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The Freshwater Mussels (Unionoida) of the World
(and other less consequential bivalves)

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Taxa known only as fossils are highlighted in gray. Those without representatives in fresh waters are highlighted in gold.

 

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Publication Information

Zhadin, V.I. 1938. Sem. Unionidae. Faune URSS, Mollusques 4(1): 169 pp.

Species Described
Anodonta sedakovi Zhadin, 1938.
‘Siemaschko’ Zhadin, 1938, Faune USSR Mollusques: 130.
Zhadin, 1965, Moll. USSR: 252.
Bogatov et al., 2005, Zool. Zhur.: 1059.
Kantor & Sysoev, 2005, Cat. Moll. Russia: 334 [as ‘Colletopterum ponderosum sedakovi’].
(error for Anodonta sedakowi Siemaschko, 1848)
= Anodonta anatina
 
Anodonta tehernaica Westerlund, 1890.
‘Bourguignat’ Of Authors.
Westerlund, 1890, Fauna Paläarct. Binnenconch.: 226.
Zhadin, 1938, Faune USSR Mollusques: 121.
(error for Anodonta tchernaica Bourguignat, 1881)
= Anodonta anatina
 
Anodonta (Haasiella) arcaeformis Zhadin, 1938.
‘Heude’ Zhadin, 1938, Faune USSR Mollusques: 147, fig. 56.
Zhadin, 1965, Moll. USSR: 260, fig. 279.
(reference to Anodon arcaeformis Heude, 1877)
= Anemina arcaeformis
 
Anodonta (Haasiella) euscaphus Zhadin, 1938.
‘Heude’ Zhadin, 1938, Faune USSR Mollusques: 149, fig. 58.
Zhadin, 1965, Moll. USSR: 260, fig. 280.
(reference to Anodon euscaphys Heude, 1879; error for Anodon euscaphys Heude, 1879)
= Anemina euscaphys
 
Anodonta anatina var. lenae Zhadin, 1938.
Zhadin, 1938, Faune USSR Mollusques: 129, fig. 45.
(intro'd as intra-sp. of Mytilus anatinus Linnaeus, 1758)
= Anodonta anatina
 
Anodonta anatina var. petshorica Zhadin, 1938.
Zhadin, 1938, Faune USSR Mollusques: 129, fig. 44.
(intro'd as intra-sp. of Mytilus anatinus Linnaeus, 1758)
= Anodonta anatina
 
Anodonta beringiana var. suifunensis Zhadin, 1938.
Zhadin, 1938, Faune USSR Mollusques: 132, fig. 48.
Zhadin, 1965, Moll. USSR: 255, fig. 272.
(intro'd as intra-sp. of Anodonta beringiana Middendorff, 1851)
= Anemina arcaeformis
 
Anodonta beringiana var. taranetzi Zhadin, 1938.
Zhadin, 1938, Faune USSR Mollusques: 133, fig. 49.
Zhadin, 1965, Moll. USSR: 255, fig. 273.
(intro'd as intra-sp. of Anodonta beringiana Middendorff, 1851)
= Sinanodonta woodiana
 
Anodonta cellensis var. sorensis Zhadin, 1938.
‘Dybowski’ Zhadin, 1938, Faune USSR Mollusques: 120.
(error for Anodonta sorensiana Dybowski, 1913)
= Anodonta anatina
 
Anodonta cyrea var. samarkadensis Zhadin, 1938.
‘Kobelt’ Zhadin, 1938, Faune USSR Mollusques: 140.
(error for Anodonta samarkandensis Kobelt, 1913)
= Anodonta cyrea
 
Anodonta piscinalis var. volgensis Zhadin, 1938.
Zhadin, 1938, Faune USSR Mollusques: 126, fig. 43.
Zhadin, 1965, Moll. USSR: 252, fig. 268.
(intro'd as intra-sp. of Anodonta piscinalis Nilsson, 1823)
= Anodonta anatina
 
Anodonta woodiana var. elliptica Zhadin, 1938.
‘Heude’ Zhadin, 1938, Faune USSR Mollusques: 144, fig. 54.
Zhadin, 1965, Moll. USSR: 256, fig. 276.
(reference to Anodon elliptica Heude, 1878)
= Sinanodonta woodiana
 
Margaritana sachalinensis Zhadin, 1938.
Zhadin, 1938, Faune USSR Mollusques: 114-115, fig. 37.
Zhadin, 1965, Moll. USSR: 238, fig. 252.
= Margaritifera laevis
 
Middendorffinaia (Pseudopotomida) shadini Moskvicheva & Starobogatov, 1973.
‘Haas’ Zhadin, 1938, Faune USSR Mollusques: 100, fig. 31 [as ‘Unio continentalis’].
‘Haas’ Zhadin, 1952, Moll. SSSR: 299, fig. 263 [as ‘Unio continentalis’].
Moskvicheva & Starobogatov, 1973, Byull. Mosk. Obshch.. Ispyt. Prir.: 31, figs. 5а-в.
= Inversidens pantoensis
 
Mytilus cellensis Menke, 1828.
‘Gmelin’ Of Authors.
Menke, 1828, Syn. Method. Moll.: 63.
Menke, 1830, Syn. Method. Moll.: 106.
Küster, 1839, Conchyl. Cab.: 16.
Mörch, 1853, Cat. Conch. Yoldi: 48.
Drouët, 1881, Unionidae Russie: 22.
Bourguignat, 1882, Natural. Sicil.: 20 [1].
Westerlund, 1890, Fauna Paläarct. Binnenconch.: 211.
Zhadin, 1938, Faune USSR Mollusques: 119.
Modell, 1945, Jena. Z. Med. Naturw.: 81.
(error for Mytilus zellensis Gmelin, 1791)
= Anodonta cygnea
 
Psilunio rothi var. komarovi Zhadin, 1938.
‘Boettg.’ Zhadin, 1938, Faune USSR Mollusques: 102.
(error for Unio komarowi O. Boettger, 1880)
= Potomida littoralis
 
Sinanodonta (Anemina) fuscoviridis Moskvicheva, 1973.
‘Heude’ Zhadin, 1938, Faune USSR Mollusques: 149, fig. 58 [as ‘Anodonta (Haasiella) euscaphus’].
Moskvicheva, 1973, Zool. Zhur.: 830.
= Anemina euscaphys
 
Sinanodonta (Anemina) shadini Moskvicheva, 1973.
‘Heude’ Zhadin, 1938, Faune USSR Mollusques: 147, fig. 56 [as ‘Anodonta arcaeformis’].
Moskvicheva, 1973, Zool. Zhur.: 828.
= Anemina arcaeformis
 
Unio lindholmi Zhadin, 1938.
Zhadin, 1938, Faune USSR Mollusques: 97, text fig. 29.
= Unio pictorum
 
Unio (Nodularia) continentalis Zhadin, 1938.
‘Haas’ Zhadin, 1938, Faune USSR Mollusques: 100, fig. 31.
‘Haas’ Zhadin, 1952, Moll. SSSR: 299, fig. 263.
(reference to Nodularia continentalis Haas, 1910)
= Inversidens pantoensis
 
Unio (Nodularia) douglasiae var. schrenki Zhadin, 1938.
‘Westerlund’ Zhadin, 1938, Faune USSR Mollusques: 99.
(error for Unio schrencki Westerlund, 1897)
= Nodularia douglasiae
 
Unio crassus consentancus Zhadin, 1938.
‘Ziegler’ Zhadin, 1938, Faune USSR Mollusques: 91.
(error for Unio consentaneus Rossmässler, 1836)
= Unio crassus
 
Unio crassus var. chlebnikovi Zhadin, 1938.
‘Kobelt’ Zhadin, 1938, Faune USSR Mollusques: 89.
(new name for Unio chlebnikowi Kobelt, 1912)
= Unio crassus
 
Unio crassus var. ishmensis Zhadin, 1938.
Zhadin, 1938, Faune USSR Mollusques: 160.
(intro'd as intra-sp. of Unio crassus Retzius, 1788)
= Unio crassus
 
Unio crassus var. stepanovi Zhadin, 1938.
‘Drouët’ Zhadin, 1938, Faune USSR Mollusques: 90.
(error for Unio stepanoffi Drouët, 1881)
= Unio crassus
 
Unio mingrelicus var. stevenianiformis Zhadin, 1938.
Zhadin, 1938, Faune USSR Mollusques: 96, text fig. 28.
(intro'd as intra-sp. of Unio mingrelicus Drouët, 1881)
= Unio crassus
 
Unio pictorum var. defectivus Zhadin, 1938.
Zhadin, 1938, Faune USSR Mollusques: 79.
(intro'd as intra-sp. of Mya pictorum Linnaeus, 1758)
= Unio pictorum
 
Unio pictorum var. okensis Zhadin, 1938.
Zhadin, 1938, Faune USSR Mollusques: 77.
(intro'd as intra-sp. of Mya pictorum Linnaeus, 1758; new name for Unio okae Kobelt & Haas, 1911)
= Unio pictorum
 
Unio pictorum var. pygmaeus Zhadin, 1938.
Zhadin, 1938, Faune USSR Mollusques: 78, text fig. 17.
(intro'd as intra-sp. of Mya pictorum Linnaeus, 1758; prim. homonym of Unio pygmaeus Lea, 1852)
= Unio pictorum
 
Unio raddei var. kutaisiana ‘Kobelt’ of Authors.
‘Kobelt’ Of Authors.
Zhadin, 1938, Faune USSR Mollusques: 94.
Zatravkin, 1987, Byull. Mosk. Obshch.. Ispyt. Prir.: 49.
(error for Unio koutaisiana Kobelt, 1886)
= Unio crassus
 
Unio tumidus var. bashkiricus Zhadin, 1938.
Zhadin, 1938, Faune USSR Mollusques: 81, text fig. 19.
(intro'd as intra-sp. of Unio tumidus Retzius, 1788)
= Unio tumidus
 
Unio tumidus var. fridmani Zhadin, 1938.
Zhadin, 1938, Faune USSR Mollusques: 86, text fig. 21.
(intro'd as intra-sp. of Unio tumidus Retzius, 1788)
= Unio tumidus
 
Unio tumidus var. ilikensis Zhadin, 1938.
‘Kobelt’ Of Authors.
Zhadin, 1938, Faune USSR Mollusques: 83.
Kantor & Sysoev, 2005, Cat. Moll. Russia: 327.
(error for Unio ilekensis Kobelt, 1912)
= Unio tumidus
 
Unio tumidus var. kobeltianus Zhadin, 1938.
Zhadin, 1938, Faune USSR Mollusques: 82.
(intro'd as intra-sp. of Unio tumidus Retzius, 1788; new name for Unio okae Kobelt & Haas, 1911; prim. homonym of Unio kobeltianus Haas, 1913)
= Unio tumidus
 
Unio tumidus var. moltshanovi Zhadin, 1938.
Zhadin, 1938, Faune USSR Mollusques: 85, text fig. 20.
(intro'd as intra-sp. of Unio tumidus Retzius, 1788)
= Unio tumidus
 
Taxonomic Opinions
Anodonta (Haasiella) arcaeformis (Heude, 1877). — Zhadin (1938: 147).
Anodonta (Haasiella) arcaeformis var. suifunica Lindholm, 1925. — Zhadin (1938: 149).
Anodonta (Haasiella) euscaphys (Heude, 1879). — Zhadin (1938: 149).
Anodonta (Pseudanodonta) complanata ‘Ziegl.’ Rossmässler, 1835. — Zhadin (1938: 145).
Anodonta (Pseudanodonta) complanata var. borealis Kobelt, 1891. — Zhadin (1938: 146).
Anodonta (Pseudanodonta) complanata var. middendorffi Siemaschko, 1848. — Zhadin (1938: 146).
Anodonta (Pseudanodonta) complanata var. nordenskiöldi Boettger, 1880 [sic]. — Zhadin (1938: 146).
Anodonta (Pseudanodonta) rossmaessleri Bourguignat, 1881. — Zhadin (1938: 147).
Anodonta (Pteranodon) woodiana (Lea, 1834). — Zhadin (1938: 140).
Anodonta (Pteranodon) woodiana var. elliptica (Heude, 1878). — Zhadin (1938: 144).
Anodonta (s.s.) anatina (Linnaeus, 1758). — Zhadin (1938: 127).
Anodonta (s.s.) anatina var. convexa Drouet, 1888. — Zhadin (1938: 128).
Anodonta (s.s.) anatina var. lenae Zhadin, 1938. — Zhadin (1938: 129).
Anodonta (s.s.) anatina var. petshorica Zhadin, 1938. — Zhadin (1938: 129).
Anodonta (s.s.) anatina var. viridiflava Drouet, 1893. — Zhadin (1938: 128).
Anodonta (s.s.) beringiana Middendorff, 1851. — Zhadin (1938: 131).
Anodonta (s.s.) beringiana var. suifunensis Zhadin, 1938. — Zhadin (1938: 132).
Anodonta (s.s.) beringiana var. taranetzi Zhadin, 1938. — Zhadin (1938: 133).
Anodonta (s.s.) cellensis [sic] (Gmelin, 1791). — Zhadin (1938: 119).
Anodonta (s.s.) cellensis [sic] var. cariosa Küster, 1876 [sic]. — Zhadin (1938: 121).
Anodonta (s.s.) cellensis [sic] var. maganica Servain, 1882. — Zhadin (1938: 120).
Anodonta (s.s.) cellensis [sic] var. selengensis Dybowski, 1913. — Zhadin (1938: 121).
Anodonta (s.s.) cellensis [sic] var. sorensis [sic] Dybowski, W., 1913. — Zhadin (1938: 120).
Anodonta (s.s.) cygnea (Linnaeus, 1758). — Zhadin (1938: 117).
Anodonta (s.s.) cyrea Drouet, 1881. — Zhadin (1938: 134).
Anodonta (s.s.) cyrea var. armenica Ovtschinnikov, 1935. — Zhadin (1938: 136).
Anodonta (s.s.) cyrea var. bactriana Rolle, 1897. — Zhadin (1938: 137).
Anodonta (s.s.) cyrea var. georgiana Drouet, 1881. — Zhadin (1938: 136).
Anodonta (s.s.) cyrea var. lenkoranensis Drouet, 1881. — Zhadin (1938: 136).
Anodonta (s.s.) cyrea var. samarkadensis [sic] Kobelt, 1913. — Zhadin (1938: 140).
Anodonta (s.s.) cyrea var. sogdiana Kobelt, 1896. — Zhadin (1938: 137).
Anodonta (s.s.) piscinalis Nilsson, 1823. — Zhadin (1938: 121).
Anodonta (s.s.) piscinalis var. crimeana Bourguignat, 1881. — Zhadin (1938: 124).
Anodonta (s.s.) piscinalis var. letourneuxi Bourguignat, 1870. — Zhadin (1938: 124).
Anodonta (s.s.) piscinalis var. ostiaria Drouet, 1881. — Zhadin (1938: 124).
Anodonta (s.s.) piscinalis var. ponderosa Pfeiffer, C., 1825. — Zhadin (1938: 125).
Anodonta (s.s.) piscinalis var. seisanensis Kobelt, 1912. — Zhadin (1938: 126).
Anodonta (s.s.) piscinalis var. sorica Dybowski, 1913. — Zhadin (1938: 127).
Anodonta (s.s.) piscinalis var. volgensis Zhadin, 1938. — Zhadin (1938: 126).
Anodonta (s.s.) sedakovi [sic] Siemaschko, 1848. — Zhadin (1938: 130).
Anodonta (s.s.) sedakowi var. nova Dybowski, 1913. — Zhadin (1938: 130).
Cristaria plicata (Leach, 1815). — Zhadin (1938: 150).
Cristaria plicata var. herculea Middendorff, 1848. — Zhadin (1938: 153).
Lanceolaria cylindrica Simpson, 1900. — Zhadin (1938: 104).
Lanceolaria grayana (Lea, 1834). — Zhadin (1938: 103).
Margaritana dahurica (Middendorff, 1850). — Zhadin (1938: 109).
Margaritana margaritifera (Linnaeus, 1758). — Zhadin (1938: 106).
Margaritana middendorffi Rosén, 1926. — Zhadin (1938: 112).
Margaritana mongolica (Middendorff, 1851). — Zhadin (1938: 115).
Margaritana sachalinensis Zhadin, 1938. — Zhadin (1938: 114).
Psilunio rothi (Bourguignat, 1863). — Zhadin (1938: 101).
Psilunio rothi var. armeniacus (Kobelt, 1912). — Zhadin (1938: 102).
Psilunio rothi var. komarovi [sic] (Boettger, O., 1880). — Zhadin (1938: 102).
Unio (Nodularia) continentalis (Haas, 1910). — Zhadin (1938: 100).
Unio (Nodularia) douglasiae Gray in Griffith & Pidgeon, 1834. — Zhadin (1938: 98).
Unio (Nodularia) douglasiae var. schrenki [sic] Westerlund, 1897. — Zhadin (1938: 99).
Unio (s.s.) ater Nilsson, 1822. — Zhadin (1938: 91).
Unio (s.s.) crassus Retzius, 1788. — Zhadin (1938: 86).
Unio (s.s.) crassus consentancus [sic] ‘Ziegler’ Rossmässler, 1836. — Zhadin (1938: 91).
Unio (s.s.) crassus kungurensis Kobelt, 1912. — Zhadin (1938: 89).
Unio (s.s.) crassus var. chlebnikovi [sic] Kobelt, 1912. — Zhadin (1938: 89).
Unio (s.s.) crassus var. ishmensis Zhadin, 1938. — Zhadin (1938: 90).
Unio (s.s.) crassus var. okae Kobelt, 1911. — Zhadin (1938: 89).
Unio (s.s.) crassus var. pseudolitoralis Clessin, 1875. — Zhadin (1938: 88).
Unio (s.s.) crassus var. roseni Kobelt, 1912. — Zhadin (1938: 91).
Unio (s.s.) crassus var. stepanovi [sic] Drouet, 1881. — Zhadin (1938: 90).
Unio (s.s.) hueti Bourguignat, 1855. — Zhadin (1938: 98).
Unio (s.s.) lindholmi Zhadin, 1938. — Zhadin (1938: 97).
Unio (s.s.) mingrelicus Drouet, 1881. — Zhadin (1938: 95).
Unio (s.s.) mingrelicus var. gregorii Kobelt, 1912. — Zhadin (1938: 96).
Unio (s.s.) mingrelicus var. stevenianiformis Zhadin, 1938. — Zhadin (1938: 96).
Unio (s.s.) pictorum (Linnaeus, 1758). — Zhadin (1938: 74).
Unio (s.s.) pictorum defectivus Zhadin, 1938. — Zhadin (1938: 79).
Unio (s.s.) pictorum gentilis Haas in Hesse, 1911. — Zhadin (1938: 79).
Unio (s.s.) pictorum var. annulatus Kobelt, 1912. — Zhadin (1938: 78).
Unio (s.s.) pictorum var. okensis Zhadin, 1938. — Zhadin (1938: 77).
Unio (s.s.) pictorum var. platyrhynchus Rossmässler, 1835. — Zhadin (1938: 77).
Unio (s.s.) pictorum var. ponderosus Spitzi in Rossmässler, 1838. — Zhadin (1938: 76).
Unio (s.s.) pictorum var. pygmaeus Zhadin, 1938. — Zhadin (1938: 78).
Unio (s.s.) pictorum var. schrenkianus Clessin, 1880. — Zhadin (1938: 77).
Unio (s.s.) sieversi Drouet, 1881. — Zhadin (1938: 93).
Unio (s.s.) sieversi var. araxenus Drouet, 1881. — Zhadin (1938: 94).
Unio (s.s.) sieversi var. colchicus Drouet, 1881. — Zhadin (1938: 95).
Unio (s.s.) sieversi var. kutaisiana [sic] Kobelt, 1886. — Zhadin (1938: 94).
Unio (s.s.) sieversi var. raddei Drouet, 1881. — Zhadin (1938: 94).
Unio (s.s.) stevenianus Drouet, 1881. — Zhadin (1938: 92).
Unio (s.s.) tumidus Retzius, 1788. — Zhadin (1938: 80).
Unio (s.s.) tumidus limicola Mörch, 1864. — Zhadin (1938: 83).
Unio (s.s.) tumidus var. bashkiricus Zhadin, 1938. — Zhadin (1938: 81).
Unio (s.s.) tumidus var. borysthenicus Kobelt, 1879. — Zhadin (1938: 83).
Unio (s.s.) tumidus var. conus Spengler, 1793. — Zhadin (1938: 86).
Unio (s.s.) tumidus var. fridmani Zhadin, 1938. — Zhadin (1938: 86).
Unio (s.s.) tumidus var. gerstfeldtianus Clessin, 1880. — Zhadin (1938: 83).
Unio (s.s.) tumidus var. ilikensis [sic] Kobelt, 1912. — Zhadin (1938: 83).
Unio (s.s.) tumidus var. kobeltianus Zhadin, 1938. — Zhadin (1938: 82).
Unio (s.s.) tumidus var. moltshanovi Zhadin, 1938. — Zhadin (1938: 85).
 
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