MUSSELpdb | valid sp. Beringiana beringiana synonymy

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Page last updated
Tue 01 Jul 2025

The Freshwater Mussels (Unionoida) of the World (and other less consequential bivalves)

FM(U)otW(aolcb) is the web version of the MUSSEL Project Database. Follow the links to browse the data or use the custom Google search field. Either way, you win!

The nominal species (available and otherwise) in this synonymy are listed by usage rather than strictly by publication date. The primary sorting order is by the most recent year (since 1969) that the species name was treated as valid, and then nominal species are sorted secondarily by publication year.

The taxonomic history of the valid species (since 1900) is listed at the bottom of the page.

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species Beringiana beringiana (Middendorff, 1851): nominal species
1 specimen available.

Anodonta cellensis var. beringiana Middendorff, 1851
Middendorff, 1851, Reise Nord. Ost. Sibiriens 2(1): 284, pl. 28, figs. 4-7; pl. 29, figs. 1-4.
(intro'd as intra-sp. of Anodonta cellensis Pfeiffer, 1821)

Anodonta cellensis var. beringiana — Middendorff (1851)
Margaron (Anodonta) cygnea — Lea (1870)
Anodonta beringiana — Simpson (1900), Simpson (1914), Burch (1975), R.I. Johnson (1980), ..., Froufe et al. (2017), Inoue et al. (2018), Zieritz et al. (2018), Tomilova et al. (2019)
Anodonta cygnea beringiana — Hannibal (1912)
Anodonta (s.s.) beringiana — Frierson (1927), Zhadin (1938), Zhadin (1965), Haas (1969)
Anodonta marginata beringiana — Modell (1945)
Anodonta (Sinanodonta) beringiana — Habe (1977)
Beringiana beringiana — Zatravkin (1983), Zatravkin & Bogatov (1987), Korniushin (1998), Bogatov & Starobogatov (2001), ..., Dai et al. (2024), Xiang et al. (2025), Dai et al. (2025), Wu et al. (2025)
Beringiana baringiana — Bogatov & Zatravkin (1988)
Sinanodonta beringiana — Williams et al. (2017), FMCS (2021)
Buldowskia beringiana beringiana — Bogatov (2022)

Anodonta youkanensis Lea, 1867
Lea, 1867, Proc. Acad. Nat. Sci. 19: 81 [as ‘youconensis’].
Lea, 1868, Jour. Acad. Nat. Sci. 6 [O. 12]: 287 [47], pl. 40, fig. 99.
(correct name for Anodonta youconensis Lea, 1867)
Type(s): FIGURED SYNTYPE USNM_86624, Upper Yukon R., Alaska.

Anodonta youkanensis — Lea (1867), B.H. Wright (1888)
Margaron (Anodonta) youkanensis — Lea (1870)
Anodonta beringiana — Simpson (1900), Simpson (1914), Burch (1975), Graf (2007), IUCN (2011)
Anodonta cygnea beringiana — Hannibal (1912)
Anodonta (s.s.) beringiana — Frierson (1927)
Anodonta (s.s.) oregonensis — Haas (1969)
Beringiana youkonensis — Zatravkin (1983), Bogatov & Starobogatov (2001), Starobogatov et al. (2004), Sayenko et al. (2009)
Beringiana youkanensis — Zatravkin & Bogatov (1987)
Beringiana yuokonensis — Bogatov & Zatravkin (1988)
Beringiana youconensis — Kantor & Sysoev (2005), Kantor et al. (2010), Vinarski & Kantor (2016)
Beringiana beringiana — Lopes-Lima et al. (2020), Bolotov et al. (2020), Bieler (2021)
Buldowskia beringiana youkonensis — Bogatov (2022)

Anodonta beringiana var. taranetzi Zhadin, 1938
Zhadin, 1938, Faune USSR Mollusques 4(1): 133, fig. 49.
Zhadin, 1965, Moll. USSR: 255, fig. 273.
(intro'd as intra-sp. of Anodonta cellensis var. beringiana Middendorff, 1851)

Anodonta beringiana var. taranetzi — Zhadin (1938)
Anodonta (s.s.) beringiana var. taranetzi — Zhadin (1938), Zhadin (1965)
Anodonta (s.s.) beringiana — Haas (1969)
Beringiana taranetzi — Zatravkin (1983), Zatravkin & Bogatov (1987), Bogatov & Zatravkin (1988), Vinarski & Kantor (2016)
Kunashiria taranetzi — Starobogatov et al. (2004), Sayenko et al. (2009)
Kunashiria taranetsi — Kantor & Sysoev (2005), Kantor et al. (2010)
Anodonta woodiana japonica — Graf (2007)
Sinanodonta woodiana — He & Zhuang (2013)
Beringiana beringiana — Lopes-Lima et al. (2020), Bolotov et al. (2020)
Kunashiria japonica taranetzi — Bogatov (2022)

Arsenievinaia compressa Bogatov & Starobogatov, 1996
Bogatov & Starobogatov, 1996, Zool. Zhur. 75: 1332. fig’d.

Arsenievinaia compressa — Bogatov & Starobogatov (1996)
Kunashiria compressa — Starobogatov et al. (2004), Kantor & Sysoev (2005), Sayenko et al. (2009), Kantor et al. (2010)
Anodonta woodiana japonica — Graf (2007)
Sinanodonta woodiana — He & Zhuang (2013)
Beringiana compressa — Vinarski & Kantor (2016)
Beringiana beringiana — Lopes-Lima et al. (2020), Bolotov et al. (2020)
Kunashiria japonica f. compressa — Bogatov (2022)

Beringiana compressa Sayenko & Bogatov, 1998
Sayenko & Bogatov, 1998, Zool. Zhur. 77(12): 1414, figs. 2-4.
(homonym of Arsenievinaia compressa Bogatov & Starobogatov, 1996)

Beringiana compressa — Sayenko & Bogatov (1998), Bogatov & Starobogatov (2001), Starobogatov et al. (2004), Kantor & Sysoev (2005), Kantor et al. (2010), Vinarski & Kantor (2016)
Anodonta beringiana — Graf (2007)
Beringiana beringiana — Lopes-Lima et al. (2020), Bolotov et al. (2020), Bulakhova et al. (2023)
Buldowskia beringiana f. compressa — Bogatov (2022)

Beringiana kamchatica Bogatov & Starobogatov, 2001
Bogatov & Starobogatov, 2001, Zool. Zhur. 80: 27, figs. 2д-е, 4г.

Beringiana kamchatica — Bogatov & Starobogatov (2001), Starobogatov et al. (2004), Kantor & Sysoev (2005), Sayenko et al. (2009), Kantor et al. (2010), Vinarski & Kantor (2016)
Anodonta beringiana — Graf (2007)
Beringiana beringiana — Lopes-Lima et al. (2020), Bolotov et al. (2020), Bulakhova et al. (2023)
Buldowskia beringiana kamchatica — Bogatov (2022)

Anodonta youconensis Lea, 1867
Lea, 1867, Proc. Acad. Nat. Sci. 19: 81.
(incorrect original spelling of Anodonta youkanensis Lea, 1867)

Anodonta youconensis — Lea (1867)
Anodonta beringiana — Simpson (1900), Simpson (1914)
Anodonta cygnea beringiana — Hannibal (1912)
Anodonta (s.s.) beringiana — Frierson (1927)
Anodonta (s.s.) oregonensis — Haas (1969)
Beringiana youkanensis — Zatravkin & Bogatov (1987)
Beringiana youconensis — Kantor & Sysoev (2005), Kantor et al. (2010), Vinarski & Kantor (2016)
Beringiana beringiana — Bieler (2021)

Arsenievinaia coptzevi Zatravkin & Bogatov, 1987
Zatravkin & Bogatov, 1987, Biv. Far East: 104, sp. 51, figs. 20 б, 24 в-г.

Arsenievinaia coptzevi — Zatravkin & Bogatov (1987), Bogatov & Zatravkin (1988)
Kunashiria coptzevi — Starobogatov et al. (2004), Kantor & Sysoev (2005), Sayenko et al. (2009), Kantor et al. (2010)
Anodonta woodiana japonica — Graf (2007)
Sinanodonta woodiana — He & Zhuang (2013)
Beringiana coptzevi — Vinarski & Kantor (2016)
Beringiana beringiana — Lopes-Lima et al. (2020), Bolotov et al. (2020)
Kunashiria japonica haconensis — Bogatov (2022)

Arsenievinaia zimini Zatravkin & Bogatov, 1987
Zatravkin & Bogatov, 1987, Biv. Far East: 102, sp. 50, figs. 24 а-б.

Arsenievinaia zimini — Zatravkin & Bogatov (1987), Bogatov & Zatravkin (1988)
Arsenievinaia sihotealinica — Martynov & Chernyshev (1992)
Kunashiria zimini — Starobogatov et al. (2004), Kantor & Sysoev (2005), Sayenko et al. (2009), Kantor et al. (2010)
Anodonta woodiana japonica — Graf (2007)
Sinanodonta woodiana — He & Zhuang (2013)
Beringiana zimini — Vinarski & Kantor (2016)
Beringiana beringiana — Lopes-Lima et al. (2020), Bolotov et al. (2020)
Kunashiria japonica haconensis — Bogatov (2022)

Arsenievinaia zarjaensis Bogatov & Zatravkin, 1988
Bogatov & Zatravkin, 1988, Trudy Zool. Inst. 187: 166, fig. 6.

Arsenievinaia zarjaensis — Bogatov & Zatravkin (1988)
Kunashiria zarjaensis — Starobogatov et al. (2004), Kantor & Sysoev (2005), Sayenko et al. (2009), Kantor et al. (2010)
Anodonta woodiana japonica — Graf (2007)
Sinanodonta woodiana — He & Zhuang (2013)
Beringiana zarjaensis — Vinarski & Kantor (2016)
Beringiana beringiana — Lopes-Lima et al. (2020), Bolotov et al. (2020)
Kunashiria japonica japonica — Bogatov (2022)

Kunashiria iturupica Bogatov et al., 1999
Bogatov et al., 1999, Ruthenica 9: 60, fig’d.

Kunashiria iturupica — Bogatov et al. (1999), Starobogatov et al. (2004), Kantor & Sysoev (2005), Sayenko et al. (2009), Kantor et al. (2010)
Anodonta woodiana japonica — Graf (2007)
Sinanodonta woodiana — He & Zhuang (2013)
Beringiana iturupica — Vinarski & Kantor (2016)
Beringiana beringiana — Lopes-Lima et al. (2020), Bolotov et al. (2020)
Kunashiria japonica taranetzi — Bogatov (2022)

Kunashiria sinanodontoides Bogatov et al., 1999
Bogatov et al., 1999, Ruthenica 9: 59, fig’d.

Kunashiria sinanodontoides — Bogatov et al. (1999), Starobogatov et al. (2004), Kantor & Sysoev (2005), Sayenko et al. (2009), Kantor et al. (2010)
Anodonta woodiana japonica — Graf (2007)
Sinanodonta woodiana — He & Zhuang (2013)
Beringiana sinanodontoides — Vinarski & Kantor (2016)
Beringiana beringiana — Lopes-Lima et al. (2020), Bolotov et al. (2020)
Kunashiria japonica taranetzi — Bogatov (2022)

Beringiana chereshnevi Bogatov & Starobogatov, 2001
Bogatov & Starobogatov, 2001, Zool. Zhur. 80: 26, figs. 2а-б, 4ж.

Beringiana chereshnevi — Bogatov & Starobogatov (2001), Starobogatov et al. (2004), Kantor & Sysoev (2005), Kantor et al. (2010), Vinarski & Kantor (2016)
Anodonta beringiana — Graf (2007)
Beringiana beringiana — Lopes-Lima et al. (2020), Bolotov et al. (2020)
Buldowskia beringiana youkonensis — Bogatov (2022)

Beringiana derzhavini Bogatov & Starobogatov, 2001
Bogatov & Starobogatov, 2001, Zool. Zhur. 80: 27, figs. 2в-г, 4а.

Beringiana derzhavini — Bogatov & Starobogatov (2001), Starobogatov et al. (2004), Kantor & Sysoev (2005), Kantor et al. (2010), Vinarski & Kantor (2016)
Anodonta beringiana — Graf (2007)
Beringiana beringiana — Lopes-Lima et al. (2020), Bolotov et al. (2020), Bulakhova et al. (2023)
Buldowskia beringiana kamchatica — Bogatov (2022)

Amuranodonta sihotealinica Zatravkin & Starobogatov, 1984
Zatravkin & Starobogatov, 1984, Zool. Zhur. 63(12): 1788, figs. 9-10.

Amuranodonta sihotealinica — Zatravkin & Starobogatov (1984)
Amuranodonta sp. nov. — Zatravkin (1983)
Arsenievinaia sihotealinica — Zatravkin & Bogatov (1987), Bogatov & Zatravkin (1988), Martynov & Chernyshev (1992), Bogatov & Sayenko (2002)
Kunashiria haconensis — Starobogatov et al. (2004), Kantor & Sysoev (2005), Sayenko et al. (2009), Kantor et al. (2010)
Sinanodonta woodiana — He & Zhuang (2013)
Beringiana haconensis — Vinarski & Kantor (2016)
Beringiana beringiana — Lopes-Lima et al. (2020), Bolotov et al. (2020)
Kunashiria japonica haconensis — Bogatov (2022)

Anodonta youcanensis Lea, 1867
‘Lea’ Lea, 1867, Obs., index I-XI: 63.
‘Lea’ Paetel, 1890, Cat. Conch.-Samml. 3: 186.
(error for Anodonta youkanensis Lea, 1867)

Anodonta youcanensis — Lea (1867)
Anodonta beringiana — Simpson (1900), Simpson (1914)
Beringiana beringiana — Bieler (2021)

Anodon youkamensis Sowerby, 1870
‘Lea’ Sowerby, 1870, Conch. Icon. 17: index.
(error for Anodonta youkanensis Lea, 1867)

Anodon youkamensis — Sowerby (1870)
Beringiana beringiana — Bieler (2021)

Anodonta youkonensis Simpson, 1893
‘Lea’ of Authors.
Simpson, 1893, Nautilus 6(12): 135.
Zatravkin, 1983, [in] I.M. Likharev, Molluscs: 42.
Bogatov & Starobogatov, 2001, Zool. Zhur. 80: 30.
Starobogatov et al., 2004, Key Freshw. Invert. Russia 6 (molluscs, polychaetes, nemerteans): 41.
Kantor & Sysoev, 2005, Cat. Moll. Russia: 332.
Sayenko et al., 2009, Zool. Zhur. 88(11): 1306.
Bogatov, 2022, Lg. Bivalve Moll. Russia’s Fresh Waters: 73.
(error for Anodonta youkanensis Lea, 1867)

Anodonta youkonensis — Simpson (1893)
Beringiana youkonensis — Bogatov & Starobogatov (2001)
Beringiana beringiana — Bieler (2021), Bulakhova et al. (2023)

Anodonta yukonensis Simpson, 1896
‘Lea’ Simpson, 1896, Proc. U.S. Nat. Mus. 18: 328.
(error for Anodonta youkanensis Lea, 1867)

Anodonta yukonensis — Simpson (1896)
Beringiana beringiana — Bieler (2021)

Arsenievinaia alimovi Bogatov & Zatravkin, 1988
Bogatov & Zatravkin, 1988, Trudy Zool. Inst. 187: 165, fig. 5.

Arsenievinaia alimovi — Bogatov & Zatravkin (1988)
Kunashiria coptzevi — Starobogatov et al. (2004), Kantor & Sysoev (2005), Sayenko et al. (2009), Kantor et al. (2010)
Sinanodonta woodiana — He & Zhuang (2013)
Beringiana coptzevi — Vinarski & Kantor (2016)
Beringiana beringiana — Lopes-Lima et al. (2020), Bolotov et al. (2020)
Kunashiria japonica taranetzi — Bogatov (2022)

Beringiana baringiana Bogatov & Zatravkin, 1988
‘Middendorff’ Bogatov & Zatravkin, 1988, Trudy Zool. Inst. 187: 167.
(error for Anodonta cellensis var. beringiana Middendorff, 1851)

Beringiana baringiana — Bogatov & Zatravkin (1988)

Beringiana yuokonensis Bogatov & Zatravkin, 1988
‘Lea’ Bogatov & Zatravkin, 1988, Trudy Zool. Inst. 187: 167.
(error for Anodonta youkanensis Lea, 1867)

Beringiana yuokonensis — Bogatov & Zatravkin (1988)

Arsenievinaia zatravkini Bogatov & Starobogatov, 1996
Bogatov & Starobogatov, 1996, Zool. Zhur. 75: 1331, fig’d.

Arsenievinaia zatravkini — Bogatov & Starobogatov (1996)
Kunashiria taranetzi — Starobogatov et al. (2004)
Kunashiria taranetsi — Kantor & Sysoev (2005), Kantor et al. (2010)
Beringiana taranetzi — Vinarski & Kantor (2016)
Beringiana beringiana — Lopes-Lima et al. (2020), Bolotov et al. (2020)
Kunashiria japonica taranetzi — Bogatov (2022)

Amuranodonta sichotealinica Korniushin, 1998
‘Zatravkin & Starobogativ’ Korniushin, 1998, Malac. Review Bivalvia 7: 79.
(error for Amuranodonta sihotealinica Zatravkin & Starobogatov, 1984)

Amuranodonta sichotealinica — Korniushin (1998)

Anemina (Buldowskia) adotymensys Labay & Shulga, 1999
Labay & Shulga, 1999, Ruthenica 9: 79, fig’d.

Anemina (Buldowskia) adotymensys — Labay & Shulga (1999)
Amuranodonta sitaensis — Starobogatov et al. (2004), Kantor & Sysoev (2005), Kantor et al. (2010)
Anemina arcaeformis — He & Zhuang (2013)
Anemina (Buldowskia) sitaensis — Vinarski & Kantor (2016)
Beringiana beringiana — Lopes-Lima et al. (2020), Bolotov et al. (2020)

Anemina (Buldowskia) lacustris Labay & Shulga, 1999
Labay & Shulga, 1999, Ruthenica 9: 78, fig’d.

Anemina (Buldowskia) lacustris — Labay & Shulga (1999)
Amuranodonta parva — Starobogatov et al. (2004), Kantor & Sysoev (2005), Kantor et al. (2010)
Anemina arcaeformis — He & Zhuang (2013)
Anemina (Buldowskia) parva — Vinarski & Kantor (2016)
Beringiana beringiana — Lopes-Lima et al. (2020), Bolotov et al. (2020)

Kunashiria japonica boreosakhalinensis Labay & Shulga, 1999
Labay & Shulga, 1999, Ruthenica 9: 77, fig’d.
(intro'd as intra-sp. of Anodonta japonica Clessin, 1874)

Kunashiria japonica boreosakhalinensis — Labay & Shulga (1999)
Kunashiria haconensis — Starobogatov et al. (2004), Kantor & Sysoev (2005), Kantor et al. (2010)
Beringiana haconensis — Vinarski & Kantor (2016)
Beringiana beringiana — Lopes-Lima et al. (2020), Bolotov et al. (2020)
Kunashiria japonica japonica — Bogatov (2022)

Beringiana georgiensis Bogatov & Starobogatov, 2001
Bogatov & Starobogatov, 2001, Zool. Zhur. 80: 28, figs. 2ж-з, 4з.

Beringiana georgiensis — Bogatov & Starobogatov (2001)
Anodonta beringiana — Graf (2007)
Buldowskia beringiana youkonensis — Bogatov (2022)

Beringiana georginesis Bogatov & Starobogatov, 2001
Bogatov & Starobogatov, 2001, Zool. Zhur. 80: 28.
(error for Beringiana georgiensis Bogatov & Starobogatov, 2001)

Beringiana georginesis — Bogatov & Starobogatov (2001)

Kunashiria taranetsi Kantor & Sysoev, 2005
‘Shadin’ Kantor & Sysoev, 2005, Cat. Moll. Russia: 339.
Kantor et al., 2010, Cat. Contin. Moll. Russia: 279.
(error for Anodonta beringiana var. taranetzi Zhadin, 1938)

Kunashiria taranetsi — Kantor & Sysoev (2005)

Beringiana georginensis Graf, 2007
‘Bogatov & Starobogatov’ Graf, 2007, Proc. Acad. Nat. Sci. 156: 86.
(error for Beringiana georgiensis Bogatov & Starobogatov, 2001)

Beringiana georginensis — Graf (2007)

Sinanodonta japonica Sano et al., 2017
‘Clessin’ of Authors.
Sano et al., 2017, J. Wat. Res. Prot. 9: Table 2. [in part]
Sano et al., 2020, Plankton Benthos Res. 15(2): Table 1. [in part]
(misidentified reference to Anodonta japonica Clessin, 1874)

Sinanodonta japonica — Sano et al. (2017)
Beringiana beringiana — Sano et al. (2022), Bulakhova et al. (2023)

Page updated: 10:00:54 Tue 01 Jul 2025

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