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Page last updated
Sun 06 Nov 2022

The Freshwater Mussels (Unionoida) of the World (and other less consequential bivalves)

FM(U)otW(aolcb) is the web version of the MUSSEL Project Database. Follow the links to browse the data or use the custom Google search field. Either way, you win!

Cladomics pages are experimental — even more so than the rest of this site!

The purpose of the cladomics pages for each genus and family-group level taxon is to provide statistics and references indicating the extent to which published phylogenetic work supports the classification used here.

The Cladomics Summary provides an overview of the number of studies that have included terminal species representing this taxon as well as an indication of how frequently it has been recovered as monophyletic.

Following the summary, the available published clades representing the taxon are listed.

For more information, visit the Cladomics FARQ (frequently asked rhetorical questions) page. Abbreviations for the various statistics are explained on the FARQ!

Search the database pages:

Unionoida | Unionoidea | Unionidae | Unioninae | Anodontini | Alasmidontina

genus Utterbackia F.C. Baker, 1927: cladomics
     3 valid species

Cladomics Summary

number of species = 3

analyzed in 87 cladograms in 47 publications
maximum number of terminal species in an analysis = 3, mean = 1.56

Basic Monophyly

number of cladograms that test basic monophyly (i.e., more than 1 included species) = 44
maximum number of included species = 3, mean = 2.11
proportion of cladograms that support basic monophyly = 0.93


Inoue et al., 2014, fig. 3A

iT tT bi riS oT roT roS iD
3 3 1 3 18 12 6 0.0

*** Top Monophyly Cladogram ***

     +-------- Pyganodon_grandis          
  +--+                                    
  |  |  +----- Utterbackia_peninsularis[2] **
  |  +--+                                 
  |     |  +-- Utterbackia_imbecillis **     
  |     +--+                              
--+        +-- Utterbackia_peninsularis[1] **
  |                                       
  |        +-- Lasmigona_compressa        
  |  +-----+                              
  |  |     +-- Lasmigona_subviridis       
  +--+                                    
     |  +----- Alasmidonta_marginata      
     +--+                                 
        |  +-- Alasmidonta_undulata       
        +--+                              
           +-- Alasmidonta_varicosa       

Smith et al., 2018, fig. 1

iT tT bi riS oT roT roS iD
1 1 0 1 16 15 7 --

*** Top Sister-Group Cladogram ***

                 +-- Utterbackia_peninsularis **        
  +--------------+                                   
  |              +-- Pyganodon_grandis               
  |                                                  
  |              +-- Anodontoides_ferussacianus      
  |     +--------+                                   
  |     |        +-- Anodontoides_denigratus         
--+  +--+                                            
  |  |  |  +-------- Strophitus_undulatus            
  |  |  +--+                                         
  |  |     |  +----- Strophitus_pascagoulaensis      
  |  |     +--+                                      
  |  |        |  +-- Strophitus_radiatus             
  +--+        +--+                                   
     |           +-- Strophitus_williamsi            
     |                                               
     |           +-- Alasmidonta_triangulata         
     |  +--------+                                   
     |  |        +-- Lasmigona_etowaensis            
     |  |                                            
     +--+        +-- Pseudodontoideus_connasaugaensis
        |  +-----+                                   
        |  |     +-- Pseudodontoideus_subvexus       
        |  |                                         
        +--+     +-- Utterbackiana_couperiana        
           |  +--+                                   
           |  |  +-- Utterbackiana_heardi            
           +--+                                      
              |  +-- Utterbackiana_hartfieldorum     
              +--+                                   
                 +-- Utterbackiana_suborbiculata     

Breton et al., 2011, fig. 3

iT tT bi riS oT roT roS iD
3 3 1 3 36 9 4 1.0
           +-- Lasmigona_compressa        
        +--+                              
        |  +-- Lasmigona_subviridis       
  +-----+                                 
  |     |  +-- Alasmidonta_marginata      
  |     +--+                              
  |        +-- Alasmidonta_undulata       
--+                                       
  |        +-- Pyganodon_lacustris        
  |  +-----+                              
  |  |     +-- Pyganodon_grandis          
  +--+                                    
     |  +----- Utterbackia_peninsularis[2] **
     +--+                                 
        |  +-- Utterbackia_peninsularis[1] **
        +--+                              
           +-- Utterbackia_imbecillis **     

Hoeh et al., 1995, fig. 2

iT tT bi riS oT roT roS iD
3 3 1 3 4 4 1 2.0
     +----- Utterbackia_peggyae **     
  +--+                              
  |  |  +-- Utterbackia_imbecillis **  
  |  +--+                           
--+     +-- Utterbackia_peninsularis **
  |                                 
  |  +----- Pyganodon_lacustris     
  |  |                              
  +--+----- Pyganodon_grandis       
     |                              
     |  +-- Pyganodon_cataracta     
     +--+                           
        +-- Pyganodon_fragilis      

Chase et al., 2018, fig. 4

iT tT bi riS oT roT roS iD
3 3 1 3 36 9 4 1.0
              +-- Pyganodon_lacustris        
        +-----+                              
        |     +-- Pyganodon_grandis          
  +-----+                                    
  |     |  +----- Utterbackia_peninsularis[2] **
  |     +--+                                 
  |        |  +-- Utterbackia_peninsularis[1] **
  |        +--+                              
--+           +-- Utterbackia_imbecillis **     
  |                                          
  |           +-- Lasmigona_costata          
  |  +--------+                              
  |  |        +-- Lasmigona_complanata       
  |  |                                       
  +--+        +-- Alasmidonta_undulata       
     |  +-----+                              
     |  |     +-- Alasmidonta_marginata      
     +--+                                    
        |  +----- Strophitus_undulatus       
        +--+                                 
           |  +-- Lasmigona_compressa        
           +--+                              
              +-- Lasmigona_subviridis       

Inoue et al., 2018, fig. 1, 3a-c

iT tT bi riS oT roT roS iD
3 21 0 3 201 14 6 19.0
  +-- Alasmidonta_marginata      
  |                              
  +-- Alasmidonta_heterodon      
  |                              
  +-- Alasmidonta_undulata       
  |                              
  +-- Beringiana_beringiana      
  |                              
  +-- Anodonta_kennerlyi         
  |                              
  +-- Anodonta_nuttalliana       
  |                              
  +-- Anodonta_oregonensis       
  |                              
  +-- Arcidens_confragosus       
  |                              
  +-- Arcidens_wheeleri          
  |                              
  +-- Lasmigona_complanata       
  |                              
--+-- Lasmigona_compressa        
  |                              
  +-- Lasmigona_costata          
  |                              
  +-- Lasmigona_decorata         
  |                              
  +-- Lasmigona_subviridis       
  |                              
  +-- Pyganodon_grandis          
  |                              
  +-- Pyganodon_lacustris        
  |                              
  +-- Pseudodontoideus_subvexus  
  |                              
  +-- Strophitus_undulatus       
  |                              
  +-- Utterbackia_imbecillis **     
  |                              
  +-- Utterbackia_peninsularis[1] **
  |                              
  +-- Utterbackia_peninsularis[2] **

Zanatta et al., 2007, fig. 1A

iT tT bi riS oT roT roS iD
2 2 0 2 14 12 3 --
     +-- Utterbackia_peggyae **   
  +--+                         
  |  +-- Utterbackia_imbecillis **
  |                            
--+  +-- Pyganodon_gibbosa     
  |  |                         
  |  +-- Pyganodon_cataracta   
  |  |                         
  +--+-- Pyganodon_fragilis    
     |                         
     +-- Pyganodon_grandis     
     |                         
     +-- Pyganodon_lacustris   

Zanatta et al., 2007, fig. 1B

iT tT bi riS oT roT roS iD
2 2 0 2 14 12 3 --
              +-- Utterbackia_peggyae **   
  +-----------+                         
  |           +-- Utterbackia_imbecillis **
--+                                     
  |  +----------- Pyganodon_lacustris   
  +--+                                  
     |  +-------- Pyganodon_grandis     
     +--+                               
        |  +----- Pyganodon_fragilis    
        +--+                            
           |  +-- Pyganodon_gibbosa     
           +--+                         
              +-- Pyganodon_cataracta   

Hoeh, 1990, fig. 2

iT tT bi riS oT roT roS iD
2 2 0 2 14 12 3 --
              +-- Utterbackia_imbecillis **
  +-----------+                         
  |           +-- Utterbackia_peggyae **   
--+                                     
  |  +----------- Pyganodon_lacustris   
  +--+                                  
     |  +-------- Pyganodon_grandis     
     +--+                               
        |  +----- Pyganodon_fragilis    
        +--+                            
           |  +-- Pyganodon_cataracta   
           +--+                         
              +-- Pyganodon_gibbosa     

Hoeh, 1990, fig. 3

iT tT bi riS oT roT roS iD
2 16 0 2 14 12 3 16.0
  +----- Pyganodon_fragilis         
  |                                 
  +----- Utterbackia_imbecillis **     
  |                                 
  +----- Utterbackia_peggyae **        
  |                                 
  +----- Anodonta_cygnea            
  |                                 
  +----- Anodonta_kennerlyi         
  |                                 
  +----- Utterbackiana_couperiana[1]
  |                                 
--+----- Utterbackiana_couperiana[2]
  |                                 
  +----- Utterbackiana_suborbiculata
  |                                 
  |  +-- Lasmigona_costata          
  +--+                              
  |  +-- Utterbackiana_implicata    
  |                                 
  |  +-- Lasmigona_complanata       
  |  |                              
  |  +-- Lasmigona_compressa        
  |  |                              
  |  +-- Pyganodon_cataracta        
  +--+                              
     +-- Pyganodon_gibbosa          
     |                              
     +-- Pyganodon_grandis          
     |                              
     +-- Pyganodon_lacustris        

Hoeh, 1990, fig. 4

iT tT bi riS oT roT roS iD
2 2 0 2 14 12 3 --
           +-- Utterbackia_imbecillis **
  +--------+                         
  |        +-- Utterbackia_peggyae **   
--+                                  
  |  +-------- Pyganodon_fragilis    
  |  |                               
  +--+     +-- Pyganodon_cataracta   
     |  +--+                         
     |  |  +-- Pyganodon_gibbosa     
     +--+                            
        |  +-- Pyganodon_lacustris   
        +--+                         
           +-- Pyganodon_grandis     

Huang et al., 2013, fig. 3

iT tT bi riS oT roT roS iD
2 2 0 2 14 2 2 --
  +-------- Lasmigona_compressa     
--+                                 
  |  +----- Pyganodon_grandis       
  +--+                              
     |  +-- Utterbackia_imbecillis **  
     +--+                           
        +-- Utterbackia_peninsularis **

Zhou et al., 2016, fig. 1

iT tT bi riS oT roT roS iD
2 2 0 2 18 2 2 --
  +-------- Lasmigona_compressa     
--+                                 
  |  +----- Pyganodon_grandis       
  +--+                              
     |  +-- Utterbackia_imbecillis **  
     +--+                           
        +-- Utterbackia_peninsularis **

Wu et al., 2016, fig. 1

iT tT bi riS oT roT roS iD
2 2 0 2 19 1 1 --
  +----- Pyganodon_grandis       
--+                              
  |  +-- Utterbackia_peninsularis **
  +--+                           
     +-- Utterbackia_imbecillis **  

Guerra et al., 2017, fig. 2a

iT tT bi riS oT roT roS iD
2 2 0 2 23 3 2 --
        +-- Lasmigona_subviridis    
  +-----+                           
  |     +-- Lasmigona_compressa     
--+                                 
  |  +----- Pyganodon_grandis       
  +--+                              
     |  +-- Utterbackia_peninsularis **
     +--+                           
        +-- Utterbackia_imbecillis **  

Guerra et al., 2017, fig. 2b

iT tT bi riS oT roT roS iD
2 2 0 2 23 3 2 --
        +-- Lasmigona_subviridis    
  +-----+                           
  |     +-- Lasmigona_compressa     
--+                                 
  |  +----- Pyganodon_grandis       
  +--+                              
     |  +-- Utterbackia_peninsularis **
     +--+                           
        +-- Utterbackia_imbecillis **  

Fonseca et al., 2016, fig. 1

iT tT bi riS oT roT roS iD
2 2 0 2 30 2 2 --
  +-------- Lasmigona_compressa     
--+                                 
  |  +----- Pyganodon_grandis       
  +--+                              
     |  +-- Utterbackia_peninsularis **
     +--+                           
        +-- Utterbackia_imbecillis **  

Amaro et al., 2017, fig. 2

iT tT bi riS oT roT roS iD
2 2 0 2 14 3 2 --
        +-- Lasmigona_costata       
  +-----+                           
  |     +-- Lasmigona_compressa     
--+                                 
  |  +----- Pyganodon_grandis       
  +--+                              
     |  +-- Utterbackia_peninsularis **
     +--+                           
        +-- Utterbackia_imbecillis **  

Amaro et al., 2017, fig. 3A

iT tT bi riS oT roT roS iD
2 2 0 2 10 2 2 --
  +-------- Lasmigona_compressa     
--+                                 
  |  +----- Pyganodon_grandis       
  +--+                              
     |  +-- Utterbackia_imbecillis **  
     +--+                           
        +-- Utterbackia_peninsularis **

Amaro et al., 2017, fig. 3B

iT tT bi riS oT roT roS iD
2 2 0 2 10 2 2 --
  +-------- Lasmigona_compressa     
--+                                 
  |  +----- Pyganodon_grandis       
  +--+                              
     |  +-- Utterbackia_imbecillis **  
     +--+                           
        +-- Utterbackia_peninsularis **

Lopes-Lima et al., 2017, fig. 2

iT tT bi riS oT roT roS iD
2 2 0 2 31 2 2 --
  +-------- Lasmigona_compressa     
--+                                 
  |  +----- Pyganodon_grandis       
  +--+                              
     |  +-- Utterbackia_peninsularis **
     +--+                           
        +-- Utterbackia_imbecillis **  

Huang et al., 2018, fig. 6

iT tT bi riS oT roT roS iD
2 2 0 2 41 2 2 --
  +-------- Lasmigona_compressa     
--+                                 
  |  +----- Pyganodon_grandis       
  +--+                              
     |  +-- Utterbackia_imbecillis **  
     +--+                           
        +-- Utterbackia_peninsularis **

Soroka & Burzynski, 2017, fig. 1

iT tT bi riS oT roT roS iD
2 2 0 2 22 2 2 --
  +-------- Lasmigona_compressa     
--+                                 
  |  +----- Pyganodon_grandis       
  +--+                              
     |  +-- Utterbackia_imbecillis **  
     +--+                           
        +-- Utterbackia_peninsularis **

Huang et al., 2019, fig. 2

iT tT bi riS oT roT roS iD
2 2 0 2 198 5 5 --
  +----- Pyganodon_grandis       
--+                              
  |  +-- Utterbackia_imbecillis **  
  +--+                           
     +-- Utterbackia_peninsularis **

Huang et al., 2019, fig. 3

iT tT bi riS oT roT roS iD
2 2 0 2 55 2 2 --
  +-------- Lasmigona_compressa     
--+                                 
  |  +----- Pyganodon_grandis       
  +--+                              
     |  +-- Utterbackia_imbecillis **  
     +--+                           
        +-- Utterbackia_peninsularis **

Huang et al., 2019, fig. S3

iT tT bi riS oT roT roS iD
2 2 0 2 196 5 5 --
  +----- Pyganodon_grandis       
--+                              
  |  +-- Utterbackia_imbecillis **  
  +--+                           
     +-- Utterbackia_peninsularis **

Huang et al., 2019, fig. S4

iT tT bi riS oT roT roS iD
2 2 0 2 199 5 5 --
  +-- Utterbackia_imbecillis **  
--+                           
  +-- Utterbackia_peninsularis **

Aunins et al., 2018, fig. 1

iT tT bi riS oT roT roS iD
2 2 0 2 39 3 2 --
        +-- Alasmidonta_heterodon   
  +-----+                           
  |     +-- Alasmidonta_varicosa    
--+                                 
  |  +----- Pyganodon_grandis       
  +--+                              
     |  +-- Utterbackia_imbecillis **  
     +--+                           
        +-- Utterbackia_peninsularis **

Wu et al., 2019, fig. 3

iT tT bi riS oT roT roS iD
2 2 0 2 45 2 2 --
  +-------- Lasmigona_compressa     
--+                                 
  |  +----- Pyganodon_grandis       
  +--+                              
     |  +-- Utterbackia_peninsularis **
     +--+                           
        +-- Utterbackia_imbecillis **  

Cha et al., 2018, fig. 1

iT tT bi riS oT roT roS iD
2 2 0 2 37 2 2 --
  +-------- Lasmigona_compressa     
--+                                 
  |  +----- Pyganodon_grandis       
  +--+                              
     |  +-- Utterbackia_peninsularis **
     +--+                           
        +-- Utterbackia_imbecillis **  

Froufe et al., 2019, fig. 3

iT tT bi riS oT roT roS iD
2 2 0 2 43 2 2 --
  +-------- Lasmigona_complanata    
--+                                 
  |  +----- Pyganodon_grandis       
  +--+                              
     |  +-- Utterbackia_peninsularis **
     +--+                           
        +-- Utterbackia_imbecillis **  

Froufe et al., 2019, fig. 6

iT tT bi riS oT roT roS iD
2 2 0 2 42 2 2 --
  +-------- Lasmigona_complanata    
--+                                 
  |  +----- Pyganodon_grandis       
  +--+                              
     |  +-- Utterbackia_peninsularis **
     +--+                           
        +-- Utterbackia_imbecillis **  

Riccardi et al., 2019, fig. 2

iT tT bi riS oT roT roS iD
2 2 0 2 18 2 2 --
  +-------- Lasmigona_compressa     
--+                                 
  |  +----- Pyganodon_grandis       
  +--+                              
     |  +-- Utterbackia_peninsularis **
     +--+                           
        +-- Utterbackia_imbecillis **  

Wu et al., 2020, fig. 3

iT tT bi riS oT roT roS iD
2 2 0 2 48 2 2 --
  +-------- Lasmigona_compressa     
--+                                 
  |  +----- Pyganodon_grandis       
  +--+                              
     |  +-- Utterbackia_peninsularis **
     +--+                           
        +-- Utterbackia_imbecillis **  

Sano et al., 2020, fig. 2

iT tT bi riS oT roT roS iD
2 2 0 2 73 4 3 --
     +----- Alasmidonta_varicosa    
  +--+                              
  |  |  +-- Alasmidonta_heterodon   
  |  +--+                           
--+     +-- Lasmigona_compressa     
  |                                 
  |  +----- Pyganodon_grandis       
  +--+                              
     |  +-- Utterbackia_imbecillis **  
     +--+                           
        +-- Utterbackia_peninsularis **

Sano et al., 2020, fig. 3

iT tT bi riS oT roT roS iD
2 2 0 2 73 4 3 --
     +----- Pyganodon_grandis       
  +--+                              
  |  |  +-- Utterbackia_imbecillis **  
  |  +--+                           
--+     +-- Utterbackia_peninsularis **
  |                                 
  |  +----- Alasmidonta_varicosa    
  +--+                              
     |  +-- Lasmigona_compressa     
     +--+                           
        +-- Alasmidonta_heterodon   

Sano et al., 2020, fig. 4

iT tT bi riS oT roT roS iD
2 2 0 2 73 4 3 --
     +----- Alasmidonta_varicosa    
  +--+                              
  |  |  +-- Alasmidonta_heterodon   
  |  +--+                           
--+     +-- Lasmigona_compressa     
  |                                 
  |  +----- Pyganodon_grandis       
  +--+                              
     |  +-- Utterbackia_imbecillis **  
     +--+                           
        +-- Utterbackia_peninsularis **

Wu et al., 2018, fig. 2

iT tT bi riS oT roT roS iD
2 2 0 2 20 1 1 --
  +----- Pyganodon_grandis       
--+                              
  |  +-- Utterbackia_peninsularis **
  +--+                           
     +-- Utterbackia_imbecillis **  

Wu et al., 2020, fig. 3

iT tT bi riS oT roT roS iD
2 2 0 2 28 1 1 --
  +----- Pyganodon_grandis       
--+                              
  |  +-- Utterbackia_peninsularis **
  +--+                           
     +-- Utterbackia_imbecillis **  

Guerra et al., 2019, fig. 1

iT tT bi riS oT roT roS iD
2 2 0 2 39 3 2 --
        +-- Lasmigona_compressa     
  +-----+                           
  |     +-- Lasmigona_subviridis    
--+                                 
  |  +----- Pyganodon_grandis       
  +--+                              
     |  +-- Utterbackia_peninsularis **
     +--+                           
        +-- Utterbackia_imbecillis **  

Guerra et al., 2019, fig. 2

iT tT bi riS oT roT roS iD
2 2 0 2 39 3 2 --
        +-- Lasmigona_compressa     
  +-----+                           
  |     +-- Lasmigona_subviridis    
--+                                 
  |  +----- Pyganodon_grandis       
  +--+                              
     |  +-- Utterbackia_peninsularis **
     +--+                           
        +-- Utterbackia_imbecillis **  

Chen et al., 2020, fig. 1

iT tT bi riS oT roT roS iD
2 27 0 2 32 4 3 26.0
                 +-------- Schistodesmus_lampreyanus
           +-----+                                  
           |     |  +----- Lepidodesma_languilati   
           |     +--+                               
           |        |  +-- Aculamprotula_coreana    
           |        +--+                            
  +--------+           +-- Aculamprotula_tortuosa   
  |        |                                        
  |        |        +----- Lasmigona_compressa      
  |        |  +-----+                               
  |        |  |     |  +-- Utterbackia_imbecillis **   
  |        |  |     +--+                            
  |        +--+        +-- Pyganodon_grandis        
  |           |                                     
  |           |  +-------- Sinanodonta_lucida       
  |           +--+                                  
  |              |  +----- Cristaria_plicata        
  |              +--+                               
--+                 |  +-- Anodonta_cygnea          
  |                 +--+                            
  |                    +-- Anodonta_anatina         
  |                                                 
  |     +----------------- Gibbosula_crassa         
  |  +--+                                           
  |  |  |  +-------------- Quadrula_quadrula        
  |  |  +--+                                        
  |  |     |  +----------- Toxolasma_parvum         
  |  |     +--+                                     
  |  |        |  +-------- Lampsilis_ornata         
  |  |        +--+                                  
  |  |           |  +----- Potamilus_leptodon       
  +--+           +--+                               
     |              |  +-- Lampsilis_siliquoidea    
     |              +--+                            
     |                 +-- Lampsilis_powellii       
     |                                              
     |                 +-- Utterbackia_peninsularis ** 
     |  +--------------+                            
     |  |              +-- Potamilus_alatus         
     +--+                                           
        |  +-------------- Lamprotula_leaii         
        |  |                                        
        +--+           +-- Sinohyriopsis_cumingii   
           |  +--------+                            
           |  |        +-- Sinohyriopsis_schlegelii 
           +--+                                     
              |  +-------- Parvasolenaia_rivularis  
              +--+                                  
                 |  +----- Sinosolenaia_carinata    
                 +--+                               
                    |  +-- Sinosolenaia_iridinea    
                    +--+                            
                       +-- Sinosolenaia_oleivora    

Zieritz et al., 2021, fig. 3-F, 5

iT tT bi riS oT roT roS iD
2 2 0 2 47 2 2 --
  +-------- Lasmigona_compressa     
--+                                 
  |  +----- Pyganodon_grandis       
  +--+                              
     |  +-- Utterbackia_imbecillis **  
     +--+                           
        +-- Utterbackia_peninsularis **

Wu et al., 2022, fig. 2d

iT tT bi riS oT roT roS iD
2 2 0 2 37 2 2 --
  +-------- Lasmigona_compressa     
--+                                 
  |  +----- Pyganodon_grandis       
  +--+                              
     |  +-- Utterbackia_imbecillis **  
     +--+                           
        +-- Utterbackia_peninsularis **

Wu et al., 2022, fig. 3

iT tT bi riS oT roT roS iD
2 2 0 2 72 2 2 --
  +-------- Lasmigona_compressa     
--+                                 
  |  +----- Pyganodon_grandis       
  +--+                              
     |  +-- Utterbackia_imbecillis **  
     +--+                           
        +-- Utterbackia_peninsularis **

Lydeard et al., 1996, fig. 3

iT tT bi riS oT roT roS iD
1 1 0 1 26 5 4 --
     +----- Utterbackia_imbecillis **  
  +--+                              
  |  |  +-- Utterbackiana_couperiana
  |  +--+                           
--+     +-- Utterbackiana           
  |                                 
  |  +----- Strophitus_radiatus     
  +--+                              
     |  +-- Lasmigona_costata       
     +--+                           
        +-- Alasmidonta_triangulata 

Lydeard et al., 1996, fig. 4

iT tT bi riS oT roT roS iD
1 1 0 1 27 4 4 --
No taxon cladogram to show.

Huang et al., 2002, fig. 1

iT tT bi riS oT roT roS iD
1 1 0 1 33 4 4 --
  +----------- Utterbackiana_couperiana
--+                                    
  |  +-------- Utterbackia_imbecillis **  
  +--+                                 
     |  +----- Strophitus_radiatus     
     +--+                              
        |  +-- Alasmidonta_triangulata 
        +--+                           
           +-- Lasmigona_costata       

Huang et al., 2002, fig. 2

iT tT bi riS oT roT roS iD
1 1 0 1 33 4 4 --
  +----- Utterbackia_imbecillis **  
  |                              
--+----- Utterbackiana_couperiana
  |                              
  |  +-- Alasmidonta_triangulata 
  +--+                           
     +-- Lasmigona_costata       

Krebs et al., 2003, fig. 3

iT tT bi riS oT roT roS iD
1 1 0 1 33 8 5 --
No taxon cladogram to show.

Boyer et al., 2011, fig. 4

iT tT bi riS oT roT roS iD
1 1 0 1 37 8 6 --
        +----- Lasmigona_complanata      
  +-----+                                
  |     |  +-- Lasmigona_costata         
  |     +--+                             
--+        +-- Arcidens_confragosus      
  |                                      
  |  +-------- Utterbackia_imbecillis **    
  |  |                                   
  +--+-------- Alasmidonta_marginata     
     |                                   
     |  +----- Pyganodon_grandis         
     +--+                                
        |  +-- Anodontoides_ferussacianus
        +--+                             
           +-- Lasmigona_compressa       

Rosenberg et al., 1994, fig. 3

iT tT bi riS oT roT roS iD
1 1 0 1 18 2 1 --
No taxon cladogram to show.

Rosenberg et al., 1997, fig. 2A

iT tT bi riS oT roT roS iD
1 1 0 1 22 2 1 --
No taxon cladogram to show.

Rosenberg et al., 1997, fig. 2B

iT tT bi riS oT roT roS iD
1 1 0 1 22 2 1 --
No taxon cladogram to show.

Campbell, 2000, fig. 1-5

iT tT bi riS oT roT roS iD
1 1 0 1 2 0 0 --
No taxon cladogram to show.

Mulvey et al., 1997, fig. 1, 3

iT tT bi riS oT roT roS iD
1 1 0 1 4 0 0 --
No taxon cladogram to show.

Huang et al., 2013, fig. 3-M

iT tT bi riS oT roT roS iD
1 1 0 1 5 1 1 --
  +-- Pyganodon_grandis       
--+                           
  +-- Utterbackia_peninsularis **

Froufe et al., 2015, fig. 1

iT tT bi riS oT roT roS iD
1 1 0 1 11 1 1 --
  +-- Utterbackia_peninsularis **
--+                           
  +-- Pyganodon_grandis       

Froufe et al., 2015, fig. 1-M

iT tT bi riS oT roT roS iD
1 1 0 1 7 1 1 --
  +-- Utterbackia_peninsularis **
--+                           
  +-- Pyganodon_grandis       

Soroka & Burzynski, 2018, fig. 3

iT tT bi riS oT roT roS iD
1 1 0 1 6 1 1 --
  +-- Pyganodon_grandis       
--+                           
  +-- Utterbackia_peninsularis **

Soroka & Burzynski, 2018, fig. 3-M

iT tT bi riS oT roT roS iD
1 1 0 1 6 1 1 --
  +-- Pyganodon_grandis       
--+                           
  +-- Utterbackia_peninsularis **

Guerra et al., 2017, fig. 2a-M

iT tT bi riS oT roT roS iD
1 1 0 1 7 1 1 --
  +-- Utterbackia_peninsularis **
--+                           
  +-- Pyganodon_grandis       

Guerra et al., 2017, fig. 2b-M

iT tT bi riS oT roT roS iD
1 1 0 1 7 1 1 --
  +-- Utterbackia_peninsularis **
--+                           
  +-- Pyganodon_grandis       

Burzynski et al., 2017, fig. 1 bottom

iT tT bi riS oT roT roS iD
1 1 0 1 12 1 1 --
  +-- Utterbackia_peninsularis **
--+                           
  +-- Pyganodon_grandis       

Burzynski et al., 2017, fig. 1 top

iT tT bi riS oT roT roS iD
1 1 0 1 12 1 1 --
  +-- Utterbackia_peninsularis **
--+                           
  +-- Pyganodon_grandis       

Fonseca et al., 2016, fig. 1-M

iT tT bi riS oT roT roS iD
1 1 0 1 8 1 1 --
  +-- Utterbackia_peninsularis **
--+                           
  +-- Pyganodon_grandis       

Amaro et al., 2017, fig. 2-M

iT tT bi riS oT roT roS iD
1 1 0 1 9 3 2 --
  +----- Pyganodon_grandis[2]    
  |                              
--+  +-- Lasmigona_costata       
  |  |                           
  +--+-- Pyganodon_grandis[1]    
     |                           
     +-- Utterbackia_peninsularis **

Amaro et al., 2017, fig. 3A-M

iT tT bi riS oT roT roS iD
1 1 0 1 6 1 1 --
  +-- Pyganodon_grandis       
--+                           
  +-- Utterbackia_peninsularis **

Amaro et al., 2017, fig. 3B-M

iT tT bi riS oT roT roS iD
1 1 0 1 6 1 1 --
  +-- Utterbackia_peninsularis **
--+                           
  +-- Pyganodon_grandis       

Lopes-Lima et al., 2017, fig. 2-M

iT tT bi riS oT roT roS iD
1 1 0 1 9 1 1 --
  +-- Utterbackia_peninsularis **
--+                           
  +-- Pyganodon_grandis       

Wen et al., 2017, fig. 3-M

iT tT bi riS oT roT roS iD
1 1 0 1 5 1 1 --
  +-- Utterbackia_peninsularis **
--+                           
  +-- Pyganodon_grandis       

Feng et al., 2016, fig. 2

iT tT bi riS oT roT roS iD
1 1 0 1 14 1 1 --
No taxon cladogram to show.

Huang et al., 2018, fig. 6-M

iT tT bi riS oT roT roS iD
1 1 0 1 16 1 1 --
  +-- Pyganodon_grandis       
--+                           
  +-- Utterbackia_peninsularis **

Huang et al., 2019, fig. 3-M

iT tT bi riS oT roT roS iD
1 1 0 1 22 1 1 --
  +-- Pyganodon_grandis       
--+                           
  +-- Utterbackia_peninsularis **

Chase et al., 2018, fig. 5

iT tT bi riS oT roT roS iD
1 1 0 1 9 0 0 --
No taxon cladogram to show.

Chase et al., 2018, fig. 1

iT tT bi riS oT roT roS iD
1 1 0 1 10 1 1 --
  +-- Utterbackia_peninsularis **
--+                           
  +-- Pyganodon_grandis       

Chase et al., 2018, fig. 1-M

iT tT bi riS oT roT roS iD
1 1 0 1 10 1 1 --
  +-- Utterbackia_peninsularis **
--+                           
  +-- Pyganodon_grandis       

Burzynski & Soroka, 2018, fig. 3

iT tT bi riS oT roT roS iD
1 1 0 1 3 1 1 --
  +-- Utterbackia_peninsularis **
--+                           
  +-- Pyganodon_grandis       

Burzynski & Soroka, 2018, fig. 5a

iT tT bi riS oT roT roS iD
1 1 0 1 19 1 1 --
  +-- Utterbackia_peninsularis **
--+                           
  +-- Pyganodon_grandis       

Burzynski & Soroka, 2018, fig. 5b

iT tT bi riS oT roT roS iD
1 1 0 1 9 1 1 --
  +-- Utterbackia_peninsularis **
--+                           
  +-- Pyganodon_grandis       

Froufe et al., 2019, fig. 3-M

iT tT bi riS oT roT roS iD
1 1 0 1 18 1 1 --
  +-- Utterbackia_peninsularis **
--+                           
  +-- Pyganodon_grandis       

Froufe et al., 2019, fig. 4

iT tT bi riS oT roT roS iD
1 1 0 1 18 1 1 --
  +-- Utterbackia_peninsularis **
--+                           
  +-- Pyganodon_grandis       

Fukata & Iigo, 2020, fig. 1

iT tT bi riS oT roT roS iD
1 1 0 1 53 3 2 --
     +-- Pyganodon_grandis       
  +--+                           
  |  +-- Utterbackia_peninsularis **
--+                              
  |  +-- Alasmidonta_varicosa    
  +--+                           
     +-- Alasmidonta_heterodon   

Guerra et al., 2019, fig. 1-M

iT tT bi riS oT roT roS iD
1 1 0 1 21 1 1 --
  +-- Utterbackia_peninsularis **
--+                           
  +-- Pyganodon_grandis       

Guerra et al., 2019, fig. 2-M

iT tT bi riS oT roT roS iD
1 1 0 1 21 1 1 --
  +-- Utterbackia_peninsularis **
--+                           
  +-- Pyganodon_grandis       

Zieritz et al., 2021, fig. 3-M, 4

iT tT bi riS oT roT roS iD
1 1 0 1 22 1 1 --
  +-- Utterbackia_peninsularis **
--+                           
  +-- Pyganodon_grandis       

Wu et al., 2022, fig. 2e

iT tT bi riS oT roT roS iD
1 1 0 1 16 1 1 --
  +-- Utterbackia_peninsularis **
--+                           
  +-- Pyganodon_grandis       

Page updated: 10:57:26 Sun 06 Nov 2022

 
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