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Page last updated
Thu 04 Apr 2024

The Freshwater Mussels (Unionoida) of the World (and other less consequential bivalves)

FM(U)otW(aolcb) is the web version of the MUSSEL Project Database. Follow the links to browse the data or use the custom Google search field. Either way, you win!

Cladomics pages are experimental — even more so than the rest of this site!

The purpose of the cladomics pages for each genus and family-group level taxon is to provide statistics and references indicating the extent to which published phylogenetic work supports the classification used here.

The Cladomics Summary provides an overview of the number of studies that have included terminal species representing this taxon as well as an indication of how frequently it has been recovered as monophyletic.

Following the summary, the available published clades representing the taxon are listed.

For more information, visit the Cladomics FARQ (frequently asked rhetorical questions) page. Abbreviations for the various statistics are explained on the FARQ!

Search the database pages:

Bivalvia | Unionoida | Unionoidea | Unionidae | Unioninae | Unionini

genus Aculamprotula Wu, Liang, Wang & Ouyang, 1999: cladomics
     9 valid species

Cladomics Summary

number of species = 9

analyzed in 86 cladograms in 51 publications
maximum number of terminals in an analysis = 5, mean = 2.52

Basic Monophyly

number of cladograms that test basic monophyly (i.e., more than 1 included species) = 49
maximum number of included terminals = 5, mean = 3.67
proportion of cladograms that support basic monophyly = 1.0

Pairwise Cladogram Clustering

The graphs below depict in two dimensions all pair-wise ingroup distances within the cladome. See the Cladmoics FARQ for more information.


PCoA Ordination


NJ tree

Lopes-Lima et al., 2020 [Publ 4080], fig. 2-4

iT tT bi riS oT roT roS iD
5 5 3 5 117 21 7 0.0

*** Top Monophyly Cladogram *** 

                 +----------- Aculamprotula_tortuosa **     
  +--------------+                                       
  |              |  +-------- Aculamprotula_scripta **      
  |              +--+                                    
  |                 |  +----- Aculamprotula_coreana **      
  |                 +--+                                 
  |                    |  +-- Aculamprotula_fibrosa **      
  |                    +--+                              
--+                       +-- Aculamprotula_tientsinensis **
  |                                                      
  |  +----------------------- Middendorffinaia_mongolica 
  |  |                                                   
  |  |                 +----- Unio_gibbus                
  |  |                 |                                 
  |  |  +--------------+----- Unio_tumidus               
  +--+  |              |                                 
     |  |              |  +-- Unio_crassus               
     |  |              +--+                              
     |  |                 +-- Unio_pictorum              
     |  |                                                
     |  |     +-------------- Nodularia_breviconcha      
     +--+  +--+                                          
        |  |  |  +----------- Nodularia[2]               
        |  |  +--+                                       
        |  |     |  +-------- Nodularia_nuxpersicae      
        |  |     +--+                                    
        |  |        |  +----- Nodularia_nipponensis      
        |  |        +--+                                 
        |  |           |  +-- Nodularia_douglasiae       
        +--+           +--+                              
           |              +-- Nodularia[1]               
           |                                             
           |              +-- Schistodesmus_lampreyanus  
           |  +-----------+                              
           |  |           +-- Schistodesmus_spinosus     
           |  |                                          
           |  |        +----- Cuneopsis_rufescens        
           |  +--------+                                 
           +--+        |  +-- Cuneopsis_heudei           
              |        +--+                              
              |           +-- Cuneopsis_celtiformis      
              |                                          
              |  +----------- Tchangsinaia_piscicula     
              +--+                                       
                 |  +-------- Inversiunio_jokohamensis   
                 +--+                                    
                    |  +----- Inversiunio_verrucosus     
                    +--+                                 
                       |  +-- Inversiunio_reinianus      
                       +--+                              
                          +-- Inversiunio_yanagawensis

Chen et al., 2023 [Publ 4302], fig. 1

iT tT bi riS oT roT roS iD
1 1 0 1 39 16 9 --

*** Top Sister-Group Cladogram ***

  +-------------------- Aculamprotula_fibrosa **       
  |                                                 
  |              +----- Middendorffinaia_mongolica  
  |  +-----------+                                  
--+  |           |  +-- Nodularia_douglasiae        
  |  |           +--+                               
  |  |              +-- Nodularia_breviconcha       
  |  |                                              
  |  |              +-- Cuneopsis_celtiformis       
  +--+  +-----------+                               
     |  |           +-- Cuneopsis_heudei            
     |  |                                           
     |  |           +-- Diaurora_aurorea            
     |  |        +--+                               
     |  |        |  +-- Diaurora_laeve              
     |  +--------+                                  
     +--+        |  +-- Schistodesmus_lampreyanus   
        |        +--+                               
        |           +-- Schistodesmus_spinosus      
        |                                           
        |           +-- Pseudocuneopsis_sichuanensis
        |  +--------+                               
        |  |        +-- Pseudocuneopsis_capitata    
        |  |                                        
        +--+        +-- Inversiunio_yanagawensis    
           |  +-----+                               
           |  |     +-- Inversiunio_reinianus       
           +--+                                     
              |  +----- Tchangsinaia_piscicula      
              +--+                                  
                 |  +-- Unio_pictorum               
                 +--+                               
                    +-- Unio_crassus

Zhou et al., 2007 [Publ 2658], fig. 1

iT tT bi riS oT roT roS iD
5 5 3 5 18 4 3 2.0 
              +-- Tchangsinaia_piscicula     
  +-----------+                              
  |           +-- Nodularia_douglasiae       
--+                                          
  |  +----------- Aculamprotula_tortuosa **     
  +--+                                       
     |  +-------- Aculamprotula_scripta **      
     +--+                                    
        |  +----- Aculamprotula_tientsinensis **
        +--+                                 
           |  +-- Aculamprotula_zonata **       
           +--+                              
              +-- Aculamprotula_fibrosa **

Zhou et al., 2007 [Publ 2658], fig. 2

iT tT bi riS oT roT roS iD
5 5 3 5 16 2 2 2.0 
  +----------- Aculamprotula_tortuosa **     
--+                                       
  |  +-------- Aculamprotula_scripta **      
  +--+                                    
     |  +----- Aculamprotula_fibrosa **      
     +--+                                 
        |  +-- Aculamprotula_zonata **       
        +--+                              
           +-- Aculamprotula_tientsinensis **

Pfeiffer & Graf, 2013 [Publ 3531], fig. 2

iT tT bi riS oT roT roS iD
5 5 2 5 30 2 2 2.0 
           +-- Unio_pictorum              
  +--------+                              
  |        +-- Nodularia_douglasiae       
--+                                       
  |  +-------- Aculamprotula_tortuosa **     
  |  |                                    
  +--+  +----- Aculamprotula_scripta **      
     |  |                                 
     +--+  +-- Aculamprotula_tientsinensis **
        |  |                              
        +--+-- Aculamprotula_fibrosa **      
           |                              
           +-- Aculamprotula_zonata **

Wu et al., 2018 [Publ 3836], fig. 2

iT tT bi riS oT roT roS iD
5 5 3 5 48 9 5 0.0 
        +----------- Aculamprotula_tortuosa **     
  +-----+                                       
  |     |  +-------- Aculamprotula_scripta **      
  |     +--+                                    
  |        |  +----- Aculamprotula_coreana **      
  |        +--+                                 
  |           |  +-- Aculamprotula_fibrosa **      
--+           +--+                              
  |              +-- Aculamprotula_tientsinensis **
  |                                             
  |              +-- Unio_crassus               
  |  +-----------+                              
  |  |           +-- Unio_pictorum              
  |  |                                          
  +--+        +----- Nodularia_douglasiae       
     |  +-----+                                 
     |  |     |  +-- Schistodesmus_lampreyanus  
     |  |     +--+                              
     +--+        +-- Schistodesmus_spinosus     
        |                                       
        |  +-------- Tchangsinaia_piscicula     
        +--+                                    
           |  +----- Cuneopsis_rufescens        
           +--+                                 
              |  +-- Cuneopsis_celtiformis      
              +--+                              
                 +-- Cuneopsis_heudei

Huang et al., 2019 [Publ 3843], fig. 1

iT tT bi riS oT roT roS iD
5 5 3 5 39 9 4 0.0 
  +----------- Aculamprotula_tortuosa **     
--+                                       
  |  +-------- Aculamprotula_scripta **      
  +--+                                    
     |  +----- Aculamprotula_coreana **      
     +--+                                 
        |  +-- Aculamprotula_fibrosa **      
        +--+                              
           +-- Aculamprotula_tientsinensis **

Huang et al., 2019 [Publ 3843], fig. 2

iT tT bi riS oT roT roS iD
5 5 2 5 199 13 5 1.0 
              +-------- Aculamprotula_tortuosa **     
              |                                    
  +-----------+  +----- Aculamprotula_coreana **      
  |           |  |                                 
  |           +--+----- Aculamprotula_scripta **      
  |              |                                 
  |              |  +-- Aculamprotula_fibrosa **      
  |              +--+                              
  |                 +-- Aculamprotula_tientsinensis **
--+                                                
  |           +-------- Unio_tumidus               
  |  +--------+                                    
  |  |        |  +----- Unio_crassus               
  |  |        +--+                                 
  |  |           |  +-- Unio_delphinus             
  |  |           +--+                              
  +--+              +-- Unio_pictorum              
     |                                             
     |              +-- Schistodesmus_lampreyanus  
     |  +-----------+                              
     |  |           +-- Schistodesmus_spinosus     
     |  |                                          
     +--+        +----- Nodularia_douglasiae[1]    
        |  +-----+                                 
        |  |     |  +-- Nodularia_douglasiae[2]    
        |  |     +--+                              
        +--+        +-- Nodularia_nuxpersicae      
           |                                       
           |  +-------- Tchangsinaia_piscicula     
           +--+                                    
              |  +----- Cuneopsis_rufescens        
              +--+                                 
                 |  +-- Cuneopsis_heudei           
                 +--+                              
                    +-- Cuneopsis_celtiformis

Huang et al., 2019 [Publ 3843], fig. S1

iT tT bi riS oT roT roS iD
5 5 3 5 70 9 4 4.0 
        +----------- Aculamprotula_tortuosa **     
  +-----+                                       
  |     |  +-------- Aculamprotula_tientsinensis **
  |     +--+                                    
  |        |  +----- Aculamprotula_fibrosa **      
  |        +--+                                 
  |           |  +-- Aculamprotula_coreana **      
--+           +--+                              
  |              +-- Aculamprotula_scripta **      
  |                                             
  |           +----- Nodularia_douglasiae[1]    
  |  +--------+                                 
  |  |        |  +-- Nodularia_nuxpersicae      
  |  |        +--+                              
  +--+           +-- Nodularia_douglasiae[2]    
     |                                          
     |  +----------- Tchangsinaia_piscicula     
     |  |                                       
     +--+        +-- Schistodesmus_spinosus     
        |  +-----+                              
        |  |     +-- Schistodesmus_lampreyanus  
        +--+                                    
           |  +----- Cuneopsis_rufescens        
           +--+                                 
              |  +-- Cuneopsis_heudei           
              +--+                              
                 +-- Cuneopsis_celtiformis

Huang et al., 2019 [Publ 3843], fig. S3

iT tT bi riS oT roT roS iD
5 5 3 5 196 12 5 2.0 
              +-------- Aculamprotula_tortuosa **     
              |                                    
  +-----------+     +-- Aculamprotula_coreana **      
  |           |  +--+                              
  |           |  |  +-- Aculamprotula_scripta **      
  |           +--+                                 
  |              |  +-- Aculamprotula_fibrosa **      
  |              +--+                              
  |                 +-- Aculamprotula_tientsinensis **
--+                                                
  |           +-------- Unio_tumidus               
  |  +--------+                                    
  |  |        |  +----- Unio_crassus               
  |  |        +--+                                 
  |  |           |  +-- Unio_delphinus             
  |  |           +--+                              
  +--+              +-- Unio_pictorum              
     |                                             
     |           +----- Nodularia_douglasiae[1]    
     |  +--------+                                 
     |  |        |  +-- Nodularia_douglasiae[2]    
     |  |        +--+                              
     +--+           +-- Nodularia_nuxpersicae      
        |                                          
        |  +----------- Schistodesmus_lampreyanus  
        +--+                                       
           |  +-------- Tchangsinaia_piscicula     
           +--+                                    
              |  +----- Cuneopsis_rufescens        
              +--+                                 
                 |  +-- Cuneopsis_heudei           
                 +--+                              
                    +-- Cuneopsis_celtiformis

Huang et al., 2019 [Publ 3843], fig. S4

iT tT bi riS oT roT roS iD
5 5 3 5 197 13 5 2.0 
  +-------- Aculamprotula_tortuosa **     
  |                                    
--+     +-- Aculamprotula_coreana **      
  |  +--+                              
  |  |  +-- Aculamprotula_scripta **      
  +--+                                 
     |  +-- Aculamprotula_fibrosa **      
     +--+                              
        +-- Aculamprotula_tientsinensis **

Lopes-Lima et al., 2020 [Publ 4080], fig. 5

iT tT bi riS oT roT roS iD
5 5 3 5 4 2 1 4.0 
              +-- Nodularia_breviconcha      
  +-----------+                              
  |           +-- Nodularia_douglasiae       
--+                                          
  |  +----------- Aculamprotula_tortuosa **     
  +--+                                       
     |  +-------- Aculamprotula_tientsinensis **
     +--+                                    
        |  +----- Aculamprotula_fibrosa **      
        +--+                                 
           |  +-- Aculamprotula_scripta **      
           +--+                              
              +-- Aculamprotula_coreana **

Wu et al., 2020 [Publ 4121], fig. 3

iT tT bi riS oT roT roS iD
5 5 3 5 45 9 6 2.0 
           +-------- Aculamprotula_tortuosa **     
           |                                    
  +--------+     +-- Aculamprotula_coreana **      
  |        |  +--+                              
  |        |  |  +-- Aculamprotula_tientsinensis **
  |        +--+                                 
  |           |  +-- Aculamprotula_polysticta **   
  |           +--+                              
--+              +-- Aculamprotula_scripta **      
  |                                             
  |        +-------- Unio_tumidus               
  |  +-----+                                    
  |  |     |  +----- Unio_crassus               
  |  |     +--+                                 
  |  |        |  +-- Unio_delphinus             
  +--+        +--+                              
     |           +-- Unio_pictorum              
     |                                          
     |  +----------- Pseudocuneopsis_capitata   
     +--+                                       
        |  +-------- Nodularia_douglasiae       
        +--+                                    
           |  +----- Tchangsinaia_piscicula     
           +--+                                 
              |  +-- Cuneopsis_heudei           
              +--+                              
                 +-- Schistodesmus_lampreyanus

Wu et al., 2022 [Publ 4238], fig. 2d

iT tT bi riS oT roT roS iD
5 5 3 5 34 18 7 2.0 
                    +-------- Aculamprotula_tortuosa **      
                    |                                     
  +-----------------+     +-- Aculamprotula_tientsinensis ** 
  |                 |  +--+                               
  |                 |  |  +-- Aculamprotula_coreana **       
  |                 +--+                                  
  |                    |  +-- Aculamprotula_polysticta **    
  |                    +--+                               
  |                       +-- Aculamprotula_scripta **       
--+                                                       
  |                 +-------- Unio_tumidus                
  |  +--------------+                                     
  |  |              |  +----- Unio_crassus                
  |  |              +--+                                  
  |  |                 |  +-- Unio_pictorum               
  |  |                 +--+                               
  |  |                    +-- Unio_delphinus              
  +--+                                                    
     |                    +-- Pseudocuneopsis_capitata    
     |  +-----------------+                               
     |  |                 +-- Pseudocuneopsis_sichuanensis
     |  |                                                 
     |  |                 +-- Nodularia_breviconcha       
     +--+  +--------------+                               
        |  |              +-- Nodularia_douglasiae        
        |  |                                              
        |  |              +-- Schistodesmus_lampreyanus   
        +--+  +-----------+                               
           |  |           +-- Schistodesmus_spinosus      
           |  |                                           
           |  |     +-------- Cuneopsis_rufescens         
           +--+  +--+                                     
              |  |  |  +----- Cuneopsis_celtiformis       
              |  |  +--+                                  
              |  |     |  +-- Cuneopsis_heudei            
              +--+     +--+                               
                 |        +-- Cuneopsis_demangei          
                 |                                        
                 |  +-------- Tchangsinaia_piscicula      
                 +--+                                     
                    |  +----- Inversiunio_jokohamensis    
                    +--+                                  
                       |  +-- Inversiunio_reinianus       
                       +--+                               
                          +-- Inversiunio_yanagawensis

Wu et al., 2022 [Publ 4238], fig. 3

iT tT bi riS oT roT roS iD
5 5 3 5 69 15 6 2.0 
                    +-------- Aculamprotula_tortuosa **      
                    |                                     
  +-----------------+     +-- Aculamprotula_tientsinensis ** 
  |                 |  +--+                               
  |                 |  |  +-- Aculamprotula_coreana **       
  |                 +--+                                  
  |                    |  +-- Aculamprotula_polysticta **    
  |                    +--+                               
  |                       +-- Aculamprotula_scripta **       
--+                                                       
  |                 +-------- Unio_tumidus                
  |  +--------------+                                     
  |  |              |  +----- Unio_crassus                
  |  |              +--+                                  
  |  |                 |  +-- Unio_pictorum               
  |  |                 +--+                               
  +--+                    +-- Unio_delphinus              
     |                                                    
     |                    +-- Pseudocuneopsis_capitata    
     |  +-----------------+                               
     |  |                 +-- Pseudocuneopsis_sichuanensis
     |  |                                                 
     +--+                 +-- Nodularia_douglasiae        
        |  +--------------+                               
        |  |              +-- Nodularia_breviconcha       
        |  |                                              
        +--+              +-- Schistodesmus_lampreyanus   
           |  +-----------+                               
           |  |           +-- Schistodesmus_spinosus      
           +--+                                           
              |  +----------- Tchangsinaia_piscicula      
              +--+                                        
                 |  +-------- Cuneopsis_rufescens         
                 +--+                                     
                    |  +----- Cuneopsis_celtiformis       
                    +--+                                  
                       |  +-- Cuneopsis_heudei            
                       +--+                               
                          +-- Cuneopsis_demangei

Wu et al., 2023 [Publ 4339], fig. 4, S5-aa

iT tT bi riS oT roT roS iD
5 5 3 5 91 17 6 2.0 
                 +-------- Aculamprotula_tortuosa **      
                 |                                     
  +--------------+     +-- Aculamprotula_coreana **       
  |              |  +--+                               
  |              |  |  +-- Aculamprotula_tientsinensis ** 
  |              +--+                                  
  |                 |  +-- Aculamprotula_polysticta **    
  |                 +--+                               
  |                    +-- Aculamprotula_scripta **       
  |                                                    
--+        +-------------- Unio_tumidus                
  |  +-----+                                           
  |  |     |  +----------- Unio_crassus                
  |  |     +--+                                        
  |  |        |  +-------- Unio_elongatulus            
  |  |        +--+                                     
  |  |           |  +----- Unio_pictorum               
  |  |           +--+                                  
  |  |              |  +-- Unio_delphinus              
  +--+              +--+                               
     |                 +-- Unio_mancus                 
     |                                                 
     |                 +-- Nodularia_breviconcha       
     |              +--+                               
     |              |  +-- Nodularia_nipponensis       
     |  +-----------+                                  
     |  |           |  +-- Pseudocuneopsis_capitata    
     |  |           +--+                               
     +--+              +-- Pseudocuneopsis_sichuanensis
        |                                              
        |  +-------------- Tchangsinaia_piscicula      
        |  |                                           
        +--+           +-- Schistodesmus_lampreyanus   
           |  +--------+                               
           |  |        +-- Schistodesmus_spinosus      
           +--+                                        
              |  +-------- Cuneopsis_rufescens         
              +--+                                     
                 |  +----- Cuneopsis_celtiformis       
                 +--+                                  
                    |  +-- Cuneopsis_demangei          
                    +--+                               
                       +-- Cuneopsis_heudei

Wu et al., 2023 [Publ 4339], fig. 4, S5-nt

iT tT bi riS oT roT roS iD
5 5 3 5 92 18 6 2.0 
                    +-------- Aculamprotula_tortuosa **      
                    |                                     
  +-----------------+     +-- Aculamprotula_coreana **       
  |                 |  +--+                               
  |                 |  |  +-- Aculamprotula_tientsinensis ** 
  |                 +--+                                  
  |                    |  +-- Aculamprotula_polysticta **    
  |                    +--+                               
  |                       +-- Aculamprotula_scripta **       
  |                                                       
--+              +----------- Unio_tumidus                
  |              |                                        
  |  +-----------+  +-------- Unio_crassus                
  |  |           |  |                                     
  |  |           +--+  +----- Unio_elongatulus            
  |  |              |  |                                  
  |  |              +--+  +-- Unio_delphinus              
  |  |                 |  |                               
  |  |                 +--+-- Unio_mancus                 
  +--+                    |                               
     |                    +-- Unio_pictorum               
     |                                                    
     |                    +-- Pseudocuneopsis_capitata    
     |  +-----------------+                               
     |  |                 +-- Pseudocuneopsis_sichuanensis
     |  |                                                 
     +--+                 +-- Nodularia_breviconcha       
        |  +--------------+                               
        |  |              +-- Nodularia_nipponensis       
        |  |                                              
        +--+           +----- Tchangsinaia_piscicula      
           |  +--------+                                  
           |  |        |  +-- Schistodesmus_lampreyanus[2]
           |  |        +--+                               
           +--+           +-- Schistodesmus_spinosus      
              |                                           
              |  +----------- Schistodesmus_lampreyanus[1]
              +--+                                        
                 |  +-------- Cuneopsis_rufescens         
                 +--+                                     
                    |  +----- Cuneopsis_celtiformis       
                    +--+                                  
                       |  +-- Cuneopsis_demangei          
                       +--+                               
                          +-- Cuneopsis_heudei

Wu et al., 2024 [Publ 4350], fig. 8a, S1

iT tT bi riS oT roT roS iD
5 5 3 5 87 17 6 2.0 
                 +-------- Aculamprotula_tortuosa **      
                 |                                     
  +--------------+     +-- Aculamprotula_coreana **       
  |              |  +--+                               
  |              |  |  +-- Aculamprotula_tientsinensis ** 
  |              +--+                                  
  |                 |  +-- Aculamprotula_polysticta **    
  |                 +--+                               
  |                    +-- Aculamprotula_scripta **       
  |                                                    
--+        +-------------- Unio_tumidus                
  |  +-----+                                           
  |  |     |  +----------- Unio_crassus                
  |  |     +--+                                        
  |  |        |  +-------- Unio_elongatulus            
  |  |        +--+                                     
  |  |           |  +----- Unio_pictorum               
  |  |           +--+                                  
  |  |              |  +-- Unio_delphinus              
  +--+              +--+                               
     |                 +-- Unio_mancus                 
     |                                                 
     |                 +-- Nodularia_breviconcha       
     |  +--------------+                               
     |  |              +-- Nodularia_douglasiae        
     |  |                                              
     |  |              +-- Pseudocuneopsis_capitata    
     |  +--------------+                               
     +--+              +-- Pseudocuneopsis_sichuanensis
        |                                              
        |              +-- Schistodesmus_lampreyanus   
        |  +-----------+                               
        |  |           +-- Schistodesmus_spinosus      
        +--+                                           
           |  +----------- Tchangsinaia_piscicula      
           +--+                                        
              |  +-------- Cuneopsis_rufescens         
              +--+                                     
                 |  +----- Cuneopsis_celtiformis       
                 +--+                                  
                    |  +-- Cuneopsis_demangei          
                    +--+                               
                       +-- Cuneopsis_heudei

Wu et al., 2024 [Publ 4350], fig. 8b, S2

iT tT bi riS oT roT roS iD
5 5 3 5 87 17 6 2.0 
                 +-------- Aculamprotula_tortuosa **      
                 |                                     
  +--------------+     +-- Aculamprotula_coreana **       
  |              |  +--+                               
  |              |  |  +-- Aculamprotula_tientsinensis ** 
  |              +--+                                  
  |                 |  +-- Aculamprotula_polysticta **    
  |                 +--+                               
  |                    +-- Aculamprotula_scripta **       
  |                                                    
--+        +-------------- Unio_tumidus                
  |  +-----+                                           
  |  |     |  +----------- Unio_crassus                
  |  |     +--+                                        
  |  |        |  +-------- Unio_elongatulus            
  |  |        +--+                                     
  |  |           |  +----- Unio_pictorum               
  |  |           +--+                                  
  |  |              |  +-- Unio_delphinus              
  +--+              +--+                               
     |                 +-- Unio_mancus                 
     |                                                 
     |                 +-- Nodularia_breviconcha       
     |  +--------------+                               
     |  |              +-- Nodularia_douglasiae        
     |  |                                              
     |  |              +-- Pseudocuneopsis_capitata    
     |  +--------------+                               
     +--+              +-- Pseudocuneopsis_sichuanensis
        |                                              
        |              +-- Schistodesmus_lampreyanus   
        |  +-----------+                               
        |  |           +-- Schistodesmus_spinosus      
        +--+                                           
           |  +----------- Tchangsinaia_piscicula      
           +--+                                        
              |  +-------- Cuneopsis_rufescens         
              +--+                                     
                 |  +----- Cuneopsis_celtiformis       
                 +--+                                  
                    |  +-- Cuneopsis_demangei          
                    +--+                               
                       +-- Cuneopsis_heudei

Huang et al., 2019 [Publ 3843], fig. 3

iT tT bi riS oT roT roS iD
4 4 2 4 53 7 4 1.0 
        +-------- Aculamprotula_tortuosa **     
  +-----+                                    
  |     |  +----- Aculamprotula_scripta **      
  |     +--+                                 
  |        |  +-- Aculamprotula_tientsinensis **
  |        +--+                              
--+           +-- Aculamprotula_coreana **      
  |                                          
  |        +----- Nodularia_douglasiae       
  |  +-----+                                 
  |  |     |  +-- Tchangsinaia_piscicula     
  |  |     +--+                              
  +--+        +-- Schistodesmus_lampreyanus  
     |                                       
     |  +-------- Unio_tumidus               
     +--+                                    
        |  +----- Unio_crassus               
        +--+                                 
           |  +-- Unio_delphinus             
           +--+                              
              +-- Unio_pictorum

Wu et al., 2019 [Publ 3864], fig. 3

iT tT bi riS oT roT roS iD
4 4 2 4 43 9 6 1.0 
           +-------- Aculamprotula_tortuosa **     
  +--------+                                    
  |        |  +----- Aculamprotula_scripta **      
  |        +--+                                 
  |           |  +-- Aculamprotula_tientsinensis **
  |           +--+                              
  |              +-- Aculamprotula_coreana **      
--+                                             
  |        +-------- Unio_tumidus               
  |  +-----+                                    
  |  |     |  +----- Unio_crassus               
  |  |     +--+                                 
  |  |        |  +-- Unio_delphinus             
  +--+        +--+                              
     |           +-- Unio_pictorum              
     |                                          
     |  +----------- Pseudocuneopsis_capitata   
     +--+                                       
        |  +-------- Nodularia_douglasiae       
        +--+                                    
           |  +----- Tchangsinaia_piscicula     
           +--+                                 
              |  +-- Cuneopsis_heudei           
              +--+                              
                 +-- Schistodesmus_lampreyanus

Wu et al., 2020 [Publ 4121], fig. 2

iT tT bi riS oT roT roS iD
4 4 2 4 30 1 1 3.0 
  +----------- Unio_pictorum              
--+                                       
  |  +-------- Aculamprotula_tortuosa **     
  +--+                                    
     |  +----- Aculamprotula_polysticta **   
     +--+                                 
        |  +-- Aculamprotula_fibrosa **      
        +--+                              
           +-- Aculamprotula_tientsinensis **

Wu et al., 2020 [Publ 4121], fig. S1

iT tT bi riS oT roT roS iD
4 4 2 4 30 1 1 3.0 
  +----------- Unio_pictorum              
--+                                       
  |  +-------- Aculamprotula_tortuosa **     
  +--+                                    
     |  +----- Aculamprotula_polysticta **   
     +--+                                 
        |  +-- Aculamprotula_tientsinensis **
        +--+                              
           +-- Aculamprotula_fibrosa **

Wu et al., 2018 [Publ 4143], fig. 2

iT tT bi riS oT roT roS iD
4 4 2 4 18 2 2 1.0 
           +-- Unio_pictorum              
  +--------+                              
  |        +-- Nodularia_douglasiae       
--+                                       
  |  +-------- Aculamprotula_tortuosa **     
  +--+                                    
     |  +----- Aculamprotula_scripta **      
     +--+                                 
        |  +-- Aculamprotula_coreana **      
        +--+                              
           +-- Aculamprotula_tientsinensis **

Wu et al., 2020 [Publ 4147], fig. 3

iT tT bi riS oT roT roS iD
4 4 2 4 26 6 6 1.0 
           +-------- Aculamprotula_tortuosa **     
  +--------+                                    
  |        |  +----- Aculamprotula_scripta **      
  |        +--+                                 
  |           |  +-- Aculamprotula_tientsinensis **
--+           +--+                              
  |              +-- Aculamprotula_coreana **      
  |                                             
  |  +-------------- Unio_pictorum              
  +--+                                          
     |  +----------- Pseudocuneopsis_capitata   
     +--+                                       
        |  +-------- Nodularia_douglasiae       
        +--+                                    
           |  +----- Schistodesmus_lampreyanus  
           +--+                                 
              |  +-- Cuneopsis_heudei           
              +--+                              
                 +-- Tchangsinaia_piscicula

Wu et al., 2020 [Publ 4147], fig. 4

iT tT bi riS oT roT roS iD
4 4 2 4 10 1 1 1.0 
  +----------- Unio_pictorum              
--+                                       
  |  +-------- Aculamprotula_tortuosa **     
  +--+                                    
     |  +----- Aculamprotula_scripta **      
     +--+                                 
        |  +-- Aculamprotula_tientsinensis **
        +--+                              
           +-- Aculamprotula_fibrosa **

Konopleva et al., 2023 [Publ 4269], fig. S1, 4

iT tT bi riS oT roT roS iD
4 4 2 4 52 8 5 1.0 
  +----------------- Middendorffinaia_mongolica 
  |                                             
  |        +-------- Aculamprotula_tortuosa **     
  |  +-----+                                    
--+  |     |  +----- Aculamprotula_scripta **      
  |  |     +--+                                 
  |  |        |  +-- Aculamprotula_tientsinensis **
  |  |        +--+                              
  +--+           +-- Aculamprotula_coreana **      
     |                                          
     |        +----- Unio_tumidus               
     |  +-----+                                 
     |  |     |  +-- Unio_pictorum              
     |  |     +--+                              
     +--+        +-- Unio_crassus               
        |                                       
        |  +-------- Nodularia_douglasiae       
        +--+                                    
           |  +----- Tchangsinaia_piscicula     
           +--+                                 
              |  +-- Cuneopsis_rufescens        
              +--+                              
                 +-- Cuneopsis_heudei

Wu et al., 2016 [Publ 3711], fig. 1

iT tT bi riS oT roT roS iD
3 3 1 3 18 2 2 2.0 
        +-- Unio_pictorum              
  +-----+                              
  |     +-- Nodularia_douglasiae       
--+                                    
  |  +----- Aculamprotula_tortuosa **     
  +--+                                 
     |  +-- Aculamprotula_coreana **      
     +--+                              
        +-- Aculamprotula_tientsinensis **

Huang et al., 2018 [Publ 3828], fig. 6

iT tT bi riS oT roT roS iD
3 3 1 3 40 6 3 2.0 
           +----- Aculamprotula_tortuosa **     
  +--------+                                 
  |        |  +-- Aculamprotula_coreana **      
  |        +--+                              
  |           +-- Aculamprotula_tientsinensis **
--+                                          
  |           +-- Tchangsinaia_piscicula     
  |  +--------+                              
  |  |        +-- Nodularia_douglasiae       
  +--+                                       
     |  +-------- Unio_tumidus               
     +--+                                    
        |  +----- Unio_crassus               
        +--+                                 
           |  +-- Unio_delphinus             
           +--+                              
              +-- Unio_pictorum

Soroka & Burzynski, 2017 [Publ 3837], fig. 1

iT tT bi riS oT roT roS iD
3 3 1 3 21 7 4 2.0 
           +----- Aculamprotula_tortuosa **     
  +--------+                                 
  |        |  +-- Aculamprotula_tientsinensis **
  |        +--+                              
  |           +-- Aculamprotula_coreana **      
--+                                          
  |        +----- Schistodesmus_lampreyanus  
  |  +-----+                                 
  |  |     |  +-- Tchangsinaia_piscicula     
  |  |     +--+                              
  +--+        +-- Nodularia_douglasiae       
     |                                       
     |  +-------- Unio_tumidus               
     +--+                                    
        |  +----- Unio_crassus               
        +--+                                 
           |  +-- Unio_pictorum              
           +--+                              
              +-- Unio_delphinus

Aunins et al., 2018 [Publ 3863], fig. 1

iT tT bi riS oT roT roS iD
3 3 1 3 38 8 5 2.0 
           +----- Aculamprotula_tortuosa **     
  +--------+                                 
  |        |  +-- Aculamprotula_coreana **      
  |        +--+                              
  |           +-- Aculamprotula_tientsinensis **
  |                                          
--+     +-------- Nodularia_douglasiae       
  |  +--+                                    
  |  |  |  +----- Tchangsinaia_piscicula     
  |  |  +--+                                 
  |  |     |  +-- Cuneopsis_heudei           
  +--+     +--+                              
     |        +-- Schistodesmus_lampreyanus  
     |                                       
     |  +-------- Unio_tumidus               
     +--+                                    
        |  +----- Unio_crassus               
        +--+                                 
           |  +-- Unio_pictorum              
           +--+                              
              +-- Unio_delphinus

Cha et al., 2018 [Publ 3916], fig. 1

iT tT bi riS oT roT roS iD
3 3 1 3 36 7 4 2.0 
           +----- Aculamprotula_tortuosa **     
  +--------+                                 
  |        |  +-- Aculamprotula_coreana **      
  |        +--+                              
  |           +-- Aculamprotula_tientsinensis **
--+                                          
  |        +----- Schistodesmus_lampreyanus  
  |  +-----+                                 
  |  |     |  +-- Nodularia_douglasiae       
  |  |     +--+                              
  +--+        +-- Tchangsinaia_piscicula     
     |                                       
     |  +-------- Unio_tumidus               
     +--+                                    
        |  +----- Unio_crassus               
        +--+                                 
           |  +-- Unio_pictorum              
           +--+                              
              +-- Unio_delphinus

Froufe et al., 2019 [Publ 3936], fig. 3

iT tT bi riS oT roT roS iD
3 3 1 3 42 7 4 2.0 
           +----- Aculamprotula_tortuosa **     
  +--------+                                 
  |        |  +-- Aculamprotula_tientsinensis **
  |        +--+                              
  |           +-- Aculamprotula_coreana **      
--+                                          
  |        +----- Nodularia_douglasiae       
  |  +-----+                                 
  |  |     |  +-- Tchangsinaia_piscicula     
  |  |     +--+                              
  +--+        +-- Schistodesmus_lampreyanus  
     |                                       
     |  +-------- Unio_tumidus               
     +--+                                    
        |  +----- Unio_crassus               
        +--+                                 
           |  +-- Unio_pictorum              
           +--+                              
              +-- Unio_delphinus

Froufe et al., 2019 [Publ 3936], fig. 6

iT tT bi riS oT roT roS iD
3 3 1 3 42 7 4 2.0 
           +----- Aculamprotula_tortuosa **     
  +--------+                                 
  |        |  +-- Aculamprotula_tientsinensis **
  |        +--+                              
  |           +-- Aculamprotula_coreana **      
--+                                          
  |        +----- Nodularia_douglasiae       
  |  +-----+                                 
  |  |     |  +-- Tchangsinaia_piscicula     
  |  |     +--+                              
  +--+        +-- Schistodesmus_lampreyanus  
     |                                       
     |  +-------- Unio_tumidus               
     +--+                                    
        |  +----- Unio_crassus               
        +--+                                 
           |  +-- Unio_pictorum              
           +--+                              
              +-- Unio_delphinus

Fukata & Iigo, 2020 [Publ 4106], fig. 1

iT tT bi riS oT roT roS iD
3 3 1 3 51 9 6 2.0 
           +----- Aculamprotula_tortuosa **     
  +--------+                                 
  |        |  +-- Aculamprotula_tientsinensis **
  |        +--+                              
  |           +-- Aculamprotula_coreana **      
  |                                          
--+     +-------- Unio_tumidus               
  |  +--+                                    
  |  |  |  +----- Unio_crassus               
  |  |  +--+                                 
  |  |     |  +-- Unio_pictorum              
  |  |     +--+                              
  +--+        +-- Unio_delphinus             
     |                                       
     |        +-- Schistodesmus_lampreyanus  
     |  +-----+                              
     |  |     +-- Cuneopsis_heudei           
     +--+                                    
        |  +----- Pseudocuneopsis_capitata   
        +--+                                 
           |  +-- Nodularia_douglasiae       
           +--+                              
              +-- Tchangsinaia_piscicula

Sano et al., 2020 [Publ 4138], fig. 2

iT tT bi riS oT roT roS iD
3 3 1 3 70 12 5 2.0 
              +----- Aculamprotula_tortuosa **     
  +-----------+                                 
  |           |  +-- Aculamprotula_tientsinensis **
  |           +--+                              
  |              +-- Aculamprotula_coreana **      
  |                                             
--+           +----- Unio_crassus               
  |  +--------+                                 
  |  |        |  +-- Unio_delphinus             
  |  |        +--+                              
  |  |           +-- Unio_pictorum              
  |  |                                          
  +--+     +-------- Inversiunio_jokohamensis   
     |  +--+                                    
     |  |  |  +----- Inversiunio_yanagawensis[2]
     |  |  +--+                                 
     |  |     |  +-- Inversiunio_reinianus      
     |  |     +--+                              
     +--+        +-- Inversiunio_yanagawensis[1]
        |                                       
        |        +-- Tchangsinaia_piscicula     
        |  +-----+                              
        |  |     +-- Schistodesmus_lampreyanus  
        +--+                                    
           |  +----- Nodularia_douglasiae[2]    
           +--+                                 
              |  +-- Nodularia_nipponensis      
              +--+                              
                 +-- Nodularia_douglasiae[1]

Sano et al., 2020 [Publ 4138], fig. 3

iT tT bi riS oT roT roS iD
3 3 1 3 69 11 5 2.0 
              +----- Aculamprotula_tortuosa **     
  +-----------+                                 
  |           |  +-- Aculamprotula_tientsinensis **
  |           +--+                              
  |              +-- Aculamprotula_coreana **      
  |                                             
--+           +----- Unio_crassus               
  |  +--------+                                 
  |  |        |  +-- Unio_delphinus             
  |  |        +--+                              
  |  |           +-- Unio_pictorum              
  |  |                                          
  +--+     +-------- Inversiunio_jokohamensis   
     |  +--+                                    
     |  |  |  +----- Inversiunio_yanagawensis[2]
     |  |  +--+                                 
     |  |     |  +-- Inversiunio_reinianus      
     |  |     +--+                              
     +--+        +-- Inversiunio_yanagawensis[1]
        |                                       
        |        +-- Nodularia_nipponensis      
        |     +--+                              
        |     |  +-- Nodularia_douglasiae       
        +-----+                                 
              |  +-- Schistodesmus_lampreyanus  
              +--+                              
                 +-- Tchangsinaia_piscicula

Sano et al., 2020 [Publ 4138], fig. 4

iT tT bi riS oT roT roS iD
3 3 1 3 69 11 5 2.0 
                 +----- Aculamprotula_tortuosa **     
  +--------------+                                 
  |              |  +-- Aculamprotula_tientsinensis **
  |              +--+                              
  |                 +-- Aculamprotula_coreana **      
  |                                                
--+              +----- Unio_crassus               
  |  +-----------+                                 
  |  |           |  +-- Unio_delphinus             
  |  |           +--+                              
  |  |              +-- Unio_pictorum              
  +--+                                             
     |              +-- Nodularia_douglasiae       
     |  +-----------+                              
     |  |           +-- Nodularia_nipponensis      
     |  |                                          
     +--+           +-- Schistodesmus_lampreyanus  
        |  +--------+                              
        |  |        +-- Tchangsinaia_piscicula     
        +--+                                       
           |  +-------- Inversiunio_jokohamensis   
           +--+                                    
              |  +----- Inversiunio_yanagawensis[2]
              +--+                                 
                 |  +-- Inversiunio_reinianus      
                 +--+                              
                    +-- Inversiunio_yanagawensis[1]

Zieritz et al., 2021 [Publ 4231], fig. 3-F, 5

iT tT bi riS oT roT roS iD
3 3 1 3 46 7 4 2.0 
           +----- Aculamprotula_tortuosa **     
  +--------+                                 
  |        |  +-- Aculamprotula_tientsinensis **
  |        +--+                              
  |           +-- Aculamprotula_coreana **      
--+                                          
  |        +----- Nodularia_douglasiae       
  |  +-----+                                 
  |  |     |  +-- Schistodesmus_lampreyanus  
  |  |     +--+                              
  +--+        +-- Tchangsinaia_piscicula     
     |                                       
     |  +-------- Unio_tumidus               
     +--+                                    
        |  +----- Unio_crassus               
        +--+                                 
           |  +-- Unio_pictorum              
           +--+                              
              +-- Unio_delphinus

Fonseca et al., 2016 [Publ 3775], fig. 1

iT tT bi riS oT roT roS iD
2 2 0 2 30 5 4 -- 
           +-- Aculamprotula_tortuosa **   
  +--------+                            
  |        +-- Aculamprotula_coreana **    
  |                                     
--+        +-- Unio_pictorum            
  |  +-----+                            
  |  |     +-- Unio_delphinus           
  +--+                                  
     |  +----- Schistodesmus_lampreyanus
     +--+                               
        |  +-- Nodularia_douglasiae     
        +--+                            
           +-- Tchangsinaia_piscicula

Zhou et al., 2017 [Publ 3788], fig. 2

iT tT bi riS oT roT roS iD
2 2 0 2 15 2 2 -- 
     +-- Nodularia_douglasiae       
  +--+                              
  |  +-- Tchangsinaia_piscicula     
--+                                 
  |  +-- Aculamprotula_tientsinensis **
  +--+                              
     +-- Aculamprotula_tortuosa **

Lopes-Lima et al., 2017 [Publ 3792], fig. 2

iT tT bi riS oT roT roS iD
2 2 0 2 31 5 4 -- 
           +-- Aculamprotula_tortuosa **   
  +--------+                            
  |        +-- Aculamprotula_coreana **    
  |                                     
--+        +-- Unio_delphinus           
  |  +-----+                            
  |  |     +-- Unio_pictorum            
  +--+                                  
     |  +----- Schistodesmus_lampreyanus
     +--+                               
        |  +-- Tchangsinaia_piscicula   
        +--+                            
           +-- Nodularia_douglasiae

Wen et al., 2017 [Publ 3795], fig. 3

iT tT bi riS oT roT roS iD
2 2 0 2 14 1 1 -- 
  +-- Aculamprotula_coreana ** 
--+                         
  +-- Aculamprotula_tortuosa **

Chen et al., 2020 [Publ 4170], fig. 1

iT tT bi riS oT roT roS iD
2 2 0 2 32 1 1 -- 
  +-- Aculamprotula_coreana ** 
--+                         
  +-- Aculamprotula_tortuosa **

Pavan-Kumar et al., 2022 [Publ 4263], fig. 4

iT tT bi riS oT roT roS iD
2 2 0 2 34 4 4 -- 
           +-- Aculamprotula_tientsinensis **
  +--------+                              
  |        +-- Aculamprotula_tortuosa **     
--+                                       
  |  +-------- Unio_pictorum              
  +--+                                    
     |  +----- Schistodesmus_lampreyanus  
     +--+                                 
        |  +-- Nodularia_douglasiae       
        +--+                              
           +-- Tchangsinaia_piscicula

Pavan-Kumar et al., 2022 [Publ 4263], fig. S5

iT tT bi riS oT roT roS iD
2 2 0 2 37 4 4 -- 
  +-- Aculamprotula_tientsinensis **
--+                              
  +-- Aculamprotula_tortuosa **

Wu et al., 2023 [Publ 4296], fig. 4

iT tT bi riS oT roT roS iD
2 2 0 2 21 11 7 -- 
  +----------------------- Unio_tumidus                 
  |                                                     
  |                    +-- Aculamprotula_scripta **        
--+  +-----------------+                                
  |  |                 +-- Aculamprotula_tientsinensis **  
  |  |                                                  
  +--+                 +-- Cuneopsis_heudei             
     |  +--------------+                                
     |  |              +-- Cuneopsis_rufescens          
     |  |                                               
     +--+           +----- Nodularia_douglasiae         
        |  +--------+                                   
        |  |        |  +-- Schistodesmus_lampreyanus    
        |  |        +--+                                
        +--+           +-- Schistodesmus_spinosus       
           |                                            
           |  +----------- Tchangsinaia_piscicula       
           +--+                                         
              |  +-------- Inversiunio_yanagawensis     
              +--+                                      
                 |  +----- Pseudocuneopsis_capitata     
                 +--+                                   
                    |  +-- Pseudocuneopsis_yangshuoensis
                    +--+                                
                       +-- Pseudocuneopsis_sichuanensis

Liu et al., 2023 [Publ 4310], fig. 4

iT tT bi riS oT roT roS iD
2 2 0 2 22 12 7 -- 
  +-------------------------- Unio_tumidus                 
  |                                                        
  |                       +-- Aculamprotula_scripta **        
--+  +--------------------+                                
  |  |                    +-- Aculamprotula_tientsinensis **  
  |  |                                                     
  +--+                    +-- Cuneopsis_heudei             
     |  +-----------------+                                
     |  |                 +-- Cuneopsis_rufescens          
     |  |                                                  
     +--+              +----- Nodularia_douglasiae         
        |  +-----------+                                   
        |  |           |  +-- Schistodesmus_lampreyanus    
        |  |           +--+                                
        +--+              +-- Schistodesmus_spinosus       
           |                                               
           |  +-------------- Tchangsinaia_piscicula       
           +--+                                            
              |  +----------- Inversiunio_yanagawensis     
              +--+                                         
                 |  +-------- Pseudocuneopsis_capitata     
                 +--+                                      
                    |  +----- Pseudocuneopsis_sichuanensis 
                    +--+                                   
                       |  +-- Pseudocuneopsis_wuana        
                       +--+                                
                          +-- Pseudocuneopsis_yangshuoensis

Liu et al., 2023 [Publ 4310], fig. 5

iT tT bi riS oT roT roS iD
2 2 0 2 22 12 7 -- 
  +----------------- Unio_tumidus                 
  |                                               
  |              +-- Aculamprotula_tientsinensis **  
  +--------------+                                
  |              +-- Aculamprotula_scripta **        
  |                                               
--+              +-- Cuneopsis_rufescens          
  |  +-----------+                                
  |  |           +-- Cuneopsis_heudei             
  |  |                                            
  |  |        +----- Nodularia_douglasiae         
  |  +--------+                                   
  +--+        |  +-- Schistodesmus_lampreyanus    
     |        +--+                                
     |           +-- Schistodesmus_spinosus       
     |                                            
     |  +----------- Tchangsinaia_piscicula       
     |  |                                         
     +--+----------- Inversiunio_yanagawensis     
        |                                         
        |  +-------- Pseudocuneopsis_capitata     
        +--+                                      
           |  +----- Pseudocuneopsis_sichuanensis 
           +--+                                   
              |  +-- Pseudocuneopsis_wuana        
              +--+                                
                 +-- Pseudocuneopsis_yangshuoensis

Dai et al., 2024 [Publ 4358], fig. 4

iT tT bi riS oT roT roS iD
2 2 0 2 50 15 6 -- 
                          +-- Aculamprotula_scripta **        
  +-----------------------+                                
  |                       +-- Aculamprotula_tientsinensis **  
  |                                                        
  |                       +-- Unio_crassus                 
--+  +--------------------+                                
  |  |                    +-- Unio_delphinus               
  |  |                                                     
  |  |              +-------- Pseudocuneopsis_capitata     
  +--+  +-----------+                                      
     |  |           |  +----- Pseudocuneopsis_yangshuoensis
     |  |           +--+                                   
     |  |              |  +-- Pseudocuneopsis_yemaoi       
     |  |              +--+                                
     +--+                 +-- Pseudocuneopsis_sichuanensis 
        |                                                  
        |                 +-- Nodularia_breviconcha        
        |  +--------------+                                
        |  |              +-- Nodularia_douglasiae         
        |  |                                               
        +--+              +-- Schistodesmus_lampreyanus    
           |  +-----------+                                
           |  |           +-- Schistodesmus_spinosus       
           +--+                                            
              |  +----------- Tchangsinaia_piscicula       
              +--+                                         
                 |  +-------- Cuneopsis_rufescens          
                 +--+                                      
                    |  +----- Cuneopsis_celtiformis        
                    +--+                                   
                       |  +-- Cuneopsis_heudei             
                       +--+                                
                          +-- Cuneopsis_demangei

Rosenberg et al., 1997 [Publ 2720], fig. 2B

iT tT bi riS oT roT roS iD
1 1 0 1 22 1 1 -- 
-- Aculamprotula_tortuosa

Ouyang et al., 2011 [Publ 3408], fig. 1

iT tT bi riS oT roT roS iD
1 1 0 1 12 2 2 -- 
-- Aculamprotula_tortuosa

Xue et al., 2016 [Publ 3659], fig. 1

iT tT bi riS oT roT roS iD
1 1 0 1 15 3 3 -- 
  +-------- Aculamprotula_tortuosa **   
--+                                  
  |  +----- Schistodesmus_lampreyanus
  +--+                               
     |  +-- Nodularia_douglasiae     
     +--+                            
        +-- Unio_pictorum

Lopes-Lima et al., 2017 [Publ 3745], fig. 1-2

iT tT bi riS oT roT roS iD
1 1 0 1 80 10 5 -- 
              +----- Unio_gibbus              
              |                               
  +-----------+----- Unio_tumidus             
  |           |                               
  |           |  +-- Unio_crassus             
  |           +--+                            
--+              +-- Unio_pictorum            
  |                                           
  |  +-------------- Aculamprotula_tortuosa **   
  |  |                                        
  +--+           +-- Nodularia_nuxpersicae    
     |  +--------+                            
     |  |        +-- Nodularia_douglasiae     
     +--+                                     
        |  +-------- Schistodesmus_lampreyanus
        +--+                                  
           |  +----- Tchangsinaia_piscicula   
           +--+                               
              |  +-- Cuneopsis_heudei         
              +--+                            
                 +-- Cuneopsis_rufescens

Bolotov et al., 2017 [Publ 3761], fig. S1

iT tT bi riS oT roT roS iD
1 1 0 1 140 9 5 -- 
           +----- Unio_tumidus             
  +--------+                               
  |        |  +-- Unio_pictorum            
  |        +--+                            
  |           +-- Unio_crassus             
--+                                        
  |  +----------- Aculamprotula_tortuosa **   
  |  |                                     
  |  |        +-- Nodularia_douglasiae     
  +--+  +-----+                            
     |  |     +-- Nodularia_nuxpersicae    
     |  |                                  
     +--+     +-- Cuneopsis_rufescens      
        |  +--+                            
        |  |  +-- Cuneopsis_heudei         
        +--+                               
           |  +-- Tchangsinaia_piscicula   
           +--+                            
              +-- Schistodesmus_lampreyanus

Bolotov et al., 2017 [Publ 3761], fig. S2

iT tT bi riS oT roT roS iD
1 1 0 1 140 9 5 -- 
        +----- Unio_tumidus             
  +-----+                               
  |     |  +-- Unio_crassus             
  |     +--+                            
  |        +-- Unio_pictorum            
--+                                     
  |  +-------- Aculamprotula_tortuosa **   
  |  |                                  
  |  +-------- Schistodesmus_lampreyanus
  |  |                                  
  +--+     +-- Nodularia_douglasiae     
     +-----+                            
     |     +-- Nodularia_nuxpersicae    
     |                                  
     |  +----- Cuneopsis_heudei         
     +--+                               
        |  +-- Tchangsinaia_piscicula   
        +--+                            
           +-- Cuneopsis_rufescens

Soroka & Burzynski, 2018 [Publ 3762], fig. 3

iT tT bi riS oT roT roS iD
1 1 0 1 6 1 1 -- 
  +-- Unio_tumidus          
--+                         
  +-- Aculamprotula_tortuosa **

Soroka & Burzynski, 2018 [Publ 3762], fig. 3-M

iT tT bi riS oT roT roS iD
1 1 0 1 6 1 1 -- 
  +-- Unio_tumidus          
--+                         
  +-- Aculamprotula_tortuosa **

Bolotov et al., 2017 [Publ 3764], fig. S1

iT tT bi riS oT roT roS iD
1 1 0 1 130 9 5 -- 
  +-- Unio_crassus             
  |                            
  +-- Unio_pictorum            
  |                            
  +-- Unio_tumidus             
  |                            
  +-- Aculamprotula_tortuosa **   
  |                            
  +-- Cuneopsis_heudei         
--+                            
  +-- Tchangsinaia_piscicula   
  |                            
  +-- Cuneopsis_rufescens      
  |                            
  +-- Nodularia_douglasiae     
  |                            
  +-- Nodularia_nuxpersicae    
  |                            
  +-- Schistodesmus_lampreyanus

Guerra et al., 2017 [Publ 3772], fig. 2a

iT tT bi riS oT roT roS iD
1 1 0 1 24 1 1 -- 
  +-- Aculamprotula_tortuosa **
--+                         
  +-- Unio_pictorum

Guerra et al., 2017 [Publ 3772], fig. 2b

iT tT bi riS oT roT roS iD
1 1 0 1 24 1 1 -- 
-- Aculamprotula_tortuosa

Burzynski et al., 2017 [Publ 3774], fig. 1 bottom

iT tT bi riS oT roT roS iD
1 1 0 1 12 2 1 -- 
  +----- Aculamprotula_tortuosa **
--+                            
  |  +-- Unio_crassus          
  +--+                         
     +-- Unio_tumidus

Burzynski et al., 2017 [Publ 3774], fig. 1 top

iT tT bi riS oT roT roS iD
1 1 0 1 12 2 1 -- 
  +----- Aculamprotula_tortuosa **
--+                            
  |  +-- Unio_crassus          
  +--+                         
     +-- Unio_tumidus

Araujo et al., 2017 [Publ 3801], fig. 2

iT tT bi riS oT roT roS iD
1 1 0 1 13 2 2 -- 
  +-- Schistodesmus_lampreyanus
--+                            
  +-- Aculamprotula_tortuosa **

Huang et al., 2018 [Publ 3828], fig. 3, 5

iT tT bi riS oT roT roS iD
1 1 0 1 28 1 1 -- 
  +-- Unio_pictorum              
--+                              
  +-- Aculamprotula_tientsinensis **

Huang et al., 2018 [Publ 3828], fig. 6-M

iT tT bi riS oT roT roS iD
1 1 0 1 16 3 1 -- 
  +-------- Aculamprotula_tortuosa **
--+                               
  |  +----- Unio_tumidus          
  +--+                            
     |  +-- Unio_crassus          
     +--+                         
        +-- Unio_delphinus

Huang et al., 2019 [Publ 3843], fig. 3-M

iT tT bi riS oT roT roS iD
1 1 0 1 22 4 2 -- 
  +----------- Schistodesmus_lampreyanus
--+                                     
  |  +-------- Aculamprotula_tortuosa **   
  +--+                                  
     |  +----- Unio_tumidus             
     +--+                               
        |  +-- Unio_crassus             
        +--+                            
           +-- Unio_delphinus

Wang et al., 2018 [Publ 3846], fig. 2

iT tT bi riS oT roT roS iD
1 1 0 1 31 7 4 -- 
  +-------------- Aculamprotula_tientsinensis **
  |                                          
  |        +----- Nodularia_douglasiae       
--+  +-----+                                 
  |  |     |  +-- Tchangsinaia_piscicula     
  |  |     +--+                              
  +--+        +-- Cuneopsis_heudei           
     |                                       
     |  +-------- Unio_tumidus               
     +--+                                    
        |  +----- Unio_crassus               
        +--+                                 
           |  +-- Unio_pictorum              
           +--+                              
              +-- Unio_delphinus

Burzynski & Soroka, 2018 [Publ 3862], fig. 5a

iT tT bi riS oT roT roS iD
1 1 0 1 19 4 1 -- 
  +----------- Aculamprotula_tortuosa **
--+                                  
  |  +-------- Unio_tumidus          
  +--+                               
     |  +----- Unio_crassus          
     +--+                            
        |  +-- Unio_pictorum         
        +--+                         
           +-- Unio_delphinus

Burzynski & Soroka, 2018 [Publ 3862], fig. 5b

iT tT bi riS oT roT roS iD
1 1 0 1 9 4 1 -- 
  +----------- Aculamprotula_tortuosa **
--+                                  
  |  +-------- Unio_tumidus          
  +--+                               
     |  +----- Unio_crassus          
     +--+                            
        |  +-- Unio_pictorum         
        +--+                         
           +-- Unio_delphinus

Wang et al., 2018 [Publ 3865], fig. 2

iT tT bi riS oT roT roS iD
1 1 0 1 31 6 3 -- 
  +-------------- Aculamprotula_tientsinensis **
  |                                          
--+           +-- Tchangsinaia_piscicula     
  |  +--------+                              
  |  |        +-- Nodularia_douglasiae       
  +--+                                       
     |  +-------- Unio_tumidus               
     +--+                                    
        |  +----- Unio_crassus               
        +--+                                 
           |  +-- Unio_pictorum              
           +--+                              
              +-- Unio_delphinus

Fukata & Iigo, 2019 [Publ 4108], fig. 1

iT tT bi riS oT roT roS iD
1 1 0 1 13 9 6 -- 
-- Aculamprotula_tientsinensis

Sano et al., 2020 [Publ 4138], fig. 1

iT tT bi riS oT roT roS iD
1 1 0 1 47 11 4 -- 
-- Aculamprotula_tortuosa

Guerra et al., 2019 [Publ 4168], fig. 1

iT tT bi riS oT roT roS iD
1 1 0 1 40 4 2 -- 
  +----------- Aculamprotula_tortuosa **   
--+                                     
  |  +-------- Schistodesmus_lampreyanus
  +--+                                  
     |  +----- Unio_tumidus             
     +--+                               
        |  +-- Unio_crassus             
        +--+                            
           +-- Unio_delphinus

Guerra et al., 2019 [Publ 4168], fig. 1-M

iT tT bi riS oT roT roS iD
1 1 0 1 21 4 2 -- 
  +----------- Schistodesmus_lampreyanus
--+                                     
  |  +-------- Aculamprotula_tortuosa **   
  +--+                                  
     |  +----- Unio_tumidus             
     +--+                               
        |  +-- Unio_crassus             
        +--+                            
           +-- Unio_delphinus

Guerra et al., 2019 [Publ 4168], fig. 2

iT tT bi riS oT roT roS iD
1 1 0 1 40 4 2 -- 
-- Aculamprotula_tortuosa

Guerra et al., 2019 [Publ 4168], fig. 2-M

iT tT bi riS oT roT roS iD
1 1 0 1 21 4 2 -- 
  +----------- Aculamprotula_tortuosa **   
--+                                     
  |  +-------- Schistodesmus_lampreyanus
  +--+                                  
     |  +----- Unio_tumidus             
     +--+                               
        |  +-- Unio_crassus             
        +--+                            
           +-- Unio_delphinus

Kim et al., 2021 [Publ 4197], fig. 1

iT tT bi riS oT roT roS iD
1 1 0 1 14 8 4 -- 
-- Aculamprotula_tientsinensis

Sonowal et al., 2021 [Publ 4221], fig. 3a

iT tT bi riS oT roT roS iD
1 1 0 1 14 2 1 -- 
-- Aculamprotula_tientsinensis

Sonowal et al., 2021 [Publ 4221], fig. 3b

iT tT bi riS oT roT roS iD
1 1 0 1 14 2 1 -- 
-- Aculamprotula_tientsinensis

Sonowal et al., 2021 [Publ 4221], fig. 4

iT tT bi riS oT roT roS iD
1 1 0 1 15 2 1 -- 
-- Aculamprotula_tientsinensis

Wu et al., 2022 [Publ 4238], fig. 2e

iT tT bi riS oT roT roS iD
1 1 0 1 16 11 6 -- 
  +-------------------- Aculamprotula_tortuosa **      
  |                                                 
  |              +----- Unio_tumidus                
--+  +-----------+                                  
  |  |           |  +-- Unio_crassus                
  |  |           +--+                               
  |  |              +-- Unio_delphinus              
  +--+                                              
     |              +-- Pseudocuneopsis_sichuanensis
     |  +-----------+                               
     |  |           +-- Pseudocuneopsis_capitata    
     +--+                                           
        |  +----------- Nodularia_douglasiae        
        |  |                                        
        +--+  +-------- Tchangsinaia_piscicula      
           |  |                                     
           +--+     +-- Cuneopsis_celtiformis       
              |  +--+                               
              |  |  +-- Cuneopsis_heudei            
              +--+                                  
                 |  +-- Schistodesmus_lampreyanus   
                 +--+                               
                    +-- Schistodesmus_spinosus

Jeratthitikul et al., 2022 [Publ 4260], fig. S1, 1

iT tT bi riS oT roT roS iD
1 1 0 1 75 9 5 -- 
              +----- Unio_tumidus             
  +-----------+                               
  |           |  +-- Unio_pictorum            
  |           +--+                            
--+              +-- Unio_crassus             
  |                                           
  |  +-------------- Aculamprotula_tortuosa **   
  |  |                                        
  +--+           +-- Nodularia_douglasiae     
     |  +--------+                            
     |  |        +-- Nodularia_nuxpersicae    
     +--+                                     
        |  +-------- Schistodesmus_lampreyanus
        +--+                                  
           |  +----- Cuneopsis_heudei         
           +--+                               
              |  +-- Tchangsinaia_piscicula   
              +--+                            
                 +-- Cuneopsis_rufescens

Jeratthitikul et al., 2022 [Publ 4260], fig. S2, 2

iT tT bi riS oT roT roS iD
1 1 0 1 75 9 5 -- 
              +----- Unio_tumidus             
  +-----------+                               
  |           |  +-- Unio_pictorum            
  |           +--+                            
--+              +-- Unio_crassus             
  |                                           
  |  +-------------- Aculamprotula_tortuosa **   
  |  |                                        
  +--+           +-- Nodularia_douglasiae     
     |  +--------+                            
     |  |        +-- Nodularia_nuxpersicae    
     +--+                                     
        |  +-------- Schistodesmus_lampreyanus
        +--+                                  
           |  +----- Cuneopsis_heudei         
           +--+                               
              |  +-- Tchangsinaia_piscicula   
              +--+                            
                 +-- Cuneopsis_rufescens

Dai et al., 2023 [Publ 4285], fig. 5

iT tT bi riS oT roT roS iD
1 1 0 1 52 6 6 -- 
  +-------------- Aculamprotula_tientsinensis **
  |                                          
--+  +----------- Unio_crassus               
  |  |                                       
  +--+        +-- Pseudocuneopsis_capitata   
     |  +-----+                              
     |  |     +-- Nodularia_douglasiae       
     +--+                                    
        |  +----- Schistodesmus_lampreyanus  
        +--+                                 
           |  +-- Cuneopsis_celtiformis      
           +--+                              
              +-- Tchangsinaia_piscicula

Chen et al., 2023 [Publ 4302], fig. 2

iT tT bi riS oT roT roS iD
1 1 0 1 39 16 9 -- 
  +-------------------- Aculamprotula_fibrosa **       
  |                                                 
  |              +----- Middendorffinaia_mongolica  
  |  +-----------+                                  
--+  |           |  +-- Nodularia_douglasiae        
  |  |           +--+                               
  |  |              +-- Nodularia_breviconcha       
  |  |                                              
  |  |              +-- Cuneopsis_celtiformis       
  +--+  +-----------+                               
     |  |           +-- Cuneopsis_heudei            
     |  |                                           
     |  |           +-- Diaurora_aurorea            
     |  |        +--+                               
     |  |        |  +-- Diaurora_laeve              
     |  +--------+                                  
     +--+        |  +-- Schistodesmus_lampreyanus   
        |        +--+                               
        |           +-- Schistodesmus_spinosus      
        |                                           
        |           +-- Pseudocuneopsis_sichuanensis
        |  +--------+                               
        |  |        +-- Pseudocuneopsis_capitata    
        |  |                                        
        +--+        +-- Inversiunio_yanagawensis    
           |  +-----+                               
           |  |     +-- Inversiunio_reinianus       
           +--+                                     
              |  +----- Tchangsinaia_piscicula      
              +--+                                  
                 |  +-- Unio_pictorum               
                 +--+                               
                    +-- Unio_crassus

Page updated: 07:04:24 Thu 04 Apr 2024
 
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