|
Bivalvia | Unionoida | Unionoidea | Unionidae | Parreysiinae tribe INDOCHINELLINI Bolotov, Pfeiffer, Vikhrev, & Konopleva in Bolotov et al., 2018: cladomics
7 genera
Cladomics Summary
number of species = 31
number of subtaxa (rank: genus) = 7
analyzed in 46 cladograms in 30 publications
maximum number of terminals in an analysis = 27, mean = 8.61
maximum number of subtaxa (genus) in an analysis = 7, mean = 3.37
Basic Monophyly
number of cladograms that test basic monophyly (i.e., more than 1 included species) = 30
maximum number of included terminals = 27, mean = 12.67
maximum number of subtaxa (genus) = 7, mean = 4.63
proportion of cladograms that support basic monophyly = 0.97
Rigorous Monophyly
number of cladograms that test rigorous monophyly = 28
(Rigorous monophyly refers to cladograms with terminal species representing more than one subtaxon (genus) taxon and at least one rigorous outgroup (tribe).
maximum number of included terminals = 27, mean = 12.64
maximum number of subtaxa (genus) = 7, mean = 4.57
proportion of cladograms that support rigorous monophyly = 0.96
Pairwise Cladogram Clustering
The graphs below depict in two dimensions all pair-wise ingroup distances within the cladome. See the Cladmoics FARQ for more information.
Bolotov et al., 2022 [Publ 4252], fig. 2
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 27 |
27 |
25 |
7 |
34 |
30 |
4 |
0.0 |
*** Top Monophyly Cladogram *** *** Top Sister-Group Cladogram ***
+-- Leoparreysia_burmana
+-----+
| +-- Leoparreysia_whitteni
+-----------------------------+
| | +----- Leoparreysia_olivacea
| +--+
| | +-- Leoparreysia_canefrii
| +--+
| +-- Leoparreysia_tavoyensis
|
| +-------- Nitia_teretiuscula
--+ +--------------------------+
| | | +----- Coelatura_aegyptiaca
| | +--+
| | | +-- Nyassunio_nyassaensis
| | +--+
| | +-- Coelatura_kunenensis
| |
| | +----------------- Indonaia_cylindrica **
| | |
+--+ +--------------+ +-------------- Indonaia_gratiosa **
| | | |
| | +--+ +----------- Indonaia_andersoniana **
| | | |
| | +--+ +-- Indonaia_caerulea **
| | | +-----+
| | | | +-- Indonaia_subclathrata **
| | +--+
| | | +----- Indonaia_rugosa **
+--+ +--+
| | +-- Indonaia_bonneaudii **
| +--+
| +-- Indonaia_shurtleffiana **
|
| +----------------------------- Indochinella_pugio **
| |
| | +----------- Radiatula_crispisulcata **
| | |
| | | +----- Radiatula_chaudhurii **
+--+ +--------------+ +--+
| | | | | +-- Radiatula_myitkyinae **
| | | | +--+
| | +--+ +-- Radiatula_kachinensis **
| | |
| | | +----- Radiatula_chindwinensis **
+--+ +--+
| | +-- Radiatula_myitthanensis **
| +--+
| +-- Radiatula_mouhoti **
|
| +----------------------- Scabiellus_songkramensis **
| |
+--+ +-------------------- Harmandia_somboriensis **
| |
+--+ +----------------- Unionetta_fabagina **
| |
+--+ +-- Scabies_crispata **
| +-----------+
| | +-- Scabies_mandarinus **
+--+
| +----------- Scabies_nucleus **
| |
+--+ +-- Scabies_humilis **
| +-----+
| | +-- Scabies_pilata **
+--+
| +----- Scabies_phaselus **
+--+
| +-- Scabies_anceps **
+--+
+-- Scabies_scobinatus **
Bolotov et al., 2022 [Publ 4252], fig. S1
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 27 |
27 |
25 |
7 |
34 |
30 |
4 |
20.0 |
+-- Leoparreysia_burmana
|
+-- Leoparreysia_canefrii
|
+--------------------------------------+-- Leoparreysia_olivacea
| |
| +-- Leoparreysia_tavoyensis
| |
| +-- Leoparreysia_whitteni
|
| +-- Coelatura_aegyptiaca
| |
| +-- Coelatura_kunenensis
--+ +-----------------------------------+
| | +-- Nitia_teretiuscula
| | |
| | +-- Nyassunio_nyassaensis
| |
| | +-- Indonaia_cylindrica **
| | +--------+
| | | +-- Indonaia_gratiosa **
| | |
+--+ +-----------------------+ +----- Indonaia_shurtleffiana **
| | | +--+
| | | | | +-- Indonaia_bonneaudii **
| | | | +--+
| | +--+ +-- Indonaia_rugosa **
| | |
| | | +----- Indonaia_andersoniana **
| | +--+
| | | +-- Indonaia_caerulea **
+--+ +--+
| +-- Indonaia_subclathrata **
|
| +-------------------------------- Indochinella_pugio **
| |
| | +----- Radiatula_chaudhurii **
| | +-----+
| | | | +-- Radiatula_kachinensis **
| | | +--+
+--+ +-----------------+ +-- Radiatula_myitkyinae **
| | |
| | | +-------- Radiatula_crispisulcata **
| | +--+
| | | +----- Radiatula_mouhoti **
| | +--+
+--+ | +-- Radiatula_myitthanensis **
| +--+
| +-- Radiatula_chindwinensis **
|
| +-------------------------- Scabiellus_songkramensis **
| |
+--+ +----------------------- Harmandia_somboriensis **
| |
+--+ +-------------------- Unionetta_fabagina **
| |
+--+ +----------------- Scabies_crispata **
| |
+--+ +-- Scabies_humilis **
| +-----------+
| | +-- Scabies_mandarinus **
+--+
| +----------- Scabies_nucleus **
+--+
| +-------- Scabies_pilata **
+--+
| +----- Scabies_anceps **
+--+
| +-- Scabies_phaselus **
+--+
+-- Scabies_scobinatus **
Bolotov et al., 2022 [Publ 4239], fig. 2
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 26 |
26 |
24 |
7 |
32 |
28 |
4 |
15.0 |
+----------- Parreysia_keralaensis
+--+
| | +-------- Parreysia_plagiosoma
| +--+
| | +----- Parreysia_nagpoorensis
| +--+
| | +-- Parreysia_corrugata
+-----------------------------+ +--+
| | +-- Parreysia_rakhinensis
| |
| | +-- Leoparreysia_burmana
| | +-----+
| | | +-- Leoparreysia_whitteni
| +-----+
| | +----- Leoparreysia_olivacea
| +--+
| | +-- Leoparreysia_canefrii
| +--+
--+ +-- Leoparreysia_tavoyensis
|
| +-------- Nitia_teretiuscula
| +--------------------------------+
| | | +----- Coelatura_aegyptiaca
| | +--+
| | | +-- Coelatura_kunenensis
| | +--+
| | +-- Nyassunio_nyassaensis
| |
| | +-- Indonaia_cylindrica **
| | +--------+
+--+ | +-- Indonaia_gratiosa **
| |
| +--------------------------+ +----- Indonaia_andersoniana **
| | | +--+
| | | | | +-- Indonaia_caerulea **
| | | | +--+
| | +--+ +-- Indonaia_subclathrata **
| | |
| | | +----- Indonaia_shurtleffiana **
| | +--+
+--+ | +-- Indonaia_bonneaudii **
| +--+
| +-- Indonaia_rugosa **
|
| +----------------------------------- Indochinella_pugio **
| |
| | +----------- Radiatula_crispisulcata **
| | |
+--+ +--------------------+ +-- Radiatula_chaudhurii **
| | | +-----+
| | | | +-- Radiatula_myitkyinae **
| | +--+
| | | +----- Radiatula_mouhoti **
+--+ +--+
| | +-- Radiatula_chindwinensis **
| +--+
| +-- Radiatula_myitthanensis **
|
| +----------------------------- Scabiellus_songkramensis **
+--+
| +-------------------------- Harmandia_somboriensis **
+--+
| +----------------------- Unionetta_fabagina **
+--+
| +-------------------- Scabies_crispata **
+--+
| +----------------- Scabies_mandarinus **
+--+
| +-------------- Scabies_nucleus **
+--+
| +----------- Scabies_pilata **
+--+
| +-------- Scabies_humilis **
+--+
| +----- Scabies_anceps **
+--+
| +-- Scabies_phaselus **
+--+
+-- Scabies_scobinatus **
Bolotov et al., 2022 [Publ 4239], fig. S1
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 26 |
26 |
23 |
7 |
32 |
28 |
4 |
2.0 |
+-- Leoparreysia_burmana
+-----+
| +-- Leoparreysia_whitteni
+-----------------------------+
| | +----- Leoparreysia_olivacea
| +--+
| | +-- Leoparreysia_canefrii
| +--+
| +-- Leoparreysia_tavoyensis
|
| +-------- Nitia_teretiuscula
--+ +--------------------------+
| | | +----- Coelatura_aegyptiaca
| | +--+
| | | +-- Coelatura_kunenensis
| | +--+
| | +-- Nyassunio_nyassaensis
| |
| | +----------------- Indonaia_cylindrica **
| | |
+--+ +--------------+ +-------------- Indonaia_gratiosa **
| | | |
| | +--+ +----------- Indonaia_andersoniana **
| | | |
| | +--+ +-- Indonaia_caerulea **
| | | +-----+
| | | | +-- Indonaia_subclathrata **
| | +--+
| | | +----- Indonaia_rugosa **
+--+ +--+
| | +-- Indonaia_bonneaudii **
| +--+
| +-- Indonaia_shurtleffiana **
|
| +----------------------------- Indochinella_pugio **
| |
| | +-------- Radiatula_crispisulcata **
| | |
| | +-----------------+ +-- Radiatula_chaudhurii **
+--+ | | +--+
| | | | +-- Radiatula_myitkyinae **
| | +--+
| | | +-- Radiatula_myitthanensis **
| | | |
+--+ +--+-- Radiatula_mouhoti **
| |
| +-- Radiatula_chindwinensis **
|
| +----------------------- Scabiellus_songkramensis **
| |
+--+ +-------------------- Harmandia_somboriensis **
| |
+--+ +----------------- Unionetta_fabagina **
| |
+--+ +-- Scabies_crispata **
| +-----------+
| | +-- Scabies_mandarinus **
+--+
| +----------- Scabies_nucleus **
| |
+--+ +-- Scabies_humilis **
| +-----+
| | +-- Scabies_pilata **
+--+
| +----- Scabies_phaselus **
+--+
| +-- Scabies_anceps **
+--+
+-- Scabies_scobinatus **
Bolotov et al., 2022 [Publ 4239], fig. S2
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 26 |
26 |
24 |
7 |
32 |
28 |
4 |
19.0 |
+----------- Leoparreysia_burmana
+-----------------------------+
| | +-------- Leoparreysia_whitteni
| +--+
| | +----- Leoparreysia_olivacea
| +--+
| | +-- Leoparreysia_canefrii
| +--+
| +-- Leoparreysia_tavoyensis
|
--+ +-------- Nitia_teretiuscula
| +-----------------------------+
| | | +----- Coelatura_aegyptiaca
| | +--+
| | | +-- Coelatura_kunenensis
| | +--+
| | +-- Nyassunio_nyassaensis
| |
| | +-- Indonaia_cylindrica **
| | +--------+
+--+ | +-- Indonaia_gratiosa **
| |
| +-----------------------+ +----- Indonaia_shurtleffiana **
| | | +--+
| | | | | +-- Indonaia_bonneaudii **
| | | | +--+
| | +--+ +-- Indonaia_rugosa **
| | |
| | | +----- Indonaia_andersoniana **
| | +--+
+--+ | +-- Indonaia_caerulea **
| +--+
| +-- Indonaia_subclathrata **
|
| +-------------------------------- Indochinella_pugio **
| |
| | +----------- Radiatula_crispisulcata **
| | |
| | +-----------------+ +-- Radiatula_chaudhurii **
+--+ | | +-----+
| | | | +-- Radiatula_myitkyinae **
| | +--+
| | | +----- Radiatula_mouhoti **
| | +--+
+--+ | +-- Radiatula_myitthanensis **
| +--+
| +-- Radiatula_chindwinensis **
|
| +-------------------------- Scabiellus_songkramensis **
| |
+--+ +----------------------- Harmandia_somboriensis **
| |
+--+ +-------------------- Unionetta_fabagina **
| |
+--+ +----------------- Scabies_crispata **
| |
+--+ +-- Scabies_humilis **
| +-----------+
| | +-- Scabies_mandarinus **
+--+
| +----------- Scabies_nucleus **
+--+
| +-------- Scabies_pilata **
+--+
| +----- Scabies_anceps **
+--+
| +-- Scabies_phaselus **
+--+
+-- Scabies_scobinatus **
Bolotov et al., 2020 [Publ 4100], fig. 1
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 21 |
21 |
19 |
7 |
99 |
20 |
4 |
16.0 |
+----------- Parreysia_keralaensis
+--+
| | +-------- Parreysia_plagiosoma
| +--+
| | +----- Parreysia_nagpoorensis[2]
| +--+
| | +-- Parreysia_nagpoorensis[1]
+--------------------+ +--+
| | +-- Parreysia_rakhinensis
| |
| | +-- Leoparreysia_burmana
| | +-----+
| | | +-- Leoparreysia_whitteni
| +-----+
| | +----- Leoparreysia_tavoyensis
| +--+
--+ | +-- Leoparreysia_olivacea
| +--+
| +-- Leoparreysia_canefrii
|
| +-- Coelatura_aegyptiaca
| +-----------------------------+
| | +-- Nitia_teretiuscula
| |
| | +----------- Indonaia_shurtleffiana **
| | +-----------------+
+--+ | | +-------- Indonaia_rugosa **
| | +--+
| | | +----- Indonaia_andersoniana **
| | +--+
| | | +-- Indonaia_subclathrata **
+--+ +--+
| +-- Indonaia_caerulea **
|
| +-------------------------- Indochinella_pugio **
| |
| | +-------- Radiatula_crispisulcata **
| | +--------------+
+--+ | | +----- Radiatula_mouhoti **
| | +--+
| | | +-- Radiatula_chaudhurii **
| | +--+
+--+ +-- Radiatula_myitkyinae **
|
| +-------------------- Scabiellus_songkramensis **
| |
| | +-- Unionetta_fabagina **
+--+ +--------------+
| | +-- Harmandia_somboriensis **
| |
+--+ +-------------- Scabies_nucleus **
| |
| | +-- Scabies_crispata **
+--+ +--------+
| | +-- Scabies_mandarinus **
| |
+--+ +-- Scabies_humilis **
| +-----+
| | +-- Scabies_pilata **
+--+
| +----- Scabies_anceps **
+--+
| +-- Scabies_phaselus **
+--+
+-- Scabies_scobinatus **
Bolotov et al., 2020 [Publ 4100], fig. S1
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 20 |
20 |
17 |
7 |
94 |
20 |
4 |
14.0 |
+----------- Parreysia_keralaensis
+--+
| | +-------- Parreysia_nagpoorensis[2]
| +--+
| | +----- Parreysia_nagpoorensis[1]
| +--+
| | +-- Parreysia_rakhinensis
+--------------------------+ +--+
| | +-- Parreysia_plagiosoma
| |
| | +-- Leoparreysia_burmana
| | +-----+
| | | +-- Leoparreysia_whitteni
| +-----+
| | +----- Leoparreysia_olivacea
| +--+
--+ | +-- Leoparreysia_tavoyensis
| +--+
| +-- Leoparreysia_canefrii
|
| +-- Coelatura_aegyptiaca
| +-----------------------------------+
| | +-- Nitia_teretiuscula
| |
| | +-------- Indonaia_shurtleffiana **
+--+ +--------------------------+
| | | +----- Indonaia_andersoniana **
| | +--+
| | | +-- Indonaia_caerulea **
| | +--+
+--+ +-- Indonaia_subclathrata **
|
| +-------------------------------- Indochinella_pugio **
| |
| | +----- Radiatula_mouhoti **
+--+ +-----------------------+
| | | +-- Radiatula_myitkyinae **
| | +--+
+--+ +-- Radiatula_chaudhurii **
|
| +-------------------------- Radiatula_crispisulcata **
| |
+--+ +----------------------- Scabiellus_songkramensis **
| |
+--+ +-------------------- Harmandia_somboriensis **
| |
+--+ +----------------- Unionetta_fabagina **
| |
+--+ +-------------- Scabies_mandarinus **
| |
+--+ +----------- Scabies_crispata **
| |
+--+ +-------- Scabies_nucleus **
| |
+--+ +-- Scabies_humilis **
| +--+
| | +-- Scabies_pilata **
+--+
| +-- Scabies_phaselus **
| |
+--+-- Scabies_scobinatus **
|
+-- Scabies_anceps **
Pfeiffer et al., 2018 [Publ 3827], fig. 2
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 18 |
18 |
15 |
7 |
11 |
10 |
4 |
14.0 |
+-- Leoparreysia_tavoyensis
+-----------------------------------+
| +-- Leoparreysia_olivacea
|
| +----- Nitia_teretiuscula
--+ +-----------------------------+
| | | +-- Coelatura_aegyptiaca
| | +--+
| | +-- Prisodontopsis_aviculaeformis
+--+
| +----- Indonaia_bonneaudii **
| +--------------------------+
| | | +-- Indonaia_andersoniana **
| | +--+
+--+ +-- Indonaia_caerulea **
|
| +----------------------------- Indochinella_pugio **
| |
+--+ +-- Radiatula_chaudhurii **
| +-----------------------+
| | +-- Radiatula_myitkyinae **
+--+
| +----------------------- Scabiellus_songkramensis **
| |
+--+ +-------------------- Unionetta_fabagina[2] **
| |
+--+ +----------------- Harmandia_somboriensis **
| |
+--+ +-------------- Unionetta_fabagina[1] **
| |
+--+ +-- Scabies_crispata **
| +--------+
| | +-- Scabies_mandarinus **
+--+
| +-------- Scabies_nucleus **
| |
+--+ +-- Scabies_pilata **
| +--+
| | +-- Scabies_humilis **
+--+
| +-- Scabies_phaselus **
| |
+--+-- Scabies_anceps **
|
+-- Scabies_scobinatus **
Jeratthitikul et al., 2019 [Publ 3922], fig. 1
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 18 |
18 |
16 |
7 |
5 |
5 |
4 |
19.0 |
+-------------------------------------------- Parreysia_keralaensis
|
--+ +----------------------------------------- Coelatura_aegyptiaca
| |
| | +----- Indonaia_rugosa **
+--+ +--------------------------------+
| | | +-- Indonaia_caerulea **
| | +--+
| | +-- Indonaia_andersoniana **
+--+
| +----------------------------------- Indochinella_pugio **
| |
| | +----- Radiatula_mouhoti **
+--+ +--------------------------+
| | | +-- Radiatula_myitkyinae **
| | +--+
+--+ +-- Radiatula_chaudhurii **
|
| +----------------------------- Scabiellus_songkramensis **
+--+
| +-------------------------- Harmandia_somboriensis **
+--+
| +----------------------- Unionetta_fabagina **
+--+
| +-------------------- Scabies_crispata **
+--+
| +----------------- Scabies_mandarinus **
+--+
| +-------------- Scabies_pilata **
+--+
| +----------- Scabies_nucleus **
+--+
| +-------- Scabies_humilis **
+--+
| +----- Scabies_scobinatus **
+--+
| +-- Scabies_phaselus **
+--+
+-- Scabies_anceps **
Konopleva et al., 2023 [Publ 4262], fig. 8
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 14 |
14 |
10 |
7 |
70 |
0 |
0 |
21.0 |
+----------------------- Indonaia_andersoniana **
|
--+ +-------------------- Indochinella_pugio **
| |
+--+ +----------------- Radiatula_mouhoti **
| |
+--+ +-------------- Scabiellus_songkramensis **
| |
| | +-- Unionetta_fabagina **
+--+ +--------+
| | +-- Harmandia_somboriensis **
| |
| | +-- Scabies_humilis **
+--+ +-----+
| | +-- Scabies_pilata **
| |
| | +----- Scabies_crispata **
| +--+
+--+ | +-- Scabies_nucleus **
| +--+
| +-- Scabies_mandarinus **
|
| +-- Scabies_phaselus **
| |
+-----+-- Scabies_anceps **
|
+-- Scabies_scobinatus **
Bolotov et al., 2017 [Publ 3764], fig. S1
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 12 |
13 |
9 |
4 |
119 |
29 |
4 |
22.0 |
+-------- Leoparreysia_olivacea
+-----------+
| | +----- Leoparreysia_tavoyensis[2]
| +--+
| | +-- Leoparreysia_canefrii
| +--+
| +-- Leoparreysia_tavoyensis[1]
|
| +-- Nitia_teretiuscula
| |
| +-- Coelatura_gabonensis
| |
| +-- Prisodontopsis_mweruensis
--+ |
| +-- Coelatura_kunenensis
| |
| +--------------+-- Coelatura_aegyptiaca[1]
| | |
| | +-- Coelatura_aegyptiaca[2]
| | |
| | +-- Prisodontopsis_aviculaeformis
| | |
| | +-- Coelatura_luapulaensis
| | |
| | +-- Coelatura_choziensis
+--+
| +-------- Indonaia_shurtleffiana **
| |
| +-----+ +----- Indonaia_rugosa **
| | | |
| | | | +-- Indonaia_andersoniana **
| | +--+--+
| | | +-- Indonaia_caerulea **
| | |
+--+ | +-- Indonaia_bonneaudii **
| +--+
| +-- Lamellidens_corrianus[2]
|
| +----------- Indochinella_pugio **
| |
| | +-- Scabies_humilis **
+--+ |
| +-----+-- Scabies_phaselus **
| | |
+--+ +-- Scabies_mandarinus **
|
| +----- Radiatula_mouhoti **
+--+
| +-- Radiatula_myitkyinae **
+--+
+-- Radiatula_chaudhurii **
Huang et al., 2019 [Publ 3843], fig. 2
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 12 |
12 |
9 |
4 |
192 |
24 |
4 |
24.0 |
+-- Leoparreysia_tavoyensis
|
+-----------------+-- Leoparreysia_canefrii
| |
| +-- Leoparreysia_olivacea
|
| +-------- Coelatura_aegyptiaca[2]
| |
| +--------+ +----- Coelatura_aegyptiaca[1]
--+ | | |
| | | +----- Nitia_teretiuscula
| | +--+
| | | +-- Coelatura_gabonensis
| | | |
| | +--+-- Coelatura_kunenensis
| | |
+--+ +-- Prisodontopsis_mweruensis
|
| +-------- Indonaia_shurtleffiana **
| |
| +-----+ +----- Indonaia_bonneaudii **
| | | |
| | +--+----- Indonaia_rugosa **
| | |
| | | +-- Indonaia_andersoniana **
+--+ +--+
| +-- Indonaia_caerulea **
|
| +----------- Indochinella_pugio **
| |
| | +----- Scabies_humilis **
+--+ +--+
| | | +-- Scabies_phaselus **
| | +--+
+--+ +-- Scabies_mandarinus **
|
| +----- Radiatula_myitkyinae **
+--+
| +-- Radiatula_chaudhurii **
+--+
+-- Radiatula_mouhoti **
Huang et al., 2019 [Publ 3843], fig. S3
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 12 |
12 |
9 |
4 |
189 |
24 |
4 |
22.0 |
+----- Leoparreysia_canefrii
+--------------+
| | +-- Leoparreysia_tavoyensis
| +--+
| +-- Leoparreysia_olivacea
|
| +----------- Coelatura_aegyptiaca[2]
| |
--+ +-----+ +-- Coelatura_aegyptiaca[1]
| | | +-----+
| | | | +-- Nitia_teretiuscula
| | +--+
| | | +----- Coelatura_kunenensis
| | +--+
| | | +-- Coelatura_gabonensis
| | +--+
+--+ +-- Prisodontopsis_mweruensis
|
| +-------- Indonaia_shurtleffiana **
| |
| +-----+ +----- Indonaia_bonneaudii **
| | | |
| | +--+----- Indonaia_rugosa **
| | |
| | | +-- Indonaia_andersoniana **
+--+ +--+
| +-- Indonaia_caerulea **
|
| +----------- Indochinella_pugio **
| |
| | +----- Scabies_phaselus **
+--+ +--+
| | | +-- Scabies_humilis **
| | +--+
+--+ +-- Scabies_mandarinus **
|
| +----- Radiatula_mouhoti **
+--+
| +-- Radiatula_myitkyinae **
+--+
+-- Radiatula_chaudhurii **
Huang et al., 2019 [Publ 3843], fig. S4
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 12 |
12 |
9 |
4 |
190 |
24 |
4 |
22.0 |
+----- Leoparreysia_tavoyensis
+--------------+
| | +-- Leoparreysia_canefrii
| +--+
| +-- Leoparreysia_olivacea
|
| +-------- Coelatura_aegyptiaca[2]
| |
| +-------- Nitia_teretiuscula
--+ +--------+
| | | +----- Coelatura_aegyptiaca[1]
| | | |
| | +--+ +-- Coelatura_gabonensis
| | | |
| | +--+-- Coelatura_kunenensis
| | |
| | +-- Prisodontopsis_mweruensis
+--+
| +----- Indonaia_shurtleffiana **
| |
| | +-- Indonaia_bonneaudii **
| +--------+--+
| | | +-- Indonaia_rugosa **
| | |
| | | +-- Indonaia_andersoniana **
| | +--+
+--+ +-- Indonaia_caerulea **
|
| +----------- Indochinella_pugio **
| |
| | +----- Scabies_humilis **
+--+ +--+
| | | +-- Scabies_phaselus **
| | +--+
+--+ +-- Scabies_mandarinus **
|
| +----- Radiatula_mouhoti **
+--+
| +-- Radiatula_myitkyinae **
+--+
+-- Radiatula_chaudhurii **
Konopleva et al., 2019 [Publ 3900], fig. 1
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 12 |
12 |
9 |
4 |
40 |
0 |
0 |
20.0 |
+----------- Indonaia_shurtleffiana **
+--+
| | +-------- Indonaia_rugosa **
| +--+
| | +----- Indonaia_andersoniana **
| +--+
| | +-- Indonaia_caerulea **
--+ +--+
| +-- Indonaia_subclathrata **
|
| +----------- Indochinella_pugio **
| |
| | +-- Scabies_humilis **
+--+ |
| +-----+-- Scabies_mandarinus **
| | |
+--+ +-- Scabies_phaselus **
|
| +----- Radiatula_mouhoti **
+--+
| +-- Radiatula_myitkyinae **
+--+
+-- Radiatula_chaudhurii **
Bolotov et al., 2022 [Publ 4250], fig. S2
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 12 |
12 |
10 |
3 |
3 |
3 |
2 |
19.0 |
+----- Parreysia_rakhinensis
+--------------+
| | +-- Leoparreysia_tavoyensis
| +--+
| +-- Leoparreysia_olivacea
|
--+ +----------- Indonaia_rugosa **
| +-----+
| | | +-------- Indonaia_shurtleffiana **
| | +--+
| | | +----- Indonaia_andersoniana **
| | +--+
+--+ | +-- Indonaia_subclathrata **
| +--+
| +-- Indonaia_caerulea **
|
| +-------------- Indochinella_pugio **
| |
+--+ +----------- Radiatula_crispisulcata **
| |
+--+ +-- Radiatula_chaudhurii **
| +-----+
| | +-- Radiatula_myitkyinae **
+--+
| +----- Radiatula_chindwinensis **
+--+
| +-- Radiatula_mouhoti **
+--+
+-- Radiatula_myitthanensis **
Bolotov et al., 2017 [Publ 3761], fig. S1
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 11 |
11 |
9 |
4 |
130 |
18 |
4 |
26.0 |
+-- Leoparreysia_tavoyensis
+-----------------+
| +-- Leoparreysia_olivacea
|
| +-------------- Nitia_teretiuscula
| +--+
| | | +----------- Coelatura_aegyptiaca[2]
--+ | +--+
| | | +-------- Coelatura_aegyptiaca[1]
| | +--+
| | | +----- Prisodontopsis_mweruensis
| | +--+
| | | +-- Coelatura_gabonensis
+--+ +--+
| +-- Coelatura_kunenensis
|
| +-------- Indonaia_shurtleffiana **
| +-----+
| | | +----- Indonaia_andersoniana **
| | +--+
| | | +-- Indonaia_rugosa **
| | +--+
+--+ +-- Indonaia_bonneaudii **
|
| +-- Indochinella_pugio **
| +--------+
| | +-- Scabies_crispata **
| |
+--+ +-- Radiatula_myitkyinae **
| +-----+
| | +-- Radiatula_chaudhurii **
+--+
| +----- Scabies_mandarinus **
+--+
| +-- Scabies_humilis **
+--+
+-- Scabies_phaselus **
Bolotov et al., 2017 [Publ 3761], fig. S2
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 11 |
11 |
9 |
4 |
130 |
18 |
4 |
26.0 |
+-- Leoparreysia_olivacea
+-----------------+
| +-- Leoparreysia_tavoyensis
|
| +-------------- Nitia_teretiuscula
| +--+
| | | +----------- Coelatura_aegyptiaca[1]
--+ | +--+
| | | +-------- Coelatura_aegyptiaca[2]
| | +--+
| | | +----- Coelatura_gabonensis
| | +--+
| | | +-- Coelatura_kunenensis
+--+ +--+
| +-- Prisodontopsis_mweruensis
|
| +-------- Indonaia_shurtleffiana **
| +-----+
| | | +----- Indonaia_andersoniana **
| | +--+
| | | +-- Indonaia_rugosa **
| | +--+
+--+ +-- Indonaia_bonneaudii **
|
| +-- Indochinella_pugio **
| +--------+
| | +-- Scabies_crispata **
| |
+--+ +-- Radiatula_myitkyinae **
| +-----+
| | +-- Radiatula_chaudhurii **
+--+
| +----- Scabies_mandarinus **
+--+
| +-- Scabies_humilis **
+--+
+-- Scabies_phaselus **
Bolotov et al., 2018 [Publ 3826], fig. 2
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 11 |
11 |
9 |
4 |
74 |
13 |
4 |
22.0 |
+----- Leoparreysia_olivacea
+--------------+
| | +-- Leoparreysia_canefrii
| +--+
| +-- Leoparreysia_tavoyensis
|
--+ +-- Coelatura_aegyptiaca
| +--------------+
| | +-- Nitia_teretiuscula
| |
| | +-------- Indonaia_shurtleffiana **
+--+ +-----+
| | | +----- Indonaia_rugosa **
| | +--+
| | | +-- Indonaia_caerulea **
| | +--+
+--+ +-- Indonaia_andersoniana **
|
| +----------- Indochinella_pugio **
| |
| | +----- Radiatula_myitkyinae **
+--+ +--+
| | | +-- Radiatula_mouhoti **
| | +--+
+--+ +-- Radiatula_chaudhurii **
|
| +----- Scabies_mandarinus **
+--+
| +-- Scabies_humilis **
+--+
+-- Scabies_phaselus **
Bolotov et al., 2022 [Publ 4250], fig. 8
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 9 |
9 |
7 |
3 |
40 |
15 |
3 |
20.0 |
+----------------------- Parreysia_rakhinensis
|
| +-- Leoparreysia_whitteni
| +-----+
| | +-- Leoparreysia_burmana
--+ +-----------+
| | | +----- Leoparreysia_olivacea
| | +--+
| | | +-- Leoparreysia_canefrii
| | +--+
+--+ +-- Leoparreysia_tavoyensis
|
| +-- Indonaia_andersoniana **
| +--------------+
| | +-- Indonaia_subclathrata **
+--+
| +-------------- Indochinella_pugio **
| |
+--+ +----------- Radiatula_crispisulcata **
| |
+--+ +-- Radiatula_chaudhurii **
| +-----+
| | +-- Radiatula_myitkyinae **
+--+
| +----- Radiatula_mouhoti **
+--+
| +-- Radiatula_chindwinensis **
+--+
+-- Radiatula_myitthanensis **
Ortiz-Sepulveda et al., 2020 [Publ 4101], fig. 1
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 8 |
8 |
3 |
4 |
28 |
28 |
2 |
22.0 |
+-------------------------------------- Leoparreysia_tavoyensis
|
| +----------- Indonaia_andersoniana **
| |
| +-----------------------+ +-------- Indochinella_pugio **
| | | |
--+ | | | +----- Radiatula_mouhoti **
| | +--+ |
| | | +----- Radiatula_myitkyinae **
| | | |
| | +--+----- Radiatula_chaudhurii **
| | |
| | | +-- Scabies_humilis **
+--+ | |
| +--+-- Scabies_phaselus **
| |
| +-- Scabies_mandarinus **
|
| +-------------------------------- Grandidieria_burtoni
| |
| | +-- Pseudospatha_tanganyicensis
| | +--------------------------+
+--+ | +-- Moncetia_anceyi
| |
| | +-- Nitia_acuminata
| | |
+--+ +-----------------------+-- Nitia_monceti
| | |
| | +-- Nitia_teretiuscula
| |
| | +-- Coelatura_aegyptiaca
| | |
+--+ +-- Coelatura_alluaudi
| |
| +--------------------+-- Coelatura_bakeri
| | |
| | +-- Coelatura_hauttecoeuri
| | |
| | +-- Coelatura_stuhlmanni
+--+
| +-- Coelatura_leopoldvillensis
| +-----------------+
| | +-- Coelatura_mesafricana
| |
| | +-- Coelatura_horei
+--+ |
| +--------------+-- Coelatura_ratidota
| | |
| | +-- Coelatura
+--+
| +-------------- Coelatura_gabonensis
| |
| +-------------- Coelatura_stanleyvillensis
+--+
| +----------- Coelatura_kunenensis
| |
| | +----- Prisodontopsis_aviculaeformis
+--+ +--+
| | | +-- Coelatura_briarti
| | +--+
| | +-- Prisodontopsis_mweruensis
+--+
| +-- Coelatura_choziensis
| |
| +-- Coelatura_hypsiprymna
| |
+-----+-- Coelatura_luapulaensis
|
+-- Coelatura_mossambicensis
|
+-- Nyassunio_nyassaensis
Ortiz-Sepulveda et al., 2020 [Publ 4101], fig. 2
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 8 |
8 |
1 |
4 |
28 |
28 |
2 |
24.0 |
+-------------------------------- Leoparreysia_tavoyensis
|
| +----- Radiatula_mouhoti **
| |
| +----- Radiatula_myitkyinae **
| |
| +----- Radiatula_chaudhurii **
| |
--+ +-----------------------+----- Indochinella_pugio **
| | |
| | +----- Indonaia_andersoniana **
| | |
| | | +-- Scabies_humilis **
| | | |
| | +--+-- Scabies_phaselus **
| | |
+--+ +-- Scabies_mandarinus **
|
| +----- Grandidieria_burtoni
| +--------------------+
| | | +-- Pseudospatha_tanganyicensis
| | +--+
| | +-- Moncetia_anceyi
| |
| | +-- Nitia_acuminata
+--+ |
| +--------------------+-- Nitia_monceti
| | |
| | +-- Nitia_teretiuscula
| |
| | +-- Coelatura_aegyptiaca
| | |
+--+ +-- Coelatura_alluaudi
| |
| +-----------------+-- Coelatura_bakeri
| | |
| | +-- Coelatura_hauttecoeuri
| | |
| | +-- Coelatura_stuhlmanni
+--+
| +-- Coelatura_leopoldvillensis
| +--------------+
| | +-- Coelatura_mesafricana
| |
| | +-- Coelatura_horei
+--+ |
| +-----------+-- Coelatura_ratidota
| | |
| | +-- Coelatura
| |
+--+ +-- Coelatura_gabonensis
| +--------+
| | +-- Coelatura_stanleyvillensis
| |
+--+ +-------- Coelatura_kunenensis
| |
| | +-- Coelatura_briarti
| | |
+--+ +--+-- Prisodontopsis_mweruensis
| | |
| | +-- Prisodontopsis_aviculaeformis
| |
+--+ +-- Coelatura_choziensis
| |
| +-- Coelatura_hypsiprymna
| |
+--+-- Coelatura_luapulaensis
|
+-- Coelatura_mossambicensis
|
+-- Nyassunio_nyassaensis
Bolotov et al., 2019 [Publ 3974], fig. 1
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 6 |
6 |
4 |
2 |
12 |
7 |
2 |
21.0 |
+-- Leoparreysia_whitteni
+-----+
| +-- Leoparreysia_burmana
+--------+
| | +----- Leoparreysia_olivacea
| +--+
| | +-- Leoparreysia_tavoyensis
| +--+
--+ +-- Leoparreysia_canefrii
|
| +-- Nitia_teretiuscula
| +-----------+
| | +-- Coelatura_aegyptiaca
+--+
| +----------- Indochinella_pugio **
| |
+--+ +-- Radiatula_myitkyinae **
| +-----+
| | +-- Radiatula_chaudhurii **
+--+
| +----- Radiatula_chindwinensis **
+--+
| +-- Radiatula_mouhoti **
+--+
+-- Radiatula_myitthanensis **
Pfeiffer et al., 2019 [Publ 3937], fig. 4-5
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 4 |
4 |
2 |
4 |
53 |
6 |
3 |
23.0 |
+-- Leoparreysia_tavoyensis
+-----------+
| +-- Leoparreysia_olivacea
|
--+ +-- Coelatura_choziensis
| +--------+
| | +-- Prisodontopsis_aviculaeformis
+--+
| +-------- Indonaia_caerulea **
+--+
| +----- Indochinella_pugio **
+--+
| +-- Scabies_phaselus **
+--+
+-- Harmandia_somboriensis **
Pfeiffer et al., 2019 [Publ 3937], fig. 6
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 4 |
4 |
2 |
4 |
53 |
6 |
3 |
23.0 |
+-- Leoparreysia_tavoyensis
+-----------+
| +-- Leoparreysia_olivacea
|
--+ +-- Coelatura_choziensis
| +--------+
| | +-- Prisodontopsis_aviculaeformis
+--+
| +-------- Indonaia_caerulea **
+--+
| +----- Indochinella_pugio **
+--+
| +-- Scabies_phaselus **
+--+
+-- Harmandia_somboriensis **
Pfeiffer & Graf, 2015 [Publ 3620], fig. 2
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 3 |
3 |
1 |
3 |
37 |
5 |
3 |
24.0 |
+-- Leoparreysia_olivacea
+--------+
| +-- Leoparreysia_tavoyensis
--+
| +-------- Coelatura_aegyptiaca
+--+
| +----- Indochinella_pugio **
+--+
| +-- Radiatula_chaudhurii **
+--+
+-- Scabies_mandarinus **
Konopleva et al., 2017 [Publ 3807], fig. 6
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 3 |
3 |
1 |
3 |
28 |
8 |
3 |
24.0 |
+-- Leoparreysia_tavoyensis
+--------+
| +-- Leoparreysia_olivacea
--+
| +-------- Coelatura_aegyptiaca
+--+
| +----- Indochinella_pugio **
+--+
| +-- Radiatula_chaudhurii **
+--+
+-- Scabies_mandarinus **
Konopleva et al., 2023 [Publ 4269], fig. S1, 4
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 3 |
3 |
1 |
3 |
53 |
4 |
3 |
24.0 |
+----------- Parreysia_rakhinensis
--+
| +-------- Leoparreysia_tavoyensis
+--+
| +----- Indonaia_andersoniana **
+--+
| +-- Indochinella_pugio **
+--+
+-- Radiatula_mouhoti **
Whelan et al., 2011 [Publ 3380], fig. 1 ML
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 2 |
2 |
0 |
2 |
35 |
8 |
3 |
-- |
+----------------- Leoparreysia_tavoyensis
|
--+ +-- Radiatula_chaudhurii **
| +-----------+
| | +-- Indochinella_pugio **
+--+
| +----------- Nitia_teretiuscula
+--+
| +-------- Coelatura_aegyptiaca[2]
+--+
| +----- Coelatura_aegyptiaca[1]
+--+
| +-- Coelatura_gabonensis
+--+
+-- Prisodontopsis_aviculaeformis
Whelan et al., 2011 [Publ 3380], fig. 1 MP
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 2 |
2 |
0 |
2 |
35 |
8 |
3 |
-- |
+-------------- Leoparreysia_tavoyensis
|
--+ +-- Radiatula_chaudhurii **
| +--------+
| | +-- Indochinella_pugio **
+--+
| +-------- Nitia_teretiuscula
| |
+--+ +----- Coelatura_aegyptiaca[1]
| |
+--+----- Coelatura_aegyptiaca[2]
|
| +-- Coelatura_gabonensis
+--+
+-- Prisodontopsis_aviculaeformis
Lopes-Lima et al., 2017 [Publ 3745], fig. 1-2
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 1 |
1 |
0 |
1 |
80 |
3 |
3 |
-- |
+----- Leoparreysia_tavoyensis
--+
| +-- Coelatura_aegyptiaca
+--+
+-- Indochinella_pugio **
Combosch et al., 2017 [Publ 3806], fig. 1
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 1 |
1 |
0 |
1 |
14 |
0 |
0 |
-- |
-- Scabies_crispata
Combosch et al., 2017 [Publ 3806], fig. 2
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 1 |
1 |
0 |
1 |
14 |
0 |
0 |
-- |
-- Scabies_crispata
Huang et al., 2018 [Publ 3828], fig. 3, 5
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 1 |
1 |
0 |
1 |
28 |
0 |
0 |
-- |
-- Scabies_crispata
Jeratthitikul et al., 2019 [Publ 3911], fig. 4
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 1 |
1 |
0 |
1 |
21 |
0 |
0 |
-- |
-- Scabies_crispata
Lopes-Lima et al., 2020 [Publ 4080], fig. 2-4
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 1 |
1 |
0 |
1 |
121 |
2 |
2 |
-- |
+-- Coelatura_aegyptiaca
|
--+-- Leoparreysia_tavoyensis
|
+-- Radiatula_mouhoti **
Wu et al., 2020 [Publ 4121], fig. 2
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 1 |
1 |
0 |
1 |
33 |
0 |
0 |
-- |
-- Scabies_crispata
Wu et al., 2020 [Publ 4121], fig. S1
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 1 |
1 |
0 |
1 |
33 |
0 |
0 |
-- |
-- Scabies_crispata
Sano et al., 2020 [Publ 4138], fig. 2
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 1 |
1 |
0 |
1 |
72 |
0 |
0 |
-- |
-- Scabies_crispata
Sano et al., 2020 [Publ 4138], fig. 3
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 1 |
1 |
0 |
1 |
71 |
0 |
0 |
-- |
-- Scabies_crispata
Sano et al., 2020 [Publ 4138], fig. 4
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 1 |
1 |
0 |
1 |
71 |
0 |
0 |
-- |
-- Scabies_crispata
Choi et al., 2020 [Publ 4169], fig. 5
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 1 |
1 |
0 |
1 |
8 |
2 |
2 |
-- |
+----- Radiatula_mouhoti **
--+
| +-- Lamellidens_generosus
+--+
+-- Leoparreysia_canefrii
Wu et al., 2021 [Publ 4215], fig. 3
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 1 |
1 |
0 |
1 |
23 |
0 |
0 |
-- |
-- Indonaia_andersoniana
Chen et al., 2023 [Publ 4302], fig. 1
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 1 |
1 |
0 |
1 |
39 |
2 |
2 |
-- |
+-- Coelatura_aegyptiaca
|
--+-- Indonaia_andersoniana **
|
+-- Parreysia_nagpoorensis
Chen et al., 2023 [Publ 4302], fig. 2
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 1 |
1 |
0 |
1 |
39 |
2 |
2 |
-- |
+----- Parreysia_nagpoorensis
--+
| +-- Coelatura_aegyptiaca
+--+
+-- Indonaia_andersoniana **
Smith, 2021 [Publ 4331], fig. 1
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 1 |
1 |
0 |
1 |
19 |
0 |
0 |
-- |
-- Scabies_phaselus
| 
