roS iD
8 8 6 1 8 3 1 21.0
                 +-- Anodonta_anatina        
                 |                           
  +--------------+-- Anodonta_cygnea         
  |              |                           
  |              +-- Pseudanodonta_complanata
--+                                          
  |  +-------------- Unio_tumidus **            
  |  |                                       
  +--+           +-- Unio_tumidiformis **       
     |  +--------+                           
     |  |        +-- Unio_crassus **            
     +--+                                    
        |  +-------- Unio **                    
        |  |                                 
        +--+     +-- Unio_delphinus **          
           |  +--+                           
           |  |  +-- Unio_mancus **             
           +--+                              
              |  +-- Unio_pictorum **           
              +--+                           
                 +-- Unio_caffer **             

Sano et al., 2017 [Publ 3799], fig. 1

iT tT bi riS oT roT roS iD
8 8 6 2 19 9 2 17.0
                       +-- Cristaria_plicata          
           +-----------+                              
           |           +-- Buldowskia_iwakawai        
  +--------+                                          
  |        |  +----------- Pletholophus_tenuis        
  |        |  |                                       
  |        +--+        +-- Sinanodonta_tumens         
  |           |  +-----+                              
  |           |  |     +-- Sinanodonta_lauta          
  |           +--+                                    
--+              |  +----- Beringiana_fukuharai       
  |              +--+                                 
  |                 |  +-- Beringiana_japonica        
  |                 +--+                              
  |                    +-- Beringiana_beringiana      
  |                                                   
  |  +-------------------- Lanceolaria_grayii         
  |  |                                                
  +--+                 +-- Nodularia_nipponensis **      
     |  +--------------+                              
     |  |              +-- Nodularia_douglasiae **       
     +--+                                             
        |  +-------------- Inversiunio_jokohamensis **   
        +--+                                          
           |  +----------- Inversiunio_yanagawensis[1] **
           +--+                                       
              |  +-------- Inversiunio_yanagawensis[2] **
              +--+                                    
                 |  +----- Inversiunio_yanagawensis[3] **
                 +--+                                 
                    |  +-- Inversiunio_yanagawensis[4] **
                    +--+                              
                       +-- Inversiunio_reinianus **      

Wang et al., 2018 [Publ 3846], fig. 2

iT tT bi riS oT roT roS iD
8 8 6 5 24 10 2 18.0
                 +----- Anodonta_anatina           
           +-----+                                 
           |     |  +-- Pyganodon_grandis          
           |     +--+                              
  +--------+        +-- Platynaias_compressa       
  |        |                                       
  |        |  +-------- Cristaria_plicata          
  |        |  |                                    
  |        +--+     +-- Anemina_euscaphys          
  |           |  +--+                              
  |           |  |  +-- Anemina_arcaeformis        
  |           +--+                                 
--+              |  +-- Sinanodonta_woodiana       
  |              +--+                              
  |                 +-- Sinanodonta_lucida         
  |                                                
  |                 +-- Lanceolaria_grayii         
  |  +--------------+                              
  |  |              +-- Lanceolaria_lanceolata     
  |  |                                             
  +--+  +-------------- Aculamprotula_tientsinensis **
     |  |                                          
     |  |        +----- Nodularia_douglasiae **       
     +--+  +-----+                                 
        |  |     |  +-- Tchangsinaia_piscicula **     
        |  |     +--+                              
        +--+        +-- Cuneopsis_heudei **           
           |                                       
           |  +-------- Unio_tumidus **               
           +--+                                    
              |  +----- Unio_crassus **               
              +--+                                 
                 |  +-- Unio_pictorum **              
                 +--+                              
                    +-- Unio_delphinus **             

Klishko et al., 2019 [Publ 3886], fig. 9

iT tT bi riS oT roT roS iD
8 8 5 3 3 2 1 15.0
              +-- Anodonta_anatina          
  +-----------+                             
  |           +-- Anodonta_cygnea           
  |                                         
  |     +-------- Middendorffinaia_mongolica **
--+  +--+                                   
  |  |  |  +----- Nodularia_breviconcha **     
  |  |  +--+                                
  |  |     |  +-- Nodularia_douglasiae **      
  +--+     +--+                             
     |        +-- Nodularia **                 
     |                                      
     |     +----- Unio_tumidus **              
     |     |                                
     +-----+----- Unio_gibbus **               
           |                                
           |  +-- Unio_crassus **              
           +--+                             
              +-- Unio_pictorum **             

Marrone et al., 2019 [Publ 3907], fig. 1

iT tT bi riS oT roT roS iD
8 8 4 1 0 0 0 23.0
  +-------------- Unio_tumidus **      
  |                                 
--+  +----------- Unio_delphinus **    
  |  |                              
  +--+  +-------- Unio_foucauldianus **
     |  |                           
     +--+  +----- Unio_mancus **       
        |  |                        
        |  +----- Unio_elongatulus **  
        +--+                        
           +----- Unio_ravoisieri **   
           |                        
           |  +-- Unio_pictorum **     
           +--+                     
              +-- Unio_tigridis **     

Klishko et al., 2018 [Publ 3945], fig. 8

iT tT bi riS oT roT roS iD
8 8 5 2 0 0 0 17.0
     +-------- Nodularia_breviconcha **
  +--+                              
  |  |  +----- Nodularia **            
  |  +--+                           
  |     |  +-- Nodularia_douglasiae ** 
--+     +--+                        
  |        +-- Nodularia_nipponensis **
  |                                 
  |     +----- Unio_tumidus **         
  |     |                           
  +-----+----- Unio_gibbus **          
        |                           
        |  +-- Unio_crassus **         
        +--+                        
           +-- Unio_pictorum **        

Lopes-Lima et al., 2020 [Publ 4080], fig. 13

iT tT bi riS oT roT roS iD
8 8 6 2 2 0 0 17.0
  +-------------------- Middendorffinaia_mongolica **
--+                                               
  |  +----------------- Nodularia_breviconcha **     
  +--+                                            
     |  +-------------- Nodularia[3] **              
     +--+                                         
        |  +----------- Nodularia[2] **              
        +--+                                      
           |  +-------- Nodularia_nuxpersicae **     
           +--+                                   
              |  +----- Nodularia_nipponensis **     
              +--+                                
                 |  +-- Nodularia_douglasiae **      
                 +--+                             
                    +-- Nodularia[1] **              

Araujo et al., 2009 [Publ 2589], fig. 7C

iT tT bi riS oT roT roS iD
7 7 4 1 4 1 1 20.0
  +----------- Anodonta_cygnea  
  |                             
  |        +-- Unio_gibbus **      
--+  +-----+                    
  |  |     +-- Unio_tumidus **     
  |  |                          
  +--+     +-- Unio_tumidiformis **
     |  +--+                    
     |  |  +-- Unio_crassus **     
     +--+                       
        |  +-- Unio_pictorum[1] ** 
        |  |                    
        +--+-- Unio_pictorum[2] ** 
           |                    
           +-- Unio_mancus **      

Zhou et al., 2007 [Publ 2658], fig. 2

iT tT bi riS oT roT roS iD
7 8 6 3 14 3 2 19.0
           +----- Acuticosta_chinensis       
  +--------+                                 
  |        |  +-- Nodularia_douglasiae **       
  |        +--+                              
--+           +-- Cuneopsis_celtiformis **      
  |                                          
  |  +----------- Aculamprotula_tortuosa **     
  +--+                                       
     |  +-------- Aculamprotula_scripta **      
     +--+                                    
        |  +----- Aculamprotula_fibrosa **      
        +--+                                 
           |  +-- Aculamprotula_zonata **       
           +--+                              
              +-- Aculamprotula_tientsinensis **

Zhang et al., 2004 [Publ 2735], fig. 1

iT tT bi riS oT roT roS iD
7 7 5 5 7 5 3 17.0
                       +-- Sinanodonta_woodiana     
  +--------------------+                            
  |                    +-- Cristaria_plicata        
  |                                                 
--+                 +----- Acuticosta_chinensis     
  |  +--------------+                               
  |  |              |  +-- Lanceolaria_grayii       
  |  |              +--+                            
  +--+                 +-- Lanceolaria_lanceolata   
     |                                              
     |  +----------------- Tchangsinaia_piscicula **   
     +--+                                           
        |  +-------------- Cuneopsis_heudei **         
        +--+                                        
           |  +----------- Nodularia_douglasiae **     
           +--+                                     
              |  +-------- Pseudocuneopsis_capitata ** 
              +--+                                  
                 |  +----- Cuneopsis_rufescens **      
                 +--+                               
                    |  +-- Schistodesmus_lampreyanus **
                    +--+                            
                       +-- Schistodesmus_spinosus **   

Pfeiffer & Graf, 2013 [Publ 3531], fig. 2

iT tT bi riS oT roT roS iD
7 7 4 3 28 6 2 18.0
                 +-- Cristaria_plicata          
           +-----+                              
           |     +-- Sinanodonta_woodiana       
  +--------+                                    
  |        |  +----- Strophitus_undulatus       
  |        +--+                                 
  |           |  +-- Platynaias_compressa       
--+           +--+                              
  |              +-- Pyganodon_grandis          
  |                                             
  |  +-------------- Acuticosta_chinensis       
  |  |                                          
  +--+           +-- Unio_pictorum **              
     |  +--------+                              
     |  |        +-- Nodularia_douglasiae **       
     +--+                                       
        |  +-------- Aculamprotula_tortuosa **     
        |  |                                    
        +--+  +----- Aculamprotula_scripta **      
           |  |                                 
           +--+  +-- Aculamprotula_tientsinensis **
              |  |                              
              +--+-- Aculamprotula_fibrosa **      
                 |                              
                 +-- Aculamprotula_zonata **       

Fonseca et al., 2016 [Publ 3775], fig. 1

iT tT bi riS oT roT roS iD
7 7 5 5 25 12 2 19.0
                 +-- Lanceolaria_lanceolata   
     +-----------+                            
     |           +-- Lanceolaria_grayii       
     |                                        
  +--+           +-- Aculamprotula_tortuosa **   
  |  |  +--------+                            
  |  |  |        +-- Aculamprotula_coreana **    
  |  |  |                                     
  |  +--+        +-- Unio_pictorum **            
  |     |  +-----+                            
  |     |  |     +-- Unio_delphinus **           
  |     +--+                                  
  |        |  +----- Schistodesmus_lampreyanus **
  |        +--+                               
--+           |  +-- Nodularia_douglasiae **     
  |           +--+                            
  |              +-- Tchangsinaia_piscicula **   
  |                                           
  |        +-------- Cristaria_plicata        
  |        |                                  
  |  +-----+     +-- Anemina_euscaphys        
  |  |     |  +--+                            
  |  |     |  |  +-- Anemina_arcaeformis      
  |  |     +--+                               
  +--+        |  +-- Sinanodonta_woodiana     
     |        +--+                            
     |           +-- Sinanodonta_lucida       
     |                                        
     |  +----------- Anodonta_anatina         
     +--+                                     
        |  +-------- Platynaias_compressa     
        +--+                                  
           |  +----- Pyganodon_grandis        
           +--+                               
              |  +-- Utterbackia_peninsularis 
              +--+                            
                 +-- Utterbackia_imbecillis   

Lopes-Lima et al., 2017 [Publ 3792], fig. 2

iT tT bi riS oT roT roS iD
7 7 5 5 26 12 2 19.0
                 +-- Lanceolaria_lanceolata   
     +-----------+                            
     |           +-- Lanceolaria_grayii       
     |                                        
  +--+           +-- Aculamprotula_tortuosa **   
  |  |  +--------+                            
  |  |  |        +-- Aculamprotula_coreana **    
  |  |  |                                     
  |  +--+        +-- Unio_delphinus **           
  |     |  +-----+                            
  |     |  |     +-- Unio_pictorum **            
  |     +--+                                  
  |        |  +----- Schistodesmus_lampreyanus **
  |        +--+                               
--+           |  +-- Tchangsinaia_piscicula **   
  |           +--+                            
  |              +-- Nodularia_douglasiae **     
  |                                           
  |     +----------- Anodonta_anatina         
  |  +--+                                     
  |  |  |  +-------- Platynaias_compressa     
  |  |  +--+                                  
  |  |     |  +----- Pyganodon_grandis        
  |  |     +--+                               
  +--+        |  +-- Utterbackia_peninsularis 
     |        +--+                            
     |           +-- Utterbackia_imbecillis   
     |                                        
     |     +-------- Cristaria_plicata        
     |     |                                  
     +-----+     +-- Sinanodonta_lucida       
           |  +--+                            
           |  |  +-- Sinanodonta_woodiana     
           +--+                               
              |  +-- Anemina_arcaeformis      
              +--+                            
                 +-- Anemina_euscaphys        

Wang et al., 2018 [Publ 3865], fig. 2

iT tT bi riS oT roT roS iD
7 7 5 4 25 11 3 17.0
     +-------------- Aculamprotula_tientsinensis **
     |                                          
  +--+           +-- Tchangsinaia_piscicula **     
  |  |  +--------+                              
  |  |  |        +-- Nodularia_douglasiae **       
  |  +--+                                       
  |     |  +-------- Unio_tumidus **               
  |     +--+                                    
  |        |  +----- Unio_crassus **               
  |        +--+                                 
--+           |  +-- Unio_pictorum **              
  |           +--+                              
  |              +-- Unio_delphinus **             
  |                                             
  |           +----- Acuticosta_chinensis       
  |  +--------+                                 
  |  |        |  +-- Lanceolaria_grayii         
  |  |        +--+                              
  |  |           +-- Lanceolaria_lanceolata     
  +--+                                          
     |        +----- Anodonta_anatina           
     |  +-----+                                 
     |  |     |  +-- Pyganodon_grandis          
     |  |     +--+                              
     +--+        +-- Platynaias_compressa       
        |                                       
        |  +-------- Cristaria_plicata          
        |  |                                    
        +--+     +-- Anemina_euscaphys          
           |  +--+                              
           |  |  +-- Anemina_arcaeformis        
           +--+                                 
              |  +-- Sinanodonta_woodiana       
              +--+                              
                 +-- Sinanodonta_lucida         

Lopes-Lima et al., 2020 [Publ 4080], fig. 12

iT tT bi riS oT roT roS iD
7 7 4 2 2 0 0 15.0
           +-- Nodularia_breviconcha **    
  +--------+                            
  |        +-- Nodularia_douglasiae **     
--+                                     
  |  +-------- Inversiunio_jokohamensis ** 
  |  |                                  
  +--+  +----- Inversiunio_verrucosus[1] **
     |  |                               
     +--+----- Inversiunio_verrucosus[2] **
        |                               
        |  +-- Inversiunio_reinianus **    
        +--+                            
           +-- Inversiunio_yanagawensis ** 

Lopes-Lima et al., 2020 [Publ 4080], fig. 5

iT tT bi riS oT roT roS iD
7 7 5 2 2 0 0 18.0
              +-- Nodularia_breviconcha **      
  +-----------+                              
  |           +-- Nodularia_douglasiae **       
--+                                          
  |  +----------- Aculamprotula_tortuosa **     
  +--+                                       
     |  +-------- Aculamprotula_tientsinensis **
     +--+                                    
        |  +----- Aculamprotula_fibrosa **      
        +--+                                 
           |  +-- Aculamprotula_scripta **      
           +--+                              
              +-- Aculamprotula_coreana **      

Lopes-Lima et al., 2021 [Publ 4217], fig. 6

iT tT bi riS oT roT roS iD
7 7 5 1 0 0 0 20.0
  +----------------- Unio_tumidus **      
--+                                    
  |  +-------------- Unio_pictorum **     
  +--+                                 
     |  +----------- Unio_sesirmensis **  
     +--+                              
        |  +-------- Unio_damascensis **  
        +--+                           
           |  +----- Unio_courtillieri ** 
           +--+                        
              |  +-- Unio_bruguierianus **
              +--+                     
                 +-- Unio_crassus **      

Dai et al., 2023 [Publ 4285], fig. 5

iT tT bi riS oT roT roS iD
7 7 5 7 46 10 4 18.0
  +-------------------- Lepidodesma_languilati     
  |                                                
  |     +-------------- Aculamprotula_tientsinensis **
  |     |                                          
  |  +--+  +----------- Unio_crassus **               
  |  |  |  |                                       
--+  |  +--+        +-- Pseudocuneopsis_capitata **   
  |  |     |  +-----+                              
  |  |     |  |     +-- Nodularia_douglasiae **       
  |  |     +--+                                    
  |  |        |  +----- Schistodesmus_lampreyanus **  
  |  |        +--+                                 
  +--+           |  +-- Cuneopsis_celtiformis **      
     |           +--+                              
     |              +-- Tchangsinaia_piscicula **     
     |                                             
     |              +-- Acuticosta_chinensis       
     |  +-----------+                              
     |  |           +-- Lanceolaria_grayii         
     |  |                                          
     +--+        +----- Cristaria_plicata          
        |  +-----+                                 
        |  |     |  +-- Anemina_arcaeformis        
        |  |     +--+                              
        +--+        +-- Sinanodonta_woodiana       
           |                                       
           |  +-------- Anodonta_cygnea            
           +--+                                    
              |  +----- Platynaias_compressa       
              +--+                                 
                 |  +-- Pyganodon_grandis          
                 +--+                              
                    +-- Utterbackia_imbecillis     

Reis et al., 2013 [Publ 3580], fig. 4

iT tT bi riS oT roT roS iD
6 6 4 1 8 3 1 19.0
           +----- Anodonta_cygnea         
  +--------+                              
  |        |  +-- Anodonta_anatina        
  |        +--+                           
--+           +-- Pseudanodonta_complanata
  |                                       
  |  +----------- Unio_tumidus **            
  |  |                                    
  +--+        +-- Unio_tumidiformis **       
     |  +-----+                           
     |  |     +-- Unio_crassus **            
     +--+                                 
        |  +----- Unio_delphinus **          
        +--+                              
           |  +-- Unio_mancus **             
           +--+                           
              +-- Unio_pictorum **           

Bolotov et al., 2018 [Publ 3826], fig. 2

iT tT bi riS oT roT roS iD
6 6 4 2 79 12 2 17.0
                    +----- Unio_tumidus **           
                 +--+                             
                 |  |  +-- Unio_crassus **           
                 |  +--+                          
  +--------------+     +-- Unio_pictorum **          
  |              |                                
  |              |  +----- Nodularia_douglasiae[1] **
  |              +--+                             
  |                 |  +-- Nodularia_nuxpersicae **  
  |                 +--+                          
--+                    +-- Nodularia_douglasiae[2] **
  |                                               
  |                    +-- Lanceolaria_gladiola   
  |  +-----------------+                          
  |  |                 +-- Lanceolaria_grayii     
  |  |                                            
  |  |           +-------- Cristaria_plicata      
  +--+  +--------+                                
     |  |        |  +----- Pletholophus_tenuis    
     |  |        +--+                             
     |  |           |  +-- Sinanodonta_jourdyi    
     +--+           +--+                          
        |              +-- Sinanodonta_lauta      
        |                                         
        |  +-------------- Anodonta_anatina       
        +--+                                      
           |  +----------- Pyganodon_grandis      
           +--+                                   
              |  +-------- Simpsonaias_ambigua    
              +--+                                
                 |  +----- Strophitus_undulatus   
                 +--+                             
                    |  +-- Alasmidonta_marginata  
                    +--+                          
                       +-- Lasmigona_costata      

Klishko et al., 2017 [Publ 3946], fig. 8

iT tT bi riS oT roT roS iD
6 6 4 2 0 0 0 15.0
           +-- Nodularia_douglasiae ** 
  +--------+                        
  |        +-- Nodularia_breviconcha **
--+                                 
  |  +-------- Unio_tumidus **         
  +--+                              
     |  +----- Unio_pictorum **        
     +--+                           
        |  +-- Unio_crassus **         
        +--+                        
           +-- Unio_courtillieri **    

Wu et al., 2018 [Publ 4143], fig. 2

iT tT bi riS oT roT roS iD
6 6 4 3 16 7 1 18.0
                    +-- Unio_pictorum **              
           +--------+                              
           |        +-- Nodularia_douglasiae **       
  +--------+                                       
  |        |  +-------- Aculamprotula_tortuosa **     
  |        +--+                                    
  |           |  +----- Aculamprotula_scripta **      
  |           +--+                                 
--+              |  +-- Aculamprotula_coreana **      
  |              +--+                              
  |                 +-- Aculamprotula_tientsinensis **
  |                                                
  |  +----------------- Sinanodonta_lucida         
  +--+                                             
     |  +-------------- Anemina_arcaeformis        
     +--+                                          
        |  +----------- Cristaria_plicata          
        +--+                                       
           |  +-------- Anodonta_anatina           
           +--+                                    
              |  +----- Pyganodon_grandis          
              +--+                                 
                 |  +-- Utterbackia_peninsularis   
                 +--+                              
                    +-- Utterbackia_imbecillis     

Choi et al., 2020 [Publ 4169], fig. 5

iT tT bi riS oT roT roS iD
6 6 4 1 3 0 0 18.0
  +----------- Nodularia_breviconcha **
  |                                 
--+        +-- Nodularia[1] **         
  |  +-----+                        
  |  |     +-- Nodularia[2] **         
  +--+                              
     |  +----- Nodularia_douglasiae ** 
     +--+                           
        |  +-- Nodularia_nipponensis **
        +--+                        
           +-- Nodularia_nuxpersicae **

Pavan-Kumar et al., 2022 [Publ 4263], fig. 4

iT tT bi riS oT roT roS iD
6 6 4 5 30 9 4 18.0
  +-------------------- Lepidodesma_languilati     
  |                                                
  |                 +-- Aculamprotula_tientsinensis **
  |        +--------+                              
  |        |        +-- Aculamprotula_tortuosa **     
--+  +-----+                                       
  |  |     |  +-------- Unio_pictorum **              
  |  |     +--+                                    
  |  |        |  +----- Schistodesmus_lampreyanus **  
  |  |        +--+                                 
  |  |           |  +-- Nodularia_douglasiae **       
  +--+           +--+                              
     |              +-- Tchangsinaia_piscicula **     
     |                                             
     |              +-- Lanceolaria_lanceolata     
     |  +-----------+                              
     |  |           +-- Lanceolaria_gladiola       
     +--+                                          
        |  +----------- Acuticosta_chinensis       
        |  |                                       
        +--+        +-- Anodonta_anatina           
           |  +-----+                              
           |  |     +-- Pyganodon_grandis          
           +--+                                    
              |  +----- Cristaria_plicata          
              +--+                                 
                 |  +-- Sinanodonta_lucida         
                 +--+                              
                    +-- Anemina_euscaphys          

Pavan-Kumar et al., 2022 [Publ 4263], fig. S5

iT tT bi riS oT roT roS iD
6 7 5 5 33 11 4 19.0
                 +-- Anodonta_anatina           
        +--------+                              
        |        +-- Pyganodon_grandis          
  +-----+                                       
  |     |  +-------- Cristaria_plicata          
  |     |  |                                    
  |     +--+     +-- Anemina_arcaeformis        
  |        |  +--+                              
  |        |  |  +-- Anemina_euscaphys          
  |        +--+                                 
  |           |  +-- Sinanodonta_lucida         
--+           +--+                              
  |              +-- Sinanodonta_woodiana       
  |                                             
  |           +----- Acuticosta_chinensis       
  |  +--------+                                 
  |  |        |  +-- Lanceolaria_lanceolata     
  |  |        +--+                              
  |  |           +-- Lanceolaria_gladiola       
  +--+                                          
     |        +----- Lepidodesma_languilati     
     |  +-----+                                 
     |  |     |  +-- Aculamprotula_tientsinensis **
     |  |     +--+                              
     +--+        +-- Aculamprotula_tortuosa **     
        |                                       
        |  +-------- Unio_pictorum **              
        +--+                                    
           |  +----- Schistodesmus_lampreyanus **  
           +--+                                 
              |  +-- Nodularia_douglasiae **       
              +--+                              
                 +-- Tchangsinaia_piscicula **     

Bashê, 2022 [Publ 4265], fig. 1

iT tT bi riS oT roT roS iD
6 6 4 1 1 0 0 21.0
  +-------------- Unio_pictorum[2] **  
--+                                 
  |  +----------- Unio_foucauldianus **
  +--+                              
     |  +-------- Unio_delphinus **    
     +--+                           
        |  +----- Unio_tigridis **     
        +--+                        
           |  +-- Unio_elongatulus **  
           +--+                     
              +-- Unio_pictorum[1] **  

Dai et al., 2024 [Publ 4358], fig. 3

iT tT bi riS oT roT roS iD
6 6 4 3 0 0 0 17.0
           +-- Schistodesmus_lampreyanus **    
  +--------+                                
  |        +-- Nodularia_douglasiae **         
--+                                         
  |  +-------- Pseudocuneopsis_capitata **     
  +--+                                      
     |  +----- Pseudocuneopsis_yangshuoensis **
     +--+                                   
        |  +-- Pseudocuneopsis_sichuanensis ** 
        +--+                                
           +-- Pseudocuneopsis_yemaoi **       

Prie et al., 2012 [Publ 3487], fig. 2

iT tT bi riS oT roT roS iD
5 5 3 1 1 0 0 18.0
  +----------- Unio_tumidus **    
--+                            
  |  +-------- Unio_crassus **    
  +--+                         
     |  +----- Unio_pictorum **   
     +--+                      
        |  +-- Unio_mancus **     
        +--+                   
           +-- Unio_elongatulus **

Wu et al., 2016 [Publ 3711], fig. 1

iT tT bi riS oT roT roS iD
5 5 3 3 16 7 1 17.0
                    +-- Unio_pictorum **              
              +-----+                              
              |     +-- Nodularia_douglasiae **       
  +-----------+                                    
  |           |  +----- Aculamprotula_tortuosa **     
  |           +--+                                 
  |              |  +-- Aculamprotula_coreana **      
--+              +--+                              
  |                 +-- Aculamprotula_tientsinensis **
  |                                                
  |  +----------------- Sinanodonta_lucida         
  +--+                                             
     |  +-------------- Anemina_arcaeformis        
     +--+                                          
        |  +----------- Cristaria_plicata          
        +--+                                       
           |  +-------- Anodonta_anatina           
           +--+                                    
              |  +----- Pyganodon_grandis          
              +--+                                 
                 |  +-- Utterbackia_peninsularis   
                 +--+                              
                    +-- Utterbackia_imbecillis     

Huang et al., 2019 [Publ 3843], fig. 3-M

iT tT bi riS oT roT roS iD
5 5 3 3 18 5 2 19.0
           +-------- Sinanodonta_woodiana     
  +--------+                                  
  |        |  +----- Anodonta_anatina         
  |        +--+                               
  |           |  +-- Pyganodon_grandis        
--+           +--+                            
  |              +-- Utterbackia_peninsularis 
  |                                           
  |  +-------------- Lanceolaria_lanceolata   
  +--+                                        
     |  +----------- Schistodesmus_lampreyanus **
     +--+                                     
        |  +-------- Aculamprotula_tortuosa **   
        +--+                                  
           |  +----- Unio_tumidus **             
           +--+                               
              |  +-- Unio_crassus **             
              +--+                            
                 +-- Unio_delphinus **           

Burzynski & Soroka, 2018 [Publ 3862], fig. 5a

iT tT bi riS oT roT roS iD
5 5 3 2 15 5 2 17.0
              +----- Sinanodonta_woodiana    
              |                              
  +-----------+----- Anodonta_anatina        
  |           |                              
  |           |  +-- Utterbackia_peninsularis
--+           +--+                           
  |              +-- Pyganodon_grandis       
  |                                          
  |  +-------------- Lanceolaria_lanceolata  
  +--+                                       
     |  +----------- Aculamprotula_tortuosa **  
     +--+                                    
        |  +-------- Unio_tumidus **            
        +--+                                 
           |  +----- Unio_crassus **            
           +--+                              
              |  +-- Unio_pictorum **           
              +--+                           
                 +-- Unio_delphinus **          

Burzynski & Soroka, 2018 [Publ 3862], fig. 5b

iT tT bi riS oT roT roS iD
5 5 3 2 5 5 2 17.0
  +----------------- Lanceolaria_lanceolata  
  |                                          
  |        +-------- Sinanodonta_woodiana    
--+  +-----+                                 
  |  |     |  +----- Anodonta_anatina        
  |  |     +--+                              
  |  |        |  +-- Utterbackia_peninsularis
  +--+        +--+                           
     |           +-- Pyganodon_grandis       
     |                                       
     |  +----------- Aculamprotula_tortuosa **  
     +--+                                    
        |  +-------- Unio_tumidus **            
        +--+                                 
           |  +----- Unio_crassus **            
           +--+                              
              |  +-- Unio_pictorum **           
              +--+                           
                 +-- Unio_delphinus **          

Bolotov et al., 2020 [Publ 4084], fig. 1

iT tT bi riS oT roT roS iD
5 5 3 3 31 14 2 16.0
              +----------- Middendorffinaia_mongolica **
  +-----------+                                      
  |           |  +-------- Nodularia_douglasiae **      
  |           +--+                                   
  |              |  +----- Unio_tumidus **              
  |              +--+                                
  |                 |  +-- Unio_pictorum **             
  |                 +--+                             
--+                    +-- Unio_crassus **              
  |                                                  
  |                    +-- Lanceolaria_grayii        
  |  +-----------------+                             
  |  |                 +-- Lanceolaria               
  |  |                                               
  |  |              +----- Anodonta_anatina          
  +--+  +-----------+                                
     |  |           |  +-- Pseudanodonta_complanata  
     |  |           +--+                             
     |  |              +-- Anodonta_cygnea           
     |  |                                            
     +--+        +-------- Amuranodonta_kijaensis    
        |  +-----+                                   
        |  |     |  +----- Anemina_arcaeformis       
        |  |     +--+                                
        |  |        |  +-- Buldowskia_suifunica      
        +--+        +--+                             
           |           +-- Buldowskia_shadini        
           |                                         
           |  +----------- Cristaria_plicata         
           +--+                                      
              |  +-------- Beringiana_beringiana     
              +--+                                   
                 |  +----- Sinanodonta_schrenkii     
                 +--+                                
                    |  +-- Sinanodonta_lauta         
                    +--+                             
                       +-- Sinanodonta_woodiana      

Wu et al., 2020 [Publ 4121], fig. 2

iT tT bi riS oT roT roS iD
5 5 3 2 29 0 0 18.0
  +----------- Unio_pictorum **              
--+                                       
  |  +-------- Aculamprotula_tortuosa **     
  +--+                                    
     |  +----- Aculamprotula_polysticta **   
     +--+                                 
        |  +-- Aculamprotula_fibrosa **      
        +--+                              
           +-- Aculamprotula_tientsinensis **

Wu et al., 2020 [Publ 4121], fig. S1

iT tT bi riS oT roT roS iD
5 5 3 2 29 0 0 18.0
  +----------- Unio_pictorum **              
--+                                       
  |  +-------- Aculamprotula_tortuosa **     
  +--+                                    
     |  +----- Aculamprotula_polysticta **   
     +--+                                 
        |  +-- Aculamprotula_tientsinensis **
        +--+                              
           +-- Aculamprotula_fibrosa **      

Wu et al., 2020 [Publ 4147], fig. 4

iT tT bi riS oT roT roS iD
5 5 3 2 9 0 0 18.0
  +----------- Unio_pictorum **              
--+                                       
  |  +-------- Aculamprotula_tortuosa **     
  +--+                                    
     |  +----- Aculamprotula_scripta **      
     +--+                                 
        |  +-- Aculamprotula_tientsinensis **
        +--+                              
           +-- Aculamprotula_fibrosa **      

Guerra et al., 2019 [Publ 4168], fig. 1

iT tT bi riS oT roT roS iD
5 5 3 3 36 8 2 17.0
     +-------------- Lanceolaria_lanceolata   
  +--+                                        
  |  |  +----------- Aculamprotula_tortuosa **   
  |  +--+                                     
  |     |  +-------- Schistodesmus_lampreyanus **
  |     +--+                                  
  |        |  +----- Unio_tumidus **             
--+        +--+                               
  |           |  +-- Unio_crassus **             
  |           +--+                            
  |              +-- Unio_delphinus **           
  |                                           
  |  +-------------- Sinanodonta_woodiana     
  |  |                                        
  +--+  +----------- Anodonta_anatina         
     |  |                                     
     +--+        +-- Platynaias_compressa     
        |  +-----+                            
        |  |     +-- Platynaias_subviridis    
        +--+                                  
           |  +----- Pyganodon_grandis        
           +--+                               
              |  +-- Utterbackia_peninsularis 
              +--+                            
                 +-- Utterbackia_imbecillis   

Guerra et al., 2019 [Publ 4168], fig. 1-M

iT tT bi riS oT roT roS iD
5 5 3 3 17 5 2 19.0
           +-------- Sinanodonta_woodiana     
  +--------+                                  
  |        |  +----- Anodonta_anatina         
  |        +--+                               
  |           |  +-- Utterbackia_peninsularis 
--+           +--+                            
  |              +-- Pyganodon_grandis        
  |                                           
  |  +-------------- Lanceolaria_lanceolata   
  +--+                                        
     |  +----------- Schistodesmus_lampreyanus **
     +--+                                     
        |  +-------- Aculamprotula_tortuosa **   
        +--+                                  
           |  +----- Unio_tumidus **             
           +--+                               
              |  +-- Unio_crassus **             
              +--+                            
                 +-- Unio_delphinus **           

Guerra et al., 2019 [Publ 4168], fig. 2

iT tT bi riS oT roT roS iD
5 13 11 3 36 8 2 25.0
  +----------------------- Aculamprotula_tortuosa **   
  |                                                 
  |              +-------- Schistodesmus_lampreyanus **
--+  +-----------+                                  
  |  |           |  +----- Unio_tumidus **             
  |  |           +--+                               
  |  |              |  +-- Unio_crassus **             
  +--+              +--+                            
     |                 +-- Unio_delphinus **           
     |                                              
     |  +----------------- Lanceolaria_lanceolata   
     |  |                                           
     +--+  +-------------- Sinanodonta_woodiana     
        |  |                                        
        +--+  +----------- Anodonta_anatina         
           |  |                                     
           +--+        +-- Platynaias_compressa     
              |  +-----+                            
              |  |     +-- Platynaias_subviridis    
              +--+                                  
                 |  +----- Pyganodon_grandis        
                 +--+                               
                    |  +-- Utterbackia_peninsularis 
                    +--+                            
                       +-- Utterbackia_imbecillis   

Guerra et al., 2019 [Publ 4168], fig. 2-M

iT tT bi riS oT roT roS iD
5 5 3 3 17 5 2 17.0
                 +-- Utterbackia_peninsularis 
              +--+                            
              |  +-- Pyganodon_grandis        
  +-----------+                               
  |           |  +-- Anodonta_anatina         
  |           +--+                            
--+              +-- Sinanodonta_woodiana     
  |                                           
  |  +-------------- Lanceolaria_lanceolata   
  +--+                                        
     |  +----------- Aculamprotula_tortuosa **   
     +--+                                     
        |  +-------- Schistodesmus_lampreyanus **
        +--+                                  
           |  +----- Unio_tumidus **             
           +--+                               
              |  +-- Unio_crassus **             
              +--+                            
                 +-- Unio_delphinus **           

Choi et al., 2020 [Publ 4169], fig. 3a

iT tT bi riS oT roT roS iD
5 5 3 3 0 0 0 14.0
        +-- Cuneopsis_heudei **     
  +-----+                        
  |     +-- Unio_pictorum **        
--+                              
  |  +----- Nodularia_breviconcha **
  +--+                           
     |  +-- Nodularia_douglasiae ** 
     +--+                        
        +-- Nodularia_nipponensis **

Mabuchi et al., 2021 [Publ 4233], fig. 1

iT tT bi riS oT roT roS iD
5 5 3 2 0 0 0 16.0
  +-------- Tchangsinaia_piscicula ** 
  |                                
--+     +-- Nodularia_douglasiae[1] **
  |  +--+                          
  |  |  +-- Nodularia_nipponensis **  
  +--+                             
     |  +-- Nodularia_douglasiae[2] **
     +--+                          
        +-- Nodularia_breviconcha **  

Pfarrer et al., 2022 [Publ 4305], fig. 2-5

iT tT bi riS oT roT roS iD
5 5 3 1 7 5 1 18.0
     +----------- Unio_tumidus **        
  +--+                                
  |  |  +-------- Unio_crassus **        
  |  +--+                             
  |     |  +----- Unio_elongatulus **    
  |     +--+                          
  |        |  +-- Unio_mancus **         
--+        +--+                       
  |           +-- Unio_pictorum **       
  |                                   
  |           +-- Cristaria_plicata   
  |     +-----+                       
  |     |     +-- Sinanodonta_woodiana
  +-----+                             
        |  +----- Anodonta_anatina    
        +--+                          
           |  +-- Anodonta_exulcerata 
           +--+                       
              +-- Anodonta_cygnea     

Nagel et al., 1998 [Publ 1946], fig. 1

iT tT bi riS oT roT roS iD
4 4 2 1 5 3 1 17.0
        +----- Pseudanodonta_complanata
  +-----+                              
  |     |  +-- Anodonta_anatina        
  |     +--+                           
--+        +-- Anodonta_cygnea         
  |                                    
  |  +-------- Unio_crassus **            
  +--+                                 
     |  +----- Unio_tumidus **            
     +--+                              
        |  +-- Unio_pictorum **           
        +--+                           
           +-- Unio_mancus **             

Nagel et al., 1998 [Publ 1946], fig. 8

iT tT bi riS oT roT roS iD
4 4 1 1 0 0 0 17.0
  +----- Unio_tumidus ** 
  |                   
--+  +-- Unio_pictorum **
  |  |                
  +--+-- Unio_crassus ** 
     |                
     +-- Unio_mancus **  

Soroka, 2005 [Publ 2726], fig. 3

iT tT bi riS oT roT roS iD
4 4 2 1 1 0 0 17.0
  +-------- Unio_tumidus ** 
--+                      
  |  +----- Unio_crassus ** 
  +--+                   
     |  +-- Unio_caffer **  
     +--+                
        +-- Unio_pictorum **

Soroka, 2010 [Publ 3404], fig. 3a

iT tT bi riS oT roT roS iD
4 4 2 1 5 5 1 17.0
              +-- Sinanodonta_woodiana    
  +-----------+                           
  |           +-- Sinanodonta_lauta       
  |                                       
--+        +----- Anodonta_anatina        
  |  +-----+                              
  |  |     |  +-- Anodonta_cygnea         
  |  |     +--+                           
  +--+        +-- Pseudanodonta_complanata
     |                                    
     |  +-------- Unio_tumidus **            
     +--+                                 
        |  +----- Unio_crassus **            
        +--+                              
           |  +-- Unio_pictorum **           
           +--+                           
              +-- Unio_mancus **             

Soroka, 2010 [Publ 3404], fig. 3b

iT tT bi riS oT roT roS iD
4 4 2 1 5 5 1 17.0
  +-------------- Anodonta_anatina        
  |                                       
  +-------------- Anodonta_cygnea         
  |                                       
--+-------------- Pseudanodonta_complanata
  |                                       
  |           +-- Sinanodonta_woodiana    
  |  +--------+                           
  |  |        +-- Sinanodonta_lauta       
  +--+                                    
     |  +-------- Unio_tumidus **            
     +--+                                 
        |  +----- Unio_crassus **            
        +--+                              
           |  +-- Unio_pictorum **           
           +--+                           
              +-- Unio_mancus **             

Xue et al., 2016 [Publ 3659], fig. 1

iT tT bi riS oT roT roS iD
4 4 2 4 12 8 2 16.0
              +-- Cristaria_plicata        
        +-----+                            
        |     +-- Platynaias_compressa     
        |                                  
  +-----+     +-- Anemina_arcaeformis      
  |     |  +--+                            
  |     |  |  +-- Anemina_euscaphys        
  |     +--+                               
  |        |  +-- Sinanodonta_lucida       
--+        +--+                            
  |           +-- Sinanodonta_woodiana     
  |                                        
  |           +-- Lanceolaria_lanceolata   
  |  +--------+                            
  |  |        +-- Lanceolaria_grayii       
  +--+                                     
     |  +-------- Aculamprotula_tortuosa **   
     +--+                                  
        |  +----- Schistodesmus_lampreyanus **
        +--+                               
           |  +-- Nodularia_douglasiae **     
           +--+                            
              +-- Unio_pictorum **            

Jadhav & Jamkhedkar, 2009 [Publ 3697], fig. 11

iT tT bi riS oT roT roS iD
4 10 7 2 22 6 1 23.0
  +-------------------- Nodularia_douglasiae ** 
  |                                          
--+  +----------------- Anodonta_cygnea      
  |  |                                       
  +--+  +-------------- Unio_pictorum[2] **     
     |  |                                    
     +--+           +-- Unio_pictorum[1] **     
        |  +--------+                        
        |  |        +-- Unio_caffer **          
        +--+                                 
           |  +-------- Anodonta             
           |  |                              
           +--+  +----- Lasmigona_costata    
              |  |                           
              +--+  +-- Strophitus_undulatus 
                 |  |                        
                 +--+-- Alasmidonta_marginata
                    |                        
                    +-- Pyganodon_grandis    

Zhou et al., 2017 [Publ 3788], fig. 2

iT tT bi riS oT roT roS iD
4 4 2 3 13 3 2 16.0
  +-------- Lanceolaria_lanceolata     
  |                                    
--+     +-- Nodularia_douglasiae **       
  |  +--+                              
  |  |  +-- Tchangsinaia_piscicula **     
  +--+                                 
     |  +-- Aculamprotula_tientsinensis **
     +--+                              
        +-- Aculamprotula_tortuosa **     

Huang et al., 2018 [Publ 3828], fig. 6-M

iT tT bi riS oT roT roS iD
4 4 2 2 13 4 1 18.0
     +-------- Sinanodonta_woodiana    
  +--+                                 
  |  |  +----- Anodonta_anatina        
  |  +--+                              
  |     |  +-- Pyganodon_grandis       
--+     +--+                           
  |        +-- Utterbackia_peninsularis
  |                                    
  |  +-------- Aculamprotula_tortuosa **  
  +--+                                 
     |  +----- Unio_tumidus **            
     +--+                              
        |  +-- Unio_crassus **            
        +--+                           
           +-- Unio_delphinus **          

Huang et al., 2002 [Publ 2657], fig. 1

iT tT bi riS oT roT roS iD
3 7 5 3 31 11 4 20.0
                 +-- Sinanodonta_woodiana     
     +-----------+                            
     |           +-- Cristaria_plicata        
  +--+                                        
  |  |  +----------- Utterbackiana_couperiana 
  |  +--+                                     
  |     |  +-------- Utterbackia_imbecillis   
  |     +--+                                  
  |        |  +----- Strophitus_radiatus      
  |        +--+                               
  |           |  +-- Alasmidonta_triangulata  
--+           +--+                            
  |              +-- Lasmigona_costata        
  |                                           
  |           +----- Acuticosta_ovata         
  |        +--+                               
  |        |  |  +-- Nodularia_douglasiae **     
  |        |  +--+                            
  |        |     +-- Tchangsinaia_piscicula **   
  +--------+                                  
           |     +-- Lanceolaria_grayii       
           |  +--+                            
           |  |  +-- Lanceolaria_lanceolata   
           +--+                               
              |  +-- Schistodesmus_lampreyanus **
              +--+                            
                 +-- Lepidodesma_languilati   

Huang et al., 2002 [Publ 2657], fig. 2

iT tT bi riS oT roT roS iD
3 14 5 3 31 11 4 27.0
  +-------- Strophitus_radiatus      
  |                                  
  +-------- Schistodesmus_lampreyanus **
  |                                  
  +-------- Lepidodesma_languilati   
  |                                  
  +-------- Acuticosta_ovata         
  |                                  
  |     +-- Sinanodonta_woodiana     
  +-----+                            
  |     +-- Cristaria_plicata        
--+                                  
  |     +-- Lanceolaria_grayii       
  +-----+                            
  |     +-- Lanceolaria_lanceolata   
  |                                  
  |     +-- Nodularia_douglasiae **     
  +-----+                            
  |     +-- Tchangsinaia_piscicula **   
  |                                  
  |  +----- Utterbackia_imbecillis   
  |  |                               
  +--+----- Utterbackiana_couperiana 
     |                               
     |  +-- Alasmidonta_triangulata  
     +--+                            
        +-- Lasmigona_costata        

Manendo et al., 2008 [Publ 2669], fig. 1

iT tT bi riS oT roT roS iD
3 3 1 1 10 5 1 16.0
        +----- Unio_crassus **            
  +-----+                              
  |     |  +-- Unio_pictorum **           
  |     +--+                           
  |        +-- Unio_tumidus **            
--+                                    
  |        +-- Anodonta_cygnea         
  |  +-----+                           
  |  |     +-- Anodonta_anatina        
  +--+                                 
     |  +----- Pseudanodonta_complanata
     +--+                              
        |  +-- Platynaias_subviridis   
        +--+                           
           +-- Pyganodon_grandis       

Källersjö et al., 2005 [Publ 2684], fig. 1A

iT tT bi riS oT roT roS iD
3 3 0 1 4 3 1 16.0
  +-- Unio_crassus ** 
  |                
--+-- Unio_pictorum **
  |                
  +-- Unio_tumidus ** 

Källersjö et al., 2005 [Publ 2684], fig. 1B-F

iT tT bi riS oT roT roS iD
3 3 1 1 4 3 1 16.0
  +----- Unio_tumidus ** 
--+                   
  |  +-- Unio_crassus ** 
  +--+                
     +-- Unio_pictorum **

Soroka, 2010 [Publ 3404], fig. 2

iT tT bi riS oT roT roS iD
3 3 0 1 4 4 1 16.0
  +----------- Sinanodonta_woodiana    
  |                                    
  |        +-- Unio_tumidus **            
--+        |                           
  |  +-----+-- Unio_pictorum **           
  |  |     |                           
  +--+     +-- Unio_crassus **            
     |                                 
     |  +----- Anodonta_cygnea         
     +--+                              
        |  +-- Anodonta_anatina        
        +--+                           
           +-- Pseudanodonta_complanata

Soroka, 2010 [Publ 3404], fig. 2-M

iT tT bi riS oT roT roS iD
3 3 1 1 2 2 1 16.0
  +----------- Sinanodonta_woodiana
--+                                
  |  +-------- Anodonta_anatina    
  +--+                             
     |  +----- Unio_crassus **        
     +--+                          
        |  +-- Unio_tumidus **        
        +--+                       
           +-- Unio_pictorum **       

Soroka, 2010 [Publ 3404], fig. 3a-M

iT tT bi riS oT roT roS iD
3 3 1 1 3 3 1 16.0
  +----------- Anodonta_anatina    
  |                                
--+        +-- Sinanodonta_woodiana
  |  +-----+                       
  |  |     +-- Sinanodonta_lauta   
  +--+                             
     |  +----- Unio_crassus **        
     +--+                          
        |  +-- Unio_tumidus **        
        +--+                       
           +-- Unio_pictorum **       

Soroka, 2010 [Publ 3404], fig. 3b-M

iT tT bi riS oT roT roS iD
3 3 1 1 3 3 1 16.0
  +----------- Anodonta_anatina    
  |                                
--+        +-- Sinanodonta_lauta   
  |  +-----+                       
  |  |     +-- Sinanodonta_woodiana
  +--+                             
     |  +----- Unio_tumidus **        
     +--+                          
        |  +-- Unio_crassus **        
        +--+                       
           +-- Unio_pictorum **       

Soroka, 2010 [Publ 3404], fig. 4

iT tT bi riS oT roT roS iD
3 3 1 1 4 4 1 16.0
        +----- Sinanodonta_woodiana    
  +-----+                              
  |     |  +-- Anodonta_cygnea         
  |     +--+                           
--+        +-- Pseudanodonta_complanata
  |                                    
  |  +-------- Anodonta_anatina        
  +--+                                 
     |  +----- Unio_tumidus **            
     +--+                              
        |  +-- Unio_pictorum **           
        +--+                           
           +-- Unio_crassus **            

Soroka, 2010 [Publ 3404], fig. 5a

iT tT bi riS oT roT roS iD
3 3 1 1 5 5 1 16.0
        +----- Unio_crassus **            
  +-----+                              
  |     |  +-- Unio_pictorum **           
  |     +--+                           
--+        +-- Unio_tumidus **            
  |                                    
  |  +-------- Anodonta_anatina        
  |  |                                 
  +--+     +-- Anodonta_cygnea         
     |  +--+                           
     |  |  +-- Pseudanodonta_complanata
     +--+                              
        |  +-- Sinanodonta_woodiana    
        +--+                           
           +-- Sinanodonta_schrenkii   

Soroka, 2010 [Publ 3404], fig. 5b

iT tT bi riS oT roT roS iD
3 3 1 1 5 5 1 16.0
  +----------- Anodonta_anatina        
  |                                    
  |        +-- Anodonta_cygnea         
--+  +-----+                           
  |  |     +-- Pseudanodonta_complanata
  |  |                                 
  |  |     +-- Sinanodonta_woodiana    
  +--+-----+                           
     |     +-- Sinanodonta_schrenkii   
     |                                 
     |  +----- Unio_tumidus **            
     +--+                              
        |  +-- Unio_pictorum **           
        +--+                           
           +-- Unio_crassus **            

Soroka, 2010 [Publ 3404], fig. 8a

iT tT bi riS oT roT roS iD
3 3 1 1 3 3 1 16.0
     +----- Unio_tumidus **        
  +--+                          
  |  |  +-- Unio_pictorum **       
  |  +--+                       
--+     +-- Unio_crassus **        
  |                             
  |  +----- Sinanodonta_woodiana
  +--+                          
     |  +-- Anodonta_anatina    
     +--+                       
        +-- Anodonta_cygnea     

Soroka, 2010 [Publ 3404], fig. 8b

iT tT bi riS oT roT roS iD
3 3 1 1 3 3 1 16.0
  +----------- Sinanodonta_woodiana
  |                                
--+        +-- Anodonta_anatina    
  |  +-----+                       
  |  |     +-- Anodonta_cygnea     
  +--+                             
     |  +----- Unio_tumidus **        
     +--+                          
        |  +-- Unio_pictorum **       
        +--+                       
           +-- Unio_crassus **        

Ouyang et al., 2011 [Publ 3408], fig. 1

iT tT bi riS oT roT roS iD
3 6 3 3 10 6 3 18.0
        +----- Anemina_arcaeformis   
  +-----+                            
  |     |  +-- Sinanodonta_elliptica 
  |     +--+                         
  |        +-- Sinanodonta_pacifica  
--+                                  
  |        +-- Nodularia_douglasiae **  
  |  +-----+                         
  |  |     +-- Cuneopsis_heudei **      
  +--+                               
     |  +----- Lanceolaria_grayii    
     |  |                            
     +--+----- Acuticosta_chinensis  
        |                            
        |  +-- Lanceolaria_gladiola  
        +--+                         
           +-- Aculamprotula_tortuosa **

Zieritz et al., 2012 [Publ 3673], fig. 4

iT tT bi riS oT roT roS iD
3 3 1 1 5 4 1 16.0
  +----- Unio_crassus ** 
--+                   
  |  +-- Unio_tumidus ** 
  +--+                
     +-- Unio_pictorum **

Plazzi et al., 2011 [Publ 3676], fig. 5

iT tT bi riS oT roT roS iD
3 3 1 1 15 6 2 16.0
  +----- Unio_tumidus ** 
--+                   
  |  +-- Unio_crassus ** 
  +--+                
     +-- Unio_pictorum **

Burzynski et al., 2017 [Publ 3774], fig. 1 bottom

iT tT bi riS oT roT roS iD
3 3 1 2 10 3 1 17.0
     +----- Aculamprotula_tortuosa **  
  +--+                              
  |  |  +-- Unio_crassus **            
  |  +--+                           
--+     +-- Unio_tumidus **            
  |                                 
  |  +----- Anodonta_anatina        
  +--+                              
     |  +-- Utterbackia_peninsularis
     +--+                           
        +-- Pyganodon_grandis       

Burzynski et al., 2017 [Publ 3774], fig. 1 top

iT tT bi riS oT roT roS iD
3 3 1 2 10 3 1 17.0
     +----- Aculamprotula_tortuosa **  
  +--+                              
  |  |  +-- Unio_crassus **            
  |  +--+                           
--+     +-- Unio_tumidus **            
  |                                 
  |  +----- Anodonta_anatina        
  +--+                              
     |  +-- Utterbackia_peninsularis
     +--+                           
        +-- Pyganodon_grandis       

Wen et al., 2017 [Publ 3795], fig. 3

iT tT bi riS oT roT roS iD
3 4 2 2 13 4 2 18.0
     +----- Anodonta_anatina      
  +--+                            
  |  |  +-- Sinanodonta_woodiana  
  |  +--+                         
  |     +-- Cristaria_plicata     
--+                               
  |     +-- Aculamprotula_coreana ** 
  |  +--+                         
  |  |  +-- Aculamprotula_tortuosa **
  +--+                            
     |  +-- Lepidodesma_languilati
     +--+                         
        +-- Unio_pictorum **         

Araujo et al., 2017 [Publ 3801], fig. 2

iT tT bi riS oT roT roS iD
3 4 2 3 11 1 1 19.0
     +-- Schistodesmus_lampreyanus **
  +--+                            
  |  +-- Aculamprotula_tortuosa **   
--+                               
  |  +-- Anodonta_anatina         
  +--+                            
     +-- Unio_delphinus **           

Froufe et al., 2017 [Publ 3802], fig. 7

iT tT bi riS oT roT roS iD
3 3 1 1 7 4 1 18.0
        +----- Unio_mancus **         
  +-----+                          
  |     |  +-- Unio_elongatulus **    
  |     +--+                       
--+        +-- Unio_pictorum **       
  |                                
  |  +-------- Sinanodonta_woodiana
  +--+                             
     |  +----- Anodonta_anatina    
     +--+                          
        |  +-- Anodonta_exulcerata 
        +--+                       
           +-- Anodonta_cygnea     

Dyduch-Falniowska & Koziol, 1989 [Publ 3814], fig. 5, 7

iT tT bi riS oT roT roS iD
3 3 1 1 2 2 1 16.0
        +-- Pseudanodonta_complanata
  +-----+                           
  |     +-- Anodonta_anatina        
--+                                 
  |  +----- Unio_pictorum **           
  +--+                              
     |  +-- Unio_tumidus **            
     +--+                           
        +-- Unio_crassus **            

Bogatov et al., 2018 [Publ 3859], fig. 1

iT tT bi riS oT roT roS iD
3 5 3 1 3 2 1 18.0
  +-------- Unio_tumidus **            
  |                                 
--+     +-- Anodonta_anatina        
  |  +--+                           
  |  |  +-- Pseudanodonta_complanata
  +--+                              
     |  +-- Unio_crassus **            
     +--+                           
        +-- Unio_pictorum **           

Bogatov et al., 2018 [Publ 3859], fig. 2

iT tT bi riS oT roT roS iD
3 5 3 1 3 2 1 18.0
  +----------- Unio_crassus **            
--+                                    
  |  +-------- Unio_pictorum **           
  +--+                                 
     |  +----- Unio_tumidus **            
     +--+                              
        |  +-- Anodonta_anatina        
        +--+                           
           +-- Pseudanodonta_complanata

Froufe et al., 2019 [Publ 3936], fig. 3-M

iT tT bi riS oT roT roS iD
3 3 1 1 16 4 1 18.0
        +----- Unio_tumidus **            
  +-----+                              
  |     |  +-- Unio_crassus **            
  |     +--+                           
--+        +-- Unio_delphinus **          
  |                                    
  |  +-------- Sinanodonta_woodiana    
  +--+                                 
     |  +----- Anodonta_anatina        
     +--+                              
        |  +-- Utterbackia_peninsularis
        +--+                           
           +-- Pyganodon_grandis       

Froufe et al., 2019 [Publ 3936], fig. 4

iT tT bi riS oT roT roS iD
3 3 1 1 16 4 1 18.0
        +----- Unio_tumidus **            
  +-----+                              
  |     |  +-- Unio_crassus **            
  |     +--+                           
--+        +-- Unio_delphinus **          
  |                                    
  |  +-------- Sinanodonta_woodiana    
  +--+                                 
     |  +----- Anodonta_anatina        
     +--+                              
        |  +-- Utterbackia_peninsularis
        +--+                           
           +-- Pyganodon_grandis       

Pfeiffer et al., 2019 [Publ 3937], fig. 4-5

iT tT bi riS oT roT roS iD
3 3 1 3 54 5 2 17.0
  +----------- Lanceolaria_lanceolata
  |                                  
  |     +----- Unio_gibbus **           
--+  +--+                            
  |  |  |  +-- Nodularia_douglasiae **  
  |  |  +--+                         
  |  |     +-- Tchangsinaia_piscicula **
  +--+                               
     |     +-- Cristaria_plicata     
     |  +--+                         
     |  |  +-- Sinanodonta_woodiana  
     +--+                            
        |  +-- Anodonta_cygnea       
        +--+                         
           +-- Pyganodon_grandis     

Pfeiffer et al., 2019 [Publ 3937], fig. 6

iT tT bi riS oT roT roS iD
3 3 1 3 54 5 2 17.0
     +-------- Lanceolaria_lanceolata
  +--+                               
  |  |  +----- Unio_gibbus **           
  |  +--+                            
  |     |  +-- Tchangsinaia_piscicula **
  |     +--+                         
--+        +-- Nodularia_douglasiae **  
  |                                  
  |        +-- Cristaria_plicata     
  |     +--+                         
  |     |  +-- Sinanodonta_woodiana  
  +-----+                            
        |  +-- Anodonta_cygnea       
        +--+                         
           +-- Pyganodon_grandis     

Chen et al., 2020 [Publ 4170], fig. 1

iT tT bi riS oT roT roS iD
3 4 2 2 31 12 2 18.0
        +-------- Schistodesmus_lampreyanus **
  +-----+                                  
  |     |  +----- Lepidodesma_languilati   
  |     +--+                               
  |        |  +-- Aculamprotula_coreana **    
  |        +--+                            
--+           +-- Aculamprotula_tortuosa **   
  |                                        
  |        +----- Platynaias_compressa     
  |  +-----+                               
  |  |     |  +-- Utterbackia_imbecillis   
  |  |     +--+                            
  +--+        +-- Pyganodon_grandis        
     |                                     
     |  +-------- Sinanodonta_lucida       
     +--+                                  
        |  +----- Cristaria_plicata        
        +--+                               
           |  +-- Anodonta_cygnea          
           +--+                            
              +-- Anodonta_anatina         

Wu et al., 2021 [Publ 4215], fig. 3

iT tT bi riS oT roT roS iD
3 3 1 2 21 1 1 16.0
  +-------- Acuticosta_chinensis    
--+                                 
  |  +----- Nodularia_douglasiae **    
  +--+                              
     |  +-- Inversiunio_jokohamensis **
     +--+                           
        +-- Inversiunio_reinianus **   

Sonowal et al., 2021 [Publ 4221], fig. 3a

iT tT bi riS oT roT roS iD
3 4 0 2 12 1 1 18.0
  +-- Anodonta_cygnea            
  |                              
  +-- Unio_ravoisieri **            
--+                              
  +-- Unio_pictorum **              
  |                              
  +-- Aculamprotula_tientsinensis **

Sonowal et al., 2021 [Publ 4221], fig. 3b

iT tT bi riS oT roT roS iD
3 9 0 2 12 1 1 23.0
  +-- Aculamprotula_tientsinensis **
  |                              
  +-- Lamprotula_leaii           
  |                              
  +-- Lampsilis_cardium          
  |                              
  +-- Potomida_littoralis        
  |                              
--+-- Unio_pictorum **              
  |                              
  +-- Anodonta_cygnea            
  |                              
  +-- Unio_ravoisieri **            
  |                              
  +-- Lamellidens_mainwaringi    
  |                              
  +-- Elliptio_complanata        

Sonowal et al., 2021 [Publ 4221], fig. 4

iT tT bi riS oT roT roS iD
3 5 3 2 13 1 1 19.0
  +----------- Aculamprotula_tientsinensis **
--+                                       
  |  +-------- Unio_pictorum **              
  +--+                                    
     |  +----- Anodonta_cygnea            
     +--+                                 
        |  +-- Unio_ravoisieri **            
        +--+                              
           +-- Lamellidens_corrianus      

Zieritz et al., 2021 [Publ 4231], fig. 3-M, 4

iT tT bi riS oT roT roS iD
3 3 1 1 20 4 1 18.0
        +----- Unio_tumidus **            
  +-----+                              
  |     |  +-- Unio_delphinus **          
  |     +--+                           
--+        +-- Unio_crassus **            
  |                                    
  |  +-------- Sinanodonta_woodiana    
  +--+                                 
     |  +----- Anodonta_anatina        
     +--+                              
        |  +-- Utterbackia_peninsularis
        +--+                           
           +-- Pyganodon_grandis       

Graf & Cummings, 2006 [Publ 1968], fig. 2

iT tT bi riS oT roT roS iD
2 2 0 1 45 4 1 --
           +-- Unio_pictorum **        
  +--------+                        
  |        +-- Unio_caffer **          
--+                                 
  |  +-------- Lasmigona_costata    
  +--+                              
     |  +----- Alasmidonta_marginata
     +--+                           
        |  +-- Pyganodon_grandis    
        +--+                        
           +-- Strophitus_undulatus 

Graf & Ó Foighil, 2000 [Publ 2648], fig. 2A

iT tT bi riS oT roT roS iD
2 2 0 1 28 6 1 --
  +-- Unio_pictorum **
--+                
  +-- Unio_caffer **  

Graf & Ó Foighil, 2000 [Publ 2648], fig. 2B

iT tT bi riS oT roT roS iD
2 2 0 1 28 6 1 --
  +-- Unio_pictorum **
--+                
  +-- Unio_caffer **  

Bogan & Hoeh, 2000 [Publ 2650], fig. 1

iT tT bi riS oT roT roS iD
2 2 0 1 32 3 1 --
  +-- Unio_pictorum **
--+                
  +-- Unio_tumidus ** 

Graf, 2002 [Publ 2651], fig. 2

iT tT bi riS oT roT roS iD
2 2 0 1 16 2 1 --
     +-- Unio_pictorum **        
  +--+                        
  |  +-- Unio_caffer **          
--+                           
  |  +-- Alasmidonta_marginata
  +--+                        
     +-- Pyganodon_grandis    

Hoeh et al., 2001 [Publ 2652], fig. 14.2

iT tT bi riS oT roT roS iD
2 2 0 1 29 3 1 --
  +-- Unio_pictorum **
--+                
  +-- Unio_tumidus ** 

Hoeh et al., 2001 [Publ 2652], fig. 14.4

iT tT bi riS oT roT roS iD
2 5 0 1 29 3 1 20.0
  +-------- Unio_pictorum **       
  |                             
--+-------- Unio_tumidus **        
  |                             
  |  +----- Anodonta_cygnea     
  +--+                          
     |  +-- Strophitus_undulatus
     +--+                       
        +-- Pyganodon_grandis   

Hoeh et al., 2002 [Publ 2653], fig. 2

iT tT bi riS oT roT roS iD
2 5 0 1 29 3 1 20.0
  +----------- Unio_pictorum **       
--+                                
  |  +-------- Unio_tumidus **        
  +--+                             
     |  +----- Anodonta_cygnea     
     +--+                          
        |  +-- Pyganodon_grandis   
        +--+                       
           +-- Strophitus_undulatus

Roe & Hoeh, 2003 [Publ 2655], fig. 4.2

iT tT bi riS oT roT roS iD
2 2 0 1 29 3 1 --
        +-- Unio_pictorum **       
  +-----+                       
  |     +-- Unio_tumidus **        
--+                             
  |  +----- Pyganodon_grandis   
  +--+                          
     |  +-- Anodonta_cygnea     
     +--+                       
        +-- Strophitus_undulatus

Roe & Hoeh, 2003 [Publ 2655], fig. 4.3

iT tT bi riS oT roT roS iD
2 2 0 1 29 3 1 --
        +-- Unio_pictorum **       
  +-----+                       
  |     +-- Unio_tumidus **        
--+                             
  |  +----- Anodonta_cygnea     
  +--+                          
     |  +-- Pyganodon_grandis   
     +--+                       
        +-- Strophitus_undulatus

Hoeh et al., 2009 [Publ 2716], fig. 3

iT tT bi riS oT roT roS iD
2 2 0 1 45 4 1 --
           +-- Unio_pictorum **        
  +--------+                        
  |        +-- Unio_caffer **          
--+                                 
  |  +-------- Lasmigona_costata    
  +--+                              
     |  +----- Alasmidonta_marginata
     +--+                           
        |  +-- Strophitus_undulatus 
        +--+                        
           +-- Pyganodon_grandis    

Hoeh et al., 2009 [Publ 2716], fig. 4

iT tT bi riS oT roT roS iD
2 2 0 1 44 4 1 --
           +-- Unio_pictorum **        
  +--------+                        
  |        +-- Unio_caffer **          
--+                                 
  |  +-------- Lasmigona_costata    
  +--+                              
     |  +----- Alasmidonta_marginata
     +--+                           
        |  +-- Strophitus_undulatus 
        +--+                        
           +-- Pyganodon_grandis    

Hoeh et al., 2009 [Publ 2716], fig. 5

iT tT bi riS oT roT roS iD
2 2 0 1 44 4 1 --
           +-- Unio_pictorum **        
  +--------+                        
  |        +-- Unio_caffer **          
--+                                 
  |  +-------- Lasmigona_costata    
  +--+                              
     |  +----- Alasmidonta_marginata
     +--+                           
        |  +-- Strophitus_undulatus 
        +--+                        
           +-- Pyganodon_grandis    

Rosenberg et al., 1997 [Publ 2720], fig. 2B

iT tT bi riS oT roT roS iD
2 17 0 2 21 4 1 31.0
  +----- Anodonta              
  |                            
  |  +-- Gonidea_angulata      
  |  |                         
  |  +-- Pyganodon_grandis     
  |  |                         
  |  +-- Utterbackia_imbecillis
  |  |                         
  |  +-- Pyganodon_cataracta   
--+  |                         
  |  +-- Lampsilis_straminea   
  |  |                         
  |  +-- Lampsilis_teres       
  |  |                         
  |  +-- Megalonaias_nervosa   
  |  |                         
  |  +-- Obliquaria_reflexa    
  |  |                         
  +--+-- Theliderma_cylindrica 
     |                         
     +-- Quadrula_quadrula     
     |                         
     +-- Uniomerus_tetralasmus 
     |                         
     +-- Amblema_plicata       
     |                         
     +-- Elliptio_complanata   
     |                         
     +-- Fusconaia_cerina      
     |                         
     +-- Plectomerus_dombeyanus
     |                         
     +-- Aculamprotula_tortuosa **
     |                         
     +-- Unio_pictorum **         

Whelan et al., 2011 [Publ 3380], fig. 1 ML

iT tT bi riS oT roT roS iD
2 2 0 1 35 2 1 --
     +-- Pyganodon_grandis
  +--+                    
  |  +-- Lasmigona_costata
--+                       
  |  +-- Unio_pictorum **    
  +--+                    
     +-- Unio_caffer **      

Whelan et al., 2011 [Publ 3380], fig. 1 MP

iT tT bi riS oT roT roS iD
2 2 0 1 35 2 1 --
     +-- Pyganodon_grandis
  +--+                    
  |  +-- Lasmigona_costata
--+                       
  |  +-- Unio_pictorum **    
  +--+                    
     +-- Unio_caffer **      

Soroka, 2010 [Publ 3404], fig. 8a-M

iT tT bi riS oT roT roS iD
2 2 0 1 1 1 1 --
  +----- Sinanodonta_woodiana
--+                          
  |  +-- Unio_pictorum **       
  +--+                       
     +-- Unio_tumidus **        

Soroka, 2010 [Publ 3404], fig. 8b-M

iT tT bi riS oT roT roS iD
2 2 0 1 1 1 1 --
  +----- Sinanodonta_woodiana
--+                          
  |  +-- Unio_pictorum **       
  +--+                       
     +-- Unio_tumidus **        

Zhou et al., 2016 [Publ 3710], fig. 1

iT tT bi riS oT roT roS iD
2 2 0 2 18 7 2 --
              +----- Lepidodesma_languilati   
  +-----------+                               
  |           |  +-- Schistodesmus_lampreyanus **
  |           +--+                            
--+              +-- Unio_pictorum **            
  |                                           
  |  +-------------- Sinanodonta_woodiana     
  +--+                                        
     |  +----------- Cristaria_plicata        
     +--+                                     
        |  +-------- Platynaias_compressa     
        +--+                                  
           |  +----- Pyganodon_grandis        
           +--+                               
              |  +-- Utterbackia_imbecillis   
              +--+                            
                 +-- Utterbackia_peninsularis 

Bolotov et al., 2016 [Publ 3743], fig. 2

iT tT bi riS oT roT roS iD
2 2 0 1 28 1 1 --
  +----- Anodonta_anatina
--+                      
  |  +-- Unio_pictorum **   
  +--+                   
     +-- Unio_ravoisieri ** 

Bolotov et al., 2016 [Publ 3743], fig. 4

iT tT bi riS oT roT roS iD
2 2 0 1 28 1 1 --
  +----- Anodonta_anatina
--+                      
  |  +-- Unio_pictorum **   
  +--+                   
     +-- Unio_ravoisieri ** 

Soroka & Burzynski, 2018 [Publ 3762], fig. 3

iT tT bi riS oT roT roS iD
2 2 0 2 5 3 1 --
        +-- Unio_tumidus **            
  +-----+                           
  |     +-- Aculamprotula_tortuosa **  
--+                                 
  |  +----- Anodonta_anatina        
  +--+                              
     |  +-- Pyganodon_grandis       
     +--+                           
        +-- Utterbackia_peninsularis

Soroka & Burzynski, 2018 [Publ 3762], fig. 3-M

iT tT bi riS oT roT roS iD
2 2 0 2 5 3 1 --
        +-- Unio_tumidus **            
  +-----+                           
  |     +-- Aculamprotula_tortuosa **  
--+                                 
  |  +----- Anodonta_anatina        
  +--+                              
     |  +-- Pyganodon_grandis       
     +--+                           
        +-- Utterbackia_peninsularis

Guerra et al., 2017 [Publ 3772], fig. 2a

iT tT bi riS oT roT roS iD
2 2 0 2 23 8 1 --
                    +-- Aculamprotula_tortuosa **  
  +-----------------+                           
  |                 +-- Unio_pictorum **           
--+                                             
  |  +----------------- Sinanodonta_woodiana    
  |  |                                          
  +--+  +-------------- Cristaria_plicata       
     |  |                                       
     +--+  +----------- Anodonta_anatina        
        |  |                                    
        +--+        +-- Platynaias_subviridis   
           |  +-----+                           
           |  |     +-- Platynaias_compressa    
           +--+                                 
              |  +----- Pyganodon_grandis       
              +--+                              
                 |  +-- Utterbackia_peninsularis
                 +--+                           
                    +-- Utterbackia_imbecillis  

Guerra et al., 2017 [Publ 3772], fig. 2b

iT tT bi riS oT roT roS iD
2 10 0 2 23 8 1 24.0
  +-------------------- Unio_pictorum **           
  |                                             
--+  +----------------- Aculamprotula_tortuosa **  
  |  |                                          
  +--+              +-- Cristaria_plicata       
     |  +-----------+                           
     |  |           +-- Sinanodonta_woodiana    
     +--+                                       
        |  +----------- Anodonta_anatina        
        |  |                                    
        +--+        +-- Platynaias_subviridis   
           |  +-----+                           
           |  |     +-- Platynaias_compressa    
           +--+                                 
              |  +----- Pyganodon_grandis       
              +--+                              
                 |  +-- Utterbackia_peninsularis
                 +--+                           
                    +-- Utterbackia_imbecillis  

Huang et al., 2018 [Publ 3828], fig. 3, 5

iT tT bi riS oT roT roS iD
2 2 0 2 27 0 0 --
  +-- Unio_pictorum **              
--+                              
  +-- Aculamprotula_tientsinensis **

Chase et al., 2018 [Publ 3847], fig. 4

iT tT bi riS oT roT roS iD
2 2 0 1 37 15 1 --
                       +-- Unio_delphinus **             
  +--------------------+                              
  |                    +-- Unio_tumidus **               
  |                                                   
--+  +-------------------- Sinanodonta_woodiana       
  |  |                                                
  |  |                 +-- Anodonta_cygnea            
  |  |  +--------------+                              
  +--+  |              +-- Anodonta_anatina           
     |  |                                             
     |  |              +-- Pyganodon_lacustris        
     |  |        +-----+                              
     +--+        |     +-- Pyganodon_grandis          
        |  +-----+                                    
        |  |     |  +----- Utterbackia_peninsularis[2]
        |  |     +--+                                 
        |  |        |  +-- Utterbackia_peninsularis[1]
        |  |        +--+                              
        +--+           +-- Utterbackia_imbecillis     
           |                                          
           |           +-- Lasmigona_costata          
           |  +--------+                              
           |  |        +-- Lasmigona_complanata       
           |  |                                       
           +--+        +-- Alasmidonta_undulata       
              |  +-----+                              
              |  |     +-- Alasmidonta_marginata      
              +--+                                    
                 |  +----- Strophitus_undulatus       
                 +--+                                 
                    |  +-- Platynaias_compressa       
                    +--+                              
                       +-- Platynaias_subviridis      

Chase et al., 2018 [Publ 3847], fig. 5

iT tT bi riS oT roT roS iD
2 2 0 1 8 2 1 --
     +-- Anodonta_anatina        
  +--+                           
  |  +-- Utterbackia_peninsularis
--+                              
  |  +-- Unio_delphinus **          
  +--+                           
     +-- Unio_tumidus **            

Chase et al., 2018 [Publ 3861], fig. 1

iT tT bi riS oT roT roS iD
2 2 0 1 9 3 1 --
        +-- Unio_delphinus **          
  +-----+                           
  |     +-- Unio_tumidus **            
--+                                 
  |  +----- Anodonta_anatina        
  +--+                              
     |  +-- Utterbackia_peninsularis
     +--+                           
        +-- Pyganodon_grandis       

Chase et al., 2018 [Publ 3861], fig. 1-M

iT tT bi riS oT roT roS iD
2 2 0 1 9 3 1 --
        +-- Unio_delphinus **          
  +-----+                           
  |     +-- Unio_tumidus **            
--+                                 
  |  +----- Anodonta_anatina        
  +--+                              
     |  +-- Utterbackia_peninsularis
     +--+                           
        +-- Pyganodon_grandis       

Lopes-Lima et al., 2020 [Publ 4080], fig. 11

iT tT bi riS oT roT roS iD
2 2 0 1 10 8 1 --
                    +-- Nodularia_breviconcha **   
  +-----------------+                           
  |                 +-- Nodularia_douglasiae **    
--+                                             
  |  +----------------- Lanceolaria_oxyrhyncha  
  |  |                                          
  +--+  +-------------- Lanceolaria_gladiola    
     |  |                                       
     +--+           +-- Lanceolaria_grayii      
        |  +--------+                           
        |  |        +-- Lanceolaria_bilirata    
        +--+                                    
           |  +-------- Lanceolaria_acrorrhyncha
           +--+                                 
              |  +----- Lanceolaria_kihirai     
              +--+                              
                 |  +-- Lanceolaria_lanceolata  
                 +--+                           
                    +-- Lanceolaria_triformis   

Riccardi et al., 2019 [Publ 4089], fig. 2

iT tT bi riS oT roT roS iD
2 2 0 1 18 16 2 --
                       +-- Lanceolaria_grayii      
                    +--+                           
                    |  +-- Lanceolaria_lanceolata  
  +-----------------+                              
  |                 |  +-- Unio_mancus **             
  |                 +--+                           
  |                    +-- Unio_elongatulus **        
  |                                                
  |              +-------- Cristaria_plicata       
--+              |                                 
  |  +-----------+     +-- Anemina_euscaphys       
  |  |           |  +--+                           
  |  |           |  |  +-- Anemina_arcaeformis     
  |  |           +--+                              
  |  |              |  +-- Sinanodonta_lucida      
  +--+              +--+                           
     |                 +-- Sinanodonta_woodiana    
     |                                             
     |  +----------------- Anodonta_anatina        
     |  |                                          
     |  |           +----- Pseudanodonta_complanata
     +--+  +--------+                              
        |  |        |  +-- Anodonta_exulcerata     
        |  |        +--+                           
        +--+           +-- Anodonta_cygnea         
           |                                       
           |  +----------- Anodonta_nuttalliana    
           +--+                                    
              |  +-------- Platynaias_compressa    
              +--+                                 
                 |  +----- Pyganodon_grandis       
                 +--+                              
                    |  +-- Utterbackia_peninsularis
                    +--+                           
                       +-- Utterbackia_imbecillis  

Tomilova et al., 2020 [Publ 4097], fig. 3

iT tT bi riS oT roT roS iD
2 2 0 1 6 6 1 --
           +-- Unio_tumidus **            
  +--------+                           
  |        +-- Unio_pictorum **           
  |                                    
  |        +-- Sinanodonta_lauta       
--+--------+                           
  |        +-- Sinanodonta_woodiana    
  |                                    
  |  +-------- Anodonta_anatina        
  +--+                                 
     |  +----- Pseudanodonta_complanata
     +--+                              
        |  +-- Anodonta_cygnea         
        +--+                           
           +-- Anodonta_exulcerata     

Mabuchi & Nishida, 2020 [Publ 4120], fig. 1

iT tT bi riS oT roT roS iD
2 3 0 2 35 31 2 16.0
              +----- Nodularia_douglasiae **    
  +-----------+                              
  |           |  +-- Lanceolaria_grayii      
  |           +--+                           
  |              +-- Inversiunio_reinianus **   
  |                                          
--+  +-------------- Cristaria_plicata       
  |  |                                       
  |  |  +----------- Sinanodonta_tumens      
  |  |  |                                    
  |  |  |        +-- Sinanodonta_lauta[5]    
  |  |  |        |                           
  +--+  |        +-- Sinanodonta_lauta[6]    
     |  |        |                           
     |  |        +-- Beringiana_japonica[15] 
     |  |        |                           
     |  |        +-- Beringiana_japonica[16] 
     |  |  +-----+                           
     |  |  |     +-- Beringiana_japonica[17] 
     +--+  |     |                           
        |  |     +-- Beringiana_japonica[18] 
        |  |     |                           
        |  |     +-- Beringiana_japonica[19] 
        |  |     |                           
        |  |     +-- Beringiana_japonica[20] 
        |  |                                 
        |  |  +----- Beringiana_japonica[1]  
        |  |  |                              
        |  |  +----- Beringiana_japonica[2]  
        |  |  |                              
        +--+  +----- Beringiana_japonica[3]  
           |  |                              
           |  +----- Beringiana_japonica[4]  
           |  |                              
           |  +----- Beringiana_japonica[5]  
           |  |                              
           |  +----- Sinanodonta_lauta[1]    
           |  |                              
           |  +----- Beringiana_japonica[6]  
           |  |                              
           |  +----- Sinanodonta_lauta[2]    
           |  |                              
           |  +----- Sinanodonta_calipygos[1]
           |  |                              
           +--+----- Beringiana_japonica[7]  
              |                              
              +----- Sinanodonta_lauta[4]    
              |                              
              +----- Beringiana_japonica[8]  
              |                              
              +----- Beringiana_japonica[9]  
              |                              
              +----- Beringiana_japonica[10] 
              |                              
              +----- Beringiana_japonica[11] 
              |                              
              +----- Beringiana_japonica[12] 
              |                              
              +----- Beringiana_japonica[13] 
              |                              
              +----- Beringiana_japonica[14] 
              |                              
              |  +-- Sinanodonta_lauta[3]    
              +--+                           
                 +-- Sinanodonta_calipygos[2]

Kano et al., 2019 [Publ 4194], fig. 6

iT tT bi riS oT roT roS iD
2 2 0 2 8 4 2 --
  +----- Lanceolaria_grayii      
--+                              
  |  +-- Inversiunio_yanagawensis **
  +--+                           
     +-- Nodularia_nipponensis **   

Nagel et al., 1998 [Publ 1946], fig. 9

iT tT bi riS oT roT roS iD
1 1 0 1 5 2 1 --
  +----- Unio **         
--+                   
  |  +-- Anodonta     
  +--+                
     +-- Pseudanodonta

Araujo et al., 2009 [Publ 2395], fig. 3

iT tT bi riS oT roT roS iD
1 1 0 1 7 0 0 --
-- Unio

Graf, 2000 [Publ 2647], fig. 1

iT tT bi riS oT roT roS iD
1 1 0 1 17 1 1 --
-- Unio

Graf & Ó Foighil, 2000 [Publ 2649], fig. 1, 2

iT tT bi riS oT roT roS iD
1 1 0 1 11 1 1 --
  +-- Unio_pictorum **    
--+                    
  +-- Pyganodon_grandis

Chong et al., 2008 [Publ 2668], fig. 3

iT tT bi riS oT roT roS iD
1 1 0 1 20 20 1 --
  +----------------- Anodonta_cygnea          
  |                                           
  +----------------- Pyganodon_fragilis       
  |                                           
  +----------------- Alasmidonta_holstonia    
  |                                           
  +----------------- Anodonta_anatina         
  |                                           
  +----------------- Unio_tumidus **             
  |                                           
  |              +-- Anodonta_nuttalliana     
  +--------------+                            
  |              +-- Anodonta_californiensis  
  |                                           
--+              +-- Beringiana_beringiana    
  +--------------+                            
  |              +-- Sinanodonta_woodiana     
  |                                           
  |              +-- Pseudodontoideus_subvexus
  |              |                            
  +--------------+-- Alasmidonta_marginata    
  |              |                            
  |              +-- Lasmigona_complanata     
  |                                           
  |  +-------------- Pseudanodonta_complanata 
  |  |                                        
  |  |        +----- Strophitus_undulatus     
  |  +--------+                               
  +--+        |  +-- Prolasmidonta_heterodon  
     |        +--+                            
     |           +-- Platynaias_compressa     
     |                                        
     |  +----------- Anodonta_oregonensis[4]  
     +--+                                     
        |  +-------- Anodonta_oregonensis[3]  
        +--+                                  
           |  +----- Anodonta_oregonensis[2]  
           +--+                               
              |  +-- Anodonta_oregonensis[1]  
              +--+                            
                 +-- Anodonta_kennerlyi       

Taylor et al., 2007 [Publ 2696], fig. 1-3

iT tT bi riS oT roT roS iD
1 1 0 1 3 1 1 --
  +-- Anodonta_cygnea
--+                  
  +-- Unio_pictorum **  

Huff et al., 2004 [Publ 2697], fig. 2

iT tT bi riS oT roT roS iD
1 1 0 1 13 1 1 --
  +-- Unio_ravoisieri **
--+                  
  +-- Anodonta       

Huff et al., 2004 [Publ 2697], fig. 4

iT tT bi riS oT roT roS iD
1 1 0 1 10 1 1 --
  +-- Unio_ravoisieri **
--+                  
  +-- Anodonta       

Huff et al., 2004 [Publ 2697], fig. 5-6

iT tT bi riS oT roT roS iD
1 1 0 1 11 1 1 --
-- Unio_ravoisieri

Rosenberg et al., 1994 [Publ 2719], fig. 3

iT tT bi riS oT roT roS iD
1 1 0 1 18 3 1 --
-- Unio_pictorum

Rosenberg et al., 1997 [Publ 2720], fig. 2A

iT tT bi riS oT roT roS iD
1 1 0 1 22 4 1 --
-- Unio_pictorum

Graf et al., 2014 [Publ 3591], fig. 3

iT tT bi riS oT roT roS iD
1 1 0 1 26 0 0 --
-- Unio_pictorum

Marshall et al., 2014 [Publ 3593], fig. 1

iT tT bi riS oT roT roS iD
1 1 0 1 13 1 1 --
  +-- Pyganodon_grandis
--+                    
  +-- Unio_pictorum **    

Graf et al., 2015 [Publ 3619], fig. 1

iT tT bi riS oT roT roS iD
1 1 0 1 19 0 0 --
-- Unio_pictorum

Pfeiffer & Graf, 2015 [Publ 3620], fig. 2

iT tT bi riS oT roT roS iD
1 1 0 1 39 2 1 --
  +----- Unio_pictorum **    
--+                       
  |  +-- Pyganodon_grandis
  +--+                    
     +-- Lasmigona_costata

Bieler et al., 2014 [Publ 3622], fig. 26, 27, 29, 30

iT tT bi riS oT roT roS iD
1 1 0 1 4 0 0 --
-- Unio_pictorum

Bieler et al., 2014 [Publ 3622], fig. 28

iT tT bi riS oT roT roS iD
1 1 0 1 4 0 0 --
-- Unio_pictorum

Huang et al., 2013 [Publ 3629], fig. 3

iT tT bi riS oT roT roS iD
1 1 0 1 15 6 1 --
  +-------------- Unio_pictorum **           
  |                                       
--+           +-- Sinanodonta_woodiana    
  |  +--------+                           
  |  |        +-- Cristaria_plicata       
  +--+                                    
     |  +-------- Platynaias_compressa    
     +--+                                 
        |  +----- Pyganodon_grandis       
        +--+                              
           |  +-- Utterbackia_imbecillis  
           +--+                           
              +-- Utterbackia_peninsularis

González & Giribet, 2015 [Publ 3632], fig. 2

iT tT bi riS oT roT roS iD
1 1 0 1 5 0 0 --
-- Unio_pictorum

González & Giribet, 2015 [Publ 3632], fig. 3

iT tT bi riS oT roT roS iD
1 1 0 1 5 0 0 --
-- Unio_pictorum

González & Giribet, 2015 [Publ 3632], fig. 4

iT tT bi riS oT roT roS iD
1 1 0 1 5 0 0 --
-- Unio_pictorum

González & Giribet, 2015 [Publ 3632], fig. 6

iT tT bi riS oT roT roS iD
1 1 0 1 5 0 0 --
-- Unio_pictorum

Bolotov et al., 2015 [Publ 3646], fig. 2A

iT tT bi riS oT roT roS iD
1 1 0 1 9 0 0 --
-- Unio_pictorum

Bolotov et al., 2015 [Publ 3646], fig. 2B

iT tT bi riS oT roT roS iD
1 1 0 1 8 0 0 --
-- Unio_pictorum

Giribet & Distel, 2003 [Publ 3670], fig. 3.5

iT tT bi riS oT roT roS iD
1 1 0 1 5 0 0 --
-- Unio_ravoisieri

Wang et al., 2016 [Publ 3714], fig. 2

iT tT bi riS oT roT roS iD
1 1 0 1 2 0 0 --
-- Unio_tumidus

Fonseca et al., 2016 [Publ 3775], fig. 1-M

iT tT bi riS oT roT roS iD
1 1 0 1 8 3 1 --
  +-------- Unio_delphinus **          
--+                                 
  |  +----- Anodonta_anatina        
  +--+                              
     |  +-- Utterbackia_peninsularis
     +--+                           
        +-- Pyganodon_grandis       

Lopes-Lima et al., 2017 [Publ 3792], fig. 2-M

iT tT bi riS oT roT roS iD
1 1 0 1 9 3 1 --
  +-------- Unio_delphinus **          
--+                                 
  |  +----- Anodonta_anatina        
  +--+                              
     |  +-- Utterbackia_peninsularis
     +--+                           
        +-- Pyganodon_grandis       

Feng et al., 2016 [Publ 3800], fig. 2

iT tT bi riS oT roT roS iD
1 1 0 1 14 7 1 --
  +----------- Tchangsinaia_piscicula **
  |                                  
--+  +-------- Pyganodon_grandis     
  |  |                               
  +--+     +-- Anemina_arcaeformis   
     |  +--+                         
     |  |  +-- Anemina_euscaphys     
     +--+                            
        |  +-- Sinanodonta_lucida    
        +--+                         
           +-- Sinanodonta_woodiana  

Combosch et al., 2017 [Publ 3806], fig. 1

iT tT bi riS oT roT roS iD
1 1 0 1 14 0 0 --
-- Unio_pictorum

Combosch et al., 2017 [Publ 3806], fig. 2

iT tT bi riS oT roT roS iD
1 1 0 1 14 0 0 --
-- Unio_pictorum

Vikhrev et al., 2018 [Publ 3812], fig. 3

iT tT bi riS oT roT roS iD
1 1 0 1 12 0 0 --
-- Unio_ravoisieri

Lopes-Lima et al., 2018 [Publ 3829], fig. 1

iT tT bi riS oT roT roS iD
1 1 0 1 27 0 0 --
-- Unio_pictorum

Lopes-Lima et al., 2018 [Publ 3829], fig. 3

iT tT bi riS oT roT roS iD
1 1 0 1 27 0 0 --
-- Unio_pictorum

Gomes-dos-Santos et al., 2019 [Publ 3943], fig. 1

iT tT bi riS oT roT roS iD
1 1 0 1 9 1 1 --
  +-- Unio_delphinus **  
--+                   
  +-- Anodonta_anatina

Gomes-dos-Santos et al., 2019 [Publ 3943], fig. 1-M

iT tT bi riS oT roT roS iD
1 1 0 1 5 1 1 --
  +-- Unio_delphinus **  
--+                   
  +-- Anodonta_anatina

Klunzinger et al., 2021 [Publ 4086], fig. 2

iT tT bi riS oT roT roS iD
1 1 0 1 5 0 0 --
-- Unio_pictorum

Bogan et al., 2023 [Publ 4295], fig. 3

iT tT bi riS oT roT roS iD
1 1 0 1 30 28 1 --
  +----------------------------------- Schistodesmus_lampreyanus ** 
  |                                                              
  |  +-------------------------------- Anodonta_cygnea           
--+  |                                                           
  |  |                       +-------- Cristaria_bellua          
  |  |  +--------------------+                                   
  |  |  |                    |  +----- Cristaria_clessini        
  +--+  |                    +--+                                
     |  |                       |  +-- Cristaria_plicata         
     |  |                       +--+                             
     |  |                          +-- Cristaria_truncata        
     |  |                                                        
     |  |              +-------------- Amuranodonta_kijaensis    
     +--+              |                                         
        |  +-----------+  +----------- Anemina_arcaeformis       
        |  |           |  |                                      
        |  |           +--+  +-------- Buldowskia_suifunica      
        |  |              |  |                                   
        |  |              +--+     +-- Buldowskia_flavotincta    
        |  |                 |  +--+                             
        |  |                 |  |  +-- Buldowskia_shadini        
        |  |                 +--+                                
        +--+                    |  +-- Buldowskia_iwakawai       
           |                    +--+                             
           |                       +-- Buldowskia_kamiyai        
           |                                                     
           |                       +-- Pletholophus_honglinhensis
           |  +--------------------+                             
           |  |                    +-- Pletholophus_tenuis       
           |  |                                                  
           |  |              +-------- Beringiana_japonica       
           +--+  +-----------+                                   
              |  |           |  +----- Beringiana_beringiana     
              |  |           +--+                                
              |  |              |  +-- Beringiana_fukuharai      
              |  |              +--+                             
              +--+                 +-- Beringiana_gosannensis    
                 |                                               
                 |                 +-- Sinanodonta_angula        
                 |  +--------------+                             
                 |  |              +-- Sinanodonta_lucida        
                 |  |                                            
                 +--+  +-------------- Sinanodonta_schrenkii     
                    |  |                                         
                    |  |           +-- Sinanodonta_calipygos     
                    +--+-----------+                             
                       |           +-- Sinanodonta_tumens        
                       |                                         
                       |  +----------- Sinanodonta_woodiana[2]   
                       +--+                                      
                          |  +-------- Sinanodonta_jourdyi       
                          +--+                                   
                             |  +----- Sinanodonta_elliptica     
                             +--+                                
                                |  +-- Sinanodonta_lauta         
                                +--+                             
                                   +-- Sinanodonta_woodiana[1]   

Page updated: 08:51:04 Mon 06 May 2024

 
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