MUSSELpdb | valid sp. Pronodularia japanensis specimens

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Page last updated
Sun 07 Sep 2025

The Freshwater Mussels (Unionoida) of the World (and other less consequential bivalves)

FM(U)otW(aolcb) is the web version of the MUSSEL Project Database. Follow the links to browse the data or use the custom Google search field. Either way, you win!

This page reports the specimens in the MUSSELpdb and provides a map if geocodes are known. The taxonomic history reports the record of different names and combinations that have been applied to this species (and its synonyms) since 2007.

The MUSSEL Project Database has moved to a new server, with a new URL! The new address for this page is: https://musselpdb.org/db.php?ty=validsp&id=4030

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species Pronodularia japanensis (Lea, 1859): specimens
3 nominal species as synonyms

Taxonomic History (since 2007)

Inversidens japanensis — Graf & Cummings (2007), Graf (2007), Chapman et al. (2008), Chapman et al. (2008), ..., Mitchell et al. (2016), Bogatov & Prozorova (2017), Guerra et al. (2017), Amaro et al. (2017)
Pronodularia japanensis — Kondo et al. (2007), Kondo (2008), Shirai et al. (2010), Whelan et al. (2011), ..., Jeratthitikul et al. (2025), Dai et al. (2025), Dai et al. (2025), Chen et al. (2025)
Unio japanensis — Huang et al. (2013), Burzynski et al. (2017)
“Unio” japanensis — Zhou et al. (2016)
Pronodluaria japanensis — Wu et al. (2016), Wu et al. (2018)
Pronodularia cf. japanensis — Lopes-Lima et al. (2020)

Range: Japan.

Map of Georeferenced Specimens
species map

Japan
ANSP_244865 Hikone, Lake Biwa, Honshu, Japan	BMNH_85-12-10-36-9 Lake Biwa	FMNH_21776 Lake Biwa, Japan
FMNH_20182 near Tokyo, Japan	FMNH_90227 near Tokyo, Japan	UMMZ_109693 Yokohama, Japan
UMMZ_109694 Ogura Yamshiro, Japan	UMMZ_109695 Yone-gawa, Uzen, Japan	UMMZ_109696 Japan
UMMZ_109697 Kisennuma, Rikuzen, Japan	UMMZ_50250 outlet to Koyama Ike, near Tottori, Japan	FMNH_120683 Higashi-tokida, Ise, Japan
FMNH_59344 Japan	FMNH_68075 Oguraike, Japan	FMNH_68297 Uzen, Japan
UMMZ_184020 Lake Otsu, Biwa Province, Japan	USNM_127634 Hitachi, Japan	USNM_346082 Uzen, Japan
BMNH_85-12-10-40-2 Lake Biwa	UMMZ_57113 Japan	SMF_13662 Umgebung von Yokohama [Japan Kanagawa]
USNM_150054 Japan	SMF_168232 loc. paddy fields near Tokyo [Japan]	USNM_152614 Japan
USNM_172691 Japan	USNM_187259 Japan	USNM_124203 paddy fields near Tokyo [Japan]
UMMZ_4438 Tokyo, Japan	INHS_21575 Japan	ANSP_43463 L. Biwa, Japan
UMMZ_23059 Japan	INHS_21576 Japan	INHS_21577 Japan
BMNH_1907-12-21-82-85 Japan	ANSP_48096 River Biwa, Yokohama, Japan	ANSP_49616 Kamakura, Japan
ANSP_49692 Japan	USNM_218088 Ogura, Yamashiro, Japan	ANSP_119929 Kamakura, Japan
MNHN_MP_2800 Japan	MNHN_MP_2801 Japan	MNHN_MP_2802 Japan
MNHN_MP_2803 Japan	UMMZ_110072 Japan	MNHN_MP_2804 Tokio [Japan]
MNHN_MP_2805 Japan	UMMZ_110073 Fongonto, Japan	MNHN_MP_2808 Yokohama, Japan
USNM_249615 Japan	BMNH_MP_0698 Japan	ANSP_80688 Ogura, Yamashiro, Japan
ANSP_80689 Ogura, Yamashiro, Japan	ANSP_80691 Yanakawa, Chikugo, Japan	ANSP_80692 Kizenmura, Rikuzen [Japan]
BMNH_MP_0704 Yamashiro, Japan	MNHN_MP_2818 Tokio [Japan]	MNHN_MP_2820 Kii, Japan
MNHN_MP_2821 Shirosaki [Japan]	UMMZ_23457 Japan	BMNH_1965181 Japan
MNHN_MP_2830 Japan

Genbank Vouchers

Japan

NCSM_27183 (COI, 16S, 28S, mtGenome) Japan

Mp_4138_Pj04-01SUTO (COI, 16S, 28S) Hiroshima, Japan

Mp_4138_Pj14-05f (COI, 16S, 28S) Gifu, Japan

Mp_4138_Pj25-06 (COI, 16S) Sakai, Yamagata, Japan

MHS_PJST-10 (COI, 28S) Arakawa River, Saitama, Japan (36.1229, 139.3433)

Mp_4138_Pj-k (COI, 16S) Sakurai, Nara, Japan

CIIMAR_BIV553 (COI, 28S) Asahi River, Okayama, Japan (34.6892, 133.9644)

MHS_MPj-ok02 (COI, 28S) Asahi River, Okayama, Japan (34.768, 133.9444)

Mp_4138_Pj04-03SUTO (COI, 16S) Hiroshima, Japan

Mp_4138_Pj14-02f (COI, 16S) Gifu, Japan

OMNH_Mo_39133_3 (COI) Yogo River, Shiga, Japan (35.5043, 136.2072)

OMNH_Mo_39130_5 (COI) Yusaka River, Tochigi, Japan (36.9067, 140.0801)

OMNH_Mo_39133_1 (COI) Yogo River, Shiga, Japan (35.5043, 136.2072)

OMNH_Mo_39125_1 (COI) Kase River, Saga, Japan (33.2749, 130.2674)

MHS_PJYG -01 (COI) Akagawa River, Yamagata, Japan (38.7612, 139.8527)

OMNH_Mo_39132_1 (COI) Asahi River, Okayama, Japan (34.6924, 133.9473)

IDEA_Shell008 (28S) Japan

MHS_PJGF-01 (COI) Ijira River, Gifu, Japan (35.5145, 136.732)

MHS_PJTT-01 (COI) Oda River, Tottori, Japan (35.5502, 134.3183)

OMNH_Mo_39124_4 (COI) Harai River, Mie, Japan (34.5573, 136.6078)

IDEA_Shell057 (28S) Japan:Fukui, Tsuruga city, Izumi

OMNH_Mo_39128_3 (COI) Yue River, Nagasaki, Japan (32.8309, 130.322)

OMNH_Mo_39132_2 (COI) Asahi River, Okayama, Japan (34.6924, 133.9473)

MHS_PJGF-02 (COI) Mimizu River, Gifu, Japan (35.4498, 136.632)

OMNH_Mo_39128_1 (COI) Yue River, Nagasaki, Japan (32.8309, 130.322)

IDEA_Shell007 (28S) Japan

OMNH_Mo_39129_3 (COI) Kiyotake River, Miyazaki, Japan (31.8464, 131.425)

CIIMAR_BIV2695 (COI, 16S, 28S)

G3813 (COI)

G3814 (COI)

G3816 (COI)

G3812 (COI)

G3811 (COI)

G3815 (COI)

Page updated: 08:28:06 Sun 07 Sep 2025

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