Unionoidea | Unionidae | Ambleminae | Quadrulini | Quadrulina | Pustulosa species Pustulosa pustulosa (Lea, 1831): specimens 31 nominal species as synonyms
Taxonomic History (since 2007)
Quadrula pustulosa — Tiemann et al. (2007), Williams et al. (2008), Bogan (2010), Boyer et al. (2011), ..., Marshall et al. (2018), Inoue et al. (2018), Gasparini et al. (2020), Klymus et al. (2020)
Amphinaias pustulosa — Graf & Cummings (2007), Cummings & Graf (2010), IUCN (2011)
Amphinaias aurea — Graf & Cummings (2007), Cummings & Graf (2010)
Amphinaias houstonensis — Graf & Cummings (2007), Cummings & Graf (2010)
Amphinaias refulgens — Graf & Cummings (2007), Cummings & Graf (2010)
Quadrula asperata — Williams et al. (2008)
Quadrula refulgens — Chapman et al. (2008), Williams et al. (2008), Chapman et al. (2008), Bogan (2010), Haag (2012), ITIS (2012), Szumowski et al. (2012), Inoue et al. (2018)
Quadrula pustulosa pustulosa — Watters et al. (2009), ITIS (2012)
Quadrula mortoni — Bogan (2010), Pfeiffer et al. (2016), Marshall et al. (2018)
Quadrula aurea — Bogan (2010), ITIS (2012), Haag (2012), Szumowski et al. (2012)
Quadrula houstonensis — Bogan (2010), Breton et al. (2011), Haag (2012), ITIS (2012), Pfeiffer et al. (2016), Chase et al. (2018)
Rotundaria pustulosa — Campbell & Lydeard (2012)
Quadrula pustulosa mortoni — Szumowski et al. (2012), ITIS (2012)
Rotundaria refulgens — Campbell & Lydeard (2012), Campbell & Lydeard (2012)
Cyclonaias pustulosa — Williams et al. (2017), Johnson et al. (2018), Lopes-Lima et al. (2019), Graf & Cummings (2021), FMCS (2021), Bieler (2021), Hopper et al. (2023)
Cyclonaias mortoni — Williams et al. (2017)
Cyclonaias aurea — Williams et al. (2017)
Cyclonaias houstonensis — Williams et al. (2017)
Cyclonaias refulgens — Williams et al. (2017)
Cyclonaias (Quadrula) refulgens — Porto-Hannes et al. (2021)
Pustulosa pustulosa — Neemuchwala et al. (2023)
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Range: Widespread in the Interior Basin and the Gulf Coastal Plain (west of the Mobile Basin), USA. Map of Georeferenced Specimens
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USA |
ANSP_10285 Jackson, La
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ANSP_10287A Bayou Teche, Louisiana
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ANSP_20250 Kentucky River
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ANSP_416329 Sangamon River, Lake of the Woods, Mahomet, Champaign Co., Illinois 40.2040° N, 88.3876° W
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USNM_84572 Texas
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USNM_84204 Cahaba R., Ala.
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USNM_84215 Wabash River, Ind.
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ANSP_20220 Green River
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USNM_84271 Pearl River, Jackson, Miss
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USNM_85763 Columbus, Miss.
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USNM_85768 Rutersville, Texas
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USNM_84275 Alabama R., Ala.
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USNM_84277 near New Orleans, La.
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USNM_84290 Enterprise, Lauderdale Co., Miss. [Alabama]
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USNM_84298 N.C.
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Genbank Vouchers
USAUF_438172 (QaurGua045) (COI, ND1, ITS1, 5.8S) USA, Texas, Dewitt County, Guadalupe River about 15 air km upstream of Cuero, TX (29.221294, -97.329695) UF_439126 (QpusOsa065) (COI, ND1, ITS1, 5.8S) USA, Missouri, Hickory, Pomme de Terre River approx. 500 , below MDC Cross Timber Access Ramp, 2.8 mi N or Hermitage (37.9815, -93.3081) UF_439157 (QpusOhi052) (COI, ND1, ITS1, 5.8S) USA, Ohio, Wood County, Ohio River; Buckley Island backchannel (39.3912312, -81.4209442) UF_438919 (QmorNec024) (COI, ND1, ITS1, 5.8S) USA, Texas, Anderson/Cherokee County, Neches River downstream of SH 79 (31.885207, -95.431684) UF_438168 (QhouBra037) (COI, ND1, ITS1, 5.8S) USA, Texas, Brazos/Burleson County, Brazos River about 15.5 km downstream from Hwy 21 (30.5811176, -96.4415058) UF_438919 (QmorNec028) (COI, ND1, ITS1, 5.8S) USA, Texas, Anderson/Cherokee County, Neches River downstream of SH 79 (31.885207, -95.431684) UF_438785 (QpusRed005) (COI, ND1, ITS1, 5.8S) USA, Texas, Lamar County, Pine Creek, ~100 m upstream from CR906 (33.854656, -95.341305) UF_438486 (QmorTri018) (COI, ND1, ITS1, 5.8S) USA, Texas, Newton County, Sabine River, 2 mi. NNW of Bancroft, LA (30.591914, -93.698572) UF_441087 (QhouCol004) (COI, ND1, ITS1, 5.8S) USA, Texas, San Saba County, San Saba River at CR 104, San Saba River Golf Course, near San Saba, TX (31.20738, -98.68761) UF_438011 (QmorTri014) (COI, ND1, ITS1, 5.8S) USA, Texas, Anderson/Freestone County, Trinity River just upstream of Hwy 79, 3.0 air mi SW of Tucker, TX (31.64941, -95.7897) UF_438741 (QaurNue086) (COI, ND1, ITS1, 5.8S) USA, Texas, Live Oak County, Nueces River upstream of Airport Rd (28.382964, -98.109617) UF_438785 (QpusRed001) (COI, ND1, ITS1, 5.8S) USA, Texas, Lamar County, Pine Creek, ~100 m upstream from CR906 (33.854656, -95.341305) UF_438785 (QpusRed002) (COI, ND1, ITS1, 5.8S) USA, Texas, Lamar County, Pine Creek, ~100 m upstream from CR906 (33.854656, -95.341305) UF_438574 (QaurNue065) (COI, ND1, ITS1, 5.8S) USA, Texas, Live Oak County, Nueces River, 400 m upstream of Hwy 59 (28.337458, -98.086636) UF_438053 (QaurNue027) (COI, ND1, ITS1, 5.8S) USA, Texas, Live Oak County, Frio River below Choke Canyon Reservoir, Tips State Rec. Area below the fall in Three Rivers (28.46104, -98.19073) UF_439127 (QpusNeo063) (COI, ND1, ITS1, 5.8S) USA, Missouri, McDonald County, Elk River, along Highway DD near Noel (36.568, -94.504) UF_438930 (QmorTri043) (COI, ND1, ITS1, 5.8S) USA, Texas, Tarrant County, Grapevine Lake, [3.4 mi. NNW of Grapevine, TX] (32.980521, -97.09841) UF_438923 (QmorNec033) (COI, ND1, ITS1, 5.8S) USA, Texas, Anderson/Cherokee County, Neches River @ CR335 (31.945598, -95.444602) UF_438919 (QmorNec026) (COI, ND1, ITS1, 5.8S) USA, Texas, Anderson/Cherokee County, Neches River downstream of SH 79 (31.885207, -95.431684) UF_439126 (QpusOsa068) (COI, ND1, ITS1, 5.8S) USA, Missouri, Hickory, Pomme de Terre River approx. 500 , below MDC Cross Timber Access Ramp, 2.8 mi N or Hermitage (37.9815, -93.3081) UF_439080 (QmorSab068) (COI, ND1, ITS1, 5.8S) USA, Texas, Wood County, Lake Fork Creek 6.7 km south of Hainesville, TX (32.65716, -95.3615) UF_438172 (QaurGua042) (COI, ND1, ITS1, 5.8S) USA, Texas, Dewitt County, Guadalupe River about 15 air km upstream of Cuero, TX (29.221294, -97.329695) UF_439081 (QrefPrl008) (COI, ND1, ITS1, 5.8S) USA, Louisiana, St. Tammany Parish, Bogue Chitto River near Lock 2 (30.62254, -89.87655) UF_441085 (QaurGua016) (COI, ND1, ITS1, 5.8S) USA, Texas, Gonzales/Guadalupe County, San Marcos River between US-90 and SR-80, south of Luling (29.67078, -97.69561) UF_438930 (QmorTri047) (COI, ND1, ITS1, 5.8S) USA, Texas, Tarrant County, Grapevine Lake, [3.4 mi. NNW of Grapevine, TX] (32.980521, -97.09841) UF_438574 (QaurNue062) (COI, ND1, ITS1, 5.8S) USA, Texas, Live Oak County, Nueces River, 400 m upstream of Hwy 59 (28.337458, -98.086636) UF_439076 (QmorTri057) (COI, ND1, ITS1, 5.8S) USA, Texas, Anderson County, Upstream of Hwy 82 near Palestina, TX (31.683591, -95.792124) UF_438974 (QmorRed051) (COI, ND1, ITS1, 5.8S) USA, Texas, Marion County, Big Cypress Bayou (just downstream from Hwy 59, near Jefferson, TX) (32.739928, -94.441666) UF_441135 (QhouBra011) (COI, ND1, ITS1, 5.8S) USA, Texas, Robertson County, Brazos River at FM 485, west of Highway 6 (30.8676502, -96.6956368) UF_439081 (QrefPrl006) (COI, ND1, ITS1, 5.8S) USA, Louisiana, St. Tammany Parish, Bogue Chitto River near Lock 2 (30.62254, -89.87655) UF_438172 (QaurGua044) (COI, ND1, ITS1, 5.8S) USA, Texas, Dewitt County, Guadalupe River about 15 air km upstream of Cuero, TX (29.221294, -97.329695) UF_439157 (QpusOhi051) (COI, ND1, ITS1, 5.8S) USA, Ohio, Wood County, Ohio River; Buckley Island backchannel (39.3912312, -81.4209442) UF_441087 (QhouCol005) (COI, ND1, ITS1, 5.8S) USA, Texas, San Saba County, San Saba River at CR 104, San Saba River Golf Course, near San Saba, TX (31.20738, -98.68761) UF_439080 (QmorSab066) (COI, ND1, ITS1, 5.8S) USA, Texas, Wood County, Lake Fork Creek 6.7 km south of Hainesville, TX (32.65716, -95.3615) UF_441171 (QmorSab001) (COI, ND1, ITS1, 5.8S) USA, Texas, Newton County, Sabine River, near river mile 82.5, Site A (30.6652604, -93.6580554) UF_439059 (QhouBra047) (COI, ND1, ITS1, 5.8S) USA, Texas, Milam County, Little River downstream of CR1915 [4 mi. S of Buckholts, TX] (30.81558, -97.13651) UF_438923 (QmorNec031) (COI, ND1, ITS1, 5.8S) USA, Texas, Anderson/Cherokee County, Neches River @ CR335 (31.945598, -95.444602) UF_439076 (QmorTri058) (COI, ND1, ITS1, 5.8S) USA, Texas, Anderson County, Upstream of Hwy 82 near Palestina, TX (31.683591, -95.792124) UF_438157 (QhouBra041) (COI, ND1, ITS1, 5.8S) USA, Texas, Williamson County, San Gabriel River about 100m downstream of Co. Road 428 bridge (30.695341, -97.27804) UF_438785 (QpusRed004) (COI, ND1, ITS1, 5.8S) USA, Texas, Lamar County, Pine Creek, ~100 m upstream from CR906 (33.854656, -95.341305) UF_438923 (QmorNec040) (COI, ND1, ITS1, 5.8S) USA, Texas, Anderson/Cherokee County, Neches River @ CR335 (31.945598, -95.444602) UF_439059 (QhouBra044) (COI, ND1, ITS1, 5.8S) USA, Texas, Milam County, Little River downstream of CR1915 [4 mi. S of Buckholts, TX] (30.81558, -97.13651) UF_438566 (QaurNue051) (COI, ND1, ITS1, 5.8S) USA, Texas, Live Oak County, Frio River about 400 m downstream of dam at Choke Canyon (28.48684, -98.23867) UF_439079 (QmorCol063) (COI, ND1, ITS1, 5.8S) USA, Texas, Wharton, Colorado River near Lane City, TX; Habitat: Run (29.173116, -96.067204) UF_438168 (QhouBra036) (COI, ND1, ITS1, 5.8S) USA, Texas, Brazos/Burleson County, Brazos River about 15.5 km downstream from Hwy 21 (30.5811176, -96.4415058) UF_439076 (QmorTri056) (COI, ND1, ITS1, 5.8S) USA, Texas, Anderson County, Upstream of Hwy 82 near Palestina, TX (31.683591, -95.792124) UF_439157 (QpusOhi056) (COI, ND1, ITS1, 5.8S) USA, Ohio, Wood County, Ohio River; Buckley Island backchannel (39.3912312, -81.4209442) UF_441173 (QmorSab005) (COI, ND1, ITS1, 5.8S) USA, Texas, Newton County, Lower Sabine River, Site 54A, 4.2 miles north of Deweyville, TX (30.3565919, -93.7606991) UF_439079 (QmorCol062) (COI, ND1, ITS1, 5.8S) USA, Texas, Wharton, Colorado River near Lane City, TX; Habitat: Run (29.173116, -96.067204) UF_440981 (QaurGua009) (COI, ND1, ITS1, 5.8S) USA, Texas, Guadalupe County, Guadalupe River at Hwy 77 crossing (28.83119, -97.06129) UF_441087 (QhouCol007) (COI, ND1, ITS1, 5.8S) USA, Texas, San Saba County, San Saba River at CR 104, San Saba River Golf Course, near San Saba, TX (31.20738, -98.68761) UF_438574 (QaurNue071) (COI, ND1, ITS1, 5.8S) USA, Texas, Live Oak County, Nueces River, 400 m upstream of Hwy 59 (28.337458, -98.086636) UF_438919 (QmorNec029) (COI, ND1, ITS1, 5.8S) USA, Texas, Anderson/Cherokee County, Neches River downstream of SH 79 (31.885207, -95.431684) UF_438172 (QaurGua043) (COI, ND1, ITS1, 5.8S) USA, Texas, Dewitt County, Guadalupe River about 15 air km upstream of Cuero, TX (29.221294, -97.329695) UF_439127 (QpusNeo061) (COI, ND1, ITS1, 5.8S) USA, Missouri, McDonald County, Elk River, along Highway DD near Noel (36.568, -94.504) UF_441135 (QhouBra014) (COI, ND1, ITS1, 5.8S) USA, Texas, Robertson County, Brazos River at FM 485, west of Highway 6 (30.8676502, -96.6956368) UF_438886 (QrefPas005) (COI, ND1, ITS1, 5.8S) USA, Mississippi, Jackson County, Pascagoula River at Paper Mill Camp slough about 2 mi. upstream of MS Hwy 614 (30.63228, -88.652401) UF_438923 (QmorNec034) (COI, ND1, ITS1, 5.8S) USA, Texas, Anderson/Cherokee County, Neches River @ CR335 (31.945598, -95.444602) UF_441257 (QaurGua021) (COI, ND1, ITS1, 5.8S) USA, Texas, Goliad County, San Antonio River 3.3 mi south of Goliad, TX, just NW of US Hwy 59 bridge; from SW shore in muddy eddy area above large drift against bridge pilings, in fairly deep, soft mud on clay substrate, ~0.5m deep (28.651169, -97.432994) UF_438886 (QrefPas004) (COI, ND1, ITS1, 5.8S) USA, Mississippi, Jackson County, Pascagoula River at Paper Mill Camp slough about 2 mi. upstream of MS Hwy 614 (30.63228, -88.652401) UF_438886 (QrefPas001) (COI, ND1, ITS1, 5.8S) USA, Mississippi, Jackson County, Pascagoula River at Paper Mill Camp slough about 2 mi. upstream of MS Hwy 614 (30.63228, -88.652401) UF_440989 (QhouCol001) (COI, ND1, ITS1, 5.8S) USA, Texas, Colorado County, Colorado River at FM 950; just east of Garwood, TX. Sampled ~30m stretch along right ascending bank nearl old bridge abutments. (29.45354, -96.39587) UF_441135 (QhouBra012) (COI, ND1, ITS1, 5.8S) USA, Texas, Robertson County, Brazos River at FM 485, west of Highway 6 (30.8676502, -96.6956368) UF_439201 (QmorNec089) (COI, ND1, ITS1, 5.8S) USA, Texas, Trinity County, Alabama Creek WMA (31.19669, -94.85869) UF_441087 (QhouCol006) (COI, ND1, ITS1, 5.8S) USA, Texas, San Saba County, San Saba River at CR 104, San Saba River Golf Course, near San Saba, TX (31.20738, -98.68761) UF_441172 (QmorSab004) (COI, ND1, ITS1, 5.8S) USA, Texas, Kaufman County, East Fork Trinity River downstream of Lewis Street, near Crandall, TX [32.592543, -96.486377] UF_439076 (QmorTri059) (COI, ND1, ITS1, 5.8S) USA, Texas, Anderson County, Upstream of Hwy 82 near Palestina, TX (31.683591, -95.792124) UF_438785 (QpusRed006) (COI, ND1, ITS1, 5.8S) USA, Texas, Lamar County, Pine Creek, ~100 m upstream from CR906 (33.854656, -95.341305) UF_440981 (QaurGua008) (COI, ND1, ITS1, 5.8S) USA, Texas, Guadalupe County, Guadalupe River at Hwy 77 crossing (28.83119, -97.06129) UF_438172 (QaurGua048) (COI, ND1, ITS1, 5.8S) USA, Texas, Dewitt County, Guadalupe River about 15 air km upstream of Cuero, TX (29.221294, -97.329695) UF_438923 (QmorNec036) (COI, ND1, ITS1, 5.8S) USA, Texas, Anderson/Cherokee County, Neches River @ CR335 (31.945598, -95.444602) UF_441256 (QmorSab008) (COI, ND1, ITS1, 5.8S) USA, Texas, Smith/Wood County, Sabine River, Hwy 14, S of Hawkins, TX, collected from a riffle 200m downstream from bridge crossing (32.5573098, -95.2056386) UF_438741 (QaurNue083) (COI, ND1, ITS1, 5.8S) USA, Texas, Live Oak County, Nueces River upstream of Airport Rd (28.382964, -98.109617) UF_438785 (QpusRed010) (COI, ND1, ITS1, 5.8S) USA, Texas, Lamar County, Pine Creek, ~100 m upstream from CR906 (33.854656, -95.341305) UF_441171 (COI, ITS1, 5.8S) USA:Texas:Newton County :Sabine River, near river mile 82.5, Site A (30.6652604, -93.6580554) UF_441135 (COI, ITS1, 5.8S) USA, Texas, Robertson County, Brazos River at FM 485, west of Highway 6 (30.8676502, -96.6956368) UF_439184 (QpusStC081) (ND1, ITS1, 5.8S) USA, Minnesota, Chisago County, St. Croix River at Interstate Park (45.3955278, -92.6627834) UF_439184 (QpusStC082) (ND1, ITS1, 5.8S) USA, Minnesota, Chisago County, St. Croix River at Interstate Park (45.3955278, -92.6627834) UF_438923 (QmorNec032) (COI, ND1) USA, Texas, Anderson/Cherokee County, Neches River @ CR335 (31.945598, -95.444602) UF_441139 (QhouBra017) (COI, ND1) USA, Texas, Comanche County, Leon River downstream from Highway 67 (31.9574036, -98.4589713) UF_438919 (QmorNec023) (COI, ND1) USA, Texas, Anderson/Cherokee County, Neches River downstream of SH 79 (31.885207, -95.431684) UF_438172 (QaurGua040) (COI, ND1) USA, Texas, Dewitt County, Guadalupe River about 15 air km upstream of Cuero, TX (29.221294, -97.329695) UF_439079 (QmorCol061) (COI, ND1) USA, Texas, Wharton, Colorado River near Lane City, TX; Habitat: Run (29.173116, -96.067204) UF_438574 (QaurNue074) (COI, ND1) USA, Texas, Live Oak County, Nueces River, 400 m upstream of Hwy 59 (28.337458, -98.086636) UF_438927 (QaurGua091) (COI, ND1) USA, Texas, Gonzales County, San Marcos River @ CR232 (29.56274, -97.543455) UF_438919 (QmorNec022) (COI, ND1) USA, Texas, Anderson/Cherokee County, Neches River downstream of SH 79 (31.885207, -95.431684) UF_438574 (QaurNue079) (COI, ND1) USA, Texas, Live Oak County, Nueces River, 400 m upstream of Hwy 59 (28.337458, -98.086636) UF_438574 (QaurNue069) (COI, ND1) USA, Texas, Live Oak County, Nueces River, 400 m upstream of Hwy 59 (28.337458, -98.086636) UF_441087 (QhouCol009) (COI, ND1) USA, Texas, San Saba County, San Saba River at CR 104, San Saba River Golf Course, near San Saba, TX (31.20738, -98.68761) UF_438486 (QmorTri017) (COI, ND1) USA, Texas, Newton County, Sabine River, 2 mi. NNW of Bancroft, LA (30.591914, -93.698572) UF_439195 (QmorSaJ070) (COI, ND1) USA, Texas, Montgomery County, West Fork of San Jacinto River, Site 7; Downstream of 242 crossing south of Conroe, TX (30.210398, -95.397995) UF_438741 (QaurNue085) (COI, ND1) USA, Texas, Live Oak County, Nueces River upstream of Airport Rd (28.382964, -98.109617) UF_438741 (QaurNue082) (COI, ND1) USA, Texas, Live Oak County, Nueces River upstream of Airport Rd (28.382964, -98.109617) UF_438168 (QhouBra035) (COI, ND1) USA, Texas, Brazos/Burleson County, Brazos River about 15.5 km downstream from Hwy 21 (30.5811176, -96.4415058) UF_439157 (QpusOhi054) (COI, ND1) USA, Ohio, Wood County, Ohio River; Buckley Island backchannel (39.3912312, -81.4209442) UF_438566 (QaurNue054) (COI, ND1) USA, Texas, Live Oak County, Frio River about 400 m downstream of dam at Choke Canyon (28.48684, -98.23867) UF_439201 (QmorNec087) (COI, ND1) USA, Texas, Trinity County, Alabama Creek WMA (31.19669, -94.85869) UF_438927 (QaurGua092) (COI, ND1) USA, Texas, Gonzales County, San Marcos River @ CR232 (29.56274, -97.543455) UF_441085 (QaurGua017) (COI, ND1) USA, Texas, Gonzales/Guadalupe County, San Marcos River between US-90 and SR-80, south of Luling (29.67078, -97.69561) UF_438930 (QmorTri046) (COI, ND1) USA, Texas, Tarrant County, Grapevine Lake, [3.4 mi. NNW of Grapevine, TX] (32.980521, -97.09841) UF_438785 (QpusRed008) (COI, ND1) USA, Texas, Lamar County, Pine Creek, ~100 m upstream from CR906 (33.854656, -95.341305) UF_441085 (QaurGua013) (COI, ND1) USA, Texas, Gonzales/Guadalupe County, San Marcos River between US-90 and SR-80, south of Luling (29.67078, -97.69561) UF_439126 (QpusOsa067) (COI, ND1) USA, Missouri, Hickory, Pomme de Terre River approx. 500 , below MDC Cross Timber Access Ramp, 2.8 mi N or Hermitage (37.9815, -93.3081) UF_438566 (QaurNue057) (COI, ND1) USA, Texas, Live Oak County, Frio River about 400 m downstream of dam at Choke Canyon (28.48684, -98.23867) UF_438741 (QaurNue081) (COI, ND1) USA, Texas, Live Oak County, Nueces River upstream of Airport Rd (28.382964, -98.109617) UF_439157 (QpusOhi053) (COI, ND1) USA, Ohio, Wood County, Ohio River; Buckley Island backchannel (39.3912312, -81.4209442) UF_438574 (QaurNue067) (COI, ND1) USA, Texas, Live Oak County, Nueces River, 400 m upstream of Hwy 59 (28.337458, -98.086636) UF_441174 (QmorSab006) (COI, ND1) USA, Louisiana, Vernon County, Anacoco Bayou upstream from Hwy 1211 (31.0163485, -93.3651647) UF_438919 (QmorNec019) (COI, ND1) USA, Texas, Anderson/Cherokee County, Neches River downstream of SH 79 (31.885207, -95.431684) UF_439193 (QmorSaJ069) (COI, ND1) USA, Texas, Montgomery County, Spring Creek, Site 2; North of Tumball, TX at FM 2978 crossing under bridge (30.134462, -95.59674) UF_439198 (QmorSaJ093) (COI, ND1) USA, Texas, San Jacinto County, East Fork San Jacinto River, upstream of FM 945, 300 m upstream from bridge (30.42912, -95.12973) UF_438574 (QaurNue078) (COI, ND1) USA, Texas, Live Oak County, Nueces River, 400 m upstream of Hwy 59 (28.337458, -98.086636) UF_439124 (QpusStF072) (COI, ND1) USA, Missouri, Wayne County, St. Francis River at Camp Lewallen (Bay Scout), 2.25 mi SW of Lodi (37.2371, -90.4857) UF_440981 (QaurGua006) (COI, ND1) USA, Texas, Guadalupe County, Guadalupe River at Hwy 77 crossing (28.83119, -97.06129) UF_438054 (QaurNue031) (COI, ND1) USA, Texas, Live Oak County, Nueces River @ Airport Road, George West (28.37436, -98.11561) UF_439124 (QpusStF074) (COI, ND1) USA, Missouri, Wayne County, St. Francis River at Camp Lewallen (Bay Scout), 2.25 mi SW of Lodi (37.2371, -90.4857) UF_440989 (QhouCol003) (COI, ND1) USA, Texas, Colorado County, Colorado River at FM 950; just east of Garwood, TX. Sampled ~30m stretch along right ascending bank near old bridge abutments. (29.45354, -96.39587) UF_438566 (QaurNue061) (COI, ND1) USA, Texas, Live Oak County, Frio River about 400 m downstream of dam at Choke Canyon (28.48684, -98.23867) UF_438950 (QmorTri049) (COI, ND1) USA, Texas, Denton County, Elm Fork of Trinity River, appx 0.25 km below Lake Lewisville Dam (33.065862, -96.964305) UF_439127 (QpusNeo064) (COI, ND1) USA, Missouri, McDonald County, Elk River, along Highway DD near Noel (36.568, -94.504) UF_438574 (QaurNue070) (COI, ND1) USA, Texas, Live Oak County, Nueces River, 400 m upstream of Hwy 59 (28.337458, -98.086636) UF_438930 (QmorTri044) (COI, ND1) USA, Texas, Tarrant County, Grapevine Lake, [3.4 mi. NNW of Grapevine, TX] (32.980521, -97.09841) UF_439127 (QpusNeo062) (COI, ND1) USA, Missouri, McDonald County, Elk River, along Highway DD near Noel (36.568, -94.504) UF_438172 (QaurGua047) (COI, ND1) USA, Texas, Dewitt County, Guadalupe River about 15 air km upstream of Cuero, TX (29.221294, -97.329695) UF_440968 (QaurGua001) (COI, ND1) USA, Texas, Refugio/Victoria Cos., San Antonio River at Hwy 77 crossing (28.531484, -97.042921) UF_438574 (QaurNue063) (COI, ND1) USA, Texas, Live Oak County, Nueces River, 400 m upstream of Hwy 59 (28.337458, -98.086636) UF_439201 (QmorNec088) (COI, ND1) USA, Texas, Trinity County, Alabama Creek WMA (31.19669, -94.85869) UF_438053 (QaurNue028) (COI, ND1) USA, Texas, Live Oak County, Frio River below Choke Canyon Reservoir, Tips State Rec. Area below the fall in Three Rivers (28.46104, -98.19073) UF_438974 (QmorRed052) (COI, ND1) USA, Texas, Marion County, Big Cypress Bayou (just downstream from Hwy 59, near Jefferson, TX) (32.739928, -94.441666) UF_438949 (QmorTri048) (COI, ND1) USA, Texas, Denton County, Elm Fork of Trinity River, 0.1 km below Lake Lewisville Dam (33.067064, -96.963914) UF_438923 (QmorNec041) (COI, ND1) USA, Texas, Anderson/Cherokee County, Neches River @ CR335 (31.945598, -95.444602) UF_438974 (QmorRed055) (COI, ND1) USA, Texas, Marion County, Big Cypress Bayou (just downstream from Hwy 59, near Jefferson, TX) (32.739928, -94.441666) UF_438974 (QmorRed054) (COI, ND1) USA, Texas, Marion County, Big Cypress Bayou (just downstream from Hwy 59, near Jefferson, TX) (32.739928, -94.441666) UF_439076 (QmorTri055) (COI, ND1) USA, Texas, Anderson County, Upstream of Hwy 82 near Palestina, TX (31.683591, -95.792124) UF_438566 (QaurNue058) (COI, ND1) USA, Texas, Live Oak County, Frio River about 400 m downstream of dam at Choke Canyon (28.48684, -98.23867) UF_441085 (QaurGua015) (COI, ND1) USA, Texas, Gonzales/Guadalupe County, San Marcos River between US-90 and SR-80, south of Luling (29.67078, -97.69561) UF_438927 (QaurGua096) (COI, ND1) USA, Texas, Gonzales County, San Marcos River @ CR232 (29.56274, -97.543455) UF_438886 (QrefPas002) (COI, ND1) USA, Mississippi, Jackson County, Pascagoula River at Paper Mill Camp slough about 2 mi. upstream of MS Hwy 614 (30.63228, -88.652401) UF_439205 (QpetCol026) (COI, ND1) USA, Texas, San Saba County, San Saba River downstream from Hwy 190 crossing near San Saba, TX (31.212419, -98.79837) UF_439081 (QrefPrl010) (COI, ND1) USA, Louisiana, St. Tammany Parish, Bogue Chitto River near Lock 2 (30.62254, -89.87655) UF_439192 (QmorSaJ073) (COI, ND1) USA, Texas, Montgomery County, West Fork of San Jacinto River NW of Conroe, TX off 105 crossing; site 9 (30.342505, -95.542981) UF_439124 (QpusStF071) (COI, ND1) USA, Missouri, Wayne County, St. Francis River at Camp Lewallen (Bay Scout), 2.25 mi SW of Lodi (37.2371, -90.4857) UF_441085 (QaurGua014) (COI, ND1) USA, Texas, Gonzales/Guadalupe County, San Marcos River between US-90 and SR-80, south of Luling (29.67078, -97.69561) UF_438927 (QaurGua095) (COI, ND1) USA, Texas, Gonzales County, San Marcos River @ CR232 (29.56274, -97.543455) UF_438923 (QmorNec042) (COI, ND1) USA, Texas, Anderson/Cherokee County, Neches River @ CR335 (31.945598, -95.444602) UF_438574 (QaurNue075) (COI, ND1) USA, Texas, Live Oak County, Nueces River, 400 m upstream of Hwy 59 (28.337458, -98.086636) UF_438785 (QpusRed009) (COI, ND1) USA, Texas, Lamar County, Pine Creek, ~100 m upstream from CR906 (33.854656, -95.341305) UF_439081 (QrefPrl009) (COI, ND1) USA, Louisiana, St. Tammany Parish, Bogue Chitto River near Lock 2 (30.62254, -89.87655) UF_438172 (QaurGua046) (COI, ND1) USA, Texas, Dewitt County, Guadalupe River about 15 air km upstream of Cuero, TX (29.221294, -97.329695) UF_438927 (QaurGua093) (COI, ND1) USA, Texas, Gonzales County, San Marcos River @ CR232 (29.56274, -97.543455) UF_439126 (QpusOsa070) (COI, ND1) USA, Missouri, Hickory, Pomme de Terre River approx. 500 , below MDC Cross Timber Access Ramp, 2.8 mi N or Hermitage (37.9815, -93.3081) UF_438785 (QpusRed007) (COI, ND1) USA, Texas, Lamar County, Pine Creek, ~100 m upstream from CR906 (33.854656, -95.341305) UF_438574 (QaurNue064) (COI, ND1) USA, Texas, Live Oak County, Nueces River, 400 m upstream of Hwy 59 (28.337458, -98.086636) UF_438172 (QaurGua049) (COI, ND1) USA, Texas, Dewitt County, Guadalupe River about 15 air km upstream of Cuero, TX (29.221294, -97.329695) UF_438157 (QhouBra042) (COI, ND1) USA, Texas, Williamson County, San Gabriel River about 100m downstream of Co. Road 428 bridge (30.695341, -97.27804) UF_438566 (QaurNue059) (COI, ND1) USA, Texas, Live Oak County, Frio River about 400 m downstream of dam at Choke Canyon (28.48684, -98.23867) UF_438574 (QaurNue076) (COI, ND1) USA, Texas, Live Oak County, Nueces River, 400 m upstream of Hwy 59 (28.337458, -98.086636) UF_438574 (QaurNue072) (COI, ND1) USA, Texas, Live Oak County, Nueces River, 400 m upstream of Hwy 59 (28.337458, -98.086636) UF_438013 (QmorTri013) (COI, ND1) USA, Texas, Leon/Houston County, Trinity River, just downstream of Hwy 7, about 12 air miles W of Crockett, TX (31.3352, -95.65641) UF_441085 (QaurGua012) (COI, ND1) USA, Texas, Gonzales/Guadalupe County, San Marcos River between US-90 and SR-80, south of Luling (29.67078, -97.69561) UF_438168 (QhouBra038) (COI, ND1) USA, Texas, Brazos/Burleson County, Brazos River about 15.5 km downstream from Hwy 21 (30.5811176, -96.4415058) UF_438574 (QaurNue077) (COI, ND1) USA, Texas, Live Oak County, Nueces River, 400 m upstream of Hwy 59 (28.337458, -98.086636) UF_438927 (QaurGua094) (COI, ND1) USA, Texas, Gonzales County, San Marcos River @ CR232 (29.56274, -97.543455) UF_441135 (QhouBra015) (COI, ND1) USA, Texas, Robertson County, Brazos River at FM 485, west of Highway 6 (30.8676502, -96.6956368) UF_438923 (QmorNec038) (COI, ND1) USA, Texas, Anderson/Cherokee County, Neches River @ CR335 (31.945598, -95.444602) UF_438566 (QaurNue060) (COI, ND1) USA, Texas, Live Oak County, Frio River about 400 m downstream of dam at Choke Canyon (28.48684, -98.23867) UF_438919 (QmorNec020) (COI, ND1) USA, Texas, Anderson/Cherokee County, Neches River downstream of SH 79 (31.885207, -95.431684) UF_439081 (QrefPrl007) (COI, ND1) USA, Louisiana, St. Tammany Parish, Bogue Chitto River near Lock 2 (30.62254, -89.87655) UF_438741 (QaurNue088) (COI, ND1) USA, Texas, Live Oak County, Nueces River upstream of Airport Rd (28.382964, -98.109617) UF_438919 (QmorNec030) (COI, ND1) USA, Texas, Anderson/Cherokee County, Neches River downstream of SH 79 (31.885207, -95.431684) UF_441135 (QhouBra013) (COI, ND1) USA, Texas, Robertson County, Brazos River at FM 485, west of Highway 6 (30.8676502, -96.6956368) UF_439198 (QmorSaJ094) (COI, ND1) USA, Texas, San Jacinto County, East Fork San Jacinto River, upstream of FM 945, 300 m upstream from bridge (30.42912, -95.12973) UF_439206 (QpetCol079) (COI, ND1) USA, Texas, Mason County, Llano River at FM1871, ~12 miles SW of Mason, TX (30.658673, -99.3245) UF_438574 (QaurNue073) (COI, ND1) USA, Texas, Live Oak County, Nueces River, 400 m upstream of Hwy 59 (28.337458, -98.086636) UF_439198 (QmorSaJ092) (COI, ND1) USA, Texas, San Jacinto County, East Fork San Jacinto River, upstream of FM 945, 300 m upstream from bridge (30.42912, -95.12973) UF_439198 (QmorSaJ091) (COI, ND1) USA, Texas, San Jacinto County, East Fork San Jacinto River, upstream of FM 945, 300 m upstream from bridge (30.42912, -95.12973) UF_438574 (QaurNue068) (COI, ND1) USA, Texas, Live Oak County, Nueces River, 400 m upstream of Hwy 59 (28.337458, -98.086636) UF_438974 (QmorRed053) (COI, ND1) USA, Texas, Marion County, Big Cypress Bayou (just downstream from Hwy 59, near Jefferson, TX) (32.739928, -94.441666) UF_438927 (QaurGua090) (COI, ND1) USA, Texas, Gonzales County, San Marcos River @ CR232 (29.56274, -97.543455) UF_438741 (QaurNue089) (COI, ND1) USA, Texas, Live Oak County, Nueces River upstream of Airport Rd (28.382964, -98.109617) UF_438886 (QrefPas003) (COI, ND1) USA, Mississippi, Jackson County, Pascagoula River at Paper Mill Camp slough about 2 mi. upstream of MS Hwy 614 (30.63228, -88.652401) UF_438566 (QaurNue050) (COI, ND1) USA, Texas, Live Oak County, Frio River about 400 m downstream of dam at Choke Canyon (28.48684, -98.23867) UF_440981 (QaurGua004) (COI, ND1) USA, Texas, Guadalupe County, Guadalupe River at Hwy 77 crossing (28.83119, -97.06129) UF_439124 (QpusStF073) (COI, ND1) USA, Missouri, Wayne County, St. Francis River at Camp Lewallen (Bay Scout), 2.25 mi SW of Lodi (37.2371, -90.4857) UF_438172 (QaurGua041) (COI, ND1) USA, Texas, Dewitt County, Guadalupe River about 15 air km upstream of Cuero, TX (29.221294, -97.329695) UF_438574 (QaurNue066) (COI, ND1) USA, Texas, Live Oak County, Nueces River, 400 m upstream of Hwy 59 (28.337458, -98.086636) UF_441085 (QaurGua018) (COI, ND1) USA, Texas, Gonzales/Guadalupe County, San Marcos River between US-90 and SR-80, south of Luling (29.67078, -97.69561) UF_439079 (QmorCol060) (COI, ND1) USA, Texas, Wharton, Colorado River near Lane City, TX; Habitat: Run (29.173116, -96.067204) UF_438566 (QaurNue052) (COI, ND1) USA, Texas, Live Oak County, Frio River about 400 m downstream of dam at Choke Canyon (28.48684, -98.23867) UF_438923 (QmorNec035) (COI, ND1) USA, Texas, Anderson/Cherokee County, Neches River @ CR335 (31.945598, -95.444602) UF_439192 (QmorSaJ072) (COI, ND1) USA, Texas, Montgomery County, West Fork of San Jacinto River NW of Conroe, TX off 105 crossing; site 9 (30.342505, -95.542981) UF_438059 (QhouCol029) (COI, ND1) USA, Texas, San Saba County, Colorado River, Burnham's property (31.2617789, -98.5947677) UF_438919 (QmorNec025) (COI, ND1) USA, Texas, Anderson/Cherokee County, Neches River downstream of SH 79 (31.885207, -95.431684) UF_438741 (QaurNue080) (COI, ND1) USA, Texas, Live Oak County, Nueces River upstream of Airport Rd (28.382964, -98.109617) UF_439157 (QpusOhi055) (COI, ND1) USA, Ohio, Wood County, Ohio River; Buckley Island backchannel (39.3912312, -81.4209442) UF_438923 (QmorNec037) (COI, ND1) USA, Texas, Anderson/Cherokee County, Neches River @ CR335 (31.945598, -95.444602) UF_438923 (QmorNec039) (COI, ND1) USA, Texas, Anderson/Cherokee County, Neches River @ CR335 (31.945598, -95.444602) UF_438919 (QmorNec021) (COI, ND1) USA, Texas, Anderson/Cherokee County, Neches River downstream of SH 79 (31.885207, -95.431684) UF_439194 (QmorSaJ071) (COI, ND1) USA, Texas, Liberty County, East Fork of San Jacinto River, Site 14; South of Cleveland at Hwy 59 crossing downstream approx. 20 m (30.29125, -95.121296) UAUC_2436 (ND1) Lake Lewisville, Denton Co., Texas UAUC_2587 (ND1) Ouachita R., Ouachita Co., Arkansas UAUC_1019 (ND1) Mississippi R.,Rock Island Co., Illinois UAUC_1077 (ND1) Big Cypress Bayou, Marion Co., Texas UAUC_866 (ND1) St. Croix R., Wisconsin UAUC_2590 (ND1) Mississippi R., Marion Co., Missouri UAUC_1083 (ND1) Lake Corpus Christi, Live Oak Co., Texas UAUC_405 (ND1) Pascagoula R.,Jackson Co., Mississippi UAUC_1085 (ND1) Lake Corpus Christi, Live Oak Co., Texas UAUC_2372 (ND1) Amite R., East Baton Rogue/Livingston Pa., Louisiana UF_439184 (QpusStC080) (ND1) USA, Minnesota, Chisago County, St. Croix River at Interstate Park (45.3955278, -92.6627834) UAUC_2441 (ND1) Ohio R., Henderson Co., Kentucky UF_439184 (QpusStC079) (ND1) USA, Minnesota, Chisago County, St. Croix River at Interstate Park (45.3955278, -92.6627834) UAUC_2591 (ND1) Ouachita R., Ouachita Co., Arkansas UAUC_658 (ND1) Wolf R., Fayette Co., Tennessee
ASU1232.7 (QpusOua036) (COI, ND1, ITS1, 5.8S) UA_3564 (QpusOhi050) (COI, ND1, ITS1, 5.8S) ASU1231.7 (QpusRed022) (COI, ND1, ITS1, 5.8S) ASU1251.3 (QpusOua085) (COI, ND1, ITS1, 5.8S) ASU1233.11 (QpusStF059) (COI, ND1, ITS1, 5.8S) UA_416 (QpusOhi047) (COI, ND1, ITS1, 5.8S) ASU1232.9 (QpusOua038) (COI, ND1, ITS1, 5.8S) ASU1232.2 (QpusOua031) (COI, ND1, ITS1, 5.8S) UA_900 (QpusStC048) (COI, ND1, ITS1, 5.8S) ASU1233.6 (QpusStF043) (COI, ND1, ITS1, 5.8S) ASU1231.22 (QpusRed029) (COI, ND1, ITS1, 5.8S) ASU1232.8 (QpusOua037) (COI, ND1, ITS1, 5.8S) ASU1231.13 (QpusRed025) (COI, ND1, ITS1, 5.8S) ASU1232.5 (QpusOua034) (COI, ND1, ITS1, 5.8S) ASU1252.1 (QspeOua024) (COI, ND1, ITS1, 5.8S) ASU1233.2 (QpusStF040) (COI, ND1, ITS1, 5.8S) ASU1233.3 (QpusStF041) (COI, ND1, ITS1, 5.8S) ASU1231.4 (QpusRed021) (COI, ND1, ITS1, 5.8S) ASU1251.6 (QpusOua088) (COI, ND1, ITS1, 5.8S) UA_3006 (QspeTri019) (COI, ND1, ITS1, 5.8S) NULL (QhouBra019) (ND1, ITS1, 5.8S) NULL (QhouBra021) (ND1, ITS1, 5.8S) NULL (QhouBra020) (ND1, ITS1, 5.8S) NULL (QhouBra023) (ND1, ITS1, 5.8S) ASU1251.5 (QpusOua087) (COI, ND1) ASU1251.4 (QpusOua086) (COI, ND1) ASU1233.10 (QpusStF058) (COI, ND1) UA_3684/UAUC1083/UAUC1085 (QaurNec112) (COI, ND1) ASU1231.16 (QpusRed027) (COI, ND1) ASU1231.1 (QpusRed020) (COI, ND1) UA_2003 (QnodRed004) (COI, ND1) ASU1233.1 (QpusStF039) (COI, ND1) ASU1251.7 (QpusOua089) (COI, ND1) ASU1231.9 (QpusRed023) (COI, ND1) ASU1231.15 (QpusRed026) (COI, ND1) ASU1232.1 (QpusOua030) (COI, ND1) ASU1232.6 (QpusOua035) (COI, ND1) UA_2001 (QpusRed049) (COI, ND1) ASU1231.18 (QpusRed028) (COI, ND1) UA_362 (QpusOhi046) (COI, ND1) ASU1232.3 (QpusOua032) (COI, ND1) ASU1233.8 (QpusStF057) (COI, ND1) ASU1231.10 (QpusRed024) (COI, ND1) ASU1232.4 (QpusOua033) (COI, ND1) ASU1251.2 (QpusOua084) (COI, ND1) BAH262 (ND1) BAH292 (ND1) GRQP05 (ND1) BAH088 (ND1) BAH261 (ND1) BAH013 (ND1) SRQP12 (ND1) SRQP17 (ND1) ZAM029 (ND1) BAH299 (ND1) BAH046 (ND1) QpusStC082 (COI) GUADL7 (ND1) ZAM016 (ND1) SRQP01 (ND1) ZAM008 (ND1) ZAM040 (ND1) BAH259 (ND1) BAH001 (ND1) BAH090 (ND1) BAH015a (ND1) BAH120 (ND1) BAH015 (ND1) BAH297 (ND1) GUADL9 (ND1) BAH290 (ND1) BAH273 (ND1) GUADL10 (ND1) BAH087 (ND1) ZAM026 (ND1) ZAM004 (ND1) BAH332 (ND1) GRQP16 (ND1) BAH324 (ND1) GUADL3 (ND1) BAH269 (ND1) BAH042 (ND1) SRQP13 (ND1) BAH326 (ND1) BAH016 (ND1) SRQP07 (ND1) TRQP04 (ND1) AT036 (ND1) BAH246 (ND1) AT205 (ND1) AT048 (ND1) BAH089 (ND1) BAH118 (ND1) BAH092 (ND1) BAH004 (ND1) BAH006 (ND1) BAH325 (ND1) BAH321 (ND1) GRQP07 (ND1) GUADL6 (ND1) GUADL12 (ND1) TRQP14 (ND1) BAH163 (ND1) GRQP03 (ND1) SRQP03 (ND1) SRQP11 (ND1) ZAM030 (ND1) AT219 (ND1) BAH351 (ND1) BAH253 (ND1) BAH254 (ND1) BAH125 (ND1) SRQP18 (ND1) BAH043 (ND1) AT028 (ND1) BAH162 (ND1) BAH012a (ND1) UA_416 (SRA) AT207 (ND1) AT216 (ND1) AT224 (ND1) AT026 (ND1) BAH271 (ND1) QhouBra023 (COI) GRQP15 (ND1) BAH255 (ND1) BAH119 (ND1) BAH301 (ND1) BAH298 (ND1) TRQP05 (ND1) SRQP02 (ND1) BAH062 (ND1) SRQP19 (ND1) ZAM009 (ND1) BAH333 (ND1) TRQP02 (ND1) AT052 (ND1) ZAM012 (ND1) BAH295 (ND1) BAH300 (ND1) SRQP20 (ND1) GRQP04 (ND1) GRQP08 (ND1) BAH304 (ND1) BAH086 (ND1) AT049 (ND1) AT042 (ND1) GRQP18 (ND1) AT218 (ND1) BAH091 (ND1) QpusStC080 (COI) BAH265 (ND1) BAH251 (ND1) AT078 (ND1) GUADL2 (ND1) ZAM024 (ND1) BAH258 (ND1) BAH131 (ND1) BAH257 (ND1) GRQP12 (ND1) BAH263 (ND1) QhouBra019 (COI) AT039 (ND1) BAH294 (ND1) TRQP13 (ND1) ZAM022 (ND1) SRQP14 (ND1) BAH124 (ND1) QpusStC081 (COI) BAH044 (ND1) BAH134 (ND1) AT044 (ND1) AT093 (ND1) QhouBra020 (COI) GRQP10 (ND1) GRQP13 (ND1) AT046 (ND1) BAH045 (ND1) BAH093 (ND1) GRQP19 (ND1) BAH264 (ND1) AT098 (ND1) BAH133 (ND1) AT054 (ND1) BAH137 (ND1) GRQP17 (ND1) TRQP01 (ND1) BAH247 (ND1) BAH095 (ND1) GRQP20 (ND1) AT053 (ND1) SRQP05 (ND1) BAH074 (ND1) BAH322 (ND1) BAH160 (ND1) QpusStC079 (COI) BAH126 (ND1) BAH012 (ND1) QhouBra021 (COI) BAH270 (ND1) TRQP03 (ND1) SRQP04 (ND1) BAH323 (ND1) GRQP01 (ND1) GRQP06 (ND1) BAH328 (ND1) GRQP09 (ND1) BAH014 (ND1) GUADL1 (ND1) GRQP14 (ND1) TRQP06 (ND1) AT217 (ND1) TRQP15 (ND1) BAH267 (ND1) AT206 (ND1) BAH291 (ND1) BAH014a (ND1) GRQP02 (ND1) BAH329 (ND1) TRQP08 (ND1)
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