|
Bivalvia | Unionoida superfamily UNIONOIDEA Rafinesque, 1820: cladomics
2 families
Cladomics Summary
number of species = 1062
number of subtaxa (rank: family) = 2
analyzed in 451 cladograms in 257 publications
maximum number of terminal species in an analysis = 204, mean = 21.69
maximum number of subtaxa (family) in an analysis = 2, mean = 1.46
Basic Monophyly
number of cladograms that test basic monophyly (i.e., more than 1 included species) = 449
maximum number of included species = 204, mean = 21.79
maximum number of subtaxa (family) = 2, mean = 1.47
proportion of cladograms that support basic monophyly = 0.93
Rigorous Monophyly
number of cladograms that test rigorous monophyly = 78
(Rigorous monophyly refers to cladograms with terminal species representing more than one subtaxon (family) taxon and at least one rigorous outgroup (superfamily).
maximum number of included species = 173, mean = 31.65
maximum number of subtaxa (family) = 2, mean = 2.0
proportion of cladograms that support rigorous monophyly = 0.65
Lopes-Lima et al., 2017, fig. 1-2
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 71 |
78 |
70 |
2 |
10 |
9 |
3 |
0.0 |
*** Top Monophyly Cladogram *** *** Top Sister-Group Cladogram ***
+-- Hyridella_depressa
+-----------------------------------+
| +-- Velesunio_ambiguus
|
| +-- Cumberlandia_monodonta **
| +--------------------------------+
| | +-- Margaritifera_margaritifera **
| |
| | +----------- Mutela_rostrata
--+ | |
| | +--------------------+ +-- Aspatharia_pfeifferiana
| | | | +-----+
| | | | | +-- Chambardia_wahlbergi
| | | +--+
| | | | +----- Etheria_elliptica
| | | +--+
| | | | +-- Anodontites_elongata
+--+ | +--+
| | +-- Mycetopoda_siliquosa
| |
| | +-------- Trapezidens_dolichorhynchus **
| | +--------------------+
| | | | +----- Leoparreysia_tavoyensis **
| | | +--+
| | | | +-- Coelatura_aegyptiaca **
| | | +--+
| | | +-- Indochinella_pugio **
| | |
| | | +----- Unio_gibbus **
+--+ | |
| | +-----------+----- Unio_tumidus **
| | | |
| | | | +-- Unio_crassus **
| | | +--+
| | +--+ +-- Unio_pictorum **
| | | |
| | | | +-------------- Aculamprotula_tortuosa **
| | | | |
| | | +--+ +-- Nodularia_nuxpersicae **
| | | | +--------+
| | | | | +-- Nodularia_douglasiae **
| | | +--+
| | | | +-------- Schistodesmus_lampreyanus **
| | | +--+
| | | | +----- Tchangsinaia_piscicula **
| | | +--+
| | +-----+ | +-- Cuneopsis_heudei **
| | | | +--+
| | | | +-- Cuneopsis_rufescens **
+--+ | |
| | | +-------- Lanceolaria_gladiola **
| | | +--------+
| | | | | +----- Lanceolaria_grayii[2] **
| | | | +--+
| | | | | +-- Lanceolaria_lanceolata **
| | | | +--+
| | | | +-- Lanceolaria_grayii[1] **
| | | |
| | +--+ +----------- Cristaria_plicata **
| | | +--+
| | | | | +-------- Anemina_arcaeformis **
| | | | +--+
| | | | | +----- Pletholophus_tenuis **
| | | | +--+
| | | | | +-- Sinanodonta_woodiana **
| | | | +--+
| | +--+ +-- Sinanodonta_lucida **
| | |
| | | +----- Pseudanodonta_complanata **
| | | |
| | | +-----+ +-- Anodonta_anatina **
| | | | | |
| | | | +--+-- Anodonta_cygnea **
| | | | |
| | +--+ +-- Anodonta_nuttalliana **
| | |
| | | +-- Pyganodon_grandis **
+--+ | +-----+
| | | +-- Simpsonaias_ambigua **
| +--+
| | +----- Strophitus_undulatus **
| +--+
| | +-- Alasmidonta_marginata **
| +--+
| +-- Lasmigona_compressa **
|
| +-- Lens_contradens **
| +--+
| | +-- Lens_comptus **
| +--+
| | | +-- Physunio_modelli **
| | +--+
| | +-- unionoid_species
| |
| +--------------+ +----- Hyriopsis_desowitzi **
| | | |
| | | +----- Rectidens_sumatrensis **
| | | |
| | | | +-- Hyriopsis_khoratensis **
| | +--+--+
| | | +-- Hyriopsis_myersiana **
| | |
| | | +-- Ensidens_ingallsianus **
| | | |
| | +--+-- Ensidens_telus **
| | |
| | +-- Ensidens_jaculus **
| |
| | +-------- Amblema_plicata **
+--+ +--+
| | | +----- Cambarunio_iris **
| | +--+
| | | +-- Lampsilis_cardium **
| | +--+
| +--------+ +-- Ortmanniana_ligamentina **
| | |
| | | +-- Quadrula_quadrula **
| | | +-----+
| | | | +-- Tritogonia_verrucosa **
| | +--+
| | | +----- Elliptio_complanata **
| | +--+
| | | +-- Pleurobema_sintoxia **
| | +--+
| | +-- Eurynia_dilatata **
+--+
| +-- Chamberlainia_hainesiana **
| +-----+
| | +-- Sinohyriopsis_cumingii **
| |
| +--------+ +-- Lamprotula_caveata **
| | | +--+
| | | | +-- Lamprotula_leaii **
| | +--+
| | | +-- Potomida_littoralis **
| | +--+
| | +-- Pronodularia_japanensis **
+--+
| +-- Sinosolenaia_carinata **
| +-----+
| | +-- Sinosolenaia_oleivora **
| +-----+
| | | +----- Gonidea_angulata **
| | +--+
| | | +-- Microcondylaea_bonellii **
+--+ +--+
| +-- Leguminaia_wheatleyi **
|
| +----------- Parvasolenaia_triangularis **
| |
+--+ +-- Pilsbryoconcha_exilis **
| +-----+
| | +-- Pilsbryoconcha_acuta **
+--+
| +----- Monodontina_cambodjensis **
+--+
| +-- Bineurus_exilis **
+--+
+-- Nyeinchanconcha_nyeinchani **
Inoue et al., 2018, fig. 1, 3a-c
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 204 |
204 |
65 |
2 |
0 |
0 |
0 |
223.0 |
+-------------------------------------------------------- Cumberlandia_monodonta **
|
| +-- Alasmidonta_marginata **
| |
| +-- Alasmidonta_heterodon **
| |
| +-- Alasmidonta_undulata **
| |
| +-- Beringiana_beringiana **
| |
| +-- Anodonta_kennerlyi **
| |
| +-- Anodonta_nuttalliana **
| |
| +-- Anodonta_oregonensis **
| |
| +-- Arcidens_confragosus **
| |
| +-- Arcidens_wheeleri **
| |
--+ +-- Lasmigona_complanata **
| |
| +--------------------------------------------------+-- Lasmigona_compressa **
| | |
| | +-- Lasmigona_costata **
| | |
| | +-- Lasmigona_decorata **
| | |
| | +-- Lasmigona_subviridis **
| | |
| | +-- Pyganodon_grandis **
| | |
| | +-- Pyganodon_lacustris **
| | |
| | +-- Pseudodontoideus_subvexus **
| | |
| | +-- Strophitus_undulatus **
| | |
| | +-- Utterbackia_imbecillis **
| | |
+--+ +-- Utterbackia_peninsularis[1] **
| |
| +-- Utterbackia_peninsularis[2] **
|
| +-------------------------------------------------- Gonidea_angulata **
| |
| | +-- Reginaia_ebenus **
| | +--------------------------------------------+
| | | +-- Reginaia_rotulata **
| | |
| | | +-- Amblema_plicata **
| | +--------------------------------------------+
| | | +-- Amblema_elliottii **
| | |
| | | +----- Popenaias_popeii **
| | | |
| | | | +-- Dromus_dromas **
| | | | |
+--+ | | +-- Epioblasma_rangiana **
| | | |
| +-----------------------------------------+ +-- Glebula_rotundata **
| | | |
| | | +-- Lampsilis_ovata **
| | | |
| | | +-- Lampsilis_virescens **
| | | |
| | | +-- Lemiox_rimosus **
| | +--+
| | +-- Obliquaria_reflexa **
| | |
| | +-- Obovaria_olivaria **
| | |
+--+ +-- Obovaria_unicolor **
| |
| +-- Plectomerus_dombeyanus **
| |
| +-- Potamilus_alatus **
| |
| +-- Potamilus_capax **
|
| +-- Uniomerus_declivis **
| +--+
| | +-- Uniomerus_tetralasmus **
| |
| | +-- Cyclonaias_tuberculata **
| +--------------------------------------+ |
| | | +-- Cyclonaias_pustulosa[1] **
| | | |
| | | +-- Theliderma_metanevra **
| | +--+
| | +-- Cyclonaias_pustulosa[2] **
| | |
| | +-- Quadrula_quadrula **
| | |
| | +-- Tritogonia_verrucosa **
| |
| | +----------------------------------------- Elliptoideus_sloatianus **
+--+ |
| | +-- Elliptio_lanceolata **
| | +-----------------------------------+
| | | +-- Parvaspina_collina **
| | |
| | | +----------------------------------- Plethobasus_cyphyus **
| | | |
| | | +----------------------------------- Eurynia_dilatata **
| | | |
| | | +----------------------------------- Hemistena_lata **
| | | |
| | | | +-- Elliptio_complanata **
+--+ | | |
| | | +-- Elliptio_hopetonensis **
| | | |
| | | +-- Elliptio_producta[1] **
| | | |
| | | +-- Elliptio_shepardiana **
| | | |
| | | +-- Elliptio_angustata **
| | | |
| | | +-- Elliptio_arca **
| | | |
| | | +-- Elliptio_arctata **
| | | |
+--+ | +-- Elliptio_jayensis **
| | |
| | +-- Elliptio_crassidens **
| | |
| +--------------------------------+-- Elliptio_dariensis **
| | |
| | +-- Elliptio_fisheriana **
| | |
| | +-- Elliptio_folliculata **
| | |
| | +-- Elliptio_fumata **
| | |
| | +-- Elliptio_icterina **
| | |
| | +-- Elliptio_mcmichaeli **
| | |
| | +-- Elliptio_producta[2] **
| | |
| | +-- Elliptio_pullata **
| | |
| | +-- Elliptio_purpurella **
| | |
| | +-- Elliptio_congaraea **
| |
+--+ +-- Pleuronaia_barnesiana **
| |
| +-----------------------------+-- Pleuronaia_dolabelloides **
| | |
| | +-- Pleuronaia_gibber **
| |
| | +-------------------------- Pleurobema_plenum **
| | |
| | | +----------------------- Pleurobema_stabile **
| | | |
| | | | +----- Pleurobema_pyriforme **
| | +--+ | +--+
| | | | | | | +-- Pleurobema_georgianum[2] **
| | | | | | +--+
| | | | | +-----------+ +-- Pleurobema_athearni **
| | | | | | |
| | | +--+ | | +----- Pleurobema_decisum **
| | | | | | |
| | | | | +--+ +-- Pleurobema_georgianum[1] **
| | | | | | |
| | | | | +--+-- Pleurobema_hanleyianum[1] **
| | | | | |
| | | | | +-- Pleurobema_hanleyianum[2] **
| | | | |
| | | +--+ +-------------- Pleurobema_beadleianum[2] **
| | | | |
| | | | +--+ +----------- Pleurobema_strodeanum[2] **
| | | | | | |
| | | | | +--+----------- Pleurobema_rubellum **
| | | | | |
| | | | | | +-------- Pleurobema_beadleianum[1] **
| | | | | +--+
| | | | | | +----- Pleurobema_taitianum **
| | | | | +--+
| | | | | | +-- Pleurobema_perovatum **
| | | +--+ +--+
| | | | +-- Pleurobema_strodeanum[1] **
| | | |
| | | | +----- Pleurobema_oviforme[6] **
| | | | |
| | | | +--+----- Pleurobema_clava[3] **
| | | | | |
| | | | | | +-- Pleurobema_oviforme[5] **
| | | | | +--+
+--+ | +--------+ +-- Pleurobema_clava[2] **
| | |
| | | +----- Pleurobema_oviforme[2] **
| | | |
| | | +----- Pleurobema_oviforme[3] **
| | +--+
| | +----- Pleurobema_oviforme[4] **
| | |
| | | +-- Pleurobema_clava[1] **
| | +--+
| | +-- Pleurobema_oviforme[1] **
| |
| | +-------------------------- Pleurobema_cordatum **
| | |
| | | +----- Fusconaia_subrotunda **
| | | +-----------------+
| +--+ | | +-- Fusconaia_cor **
| | | | +--+
| | | | +-- Fusconaia_cuneolus **
| | | |
| | | | +-------------------- Fusconaia_ozarkensis **
| | +--+ |
| | | | +-- Fusconaia_escambia **
| | | | +--------------+
| | | | | +-- Fusconaia_burkei **
| | | | |
| | | | | +----- Fusconaia_mitchelli **
| | +--+ | |
| | | | +----- Fusconaia_flava[27] **
| | | | |
| | | +-----------+ +-- Fusconaia_askewi[1] **
| | | | | |
| | | | | +-- Fusconaia_askewi[2] **
| | | | | |
| | | | | +-- Fusconaia_askewi[3] **
| | +--+ +--+
| | | +-- Fusconaia_askewi[4] **
| | | |
| | | +-- Fusconaia_askewi[5] **
| | | |
| | | +-- Fusconaia_chunii **
| | |
| | | +-------------- Fusconaia_masoni **
+--+ | |
| | | +----------- Fusconaia_flava[26] **
| | | |
| | | | +-- Fusconaia_cerina[8] **
| +--+ | +-----+
| | | | +-- Fusconaia_flava[25] **
| | | |
| | | | +----- Fusconaia_flava[5] **
| | | | |
| | | | +----- Fusconaia_flava[6] **
| +--+ | |
| | | +----- Fusconaia_flava[7] **
| | | |
| | | +----- Fusconaia_flava[8] **
| | | |
| | | +----- Fusconaia_cerina[4] **
| | | |
| | | +----- Fusconaia_flava[9] **
| | | |
| +--+ +----- Fusconaia_flava[10] **
| | |
| | +----- Fusconaia_flava[11] **
| | |
| | +----- Fusconaia_flava[12] **
| | |
| | +----- Fusconaia_flava[13] **
| | |
| | +----- Fusconaia_flava[14] **
| | |
| | +----- Fusconaia_flava[15] **
| | |
| | +----- Fusconaia_flava[16] **
| | |
| | +----- Fusconaia_flava[17] **
| +--+
| +----- Fusconaia_flava[18] **
| |
| +----- Fusconaia_flava[19] **
| |
| +----- Fusconaia_flava[20] **
| |
| +----- Fusconaia_flava[21] **
| |
| +----- Fusconaia_flava[22] **
| |
| +----- Fusconaia_flava[23] **
| |
| +----- Fusconaia_flava[24] **
| |
| +----- Fusconaia_cerina[5] **
| |
| +----- Fusconaia_cerina[6] **
| |
| +----- Fusconaia_cerina[7] **
| |
| | +-- Fusconaia_flava[1] **
| | |
| | +-- Fusconaia_flava[2] **
| | |
| | +-- Fusconaia_flava[3] **
| | |
| +--+-- Fusconaia_flava[4] **
| |
| +-- Fusconaia_cerina[1] **
| |
| +-- Fusconaia_cerina[2] **
| |
| +-- Fusconaia_cerina[3] **
|
| +----------------- Pleurobema_riddellii **
| |
| | +-------------- Pleurobema_sintoxia[21] **
| | |
| | | +----------- Pleurobema_sintoxia[19] **
| | | |
| | | +----------- Pleurobema_sintoxia[20] **
| | | |
+-----------+ | | +----- Pleurobema_sintoxia[18] **
| | | |
| | | +----- Pleurobema_rubrum[19] **
| | | |
| | | +----- Pleurobema_rubrum[20] **
| | +-----+
| | | +----- Pleurobema_rubrum[21] **
| | | |
| | | | +-- Pleurobema_rubrum[17] **
+--+ | | |
| | | +-- Pleurobema_sintoxia[16] **
| | +--+
| | +-- Pleurobema_sintoxia[17] **
| | |
| | +-- Pleurobema_rubrum[18] **
| |
| | +-------- Pleurobema[4] **
| | |
| | +-------- Pleurobema[5] **
| | |
| | +-------- Pleurobema_sintoxia[1] **
| | |
| | +-------- Pleurobema[8] **
| | |
| | +-------- Pleurobema[9] **
+--+ |
| +-------- Pleurobema[10] **
| |
| +-------- Pleurobema_sintoxia[3] **
| |
| +-------- Pleurobema_sintoxia[4] **
| |
| +-------- Pleurobema_sintoxia[5] **
| |
| +-------- Pleurobema_sintoxia[6] **
| |
| +-------- Pleurobema_sintoxia[7] **
| |
| +-------- Pleurobema_sintoxia[8] **
| |
| +-------- Pleurobema_sintoxia[9] **
| |
| +-------- Pleurobema_sintoxia[10] **
| |
| +-------- Pleurobema_sintoxia[11] **
| |
| +-------- Pleurobema_sintoxia[12] **
| |
| +-------- Pleurobema_sintoxia[13] **
| |
| +-------- Pleurobema_sintoxia[14] **
| |
| +-------- Pleurobema_sintoxia[15] **
| |
+--+-------- Pleurobema_rubrum[4] **
|
+-------- Pleurobema_rubrum[5] **
|
+-------- Pleurobema_rubrum[6] **
|
+-------- Pleurobema_rubrum[7] **
|
+-------- Pleurobema_rubrum[8] **
|
+-------- Pleurobema_rubrum[9] **
|
+-------- Pleurobema_rubrum[10] **
|
+-------- Pleurobema_rubrum[11] **
|
+-------- Pleurobema_rubrum[12] **
|
+-------- Pleurobema_rubrum[13] **
|
+-------- Pleurobema_rubrum[14] **
|
+-------- Pleurobema_rubrum[15] **
|
+-------- Pleurobema_rubrum[16] **
|
| +-- Pleurobema[6] **
+-----+
| +-- Pleurobema_rubrum[1] **
|
| +-- Pleurobema_sintoxia[2] **
+-----+
| +-- Pleurobema_rubrum[2] **
|
| +-- Pleurobema[7] **
+-----+
| +-- Pleurobema_rubrum[3] **
|
| +----- Pleurobema[3] **
+--+
| +-- Pleurobema[1] **
+--+
+-- Pleurobema[2] **
Huang et al., 2019, fig. 2
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 173 |
173 |
147 |
2 |
26 |
24 |
2 |
136.0 |
+----- Alathyria_jacksoni
+--+
| | +-- Alathyria_profuga
| +--+
+--------+ +-- Velesunio_ambiguus
| |
| | +----- Alathyria_pertexta
| +--+
| | +-- Lortiella_froggatti
+--------------------------------+ +--+
| | +-- Microdontia_anodontaeformis
| |
| | +-------------- Westralunio_carteri
| | |
| +--+ +----------- Echyridella_menziesii
| | |
| | | +----- Diplodon_suavidicus
| +--+ +--+
| | | | +-- Castalia_ambigua
| | | +--+
| +--+ +-- Triplodon_corrugatus
| |
| | +----- Hyridella_australis
| +--+
| | +-- Hyridella_depressa
| +--+
| +-- Hyridella_drapeta
|
| +-------- Anodontites_trapesialis
| +--+
| | | +----- Mutela_rostrata
| | +--+
| | | +-- Mutela_dubia
| | +--+
| +-----------------------------------+ +-- Mutela_hargeri
| | |
| | | +----- Mycetopoda_siliquosa
| | | +--+
--+ | | | | +-- Anodontites_elongata
| | | | +--+
| | +--+ +-- Lamproscapha_ensiformis
| | |
| | | +----- Etheria_elliptica
| | +--+
| | | +-- Aspatharia_pfeifferiana
| | +--+
| | +-- Chambardia_wahlbergi
| |
| | +----------------- Gibbosula_rochechouartii **
| | |
| | | +-------------- Gibbosula_laosensis **
| | +--------------------------+ |
| | | | | +----------- Cumberlandia_monodonta **
| | | | | |
| | | +--+ | +----- Pseudunio_marocanus **
| | | | +-----+
| | | | | | +-- Pseudunio_auricularius **
| | | | | +--+
| | | +--+ +-- Pseudunio_homsensis **
| | | |
| | | | +-- Margaritifera_dahurica **
| | | | +-----+
| | | | | +-- Margaritifera_margaritifera **
| | | | |
| | | +--+ +----- Margaritifera_laevis **
| | | | |
| | | | | +-- Margaritifera_falcata **
| | | +--+--+
+--+ | | +-- Margaritifera_middendorffi **
| | |
| | | +-- Margaritifera_hembeli **
| | +--+
| | +-- Margaritifera_marrianae **
| |
| | +----- Trapezidens_exolescens **
| | +-----------------+
| | | | +-- Trapezidens_dolichorhynchus **
| | | +--+
| | +--+ +-- Trapezidens_angustior **
| | | |
| | | | +-------------------- Lamellidens_corrianus **
| | | | |
| | | +--+ +----------------- Lamellidens_savadiensis **
| | | | |
| | | +--+ +-- Lamellidens_lamellatus **
| | | | +-----------+
| | | | | +-- Lamellidens_jenkinsianus **
| | | +--+
| | | | +----------- Lamellidens_generosus **
| | | +--+
| | | | +-------- Lamellidens_mainwaringi[2] **
| | | +--+
| | +--------------+ | +----- Lamellidens_mainwaringi[1] **
| | | | +--+
| | | | | +-- Lamellidens_ferrugineus **
| | | | +--+
| | | | +-- Lamellidens_marginalis **
| | | |
| | | | +----- Parreysia_keralaensis **
| | | | +-----------------+
| | | | | | +-- Parreysia_nagpoorensis **
| | | | | +--+
| | | | | +-- Parreysia_plagiosoma **
+--+ | | |
| | | | +-- Leoparreysia_tavoyensis **
| | | | |
| | +--+ +-----------------+-- Leoparreysia_canefrii **
| | | | |
| | | | +-- Leoparreysia_olivacea **
| | | |
| | | | +-------- Coelatura_aegyptiaca[2] **
| | | | |
| | | | +--------+ +----- Coelatura_aegyptiaca[1] **
| | +--+ | | |
| | | | | +----- Nitia_teretiuscula **
| | | | +--+
| | | | | +-- Coelatura_gabonensis **
| | | | | |
| | | | +--+-- Coelatura_kunenensis **
| | | | |
| | +--+ +-- Prisodontopsis_mweruensis **
| | |
| | | +-------- Indonaia_shurtleffiana **
| | | |
| | | +-----+ +----- Indonaia_bonneaudii **
| | | | | |
| | | | +--+----- Indonaia_rugosa **
| | | | |
| | | | | +-- Indonaia_andersoniana **
| | +--+ +--+
| | | +-- Indonaia_caerulea **
| | |
| | | +----------- Indochinella_pugio **
| | | |
| | | | +----- Scabies_humilis **
| | +--+ +--+
| | | | | +-- Scabies_phaselus **
| | | | +--+
| | +--+ +-- Scabies_mandarinus **
| | |
| | | +----- Radiatula_myitkyinae **
| | +--+
| | | +-- Radiatula_chaudhurii **
| | +--+
| | +-- Radiatula_mouhoti **
| |
| | +-------------------------------- Acuticosta_chinensis **
| | |
| | | +----------------------------- Lepidodesma_languilati **
| | | |
| | | | +-------- Aculamprotula_tortuosa **
| | | | |
| | | | +-----------+ +----- Aculamprotula_coreana **
| | | | | | |
| | | | | +--+----- Aculamprotula_scripta **
| | | | | |
+--+ | | | | +-- Aculamprotula_fibrosa **
| | | | +--+
| | | | +-- Aculamprotula_tientsinensis **
| +-----+ +--------+
| | | | | +-------- Unio_tumidus **
| | | | | +--------+
| | | | | | | +----- Unio_crassus **
| | | | | | +--+
| | | | | | | +-- Unio_delphinus **
| | | | | | +--+
| | | | +--+ +-- Unio_pictorum **
| | | | |
| | | | | +-- Schistodesmus_lampreyanus **
| | | | | +-----------+
| | | | | | +-- Schistodesmus_spinosus **
| | | | | |
| | | | +--+ +----- Nodularia_douglasiae[1] **
| | +--+ | +-----+
| | | | | | +-- Nodularia_douglasiae[2] **
| | | | | +--+
| | | +--+ +-- Nodularia_nuxpersicae **
| | | |
| | | | +-------- Tchangsinaia_piscicula **
| | | +--+
| | | | +----- Cuneopsis_rufescens **
| | | +--+
| | | | +-- Cuneopsis_heudei **
| | | +--+
| | | +-- Cuneopsis_celtiformis **
| | |
| | | +-------- Lanceolaria_gladiola **
| | | |
| | | +-----------------+ +----- Lanceolaria_grayii[1] **
| | | | | |
| | | | +--+----- Lanceolaria_grayii[2] **
| | | | |
| | | | | +-- Lanceolaria_lanceolata **
| | | | +--+
| | | | +-- Lanceolaria_triformis **
| | | |
| | | | +-------------------- Anemina_arcaeformis **
| | | | |
| | | | +--+ +----------------- Cristaria_plicata **
| | +--+ | | |
| | | | +--+ +-------------- Pletholophus_tenuis **
| | | | | |
| | | | +--+ +-- Sinanodonta_angula **
| | | | | +--------+
| | | | | | +-- Sinanodonta_lucida **
| | | | +--+
| | | | | +-------- Sinanodonta_woodiana[2] **
| | | | +--+
| | | | | +----- Sinanodonta[2] **
| | | | +--+
| | +--+ | +-- Sinanodonta_woodiana[1] **
| | | +--+
| | | +-- Sinanodonta[1] **
| | |
| | | +-------- Alasmidonta_marginata **
| | | |
| | | +-------- Lasmigona_compressa **
| | | |
| | | +-------- Pseudanodonta_complanata **
| | | |
| | | +-------- Simpsonaias_ambigua **
+--+ | |
| +--------------+-------- Strophitus_undulatus **
| |
| | +----- Anodonta_anatina **
| +--+
| | | +-- Anodonta_cygnea **
| | +--+
| | +-- Anodonta_nuttalliana **
| |
| | +----- Pyganodon_grandis **
| +--+
| | +-- Utterbackia_imbecillis **
| +--+
| +-- Utterbackia_peninsularis **
|
| +----------- Hyriopsis_desowitzi **
| |
| +----------- Rectidens_sumatrensis **
| |
| | +-- Hyriopsis_bialata **
| | |
| | +-- Hyriopsis_myersiana **
| +-----+--------+
| | | +-- Hyriopsis_panhai **
| | | |
| | | +-- Hyriopsis_khoratensis **
| | |
| | | +-- Ensidens_ingallsianus **
| | | +-----+
| | | | +-- Ensidens_telus **
| | +--+
| | | +----- Ensidens_sagittarius **
| +-----------------+ +--+
| | | | +-- Ensidens_spiculus **
| | | +--+
| | | +-- Ensidens_jaculus **
| | |
| | | +----- Physunio **
| | | +--------+
| | | | | +-- Physunio_modelli **
| | | | +--+
| | | | +-- Physunio_superbus **
| | | |
| | +--+ +-------- Yaukthwa_inlenensis **
| | | +--+
| | | | | +----- Yaukthwa_panhai **
| | | | +--+
| | | | | +-- Yaukthwa_nesemanni **
| | | | +--+
| | +--+ +-- Yaukthwa_rectangularis **
| | |
| | | +-- Lens_contradens **
| | | +-----+
| | | | +-- Lens_micropterus **
| | +--+
| | | +----- Lens_eximius **
| | +--+
| | | +-- Lens_comptus **
| | +--+
| | +-- Lens_novoselovi **
| |
| | +-- Megalonaias_nervosa **
| | +--+
+--+ | +-- Uniomerus_carolinianus **
| +-----------------------+
| | | +-- Quadrula_quadrula **
| | +--+
| | +-- Tritogonia_verrucosa **
| |
| | +-------- Elliptoideus_sloatianus **
| +--+ |
| | | +-----------------+ +-- Elliptio_complanata **
| | | | | +--+
| | | | | | +-- Eurynia_dilatata **
| | | | +--+
| | | | | +-- Fusconaia_flava **
| | +--+ +--+
| | | +-- Pleurobema_sintoxia **
| | |
| | | +----------------------- Amblema_plicata **
| | | |
| | | | +----- Toxolasma_parvum **
| | +--+ +--------------+
| | | | | +-- Obliquaria_reflexa **
| | | | +--+
| | | | +-- Plectomerus_dombeyanus **
| | +--+
| | | +----------------- Potamilus_leptodon **
| | | |
| | | | +-------------- Potamilus_alatus **
| | +--+ |
| | | | +----------- Venustaconcha_ellipsiformis **
| | | | |
| | +--+ +----------- Leaunio_lienosus **
| | | |
+--+ | | +-- Ellipsaria_lineolata **
| | +--------+
| +--+ +-- Truncilla_truncata **
| |
| | +-- Medionidus_conradicus **
| +--------+
| | +-- Ptychobranchus_subtentus **
| |
| | +-------- Cambarunio_iris **
| +--+
| | +----- Ortmanniana_ligamentina **
| +--+
| | +-- Lampsilis_cardium **
| +--+
| +-- Lampsilis_ornata **
|
| +----- Chamberlainia_hainesiana **
| +-----------------------+
| | | +-- Sinohyriopsis_cumingii **
| | +--+
| | +-- Sinohyriopsis_schlegelii **
| |
| | +-- Potomida_littoralis **
| | +-----+
| | | +-- Pronodularia_japanensis **
| | +-----------------+
+--+ | | +----- Lamprotula_leaii **
| | +--+
| | | +-- Lamprotula_caveata **
| | +--+
| | +-- Lamprotula_cornuumlunae **
| |
| | +----- Sinosolenaia_carinata **
| | +-----+
| | | | +-- Sinosolenaia_oleivora **
+--+ | +--+
| | +-- Sinosolenaia_recognita **
| +-----------+
| | | +----- Gonidea_angulata **
| | | +--+
| | | | | +-- Leguminaia_wheatleyi **
| | | | +--+
| | +--+ +-- Microcondylaea_bonellii **
| | |
| | | +-- Ptychorhynchus_pfisteri **
| | | |
+--+ +-----+-- Parvasolenaia_rivularis **
| |
| +-- Parvasolenaia_triangularis **
|
| +----- Pseudodon_manueli **
| +--------------+
| | | +-- Pseudodon_inoscularis **
| | +--+
| | +-- Pseudodon_bogani **
+--+
| +----------------- Pilsbryoconcha_exilis **
| |
| | +----------- Nyeinchanconcha_nyeinchani **
| | +--+
+--+ | | +-------- Thaiconcha_callifera **
| | +--+
| | | +----- Bineurus_loeiensis **
| | +--+
| | | +-- Bineurus_mouhotii **
+--+ +--+
| +-- Bineurus_exilis **
|
| +-- Monodontina_vondembuschiana **
| +--+
| | +-- Monodontina_cambodjensis **
+--------+
| +-- Sundadontina_cumingii **
| |
+--+-- Sundadontina_tumida[1] **
|
+-- Sundadontina_tumida[2] **
Huang et al., 2019, fig. S4
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 173 |
173 |
104 |
2 |
26 |
24 |
2 |
152.0 |
+----------- Etheria_elliptica
|
| +-- Aspatharia_pfeifferiana
+--------+
| +-- Chambardia_wahlbergi
|
+-----------------------+ +----- Mycetopoda_siliquosa
| +-----+
| | | +-- Anodontites_elongata
| | +--+
| | +-- Lamproscapha_ensiformis
| |
| | +-------- Anodontites_trapesialis
| +--+
| | +----- Mutela_hargeri
| +--+
| | +-- Mutela_dubia
| +--+
| +-- Mutela_rostrata
|
| +----------- Echyridella_menziesii
| |
| +----------- Westralunio_carteri
| |
| | +----- Diplodon_suavidicus
| +-----+
| | | +-- Castalia_ambigua
| | +--+
| | +-- Triplodon_corrugatus
| |
| +--------------------+ +----- Hyridella_australis
| | +-----+
| | | | +-- Hyridella_depressa
| | | +--+
| | | +-- Hyridella_drapeta
| | |
| | | +----- Velesunio_ambiguus
--+ | | +--+
| | | | | +-- Alathyria_jacksoni
| | | | +--+
| | +--+ +-- Alathyria_profuga
| | |
| | | +----- Alathyria_pertexta
| | +--+
| | | +-- Lortiella_froggatti
| | +--+
| | +-- Microdontia_anodontaeformis
| |
| | +-------------------- Gibbosula_rochechouartii **
| | |
| | +--------+ +----------------- Gibbosula_laosensis **
| | | | |
| | | +--+ +-------------- Cumberlandia_monodonta **
| | | | |
| | | | | +----- Pseudunio_marocanus **
| | | +--+ +-----+
| | | | | | +-- Pseudunio_auricularius **
| | | | | +--+
| | | | | +-- Pseudunio_homsensis **
| | | +--+
| | | | +-------- Margaritifera_falcata **
| | | | |
| | | | | +-- Margaritifera_dahurica **
| | | | +-----+
| | | +--+ +-- Margaritifera_margaritifera **
| | | |
| | | | +----- Margaritifera_laevis **
| | | | |
| | | +--+----- Margaritifera_middendorffi **
+--+ | |
| | | +-- Margaritifera_hembeli **
| | +--+
| | +-- Margaritifera_marrianae **
| |
| | +-- Parreysia_keralaensis **
| | |
| | +--------------------+-- Parreysia_nagpoorensis **
| | | |
| | | +-- Parreysia_plagiosoma **
| | |
| | | +----- Trapezidens_exolescens **
| | | +--------+
| | | | | +-- Trapezidens_angustior **
| | | | +--+
| | +--------+ +-- Trapezidens_dolichorhynchus **
| | | |
| | | | +----------- Lamellidens_corrianus **
| | | | |
| | | +--+ +-------- Lamellidens_savadiensis **
| | | | |
| | | | | +----- Lamellidens_marginalis **
| | | +--+ |
| | | | +----- Lamellidens_ferrugineus **
| | | | |
| | +--+ +--+ +-- Lamellidens_mainwaringi[1] **
| | | | | |
| | | | | +-- Lamellidens_mainwaringi[2] **
| | | | | |
| | | | +--+-- Lamellidens_lamellatus **
| | | | |
| | | | +-- Lamellidens_generosus **
| | | | |
| | | | +-- Lamellidens_jenkinsianus **
| | | |
| | | | +----- Leoparreysia_tavoyensis **
| | | | +--------------+
| | | | | | +-- Leoparreysia_canefrii **
| | | | | +--+
| | | | | +-- Leoparreysia_olivacea **
+--+ | | |
| | | | +-------- Coelatura_aegyptiaca[2] **
| | | | |
| | | | +-------- Nitia_teretiuscula **
| | +--+ +--------+
| | | | | +----- Coelatura_aegyptiaca[1] **
| | | | | |
| | | | +--+ +-- Coelatura_gabonensis **
| | | | | |
| | | | +--+-- Coelatura_kunenensis **
| | | | |
| | | | +-- Prisodontopsis_mweruensis **
| | +--+
| | | +----- Indonaia_shurtleffiana **
| | | |
| | | | +-- Indonaia_bonneaudii **
| | | +--------+--+
| | | | | +-- Indonaia_rugosa **
| | | | |
| | | | | +-- Indonaia_andersoniana **
| | | | +--+
| | +--+ +-- Indonaia_caerulea **
| | |
| | | +----------- Indochinella_pugio **
| | | |
| | | | +----- Scabies_humilis **
| | +--+ +--+
| | | | | +-- Scabies_phaselus **
| | | | +--+
| | +--+ +-- Scabies_mandarinus **
| | |
| | | +----- Radiatula_mouhoti **
| | +--+
| | | +-- Radiatula_myitkyinae **
| | +--+
| | +-- Radiatula_chaudhurii **
| |
| | +-------------------- Chamberlainia_hainesiana **
| | |
| | +-------------------- Megalonaias_nervosa **
| | |
| | +-------------------- Uniomerus_carolinianus **
| | |
| | | +-- Quadrula_quadrula **
| | +-----------------+
| | | +-- Tritogonia_verrucosa **
| | |
| | | +-- Sinohyriopsis_cumingii **
| | +-----------------+
| | | +-- Sinohyriopsis_schlegelii **
| | |
| | | +-- Potomida_littoralis **
| | | +-----+
| | | | +-- Pronodularia_japanensis **
| | +-----------+
| | | | +----- Lamprotula_leaii **
| | | +--+
| | | | +-- Lamprotula_caveata **
| | | +--+
| | | +-- Lamprotula_cornuumlunae **
+--+ |
| | +-------- Sinosolenaia_carinata **
| | |
| | | +-- Sinosolenaia_recognita **
| | +-----+
| | | +-- Sinosolenaia_oleivora **
| | |
| +-----------+ +----- Gonidea_angulata **
| | +--+
| | | | +-- Leguminaia_wheatleyi **
| | | +--+
| | | +-- Microcondylaea_bonellii **
| | |
| | | +----- Ptychorhynchus_pfisteri **
| | +--+
| | | +-- Parvasolenaia_rivularis **
| | +--+
| | +-- Parvasolenaia_triangularis **
| |
| | +----- Pseudodon_manueli **
| | +-----+
| | | | +-- Pseudodon_inoscularis **
| | | +--+
| | | +-- Pseudodon_bogani **
| | |
| | | +-------- Pilsbryoconcha_exilis **
| | | |
| +--------+ +-------- Nyeinchanconcha_nyeinchani **
| | | |
| | | +-------- Thaiconcha_callifera **
| | | |
| | | +-------- Sundadontina_cumingii **
| | | |
| | | +-------- Sundadontina_tumida[1] **
| | +--+
| | +-------- Sundadontina_tumida[2] **
| | |
| | | +----- Monodontina_cambodjensis **
| | +--+
| | | | +-- Monodontina_mekongi **
| | | +--+
| | | +-- Monodontina_vondembuschiana **
| | |
| | | +----- Bineurus_loeiensis **
| | +--+
| | | +-- Bineurus_mouhotii **
| | +--+
| | +-- Bineurus_exilis **
| |
| | +-------------- Amblema_plicata **
| | |
| | +-------------- Toxolasma_parvum **
| | |
| | | +-- Obliquaria_reflexa **
| | +-----------+
| | | +-- Plectomerus_dombeyanus **
| | |
| | | +----------- Elliptoideus_sloatianus **
| | +--+
| | | | +-------- Elliptio_complanata **
| | | +--+
| | | | +----- Eurynia_dilatata **
| +-----+ +--+
| | | | +-- Fusconaia_flava **
| | | +--+
| | | +-- Pleurobema_sintoxia **
| | |
| | | +-------- Ellipsaria_lineolata **
| | | |
| | | +-------- Potamilus_leptodon **
| | | |
| | | +-------- Medionidus_conradicus **
| | | |
| | | +-------- Potamilus_alatus **
| | | |
| | | +-------- Ptychobranchus_subtentus **
+-----+ +-----+
| +-------- Truncilla_truncata **
| |
| +-------- Venustaconcha_ellipsiformis **
| |
| +-------- Cambarunio_iris **
| |
| +-------- Leaunio_lienosus **
| |
| | +----- Ortmanniana_ligamentina **
| +--+
| | +-- Lampsilis_cardium **
| +--+
| +-- Lampsilis_ornata **
|
| +-------------- Hyriopsis_desowitzi **
| |
| +-------------- Hyriopsis_myersiana **
| |
| +-------------- Hyriopsis_panhai **
| |
| +--+ +-- Hyriopsis_bialata **
| | +-----------+
| | | +-- Hyriopsis_khoratensis **
| | |
| | | +----------- Rectidens_sumatrensis **
| | | |
| | +--+ +-- Ensidens_telus **
| | | +-----+
| | | | +-- Ensidens_ingallsianus **
| | +--+
| | | +----- Ensidens_sagittarius **
| | +--+
+--+ | +-- Ensidens_spiculus **
| | +--+
| | +-- Ensidens_jaculus **
| |
| | +----------- Physunio_modelli **
| | |
| | +----------- Lens **
| | |
| | +----------- Physunio_superbus **
| | |
| | | +-------- Yaukthwa_inlenensis **
| | +--+
| +-----+ | +----- Yaukthwa_panhai **
| | +--+
| | | +-- Yaukthwa_nesemanni **
| | +--+
| | +-- Yaukthwa_rectangularis **
| |
| | +-- Lens_comptus **
| | +-----+
| | | +-- Lens_novoselovi **
| +--+
| | +----- Lens_eximius **
| +--+
| | +-- Lens_contradens **
| +--+
| +-- Lens_micropterus **
|
| +----------------- Acuticosta_chinensis **
| |
| +----------------- Alasmidonta_marginata **
| |
| +----------------- Anodonta_anatina **
| |
| +----------------- Anemina_arcaeformis **
| |
| +----------------- Anodonta_cygnea **
| |
| +----------------- Cristaria_plicata **
| |
| +----------------- Anodonta_nuttalliana **
| |
| +----------------- Lasmigona_compressa **
| |
| +----------------- Lepidodesma_languilati **
| |
| +----------------- Pseudanodonta_complanata **
| |
| +----------------- Pyganodon_grandis **
| |
| +----------------- Simpsonaias_ambigua **
| |
| +----------------- Strophitus_undulatus **
| |
| | +-- Schistodesmus_lampreyanus **
| +--------------+
| | +-- Schistodesmus_spinosus **
| |
| | +-- Utterbackia_imbecillis **
| +--------------+
| | +-- Utterbackia_peninsularis **
| |
| | +-------- Lanceolaria_gladiola **
| +--------+
| | | +----- Lanceolaria_grayii **
| | +--+
+--+ | +-- Lanceolaria_lanceolata **
| +--+
| +-- Lanceolaria_triformis **
|
| +-------- Unio_tumidus **
+--------+
| | +----- Unio_crassus **
| +--+
| | +-- Unio_delphinus **
| +--+
| +-- Unio_pictorum **
|
| +-------- Aculamprotula_tortuosa **
| |
+--------+ +-- Aculamprotula_coreana **
| | +--+
| | | +-- Aculamprotula_scripta **
| +--+
| | +-- Aculamprotula_fibrosa **
| +--+
| +-- Aculamprotula_tientsinensis **
|
| +-------- Cuneopsis_heudei **
| |
| +-------- Cuneopsis_celtiformis **
| |
+--------+-------- Tchangsinaia_piscicula **
| |
| +-------- Cuneopsis_rufescens **
| |
| | +----- Nodularia_douglasiae[2] **
| +--+
| | +-- Nodularia_douglasiae[1] **
| +--+
| +-- Nodularia_nuxpersicae **
|
| +-------------- Pletholophus_tenuis **
| |
+--+ +-- Sinanodonta_angula **
| +--------+
| | +-- Sinanodonta_lucida **
+--+
| +-------- Sinanodonta_woodiana[1] **
+--+
| +----- Sinanodonta[1] **
+--+
| +-- Sinanodonta_woodiana[2] **
+--+
+-- Sinanodonta[2] **
Huang et al., 2019, fig. S3
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 171 |
171 |
163 |
2 |
26 |
24 |
2 |
148.0 |
+----- Velesunio_ambiguus
+--+
| | +-- Alathyria_jacksoni
| +--+
+--------+ +-- Alathyria_profuga
| |
| | +----- Alathyria_pertexta
| +--+
| | +-- Lortiella_froggatti
+-----------------------------------------------+ +--+
| | +-- Microdontia_anodontaeformis
| |
| | +-------------- Westralunio_carteri
| | |
| +--+ +----------- Echyridella_menziesii
| | |
| | | +----- Diplodon_suavidicus
| +--+ +--+
| | | | +-- Castalia_ambigua
| | | +--+
| +--+ +-- Triplodon_corrugatus
| |
| | +----- Hyridella_australis
| +--+
| | +-- Hyridella_depressa
| +--+
| +-- Hyridella_drapeta
|
| +-------- Anodontites_trapesialis
| +--+
| | | +----- Mutela_rostrata
| | +--+
| | | +-- Mutela_dubia
| | +--+
| +--------------------------------------------------+ +-- Mutela_hargeri
| | |
| | | +----- Mycetopoda_siliquosa
--+ | | +--+
| | | | | +-- Anodontites_elongata
| | | | +--+
| | +--+ +-- Lamproscapha_ensiformis
| | |
| | | +----- Etheria_elliptica
| | +--+
| | | +-- Aspatharia_pfeifferiana
| | +--+
| | +-- Chambardia_wahlbergi
| |
| | +-------------------- Gibbosula_rochechouartii **
| | |
| | | +----------------- Gibbosula_laosensis **
| | +--------------------------------------+ |
| | | | | +-------- Cumberlandia_monodonta **
| | | | | +-----+
| | | +--+ | | +----- Pseudunio_marocanus **
| | | | | +--+
| | | | | | +-- Pseudunio_auricularius **
| | | | | +--+
| | | +--+ +-- Pseudunio_homsensis **
| | | |
| | | | +-- Margaritifera_dahurica **
| | | | +--------+
| | | | | +-- Margaritifera_margaritifera **
| | | +--+
| | | | +-------- Margaritifera_falcata **
+--+ | | |
| | +--+ +-- Margaritifera_hembeli **
| | | +--+
| | | | +-- Margaritifera_marrianae **
| | +--+
| | | +-- Margaritifera_laevis **
| | +--+
| | +-- Margaritifera_middendorffi **
| |
| | +----- Trapezidens_exolescens **
| | +-----+
| | | | +-- Trapezidens_angustior **
| | | +--+
| | | +-- Trapezidens_dolichorhynchus **
| | |
| | +--------------+ +-- Lamellidens_corrianus **
| | | | +-----+
| | | | | +-- Lamellidens_savadiensis **
| | | | |
| | | | | +----- Lamellidens_lamellatus **
| | | +--+ |
| | | | +----- Lamellidens_jenkinsianus **
| | | | |
| | | | | +-- Lamellidens_marginalis **
| | | +--+--+
| | | | +-- Lamellidens_ferrugineus **
| | | |
| | +-----------------------------+ | +-- Lamellidens_mainwaringi[1] **
| | | | | |
| | | | +--+-- Lamellidens_mainwaringi[2] **
| | | | |
+--+ | | +-- Lamellidens_generosus **
| | |
| | | +----- Parreysia_keralaensis **
| | | +-----------------+
| | | | | +-- Parreysia_nagpoorensis **
| | | | +--+
| | | | +-- Parreysia_plagiosoma **
| | | |
| | | | +----- Leoparreysia_canefrii **
| | +--+ +--------------+
| | | | | +-- Leoparreysia_tavoyensis **
| | | | +--+
| | | | +-- Leoparreysia_olivacea **
| | | |
| | | | +----------- Coelatura_aegyptiaca[2] **
| | | | |
| | +--+ +-----+ +-- Coelatura_aegyptiaca[1] **
| | | | | +-----+
| | | | | | +-- Nitia_teretiuscula **
| | | | +--+
| | | | | +----- Coelatura_kunenensis **
| | | | +--+
| | | | | +-- Coelatura_gabonensis **
| | | | +--+
| | +--+ +-- Prisodontopsis_mweruensis **
| | |
| | | +-------- Indonaia_shurtleffiana **
| | | |
| | | +-----+ +----- Indonaia_bonneaudii **
| | | | | |
| | | | +--+----- Indonaia_rugosa **
| | | | |
| | | | | +-- Indonaia_andersoniana **
| | +--+ +--+
| | | +-- Indonaia_caerulea **
| | |
| | | +----------- Indochinella_pugio **
| | | |
| | | | +----- Scabies_phaselus **
| | +--+ +--+
| | | | | +-- Scabies_humilis **
| | | | +--+
| | +--+ +-- Scabies_mandarinus **
| | |
| | | +----- Radiatula_mouhoti **
| | +--+
+--+ | +-- Radiatula_myitkyinae **
| +--+
| +-- Radiatula_chaudhurii **
|
| +----- Pseudodon_inoscularis **
| +--------------+
| | | +-- Pseudodon_bogani **
| | +--+
| | +-- Pseudodon_manueli **
| +--------------------------------+
| | | +----------------- Pilsbryoconcha_exilis **
| | | |
| | | | +----------- Sundadontina_tumida **
| | | | +--+
| | +--+ | | +-------- Monodontina_cambodjensis **
| | | | +--+
| | | | | +----- Monodontina_vondembuschiana[1] **
| | | | +--+
| | | | | +-- Monodontina_mekongi **
| | +--+ +--+
| | | +-- Monodontina_vondembuschiana[2] **
| | |
| | | +-- Nyeinchanconcha_nyeinchani **
| | | +--------+
| | | | +-- Sundadontina_cumingii **
| | +--+
| | | +-------- Thaiconcha_callifera **
| | +--+
| | | +----- Bineurus_loeiensis **
| | +--+
| | | +-- Bineurus_mouhotii **
| | +--+
| | +-- Bineurus_exilis **
| |
| | +-- Ensidens_telus **
| | +-----+
| | | +-- Ensidens_ingallsianus **
| | +-----+
| | | | +----- Ensidens_sagittarius **
| | | +--+
| | | | +-- Ensidens_spiculus **
| | +--+ +--+
| | | | +-- Ensidens_jaculus **
| | | |
| | | | +----------- Hyriopsis_desowitzi **
| | | | |
| | | +--+ +-------- Rectidens_sumatrensis **
| | | | |
| | | +--+ +-- Hyriopsis_panhai **
| | | | +--+
+--+ | | | +-- Hyriopsis_khoratensis **
| +--------------------------------+ +--+
| | | | +-- Hyriopsis_bialata **
| | | +--+
| | | +-- Hyriopsis_myersiana **
| | |
| | | +----- Physunio **
| | | +--------+
| | | | | +-- Physunio_superbus **
| | | | +--+
| | | | +-- Physunio_modelli **
| | | |
| | +--+ +-------- Yaukthwa_inlenensis **
| | | +--+
| | | | | +----- Yaukthwa_panhai **
| | | | +--+
| | | | | +-- Yaukthwa_nesemanni **
| | | | +--+
| | +--+ +-- Yaukthwa_rectangularis **
| | |
| | | +-- Lens_contradens **
| | | +-----+
| | | | +-- Lens_micropterus **
| | +--+
| | | +----- Lens_eximius **
| | +--+
| | | +-- Lens_novoselovi **
| | +--+
| | +-- Lens_comptus **
| |
| | +----- Chamberlainia_hainesiana **
| | +--------------------------------------+
| | | | +-- Sinohyriopsis_cumingii **
| | | +--+
| | | +-- Sinohyriopsis_schlegelii **
| | |
| | | +-- Potomida_littoralis **
| | | +-----+
| | | | +-- Pronodularia_japanensis **
| | | +-----+
| | | | | +----- Lamprotula_leaii **
| | | | +--+
+--+ | | | +-- Lamprotula_caveata **
| | | +--+
| +--+ +--------------------------+ +-- Lamprotula_cornuumlunae **
| | | | |
| | | | | +-- Sinosolenaia_recognita **
| | | | | +--------+
| | | | | | +-- Sinosolenaia_oleivora **
| | | | | |
| | | | +--+ +----- Gonidea_angulata **
| | | | | +--+
| | | | | | | +-- Leguminaia_wheatleyi **
| | | | | | +--+
| | | | +--+ +-- Microcondylaea_bonellii **
| | | | |
| | | | | +----- Ptychorhynchus_pfisteri **
| | +--+ +--+
| | | | +-- Parvasolenaia_rivularis **
| | | +--+
| | | +-- Parvasolenaia_triangularis **
| | |
| | | +-- Megalonaias_nervosa **
| | | +--+
| | | | +-- Uniomerus_carolinianus **
| | | +--------------------------------+
| | | | | +-- Quadrula_quadrula **
| | | | +--+
| | | | +-- Tritogonia_verrucosa **
| | | |
| | | | +-------- Elliptoideus_sloatianus **
| | +--+ |
| | | +--------------------------+ +-- Elliptio_complanata **
| | | | | +--+
| | | | | | +-- Eurynia_dilatata **
| | | | +--+
| | | | | +-- Fusconaia_flava **
| | +--+ +--+
| | | +-- Pleurobema_sintoxia **
| | |
| | | +-------------------------------- Amblema_plicata **
| | | |
| | +--+ +----------------------------- Toxolasma_parvum **
| | | |
+--+ +--+ +-- Obliquaria_reflexa **
| | +-----------------------+
| | | +-- Plectomerus_dombeyanus **
| +--+
| | +----------------------- Potamilus_leptodon **
| | |
| +--+ +-------------------- Potamilus_alatus **
| | |
| | | +-- Medionidus_conradicus **
| +--+ +--------------+
| | | +-- Ptychobranchus_subtentus **
| | |
| +--+ +-- Ellipsaria_lineolata **
| | +-----------+
| | | +-- Truncilla_truncata **
| +--+
| | +----------- Leaunio_lienosus **
| | |
| +--+ +-- Venustaconcha_ellipsiformis **
| | +-----+
| | | +-- Cambarunio_iris **
| +--+
| | +----- Ortmanniana_ligamentina **
| +--+
| | +-- Lampsilis_cardium **
| +--+
| +-- Lampsilis_ornata **
|
| +----------------------------------- Acuticosta_chinensis **
| |
| | +-------- Aculamprotula_tortuosa **
| | |
| | +-----------+ +-- Aculamprotula_coreana **
| | | | +--+
| | | | | +-- Aculamprotula_scripta **
| | | +--+
| | | | +-- Aculamprotula_fibrosa **
| | | +--+
| | | +-- Aculamprotula_tientsinensis **
| | +-----------+
| | | | +-------- Unio_tumidus **
+-----------+ | | +--------+
| | | | | +----- Unio_crassus **
| | | | +--+
| | | | | +-- Unio_delphinus **
| | | | +--+
| | +--+ +-- Unio_pictorum **
| | |
| | | +----- Nodularia_douglasiae[1] **
| | | +--------+
| | | | | +-- Nodularia_douglasiae[2] **
| | | | +--+
| | +--+ +-- Nodularia_nuxpersicae **
| | |
+--+ | +----------- Schistodesmus_lampreyanus **
| +--+
| | +-------- Tchangsinaia_piscicula **
| +--+
| | +----- Cuneopsis_rufescens **
| +--+
| | +-- Cuneopsis_heudei **
| +--+
| +-- Cuneopsis_celtiformis **
|
| +----------------------------- Lepidodesma_languilati **
| |
| | +-------- Lanceolaria_gladiola **
| | +-----------------+
| | | | +----- Lanceolaria_grayii **
| | | +--+
+--+ | | +-- Lanceolaria_lanceolata **
| | +--+
| | +-- Lanceolaria_triformis **
| |
| | +-------------------- Anemina_arcaeformis **
| | |
| | +--+ +----------------- Cristaria_plicata **
+--+ | | |
| | +--+ +-------------- Pletholophus_tenuis **
| | | |
| | +--+ +-- Sinanodonta_angula **
| | | +--------+
| | | | +-- Sinanodonta_lucida **
| | +--+
| | | +-------- Sinanodonta_woodiana[2] **
| | +--+
| | | +----- Sinanodonta[2] **
+--+ +--+
| | +-- Sinanodonta_woodiana[1] **
| +--+
| +-- Sinanodonta[1] **
|
| +-- Alasmidonta_marginata **
| +--------+
| | +-- Lasmigona_compressa **
| |
| | +----- Anodonta_anatina **
| +-----+
| | | +-- Anodonta_cygnea **
+-----------+ +--+
| +-- Anodonta_nuttalliana **
|
| +----- Pseudanodonta_complanata **
| +--+
| | | +-- Simpsonaias_ambigua **
| | +--+
+--+ +-- Strophitus_undulatus **
|
| +----- Pyganodon_grandis **
+--+
| +-- Utterbackia_imbecillis **
+--+
+-- Utterbackia_peninsularis **
Bolotov et al., 2017, fig. S1
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 125 |
125 |
68 |
1 |
0 |
0 |
0 |
124.5 |
+----- Trapezidens_exolescens **
+--------+
| | +-- Trapezidens_dolichorhynchus **
| +--+
+-----------+ +-- Trapezidens_angustior **
| |
| | +----------- Lamellidens_lamellatus **
| | |
| +--+ +-------- Lamellidens_jenkinsianus **
| | |
| | | +----- Lamellidens_generosus **
| +--+ |
| | +----- Lamellidens_mainwaringi **
| | |
| +--+ +-- Lamellidens_ferrugineus **
| +--+
| | +-- Lamellidens_marginalis **
+--+ |
| | | +-- Lamellidens_savadiensis **
| | +--+
| | +-- Lamellidens_corrianus[1] **
| |
| | +-------- Parreysia_keralaensis **
| | +--------------+
| | | | +----- Parreysia_plagiosoma **
| | | +--+
| | | | +-- Parreysia_corrugata **
| | | +--+
| | | +-- Parreysia_nagpoorensis **
| | |
| | | +-------- Leoparreysia_olivacea **
| +--+ +-----------+
| | | | +----- Leoparreysia_tavoyensis[2] **
| | | +--+
| | | | +-- Leoparreysia_canefrii **
| | | +--+
| | | +-- Leoparreysia_tavoyensis[1] **
| | |
| | | +-- Nitia_teretiuscula **
| | | |
| +--+ +-- Coelatura_gabonensis **
| | |
| | +-- Prisodontopsis_mweruensis **
| | +--------------+
| | | +-- Coelatura_kunenensis **
| | | |
| | | +-- Coelatura_aegyptiaca[1] **
| | | |
| | | +-- Coelatura_aegyptiaca[2] **
| | |
| +--+ +-------- Indonaia_shurtleffiana **
| | |
| | +-----+ +----- Indonaia_rugosa **
| | | | |
| | | | | +-- Indonaia_andersoniana **
| | | +--+--+
| | | | +-- Indonaia_caerulea **
| | | |
| +--+ | +-- Indonaia_bonneaudii **
| | +--+
| | +-- Lamellidens_corrianus[2] **
| |
| | +----------- Indochinella_pugio **
| | |
| | | +-- Scabies_humilis **
| +--+ |
| | +-----+-- Scabies_phaselus **
| | | |
--+ +--+ +-- Scabies_mandarinus **
| |
| | +----- Radiatula_mouhoti **
| +--+
| | +-- Radiatula_myitkyinae **
| +--+
| +-- Radiatula_chaudhurii **
|
| +----- Pseudodon_inoscularis **
| +--------+
| | | +-- Pseudodon_manueli **
| | +--+
| | +-- Pseudodon_bogani **
| |
| | +-- Thaiconcha_callifera **
| +--------+ +--------+
| | | | +-- Pilsbryoconcha_exilis **
| | | |
| | | | +-- Nyeinchanconcha_nyeinchani **
| | | +--------+
| | | | +-- Sundadontina_cumingii **
| | | |
| | +--+ +----- Bineurus_exilis **
| | +-----+
| | | | +-- Bineurus_mouhotii **
| | | +--+
| | | +-- Bineurus_loeiensis **
| | |
| | | +-------- Sundadontina_tumida **
| | +--+
| | | +----- Monodontina_cambodjensis **
| | +--+
| | | +-- Monodontina_mekongi **
| | +--+
| | +-- Monodontina_vondembuschiana **
| |
| | +-- Ensidens_telus **
| | +-----+
| | | +-- Ensidens_ingallsianus **
| | +--+
| | | | +----- Ensidens_jaculus **
| | | +--+
| | | | +-- Ensidens_spiculus **
| | | +--+
| | +-----+ +-- Ensidens_sagittarius **
| | | |
| | | | +-------- Hyriopsis_desowitzi **
| | | | |
| | | | +-------- Rectidens_sumatrensis **
| | | +--+
| | | | +----- Hyriopsis_panhai **
| | | | |
+-----+ | +--+ +-- Hyriopsis_myersiana **
| | | |
| +--+ +--+-- Hyriopsis_bialata **
| | | |
| | | +-- Hyriopsis_khoratensis **
| | |
| | | +----- Physunio **
| | | +--------+
| | | | | +-- Physunio_superbus **
| | | | +--+
| | | | +-- Physunio_modelli **
| | | |
| | +--+ +-------- Yaukthwa_inlenensis **
| | | +--+
| | | | | +----- Yaukthwa_panhai **
| | | | +--+
| | | | | +-- Yaukthwa_nesemanni **
| | | | +--+
| | +--+ +-- Yaukthwa_rectangularis **
| | |
| | | +-- Lens_contradens **
| | | +-----+
| | | | +-- Lens_micropterus **
| | +--+
| | | +----- Lens_eximius **
| | +--+
| | | +-- Lens_comptus **
| | +--+
| | +-- Lens_novoselovi **
| |
| | +-- Amblema_plicata **
| | |
| | +-- Ortmanniana_ligamentina **
| | |
| | +-- Lampsilis_cardium **
| | |
| | +-- Cambarunio_iris **
+--+ |
| +--+-- Elliptio_complanata **
| | |
| | +-- Eurynia_dilatata **
| | |
| | +-- Pleurobema_sintoxia **
| | |
| | +-- Quadrula_quadrula **
| | |
| | +-- Tritogonia_verrucosa **
| |
| | +-- Chamberlainia_hainesiana **
| +--+ |
| | | +-- Sinohyriopsis_cumingii **
| | | |
| | | +-- Lamprotula_caveata **
| | | |
| | | +-- Lamprotula_leaii **
| | | |
| | | +-- Potomida_littoralis **
| | | |
| | | +-- Pronodularia_japanensis **
| | | |
| | +--+-- Gonidea_angulata **
| | |
| | +-- Leguminaia_wheatleyi **
| | |
| | +-- Leguminaia **
| | |
| | +-- Microcondylaea_bonellii **
| | |
| | +-- Sinosolenaia_carinata **
| | |
| | +-- Sinosolenaia_oleivora **
| | |
| | +-- Sinosolenaia_recognita **
+-----------+
| +-- Unio_crassus **
| |
| +-- Unio_pictorum **
| |
| +-- Unio_tumidus **
| |
| +-- Aculamprotula_tortuosa **
| |
| +-- Cuneopsis_heudei **
| +--+
| | +-- Tchangsinaia_piscicula **
| | |
| | +-- Cuneopsis_rufescens **
| | |
| | +-- Nodularia_douglasiae **
| | |
| | +-- Nodularia_nuxpersicae **
| | |
| | +-- Schistodesmus_lampreyanus **
| |
| | +-- Alasmidonta_marginata **
| | |
| | +-- Anodonta_anatina **
+--+ |
| +-- Anodonta_cygnea **
| |
| +-- Anodonta_nuttalliana **
| |
| +-- Lasmigona_compressa **
| |
| +-- Pseudanodonta_complanata **
| |
| +-- Pyganodon_grandis **
| |
| +-- Simpsonaias_ambigua **
| |
| +-- Strophitus_undulatus **
+--+
+-- Anemina **
|
+-- Cristaria_plicata **
|
+-- Cristaria **
|
+-- Pletholophus_tenuis **
|
+-- Sinanodonta_lucida **
|
+-- Sinanodonta **
|
+-- Lanceolaria_gladiola **
|
+-- Lanceolaria_grayii[1] **
|
+-- Lanceolaria_grayii[2] **
Bolotov et al., 2017, fig. S1
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 123 |
136 |
133 |
2 |
15 |
13 |
2 |
108.5 |
+----------------- Cumberlandia_monodonta **
|
| +----- Gibbosula_laosensis **
+-----------------------+ +--------+
| | | | +-- Pseudunio_auricularius **
| | | +--+
| | | +-- Pseudunio_marocanus **
| +--+
| | +-- Margaritifera_dahurica **
| | +--------+
| | | +-- Margaritifera_margaritifera **
| +--+
| | +-------- Margaritifera_falcata **
| +--+
| | +----- Margaritifera_middendorffi **
| +--+
| | +-- Margaritifera_laevis **
| +--+
| +-- Margaritifera_marrianae **
|
| +-------- Anodontites_elongata
| +-----+
| | | +----- Etheria_elliptica
| | +--+
| | | +-- Chambardia_wahlbergi
| | +--+
| | +-- Aspatharia_pfeifferiana
| +-----------------------+
| | | +----- Diplodon_suavidicus
| | | +-----+
| | | | | +-- Castalia_ambigua
| | | | +--+
| | | | +-- Triplodon_corrugatus
| | +--+
--+ | | +----- Alathyria_pertexta
| | | +--+
| | | | | +-- Microdontia_anodontaeformis
| | | | +--+
| | +--+ +-- Lortiella_froggatti
| | |
| | | +----- Velesunio_ambiguus
| | +--+
| | | +-- Alathyria_jacksoni
| | +--+
| | +-- Alathyria_profuga
| |
| | +-- Trapezidens_dolichorhynchus **
| | +-----------+
| | | +-- Trapezidens_exolescens **
| | +-----------+
| | | | +----------- Lamellidens_corrianus **
| | | | |
| | | +--+ +-------- Lamellidens_savadiensis **
| | | | |
| | | +--+ +-- Lamellidens_jenkinsianus **
| | | | +--+
| | | | | +-- Lamellidens_mainwaringi **
| | | +--+
| | +--------+ | +-- Lamellidens_ferrugineus **
| | | | +--+
| | | | +-- Lamellidens_marginalis **
| | | |
| | | | +-- Parreysia_nagpoorensis **
| | | | +--------------------+
| | | | | +-- Parreysia_plagiosoma **
| | | | |
+--+ | | | +-- Leoparreysia_tavoyensis **
| | | | +-----------------+
| | +--+ | +-- Leoparreysia_olivacea **
| | | |
| | | | +-------------- Nitia_teretiuscula **
| | | | +--+
| | | | | | +----------- Coelatura_aegyptiaca[2] **
| | +--+ | +--+
| | | | | +-------- Coelatura_aegyptiaca[1] **
| | | | +--+
| | | | | +----- Prisodontopsis_mweruensis **
| | | | +--+
| | | | | +-- Coelatura_gabonensis **
| | +--+ +--+
| | | +-- Coelatura_kunenensis **
| | |
| | | +-------- Indonaia_shurtleffiana **
| | | +-----+
| | | | | +----- Indonaia_andersoniana **
| | | | +--+
| | | | | +-- Indonaia_rugosa **
| | | | +--+
| | +--+ +-- Indonaia_bonneaudii **
| | |
| | | +-- Indochinella_pugio **
| | | +--------+
| | | | +-- Scabies_crispata **
| | | |
| | +--+ +-- Radiatula_myitkyinae **
| | | +-----+
| | | | +-- Radiatula_chaudhurii **
| | +--+
| | | +----- Scabies_mandarinus **
| | +--+
| | | +-- Scabies_humilis **
| | +--+
| | +-- Scabies_phaselus **
| |
| | +-- Ensidens_telus **
+--+ +-----+
| | +-- Ensidens_ingallsianus **
| +-----+
| | | +----- Ensidens_sagittarius **
| | +--+
| | | +-- Ensidens_spiculus **
| | +--+
| +--+ +-- Ensidens_jaculus **
| | |
| | | +-- Hyriopsis_bialata **
| | | +--------+
| | | | +-- Hyriopsis_khoratensis **
| | +--+
| | | +-------- Hyriopsis_panhai **
| | +--+
| | | +----- Hyriopsis_myersiana **
| | +--+
| +--------------+ | +-- Rectidens_sumatrensis **
| | | +--+
| | | +-- Hyriopsis_desowitzi **
| | |
| | | +----- Physunio **
| | | +--------+
| | | | | +-- Physunio_modelli **
| | | | +--+
| | | | +-- Physunio_superbus **
| | | |
| | +--+ +-------- Yaukthwa_inlenensis **
| | | +--+
| | | | | +----- Yaukthwa_panhai **
| | | | +--+
| | | | | +-- Yaukthwa_rectangularis **
| | | | +--+
| | +--+ +-- Yaukthwa_nesemanni **
| | |
| | | +-- Lens_contradens **
| | | +-----+
| | | | +-- Lens_micropterus **
| | +--+
| | | +----- Lens_eximius **
| | +--+
| | | +-- Lens_comptus **
| | +--+
| | +-- Lens_novoselovi **
| |
| | +-------------------- Pseudodon_inoscularis **
| | |
+--+ | +----------------- Pilsbryoconcha_exilis **
| +--------+ |
| | | | +-------- Sundadontina_tumida **
| | | | +-----+
| | +--+ | | +----- Monodontina_cambodjensis **
| | | | +--+
| | | | | +-- Monodontina_mekongi **
| | | | +--+
| | +--+ +-- Monodontina_vondembuschiana **
| | |
| | | +-- Nyeinchanconcha_nyeinchani **
| | | +--------+
| | | | +-- Sundadontina_cumingii **
| | +--+
| | | +-------- Thaiconcha_callifera **
| | +--+
| | | +----- Bineurus_exilis **
| | +--+
| | | +-- Bineurus_mouhotii **
| | +--+
| | +-- Bineurus_loeiensis **
| |
| | +-------- Amblema_plicata **
| | +--+
| | | | +----- Ortmanniana_ligamentina **
| | | +--+
| | | | +-- Lampsilis_cardium **
| | | +--+
| | +-----+ +-- Cambarunio_iris **
| | | |
+--+ | | +-- Quadrula_quadrula **
| | | +-----+
| | | | +-- Tritogonia_verrucosa **
| | +--+
| | | +----- Pleurobema_sintoxia **
| | +--+
| | | +-- Eurynia_dilatata **
| | +--+
| | +-- Elliptio_complanata **
| +--------+
| | | +-- Sinohyriopsis_cumingii **
| | | +-----+
| | | | +-- Chamberlainia_hainesiana **
| | | |
| | | +-----+ +-- Potomida_littoralis **
| | | | | +--+
| | | | | | +-- Pronodularia_japanensis **
| | | | +--+
| | | | | +-- Lamprotula_caveata **
| | | | +--+
| | +--+ +-- Lamprotula_leaii **
| | |
| | | +----- Sinosolenaia_carinata **
| | | +-----+
| | | | | +-- Sinosolenaia_oleivora **
| | | | +--+
| | +--+ +-- Sinosolenaia_recognita **
| | |
| | | +-------- Gonidea_angulata **
| | +--+
+--+ | +----- Microcondylaea_bonellii **
| +--+
| | +-- Leguminaia_wheatleyi **
| +--+
| +-- Leguminaia **
|
| +----- Unio_tumidus **
| +--------+
| | | +-- Unio_pictorum **
| | +--+
| | +-- Unio_crassus **
| +--------+
| | | +----------- Aculamprotula_tortuosa **
| | | |
| | | | +-- Nodularia_douglasiae **
| | +--+ +-----+
| | | | +-- Nodularia_nuxpersicae **
| | | |
| | +--+ +-- Cuneopsis_rufescens **
| | | +--+
| | | | +-- Cuneopsis_heudei **
| | +--+
+--+ | +-- Tchangsinaia_piscicula **
| +--+
| +-- Schistodesmus_lampreyanus **
|
| +-- Lanceolaria_grayii **
| +-----------------+
| | +-- Lanceolaria_gladiola **
| |
| | +-- Cristaria_plicata **
| | +--------+
| | | +-- Cristaria **
+--+ +-----+
| | | +-------- Pletholophus_tenuis **
| | +--+
| | | +----- Sinanodonta_lucida **
| | +--+
| | | +-- Sinanodonta **
+--+ +--+
| +-- Anemina **
|
| +-------------- Anodonta_anatina **
| |
| | +----- Pseudanodonta_complanata **
+--+ +-----+
| | | +-- Anodonta_cygnea **
| | +--+
| | +-- Anodonta_nuttalliana **
+--+
| +-- Lasmigona_compressa **
| +-----+
| | +-- Alasmidonta_marginata **
+--+
| +----- Simpsonaias_ambigua **
+--+
| +-- Pyganodon_grandis **
+--+
+-- Strophitus_undulatus **
Bolotov et al., 2017, fig. S2
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 123 |
136 |
120 |
2 |
15 |
13 |
2 |
111.5 |
+-------------------- Chamberlainia_hainesiana **
|
+-----------+ +----------------- Cumberlandia_monodonta **
| | |
| | | +-------------- Gibbosula_laosensis **
| +--+ |
| | | +-- Pseudunio_auricularius **
| | +-----------+
| +--+ +-- Pseudunio_marocanus **
| |
| | +-- Margaritifera_dahurica **
| | +--------+
| | | +-- Margaritifera_margaritifera **
| +--+
| | +-------- Margaritifera_falcata **
| +--+
| | +----- Margaritifera_middendorffi **
| +--+
| | +-- Margaritifera_laevis **
| +--+
| +-- Margaritifera_marrianae **
|
| +-- Anodontites_elongata
| +--+
| | +-- Etheria_elliptica
| +--------+
| | | +-- Aspatharia_pfeifferiana
| | +--+
| | +-- Chambardia_wahlbergi
| |
+-----------------+ +----- Diplodon_suavidicus
| | +-----+
| | | | +-- Castalia_ambigua
| | | +--+
| | | +-- Triplodon_corrugatus
| +--+
| | +----- Velesunio_ambiguus
| | +--+
| | | | +-- Alathyria_jacksoni
| | | +--+
| +--+ +-- Alathyria_profuga
| |
| | +----- Microdontia_anodontaeformis
| +--+
| | +-- Alathyria_pertexta
| +--+
| +-- Lortiella_froggatti
|
| +-- Parreysia_nagpoorensis **
| +-----------+
| | +-- Parreysia_plagiosoma **
| |
| +--------+ +-- Trapezidens_dolichorhynchus **
| | | +--------+
| | | | +-- Trapezidens_exolescens **
| | | |
| | +--+ +-------- Lamellidens_savadiensis **
--+ | | |
| | | +-------- Lamellidens_corrianus **
| | +--+
| | | +-- Lamellidens_marginalis **
| | | +--+
| | | | +-- Lamellidens_ferrugineus **
| +-----+ +--+
| | | | +-- Lamellidens_jenkinsianus **
| | | +--+
| | | +-- Lamellidens_mainwaringi **
| | |
| | | +-- Leoparreysia_olivacea **
| | | +-----------------+
| | | | +-- Leoparreysia_tavoyensis **
| | | |
| | | | +-------------- Nitia_teretiuscula **
| | | | +--+
| | | | | | +----------- Coelatura_aegyptiaca[1] **
| | +--+ | +--+
| | | | | +-------- Coelatura_aegyptiaca[2] **
| | | | +--+
| | | | | +----- Coelatura_gabonensis **
| | | | +--+
| | | | | +-- Coelatura_kunenensis **
| | +--+ +--+
| | | +-- Prisodontopsis_mweruensis **
| | |
| | | +-------- Indonaia_shurtleffiana **
| | | +-----+
| | | | | +----- Indonaia_andersoniana **
| | | | +--+
| | | | | +-- Indonaia_rugosa **
| | | | +--+
| | +--+ +-- Indonaia_bonneaudii **
| | |
| | | +-- Indochinella_pugio **
| | | +--------+
| | | | +-- Scabies_crispata **
| | | |
| | +--+ +-- Radiatula_myitkyinae **
| | | +-----+
| | | | +-- Radiatula_chaudhurii **
| | +--+
| | | +----- Scabies_mandarinus **
| | +--+
| | | +-- Scabies_humilis **
| | +--+
| | +-- Scabies_phaselus **
| |
| | +----- Unio_tumidus **
| | +-----+
| | | | +-- Unio_crassus **
| | | +--+
| | | +-- Unio_pictorum **
| | +-----+
+--+ | | +-------- Aculamprotula_tortuosa **
| | | |
| | | +-------- Schistodesmus_lampreyanus **
| | | |
| | +--+ +-- Nodularia_douglasiae **
| | +-----+
| | | +-- Nodularia_nuxpersicae **
| | |
| | | +----- Cuneopsis_heudei **
| | +--+
| | | +-- Tchangsinaia_piscicula **
| | +--+
| +--------+ +-- Cuneopsis_rufescens **
| | |
| | | +-- Lanceolaria_grayii **
| | | +-----------+
| | | | +-- Lanceolaria_gladiola **
| | | |
| | | | +-- Cristaria_plicata **
| | | | +--------+
| | | | | +-- Cristaria **
| | | +--+
| | | | | +-------- Pletholophus_tenuis **
| | | | +--+
| | | | | +----- Sinanodonta_lucida **
| | +--+ +--+
| | | | +-- Anemina **
| | | +--+
| | | +-- Sinanodonta **
| | |
| | | +-------- Anodonta_anatina **
| | | |
| | | +-------- Pseudanodonta_complanata **
| | | |
| | | | +-- Anodonta_cygnea **
| | +-----+-----+
| | | +-- Anodonta_nuttalliana **
| | |
| | | +----- Pyganodon_grandis **
| | | |
| | | +----- Simpsonaias_ambigua **
| | +--+
| | +----- Strophitus_undulatus **
| | |
| | | +-- Alasmidonta_marginata **
| | +--+
| | +-- Lasmigona_compressa **
| |
| | +-------- Lens_contradens **
| | +-----+
+--+ | | +----- Lens_eximius **
| | +--+
| | | +-- Lens_comptus **
| | +--+
| +--+ +-- Lens_novoselovi **
| | |
| | | +----- Physunio **
| | | +-----+
| | | | | +-- Physunio_modelli **
| | | | +--+
| | +--+ +-- Physunio_superbus **
| | |
| | | +-------- Yaukthwa_inlenensis **
| | +--+
| | | +----- Yaukthwa_panhai **
| +-----+ +--+
| | | | +-- Yaukthwa_rectangularis **
| | | +--+
| | | +-- Yaukthwa_nesemanni **
| | |
| | | +----- Hyriopsis_panhai **
| | | |
| | | +--------+----- Hyriopsis_khoratensis **
| | | | |
| | | | | +-- Hyriopsis_myersiana **
| | | | +--+
| | +--+ +-- Hyriopsis_bialata **
| | |
| | | +----------- Hyriopsis_desowitzi **
| | | |
| | | +----------- Rectidens_sumatrensis **
| | +--+
| | | +-- Ensidens_telus **
| | | +-----+
| | | | +-- Ensidens_ingallsianus **
| | +--+
| | | +----- Ensidens_sagittarius **
| | +--+
+--+ | +-- Ensidens_spiculus **
| +--+
| +-- Ensidens_jaculus **
|
| +-------- Amblema_plicata **
| +--+
| | | +----- Cambarunio_iris **
| | +--+
| | | +-- Ortmanniana_ligamentina **
| | +--+
| +--------+ +-- Lampsilis_cardium **
| | |
| | | +-- Quadrula_quadrula **
| | | +-----+
| | | | +-- Tritogonia_verrucosa **
| | +--+
| | | +----- Pleurobema_sintoxia **
| | +--+
| | | +-- Elliptio_complanata **
| | +--+
| | +-- Eurynia_dilatata **
| |
| | +-------- Sinohyriopsis_cumingii **
| | |
+--+ +-----+ +-- Lamprotula_caveata **
| | | +--+
| | | | +-- Lamprotula_leaii **
| | +--+
| | | +-- Potomida_littoralis **
| +--+ +--+
| | | +-- Pronodularia_japanensis **
| | |
| | | +----------- Sinosolenaia_carinata **
| | | |
| | | | +-- Sinosolenaia_oleivora **
| | +--+--------+
| | | +-- Sinosolenaia_recognita **
| | |
| | | +-------- Gonidea_angulata **
+--+ +--+
| | +----- Microcondylaea_bonellii **
| +--+
| | +-- Leguminaia_wheatleyi **
| +--+
| +-- Leguminaia **
|
| +-------------- Pilsbryoconcha_exilis **
| |
| | +----------- Pseudodon_inoscularis[2] **
| | |
+--+ | +-------- Thaiconcha_callifera **
| +--+
| | | +----- Bineurus_exilis **
| | +--+
| | | +-- Bineurus_mouhotii **
+--+ +--+
| +-- Bineurus_loeiensis **
|
| +----- Monodontina_cambodjensis **
| +--+
| | | +-- Monodontina_mekongi **
| | +--+
| | +-- Monodontina_vondembuschiana **
+--+
| +-- Pseudodon_inoscularis[1] **
| +--+
| | +-- Sundadontina_tumida **
+--+
| +-- Nyeinchanconcha_nyeinchani **
+--+
+-- Sundadontina_cumingii **
Bolotov et al., 2020, fig. 1
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 121 |
121 |
100 |
2 |
0 |
0 |
0 |
156.0 |
+-- Gibbosula_laosensis **
|
+-- Margaritifera_dahurica **
|
+-----------------------------------------+-- Margaritifera_margaritifera **
| |
| +-- Margaritifera_laevis **
| |
| +-- Margaritifera_middendorffi **
|
| +-------- Trapezidens_exolescens **
| +--+
| | | +----- Trapezidens_scutum **
| | +--+
| | | +-- Trapezidens_dolichorhynchus **
| +--------------------------+ +--+
| | | +-- Trapezidens_angustior **
| | |
| | | +-------- Lamellidens_savadiensis **
| | +--+
| | | +----- Lamellidens_generosus **
| | +--+
| | | +-- Lamellidens_ferrugineus **
| | +--+
| | +-- Lamellidens_mainwaringi **
| |
| | +----------- Parreysia_keralaensis **
| | +--+
| +--+ | | +-------- Parreysia_plagiosoma **
| | | | +--+
| | | | | +----- Parreysia_nagpoorensis[2] **
| | | | +--+
| | | | | +-- Parreysia_nagpoorensis[1] **
| | | +--------------------+ +--+
| | | | | +-- Parreysia_rakhinensis **
| | | | |
| | | | | +-- Leoparreysia_burmana **
| | | | | +-----+
| | | | | | +-- Leoparreysia_whitteni **
| | | | +-----+
| | | | | +----- Leoparreysia_tavoyensis **
| | | | +--+
| | +--+ | +-- Leoparreysia_olivacea **
| | | +--+
| | | +-- Leoparreysia_canefrii **
| | |
| | | +-- Coelatura_aegyptiaca **
--+ | | +-----------------------------+
| | | | +-- Nitia_teretiuscula **
| | | |
| | | | +----------- Indonaia_shurtleffiana **
| | | | +-----------------+
| | +--+ | | +-------- Indonaia_rugosa **
| | | | +--+
| | | | | +----- Indonaia_andersoniana **
| | | | +--+
| | | | | +-- Indonaia_subclathrata **
| | +--+ +--+
| | | +-- Indonaia_caerulea **
| | |
| | | +-------------------------- Indochinella_pugio **
| | | |
| | | | +-------- Radiatula_crispisulcata **
| | | | +--------------+
| | +--+ | | +----- Radiatula_mouhoti **
| | | | +--+
| | | | | +-- Radiatula_chaudhurii **
| | | | +--+
| | +--+ +-- Radiatula_myitkyinae **
| | |
| | | +-------------------- Scabiellus_songkramensis **
| | | |
| | | | +-- Unionetta_fabagina **
| | +--+ +--------------+
| | | | +-- Harmandia_somboriensis **
| | | |
| | +--+ +-------------- Scabies_nucleus **
| | | |
| | | | +-- Scabies_crispata **
| | +--+ +--------+
| | | | +-- Scabies_mandarinus **
| | | |
| | +--+ +-- Scabies_humilis **
| | | +-----+
| | | | +-- Scabies_pilata **
| | +--+
| | | +----- Scabies_anceps **
| | +--+
| | | +-- Scabies_phaselus **
| | +--+
+--+ +-- Scabies_scobinatus **
|
| +----------- Hyriopsis_desowitzi **
| |
| +--+ +-------- Rectidens_sumatrensis **
| | | |
| | +--+ +-- Hyriopsis_khoratensis **
| | | +--+
| | | | +-- Hyriopsis_bialata **
| | +--+
| | | +-- Hyriopsis_myersiana **
| +--------+ +--+
| | | +-- Hyriopsis_panhai **
| | |
| | | +-- Ensidens_telus **
| | | +-----+
| | | | +-- Ensidens_ingallsianus **
| | | |
| | +-----+ +-- Ensidens_jaculus **
| | | +--+
| | | | +-- Ensidens_spiculus **
| | +--+
| | | +-- Ensidens_dugasti **
| | +--+
| | +-- Ensidens_sagittarius **
| +-----+
| | | +-- Trapezoideus_lenya **
| | | +-----+
| | | | +-- Trapezoideus_foliaceus **
| | | +-----------+
| | | | | +----- Physunio_superbus **
| | | | +--+
| | | | | +-- Physunio_modelli **
| | | | +--+
| | | | +-- Physunio **
| | | |
| | | | +-- Yaukthwa_elongatula **
| | | | +-----------+
| | | | | +-- Yaukthwa_baniensis **
| | +--+ |
| | | +--+ +----- Yaukthwa_inlenensis **
| | | | | +-----+
| | | | | | | +-- Yaukthwa_avaensis **
| | | | | | +--+
| | | | +--+ +-- Yaukthwa_paiensis **
| | | | |
| | | | | +-------- Yaukthwa_panhai **
| | | | +--+
| | | | | +----- Yaukthwa_dalliana **
| | +--+ +--+
| | | | +-- Yaukthwa_zayleymanensis **
| | | +--+
| | | +-- Yaukthwa_nesemanni **
| | |
| | | +----- Lens_misellus **
| | | +-----+
| | | | | +-- Lens_contradens **
| | | | +--+
| | | | +-- Lens_micropterus **
| | +-----+
| | | +----- Lens_pallegoixi **
| | | +--+
| | | | | +-- Lens_comptus **
+-----------+ | | +--+
| +--+ +-- Lens_novoselovi **
| |
| | +----- Lens_maenamensis **
| +--+
| | +-- Lens_eximius **
| +--+
| +-- Lens_rolfbrandti **
|
| +-- Sinosolenaia_iridinea **
| |
| +-- Sinosolenaia_recognita[1] **
| |
| +-- Sinosolenaia_carinata **
| |
| +-- Leguminaia_wheatleyi **
| |
| +-- Microcondylaea_bonellii **
| |
| +-- Gonidea_angulata **
| |
| +-- Ptychorhynchus_pfisteri **
| |
| +-- Parvasolenaia_rivularis **
| |
| +-----------------------+-- Lamprotula_leaii **
| | |
| | +-- Lamprotula **
| | |
| | +-- Lamprotula_caveata **
| | |
| | +-- Potomida_littoralis **
| | |
| | +-- Pronodularia_japanensis **
| | |
| | +-- Sinohyriopsis_cumingii **
| | |
| | +-- Chamberlainia_hainesiana **
+--+ |
| +-- Sinosolenaia_recognita[2] **
| |
| +-- Sinosolenaia_oleivora **
|
| +----------- Pseudodon_kayinensis **
| +-----------+
| | | +-------- Pseudodon_inoscularis **
| | +--+
| | | +----- Pseudodon_bogani **
| | +--+
| | | +-- Pseudodon_salwenianus **
| | +--+
+--+ +-- Pseudodon_manueli **
|
| +----- Bineurus_loeiensis **
| +--------------+
| | | +-- Bineurus_exilis **
| | +--+
| | +-- Bineurus_mouhotii **
| |
+--+ +----------------- Nyeinchanconcha_nyeinchani **
| |
| | +-- Thaiconcha_callifera **
| | +-----+
| | | +-- Thaiconcha_munelliptica **
+--+ +-----+
| | | +----- Pilsbryoconcha_exilis[2] **
| | +--+
| | | +-- Pilsbryoconcha_exilis[1] **
| | +--+
| | +-- Pilsbryoconcha **
+--+
| +-------- Monodontina_lenyanensis **
| +--+
| | | +----- Monodontina_cambodjensis **
| | +--+
| | | +-- Monodontina_vondembuschiana **
| | +--+
+--+ +-- Monodontina_mekongi **
|
| +-- Sundadontina_taskaevi **
| +-----+
| | +-- Sundadontina_brandti **
+--+
| +----- Sundadontina_tumida **
+--+
| +-- Sundadontina_cumingii **
+--+
+-- Sundadontina_tanintharyiensis **
Campbell et al., 2005, fig. 1
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 117 |
117 |
103 |
2 |
0 |
0 |
0 |
157.0 |
+-- Margaritifera_margaritifera **
+--------------------------------------------------+
| +-- Cumberlandia_monodonta **
|
| +-------------------------------------------------- Sinohyriopsis_cumingii **
| |
| | +----------- Anodonta_oregonensis **
| | |
| +--------------------------------------+ +-- Anodonta_cygnea **
| | | +-----+
| | | | +-- Lasmigona_etowaensis **
--+ | +--+
| | | +----- Pyganodon_grandis **
| | +--+
| | | +-- Lasmigona_costata **
| | +--+
| | +-- Pseudodontoideus_subvexus **
| |
| | +----- Gonidea_angulata **
| | +-----------------------------------------+
| | | | +-- Potomida_littoralis **
+--+ | +--+
| | +-- Pronodularia_japanensis **
| |
| | +-- Megalonaias_nervosa **
| | +-----------+
| | | +-- Uniomerus_declivis **
| | |
| | +-----------------------------+ +----------- Theliderma_metanevra **
| | | | |
| | | | | +----- Tritogonia_verrucosa **
| | | +--+ +--+
| | | | | | +-- Quadrula_quadrula **
| | | | | +--+
| | | | | +-- Tritogonia_nobilis **
| | | +--+
| | | | +----- Cyclonaias_kieneriana **
| | | | |
| | | | | +-- Cyclonaias_tuberculata **
+--+ | +--+--+
| | | +-- Cyclonaias_succissa **
| | |
| | | +-- Cyclonaias_infucata **
| | +--+
| | +-- Cyclonaias_kleiniana **
| |
| | +----------------------------------- Elliptoideus_sloatianus **
| | |
| | | +-------------------------------- Hemistena_lata **
| | | |
| | +-----+ | +----- Plethobasus_cyphyus **
| | | | | +-----------------------+
| | | | | | | +-- Elliptio_arca **
| | | | | | +--+
| | | | | | +-- Elliptio_crassidens **
| | | +--+ |
| | | | | +-- Eurynia_dilatata **
| | | | | +-----------------------+
| | | | | | +-- Parvaspina_collina **
+--+ | | | |
| | | | | +----- Pleuronaia_gibber **
| | | | +--------------------+
| | | | | | +-- Pleuronaia_barnesiana **
| | +--+ | +--+
| | | | +-- Pleuronaia_dolabelloides **
| | | |
| | | | +----------- Fusconaia_subrotunda **
| | | | |
| | | | | +-- Fusconaia_cor **
| | | +--------------+ +-----+
| | | | | | +-- Fusconaia_cuneolus **
| | | | | |
| | | | | | +-- Fusconaia_cerina[1] **
| | | | +--+ |
| | | | | +--+-- Fusconaia_cerina[2] **
| | +--+ | | |
| | | | | +-- Fusconaia_flava **
| | | +--+
| | | | +-- Fusconaia_burkei[1] **
| | | | |
| | | | +-- Fusconaia_burkei[2] **
| | | +--+
| | | +-- Fusconaia_burkei[3] **
| | | |
| | | +-- Fusconaia_escambia **
| | |
| | | +-- Pleurobema_rubrum[2] **
| | | +--+
| | | | +-- Pleurobema_cordatum **
| | | +-----------------+
| | | | | +-- Pleurobema_rubrum[1] **
+--+ | | +--+
| | | +-- Pleurobema_sintoxia **
| +--+
| | +-------------------- Pleurobema_pyriforme **
| | |
| | | +-- Pleurobema_georgianum[2] **
| | | |
| +--+ +--------------+-- Pleurobema_georgianum[3] **
| | | |
| | | +-- Pleurobema_athearni **
| | |
| | | +----- Pleurobema_decisum **
| +--+ |
| | +--------+ +-- Pleurobema_hanleyianum[1] **
| | | | |
| | | +--+-- Pleurobema_hanleyianum[2] **
| | | |
| | | +-- Pleurobema_georgianum[1] **
| +--+
| | +-- Pleurobema_oviforme[1] **
| | |
| | +--------+-- Pleurobema_oviforme[2] **
| | | |
| +--+ +-- Pleurobema_clava **
| |
| | +-------- Pleurobema_strodeanum **
| +--+
| | +----- Pleurobema_rubellum **
| +--+
| | +-- Pleurobema_perovatum **
| +--+
| +-- Pleurobema_taitianum **
|
| +----- Plectomerus_dombeyanus **
| +--------------------------------+
| | | +-- Reginaia_ebenus **
| | +--+
| | +-- Reginaia_rotulata **
| |
| | +----- Popenaias_popeii **
| | +-----------------------------+
| | | | +-- Amblema_elliottii **
+--+ | +--+
| | +-- Amblema_plicata **
| |
| | +-- Toxolasma_parvum **
| | +-----+
| | | +-- Toxolasma_texasiense **
| | +-----------------------+
| | | | +----- Obliquaria_reflexa **
+--+ | +--+
| | | +-- Cyrtonaias_tampicoensis **
| | +--+
| | +-- Glebula_rotundata **
| |
| | +-- Potamilus_fragilis **
| | +--+
| | | +-- Potamilus_leptodon **
| | +-----------+
| | | | +-- Potamilus_alatus **
| | | +--+
+--+ | +-- Potamilus_purpuratus **
| |
| +-----------+ +-- Sagittunio_nasutus **
| | | +--+
| | | | +-- Medionidus_conradicus **
| | | +--------+
| | | | | +-- Ptychobranchus_fasciolaris **
| | | | +--+
| | +--+ +-- Venustaconcha_ellipsiformis **
| | |
| | | +----------- Medionidus_acutissimus **
| | +--+
| | | +-------- Lemiox_rimosus **
| | +--+
+--+ | +----- Dromus_dromas **
| +--+
| | +-- Cyprogenia_aberti **
| +--+
| +-- Cyprogenia_stegaria **
|
| +-- Ellipsaria_lineolata **
| +-----------------------+
| | +-- Truncilla_truncata **
| |
| | +----------------------- Villosa_villosa **
| | |
| | | +-- Obovaria_olivaria **
+--+ | +--------------+
| | | +-- Venustaconcha_pleasii **
| | |
| | +--+ +----- Obovaria_unicolor **
| | | | +--------+
| | | | | | +-- Obovaria_arkansasensis **
+--+ | | | +--+
| | +--+ +-- Obovaria_subrotunda **
| | |
| | | +----------- Epioblasma_brevidens **
| | +--+
| | | +-------- Epioblasma_triquetra **
| | +--+
| | | +----- Epioblasma_walkeri[2] **
| | +--+
| | | +-- Epioblasma_capsaeformis **
+--+ +--+
| +-- Epioblasma_walkeri[1] **
|
| +-- Ligumia_recta **
| +--------+
| | +-- Cambarunio_iris **
| +--+
| | | +-------- Lampsilis_teres **
| | +--+
| | | +----- Ortmanniana_ligamentina **
| | +--+
| | | +-- Lampsilis_siliquoidea **
| | +--+
| | +-- Leaunio_vanuxemensis **
+-----+
| +-- Lampsilis_cardium **
| +--+
| | +-- Lampsilis_ovata **
| +-----+
| | | +-- Ortmanniana_pectorosa **
| | +--+
| | +-- Lampsilis_ornata **
| |
+--+ +----- Hamiota_australis[2] **
| +--+
| | | +-- Hamiota_australis[1] **
| | +--+
| | +-- Hamiota_subangulata **
+--+
| +----- Hamiota_altilis[2] **
| |
| | +-- Hamiota_altilis[1] **
+--+--+
| +-- Hamiota_perovalis[1] **
|
| +-- Hamiota_altilis[3] **
+--+
+-- Hamiota_perovalis[2] **
Campbell et al., 2005, fig. 2
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 115 |
115 |
92 |
2 |
0 |
0 |
0 |
159.0 |
+-- Cumberlandia_monodonta **
+--------------------------------------------+
| +-- Margaritifera_margaritifera **
|
| +----------- Sinohyriopsis_cumingii **
| |
| +--------------------------------+ +-------- Pyganodon_grandis **
| | | |
| | | | +-- Lasmigona_costata **
| | +--+-----+
| | | +-- Pseudodontoideus_subvexus **
--+ | |
| | | +----- Lasmigona_etowaensis **
| | +--+
| | | +-- Anodonta_cygnea **
| | +--+
| | +-- Anodonta_oregonensis **
| |
| | +----- Gonidea_angulata **
| | +-----------------------------------+
| | | | +-- Potomida_littoralis **
| | | +--+
+--+ | +-- Pronodularia_japanensis **
| |
| | +-------------------- Megalonaias_nervosa **
| | |
| | +-----------------+ +----------------- Uniomerus_declivis **
| | | | |
| | | +--+ +-------------- Theliderma_metanevra **
| | | | |
| | | +--+ +----------- Quadrula_quadrula **
| | | | |
| | | | | +-- Tritogonia_nobilis **
| | | +--+ +-----+
| | | | | +-- Tritogonia_verrucosa **
| | | | |
| | | +--+ +----- Cyclonaias_kieneriana **
| | | | |
+--+ | | | +-- Cyclonaias_tuberculata **
| | +--+--+
| | | +-- Cyclonaias_succissa **
| | |
| | | +-- Cyclonaias_infucata **
| | +--+
| | +-- Cyclonaias_kleiniana **
| |
| | +-------------------------------- Elliptoideus_sloatianus **
| | |
| | | +----------------------------- Plethobasus_cyphyus **
| | | |
| | | +----------------------------- Parvaspina_collina **
| | +--+ |
| | | | | +-- Elliptio_arca **
| | | | +--------------------------+
| | | | | +-- Elliptio_crassidens **
| | | | |
| | | | | +-------------------------- Eurynia_dilatata **
| | | +--+ |
+--+ | | +-------------------------- Hemistena_lata **
| | | |
| | | | +-- Pleurobema_rubrum[1] **
| | | | +--+
| | | | | +-- Pleurobema_sintoxia **
| | | | +-----------------+
| | | | | | +-- Pleurobema_rubrum[2] **
| | | | | +--+
| | +--+ | +-- Pleurobema_cordatum **
| | | |
| | | | +-------------- Fusconaia_subrotunda **
| | | | |
| | | | | +-- Fusconaia_cor **
| | | +--------+ +--------+
| | | | | | +-- Fusconaia_cuneolus **
| | | | | |
| | | | +--+ +-- Fusconaia_cerina **
| | | | | +-----+
| | | | | | +-- Fusconaia_flava **
| | +--+ +--+
| | | | +----- Fusconaia_burkei[2] **
| | | +--+
| | | | +-- Fusconaia_escambia **
| | | +--+
| | | +-- Fusconaia_burkei[1] **
| | |
| | | +----- Pleuronaia_gibber **
| | | +--------------+
| | | | | +-- Pleuronaia_barnesiana **
| | | | +--+
| | | | +-- Pleuronaia_dolabelloides **
| | | |
| | | | +-------- Pleurobema_rubellum **
+--+ +--+ +--------+
| | | | +----- Pleurobema_strodeanum **
| | | +--+
| | | | +-- Pleurobema_perovatum **
| | | +--+
| | | +-- Pleurobema_taitianum **
| +--+
| | +-- Pleurobema_oviforme[1] **
| | |
| | +-----------+-- Pleurobema_oviforme[2] **
| | | |
| | | +-- Pleurobema_clava **
| +--+
| | +----------- Pleurobema_pyriforme **
| | |
| +--+ +-- Pleurobema_athearni **
| | +-----+
| | | +-- Pleurobema_georgianum[2] **
| +--+
| | +----- Pleurobema_decisum **
| | |
| +--+ +-- Pleurobema_hanleyianum[1] **
| | |
| +--+-- Pleurobema_hanleyianum[2] **
| |
| +-- Pleurobema_georgianum[1] **
|
| +-------- Popenaias_popeii **
| |
| +--------------+ +-- Amblema_elliottii **
| | | +--+
| | | | +-- Amblema_plicata **
| | +--+
| | | +-- Reginaia_ebenus **
| | | |
| | +--+-- Reginaia_rotulata[1] **
| | |
| | +-- Reginaia_rotulata[2] **
| |
| | +----- Plectomerus_dombeyanus **
| | +--------------+
+-----------+ | | +-- Toxolasma_parvum **
| | +--+
| | +-- Toxolasma_texasiense **
| |
| | +----- Obliquaria_reflexa **
| | +-----------+
| | | | +-- Cyrtonaias_tampicoensis **
| | | +--+
| | | +-- Glebula_rotundata **
| | |
| | | +-------------- Sagittunio_nasutus **
| | | |
+--+ | +-------------- Leaunio_vanuxemensis **
| | |
| | | +-- Ellipsaria_lineolata **
| | +-----------+
| | | +-- Truncilla_truncata **
| | |
| | | +----- Lampsilis_teres **
| | +--------+
| | | | +-- Ortmanniana_ligamentina **
| | | +--+
| | | +-- Lampsilis_siliquoidea **
| | |
| | | +----- Villosa_villosa **
| | +--------+
+--+ | | +-- Ligumia_recta **
| | +--+
| | +-- Cambarunio_iris **
| |
| | +-------- Ortmanniana_pectorosa **
| +-----+
| | | +----- Lampsilis_ornata **
| | +--+
| | | +-- Lampsilis_cardium **
| | +--+
| | +-- Lampsilis_ovata **
| |
| | +-- Potamilus_fragilis **
| | +--+
| | | +-- Potamilus_leptodon **
| +--------+
| | | +-- Potamilus_alatus **
| | +--+
| | +-- Potamilus_purpuratus **
| |
| | +-- Ptychobranchus_fasciolaris **
+--+ +--------+
| | +-- Venustaconcha_ellipsiformis **
| |
+--+ +-------- Medionidus_conradicus **
| | |
| | | +-- Lemiox_rimosus **
| +--+-----+
| | +-- Medionidus_acutissimus **
| |
| | +----- Dromus_dromas **
| +--+
| | +-- Cyprogenia_aberti **
| +--+
| +-- Cyprogenia_stegaria **
|
| +----- Hamiota_australis[2] **
| +-----+
| | | +-- Hamiota_australis[1] **
| | +--+
| | +-- Hamiota_subangulata **
+--+
| | +-- Hamiota_altilis[3] **
| | +-----+
| | | +-- Hamiota_perovalis[2] **
| +--+
| | +----- Hamiota_altilis[2] **
| +--+
| | +-- Hamiota_altilis[1] **
| +--+
| +-- Hamiota_perovalis[1] **
|
| +----------- Obovaria_olivaria **
| |
| +----------- Venustaconcha_pleasii **
| |
| | +----- Obovaria_unicolor **
| +-----+
+--+ | +-- Obovaria_arkansasensis **
| +--+
| +-- Obovaria_subrotunda **
|
| +-- Epioblasma_brevidens **
| +-----+
| | +-- Epioblasma_triquetra **
+--+
| +----- Epioblasma_walkeri[1] **
+--+
| +-- Epioblasma_walkeri[2] **
+--+
+-- Epioblasma_capsaeformis **
Bolotov et al., 2020, fig. S1
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 115 |
115 |
100 |
1 |
0 |
0 |
0 |
150.0 |
+-------- Trapezidens_exolescens **
+--+
| | +----- Trapezidens_scutum **
| +--+
| | +-- Trapezidens_dolichorhynchus **
+--------------------------------+ +--+
| | +-- Trapezidens_angustior **
| |
| | +-------- Lamellidens_savadiensis **
| +--+
| | +----- Lamellidens_generosus **
| +--+
| | +-- Lamellidens_ferrugineus **
| +--+
| +-- Lamellidens_mainwaringi **
|
| +----------- Parreysia_keralaensis **
| +--+
+--+ | | +-------- Parreysia_nagpoorensis[2] **
| | | +--+
| | | | +----- Parreysia_nagpoorensis[1] **
| | | +--+
| | | | +-- Parreysia_rakhinensis **
| | +--------------------------+ +--+
| | | | +-- Parreysia_plagiosoma **
| | | |
| | | | +-- Leoparreysia_burmana **
| | | | +-----+
| | | | | +-- Leoparreysia_whitteni **
| | | +-----+
| | | | +----- Leoparreysia_olivacea **
| | | +--+
| +--+ | +-- Leoparreysia_tavoyensis **
| | +--+
| | +-- Leoparreysia_canefrii **
| |
| | +-- Coelatura_aegyptiaca **
| | +-----------------------------------+
| | | +-- Nitia_teretiuscula **
| | |
| | | +-------- Indonaia_shurtleffiana **
| +--+ +--------------------------+
| | | | +----- Indonaia_andersoniana **
| | | +--+
| | | | +-- Indonaia_caerulea **
| | | +--+
| +--+ +-- Indonaia_subclathrata **
| |
| | +-------------------------------- Indochinella_pugio **
| | |
| | | +----- Radiatula_mouhoti **
| +--+ +-----------------------+
| | | | +-- Radiatula_myitkyinae **
| | | +--+
| +--+ +-- Radiatula_chaudhurii **
| |
| | +-------------------------- Radiatula_crispisulcata **
| | |
| +--+ +----------------------- Scabiellus_songkramensis **
| | |
| +--+ +-------------------- Harmandia_somboriensis **
| | |
| +--+ +----------------- Unionetta_fabagina **
| | |
| +--+ +-------------- Scabies_mandarinus **
| | |
| +--+ +----------- Scabies_crispata **
| | |
| +--+ +-------- Scabies_nucleus **
| | |
| +--+ +-- Scabies_humilis **
| | +--+
| | | +-- Scabies_pilata **
| +--+
| | +-- Scabies_phaselus **
| | |
| +--+-- Scabies_scobinatus **
| |
--+ +-- Scabies_anceps **
|
| +-------------- Hyriopsis_desowitzi **
| +--+
| | | +----------- Rectidens_sumatrensis **
| | +--+
| | | +-------- Hyriopsis_panhai **
| | +--+
| | | +----- Hyriopsis_myersiana **
| | +--+
| +--------+ | +-- Hyriopsis_khoratensis **
| | | +--+
| | | +-- Hyriopsis_bialata **
| | |
| | | +-- Ensidens_telus **
| | | +-----+
| | | | +-- Ensidens_ingallsianus **
| | | |
| | +--------+ +-- Ensidens_dugasti **
| | | +--+
| | | | +-- Ensidens_sagittarius **
| | +--+
| +--+ | +-- Ensidens_spiculus **
| | | +--+
| | | +-- Ensidens_jaculus **
| | |
| | | +----- Physunio **
| | | +-----------------+
| | | | | +-- Physunio_modelli **
| | | | +--+
| | | | +-- Physunio_superbus **
| | | |
| | | | +-- Trapezoideus_lenya **
| | | | +-----------------+
| | +--+ | +-- Trapezoideus_foliaceus **
| | | |
| | | | +----- Lens_misellus **
| | | | +-----+
| | | | | | +-- Lens_contradens **
| | | | | +--+
| | | | | +-- Lens_micropterus **
| | +--+ +-----+
| | | | | +----- Lens_maenamensis **
| | | | | +--+
| | | | | | | +-- Lens_eximius **
| | | | | | +--+
| | | | +--+ +-- Lens_rolfbrandti **
| | | | |
| | | | | +----- Lens_pallegoixi **
| | +--+ +--+
| | | | +-- Lens_novoselovi **
| | | +--+
| | | +-- Lens_comptus **
| | |
| | | +-- Yaukthwa_elongatula **
| | | +-----------+
| | | | +-- Yaukthwa_baniensis **
| | | |
| | +--+ +----- Yaukthwa_inlenensis **
| | | +-----+
+-----------------+ | | | +-- Yaukthwa_avaensis **
| | | +--+
| +--+ +-- Yaukthwa_paiensis **
| |
| | +-------- Yaukthwa_panhai **
| +--+
| | +----- Yaukthwa_dalliana **
| +--+
| | +-- Yaukthwa_zayleymanensis **
| +--+
| +-- Yaukthwa_nesemanni **
|
| +-- Sinohyriopsis_cumingii **
| +--+
| | +-- Chamberlainia_hainesiana **
| |
| +--+ +-- Lamprotula_leaii **
| | | |
| | | +-- Lamprotula **
| | | |
| | +--+-- Lamprotula_caveata **
| | |
| | +-- Potomida_littoralis **
| | |
| | +-- Pronodularia_japanensis **
| +-----------------+
| | | +-- Sinosolenaia_iridinea **
| | | |
| | | +-- Sinosolenaia_recognita[1] **
| | | |
| | | +-- Sinosolenaia_carinata **
| | | |
| | | +-- Leguminaia_wheatleyi **
| | | |
| | | +-- Microcondylaea_bonellii **
| | +-----+
| | +-- Gonidea_angulata **
| | |
| | +-- Ptychorhynchus_pfisteri **
+--+ |
| +-- Parvasolenaia_rivularis **
| |
| +-- Sinosolenaia_recognita[2] **
| |
| +-- Sinosolenaia_oleivora **
|
| +----------- Pseudodon_kayinensis **
| +-----------+
| | | +-------- Pseudodon_inoscularis **
| | +--+
| | | +----- Pseudodon_bogani **
| | +--+
| | | +-- Pseudodon_salwenianus **
| | +--+
+--+ +-- Pseudodon_manueli **
|
| +----- Bineurus_loeiensis **
| +--------------+
| | | +-- Bineurus_exilis **
| | +--+
| | +-- Bineurus_mouhotii **
| |
+--+ +----------------- Nyeinchanconcha_nyeinchani **
| |
| | +-- Thaiconcha_callifera **
| | +-----+
| | | +-- Thaiconcha_munelliptica **
+--+ +-----+
| | | +----- Pilsbryoconcha_exilis[2] **
| | +--+
| | | +-- Pilsbryoconcha_exilis[1] **
| | +--+
| | +-- Pilsbryoconcha **
+--+
| +-------- Monodontina_lenyanensis **
| +--+
| | | +----- Monodontina_cambodjensis **
| | +--+
| | | +-- Monodontina_vondembuschiana **
| | +--+
+--+ +-- Monodontina_mekongi **
|
| +-- Sundadontina_tanintharyiensis **
| +-----+
| | +-- Sundadontina_cumingii **
+--+
| +----- Sundadontina_tumida **
+--+
| +-- Sundadontina_taskaevi **
+--+
+-- Sundadontina_brandti **
Lopes-Lima et al., 2020, fig. 2-4
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 113 |
118 |
97 |
2 |
9 |
7 |
2 |
100.0 |
+-- Hyridella_depressa
+--------------------------------------------+
| +-- Velesunio_ambiguus
|
| +-- Margaritifera_laevis **
| +-----------------------------------------+
| | +-- Margaritifera_middendorffi **
| |
| | +-- Lamproscapha_ensiformis
--+ | +-----+
| | | +-- Mycetopoda_siliquosa
| | +--------------------------------+
| | | | +----- Etheria_elliptica
| | | +--+
| | | | +-- Aspatharia_pfeifferiana
| | | +--+
| | | +-- Chambardia_wahlbergi
| | |
+--+ | +-- Coelatura_aegyptiaca **
| | |
| | +-----------------------------------+-- Leoparreysia_tavoyensis **
| | | |
| | | +-- Radiatula_mouhoti **
| | |
| | | +-- Lens_contradens **
| | | |
| | | +-- Ensidens_sagittarius **
| | | |
| | | +-----------------------+-- Physunio_modelli **
| | | | |
| | | | +-- Rectidens_sumatrensis **
| | | | |
+--+ | | +-- Yaukthwa_nesemanni **
| | |
| | | +-- Amblema_plicata **
| | | |
| | | +-- Elliptio_complanata **
| | | +--------------------+
| | +--------+ | +-- Lampsilis_cardium **
| | | | | |
| | | | | +-- Quadrula_quadrula **
| | | | |
| | | | | +----- Chamberlainia_hainesiana **
| | | | | +-----------+
| | | | | | | +-- Sinohyriopsis_cumingii **
| | | | | | +--+
| | | | | +--+ +-- Sinohyriopsis_schlegelii **
| | | | | | |
| | | +--+ | | +-------------- Pronodularia_japanensis **
| | | | | | |
| | | | | +--+ +----------- Potomida_littoralis **
| | | | | | |
| | | | | +--+ +-------- Lamprotula_caveata **
+--+ | | | | |
| | | | +--+ +----- Lamprotula_leaii[1] **
| | | | | |
| | | | +--+----- Lamprotula_leaii[2] **
| | | | |
| | | | | +-- Lamprotula_seomjinensis **
| | | | +--+
| | +--+ +-- Lamprotula **
| | |
| | | +-- Pseudodon_inoscularis **
| | | +--------+
| | | | +-- Pseudodon_manueli **
| | | +-----+
| | | | | +-------- Pilsbryoconcha_exilis **
| | | | | |
| | | | +--+ +----- Monodontina_mekongi **
| | | | | |
| | | | +--+ +-- Nyeinchanconcha_nyeinchani **
| | | | | |
| | | | +--+-- Bineurus_mouhotii **
| | | | |
| | +--+ +-- Sundadontina_tumida **
| | |
| | | +-------- Inversidens_brandtii **
| | | |
| | | +-----+ +-- Obovalis_omiensis **
| | | | | +--+
| | | | | | +-- Ptychorhynchus_pfisteri **
| | | | +--+
| | | | | +-- Parvasolenaia_rivularis **
| | | | +--+
| | +--+ +-- Koreosolenaia_sitgyensis **
| | |
| | | +----- Gonidea_angulata **
+--+ | +-----+
| | | | +-- Leguminaia_wheatleyi **
| | | +--+
| +--+ +-- Microcondylaea_bonellii **
| |
| | +-------- Sinosolenaia_carinata **
| +--+
| | +----- Sinosolenaia_recognita **
| +--+
| | +-- Sinosolenaia_iridinea **
| +--+
| +-- Sinosolenaia_oleivora **
|
| +-------------------------------- Acuticosta_chinensis **
| |
| +-------------------------------- Lepidodesma_languilati **
| |
| | +----------- Aculamprotula_tortuosa **
| | +--------------+
| | | | +-------- Aculamprotula_scripta **
| | | +--+
| | | | +----- Aculamprotula_coreana **
| | | +--+
| | | | +-- Aculamprotula_fibrosa **
| | | +--+
| +-----+ +-- Aculamprotula_tientsinensis **
| | |
| | | +----------------------- Middendorffinaia_mongolica **
| | | |
| | | | +----- Unio_gibbus **
| | | | |
| | | | +--------------+----- Unio_tumidus **
| | | | | |
| | +--+ | | +-- Unio_crassus **
| | | | +--+
| | | | +-- Unio_pictorum **
| | | |
| | | | +-------------- Nodularia_breviconcha **
| | | | +--+
| | +--+ | | +----------- Nodularia[2] **
| | | | +--+
| | | | | +-------- Nodularia_nuxpersicae **
| | | | +--+
| | | | | +----- Nodularia_nipponensis **
| | | | +--+
| | | | | +-- Nodularia_douglasiae **
| | | | +--+
| | +--+ +-- Nodularia[1] **
| | |
| | | +----- Cuneopsis_rufescens **
+--+ | +--------+
| | | | +-- Cuneopsis_heudei **
| | | +--+
| | | +-- Cuneopsis_celtiformis **
| | |
| | | +----- Schistodesmus_spinosus **
| +--+--------+
| | | +-- Schistodesmus_lampreyanus **
| | +--+
| | +-- Schistodesmus **
| |
| | +----------- Tchangsinaia_piscicula **
| +--+
| | +-------- Inversiunio_jokohamensis **
| +--+
| | +----- Inversiunio_verrucosus **
| +--+
| | +-- Inversiunio_reinianus **
| +--+
| +-- Inversiunio_yanagawensis **
|
| +----------------- Lanceolaria_oxyrhyncha **
| |
| +-----------+ +-------------- Lanceolaria_gladiola **
| | | |
| | +--+ +-- Lanceolaria_bilirata **
| | | +--------+
| | | | +-- Lanceolaria_grayii **
| | +--+
| | | +-------- Lanceolaria_acrorrhyncha **
| | +--+
| | | +----- Lanceolaria_kihirai **
| | +--+
| | | +-- Lanceolaria_lanceolata **
| | +--+
| | +-- Lanceolaria_triformis **
| |
| | +-------- Pseudanodonta_complanata **
+--+ |
| | +----- Anodonta_cygnea **
| +--+
| | | +-- Anodonta_anatina **
| +-----------------+ +--+
| | | +-- Anodonta_nuttalliana **
| | |
| | | +----- Pyganodon_grandis **
| | | |
| | | +----- Simpsonaias_ambigua **
| | +--+
| | +----- Strophitus_undulatus **
| | |
| | | +-- Alasmidonta_marginata **
| | +--+
| | +-- Lasmigona_compressa **
+--+
| +-- Cristaria_plicata **
| +--------------------+
| | +-- Cristaria_clessini **
| |
| | +-------------- Anemina_arcaeformis **
| | |
| +--------+ +----------- Amuranodonta_kijaensis **
| | | |
| | +--+ +-------- Buldowskia_suifunica **
| | | |
| | +--+ +-- Buldowskia_flavotincta **
+--+ | +--+
| | | +-- Buldowskia_shadini **
| +--+
| | +-- Buldowskia_iwakawai **
| +--+
| +-- Buldowskia_kamiyai **
|
| +-------------------- Pletholophus_tenuis **
| |
| | +-------- Beringiana_japonica **
| | +--------+
+--+ | | +----- Beringiana_beringiana **
| | +--+
| | | +-- Beringiana_fukuharai **
| | +--+
+--+ +-- Beringiana_gosannensis **
|
| +-- Sinanodonta_angula **
| +-----------+
| | +-- Sinanodonta_lucida **
| |
+--+ +----------- Sinanodonta_schrenkii **
| |
| | +-- Sinanodonta_calipygos **
| +--------+
+--+ +-- Sinanodonta_tumens **
|
| +-------- Sinanodonta_woodiana[2] **
| |
+--+ +----- Sinanodonta_elliptica **
| |
+--+----- Sinanodonta_jourdyi **
|
| +-- Sinanodonta_lauta **
+--+
+-- Sinanodonta_woodiana[1] **
Huang et al., 2019, fig. S1
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 108 |
108 |
75 |
2 |
0 |
0 |
0 |
127.0 |
+-- Gibbosula_rochechouartii **
+-----------------------------------+
| +-- Margaritifera_dahurica **
|
| +----- Sinohyriopsis_cumingii[2] **
| +--------------+
| | | +-- Sinohyriopsis_cumingii[1] **
| | +--+
| | +-- Sinohyriopsis_schlegelii **
| |
| | +----- Parvasolenaia_rivularis **
| +--------------+ +-----------+
| | | | | +-- Parvasolenaia_triangularis **
| | | | +--+
| | | | +-- Ptychorhynchus_pfisteri **
| | | |
| | | | +-------- Sinosolenaia_recognita **
| | +--+ |
| | | +-----+ +----- Sinosolenaia_oleivora[2] **
| | | | | |
| | | | +--+ +-- Sinosolenaia_oleivora[1] **
| | | | | |
| | | | +--+-- Sinosolenaia_iridinea **
| | +--+ |
--+ | | +-- Sinosolenaia_oleivora[3] **
| | |
| | | +----------- Sinosolenaia_carinata **
| | | |
| | +--+ +-------- Lamprotula_leaii **
| | | |
| | +--+ +----- Lamprotula_cornuumlunae[1] **
| | | |
| | | +----- Lamprotula_cornuumlunae[3] **
| | +--+
| | +----- Lamprotula_cornuumlunae[4] **
| | |
| | | +-- Lamprotula_cornuumlunae[2] **
| | +--+
| | +-- Lamprotula_caveata **
| |
| | +-------------------------- Anemina_arcaeformis[5] **
| | |
| | | +----------------------- Anemina_euscaphys[2] **
| | +-----+ |
| | | | | +-------------------- Anemina_arcaeformis[8] **
| | | | | |
| | | +--+ +-------------------- Anemina_globosula[1] **
+--+ | | |
| | | | +----------------- Anemina_arcaeformis[9] **
| | | | |
| | +--+ +----------------- Anemina_euscaphys[3] **
| | | |
| | | +----------------- Anemina_globosula[2] **
| | | |
| | | | +-- Anemina_arcaeformis[1] **
| | | +--------------+
| | +--+ +-- Anemina_euscaphys[1] **
| | |
| | | +-- Anemina_arcaeformis[6] **
| | | +-----------+
| | | | +-- Anemina_euscaphys[4] **
| | | |
| | | | +----------- Anemina_arcaeformis[4] **
| | | | |
| | +--+ +----------- Anemina_euscaphys[7] **
| | | |
| | | | +-------- Anemina_arcaeformis[12] **
| | | +--+
| | | | | +----- Anemina_arcaeformis[7] **
| | +--+ +--+
| | | | +-- Anemina_arcaeformis[10] **
| | | +--+
| | | +-- Anemina_euscaphys[6] **
| | |
| | | +----- Anemina_arcaeformis[3] **
| | | |
| | +-----+----- Anemina_arcaeformis[11] **
| | |
| | | +-- Anemina_arcaeformis[2] **
| | +--+
| | +-- Anemina_euscaphys[5] **
| |
| | +-- Sinanodonta_angula **
+--+ +--------------------+
| | +-- Sinanodonta_lucida **
| |
| | +-------- Sinanodonta_woodiana[11] **
| | |
| | +-----------+ +----- Sinanodonta_woodiana[5] **
| | | | |
| +-----+ | | | +-- Sinanodonta_elliptica[1] **
| | | | +--+ |
| | | | | +-- Sinanodonta_woodiana[3] **
| | | | | |
| | | | | +-- Sinanodonta_woodiana[4] **
| | | | +--+
| | | | +-- Sinanodonta_woodiana[8] **
| | +--+ |
| | | +-- Sinanodonta_woodiana[9] **
| | | |
| | | +-- Sinanodonta_woodiana[14] **
| | |
| | | +----------------- Sinanodonta[6] **
| | | |
| | | | +----------- Sinanodonta[7] **
| | | | |
| | | | +--+ +-------- Sinanodonta[5] **
| | +--+ | | |
| | | | +--+ +----- Sinanodonta[1] **
| | | | | |
| | | | +--+----- Sinanodonta[3] **
| | | | |
| | | | | +-- Sinanodonta[2] **
| | +--+ +--+
| | | +-- Sinanodonta[4] **
| | |
| | | +----------- Sinanodonta_woodiana[2] **
| | | |
| | | | +-------- Sinanodonta_elliptica[2] **
| | | | |
| | | | +-------- Sinanodonta_woodiana[1] **
| | +--+ |
| | | +-------- Sinanodonta_woodiana[7] **
+--+ | |
| | +-------- Sinanodonta_woodiana[10] **
| | |
| +--+-------- Sinanodonta_woodiana[13] **
| |
| | +-- Sinanodonta_elliptica[3] **
| +-----+
| | +-- Sinanodonta_woodiana[12] **
| |
| | +----- Sinanodonta_elliptica[4] **
| | |
| +--+----- Sinanodonta_elliptica[6] **
| |
| | +-- Sinanodonta_elliptica[5] **
| +--+
| +-- Sinanodonta_woodiana[6] **
|
| +-------------- Acuticosta_ovata[1] **
| |
| | +----------- Acuticosta_chinensis[3] **
| +-----------+ |
| | | +----------- Acuticosta_chinensis[4] **
| | | |
| | +--+----------- Acuticosta_chinensis[6] **
| | |
| | | +-- Acuticosta_chinensis[1] **
| | | +-----+
| | | | +-- Acuticosta_ovata[2] **
| | +--+
| | | +----- Acuticosta_chinensis[5] **
| | | |
| | +--+----- Acuticosta_chinensis[7] **
| | |
| | | +-- Acuticosta_chinensis[2] **
+--+ +--+
| +-- Acuticosta_ovata[3] **
|
| +-- Lepidodesma_languilati **
| +--------------------+
| | +-- Cristaria_plicata **
| |
| | +----------- Lanceolaria_gladiola **
| | |
| | | +-------- Lanceolaria_grayii[5] **
| | +--------+ |
| | | | | +-- Lanceolaria_lanceolata **
| | | | +-----+
| | | | | +-- Lanceolaria_triformis **
+--+ | +--+
| | | +----- Lanceolaria_grayii[4] **
| | | |
| | | | +-- Lanceolaria_eucylindrica[1] **
| | | | |
| | +--+ +-- Lanceolaria_eucylindrica[2] **
| | | |
| | | +-- Lanceolaria_eucylindrica[3] **
| | | |
| | +--+-- Lanceolaria_eucylindrica[4] **
+--+ |
| +-- Lanceolaria_grayii[1] **
| |
| +-- Lanceolaria_grayii[2] **
| |
| +-- Lanceolaria_grayii[3] **
|
| +----------- Aculamprotula_tortuosa **
| +-----+
| | | +-------- Aculamprotula_tientsinensis **
| | +--+
| | | +----- Aculamprotula_fibrosa **
| | +--+
| | | +-- Aculamprotula_coreana **
+--+ +--+
| +-- Aculamprotula_scripta **
|
| +----- Nodularia_douglasiae[1] **
| +--------+
| | | +-- Nodularia_nuxpersicae **
| | +--+
+--+ +-- Nodularia_douglasiae[2] **
|
| +----------- Tchangsinaia_piscicula **
| |
+--+ +-- Schistodesmus_spinosus **
| +-----+
| | +-- Schistodesmus_lampreyanus **
+--+
| +----- Cuneopsis_rufescens **
+--+
| +-- Cuneopsis_heudei **
+--+
+-- Cuneopsis_celtiformis **
Bolotov et al., 2018, fig. 2
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 85 |
85 |
49 |
1 |
0 |
0 |
0 |
97.0 |
+-- Trapezidens_exolescens **
+--------+
| +-- Trapezidens_dolichorhynchus **
+--------------+
| | +-------- Lamellidens_savadiensis **
| +--+
| | +----- Lamellidens_ferrugineus **
| +--+
| | +-- Lamellidens_generosus **
| +--+
+--------+ +-- Lamellidens_mainwaringi **
| |
| | +-- Parreysia_keralaensis **
| | +--------------------+
| | | +-- Parreysia_nagpoorensis **
| | |
| | | +----- Leoparreysia_olivacea **
| +--+ +--------------+
| | | | +-- Leoparreysia_canefrii **
| | | +--+
| | | +-- Leoparreysia_tavoyensis **
| | |
| +--+ +-- Coelatura_aegyptiaca **
| | +--------------+
| | | +-- Nitia_teretiuscula **
| | |
| | | +-------- Indonaia_shurtleffiana **
| +--+ +-----+
| | | | +----- Indonaia_rugosa **
| | | +--+
| | | | +-- Indonaia_caerulea **
| | | +--+
| +--+ +-- Indonaia_andersoniana **
| |
| | +----------- Indochinella_pugio **
| | |
| | | +----- Radiatula_myitkyinae **
| +--+ +--+
| | | | +-- Radiatula_mouhoti **
| | | +--+
| +--+ +-- Radiatula_chaudhurii **
| |
| | +----- Scabies_mandarinus **
| +--+
| | +-- Scabies_humilis **
| +--+
--+ +-- Scabies_phaselus **
|
| +-- Pseudodon_inoscularis **
| |
| +--+-- Pseudodon_bogani **
| | |
| | +-- Pseudodon_manueli **
| |
| | +-- Bineurus_mouhotii **
| +--------------------------+ |
| | | +-- Monodontina_cambodjensis **
| | | |
| | | +-- Monodontina_mekongi **
| | | |
| | | +-- Pilsbryoconcha_exilis[1] **
| | +--+
| | +-- Pilsbryoconcha_exilis[2] **
| | |
| | +-- Bineurus_loeiensis **
| | |
| | +-- Bineurus_exilis **
| | |
| | +-- Monodontina_vondembuschiana **
| |
| | +-- Ensidens_telus **
| | |
| | +-- Ensidens_ingallsianus **
| | |
| | +-- Ensidens_jaculus **
| | |
| | +-- Hyriopsis_desowitzi **
| | +--+
| | | +-- Hyriopsis_myersiana **
| | | |
| | | +-- Hyriopsis_khoratensis **
| | | |
+--+ | +-- Ensidens_spiculus **
| | |
| | +-- Ensidens_sagittarius **
| |
| +-----------------------+ +-- Lens_eximius **
| | | |
| | | +-- Physunio_modelli[1] **
| | | |
| | | +-- Yaukthwa_nesemanni **
| | | |
| | | +-- Yaukthwa_panhai **
| | | |
| | | +-- Lens_comptus **
| | | |
| | +--+-- Lens_novoselovi **
| | |
| | +-- Physunio_modelli[2] **
| | |
| | +-- Physunio **
| | |
| | +-- Yaukthwa_inlenensis **
| | |
| | +-- Lens_contradens **
| | |
| | +-- Lens_micropterus **
| |
| | +-- Lamprotula_leaii **
+--+ |
| +-- Potomida_littoralis **
| |
| +-- Pronodularia_japanensis **
| |
| +-- Gonidea_angulata **
| |
| +-- Leguminaia_wheatleyi **
| |
| +-- Microcondylaea_bonellii **
| |
| +-----------------------+-- Amblema_plicata **
| | |
| | +-- Ortmanniana_ligamentina **
| | |
| | +-- Lampsilis_cardium **
| | |
| | +-- Cambarunio_iris **
| | |
| | +-- Elliptio_complanata **
| | |
| | +-- Pleurobema_sintoxia **
| | |
+--+ +-- Quadrula_quadrula **
|
| +----- Unio_tumidus **
| +--+
| | | +-- Unio_crassus **
| | +--+
| +--------------+ +-- Unio_pictorum **
| | |
| | | +----- Nodularia_douglasiae[1] **
| | +--+
| | | +-- Nodularia_nuxpersicae **
| | +--+
+--+ +-- Nodularia_douglasiae[2] **
|
| +-- Lanceolaria_gladiola **
| +-----------------+
| | +-- Lanceolaria_grayii **
| |
| | +-------- Cristaria_plicata **
+--+ +--------+
| | | +----- Pletholophus_tenuis **
| | +--+
| | | +-- Sinanodonta **
+--+ +--+
| +-- Anemina **
|
| +-------------- Anodonta_anatina **
+--+
| +----------- Pyganodon_grandis **
+--+
| +-------- Simpsonaias_ambigua **
+--+
| +----- Strophitus_undulatus **
+--+
| +-- Alasmidonta_marginata **
+--+
+-- Lasmigona_compressa **
Konopleva et al., 2023, fig. 8
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 83 |
83 |
78 |
1 |
0 |
0 |
0 |
129.0 |
+----------------------- Indonaia_andersoniana **
|
+--------------+ +-------------------- Indochinella_pugio **
| | |
| +--+ +----------------- Radiatula_mouhoti **
| | |
| +--+ +-------------- Scabiellus_songkramensis **
| | |
| | | +-- Unionetta_fabagina **
| +--+ +--------+
| | | +-- Harmandia_somboriensis **
| | |
| | | +-- Scabies_humilis **
| +--+ +-----+
| | | +-- Scabies_pilata **
| | |
| | | +----- Scabies_crispata **
| | +--+
| +--+ | +-- Scabies_nucleus **
| | +--+
| | +-- Scabies_mandarinus **
| |
| | +-- Scabies_phaselus **
| | |
| +-----+-- Scabies_anceps **
| |
| +-- Scabies_scobinatus **
|
| +----------- Pseudodon_kayinensis **
| +--------------------+
| | | +-------- Pseudodon_inoscularis **
| | +--+
| | | +----- Pseudodon_bogani **
| | +--+
| | | +-- Pseudodon_salwenianus **
| | +--+
| | +-- Pseudodon_manueli **
--+ +--+
| | | +-- Pilsbryoconcha_schomburgki **
| | | +-----+
| | | | +-- Pilsbryoconcha_exilis[1] **
| | | +--------------------+
| | | | | +----- Pilsbryoconcha **
| | | | +--+
| | | | | +-- Pilsbryoconcha_exilis[2] **
| | | | +--+
| | +--+ +-- Pilsbryoconcha_linguaeformis **
| | |
| | | +-------------------------- Songkhlanaia_tamodienica **
| | | |
| | | | +-- Namkongnaia_inkhavilayi **
| | | | +--------------------+
| | +--+ | +-- Namkongnaia_lemeslei **
| | | |
| | | | +-------- Monodontina_lenyanensis **
| | | | +-----------+
| | | | | | +----- Monodontina_cambodjensis **
| | +--+ | +--+
| | | | | +-- Monodontina_vondembuschiana **
| | | | +--+
| | | | +-- Monodontina_mekongi **
| | | |
| | | | +-- Sundadontina_taskaevi **
| | +--+ +--------+
| | | | +-- Sundadontina_brandti **
| | | +-----+
| | | | | +-------- Sundadontina_tumida **
| | | | +--+
| | | | | +----- Sundadontina_tanintharyiensis **
| | | | +--+
| | | | | +-- Sundadontina_plugpomenica **
| | +--+ +--+
+--+ | +-- Sundadontina_cumingii **
| |
| | +-------------- Nyeinchanconcha_nyeinchani **
| | |
| | | +----- Thaiconcha_thaiensis **
| +--+ +-----+
| | | | +-- Thaiconcha_munelliptica **
| | | +--+
| +--+ +-- Thaiconcha_callifera **
| |
| | +-------- Bineurus_loeiensis **
| +--+
| | +----- Bineurus_mouhotii **
| +--+
| | +-- Bineurus_anodontinum **
| +--+
| +-- Bineurus_exilis **
|
| +-- Gonidea_angulata **
| +--+
| | +-- Leguminaia_wheatleyi **
| +-----------------------+
| | | +-- Lamprotula_leaii **
| | +--+
| | +-- Potomida_littoralis **
| |
| | +-- Ensidens_ingallsianus **
| | +-----+
| | | +-- Ensidens_telus **
| | |
| | +--------------+ +-- Ensidens_jaculus **
| | | | +--+
| | | | | +-- Ensidens **
| | | +--+
| | | | +-- Ensidens_spiculus **
| | +--+ +--+
+-----+ | | +-- Ensidens_dugasti **
| | |
| | | +-- Hyriopsis_desowitzi **
| | | +-----------------+
| | | | +-- Hyriopsis_delaportei **
| | +--+
| | | +----------------- Rectidens_sumatrensis **
| | | |
| | +--+ +-------------- Hyriopsis_panhai **
| | | |
| | +--+ +----------- Hyriopsis_myersiana **
| | | |
| | +--+ +-- Hyriopsis_khoratensis **
| | | +-----+
| | | | +-- Hyriopsis_bialata **
| | +--+
+--+ | +----- Hyriopsis_sakhonensis **
| +--+
| | +-- Hyriopsis_phuphaniensis **
| +--+
| +-- Hyriopsis_kratiensis **
|
| +----- Physunio_modelli **
| +-----+
| | | +-- Physunio_superbus **
| | +--+
| | +-- Physunio_massini **
| +-----+
| | | +-- Trapezoideus_thachiadensis **
| | | +-----+
| | | | +-- Trapezoideus_subviridis **
| | +--+
| | | +----- Trapezoideus_mitanensis **
| | +--+
+--------+ | +-- Trapezoideus_foliaceus **
| +--+
| +-- Trapezoideus_lenya **
|
| +-- Yaukthwa_nesemanni **
| +-----------+
| | +-- Yaukthwa_elongatula **
| |
+--+ +-- Lens_contradens **
| +--------+
| | +-- Lens_misellus **
| |
+--+ +----- Lens_pallegoixi **
| +--+
| | | +-- Lens_novoselovi **
| | +--+
+--+ +-- Lens_comptus **
|
| +----- Lens_maenamensis **
+--+
| +-- Lens_eximius **
+--+
+-- Lens_rolfbrandti **
Jeratthitikul et al., 2022, fig. S1, 1
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 76 |
76 |
72 |
2 |
0 |
0 |
0 |
66.0 |
+-------------------- Cumberlandia_monodonta **
|
+-----------------+ +----------------- Gibbosula_laosensis **
| | |
| | | +-- Pseudunio_auricularius **
| +--+ +-----------+
| | | +-- Pseudunio_marocanus **
| | |
| +--+ +-- Margaritifera_margaritifera **
| | +--------+
| | | +-- Margaritifera_dahurica **
| +--+
| | +-------- Margaritifera_falcata **
| +--+
| | +----- Margaritifera_middendorffi **
| +--+
| | +-- Margaritifera_laevis **
| +--+
| +-- Margaritifera_marrianae **
|
| +-- Pseudodon_inoscularis **
| +--------------------+
| | +-- Pseudodon_kayinensis **
| |
--+ | +-- Nyeinchanconcha_nyeinchani **
| | +-----+
| +-----------+ | +-- Sundadontina_cumingii **
| | | +-----------+
| | | | | +----- Thaiconcha_callifera **
| | | | +--+
| | | | | +-- Bineurus_exilis **
| | | | +--+
| | +--+ +-- Bineurus_mouhotii **
| | |
| | | +-- Namkongnaia_inkhavilayi **
| | | +--------------+
| | | | +-- Monodontina_vondembuschiana **
| | | |
| | +--+ +-- Pilsbryoconcha_exilis **
| | | +-----------+
| | | | +-- Pilsbryoconcha_schomburgki **
| | | |
| | +--+ +-- Pilsbryoconcha_carinifera **
| | | +--------+
| | | | +-- Pilsbryoconcha_linguaeformis **
| | +--+
+--+ | +-------- Pilsbryoconcha_hoikaab **
| +--+
| | +----- Pilsbryoconcha_mekongiana **
| +--+
| | +-- Pilsbryoconcha_acuta **
| +--+
| +-- Pilsbryoconcha_kittitati **
|
| +----- Lens_contradens **
| +--+
| | | +-- Trapezoideus_foliaceus **
| | +--+
| +-----------------------+ +-- Physunio_superbus **
| | |
| | | +-- Ensidens_ingallsianus **
| | | |
| | +-----+-- Hyriopsis_bialata **
| | |
| | +-- Rectidens_sumatrensis **
| |
| | +----- Sinosolenaia_carinata **
+--+ +-----------------------+
| | | +-- Leguminaia_wheatleyi **
| | +--+
| | +-- Gonidea_angulata **
| |
| | +-- Chamberlainia_hainesiana **
| | +-----+
| | | +-- Sinohyriopsis_cumingii **
| | +-----------------+
| | | | +----- Lamprotula_leaii **
+--+ | +--+
| | | +-- Potomida_littoralis **
| | +--+
| | +-- Pronodularia_japanensis **
| |
| | +-------- Amblema_plicata **
| | +--+
| | | | +----- Ortmanniana_ligamentina **
| | | +--+
| | | | +-- Cambarunio_iris **
| | | +--+
+--+ +-----------+ +-- Lampsilis_cardium **
| | |
| | | +-- Quadrula_quadrula **
| | | +-----+
| | | | +-- Tritogonia_verrucosa **
| | +--+
| | | +----- Elliptio_complanata **
| | +--+
| | | +-- Pleurobema_sintoxia **
| | +--+
| | +-- Eurynia_dilatata **
| |
| | +----- Unio_tumidus **
+--+ +-----------+
| | | +-- Unio_pictorum **
| | +--+
| +--+ +-- Unio_crassus **
| | |
| | | +-------------- Aculamprotula_tortuosa **
| | | |
| | +--+ +-- Nodularia_douglasiae **
| | | +--------+
| | | | +-- Nodularia_nuxpersicae **
| | +--+
| | | +-------- Schistodesmus_lampreyanus **
| | +--+
| | | +----- Cuneopsis_heudei **
+--+ +--+
| | +-- Tchangsinaia_piscicula **
| +--+
| +-- Cuneopsis_rufescens **
|
| +-- Lanceolaria_grayii **
| +--------------+
| | +-- Lanceolaria_gladiola **
| |
| | +----------- Cristaria_plicata **
| | +--+
+--+ | | +-------- Pletholophus_tenuis **
| | +--+
| | | +----- Sinanodonta_lucida **
| | +--+
| | | +-- Sinanodonta_lauta **
| | +--+
+--+ +-- Sinanodonta_jourdyi **
|
| +-------- Pseudanodonta_complanata **
| +--+
| | | +----- Anodonta_nuttalliana **
| | +--+
| | | +-- Anodonta_anatina **
| | +--+
+--+ +-- Anodonta_cygnea **
|
| +-- Pyganodon_grandis **
| +-----+
| | +-- Simpsonaias_ambigua **
+--+
| +----- Strophitus_undulatus **
+--+
| +-- Alasmidonta_marginata **
+--+
+-- Lasmigona_compressa **
Jeratthitikul et al., 2022, fig. S2, 2
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 76 |
76 |
73 |
2 |
0 |
0 |
0 |
75.0 |
+----------------- Cumberlandia_monodonta **
|
| +----- Gibbosula_laosensis **
+--------------------+ +--------+
| | | | +-- Pseudunio_auricularius **
| | | +--+
| | | +-- Pseudunio_marocanus **
| +--+
| | +-- Margaritifera_dahurica **
| | +--------+
| | | +-- Margaritifera_margaritifera **
| +--+
| | +-------- Margaritifera_falcata **
| +--+
| | +----- Margaritifera_marrianae **
| +--+
| | +-- Margaritifera_middendorffi **
| +--+
| +-- Margaritifera_laevis **
|
| +-- Pseudodon_inoscularis **
| +--------------------+
| | +-- Pseudodon_kayinensis **
| |
--+ | +-- Nyeinchanconcha_nyeinchani **
| | +-----+
| +-----------+ | +-- Sundadontina_cumingii **
| | | +-----------+
| | | | | +----- Thaiconcha_callifera **
| | | | +--+
| | | | | +-- Bineurus_exilis **
| | | | +--+
| | +--+ +-- Bineurus_mouhotii **
| | |
| | | +-- Namkongnaia_inkhavilayi **
| | | +--------------+
| | | | +-- Monodontina_vondembuschiana **
| | | |
| | +--+ +-- Pilsbryoconcha_exilis **
| | | +-----------+
| | | | +-- Pilsbryoconcha_schomburgki **
| | | |
| | +--+ +-- Pilsbryoconcha_carinifera **
| | | +--------+
| | | | +-- Pilsbryoconcha_linguaeformis **
| | +--+
+--+ | +-------- Pilsbryoconcha_hoikaab **
| +--+
| | +----- Pilsbryoconcha_mekongiana **
| +--+
| | +-- Pilsbryoconcha_acuta **
| +--+
| +-- Pilsbryoconcha_kittitati **
|
| +----- Ensidens_ingallsianus **
| +--+
| | | +-- Rectidens_sumatrensis **
| | +--+
| +-----------------------+ +-- Hyriopsis_bialata **
| | |
| | | +----- Lens_contradens **
| | +--+
| | | +-- Trapezoideus_foliaceus **
| | +--+
| | +-- Physunio_superbus **
| |
| | +----- Sinosolenaia_carinata **
+--+ +-----------------------+
| | | +-- Gonidea_angulata **
| | +--+
| | +-- Leguminaia_wheatleyi **
| |
| | +-- Chamberlainia_hainesiana **
| | +-----+
| | | +-- Sinohyriopsis_cumingii **
| | +-----------------+
| | | | +----- Potomida_littoralis **
+--+ | +--+
| | | +-- Lamprotula_leaii **
| | +--+
| | +-- Pronodularia_japanensis **
| |
| | +-------- Amblema_plicata **
| | +--+
| | | | +----- Ortmanniana_ligamentina **
| | | +--+
| | | | +-- Lampsilis_cardium **
| | | +--+
+--+ +-----------+ +-- Cambarunio_iris **
| | |
| | | +-- Quadrula_quadrula **
| | | +-----+
| | | | +-- Tritogonia_verrucosa **
| | +--+
| | | +----- Pleurobema_sintoxia **
| | +--+
| | | +-- Eurynia_dilatata **
| | +--+
| | +-- Elliptio_complanata **
| |
| | +----- Unio_tumidus **
+--+ +-----------+
| | | +-- Unio_pictorum **
| | +--+
| +--+ +-- Unio_crassus **
| | |
| | | +-------------- Aculamprotula_tortuosa **
| | | |
| | +--+ +-- Nodularia_douglasiae **
| | | +--------+
| | | | +-- Nodularia_nuxpersicae **
| | +--+
| | | +-------- Schistodesmus_lampreyanus **
| | +--+
| | | +----- Cuneopsis_heudei **
+--+ +--+
| | +-- Tchangsinaia_piscicula **
| +--+
| +-- Cuneopsis_rufescens **
|
| +-- Lanceolaria_grayii **
| +--------------+
| | +-- Lanceolaria_gladiola **
| |
| | +----------- Cristaria_plicata **
| | +--+
+--+ | | +-------- Pletholophus_tenuis **
| | +--+
| | | +----- Sinanodonta_lucida **
| | +--+
| | | +-- Sinanodonta_jourdyi **
| | +--+
+--+ +-- Sinanodonta_lauta **
|
| +-------- Pseudanodonta_complanata **
| +--+
| | | +----- Anodonta_anatina **
| | +--+
| | | +-- Anodonta_nuttalliana **
| | +--+
+--+ +-- Anodonta_cygnea **
|
| +-- Alasmidonta_marginata **
| +-----+
| | +-- Lasmigona_compressa **
+--+
| +----- Pyganodon_grandis **
+--+
| +-- Strophitus_undulatus **
+--+
+-- Simpsonaias_ambigua **
Campbell & Lydeard, 2012, fig. 1-2
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 74 |
74 |
63 |
1 |
0 |
0 |
0 |
140.0 |
+-- Uniomerus_declivis **
+-----------------+
| +-- Uniomerus_tetralasmus **
+--------------------+
| | +----------------- Theliderma_metanevra **
| | |
| +--+ +-------------- Tritogonia_verrucosa **
| | |
| | | +-- Quadrula_quadrula **
| +--+ +--------+
| | | +-- Tritogonia_nobilis **
| | |
| +--+ +-- Cyclonaias_infucata **
| | +-----+
| | | +-- Cyclonaias_kleiniana **
| +--+
| | +----- Cyclonaias_tuberculata **
| +--+
| | +-- Cyclonaias_pustulosa **
| +--+
| +-- Cyclonaias_succissa **
|
| +----- Plectomerus_dombeyanus **
| +-----------------+
| | | +-- Reginaia_ebenus **
| | +--+
--+ | +-- Reginaia_rotulata **
| |
| | +----- Popenaias_popeii **
| +--------------+ +--------------+
| | | | | +-- Amblema_elliottii **
| | | | +--+
| | | | +-- Amblema_plicata **
| | | |
| | | | +----------------- Potamilus_alatus **
| | +--+ |
| | | +----------------- Potamilus_capax **
| | | |
| | | +----------------- Glebula_rotundata **
| | | |
| | | | +-- Toxolasma_lividum **
| | | +--------------+
| | +--+ +-- Paetulunio_fabalis **
| | |
| | | +-- Dromus_dromas **
| | +--------------+
| | | +-- Lemiox_rimosus **
| | |
| | | +-- Lampsilis_ovata **
+--+ | +-----------+
| | | +-- Obliquaria_reflexa **
| +--+
| | +----------- Lampsilis_virescens **
| | |
| +--+ +-- Epioblasma_rangiana **
| | +-----+
| | | +-- Obovaria_arkansasensis **
| +--+
| | +----- Obovaria_olivaria **
| +--+
| | +-- Venustaconcha_ellipsiformis **
| +--+
| +-- Venustaconcha_pleasii **
|
| +----------------------------------- Elliptoideus_sloatianus **
| |
| | +-------------------------------- Pleurobema_stabile **
| | |
| | | +-- Pleurobema_clava **
| | | +-----------+
+--+ | | +-- Pleurobema_oviforme **
| | |
| | +--------------+ +----------- Pleurobema_pyriforme **
| | | | |
| | | | | +-- Pleurobema_athearni **
| | | | | +--+
+--+ | +--+ | +-- Pleurobema_georgianum **
| | | +--+
| | | | | +-- Pleurobema_decisum **
| | | | +--+
| | | | +-- Pleurobema_hanleyianum **
| | +--+
| | | +----- Pleurobema_beadleianum **
| | | |
| | | +----- Pleurobema_strodeanum **
| | +--+
| | +----- Pleurobema_rubellum **
+--+ |
| | +-- Pleurobema_perovatum **
| +--+
| +-- Pleurobema_taitianum **
|
| +-------- Pleurobema_riddellii **
| +-----------------+
| | | +----- Pleurobema_cordatum **
| | +--+
| | | +-- Pleurobema_rubrum **
| | +--+
| | +-- Pleurobema_sintoxia **
| |
| | +----------------- Fusconaia_subrotunda **
| | |
+--+ +-----+ +-- Fusconaia_cor **
| | | +-----------+
| | | | +-- Fusconaia_cuneolus **
| | +--+
| | | +----------- Fusconaia_ozarkensis **
| | | |
| | +--+ +-- Fusconaia_escambia **
| | | +-----+
| | | | +-- Fusconaia_burkei **
| | +--+
+--+ | +----- Fusconaia_masoni **
| +--+
| | +-- Fusconaia_flava **
| +--+
| +-- Fusconaia_cerina **
|
| +----- Pleuronaia_gibber **
| +--------------+
| | | +-- Pleuronaia_barnesiana **
| | +--+
| | +-- Pleuronaia_dolabelloides **
+--+
| +-- Eurynia_dilatata **
| +--------------+
| | +-- Parvaspina_collina **
| |
+--+ +-- Hemistena_lata **
| +-----------+
| | +-- Plethobasus_cyphyus **
| |
+--+ +----------- Elliptio_shepardiana **
| |
| | +-------- Elliptio_complanata **
+--+ |
| | +----- Elliptio_purpurella **
| +--+
| | | +-- Elliptio_arca **
+--+ +--+
| +-- Elliptio_arctata **
|
| +-- Elliptio_mcmichaeli **
| |
+-----+-- Elliptio_pullata **
|
+-- Elliptio_crassidens **
Campbell & Lydeard, 2012, fig. 3
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 74 |
74 |
67 |
1 |
0 |
0 |
0 |
139.0 |
+-- Uniomerus_declivis **
+--------------+
| +-- Uniomerus_tetralasmus **
|
+--------------------+ +-- Tritogonia_verrucosa **
| | +-----------+
| | | +-- Tritogonia_nobilis **
| | |
| +--+ +----------- Quadrula_quadrula **
| | |
| | +----------- Theliderma_metanevra **
| +--+
| | +-- Cyclonaias_infucata **
| | +-----+
| | | +-- Cyclonaias_kleiniana **
| +--+
| | +----- Cyclonaias_tuberculata **
| +--+
| | +-- Cyclonaias_pustulosa **
| +--+
| +-- Cyclonaias_succissa **
|
| +-- Amblema_elliottii **
| +--+
| | +-- Amblema_plicata **
| +--------------------------+
| | | +-- Reginaia_ebenus **
--+ | +--+
| | +-- Reginaia_rotulata **
| +--+
| | | +----------------------------- Popenaias_popeii **
| | | |
| | | | +-- Glebula_rotundata **
| | +--+ +-----------------------+
| | | | +-- Paetulunio_fabalis **
| | | |
| | +--+ +-- Toxolasma_lividum **
| | | +--------------------+
| | | | +-- Plectomerus_dombeyanus **
| | | |
| | +--+ +-------------------- Obliquaria_reflexa **
| | | |
| | | | +-- Dromus_dromas **
| | | | +--+
| | +--+ | +-- Lemiox_rimosus **
| | | +-----------+
| | | | | +-- Potamilus_alatus **
| | | | +--+
| | +--+ +-- Potamilus_capax **
| | |
| | | +-------------- Lampsilis_ovata **
+--+ | |
| +--+ +----------- Lampsilis_virescens **
| | |
| +--+ +-- Epioblasma_rangiana **
| | +-----+
| | | +-- Obovaria_arkansasensis **
| +--+
| | +----- Obovaria_olivaria **
| +--+
| | +-- Venustaconcha_ellipsiformis **
| +--+
| +-- Venustaconcha_pleasii **
|
| +-------------------------------- Elliptoideus_sloatianus **
| |
| | +-- Eurynia_dilatata **
| | +--------------------------+
| | | +-- Parvaspina_collina **
| | |
| | | +-- Hemistena_lata **
| | +--------------------------+
| | | +-- Plethobasus_cyphyus **
+--+ |
| | +----------- Elliptio_shepardiana **
| | |
| | | +----- Elliptio_crassidens **
| +-----------------+ +--+
| | | | | +-- Elliptio_mcmichaeli **
| | | | +--+
| | +--+ +-- Elliptio_pullata **
| | |
| | | +----- Elliptio_complanata **
+--+ | |
| +--+ +-- Elliptio_purpurella **
| | |
| +--+-- Elliptio_arca **
| |
| +-- Elliptio_arctata **
|
| +----- Pleuronaia_gibber **
| +--------------------+
| | | +-- Pleuronaia_barnesiana **
| | +--+
| | +-- Pleuronaia_dolabelloides **
| |
| | +-------------- Fusconaia_ozarkensis **
| | |
| | | +----- Fusconaia_subrotunda **
| | +--------+ +-----+
+--+ | | | | +-- Fusconaia_cor **
| | | | +--+
| | | | +-- Fusconaia_cuneolus **
| | +--+
| | | +-- Fusconaia_escambia **
| | | +-----+
| | | | +-- Fusconaia_burkei **
| | +--+
| | | +----- Fusconaia_masoni **
+--+ +--+
| | +-- Fusconaia_cerina **
| +--+
| +-- Fusconaia_flava **
|
| +-------- Pleurobema_riddellii **
| +-----------+
| | | +----- Pleurobema_cordatum **
| | +--+
| | | +-- Pleurobema_rubrum **
| | +--+
+--+ +-- Pleurobema_sintoxia **
|
| +----------------- Pleurobema_stabile **
| |
| | +----------- Pleurobema_strodeanum **
| | +--+
+--+ | | +-------- Pleurobema_rubellum **
| | +--+
| | | +----- Pleurobema_beadleianum **
| | +--+
| | | +-- Pleurobema_perovatum **
+--+ +--+
| +-- Pleurobema_taitianum **
|
| +-- Pleurobema_clava **
| +--------+
| | +-- Pleurobema_oviforme **
+--+
| +-------- Pleurobema_pyriforme **
| |
+--+ +-- Pleurobema_athearni **
| +--+
| | +-- Pleurobema_georgianum **
+--+
| +-- Pleurobema_decisum **
+--+
+-- Pleurobema_hanleyianum **
Wu et al., 2022, fig. 3
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 71 |
71 |
68 |
2 |
3 |
3 |
2 |
88.0 |
+-- Mutela_dubia
+--------------------------------------------+
| +-- Anodontites_trapesialis
|
| +-------------------------------------------- Echyridella_menziesii
| |
| | +-------------- Gibbosula_rochechouartii **
| | +--------------------------+
--+ | | | +----------- Cumberlandia_monodonta **
| | | +--+
| | | | +-------- Pseudunio_marocanus **
| | | +--+
| | | | +----- Margaritifera_falcata **
| | | +--+
| | | | +-- Margaritifera_dahurica **
+--+ | +--+
| | +-- Margaritifera_margaritifera **
| |
| | +-- Uniomerus_tetralasmus **
| | +--------------------------+
| | | +-- Quadrula_quadrula **
| | |
| | +--------+ +-- Pleurobema_oviforme **
| | | | +-----------------------+
| | | | | +-- Elliptio_complanata **
| | | +--+
+--+ | | +----------------------- Amblema_plicata **
| | | |
| | +--+ +-------------------- Popenaias_popeii **
| | | |
| | +--+ +----------------- Toxolasma_parvum **
| | | |
| | +--+ +-- Potamilus_alatus **
| | | +-----------+
| | | | +-- Potamilus_leptodon **
| | +--+
| | | +----------- Leaunio_lienosus **
| | | |
| | +--+ +-- Lampsilis_cardium **
| | | +-----+
| | | | +-- Lampsilis_ornata **
| | +--+
| | | +----- Venustaconcha_ellipsiformis **
| | +--+
| | | +-- Lampsilis_siliquoidea **
| | +--+
+--+ +-- Lampsilis_powellii **
|
| +----- Chamberlainia_hainesiana **
| +-----------+
| | | +-- Sinohyriopsis_schlegelii **
| | +--+
| | +-- Sinohyriopsis_cumingii **
| |
| +-----------------+ +-------- Potomida_littoralis **
| | | +-----+
| | | | | +----- Pronodularia_japanensis **
| | | | +--+
| | | | | +-- Lamprotula_caveata **
| | | | +--+
| | +--+ +-- Lamprotula_leaii **
| | |
| | | +-- Pilsbryoconcha_exilis **
| | | +--------+
| | | | +-- Monodontina_vondembuschiana **
| | | |
| | +--+ +-- Ptychorhynchus_pfisteri **
| | | +-----+
| | | | +-- Parvasolenaia_rivularis **
| | +--+
| | | +----- Microcondylaea_bonellii **
+--+ +--+
| | +-- Sinosolenaia_carinata **
| +--+
| +-- Sinosolenaia_recognita **
|
| +-------------------------------- Lepidodesma_languilati **
| |
| | +-------- Acuticosta_chinensis **
| | +--------+
| | | | +----- Lanceolaria_gladiola **
| | | +--+
| | | | +-- Lanceolaria_grayii **
| | | +--+
| | | +-- Lanceolaria_lanceolata **
| | |
| | +-----------+ +-------- Cristaria_plicata **
| | | | |
+--+ | | +-----+ +-- Sinanodonta_woodiana **
| | | | | +--+
| | | | | | +-- Sinanodonta_lucida **
| | | | +--+
| | | | | +-- Anemina_arcaeformis **
| | +--+ +--+
| | | +-- Anemina_euscaphys **
| | |
| | | +-- Anodonta_cygnea **
| | | +--------+
| | | | +-- Anodonta_anatina **
| | +--+
| | | +-------- Lasmigona_compressa **
+--+ +--+
| | +----- Pyganodon_grandis **
| +--+
| | +-- Utterbackia_imbecillis **
| +--+
| +-- Utterbackia_peninsularis **
|
| +-------- Aculamprotula_tortuosa **
| |
| +-----------------+ +-- Aculamprotula_tientsinensis **
| | | +--+
| | | | +-- Aculamprotula_coreana **
| | +--+
| | | +-- Aculamprotula_polysticta **
| | +--+
| | +-- Aculamprotula_scripta **
+--+
| +-------- Unio_tumidus **
| +--------------+
| | | +----- Unio_crassus **
| | +--+
| | | +-- Unio_pictorum **
| | +--+
+--+ +-- Unio_delphinus **
|
| +-- Pseudocuneopsis_capitata **
| +-----------------+
| | +-- Pseudocuneopsis_sichuanensis **
| |
+--+ +-- Nodularia_douglasiae **
| +--------------+
| | +-- Nodularia_breviconcha **
| |
+--+ +-- Schistodesmus_lampreyanus **
| +-----------+
| | +-- Schistodesmus_spinosus **
+--+
| +----------- Tchangsinaia_piscicula **
+--+
| +-------- Cuneopsis_rufescens **
+--+
| +----- Cuneopsis_celtiformis **
+--+
| +-- Cuneopsis_heudei **
+--+
+-- Cuneopsis_demangei **
Sano et al., 2020, fig. 2
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 64 |
64 |
61 |
2 |
11 |
9 |
2 |
96.0 |
+----- Triplodon_corrugatus
+--------------------------------------------+
| | +-- Hyridella_australis
| +--+
| +-- Velesunio_ambiguus
|
| +----------- Mutela_hargeri
| |
| +-----------------------------------+ +-- Aspatharia_pfeifferiana
| | | +-----+
| | | | +-- Chambardia_wahlbergi
--+ | +--+
| | | +----- Etheria_elliptica
| | +--+
| | | +-- Anodontites_elongata
| | +--+
| | +-- Lamproscapha_ensiformis
| |
| | +-------------------- Cumberlandia_monodonta **
| | |
| | | +-- Gibbosula_rochechouartii **
| | | +--+
+--+ +-----------------------+ | +-- Gibbosula_laosensis **
| | | +-----------+
| | | | | +-- Pseudunio_auricularius **
| | | | +--+
| | | | +-- Pseudunio_marocanus **
| | +--+
| | | +-- Margaritifera_dahurica **
| | | +-----------+
| | | | +-- Margaritifera_margaritifera **
| | +--+
| | | +----------- Margaritifera_falcata **
| | +--+
| | | +-------- Margaritifera_middendorffi **
| | +--+
+--+ | +----- Margaritifera_laevis **
| +--+
| | +-- Margaritifera_hembeli **
| +--+
| +-- Margaritifera_marrianae **
|
| +----------------------------------------- Scabies_crispata **
| |
| | +----------------- Lampsilis_cardium **
| | |
| | | +----- Sinohyriopsis_cumingii[2] **
| | +--------------------+ +--------+
| | | | | | +-- Sinohyriopsis_schlegelii **
| | | | | +--+
| | | +--+ +-- Sinohyriopsis_cumingii[1] **
+--+ | |
| | | +----------- Inversidens_brandtii **
| | | |
| | +--+ +-- Ptychorhynchus_pfisteri **
| | | +-----+
| | | | +-- Obovalis_omiensis **
| | +--+
| | | +----- Lamprotula_leaii **
| | +--+
| | | +-- Pronodularia_japanensis **
+--+ +--+
| +-- Potomida_littoralis **
|
| +----- Anemina_arcaeformis[2] **
| +-----------------------------+
| | | +-- Anemina_arcaeformis[1] **
| | +--+
| | +-- Anemina_euscaphys **
| |
| | +----------------- Pletholophus_tenuis **
| | |
| | | +-------- Sinanodonta_lucida **
| | | +-----+
| | +--------------+ | | +----- Sinanodonta_tumens **
| | | | | +--+
+--+ | | | | +-- Sinanodonta_lauta[1] **
| | | | +--+
| | +--+ +-- Sinanodonta_woodiana **
| | |
| | | +-- Sinanodonta_lauta[3] **
| | | +--------+
| | | | +-- Beringiana_japonica[3] **
| | +--+
| | | +-------- Beringiana_japonica[2] **
| | | |
| | +--+ +-- Sinanodonta_lauta[2] **
+--+ | +--+
| | | +-- Sinanodonta_calipygos[1] **
| +--+
| | +-- Beringiana_japonica[1] **
| +--+
| +-- Sinanodonta_calipygos[2] **
|
| +----------------------------- Cristaria_plicata **
| |
| | +-- Anodonta_cygnea **
| | +--------+
| | | +-- Anodonta_anatina **
| | |
| | +--------------+ +----- Alasmidonta_varicosa **
+--+ | | +--+
| | | | | +-- Alasmidonta_heterodon **
| | | | +--+
| | +--+ +-- Lasmigona_compressa **
| | |
| | | +----- Pyganodon_grandis **
| | +--+
+--+ | +-- Utterbackia_imbecillis **
| +--+
| +-- Utterbackia_peninsularis **
|
| +----- Lanceolaria_grayii[1] **
| +-----------------+
| | | +-- Lanceolaria_lanceolata **
| | +--+
| | +-- Lanceolaria_grayii[2] **
+--+
| +-------------------- Lepidodesma_languilati **
| |
| | +----- Aculamprotula_tortuosa **
| | +-----------+
+--+ | | +-- Aculamprotula_tientsinensis **
| | +--+
| | +-- Aculamprotula_coreana **
| |
+--+ +----- Unio_crassus **
| +--------+
| | | +-- Unio_delphinus **
| | +--+
| | +-- Unio_pictorum **
+--+
| +----- Nodularia_douglasiae **
| +-----+
| | | +-- Tchangsinaia_piscicula **
| | +--+
+--+ +-- Schistodesmus_lampreyanus **
|
| +-------- Inversiunio_jokohamensis **
+--+
| +----- Inversiunio_yanagawensis[2] **
+--+
| +-- Inversiunio_reinianus **
+--+
+-- Inversiunio_yanagawensis[1] **
Sano et al., 2020, fig. 3
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 64 |
64 |
61 |
2 |
11 |
9 |
2 |
98.0 |
+----- Triplodon_corrugatus
+--------------------------------------------+
| | +-- Hyridella_australis
| +--+
| +-- Velesunio_ambiguus
|
| +----------- Mutela_hargeri
| |
| +-----------------------------------+ +-- Aspatharia_pfeifferiana
| | | +-----+
| | | | +-- Chambardia_wahlbergi
--+ | +--+
| | | +----- Etheria_elliptica
| | +--+
| | | +-- Anodontites_elongata
| | +--+
| | +-- Lamproscapha_ensiformis
| |
| | +----------------------- Cumberlandia_monodonta **
| | |
| | | +-- Pseudunio_auricularius **
| | +--------------------+ +-----------------+
+--+ | | | +-- Pseudunio_marocanus **
| | | |
| | +--+ +-- Gibbosula_laosensis **
| | | +--------------+
| | | | +-- Gibbosula_rochechouartii **
| | | |
| | +--+ +-- Margaritifera_margaritifera **
| | | +-----------+
| | | | +-- Margaritifera_dahurica **
| | +--+
| | | +----------- Margaritifera_falcata **
| | +--+
| | | +-------- Margaritifera_middendorffi **
| | +--+
+--+ | +----- Margaritifera_laevis **
| +--+
| | +-- Margaritifera_hembeli **
| +--+
| +-- Margaritifera_marrianae **
|
| +----------------------------------------- Scabies_crispata **
| |
| | +----- Sinohyriopsis_cumingii[2] **
| | +--------+
| | | | +-- Sinohyriopsis_schlegelii **
| | | +--+
| | | +-- Sinohyriopsis_cumingii[1] **
| | +-----------------------+
| | | | +----------- Lampsilis_cardium **
| | | | |
+--+ | | | +----- Lamprotula_leaii **
| | +--+ +--+
| | | | | +-- Potomida_littoralis **
| | | | +--+
| | +--+ +-- Pronodularia_japanensis **
| | |
| | | +----- Inversidens_brandtii **
| | +--+
| | | +-- Obovalis_omiensis **
| | +--+
+--+ +-- Ptychorhynchus_pfisteri **
|
| +----- Anemina_arcaeformis[2] **
| +-----------------------------+
| | | +-- Anemina_arcaeformis[1] **
| | +--+
| | +-- Anemina_euscaphys **
| |
| | +-------- Sinanodonta_lucida **
| | +--------+
| | | | +----- Sinanodonta_tumens **
| | | +--+
| | | | +-- Sinanodonta_woodiana **
| | +--------------+ +--+
+--+ | | +-- Sinanodonta_lauta[3] **
| | |
| | | +-------------- Pletholophus_tenuis **
| | | |
| | +--+ +-- Sinanodonta_lauta[1] **
| | | +--------+
| | | | +-- Beringiana_japonica[1] **
| | +--+
| | | +-------- Beringiana_japonica[3] **
| | | |
| | +--+ +-- Beringiana_japonica[2] **
| | | +--+
+--+ | | +-- Sinanodonta_calipygos[2] **
| +--+
| | +-- Sinanodonta_lauta[2] **
| +--+
| +-- Sinanodonta_calipygos[1] **
|
| +----------------------------- Cristaria_plicata **
| |
| | +-- Anodonta_anatina **
| | +--------+
| | | +-- Anodonta_cygnea **
| | |
| | +--------------+ +----- Pyganodon_grandis **
+--+ | | +--+
| | | | | +-- Utterbackia_imbecillis **
| | | | +--+
| | +--+ +-- Utterbackia_peninsularis **
| | |
| | | +----- Alasmidonta_varicosa **
| | +--+
+--+ | +-- Lasmigona_compressa **
| +--+
| +-- Alasmidonta_heterodon **
|
| +----- Lanceolaria_grayii[1] **
| +-----------------+
| | | +-- Lanceolaria_lanceolata **
| | +--+
| | +-- Lanceolaria_grayii[2] **
+--+
| +-------------------- Lepidodesma_languilati **
| |
| | +----- Aculamprotula_tortuosa **
| | +-----------+
+--+ | | +-- Aculamprotula_tientsinensis **
| | +--+
| | +-- Aculamprotula_coreana **
| |
+--+ +----- Unio_crassus **
| +--------+
| | | +-- Unio_delphinus **
| | +--+
| | +-- Unio_pictorum **
+--+
| +----- Nodularia_douglasiae **
| +-----+
| | | +-- Schistodesmus_lampreyanus **
| | +--+
+--+ +-- Tchangsinaia_piscicula **
|
| +-------- Inversiunio_jokohamensis **
+--+
| +----- Inversiunio_yanagawensis[2] **
+--+
| +-- Inversiunio_reinianus **
+--+
+-- Inversiunio_yanagawensis[1] **
Sano et al., 2020, fig. 4
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 64 |
64 |
61 |
2 |
11 |
9 |
2 |
96.0 |
+----- Triplodon_corrugatus
+-----------------------------------------------+
| | +-- Hyridella_australis
| +--+
| +-- Velesunio_ambiguus
|
| +----------- Mutela_hargeri
| |
| +--------------------------------------+ +-- Aspatharia_pfeifferiana
| | | +-----+
| | | | +-- Chambardia_wahlbergi
--+ | +--+
| | | +----- Etheria_elliptica
| | +--+
| | | +-- Anodontites_elongata
| | +--+
| | +-- Lamproscapha_ensiformis
| |
| | +-------------------- Cumberlandia_monodonta **
| | |
| | | +-- Pseudunio_auricularius **
| | +--------------------------+ +--------------+
+--+ | | | +-- Pseudunio_marocanus **
| | | |
| | +--+ +-- Gibbosula_rochechouartii **
| | | +-----------+
| | | | +-- Gibbosula_laosensis **
| | | |
| | +--+ +-- Margaritifera_margaritifera **
| | | +--------+
| | | | +-- Margaritifera_dahurica **
| | +--+
| | | +-------- Margaritifera_falcata **
| | | |
| | +--+ +-- Margaritifera_laevis **
| | | +--+
+--+ | | +-- Margaritifera_middendorffi **
| +--+
| | +-- Margaritifera_hembeli **
| +--+
| +-- Margaritifera_marrianae **
|
| +-------------------------------------------- Scabies_crispata **
| |
| | +----------------- Lampsilis_cardium **
| | |
| | | +----- Sinohyriopsis_cumingii[2] **
| | +-----------------------+ +--------+
| | | | | | +-- Sinohyriopsis_schlegelii **
| | | | | +--+
| | | +--+ +-- Sinohyriopsis_cumingii[1] **
+--+ | |
| | | +----------- Inversidens_brandtii **
| | | |
| | +--+ +-- Obovalis_omiensis **
| | | +-----+
| | | | +-- Ptychorhynchus_pfisteri **
| | +--+
| | | +----- Lamprotula_leaii **
| | +--+
| | | +-- Pronodularia_japanensis **
+--+ +--+
| +-- Potomida_littoralis **
|
| +----- Anemina_arcaeformis[2] **
| +--------------------------------+
| | | +-- Anemina_arcaeformis[1] **
| | +--+
| | +-- Anemina_euscaphys **
| |
| | +-------- Sinanodonta_lucida **
| | +--------+
| | | | +----- Sinanodonta_tumens **
| | | +--+
| | | | +-- Sinanodonta_woodiana **
| | | +--+
+--+ +-----------------+ +-- Sinanodonta_lauta[1] **
| | |
| | | +-------------- Pletholophus_tenuis **
| | | |
| | | | +-- Sinanodonta_lauta[2] **
| | +--+ +--------+
| | | | +-- Beringiana_japonica[1] **
| | | |
| | +--+ +-- Beringiana_japonica[3] **
| | | +-----+
| | | | +-- Sinanodonta_calipygos[2] **
| | +--+
+--+ | +----- Beringiana_japonica[2] **
| +--+
| | +-- Sinanodonta_lauta[3] **
| +--+
| +-- Sinanodonta_calipygos[1] **
|
| +-------------------------------- Cristaria_plicata **
| |
| | +-- Anodonta_cygnea **
| | +--------+
| | | +-- Anodonta_anatina **
| | |
| | +-----------------+ +----- Alasmidonta_varicosa **
+--+ | | +--+
| | | | | +-- Alasmidonta_heterodon **
| | | | +--+
| | +--+ +-- Lasmigona_compressa **
| | |
| | | +----- Pyganodon_grandis **
| | +--+
+--+ | +-- Utterbackia_imbecillis **
| +--+
| +-- Utterbackia_peninsularis **
|
| +-------------------------- Lepidodesma_languilati **
| |
| | +----- Lanceolaria_grayii[1] **
| | +-----------------+
+--+ | | +-- Lanceolaria_lanceolata **
| | +--+
| | +-- Lanceolaria_grayii[2] **
| |
+--+ +----- Aculamprotula_tortuosa **
| +--------------+
| | | +-- Aculamprotula_tientsinensis **
| | +--+
| | +-- Aculamprotula_coreana **
+--+
| +----- Unio_crassus **
| +-----------+
| | | +-- Unio_delphinus **
| | +--+
+--+ +-- Unio_pictorum **
|
| +-------------- Nodularia_douglasiae **
| |
+--+ +-- Schistodesmus_lampreyanus **
| +--------+
| | +-- Tchangsinaia_piscicula **
+--+
| +-------- Inversiunio_jokohamensis **
+--+
| +----- Inversiunio_yanagawensis[2] **
+--+
| +-- Inversiunio_reinianus **
+--+
+-- Inversiunio_yanagawensis[1] **
Bolotov et al., 2022, fig. 2
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 61 |
61 |
54 |
1 |
0 |
0 |
0 |
127.0 |
+-- Monodontina_lenyanensis **
|
+-- Nyeinchanconcha_nyeinchani **
+--------------------------------------------+
| +-- Pseudodon_kayinensis **
| |
| +-- Sundadontina_tanintharyiensis **
|
| +----------- Trapezidens_mogaungensis **
| |
| +-----+ +-------- Trapezidens_exolescens **
| | | |
| | +--+ +----- Trapezidens_scutum **
| | | |
| | +--+ +-- Trapezidens_angustior **
| | | |
| +--------------------------+ +--+-- Trapezidens_dolichorhynchus **
| | | |
--+ | | +-- Trapezidens_yeti **
| | |
| | | +-------------- Lamellidens_marginalis **
| | | |
| | | | +----------- Lamellidens_ferrugineus **
| | +--+ |
| | | | +-------- Lamellidens_generosus **
| | | +--+
| | | | | +----- Lamellidens_mainwaringi **
| | +--+ +--+
| | | | +-- Lamellidens_lamellatus **
| | | +--+
| | | +-- Lamellidens_jenkinsianus **
| | |
| | | +-------- Lamellidens_nongyangensis **
+--+ +--+
| | +----- Lamellidens_savadiensis **
| +--+
| | +-- Lamellidens_corrianus **
| +--+
| +-- Lamellidens_chindwinensis **
|
| +----------- Parreysia_keralaensis **
| +-----------------------------+
| | | +-------- Parreysia_corrugata **
| | +--+
| | | +----- Parreysia_rakhinensis **
| | +--+
| | | +-- Parreysia_nagpoorensis **
| | +--+
| | +-- Parreysia_plagiosoma **
| |
| | +-- Leoparreysia_burmana **
+--+ +-----+
| | +-- Leoparreysia_whitteni **
| +-----------------------------+
| | | +----- Leoparreysia_olivacea **
| | +--+
| | | +-- Leoparreysia_canefrii **
| | +--+
| | +-- Leoparreysia_tavoyensis **
| |
| | +-------- Nitia_teretiuscula **
+--+ +--------------------------+
| | | +----- Coelatura_aegyptiaca **
| | +--+
| | | +-- Nyassunio_nyassaensis **
| | +--+
| | +-- Coelatura_kunenensis **
| |
| | +----------------- Indonaia_cylindrica **
| | |
+--+ +--------------+ +-------------- Indonaia_gratiosa **
| | | |
| | +--+ +----------- Indonaia_andersoniana **
| | | |
| | +--+ +-- Indonaia_caerulea **
| | | +-----+
| | | | +-- Indonaia_subclathrata **
| | +--+
| | | +----- Indonaia_rugosa **
+--+ +--+
| | +-- Indonaia_bonneaudii **
| +--+
| +-- Indonaia_shurtleffiana **
|
| +----------------------------- Indochinella_pugio **
| |
| | +----------- Radiatula_crispisulcata **
| | |
| | | +----- Radiatula_chaudhurii **
+--+ +--------------+ +--+
| | | | | +-- Radiatula_myitkyinae **
| | | | +--+
| | +--+ +-- Radiatula_kachinensis **
| | |
| | | +----- Radiatula_chindwinensis **
+--+ +--+
| | +-- Radiatula_myitthanensis **
| +--+
| +-- Radiatula_mouhoti **
|
| +----------------------- Scabiellus_songkramensis **
| |
+--+ +-------------------- Harmandia_somboriensis **
| |
+--+ +----------------- Unionetta_fabagina **
| |
+--+ +-- Scabies_crispata **
| +-----------+
| | +-- Scabies_mandarinus **
+--+
| +----------- Scabies_nucleus **
| |
+--+ +-- Scabies_humilis **
| +-----+
| | +-- Scabies_pilata **
+--+
| +----- Scabies_phaselus **
+--+
| +-- Scabies_anceps **
+--+
+-- Scabies_scobinatus **
Bolotov et al., 2022, fig. S1
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 61 |
61 |
48 |
1 |
0 |
0 |
0 |
127.0 |
+-- Monodontina_lenyanensis **
|
+-- Nyeinchanconcha_nyeinchani **
+-----------------------------------------------+
| +-- Pseudodon_kayinensis **
| |
| +-- Sundadontina_tanintharyiensis **
|
| +-- Parreysia_keralaensis **
| |
| +-- Parreysia_corrugata **
| |
| +--------------------------------------------+-- Parreysia_plagiosoma **
| | |
| | +-- Parreysia_rakhinensis **
--+ | |
| | +-- Parreysia_nagpoorensis **
| |
| | +-------------- Trapezidens_mogaungensis **
| | +--+
| | | | +----------- Trapezidens_scutum **
| | | +--+
| | | | +-------- Trapezidens_exolescens **
| | | +--+
| | | | +----- Trapezidens_yeti **
| | | +--+
| | | | +-- Trapezidens_dolichorhynchus **
| | +--------------------------+ +--+
+--+ | | +-- Trapezidens_angustior **
| | |
| | | +-------- Lamellidens_savadiensis **
| | | +-----+
| | | | | +----- Lamellidens_chindwinensis **
| | | | +--+
| | | | | +-- Lamellidens_corrianus **
| | | | +--+
| | +--+ +-- Lamellidens_nongyangensis **
| | |
| | | +-- Lamellidens_marginalis **
| | | +--------+
| | | | +-- Lamellidens_ferrugineus **
| | +--+
| | | +-------- Lamellidens_generosus **
| | +--+
| | | +----- Lamellidens_mainwaringi **
+--+ +--+
| | +-- Lamellidens_jenkinsianus **
| +--+
| +-- Lamellidens_lamellatus **
|
| +-- Leoparreysia_burmana **
| |
| +-- Leoparreysia_canefrii **
| |
| +--------------------------------------+-- Leoparreysia_olivacea **
| | |
| | +-- Leoparreysia_tavoyensis **
| | |
| | +-- Leoparreysia_whitteni **
| |
| | +-- Coelatura_aegyptiaca **
| | |
| | +-- Coelatura_kunenensis **
+--+ +-----------------------------------+
| | +-- Nitia_teretiuscula **
| | |
| | +-- Nyassunio_nyassaensis **
| |
| | +-- Indonaia_cylindrica **
| | +--------+
| | | +-- Indonaia_gratiosa **
| | |
+--+ +-----------------------+ +----- Indonaia_shurtleffiana **
| | | +--+
| | | | | +-- Indonaia_bonneaudii **
| | | | +--+
| | +--+ +-- Indonaia_rugosa **
| | |
| | | +----- Indonaia_andersoniana **
| | +--+
| | | +-- Indonaia_caerulea **
+--+ +--+
| +-- Indonaia_subclathrata **
|
| +-------------------------------- Indochinella_pugio **
| |
| | +----- Radiatula_chaudhurii **
| | +-----+
| | | | +-- Radiatula_kachinensis **
| | | +--+
+--+ +-----------------+ +-- Radiatula_myitkyinae **
| | |
| | | +-------- Radiatula_crispisulcata **
| | +--+
| | | +----- Radiatula_mouhoti **
| | +--+
+--+ | +-- Radiatula_myitthanensis **
| +--+
| +-- Radiatula_chindwinensis **
|
| +-------------------------- Scabiellus_songkramensis **
| |
+--+ +----------------------- Harmandia_somboriensis **
| |
+--+ +-------------------- Unionetta_fabagina **
| |
+--+ +----------------- Scabies_crispata **
| |
+--+ +-- Scabies_humilis **
| +-----------+
| | +-- Scabies_mandarinus **
+--+
| +----------- Scabies_nucleus **
+--+
| +-------- Scabies_pilata **
+--+
| +----- Scabies_anceps **
+--+
| +-- Scabies_phaselus **
+--+
+-- Scabies_scobinatus **
Zanatta & Murphy, 2006, fig. 2
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 58 |
58 |
49 |
2 |
0 |
0 |
0 |
123.0 |
+----------------------------------------------------- Pyganodon_grandis **
|
+----------------------------------------------------- Cumberlandia_monodonta **
|
| +-------- Pleuronaia_dolabelloides **
| |
--+ +-----------------------------------------+-------- Pleurobema_clava **
| | |
| | | +----- Fusconaia_flava **
| | +--+
| | | +-- Hemistena_lata **
| | +--+
| | +-- Eurynia_dilatata **
+--+
| +----------------------------------------------- Amblema_plicata **
| |
| +----------------------------------------------- Popenaias_popeii **
| |
| | +----- Paetulunio_fabalis **
| | |
| | +--------------------------------------+ +-- Toxolasma_parvum **
+--+ | | |
| | +--+-- Toxolasma_lividum **
| | |
| | +-- Toxolasma_texasiense **
| |
| | +----- Obliquaria_reflexa **
| | +-----------------------------------+
+--+ | | +-- Glebula_rotundata **
| | +--+
| | +-- Cyrtonaias_tampicoensis **
| |
| | +----- Ptychobranchus_fasciolaris **
| | +--+
| | | | +-- Dromus_dromas **
| | | +--+
+--+ +-----------------------------+ +-- Cyprogenia_stegaria **
| | |
| | | +----- Medionidus_conradicus **
| | +--+
| | | +-- Medionidus_acutissimus **
| | +--+
| | +-- Lemiox_rimosus **
| |
| | +----------------------------------- Sagittunio_nasutus **
| | |
+--+ | +-------------- Potamilus_leptodon **
| | |
| +--------------------+ +----------- Potamilus_fragilis **
| | | |
| | +--+ +-- Potamilus_alatus **
| | | +-----+
| | | | +-- Potamilus_purpuratus **
| | +--+
| | | +----- Potamilus_inflatus **
| | +--+
+--+ | +-- Potamilus_ohiensis **
| +--+
| +-- Potamilus_amphichaenus **
|
| +-- Truncilla_truncata **
| +-----------------------------+
| | +-- Ellipsaria_lineolata **
| |
| | +-- Ortmanniana_pectorosa **
| | +-----+
| | | +-- Lampsilis_fasciola **
| | +--------------------+
+--+ | | +----- Lampsilis_ornata **
| | +--+
| | | +-- Lampsilis_ovata **
| | +--+
| | +-- Lampsilis_cardium **
| |
| | +-- Ligumia_recta **
+--+ +--------+
| | +-- Cambarunio_iris **
| +--------------+
| | | +-------- Lampsilis_teres **
| | | |
| | +--+ +-- Hamiota_australis **
| | | +--+
| | | | +-- Hamiota_subangulata **
+--+ +--+
| | +-- Hamiota_altilis **
| +--+
| +-- Hamiota_perovalis **
|
| +----------------------- Leaunio_vanuxemensis **
| |
+--+ +-------------------- Ortmanniana_ligamentina **
| |
| | +----------------- Lampsilis_siliquoidea **
+--+ |
| | +-------------- Obovaria_olivaria **
| | |
+--+ | +-- Venustaconcha_ellipsiformis **
| +-----------+
| | +-- Venustaconcha_pleasii **
| |
+--+ +----- Obovaria_unicolor **
+--------+
| | +-- Obovaria_subrotunda **
| +--+
| +-- Obovaria_arkansasensis **
|
| +----------- Epioblasma_brevidens **
+--+
| +-------- Epioblasma_triquetra **
+--+
| +----- Epioblasma_rangiana **
+--+
| +-- Epioblasma_walkeri **
+--+
+-- Epioblasma_capsaeformis **
Wu et al., 2018, fig. 2
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 58 |
58 |
46 |
2 |
0 |
0 |
0 |
87.0 |
+-- Margaritifera_dahurica **
+--------------------------------+
| +-- Margaritifera_falcata **
|
| +-------------------------------- Quadrula_quadrula **
| |
| | +-- Amblema_plicata **
| +-----------------------------+
| | +-- Obovaria_arkansasensis **
| |
| | +----------- Pleurobema_sintoxia **
--+ | |
| +--------------------+ +-- Elliptio_complanata **
| | | +-----+
| | | | +-- Eurynia_dilatata **
| | +--+
| | | +----- Fusconaia_askewi **
| | +--+
| | | +-- Fusconaia_cerina **
| | +--+
| | +-- Fusconaia_burkei **
+--+
| +----- Potomida_littoralis **
| +--------+
| | | +-- Lamprotula_caveata **
| | +--+
| +--------------+ +-- Lamprotula_leaii **
| | |
| | | +----------- Sinosolenaia_carinata **
| | | |
| | +--+ +-------- Gonidea_angulata **
| | | |
| | +--+-------- Sinosolenaia_oleivora **
| | |
| | | +----- Ptychorhynchus_pfisteri **
| | +--+
| | | +-- Parvasolenaia_rivularis **
| | +--+
+--+ +-- Parvasolenaia_triangularis **
|
| +-------------------------- Sinohyriopsis_cumingii **
| |
| | +----------------------- Lepidodesma_languilati **
| | |
| | | +----------- Aculamprotula_tortuosa **
| | | +-----+
| | | | | +-------- Aculamprotula_scripta **
| | | | +--+
| | | | | +----- Aculamprotula_coreana **
| | | | +--+
| | | | | +-- Aculamprotula_fibrosa **
+--+ +-----+ +--+
| | | +-- Aculamprotula_tientsinensis **
| | |
| | | +-- Unio_crassus **
| | | +-----------+
| | | | +-- Unio_pictorum **
| | | |
| | +--+ +----- Nodularia_douglasiae **
| | | +-----+
| | | | | +-- Schistodesmus_lampreyanus **
| | | | +--+
+--+ +--+ +-- Schistodesmus_spinosus **
| |
| | +-------- Tchangsinaia_piscicula **
| +--+
| | +----- Cuneopsis_rufescens **
| +--+
| | +-- Cuneopsis_celtiformis **
| +--+
| +-- Cuneopsis_heudei **
|
| +----------- Acuticosta_chinensis **
| +--------+
| | | +-------- Lanceolaria_gladiola **
| | +--+
| | | +----- Lanceolaria_grayii **
| | +--+
| | | +-- Lanceolaria_lanceolata **
| | +--+
| | +-- Lanceolaria_triformis **
| |
| | +-------------- Anodonta_cygnea **
+--+ |
| +--+ +----------- Anodonta_anatina **
| | | |
| | +--+ +-------- Strophitus_undulatus **
| | | |
| | +--+ +----- Lasmigona_compressa **
| | | |
| | +--+----- Alasmidonta_marginata **
| | |
| | | +-- Pyganodon_cataracta **
+--+ +--+
| +-- Pyganodon_grandis **
|
| +-------- Cristaria_plicata **
| |
| | +-- Sinanodonta_woodiana **
| | +--+
| | | +-- Sinanodonta_elliptica **
| +--+
| | | +-- Sinanodonta_lucida **
+--------+ +--+
| +-- Sinanodonta_angula **
|
| +-- Anemina_arcaeformis[1] **
| |
| +-- Anemina_arcaeformis[2] **
| |
+-----+-- Anemina_globosula[1] **
|
+-- Anemina_globosula[2] **
|
+-- Anemina_globosula[3] **
Bolotov et al., 2022, fig. 2
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 58 |
58 |
53 |
1 |
0 |
0 |
0 |
124.0 |
+-- Monodontina_lenyanensis **
|
+-- Nyeinchanconcha_nyeinchani **
+-----------------------------------------------+
| +-- Pseudodon_kayinensis **
| |
| +-- Sundadontina_tanintharyiensis **
|
| +-- Trapezidens_exolescens **
| +-----+
| | +-- Trapezidens_scutum **
| +-----------+
| | | +----- Trapezidens_angustior **
| | +--+
| | | +-- Trapezidens_dolichorhynchus **
| +--------------------------+ +--+
| | | +-- Trapezidens_yeti **
| | |
--+ | | +----------------- Lamellidens_corrianus **
| | | |
| | +--+ +-------------- Lamellidens_nongyangensis **
| | | |
| | +--+ +----------- Lamellidens_savadiensis **
| | | |
| | | | +-- Lamellidens_ferrugineus **
| | +--+ +-----+
| | | | +-- Lamellidens_marginalis **
| | | |
| | +--+ +-- Lamellidens_lamellatus **
| | | +--+
| | | | +-- Lamellidens_jenkinsianus **
| | +--+
| | | +-- Lamellidens_generosus **
| | +--+
+--+ +-- Lamellidens_mainwaringi **
|
| +----------- Parreysia_keralaensis **
| +--+
| | | +-------- Parreysia_plagiosoma **
| | +--+
| | | +----- Parreysia_nagpoorensis **
| | +--+
| | | +-- Parreysia_corrugata **
| +-----------------------------+ +--+
| | | +-- Parreysia_rakhinensis **
| | |
| | | +-- Leoparreysia_burmana **
| | | +-----+
| | | | +-- Leoparreysia_whitteni **
| | +-----+
| | | +----- Leoparreysia_olivacea **
| | +--+
| | | +-- Leoparreysia_canefrii **
| | +--+
+--+ +-- Leoparreysia_tavoyensis **
|
| +-------- Nitia_teretiuscula **
| +--------------------------------+
| | | +----- Coelatura_aegyptiaca **
| | +--+
| | | +-- Coelatura_kunenensis **
| | +--+
| | +-- Nyassunio_nyassaensis **
| |
| | +-- Indonaia_cylindrica **
| | +--------+
+--+ | +-- Indonaia_gratiosa **
| |
| +--------------------------+ +----- Indonaia_andersoniana **
| | | +--+
| | | | | +-- Indonaia_caerulea **
| | | | +--+
| | +--+ +-- Indonaia_subclathrata **
| | |
| | | +----- Indonaia_shurtleffiana **
| | +--+
+--+ | +-- Indonaia_bonneaudii **
| +--+
| +-- Indonaia_rugosa **
|
| +----------------------------------- Indochinella_pugio **
| |
| | +----------- Radiatula_crispisulcata **
| | |
+--+ +--------------------+ +-- Radiatula_chaudhurii **
| | | +-----+
| | | | +-- Radiatula_myitkyinae **
| | +--+
| | | +----- Radiatula_mouhoti **
+--+ +--+
| | +-- Radiatula_chindwinensis **
| +--+
| +-- Radiatula_myitthanensis **
|
| +----------------------------- Scabiellus_songkramensis **
+--+
| +-------------------------- Harmandia_somboriensis **
+--+
| +----------------------- Unionetta_fabagina **
+--+
| +-------------------- Scabies_crispata **
+--+
| +----------------- Scabies_mandarinus **
+--+
| +-------------- Scabies_nucleus **
+--+
| +----------- Scabies_pilata **
+--+
| +-------- Scabies_humilis **
+--+
| +----- Scabies_anceps **
+--+
| +-- Scabies_phaselus **
+--+
+-- Scabies_scobinatus **
Bolotov et al., 2022, fig. S1
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 58 |
58 |
51 |
1 |
0 |
0 |
0 |
124.0 |
+-- Monodontina_lenyanensis **
|
+-- Nyeinchanconcha_nyeinchani **
+--------------------------------------------+
| +-- Pseudodon_kayinensis **
| |
| +-- Sundadontina_tanintharyiensis **
|
| +-------- Trapezidens_exolescens **
| |
| +--------+ +----- Trapezidens_scutum **
| | | |
| | +--+ +-- Trapezidens_angustior **
| | | |
| | +--+-- Trapezidens_dolichorhynchus **
| | |
| +--------------------------+ +-- Trapezidens_yeti **
--+ | |
| | | +----- Lamellidens_savadiensis **
| | | +--------+
| | | | | +-- Lamellidens_corrianus **
| | | | +--+
| | | | +-- Lamellidens_nongyangensis **
| | +--+
| | | +-- Lamellidens_ferrugineus **
| | | +--------+
| | | | +-- Lamellidens_marginalis **
| | +--+
| | | +-------- Lamellidens_lamellatus **
| | +--+
| | | +----- Lamellidens_jenkinsianus **
+--+ +--+
| | +-- Lamellidens_generosus **
| +--+
| +-- Lamellidens_mainwaringi **
|
| +----------- Parreysia_keralaensis **
| +-----------------------------+
| | | +-------- Parreysia_corrugata **
| | +--+
| | | +----- Parreysia_rakhinensis **
| | +--+
| | | +-- Parreysia_plagiosoma **
| | +--+
| | +-- Parreysia_nagpoorensis **
| |
| | +-- Leoparreysia_burmana **
+--+ +-----+
| | +-- Leoparreysia_whitteni **
| +-----------------------------+
| | | +----- Leoparreysia_olivacea **
| | +--+
| | | +-- Leoparreysia_canefrii **
| | +--+
| | +-- Leoparreysia_tavoyensis **
| |
| | +-------- Nitia_teretiuscula **
+--+ +--------------------------+
| | | +----- Coelatura_aegyptiaca **
| | +--+
| | | +-- Coelatura_kunenensis **
| | +--+
| | +-- Nyassunio_nyassaensis **
| |
| | +----------------- Indonaia_cylindrica **
| | |
+--+ +--------------+ +-------------- Indonaia_gratiosa **
| | | |
| | +--+ +----------- Indonaia_andersoniana **
| | | |
| | +--+ +-- Indonaia_caerulea **
| | | +-----+
| | | | +-- Indonaia_subclathrata **
| | +--+
| | | +----- Indonaia_rugosa **
+--+ +--+
| | +-- Indonaia_bonneaudii **
| +--+
| +-- Indonaia_shurtleffiana **
|
| +----------------------------- Indochinella_pugio **
| |
| | +-------- Radiatula_crispisulcata **
| | |
| | +-----------------+ +-- Radiatula_chaudhurii **
+--+ | | +--+
| | | | +-- Radiatula_myitkyinae **
| | +--+
| | | +-- Radiatula_myitthanensis **
| | | |
+--+ +--+-- Radiatula_mouhoti **
| |
| +-- Radiatula_chindwinensis **
|
| +----------------------- Scabiellus_songkramensis **
| |
+--+ +-------------------- Harmandia_somboriensis **
| |
+--+ +----------------- Unionetta_fabagina **
| |
+--+ +-- Scabies_crispata **
| +-----------+
| | +-- Scabies_mandarinus **
+--+
| +----------- Scabies_nucleus **
| |
+--+ +-- Scabies_humilis **
| +-----+
| | +-- Scabies_pilata **
+--+
| +----- Scabies_phaselus **
+--+
| +-- Scabies_anceps **
+--+
+-- Scabies_scobinatus **
Bolotov et al., 2022, fig. S2
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 58 |
58 |
53 |
1 |
0 |
0 |
0 |
124.0 |
+-- Monodontina_lenyanensis **
|
+-- Nyeinchanconcha_nyeinchani **
+-----------------------------------------------+
| +-- Pseudodon_kayinensis **
| |
| +-- Sundadontina_tanintharyiensis **
|
| +----------- Parreysia_keralaensis **
| +-----------------------------------+
| | | +-------- Parreysia_corrugata **
| | +--+
| | | +----- Parreysia_nagpoorensis **
| | +--+
--+ | | +-- Parreysia_plagiosoma **
| | +--+
| | +-- Parreysia_rakhinensis **
| |
| | +----------- Trapezidens_scutum **
| | +-----+
| | | | +-------- Trapezidens_exolescens **
| | | +--+
| | | | +----- Trapezidens_yeti **
| | | +--+
| | | | +-- Trapezidens_dolichorhynchus **
+--+ | +--+
| +--------------------------+ +-- Trapezidens_angustior **
| | |
| | | +----- Lamellidens_savadiensis **
| | | +--------+
| | | | | +-- Lamellidens_corrianus **
| | | | +--+
| | | | +-- Lamellidens_nongyangensis **
| | +--+
| | | +-- Lamellidens_marginalis **
| | | +--------+
| | | | +-- Lamellidens_ferrugineus **
| | +--+
| | | +-------- Lamellidens_lamellatus **
| | +--+
| | | +----- Lamellidens_jenkinsianus **
+--+ +--+
| | +-- Lamellidens_mainwaringi **
| +--+
| +-- Lamellidens_generosus **
|
| +----------- Leoparreysia_burmana **
| +-----------------------------+
| | | +-------- Leoparreysia_whitteni **
| | +--+
| | | +----- Leoparreysia_olivacea **
| | +--+
| | | +-- Leoparreysia_canefrii **
| | +--+
| | +-- Leoparreysia_tavoyensis **
| |
+--+ +-------- Nitia_teretiuscula **
| +-----------------------------+
| | | +----- Coelatura_aegyptiaca **
| | +--+
| | | +-- Coelatura_kunenensis **
| | +--+
| | +-- Nyassunio_nyassaensis **
| |
| | +-- Indonaia_cylindrica **
| | +--------+
+--+ | +-- Indonaia_gratiosa **
| |
| +-----------------------+ +----- Indonaia_shurtleffiana **
| | | +--+
| | | | | +-- Indonaia_bonneaudii **
| | | | +--+
| | +--+ +-- Indonaia_rugosa **
| | |
| | | +----- Indonaia_andersoniana **
| | +--+
+--+ | +-- Indonaia_caerulea **
| +--+
| +-- Indonaia_subclathrata **
|
| +-------------------------------- Indochinella_pugio **
| |
| | +----------- Radiatula_crispisulcata **
| | |
| | +-----------------+ +-- Radiatula_chaudhurii **
+--+ | | +-----+
| | | | +-- Radiatula_myitkyinae **
| | +--+
| | | +----- Radiatula_mouhoti **
| | +--+
+--+ | +-- Radiatula_myitthanensis **
| +--+
| +-- Radiatula_chindwinensis **
|
| +-------------------------- Scabiellus_songkramensis **
| |
+--+ +----------------------- Harmandia_somboriensis **
| |
+--+ +-------------------- Unionetta_fabagina **
| |
+--+ +----------------- Scabies_crispata **
| |
+--+ +-- Scabies_humilis **
| +-----------+
| | +-- Scabies_mandarinus **
+--+
| +----------- Scabies_nucleus **
+--+
| +-------- Scabies_pilata **
+--+
| +----- Scabies_anceps **
+--+
| +-- Scabies_phaselus **
+--+
+-- Scabies_scobinatus **
Zanatta & Murphy, 2006, fig. 1
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 57 |
57 |
52 |
2 |
0 |
0 |
0 |
123.0 |
+----------------------------------------------------- Cumberlandia_monodonta **
|
| +-------------------------------------------------- Pyganodon_grandis **
| |
--+ | +-- Amblema_plicata **
| | +--+
| | | +-- Popenaias_popeii **
| | |
+--+ +-----------------------------------------+ +-- Pleurobema_clava **
| | | |
| | | +-- Eurynia_dilatata **
| | +--+
| | +-- Pleuronaia_dolabelloides **
| | |
+--+ +-- Fusconaia_flava **
|
| +-- Cyrtonaias_tampicoensis **
| +-----------------------------------------+
| | +-- Glebula_rotundata **
| |
| | +----------------------------------------- Paetulunio_fabalis **
+--+ |
| | +-------- Obliquaria_reflexa **
| | +-----------------------------+
| | | | +----- Toxolasma_texasiense **
| | | +--+
+--+ | | +-- Toxolasma_parvum **
| | +--+
| | +-- Toxolasma_lividum **
| |
| | +----------------------------------- Sagittunio_nasutus **
| | |
+--+ | +-- Medionidus_conradicus **
| | +-----+
| | | +-- Ptychobranchus_fasciolaris **
| | +--------+
| | | | +----- Medionidus_acutissimus **
| | | | |
| | | +--+----- Lemiox_rimosus **
| | | |
| | +--------------+ | +-- Dromus_dromas **
+--+ | | +--+
| | | +-- Cyprogenia_stegaria **
| | |
| | | +-------------- Potamilus_leptodon **
| | | |
| | +--+ +----------- Potamilus_fragilis **
| | | |
| | +--+ +-- Potamilus_alatus **
| | | +-----+
| | | | +-- Potamilus_purpuratus **
| | +--+
| | | +----- Potamilus_inflatus **
+--+ +--+
| | +-- Potamilus_ohiensis **
| +--+
| +-- Potamilus_amphichaenus **
|
| +----- Lampsilis_teres **
| +-----------------------+
| | | +-- Truncilla_truncata **
| | +--+
| | +-- Ellipsaria_lineolata **
| |
| | +-------------------------- Leaunio_vanuxemensis **
| | |
+--+ | +-- Hamiota_australis **
| | +--+
| | | +-- Hamiota_subangulata **
| | +-----+
| | | | +-- Hamiota_altilis **
| | | +--+
| | | +-- Hamiota_perovalis **
+--+ +-----------+
| | | +-- Lampsilis_fasciola **
| | | +-----+
| | | | +-- Ortmanniana_pectorosa **
| | +--+
| | | +----- Lampsilis_ornata **
| | +--+
| | | +-- Lampsilis_ovata **
| | +--+
+--+ +-- Lampsilis_cardium **
|
| +-------- Lampsilis_siliquoidea **
| +-----------+
| | | +----- Ortmanniana_ligamentina **
| | +--+
| | | +-- Cambarunio_iris **
| | +--+
| | +-- Ligumia_recta **
+--+
| +-- Venustaconcha_ellipsiformis **
| +--------------+
| | +-- Venustaconcha_pleasii **
| |
| | +-------- Obovaria_olivaria **
+--+ +-----+
| | | +----- Obovaria_unicolor **
| | +--+
| | | +-- Obovaria_subrotunda **
+--+ +--+
| +-- Obovaria_arkansasensis **
|
| +----------- Epioblasma_brevidens **
+--+
| +-------- Epioblasma_triquetra **
+--+
| +----- Epioblasma_rangiana **
+--+
| +-- Epioblasma_walkeri **
+--+
+-- Epioblasma_capsaeformis **
Konopleva et al., 2023, fig. S1, 4
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 56 |
56 |
54 |
2 |
0 |
0 |
0 |
83.0 |
+----------------------------------- Gibbosula_laosensis **
|
| +-- Trapezidens_exolescens **
| +-----------+
| | +-- Lamellidens_savadiensis **
| +-----------------+
| | | +----------- Parreysia_rakhinensis **
| | +--+
| | | +-------- Leoparreysia_tavoyensis **
| | +--+
| | | +----- Indonaia_andersoniana **
| | +--+
--+ | | +-- Indochinella_pugio **
| | +--+
| | +-- Radiatula_mouhoti **
| |
| | +-- Lampsilis_ovata **
| | +--------------+
| | | +-- Amblema_plicata **
| | |
| | | +----------- Sinosolenaia_recognita **
| | | |
| | | +--+ +-- Ptychorhynchus_pfisteri **
| | +-----------+ | | +-----+
+--+ | | | | | +-- Parvasolenaia_rivularis **
| | | | +--+
| | | | | +----- Gonidea_angulata **
| | | | +--+
| | | | | +-- Microcondylaea_bonellii **
| | +--+ +--+
| | | +-- Leguminaia_wheatleyi **
| | |
| | | +----- Chamberlainia_hainesiana **
| | | +-----+
| | | | | +-- Sinohyriopsis_schlegelii **
| | | | +--+
| | +--+ +-- Sinohyriopsis_cumingii **
| | |
| | | +-------- Potomida_littoralis **
| | +--+
| | | +----- Pronodularia_japanensis **
| | +--+
| | | +-- Lamprotula_caveata **
+--+ +--+
| +-- Lamprotula_leaii **
|
| +----------------- Middendorffinaia_mongolica **
| |
| | +-------- Aculamprotula_tortuosa **
| | +-----+
| +--------+ | | +----- Aculamprotula_scripta **
| | | | +--+
| | | | | +-- Aculamprotula_tientsinensis **
| | | | +--+
| | +--+ +-- Aculamprotula_coreana **
| | |
| | | +----- Unio_tumidus **
| | | +-----+
| | | | | +-- Unio_pictorum **
| | | | +--+
| | +--+ +-- Unio_crassus **
| | |
| | | +-------- Nodularia_douglasiae **
+--+ +--+
| | +----- Tchangsinaia_piscicula **
| +--+
| | +-- Cuneopsis_rufescens **
| +--+
| +-- Cuneopsis_heudei **
|
| +-------- Lepidodesma_languilati **
| +--------------+
| | | +----- Lanceolaria_gladiola **
| | +--+
| | | +-- Lanceolaria_grayii **
| | +--+
+--+ +-- Lanceolaria_triformis **
|
| +-------------------- Acuticosta_chinensis **
| |
| | +----- Anodonta_anatina **
| | +-----------+
+--+ | | +-- Pseudanodonta_complanata **
| | +--+
| | +-- Anodonta_cygnea **
| |
| | +-------- Anemina_arcaeformis **
+--+ +-----+
| | | +----- Amuranodonta_kijaensis **
| | +--+
| | | +-- Buldowskia_suifunica **
| | +--+
| | +-- Buldowskia_shadini **
+--+
| +-------- Beringiana_beringiana **
| +--+
| | | +----- Sinanodonta_schrenkii **
| | +--+
| | | +-- Sinanodonta_woodiana **
+--+ +--+
| +-- Sinanodonta_lauta **
|
| +-------- Cristaria **
| |
+--+ +-- Cristaria_truncata **
| +--+
| | +-- Cristaria_plicata **
+--+
| +-- Cristaria_bellua **
+--+
+-- Cristaria_clessini **
Fukata & Iigo, 2020, fig. 1
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 54 |
54 |
51 |
2 |
0 |
0 |
0 |
85.0 |
+----- Gibbosula_crassa **
+-----------------------+
| | +-- Pseudunio_marocanus **
| +--+
| +-- Cumberlandia_monodonta **
|
| +-- Quadrula_quadrula **
| +-----------------+
| | +-- Uniomerus_tetralasmus **
| +--+
| | | +----------------- Elliptio_complanata **
| | | |
| | +--+ +-- Potamilus_leptodon **
| | | +-----------+
| | | | +-- Potamilus_alatus **
| | +--+
| | | +----------- Leaunio_lienosus **
| | | |
| | +--+ +-- Lampsilis_cardium **
| | | +-----+
--+ | | | +-- Lampsilis_ornata **
| +--+ +--+
| | | | +----- Venustaconcha_ellipsiformis **
| | | +--+
| | | | +-- Lampsilis_powellii **
| | | +--+
| | | +-- Lampsilis_siliquoidea **
| | |
| | | +-- Sinohyriopsis_cumingii **
| | | +-----------+
| | | | +-- Sinohyriopsis_schlegelii **
| | | |
| | | | +-- Pilsbryoconcha_exilis **
| | +--------+ +--------+
| | | | +-- Monodontina_vondembuschiana **
| | | |
| | | | +-- Lamprotula_caveata **
| | | | +--+
| | +--+ | +-- Lamprotula_leaii **
| | | +--+
+--+ | | | +-- Pronodularia_japanensis **
| | | +--+
| | | +-- Potomida_littoralis **
| +--+
| | +-- Sinosolenaia_carinata **
| | +--+
| | | +-- Sinosolenaia_oleivora **
| +--+
| | +-- Parvasolenaia_rivularis **
| +--+
| +-- Ptychorhynchus_pfisteri **
|
| +----------------------- Lepidodesma_languilati **
| |
| | +----- Lanceolaria_gladiola **
| | +--------------+
| | | | +-- Lanceolaria_grayii **
| | | +--+
| | | +-- Lanceolaria_lanceolata **
+--+ |
| | +-------------- Acuticosta_chinensis **
| | |
| | | +-------- Cristaria_plicata **
| | | |
| | | +--+ +-- Sinanodonta_woodiana **
| | +--+ | | +--+
+--+ | | | | | +-- Sinanodonta_lucida **
| | | | +--+
| | | | | +-- Anemina_arcaeformis **
| | | | +--+
| | +--+ +-- Anemina_euscaphys **
| | |
| | | +-- Anodonta_anatina **
| | | +-----+
| | | | +-- Anodonta_cygnea **
| | | |
| | +--+ +-- Pyganodon_grandis **
+--+ | +--+
| | | +-- Utterbackia_peninsularis **
| +--+
| | +-- Alasmidonta_varicosa **
| +--+
| +-- Alasmidonta_heterodon **
|
| +----- Aculamprotula_tortuosa **
| +--------+
| | | +-- Aculamprotula_tientsinensis **
| | +--+
| | +-- Aculamprotula_coreana **
| |
+--+ +-------- Unio_tumidus **
| +--+
| | | +----- Unio_crassus **
| | +--+
| | | +-- Unio_pictorum **
| | +--+
+--+ +-- Unio_delphinus **
|
| +-- Schistodesmus_lampreyanus **
| +-----+
| | +-- Cuneopsis_heudei **
+--+
| +----- Pseudocuneopsis_capitata **
+--+
| +-- Nodularia_douglasiae **
+--+
+-- Tchangsinaia_piscicula **
Huang et al., 2019, fig. 3
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 53 |
53 |
50 |
2 |
4 |
3 |
2 |
80.0 |
+-- Anodontites_trapesialis
+--------------------------------------+
| +-- Mutela_dubia
|
| +-------------------------------------- Echyridella_menziesii
| |
| | +-------------- Gibbosula_rochechouartii **
| | +--------------------+
--+ | | | +----------- Cumberlandia_monodonta **
| | | +--+
| | | | +-------- Pseudunio_marocanus **
| | | +--+
| | | | +----- Margaritifera_falcata **
| | | +--+
| | | | +-- Margaritifera_margaritifera **
| | | +--+
+--+ | +-- Margaritifera_dahurica **
| |
| | +-- Sinohyriopsis_cumingii **
| | +-----------+
| | | +-- Sinohyriopsis_schlegelii **
| | |
| | | +-------- Potomida_littoralis **
| | +--------+ +--+
| | | | | | +----- Pronodularia_japanensis **
| | | | | +--+
| | | | | | +-- Lamprotula_caveata **
| | | | | +--+
+--+ | +--+ +-- Lamprotula_leaii **
| | |
| | | +-- Ptychorhynchus_pfisteri **
| | | +--+
| +--------+ | | +-- Parvasolenaia_rivularis **
| | | +-----+
| | | | +-- Sinosolenaia_oleivora **
| | | +--+
| | | +-- Sinosolenaia_carinata **
| | |
| | | +-- Quadrula_quadrula **
| | | +-----------------+
| | | | +-- Uniomerus_tetralasmus **
| | +--+
| | | +----------------- Elliptio_complanata **
| | | |
| | +--+ +-------------- Toxolasma_parvum **
| | | |
| | +--+ +-- Potamilus_leptodon **
| | | +--------+
| | | | +-- Potamilus_alatus **
+--+ +--+
| | +-------- Leaunio_lienosus **
| +--+
| | +----- Venustaconcha_ellipsiformis **
| +--+
| | +-- Lampsilis_cardium **
| +--+
| +-- Lampsilis_ornata **
|
| +----------------------------- Lepidodesma_languilati **
| |
| | +-------- Aculamprotula_tortuosa **
| | +-----+
| | | | +----- Aculamprotula_scripta **
| | | +--+
| | | | +-- Aculamprotula_tientsinensis **
| | | +--+
| | +-----------+ +-- Aculamprotula_coreana **
+--+ | |
| | | +----- Nodularia_douglasiae **
| | | +-----+
| | | | | +-- Tchangsinaia_piscicula **
| | | | +--+
| | +--+ +-- Schistodesmus_lampreyanus **
| | |
| | | +-------- Unio_tumidus **
| | +--+
+--+ | +----- Unio_crassus **
| +--+
| | +-- Unio_delphinus **
| +--+
| +-- Unio_pictorum **
|
| +----- Lanceolaria_gladiola **
| +-----------------+
| | | +-- Lanceolaria_lanceolata **
| | +--+
| | +-- Lanceolaria_grayii **
+--+
| +-------------------- Acuticosta_chinensis **
| |
| | +-------- Cristaria_plicata **
| | |
+--+ +--------+ +-- Sinanodonta_lucida **
| | | +--+
| | | | +-- Sinanodonta_woodiana **
| | +--+
+--+ | +-- Anemina_arcaeformis **
| +--+
| +-- Anemina_euscaphys **
|
| +-------------- Anodonta_cygnea **
+--+
| +----------- Anodonta_anatina **
+--+
| +-------- Lasmigona_compressa **
+--+
| +----- Pyganodon_grandis **
+--+
| +-- Utterbackia_imbecillis **
+--+
+-- Utterbackia_peninsularis **
Konopleva et al., 2019, fig. 1
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 52 |
52 |
40 |
2 |
0 |
0 |
0 |
106.0 |
+-- Gibbosula_laosensis **
+--------------------------------+
| +-- Margaritifera_dahurica **
|
| +-- Pseudodon_inoscularis **
| |
| +--+-- Pseudodon_bogani **
| | |
| | +-- Pseudodon_manueli **
| |
| | +-- Bineurus_mouhotii **
| +--------------------------+ |
| | | +-- Monodontina_cambodjensis **
| | | |
| | | +-- Monodontina_mekongi **
--+ | | |
| | | +-- Pilsbryoconcha_exilis[1] **
| | +--+
| | +-- Pilsbryoconcha_exilis[2] **
| | |
| | +-- Bineurus_loeiensis **
| | |
| | +-- Bineurus_exilis **
| | |
| | +-- Monodontina_vondembuschiana **
| |
| | +----------- Indonaia_shurtleffiana **
| | +--+
| | | | +-------- Indonaia_rugosa **
| | | +--+
+--+ | | +----- Indonaia_andersoniana **
| | +--+
| | | +-- Indonaia_caerulea **
| +--------------+ +--+
| | | +-- Indonaia_subclathrata **
| | |
| | | +----------- Indochinella_pugio **
| | | |
| | | | +-- Scabies_humilis **
| | +--+ |
| | | +-----+-- Scabies_mandarinus **
| | | | |
| | +--+ +-- Scabies_phaselus **
| | |
| | | +----- Radiatula_mouhoti **
| | +--+
| | | +-- Radiatula_myitkyinae **
| | +--+
| | +-- Radiatula_chaudhurii **
| |
+--+ +-- Ensidens_ingallsianus **
| +-----+
| | +-- Ensidens_telus **
| +-----+
| | | +----- Ensidens_sagittarius **
| | +--+
| | | +-- Ensidens_jaculus **
| +-----------+ +--+
| | | +-- Ensidens_spiculus **
| | |
| | | +----------- Hyriopsis_desowitzi **
| | | |
| | +--+----------- Rectidens_sumatrensis **
| | |
| | | +-------- Hyriopsis_panhai **
| | +--+
| | | +----- Hyriopsis_myersiana **
| | +--+
+--+ | +-- Hyriopsis_khoratensis **
| +--+
| +-- Hyriopsis_bialata **
|
| +-- Yaukthwa_paiensis **
| +-----+
| | +-- Yaukthwa_inlenensis **
| +--------------+
| | | +----- Yaukthwa_panhai **
| | +--+
| | | +-- Yaukthwa_nesemanni **
| | +--+
+--+ +-- Yaukthwa_dalliana **
|
| +----- Physunio_superbus **
| +--------------+
| | | +-- Physunio_modelli **
| | +--+
+--+ +-- Physunio **
|
| +----------------- Trapezoideus_foliaceus **
| |
+--+ +-- Lens_contradens **
| +-----------+
| | +-- Lens_micropterus **
+--+
| +----------- Lens_misellus **
+--+
| +-------- Lens_eximius **
+--+
| +----- Lens_pallegoixi **
+--+
| +-- Lens_comptus **
+--+
+-- Lens_novoselovi **
Pfeiffer et al., 2019, fig. 4-5
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 52 |
52 |
49 |
2 |
5 |
4 |
2 |
94.0 |
+----- Anodontites_elongata
+--------------------------------+
| | +-- Etheria_elliptica
| +--+
| +-- Aspatharia_pfeifferiana
|
| +----------------------------------- Hyridella_australis
| |
--+ | +-- Margaritifera_margaritifera **
| | +-----------------------------+
| | | +-- Margaritifera_hembeli **
| | |
| | | +-- Quadrula_quadrula **
| | | +--------------------+
| | | | +-- Uniomerus_tetralasmus **
+--+ | |
| | | +----- Elliptoideus_sloatianus **
| | +-----+ +--------------+
| | | | | | +-- Elliptio_complanata **
| | | | | +--+
| | | | | +-- Pleurobema_strodeanum **
| | | | |
| | | +--+ +----- Reginaia_ebenus **
| | | | +-----------+
| | | | | | +-- Plectomerus_dombeyanus **
+--+ | | | +--+
| | | | +-- Amblema_plicata **
| | +--+
| | | +-- Psoronaias_semigranosa **
| | | +-----------+
| | | | +-- Popenaias_popeii **
| | +--+
| | | +----------- Obliquaria_reflexa **
| | +--+
| | | +-------- Cyrtonaias_tampicoensis **
| | +--+
| | | +----- Leaunio_lienosus **
| | +--+
| | | +-- Lampsilis_cardium **
| | +--+
| | +-- Pachynaias_spheniopsis **
| |
+--+ +-- Chamberlainia_hainesiana **
| +--------------------+
| | +-- Sinohyriopsis_cumingii **
| |
| | +----------------- Lamprotula_cornuumlunae **
| | +--+
| | | | +-------------- Gonidea_angulata **
| +--+ | +--+
| | | | | +----------- Pseudodon_crebristriatus **
| | | | +--+
| | | | | +-------- Bineurus_anodontinum **
| | | | +--+
| | | | | +----- Pilsbryoconcha_exilis **
| | | | +--+
| | +--+ | +-- Monodontina_vondembuschiana **
| | | +--+
| | | +-- Pseudodon_inoscularis **
| | |
| | | +----- Ensidens_ingallsianus **
| | | +-----+
| | | | | +-- Hyriopsis_bialata **
| | | | +--+
| | | | +-- Rectidens_sumatrensis **
| | +--------+
| | | +-- Trapezoideus_foliaceus **
| | | +-----+
+--+ | | +-- Physunio_superbus **
| +--+
| | +----- Yaukthwa_paiensis **
| +--+
| | +-- Lens_contradens **
| +--+
| +-- Solenaia_emarginata **
|
| +----------- Lanceolaria_lanceolata **
| |
| | +----- Unio_gibbus **
| +--------+ +--+
| | | | | +-- Nodularia_douglasiae **
| | | | +--+
| | | | +-- Tchangsinaia_piscicula **
| | +--+
| | | +-- Cristaria_plicata **
| | | +--+
| | | | +-- Sinanodonta_woodiana **
+-----+ +--+
| | +-- Anodonta_cygnea **
| +--+
| +-- Pyganodon_grandis **
|
| +-- Trapezidens_dolichorhynchus **
| +--------------+
| | +-- Lamellidens_savadiensis **
| |
+--+ +-- Leoparreysia_tavoyensis **
| +-----------+
| | +-- Leoparreysia_olivacea **
| |
+--+ +-- Coelatura_choziensis **
| +--------+
| | +-- Prisodontopsis_aviculaeformis **
+--+
| +-------- Indonaia_caerulea **
+--+
| +----- Indochinella_pugio **
+--+
| +-- Scabies_phaselus **
+--+
+-- Harmandia_somboriensis **
Pfeiffer et al., 2019, fig. 6
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 52 |
53 |
49 |
2 |
5 |
4 |
2 |
96.0 |
+----- Anodontites_elongata
+-----------------------+
| | +-- Etheria_elliptica
| +--+
| +-- Aspatharia_pfeifferiana
|
| +----- Hyridella_australis
| +--------------------+
| | | +-- Margaritifera_margaritifera **
| | +--+
| | +-- Margaritifera_hembeli **
| |
| | +-- Quadrula_quadrula **
| | +-----------------+
| | | +-- Uniomerus_tetralasmus **
--+ | |
| | | +----- Elliptoideus_sloatianus **
| | | +--+
| | | | | +-- Elliptio_complanata **
| | +--+ | +--+
| | | | +--------+ +-- Pleurobema_strodeanum **
| | | | | |
| | | | | | +----- Reginaia_ebenus **
| | | | | +--+
| | | | | | +-- Plectomerus_dombeyanus **
| | | +--+ +--+
| | | | +-- Amblema_plicata **
| | | |
| | | | +-- Psoronaias_semigranosa **
| | | | +-----------+
+--+ | | | +-- Popenaias_popeii **
| | +--+
| | | +----------- Obliquaria_reflexa **
| | +--+
| | | +-------- Cyrtonaias_tampicoensis **
| | +--+
| | | +----- Leaunio_lienosus **
| | +--+
| | | +-- Lampsilis_cardium **
| | +--+
| | +-- Pachynaias_spheniopsis **
| |
| | +-- Chamberlainia_hainesiana **
| | +-----------------+
| | | +-- Sinohyriopsis_cumingii **
| | |
| | | +-- Gonidea_angulata **
| | | +-----------+
| | | | +-- Lamprotula_cornuumlunae **
| | | +--+
| +--+ | | +----------- Pseudodon_crebristriatus **
| | | | +--+
| | | | | +-------- Pilsbryoconcha_exilis **
| | | | +--+
+--+ | | | +----- Bineurus_anodontinum **
| | | +--+
| | | | +-- Monodontina_vondembuschiana **
| +--+ +--+
| | +-- Pseudodon_inoscularis **
| |
| | +----- Ensidens_ingallsianus **
| | +-----+
| | | | +-- Hyriopsis_bialata **
| | | +--+
| | | +-- Rectidens_sumatrensis **
| +-----+
| | +-- Trapezoideus_foliaceus **
| | +-----+
| | | +-- Physunio_superbus **
| +--+
| | +----- Yaukthwa_paiensis **
| +--+
| | +-- Lens_contradens **
| +--+
| +-- Solenaia_emarginata **
|
| +-------- Lanceolaria_lanceolata **
| +--+
| | | +----- Unio_gibbus **
| | +--+
| | | +-- Tchangsinaia_piscicula **
| | +--+
| +--------+ +-- Nodularia_douglasiae **
| | |
| | | +-- Cristaria_plicata **
| | | +--+
| | | | +-- Sinanodonta_woodiana **
| | +-----+
| | | +-- Anodonta_cygnea **
+--+ +--+
| +-- Pyganodon_grandis **
|
| +-- Trapezidens_dolichorhynchus **
| +--------------+
| | +-- Lamellidens_savadiensis **
| |
+--+ +-- Leoparreysia_tavoyensis **
| +-----------+
| | +-- Leoparreysia_olivacea **
| |
+--+ +-- Coelatura_choziensis **
| +--------+
| | +-- Prisodontopsis_aviculaeformis **
+--+
| +-------- Indonaia_caerulea **
+--+
| +----- Indochinella_pugio **
+--+
| +-- Scabies_phaselus **
+--+
+-- Harmandia_somboriensis **
Campbell & Lydeard, 2012, fig. 3
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 51 |
51 |
41 |
1 |
0 |
0 |
0 |
114.0 |
+-------------- Uniomerus_declivis **
|
+--------------------------------+ +-- Cyclonaias_infucata **
| | +--------+
| | | +-- Cyclonaias_kleiniana **
| +--+
| | +-------- Quadrula_quadrula **
| +--+
| | +----- Cyclonaias_tuberculata **
| +--+
| | +-- Cyclonaias_succissa **
| +--+
| +-- Cyclonaias_pustulosa **
--+
| +-------------- Elliptoideus_sloatianus **
| +-----------------------------+
| | | +----------- Plectomerus_dombeyanus **
| | +--+
| | | +-------- Obovaria_unicolor **
| | +--+
| | | +----- Obovaria_olivaria **
| | +--+
| | | +-- Lampsilis_ovata **
| | +--+
| | +-- Lemiox_rimosus **
+--+
| +----------- Cyrtonaias_tampicoensis **
| |
| | +----- Popenaias_popeii **
| +-----------------------------+ +--+
| | | | | +-- Amblema_elliottii **
| | | | +--+
| | +--+ +-- Amblema_plicata **
| | |
| | | +-- Reginaia_ebenus[1] **
| | | |
+--+ +-----+-- Reginaia_ebenus[2] **
| |
| +-- Reginaia_rotulata **
|
| +-- Elliptio_arca **
| +-----------------------------------+
| | +-- Elliptio_crassidens **
| |
+--+ +----------------------------------- Eurynia_dilatata **
| |
| | +-------------------------------- Parvaspina_collina **
| | |
+--+ | +-- Hemistena_lata **
| +-----------------------------+
| | +-- Plethobasus_cyphyus **
| |
+--+ +----- Pleuronaia_barnesiana **
| +-----------------------+
| | | +-- Pleuronaia_dolabelloides **
| | +--+
| | +-- Pleuronaia_gibber **
| |
| | +-- Pleurobema_stabile **
+--+ +-----+
| | +-- Pleurobema_sintoxia **
| +-----------------+
| | | +----- Pleurobema_decisum **
| | +--+
| | | +-- Pleurobema_clava **
+--+ +--+
| +-- Pleurobema_pyriforme **
|
| +----------------------- Pleurobema_cordatum **
| |
+--+ +-------------------- Fusconaia_subrotunda **
| |
| | +-- Fusconaia_cor **
+--+ +--------------+
| | +-- Fusconaia_cuneolus **
| |
+--+ +-------------- Fusconaia_ozarkensis **
| |
| | +-- Fusconaia_flava[4] **
| | +--------+
+--+ | +-- Fusconaia_masoni **
| |
| | +-- Fusconaia_escambia **
| | |
+--+ +-- Fusconaia_burkei[1] **
| +-----+
| | +-- Fusconaia_burkei[2] **
| | |
| | +-- Fusconaia_burkei[3] **
+--+
| +----- Fusconaia_flava[3] **
| |
| | +-- Fusconaia_cerina **
+--+ |
| +-- Fusconaia_askewi[1] **
| |
+--+-- Fusconaia_askewi[2] **
|
+-- Fusconaia_flava[1] **
|
+-- Fusconaia_flava[2] **
Bolotov et al., 2022, fig. 8
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 50 |
50 |
42 |
2 |
0 |
0 |
0 |
118.0 |
+-- Gibbosula_laosensis **
|
+-- Margaritifera_dahurica **
|
+-- Sinosolenaia_oleivora **
|
+-----------------------------+-- Leguminaia_wheatleyi **
| |
| +-- Gonidea_angulata **
| |
| +-- Lamprotula_leaii **
| |
| +-- Potomida_littoralis **
|
| +-- Monodontina_lenyanensis **
| +-----------+
| | +-- Sundadontina_tanintharyiensis **
| +-----+
| | | +----------- Pseudodon_kayinensis **
| | +--+
| | | +-------- Pseudodon_inoscularis **
| | +--+
| | | +----- Pseudodon_bogani **
| | +--+
| | | +-- Pseudodon_salwenianus **
--+ +--------+ +--+
| | | +-- Pseudodon_manueli **
| | |
| | | +----- Trapezoideus_mitanensis **
| | | +-----------+
| | | | | +-- Trapezoideus_foliaceus **
| | | | +--+
| | | | +-- Trapezoideus_lenya **
| | +--+
| | | +-- Yaukthwa_elongatula **
| | | +-----------+
| | | | +-- Yaukthwa_baniensis **
| | | |
| | +--+ +----- Yaukthwa_inlenensis **
| | | +-----+
| | | | | +-- Yaukthwa_paiensis **
| | | | +--+
| | +--+ +-- Yaukthwa_avaensis **
| | |
| | | +-------- Yaukthwa_panhai **
+--+ +--+
| | +----- Yaukthwa_rectangularis **
| +--+
| | +-- Yaukthwa_nesemanni **
| +--+
| +-- Yaukthwa_zayleymanensis **
|
| +-------- Lamellidens_savadiensis **
| +-----+
| | | +----- Lamellidens_ferrugineus **
| | +--+
| | | +-- Lamellidens_generosus **
| +-----------+ +--+
| | | +-- Lamellidens_mainwaringi **
| | |
| | | +----------- Trapezidens_exolescens **
| | +--+
| | | +-------- Trapezidens_scutum **
| | +--+
| | | +----- Trapezidens_yeti **
| | +--+
+--+ | +-- Trapezidens_angustior **
| +--+
| +-- Trapezidens_dolichorhynchus **
|
| +----------------------- Parreysia_rakhinensis **
| |
| | +-- Leoparreysia_whitteni **
| | +-----+
| | | +-- Leoparreysia_burmana **
+--+ +-----------+
| | | +----- Leoparreysia_olivacea **
| | +--+
| | | +-- Leoparreysia_canefrii **
| | +--+
+--+ +-- Leoparreysia_tavoyensis **
|
| +-- Indonaia_andersoniana **
| +--------------+
| | +-- Indonaia_subclathrata **
+--+
| +-------------- Indochinella_pugio **
| |
+--+ +----------- Radiatula_crispisulcata **
| |
+--+ +-- Radiatula_chaudhurii **
| +-----+
| | +-- Radiatula_myitkyinae **
+--+
| +----- Radiatula_mouhoti **
+--+
| +-- Radiatula_chindwinensis **
+--+
+-- Radiatula_myitthanensis **
Zieritz et al., 2021, fig. 3-F, 5
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 49 |
49 |
46 |
2 |
0 |
0 |
0 |
78.0 |
+----- Pseudunio_marocanus **
+--------------------------+
| | +-- Margaritifera_falcata **
| +--+
| +-- Margaritifera_dahurica **
|
| +----------- Quadrula_quadrula **
| |
| +-----------------+ +-------- Toxolasma_parvum **
| | | |
| | +--+ +-- Potamilus_alatus **
| | | +--+
| | | | +-- Potamilus_leptodon **
--+ | +--+
| | | +-- Venustaconcha_ellipsiformis **
| | +--+
| | +-- Lampsilis_ornata **
| |
| | +----- Chamberlainia_hainesiana **
| | +--------------+
| | | | +-- Sinohyriopsis_cumingii **
| | | +--+
| | | +-- Sinohyriopsis_schlegelii **
| | |
| | | +-- Lens_contradens **
+--+ +-----+ +--+
| | | | +-- Physunio_superbus **
| | | +-----------+
| | | | | +-- Hyriopsis_bialata **
| | | | +--+
| | | | +-- Rectidens_sumatrensis **
| | | |
| | +--+ +-------- Potomida_littoralis **
| | | +-----+
| | | | | +----- Pronodularia_japanensis **
| | | | +--+
| | | | | +-- Lamprotula_caveata **
| | | | +--+
| | +--+ +-- Lamprotula_leaii **
| | |
| | | +-- Pilsbryoconcha_exilis **
| | | +--------+
+--+ | | +-- Monodontina_vondembuschiana **
| +--+
| | +-------- Ptychorhynchus_pfisteri **
| +--+
| | +----- Microcondylaea_bonellii **
| +--+
| | +-- Sinosolenaia_recognita **
| +--+
| +-- Sinosolenaia_carinata **
|
| +----------------------- Lepidodesma_languilati **
| |
| | +----- Aculamprotula_tortuosa **
| | +--------+
| | | | +-- Aculamprotula_tientsinensis **
| | | +--+
| | | +-- Aculamprotula_coreana **
| | +-----+
+--+ | | +----- Nodularia_douglasiae **
| | | +-----+
| | | | | +-- Schistodesmus_lampreyanus **
| | | | +--+
| | +--+ +-- Tchangsinaia_piscicula **
| | |
| | | +-------- Unio_tumidus **
| | +--+
| | | +----- Unio_crassus **
+--+ +--+
| | +-- Unio_pictorum **
| +--+
| +-- Unio_delphinus **
|
| +-- Lanceolaria_grayii **
| +--------------+
| | +-- Lanceolaria_lanceolata **
| |
| | +----------- Anodonta_anatina **
+--+ +--+
| | | +-------- Lasmigona_compressa **
| | +--+
| | | +----- Pyganodon_grandis **
| | +--+
+--+ | +-- Utterbackia_imbecillis **
| +--+
| +-- Utterbackia_peninsularis **
|
| +-------- Cristaria_plicata **
| |
+-----+ +-- Sinanodonta_lucida **
| +--+
| | +-- Sinanodonta_woodiana **
+--+
| +-- Anemina_arcaeformis **
+--+
+-- Anemina_euscaphys **
Wu et al., 2020, fig. 3
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 46 |
46 |
43 |
2 |
4 |
3 |
2 |
84.0 |
+-------------------------------------- Echyridella_menziesii
|
| +-- Anodontites_trapesialis
| +--------------------------------+
| | +-- Mutela_dubia
--+ |
| | +-------- Gibbosula_rochechouartii **
| | +-----------------------+
| | | | +----- Cumberlandia_monodonta **
| | | +--+
+--+ | | +-- Margaritifera_falcata **
| | +--+
| | +-- Margaritifera_dahurica **
| |
| | +----------- Quadrula_quadrula **
| | |
| | +-----------------+ +-------- Toxolasma_parvum **
+--+ | | |
| | +--+ +-- Venustaconcha_ellipsiformis **
| | | +--+
| | | | +-- Lampsilis_ornata **
| | +--+
| | | +-- Potamilus_alatus **
| | +--+
| | +-- Potamilus_leptodon **
| |
| | +-- Sinohyriopsis_schlegelii **
| | +--------+
+--+ | +-- Sinohyriopsis_cumingii **
| |
| +--------------+ +----- Potomida_littoralis **
| | | +--+
| | | | | +-- Pronodularia_japanensis **
| | | | +--+
| | +--+ +-- Lamprotula_leaii **
| | |
| | | +----- Ptychorhynchus_pfisteri **
| | +--+
| | | +-- Sinosolenaia_oleivora **
| | +--+
| | +-- Sinosolenaia_carinata **
+--+
| +----------------------- Lepidodesma_languilati **
| |
| | +----- Acuticosta_chinensis **
| | +-----------+
| | | | +-- Lanceolaria_lanceolata **
| | | +--+
| | | +-- Lanceolaria_grayii **
| | |
| | +--+ +----------- Anodonta_anatina **
| | | | +--+
| | | | | | +-------- Lasmigona_compressa **
+--+ | | | +--+
| | | | | +----- Pyganodon_grandis **
| | | | +--+
| | +--+ | +-- Utterbackia_peninsularis **
| | | +--+
| | | +-- Utterbackia_imbecillis **
| | |
| | | +-------- Cristaria_plicata **
| | | |
| | +-----+ +-- Sinanodonta_woodiana **
| | | +--+
+--+ | | +-- Sinanodonta_lauta **
| +--+
| | +-- Anemina_arcaeformis **
| +--+
| +-- Anemina_euscaphys **
|
| +-------- Aculamprotula_tortuosa **
| |
| +--------+ +-- Aculamprotula_coreana **
| | | +--+
| | | | +-- Aculamprotula_tientsinensis **
| | +--+
| | | +-- Aculamprotula_polysticta **
| | +--+
+--+ +-- Aculamprotula_scripta **
|
| +-------- Unio_tumidus **
| +-----+
| | | +----- Unio_crassus **
| | +--+
| | | +-- Unio_delphinus **
+--+ +--+
| +-- Unio_pictorum **
|
| +----------- Pseudocuneopsis_capitata **
+--+
| +-------- Nodularia_douglasiae **
+--+
| +----- Tchangsinaia_piscicula **
+--+
| +-- Cuneopsis_heudei **
+--+
+-- Schistodesmus_lampreyanus **
Burlakova et al., 2012, fig. 1
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 45 |
45 |
42 |
1 |
0 |
0 |
0 |
113.0 |
+-- Amblema_elliottii **
+--------------------------------------------+
| +-- Amblema_plicata **
|
--+ +-- Lampsilis_ovata **
| +-----------------------------------------+
| | +-- Plectomerus_dombeyanus **
| |
+--+ +----------------------------------------- Popenaias_popeii **
| |
| | +-------------------------------------- Cyrtonaias_tampicoensis **
+--+ |
| | +----------------------------------- Parvaspina_collina **
| | |
+--+ | +----- Plethobasus_cyphyus **
| | +--------------------------+
| | | | +-- Elliptio_arca **
| | | +--+
| | | +-- Elliptio_crassidens **
+--+ |
| | +----------------------- Fusconaia_subrotunda **
| | |
| | | +-- Fusconaia_cor **
| | +-----+ +-----------------+
| | | | | +-- Fusconaia_cuneolus **
| | | | |
| | | +--+ +----------------- Fusconaia_ozarkensis **
+--+ | | |
| | | | +-------- Fusconaia_askewi[4] **
| | | | +-----+
| | +--+ | | +----- Fusconaia_askewi[3] **
| | | | +--+
| | | | | +-- Fusconaia_lananensis **
| | | | +--+
| | +--+ +-- Fusconaia_askewi[2] **
| | |
| | | +-- Fusconaia_burkei **
| | | +--------+
| | | | +-- Fusconaia_escambia **
+--+ +--+
| | +-------- Fusconaia_masoni **
| +--+
| | +----- Fusconaia_cerina **
| +--+
| | +-- Fusconaia_askewi[1] **
| +--+
| +-- Fusconaia_flava **
|
| +-- Eurynia_dilatata **
| +-----------------------+
| | +-- Hemistena_lata **
| |
| | +----- Pleuronaia_gibber **
| | +-----+
| | | | +-- Pleuronaia_dolabelloides **
+--+ | +--+
| +-----------+ +-- Pleuronaia_barnesiana **
| | |
| | | +-------- Pleurobema_riddellii **
| | +--+
| | | +----- Pleurobema_sintoxia **
| | +--+
+--+ | +-- Pleurobema_rubrum **
| +--+
| +-- Pleurobema_cordatum **
|
| +-------------------- Pleurobema_stabile **
| |
+--+ +----------------- Pleurobema_pyriforme **
| |
| | +----- Pleurobema_georgianum **
+--+ +--------+
| | | +-- Pleurobema_hanleyianum **
| | +--+
| | +-- Pleurobema_decisum **
+--+
| +-- Pleurobema_clava **
| +--------+
| | +-- Pleurobema_oviforme **
+--+
| +-------- Pleurobema_beadleianum **
| |
+--+ +-- Pleurobema_rubellum **
| +--+
| | +-- Pleurobema_strodeanum **
+--+
| +-- Pleurobema_perovatum **
+--+
+-- Pleurobema_taitianum **
Wu et al., 2019, fig. 3
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 45 |
47 |
44 |
2 |
2 |
2 |
2 |
82.0 |
+-- Echyridella_menziesii
+--------+
| +-- Mutela_dubia
+--------------------+
| | +-------- Gibbosula_rochechouartii **
| +--+
| | +----- Cumberlandia_monodonta **
| +--+
| | +-- Margaritifera_dahurica **
| +--+
| +-- Margaritifera_falcata **
|
| +----------- Quadrula_quadrula **
| |
--+ +-----------------+ +-------- Toxolasma_parvum **
| | | |
| | +--+ +-- Lampsilis_ornata **
| | | +--+
| | | | +-- Venustaconcha_ellipsiformis **
| | +--+
| | | +-- Potamilus_leptodon **
| | +--+
| | +-- Potamilus_alatus **
| |
| | +-- Sinohyriopsis_cumingii **
| | +--------+
+--+ | +-- Sinohyriopsis_schlegelii **
| |
| +--------------+ +----- Potomida_littoralis **
| | | +--+
| | | | | +-- Lamprotula_leaii **
| | | | +--+
| | +--+ +-- Pronodularia_japanensis **
| | |
| | | +----- Ptychorhynchus_pfisteri **
| | +--+
| | | +-- Sinosolenaia_oleivora **
| | +--+
| | +-- Sinosolenaia_carinata **
+--+
| +----------------------- Lepidodesma_languilati **
| |
| | +-------- Aculamprotula_tortuosa **
| | +--------+
| | | | +----- Aculamprotula_scripta **
| | | +--+
| | | | +-- Aculamprotula_tientsinensis **
| | | +--+
| | | +-- Aculamprotula_coreana **
| | +--+
+--+ | | +-------- Unio_tumidus **
| | | +-----+
| | | | | +----- Unio_crassus **
| | | | +--+
| | | | | +-- Unio_delphinus **
| | +--+ +--+
| | | +-- Unio_pictorum **
| | |
| | | +----------- Pseudocuneopsis_capitata **
| | +--+
| | | +-------- Nodularia_douglasiae **
+--+ +--+
| | +----- Tchangsinaia_piscicula **
| +--+
| | +-- Cuneopsis_heudei **
| +--+
| +-- Schistodesmus_lampreyanus **
|
| +----- Acuticosta_chinensis **
| +-----------+
| | | +-- Lanceolaria_grayii **
| | +--+
| | +-- Lanceolaria_lanceolata **
| |
+--+ +----------- Anodonta_anatina **
| +--+
| | | +-------- Lasmigona_compressa **
| | +--+
| | | +----- Pyganodon_grandis **
| | +--+
+--+ | +-- Utterbackia_peninsularis **
| +--+
| +-- Utterbackia_imbecillis **
|
| +-------- Cristaria_plicata **
| |
+-----+ +-- Anemina_arcaeformis **
| +--+
| | +-- Anemina_euscaphys **
+--+
| +-- Sinanodonta_lucida **
+--+
+-- Sinanodonta_woodiana **
Froufe et al., 2019, fig. 3
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 45 |
45 |
42 |
2 |
0 |
0 |
0 |
75.0 |
+----- Pseudunio_marocanus **
+--------------------------+
| | +-- Margaritifera_dahurica **
| +--+
| +-- Margaritifera_falcata **
|
| +----------- Quadrula_quadrula **
| |
| +-----------------+ +-------- Toxolasma_parvum **
| | | |
| | +--+ +-- Lampsilis_ornata **
| | | +--+
--+ | | | +-- Venustaconcha_ellipsiformis **
| | +--+
| | | +-- Potamilus_leptodon **
| | +--+
| | +-- Potamilus_alatus **
| |
| | +----- Chamberlainia_hainesiana **
| | +-----------+
| | | | +-- Sinohyriopsis_cumingii **
| | | +--+
| | | +-- Sinohyriopsis_schlegelii **
+--+ |
| +--------+ +-------- Potomida_littoralis **
| | | +-----+
| | | | | +----- Pronodularia_japanensis **
| | | | +--+
| | | | | +-- Lamprotula_leaii **
| | | | +--+
| | +--+ +-- Lamprotula_caveata **
| | |
| | | +-- Pilsbryoconcha_exilis **
| | | +--------+
| | | | +-- Monodontina_vondembuschiana **
| | +--+
| | | +-------- Ptychorhynchus_pfisteri **
| | +--+
+--+ | +----- Microcondylaea_bonellii **
| +--+
| | +-- Sinosolenaia_carinata **
| +--+
| +-- Sinosolenaia_oleivora **
|
| +----------------------- Lepidodesma_languilati **
| |
| | +----- Aculamprotula_tortuosa **
| | +--------+
| | | | +-- Aculamprotula_tientsinensis **
| | | +--+
| | | +-- Aculamprotula_coreana **
| | +-----+
+--+ | | +----- Nodularia_douglasiae **
| | | +-----+
| | | | | +-- Tchangsinaia_piscicula **
| | | | +--+
| | +--+ +-- Schistodesmus_lampreyanus **
| | |
| | | +-------- Unio_tumidus **
| | +--+
| | | +----- Unio_crassus **
+--+ +--+
| | +-- Unio_pictorum **
| +--+
| +-- Unio_delphinus **
|
| +-- Lanceolaria_lanceolata **
| +--------------+
| | +-- Lanceolaria_grayii **
| |
| | +-------- Cristaria_plicata **
+--+ |
| +-----+ +-- Anemina_arcaeformis **
| | | +--+
| | | | +-- Anemina_euscaphys **
| | +--+
+--+ | +-- Sinanodonta_lucida **
| +--+
| +-- Sinanodonta_woodiana **
|
| +----------- Anodonta_anatina **
+--+
| +-------- Lasmigona_complanata **
+--+
| +----- Pyganodon_grandis **
+--+
| +-- Utterbackia_peninsularis **
+--+
+-- Utterbackia_imbecillis **
Sano et al., 2020, fig. 1
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 45 |
45 |
14 |
2 |
1 |
0 |
0 |
92.0 |
+----- Gibbosula_rochechouartii **
+-----------+
| | +-- Margaritifera_middendorffi **
| +--+
| +-- Margaritifera_laevis **
|
| +-------- Sinosolenaia_oleivora **
| |
| +--+ +----- Inversidens_brandtii **
| | | |
| | +--+ +-- Obovalis_omiensis[1] **
| | | |
| | +--+-- Obovalis_omiensis[2] **
| +--+ |
| | | +-- Ptychorhynchus_pfisteri **
| | |
--+ | | +-- Lamprotula_caveata **
| | | +--+
| | | | +-- Pronodularia_japanensis **
| | +-----+
| | | +-- Sinohyriopsis_schlegelii[1] **
| | | |
| | +--+-- Sinohyriopsis_schlegelii[2] **
| | |
| | +-- Sinohyriopsis_cumingii **
| |
| | +-------- Lanceolaria_grayii[1] **
| | |
| | +-------- Lanceolaria_grayii[2] **
| | |
| | +-------- Cristaria_plicata[1] **
| | |
+--+ +-------- Cristaria_plicata[2] **
| |
| +-------- Sinanodonta_lauta[1] **
| |
| +-------- Sinanodonta_lauta[2] **
| |
| +-------- Beringiana_japonica[1] **
| |
| +-------- Beringiana_japonica[2] **
| |
| +-------- Sinanodonta_calipygos[1] **
| |
| +-------- Sinanodonta_calipygos[2] **
| |
| +-------- Anemina_arcaeformis[1] **
| |
| +-------- Anemina_arcaeformis[2] **
| |
| +-------- Sinanodonta_tumens[1] **
| |
+-----+-------- Sinanodonta_tumens[2] **
|
+-------- Pletholophus_tenuis **
|
+-------- Anemina_arcaeformis[3] **
|
+-------- Lanceolaria_lanceolata **
|
+-------- Cristaria_plicata[3] **
|
+-------- Lanceolaria_gladiola **
|
+-------- Sinanodonta_angula **
|
+-------- Lepidodesma_languilati **
|
| +-- Nodularia_douglasiae **
+-----+
| +-- Cuneopsis_celtiformis **
|
| +-- Aculamprotula_tortuosa **
+-----+
| +-- Acuticosta_chinensis **
|
| +----- Schistodesmus_lampreyanus **
| |
| | +-- Inversiunio_reinianus[1] **
+--+ |
| +-- Inversiunio_reinianus[2] **
| |
| +-- Inversiunio_jokohamensis[1] **
+--+
+-- Inversiunio_jokohamensis[2] **
|
+-- Inversiunio_jokohamensis[3] **
|
+-- Inversiunio_jokohamensis[4] **
Campbell & Lydeard, 2012, fig. 1
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 44 |
44 |
41 |
1 |
0 |
0 |
0 |
114.0 |
+----------- Quadrula_quadrula **
|
+--------------------------------------+ +-- Cyclonaias_infucata **
| | +-----+
| | | +-- Cyclonaias_kleiniana **
| +--+
| | +----- Cyclonaias_tuberculata **
| +--+
| | +-- Cyclonaias_succissa **
| +--+
| +-- Cyclonaias_pustulosa **
|
| +----- Popenaias_popeii **
| +-----------+
--+ | | +-- Amblema_elliottii **
| | +--+
| | +-- Amblema_plicata **
| +-----------------------------+
| | | +----- Cyrtonaias_tampicoensis **
| | | +--------+
| | | | | +-- Reginaia_ebenus **
| | | | +--+
| | +--+ +-- Reginaia_rotulata **
| | |
| | | +----------- Plectomerus_dombeyanus **
| | +--+
| | | +-------- Lemiox_rimosus **
+--+ +--+
| | +----- Lampsilis_ovata **
| +--+
| | +-- Obovaria_unicolor **
| +--+
| +-- Obovaria_olivaria **
|
| +-------------------------------------------- Elliptoideus_sloatianus **
| |
| | +----------------------------------------- Pleurobema_stabile **
+--+ |
| | +----- Pleurobema_pyriforme **
| | +--------------------------------+
+--+ | | +-- Pleurobema_clava **
| | +--+
| | +-- Pleurobema_decisum **
| |
+--+ +-- Pleurobema_cordatum **
| +--------------------------------+
| | +-- Pleurobema_sintoxia **
| |
| | +-- Eurynia_dilatata **
| | +-----+
+--+ | +-- Hemistena_lata **
| +-----------------------+
| | | +----- Plethobasus_cyphyus **
| | +--+
| | | +-- Elliptio_arca **
| | +--+
+--+ +-- Elliptio_crassidens **
|
| +----- Pleuronaia_gibber **
| +-----------------------+
| | | +-- Pleuronaia_barnesiana **
| | +--+
+--+ +-- Pleuronaia_dolabelloides **
|
| +-------------------------- Parvaspina_collina **
| |
+--+ +----------------------- Fusconaia_subrotunda **
| |
+--+ +-- Fusconaia_cor **
| +-----------------+
| | +-- Fusconaia_cuneolus **
+--+
| +----------------- Fusconaia_ozarkensis **
| |
+--+ +-- Fusconaia_burkei **
| +-----------+
| | +-- Fusconaia_escambia **
+--+
| +----------- Fusconaia_masoni **
+--+
| +-------- Fusconaia_cerina[2] **
+--+
| +----- Fusconaia_cerina[1] **
+--+
| +-- Fusconaia_flava **
+--+
+-- Fusconaia_askewi **
Campbell & Lydeard, 2012, fig. 2
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 44 |
44 |
36 |
1 |
0 |
0 |
0 |
113.0 |
+----------- Quadrula_quadrula **
|
+-----------------------+ +-- Cyclonaias_infucata **
| | +-----+
| | | +-- Cyclonaias_kleiniana **
| +--+
| | +----- Cyclonaias_tuberculata **
| +--+
| | +-- Cyclonaias_succissa **
| +--+
| +-- Cyclonaias_pustulosa **
|
| +-- Reginaia_ebenus **
| +--+
| | +-- Reginaia_rotulata **
--+ +-----------+
| | | +-- Amblema_elliottii **
| | +--+
| +--------------+ +-- Amblema_plicata **
| | |
| | | +-------------- Popenaias_popeii **
| | | |
| | +--+ +----------- Cyrtonaias_tampicoensis **
| | | |
| | +--+----------- Plectomerus_dombeyanus **
| | |
| | | +-------- Lemiox_rimosus **
| | +--+
| | | +----- Lampsilis_ovata **
+--+ +--+
| | +-- Obovaria_unicolor **
| +--+
| +-- Obovaria_olivaria **
|
| +----------------------------- Elliptoideus_sloatianus **
| |
| | +-------------------------- Hemistena_lata **
| | |
| | +-------------------------- Plethobasus_cyphyus **
| | |
+--+ | +-- Elliptio_arca **
| +-----------------------+
| | +-- Elliptio_crassidens **
| |
| | +-- Eurynia_dilatata **
| +-----------------------+
| | +-- Parvaspina_collina **
+--+
| +----- Pleuronaia_gibber **
| +-----------------+
| | | +-- Pleuronaia_barnesiana **
| | +--+
| | +-- Pleuronaia_dolabelloides **
| |
| | +-- Pleurobema_cordatum **
| | +--------+
| | | +-- Pleurobema_sintoxia **
| +-----------+
| | | +-------- Pleurobema_stabile **
+--+ +--+
| | +----- Pleurobema_pyriforme **
| +--+
| | +-- Pleurobema_clava **
| +--+
| +-- Pleurobema_decisum **
|
| +----- Fusconaia_subrotunda **
| +--------------+
| | | +-- Fusconaia_cor **
| | +--+
+--+ +-- Fusconaia_cuneolus **
|
| +----------------- Fusconaia_ozarkensis **
| |
+--+ +-- Fusconaia_burkei **
| +-----------+
| | +-- Fusconaia_escambia **
+--+
| +----------- Fusconaia_masoni **
+--+
| +-------- Fusconaia_cerina[2] **
+--+
| +----- Fusconaia_cerina[1] **
+--+
| +-- Fusconaia_flava **
+--+
+-- Fusconaia_askewi **
Campbell & Lydeard, 2012, fig. 4
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 44 |
44 |
40 |
1 |
0 |
0 |
0 |
114.0 |
+----------- Quadrula_quadrula **
|
+-----------------------------------------+ +-- Cyclonaias_infucata **
| | +-----+
| | | +-- Cyclonaias_kleiniana **
| +--+
| | +----- Cyclonaias_tuberculata **
| +--+
| | +-- Cyclonaias_succissa **
| +--+
| +-- Cyclonaias_pustulosa **
|
| +----- Popenaias_popeii **
| +--------+
--+ | | +-- Amblema_elliottii **
| | +--+
| | +-- Amblema_plicata **
| +-----------------------------------+
| | | +----------- Plectomerus_dombeyanus **
| | | |
| | | | +----- Cyrtonaias_tampicoensis **
| | | +-----+
| | +--+ | +-- Reginaia_ebenus **
| | | +--+
| | | +-- Reginaia_rotulata **
| | |
+--+ | +-------- Lemiox_rimosus **
| +--+
| | +----- Lampsilis_ovata **
| +--+
| | +-- Obovaria_unicolor **
| +--+
| +-- Obovaria_olivaria **
|
| +----------------------------------------------- Elliptoideus_sloatianus **
| |
+--+ +-------------------------------------------- Pleurobema_stabile **
| |
+--+ +----------------------------------------- Hemistena_lata **
| |
+--+ +-------------------------------------- Eurynia_dilatata **
| |
| | +-- Pleurobema_cordatum **
+--+ +--------------------------------+
| | +-- Pleurobema_sintoxia **
| |
| | +----- Pleurobema_clava **
+--+ +--------------------------+
| | | +-- Pleurobema_decisum **
| | +--+
| | +-- Pleurobema_pyriforme **
| |
+--+ +-------- Parvaspina_collina **
| +--------------------+
| | | +----- Pleuronaia_gibber **
| | +--+
| | | +-- Pleuronaia_barnesiana **
| | +--+
+--+ +-- Pleuronaia_dolabelloides **
|
| +----- Plethobasus_cyphyus **
| +--------------------+
| | | +-- Elliptio_arca **
| | +--+
+--+ +-- Elliptio_crassidens **
|
| +----------------------- Fusconaia_subrotunda **
| |
+--+ +-- Fusconaia_cor **
| +-----------------+
| | +-- Fusconaia_cuneolus **
+--+
| +----------------- Fusconaia_ozarkensis **
| |
+--+ +-- Fusconaia_burkei **
| +-----------+
| | +-- Fusconaia_escambia **
+--+
| +----------- Fusconaia_masoni **
+--+
| +-------- Fusconaia_cerina[2] **
+--+
| +----- Fusconaia_cerina[1] **
+--+
| +-- Fusconaia_askewi **
+--+
+-- Fusconaia_flava **
Burlakova et al., 2012, fig. 2
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 44 |
44 |
31 |
1 |
0 |
0 |
0 |
112.0 |
+----------------------------------- Parvaspina_collina **
|
| +-------------------------------- Pleurobema_plenum **
| |
--+ | +-- Plethobasus_cyphyus **
| | +-----+
| | | +-- Hemistena_lata **
| | +--------------------+
| | | | +----- Eurynia_dilatata **
+--+ | +--+
| | | +-- Elliptio_arca **
| | +--+
| | +-- Elliptio_crassidens **
| |
| | +-------------------------- Pleurobema_cordatum[2] **
| | |
+--+ | +-------------- Fusconaia_subrotunda **
| | |
| | | +-- Fusconaia_cor **
| | | +--------+
| | +--------+ | +-- Fusconaia_cuneolus **
| | | | |
| | | | | +-------- Fusconaia_flava[4] **
| | | | | |
| | | +--+ +-------- Fusconaia_ozarkensis **
| | | | |
+--+ | | +-------- Fusconaia_masoni **
| | | |
| | | | +----- Fusconaia_askewi[1] **
| | | +--+
| | +--+ | +-- Fusconaia_burkei **
| | | +--+
| | | +-- Fusconaia_escambia **
| | |
| | | +----- Fusconaia_cerina[1] **
| | | |
| | | +----- Fusconaia_cerina[2] **
| | | |
+--+ +--+ +-- Fusconaia_flava[1] **
| | |
| | +-- Fusconaia_flava[2] **
| | |
| +--+-- Fusconaia_flava[3] **
| |
| +-- Fusconaia_askewi[2] **
| |
| +-- Fusconaia_askewi[3] **
|
| +-------- Pleurobema_stabile **
| |
| +-----------+ +----- Pleurobema_riddellii **
| | | |
| | +--+ +-- Pleurobema_cordatum[1] **
| | | |
| | | +-- Pleurobema_rubrum **
| | +--+
| | +-- Pleurobema_sintoxia[1] **
+--+ |
| +-- Pleurobema_sintoxia[2] **
|
| +----- Pleuronaia_barnesiana **
| +-----------+
| | | +-- Pleuronaia_dolabelloides **
| | +--+
| | +-- Pleuronaia_gibber **
+--+
| +-- Pleurobema_georgianum **
| +-----------+
| | +-- Pleurobema_pyriforme **
| |
+--+ +-- Pleurobema_decisum **
| +--------+
| | +-- Pleurobema_hanleyianum **
| |
+--+ +-- Pleurobema_clava **
| +-----+
| | +-- Pleurobema_oviforme **
| |
+--+ +----- Pleurobema_rubellum **
| |
| +----- Pleurobema_beadleianum **
+--+
+----- Pleurobema_strodeanum **
|
| +-- Pleurobema_perovatum **
+--+
+-- Pleurobema_taitianum **
Froufe et al., 2019, fig. 6
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 44 |
44 |
41 |
2 |
0 |
0 |
0 |
74.0 |
+-- Pseudunio_marocanus **
+-----------------------------+
| +-- Margaritifera_falcata **
|
| +----------- Quadrula_quadrula **
| |
| +-----------------+ +-------- Toxolasma_parvum **
| | | |
| | +--+ +-- Potamilus_leptodon **
| | | +--+
| | | | +-- Potamilus_alatus **
--+ | +--+
| | | +-- Lampsilis_ornata **
| | +--+
| | +-- Venustaconcha_ellipsiformis **
| |
| | +----- Chamberlainia_hainesiana **
| | +-----------+
| | | | +-- Sinohyriopsis_cumingii **
| | | +--+
| | | +-- Sinohyriopsis_schlegelii **
+--+ |
| +--------+ +-------- Potomida_littoralis **
| | | +-----+
| | | | | +----- Pronodularia_japanensis **
| | | | +--+
| | | | | +-- Lamprotula_leaii **
| | | | +--+
| | +--+ +-- Lamprotula_caveata **
| | |
| | | +-- Pilsbryoconcha_exilis **
| | | +--------+
| | | | +-- Monodontina_vondembuschiana **
| | +--+
| | | +-------- Ptychorhynchus_pfisteri **
| | +--+
+--+ | +----- Microcondylaea_bonellii **
| +--+
| | +-- Sinosolenaia_carinata **
| +--+
| +-- Sinosolenaia_oleivora **
|
| +----------------------- Lepidodesma_languilati **
| |
| | +----- Aculamprotula_tortuosa **
| | +--------+
| | | | +-- Aculamprotula_tientsinensis **
| | | +--+
| | | +-- Aculamprotula_coreana **
| | +-----+
+--+ | | +----- Nodularia_douglasiae **
| | | +-----+
| | | | | +-- Tchangsinaia_piscicula **
| | | | +--+
| | +--+ +-- Schistodesmus_lampreyanus **
| | |
| | | +-------- Unio_tumidus **
| | +--+
| | | +----- Unio_crassus **
+--+ +--+
| | +-- Unio_pictorum **
| +--+
| +-- Unio_delphinus **
|
| +-- Lanceolaria_lanceolata **
| +--------------+
| | +-- Lanceolaria_grayii **
| |
| | +----------- Anodonta_anatina **
+--+ +--+
| | | +-------- Lasmigona_complanata **
| | +--+
| | | +----- Pyganodon_grandis **
| | +--+
+--+ | +-- Utterbackia_peninsularis **
| +--+
| +-- Utterbackia_imbecillis **
|
| +-------- Cristaria_plicata **
| |
+-----+ +-- Sinanodonta_lucida **
| +--+
| | +-- Sinanodonta_woodiana **
+--+
| +-- Anemina_arcaeformis **
+--+
+-- Anemina_euscaphys **
Inoue et al., 2020, fig. 3
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 44 |
44 |
42 |
1 |
0 |
0 |
0 |
110.0 |
+----------------------------------------------- Arcidens_confragosus **
|
| +-- Cyclonaias_petrina **
| +-----------------------------------------+
| | +-- Quadrula_quadrula **
--+ |
| | +-------------- Parvaspina_collina **
| | |
| | | +-- Eurynia_dilatata **
| | +--------------------------+ +--------+
| | | | | +-- Hemistena_lata **
+--+ | | |
| | +--+ +-- Pleurobema_clava **
| | | +-----+
| | | | +-- Pleurobema_sintoxia **
| | +--+
| | | +----- Pleuronaia_dolabelloides **
| | +--+
| | | +-- Fusconaia_flava **
+--+ +--+
| +-- Fusconaia_subrotunda **
|
| +-- Amblema_plicata **
| +--+
| | +-- Amblema_elliottii **
| +--------------------------------+
| | | +-- Reginaia_ebenus **
| | +--+
| | +-- Reginaia_rotulata **
| |
+--+ +----------------------------------- Toxolasma_lividum **
| |
| | +-------- Potamilus_fragilis **
| | |
| | +-----------------------+ +-- Potamilus_ohiensis **
+--+ | | +--+
| | | | +-- Potamilus_amphichaenus **
| | +--+
| | | +-- Potamilus_metnecktayi **
| | +--+
| | +-- Potamilus_purpuratus **
+--+
| +-------- Ellipsaria_lineolata **
| +--------------------+
| | | +----- Truncilla_macrodon **
| | +--+
| | | +-- Truncilla_donaciformis **
| | +--+
| | +-- Truncilla_cognata **
+--+
| +-------------------------- Leaunio_lienosus **
| |
| | +-------------------- Lampsilis_bracteata **
| | |
| | +--+ +----------------- Lampsilis_fasciola **
| | | | |
+--+ | +--+ +-------------- Lampsilis_ornata **
| | | |
| | +--+ +-- Lampsilis_ovata **
| | | +--------+
| | | | +-- Lampsilis_cariosa **
| | +--+
| | | +-------- Lampsilis_cardium[3] **
+--+ +--+
| | +----- Lampsilis_cardium[2] **
| +--+
| | +-- Lampsilis_satura **
| +--+
| +-- Lampsilis_cardium[1] **
|
| +-- Lampsilis_higginsii **
| +--------------+
| | +-- Ortmanniana_ligamentina **
+-----+
| +-------------- Lampsilis_teres **
| |
+--+ +-- Lampsilis_radiata **
| +--------+
| | +-- Lampsilis_siliquoidea **
+--+
| +-------- Lampsilis_bergmanni **
+--+
| +----- Lampsilis_hydiana **
+--+
| +-- Lampsilis_virescens **
+--+
+-- Lampsilis_straminea **
Burlakova et al., 2012, fig. 3
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 43 |
43 |
26 |
1 |
0 |
0 |
0 |
111.0 |
+----------------------------------- Plectomerus_dombeyanus **
|
| +-------------------------------- Hemistena_lata **
| |
--+ +-------------------------------- Pleurobema_stabile **
| |
| | +----------------------------- Eurynia_dilatata **
| | |
+--+ | +-------- Pleurobema_riddellii **
| | |
| | +-----------------+ +----- Pleurobema_sintoxia **
| | | | |
| | | +--+----- Pleurobema_rubrum[2] **
| | | |
+--+ | | +-- Pleurobema_cordatum **
| | +--+
| | +-- Pleurobema_rubrum[1] **
| |
| | +----------------------- Parvaspina_collina **
| | |
| | | +----- Pleuronaia_gibber **
| | +-----------------+
| | | | +-- Pleuronaia_dolabelloides **
+--+ | +--+
| | +-- Pleuronaia_barnesiana **
| |
| | +----- Plethobasus_cyphyus **
| +-----------------+
| | | +-- Elliptio_arca **
| | +--+
| | +-- Elliptio_crassidens **
| |
| | +-- Pleurobema_clava **
| | +--------+
| | | +-- Pleurobema_oviforme **
| | |
| | | +-------- Pleurobema_decisum **
+--+-----------+ |
| | +-------- Pleurobema_pyriforme **
| | |
| | +-------- Pleurobema_hanleyianum **
| +--+
| +-------- Pleurobema_georgianum **
| |
| | +----- Pleurobema_rubellum **
| | |
| | +----- Pleurobema_beadleianum **
| +--+
| +----- Pleurobema_strodeanum **
| |
| | +-- Pleurobema_perovatum **
| +--+
| +-- Pleurobema_taitianum **
|
| +-------------------- Fusconaia_subrotunda **
| |
| | +-- Fusconaia_cor **
+--+ +--------------+
| | +-- Fusconaia_cuneolus **
| |
| | +-------------- Fusconaia_ozarkensis **
+--+ |
| | +-------- Fusconaia_masoni **
| | |
| | | +-- Fusconaia_burkei **
| | +--+-----+
+--+ | | +-- Fusconaia_escambia **
| | |
| | | +----- Fusconaia_cerina **
| | +--+
| | | +-- Fusconaia_askewi[1] **
| | +--+
+--+ +-- Fusconaia_flava **
|
| +-- Fusconaia_askewi[2] **
| |
| +-- Fusconaia_askewi[3] **
| |
| +-- Fusconaia_askewi[4] **
+--------+
+-- Fusconaia_lananensis[1] **
|
+-- Fusconaia_lananensis[2] **
|
+-- Fusconaia_askewi[5] **
Inoue et al., 2020, fig. 4
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 43 |
43 |
41 |
1 |
0 |
0 |
0 |
109.0 |
+-------------------------------------------------- Arcidens_confragosus **
|
| +-- Cyclonaias_petrina **
| +--------------------------------------------+
--+ | +-- Quadrula_quadrula **
| |
| | +-- Reginaia_ebenus **
| | +-----------------------------------------+
+--+ | +-- Reginaia_rotulata **
| |
| | +-------------- Parvaspina_collina **
| | |
| | | +-- Eurynia_dilatata **
| | +--------------------------+ +--------+
+--+ | | | +-- Hemistena_lata **
| | | |
| | +--+ +-- Pleurobema_clava **
| | | +-----+
| | | | +-- Pleurobema_sintoxia **
| | +--+
| | | +----- Pleuronaia_dolabelloides **
+--+ +--+
| | +-- Fusconaia_flava **
| +--+
| +-- Fusconaia_subrotunda **
|
| +-- Amblema_plicata **
| +-----------------------------------+
| | +-- Amblema_elliottii **
| |
+--+ +----------------------------------- Toxolasma_lividum **
| |
| | +-------------------------------- Ellipsaria_lineolata **
+--+ |
| | +----- Truncilla_cognata **
| | +-----------------------+
| | | | +-- Truncilla_macrodon **
+--+ | +--+
| | +-- Truncilla_donaciformis **
| |
| | +-------- Potamilus_fragilis **
| | |
+--+ +-----------------+ +-- Potamilus_ohiensis **
| | | +--+
| | | | +-- Potamilus_amphichaenus **
| | +--+
| | | +-- Potamilus_metnecktayi **
| | +--+
| | +-- Potamilus_purpuratus **
+--+
| +----------------------- Leaunio_lienosus **
| |
| | +----------------- Lampsilis_bracteata **
| | |
| | +--+ +-- Lampsilis_ornata **
| | | | +-----------+
+--+ | | | +-- Lampsilis_fasciola **
| | +--+
| | | +----------- Lampsilis_ovata **
| | +--+
| | | +-------- Lampsilis_cardium[3] **
| | +--+
+--+ | +----- Lampsilis_cardium[2] **
| +--+
| | +-- Lampsilis_satura **
| +--+
| +-- Lampsilis_cardium[1] **
|
| +-------------- Lampsilis_teres **
| |
| | +-- Lampsilis_higginsii **
+-----+ +--------+
| | +-- Ortmanniana_ligamentina **
| |
+--+ +-- Lampsilis_radiata **
| +-----+
| | +-- Lampsilis_siliquoidea **
| |
+--+ +-- Lampsilis_hydiana **
| +--+
| | +-- Lampsilis_bergmanni **
+--+
| +-- Lampsilis_virescens **
+--+
+-- Lampsilis_straminea **
Smith et al., 2019, fig. 2
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 42 |
42 |
40 |
1 |
0 |
0 |
0 |
111.0 |
+----------------------------------------------- Quadrula_quadrula **
|
| +-- Amblema_plicata **
--+ +-----------------------------------------+
| | +-- Reginaia_ebenus **
| |
+--+ +----------------------------------------- Plectomerus_dombeyanus **
| |
| | +-------------------------------------- Toxolasma_lividum **
+--+ |
| | +----- Popenaias_popeii **
| | +-----------------------------+
+--+ | | +-- Glebula_rotundata **
| | +--+
| | +-- Cyrtonaias_tampicoensis **
| |
| | +-------------------------------- Obliquaria_reflexa **
+--+ |
| | +-- Medionidus_conradicus **
| | +--+
| | | +-- Lemiox_rimosus **
| | +-----------------------+
| | | | +-- Cyprogenia_stegaria **
| | | +--+
+--+ | +-- Ptychobranchus_fasciolaris **
| |
| | +-- Hamiota_subangulata **
| | +--------------------+
| | | +-- Villosa_villosa **
| | |
| | +--+ +-------------------- Ligumia_recta **
| | | | |
+--+ | | | +----- Venustaconcha_ellipsiformis **
| | +--+ +-----------+
| | | | | +-- Obovaria_subrotunda **
| | | | +--+
| | +--+ +-- Epioblasma_rangiana **
| | |
| | | +-------------- Ortmanniana_pectorosa **
| | +--+
| | | +----------- Ortmanniana_ligamentina **
| | +--+
| | | +-------- Lampsilis_ovata **
| | +--+
+--+ | +----- Lampsilis_dolabraeformis **
| +--+
| | +-- Lampsilis_satura **
| +--+
| +-- Lampsilis_cardium **
|
| +-------------- Atlanticoncha_ochracea **
| +-----+
| | | +----------- Ellipsaria_lineolata **
| | +--+
| | | +-------- Truncilla_truncata **
| | +--+
| | | +----- Truncilla_cognata **
| | +--+
+-----+ | +-- Truncilla_macrodon **
| +--+
| +-- Truncilla_donaciformis **
|
| +----------------- Potamilus_capax **
| |
| | +----- Potamilus_inflatus **
+--+ +--------+
| | | +-- Potamilus_fragilis **
| | +--+
| | +-- Potamilus_leptodon **
+--+
| +----- Potamilus_ohiensis **
| +-----+
| | | +-- Potamilus_amphichaenus **
| | +--+
+--+ +-- Potamilus_streckersoni **
|
| +-------- Potamilus_metnecktayi **
+--+
| +----- Potamilus_purpuratus[1] **
+--+
| +-- Potamilus_purpuratus[2] **
+--+
+-- Potamilus_alatus **
Aunins et al., 2018, fig. 1
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 41 |
41 |
38 |
2 |
0 |
0 |
0 |
76.0 |
+----- Cumberlandia_monodonta **
+--------------------------+
| | +-- Margaritifera_falcata **
| +--+
| +-- Margaritifera_dahurica **
|
| +-------- Quadrula_quadrula **
| +--------------------+
| | | +----- Toxolasma_parvum **
--+ | +--+
| | | +-- Lampsilis_ornata **
| | +--+
| | +-- Venustaconcha_ellipsiformis **
| |
| | +-- Sinohyriopsis_cumingii **
| | +--------+
| | | +-- Sinohyriopsis_schlegelii **
+--+ |
| +--------------+ +-- Lamprotula_caveata **
| | | +-----+
| | | | +-- Potomida_littoralis **
| | +--+
| | | +----- Sinosolenaia_oleivora **
| | +--+
| | | +-- Ptychorhynchus_pfisteri **
| | +--+
| | +-- Sinosolenaia_carinata **
+--+
| +----------------------- Lepidodesma_languilati **
| |
| | +----- Aculamprotula_tortuosa **
| | +--------+
| | | | +-- Aculamprotula_coreana **
| | | +--+
| | | +-- Aculamprotula_tientsinensis **
| | |
| | +-----+ +-------- Nodularia_douglasiae **
+--+ | | +--+
| | | | | +----- Tchangsinaia_piscicula **
| | | | +--+
| | | | | +-- Cuneopsis_heudei **
| | +--+ +--+
| | | +-- Schistodesmus_lampreyanus **
| | |
| | | +-------- Unio_tumidus **
| | +--+
| | | +----- Unio_crassus **
+--+ +--+
| | +-- Unio_pictorum **
| +--+
| +-- Unio_delphinus **
|
| +----- Acuticosta_chinensis **
| +-----------+
| | | +-- Lanceolaria_grayii **
| | +--+
| | +-- Lanceolaria_lanceolata **
| |
| | +-------- Cristaria_plicata **
+--+ |
| +-----+ +-- Anemina_arcaeformis **
| | | +--+
| | | | +-- Anemina_euscaphys **
| | +--+
| | | +-- Sinanodonta_woodiana **
| | +--+
+--+ +-- Sinanodonta_lucida **
|
| +-- Anodonta_anatina **
| +--------+
| | +-- Anodonta_cygnea **
| |
+--+ +-- Alasmidonta_heterodon **
| +-----+
| | +-- Alasmidonta_varicosa **
+--+
| +----- Pyganodon_grandis **
+--+
| +-- Utterbackia_imbecillis **
+--+
+-- Utterbackia_peninsularis **
Burdick & White, 2007, fig. 3
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 40 |
40 |
32 |
1 |
0 |
0 |
0 |
106.0 |
+----------------------------------------- Lampsilis_ornata **
|
--+ +-------------------------------------- Parvaspina_collina **
| |
| | +----------------------------------- Reginaia_ebenus **
+--+ |
| | +----- Eurynia_dilatata **
| | +--------------------------+
| | | | +-- Plethobasus_cyphyus **
+--+ | +--+
| | +-- Elliptio_crassidens **
| |
| | +----------------------------- Pleurobema_plenum **
| | |
| | | +-- Pleuronaia_barnesiana **
+--+ +--------------------------+
| | +-- Pleuronaia_gibber **
| |
| | +-------------- Pleurobema_stabile **
| | |
| | | +-- Pleurobema_rubellum **
| | | +-----+
| | | | +-- Pleurobema_strodeanum **
| | +-----------+ +--+
+--+ | | | | +----- Pleurobema_beadleianum **
| | | | +--+
| | | | | +-- Pleurobema_perovatum **
| | | | +--+
| | +--+ +-- Pleurobema_taitianum **
| | |
| | | +-- Pleurobema_clava **
| | | +-----+
| | | | +-- Pleurobema_oviforme **
| | | |
| | +--+ +-- Pleurobema_decisum **
| | | +--+
+--+ | | +-- Pleurobema_hanleyianum **
| +--+
| | +-- Pleurobema_pyriforme **
| +--+
| +-- Pleurobema_georgianum **
|
| +----- Pleurobema_sintoxia[1] **
| |
| +-----------------+ +-- Pleurobema_cordatum[3] **
| | | |
| | +--+-- Pleurobema_sintoxia[2] **
| | |
| | +-- Pleurobema_rubrum **
+--+
| +-- Fusconaia_flava[4] **
| |
| +-----------------+-- Pleurobema_cordatum[1] **
| | |
| | +-- Pleurobema_cordatum[2] **
+--+
| +----------------- Fusconaia_subrotunda **
| |
| | +-- Fusconaia_cor **
+--+ +-----------+
| | +-- Fusconaia_cuneolus **
| |
+--+ +----------- Fusconaia_ozarkensis **
| |
| | +-- Fusconaia_masoni **
+--+ +-----+
| | +-- Fusconaia_flava[3] **
| |
+--+ +----- Fusconaia_flava[1] **
| |
| +----- Fusconaia_cerina[4] **
+--+
| +-- Fusconaia_cerina[1] **
| |
| +-- Fusconaia_cerina[2] **
+--+
+-- Fusconaia_cerina[3] **
|
+-- Fusconaia_flava[2] **
Sano et al., 2017, fig. 2
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 40 |
40 |
32 |
2 |
2 |
0 |
0 |
89.0 |
+-- Margaritifera_laevis **
+-----------------------+
| +-- Margaritifera_middendorffi **
|
| +-- Obovalis_omiensis **
| +--------+
| | +-- Ptychorhynchus_pfisteri **
| |
| +-----------+ +-- Inversidens_brandtii **
| | | +-----+
| | | | +-- Sinosolenaia_oleivora **
| | | |
--+ | +--+ +-- Pronodularia_japanensis **
| | | +--+
| | | | +-- Lamprotula_leaii **
| | +--+
| | | +-- Sinohyriopsis_schlegelii **
| | +--+
| | +-- Sinohyriopsis_cumingii **
| |
| | +-- Cristaria_plicata **
| | +-----------+
| | | +-- Anemina_arcaeformis **
+--+ |
| | +-------- Pletholophus_tenuis **
| +-----+ +--+
| | | | | +----- Sinanodonta_woodiana **
| | | | +--+
| | | | | +-- Sinanodonta_lauta[3] **
| | | | +--+
| | +--+ +-- Sinanodonta_tumens **
| | |
| | | +-- Sinanodonta_lauta[2] **
| | | +-----+
| | | | +-- Beringiana_japonica[2] **
| | +--+
| | | +----- Sinanodonta_lauta[1] **
| | +--+
| | | +-- Beringiana_japonica[1] **
+--+ +--+
| +-- Sinanodonta_calipygos **
|
| +-------- Tchangsinaia_piscicula **
| |
| +--------+ +----- Nodularia_nipponensis[2] **
| | | |
| | | | +-- Nodularia_douglasiae[1] **
| | +--+ |
| | | +-- Nodularia_douglasiae[2] **
| | | |
| | | +-- Nodularia_douglasiae[3] **
| | +--+
| | +-- Nodularia_douglasiae[4] **
+--+ |
| +-- Nodularia_douglasiae[5] **
| |
| +-- Nodularia_nipponensis[1] **
|
| +-- Schistodesmus_lampreyanus **
| +-----------+
| | +-- Acuticosta_ovata **
| |
| | +-------- Lepidodesma_languilati **
+--+ +--+
| | | +----- Lanceolaria_grayii[1] **
| | +--+
| | | +-- Lanceolaria_lanceolata **
+--+ +--+
| +-- Lanceolaria_grayii[2] **
|
| +-------- Inversiunio_jokohamensis **
| |
+--+ +----- Inversiunio_yanagawensis[3] **
| |
+--+ +-- Inversiunio_yanagawensis[1] **
| |
+--+-- Inversiunio_yanagawensis[2] **
|
+-- Inversiunio_reinianus **
Huang et al., 2019, fig. 1
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 40 |
40 |
36 |
2 |
0 |
0 |
0 |
93.0 |
+-- Margaritifera_dahurica **
+--------------------------------------+
| +-- Gibbosula_rochechouartii **
|
| +----- Lamprotula_leaii **
| |
| +--------+ +-- Lamprotula_caveata[1] **
| | | |
| | +--+-- Lamprotula_cornuumlunae **
| | |
--+ +-----------------------+ +-- Lamprotula_caveata[2] **
| | |
| | | +-- Parvasolenaia_rivularis **
| | | +--------+
| | | | +-- Parvasolenaia_triangularis **
| | +--+
| | | +-------- Ptychorhynchus_pfisteri **
| | +--+
| | | +----- Sinosolenaia_carinata **
+--+ +--+
| | +-- Sinosolenaia_recognita **
| +--+
| +-- Sinosolenaia_oleivora **
|
| +----- Sinohyriopsis_cumingii[1] **
| +-----------------------------+
| | | +-- Sinohyriopsis_cumingii[2] **
| | +--+
| | +-- Sinohyriopsis_schlegelii **
+--+
| +-------------------------------- Anemina_arcaeformis **
| |
| | +----------------------------- Acuticosta_chinensis **
+--+ |
| | +----- Sinanodonta_woodiana **
| | +--------------------+
| | | | +-- Sinanodonta_angula **
+--+ | +--+
| | +-- Sinanodonta_lucida **
| |
| | +----------- Aculamprotula_tortuosa **
| | +-----------+
+--+ | | +-------- Aculamprotula_scripta **
| | +--+
| | | +----- Aculamprotula_coreana **
| | +--+
| | | +-- Aculamprotula_fibrosa **
| | +--+
| | +-- Aculamprotula_tientsinensis **
+--+
| +----------- Cristaria_plicata **
| +--------+
| | | +-------- Lanceolaria_gladiola **
| | +--+
| | | +----- Lanceolaria_grayii **
| | +--+
| | | +-- Lanceolaria_lanceolata **
+--+ +--+
| +-- Lanceolaria_triformis **
|
| +----------------- Lepidodesma_languilati **
| |
| | +-- Schistodesmus_lampreyanus **
+--+ +-----------+
| | +-- Schistodesmus_spinosus **
| |
+--+ +----- Nodularia_douglasiae[1] **
| +-----+
| | | +-- Nodularia_douglasiae[2] **
| | +--+
+--+ +-- Nodularia_nuxpersicae **
|
| +-------- Cuneopsis_rufescens **
+--+
| +----- Tchangsinaia_piscicula **
+--+
| +-- Cuneopsis_heudei **
+--+
+-- Cuneopsis_celtiformis **
Huang et al., 2018, fig. 6
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 39 |
39 |
36 |
2 |
4 |
3 |
2 |
76.0 |
+-- Anodontites_trapesialis
+--------------------------+
| +-- Mutela_dubia
|
| +-------------------------- Echyridella_menziesii
| |
| | +-------------- Gibbosula_rochechouartii **
--+ | +--------+
| | | | +----------- Cumberlandia_monodonta **
| | | +--+
| | | | +-------- Pseudunio_marocanus **
| | | +--+
| | | | +----- Margaritifera_falcata **
| | | +--+
+--+ | | +-- Margaritifera_margaritifera **
| | +--+
| | +-- Margaritifera_dahurica **
| |
| | +----------- Quadrula_quadrula **
| | |
| | +-----+ +-------- Toxolasma_parvum **
| | | | |
| | | +--+ +-- Lampsilis_ornata **
| | | | +--+
| | | | | +-- Venustaconcha_ellipsiformis **
+--+ | +--+
| | | +-- Potamilus_leptodon **
| +--+ +--+
| | | +-- Potamilus_alatus **
| | |
| | | +-- Sinohyriopsis_cumingii **
| | | +-----------+
| | | | +-- Sinohyriopsis_schlegelii **
| | | |
| | +--+ +-- Sinosolenaia_oleivora **
| | | +--------+
| | | | +-- Sinosolenaia_carinata **
| | +--+
| | | +-------- Potomida_littoralis **
| | +--+
| | | +----- Pronodularia_japanensis **
| | +--+
| | | +-- Lamprotula_caveata **
+--+ +--+
| +-- Lamprotula_leaii **
|
| +----- Aculamprotula_tortuosa **
| +--------+
| | | +-- Aculamprotula_coreana **
| | +--+
| | +-- Aculamprotula_tientsinensis **
| +--+
| | | +-- Tchangsinaia_piscicula **
| | | +--------+
| | | | +-- Nodularia_douglasiae **
| | +--+
| | | +-------- Unio_tumidus **
| | +--+
| | | +----- Unio_crassus **
| | +--+
+--+ | +-- Unio_delphinus **
| +--+
| +-- Unio_pictorum **
|
| +----------- Anodonta_anatina **
| +--+
| | | +-------- Lasmigona_compressa **
| | +--+
| | | +----- Pyganodon_grandis **
| | +--+
+--+ | +-- Utterbackia_imbecillis **
| +--+
| +-- Utterbackia_peninsularis **
|
| +-------- Cristaria_plicata **
| |
+-----+ +-- Anemina_arcaeformis **
| +--+
| | +-- Anemina_euscaphys **
+--+
| +-- Sinanodonta_woodiana **
+--+
+-- Sinanodonta_lucida **
Pfeiffer et al., 2019, fig. 1
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 39 |
39 |
34 |
1 |
0 |
0 |
0 |
99.0 |
+-------------------------------------- Gonidea_angulata **
|
| +----------- Megalonaias_nervosa **
| +-----------------------+
| | | +-------- Uniomerus_declivis **
| | +--+
--+ | | +----- Tritogonia_verrucosa **
| | +--+
| | | +-- Cyclonaias_tuberculata **
| | +--+
| | +-- Quadrula_quadrula **
| |
| | +-------------- Elliptoideus_sloatianus **
+--+ +-----------------+
| | | +----------- Elliptio_fumata **
| | +--+
| | | +-------- Pleuronaia_barnesiana **
| | +--+
| | | +----- Pleurobema_riddellii **
| | +--+
| | | +-- Fusconaia_flava **
| | +--+
| | +-- Pleurobema_strodeanum **
| |
+--+ +-- Amblema_plicata **
| +--+
| | +-- Amblema_neislerii **
| +-----------------------+
| | | +-- Reginaia_ebenus **
| | +--+
| | +-- Reginaia_rotulata **
| |
| | +----------------- Popenaias_popeii **
| | |
| | +--------+ +-------------- Psoronaias_guatemalensis[2] **
| | | | |
+--+ | +--+ +----------- Psoronaias_guatemalensis[1] **
| | | |
| | +--+ +-------- Psoronaias_semigranosa[3] **
| | | |
| | | | +----- Psoronaias_semigranosa[2] **
| | +--+ |
| | | +----- Psoronaias_usumasintae[2] **
| | | |
| | +--+----- Psoronaias_percompressa **
+--+ |
| | +-- Psoronaias_semigranosa[1] **
| | |
| +--+-- Psoronaias_profunda **
| |
| +-- Psoronaias_usumasintae[1] **
|
| +-- Cyrtonaias_tampicoensis **
| +--------------------+
| | +-- Glebula_rotundata **
| |
+--+ +-------------------- Potamilus_alatus **
| |
| | +-- Truncilla_truncata **
+--+ +--------------+
| | +-- Ptychobranchus_fasciolaris **
| |
+--+ +-------------- Pachynaias_spheniopsis **
| |
| | +-- Cyrtonaias_explicata **
+--+ +--------+
| | +-- Sphenonaias_microdon **
| |
+--+ +-- Lampsilis_cardium **
| +-----+
| | +-- Ortmanniana_ligamentina **
+--+
| +----- Cambarunio_iris **
+--+
| +-- Epioblasma_triquetra **
+--+
+-- Ligumia_recta **
Cha et al., 2018, fig. 1
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 39 |
39 |
36 |
2 |
0 |
0 |
0 |
79.0 |
+----- Pseudunio_marocanus **
+--------------------------+
| | +-- Margaritifera_falcata **
| +--+
| +-- Margaritifera_dahurica **
|
| +----------- Quadrula_quadrula **
| +-----------------+
| | | +-------- Toxolasma_parvum **
| | +--+
--+ | | +----- Lampsilis_ornata **
| | +--+
| | | +-- Potamilus_alatus **
| | +--+
| | +-- Potamilus_leptodon **
| |
| | +-- Sinohyriopsis_cumingii **
| | +-----------+
| | | +-- Sinohyriopsis_schlegelii **
+--+ |
| +-----------+ +----- Ptychorhynchus_pfisteri **
| | | +-----+
| | | | | +-- Sinosolenaia_carinata **
| | | | +--+
| | +--+ +-- Sinosolenaia_oleivora **
| | |
| | | +-------- Potomida_littoralis **
| | +--+
| | | +----- Pronodularia_japanensis **
| | +--+
| | | +-- Lamprotula_caveata **
+--+ +--+
| +-- Lamprotula_leaii **
|
| +----------------------- Lepidodesma_languilati **
| |
| | +----- Aculamprotula_tortuosa **
| | +--------+
| | | | +-- Aculamprotula_coreana **
| | | +--+
| | | +-- Aculamprotula_tientsinensis **
| | +-----+
+--+ | | +----- Schistodesmus_lampreyanus **
| | | +-----+
| | | | | +-- Nodularia_douglasiae **
| | | | +--+
| | +--+ +-- Tchangsinaia_piscicula **
| | |
| | | +-------- Unio_tumidus **
| | +--+
+--+ | +----- Unio_crassus **
| +--+
| | +-- Unio_pictorum **
| +--+
| +-- Unio_delphinus **
|
| +----------------- Lanceolaria_grayii **
| |
| | +----------- Anodonta_anatina **
| | +--+
+--+ | | +-------- Lasmigona_compressa **
| | +--+
| | | +----- Pyganodon_grandis **
| | +--+
+--+ | +-- Utterbackia_peninsularis **
| +--+
| +-- Utterbackia_imbecillis **
|
| +-------- Cristaria_plicata **
| |
+-----+ +-- Anemina_arcaeformis **
| +--+
| | +-- Anemina_euscaphys **
+--+
| +-- Sinanodonta_lucida **
+--+
+-- Sinanodonta_woodiana **
Wu et al., 2022, fig. 2d
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 39 |
39 |
37 |
1 |
0 |
0 |
0 |
79.0 |
+-------------------------------- Lepidodesma_languilati **
|
| +----- Lanceolaria_gladiola **
| +--------------+
| | | +-- Lanceolaria_grayii **
| | +--+
| | +-- Lanceolaria_lanceolata **
| +--------+
| | | +----------------- Acuticosta_chinensis **
| | | |
| | | | +-------- Cristaria_plicata **
| | | | |
--+ | +--+ +-----+ +-- Sinanodonta_woodiana **
| | | | | +--+
| | | | | | +-- Sinanodonta_lucida **
| | | | +--+
| | | | | +-- Anemina_arcaeformis **
| | +--+ +--+
| | | +-- Anemina_euscaphys **
| | |
| | | +-- Anodonta_cygnea **
| | | +--------+
| | | | +-- Anodonta_anatina **
| | +--+
+--+ | +-------- Lasmigona_compressa **
| +--+
| | +----- Pyganodon_grandis **
| +--+
| | +-- Utterbackia_imbecillis **
| +--+
| +-- Utterbackia_peninsularis **
|
| +-------- Aculamprotula_tortuosa **
| |
| +-----------------+ +-- Aculamprotula_tientsinensis **
| | | +--+
| | | | +-- Aculamprotula_coreana **
| | +--+
| | | +-- Aculamprotula_polysticta **
| | +--+
| | +-- Aculamprotula_scripta **
+--+
| +-------- Unio_tumidus **
| +--------------+
| | | +----- Unio_crassus **
| | +--+
| | | +-- Unio_pictorum **
| | +--+
| | +-- Unio_delphinus **
+--+
| +-- Pseudocuneopsis_capitata **
| +-----------------+
| | +-- Pseudocuneopsis_sichuanensis **
| |
| | +-- Nodularia_breviconcha **
+--+ +--------------+
| | +-- Nodularia_douglasiae **
| |
| | +-- Schistodesmus_lampreyanus **
+--+ +-----------+
| | +-- Schistodesmus_spinosus **
| |
| | +-------- Cuneopsis_rufescens **
+--+ +--+
| | | +----- Cuneopsis_celtiformis **
| | +--+
| | | +-- Cuneopsis_heudei **
+--+ +--+
| +-- Cuneopsis_demangei **
|
| +-------- Tchangsinaia_piscicula **
+--+
| +----- Inversiunio_jokohamensis **
+--+
| +-- Inversiunio_reinianus **
+--+
+-- Inversiunio_yanagawensis **
Pavan-Kumar et al., 2022, fig. S5
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 39 |
39 |
36 |
2 |
0 |
0 |
0 |
83.0 |
+-- Margaritifera_dahurica **
+-----------------------+
| +-- Margaritifera_falcata **
|
| +-------------------- Lamellidens_marginalis **
| |
| | +-- Anodonta_anatina **
| | +--------+
| | | +-- Pyganodon_grandis **
| | +-----+
| +--+ | | +-------- Cristaria_plicata **
| | | | | |
| | | | +--+ +-- Anemina_arcaeformis **
| | | | | +--+
| | | | | | +-- Anemina_euscaphys **
--+ | | | +--+
| | | | | +-- Sinanodonta_lucida **
| | +--+ +--+
| | | +-- Sinanodonta_woodiana **
| | |
| | | +----- Acuticosta_chinensis **
| | | +--------+
| | | | | +-- Lanceolaria_lanceolata **
| | | | +--+
| | | | +-- Lanceolaria_gladiola **
| | +--+
| | | +----- Lepidodesma_languilati **
| | | +-----+
| | | | | +-- Aculamprotula_tientsinensis **
| | | | +--+
+--+ +--+ +-- Aculamprotula_tortuosa **
| |
| | +-------- Unio_pictorum **
| +--+
| | +----- Schistodesmus_lampreyanus **
| +--+
| | +-- Nodularia_douglasiae **
| +--+
| +-- Tchangsinaia_piscicula **
|
| +-------------- Quadrula_quadrula **
| |
| +--+ +----------- Toxolasma_parvum **
| | | |
| | +--+ +-------- Potamilus_leptodon **
| | | |
| | +--+ +-- Lampsilis_cardium **
| | | +--+
| | | | +-- Lampsilis_ornata **
| | +--+
| | | +-- Lampsilis_siliquoidea **
+-----+ +--+
| +-- Venustaconcha_ellipsiformis **
|
| +----- Chamberlainia_hainesiana **
| +--------+
| | | +-- Sinohyriopsis_schlegelii **
| | +--+
| | +-- Sinohyriopsis_cumingii **
| |
| | +-- Sinosolenaia_carinata **
+--+ +--+
| | +-- Sinosolenaia_oleivora **
| +-----+
| | | +-- Parvasolenaia_rivularis **
| | +--+
| | +-- Microcondylaea_bonellii **
+--+
| +-- Monodontina_vondembuschiana **
| +-----+
| | +-- Pilsbryoconcha_exilis **
+--+
| +----- Potomida_littoralis **
+--+
| +-- Lamprotula_leaii **
+--+
+-- Pronodularia_japanensis **
Boyer et al., 2011, fig. 4
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 38 |
38 |
32 |
1 |
0 |
0 |
0 |
103.0 |
+-------------- Strophitus_undulatus **
|
| +----- Lasmigona_complanata **
+--------------------+ +-----+
| | | | +-- Lasmigona_costata **
| | | +--+
| +--+ +-- Arcidens_confragosus **
| |
| | +-------- Utterbackia_imbecillis **
| | |
| +--+-------- Alasmidonta_marginata **
| |
| | +----- Pyganodon_grandis **
| +--+
--+ | +-- Anodontoides_ferussacianus **
| +--+
| +-- Lasmigona_compressa **
|
| +----------- Theliderma_metanevra **
| |
| +--------------------+ +-- Tritogonia_verrucosa **
| | | +-----+
| | | | +-- Quadrula_quadrula **
| | +--+
| | | +----- Cyclonaias_pustulosa **
| | +--+
+--+ | +-- Cyclonaias_tuberculata **
| +--+
| +-- Cyclonaias_nodulata **
|
| +----------------------------- Megalonaias_nervosa **
| |
+--+ +-------------------------- Toxolasma_parvum **
| |
| | +----------------------- Amblema_plicata **
+--+ |
| | +-- Reginaia_ebenus **
| | +-----------------+
| | | +-- Obliquaria_reflexa **
+--+ |
| | +-------- Pleurobema_sintoxia **
| +-----------+
| | | +----- Elliptio_crassidens **
| | +--+
| | | +-- Fusconaia_flava **
+--+ +--+
| +-- Eurynia_dilatata **
|
| +----- Potamilus_fragilis **
| +-----------+
| | | +-- Potamilus_ohiensis **
| | +--+
| | +-- Potamilus_alatus **
| |
+--+ +----- Ellipsaria_lineolata **
| +--------+
| | | +-- Truncilla_donaciformis **
| | +--+
| | +-- Truncilla_truncata **
+--+
| +----------- Lampsilis_cardium **
| |
| | +----- Ligumia_recta **
| +-----+
+--+ | +-- Obovaria_olivaria **
| +--+
| +-- Epioblasma_triquetra **
|
| +-------- Lampsilis_teres **
+--+
| +----- Lampsilis_siliquoidea **
+--+
| +-- Ortmanniana_ligamentina **
+--+
+-- Lampsilis_higginsii **
Breton et al., 2011, fig. 3
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 38 |
38 |
34 |
2 |
1 |
1 |
1 |
99.0 |
+-------------------------------------- Echyridella_menziesii
|
| +-------- Cumberlandia_monodonta **
| +--------------------------+
| | | +----- Margaritifera_margaritifera **
| | +--+
--+ | | +-- Margaritifera_falcata **
| | +--+
| | +-- Margaritifera_marrianae **
| |
| | +----- Quadrula_quadrula **
| | +--------------------------+
+--+ | | +-- Cyclonaias_tuberculata **
| | +--+
| | +-- Cyclonaias_pustulosa **
| |
| | +-------------------------- Pronodularia_japanensis **
| | |
| | +--+ +----------------------- Sinohyriopsis_cumingii **
| | | | |
| | | +--+ +-------------------- Cristaria_plicata **
| | | | |
+--+ | | | +-- Lasmigona_complanata **
| | +--+ +--------------+
| | | | +-- Lasmigona_costata **
| | | |
| | +--+ +-------------- Strophitus_undulatus **
| | | |
| | | | +-- Lasmigona_compressa **
| | | | +--+
| | +--+ | +-- Lasmigona_subviridis **
| | | +-----+
| | | | | +-- Alasmidonta_marginata **
+--+ | | +--+
| | | +-- Alasmidonta_undulata **
| +--+
| | +-- Pyganodon_lacustris **
| | +-----+
| | | +-- Pyganodon_grandis **
| +--+
| | +----- Utterbackia_peninsularis[2] **
| +--+
| | +-- Utterbackia_peninsularis[1] **
| +--+
| +-- Utterbackia_imbecillis **
|
| +-------------------------- Megalonaias_nervosa **
| |
+--+ +----------------------- Fusconaia_flava **
| |
+--+ +-------------------- Reginaia_ebenus **
| |
| | +-------- Toxolasma_paulum[2] **
+--+ +--------+
| | | +----- Toxolasma_lividum **
| | +--+
| | | +-- Toxolasma_parvum **
+--+ +--+
| +-- Toxolasma_paulum[1] **
|
| +-------------- Ellipsaria_lineolata **
| |
+--+ +----------- Truncilla_macrodon **
| |
| | +-- Lemiox_rimosus **
+--+ +-----+
| | +-- Potamilus_metnecktayi **
| |
+--+ +-- Cambarunio_iris **
| +--+
| | +-- Venustaconcha_ellipsiformis **
+--+
| +-- Lampsilis_powellii **
| |
+--+-- Lampsilis_ornata **
|
+-- Lampsilis_siliquoidea **
Pfeiffer et al., 2016, fig. 2
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 38 |
38 |
33 |
1 |
0 |
0 |
0 |
104.0 |
+-------------------------------------- Utterbackiana_suborbiculata **
|
| +-- Amblema_plicata **
| +--+
| | +-- Amblema_neislerii **
| +-----------------------------+
--+ | | +-- Reginaia_rotulata **
| | +--+
| | +-- Reginaia_ebenus **
| |
| | +-- Cyrtonaias_tampicoensis **
| | +-----+
| | | +-- Glebula_rotundata **
+--+ |
| +-----------------------+ +-- Lampsilis_teres **
| | | +--+
| | | | +-- Leaunio_lienosus **
| | +--+
| | | +-- Truncilla_macrodon **
| | +--+
| | +-- Potamilus_fragilis **
+--+
| +----------------------------- Elliptoideus_sloatianus **
| |
| | +-------------- Uniomerus_declivis **
| | +-----------+
| | | | +----------- Quadrula_quadrula **
| | | +--+
+--+ | | +-------- Tritogonia_verrucosa **
| | +--+
| | | +----- Cyclonaias_pustulosa **
| | +--+
| | | +-- Cyclonaias_petrina **
| | +--+
+--+ +-- Cyclonaias_tuberculata **
|
| +-- Elliptio_fumata **
| +--------------------+
| | +-- Elliptio_crassidens **
| |
| | +-------------------- Parvaspina_collina **
| | |
| | +-------------------- Plethobasus_cyphyus **
+--+ |
| | +-- Hemistena_lata **
| +-----------------+
| | +-- Eurynia_dilatata **
| |
| | +-------------- Fusconaia_askewi **
+--+ |
| +--+ +----------- Fusconaia_mitchelli **
| | | |
| | +--+ +-- Fusconaia_escambia **
| | | +-----+
| | | | +-- Fusconaia_burkei **
| | +--+
| | | +----- Fusconaia_flava[2] **
| | +--+
+--+ | +-- Fusconaia_cerina **
| +--+
| +-- Fusconaia_flava[1] **
|
| +----- Pleurobema_riddellii **
| +--+
| | | +-- Pleurobema_sintoxia **
| | +--+
+--------+ +-- Pleurobema_cordatum **
|
| +----- Pleuronaia_dolabelloides **
| |
+--+ +-- Pleurobema_pyriforme **
| |
+--+-- Pleurobema_strodeanum **
|
+-- Pleurobema_clava **
Chase et al., 2018, fig. 4
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 38 |
39 |
36 |
2 |
1 |
1 |
1 |
94.0 |
+-------- Cumberlandia_monodonta **
+--------------------+
| | +----- Margaritifera_falcata **
| +--+
| | +-- Margaritifera_marrianae **
| +--+
| +-- Margaritifera_margaritifera **
|
| +-- Unio_delphinus **
| +--------------------+
| | +-- Unio_tumidus **
| |
| +--+ +-------------------- Sinanodonta_woodiana **
| | | |
| | | | +-- Anodonta_cygnea **
| | | | +--------------+
--+ | +--+ | +-- Anodonta_anatina **
| | | |
| | | | +-- Pyganodon_lacustris **
| | | | +-----+
| | +--+ | +-- Pyganodon_grandis **
| | | +-----+
| | | | | +----- Utterbackia_peninsularis[2] **
| | | | +--+
| | | | | +-- Utterbackia_peninsularis[1] **
| | | | +--+
| | +--+ +-- Utterbackia_imbecillis **
| | |
| | | +-- Lasmigona_costata **
| | | +--------+
| | | | +-- Lasmigona_complanata **
| | | |
+--+ +--+ +-- Alasmidonta_undulata **
| | +-----+
| | | +-- Alasmidonta_marginata **
| +--+
| | +----- Strophitus_undulatus **
| +--+
| | +-- Lasmigona_compressa **
| +--+
| +-- Lasmigona_subviridis **
|
| +-- Echyridella_menziesii
| +--------+
| | +-- Reginaia_ebenus **
| +--------+
| | | +-------- Fusconaia_flava **
| | +--+
| | | +----- Megalonaias_nervosa **
| | +--+
| | | +-- Cyclonaias_tuberculata **
| | +--+
+-----+ +-- Cyclonaias_pustulosa **
|
| +-------- Toxolasma_parvum **
| +--------+
| | | +----- Toxolasma_lividum **
| | +--+
| | | +-- Toxolasma_paulum **
| | +--+
+--+ +-- Toxolasma_texasiense **
|
| +-------------- Venustaconcha_ellipsiformis **
| |
| | +----- Lemiox_rimosus **
+--+ +-----+
| | | +-- Lampsilis_powellii **
| | +--+
+--+ +-- Lampsilis_siliquoidea **
|
| +-------- Potamilus_metnecktayi **
+--+
| +----- Cambarunio_iris **
+--+
| +-- Ellipsaria_lineolata **
+--+
+-- Truncilla_macrodon **
Perkins et al., 2017, fig. 2
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 37 |
37 |
34 |
1 |
0 |
0 |
0 |
109.0 |
+----------------------------------------- Amblema_plicata **
|
+----------------------------------------- Plectomerus_dombeyanus **
|
| +-- Pleurobema_clava **
| +-----------+
| | +-- Pleurobema_oviforme **
| |
| | +-------- Pleurobema_pyriforme **
| +-----------------------+ +--+
--+ | | | | +----- Pleurobema_georgianum **
| | | | +--+
| | | | | +-- Pleurobema_hanleyianum **
| | +--+ +--+
| | | +-- Pleurobema_decisum **
| | |
| | | +-------- Pleurobema_rubellum **
| | +--+
| | | +----- Pleurobema_perovatum **
| | +--+
+--+ | +-- Pleurobema_beadleianum **
| +--+
| +-- Pleurobema_strodeanum **
|
| +----- Pleuronaia_gibber **
| +-----------------------------+
| | | +-- Pleuronaia_barnesiana **
| | +--+
| | +-- Pleuronaia_dolabelloides **
| |
| | +-- Fusconaia_masoni **
+--+ +--+
| | +-- Fusconaia_flava **
| +--------------------------+
| | | +-- Fusconaia_subrotunda **
| | +--+
| | +-- Fusconaia_cor **
+--+
| +-- Parvaspina_collina **
| +--------------------------+
| | +-- Parvaspina_steinstansana **
| |
+--+ +-- Eurynia_dilatata **
| +-----------------------+
| | +-- Hemistena_lata **
+--+
| +----------------------- Elliptio_producta[2] **
| |
+--+ +-------------------- Elliptio_producta[1] **
| |
+--+ +----------------- Elliptio_spinosa **
| |
| | +-------------- Elliptio_icterina **
+--+ |
| | +-------- Elliptio_roanokensis **
| | +--+
+--+ | | +----- Elliptio_complanata **
| | +--+
| | | +-- Elliptio_crassidens **
| | +--+
+--+ +-- Elliptio_pullata **
|
| +-- Elliptio_congaraea **
| +-----+
| | +-- Elliptio_hopetonensis **
| |
+--+ +-- Elliptio_folliculata **
| +--+
| | +-- Elliptio_jayensis **
+--+
| +-- Elliptio_arca **
+--+
+-- Elliptio_arctata **
Pieri et al., 2018, fig. 3
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 37 |
37 |
23 |
1 |
0 |
0 |
0 |
91.0 |
+----------------------------------------- Arcidens_confragosus **
|
| +-- Quadrula_quadrula **
| +-----------------------------------+
--+ | +-- Cyclonaias_petrina **
| |
| | +----------------------------------- Eurynia_dilatata **
| | |
+--+ | +----- Pleurobema_clava **
| +-----------------------------+
| | | +-- Pleurobema_riddellii **
| | +--+
| | +-- Pleurobema_sintoxia **
+--+
| +----- Fusconaia_subrotunda **
| +--------------------------+
| | | +-- Fusconaia_cor **
| | +--+
| | +-- Fusconaia_cuneolus **
| |
+--+ +----------------------------- Fusconaia_ozarkensis **
| |
| | +----------- Fusconaia_cerina[3] **
| | |
| | +--------------+ +-------- Fusconaia_flava[1] **
+--+ | | |
| | +--+ +----- Fusconaia_flava[3] **
| | | |
| | +--+ +-- Fusconaia_cerina[1] **
| | | |
| | +--+-- Fusconaia_cerina[2] **
+--+ |
| +-- Fusconaia_flava[2] **
|
| +----------------------- Fusconaia_masoni **
| |
| | +-- Fusconaia_burkei **
| +--------------------+
+--+ +-- Fusconaia_escambia **
|
| +-------------------- Fusconaia_mitchelli **
| |
+--+ +----------------- Fusconaia_chunii **
| |
| | +-------------- Fusconaia_askewi[9] **
+--+ |
| | +----------- Fusconaia_askewi[6] **
| | |
+--+ +----------- Fusconaia_askewi[7] **
| |
| +----------- Fusconaia_askewi[8] **
| |
| +----------- Fusconaia_lananensis[5] **
+--+
+----------- Fusconaia_lananensis[6] **
|
| +-------- Fusconaia_askewi[5] **
| |
| +-------- Fusconaia_lananensis[4] **
| |
+--+ +----- Fusconaia_askewi[2] **
| |
| +----- Fusconaia_askewi[3] **
| |
+--+----- Fusconaia_askewi[4] **
|
| +-- Fusconaia_askewi[1] **
| |
| +-- Fusconaia_lananensis[1] **
+--+
+-- Fusconaia_lananensis[2] **
|
+-- Fusconaia_lananensis[3] **
Mabuchi & Nishida, 2020, fig. 1
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 37 |
37 |
11 |
1 |
0 |
0 |
0 |
82.0 |
+-------- Obovalis_omiensis **
+-----------+
| | +----- Inversidens_brandtii **
| +--+
| | +-- Sinohyriopsis_schlegelii **
| +--+
| +-- Pronodularia_japanensis **
--+
| +----- Nodularia_douglasiae **
| +-----------+
| | | +-- Lanceolaria_grayii **
| | +--+
| | +-- Inversiunio_reinianus **
| |
+--+ +-------------- Cristaria_plicata **
| |
| | +----------- Sinanodonta_tumens **
| | |
| | | +-- Sinanodonta_lauta[5] **
| | | |
+--+ | +-- Sinanodonta_lauta[6] **
| | |
| | +-- Beringiana_japonica[15] **
| | |
| | +-- Beringiana_japonica[16] **
| | +-----+
| | | +-- Beringiana_japonica[17] **
+--+ | |
| | +-- Beringiana_japonica[18] **
| | |
| | +-- Beringiana_japonica[19] **
| | |
| | +-- Beringiana_japonica[20] **
| |
| | +----- Beringiana_japonica[1] **
| | |
| | +----- Beringiana_japonica[2] **
| | |
+--+ +----- Beringiana_japonica[3] **
| |
| +----- Beringiana_japonica[4] **
| |
| +----- Beringiana_japonica[5] **
| |
| +----- Sinanodonta_lauta[1] **
| |
| +----- Beringiana_japonica[6] **
| |
| +----- Sinanodonta_lauta[2] **
| |
| +----- Sinanodonta_calipygos[1] **
| |
+--+----- Beringiana_japonica[7] **
|
+----- Sinanodonta_lauta[4] **
|
+----- Beringiana_japonica[8] **
|
+----- Beringiana_japonica[9] **
|
+----- Beringiana_japonica[10] **
|
+----- Beringiana_japonica[11] **
|
+----- Beringiana_japonica[12] **
|
+----- Beringiana_japonica[13] **
|
+----- Beringiana_japonica[14] **
|
| +-- Sinanodonta_lauta[3] **
+--+
+-- Sinanodonta_calipygos[2] **
Jeratthitikul et al., 2021, fig. 1
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 37 |
37 |
33 |
1 |
0 |
0 |
0 |
88.0 |
+----- Lens_contradens **
+--+
| | +-- Trapezoideus_foliaceus **
| +--+
+--------------------+ +-- Physunio_superbus **
| |
| | +----- Ensidens_ingallsianus **
| +--+
| | +-- Hyriopsis_bialata **
| +--+
| +-- Rectidens_sumatrensis **
|
| +----- Sinosolenaia_carinata **
| +-----+
| | | +-- Leguminaia_wheatleyi **
--+ | +--+
| | +-- Gonidea_angulata **
| +--------------+
| | | +-- Chamberlainia_hainesiana **
| | | +-----+
| | | | +-- Sinohyriopsis_cumingii **
| | +--+
| | | +----- Potomida_littoralis **
| 
