Unionoida | Unionoidea | Unionidae | Unioninae | Anodontini | Alasmidontina genus Utterbackia F.C. Baker, 1927: cladomics 3 valid species
Cladomics Summary
number of species = 3
analyzed in 102 cladograms in 56 publications
maximum number of terminals in an analysis = 3, mean = 1.53
Basic Monophyly
number of cladograms that test basic monophyly (i.e., more than 1 included species) = 53
maximum number of included terminals = 3, mean = 2.02
proportion of cladograms that support basic monophyly = 0.94
Pairwise Cladogram Clustering
The graphs below depict in two dimensions all pair-wise ingroup distances within the cladome. See the Cladmoics FARQ for more information.
Whelan et al., 2023 [Publ 4287], fig. 2
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
2 |
2 |
0 |
3 |
39 |
31 |
12 |
-- |
*** Top Monophyly Cladogram *** *** Top Sister-Group Cladogram ***
+-------- Pyganodon
+-----+
| | +----- Utterbackia_peninsularis__2 **
| +--+
| | +-- Utterbackia_peninsularis__1 **
| +--+
| +-- Utterbackia_imbecillis **
|
--+ +-------- Utterbackiana
| |
| +--+ +-- Lasmigona_costata
| | | +--+
| | | | +-- Pseudodontoideus
| | +--+
| | | +-- Lasmigona_complanata
| | +--+
+--+ +-- Arcidens
|
| +----- Pressodonta
| +--+
| | | +-- Anodontoides
| | +--+
| | +-- Strophitus
+--+
| +-- Prolasmidonta
| +--+
| | +-- Platynaias
+--+
| +-- Alasminota
+--+
+-- Alasmidonta
Hoeh et al., 1995 [Publ 3391], fig. 2
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
3 |
3 |
1 |
3 |
4 |
4 |
1 |
-- |
+-------- Pyganodon
--+
| +----- Utterbackia_peggyae **
+--+
| +-- Utterbackia_imbecillis **
+--+
+-- Utterbackia_peninsularis **
Wu et al., 2023 [Publ 4339], fig. 4_S5-aa
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
2 |
2 |
0 |
2 |
94 |
5 |
5 |
-- |
+----- Pyganodon
+--+
| | +-- Utterbackia_peninsularis **
| +--+
| +-- Utterbackia_imbecillis **
--+
| +-- Alasmidonta
| +--+
| | +-- Platynaias
+--+
| +-- Prolasmidonta
+--+
+-- Lasmigona
Amaro et al., 2017 [Publ 3791], fig. 3A
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
2 |
2 |
0 |
2 |
10 |
2 |
2 |
-- |
+-------- Platynaias
--+
| +----- Pyganodon
+--+
| +-- Utterbackia_imbecillis **
+--+
+-- Utterbackia_peninsularis **
Wu et al., 2021 [Publ 4366], fig. 4
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
2 |
2 |
0 |
2 |
67 |
4 |
4 |
-- |
+----- Pyganodon
+--+
| | +-- Utterbackia_imbecillis **
| +--+
--+ +-- Utterbackia_peninsularis **
|
| +----- Prolasmidonta
+--+
| +-- Platynaias
+--+
+-- Alasmidonta
Huang et al., 2019 [Publ 3843], fig. 2
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
2 |
2 |
0 |
2 |
196 |
5 |
5 |
-- |
+----- Pyganodon
--+
| +-- Utterbackia_imbecillis **
+--+
+-- Utterbackia_peninsularis **
Huang et al., 2019 [Publ 3843], fig. S3
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
2 |
2 |
0 |
2 |
194 |
5 |
5 |
-- |
+----- Pyganodon
--+
| +-- Utterbackia_imbecillis **
+--+
+-- Utterbackia_peninsularis **
Fonseca et al., 2016 [Publ 3775], fig. 1
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
2 |
2 |
0 |
2 |
30 |
2 |
2 |
-- |
+-------- Platynaias
--+
| +----- Pyganodon
+--+
| +-- Utterbackia_peninsularis **
+--+
+-- Utterbackia_imbecillis **
Huang et al., 2018 [Publ 3828], fig. 6
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
2 |
2 |
0 |
2 |
41 |
2 |
2 |
-- |
+-------- Platynaias
--+
| +----- Pyganodon
+--+
| +-- Utterbackia_imbecillis **
+--+
+-- Utterbackia_peninsularis **
Breton et al., 2011 [Publ 3376], fig. 3
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
2 |
2 |
0 |
3 |
35 |
9 |
5 |
-- |
+----------------- Lasmigona
|
--+ +-------------- Strophitus
| |
+--+ +-- Platynaias
| +--------+
| | +-- Alasmidonta
+--+
| +-------- Pyganodon
+--+
| +----- Utterbackia_peninsularis__2 **
+--+
| +-- Utterbackia_peninsularis__1 **
+--+
+-- Utterbackia_imbecillis **
Wu et al., 2023 [Publ 4339], fig. 4_S5-nt
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
2 |
2 |
0 |
2 |
94 |
5 |
5 |
-- |
+----- Pyganodon
+-----+
| | +-- Utterbackia_imbecillis **
| +--+
--+ +-- Utterbackia_peninsularis **
|
| +-------- Lasmigona
+--+
| +----- Prolasmidonta
+--+
| +-- Alasmidonta
+--+
+-- Platynaias
Lopes-Lima et al., 2017 [Publ 3792], fig. 2
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
2 |
2 |
0 |
2 |
31 |
2 |
2 |
-- |
+-------- Platynaias
--+
| +----- Pyganodon
+--+
| +-- Utterbackia_peninsularis **
+--+
+-- Utterbackia_imbecillis **
Zieritz et al., 2021 [Publ 4231], fig. 3-F_5
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
2 |
2 |
0 |
2 |
47 |
2 |
2 |
-- |
+-------- Platynaias
--+
| +----- Pyganodon
+--+
| +-- Utterbackia_imbecillis **
+--+
+-- Utterbackia_peninsularis **
Amaro et al., 2017 [Publ 3791], fig. 3B
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
2 |
2 |
0 |
2 |
10 |
2 |
2 |
-- |
+-------- Platynaias
--+
| +----- Pyganodon
+--+
| +-- Utterbackia_imbecillis **
+--+
+-- Utterbackia_peninsularis **
Soroka & Burzynski, 2017 [Publ 3837], fig. 1
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
2 |
2 |
0 |
2 |
22 |
2 |
2 |
-- |
+-------- Platynaias
--+
| +----- Pyganodon
+--+
| +-- Utterbackia_imbecillis **
+--+
+-- Utterbackia_peninsularis **
Cha et al., 2018 [Publ 3916], fig. 1
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
2 |
2 |
0 |
2 |
37 |
2 |
2 |
-- |
+-------- Platynaias
--+
| +----- Pyganodon
+--+
| +-- Utterbackia_peninsularis **
+--+
+-- Utterbackia_imbecillis **
Wu et al., 2018 [Publ 4143], fig. 2
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
2 |
2 |
0 |
2 |
20 |
1 |
1 |
-- |
+----- Pyganodon
--+
| +-- Utterbackia_peninsularis **
+--+
+-- Utterbackia_imbecillis **
Huang et al., 2019 [Publ 3843], fig. S4
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
2 |
2 |
0 |
2 |
196 |
5 |
5 |
-- |
+-- Utterbackia_imbecillis **
--+
+-- Utterbackia_peninsularis **
Wu et al., 2024 [Publ 4350], fig. 8b_S2
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
2 |
2 |
0 |
2 |
90 |
5 |
5 |
-- |
+----- Pyganodon
+-----+
| | +-- Utterbackia_imbecillis **
| +--+
--+ +-- Utterbackia_peninsularis **
|
| +-------- Lasmigona
+--+
| +----- Prolasmidonta
+--+
| +-- Alasmidonta
+--+
+-- Platynaias
Guerra et al., 2017 [Publ 3772], fig. 2b
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
2 |
2 |
0 |
2 |
23 |
3 |
2 |
-- |
+-------- Platynaias
--+
| +----- Pyganodon
+--+
| +-- Utterbackia_peninsularis **
+--+
+-- Utterbackia_imbecillis **
Wu et al., 2024 [Publ 4350], fig. 8a_S1
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
2 |
2 |
0 |
2 |
90 |
5 |
5 |
-- |
+----- Pyganodon
+-----+
| | +-- Utterbackia_imbecillis **
| +--+
--+ +-- Utterbackia_peninsularis **
|
| +-------- Lasmigona
+--+
| +----- Prolasmidonta
+--+
| +-- Alasmidonta
+--+
+-- Platynaias
Zhou et al., 2016 [Publ 3710], fig. 1
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
2 |
2 |
0 |
2 |
18 |
2 |
2 |
-- |
+-------- Platynaias
--+
| +----- Pyganodon
+--+
| +-- Utterbackia_imbecillis **
+--+
+-- Utterbackia_peninsularis **
Wu et al., 2021 [Publ 4366], fig. 3-BI
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
2 |
2 |
0 |
2 |
72 |
5 |
4 |
-- |
+----- Pyganodon
+--+
| | +-- Utterbackia_peninsularis **
| +--+
--+ +-- Utterbackia_imbecillis **
|
| +----- Prolasmidonta
+--+
| +-- Alasmidonta
+--+
+-- Platynaias
Huang et al., 2013 [Publ 3629], fig. 3
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
2 |
2 |
0 |
2 |
14 |
2 |
2 |
-- |
+-------- Platynaias
--+
| +----- Pyganodon
+--+
| +-- Utterbackia_imbecillis **
+--+
+-- Utterbackia_peninsularis **
Inoue et al., 2014 [Publ 3635], fig. 3A
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
2 |
2 |
0 |
3 |
18 |
12 |
7 |
-- |
+-------- Pseudodontoideus
+--------+
| | +----- Lasmigona_costata
| +--+
| | +-- Lasmigona_complanata
--+ +--+
| +-- Arcidens
|
| +-------------- Strophitus
| |
+--+ +-- Platynaias
| +--------+
| | +-- Alasmidonta
+--+
| +-------- Pyganodon
+--+
| +----- Utterbackia_peninsularis__2 **
+--+
| +-- Utterbackia_imbecillis **
+--+
+-- Utterbackia_peninsularis__1 **
Wu et al., 2022 [Publ 4238], fig. 3
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
2 |
2 |
0 |
2 |
72 |
2 |
2 |
-- |
+-------- Platynaias
--+
| +----- Pyganodon
+--+
| +-- Utterbackia_imbecillis **
+--+
+-- Utterbackia_peninsularis **
Inoue et al., 2018 [Publ 3858], fig. 1_3a-c
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
2 |
20 |
0 |
3 |
102 |
14 |
8 |
-- |
+-- Alasmidonta_marginata
|
+-- Prolasmidonta
|
+-- Alasmidonta_undulata
|
+-- Beringiana
|
+-- Anodonta_kennerlyi
|
+-- Anodonta_nuttalliana
|
+-- Anodonta_oregonensis
|
+-- Arcidens_confragosus
|
+-- Arcidens_wheeleri
|
+-- Lasmigona_complanata
|
--+-- Platynaias_compressa
|
+-- Lasmigona_costata
|
+-- Platynaias_decorata
|
+-- Platynaias_subviridis
|
+-- Pyganodon_grandis
|
+-- Pyganodon_lacustris
|
+-- Pseudodontoideus
|
+-- Strophitus
|
+-- Utterbackia_imbecillis **
|
+-- Utterbackia_peninsularis__1 **
|
+-- Utterbackia_peninsularis__2 **
Hoeh, 1990 [Publ 2717], fig. 3
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
2 |
15 |
0 |
2 |
13 |
11 |
4 |
-- |
+----- Pyganodon_fragilis
|
+----- Utterbackia_imbecillis **
|
+----- Utterbackia_peggyae **
|
+----- Anodonta_cygnea
|
+----- Anodonta_kennerlyi
|
+----- Utterbackiana_couperiana__1
|
--+----- Utterbackiana_couperiana__2
|
+----- Utterbackiana_suborbiculata
|
| +-- Lasmigona_costata
+--+
| +-- Utterbackiana_implicata
|
| +-- Lasmigona_complanata
| |
| +-- Platynaias
| |
| +-- Pyganodon_cataracta
+--+
+-- Pyganodon_gibbosa
|
+-- Pyganodon_grandis
|
+-- Pyganodon_lacustris
Guerra et al., 2017 [Publ 3772], fig. 2a
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
2 |
2 |
0 |
2 |
23 |
3 |
2 |
-- |
+-------- Platynaias
--+
| +----- Pyganodon
+--+
| +-- Utterbackia_peninsularis **
+--+
+-- Utterbackia_imbecillis **
Amaro et al., 2017 [Publ 3791], fig. 2
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
2 |
2 |
0 |
2 |
14 |
3 |
3 |
-- |
+-- Lasmigona
+-----+
| +-- Platynaias
--+
| +----- Pyganodon
+--+
| +-- Utterbackia_peninsularis **
+--+
+-- Utterbackia_imbecillis **
Wu et al., 2020 [Publ 4121], fig. 3
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
2 |
2 |
0 |
2 |
48 |
2 |
2 |
-- |
+-------- Platynaias
--+
| +----- Pyganodon
+--+
| +-- Utterbackia_peninsularis **
+--+
+-- Utterbackia_imbecillis **
Froufe et al., 2019 [Publ 3936], fig. 3
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
2 |
2 |
0 |
2 |
43 |
2 |
2 |
-- |
+-------- Lasmigona
--+
| +----- Pyganodon
+--+
| +-- Utterbackia_peninsularis **
+--+
+-- Utterbackia_imbecillis **
Aunins et al., 2018 [Publ 3863], fig. 1
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
2 |
2 |
0 |
2 |
39 |
3 |
3 |
-- |
+-- Prolasmidonta
+-----+
| +-- Alasmidonta
--+
| +----- Pyganodon
+--+
| +-- Utterbackia_imbecillis **
+--+
+-- Utterbackia_peninsularis **
Zanatta et al., 2007 [Publ 2698], fig. 1B
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
2 |
2 |
0 |
2 |
14 |
12 |
4 |
-- |
+----- Pyganodon
--+
| +-- Utterbackia_peggyae **
+--+
+-- Utterbackia_imbecillis **
Sano et al., 2020 [Publ 4138], fig. 3
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
2 |
2 |
0 |
2 |
67 |
4 |
4 |
-- |
+----- Pyganodon
+--+
| | +-- Utterbackia_imbecillis **
| +--+
--+ +-- Utterbackia_peninsularis **
|
| +----- Alasmidonta
+--+
| +-- Platynaias
+--+
+-- Prolasmidonta
Riccardi et al., 2019 [Publ 4089], fig. 2
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
2 |
2 |
0 |
2 |
18 |
2 |
2 |
-- |
+-------- Platynaias
--+
| +----- Pyganodon
+--+
| +-- Utterbackia_peninsularis **
+--+
+-- Utterbackia_imbecillis **
Wu et al., 2019 [Publ 3864], fig. 3
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
2 |
2 |
0 |
2 |
45 |
2 |
2 |
-- |
+-------- Platynaias
--+
| +----- Pyganodon
+--+
| +-- Utterbackia_peninsularis **
+--+
+-- Utterbackia_imbecillis **
Wu et al., 2021 [Publ 4366], fig. 3-ML
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
2 |
2 |
0 |
2 |
72 |
5 |
4 |
-- |
+----- Pyganodon
+--+
| | +-- Utterbackia_peninsularis **
| +--+
--+ +-- Utterbackia_imbecillis **
|
| +----- Prolasmidonta
+--+
| +-- Alasmidonta
+--+
+-- Platynaias
Chen et al., 2020 [Publ 4170], fig. 1
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
2 |
27 |
0 |
2 |
32 |
4 |
4 |
-- |
+----- Schistodesmus
+-----+
| | +-- Lepidodesma
| +--+
| +-- Aculamprotula
+-----------+
| | +----- Platynaias
| | +--+
| | | | +-- Utterbackia_imbecillis **
| | | +--+
| +--+ +-- Pyganodon
| |
| | +----- Sinanodonta
| +--+
| | +-- Cristaria
--+ +--+
| +-- Anodonta
|
| +-- Utterbackia_peninsularis **
| +--------+
| | +-- Potamilus_alatus
| +--------+
| | | +-------- Lamprotula
| | +--+
| | | +----- Sinohyriopsis
| | +--+
+--+ | +-- Parvasolenaia
| +--+
| +-- Sinosolenaia
|
| +----------------- Gibbosula
+--+
| +-------------- Quadrula
+--+
| +----------- Toxolasma
+--+
| +-------- Lampsilis_ornata
+--+
| +----- Potamilus_leptodon
+--+
| +-- Lampsilis_siliquoidea
+--+
+-- Lampsilis_powellii
Zanatta et al., 2007 [Publ 2698], fig. 1A
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
2 |
2 |
0 |
2 |
14 |
12 |
4 |
-- |
+----- Pyganodon
--+
| +-- Utterbackia_peggyae **
+--+
+-- Utterbackia_imbecillis **
Hoeh, 1990 [Publ 2717], fig. 2
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
2 |
2 |
0 |
2 |
13 |
11 |
4 |
-- |
+----- Pyganodon
--+
| +-- Utterbackia_imbecillis **
+--+
+-- Utterbackia_peggyae **
Wu et al., 2022 [Publ 4238], fig. 2d
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
2 |
2 |
0 |
2 |
37 |
2 |
2 |
-- |
+-------- Platynaias
--+
| +----- Pyganodon
+--+
| +-- Utterbackia_imbecillis **
+--+
+-- Utterbackia_peninsularis **
Wu et al., 2024 [Publ 4379], fig. 5
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
2 |
2 |
0 |
2 |
52 |
5 |
5 |
-- |
+----- Pyganodon
+-----+
| | +-- Utterbackia_imbecillis **
| +--+
--+ +-- Utterbackia_peninsularis **
|
| +-------- Lasmigona
+--+
| +----- Prolasmidonta
+--+
| +-- Alasmidonta
+--+
+-- Platynaias
Wu et al., 2016 [Publ 3711], fig. 1
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
2 |
2 |
0 |
2 |
19 |
1 |
1 |
-- |
+----- Pyganodon
--+
| +-- Utterbackia_peninsularis **
+--+
+-- Utterbackia_imbecillis **
Huang et al., 2019 [Publ 3843], fig. 3
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
2 |
2 |
0 |
2 |
55 |
2 |
2 |
-- |
+-------- Lasmigona
--+
| +----- Pyganodon
+--+
| +-- Utterbackia_imbecillis **
+--+
+-- Utterbackia_peninsularis **
Sano et al., 2020 [Publ 4138], fig. 2
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
2 |
2 |
0 |
2 |
67 |
4 |
4 |
-- |
+----- Alasmidonta
+--+
| | +-- Prolasmidonta
| +--+
--+ +-- Platynaias
|
| +----- Pyganodon
+--+
| +-- Utterbackia_imbecillis **
+--+
+-- Utterbackia_peninsularis **
Wu et al., 2020 [Publ 4147], fig. 3
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
2 |
2 |
0 |
2 |
27 |
1 |
1 |
-- |
+----- Pyganodon
--+
| +-- Utterbackia_peninsularis **
+--+
+-- Utterbackia_imbecillis **
Hoeh, 1990 [Publ 2717], fig. 4
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
2 |
2 |
0 |
2 |
13 |
11 |
4 |
-- |
+----- Pyganodon
--+
| +-- Utterbackia_imbecillis **
+--+
+-- Utterbackia_peggyae **
Froufe et al., 2019 [Publ 3936], fig. 6
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
2 |
2 |
0 |
2 |
43 |
2 |
2 |
-- |
+-------- Lasmigona
--+
| +----- Pyganodon
+--+
| +-- Utterbackia_peninsularis **
+--+
+-- Utterbackia_imbecillis **
Guerra et al., 2019 [Publ 4168], fig. 1
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
2 |
2 |
0 |
2 |
39 |
3 |
2 |
-- |
+-------- Platynaias
--+
| +----- Pyganodon
+--+
| +-- Utterbackia_peninsularis **
+--+
+-- Utterbackia_imbecillis **
Sano et al., 2020 [Publ 4138], fig. 4
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
2 |
2 |
0 |
2 |
67 |
4 |
4 |
-- |
+----- Alasmidonta
+--+
| | +-- Prolasmidonta
| +--+
--+ +-- Platynaias
|
| +----- Pyganodon
+--+
| +-- Utterbackia_imbecillis **
+--+
+-- Utterbackia_peninsularis **
Guerra et al., 2019 [Publ 4168], fig. 2
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
2 |
2 |
0 |
2 |
39 |
3 |
2 |
-- |
+-------- Platynaias
--+
| +----- Pyganodon
+--+
| +-- Utterbackia_peninsularis **
+--+
+-- Utterbackia_imbecillis **
Chase et al., 2018 [Publ 3847], fig. 4
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
2 |
2 |
0 |
3 |
36 |
9 |
5 |
-- |
+-------- Pyganodon
+--+
| | +----- Utterbackia_peninsularis__2 **
| +--+
| | +-- Utterbackia_peninsularis__1 **
--+ +--+
| +-- Utterbackia_imbecillis **
|
| +-------- Lasmigona
+--+
| +----- Alasmidonta
+--+
| +-- Strophitus
+--+
+-- Platynaias
Campbell, 2000 [Publ 2724], fig. 1-5
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
1 |
1 |
0 |
1 |
2 |
0 |
0 |
-- |
-- Utterbackia_imbecillis
Huang et al., 2019 [Publ 3843], fig. 3-M
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
1 |
1 |
0 |
1 |
22 |
1 |
1 |
-- |
+-- Pyganodon
--+
+-- Utterbackia_peninsularis **
Whelan et al., 2023 [Publ 4287], fig. 4
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
1 |
1 |
0 |
1 |
26 |
24 |
10 |
-- |
+-------------------- Pressodonta
|
| +-------- Utterbackiana
--+ +--------+
| | | +----- Alasminota
| | +--+
| | | +-- Arcidens
+--+ +--+
| +-- Pseudodontoideus
|
| +-------------- Alasmidonta
+--+
| +----------- Platynaias
+--+
| +-------- Strophitus
+--+
| +----- Pyganodon
+--+
| +-- Utterbackia_peninsularis **
+--+
+-- Anodontoides
Amaro et al., 2017 [Publ 3791], fig. 2-M
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
1 |
1 |
0 |
1 |
9 |
2 |
2 |
-- |
-- Utterbackia_peninsularis
Rosenberg et al., 1997 [Publ 2720], fig. 2A
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
1 |
1 |
0 |
1 |
21 |
2 |
1 |
-- |
-- Utterbackia_imbecillis
Smith et al., 2018 [Publ 3790], fig. 1
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
1 |
1 |
0 |
1 |
17 |
16 |
7 |
-- |
+-- Utterbackia_peninsularis **
+--------+
| +-- Pyganodon
|
--+ +-- Anodontoides
| +-----+
| | +-- Strophitus
| |
+--+ +-- Alasmidonta
| +--+
| | +-- Alasminota
+--+
| +-- Pseudodontoideus
+--+
+-- Utterbackiana
Feng et al., 2016 [Publ 3800], fig. 2
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
1 |
1 |
0 |
1 |
14 |
1 |
1 |
-- |
-- Utterbackia_peninsularis
Chase et al., 2018 [Publ 3861], fig. 1-M
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
1 |
1 |
0 |
1 |
10 |
1 |
1 |
-- |
+-- Utterbackia_peninsularis **
--+
+-- Pyganodon
Amaro et al., 2017 [Publ 3791], fig. 3B-M
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
1 |
1 |
0 |
1 |
6 |
1 |
1 |
-- |
+-- Utterbackia_peninsularis **
--+
+-- Pyganodon
Krebs et al., 2003 [Publ 2691], fig. 3
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
1 |
1 |
0 |
1 |
32 |
8 |
6 |
-- |
-- Utterbackia_imbecillis
Dai et al., 2024 [Publ 4358], fig. 4
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
1 |
1 |
0 |
1 |
51 |
2 |
2 |
-- |
+----- Platynaias
--+
| +-- Pyganodon
+--+
+-- Utterbackia_imbecillis **
Dai et al., 2024 [Publ 4386], fig. 6
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
1 |
1 |
0 |
1 |
39 |
1 |
1 |
-- |
+-- Pyganodon
--+
+-- Utterbackia_peninsularis **
Boyer et al., 2011 [Publ 2718], fig. 4
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
1 |
1 |
0 |
1 |
37 |
8 |
7 |
-- |
+-------------- Strophitus
|
| +----- Lasmigona_complanata
--+ +-----+
| | | +-- Lasmigona_costata
| | +--+
+--+ +-- Arcidens
|
| +-------- Utterbackia_imbecillis **
| |
+--+-------- Alasmidonta
|
| +----- Pyganodon
+--+
| +-- Anodontoides
+--+
+-- Platynaias
Lydeard et al., 1996 [Publ 2656], fig. 4
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
1 |
1 |
0 |
1 |
27 |
4 |
4 |
-- |
-- Utterbackia_imbecillis
Huang et al., 2002 [Publ 2657], fig. 1
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
1 |
1 |
0 |
1 |
33 |
4 |
4 |
-- |
+----------- Utterbackiana
--+
| +-------- Utterbackia_imbecillis **
+--+
| +----- Strophitus
+--+
| +-- Alasmidonta
+--+
+-- Lasmigona
Burzynski & Soroka, 2018 [Publ 3862], fig. 5b
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
1 |
1 |
0 |
1 |
9 |
1 |
1 |
-- |
+-- Utterbackia_peninsularis **
--+
+-- Pyganodon
Huang et al., 2002 [Publ 2657], fig. 2
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
1 |
1 |
0 |
1 |
33 |
4 |
4 |
-- |
+----- Utterbackia_imbecillis **
|
--+----- Utterbackiana
|
| +-- Alasmidonta
+--+
+-- Lasmigona
Amaro et al., 2017 [Publ 3791], fig. 3A-M
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
1 |
1 |
0 |
1 |
6 |
1 |
1 |
-- |
+-- Pyganodon
--+
+-- Utterbackia_peninsularis **
Guerra et al., 2017 [Publ 3772], fig. 2a-M
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
1 |
1 |
0 |
1 |
7 |
1 |
1 |
-- |
+-- Utterbackia_peninsularis **
--+
+-- Pyganodon
Whelan et al., 2023 [Publ 4287], fig. 3
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
1 |
1 |
0 |
1 |
27 |
26 |
11 |
-- |
+-------------------- Utterbackiana
|
| +-- Utterbackia_peninsularis **
--+ +--------------+
| | +-- Pyganodon
| |
+--+ +----- Alasmidonta
| +--------+
| | | +-- Arcidens
| | +--+
+--+ +-- Pseudodontoideus
|
| +----------- Alasminota
| |
+--+ +-- Anodontoides
| +-----+
| | +-- Strophitus
+--+
| +----- Pressodonta
+--+
| +-- Platynaias
+--+
+-- Prolasmidonta
Smith, 2021 [Publ 4331], fig. 1
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
1 |
1 |
0 |
1 |
19 |
1 |
1 |
-- |
+-- Pyganodon
--+
+-- Utterbackia_peninsularis **
Burzynski et al., 2017 [Publ 3774], fig. 1-bottom
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
1 |
1 |
0 |
1 |
12 |
1 |
1 |
-- |
+-- Utterbackia_peninsularis **
--+
+-- Pyganodon
Wu et al., 2022 [Publ 4238], fig. 2e
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
1 |
1 |
0 |
1 |
16 |
1 |
1 |
-- |
+-- Utterbackia_peninsularis **
--+
+-- Pyganodon
Fukata & Iigo, 2020 [Publ 4106], fig. 1
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
1 |
1 |
0 |
1 |
53 |
3 |
3 |
-- |
+-- Pyganodon
+--+
| +-- Utterbackia_peninsularis **
--+
| +-- Alasmidonta
+--+
+-- Prolasmidonta
Rosenberg et al., 1997 [Publ 2720], fig. 2B
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
1 |
1 |
0 |
1 |
21 |
2 |
1 |
-- |
-- Utterbackia_imbecillis
Guerra et al., 2019 [Publ 4168], fig. 2-M
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
1 |
1 |
0 |
1 |
21 |
1 |
1 |
-- |
+-- Utterbackia_peninsularis **
--+
+-- Pyganodon
Wen et al., 2017 [Publ 3795], fig. 3-M
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
1 |
1 |
0 |
1 |
5 |
1 |
1 |
-- |
+-- Utterbackia_peninsularis **
--+
+-- Pyganodon
Chase et al., 2018 [Publ 3861], fig. 1
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
1 |
1 |
0 |
1 |
10 |
1 |
1 |
-- |
+-- Utterbackia_peninsularis **
--+
+-- Pyganodon
Froufe et al., 2015 [Publ 3664], fig. 1-M
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
1 |
1 |
0 |
1 |
7 |
1 |
1 |
-- |
+-- Utterbackia_peninsularis **
--+
+-- Pyganodon
Rosenberg et al., 1994 [Publ 2719], fig. 3
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
1 |
1 |
0 |
1 |
18 |
2 |
1 |
-- |
-- Utterbackia_imbecillis
Burzynski et al., 2017 [Publ 3774], fig. 1-top
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
1 |
1 |
0 |
1 |
12 |
1 |
1 |
-- |
+-- Utterbackia_peninsularis **
--+
+-- Pyganodon
Guerra et al., 2019 [Publ 4168], fig. 1-M
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
1 |
1 |
0 |
1 |
21 |
1 |
1 |
-- |
+-- Utterbackia_peninsularis **
--+
+-- Pyganodon
Huang et al., 2013 [Publ 3629], fig. 3-M
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
1 |
1 |
0 |
1 |
5 |
1 |
1 |
-- |
+-- Pyganodon
--+
+-- Utterbackia_peninsularis **
Guerra et al., 2017 [Publ 3772], fig. 2b-M
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
1 |
1 |
0 |
1 |
7 |
1 |
1 |
-- |
+-- Utterbackia_peninsularis **
--+
+-- Pyganodon
Froufe et al., 2015 [Publ 3664], fig. 1
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
1 |
1 |
0 |
1 |
11 |
1 |
1 |
-- |
+-- Utterbackia_peninsularis **
--+
+-- Pyganodon
Lopes-Lima et al., 2017 [Publ 3792], fig. 2-M
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
1 |
1 |
0 |
1 |
9 |
1 |
1 |
-- |
+-- Utterbackia_peninsularis **
--+
+-- Pyganodon
Burzynski & Soroka, 2018 [Publ 3862], fig. 5a
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
1 |
1 |
0 |
1 |
19 |
1 |
1 |
-- |
+-- Utterbackia_peninsularis **
--+
+-- Pyganodon
Fonseca et al., 2016 [Publ 3775], fig. 1-M
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
1 |
1 |
0 |
1 |
8 |
1 |
1 |
-- |
+-- Utterbackia_peninsularis **
--+
+-- Pyganodon
Chase et al., 2018 [Publ 3847], fig. 5
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
1 |
1 |
0 |
1 |
9 |
0 |
0 |
-- |
-- Utterbackia_peninsularis
Burzynski & Soroka, 2018 [Publ 3862], fig. 3
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
1 |
1 |
0 |
1 |
3 |
1 |
1 |
-- |
+-- Utterbackia_peninsularis **
--+
+-- Pyganodon
Froufe et al., 2019 [Publ 3936], fig. 3-M
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
1 |
1 |
0 |
1 |
18 |
1 |
1 |
-- |
+-- Utterbackia_peninsularis **
--+
+-- Pyganodon
Dai et al., 2023 [Publ 4285], fig. 5
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
1 |
1 |
0 |
1 |
52 |
2 |
2 |
-- |
+----- Platynaias
--+
| +-- Pyganodon
+--+
+-- Utterbackia_imbecillis **
Huang et al., 2018 [Publ 3828], fig. 6-M
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
1 |
1 |
0 |
1 |
16 |
1 |
1 |
-- |
+-- Pyganodon
--+
+-- Utterbackia_peninsularis **
Soroka & Burzynski, 2018 [Publ 3762], fig. 3-M
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
1 |
1 |
0 |
1 |
6 |
1 |
1 |
-- |
+-- Pyganodon
--+
+-- Utterbackia_peninsularis **
Soroka & Burzynski, 2018 [Publ 3762], fig. 3
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
1 |
1 |
0 |
1 |
6 |
1 |
1 |
-- |
+-- Pyganodon
--+
+-- Utterbackia_peninsularis **
Mulvey et al., 1997 [Publ 3390], fig. 1_3
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
1 |
1 |
0 |
1 |
4 |
0 |
0 |
-- |
-- Utterbackia_imbecillis
Froufe et al., 2019 [Publ 3936], fig. 4
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
1 |
1 |
0 |
1 |
18 |
1 |
1 |
-- |
+-- Utterbackia_peninsularis **
--+
+-- Pyganodon
Lydeard et al., 1996 [Publ 2656], fig. 3
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
1 |
1 |
0 |
1 |
26 |
5 |
4 |
-- |
+-- Utterbackia_imbecillis **
+-----+
| +-- Utterbackiana_couperiana
--+
| +----- Strophitus
+--+
| +-- Lasmigona
+--+
+-- Alasmidonta
Zieritz et al., 2021 [Publ 4231], fig. 3-M_4
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
1 |
1 |
0 |
1 |
22 |
1 |
1 |
-- |
+-- Utterbackia_peninsularis **
--+
+-- Pyganodon
|