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Tue 01 Jul 2025 |
The Freshwater Mussels (Unionoida) of the World (and other less consequential bivalves)
FM(U)otW(aolcb) is the web version of the MUSSEL Project Database. Follow the links to browse the data or use the custom Google search field. Either way, you win!
This page documents data for Lopes-Lima et al. (2020), such as genera and species attributed to that publication (including known misspellings, misidentifications, and other unavailable names) as well as taxonomic opinions, if available. Taxonomic opinions refer to uses of genera and species as valid and invalid in that work.
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Publication information
Lopes-Lima, M., A. Hattori, T. Kondo, J.H. Lee, S.K. Kim et al. 2020. Freshwater mussels (Bivalvia: Unionidae) from the rising sun (Far East Asia): phylogeny, systematics, and distribution. Molecular Phylogenetics and Evolution 146: 1-27.
Click here for online access.
Cladogram Figures
(click the figure number to see the cladogram) fig. 10
fig. 11
fig. 12
fig. 13
fig. 14
fig. 16
fig. 17
fig. 2-4
fig. 5
fig. 6
fig. 7
fig. 8
fig. 9
Attributed Genus Names
Koreosolenaia Lopes-Lima et al., 2020
Lee, Kim, Lopes-Lima & Bogan in Lopes-Lima et al., 2020, MPE 146: [15, 23].
Type species: Koreosolenaia sitgyensis Lopes-Lima et al., 2020 (orig. desig.) = Koreosolenaia sitgyensis |
Apppius Lopes-Lima et al., 2020
‘Menke’ Lopes-Lima et al., 2020, MPE 146: Appendix 1.
(error for Appius Menke, 1830)
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Nipponihyra Lopes-Lima et al., 2020
‘Starobogatov’ Lopes-Lima et al., 2020, MPE 146: Appendix 1.
(error for Nipponihyria Starobogatov, 1970)
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Attributed Species Names
Anodon randalii Conrad, 1857
‘Trask’ Conrad, 1857, Proc. Acad. Nat. Sci. 9: 165.
‘Trask’ Lopes-Lima et al., 2020, MPE 146: Appendix 1.
(error for Anodonta randalli Trask, 1854)
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Lasmigona compressa Graf & Ó Foighil, 2000
‘Lea’ of Authors.
Graf & Ó Foighil, 2000, J. Moll. Studies 66: 160.
Graf & Cummings, 2006, Zool. J. Linn. Soc. 148: 348.
Hoeh et al., 2009, Malacologia 51(2): 310.
Jadhav & Jamkhedkar, 2009, Res. J. BioTech. 4(1): 43.
Whelan et al., 2011, MPE 61: 506.
Lopes-Lima et al., 2017, MPE 106: 177.
Bolotov et al., 2017, Sci. Repts. 7: Table S1.
Bolotov et al., 2017, Sci. Repts. 7: Table S1.
Bolotov et al., 2018, Sci. Repts. 8: Table S1.
Huang et al., 2019, MPE 130: Table S4.
Lopes-Lima et al., 2020, MPE 146: Table 1.
Wu et al., 2021, J. Lake Sci. 33(6): attached Table 1.
Jeratthitikul et al., 2022, Sci. Repts. 12: Table S1.
Wu et al., 2023, ZooKeys 1166: 263, Table 1.
Liu et al., 2023, ZooKeys 1179: Table 1.
Dai et al., 2025, Zoosyst. Evol. 101(2): Table 1. [in part]
(misidentified reference to Symphynota compressa Lea, 1829)
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Middendorffinaia suifunensis Graf, 2007
‘Moskvicheva & Starobogatov’ of Authors.
Graf, 2007, Proc. Acad. Nat. Sci. 156: 85.
Lopes-Lima et al., 2020, MPE 146: Appendix 1.
Bolotov et al., 2020, Sci. Repts. 10: S32.
Bogatov, 2022, Lg. Bivalve Moll. Russia’s Fresh Waters: 59, 61.
(error for Middendorffinaia (Suifununio) sujfunensis Moskvicheva & Starobogatov, 1973)
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Lamellidens corrianus Whelan et al., 2011
‘Lea’ of Authors.
Whelan et al., 2011, MPE 61: 506.
Pfeiffer & Graf, 2015, Zool. J. Linn. Soc. 175(2): 310.
Bolotov et al., 2017, Sci. Repts. 7: Table S1 [as ‘Lamellidens corrianus sp. 1].
Konopleva et al., 2017, Systemat. Biodiv. 15(3): 207.
Pfeiffer et al., 2018, J. Moll. Studies 84(4): [3].
Pfeiffer et al., 2019, MPE 137: 117.
Lopes-Lima et al., 2020, MPE 146: Table 1.
(misidentified reference to Unio corrianus Lea, 1834)
Type(s): SEQUENCED SPECIMEN UMMZ_304642, Ayeryarwaddy River, mud bank on E side of river, Mandalay Division, Burma. 21.48187, 95.30501.
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Pseudodon inoscularis Pfeiffer & Graf, 2013
‘Gould’ of Authors.
Pfeiffer & Graf, 2013, J. Moll. Studies 79: 251.
Pfeiffer & Graf, 2015, Zool. J. Linn. Soc. 175(2): 310.
Konopleva et al., 2017, Systemat. Biodiv. 15(3): 207.
Bolotov et al., 2017, Sci. Repts. 7: Table S1 [as ‘Pseudodon cf. inoscularis’].
Bolotov et al., 2017, Sci. Repts. 7: Table S1 [as ‘ “Monodontina” cf. inoscularis sp. 2’].
Huang et al., 2019, MPE 130: Table S4 [as ‘ “Monodontina” cf. inoscularis sp. 2’].
Lopes-Lima et al., 2020, MPE 146: Table 1 [as ‘ “Monodontina” cf. inoscularis sp. 2’].
(misidentified reference to Anodon inoscularis Gould, 1844)
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Pilsbryoconcha exilis Pfeiffer & Graf, 2015
‘Lea’ of Authors.
Pfeiffer & Graf, 2015, Zool. J. Linn. Soc. 175(2): 310.
Ng et al., 2016, PLoS One 11(8), e0161130: [5]. [in part]
Bolotov et al., 2017, Sci. Repts. 7: Table S1 [as ‘P. aff. exilis sp. 1’].
Bolotov et al., 2017, Sci. Repts. 7: Table S1 [as ‘P. aff. exilis sp. 1’].
Konopleva et al., 2017, Systemat. Biodiv. 15(3): 207.
Bolotov et al., 2018, Sci. Repts. 8: Table S1 [as ‘P. aff. exilis sp. 1’].
Huang et al., 2019, MPE 130: Table S4 [as ‘P. aff. exilis sp. 1’].
Konopleva et al., 2019, Sci. Repts. 9: Table S1 [as ‘P. aff. exilis sp. 1’].
Lopes-Lima et al., 2020, MPE 146: Table 1 [as ‘P. aff. exilis sp. 1’].
Zieritz et al., 2020, Biodivers. Conserv. 29: 2244.
Wu et al., 2021, ZooKeys 1054: 87.
Wu et al., 2021, J. Lake Sci. 33(6): attached Table 1.
Wu et al., 2021, J. Lake Sci. 33(6): attached Table 1.
Konopleva et al., 2023, Diversity 15(1), 10: Table S2 [as ‘P. aff. exilis’].
Dai et al., 2023, Invert. Syst. 37(2): Tables 1-2. [in part]
(misidentified reference to Anodonta exilis Lea, 1838)
Type(s): SEQUENCED SPECIMEN UMMZ_304647, Beung Dang Teuk Leach Community Fishery Pursat Province, Cambodia.
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Ensidens sagittarius Pfeiffer & Graf, 2015
‘Lea’ of Authors.
Pfeiffer & Graf, 2015, Zool. J. Linn. Soc. 175(2): 310.
Konopleva et al., 2017, Systemat. Biodiv. 15(3): 207.
Lopes-Lima et al., 2017, MPE 106: 177.
Bolotov et al., 2017, Sci. Repts. 7: Table S1 [as ‘Ensidens aff. sagittarius sp. 1].
Bolotov et al., 2017, Sci. Repts. 7: Table S1 [as ‘Ensidens aff. sagittarius sp. 1].
Bolotov et al., 2018, Sci. Repts. 8: Table S1 [as ‘Ensidens aff. sagittarius sp. 1].
Konopleva et al., 2019, Sci. Repts. 9: Table S1 [as ‘E. aff. sagittarius sp. 1].
Huang et al., 2019, MPE 130: Table S4 [as ‘Ensidens aff. sagittarius sp. 1].
Zieritz et al., 2020, Biodivers. Conserv. 29: 2242.
Lopes-Lima et al., 2020, MPE 146: Table 1.
Wu et al., 2021, J. Lake Sci. 33(6): attached Table 1.
(misidentified reference to Unio sagittarius Lea, 1856)
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Pseudodon cumingii Pfeiffer & Graf, 2015
‘Lea’ of Authors.
Pfeiffer & Graf, 2015, Zool. J. Linn. Soc. 175(2): 310.
Lopes-Lima et al., 2017, MPE 106: 177.
Bolotov et al., 2017, Sci. Repts. 7: Table S1.
Konopleva et al., 2017, Systemat. Biodiv. 15(3): 207.
Bolotov et al., 2017, Sci. Repts. 7: Table S1 [as ‘”Bineurus” cumingii’].
Huang et al., 2019, MPE 130: Table S4 [as ‘”Bineurus” cumingii’].
Lopes-Lima et al., 2020, MPE 146: Table 1.
(misidentified reference to Anodonta cumingii Lea, 1851)
Type(s): SPECIMEN UMMZ_304648, Nam Phiat, Phonethong, ca. 2 km from confluence, Bolikhamxai Province, Laos. 18.08389, 104.97806.
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Chamberlainia hainesiana Kongim et al., 2015
‘Lea’ of Authors.
Kongim et al., 2015, ZooKeys 514: 98, fig. 2A.
Lopes-Lima et al., 2017, MPE 106: 177.
Bolotov et al., 2017, Sci. Repts. 7: Table S1.
Bolotov et al., 2017, Sci. Repts. 7: Table S1.
Froufe et al., 2019, Heredity 124: 185.
Huang et al., 2019, MPE 130: Table S4.
Chen et al., 2020, mtDNA B 5(3): 2800.
Teiga-Teixeira et al., 2020, mtDNA B 5(3): 2978.
Zieritz et al., 2020, Biodivers. Conserv. 29: 2244.
Bolotov et al., 2020, Sci. Repts. 10: Table S1.
Lopes-Lima et al., 2020, MPE 146: Table 1.
Zieritz et al., 2021, Zool. J. Linn. Soc. 193(3): Table 1.
Zieritz et al., 2021, Aquatic Conserv.: Mar. Freshw. Ecosyst. 31: 3176.
Jeratthitikul et al., 2021, Eur. J. Taxonomy 775: 125.
Wu et al., 2021, ZooKeys 1054: 87.
Wu et al., 2021, J. Lake Sci. 33(6): attached Table 1.
Wu et al., 2022, Zool. Scripta 51: Table S2.
Pavan-Kumar et al., 2022, Mol. Biol. Rep. 49: Table S1.
Jeratthitikul et al., 2022, Sci. Repts. 12: Table S1.
Dai et al., 2023, Invert. Syst. 37(2): Tables 1-2.
Konopleva et al., 2023, Zoosyst. Evol. 99(1): Table S1.
Wu et al., 2023, Org Divers Evol (2023): Table 2.
Dai et al., 2024, Zool. Scripta 53: Table 1.
Dai et al., 2024, Invert. Syst. 38(6): Table 3.
(misidentified reference to Unio hainesianus Lea, 1856)
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Sinanodonta woodiana Ng et al., 2016
‘Lea’ of Authors.
Ng et al., 2016, PLoS One 11(8), e0161130: Table S2.
Bolotov et al., 2016, Biochem. Syst. & Ecol. 67: 60 [as ‘lineage A’ & ‘lineage B’]. [in part]
Zieritz et al., 2016, Sci. Total Environ. 571: 1071.
Vikhrev et al., 2017, Trop. Conserv. Sci. 10: Table 1 [as ‘lineage A’ & ‘lineage B’]. [in part]
Bespalaya et al., 2018, Limnologica 69: Table 1 [as ‘lineage A’ & ‘lineage B’]. [in part]
Konečný et al., 2018, Evol. Appl. 11: Table S1 [as ‘lineage A’ & ‘lineage B’]. [in part]
Zieritz et al., 2018, Biol. Conserv. 219: 131.
Kondakov et al., 2018, Zootaxa 4462(4): 517, Table 1.
Lopes-Lima et al., 2020, MPE 146: Table S7 [as ‘Sinanodonta cf. woodiana 1’]. [in part]
Bauer et al., 2021, Neotrop. Natur. 5: 1, figs. 1-2.
Sano et al., 2022, MPE 175: Table S1 [as ‘S. woodiana sp. 1’]. [in part]
Bolotov et al., 2022, Aquatic Conserv.: Mar. Freshw. Ecosyst. 32(9): 1498, Table S3. [in part]
Bogan et al., 2023, Diversity 15(6), 710: Tables 1, S3 [as ‘Sinanodonta cf. woodiana 1’]. [in part]
(misidentified reference to Symphynota woodiana Lea, 1834)
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Cuneopsis heudei Lopes-Lima et al., 2017
‘Heude’ of Authors.
Liu et al., 2017, PLoS One 12(12 ), e0189737: Table S4.
Lopes-Lima et al., 2017, MPE 106: Table 2.
Bolotov et al., 2017, Sci. Repts. 7: Table S1.
Bolotov et al., 2017, Sci. Repts. 7: Table S1.
Wu et al., 2018, PLoS One 13(8), e0200956: 5.
Huang et al., 2019, MPE 130: Tables S1, S4. [in part]
Lopes-Lima et al., 2020, MPE 146: Table 1. [in part]
Wu et al., 2021, J. Lake Sci. 33(6): attached Table 1. [in part]
Wu et al., 2022, Zool. Scripta 51: Table S1. [in part]
Jeratthitikul et al., 2022, Sci. Repts. 12: Table S1.
Konopleva et al., 2023, Zoosyst. Evol. 99(1): Table S1.
Chen et al., 2023, ZooKeys 1173: Table 1.
Wu et al., 2023, ZooKeys 1166: Table 1.
Liu et al., 2023, ZooKeys 1179: Table 1.
(misidentified reference to Unio heudei Heude, 1874)
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Hyriopsis bialata Lopes-Lima et al., 2017
‘Simpson’ of Authors.
Lopes-Lima et al., 2017, MPE 106: 177.
Muanta et al., 2019, J. Moll. Studies 85(2): 225.
Lopes-Lima et al., 2020, MPE 146: Table 1.
(misidentified reference to Hyriopsis bialatus Simpson, 1900)
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Pseudodon vondembuschianus Bolotov et al., 2017
‘Lea’ of Authors.
Bolotov et al., 2017, Sci. Repts. 7: Table S1 [as ‘Pseudodon aff. vondembuschianus sp. 1’].
Bolotov et al., 2017, Sci. Repts. 7: Table S1 [as ‘Monodontina aff. vondembuschiana sp. 1’].
Bolotov et al., 2018, Sci. Repts. 8: Table S1 [as ‘Monodontina aff. vondembuschiana sp. 1’].
Huang et al., 2019, MPE 130: Table S4 [as ‘Monodontina aff. vondembuschiana sp. 1’].
Konopleva et al., 2019, Sci. Repts. 9: Table S1 [as ‘Monodontina aff. vondembuschiana sp. 1’].
Lopes-Lima et al., 2020, MPE 146: Table 1 [as ‘Monodontina aff. vondembuschiana sp. 1’].
(misidentified reference to Margaritana vondembuschiana Lea, 1840)
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Pseudodon avae Bolotov et al., 2017
‘Theobald’ of Authors.
Bolotov et al., 2017, Sci. Repts. 7: Table S1 [as ‘Pseudodon cf. avae’].
Bolotov et al., 2017, Sci. Repts. 7: Table S1.
Bolotov et al., 2018, Sci. Repts. 8: Table S1.
Konopleva et al., 2019, Ecologica Montenegrina 24: Appendix 1 [as ‘Pseudodon cf. avae’].
Huang et al., 2019, MPE 130: Table S4.
Konopleva et al., 2019, Sci. Repts. 9: Table S1.
Zieritz et al., 2020, Biodivers. Conserv. 29: 2244.
Lopes-Lima et al., 2020, MPE 146: Table 1.
Dai et al., 2023, Invert. Syst. 37(2): 155, Table 1.
(misidentified reference to Monocondylaea avae Theobald, 1873)
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Solenaia oleivora Huang et al., 2019
‘Heude’ of Authors.
Huang et al., 2019, MPE 130: Table S4 [as ‘Solenaia oleivora sp. 2’].
Lopes-Lima et al., 2020, MPE 146: Table 1 [as ‘Solenaia cf. oleivora 3’].
Bolotov et al., 2020, Sci. Repts. 10: Table S1 [as ‘Solenaia oleivora sp. 2’].
(misidentified reference to Mycetopus oleivorus Heude, 1877)
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Solenaia oleivora Huang et al., 2019
‘Heude’ of Authors.
Huang et al., 2019, MPE 130: Table S4 [as ‘Solenaia oleivora SDCFM189’].
Chen et al., 2020, mtDNA B 5(3): 2800 [as ‘Solenaia oleivora MG’]. [in part]
Lopes-Lima et al., 2020, MPE 146: Table 1 [as ‘Solenaia cf. oleivora 1’].
Bolotov et al., 2020, Sci. Repts. 10: Table S1 [as ‘Solenaia oleivora sp. 1’].
Sano et al., 2020, Plankton Benthos Res. 15(2): 101.
Bolotov et al., 2022, Hindawi 6600359: Table S1.
(misidentified reference to Mycetopus oleivorus Heude, 1877)
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Koreosolenaia sitgyensis Lopes-Lima et al., 2020
Lee, Kim, Lopes-Lima & Bogan in Lopes-Lima et al., 2020, MPE 146: [23], fig. 18D.
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Unio osbreckii Lopes-Lima et al., 2020
‘Philippi’ Lopes-Lima et al., 2020, MPE 146: Appendix 1.
(error for Unio osbeckii Philippi, 1844)
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Pronodularia japanensis Lopes-Lima et al., 2020
‘Lea’ of Authors.
Lopes-Lima et al., 2020, MPE 146: fig. 16 [as ‘Pronodularia cf. japanensis 3]. [in part]
(reference to Unio japanensis Lea, 1859)
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Beringiana fukuharai Lopes-Lima et al., 2020
San, Hattori & Kondo in Lopes-Lima et al., 2020, MPE 146: [20], fig. 18A.
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Beringiana gosannensis Lopes-Lima et al., 2020
San, Hattori & Kondo in Lopes-Lima et al., 2020, MPE 146: [20], fig. 18B.
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Buldowskia kamiyai Lopes-Lima et al., 2020
San, Hattori & Kondo in Lopes-Lima et al., 2020, MPE 146: [4, 22], fig. 18C.
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Nodularia breviconcha Lopes-Lima et al., 2020
Lee, Kim, Bogan & Kondo in Lopes-Lima et al., 2020, MPE 146: [5, 23].
(new name for Unio douglasiae var. sinuolatus Martens, 1905)
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Taxonomic Opinions
Aculamprotula cf. fibrosa (Heude, 1877)
Aculamprotula coreana (Martens, 1886)
Aculamprotula scripta (Heude, 1875)
Aculamprotula tientsinensis
Aculamprotula tortuosa (Lea, 1865)
Acuticosta chinensis
Alasmidonta marginata
Amblema plicata
Amuranodonta kijaensis Moskvicheva, 1973
+ Amuranodonta parva Moskvicheva, 1973
+ Amuranodonta minima Moskvicheva, 1973
+ Buldowskia (Amuranodonta) lomakini Zatravkin & Bogatov, 1987
+ Buldowskia (Amuranodonta) boloniensis Zatravkin & Bogatov, 1987
+ Amuranodonta inflata Bogatov & Starobogatov, 1996
+ Amuranodonta pulchra Bogatov & Starobogatov, 1996
+ Anemina ussuriensis Qian et al., 2015
Anemina arcaeformis (Heude, 1877)
+ Anodon rivularis Heude, 1877
+ Anodonta heudei Neumayr, 1899
+ Anemina globosula Wu et al., 2018
+ Anemina euscaphys Huang et al., 2019
Anemina euscaphys
Anodonta anatina
Anodonta cygnea
Anodonta nuttalliana
Aspatharia pfeifferiana
Beringiana beringiana (Middendorff, 1851)
+ Anodonta youkanensis Lea, 1867
+ Anodonta beringiana var. taranetzi Zhadin, 1938
+ Amuranodonta sihotealinica Zatravkin & Starobogatov, 1984
+ Arsenievinaia coptzevi Zatravkin & Bogatov, 1987
+ Arsenievinaia zimini Zatravkin & Bogatov, 1987
+ Arsenievinaia zarjaensis Bogatov & Zatravkin, 1988
+ Arsenievinaia alimovi Bogatov & Zatravkin, 1988
+ Arsenievinaia compressa Bogatov & Starobogatov, 1996
+ Arsenievinaia zatravkini Bogatov & Starobogatov, 1996
+ Beringiana compressa Sayenko & Bogatov, 1998
+ Kunashiria japonica boreosakhalinensis Labay & Shulga, 1999
+ Anemina (Buldowskia) lacustris Labay & Shulga, 1999
+ Anemina (Buldowskia) adotymensys Labay & Shulga, 1999
+ Kunashiria iturupica Bogatov et al., 1999
+ Kunashiria sinanodontoides Bogatov et al., 1999
+ Beringiana kamchatica Bogatov & Starobogatov, 2001
+ Beringiana derzhavini Bogatov & Starobogatov, 2001
+ Beringiana chereshnevi Bogatov & Starobogatov, 2001
Beringiana fukuharai San, Hattori & Kondo sp. nov.
Beringiana gosannensis San, Hattori & Kondo sp. nov.
Beringiana japonica (Clessin, 1874)
Bineurus aff. mouhotii sp. 1
Bineurus cumingii
Buldowskia flavotincta (Martens, 1905)
Buldowskia iwakawai (Suzuki, 1939)
Buldowskia kamiyai San, Hattori & Kondo sp. nov.
Buldowskia shadini (Moskvicheva, 1973)
+ Anodonta (Haasiella) euscaphus Zhadin, 1938
+ Sinanodonta (Anemina) fuscoviridis Moskvicheva, 1973
+ Sinanodonta (Anemina) buldowskii Moskvicheva, 1973
+ Buldowskia (Amurbuldowskia) sitaensis Bogatov & Starobogatov, 1996
Buldowskia suifunica (Lindholm, 1925)
+ Anodonta beringiana var. suifunensis Zhadin, 1938
+ Anodonta woodiana var. elliptica Zhadin, 1938
+ Anodonta (Haasiella) arcaeformis Zhadin, 1938
+ Buldowskia suputinensis Moskvicheva, 1973
+ Amuranodonta starobogatovi Moskvicheva, 1973
+ Buldowskia cylindrica Moskvicheva, 1973
+ Anemina (Buldowskia) zatrawkini Martynov & Chernyshev, 1992
+ Anemina (s.s.) shadini deflexa Martynov & Chernyshev, 1992
+ Buldowskia (Buldowskia) possietica Bogatov & Starobogatov, 1996
+ Buldowskia (Buldowskia) koreana Bogatov & Starobogatov, 1996
Chambardia wahlbergi
Chamberlainia hainesiana
Coelatura aegyptiaca
Contradens contradens
Cristaria cf. radiata
Cristaria clessini (Kobelt, 1879)
+ Dipsas plicata var. japonica Kobelt, 1879
Cristaria plicata (Leach, 1814)
+ Cristaria tuberculata Schumacher, 1817
+ Anodonta cristata Blainville, 1825
+ Anodonta dipsas Blainville, 1825
+ Symphynota bialata Lea, 1829
+ Dianisotis chinensis Rafinesque, 1831
+ Anodonta (Dispsas) herculea Middendorff, 1848
+ Anodon gigantea Gould, 1849
+ Margaron (Anodonta) onowensis Lea, 1852
+ Anodonta bellua Morelet, 1866
+ Anodon herculeus Sowerby, 1867
+ Anodonta magnifica Clessin, 1873
+ Anodonta (Dipsas) spatiosa Clessin, 1876
+ Dipsas occidentalis Heude, 1885
+ Barbala plicatula Paetel, 1890
Cuneopsis celtiformis
Cuneopsis cf. heudei
Cuneopsis pisciculus
Cuneopsis rufescens
Elliptio complanata
Ensidens aff. sagittarius sp. 1
Etheria elliptica
Gonidea angulata (Lea, 1838)
Hyridella depressa
Hyriopsis bialata
Inversidens brandtii (Kobelt, 1879)
Inversidens pantoensis (Neumayr, 1899)
Inversiunio jokohamensis (Ihering, 1893)
+ Unio japanensis Kobelt, 1879
+ Unio haconensis Ihering, 1893
Inversiunio reinianus (Kobelt, 1879)
+ Nodularia hirasei Haas, 1911
Inversiunio verrucosus Kondo, Yang & Choi, 2007
Inversiunio yanagawensis (Kondo, 1982)
Koreosolenaia sitgyensis Lee, Kim, Lopes-Lima & Bogan gen. & sp. nov.
+ Solenaia triangularis Wu et al., 2018
Lamellidens corrianus
Lamproscapha ensiformis
Lamprotula caveata (Heude, 1877)
+ Lamprotula scripta Wang et al., 2018
Lamprotula leaii (Gray in Griffith & Pidgeon, 1833)
+ Unio nodulosus Reeve, 1864
+ Unio leleci Heude, 1875
+ Unio richthofeni Martens, 1875
+ Unio gottschei Martens, 1894
+ Unio leai var. truncatula Neumayr, 1899
+ Unio (Quadrula) leai var. ponderosa Dautzenberg & Fischer, 1905
Lamprotula seomjinensis (Kondo, Yang & Choi, 2007)
Lampsilis cardium
Lanceolaria acrorrhyncha (Martens, 1894)
Lanceolaria cf. bilirata (Martens, 1902)
Lanceolaria gladiola
Lanceolaria grayii (Griffith & Pidgeon, 1833)
+ Unio grayanus Schrenck, 1867
+ Unio grayanus var. decurvata Schrenck, 1867
+ Nodularia cylindracea Simpson, 1900
+ Nodularia cylindrica Simpson, 1900
+ Lanceolaria ussuriensis Moskvicheva, 1973
+ Lanceolaria chankensis Moskvicheva, 1973
+ Lanceolaria maacki Moskvicheva, 1973
+ Lanceolaria (Pericylindrica) bogatovi Zatravkin & Starobogatov, 1984
Lanceolaria kihirai (Kondo & Hattori, 2019)
Lanceolaria lanceolata (Lea, 1856)
Lanceolaria oxyrhyncha (Martens, 1861)
Lanceolaria triformis (Heude, 1877)
Lasmigona compressa
Leguminaia wheatleyi
Leoparreysia bhamoensis
Lepidodesma languilati
Margaritifera dahurica
Margaritifera laevis
Margaritifera middendorffi
Microcondylaea bonellii
Middendorffinaia mongolica (Middendorff, 1851)
+ Unio (Nodularia) continentalis Zhadin, 1938
+ Middendorffinaia (Suifununio) sujfunensis Moskvicheva & Starobogatov, 1973
+ Middendorffinaia (Pseudopotomida) shadini Moskvicheva & Starobogatov, 1973
+ Middendorffinaia (Pseudopotomida) dulkeitiana Moskvicheva & Starobogatov, 1973
+ Middendorffinaia (Pseudopotomida) weliczkowskii Moskvicheva & Starobogatov, 1973
+ Middendorffinaia (Pseudopotomida) hassanica Moskvicheva & Starobogatov, 1973
+ Middendorffinaia (Pseudopotomida) martensi Moskvicheva & Starobogatov, 1973
+ Middendorffinaia maihensis Moskvicheva, 1973
+ Middendorffinaia alimovi Bogatov, 2012
Monodontina aff. vondembuschiana sp. 1
Monodontina aff. “inoscularis” sp. 2
Mycetopoda siliquosa
Neotrigonia lamarckii
Neotrigonia margaritacea
Nodularia breviconcha Lee, Kim, Bogan & Kondo nom. nov.
+ Unio douglasiae var. sinuolatus Martens, 1905
Nodularia douglasiae (Griffith & Pidgeon, 1833)
+ Unio murchisonianus Lea, 1834
+ Unio osbeckii Philippi, 1844
+ Unio pictorum var. longirostris Middendorff, 1851
+ Unio shanghaiensis Lea, 1859
+ Unio wrightii Lea, 1865
+ Unio sculptus Deshayes, 1873
+ Unio biwae Kobelt, 1879
+ Unio douglasiae chloreus Heude, 1885
+ Unio douglasiae dactylinus Heude, 1885
+ Unio douglasiae deltae Heude, 1885
+ Unio douglasiae firmus Heude, 1885
+ Unio douglasiae hainanensis Heude, 1885
+ Unio pictorum var. amurensis Mousson, 1887
+ Unio middendorffi Westerlund, 1890
+ Unio verrucifer Martens, 1894
+ Unio pliculosus Martens, 1894
+ Unio abbreviatus Westerlund, 1897
+ Unio schrencki Westerlund, 1897
+ Nodularia douglasiae flavoviridis Haas, 1910
+ Nodularia sujfunica Moskvicheva, 1973
+ Nodularia vladivostokensis Moskvicheva, 1973
+ Middendorffinaia (s.s.) ussuriensis Moskvicheva & Starobogatov, 1973
+ Middendorffinaia (s.s.) arsenievi Moskvicheva & Starobogatov, 1973
+ Nodularia lebedevi Zatravkin & Starobogatov, 1984
+ Nodularia (Magadaninaia) extremalis Martynov & Chernyshev, 1992
+ Nodularia (s.s.) moskvichevae Bogatov & Starobogatov, 1992
+ Middendorffinaia (Middendorffinaia) ochotica Bogatov, 2000
+ Nodularia sakhalinensis Bogatov, 2001
+ Nodularia jourdyi Bolotov et al., 2018
Nodularia nipponensis (Martens, 1877)
Nodularia nuxpersicae
Obovalis omiensis (Heimburg, 1884)
Parvasolenaia neotriangularis He & Zhuang, 2013
Parvasolenaia rivularis (Heude, 1877)
+ Solenaia triangularis Lopes-Lima et al., 2017
Parvasolenaia triangularis (Heude, 1885)
Physunio modelli
Pilsbryoconcha aff. compressa sp. 1
Pilsbryoconcha aff. exilis sp. 1
Pletholophus reinianus (Martens, 1875)
Pletholophus tenuis (Gray in Griffith & Pidgeon, 1833)
+ Symphynota discoidea Lea, 1834
Potomida littoralis
Pronodularia cf. japanensis (Lea, 1859)
Pseudanodonta complanata
Pseudodon avae
Pseudodon manueli
Ptychorhynchus pfisteri (Heude, 1874)
Pyganodon grandis
Quadrula quadrula
Radiatula mouhoti
Rectidens sumatrensis
Schistodesmus lampreyanus
Schistodesmus spinosus
Simpsonaias ambigua
Sinanodonta angula (Tchang, Li & Liu, 1965)
Sinanodonta calipygos (Kobelt, 1879)
Sinanodonta cf. elliptica
+ Sinanodonta woodiana Bolotov et al., 2016
+ Sinanodonta joreti Huang et al., 2019
+ Sinanodonta pacifica Huang et al., 2019
Sinanodonta cf. woodiana 2 (Lea, 1834)
+ Sinanodonta elliptica Huang et al., 2019
Sinanodonta jourdyi
+ Sinanodonta woodiana Lopes-Lima et al., 2017
Sinanodonta lauta (Martens, 1877)
+ Sinanodonta woodiana fukudai Modell, 1945
+ Sinanodonta (Ellipsanodon) ovata Bogatov & Starobogatov, 1996
+ Sinanodonta (Ellipsanodon) manchurica Bogatov & Starobogatov, 1996
Sinanodonta lucida
Sinanodonta schrenkii (Lea, 1870)
+ Anodonta magnifica Schrenck, 1867
+ Sinanodonta (s.s.) schrencki Moskvicheva, 1973
+ Sinanodonta (s.s.) amurensis Moskvicheva, 1973
+ Sinanodonta (Cristariopsis) crassitesta Moskvicheva, 1973
+ Sinanodonta (s.s.) likharevi Moskvicheva, 1973
+ Sinanodonta (s.s.) renzini Bogatov & Zatravkin, 1988
+ Sinanodonta (Cristariopsis) primorjensis Bogatov & Zatravkin, 1988
+ Sinanodonta woodiana Soroka, 2010
Sinanodonta tumens (Haas, 1910)
+ Oguranodonta ogurae Kuroda & Habe, 1987
Sinohyriopsis cf. cumingii (Lea, 1852)
Sinohyriopsis goliath (Rolle, 1904)
Sinohyriopsis schlegelii (Martens, 1861)
Solenaia carinata
Solenaia cf. oleivora 2
Solenaia sp.
Strophitus undulatus
Trapezoideus nesemanni
Unio crassus
Unio gibbus
Unio pictorum
Unio tumidus
Velesunio ambiguus
|
Cladogram Figure 10
+-- Margaritifera_middendorffi
+-----------------+
| +-- Margaritifera_laevis
|
--+ +-- Beringiana_japonica
| +--------------+
| | +-- Beringiana_beringiana
| |
+--+ +-- Sinanodonta_angula
| +-----------+
| | +-- Sinanodonta_lucida
| |
+--+ +----------- Sinanodonta_schrenkii
| |
| | +-- Sinanodonta_calipygos
| +--------+
+--+ +-- Sinanodonta_tumens
|
| +-------- Sinanodonta_woodiana
| |
+--+-------- Sinanodonta_jourdyi
|
| +----- Sinanodonta_elliptica
+--+
| +-- Sinanodonta_lauta
+--+
+-- Sinanodonta
Cladogram Figure 11
+-- Margaritifera_laevis
+--------------------+
| +-- Margaritifera_middendorffi
|
--+ +-- Nodularia_breviconcha
| +-----------------+
| | +-- Nodularia_douglasiae
+--+
| +----------------- Lanceolaria_oxyrhyncha
| |
+--+ +-------------- Lanceolaria_gladiola
| |
+--+ +-- Lanceolaria_grayii
| +--------+
| | +-- Lanceolaria_bilirata
+--+
| +-------- Lanceolaria_acrorrhyncha
+--+
| +----- Lanceolaria_kihirai
+--+
| +-- Lanceolaria_lanceolata
+--+
+-- Lanceolaria_triformis
Cladogram Figure 12
+-- Margaritifera_laevis
+-----------+
| +-- Margaritifera_middendorffi
|
--+ +-- Nodularia_breviconcha
| +--------+
| | +-- Nodularia_douglasiae
+--+
| +-------- Inversiunio_jokohamensis
| |
+--+ +----- Inversiunio_verrucosus__1
| |
+--+----- Inversiunio_verrucosus__2
|
| +-- Inversiunio_reinianus
+--+
+-- Inversiunio_yanagawensis
Cladogram Figure 13
+-- Margaritifera_laevis
+--------------------+
| +-- Margaritifera_middendorffi
--+
| +-------------------- Middendorffinaia_mongolica
+--+
| +----------------- Nodularia_breviconcha
+--+
| +-------------- Nodularia_fusiformans
+--+
| +----------- Nodularia_huana
+--+
| +-------- Nodularia_nuxpersicae
+--+
| +----- Nodularia_nipponensis
+--+
| +-- Nodularia_douglasiae
+--+
+-- Nodularia_dualobtusa
Cladogram Figure 14
+-- Margaritifera_laevis
+-----+
| +-- Margaritifera_middendorffi
|
--+ +-- Parvasolenaia_rivularis
| +--+
| | +-- Koreosolenaia_sitgyensis
+--+
| +-- Lamprotula_leaii
+--+
+-- Lamprotula_seomjinensis
Cladogram Figure 16
+-- Margaritifera_laevis
+-----------------+
| +-- Margaritifera_middendorffi
--+
| +----------------- Chamberlainia_somsakpanhai
+--+
| +-------------- Sinohyriopsis_schlegelii
+--+
| +----------- Pronodularia_japanensis
+--+
| +-------- Lamprotula_caveata
+--+
| +----- Lamprotula_leaii__2
+--+
| +-- Lamprotula_seomjinensis
+--+
+-- Lamprotula_leaii__1
Cladogram Figure 17
+-- Margaritifera_laevis
+-----------+
| +-- Margaritifera_middendorffi
|
--+ +----------- Chamberlainia_somsakpanhai
| |
| | +-------- Sinohyriopsis_cumingii__10
+--+ |
| | +----- Sinohyriopsis_cumingii__1
| | |
| | +----- Sinohyriopsis_cumingii__2
+--+ |
| +----- Sinohyriopsis_cumingii__3
| |
| +----- Sinohyriopsis_cumingii__4
| |
| +----- Sinohyriopsis_cumingii__5
+--+
+----- Sinohyriopsis_cumingii__6
|
+----- Sinohyriopsis_cumingii__7
|
+----- Sinohyriopsis_cumingii__8
|
+----- Sinohyriopsis_cumingii__9
|
| +-- Sinohyriopsis_schlegelii
+--+
+-- Sinohyriopsis_goliath
Cladogram Figure 2-4
+-- Neotrigonia_lamarckii
+-----------------------------------------------+
| +-- Neotrigonia_margaritacea
|
| +-- Hyridella_depressa
| +--------------------------------------------+
| | +-- Velesunio_ambiguus
--+ |
| | +-- Margaritifera_laevis
| | +-----------------------------------------+
| | | +-- Margaritifera_middendorffi
| | |
| | | +-- Lamproscapha_ensiformis
+--+ | +-----+
| | | +-- Mycetopoda_siliquosa
| | +--------------------------------+
| | | | +----- Etheria_elliptica
| | | +--+
| | | | +-- Aspatharia_pfeifferiana
| | | +--+
| | | +-- Chambardia_wahlbergi
| | |
+--+ | +-- Coelatura_aegyptiaca
| | |
| | +-----------------------------------+-- Leoparreysia_tavoyensis
| | | |
| | | +-- Radiatula_mouhoti
| | |
| | | +-- Lens_contradens
| | | |
| | | +-- Ensidens_sagittarius
| | | |
| | | +-----------------------+-- Physunio_modelli
| | | | |
| | | | +-- Rectidens_sumatrensis
| | | | |
+--+ | | +-- Yaukthwa_nesemanni
| | |
| | | +-- Amblema_plicata
| | | |
| | | +-- Elliptio_complanata
| | | +--------------------+
| | +--------+ | +-- Lampsilis_cardium
| | | | | |
| | | | | +-- Quadrula_quadrula
| | | | |
| | | | | +----- Chamberlainia_somsakpanhai
| | | | | +-----------+
| | | | | | | +-- Sinohyriopsis_cumingii
| | | | | | +--+
| | | | | +--+ +-- Sinohyriopsis_schlegelii
| | | | | | |
| | | | | | | +-------------- Pronodularia_japanensis
| | | +--+ | | |
| | | | | +--+ +----------- Potomida_littoralis
| | | | | | |
| | | | | +--+ +-------- Lamprotula_caveata
| | | | | | |
+--+ | | | +--+ +----- Lamprotula_leaii__2
| | | | | |
| | | | +--+----- Lamprotula_leaii__3
| | | | |
| | | | | +-- Lamprotula_seomjinensis
| | | | +--+
| | | | +-- Lamprotula_leaii__1
| | +--+
| | | +-- Indopseudodon_indawgyiensis
| | | +--------+
| | | | +-- Indopseudodon_salwenianus
| | | |
| | | +-----+ +-- Pilsbryoconcha_exilis
| | | | | +-----+
| | | | | | +-- Pilsbryoconcha_carinifera
| | | | +--+
| | | | | +----- Pseudodon_mekongi
| | | | | |
| | | | +--+ +-- Nyeinchanconcha_nyeinchani
| | | | | |
| | | | +--+-- Bineurus_mouhotii
| | | | |
| | +--+ +-- Sundadontina_tumida
| | |
| | | +-------- Inversidens_brandtii
| | | |
| | | +-----+ +-- Obovalis_omiensis
| | | | | +--+
| | | | | | +-- Ptychorhynchus_pfisteri
| | | | +--+
| | | | | +-- Parvasolenaia_rivularis
| | | | +--+
| | +--+ +-- Koreosolenaia_sitgyensis
| | |
+--+ | +----- Gonidea_angulata
| | +-----+
| | | | +-- Leguminaia_wheatleyi
| | | +--+
| +--+ +-- Microcondylaea_bonellii
| |
| | +-------- Sinosolenaia_carinata
| +--+
| | +----- Sinosolenaia_recognita
| +--+
| | +-- Sinosolenaia_iridinea
| +--+
| +-- Sinosolenaia_oleivora
|
| +-------------------------------- Acuticosta_chinensis
| |
| +-------------------------------- Lepidodesma_languilati
| |
| | +----------- Aculamprotula_tortuosa
| | +--------------+
| | | | +-------- Aculamprotula_scripta
| | | +--+
| | | | +----- Aculamprotula_coreana
| | | +--+
| | | | +-- Aculamprotula_fibrosa
| | | +--+
| +-----+ +-- Aculamprotula_tientsinensis
| | |
| | | +----------------------- Middendorffinaia_mongolica
| | | |
| | | | +----- Unio_gibbus
| | | | |
| | | | +--------------+----- Unio_tumidus
| | | | | |
| | +--+ | | +-- Unio_crassus
| | | | +--+
| | | | +-- Unio_pictorum
| | | |
| | | | +-------------- Nodularia_breviconcha
| | | | +--+
| | +--+ | | +----------- Nodularia_huana
| | | | +--+
| | | | | +-------- Nodularia_nuxpersicae
| | | | +--+
| | | | | +----- Nodularia_nipponensis
| | | | +--+
| | | | | +-- Nodularia_douglasiae
| | | | +--+
| | +--+ +-- Nodularia_dualobtusa
| | |
| | | +-- Schistodesmus_lampreyanus
+--+ | +-----------+
| | | +-- Schistodesmus_spinosus
| | |
| | | +-------- Cuneopsis_rufescens
| | +-----+
| | | | +----- Cuneopsis_celtiformis
| +--+ +--+
| | | +-- Cuneopsis_szechenyii
| | +--+
| | +-- Cuneopsis_heudei
| |
| | +----------- Tchangsinaia_piscicula
| +--+
| | +-------- Inversiunio_jokohamensis
| +--+
| | +----- Inversiunio_verrucosus
| +--+
| | +-- Inversiunio_reinianus
| +--+
| +-- Inversiunio_yanagawensis
|
| +----------------- Lanceolaria_oxyrhyncha
| |
| +-----------+ +-------------- Lanceolaria_gladiola
| | | |
| | +--+ +-- Lanceolaria_bilirata
| | | +--------+
| | | | +-- Lanceolaria_grayii
| | +--+
| | | +-------- Lanceolaria_acrorrhyncha
| | +--+
| | | +----- Lanceolaria_kihirai
| | +--+
| | | +-- Lanceolaria_lanceolata
| | +--+
| | +-- Lanceolaria_triformis
| |
| | +-------- Pseudanodonta_complanata
+--+ |
| | +----- Anodonta_cygnea
| +--+
| | | +-- Anodonta_anatina
| +-----------------+ +--+
| | | +-- Anodonta_nuttalliana
| | |
| | | +----- Pyganodon_grandis
| | | |
| | | +----- Simpsonaias_ambigua
| | +--+
| | +----- Strophitus_undulatus
| | |
| | | +-- Alasmidonta_marginata
| | +--+
| | +-- Lasmigona_costata
+--+
| +-- Cristaria_plicata
| +--------------------+
| | +-- Cristaria_clessini
| |
| | +-------------- Anemina_arcaeformis
| | |
| +--------+ +----------- Amuranodonta_kijaensis
| | | |
| | +--+ +-------- Buldowskia_suifunica
| | | |
| | +--+ +-- Buldowskia_flavotincta
+--+ | +--+
| | | +-- Buldowskia_shadini
| +--+
| | +-- Buldowskia_iwakawai
| +--+
| +-- Buldowskia_kamiyai
|
| +-------------------- Pletholophus_tenuis
| |
| | +-------- Beringiana_japonica
| | +--------+
+--+ | | +----- Beringiana_beringiana
| | +--+
| | | +-- Beringiana_fukuharai
| | +--+
+--+ +-- Beringiana_gosannensis
|
| +-- Sinanodonta_angula
| +-----------+
| | +-- Sinanodonta_lucida
| |
+--+ +----------- Sinanodonta_schrenkii
| |
| | +-- Sinanodonta_calipygos
| +--------+
+--+ +-- Sinanodonta_tumens
|
| +-------- Sinanodonta_woodiana
| |
+--+ +----- Sinanodonta_elliptica
| |
+--+----- Sinanodonta_jourdyi
|
| +-- Sinanodonta_lauta
+--+
+-- Sinanodonta_pacifica
Cladogram Figure 5
+-- Margaritifera_laevis
+--------------+
| +-- Margaritifera_middendorffi
|
--+ +-- Nodularia_breviconcha
| +-----------+
| | +-- Nodularia_douglasiae
+--+
| +----------- Aculamprotula_tortuosa
+--+
| +-------- Aculamprotula_tientsinensis
+--+
| +----- Aculamprotula_fibrosa
+--+
| +-- Aculamprotula_scripta
+--+
+-- Aculamprotula_coreana
Cladogram Figure 6
+-- Margaritifera_laevis
+-----------------+
| +-- Margaritifera_middendorffi
|
--+ +-- Sinanodonta_lauta
| +--------------+
| | +-- Sinanodonta_woodiana
+--+
| +-------------- Amuranodonta_kijaensis
| |
+--+ +----------- Anemina_arcaeformis
| |
+--+ +-- Buldowskia_iwakawai
| +-----+
| | +-- Buldowskia_kamiyai
+--+
| +----- Buldowskia_shadini
+--+
| +-- Buldowskia_suifunica
+--+
+-- Buldowskia_flavotincta
Cladogram Figure 7
+-- Margaritifera_laevis
+-----------+
| +-- Margaritifera_dahurica
|
--+ +-- Sinanodonta_woodiana
| +--------+
| | +-- Sinanodonta_tumens
+--+
| +-------- Beringiana_gosannensis
+--+
| +----- Beringiana_fukuharai
+--+
| +-- Beringiana_beringiana
+--+
+-- Beringiana_japonica
Cladogram Figure 8
+-- Margaritifera_laevis
+--------+
| +-- Margaritifera_middendorffi
|
--+ +-- Sinanodonta_woodiana
| +-----+
| | +-- Sinanodonta_lucida
+--+
| +----- Cristaria_clessini
+--+
| +-- Cristaria_plicata
+--+
+-- Cristaria_radiata
Cladogram Figure 9
+-- Margaritifera_laevis
+--------+
| +-- Margaritifera_middendorffi
--+
| +-------- Cristaria_plicata
+--+
| +----- Sinanodonta_schrenkii
+--+
| +-- Pletholophus_tenuis
+--+
+-- Pletholophus_reinianus
|
Page updated: 07:59:57 Tue 01 Jul 2025 |
|
