| Page last updated
Sun 07 Sep 2025 |
The Freshwater Mussels (Unionoida) of the World (and other less consequential bivalves)
FM(U)otW(aolcb) is the web version of the MUSSEL Project Database. Follow the links to browse the data or use the custom Google search field. Either way, you win!
This page documents data for Dai et al. (2025), such as genera and species attributed to that publication (including known misspellings, misidentifications, and other unavailable names) as well as taxonomic opinions, if available. Taxonomic opinions refer to uses of genera and species as valid and invalid in that work. The MUSSEL Project Database has moved to a new server, with a new URL! The new address for this page is: https://musselpdb.org/db.php?ty=publ&id=4440
Search the database pages:
|
|
Publication information
Dai, Y.-T., Z.-G. Chen, S. Ouyang, X.-C. Huang & X.-P. Wu. 2025. A new tribe, genus, and species of freshwater mussel from the Changjiang River Basin in China (Bivlavia, Unionidae, Unioninae). Zoosystematics and Evolution 101(2): 779-790.
Click here for online access.
Cladogram Figures
(click the figure number to see the cladogram) fig. 1
fig. 2
fig. 3
Attributed Genus Names
Globunio Dai et al., 2025
Dai, Chen, Huang & Wu in Dai et al., 2025, Zoosyst. Evol. 101(2): 786.
Type species: Globunio mirificus Dai et al., 2025 (orig. desig.) = Globunio mirificus |
Attributed Species Names
Lasmigona compressa Graf & Ó Foighil, 2000
‘Lea’ of Authors.
Graf & Ó Foighil, 2000, J. Moll. Studies 66: 160.
Graf & Cummings, 2006, Zool. J. Linn. Soc. 148: 348.
Hoeh et al., 2009, Malacologia 51(2): 310.
Jadhav & Jamkhedkar, 2009, Res. J. BioTech. 4(1): 43.
Whelan et al., 2011, MPE 61: 506.
Lopes-Lima et al., 2017, MPE 106: 177.
Bolotov et al., 2017, Sci. Repts. 7: Table S1.
Bolotov et al., 2017, Sci. Repts. 7: Table S1.
Bolotov et al., 2018, Sci. Repts. 8: Table S1.
Huang et al., 2019, MPE 130: Table S4.
Lopes-Lima et al., 2020, MPE 146: Table 1.
Wu et al., 2021, J. Lake Sci. 33(6): attached Table 1.
Jeratthitikul et al., 2022, Sci. Repts. 12: Table S1.
Wu et al., 2023, ZooKeys 1166: 263, Table 1.
Liu et al., 2023, ZooKeys 1179: Table 1.
Dai et al., 2025, Zoosyst. Evol. 101(2): Table 1. [in part]
(misidentified reference to Symphynota compressa Lea, 1829)
|
Unio crassus Soroka, 2005
‘Philippson’ of Authors.
Soroka, 2005, Zool. Sci. 22: 1138, Table 1.
Soroka, 2010, Folia Malac. 18(4): Table 2. [in part]
Reis et al., 2013, Graellsia 69(1): 22, Table 2.
Prié & Puillandre, 2014, Hydrobiologia 735(1): online resource 1 [as ‘Unio crassus crassus’].
Douda et al., 2014, J. Appl. Ecol. 51: 1087. [in part]
Froufe et al., 2016, Org Divers Evol 16(3): 606, fig. 3. [in part]
Klishko et al., 2017, Zootaxa 4286(1): 98, Table 3.
Soroka & Burzynski, 2017, mtDNA B 2(2): 863.
Burzynski et al., 2017, MPE 107: 606.
Lopes-Lima et al., 2017, Biol. Rev. 92(1): Table S1 [as ‘Unio crassus crassus’]. [in part]
Araujo et al., 2018, J. Linn. Soc., Zool. 182(2): supplementary table. [in part]
Klishko et al., 2018, Systemat. Biodiv. 16(3): 291.
Burzynski & Soroka, 2018, PeerJ 6, e5573: 3.
Wang et al., 2018, Conserv. Genet. Resour. 10(4): 611.
Wang et al., 2018, Conserv. Genet. Resour. 11: 140.
Cha et al., 2018, mtDNA B 3(2): 946.
Aunins et al., 2018, mtDNA B 3(2): 1125.
Wu et al., 2018, PLoS One 13(8), e0200956: 6. [in part]
Klishko et al., 2019, Ruthenica 29(1): 57, Table 1.
Wu et al., 2019, ZooKeys 812: 29.
Huang et al., 2019, MPE 130: Table S5. [in part]
Fukata & Iigo, 2019, mtDNA B 4(2): 3489.
Guerra et al., 2019, BMC Evol. Biol. 19 (229): Table S1.
Froufe et al., 2019, Heredity 124: 185.
Soroka, 2020, J. Zool. Syst. Evol. Res. 58(2): Table 2.
Fukata & Iigo, 2020, mtDNA B 5(2): 1216.
Sano et al., 2020, Plankton Benthos Res. 15(2): 99.
Wu et al., 2020, Invert. Syst. 34: Table S2.
Kilikowska et al., 2020, Life 10(7): 1. [in part]
Lopes-Lima et al., 2021, MPE 163: fig. 6 [by voucher number]. [in part]
Sayenko et al., 2021, Systemat. Biodiv. 19: 58, Table 1.
Wu et al., 2021, J. Lake Sci. 33(6): attached Table 2.
Zieritz et al., 2021, Zool. J. Linn. Soc. 193(3): 1092.
Kim et al., 2021, mtDNA B 6(1): 80.
Pfarrer et al., 2022, Hydrobiologia 849: ESM1.
Wu et al., 2022, Zool. Scripta 51: Table S2.
Dai et al., 2023, Invert. Syst. 37(2): Table 2.
Chen et al., 2023, ZooKeys 1173: Table 1.
Wu et al., 2023, Org Divers Evol (2023): Table 2.
Dai et al., 2024, Zool. Scripta 53: Table 1.
Wu et al., 2024, J. Zool. Syst. Evol. Res. 2024, 1499508: Table 2.
Wu et al., 2024, Zool. Scripta early online: Table S3.
Dai et al., 2024, Invert. Syst. 38(6): Table 3. [in part]
Chen et al., 2025, Zoosyst. Evol. 101(4): Tables S2-S3.
Dai et al., 2025, Zoosyst. Evol. 101(2): Tables 1-2.
(misidentified reference to Unio crassus Retzius, 1788)
|
Chamberlainia hainesiana Kongim et al., 2015
‘Lea’ of Authors.
Kongim et al., 2015, ZooKeys 514: 98, fig. 2A.
Lopes-Lima et al., 2017, MPE 106: 177.
Bolotov et al., 2017, Sci. Repts. 7: Table S1.
Bolotov et al., 2017, Sci. Repts. 7: Table S1.
Froufe et al., 2019, Heredity 124: 185.
Huang et al., 2019, MPE 130: Table S4.
Chen et al., 2020, mtDNA B 5(3): 2800.
Teiga-Teixeira et al., 2020, mtDNA B 5(3): 2978.
Zieritz et al., 2020, Biodivers. Conserv. 29: 2244.
Bolotov et al., 2020, Sci. Repts. 10: Table S1.
Lopes-Lima et al., 2020, MPE 146: Table 1.
Zieritz et al., 2021, Zool. J. Linn. Soc. 193(3): Table 1.
Zieritz et al., 2021, Aquatic Conserv.: Mar. Freshw. Ecosyst. 31: 3176.
Jeratthitikul et al., 2021, Eur. J. Taxonomy 775: 125.
Wu et al., 2021, ZooKeys 1054: 87.
Wu et al., 2021, J. Lake Sci. 33(6): attached Table 1.
Wu et al., 2022, Zool. Scripta 51: Table S2.
Pavan-Kumar et al., 2022, Mol. Biol. Rep. 49: Table S1.
Jeratthitikul et al., 2022, Sci. Repts. 12: Table S1.
Dai et al., 2023, Invert. Syst. 37(2): Tables 1-2.
Konopleva et al., 2023, Zoosyst. Evol. 99(1): Table S1.
Wu et al., 2023, Org Divers Evol (2023): Table 2.
Dai et al., 2024, Zool. Scripta 53: Table 1.
Dai et al., 2024, Invert. Syst. 38(6): Table 3.
Dai et al., 2024, Zoosyst. Evol. 100(4): Table 2.
Chen et al., 2025, Zoosyst. Evol. 101(4): Table S3.
Dai et al., 2025, Zoosyst. Evol. 101(2): Table 1.
Dai et al., 2025, Zoosyst. Evol. 101(2): Table 2.
(misidentified reference to Unio hainesianus Lea, 1856)
|
Cuneopsis heudei Lopes-Lima et al., 2017
‘Heude’ of Authors.
Liu et al., 2017, PLoS One 12(12 ), e0189737: Table S4.
Lopes-Lima et al., 2017, MPE 106: Table 2.
Bolotov et al., 2017, Sci. Repts. 7: Table S1.
Bolotov et al., 2017, Sci. Repts. 7: Table S1.
Wu et al., 2018, PLoS One 13(8), e0200956: 5.
Huang et al., 2019, MPE 130: Tables S1, S4. [in part]
Lopes-Lima et al., 2020, MPE 146: Table 1. [in part]
Wu et al., 2021, J. Lake Sci. 33(6): attached Table 1. [in part]
Wu et al., 2022, Zool. Scripta 51: Table S1. [in part]
Jeratthitikul et al., 2022, Sci. Repts. 12: Table S1.
Konopleva et al., 2023, Zoosyst. Evol. 99(1): Table S1.
Chen et al., 2023, ZooKeys 1173: Table 1.
Wu et al., 2023, ZooKeys 1166: Table 1.
Liu et al., 2023, ZooKeys 1179: Table 1.
Chen et al., 2025, Zoosyst. Evol. 101(4): Table S2. [in part]
Dai et al., 2025, Zoosyst. Evol. 101(2): Table 1. [in part]
(misidentified reference to Unio heudei Heude, 1874)
|
Globunio mirificus Dai et al., 2025
Chen, Dai, Huang & Wu in Dai et al., 2025, Zoosyst. Evol. 101(2): 786, fig. 4.
Type(s): 24_NCU_XPWU_GM01, Honggutan District [红谷滩区], Nanchang City [南昌市], Jiangxi Province [江西省], China.
|
Taxonomic Opinions
Aculamprotula coreana (Martens, 1886)
Aculamprotula fibrosa (Heude, 1877)
Aculamprotula polysticta (Heude, 1877)
Aculamprotula scripta (Heude, 1875)
Aculamprotula tientsinensis (Crosse & Debeaux, 1863)
Aculamprotula tortuosa (Lea, 1865)
Acuticosta chinensis (Lea, 1868)
Alasmidonta marginata Say, 1818
Amblema plicata (Say, 1817)
Amuranodonta kijaensis Moskvicheva, 1973
Anemina arcaeformis (Heude, 1877)
Anemina euscaphys (Heude, 1879)
Anodonta anatina (Linnaeus, 1758)
Anodonta cygnea (Linnaeus, 1758)
Anodontites trapesialis (Lamarck, 1819)
Beringiana beringiana (Middendorff, 1851)
Chamberlainia hainesiana (Lea, 1856)
Coelatura aegyptiaca (Cailliaud, 1823)
Cristaria plicata (Leach, 1814)
Cumberlandia monodonta (Say, 1829)
Cuneopsis celtiformis (Heude, 1874)
Cuneopsis demangei Haas, 1929
Cuneopsis heudei (Heude, 1874)
Cuneopsis rufescens (Heude, 1874)
Diaurora aurorea (Heude, 1883)
Echyridella menziesii (Gray, 1843)
Elliptio complanata (Lightfoot, 1786)
Gibbosula rochechouartii (Heude, 1875)
Globunio mirificus Chen, Dai, Huang & Wu, sp. nov.
Hyriopsis bialata Simpson, 1900
Indonaia andersoniana (Nevill, 1877)
Inversidens rentianensis Wu & Wu, 2021
Inversiunio reinianus (Kobelt, 1879)
Inversiunio yanagawensis (Kondo, 1982)
Lamprotula caveata (Heude, 1877)
Lamprotula gottschei-Schistodesmus.sp [sic]
Lamprotula leaii (Gray, 1833)
Lampsilis cardium Rafinesque, 1820
Lampsilis ornata (Conrad, 1835)
Lampsilis powellii (Lea, 1852)
Lampsilis siliquoideus (Barnes, 1823)
Lanceolaria gladiola (Heude, 1877)
Lanceolaria grayii (Gray, 1833)
+ Unio grayanus Lea, 1834
Lanceolaria lanceolata (Lea, 1856)
Lanceolaria oxyrhyncha (Martens, 1861)
Lasmigona compressa (Lea, 1829)
Leaunio lienosa (Conrad, 1834)
Lens contradens (Lea, 1838)
Lepidodesma languilati (Heude, 1874)
Margaritifera dahurica (Middendorff, 1850)
Margaritifera falcata (Gould, 1850)
Margaritifera margaritifera (Linnaeus, 1758)
Microcondylaea bonellii (Férussac, 1827)
Monodontina vondembuschiana (Lea, 1840)
Mutela dubia (Gmelin, 1791)
Nodularia breviconcha Lee, Kim, Bogan & Kondo, 2020
Nodularia douglasiae (Gray, 1833)
Nodularia fusiformans Wu & Liu, 2024
Nodularia nuxpersicae (Dunker, 1848)
Parreysia nagpoorensis (Lea, 1860)
Parvasolenaia rivularis (Heude, 1877)
Physunio superbus (Lea, 1843)
Pilsbryoconcha exilis (Lea, 1838)
Pletholophus tenuis (Griffith & Pidgeon, 1833)
Pleurobema oviforme (Conrad, 1834)
Popenaias popeii (Lea, 1857)
Postolata guangxiensis Dai, Huang, Guo & Wu, 2023
Potamilus alatus (Say, 1817)
Potamilus leptodon (Rafinesque, 1820)
Potamilus streckersoni
Potomida littoralis (Cuvier, 1798)
Pronodularia japanensis (Lea, 1859)
Pseudanodonta complanata (Rossmässler, 1835)
Pseudocuneopsis capitata (Heude, 1874)
Pseudocuneopsis sichuanensis Huang, Dai, Chen & Wu , 2022
Pseudocuneopsis yangshuoensis Wu & Liu, 2023
Pseudocuneopsis yemaoi Dai, Chen, Huang & Wu, 2024
Pseudunio marocanus (Pallary, 1918)
Ptychorhynchus pfisteri (Heude, 1874)
Pyganodon grandis (Say, 1829)
Quadrula quadrula (Rafinesque, 1820)
Rectidens sumatrensis (Dunker, 1852)
Schistodesmus lampreyanus (Baird & Adams, 1867)
Schistodesmus spinosus (Simpson, 1900)
Sinanodonta lucida (Heude, 1877)
Sinanodonta woodiana (Lea, 1834)
Sinohyriopsis cumingii (Lea, 1852)
Sinohyriopsis schlegelii (Martens, 1861)
Sinosolenaia carinata (Heude, 1877)
Sinosolenaia oleivora (Heude, 1877)
Tchangsinaia piscicula (Heude, 1874)
Toxolasma parvum (Barnes, 1823)
Unio crassus Philipsson, 1788
Unio delphinus Spengler, 1793
Unio elongatulus
Unio mancus
Unio pictorum (Linnaeus, 1758)
Unio tumidus Philipsson, 1788
Uniomerus tetralasmus (Say, 1831)
Utterbackia imbecillis (Say, 1829)
Utterbackia peninsularis Bogan & Hoeh, 1995
Venustaconcha ellipsiformis (Conrad, 1836)
|
Cladogram Figure 1
+-- Gibbosula_rochechouartii
+--------------------------------------------------+
| +-- Margaritifera_dahurica
|
| +-- Pronodularia_japanensis
--+ +-----------------------------------------------+
| | +-- Lamprotula_leaii
| |
| | +-- Lampsilis_siliquoidea
+--+ +--------------------------------------------+
| | +-- Quadrula_quadrula
| |
| | +-- Anemina_arcaeformis
| | +-----------------------------------------+
+--+ | +-- Amuranodonta_kijaensis
| |
| | +----- Parreysia_nagpoorensis
| +--------------------------------------+
| | | +-- Coelatura_aegyptiaca
| | +--+
+--+ +-- Indonaia_andersoniana
|
| +----- Pletholophus_reinianus
| +-----------------------------------+
| | | +-- Sinanodonta_woodiana
| | +--+
| | +-- Beringiana_beringiana
| |
+--+ +-------------------------------------- Cristaria_plicata
| |
| | +-- Alasmidonta_marginata
| | +--+
| | | +-- Lasmigona_costata
| +--------------------------------+
+--+ | +-- Anodonta_anatina
| +--+
| +-- Pseudanodonta_complanata
|
| +-- Lanceolaria_gladiola
| +--------------------------------+
| | +-- Lanceolaria_oxyrhyncha
+--+
| +-------------------------------- Lepidodesma_languilati
| |
+--+ +----------------------------- Acuticosta_chinensis
| |
| | +-------------------------- Globunio_mirificus
+--+ |
| | +-------- Aculamprotula_tortuosa
| | |
| | +--------------+ +----- Aculamprotula_coreana
+--+ | | |
| | +--+----- Aculamprotula_fibrosa
| | |
| | | +-- Aculamprotula_polysticta
| | +--+
+--+ +-- Aculamprotula_scripta
|
| +-------------------- Middendorffinaia_mongolica
| |
| | +-- Nodularia_douglasiae
| | +--------------+
+--+ | +-- Nodularia_breviconcha
| |
| | +-- Cuneopsis_celtiformis
| | +-----------+
+--+ | +-- Cuneopsis_heudei
| |
| | +----- Diaurora_aurorea
| | |
| +--------+ +-- Schistodesmus_lampreyanus
| | | |
+--+ +--+-- Schistodesmus_spinosus
| |
| +-- Schistodesmus
|
| +-- Pseudocuneopsis_sichuanensis
| +--------+
| | +-- Pseudocuneopsis_capitata
| |
+--+ +-- Unio_pictorum
| +-----+
| | +-- Unio_crassus
+--+
| +----- Tchangsinaia_piscicula
+--+
| +-- Inversiunio_yanagawensis
+--+
+-- Inversiunio_reinianus
Cladogram Figure 2
+-- Gibbosula_rochechouartii
+--------------------------------------------+
| +-- Margaritifera_dahurica
|
| +-- Pronodularia_japanensis
| +--+
| | +-- Lamprotula_leaii
--+ +--------------------------------------+
| | | +-- Lampsilis_siliquoidea
| | +--+
| | +-- Quadrula_quadrula
| |
| | +-- Coelatura_aegyptiaca
+--+ |
| +--------------------------------------+-- Indonaia_andersoniana
| | |
| | +-- Parreysia_nagpoorensis
| |
| | +-- Anemina_arcaeformis
+--+ +-----------------------------------+
| | +-- Amuranodonta_kijaensis
| |
| | +----- Pletholophus_reinianus
| | +-----------------------------+
+--+ | | +-- Sinanodonta_woodiana
| | +--+
| | +-- Beringiana_beringiana
| |
| | +-- Alasmidonta_marginata
+--+ +--+
| | +-- Lasmigona_costata
| +--------------------------+
| | | +-- Anodonta_anatina
| | +--+
| | +-- Pseudanodonta_complanata
| |
+--+ +----------------------------- Lanceolaria_gladiola
| |
| +----------------------------- Lanceolaria_oxyrhyncha
| |
| +----------------------------- Cristaria_plicata
+--+
| +-------------------------- Acuticosta_chinensis
| |
| +-------------------------- Lepidodesma_languilati
| |
+--+ +----------------------- Globunio_mirificus
| |
| | +-------- Aculamprotula_tortuosa
| | |
| | +-----------+ +-- Aculamprotula_fibrosa
+--+ | | +--+
| | | | +-- Aculamprotula_coreana
| | +--+
| | | +-- Aculamprotula_scripta
| | +--+
| | +-- Aculamprotula_polysticta
+--+
| +----- Middendorffinaia_mongolica
| +-----------+
| | | +-- Nodularia_douglasiae
| | +--+
| | +-- Nodularia_breviconcha
| |
| | +-- Cuneopsis_celtiformis
+--+ +-----------+
| | +-- Cuneopsis_heudei
| |
| | +-------- Diaurora_aurorea
| +-----+
| | | +----- Schistodesmus_spinosus
| | +--+
+--+ | +-- Schistodesmus_lampreyanus
| +--+
| +-- Schistodesmus
|
| +-- Pseudocuneopsis_sichuanensis
| +--------+
| | +-- Pseudocuneopsis_capitata
| |
+--+ +-- Inversiunio_yanagawensis
| +-----+
| | +-- Inversiunio_reinianus
+--+
| +----- Tchangsinaia_piscicula
+--+
| +-- Unio_pictorum
+--+
+-- Unio_crassus
Cladogram Figure 3
+-- Anodontites_trapesialis
+--------------------------------------------+
| +-- Mutela_dubia
|
| +-------------------------------------------- Echyridella_menziesii
| |
| | +-------------- Gibbosula_rochechouartii
| | +--------------------------+
--+ | | | +----------- Cumberlandia_monodonta
| | | +--+
| | | | +-------- Pseudunio_marocanus
| | | +--+
| | | | +----- Margaritifera_falcata
| | | +--+
| | | | +-- Margaritifera_dahurica
| | | +--+
+--+ | +-- Margaritifera_margaritifera
| |
| | +-- Quadrula_quadrula
| | +--------------------------+
| | | +-- Uniomerus_tetralasmus
| | |
| | +--------+ +-- Elliptio_complanata
| | | | +-----------------------+
| | | | | +-- Pleurobema_oviforme
| | | +--+
| | | | +----------------------- Amblema_plicata
| | | | |
+--+ | +--+ +-------------------- Popenaias_popeii
| | | |
| | +--+ +----------------- Toxolasma_parvum
| | | |
| | | | +----- Potamilus_leptodon
| | +--+ +--------+
| | | | | +-- Potamilus_streckersoni
| | | | +--+
| | +--+ +-- Potamilus_alatus
| | |
| | | +----------- Leaunio_lienosus
| | | |
| | +--+ +-- Lampsilis_cardium
| | | +-----+
| | | | +-- Lampsilis_ornata
| | +--+
| | | +----- Venustaconcha_ellipsiformis
| | +--+
| | | +-- Lampsilis_powellii
| | +--+
| | +-- Lampsilis_siliquoidea
| |
+--+ +----- Chamberlainia_hainesiana
| +-----------------+
| | | +-- Sinohyriopsis_cumingii
| | +--+
| | +-- Sinohyriopsis_schlegelii
| |
| | +-- Lens_contradens
| +-----------+ +--+
| | | | +-- Physunio_superbus
| | | +--------------+
| | | | | +-- Hyriopsis_bialata
| | | | +--+
| | | | +-- Rectidens_sumatrensis
| | | |
| | +--+ +-------- Potomida_semirugata
| | | +--------+
| | | | | +----- Pronodularia_japanensis
| | | | +--+
| | | | | +-- Lamprotula_caveata
| | | | +--+
| | +--+ +-- Lamprotula_leaii
| | |
| | | +-- Pseudodon_vondembuschianus
| | | +-----------+
| | | | +-- Pilsbryoconcha_exilis
| | | |
| | +--+ +----------- Inversiunio_reinianus
| | | |
| | | | +----- Parvasolenaia_rivularis
+--+ +--+ +--+
| | | | +-- Postolata_guangxiensis
| | | +--+
| +--+ +-- Ptychorhynchus_pfisteri
| |
| | +----- Microcondylaea_bonellii
| +--+
| | +-- Sinosolenaia_carinata
| +--+
| +-- Sinosolenaia_oleivora
|
| +-------------------------------- Lepidodesma_languilati
| |
| | +-------- Acuticosta_chinensis
| | +--------+
| | | | +----- Lanceolaria_gladiola
| | | +--+
| | | | +-- Lanceolaria_lanceolata
| | | +--+
| | | +-- Lanceolaria_grayii
| | +-----------+
| | | | +-------- Cristaria_plicata
+--+ | | +-----+
| | | | | +----- Anemina_arcaeformis
| | | | +--+
| | | | | +-- Sinanodonta_lucida
| | | | +--+
| | +--+ +-- Sinanodonta_woodiana
| | |
| | | +-- Anodonta_anatina
| | | +--------+
| | | | +-- Anodonta_cygnea
| | +--+
| | | +-------- Platynaias_compressa
+--+ +--+
| | +----- Pyganodon_grandis
| +--+
| | +-- Utterbackia_imbecillis
| +--+
| +-- Utterbackia_peninsularis
|
| +-------------------------- Globunio_mirificus
| |
| | +-------- Aculamprotula_tortuosa
| | |
| | +--------------+ +-- Aculamprotula_polysticta
| | | | +--+
| | | | | +-- Aculamprotula_scripta
+--+ | +--+
| | | +-- Aculamprotula_coreana
| | +--+
| | +-- Aculamprotula_tientsinensis
| |
| | +-------------- Unio_tumidus
| | +-----+
+--+ | | +----------- Unio_crassus
| | +--+
| | | +-------- Unio_delphinus
| | +--+
| | | +----- Unio_pictorum
| | +--+
| | | +-- Unio_elongatulus
| | +--+
| | +-- Unio_mancus
| |
+--+ +-------- Nodularia_breviconcha
| +--+
| | | +----- Nodularia_fusiformans
| | +--+
| | | +-- Nodularia_nuxpersicae
| +-----+ +--+
| | | +-- Nodularia_douglasiae
| | |
| | | +-------- Pseudocuneopsis_capitata
| | +--+
| | | +----- Pseudocuneopsis_yangshuoensis
| | +--+
+--+ | +-- Pseudocuneopsis_sichuanensis
| +--+
| +-- Pseudocuneopsis_yemaoi
|
| +----- Schistodesmus_spinosus
| +--------+
| | | +-- Schistodesmus_lampreyanus
| | +--+
+--+ +-- Schistodesmus
|
| +----------- Tchangsinaia_piscicula
+--+
| +-------- Cuneopsis_rufescens
+--+
| +----- Cuneopsis_celtiformis
+--+
| +-- Cuneopsis_heudei
+--+
+-- Cuneopsis_demangei
|
Page updated: 08:31:00 Sun 07 Sep 2025 |
|
