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Page last updated
Sun 31 Oct 2021

The Freshwater Mussels (Unionoida) of the World (and other less consequential bivalves)

FM(U)otW(aolcb) is the web version of the MUSSEL Project Database. Follow the links to browse the data or use the custom Google search field. Either way, you win!

The nominal species (available and otherwise) in this synonymy are listed by usage rather than strictly by publication date. The primary sorting order is by the most recent year (since 1969) that the species name was treated as valid, and then nominal species are sorted secondarily by publication year.

The taxonomic history of the valid species (since 1900) is listed at the bottom of the page.

Search the database pages:

Unionoidea | Unionidae | Unioninae | Anodontini | Cristariina | Beringiana

species Beringiana beringiana (Middendorff, 1851): nominal species
     1 specimen available.

Anodonta cellensis var. beringiana Middendorff, 1851
Middendorff, 1851, Reise Nord. Ost. Sibiriens 2(1): 284, pl. 28, figs. 4-7; pl. 29, figs. 1-4.
(intro'd as intra-sp. of Anodonta cellensis Pfeiffer, 1821)

Anodonta cellensis var. beringianaMiddendorff (1851)
Margaron (Anodonta) cygneaLea (1870)
Anodonta beringianaSimpson (1900), Simpson (1914), Burch (1975), R.I. Johnson (1980), ..., Klishko et al. (2016), Froufe et al. (2017), Inoue et al. (2018), Zieritz et al. (2018)
Anodonta cygnea beringianaHannibal (1912)
Anodonta (s.s.) beringianaFrierson (1927), Zhadin (1938), Zhadin (1965), Haas (1969)
Anodonta marginata beringianaModell (1945)
Anodonta (Sinanodonta) beringianaHabe (1977)
Beringiana beringianaZatravkin (1983), Zatravkin & Bogatov (1987), Korniushin (1998), Bogatov & Starobogatov (2001), ..., Lopes-Lima et al. (2020), Bolotov et al. (2020), Graf & Cummings (2021), Bieler (2021)
Beringiana baringianaBogatov & Zatravkin (1988)
Sinanodonta beringianaWilliams et al. (2017)

Anodonta youconensis Lea, 1867
Lea, 1867, Proc. Acad. Nat. Sci. 19: 81.
(incorrect original spelling of Anodonta youkanensis Lea, 1867)

Anodonta youconensisLea (1867)
Anodonta beringianaSimpson (1900), Simpson (1914)
Anodonta cygnea beringianaHannibal (1912)
Anodonta (s.s.) beringianaFrierson (1927)
Anodonta (s.s.) oregonensisHaas (1969)
Beringiana youkanensisZatravkin & Bogatov (1987)
Beringiana youconensisKantor & Sysoev (2005), Kantor et al. (2010), Vinarski & Kantor (2016)
Beringiana beringianaBieler (2021)

Anodonta beringiana var. taranetzi Zhadin, 1938
Zhadin, 1938, Faune USSR Mollusques 4(1): 133, fig. 49.
Zhadin, 1965, Moll. USSR: 255, fig. 273.
(intro'd as intra-sp. of Anodonta cellensis var. beringiana Middendorff, 1851)

Anodonta beringiana var. taranetziZhadin (1938)
Anodonta (s.s.) beringiana var. taranetziZhadin (1938), Zhadin (1965)
Anodonta (s.s.) beringianaHaas (1969)
Beringiana taranetziZatravkin (1983), Zatravkin & Bogatov (1987), Bogatov & Zatravkin (1988), Vinarski & Kantor (2016)
Kunashiria taranetziStarobogatov et al. (2004), Sayenko et al. (2009)
Kunashiria taranetsiKantor & Sysoev (2005), Kantor et al. (2010)
Anodonta woodiana japonicaGraf (2007)
Sinanodonta woodianaHe & Zhuang (2013)
Beringiana beringianaLopes-Lima et al. (2020), Bolotov et al. (2020)

Arsenievinaia coptzevi Zatravkin & Bogatov, 1987
Zatravkin & Bogatov, 1987, Biv. Far East: 104, sp. 51, figs. 20 б, 24 в-г.

Arsenievinaia coptzeviZatravkin & Bogatov (1987), Bogatov & Zatravkin (1988)
Kunashiria coptzeviStarobogatov et al. (2004), Kantor & Sysoev (2005), Sayenko et al. (2009), Kantor et al. (2010)
Anodonta woodiana japonicaGraf (2007)
Sinanodonta woodianaHe & Zhuang (2013)
Beringiana coptzeviVinarski & Kantor (2016)
Beringiana beringianaLopes-Lima et al. (2020), Bolotov et al. (2020)

Arsenievinaia zimini Zatravkin & Bogatov, 1987
Zatravkin & Bogatov, 1987, Biv. Far East: 102, sp. 50, figs. 24 а-б.

Arsenievinaia ziminiZatravkin & Bogatov (1987), Bogatov & Zatravkin (1988)
Arsenievinaia sihotealinicaMartynov & Chernyshev (1992)
Kunashiria ziminiStarobogatov et al. (2004), Kantor & Sysoev (2005), Sayenko et al. (2009), Kantor et al. (2010)
Anodonta woodiana japonicaGraf (2007)
Sinanodonta woodianaHe & Zhuang (2013)
Beringiana ziminiVinarski & Kantor (2016)
Beringiana beringianaLopes-Lima et al. (2020), Bolotov et al. (2020)

Arsenievinaia zarjaensis Bogatov & Zatravkin, 1988
Bogatov & Zatravkin, 1988, Trudy Zool. Inst. 187: 166, fig. 6.

Arsenievinaia zarjaensisBogatov & Zatravkin (1988)
Kunashiria zarjaensisStarobogatov et al. (2004), Kantor & Sysoev (2005), Sayenko et al. (2009), Kantor et al. (2010)
Anodonta woodiana japonicaGraf (2007)
Sinanodonta woodianaHe & Zhuang (2013)
Beringiana zarjaensisVinarski & Kantor (2016)
Beringiana beringianaLopes-Lima et al. (2020), Bolotov et al. (2020)

Arsenievinaia compressa Bogatov & Starobogatov, 1996
Bogatov & Starobogatov, 1996, Zool. Zhur. 75: 1332. fig’d.

Arsenievinaia compressaBogatov & Starobogatov (1996)
Kunashiria compressaStarobogatov et al. (2004), Kantor & Sysoev (2005), Sayenko et al. (2009), Kantor et al. (2010)
Anodonta woodiana japonicaGraf (2007)
Sinanodonta woodianaHe & Zhuang (2013)
Beringiana compressaVinarski & Kantor (2016)
Beringiana beringianaLopes-Lima et al. (2020), Bolotov et al. (2020)

Beringiana compressa Sayenko & Bogatov, 1998
Sayenko & Bogatov, 1998, Zool. Zhur. 77(12): 1414, figs. 2-4.
(homonym of Arsenievinaia compressa Bogatov & Starobogatov, 1996)

Beringiana compressaSayenko & Bogatov (1998), Bogatov & Starobogatov (2001), Starobogatov et al. (2004), Kantor & Sysoev (2005), Kantor et al. (2010), Vinarski & Kantor (2016)
Anodonta beringianaGraf (2007)
Beringiana beringianaLopes-Lima et al. (2020), Bolotov et al. (2020)

Kunashiria iturupica Bogatov et al., 1999
Bogatov et al., 1999, Ruthenica 9: 60, fig’d.

Kunashiria iturupicaBogatov et al. (1999), Starobogatov et al. (2004), Kantor & Sysoev (2005), Sayenko et al. (2009), Kantor et al. (2010)
Anodonta woodiana japonicaGraf (2007)
Sinanodonta woodianaHe & Zhuang (2013)
Beringiana iturupicaVinarski & Kantor (2016)
Beringiana beringianaLopes-Lima et al. (2020), Bolotov et al. (2020)

Kunashiria sinanodontoides Bogatov et al., 1999
Bogatov et al., 1999, Ruthenica 9: 59, fig’d.

Kunashiria sinanodontoidesBogatov et al. (1999), Starobogatov et al. (2004), Kantor & Sysoev (2005), Sayenko et al. (2009), Kantor et al. (2010)
Anodonta woodiana japonicaGraf (2007)
Sinanodonta woodianaHe & Zhuang (2013)
Beringiana sinanodontoidesVinarski & Kantor (2016)
Beringiana beringianaLopes-Lima et al. (2020), Bolotov et al. (2020)

Beringiana chereshnevi Bogatov & Starobogatov, 2001
Bogatov & Starobogatov, 2001, Zool. Zhur. 80: 26, figs. 2а-б, 4ж.

Beringiana chereshneviBogatov & Starobogatov (2001), Starobogatov et al. (2004), Kantor & Sysoev (2005), Kantor et al. (2010), Vinarski & Kantor (2016)
Anodonta beringianaGraf (2007)
Beringiana beringianaLopes-Lima et al. (2020), Bolotov et al. (2020)

Beringiana derzhavini Bogatov & Starobogatov, 2001
Bogatov & Starobogatov, 2001, Zool. Zhur. 80: 27, figs. 2в-г, 4а.

Beringiana derzhaviniBogatov & Starobogatov (2001), Starobogatov et al. (2004), Kantor & Sysoev (2005), Kantor et al. (2010), Vinarski & Kantor (2016)
Anodonta beringianaGraf (2007)
Beringiana beringianaLopes-Lima et al. (2020), Bolotov et al. (2020)

Beringiana kamchatica Bogatov & Starobogatov, 2001
Bogatov & Starobogatov, 2001, Zool. Zhur. 80: 27, figs. 2д-е, 4г.

Beringiana kamchaticaBogatov & Starobogatov (2001), Starobogatov et al. (2004), Kantor & Sysoev (2005), Sayenko et al. (2009), Kantor et al. (2010), Vinarski & Kantor (2016)
Anodonta beringianaGraf (2007)
Beringiana beringianaLopes-Lima et al. (2020), Bolotov et al. (2020)

Anodonta youkanensis Lea, 1867
Lea, 1867, Proc. Acad. Nat. Sci. 19: 81 [as ‘youconensis’].
Lea, 1868, Jour. Acad. Nat. Sci. 6 [O. 12]: 287 [47], pl. 40, fig. 99.
(correct name for Anodonta youconensis Lea, 1867)
Type(s): FIGURED SYNTYPE USNM_86624, Upper Yukon R., Alaska.

Anodonta youkanensisLea (1867), B.H. Wright (1888)
Margaron (Anodonta) youkanensisLea (1870)
Anodonta beringianaSimpson (1900), Simpson (1914), Burch (1975), Graf (2007), IUCN (2011)
Anodonta cygnea beringianaHannibal (1912)
Anodonta (s.s.) beringianaFrierson (1927)
Anodonta (s.s.) oregonensisHaas (1969)
Beringiana youkonensisZatravkin (1983), Bogatov & Starobogatov (2001), Starobogatov et al. (2004), Sayenko et al. (2009)
Beringiana youkanensisZatravkin & Bogatov (1987)
Beringiana yuokonensisBogatov & Zatravkin (1988)
Beringiana youconensisKantor & Sysoev (2005), Kantor et al. (2010), Vinarski & Kantor (2016)
Beringiana beringianaLopes-Lima et al. (2020), Bolotov et al. (2020), Bieler (2021)

Amuranodonta sihotealinica Zatravkin & Starobogatov, 1984
Zatravkin & Starobogatov, 1984, Zool. Zhur. 63(12): 1788, figs. 9-10.

Amuranodonta sihotealinicaZatravkin & Starobogatov (1984)
Amuranodonta sp. nov.Zatravkin (1983)
Arsenievinaia sihotealinicaZatravkin & Bogatov (1987), Bogatov & Zatravkin (1988), Martynov & Chernyshev (1992), Bogatov & Sayenko (2002)
Kunashiria haconensisStarobogatov et al. (2004), Kantor & Sysoev (2005), Sayenko et al. (2009), Kantor et al. (2010)
Sinanodonta woodianaHe & Zhuang (2013)
Beringiana haconensisVinarski & Kantor (2016)
Beringiana beringianaLopes-Lima et al. (2020), Bolotov et al. (2020)

Anodonta youcanensis Lea, 1867
‘Lea’ Lea, 1867, Obs., index I-XI: 63.
‘Lea’ Paetel, 1890, Cat. Conch.-Samml. 3: 186.
(error for Anodonta youkanensis Lea, 1867)

Anodonta youcanensisLea (1867)
Anodonta beringianaSimpson (1900), Simpson (1914)
Beringiana beringianaBieler (2021)

Anodon youkamensis Sowerby, 1870
‘Lea’ Sowerby, 1870, Conch. Icon. 17: index.
(error for Anodonta youkanensis Lea, 1867)

Anodon youkamensisSowerby (1870)
Beringiana beringianaBieler (2021)

Anodonta youkonensis Simpson, 1893
‘Lea’ of Authors.
Simpson, 1893, Nautilus 6(12): 135.
Zatravkin, 1983, [in] I.M. Likharev, Molluscs: 42.
Bogatov & Starobogatov, 2001, Zool. Zhur. 80: 30.
Starobogatov et al., 2004, Key Freshw. Invert. Russia 6 (molluscs, polychaetes, nemerteans): 41.
Kantor & Sysoev, 2005, Cat. Moll. Russia: 332.
Sayenko et al., 2009, Zool. Zhur. 88(11): 1306.
(error for Anodonta youkanensis Lea, 1867)

Anodonta youkonensisSimpson (1893)
Beringiana youkonensisBogatov & Starobogatov (2001)
Beringiana beringianaBieler (2021)

Anodonta yukonensis Simpson, 1896
‘Lea’ Simpson, 1896, Proc. U.S. Nat. Mus. 18: 328.
(error for Anodonta youkanensis Lea, 1867)

Anodonta yukonensisSimpson (1896)
Beringiana beringianaBieler (2021)

Arsenievinaia alimovi Bogatov & Zatravkin, 1988
Bogatov & Zatravkin, 1988, Trudy Zool. Inst. 187: 165, fig. 5.

Arsenievinaia alimoviBogatov & Zatravkin (1988)
Kunashiria coptzeviStarobogatov et al. (2004), Kantor & Sysoev (2005), Sayenko et al. (2009), Kantor et al. (2010)
Sinanodonta woodianaHe & Zhuang (2013)
Beringiana coptzeviVinarski & Kantor (2016)
Beringiana beringianaLopes-Lima et al. (2020), Bolotov et al. (2020)

Beringiana baringiana Bogatov & Zatravkin, 1988
‘Middendorff’ Bogatov & Zatravkin, 1988, Trudy Zool. Inst. 187: 167.
(error for Anodonta cellensis var. beringiana Middendorff, 1851)

Beringiana baringianaBogatov & Zatravkin (1988)

Beringiana yuokonensis Bogatov & Zatravkin, 1988
‘Lea’ Bogatov & Zatravkin, 1988, Trudy Zool. Inst. 187: 167.
(error for Anodonta youkanensis Lea, 1867)

Beringiana yuokonensisBogatov & Zatravkin (1988)

Arsenievinaia zatravkini Bogatov & Starobogatov, 1996
Bogatov & Starobogatov, 1996, Zool. Zhur. 75: 1331, fig’d.

Arsenievinaia zatravkiniBogatov & Starobogatov (1996)
Kunashiria taranetziStarobogatov et al. (2004)
Kunashiria taranetsiKantor & Sysoev (2005), Kantor et al. (2010)
Beringiana taranetziVinarski & Kantor (2016)
Beringiana beringianaLopes-Lima et al. (2020), Bolotov et al. (2020)

Amuranodonta sichotealinica Korniushin, 1998
‘Zatravkin & Starobogativ’ Korniushin, 1998, Malac. Review Bivalvia 7: 79.
(error for Amuranodonta sihotealinica Zatravkin & Starobogatov, 1984)

Amuranodonta sichotealinicaKorniushin (1998)

Anemina (Buldowskia) adotymensys Labay & Shulga, 1999
Labay & Shulga, 1999, Ruthenica 9: 79, fig’d.

Anemina (Buldowskia) adotymensysLabay & Shulga (1999)
Amuranodonta sitaensisStarobogatov et al. (2004), Kantor & Sysoev (2005), Kantor et al. (2010)
Anemina arcaeformisHe & Zhuang (2013)
Anemina (Buldowskia) sitaensisVinarski & Kantor (2016)
Beringiana beringianaLopes-Lima et al. (2020), Bolotov et al. (2020)

Anemina (Buldowskia) lacustris Labay & Shulga, 1999
Labay & Shulga, 1999, Ruthenica 9: 78, fig’d.

Anemina (Buldowskia) lacustrisLabay & Shulga (1999)
Amuranodonta parvaStarobogatov et al. (2004), Kantor & Sysoev (2005), Kantor et al. (2010)
Anemina arcaeformisHe & Zhuang (2013)
Anemina (Buldowskia) parvaVinarski & Kantor (2016)
Beringiana beringianaLopes-Lima et al. (2020), Bolotov et al. (2020)

Kunashiria japonica boreosakhalinensis Labay & Shulga, 1999
Labay & Shulga, 1999, Ruthenica 9: 77, fig’d.
(intro'd as intra-sp. of Anodonta japonica Clessin, 1874)

Kunashiria japonica boreosakhalinensisLabay & Shulga (1999)
Kunashiria haconensisStarobogatov et al. (2004), Kantor & Sysoev (2005), Kantor et al. (2010)
Beringiana haconensisVinarski & Kantor (2016)
Beringiana beringianaLopes-Lima et al. (2020), Bolotov et al. (2020)

Beringiana georgiensis Bogatov & Starobogatov, 2001
Bogatov & Starobogatov, 2001, Zool. Zhur. 80: 28, figs. 2ж-з, 4з.

Beringiana georgiensisBogatov & Starobogatov (2001)
Anodonta beringianaGraf (2007)

Beringiana georginesis Bogatov & Starobogatov, 2001
Bogatov & Starobogatov, 2001, Zool. Zhur. 80: 28.
(error for Beringiana georgiensis Bogatov & Starobogatov, 2001)

Beringiana georginesisBogatov & Starobogatov (2001)

Kunashiria taranetsi Kantor & Sysoev, 2005
‘Shadin’ Kantor & Sysoev, 2005, Cat. Moll. Russia: 339.
Kantor et al., 2010, Cat. Contin. Moll. Russia: 279.
(error for Anodonta beringiana var. taranetzi Zhadin, 1938)

Kunashiria taranetsiKantor & Sysoev (2005)

Beringiana georginensis Graf, 2007
‘Bogatov & Starobogatov’ Graf, 2007, Proc. Acad. Nat. Sci. 156: 86.
(error for Beringiana georgiensis Bogatov & Starobogatov, 2001)

Beringiana georginensisGraf (2007)

Taxonomic History (since 1900)

Anodonta beringianaSimpson (1900), Simpson (1914), Burch (1975), R.I. Johnson (1980), ..., Klishko et al. (2016), Froufe et al. (2017), Inoue et al. (2018), Zieritz et al. (2018)
Anodonta cygnea beringianaHannibal (1912)
Anodonta (s.s.) beringianaFrierson (1927), Zhadin (1938), Zhadin (1965), Haas (1969)
Anodonta beringiana var. taranetziZhadin (1938)
Anodonta (s.s.) beringiana var. taranetziZhadin (1938), Zhadin (1965)
Anodonta marginata beringianaModell (1945)
Anodonta (s.s.) oregonensisHaas (1969)
Anodonta (Sinanodonta) beringianaHabe (1977)
Beringiana beringianaZatravkin (1983), Zatravkin & Bogatov (1987), Korniushin (1998), Bogatov & Starobogatov (2001), ..., Lopes-Lima et al. (2020), Bolotov et al. (2020), Graf & Cummings (2021), Bieler (2021)
Beringiana youkonensisZatravkin (1983), Bogatov & Starobogatov (2001), Starobogatov et al. (2004), Sayenko et al. (2009)
Beringiana taranetziZatravkin (1983), Zatravkin & Bogatov (1987), Bogatov & Zatravkin (1988), Vinarski & Kantor (2016)
Amuranodonta sp. nov.Zatravkin (1983)
Amuranodonta sihotealinicaZatravkin & Starobogatov (1984)
Beringiana youkanensisZatravkin & Bogatov (1987)
Arsenievinaia sihotealinicaZatravkin & Bogatov (1987), Bogatov & Zatravkin (1988), Martynov & Chernyshev (1992), Bogatov & Sayenko (2002)
Arsenievinaia coptzeviZatravkin & Bogatov (1987), Bogatov & Zatravkin (1988)
Arsenievinaia ziminiZatravkin & Bogatov (1987), Bogatov & Zatravkin (1988)
Beringiana baringianaBogatov & Zatravkin (1988)
Beringiana yuokonensisBogatov & Zatravkin (1988)
Arsenievinaia alimoviBogatov & Zatravkin (1988)
Arsenievinaia zarjaensisBogatov & Zatravkin (1988)
Arsenievinaia compressaBogatov & Starobogatov (1996)
Arsenievinaia zatravkiniBogatov & Starobogatov (1996)
Beringiana compressaSayenko & Bogatov (1998), Bogatov & Starobogatov (2001), Starobogatov et al. (2004), Kantor & Sysoev (2005), Kantor et al. (2010), Vinarski & Kantor (2016)
Amuranodonta sichotealinicaKorniushin (1998)
Anemina (Buldowskia) adotymensysLabay & Shulga (1999)
Anemina (Buldowskia) lacustrisLabay & Shulga (1999)
Kunashiria iturupicaBogatov et al. (1999), Starobogatov et al. (2004), Kantor & Sysoev (2005), Sayenko et al. (2009), Kantor et al. (2010)
Kunashiria sinanodontoidesBogatov et al. (1999), Starobogatov et al. (2004), Kantor & Sysoev (2005), Sayenko et al. (2009), Kantor et al. (2010)
Kunashiria japonica boreosakhalinensisLabay & Shulga (1999)
Beringiana chereshneviBogatov & Starobogatov (2001), Starobogatov et al. (2004), Kantor & Sysoev (2005), Kantor et al. (2010), Vinarski & Kantor (2016)
Beringiana derzhaviniBogatov & Starobogatov (2001), Starobogatov et al. (2004), Kantor & Sysoev (2005), Kantor et al. (2010), Vinarski & Kantor (2016)
Beringiana georgiensisBogatov & Starobogatov (2001)
Beringiana georginesisBogatov & Starobogatov (2001)
Beringiana kamchaticaBogatov & Starobogatov (2001), Starobogatov et al. (2004), Kantor & Sysoev (2005), Sayenko et al. (2009), Kantor et al. (2010), Vinarski & Kantor (2016)
Kunashiria taranetziStarobogatov et al. (2004), Sayenko et al. (2009)
Kunashiria haconensisStarobogatov et al. (2004), Kantor & Sysoev (2005), Sayenko et al. (2009), Kantor et al. (2010)
Kunashiria coptzeviStarobogatov et al. (2004), Kantor & Sysoev (2005), Sayenko et al. (2009), Kantor et al. (2010)
Kunashiria ziminiStarobogatov et al. (2004), Kantor & Sysoev (2005), Sayenko et al. (2009), Kantor et al. (2010)
Kunashiria zarjaensisStarobogatov et al. (2004), Kantor & Sysoev (2005), Sayenko et al. (2009), Kantor et al. (2010)
Kunashiria compressaStarobogatov et al. (2004), Kantor & Sysoev (2005), Sayenko et al. (2009), Kantor et al. (2010)
Amuranodonta sitaensisStarobogatov et al. (2004), Kantor & Sysoev (2005), Kantor et al. (2010)
Amuranodonta parvaStarobogatov et al. (2004), Kantor & Sysoev (2005), Kantor et al. (2010)
Beringiana youconensisKantor & Sysoev (2005), Kantor et al. (2010), Vinarski & Kantor (2016)
Kunashiria taranetsiKantor & Sysoev (2005), Kantor et al. (2010)
Anodonta woodiana japonicaGraf (2007)
Beringiana georginensisGraf (2007)
Sinanodonta woodianaHe & Zhuang (2013)
Anemina arcaeformisHe & Zhuang (2013)
Beringiana haconensisVinarski & Kantor (2016)
Beringiana coptzeviVinarski & Kantor (2016)
Beringiana ziminiVinarski & Kantor (2016)
Beringiana zarjaensisVinarski & Kantor (2016)
Anemina (Buldowskia) sitaensisVinarski & Kantor (2016)
Anemina (Buldowskia) parvaVinarski & Kantor (2016)
Beringiana iturupicaVinarski & Kantor (2016)
Beringiana sinanodontoidesVinarski & Kantor (2016)
Sinanodonta beringianaWilliams et al. (2017)

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