|
Bivalvia | Unionoida | Unionoidea | Unionidae | Unioninae tribe ANODONTINI Rafinesque, 1820: cladomics
3 subtribes, 1 genus
Cladomics Summary
number of species = 111
number of subtaxa (rank: subtribe) = 4
analyzed in 307 cladograms in 172 publications
maximum number of terminals in an analysis = 44, mean = 6.18
maximum number of subtaxa (subtribe) in an analysis = 3, mean = 1.78
Basic Monophyly
number of cladograms that test basic monophyly (i.e., more than 1 included species) = 259
maximum number of included terminals = 44, mean = 7.14
maximum number of subtaxa (subtribe) = 3, mean = 1.92
proportion of cladograms that support basic monophyly = 0.86
Rigorous Monophyly
number of cladograms that test rigorous monophyly = 118
(Rigorous monophyly refers to cladograms with terminal species representing more than one subtaxon (subtribe) taxon and at least one rigorous outgroup (tribe).
maximum number of included terminals = 33, mean = 8.35
maximum number of subtaxa (subtribe) = 3, mean = 2.58
proportion of cladograms that support rigorous monophyly = 0.78
Pairwise Cladogram Clustering
The graphs below depict in two dimensions all pair-wise ingroup distances within the cladome. See the Cladmoics FARQ for more information.
Chen et al., 2023 [Publ 4302], fig. 1
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 10 |
31 |
23 |
3 |
30 |
21 |
4 |
0.0 |
*** Top Monophyly Cladogram ***
+-- Anemina_arcaeformis **
+--------------------------+
| +-- Amuranodonta_kijaensis **
|
| +----- Pletholophus_tenuis **
| +--------------------+
| | | +-- Sinanodonta_jourdyi **
--+ | +--+
| | +-- Beringiana_beringiana **
| |
| | +----------------------- Lanceolaria_gladiola
| | |
| | +----------------------- Acuticosta_chinensis
+--+ |
| +----------------------- Lepidodesma_languilati
| |
| +----------------------- Lanceolaria_oxyrhyncha
| |
| +----------------------- Cristaria_plicata **
| |
| | +-- Alasmidonta_marginata **
+--+ +--+
| | +-- Lasmigona_costata **
+-----------------+
| | +-- Anodonta_anatina **
| +--+
| +-- Pseudanodonta_complanata **
|
| +-------------------- Aculamprotula_fibrosa
| |
| | +----- Middendorffinaia_mongolica
| | +-----------+
+--+ | | +-- Nodularia_douglasiae
| | +--+
| | +-- Nodularia_breviconcha
| |
| | +-- Cuneopsis_celtiformis
+--+ +-----------+
| | +-- Cuneopsis_heudei
| |
| | +-- Diaurora_aurorea
| | +--+
| | | +-- Diaurora_laeve
| +--------+
+--+ | +-- Schistodesmus_lampreyanus
| +--+
| +-- Schistodesmus_spinosus
|
| +-- Pseudocuneopsis_sichuanensis
| +--------+
| | +-- Pseudocuneopsis_capitata
| |
+--+ +-- Inversiunio_yanagawensis
| +-----+
| | +-- Inversiunio_reinianus
+--+
| +----- Tchangsinaia_piscicula
+--+
| +-- Unio_pictorum
+--+
+-- Unio_crassus
Huang et al., 2019 [Publ 3843], fig. S1
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 18 |
55 |
40 |
1 |
57 |
37 |
4 |
59.0 |
*** Top Sister-Group Cladogram ***
+-------------- Anemina_arcaeformis[2] **
|
+-----------------+ +----------- Anemina_arcaeformis[5] **
| | |
| | | +-------- Anemina_arcaeformis[3] **
| +--+ |
| | +-------- Anemina_arcaeformis[7] **
| | |
| +--+ +----- Anemina_arcaeformis[4] **
| | |
| | +----- Anemina_arcaeformis[6] **
| | |
| +--+----- Anemina_arcaeformis[8] **
| |
| +----- Anemina_arcaeformis[9] **
| |
| | +-- Anemina_arcaeformis[1] **
| +--+
| +-- Anemina_euscaphys **
--+
| +-- Sinanodonta_angula **
| +-----------+
| | +-- Sinanodonta_lucida **
| +--------------+
| | | +----------- Sinanodonta_woodiana[2] **
| | +--+
| | | +-------- Sinanodonta_jourdyi **
| | +--+
| | | +----- Sinanodonta_elliptica **
| | +--+
| | | +-- Sinanodonta_woodiana[1] **
| | +--+
| | +-- Sinanodonta_lauta **
| |
| | +-------------- Acuticosta_ovata[1]
| | |
| | | +----------- Acuticosta_chinensis[3]
+--+ +-----------+ |
| | | +----------- Acuticosta_chinensis[4]
| | | |
| | +--+----------- Acuticosta_chinensis[6]
| | |
| | | +-- Acuticosta_chinensis[1]
| | | +-----+
| | | | +-- Acuticosta_ovata[2]
| | +--+
| | | +----- Acuticosta_chinensis[5]
| | | |
| | +--+----- Acuticosta_chinensis[7]
| | |
| | | +-- Acuticosta_chinensis[2]
+--+ +--+
| +-- Acuticosta_ovata[3]
|
| +-- Lepidodesma_languilati
| +--------------------+
| | +-- Cristaria_plicata **
| |
| | +----------- Lanceolaria_gladiola
| | |
| | | +-------- Lanceolaria_grayii[5]
| | +--------+ |
| | | | | +-- Lanceolaria_lanceolata
| | | | +-----+
| | | | | +-- Lanceolaria_triformis
+--+ | +--+
| | | +----- Lanceolaria_grayii[4]
| | | |
| | | | +-- Lanceolaria_eucylindrica[1]
| | | | |
| | +--+ +-- Lanceolaria_eucylindrica[2]
| | | |
| | | +-- Lanceolaria_eucylindrica[3]
| | | |
| | +--+-- Lanceolaria_eucylindrica[4]
+--+ |
| +-- Lanceolaria_grayii[1]
| |
| +-- Lanceolaria_grayii[2]
| |
| +-- Lanceolaria_grayii[3]
|
| +----------- Aculamprotula_tortuosa
| +-----+
| | | +-------- Aculamprotula_tientsinensis
| | +--+
| | | +----- Aculamprotula_fibrosa
| | +--+
| | | +-- Aculamprotula_coreana
+--+ +--+
| +-- Aculamprotula_scripta
|
| +----- Nodularia_douglasiae[1]
| +--------+
| | | +-- Nodularia_nuxpersicae
| | +--+
+--+ +-- Nodularia_douglasiae[2]
|
| +----------- Tchangsinaia_piscicula
| |
+--+ +-- Schistodesmus_spinosus
| +-----+
| | +-- Schistodesmus_lampreyanus
+--+
| +----- Cuneopsis_rufescens
+--+
| +-- Cuneopsis_heudei
+--+
+-- Cuneopsis_celtiformis
Whelan et al., 2023 [Publ 4287], fig. 2
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 44 |
44 |
42 |
3 |
0 |
0 |
0 |
67.0 |
+-------- Anemina_arcaeformis **
+-----------------------------+
| | +----- Sinanodonta_lucida **
| +--+
| | +-- Sinanodonta_woodiana **
| +--+
--+ +-- Sinanodonta_tumens **
|
| +----------------------------------- Anodonta_cygnea **
| |
| | +-- Anodonta_nuttalliana **
| | +-----------------------------+
+--+ | +-- Pseudanodonta_complanata **
| |
| | +----------------------------- Anodonta_anatina **
| | |
| | | +----- Utterbackia_peninsularis[2] **
+--+ | +--+
| | | | +-- Utterbackia_peninsularis[1] **
| | | +--+
| | +-----------------+ +-- Utterbackia_imbecillis **
| | | |
| | | | +----- Pyganodon_grandis[2] **
+--+ | +--+
| | | +-- Pyganodon_grandis[1] **
| | +--+
| | +-- Pyganodon_lacustris **
| |
| | +-- Utterbackiana_couperiana **
| | +--+
| | | +-- Utterbackiana_heardi **
| | +-----+
| | | | +-- Utterbackiana_suborbiculata **
+--+ | +--+
| | +-- Utterbackiana_hartfieldorum **
| +-----------+
| | | +----- Lasmigona_costata **
| | | +--+
| | | | | +-- Pseudodontoideus_subvexus **
| | | | +--+
| | +--+ +-- Pseudodontoideus_connasaugaensis **
| | |
| | | +----- Lasmigona_complanata **
| | +--+
| | | +-- Arcidens_confragosus **
| | +--+
| | +-- Arcidens_wheeleri **
+--+
| +----------------- Pressodonta_viridis **
| |
| +--+ +-- Anodontoides_denigratus **
| | | +-----------+
| | | | +-- Anodontoides_ferussacianus **
| | +--+
| | | +----------- Strophitus_undulatus **
| | +--+
| | | +-------- Strophitus_pascagoulaensis[2] **
| | +--+
| | | +----- Strophitus_radiatus **
+--+ +--+
| | +-- Strophitus_pascagoulaensis[1] **
| +--+
| +-- Strophitus_williamsi **
|
| +----- Prolasmidonta_heterodon **
| +-----------+
| | | +-- Platynaias_subviridis **
| | +--+
| | +-- Platynaias_compressa **
+--+
| +-- Alasmidonta_holstonia **
| +-----------+
| | +-- Alasminota_etowaensis **
| |
+--+ +-- Alasmidonta_raveneliana **
| +--------+
| | +-- Alasmidonta_marginata **
| |
+--+ +-- Alasmidonta_varicosa **
| +-----+
| | +-- Alasmidonta_uwharriensis **
+--+
| +----- Alasmidonta_undulata **
+--+
| +-- Alasmidonta_triangulata **
+--+
+-- Alasmidonta_arcula **
Lopes-Lima et al., 2020 [Publ 4080], fig. 2-4
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 33 |
33 |
25 |
3 |
89 |
36 |
4 |
47.0 |
+-------------------------------- Acuticosta_chinensis
|
+-------------------------------- Lepidodesma_languilati
|
| +----------- Aculamprotula_tortuosa
| +--------------+
| | | +-------- Aculamprotula_scripta
| | +--+
| | | +----- Aculamprotula_coreana
| | +--+
| | | +-- Aculamprotula_fibrosa
| | +--+
+-----+ +-- Aculamprotula_tientsinensis
| |
| | +----------------------- Middendorffinaia_mongolica
| | |
| | | +----- Unio_gibbus
| | | |
| | | +--------------+----- Unio_tumidus
| +--+ | |
| | | | +-- Unio_crassus
| | | +--+
| | | +-- Unio_pictorum
| | |
| | | +-------------- Nodularia_breviconcha
| +--+ +--+
| | | | +----------- Nodularia[2]
| | | +--+
| | | | +-------- Nodularia_nuxpersicae
| | | +--+
| | | | +----- Nodularia_nipponensis
| | | +--+
| | | | +-- Nodularia_douglasiae
| +--+ +--+
| | +-- Nodularia[1]
| |
--+ | +-- Schistodesmus_lampreyanus
| | +-----------+
| | | +-- Schistodesmus_spinosus
| | |
| | | +----- Cuneopsis_rufescens
| | +--------+
| +--+ | +-- Cuneopsis_heudei
| | +--+
| | +-- Cuneopsis_celtiformis
| |
| | +----------- Tchangsinaia_piscicula
| +--+
| | +-------- Inversiunio_jokohamensis
| +--+
| | +----- Inversiunio_verrucosus
| +--+
| | +-- Inversiunio_reinianus
| +--+
| +-- Inversiunio_yanagawensis
|
| +----------------- Lanceolaria_oxyrhyncha
| |
| +-----------+ +-------------- Lanceolaria_gladiola
| | | |
| | +--+ +-- Lanceolaria_bilirata
| | | +--------+
| | | | +-- Lanceolaria_grayii
| | +--+
| | | +-------- Lanceolaria_acrorrhyncha
| | +--+
| | | +----- Lanceolaria_kihirai
| | +--+
| | | +-- Lanceolaria_lanceolata
| | +--+
| | +-- Lanceolaria_triformis
| |
| | +-------- Pseudanodonta_complanata **
+--+ |
| | +----- Anodonta_cygnea **
| +--+
| | | +-- Anodonta_anatina **
| +-----------------+ +--+
| | | +-- Anodonta_nuttalliana **
| | |
| | | +----- Pyganodon_grandis **
| | | |
| | | +----- Simpsonaias_ambigua **
| | +--+
| | +----- Strophitus_undulatus **
| | |
| | | +-- Alasmidonta_marginata **
| | +--+
| | +-- Lasmigona_costata **
+--+
| +-- Cristaria_plicata **
| +--------------------+
| | +-- Cristaria_clessini **
| |
| | +-------------- Anemina_arcaeformis **
| | |
| +--------+ +----------- Amuranodonta_kijaensis **
| | | |
| | +--+ +-------- Buldowskia_suifunica **
| | | |
| | +--+ +-- Buldowskia_flavotincta **
+--+ | +--+
| | | +-- Buldowskia_shadini **
| +--+
| | +-- Buldowskia_iwakawai **
| +--+
| +-- Buldowskia_kamiyai **
|
| +-------------------- Pletholophus_tenuis **
| |
| | +-------- Beringiana_japonica **
| | +--------+
+--+ | | +----- Beringiana_beringiana **
| | +--+
| | | +-- Beringiana_fukuharai **
| | +--+
+--+ +-- Beringiana_gosannensis **
|
| +-- Sinanodonta_angula **
| +-----------+
| | +-- Sinanodonta_lucida **
| |
+--+ +----------- Sinanodonta_schrenkii **
| |
| | +-- Sinanodonta_calipygos **
| +--------+
+--+ +-- Sinanodonta_tumens **
|
| +-------- Sinanodonta_woodiana[2] **
| |
+--+ +----- Sinanodonta_elliptica **
| |
+--+----- Sinanodonta_jourdyi **
|
| +-- Sinanodonta_lauta **
+--+
+-- Sinanodonta_woodiana[1] **
Mabuchi & Nishida, 2020 [Publ 4120], fig. 1
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 30 |
30 |
5 |
1 |
7 |
3 |
2 |
34.0 |
+----- Nodularia_douglasiae
+-----------+
| | +-- Lanceolaria_grayii
| +--+
| +-- Inversiunio_reinianus
|
--+ +-------------- Cristaria_plicata **
| |
| | +----------- Sinanodonta_tumens **
| | |
| | | +-- Sinanodonta_lauta[5] **
| | | |
+--+ | +-- Sinanodonta_lauta[6] **
| | |
| | +-- Beringiana_japonica[15] **
| | |
| | +-- Beringiana_japonica[16] **
| | +-----+
| | | +-- Beringiana_japonica[17] **
+--+ | |
| | +-- Beringiana_japonica[18] **
| | |
| | +-- Beringiana_japonica[19] **
| | |
| | +-- Beringiana_japonica[20] **
| |
| | +----- Beringiana_japonica[1] **
| | |
| | +----- Beringiana_japonica[2] **
| | |
+--+ +----- Beringiana_japonica[3] **
| |
| +----- Beringiana_japonica[4] **
| |
| +----- Beringiana_japonica[5] **
| |
| +----- Sinanodonta_lauta[1] **
| |
| +----- Beringiana_japonica[6] **
| |
| +----- Sinanodonta_lauta[2] **
| |
| +----- Sinanodonta_calipygos[1] **
| |
+--+----- Beringiana_japonica[7] **
|
+----- Sinanodonta_lauta[4] **
|
+----- Beringiana_japonica[8] **
|
+----- Beringiana_japonica[9] **
|
+----- Beringiana_japonica[10] **
|
+----- Beringiana_japonica[11] **
|
+----- Beringiana_japonica[12] **
|
+----- Beringiana_japonica[13] **
|
+----- Beringiana_japonica[14] **
|
| +-- Sinanodonta_lauta[3] **
+--+
+-- Sinanodonta_calipygos[2] **
Whelan et al., 2023 [Publ 4287], fig. 3
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 29 |
29 |
27 |
2 |
0 |
0 |
0 |
55.0 |
+-------------------------------- Anodonta_anatina **
|
| +-- Utterbackiana_heardi **
| +--+
| | +-- Utterbackiana_couperiana **
| +-----------------------+
--+ | | +-- Utterbackiana_hartfieldorum **
| | +--+
| | +-- Utterbackiana_suborbiculata **
| |
| | +-- Utterbackia_peninsularis **
| | +-----------------------+
+--+ | +-- Pyganodon_grandis **
| |
| | +-------------------- Alasminota_etowaensis **
| | |
| | | +----- Pressodonta_viridis **
| | +--+ +-----------+
| | | | | | +-- Platynaias_subviridis **
| | | | | +--+
+--+ | | | +-- Prolasmidonta_heterodon **
| | +--+
| | | +-- Anodontoides_ferussacianus **
| | | +-----------+
| | | | +-- Anodontoides_denigratus **
| | +--+
| | | +----------- Strophitus_undulatus **
| | +--+
| | | +-------- Strophitus_pascagoulaensis[2] **
| | +--+
+--+ | +----- Strophitus_pascagoulaensis[1] **
| +--+
| | +-- Strophitus_radiatus **
| +--+
| +-- Strophitus_williamsi **
|
| +-- Arcidens_confragosus **
| +--+
| | +-- Arcidens_wheeleri **
| +--------+
| | | +-- Pseudodontoideus_connasaugaensis **
| | +--+
| | +-- Pseudodontoideus_subvexus **
+--------+
| +-- Alasmidonta_marginata **
| +--------+
| | +-- Alasmidonta_raveneliana **
| |
+--+ +-- Alasmidonta_varicosa **
| +-----+
| | +-- Alasmidonta_uwharriensis **
+--+
| +----- Alasmidonta_undulata **
+--+
| +-- Alasmidonta_triangulata **
+--+
+-- Alasmidonta_arcula **
Whelan et al., 2023 [Publ 4287], fig. 4
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 29 |
29 |
25 |
2 |
0 |
0 |
0 |
55.0 |
+-- Anodonta_anatina **
+-----------------------+
| +-- Anodonta_cygnea **
|
| +----------------------- Pressodonta_viridis **
| |
| | +----- Utterbackiana_couperiana **
--+ | |
| | +-----------+ +-- Utterbackiana_heardi **
| | | | |
| | | +--+-- Utterbackiana_hartfieldorum **
| | | |
| | +--+ +-- Utterbackiana_suborbiculata **
| | | |
+--+ | | +-------------- Alasminota_etowaensis **
| | | |
| | +--+ +-- Arcidens_confragosus **
| | | +--------+
| | | | +-- Arcidens_wheeleri **
| | +--+
| | | +-------- Pseudodontoideus_connasaugaensis[3] **
| | +--+
| | | +----- Pseudodontoideus_connasaugaensis[2] **
| | +--+
+--+ | +-- Pseudodontoideus_subvexus **
| +--+
| +-- Pseudodontoideus_connasaugaensis[1] **
|
| +----- Alasmidonta_raveneliana **
| |
| | +-- Alasmidonta_uwharriensis **
| +-----------+--+
| | | +-- Alasmidonta_varicosa **
| | |
| | | +-- Alasmidonta_arcula **
| | +--+
+--+ +-- Alasmidonta_triangulata **
|
| +-------------- Platynaias_subviridis **
| |
| | +-------- Strophitus_pascagoulaensis **
| | +--+
+--+ | | +----- Strophitus_undulatus **
| | +--+
| | | +-- Strophitus_radiatus **
| | +--+
+--+ +-- Strophitus_williamsi **
|
| +-- Pyganodon_grandis **
| +-----+
| | +-- Pyganodon_cataracta **
+--+
| +----- Utterbackia_peninsularis **
+--+
| +-- Anodontoides_ferussacianus **
+--+
+-- Anodontoides_denigratus **
Bogan et al., 2023 [Publ 4295], fig. 3
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 28 |
28 |
25 |
2 |
3 |
1 |
1 |
49.0 |
+----------------------------------- Schistodesmus_lampreyanus
|
| +-------------------------------- Anodonta_cygnea **
--+ |
| | +-------- Cristaria_bellua **
| | +--------------------+
| | | | +----- Cristaria_clessini **
+--+ | +--+
| | | +-- Cristaria_plicata **
| | +--+
| | +-- Cristaria_truncata **
| |
| | +-------------- Amuranodonta_kijaensis **
+--+ |
| +-----------+ +----------- Anemina_arcaeformis **
| | | |
| | +--+ +-------- Buldowskia_suifunica **
| | | |
| | +--+ +-- Buldowskia_flavotincta **
| | | +--+
| | | | +-- Buldowskia_shadini **
| | +--+
+--+ | +-- Buldowskia_iwakawai **
| +--+
| +-- Buldowskia_kamiyai **
|
| +-- Pletholophus_honglinhensis **
| +--------------------+
| | +-- Pletholophus_tenuis **
| |
| | +-------- Beringiana_japonica **
+--+ +-----------+
| | | +----- Beringiana_beringiana **
| | +--+
| | | +-- Beringiana_fukuharai **
| | +--+
+--+ +-- Beringiana_gosannensis **
|
| +-- Sinanodonta_angula **
| +--------------+
| | +-- Sinanodonta_lucida **
| |
+--+ +-------------- Sinanodonta_schrenkii **
| |
| | +-- Sinanodonta_calipygos **
+--+-----------+
| +-- Sinanodonta_tumens **
|
| +----------- Sinanodonta_woodiana[2] **
+--+
| +-------- Sinanodonta_jourdyi **
+--+
| +----- Sinanodonta_elliptica **
+--+
| +-- Sinanodonta_lauta **
+--+
+-- Sinanodonta_woodiana[1] **
Inoue et al., 2018 [Publ 3858], fig. 1, 3a-c
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 21 |
21 |
0 |
3 |
184 |
0 |
0 |
46.0 |
+-- Alasmidonta_marginata **
|
+-- Prolasmidonta_heterodon **
|
+-- Alasmidonta_undulata **
|
+-- Beringiana_beringiana **
|
+-- Anodonta_kennerlyi **
|
+-- Anodonta_nuttalliana **
|
+-- Anodonta_oregonensis **
|
+-- Arcidens_confragosus **
|
+-- Arcidens_wheeleri **
|
+-- Lasmigona_complanata **
|
--+-- Platynaias_compressa **
|
+-- Lasmigona_costata **
|
+-- Platynaias_decorata **
|
+-- Platynaias_subviridis **
|
+-- Pyganodon_grandis **
|
+-- Pyganodon_lacustris **
|
+-- Pseudodontoideus_subvexus **
|
+-- Strophitus_undulatus **
|
+-- Utterbackia_imbecillis **
|
+-- Utterbackia_peninsularis[1] **
|
+-- Utterbackia_peninsularis[2] **
Chong et al., 2008 [Publ 2668], fig. 3
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 20 |
21 |
10 |
3 |
1 |
1 |
1 |
41.0 |
+----------------- Anodonta_cygnea **
|
+----------------- Pyganodon_fragilis **
|
+----------------- Alasmidonta_holstonia **
|
+----------------- Anodonta_anatina **
|
+----------------- Unio_tumidus
|
| +-- Anodonta_nuttalliana **
+--------------+
| +-- Anodonta_californiensis **
|
--+ +-- Beringiana_beringiana **
+--------------+
| +-- Sinanodonta_woodiana **
|
| +-- Pseudodontoideus_subvexus **
| |
+--------------+-- Alasmidonta_marginata **
| |
| +-- Lasmigona_complanata **
|
| +-------------- Pseudanodonta_complanata **
| |
| | +----- Strophitus_undulatus **
| +--------+
+--+ | +-- Prolasmidonta_heterodon **
| +--+
| +-- Platynaias_compressa **
|
| +----------- Anodonta_oregonensis[4] **
+--+
| +-------- Anodonta_oregonensis[3] **
+--+
| +----- Anodonta_oregonensis[2] **
+--+
| +-- Anodonta_oregonensis[1] **
+--+
+-- Anodonta_kennerlyi **
Huang et al., 2019 [Publ 3843], fig. 2
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 20 |
20 |
13 |
3 |
184 |
25 |
4 |
40.0 |
+----------------------------- Lepidodesma_languilati
|
| +-------- Aculamprotula_tortuosa
| |
| +-----------+ +----- Aculamprotula_coreana
| | | |
| | +--+----- Aculamprotula_scripta
| | |
| | | +-- Aculamprotula_fibrosa
| | +--+
| | +-- Aculamprotula_tientsinensis
+--------+
| | +-------- Unio_tumidus
| | +--------+
| | | | +----- Unio_crassus
| | | +--+
| | | | +-- Unio_delphinus
| | | +--+
| +--+ +-- Unio_pictorum
| |
| | +-- Schistodesmus_lampreyanus
| | +-----------+
| | | +-- Schistodesmus_spinosus
| | |
| +--+ +----- Nodularia_douglasiae[1]
--+ | +-----+
| | | | +-- Nodularia_douglasiae[2]
| | | +--+
| +--+ +-- Nodularia_nuxpersicae
| |
| | +-------- Tchangsinaia_piscicula
| +--+
| | +----- Cuneopsis_rufescens
| +--+
| | +-- Cuneopsis_heudei
| +--+
| +-- Cuneopsis_celtiformis
|
| +-------- Lanceolaria_gladiola
| |
| +-----------------+ +----- Lanceolaria_grayii[1]
| | | |
| | +--+----- Lanceolaria_grayii[2]
| | |
| | | +-- Lanceolaria_lanceolata
| | +--+
| | +-- Lanceolaria_triformis
| |
| | +-------------------- Anemina_arcaeformis **
| | |
| | +--+ +----------------- Cristaria_plicata **
+--+ | | |
| | +--+ +-------------- Pletholophus_tenuis **
| | | |
| | +--+ +-- Sinanodonta_angula **
| | | +--------+
| | | | +-- Sinanodonta_lucida **
| | +--+
| | | +-------- Sinanodonta_woodiana[2] **
| | +--+
| | | +----- Sinanodonta_jourdyi **
| | +--+
+--+ | +-- Sinanodonta_elliptica **
| +--+
| +-- Sinanodonta_woodiana[1] **
|
| +-------- Alasmidonta_marginata **
| |
| +-------- Lasmigona_costata **
| |
| +-------- Pseudanodonta_complanata **
| |
| +-------- Simpsonaias_ambigua **
| |
+--------------+-------- Strophitus_undulatus **
|
| +----- Anodonta_anatina **
+--+
| | +-- Anodonta_cygnea **
| +--+
| +-- Anodonta_nuttalliana **
|
| +----- Pyganodon_grandis **
+--+
| +-- Utterbackia_imbecillis **
+--+
+-- Utterbackia_peninsularis **
Huang et al., 2019 [Publ 3843], fig. S3
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 20 |
20 |
17 |
3 |
181 |
23 |
4 |
39.0 |
+----------------------------- Lepidodesma_languilati
|
| +-------- Lanceolaria_gladiola
| +-----------------+
| | | +----- Lanceolaria_grayii
| | +--+
--+ | | +-- Lanceolaria_lanceolata
| | +--+
| | +-- Lanceolaria_triformis
| |
| | +-------------------- Anemina_arcaeformis **
| | |
| | +--+ +----------------- Cristaria_plicata **
+--+ | | |
| | +--+ +-------------- Pletholophus_tenuis **
| | | |
| | +--+ +-- Sinanodonta_angula **
| | | +--------+
| | | | +-- Sinanodonta_lucida **
| | +--+
| | | +-------- Sinanodonta_woodiana[2] **
| | +--+
| | | +----- Sinanodonta_jourdyi **
+--+ +--+
| | +-- Sinanodonta_elliptica **
| +--+
| +-- Sinanodonta_woodiana[1] **
|
| +-- Alasmidonta_marginata **
| +--------+
| | +-- Lasmigona_costata **
| |
| | +----- Anodonta_anatina **
| +-----+
| | | +-- Anodonta_cygnea **
+-----------+ +--+
| +-- Anodonta_nuttalliana **
|
| +----- Pseudanodonta_complanata **
| +--+
| | | +-- Simpsonaias_ambigua **
| | +--+
+--+ +-- Strophitus_undulatus **
|
| +----- Pyganodon_grandis **
+--+
| +-- Utterbackia_imbecillis **
+--+
+-- Utterbackia_peninsularis **
Huang et al., 2019 [Publ 3843], fig. S4
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 20 |
44 |
21 |
3 |
182 |
24 |
4 |
46.0 |
+----------------- Acuticosta_chinensis
|
+----------------- Alasmidonta_marginata **
|
+----------------- Anodonta_anatina **
|
+----------------- Anemina_arcaeformis **
|
+----------------- Anodonta_cygnea **
|
+----------------- Cristaria_plicata **
|
+----------------- Anodonta_nuttalliana **
|
+----------------- Lasmigona_costata **
|
+----------------- Lepidodesma_languilati
|
+----------------- Pseudanodonta_complanata **
|
+----------------- Pyganodon_grandis **
|
+----------------- Simpsonaias_ambigua **
|
+----------------- Strophitus_undulatus **
|
| +-- Schistodesmus_lampreyanus
+--------------+
| +-- Schistodesmus_spinosus
|
| +-- Utterbackia_imbecillis **
+--------------+
| +-- Utterbackia_peninsularis **
|
| +-------- Lanceolaria_gladiola
+--------+
| | +----- Lanceolaria_grayii
| +--+
--+ | +-- Lanceolaria_lanceolata
| +--+
| +-- Lanceolaria_triformis
|
| +-------- Unio_tumidus
+--------+
| | +----- Unio_crassus
| +--+
| | +-- Unio_delphinus
| +--+
| +-- Unio_pictorum
|
| +-------- Aculamprotula_tortuosa
| |
+--------+ +-- Aculamprotula_coreana
| | +--+
| | | +-- Aculamprotula_scripta
| +--+
| | +-- Aculamprotula_fibrosa
| +--+
| +-- Aculamprotula_tientsinensis
|
| +-------- Cuneopsis_heudei
| |
| +-------- Cuneopsis_celtiformis
| |
+--------+-------- Tchangsinaia_piscicula
| |
| +-------- Cuneopsis_rufescens
| |
| | +----- Nodularia_douglasiae[2]
| +--+
| | +-- Nodularia_douglasiae[1]
| +--+
| +-- Nodularia_nuxpersicae
|
| +-------------- Pletholophus_tenuis **
| |
+--+ +-- Sinanodonta_angula **
| +--------+
| | +-- Sinanodonta_lucida **
+--+
| +-------- Sinanodonta_woodiana[1] **
+--+
| +----- Sinanodonta_jourdyi **
+--+
| +-- Sinanodonta_elliptica **
+--+
+-- Sinanodonta_woodiana[2] **
Sano et al., 2020 [Publ 4138], fig. 2
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 20 |
39 |
37 |
3 |
53 |
19 |
3 |
49.0 |
+----- Buldowskia_iwakawai **
+-----------------------------+
| | +-- Anemina_arcaeformis **
| +--+
| +-- Anemina_euscaphys **
|
| +-------------- Pletholophus_tenuis **
| |
| | +----- Beringiana_fukuharai **
| +-----------------+ +-----+
--+ | | | | +-- Beringiana_japonica **
| | | | +--+
| | +--+ +-- Beringiana_beringiana **
| | |
| | | +-------- Sinanodonta_lucida **
| | +--+
| | | +----- Sinanodonta_tumens **
| | +--+
| | | +-- Sinanodonta_lauta **
+--+ +--+
| +-- Sinanodonta_woodiana **
|
| +----------------------------- Cristaria_plicata **
| |
| | +-- Anodonta_cygnea **
| | +--------+
| | | +-- Anodonta_anatina **
| | |
| | +--------------+ +----- Alasmidonta_varicosa **
+--+ | | +--+
| | | | | +-- Prolasmidonta_heterodon **
| | | | +--+
| | +--+ +-- Platynaias_compressa **
| | |
| | | +----- Pyganodon_grandis **
| | +--+
+--+ | +-- Utterbackia_imbecillis **
| +--+
| +-- Utterbackia_peninsularis **
|
| +----- Lanceolaria_grayii[1]
| +-----------------+
| | | +-- Lanceolaria_lanceolata
| | +--+
| | +-- Lanceolaria_grayii[2]
+--+
| +-------------------- Lepidodesma_languilati
| |
| | +----- Aculamprotula_tortuosa
| | +-----------+
+--+ | | +-- Aculamprotula_tientsinensis
| | +--+
| | +-- Aculamprotula_coreana
| |
+--+ +----- Unio_crassus
| +--------+
| | | +-- Unio_delphinus
| | +--+
| | +-- Unio_pictorum
| |
+--+ +-------- Inversiunio_jokohamensis
| +--+
| | | +----- Inversiunio_yanagawensis[2]
| | +--+
| | | +-- Inversiunio_reinianus
| | +--+
+--+ +-- Inversiunio_yanagawensis[1]
|
| +-- Tchangsinaia_piscicula
| +-----+
| | +-- Schistodesmus_lampreyanus
+--+
| +----- Nodularia_douglasiae[2]
+--+
| +-- Nodularia_nipponensis
+--+
+-- Nodularia_douglasiae[1]
Sano et al., 2020 [Publ 4138], fig. 3
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 20 |
38 |
36 |
3 |
52 |
18 |
3 |
50.0 |
+----- Buldowskia_iwakawai **
+-----------------------------+
| | +-- Anemina_arcaeformis **
| +--+
| +-- Anemina_euscaphys **
|
| +-------- Pletholophus_tenuis **
| +--+
| | | +----- Beringiana_fukuharai **
| | +--+
--+ | | +-- Beringiana_japonica **
| +--------------------+ +--+
| | | +-- Beringiana_beringiana **
| | |
| | | +-------- Sinanodonta_lucida **
| | +--+
| | | +----- Sinanodonta_tumens **
| | +--+
| | | +-- Sinanodonta_woodiana **
| | +--+
+--+ +-- Sinanodonta_lauta **
|
| +----------------------------- Cristaria_plicata **
| |
| | +-- Anodonta_anatina **
| | +--------+
| | | +-- Anodonta_cygnea **
| | |
| | +--------------+ +----- Pyganodon_grandis **
+--+ | | +--+
| | | | | +-- Utterbackia_imbecillis **
| | | | +--+
| | +--+ +-- Utterbackia_peninsularis **
| | |
| | | +----- Alasmidonta_varicosa **
| | +--+
+--+ | +-- Platynaias_compressa **
| +--+
| +-- Prolasmidonta_heterodon **
|
| +----- Lanceolaria_grayii[1]
| +-----------------+
| | | +-- Lanceolaria_lanceolata
| | +--+
| | +-- Lanceolaria_grayii[2]
+--+
| +-------------------- Lepidodesma_languilati
| |
| | +----- Aculamprotula_tortuosa
| | +-----------+
+--+ | | +-- Aculamprotula_tientsinensis
| | +--+
| | +-- Aculamprotula_coreana
| |
+--+ +----- Unio_crassus
| +--------+
| | | +-- Unio_delphinus
| | +--+
| | +-- Unio_pictorum
| |
+--+ +-------- Inversiunio_jokohamensis
| +--+
| | | +----- Inversiunio_yanagawensis[2]
| | +--+
| | | +-- Inversiunio_reinianus
| | +--+
+--+ +-- Inversiunio_yanagawensis[1]
|
| +-- Nodularia_nipponensis
| +--+
| | +-- Nodularia_douglasiae
+-----+
| +-- Schistodesmus_lampreyanus
+--+
+-- Tchangsinaia_piscicula
Sano et al., 2020 [Publ 4138], fig. 4
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 20 |
38 |
36 |
3 |
52 |
18 |
3 |
50.0 |
+----- Buldowskia_iwakawai **
+--------------------------------+
| | +-- Anemina_arcaeformis **
| +--+
| +-- Anemina_euscaphys **
|
| +-------- Sinanodonta_lucida **
| +--+
| | | +----- Sinanodonta_tumens **
| | +--+
--+ | | +-- Sinanodonta_woodiana **
| +-----------------------+ +--+
| | | +-- Sinanodonta_lauta **
| | |
| | | +-------- Pletholophus_tenuis **
| | +--+
| | | +----- Beringiana_fukuharai **
| | +--+
| | | +-- Beringiana_japonica **
| | +--+
+--+ +-- Beringiana_beringiana **
|
| +-------------------------------- Cristaria_plicata **
| |
| | +-- Anodonta_cygnea **
| | +--------+
| | | +-- Anodonta_anatina **
| | |
| | +-----------------+ +----- Alasmidonta_varicosa **
+--+ | | +--+
| | | | | +-- Prolasmidonta_heterodon **
| | | | +--+
| | +--+ +-- Platynaias_compressa **
| | |
| | | +----- Pyganodon_grandis **
| | +--+
+--+ | +-- Utterbackia_imbecillis **
| +--+
| +-- Utterbackia_peninsularis **
|
| +-------------------------- Lepidodesma_languilati
| |
| | +----- Lanceolaria_grayii[1]
| | +-----------------+
+--+ | | +-- Lanceolaria_lanceolata
| | +--+
| | +-- Lanceolaria_grayii[2]
| |
+--+ +----- Aculamprotula_tortuosa
| +--------------+
| | | +-- Aculamprotula_tientsinensis
| | +--+
| | +-- Aculamprotula_coreana
| |
+--+ +----- Unio_crassus
| +-----------+
| | | +-- Unio_delphinus
| | +--+
| | +-- Unio_pictorum
+--+
| +-- Nodularia_douglasiae
| +-----------+
| | +-- Nodularia_nipponensis
| |
+--+ +-- Schistodesmus_lampreyanus
| +--------+
| | +-- Tchangsinaia_piscicula
+--+
| +-------- Inversiunio_jokohamensis
+--+
| +----- Inversiunio_yanagawensis[2]
+--+
| +-- Inversiunio_reinianus
+--+
+-- Inversiunio_yanagawensis[1]
Smith et al., 2018 [Publ 3790], fig. 1
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 19 |
19 |
17 |
2 |
0 |
0 |
0 |
46.0 |
+-------------------- Cristaria_plicata **
|
| +-- Utterbackia_peninsularis **
| +--------------+
| | +-- Pyganodon_grandis **
--+ |
| | +-- Anodontoides_ferussacianus **
| | +--------+
| | | +-- Anodontoides_denigratus **
| | |
+--+ +--+ +----- Strophitus_pascagoulaensis **
| | | +--+
| | | | | +-- Strophitus_radiatus **
| | | | +--+
| | +--+ +-- Strophitus_williamsi **
| | |
| | | +----- Strophitus_howellsi[2] **
| | +--+
+--+ | +-- Strophitus_undulatus **
| +--+
| +-- Strophitus_howellsi[1] **
|
| +-- Alasmidonta_triangulata **
| +--------+
| | +-- Alasminota_etowaensis **
| |
+--+ +-- Pseudodontoideus_connasaugaensis **
| +-----+
| | +-- Pseudodontoideus_subvexus **
| |
+--+ +-- Utterbackiana_couperiana **
| +--+
| | +-- Utterbackiana_heardi **
+--+
| +-- Utterbackiana_hartfieldorum **
+--+
+-- Utterbackiana_suborbiculata **
Wu et al., 2023 [Publ 4339], fig. 4, S5-aa
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 19 |
19 |
17 |
3 |
77 |
28 |
4 |
42.0 |
+-------- Aculamprotula_tortuosa
|
+--------------+ +-- Aculamprotula_coreana
| | +--+
| | | +-- Aculamprotula_tientsinensis
| +--+
| | +-- Aculamprotula_polysticta
| +--+
| +-- Aculamprotula_scripta
|
+--+ +-------------- Unio_tumidus
| | +-----+
| | | | +----------- Unio_crassus
| | | +--+
| | | | +-------- Unio_elongatulus
| | | +--+
| | | | +----- Unio_pictorum
| | | +--+
| | | | +-- Unio_delphinus
| +--+ +--+
| | +-- Unio_mancus
| |
| | +-- Nodularia_breviconcha
| | +--+
| | | +-- Nodularia_nipponensis
| | +-----------+
| | | | +-- Pseudocuneopsis_capitata
| | | +--+
| +--+ +-- Pseudocuneopsis_sichuanensis
| |
| | +-------------- Tchangsinaia_piscicula
| | |
--+ +--+ +-- Schistodesmus_lampreyanus
| | +--------+
| | | +-- Schistodesmus_spinosus
| +--+
| | +-------- Cuneopsis_rufescens
| +--+
| | +----- Cuneopsis_celtiformis
| +--+
| | +-- Cuneopsis_demangei
| +--+
| +-- Cuneopsis_heudei
|
| +-------- Acuticosta_chinensis
| +--------+
| | | +----- Lanceolaria_gladiola
| | +--+
| | | +-- Lanceolaria_grayii
| | +--+
| | +-- Lanceolaria_lanceolata
| |
| | +----- Cristaria_plicata **
| | +-----+
| | | | +-- Anemina_arcaeformis **
+--------+ | +--+
| +--+ +-- Anemina_euscaphys **
| | |
| | | +-------- Beringiana_fukuharai **
| | +--+
| | | +----- Sinanodonta_lucida **
| | +--+
| | | +-- Sinanodonta_tumens **
| | +--+
| | +-- Sinanodonta_woodiana **
+--+
| +-- Anodonta_nuttalliana **
| +-----+
| | +-- Pseudanodonta_complanata **
| +--+
| | | +----- Anodonta_anatina **
| | +--+
| | | +-- Anodonta_cygnea **
| | +--+
| | +-- Anodonta_exulcerata **
+--+
| +----- Pyganodon_grandis **
| +--+
| | | +-- Utterbackia_peninsularis **
| | +--+
| | +-- Utterbackia_imbecillis **
+--+
| +-- Alasmidonta_varicosa **
| +--+
| | +-- Platynaias_compressa **
+--+
| +-- Prolasmidonta_heterodon **
+--+
+-- Lasmigona_complanata **
Wu et al., 2023 [Publ 4339], fig. 4, S5-nt
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 19 |
19 |
15 |
3 |
78 |
29 |
4 |
44.0 |
+-------- Acuticosta_chinensis
+--------------+
| | +----- Lanceolaria_gladiola
| +--+
| | +-- Lanceolaria_grayii
| +--+
| +-- Lanceolaria_lanceolata
|
| +-------------- Cristaria_plicata **
+-----+ |
| | +-----+ +-- Anemina_arcaeformis **
| | | | +--------+
| | | | | +-- Anemina_euscaphys **
| | | +--+
| | | | +-------- Beringiana_fukuharai **
| | | +--+
| | | | +----- Sinanodonta_lucida **
| | | +--+
| +--+ | +-- Sinanodonta_tumens **
| | +--+
| | +-- Sinanodonta_woodiana **
| |
| | +----------------- Anodonta_nuttalliana **
| | |
| | +----------------- Pseudanodonta_complanata **
| | |
| +--+ +-------------- Anodonta_cygnea **
| | |
| | | +-- Anodonta_exulcerata **
| | +-----------+
| | | +-- Anodonta_anatina **
| +--+
--+ | +----- Pyganodon_grandis **
| | +-----+
| | | | +-- Utterbackia_imbecillis **
| | | +--+
| +--+ +-- Utterbackia_peninsularis **
| |
| | +-------- Lasmigona_complanata **
| +--+
| | +----- Prolasmidonta_heterodon **
| +--+
| | +-- Alasmidonta_varicosa **
| +--+
| +-- Platynaias_compressa **
|
| +-------- Aculamprotula_tortuosa
| |
| +-----------------+ +-- Aculamprotula_coreana
| | | +--+
| | | | +-- Aculamprotula_tientsinensis
| | +--+
| | | +-- Aculamprotula_polysticta
| | +--+
| | +-- Aculamprotula_scripta
| |
+--+ +----------- Unio_tumidus
| |
| +-----------+ +-------- Unio_crassus
| | | |
| | +--+ +----- Unio_elongatulus
| | | |
| | +--+ +-- Unio_delphinus
| | | |
| | +--+-- Unio_mancus
+--+ |
| +-- Unio_pictorum
|
| +-- Pseudocuneopsis_capitata
| +-----------------+
| | +-- Pseudocuneopsis_sichuanensis
| |
+--+ +-- Nodularia_breviconcha
| +--------------+
| | +-- Nodularia_nipponensis
| |
+--+ +----- Tchangsinaia_piscicula
| +--------+
| | | +-- Schistodesmus_lampreyanus[2]
| | +--+
+--+ +-- Schistodesmus_spinosus
|
| +----------- Schistodesmus_lampreyanus[1]
+--+
| +-------- Cuneopsis_rufescens
+--+
| +----- Cuneopsis_celtiformis
+--+
| +-- Cuneopsis_demangei
+--+
+-- Cuneopsis_heudei
Wu et al., 2024 [Publ 4350], fig. 8a, S1
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 19 |
19 |
17 |
3 |
73 |
28 |
4 |
42.0 |
+-------- Aculamprotula_tortuosa
|
+--------------+ +-- Aculamprotula_coreana
| | +--+
| | | +-- Aculamprotula_tientsinensis
| +--+
| | +-- Aculamprotula_polysticta
| +--+
| +-- Aculamprotula_scripta
|
+--+ +-------------- Unio_tumidus
| | +-----+
| | | | +----------- Unio_crassus
| | | +--+
| | | | +-------- Unio_elongatulus
| | | +--+
| | | | +----- Unio_pictorum
| | | +--+
| | | | +-- Unio_delphinus
| +--+ +--+
| | +-- Unio_mancus
| |
| | +-- Nodularia_breviconcha
| | +--------------+
| | | +-- Nodularia_douglasiae
| | |
| | | +-- Pseudocuneopsis_capitata
| | +--------------+
| +--+ +-- Pseudocuneopsis_sichuanensis
| |
| | +-- Schistodesmus_lampreyanus
| | +-----------+
--+ | | +-- Schistodesmus_spinosus
| +--+
| | +----------- Tchangsinaia_piscicula
| +--+
| | +-------- Cuneopsis_rufescens
| +--+
| | +----- Cuneopsis_celtiformis
| +--+
| | +-- Cuneopsis_demangei
| +--+
| +-- Cuneopsis_heudei
|
| +-------- Acuticosta_chinensis
| +--------------+
| | | +----- Lanceolaria_gladiola
| | +--+
| | | +-- Lanceolaria_grayii
| | +--+
| | +-- Lanceolaria_lanceolata
| |
| | +-------------- Cristaria_plicata **
| | |
+--+ +-----+ +-- Anemina_arcaeformis **
| | | +--------+
| | | | +-- Anemina_euscaphys **
| | +--+
| | | +-------- Beringiana_fukuharai **
| | +--+
| | | +----- Sinanodonta_lucida **
| | +--+
| | | +-- Sinanodonta_tumens **
+--+ +--+
| +-- Sinanodonta_woodiana **
|
| +-------- Pseudanodonta_complanata **
| +--------+
| | | +----- Anodonta_anatina **
| | +--+
| | | +-- Anodonta_cygnea **
| | +--+
+--+ +-- Anodonta_exulcerata **
|
| +-------------- Anodonta_nuttalliana **
| |
| | +----- Pyganodon_grandis **
+--+ +-----+
| | | +-- Utterbackia_imbecillis **
| | +--+
+--+ +-- Utterbackia_peninsularis **
|
| +-------- Lasmigona_complanata **
+--+
| +----- Prolasmidonta_heterodon **
+--+
| +-- Alasmidonta_varicosa **
+--+
+-- Platynaias_compressa **
Wu et al., 2024 [Publ 4350], fig. 8b, S2
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 19 |
19 |
17 |
3 |
73 |
28 |
4 |
42.0 |
+-------- Aculamprotula_tortuosa
|
+--------------+ +-- Aculamprotula_coreana
| | +--+
| | | +-- Aculamprotula_tientsinensis
| +--+
| | +-- Aculamprotula_polysticta
| +--+
| +-- Aculamprotula_scripta
|
+--+ +-------------- Unio_tumidus
| | +-----+
| | | | +----------- Unio_crassus
| | | +--+
| | | | +-------- Unio_elongatulus
| | | +--+
| | | | +----- Unio_pictorum
| | | +--+
| | | | +-- Unio_delphinus
| +--+ +--+
| | +-- Unio_mancus
| |
| | +-- Nodularia_breviconcha
| | +--------------+
| | | +-- Nodularia_douglasiae
| | |
| | | +-- Pseudocuneopsis_capitata
| | +--------------+
| +--+ +-- Pseudocuneopsis_sichuanensis
| |
| | +-- Schistodesmus_lampreyanus
--+ | +-----------+
| | | +-- Schistodesmus_spinosus
| +--+
| | +----------- Tchangsinaia_piscicula
| +--+
| | +-------- Cuneopsis_rufescens
| +--+
| | +----- Cuneopsis_celtiformis
| +--+
| | +-- Cuneopsis_demangei
| +--+
| +-- Cuneopsis_heudei
|
| +-------- Acuticosta_chinensis
| +--------------+
| | | +----- Lanceolaria_gladiola
| | +--+
| | | +-- Lanceolaria_grayii
| | +--+
| | +-- Lanceolaria_lanceolata
| |
| | +-------------- Cristaria_plicata **
+--+ |
| +-----+ +-- Anemina_arcaeformis **
| | | +--------+
| | | | +-- Anemina_euscaphys **
| | +--+
| | | +-------- Beringiana_fukuharai **
| | +--+
| | | +----- Sinanodonta_lucida **
| | +--+
+--+ | +-- Sinanodonta_tumens **
| +--+
| +-- Sinanodonta_woodiana **
|
| +----------------- Anodonta_nuttalliana **
| |
| | +-------- Pseudanodonta_complanata **
| | +-----+
+--+ | | +----- Anodonta_anatina **
| | +--+
| | | +-- Anodonta_cygnea **
| | +--+
+--+ +-- Anodonta_exulcerata **
|
| +----- Pyganodon_grandis **
| +-----+
| | | +-- Utterbackia_imbecillis **
| | +--+
+--+ +-- Utterbackia_peninsularis **
|
| +-------- Lasmigona_complanata **
+--+
| +----- Prolasmidonta_heterodon **
+--+
| +-- Alasmidonta_varicosa **
+--+
+-- Platynaias_compressa **
Sano et al., 2022 [Publ 4311], fig. 3
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 18 |
18 |
16 |
1 |
0 |
0 |
0 |
42.0 |
+-------- Pletholophus_tenuis **
+--------------+
| | +----- Cristaria_plicata **
| +--+
| | +-- Buldowskia_kamiyai **
| +--+
| +-- Buldowskia_iwakawai **
--+
| +-------- Beringiana_fukuharai **
| +-----------+
| | | +----- Beringiana_gosannensis **
| | +--+
| | | +-- Beringiana_japonica **
| | +--+
+--+ +-- Beringiana_beringiana **
|
| +-- Sinanodonta_angula **
| +--------------+
| | +-- Sinanodonta_lucida **
| |
+--+ +----- Sinanodonta_schrenkii **
| +--------+
| | | +-- Sinanodonta_tumens **
| | +--+
+--+ +-- Sinanodonta_calipygos **
|
| +----------- Sinanodonta_woodiana[2] **
+--+
| +-------- Sinanodonta_jourdyi **
+--+
| +----- Sinanodonta_elliptica **
+--+
| +-- Sinanodonta_woodiana[1] **
+--+
+-- Sinanodonta_lauta **
Zanatta et al., 2007 [Publ 2698], fig. 1A
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 16 |
16 |
10 |
2 |
0 |
0 |
0 |
44.0 |
+-- Platynaias_compressa **
|
+-----------------+-- Lasmigona_costata **
| |
| +-- Lasmigona_complanata **
|
--+ +----------------- Anodonta_cygnea **
| |
| | +----------- Anodonta_kennerlyi **
| | +--+
+--+ | | +-------- Utterbackiana_implicata **
| | +--+
| | | +----- Utterbackiana_suborbiculata **
| | +--+
| | | +-- Utterbackiana_heardi **
+--+ +--+
| +-- Utterbackiana_couperiana **
|
| +-- Utterbackia_peggyae **
| +--+
| | +-- Utterbackia_imbecillis **
| |
+--------+ +-- Pyganodon_gibbosa **
| |
| +-- Pyganodon_cataracta **
| |
+--+-- Pyganodon_fragilis **
|
+-- Pyganodon_grandis **
|
+-- Pyganodon_lacustris **
Zanatta et al., 2007 [Publ 2698], fig. 1B
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 16 |
16 |
13 |
2 |
0 |
0 |
0 |
44.0 |
+-- Lasmigona_costata **
|
+-----------------+-- Platynaias_compressa **
| |
| +-- Lasmigona_complanata **
|
| +-------------- Anodonta_cygnea **
| +--+
--+ | | +----------- Anodonta_kennerlyi **
| | +--+
| | | +-------- Utterbackiana_implicata **
| | +--+
| | | +----- Utterbackiana_heardi **
| | +--+
+--+ | +-- Utterbackiana_suborbiculata **
| +--+
| +-- Utterbackiana_couperiana **
|
| +-- Utterbackia_peggyae **
| +-----------+
| | +-- Utterbackia_imbecillis **
+--+
| +----------- Pyganodon_lacustris **
+--+
| +-------- Pyganodon_grandis **
+--+
| +----- Pyganodon_fragilis **
+--+
| +-- Pyganodon_gibbosa **
+--+
+-- Pyganodon_cataracta **
Hoeh, 1990 [Publ 2717], fig. 2
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 16 |
16 |
13 |
2 |
0 |
0 |
0 |
43.0 |
+-- Lasmigona_complanata **
|
+-----------------+-- Platynaias_compressa **
| |
| +-- Lasmigona_costata **
|
| +-------------- Anodonta_cygnea **
| +--+
--+ | | +----------- Anodonta_kennerlyi **
| | +--+
| | | +-------- Utterbackiana_implicata **
| | +--+
| | | +----- Utterbackiana_couperiana[2] **
| | +--+
+--+ | +-- Utterbackiana_suborbiculata **
| +--+
| +-- Utterbackiana_couperiana[1] **
|
| +-- Utterbackia_imbecillis **
| +-----------+
| | +-- Utterbackia_peggyae **
+--+
| +----------- Pyganodon_lacustris **
+--+
| +-------- Pyganodon_grandis **
+--+
| +----- Pyganodon_fragilis **
+--+
| +-- Pyganodon_cataracta **
+--+
+-- Pyganodon_gibbosa **
Hoeh, 1990 [Publ 2717], fig. 3
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 16 |
16 |
2 |
2 |
0 |
0 |
0 |
43.0 |
+----- Pyganodon_fragilis **
|
+----- Utterbackia_imbecillis **
|
+----- Utterbackia_peggyae **
|
+----- Anodonta_cygnea **
|
+----- Anodonta_kennerlyi **
|
+----- Utterbackiana_couperiana[1] **
|
--+----- Utterbackiana_couperiana[2] **
|
+----- Utterbackiana_suborbiculata **
|
| +-- Lasmigona_costata **
+--+
| +-- Utterbackiana_implicata **
|
| +-- Lasmigona_complanata **
| |
| +-- Platynaias_compressa **
| |
| +-- Pyganodon_cataracta **
+--+
+-- Pyganodon_gibbosa **
|
+-- Pyganodon_grandis **
|
+-- Pyganodon_lacustris **
Hoeh, 1990 [Publ 2717], fig. 4
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 16 |
16 |
13 |
2 |
0 |
0 |
0 |
43.0 |
+-- Lasmigona_complanata **
|
+-----------------+-- Platynaias_compressa **
| |
| +-- Lasmigona_costata **
|
| +-------------- Anodonta_cygnea **
| +--+
--+ | | +----------- Anodonta_kennerlyi **
| | +--+
| | | +-------- Utterbackiana_implicata **
| | +--+
| | | +----- Utterbackiana_couperiana[2] **
| | +--+
+--+ | +-- Utterbackiana_suborbiculata **
| +--+
| +-- Utterbackiana_couperiana[1] **
|
| +-- Utterbackia_imbecillis **
| +--------+
| | +-- Utterbackia_peggyae **
+-----+
| +-------- Pyganodon_fragilis **
| |
+--+ +-- Pyganodon_cataracta **
| +--+
| | +-- Pyganodon_gibbosa **
+--+
| +-- Pyganodon_lacustris **
+--+
+-- Pyganodon_grandis **
Sano et al., 2020 [Publ 4138], fig. 1
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 16 |
34 |
5 |
1 |
32 |
18 |
4 |
43.0 |
+-------- Lanceolaria_grayii[1]
|
+-------- Lanceolaria_grayii[2]
|
+-------- Cristaria_plicata[1] **
|
+-------- Cristaria_plicata[2] **
|
+-------- Sinanodonta_lauta **
|
+-------- Beringiana_japonica **
|
+-------- Beringiana_fukuharai[1] **
|
+-------- Beringiana_beringiana **
|
+-------- Beringiana_fukuharai[2] **
|
+-------- Beringiana_fukuharai[3] **
|
+-------- Buldowskia_iwakawai[1] **
|
+-------- Buldowskia_iwakawai[2] **
|
+-------- Sinanodonta_tumens[1] **
|
+-------- Sinanodonta_tumens[2] **
|
--+-------- Pletholophus_tenuis **
|
+-------- Anemina_arcaeformis **
|
+-------- Lanceolaria_lanceolata
|
+-------- Cristaria_plicata[3] **
|
+-------- Lanceolaria_gladiola
|
+-------- Sinanodonta_angula **
|
+-------- Lepidodesma_languilati
|
| +-- Aculamprotula_tortuosa
+-----+
| +-- Acuticosta_chinensis
|
| +----- Cuneopsis_celtiformis
| |
+--+ +-- Nodularia_douglasiae[1]
| | |
| +--+-- Nodularia_douglasiae[2]
| |
| +-- Nodularia_nipponensis
|
| +----- Schistodesmus_lampreyanus
| |
| | +-- Inversiunio_reinianus[1]
+--+ |
| +-- Inversiunio_reinianus[2]
| |
| +-- Inversiunio_jokohamensis[1]
+--+
+-- Inversiunio_jokohamensis[2]
|
+-- Inversiunio_jokohamensis[3]
|
+-- Inversiunio_jokohamensis[4]
Konopleva et al., 2023 [Publ 4269], fig. S1, 4
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 16 |
16 |
14 |
2 |
40 |
17 |
4 |
38.0 |
+-------- Lepidodesma_languilati
+--------------+
| | +----- Lanceolaria_gladiola
| +--+
| | +-- Lanceolaria_grayii
| +--+
--+ +-- Lanceolaria_triformis
|
| +-------------------- Acuticosta_chinensis
| |
| | +----- Anodonta_anatina **
| | +-----------+
+--+ | | +-- Pseudanodonta_complanata **
| | +--+
| | +-- Anodonta_cygnea **
| |
| | +-------- Anemina_arcaeformis **
+--+ +-----+
| | | +----- Amuranodonta_kijaensis **
| | +--+
| | | +-- Buldowskia_suifunica **
| | +--+
| | +-- Buldowskia_shadini **
+--+
| +-------- Beringiana_beringiana **
| +--+
| | | +----- Sinanodonta_schrenkii **
| | +--+
| | | +-- Sinanodonta_woodiana **
+--+ +--+
| +-- Sinanodonta_lauta **
|
| +-------- Cristaria_radiata **
| |
+--+ +-- Cristaria_truncata **
| +--+
| | +-- Cristaria_plicata **
+--+
| +-- Cristaria_bellua **
+--+
+-- Cristaria_clessini **
Inoue et al., 2014 [Publ 3635], fig. 3A
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 15 |
15 |
13 |
1 |
6 |
0 |
0 |
41.0 |
+----------- Pseudodontoideus_subvexus **
+-----+
| | +-------- Lasmigona_costata **
| +--+
| | +----- Lasmigona_complanata **
| +--+
| | +-- Arcidens_confragosus **
--+ +--+
| +-- Arcidens_wheeleri **
|
| +-------------- Strophitus_undulatus **
| |
| | +-------- Pyganodon_grandis **
| | +--+
+--+ | | +----- Utterbackia_peninsularis[2] **
| | +--+
| | | +-- Utterbackia_imbecillis **
| | +--+
+--+ +-- Utterbackia_peninsularis[1] **
|
| +-- Platynaias_compressa **
| +-----+
| | +-- Platynaias_subviridis **
+--+
| +----- Alasmidonta_marginata **
+--+
| +-- Alasmidonta_undulata **
+--+
+-- Alasmidonta_varicosa **
Bolotov et al., 2017 [Publ 3761], fig. S1
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 15 |
15 |
13 |
3 |
126 |
12 |
2 |
36.0 |
+----- Unio_tumidus
+--------+
| | +-- Unio_pictorum
| +--+
| +-- Unio_crassus
+--------+
| | +----------- Aculamprotula_tortuosa
| | |
| | | +-- Nodularia_douglasiae
| +--+ +-----+
| | | +-- Nodularia_nuxpersicae
| | |
| +--+ +-- Cuneopsis_rufescens
| | +--+
| | | +-- Cuneopsis_heudei
| +--+
--+ | +-- Tchangsinaia_piscicula
| +--+
| +-- Schistodesmus_lampreyanus
|
| +-- Lanceolaria_grayii
| +-----------------+
| | +-- Lanceolaria_gladiola
| |
| | +-- Cristaria_plicata **
| | +--------+
| | | +-- Cristaria_radiata **
+--+ +-----+
| | | +-------- Pletholophus_tenuis **
| | +--+
| | | +----- Sinanodonta_lucida **
| | +--+
| | | +-- Sinanodonta_jourdyi **
+--+ +--+
| +-- Sinanodonta_lauta **
|
| +-------------- Anodonta_anatina **
| |
| | +----- Pseudanodonta_complanata **
+--+ +-----+
| | | +-- Anodonta_cygnea **
| | +--+
| | +-- Anodonta_nuttalliana **
+--+
| +-- Lasmigona_costata **
| +-----+
| | +-- Alasmidonta_marginata **
+--+
| +----- Simpsonaias_ambigua **
+--+
| +-- Pyganodon_grandis **
+--+
+-- Strophitus_undulatus **
Bolotov et al., 2017 [Publ 3761], fig. S2
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 15 |
17 |
10 |
3 |
126 |
12 |
2 |
35.0 |
+----- Unio_tumidus
+-----+
| | +-- Unio_crassus
| +--+
| +-- Unio_pictorum
+-----+
| | +-------- Aculamprotula_tortuosa
| | |
| | +-------- Schistodesmus_lampreyanus
| | |
| +--+ +-- Nodularia_douglasiae
| +-----+
| | +-- Nodularia_nuxpersicae
| |
| | +----- Cuneopsis_heudei
| +--+
| | +-- Tchangsinaia_piscicula
| +--+
--+ +-- Cuneopsis_rufescens
|
| +-- Lanceolaria_grayii
| +-----------+
| | +-- Lanceolaria_gladiola
| |
| | +-- Cristaria_plicata **
| | +--------+
| | | +-- Cristaria_radiata **
| +--+
| | | +-------- Pletholophus_tenuis **
| | +--+
| | | +----- Sinanodonta_lucida **
+--+ +--+
| | +-- Sinanodonta_lauta **
| +--+
| +-- Sinanodonta_jourdyi **
|
| +-------- Anodonta_anatina **
| |
| +-------- Pseudanodonta_complanata **
| |
| | +-- Anodonta_cygnea **
+-----+-----+
| +-- Anodonta_nuttalliana **
|
| +----- Pyganodon_grandis **
| |
| +----- Simpsonaias_ambigua **
+--+
+----- Strophitus_undulatus **
|
| +-- Alasmidonta_marginata **
+--+
+-- Lasmigona_costata **
Bolotov et al., 2017 [Publ 3764], fig. S1
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 15 |
18 |
0 |
3 |
116 |
13 |
2 |
37.0 |
+-- Unio_crassus
|
+-- Unio_pictorum
|
+-- Unio_tumidus
|
+-- Aculamprotula_tortuosa
|
+-- Cuneopsis_heudei
+--+
| +-- Tchangsinaia_piscicula
| |
| +-- Cuneopsis_rufescens
| |
| +-- Nodularia_douglasiae
| |
| +-- Nodularia_nuxpersicae
| |
| +-- Schistodesmus_lampreyanus
|
| +-- Alasmidonta_marginata **
| |
| +-- Anodonta_anatina **
--+ |
| +-- Anodonta_cygnea **
| |
| +-- Anodonta_nuttalliana **
| |
| +-- Lasmigona_costata **
| |
| +-- Pseudanodonta_complanata **
| |
| +-- Pyganodon_grandis **
| |
| +-- Simpsonaias_ambigua **
| |
| +-- Strophitus_undulatus **
+--+
+-- Sinanodonta_lauta **
|
+-- Cristaria_plicata **
|
+-- Cristaria_radiata **
|
+-- Pletholophus_tenuis **
|
+-- Sinanodonta_lucida **
|
+-- Sinanodonta_jourdyi **
|
+-- Lanceolaria_gladiola
|
+-- Lanceolaria_grayii[1]
|
+-- Lanceolaria_grayii[2]
Chase et al., 2018 [Publ 3847], fig. 4
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 15 |
15 |
13 |
3 |
24 |
2 |
1 |
38.0 |
+-- Unio_delphinus
+--------------------+
| +-- Unio_tumidus
|
--+ +-------------------- Sinanodonta_woodiana **
| |
| | +-- Anodonta_cygnea **
| | +--------------+
+--+ | +-- Anodonta_anatina **
| |
| | +-- Pyganodon_lacustris **
| | +-----+
+--+ | +-- Pyganodon_grandis **
| +-----+
| | | +----- Utterbackia_peninsularis[2] **
| | +--+
| | | +-- Utterbackia_peninsularis[1] **
| | +--+
+--+ +-- Utterbackia_imbecillis **
|
| +-- Lasmigona_costata **
| +--------+
| | +-- Lasmigona_complanata **
| |
+--+ +-- Alasmidonta_undulata **
| +-----+
| | +-- Alasmidonta_marginata **
+--+
| +----- Strophitus_undulatus **
+--+
| +-- Platynaias_compressa **
+--+
+-- Platynaias_subviridis **
Lopes-Lima et al., 2017 [Publ 3745], fig. 1-2
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 14 |
14 |
11 |
3 |
67 |
15 |
2 |
33.0 |
+----- Unio_gibbus
|
+-----------+----- Unio_tumidus
| |
| | +-- Unio_crassus
| +--+
+--+ +-- Unio_pictorum
| |
| | +-------------- Aculamprotula_tortuosa
| | |
| +--+ +-- Nodularia_nuxpersicae
| | +--------+
| | | +-- Nodularia_douglasiae
| +--+
| | +-------- Schistodesmus_lampreyanus
| +--+
| | +----- Tchangsinaia_piscicula
| +--+
--+ | +-- Cuneopsis_heudei
| +--+
| +-- Cuneopsis_rufescens
|
| +-------- Lanceolaria_gladiola
| +--------+
| | | +----- Lanceolaria_grayii[2]
| | +--+
| | | +-- Lanceolaria_lanceolata
| | +--+
| | +-- Lanceolaria_grayii[1]
| |
+--+ +----------- Cristaria_plicata **
| +--+
| | | +-------- Anemina_arcaeformis **
| | +--+
| | | +----- Pletholophus_tenuis **
| | +--+
| | | +-- Sinanodonta_jourdyi **
| | +--+
+--+ +-- Sinanodonta_lucida **
|
| +----- Pseudanodonta_complanata **
| |
| +-----+ +-- Anodonta_anatina **
| | | |
| | +--+-- Anodonta_cygnea **
| | |
+--+ +-- Anodonta_nuttalliana **
|
| +-- Pyganodon_grandis **
| +-----+
| | +-- Simpsonaias_ambigua **
+--+
| +----- Strophitus_undulatus **
+--+
| +-- Alasmidonta_marginata **
+--+
+-- Lasmigona_costata **
Riccardi et al., 2019 [Publ 4089], fig. 2
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 14 |
14 |
12 |
3 |
6 |
4 |
2 |
41.0 |
+-- Lanceolaria_grayii
+--+
| +-- Lanceolaria_lanceolata
+-----------------+
| | +-- Unio_mancus
| +--+
| +-- Unio_elongatulus
|
| +-------- Cristaria_plicata **
--+ |
| +-----------+ +-- Anemina_euscaphys **
| | | +--+
| | | | +-- Anemina_arcaeformis **
| | +--+
| | | +-- Sinanodonta_lucida **
+--+ +--+
| +-- Sinanodonta_woodiana **
|
| +----------------- Anodonta_anatina **
| |
| | +----- Pseudanodonta_complanata **
+--+ +--------+
| | | +-- Anodonta_exulcerata **
| | +--+
+--+ +-- Anodonta_cygnea **
|
| +----------- Anodonta_nuttalliana **
+--+
| +-------- Platynaias_compressa **
+--+
| +----- Pyganodon_grandis **
+--+
| +-- Utterbackia_peninsularis **
+--+
+-- Utterbackia_imbecillis **
Jeratthitikul et al., 2022 [Publ 4260], fig. S1, 1
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 14 |
14 |
12 |
3 |
62 |
12 |
2 |
33.0 |
+----- Unio_tumidus
+-----------+
| | +-- Unio_pictorum
| +--+
+--+ +-- Unio_crassus
| |
| | +-------------- Aculamprotula_tortuosa
| | |
| +--+ +-- Nodularia_douglasiae
| | +--------+
| | | +-- Nodularia_nuxpersicae
| +--+
| | +-------- Schistodesmus_lampreyanus
| +--+
| | +----- Cuneopsis_heudei
--+ +--+
| | +-- Tchangsinaia_piscicula
| +--+
| +-- Cuneopsis_rufescens
|
| +-- Lanceolaria_grayii
| +--------------+
| | +-- Lanceolaria_gladiola
| |
| | +----------- Cristaria_plicata **
| | +--+
+--+ | | +-------- Pletholophus_tenuis **
| | +--+
| | | +----- Sinanodonta_lucida **
| | +--+
| | | +-- Sinanodonta_lauta **
| | +--+
+--+ +-- Sinanodonta_jourdyi **
|
| +-------- Pseudanodonta_complanata **
| +--+
| | | +----- Anodonta_nuttalliana **
| | +--+
| | | +-- Anodonta_anatina **
| | +--+
+--+ +-- Anodonta_cygnea **
|
| +-- Pyganodon_grandis **
| +-----+
| | +-- Simpsonaias_ambigua **
+--+
| +----- Strophitus_undulatus **
+--+
| +-- Alasmidonta_marginata **
+--+
+-- Lasmigona_costata **
Jeratthitikul et al., 2022 [Publ 4260], fig. S2, 2
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 14 |
14 |
12 |
3 |
62 |
12 |
2 |
33.0 |
+----- Unio_tumidus
+-----------+
| | +-- Unio_pictorum
| +--+
+--+ +-- Unio_crassus
| |
| | +-------------- Aculamprotula_tortuosa
| | |
| +--+ +-- Nodularia_douglasiae
| | +--------+
| | | +-- Nodularia_nuxpersicae
| +--+
| | +-------- Schistodesmus_lampreyanus
| +--+
| | +----- Cuneopsis_heudei
--+ +--+
| | +-- Tchangsinaia_piscicula
| +--+
| +-- Cuneopsis_rufescens
|
| +-- Lanceolaria_grayii
| +--------------+
| | +-- Lanceolaria_gladiola
| |
| | +----------- Cristaria_plicata **
| | +--+
+--+ | | +-------- Pletholophus_tenuis **
| | +--+
| | | +----- Sinanodonta_lucida **
| | +--+
| | | +-- Sinanodonta_jourdyi **
| | +--+
+--+ +-- Sinanodonta_lauta **
|
| +-------- Pseudanodonta_complanata **
| +--+
| | | +----- Anodonta_anatina **
| | +--+
| | | +-- Anodonta_nuttalliana **
| | +--+
+--+ +-- Anodonta_cygnea **
|
| +-- Alasmidonta_marginata **
| +-----+
| | +-- Lasmigona_costata **
+--+
| +----- Pyganodon_grandis **
+--+
| +-- Strophitus_undulatus **
+--+
+-- Simpsonaias_ambigua **
Sano et al., 2022 [Publ 4311], fig. 2
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 14 |
14 |
12 |
1 |
0 |
0 |
0 |
40.0 |
+-------- Pletholophus_tenuis **
+-----------+
| | +----- Cristaria_plicata **
| +--+
| | +-- Buldowskia_iwakawai **
| +--+
| +-- Buldowskia_kamiyai **
--+
| +-------- Beringiana_fukuharai **
| +--------+
| | | +----- Beringiana_japonica **
| | +--+
| | | +-- Beringiana_gosannensis **
+--+ +--+
| +-- Beringiana_beringiana **
|
| +-------------- Sinanodonta_tumens **
+--+
| +----------- Sinanodonta[2] **
+--+
| +-------- Sinanodonta_calipygos **
+--+
| +----- Sinanodonta_lauta **
+--+
| +-- Sinanodonta_woodiana **
+--+
+-- Sinanodonta[1] **
Breton et al., 2011 [Publ 3376], fig. 3
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 13 |
13 |
11 |
2 |
26 |
0 |
0 |
37.0 |
+-------------------- Cristaria_plicata **
|
| +-- Lasmigona_complanata **
--+ +--------------+
| | +-- Lasmigona_costata **
| |
+--+ +-------------- Strophitus_undulatus **
| |
| | +-- Platynaias_compressa **
| | +--+
+--+ | +-- Platynaias_subviridis **
| +-----+
| | | +-- Alasmidonta_marginata **
| | +--+
| | +-- Alasmidonta_undulata **
+--+
| +-- Pyganodon_lacustris **
| +-----+
| | +-- Pyganodon_grandis **
+--+
| +----- Utterbackia_peninsularis[2] **
+--+
| +-- Utterbackia_peninsularis[1] **
+--+
+-- Utterbackia_imbecillis **
Wu et al., 2018 [Publ 3836], fig. 2
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 13 |
13 |
9 |
3 |
40 |
20 |
4 |
38.0 |
+----------------------- Lepidodesma_languilati
|
| +----------- Aculamprotula_tortuosa
| +-----+
| | | +-------- Aculamprotula_scripta
| | +--+
| | | +----- Aculamprotula_coreana
| | +--+
| | | +-- Aculamprotula_fibrosa
+-----+ +--+
| | +-- Aculamprotula_tientsinensis
| |
| | +-- Unio_crassus
| | +-----------+
| | | +-- Unio_pictorum
| | |
| +--+ +----- Nodularia_douglasiae
| | +-----+
| | | | +-- Schistodesmus_lampreyanus
--+ | | +--+
| +--+ +-- Schistodesmus_spinosus
| |
| | +-------- Tchangsinaia_piscicula
| +--+
| | +----- Cuneopsis_rufescens
| +--+
| | +-- Cuneopsis_celtiformis
| +--+
| +-- Cuneopsis_heudei
|
| +----------- Acuticosta_chinensis
| +--------+
| | | +-------- Lanceolaria_gladiola
| | +--+
| | | +----- Lanceolaria_grayii
| | +--+
| | | +-- Lanceolaria_lanceolata
| | +--+
| | +-- Lanceolaria_triformis
+--+
| +-------- Anemina_arcaeformis **
| |
| +-------- Cristaria_plicata **
| +--------+
| | | +-- Sinanodonta_woodiana **
| | | +--+
| | | | +-- Sinanodonta_elliptica **
| | +--+
+--+ | +-- Sinanodonta_lucida **
| +--+
| +-- Sinanodonta_angula **
|
| +-------------- Anodonta_cygnea **
| |
+--+ +----------- Anodonta_anatina **
| |
+--+ +-------- Strophitus_undulatus **
| |
+--+ +----- Platynaias_compressa **
| |
+--+----- Alasmidonta_marginata **
|
| +-- Pyganodon_cataracta **
+--+
+-- Pyganodon_grandis **
Froufe et al., 2017 [Publ 3802], fig. 6
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 12 |
12 |
9 |
2 |
0 |
0 |
0 |
37.0 |
+-- Anodonta_anatina **
+--------------------+
| +-- Anodonta_cygnea **
--+
| +-------------------- Anemina_arcaeformis **
| |
+--+ +-- Cristaria_plicata **
| +--------------+
| | +-- Cristaria_radiata **
+--+
| +-------------- Beringiana_beringiana **
| |
+--+ +----------- Sinanodonta_lucida **
| |
+--+ +-------- Sinanodonta_schrenkii **
| |
+--+ +----- Sinanodonta_woodiana[1] **
| |
+--+ +-- Sinanodonta_lauta **
| |
+--+-- Sinanodonta_elliptica **
|
+-- Sinanodonta_woodiana[2] **
Aunins et al., 2018 [Publ 3863], fig. 1
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 12 |
12 |
10 |
3 |
29 |
15 |
4 |
39.0 |
+----- Aculamprotula_tortuosa
+--------+
| | +-- Aculamprotula_coreana
| +--+
| +-- Aculamprotula_tientsinensis
|
+-----+ +-------- Nodularia_douglasiae
| | +--+
| | | | +----- Tchangsinaia_piscicula
| | | +--+
| | | | +-- Cuneopsis_heudei
| +--+ +--+
| | +-- Schistodesmus_lampreyanus
| |
| | +-------- Unio_tumidus
| +--+
| | +----- Unio_crassus
--+ +--+
| | +-- Unio_pictorum
| +--+
| +-- Unio_delphinus
|
| +----- Acuticosta_chinensis
| +-----------+
| | | +-- Lanceolaria_grayii
| | +--+
| | +-- Lanceolaria_lanceolata
| |
| | +-------- Cristaria_plicata **
+--+ |
| +-----+ +-- Anemina_arcaeformis **
| | | +--+
| | | | +-- Anemina_euscaphys **
| | +--+
| | | +-- Sinanodonta_woodiana **
| | +--+
+--+ +-- Sinanodonta_lucida **
|
| +-- Anodonta_anatina **
| +--------+
| | +-- Anodonta_cygnea **
| |
+--+ +-- Prolasmidonta_heterodon **
| +-----+
| | +-- Alasmidonta_varicosa **
+--+
| +----- Pyganodon_grandis **
+--+
| +-- Utterbackia_imbecillis **
+--+
+-- Utterbackia_peninsularis **
Lopes-Lima et al., 2020 [Publ 4080], fig. 10
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 12 |
12 |
8 |
1 |
2 |
0 |
0 |
38.0 |
+-- Beringiana_japonica **
+--------------+
| +-- Beringiana_beringiana **
|
--+ +-- Sinanodonta_angula **
| +-----------+
| | +-- Sinanodonta_lucida **
| |
+--+ +----------- Sinanodonta_schrenkii **
| |
| | +-- Sinanodonta_calipygos **
| +--------+
+--+ +-- Sinanodonta_tumens **
|
| +-------- Sinanodonta_woodiana[2] **
| |
+--+-------- Sinanodonta_jourdyi **
|
| +----- Sinanodonta_elliptica **
+--+
| +-- Sinanodonta_lauta **
+--+
+-- Sinanodonta_woodiana[1] **
Bolotov et al., 2020 [Publ 4084], fig. 1
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 12 |
12 |
10 |
2 |
24 |
7 |
2 |
34.0 |
+----------- Middendorffinaia_mongolica
+-----------+
| | +-------- Nodularia_douglasiae
| +--+
| | +----- Unio_tumidus
| +--+
| | +-- Unio_pictorum
| +--+
--+ +-- Unio_crassus
|
| +-- Lanceolaria_grayii
| +-----------------+
| | +-- Lanceolaria
| |
| | +----- Anodonta_anatina **
+--+ +-----------+
| | | +-- Pseudanodonta_complanata **
| | +--+
| | +-- Anodonta_cygnea **
| |
+--+ +-------- Amuranodonta_kijaensis **
| +-----+
| | | +----- Anemina_arcaeformis **
| | +--+
| | | +-- Buldowskia_suifunica **
+--+ +--+
| +-- Buldowskia_shadini **
|
| +----------- Cristaria_plicata **
+--+
| +-------- Beringiana_beringiana **
+--+
| +----- Sinanodonta_schrenkii **
+--+
| +-- Sinanodonta_lauta **
+--+
+-- Sinanodonta_woodiana **
Bogan et al., 2023 [Publ 4295], fig. 6
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 12 |
12 |
9 |
1 |
1 |
0 |
0 |
40.0 |
+----------------------- Pletholophus_honglinhensis **
|
--+ +-------------------- Cristaria_truncata **
| |
+--+ +-- Sinanodonta_angula **
| +--------------+
| | +-- Sinanodonta_lucida **
+--+
| +-------------- Sinanodonta_schrenkii **
| |
+--+ +-- Sinanodonta_calipygos **
| +--------+
| | +-- Sinanodonta_tumens **
+--+
| +-------- Sinanodonta_woodiana[2] **
| |
+--+-------- Sinanodonta_jourdyi **
|
| +----- Sinanodonta_elliptica **
+--+
| +-- Sinanodonta_lauta **
+--+
+-- Sinanodonta_woodiana[1] **
Soroka & Burzynski, 2017 [Publ 3837], fig. 1
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 11 |
11 |
9 |
3 |
13 |
13 |
3 |
38.0 |
+-------- Cristaria_plicata **
|
+-----+ +-- Anemina_arcaeformis **
| | +--+
| | | +-- Anemina_euscaphys **
| +--+
| | +-- Sinanodonta_woodiana **
+--------+ +--+
| | +-- Sinanodonta_lucida **
| |
| | +-- Anodonta_cygnea **
| | +--------+
| | | +-- Anodonta_anatina **
| +--+
| | +-------- Platynaias_compressa **
| +--+
--+ | +----- Pyganodon_grandis **
| +--+
| | +-- Utterbackia_imbecillis **
| +--+
| +-- Utterbackia_peninsularis **
|
| +-- Lanceolaria_lanceolata
| +-----------------+
| | +-- Lanceolaria_grayii
| |
+--+ +----------------- Lepidodesma_languilati
| |
| | +----- Aculamprotula_tortuosa
+--+ +--------+
| | | +-- Aculamprotula_tientsinensis
| | +--+
| | +-- Aculamprotula_coreana
+--+
| +----- Schistodesmus_lampreyanus
| +-----+
| | | +-- Tchangsinaia_piscicula
| | +--+
+--+ +-- Nodularia_douglasiae
|
| +-------- Unio_tumidus
+--+
| +----- Unio_crassus
+--+
| +-- Unio_pictorum
+--+
+-- Unio_delphinus
Huang et al., 2019 [Publ 3843], fig. 3
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 11 |
11 |
9 |
3 |
46 |
16 |
4 |
37.0 |
+----- Lanceolaria_gladiola
+-----------------+
| | +-- Lanceolaria_lanceolata
| +--+
| +-- Lanceolaria_grayii
--+
| +-------------------- Acuticosta_chinensis
| |
| | +-------- Cristaria_plicata **
| | |
+--+ +--------+ +-- Sinanodonta_lucida **
| | | +--+
| | | | +-- Sinanodonta_woodiana **
| | +--+
+--+ | +-- Anemina_arcaeformis **
| +--+
| +-- Anemina_euscaphys **
|
| +-------------- Anodonta_cygnea **
+--+
| +----------- Anodonta_anatina **
+--+
| +-------- Lasmigona_costata **
+--+
| +----- Pyganodon_grandis **
+--+
| +-- Utterbackia_imbecillis **
+--+
+-- Utterbackia_peninsularis **
Fukata & Iigo, 2020 [Publ 4106], fig. 1
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 11 |
11 |
9 |
3 |
43 |
17 |
4 |
38.0 |
+-------------- Acuticosta_chinensis
|
| +-------- Cristaria_plicata **
| |
| +--+ +-- Sinanodonta_woodiana **
+--+ | | +--+
| | | | | +-- Sinanodonta_lucida **
| | | +--+
| | | | +-- Anemina_arcaeformis **
| | | +--+
| +--+ +-- Anemina_euscaphys **
| |
| | +-- Anodonta_anatina **
| | +-----+
| | | +-- Anodonta_cygnea **
| | |
| +--+ +-- Pyganodon_grandis **
--+ | +--+
| | | +-- Utterbackia_peninsularis **
| +--+
| | +-- Alasmidonta_varicosa **
| +--+
| +-- Prolasmidonta_heterodon **
|
| +----- Aculamprotula_tortuosa
| +--------+
| | | +-- Aculamprotula_tientsinensis
| | +--+
| | +-- Aculamprotula_coreana
| |
+--+ +-------- Unio_tumidus
| +--+
| | | +----- Unio_crassus
| | +--+
| | | +-- Unio_pictorum
| | +--+
+--+ +-- Unio_delphinus
|
| +-- Schistodesmus_lampreyanus
| +-----+
| | +-- Cuneopsis_heudei
+--+
| +----- Pseudocuneopsis_capitata
+--+
| +-- Nodularia_douglasiae
+--+
+-- Tchangsinaia_piscicula
Chen et al., 2020 [Publ 4170], fig. 1
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 11 |
34 |
32 |
3 |
23 |
4 |
2 |
63.0 |
+----- Prolasmidonta_heterodon **
+--+
| | +-- Alasmidonta_varicosa **
| +--+
| +-- Anemina_euscaphys **
+-----------------+
| | +-- Lamprotula_caveata
| | +--+
| | | +-- Chamberlainia_somsakpanhai
| +--+
| | +-- Pseudodon_vondembuschianus
| +--+
| +-- Pilsbryoconcha_exilis
|
| +-------- Schistodesmus_lampreyanus
| +-----+
| | | +----- Lepidodesma_languilati
| | +--+
--+ | | +-- Aculamprotula_coreana
| | +--+
| +--------+ +-- Aculamprotula_tortuosa
| | |
| | | +----- Platynaias_compressa **
| | | +-----+
| | | | | +-- Utterbackia_imbecillis **
| | | | +--+
| | +--+ +-- Pyganodon_grandis **
| | |
| | | +-------- Sinanodonta_lucida **
| | +--+
| | | +----- Cristaria_plicata **
| | +--+
+--+ | +-- Anodonta_cygnea **
| +--+
| +-- Anodonta_anatina **
|
| +----------------- Gibbosula_crassa
| +--+
| | | +-------------- Quadrula_quadrula
| | +--+
| | | +----------- Toxolasma_parvum
| | +--+
| | | +-------- Lampsilis_ornata
| | +--+
| | | +----- Potamilus_leptodon
+--+ +--+
| | +-- Lampsilis_siliquoidea
| +--+
| +-- Lampsilis_powellii
|
| +-- Utterbackia_peninsularis **
| +--------------+
| | +-- Potamilus_alatus
+--+
| +-------------- Lamprotula_leaii
| |
+--+ +-- Sinohyriopsis_cumingii
| +--------+
| | +-- Sinohyriopsis_schlegelii
+--+
| +-------- Parvasolenaia_rivularis
+--+
| +----- Sinosolenaia_carinata
+--+
| +-- Sinosolenaia_iridinea
+--+
+-- Sinosolenaia_oleivora
Wu et al., 2022 [Publ 4238], fig. 2d
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 11 |
11 |
9 |
3 |
28 |
28 |
4 |
38.0 |
+----- Lanceolaria_gladiola
+--------------+
| | +-- Lanceolaria_grayii
| +--+
| +-- Lanceolaria_lanceolata
--+
| +----------------- Acuticosta_chinensis
| |
| | +-------- Cristaria_plicata **
| | |
+--+ +-----+ +-- Sinanodonta_woodiana **
| | | +--+
| | | | +-- Sinanodonta_lucida **
| | +--+
| | | +-- Anemina_arcaeformis **
+--+ +--+
| +-- Anemina_euscaphys **
|
| +-- Anodonta_cygnea **
| +--------+
| | +-- Anodonta_anatina **
+--+
| +-------- Platynaias_compressa **
+--+
| +----- Pyganodon_grandis **
+--+
| +-- Utterbackia_imbecillis **
+--+
+-- Utterbackia_peninsularis **
Wu et al., 2022 [Publ 4238], fig. 3
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 11 |
11 |
9 |
3 |
63 |
25 |
4 |
38.0 |
+-------- Acuticosta_chinensis
+--------+
| | +----- Lanceolaria_gladiola
| +--+
| | +-- Lanceolaria_grayii
| +--+
| +-- Lanceolaria_lanceolata
|
+-----------+ +-------- Cristaria_plicata **
| | |
| | +-----+ +-- Sinanodonta_woodiana **
| | | | +--+
| | | | | +-- Sinanodonta_lucida **
| | | +--+
| | | | +-- Anemina_arcaeformis **
| +--+ +--+
| | +-- Anemina_euscaphys **
| |
| | +-- Anodonta_cygnea **
| | +--------+
| | | +-- Anodonta_anatina **
| +--+
| | +-------- Platynaias_compressa **
--+ +--+
| | +----- Pyganodon_grandis **
| +--+
| | +-- Utterbackia_imbecillis **
| +--+
| +-- Utterbackia_peninsularis **
|
| +-------- Aculamprotula_tortuosa
| |
| +-----------------+ +-- Aculamprotula_tientsinensis
| | | +--+
| | | | +-- Aculamprotula_coreana
| | +--+
| | | +-- Aculamprotula_polysticta
| | +--+
| | +-- Aculamprotula_scripta
+--+
| +-------- Unio_tumidus
| +--------------+
| | | +----- Unio_crassus
| | +--+
| | | +-- Unio_pictorum
| | +--+
+--+ +-- Unio_delphinus
|
| +-- Pseudocuneopsis_capitata
| +-----------------+
| | +-- Pseudocuneopsis_sichuanensis
| |
+--+ +-- Nodularia_douglasiae
| +--------------+
| | +-- Nodularia_breviconcha
| |
+--+ +-- Schistodesmus_lampreyanus
| +-----------+
| | +-- Schistodesmus_spinosus
+--+
| +----------- Tchangsinaia_piscicula
+--+
| +-------- Cuneopsis_rufescens
+--+
| +----- Cuneopsis_celtiformis
+--+
| +-- Cuneopsis_heudei
+--+
+-- Cuneopsis_demangei
Krebs et al., 2003 [Publ 2691], fig. 3
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 10 |
10 |
5 |
2 |
24 |
0 |
0 |
37.5 |
+----------- Utterbackiana_couperiana **
|
+----------- Platynaias_compressa **
|
+----------- Alasmidonta_triangulata **
|
--+ +-- Lasmigona_complanata **
+--------+
| +-- Lasmigona_costata **
|
| +-- Strophitus_radiatus **
| +-----+
| | +-- Strophitus_undulatus **
+--+
| +----- Utterbackia_imbecillis **
+--+
| +-- Pyganodon_grandis **
+--+
+-- Anodonta **
Fonseca et al., 2016 [Publ 3775], fig. 1
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 10 |
10 |
8 |
3 |
22 |
9 |
2 |
37.0 |
+-- Lanceolaria_lanceolata
+-----------+
| +-- Lanceolaria_grayii
|
+--+ +-- Aculamprotula_tortuosa
| | +--------+
| | | +-- Aculamprotula_coreana
| | |
| +--+ +-- Unio_pictorum
| | +-----+
| | | +-- Unio_delphinus
| +--+
| | +----- Schistodesmus_lampreyanus
| +--+
--+ | +-- Nodularia_douglasiae
| +--+
| +-- Tchangsinaia_piscicula
|
| +-------- Cristaria_plicata **
| |
| +-----+ +-- Anemina_euscaphys **
| | | +--+
| | | | +-- Anemina_arcaeformis **
| | +--+
+--+ | +-- Sinanodonta_woodiana **
| +--+
| +-- Sinanodonta_lucida **
|
| +----------- Anodonta_anatina **
+--+
| +-------- Platynaias_compressa **
+--+
| +----- Pyganodon_grandis **
+--+
| +-- Utterbackia_peninsularis **
+--+
+-- Utterbackia_imbecillis **
Lopes-Lima et al., 2017 [Publ 3792], fig. 2
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 10 |
10 |
8 |
3 |
23 |
9 |
2 |
37.0 |
+-- Lanceolaria_lanceolata
+-----------+
| +-- Lanceolaria_grayii
|
+--+ +-- Aculamprotula_tortuosa
| | +--------+
| | | +-- Aculamprotula_coreana
| | |
| +--+ +-- Unio_delphinus
| | +-----+
| | | +-- Unio_pictorum
| +--+
| | +----- Schistodesmus_lampreyanus
| +--+
--+ | +-- Tchangsinaia_piscicula
| +--+
| +-- Nodularia_douglasiae
|
| +----------- Anodonta_anatina **
| +--+
| | | +-------- Platynaias_compressa **
| | +--+
| | | +----- Pyganodon_grandis **
| | +--+
+--+ | +-- Utterbackia_peninsularis **
| +--+
| +-- Utterbackia_imbecillis **
|
| +-------- Cristaria_plicata **
| |
+-----+ +-- Sinanodonta_lucida **
| +--+
| | +-- Sinanodonta_woodiana **
+--+
| +-- Anemina_arcaeformis **
+--+
+-- Anemina_euscaphys **
Bolotov et al., 2018 [Publ 3826], fig. 2
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 10 |
10 |
8 |
3 |
75 |
8 |
2 |
31.0 |
+----- Unio_tumidus
+--+
| | +-- Unio_crassus
| +--+
+--------------+ +-- Unio_pictorum
| |
| | +----- Nodularia_douglasiae[1]
| +--+
| | +-- Nodularia_nuxpersicae
| +--+
--+ +-- Nodularia_douglasiae[2]
|
| +-- Lanceolaria_gladiola
| +-----------------+
| | +-- Lanceolaria_grayii
| |
| | +-------- Cristaria_plicata **
+--+ +--------+
| | | +----- Pletholophus_tenuis **
| | +--+
| | | +-- Sinanodonta_jourdyi **
+--+ +--+
| +-- Sinanodonta_lauta **
|
| +-------------- Anodonta_anatina **
+--+
| +----------- Pyganodon_grandis **
+--+
| +-------- Simpsonaias_ambigua **
+--+
| +----- Strophitus_undulatus **
+--+
| +-- Alasmidonta_marginata **
+--+
+-- Lasmigona_costata **
Huang et al., 2018 [Publ 3828], fig. 6
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 10 |
10 |
8 |
3 |
33 |
9 |
1 |
37.0 |
+----- Aculamprotula_tortuosa
+--------+
| | +-- Aculamprotula_coreana
| +--+
| +-- Aculamprotula_tientsinensis
+--+
| | +-- Tchangsinaia_piscicula
| | +--------+
| | | +-- Nodularia_douglasiae
| +--+
| | +-------- Unio_tumidus
| +--+
| | +----- Unio_crassus
| +--+
--+ | +-- Unio_delphinus
| +--+
| +-- Unio_pictorum
|
| +----------- Anodonta_anatina **
| +--+
| | | +-------- Platynaias_compressa **
| | +--+
| | | +----- Pyganodon_grandis **
| | +--+
+--+ | +-- Utterbackia_imbecillis **
| +--+
| +-- Utterbackia_peninsularis **
|
| +-------- Cristaria_plicata **
| |
+-----+ +-- Anemina_arcaeformis **
| +--+
| | +-- Anemina_euscaphys **
+--+
| +-- Sinanodonta_woodiana **
+--+
+-- Sinanodonta_lucida **
Wu et al., 2019 [Publ 3864], fig. 3
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 10 |
10 |
8 |
3 |
37 |
17 |
4 |
37.0 |
+-------- Aculamprotula_tortuosa
+--------+
| | +----- Aculamprotula_scripta
| +--+
| | +-- Aculamprotula_tientsinensis
| +--+
| +-- Aculamprotula_coreana
+--+
| | +-------- Unio_tumidus
| | +-----+
| | | | +----- Unio_crassus
| | | +--+
| | | | +-- Unio_delphinus
| +--+ +--+
| | +-- Unio_pictorum
| |
| | +----------- Pseudocuneopsis_capitata
| +--+
| | +-------- Nodularia_douglasiae
--+ +--+
| | +----- Tchangsinaia_piscicula
| +--+
| | +-- Cuneopsis_heudei
| +--+
| +-- Schistodesmus_lampreyanus
|
| +----- Acuticosta_chinensis
| +-----------+
| | | +-- Lanceolaria_grayii
| | +--+
| | +-- Lanceolaria_lanceolata
| |
+--+ +----------- Anodonta_anatina **
| +--+
| | | +-------- Platynaias_compressa **
| | +--+
| | | +----- Pyganodon_grandis **
| | +--+
+--+ | +-- Utterbackia_peninsularis **
| +--+
| +-- Utterbackia_imbecillis **
|
| +-------- Cristaria_plicata **
| |
+-----+ +-- Anemina_arcaeformis **
| +--+
| | +-- Anemina_euscaphys **
+--+
| +-- Sinanodonta_lucida **
+--+
+-- Sinanodonta_woodiana **
Cha et al., 2018 [Publ 3916], fig. 1
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 10 |
10 |
8 |
3 |
29 |
12 |
3 |
37.0 |
+----- Aculamprotula_tortuosa
+--------+
| | +-- Aculamprotula_coreana
| +--+
| +-- Aculamprotula_tientsinensis
+-----+
| | +----- Schistodesmus_lampreyanus
| | +-----+
| | | | +-- Nodularia_douglasiae
| | | +--+
| +--+ +-- Tchangsinaia_piscicula
| |
| | +-------- Unio_tumidus
| +--+
--+ | +----- Unio_crassus
| +--+
| | +-- Unio_pictorum
| +--+
| +-- Unio_delphinus
|
| +----------------- Lanceolaria_grayii
| |
| | +----------- Anodonta_anatina **
| | +--+
+--+ | | +-------- Platynaias_compressa **
| | +--+
| | | +----- Pyganodon_grandis **
| | +--+
+--+ | +-- Utterbackia_peninsularis **
| +--+
| +-- Utterbackia_imbecillis **
|
| +-------- Cristaria_plicata **
| |
+-----+ +-- Anemina_arcaeformis **
| +--+
| | +-- Anemina_euscaphys **
+--+
| +-- Sinanodonta_lucida **
+--+
+-- Sinanodonta_woodiana **
Froufe et al., 2019 [Publ 3936], fig. 3
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 10 |
10 |
8 |
3 |
35 |
13 |
3 |
37.0 |
+----- Aculamprotula_tortuosa
+--------+
| | +-- Aculamprotula_tientsinensis
| +--+
| +-- Aculamprotula_coreana
+-----+
| | +----- Nodularia_douglasiae
| | +-----+
| | | | +-- Tchangsinaia_piscicula
| | | +--+
| +--+ +-- Schistodesmus_lampreyanus
| |
| | +-------- Unio_tumidus
| +--+
| | +----- Unio_crassus
--+ +--+
| | +-- Unio_pictorum
| +--+
| +-- Unio_delphinus
|
| +-- Lanceolaria_lanceolata
| +--------------+
| | +-- Lanceolaria_grayii
| |
| | +-------- Cristaria_plicata **
+--+ |
| +-----+ +-- Anemina_arcaeformis **
| | | +--+
| | | | +-- Anemina_euscaphys **
| | +--+
+--+ | +-- Sinanodonta_lucida **
| +--+
| +-- Sinanodonta_woodiana **
|
| +----------- Anodonta_anatina **
+--+
| +-------- Lasmigona_complanata **
+--+
| +----- Pyganodon_grandis **
+--+
| +-- Utterbackia_peninsularis **
+--+
+-- Utterbackia_imbecillis **
Froufe et al., 2019 [Publ 3936], fig. 6
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 10 |
10 |
8 |
3 |
35 |
13 |
3 |
37.0 |
+----- Aculamprotula_tortuosa
+--------+
| | +-- Aculamprotula_tientsinensis
| +--+
| +-- Aculamprotula_coreana
+-----+
| | +----- Nodularia_douglasiae
| | +-----+
| | | | +-- Tchangsinaia_piscicula
| | | +--+
| +--+ +-- Schistodesmus_lampreyanus
| |
| | +-------- Unio_tumidus
| +--+
| | +----- Unio_crassus
--+ +--+
| | +-- Unio_pictorum
| +--+
| +-- Unio_delphinus
|
| +-- Lanceolaria_lanceolata
| +--------------+
| | +-- Lanceolaria_grayii
| |
| | +----------- Anodonta_anatina **
+--+ +--+
| | | +-------- Lasmigona_complanata **
| | +--+
| | | +----- Pyganodon_grandis **
| | +--+
+--+ | +-- Utterbackia_peninsularis **
| +--+
| +-- Utterbackia_imbecillis **
|
| +-------- Cristaria_plicata **
| |
+-----+ +-- Sinanodonta_lucida **
| +--+
| | +-- Sinanodonta_woodiana **
+--+
| +-- Anemina_arcaeformis **
+--+
+-- Anemina_euscaphys **
Lopes-Lima et al., 2020 [Publ 4080], fig. 6
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 10 |
10 |
8 |
1 |
2 |
0 |
0 |
36.0 |
+-- Sinanodonta_lauta **
+--------------+
| +-- Sinanodonta_woodiana **
|
--+ +-------------- Amuranodonta_kijaensis **
| |
| | +-- Anemina_arcaeformis **
+--+ +--------+
| | +-- Anemina_euscaphys **
| |
+--+ +-- Buldowskia_iwakawai **
| +-----+
| | +-- Buldowskia_kamiyai **
+--+
| +----- Buldowskia_shadini **
+--+
| +-- Buldowskia_suifunica **
+--+
+-- Buldowskia_flavotincta **
Wu et al., 2020 [Publ 4121], fig. 3
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 10 |
10 |
8 |
3 |
40 |
18 |
4 |
37.0 |
+----- Acuticosta_chinensis
+-----------+
| | +-- Lanceolaria_lanceolata
| +--+
| +-- Lanceolaria_grayii
|
+--+ +----------- Anodonta_anatina **
| | +--+
| | | | +-------- Platynaias_compressa **
| | | +--+
| | | | +----- Pyganodon_grandis **
| | | +--+
| +--+ | +-- Utterbackia_peninsularis **
| | +--+
| | +-- Utterbackia_imbecillis **
| |
| | +-------- Cristaria_plicata **
| | |
| +-----+ +-- Sinanodonta_woodiana **
| | +--+
--+ | | +-- Sinanodonta_lauta **
| +--+
| | +-- Anemina_arcaeformis **
| +--+
| +-- Anemina_euscaphys **
|
| +-------- Aculamprotula_tortuosa
| |
| +--------+ +-- Aculamprotula_coreana
| | | +--+
| | | | +-- Aculamprotula_tientsinensis
| | +--+
| | | +-- Aculamprotula_polysticta
| | +--+
+--+ +-- Aculamprotula_scripta
|
| +-------- Unio_tumidus
| +-----+
| | | +----- Unio_crassus
| | +--+
| | | +-- Unio_delphinus
+--+ +--+
| +-- Unio_pictorum
|
| +----------- Pseudocuneopsis_capitata
+--+
| +-------- Nodularia_douglasiae
+--+
| +----- Tchangsinaia_piscicula
+--+
| +-- Cuneopsis_heudei
+--+
+-- Schistodesmus_lampreyanus
Zieritz et al., 2021 [Publ 4231], fig. 3-F, 5
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 10 |
10 |
8 |
3 |
39 |
13 |
3 |
37.0 |
+----- Aculamprotula_tortuosa
+--------+
| | +-- Aculamprotula_tientsinensis
| +--+
| +-- Aculamprotula_coreana
+-----+
| | +----- Nodularia_douglasiae
| | +-----+
| | | | +-- Schistodesmus_lampreyanus
| | | +--+
| +--+ +-- Tchangsinaia_piscicula
| |
| | +-------- Unio_tumidus
| +--+
| | +----- Unio_crassus
--+ +--+
| | +-- Unio_pictorum
| +--+
| +-- Unio_delphinus
|
| +-- Lanceolaria_grayii
| +--------------+
| | +-- Lanceolaria_lanceolata
| |
| | +----------- Anodonta_anatina **
+--+ +--+
| | | +-------- Platynaias_compressa **
| | +--+
| | | +----- Pyganodon_grandis **
| | +--+
+--+ | +-- Utterbackia_imbecillis **
| +--+
| +-- Utterbackia_peninsularis **
|
| +-------- Cristaria_plicata **
| |
+-----+ +-- Sinanodonta_lucida **
| +--+
| | +-- Sinanodonta_woodiana **
+--+
| +-- Anemina_arcaeformis **
+--+
+-- Anemina_euscaphys **
Chen et al., 2023 [Publ 4302], fig. 2
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 10 |
10 |
8 |
3 |
30 |
21 |
4 |
25.0 |
+----------------------- Lepidodesma_languilati
|
| +-- Lanceolaria_gladiola
| +--------------+
| | +-- Lanceolaria_oxyrhyncha
| |
| | +-- Alasmidonta_marginata **
+-----+ +--+
| | | +-- Lasmigona_costata **
| | +--------+
| | | | +-- Anodonta_anatina **
| | | +--+
| +--+ +-- Pseudanodonta_complanata **
| |
| | +----------- Cristaria_plicata **
--+ | |
| +--+ +-- Anemina_arcaeformis **
| | +-----+
| | | +-- Amuranodonta_kijaensis **
| +--+
| | +----- Pletholophus_tenuis **
| +--+
| | +-- Sinanodonta_jourdyi **
| +--+
| +-- Beringiana_beringiana **
|
| +-------------------- Aculamprotula_fibrosa
| |
| | +----- Middendorffinaia_mongolica
| | +-----------+
+--+ | | +-- Nodularia_douglasiae
| | +--+
| | +-- Nodularia_breviconcha
| |
| | +-- Cuneopsis_celtiformis
+--+ +-----------+
| | +-- Cuneopsis_heudei
| |
| | +-- Diaurora_aurorea
| | +--+
| | | +-- Diaurora_laeve
| +--------+
+--+ | +-- Schistodesmus_lampreyanus
| +--+
| +-- Schistodesmus_spinosus
|
| +-- Pseudocuneopsis_sichuanensis
| +--------+
| | +-- Pseudocuneopsis_capitata
| |
+--+ +-- Inversiunio_yanagawensis
| +-----+
| | +-- Inversiunio_reinianus
+--+
| +----- Tchangsinaia_piscicula
+--+
| +-- Unio_pictorum
+--+
+-- Unio_crassus
King et al., 1999 [Publ 2699], fig. 2
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 9 |
9 |
7 |
1 |
0 |
0 |
0 |
38.0 |
+-- Pyganodon_grandis **
+-----------+
| +-- Pyganodon_fragilis **
|
--+ +----- Lasmigona_costata **
| +-----+
| | | +-- Lasmigona_complanata **
| | +--+
+--+ +-- Lasmigona_alabamensis **
|
| +-------- Strophitus_undulatus **
+--+
| +----- Prolasmidonta_heterodon **
+--+
| +-- Platynaias_subviridis **
+--+
+-- Platynaias_compressa **
King et al., 1999 [Publ 2699], fig. 4
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 9 |
9 |
3 |
1 |
0 |
0 |
0 |
35.0 |
+-- Lasmigona_complanata **
+-----+
| +-- Lasmigona_costata **
|
--+ +----- Prolasmidonta_heterodon **
| |
| +----- Strophitus_undulatus **
| |
+--+ +-- Platynaias_compressa **
| |
| +-- Platynaias_subviridis[1] **
| |
+--+-- Platynaias_subviridis[2] **
|
+-- Platynaias_subviridis[3] **
|
+-- Platynaias_subviridis[4] **
Boyer et al., 2011 [Publ 2718], fig. 4
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 9 |
9 |
6 |
1 |
29 |
0 |
0 |
36.0 |
+-------------- Strophitus_undulatus **
|
| +----- Lasmigona_complanata **
--+ +-----+
| | | +-- Lasmigona_costata **
| | +--+
+--+ +-- Arcidens_confragosus **
|
| +-------- Utterbackia_imbecillis **
| |
+--+-------- Alasmidonta_marginata **
|
| +----- Pyganodon_grandis **
+--+
| +-- Anodontoides_ferussacianus **
+--+
+-- Platynaias_compressa **
Cyr et al., 2007 [Publ 2745], fig. 2C
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 9 |
9 |
4 |
2 |
0 |
0 |
0 |
34.0 |
+----------- Pseudanodonta_complanata **
|
| +-- Pyganodon_grandis **
--+ +-----+
| | +-- Pyganodon[1] **
| |
+--+ +----- Pyganodon[3] **
| |
| | +-- Pyganodon_cataracta[1] **
+--+ |
| +-- Pyganodon_fragilis[1] **
| |
+--+-- Pyganodon[2] **
|
+-- Pyganodon_cataracta[2] **
|
+-- Pyganodon_fragilis[2] **
Sano et al., 2017 [Publ 3799], fig. 2
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 9 |
9 |
7 |
1 |
27 |
15 |
4 |
35.0 |
+-- Cristaria_plicata **
+-----------+
| +-- Buldowskia_iwakawai **
|
+-----+ +----- Beringiana_fukuharai **
| | +-----+
| | | | +-- Beringiana_japonica **
| | | +--+
| +--+ +-- Beringiana_beringiana **
| |
| | +-------- Pletholophus_tenuis **
| +--+
| | +----- Sinanodonta_woodiana **
| +--+
--+ | +-- Sinanodonta_lauta **
| +--+
| +-- Sinanodonta_tumens **
|
| +-------- Tchangsinaia_piscicula
| +--------+
| | | +----- Nodularia_nipponensis
| | +--+
| | | +-- Nodularia_douglasiae
| | +--+
| | +-- Nodularia_breviconcha
+--+
| +-- Schistodesmus_lampreyanus
| +-----------+
| | +-- Acuticosta_ovata
| |
| | +-------- Lepidodesma_languilati
+--+ +--+
| | | +----- Lanceolaria_grayii[1]
| | +--+
| | | +-- Lanceolaria_lanceolata
+--+ +--+
| +-- Lanceolaria_grayii[2]
|
| +-------- Inversiunio_jokohamensis
| |
+--+ +----- Inversiunio_yanagawensis[3]
| |
+--+ +-- Inversiunio_yanagawensis[1]
| |
+--+-- Inversiunio_yanagawensis[2]
|
+-- Inversiunio_reinianus
Wu et al., 2023 [Publ 4339], fig. 5
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 9 |
9 |
7 |
3 |
10 |
2 |
1 |
34.0 |
+-- Lepidodesma_languilati
+-----------+
| +-- Lepidodesma_aligera
|
| +-------- Anemina_arcaeformis **
--+ +--+
| | | +----- Cristaria_plicata **
| | +--+
| | | +-- Sinanodonta_tumens **
| | +--+
+--+ +-- Sinanodonta_woodiana **
|
| +-- Platynaias **
| +-----+
| | +-- Platynaias_compressa **
+--+
| +----- Anodonta_anatina **
+--+
| +-- Pseudanodonta_complanata **
+--+
+-- Anodonta_cygnea **
Guerra et al., 2017 [Publ 3772], fig. 2a
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 8 |
8 |
6 |
3 |
17 |
2 |
1 |
33.0 |
+-- Aculamprotula_tortuosa
+-----------------+
| +-- Unio_pictorum
--+
| +----------------- Sinanodonta_woodiana **
| |
+--+ +-------------- Cristaria_plicata **
| |
+--+ +----------- Anodonta_anatina **
| |
+--+ +-- Platynaias_subviridis **
| +-----+
| | +-- Platynaias_compressa **
+--+
| +----- Pyganodon_grandis **
+--+
| +-- Utterbackia_peninsularis **
+--+
+-- Utterbackia_imbecillis **
Guerra et al., 2017 [Publ 3772], fig. 2b
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 8 |
8 |
6 |
3 |
17 |
2 |
1 |
35.0 |
+-------------------- Unio_pictorum
|
--+ +----------------- Aculamprotula_tortuosa
| |
+--+ +-- Cristaria_plicata **
| +-----------+
| | +-- Sinanodonta_woodiana **
+--+
| +----------- Anodonta_anatina **
| |
+--+ +-- Platynaias_subviridis **
| +-----+
| | +-- Platynaias_compressa **
+--+
| +----- Pyganodon_grandis **
+--+
| +-- Utterbackia_peninsularis **
+--+
+-- Utterbackia_imbecillis **
Sano et al., 2017 [Publ 3799], fig. 1
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 8 |
8 |
6 |
1 |
19 |
9 |
2 |
32.0 |
+-- Cristaria_plicata **
+-----------+
| +-- Buldowskia_iwakawai **
+--------+
| | +----------- Pletholophus_tenuis **
| | |
| +--+ +-- Sinanodonta_tumens **
| | +-----+
| | | +-- Sinanodonta_lauta **
| +--+
--+ | +----- Beringiana_fukuharai **
| +--+
| | +-- Beringiana_japonica **
| +--+
| +-- Beringiana_beringiana **
|
| +-------------------- Lanceolaria_grayii
| |
+--+ +-- Nodularia_nipponensis
| +--------------+
| | +-- Nodularia_douglasiae
+--+
| +-------------- Inversiunio_jokohamensis
+--+
| +----------- Inversiunio_yanagawensis[1]
+--+
| +-------- Inversiunio_yanagawensis[2]
+--+
| +----- Inversiunio_yanagawensis[3]
+--+
| +-- Inversiunio_yanagawensis[4]
+--+
+-- Inversiunio_reinianus
Vikhrev et al., 2017 [Publ 3810], fig. 4
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 8 |
8 |
6 |
1 |
2 |
0 |
0 |
34.0 |
+-------------------- Anemina_arcaeformis **
--+
| +----------------- Sinanodonta_lucida **
+--+
| +-------------- Sinanodonta_schrenkii **
+--+
| +----------- Sinanodonta_woodiana[1] **
+--+
| +-------- Sinanodonta_jourdyi **
+--+
| +----- Sinanodonta_elliptica **
+--+
| +-- Sinanodonta_lauta **
+--+
+-- Sinanodonta_woodiana[2] **
Bespalaya et al., 2018 [Publ 3811], fig. 3
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 8 |
8 |
6 |
1 |
2 |
0 |
0 |
34.0 |
+-------------------- Anemina_arcaeformis **
--+
| +----------------- Sinanodonta_lucida **
+--+
| +-------------- Sinanodonta_schrenkii **
+--+
| +----------- Sinanodonta_woodiana[2] **
+--+
| +-------- Sinanodonta_jourdyi **
+--+
| +----- Sinanodonta_elliptica **
+--+
| +-- Sinanodonta_woodiana[1] **
+--+
+-- Sinanodonta_lauta **
Huang et al., 2019 [Publ 3843], fig. 1
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 8 |
28 |
26 |
1 |
36 |
20 |
4 |
42.0 |
+-------------------------------- Anemina_arcaeformis **
|
| +----------------------------- Acuticosta_chinensis
| |
--+ | +-- Sinanodonta_angula **
| | +--------+
| | | +-- Sinanodonta_lucida **
| | +--------------+
| | | | +-------- Sinanodonta_jourdyi **
+--+ | +--+
| | | +----- Sinanodonta_woodiana[2] **
| | +--+
| | | +-- Sinanodonta_elliptica **
| | +--+
| | +-- Sinanodonta_woodiana[1] **
| |
+--+ +----------- Aculamprotula_tortuosa
| +-----------+
| | | +-------- Aculamprotula_scripta
| | +--+
| | | +----- Aculamprotula_coreana
| | +--+
| | | +-- Aculamprotula_fibrosa
| | +--+
| | +-- Aculamprotula_tientsinensis
+--+
| +----------- Cristaria_plicata **
| +--------+
| | | +-------- Lanceolaria_gladiola
| | +--+
| | | +----- Lanceolaria_grayii
| | +--+
| | | +-- Lanceolaria_lanceolata
+--+ +--+
| +-- Lanceolaria_triformis
|
| +----------------- Lepidodesma_languilati
| |
| | +-- Schistodesmus_lampreyanus
+--+ +-----------+
| | +-- Schistodesmus_spinosus
| |
+--+ +----- Nodularia_douglasiae[1]
| +-----+
| | | +-- Nodularia_douglasiae[2]
| | +--+
+--+ +-- Nodularia_nuxpersicae
|
| +-------- Cuneopsis_rufescens
+--+
| +----- Tchangsinaia_piscicula
+--+
| +-- Cuneopsis_heudei
+--+
+-- Cuneopsis_celtiformis
Wang et al., 2018 [Publ 3846], fig. 2
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 8 |
8 |
6 |
3 |
24 |
10 |
2 |
35.0 |
+----- Anodonta_anatina **
+-----+
| | +-- Pyganodon_grandis **
| +--+
+--------+ +-- Platynaias_compressa **
| |
| | +-------- Cristaria_plicata **
| | |
| +--+ +-- Anemina_euscaphys **
| | +--+
| | | +-- Anemina_arcaeformis **
| +--+
--+ | +-- Sinanodonta_woodiana **
| +--+
| +-- Sinanodonta_lucida **
|
| +-- Lanceolaria_grayii
| +--------------+
| | +-- Lanceolaria_lanceolata
| |
+--+ +-------------- Aculamprotula_tientsinensis
| |
| | +----- Nodularia_douglasiae
+--+ +-----+
| | | +-- Tchangsinaia_piscicula
| | +--+
+--+ +-- Cuneopsis_heudei
|
| +-------- Unio_tumidus
+--+
| +----- Unio_crassus
+--+
| +-- Unio_pictorum
+--+
+-- Unio_delphinus
Wang et al., 2018 [Publ 3865], fig. 2
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 8 |
8 |
6 |
3 |
24 |
10 |
3 |
35.0 |
+-------------- Aculamprotula_tientsinensis
|
+--+ +-- Tchangsinaia_piscicula
| | +--------+
| | | +-- Nodularia_douglasiae
| +--+
| | +-------- Unio_tumidus
| +--+
| | +----- Unio_crassus
| +--+
--+ | +-- Unio_pictorum
| +--+
| +-- Unio_delphinus
|
| +----- Acuticosta_chinensis
| +--------+
| | | +-- Lanceolaria_grayii
| | +--+
| | +-- Lanceolaria_lanceolata
+--+
| +----- Anodonta_anatina **
| +-----+
| | | +-- Pyganodon_grandis **
| | +--+
+--+ +-- Platynaias_compressa **
|
| +-------- Cristaria_plicata **
| |
+--+ +-- Anemina_euscaphys **
| +--+
| | +-- Anemina_arcaeformis **
+--+
| +-- Sinanodonta_woodiana **
+--+
+-- Sinanodonta_lucida **
Kondakov et al., 2018 [Publ 3914], fig. 5
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 8 |
8 |
6 |
1 |
0 |
0 |
0 |
34.0 |
+-------------------- Anemina_arcaeformis **
--+
| +----------------- Sinanodonta_lucida **
+--+
| +-------------- Sinanodonta_schrenkii **
+--+
| +----------- Sinanodonta_woodiana[2] **
+--+
| +-------- Sinanodonta_jourdyi **
+--+
| +----- Sinanodonta_elliptica **
+--+
| +-- Sinanodonta_woodiana[1] **
+--+
+-- Sinanodonta_lauta **
Huang et al., 2002 [Publ 2657], fig. 1
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 7 |
7 |
5 |
2 |
27 |
7 |
4 |
34.0 |
+-- Sinanodonta_woodiana **
+-----------+
| +-- Cristaria_plicata **
+--+
| | +----------- Utterbackiana_couperiana **
| +--+
| | +-------- Utterbackia_imbecillis **
| +--+
| | +----- Strophitus_radiatus **
| +--+
| | +-- Alasmidonta_triangulata **
--+ +--+
| +-- Lasmigona_costata **
|
| +----- Acuticosta_ovata
| +--+
| | | +-- Nodularia_douglasiae
| | +--+
| | +-- Tchangsinaia_piscicula
+--------+
| +-- Lanceolaria_grayii
| +--+
| | +-- Lanceolaria_lanceolata
+--+
| +-- Schistodesmus_lampreyanus
+--+
+-- Lepidodesma_languilati
Huang et al., 2002 [Publ 2657], fig. 2
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 7 |
14 |
5 |
2 |
27 |
7 |
4 |
34.0 |
+-------- Strophitus_radiatus **
|
+-------- Schistodesmus_lampreyanus
|
+-------- Lepidodesma_languilati
|
+-------- Acuticosta_ovata
|
| +-- Sinanodonta_woodiana **
+-----+
| +-- Cristaria_plicata **
--+
| +-- Lanceolaria_grayii
+-----+
| +-- Lanceolaria_lanceolata
|
| +-- Nodularia_douglasiae
+-----+
| +-- Tchangsinaia_piscicula
|
| +----- Utterbackia_imbecillis **
| |
+--+----- Utterbackiana_couperiana **
|
| +-- Alasmidonta_triangulata **
+--+
+-- Lasmigona_costata **
Hoeh et al., 1995 [Publ 3391], fig. 2
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 7 |
7 |
4 |
1 |
0 |
0 |
0 |
37.0 |
+----- Utterbackia_peggyae **
+--+
| | +-- Utterbackia_imbecillis **
| +--+
--+ +-- Utterbackia_peninsularis **
|
| +----- Pyganodon_lacustris **
| |
+--+----- Pyganodon_grandis **
|
| +-- Pyganodon_cataracta **
+--+
+-- Pyganodon_fragilis **
Inoue et al., 2014 [Publ 3635], fig. 3B
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 7 |
7 |
2 |
1 |
1 |
0 |
0 |
37.0 |
+-------- Pyganodon_grandis **
|
+-------- Pyganodon_cataracta **
|
--+-------- Pyganodon_fragilis **
|
+-------- Pyganodon **
|
| +----- Platynaias_subviridis **
+--+
| +-- Arcidens_confragosus **
+--+
+-- Arcidens_wheeleri **
Wu et al., 2016 [Publ 3711], fig. 1
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 7 |
7 |
5 |
3 |
14 |
5 |
1 |
32.0 |
+-- Unio_pictorum
+-----+
| +-- Nodularia_douglasiae
+-----------+
| | +----- Aculamprotula_tortuosa
| +--+
| | +-- Aculamprotula_coreana
--+ +--+
| +-- Aculamprotula_tientsinensis
|
| +----------------- Sinanodonta_lucida **
+--+
| +-------------- Anemina_arcaeformis **
+--+
| +----------- Cristaria_plicata **
+--+
| +-------- Anodonta_anatina **
+--+
| +----- Pyganodon_grandis **
+--+
| +-- Utterbackia_peninsularis **
+--+
+-- Utterbackia_imbecillis **
Feng et al., 2016 [Publ 3800], fig. 2
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 7 |
13 |
11 |
3 |
8 |
1 |
1 |
42.0 |
+-- Anodonta_anatina **
+--------------+
| +-- Utterbackia_peninsularis **
|
| +-- Sinosolenaia_carinata
| +-----+
--+ | +-- Sinosolenaia_oleivora
| +-----+
| | | +----- Quadrula_quadrula
| | +--+
| | | +-- Potamilus_leptodon
+--+ +--+
| +-- Lampsilis_ornata
|
| +----------- Tchangsinaia_piscicula
| |
+--+ +-------- Pyganodon_grandis **
| |
+--+ +-- Anemina_arcaeformis **
| +--+
| | +-- Anemina_euscaphys **
+--+
| +-- Sinanodonta_lucida **
+--+
+-- Sinanodonta_woodiana **
Wu et al., 2018 [Publ 4143], fig. 2
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 7 |
7 |
5 |
3 |
15 |
6 |
1 |
32.0 |
+-- Unio_pictorum
+--------+
| +-- Nodularia_douglasiae
+--------+
| | +-------- Aculamprotula_tortuosa
| +--+
| | +----- Aculamprotula_scripta
| +--+
--+ | +-- Aculamprotula_coreana
| +--+
| +-- Aculamprotula_tientsinensis
|
| +----------------- Sinanodonta_lucida **
+--+
| +-------------- Anemina_arcaeformis **
+--+
| +----------- Cristaria_plicata **
+--+
| +-------- Anodonta_anatina **
+--+
| +----- Pyganodon_grandis **
+--+
| +-- Utterbackia_peninsularis **
+--+
+-- Utterbackia_imbecillis **
Wu et al., 2020 [Publ 4147], fig. 3
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 7 |
7 |
5 |
3 |
23 |
10 |
1 |
34.0 |
+----- Cristaria_plicata **
+-----+
| | +-- Sinanodonta_lucida **
| +--+
+--------+ +-- Anemina_arcaeformis **
| |
| | +-------- Anodonta_anatina **
| +--+
| | +----- Pyganodon_grandis **
| +--+
| | +-- Utterbackia_peninsularis **
--+ +--+
| +-- Utterbackia_imbecillis **
|
| +-------- Aculamprotula_tortuosa
| +--------+
| | | +----- Aculamprotula_scripta
| | +--+
| | | +-- Aculamprotula_tientsinensis
+--+ +--+
| +-- Aculamprotula_coreana
|
| +-------------- Unio_pictorum
+--+
| +----------- Pseudocuneopsis_capitata
+--+
| +-------- Nodularia_douglasiae
+--+
| +----- Schistodesmus_lampreyanus
+--+
| +-- Cuneopsis_heudei
+--+
+-- Tchangsinaia_piscicula
Guerra et al., 2019 [Publ 4168], fig. 1
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 7 |
7 |
5 |
3 |
34 |
6 |
2 |
34.0 |
+-------------- Lanceolaria_lanceolata
+--+
| | +----------- Aculamprotula_tortuosa
| +--+
| | +-------- Schistodesmus_lampreyanus
| +--+
| | +----- Unio_tumidus
--+ +--+
| | +-- Unio_crassus
| +--+
| +-- Unio_delphinus
|
| +-------------- Sinanodonta_woodiana **
| |
+--+ +----------- Anodonta_anatina **
| |
+--+ +-- Platynaias_compressa **
| +-----+
| | +-- Platynaias_subviridis **
+--+
| +----- Pyganodon_grandis **
+--+
| +-- Utterbackia_peninsularis **
+--+
+-- Utterbackia_imbecillis **
Guerra et al., 2019 [Publ 4168], fig. 2
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 7 |
7 |
5 |
3 |
34 |
6 |
2 |
34.0 |
+-------- Schistodesmus_lampreyanus
+-----------+
| | +----- Unio_tumidus
| +--+
| | +-- Unio_crassus
--+ +--+
| +-- Unio_delphinus
|
| +----------------- Lanceolaria_lanceolata
| |
+--+ +-------------- Sinanodonta_woodiana **
| |
+--+ +----------- Anodonta_anatina **
| |
+--+ +-- Platynaias_compressa **
| +-----+
| | +-- Platynaias_subviridis **
+--+
| +----- Pyganodon_grandis **
+--+
| +-- Utterbackia_peninsularis **
+--+
+-- Utterbackia_imbecillis **
Pavan-Kumar et al., 2022 [Publ 4263], fig. S5
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 7 |
7 |
5 |
3 |
32 |
10 |
4 |
34.0 |
+-- Anodonta_anatina **
+--------+
| +-- Pyganodon_grandis **
+-----+
| | +-------- Cristaria_plicata **
| | |
| +--+ +-- Anemina_arcaeformis **
| | +--+
| | | +-- Anemina_euscaphys **
| +--+
| | +-- Sinanodonta_lucida **
--+ +--+
| +-- Sinanodonta_woodiana **
|
| +----- Acuticosta_chinensis
| +--------+
| | | +-- Lanceolaria_lanceolata
| | +--+
| | +-- Lanceolaria_gladiola
+--+
| +----- Lepidodesma_languilati
| +-----+
| | | +-- Aculamprotula_tientsinensis
| | +--+
+--+ +-- Aculamprotula_tortuosa
|
| +-------- Unio_pictorum
+--+
| +----- Schistodesmus_lampreyanus
+--+
| +-- Nodularia_douglasiae
+--+
+-- Tchangsinaia_piscicula
Dai et al., 2023 [Publ 4285], fig. 5
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 7 |
7 |
5 |
3 |
46 |
10 |
4 |
35.0 |
+-------------- Aculamprotula_tientsinensis
|
+--+ +----------- Unio_crassus
| | |
| +--+ +-- Pseudocuneopsis_capitata
| | +-----+
| | | +-- Nodularia_douglasiae
| +--+
| | +----- Schistodesmus_lampreyanus
| +--+
--+ | +-- Cuneopsis_celtiformis
| +--+
| +-- Tchangsinaia_piscicula
|
| +-- Acuticosta_chinensis
| +-----------+
| | +-- Lanceolaria_grayii
| |
+--+ +----- Cristaria_plicata **
| +-----+
| | | +-- Anemina_arcaeformis **
| | +--+
+--+ +-- Sinanodonta_woodiana **
|
| +-------- Anodonta_cygnea **
+--+
| +----- Platynaias_compressa **
+--+
| +-- Pyganodon_grandis **
+--+
+-- Utterbackia_imbecillis **
Smith et al., 2023 [Publ 4291], fig. 2
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 7 |
7 |
5 |
1 |
0 |
0 |
0 |
37.0 |
+-- Anodontoides_ferussacianus **
+--------+
| +-- Anodontoides_denigratus **
|
--+ +-- Strophitus_undulatus **
| +-----+
| | +-- Strophitus_howellsi **
+--+
| +----- Strophitus_pascagoulaensis **
+--+
| +-- Strophitus_radiatus **
+--+
+-- Strophitus_williamsi **
Dai et al., 2024 [Publ 4358], fig. 4
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 7 |
7 |
5 |
3 |
45 |
20 |
4 |
34.0 |
+----------------- Lanceolaria_gladiola
|
--+ +-------------- Acuticosta_chinensis
| |
| | +----- Cristaria_plicata **
+--+ +-----+
| | | +-- Anemina_arcaeformis **
| | +--+
+--+ +-- Sinanodonta_woodiana **
|
| +-------- Anodonta_anatina **
+--+
| +----- Platynaias_compressa **
+--+
| +-- Pyganodon_grandis **
+--+
+-- Utterbackia_imbecillis **
Nagel et al., 1998 [Publ 1946], fig. 7
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 6 |
6 |
4 |
2 |
0 |
0 |
0 |
33.0 |
+-------------- Pseudanodonta_complanata **
--+
| +----------- Anodonta_cygnea **
+--+
| +-------- Sinanodonta_woodiana **
+--+
| +----- Anodonta[2] **
+--+
| +-- Anodonta_anatina **
+--+
+-- Anodonta[1] **
Graf & Ó Foighil, 2000 [Publ 2648], fig. 2A
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 6 |
6 |
4 |
2 |
24 |
2 |
1 |
32.0 |
+----------- Anodonta_cygnea **
|
--+ +-- Alasmidonta_marginata **
| +-----+
| | +-- Lasmigona_costata **
+--+
| +----- Strophitus_undulatus **
+--+
| +-- Pyganodon_grandis **
+--+
+-- Pyganodon_fragilis **
Graf & Ó Foighil, 2000 [Publ 2648], fig. 2B
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 6 |
6 |
4 |
2 |
24 |
2 |
1 |
32.0 |
+----------- Anodonta_cygnea **
|
--+ +-- Alasmidonta_marginata **
| +-----+
| | +-- Lasmigona_costata **
+--+
| +----- Strophitus_undulatus **
+--+
| +-- Pyganodon_grandis **
+--+
+-- Pyganodon_fragilis **
Lydeard et al., 1996 [Publ 2656], fig. 3
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 6 |
6 |
4 |
1 |
21 |
0 |
0 |
34.0 |
+----- Utterbackia_imbecillis **
+--+
| | +-- Utterbackiana_couperiana **
| +--+
--+ +-- Utterbackiana **
|
| +----- Strophitus_radiatus **
+--+
| +-- Lasmigona_costata **
+--+
+-- Alasmidonta_triangulata **
Lydeard et al., 1996 [Publ 2656], fig. 4
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 6 |
6 |
4 |
2 |
22 |
0 |
0 |
32.5 |
+----- Utterbackia_imbecillis **
+--+
| | +-- Utterbackiana_couperiana **
| +--+
--+ +-- Anodonta **
|
| +----- Strophitus_radiatus **
+--+
| +-- Lasmigona_costata **
+--+
+-- Alasmidonta_triangulata **
Campbell et al., 2005 [Publ 2660], fig. 1
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 6 |
6 |
4 |
2 |
111 |
0 |
0 |
35.0 |
+----------- Anodonta_oregonensis **
|
--+ +-- Anodonta_cygnea **
| +-----+
| | +-- Alasmidonta_holstonia **
+--+
| +----- Pyganodon_grandis **
+--+
| +-- Lasmigona_costata **
+--+
+-- Pseudodontoideus_subvexus **
Campbell et al., 2005 [Publ 2660], fig. 2
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 6 |
6 |
3 |
2 |
109 |
0 |
0 |
35.0 |
+-------- Pyganodon_grandis **
|
| +-- Lasmigona_costata **
--+-----+
| +-- Pseudodontoideus_subvexus **
|
| +----- Alasmidonta_holstonia **
+--+
| +-- Anodonta_cygnea **
+--+
+-- Anodonta_oregonensis **
Boyer et al., 2011 [Publ 2718], fig. 3
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 6 |
6 |
2 |
1 |
29 |
0 |
0 |
33.0 |
+----- Alasmidonta_marginata **
|
+----- Pyganodon_grandis **
|
--+ +-- Lasmigona_complanata **
+--+
| +-- Lasmigona_costata **
|
| +-- Platynaias_compressa **
+--+
+-- Strophitus_undulatus **
Cyr et al., 2007 [Publ 2745], fig. 2B
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 6 |
6 |
4 |
2 |
0 |
0 |
0 |
34.0 |
+----------- Anodonta_nuttalliana **
|
--+ +-- Pyganodon_grandis **
| +-----+
| | +-- Pyganodon[1] **
+--+
| +----- Pyganodon[2] **
+--+
| +-- Pyganodon_cataracta **
+--+
+-- Pyganodon_fragilis **
Nagel et al., 1996 [Publ 3400], fig. 351
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 6 |
6 |
4 |
2 |
0 |
0 |
0 |
33.0 |
+-------------- Pseudanodonta_complanata **
--+
| +----------- Anodonta_cygnea **
+--+
| +-------- Sinanodonta_woodiana **
+--+
| +----- Anodonta[2] **
+--+
| +-- Anodonta_anatina **
+--+
+-- Anodonta[1] **
Huang et al., 2013 [Publ 3629], fig. 3
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 6 |
6 |
4 |
2 |
10 |
1 |
1 |
35.0 |
+-------------- Unio_pictorum
|
--+ +-- Sinanodonta_woodiana **
| +--------+
| | +-- Cristaria_plicata **
+--+
| +-------- Platynaias_compressa **
+--+
| +----- Pyganodon_grandis **
+--+
| +-- Utterbackia_imbecillis **
+--+
+-- Utterbackia_peninsularis **
Xue et al., 2016 [Publ 3659], fig. 1
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 6 |
6 |
4 |
2 |
10 |
6 |
2 |
33.0 |
+-- Cristaria_plicata **
+-----+
| +-- Platynaias_compressa **
|
+-----+ +-- Anemina_arcaeformis **
| | +--+
| | | +-- Anemina_euscaphys **
| +--+
| | +-- Sinanodonta_lucida **
--+ +--+
| +-- Sinanodonta_woodiana **
|
| +-- Lanceolaria_lanceolata
| +--------+
| | +-- Lanceolaria_grayii
+--+
| +-------- Aculamprotula_tortuosa
+--+
| +----- Schistodesmus_lampreyanus
+--+
| +-- Nodularia_douglasiae
+--+
+-- Unio_pictorum
Jadhav & Jamkhedkar, 2009 [Publ 3697], fig. 11
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 6 |
9 |
6 |
2 |
20 |
4 |
1 |
32.5 |
+-------------------- Nodularia_douglasiae
|
--+ +----------------- Anodonta_cygnea **
| |
+--+ +-------------- Unio_pictorum[2]
| |
+--+ +-- Unio_pictorum[1]
| +--------+
| | +-- Unio_caffer
+--+
| +-------- Anodonta **
| |
+--+ +----- Lasmigona_costata **
| |
+--+ +-- Strophitus_undulatus **
| |
+--+-- Alasmidonta_marginata **
|
+-- Pyganodon_grandis **
Zhou et al., 2016 [Publ 3710], fig. 1
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 6 |
6 |
4 |
2 |
14 |
3 |
2 |
33.0 |
+----- Lepidodesma_languilati
+-----------+
| | +-- Schistodesmus_lampreyanus
| +--+
--+ +-- Unio_pictorum
|
| +-------------- Sinanodonta_woodiana **
+--+
| +----------- Cristaria_plicata **
+--+
| +-------- Platynaias_compressa **
+--+
| +----- Pyganodon_grandis **
+--+
| +-- Utterbackia_imbecillis **
+--+
+-- Utterbackia_peninsularis **
Bolotov et al., 2016 [Publ 3732], fig. 2
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 6 |
6 |
4 |
1 |
2 |
0 |
0 |
36.0 |
+-------------- Sinanodonta_lucida **
--+
| +----------- Sinanodonta_schrenkii **
+--+
| +-------- Sinanodonta_woodiana[2] **
+--+
| +----- Sinanodonta_elliptica **
+--+
| +-- Sinanodonta_woodiana[1] **
+--+
+-- Sinanodonta_lauta **
Hewitt et al., 2018 [Publ 3758], fig. 5
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 6 |
6 |
3 |
1 |
3 |
0 |
0 |
35.0 |
+-------- Pyganodon_grandis **
|
| +-- Platynaias_subviridis **
--+-----+
| +-- Strophitus_undulatus **
|
| +----- Lasmigona_costata **
+--+
| +-- Arcidens_wheeleri **
+--+
+-- Lasmigona_complanata **
Gomes-dos-Santos et al., 2019 [Publ 4042], fig. 8
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 6 |
6 |
4 |
2 |
0 |
0 |
0 |
35.0 |
+----- Anodonta_anatina **
+--+
| | +-- Anodonta_exulcerata **
| +--+
--+ +-- Anodonta_cygnea **
|
| +----- Cristaria_radiata **
+--+
| +-- Anemina_arcaeformis **
+--+
+-- Sinanodonta_woodiana **
Lopes-Lima et al., 2020 [Publ 4080], fig. 7
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 6 |
6 |
4 |
1 |
2 |
0 |
0 |
34.0 |
+-- Sinanodonta_woodiana **
+--------+
| +-- Sinanodonta_tumens **
--+
| +-------- Beringiana_gosannensis **
+--+
| +----- Beringiana_fukuharai **
+--+
| +-- Beringiana_beringiana **
+--+
+-- Beringiana_japonica **
Riccardi et al., 2019 [Publ 4089], fig. 1
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 6 |
6 |
4 |
2 |
0 |
0 |
0 |
32.0 |
+-- Sinanodonta_woodiana **
+-----+
| +-- Anemina_arcaeformis **
|
--+ +-- Anodonta_anatina **
| +--+
| | +-- Pseudanodonta_complanata **
+--+
| +-- Anodonta_exulcerata **
+--+
+-- Anodonta_cygnea **
Tomilova et al., 2020 [Publ 4097], fig. 3
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 6 |
8 |
5 |
2 |
2 |
2 |
1 |
33.0 |
+-- Unio_tumidus
+--------+
| +-- Unio_pictorum
|
| +-- Sinanodonta_lauta **
--+--------+
| +-- Sinanodonta_woodiana **
|
| +-------- Anodonta_anatina **
+--+
| +----- Pseudanodonta_complanata **
+--+
| +-- Anodonta_cygnea **
+--+
+-- Anodonta_exulcerata **
Bogan et al., 2023 [Publ 4295], fig. 4
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 6 |
6 |
4 |
1 |
1 |
0 |
0 |
32.0 |
+-- Sinanodonta_jourdyi **
+--------+
| +-- Pletholophus_honglinhensis **
--+
| +-------- Cristaria_radiata **
+--+
| +----- Cristaria_clessini **
+--+
| +-- Cristaria_plicata **
+--+
+-- Cristaria_truncata **
Manendo et al., 2008 [Publ 2669], fig. 1
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 5 |
5 |
3 |
2 |
8 |
3 |
1 |
33.0 |
+----- Unio_crassus
+-----+
| | +-- Unio_pictorum
| +--+
| +-- Unio_tumidus
--+
| +-- Anodonta_cygnea **
| +-----+
| | +-- Anodonta_anatina **
+--+
| +----- Pseudanodonta_complanata **
+--+
| +-- Platynaias_subviridis **
+--+
+-- Pyganodon_grandis **
Cyr et al., 2007 [Publ 2745], fig. 2A
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 5 |
5 |
2 |
1 |
1 |
0 |
0 |
34.0 |
+-------- Pyganodon_grandis **
|
--+-------- Pyganodon[2] **
|
| +----- Pyganodon[1] **
+--+
| +-- Pyganodon_cataracta **
+--+
+-- Pyganodon_fragilis **
Cyr et al., 2007 [Publ 2745], fig. 2D
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 5 |
5 |
2 |
1 |
0 |
0 |
0 |
34.0 |
+-------- Pyganodon_grandis **
|
--+-------- Pyganodon[2] **
|
| +----- Pyganodon[1] **
+--+
| +-- Pyganodon_cataracta **
+--+
+-- Pyganodon_fragilis **
Soroka, 2010 [Publ 3404], fig. 3a
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 5 |
9 |
7 |
2 |
4 |
4 |
1 |
31.0 |
+-- Sinanodonta_woodiana **
+-----------+
| +-- Sinanodonta_lauta **
|
--+ +----- Anodonta_anatina **
| +-----+
| | | +-- Anodonta_cygnea **
| | +--+
+--+ +-- Pseudanodonta_complanata **
|
| +-------- Unio_tumidus
+--+
| +----- Unio_crassus
+--+
| +-- Unio_pictorum
+--+
+-- Unio_mancus
Soroka, 2010 [Publ 3404], fig. 3b
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 5 |
9 |
5 |
2 |
4 |
4 |
1 |
34.0 |
+-------------- Anodonta_anatina **
|
+-------------- Anodonta_cygnea **
|
--+-------------- Pseudanodonta_complanata **
|
| +-- Sinanodonta_woodiana **
| +--------+
| | +-- Sinanodonta_lauta **
+--+
| +-------- Unio_tumidus
+--+
| +----- Unio_crassus
+--+
| +-- Unio_pictorum
+--+
+-- Unio_mancus
Soroka, 2010 [Publ 3404], fig. 5a
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 5 |
5 |
3 |
2 |
3 |
3 |
1 |
33.0 |
+----- Unio_crassus
+-----+
| | +-- Unio_pictorum
| +--+
--+ +-- Unio_tumidus
|
| +-------- Anodonta_anatina **
| |
+--+ +-- Anodonta_cygnea **
| +--+
| | +-- Pseudanodonta_complanata **
+--+
| +-- Sinanodonta_woodiana **
+--+
+-- Sinanodonta_schrenkii **
Soroka, 2010 [Publ 3404], fig. 5b
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 5 |
8 |
5 |
2 |
3 |
3 |
1 |
33.0 |
+----------- Anodonta_anatina **
|
| +-- Anodonta_cygnea **
--+ +-----+
| | +-- Pseudanodonta_complanata **
| |
| | +-- Sinanodonta_woodiana **
+--+-----+
| +-- Sinanodonta_schrenkii **
|
| +----- Unio_tumidus
+--+
| +-- Unio_pictorum
+--+
+-- Unio_crassus
Soroka, 2010 [Publ 3404], fig. 9a
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 5 |
5 |
3 |
1 |
0 |
0 |
0 |
33.0 |
+----------- Buldowskia_flavotincta **
--+
| +-------- Beringiana_beringiana **
+--+
| +----- Sinanodonta_schrenkii **
+--+
| +-- Sinanodonta_lauta **
+--+
+-- Sinanodonta_woodiana **
Soroka, 2010 [Publ 3404], fig. 9c
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 5 |
5 |
3 |
1 |
0 |
0 |
0 |
34.0 |
+----------- Buldowskia_flavotincta **
--+
| +-------- Sinanodonta_woodiana[2] **
+--+
| +----- Sinanodonta_woodiana[1] **
+--+
| +-- Sinanodonta_lauta **
+--+
+-- Sinanodonta_schrenkii **
Pfeiffer & Graf, 2013 [Publ 3531], fig. 2
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 5 |
5 |
3 |
2 |
30 |
8 |
2 |
34.0 |
+-- Cristaria_plicata **
+-----+
| +-- Sinanodonta_woodiana **
+--------+
| | +----- Strophitus_undulatus **
| +--+
| | +-- Platynaias_compressa **
--+ +--+
| +-- Pyganodon_grandis **
|
| +-------------- Acuticosta_chinensis
| |
+--+ +-- Unio_pictorum
| +--------+
| | +-- Nodularia_douglasiae
+--+
| +-------- Aculamprotula_tortuosa
| |
+--+ +----- Aculamprotula_scripta
| |
+--+ +-- Aculamprotula_tientsinensis
| |
+--+-- Aculamprotula_fibrosa
|
+-- Aculamprotula_zonata
Plazzi et al., 2011 [Publ 3676], fig. 5
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 5 |
18 |
11 |
3 |
13 |
4 |
2 |
41.0 |
+----------------- Cristaria_plicata **
|
| +-------------- Lanceolaria_grayii
| |
| | +-- Pyganodon_grandis **
| +-----------+
--+ | +-- Pseudanodonta_complanata **
| |
| | +-- Anodonta_anatina **
| +-----------+
| | +-- Anodonta_cygnea **
| |
| | +----- Unio_tumidus
+--+--------+
| | +-- Unio_crassus
| +--+
| +-- Unio_pictorum
|
| +----- Alathyria_jacksoni
| |
| +-----+----- Pronodularia_japanensis
| | |
| | | +-- Sinohyriopsis_cumingii
| | +--+
+--+ +-- Sinohyriopsis_schlegelii
|
| +-------- Quadrula_quadrula
| |
+--+-------- Parvaspina_collina
|
| +----- Lampsilis_ornata
+--+
| +-- Epioblasma_rangiana
+--+
+-- Venustaconcha_ellipsiformis
Amaro et al., 2017 [Publ 3791], fig. 2
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 5 |
5 |
3 |
1 |
13 |
0 |
0 |
34.0 |
+-- Lasmigona_costata **
+-----+
| +-- Platynaias_compressa **
--+
| +----- Pyganodon_grandis **
+--+
| +-- Utterbackia_peninsularis **
+--+
+-- Utterbackia_imbecillis **
Lopes-Lima et al., 2020 [Publ 4080], fig. 8
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 5 |
5 |
3 |
1 |
2 |
0 |
0 |
33.0 |
+-- Sinanodonta_woodiana **
+-----+
| +-- Sinanodonta_lucida **
--+
| +----- Cristaria_clessini **
+--+
| +-- Cristaria_plicata **
+--+
+-- Cristaria_radiata **
Lyubas et al., 2022 [Publ 4255], fig. 7
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 5 |
5 |
3 |
1 |
26 |
23 |
1 |
32.0 |
+----------- Pseudanodonta_complanata **
+--------------------+
| | +-------- Anodonta_anatina **
| +--+
| | +----- Anodonta_cygnea **
| +--+
| | +-- Anodonta_exulcerata **
| +--+
--+ +-- Anodonta_seddoni **
|
| +----------------------------- Unio_tumidus
| |
| | +-- Unio_durieui
| | +-----------------------+
| | | +-- Unio_gibbus
+--+ |
| | +-------------- Unio_damascensis
| | |
| | +-------------- Unio_sesirmensis
| | +--------+
+--+ | | +----------- Unio_ionicus
| | | |
| | +--+ +-- Unio_bruguierianus
| | | +-----+
| | | | +-- Unio_tumidiformis
| | +--+
| | | +----- Unio_crassus[2]
+--+ +--+
| | +-- Unio_carneus
| +--+
| +-- Unio_crassus[1]
|
| +----- Unio_caffer
| +--------------+
| | | +-- Unio_terminalis
| | +--+
+--+ +-- Unio_delicatus
|
| +----------------- Unio_pictorum
| |
+--+ +-------------- Unio_eucirrus
| |
+--+ +-- Unio_elongatulus
| +--------+
| | +-- Unio_mancus
+--+
| +-------- Unio_ravoisieri
| |
+--+ +-- Unio_delphinus
| +--+
| | +-- Unio_foucauldianus
+--+
| +-- Unio_hueti
+--+
+-- Unio_tigridis
Lyubas et al., 2022 [Publ 4255], fig. S1
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 5 |
5 |
3 |
1 |
26 |
23 |
1 |
32.0 |
+----------- Anodonta_anatina **
+--------+
| | +-------- Pseudanodonta_complanata **
| +--+
| | +----- Anodonta_exulcerata **
| +--+
| | +-- Anodonta_seddoni **
| +--+
| +-- Anodonta_cygnea **
|
--+ +----------------- Unio_tumidus
| |
| | +-- Unio_durieui
| +--------------+
| | +-- Unio_gibbus
| |
| | +----------- Unio_sesirmensis
| | |
| | +--+ +-------- Unio_damascensis
+--+ | | |
| | | | +----- Unio_ionicus
| | +--+ |
| | | | +-- Unio_bruguierianus
| | | +--+
| | +--+ +-- Unio_tumidiformis
| | |
| | | +-- Unio_carneus
| | | |
| | +--+-- Unio_crassus[1]
+--+ |
| +-- Unio_crassus[2]
|
| +-------- Unio_elongatulus
| |
| +-------- Unio_mancus
| |
| | +-- Unio_pictorum
| +-----+
| | +-- Unio_eucirrus
| |
+-----+ +----- Unio_caffer
+--+
| | +-- Unio_terminalis
| +--+
| +-- Unio_delicatus
|
| +----- Unio_ravoisieri
| |
| | +-- Unio_delphinus
+--+--+
| +-- Unio_foucauldianus
|
| +-- Unio_hueti
+--+
+-- Unio_tigridis
Pavan-Kumar et al., 2022 [Publ 4263], fig. 4
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 5 |
5 |
3 |
3 |
31 |
10 |
4 |
33.0 |
+-- Lanceolaria_lanceolata
+-----------+
| +-- Lanceolaria_gladiola
--+
| +----------- Acuticosta_chinensis
| |
+--+ +-- Anodonta_anatina **
| +-----+
| | +-- Pyganodon_grandis **
+--+
| +----- Cristaria_plicata **
+--+
| +-- Sinanodonta_lucida **
+--+
+-- Anemina_euscaphys **
Bogan et al., 2023 [Publ 4295], fig. 5
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 5 |
5 |
3 |
1 |
1 |
0 |
0 |
31.0 |
+----------- Cristaria_truncata **
--+
| +-------- Sinanodonta_jourdyi **
+--+
| +----- Pletholophus_honglinhensis **
+--+
| +-- Pletholophus_tenuis **
+--+
+-- Pletholophus_reinianus **
Pfarrer et al., 2022 [Publ 4305], fig. 2-5
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 5 |
5 |
3 |
2 |
7 |
5 |
1 |
33.0 |
+----------- Unio_tumidus
+--+
| | +-------- Unio_crassus
| +--+
| | +----- Unio_elongatulus
| +--+
| | +-- Unio_mancus
--+ +--+
| +-- Unio_pictorum
|
| +-- Cristaria_plicata **
| +-----+
| | +-- Sinanodonta_woodiana **
+-----+
| +----- Anodonta_anatina **
+--+
| +-- Anodonta_exulcerata **
+--+
+-- Anodonta_cygnea **
Graf & Cummings, 2006 [Publ 1968], fig. 2
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 4 |
4 |
2 |
1 |
43 |
2 |
1 |
31.0 |
+-- Unio_pictorum
+--------+
| +-- Unio_caffer
--+
| +-------- Lasmigona_costata **
+--+
| +----- Alasmidonta_marginata **
+--+
| +-- Pyganodon_grandis **
+--+
+-- Strophitus_undulatus **
Hoeh et al., 2009 [Publ 2716], fig. 3
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 4 |
4 |
2 |
1 |
43 |
2 |
1 |
31.0 |
+-- Unio_pictorum
+--------+
| +-- Unio_caffer
--+
| +-------- Lasmigona_costata **
+--+
| +----- Alasmidonta_marginata **
+--+
| +-- Strophitus_undulatus **
+--+
+-- Pyganodon_grandis **
Hoeh et al., 2009 [Publ 2716], fig. 4
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 4 |
4 |
2 |
1 |
42 |
2 |
1 |
31.0 |
+-- Unio_pictorum
+--------+
| +-- Unio_caffer
--+
| +-------- Lasmigona_costata **
+--+
| +----- Alasmidonta_marginata **
+--+
| +-- Strophitus_undulatus **
+--+
+-- Pyganodon_grandis **
Hoeh et al., 2009 [Publ 2716], fig. 5
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 4 |
4 |
2 |
1 |
42 |
2 |
1 |
31.0 |
+-- Unio_pictorum
+--------+
| +-- Unio_caffer
--+
| +-------- Lasmigona_costata **
+--+
| +----- Alasmidonta_marginata **
+--+
| +-- Strophitus_undulatus **
+--+
+-- Pyganodon_grandis **
Rosenberg et al., 1997 [Publ 2720], fig. 2A
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 4 |
18 |
1 |
2 |
19 |
1 |
1 |
46.0 |
+----- Anodonta_anatina **
|
| +-- Gonidea_angulata
| |
| +-- Pyganodon_grandis **
| |
| +-- Utterbackia_imbecillis **
| |
| +-- Pyganodon_cataracta **
--+ |
| +-- Lampsilis_straminea
| |
| +-- Lampsilis_teres
| |
| +-- Megalonaias_nervosa
| |
| +-- Obliquaria_reflexa
| |
+--+-- Theliderma_cylindrica
|
+-- Quadrula_quadrula
|
+-- Uniomerus_tetralasmus
|
+-- Amblema_plicata
|
+-- Elliptio_complanata
|
+-- Fusconaia_cerina
|
+-- Plectomerus_dombeyanus
|
+-- Pleurobema_cordatum
|
+-- Unio_pictorum
Rosenberg et al., 1997 [Publ 2720], fig. 2B
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 4 |
18 |
1 |
2 |
19 |
2 |
1 |
46.5 |
+----- Anodonta **
|
| +-- Gonidea_angulata
| |
| +-- Pyganodon_grandis **
| |
| +-- Utterbackia_imbecillis **
| |
| +-- Pyganodon_cataracta **
--+ |
| +-- Lampsilis_straminea
| |
| +-- Lampsilis_teres
| |
| +-- Megalonaias_nervosa
| |
| +-- Obliquaria_reflexa
| |
+--+-- Theliderma_cylindrica
|
+-- Quadrula_quadrula
|
+-- Uniomerus_tetralasmus
|
+-- Amblema_plicata
|
+-- Elliptio_complanata
|
+-- Fusconaia_cerina
|
+-- Plectomerus_dombeyanus
|
+-- Aculamprotula_tortuosa
|
+-- Unio_pictorum
Soroka, 2010 [Publ 3404], fig. 2
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 4 |
7 |
4 |
2 |
3 |
3 |
1 |
29.0 |
+----------- Sinanodonta_woodiana **
|
| +-- Unio_tumidus
--+ |
| +-----+-- Unio_pictorum
| | |
+--+ +-- Unio_crassus
|
| +----- Anodonta_cygnea **
+--+
| +-- Anodonta_anatina **
+--+
+-- Pseudanodonta_complanata **
Soroka, 2010 [Publ 3404], fig. 4
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 4 |
7 |
5 |
2 |
3 |
3 |
1 |
33.0 |
+----- Sinanodonta_woodiana **
+-----+
| | +-- Anodonta_cygnea **
| +--+
--+ +-- Pseudanodonta_complanata **
|
| +-------- Anodonta_anatina **
+--+
| +----- Unio_tumidus
+--+
| +-- Unio_pictorum
+--+
+-- Unio_crassus
Soroka, 2010 [Publ 3404], fig. 9b
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 4 |
4 |
2 |
1 |
0 |
0 |
0 |
34.0 |
+-------- Buldowskia_flavotincta **
--+
| +----- Sinanodonta_schrenkii **
+--+
| +-- Sinanodonta_lauta **
+--+
+-- Sinanodonta_woodiana **
Klishko et al., 2014 [Publ 3666], fig. 8
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 4 |
4 |
2 |
1 |
0 |
0 |
0 |
30.0 |
+-- Cristaria_plicata **
+--+
| +-- Cristaria_radiata **
--+
| +-- Beringiana_beringiana **
+--+
+-- Sinanodonta_woodiana **
Zieritz et al., 2012 [Publ 3673], fig. 4
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 4 |
8 |
4 |
2 |
4 |
3 |
1 |
32.0 |
+-- Pseudanodonta_complanata **
|
+--------+-- Anodonta_anatina **
| |
| +-- Anodonta_cygnea **
--+
| +-------- Sinanodonta_woodiana **
| |
+--+-------- Margaritifera_margaritifera
|
| +----- Unio_crassus
+--+
| +-- Unio_tumidus
+--+
+-- Unio_pictorum
Mezhzherin et al., 2014 [Publ 3705], fig. 2
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 4 |
4 |
2 |
2 |
0 |
0 |
0 |
30.0 |
+-------- Sinanodonta_woodiana **
--+
| +----- Pseudanodonta_complanata **
+--+
| +-- Anodonta_anatina **
+--+
+-- Anodonta_cygnea **
Mezhzherin et al., 2014 [Publ 3705], fig. 4
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 4 |
4 |
2 |
2 |
0 |
0 |
0 |
30.0 |
+-------- Sinanodonta_woodiana **
--+
| +----- Anodonta_anatina **
+--+
| +-- Anodonta_cygnea **
+--+
+-- Pseudanodonta_complanata **
Klishko et al., 2016 [Publ 3733], fig. 1
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 4 |
4 |
2 |
1 |
0 |
0 |
0 |
30.0 |
+-- Cristaria_plicata **
+--+
| +-- Cristaria_radiata **
--+
| +-- Sinanodonta_woodiana **
+--+
+-- Beringiana_beringiana **
Sayenko et al., 2017 [Publ 3760], fig. 3
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 4 |
4 |
2 |
1 |
0 |
0 |
0 |
32.0 |
+-------- Beringiana_beringiana **
--+
| +----- Sinanodonta_schrenkii **
+--+
| +-- Sinanodonta_woodiana **
+--+
+-- Sinanodonta_lauta **
Amaro et al., 2017 [Publ 3791], fig. 2-M
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 4 |
4 |
1 |
2 |
6 |
0 |
0 |
32.0 |
+----- Pyganodon_grandis **
|
--+ +-- Lasmigona_costata **
| |
+--+-- Sinanodonta_woodiana **
|
+-- Utterbackia_peninsularis **
Amaro et al., 2017 [Publ 3791], fig. 3A
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 4 |
4 |
2 |
1 |
8 |
0 |
0 |
34.0 |
+-------- Platynaias_compressa **
--+
| +----- Pyganodon_grandis **
+--+
| +-- Utterbackia_imbecillis **
+--+
+-- Utterbackia_peninsularis **
Amaro et al., 2017 [Publ 3791], fig. 3B
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 4 |
4 |
2 |
1 |
8 |
0 |
0 |
34.0 |
+-------- Platynaias_compressa **
--+
| +----- Pyganodon_grandis **
+--+
| +-- Utterbackia_imbecillis **
+--+
+-- Utterbackia_peninsularis **
Froufe et al., 2017 [Publ 3802], fig. 2
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 4 |
4 |
2 |
2 |
0 |
0 |
0 |
31.0 |
+-------- Cristaria_plicata **
--+
| +----- Anodonta_anatina **
+--+
| +-- Anodonta_cygnea **
+--+
+-- Anodonta_exulcerata **
Froufe et al., 2017 [Publ 3802], fig. 7
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 4 |
4 |
2 |
2 |
6 |
3 |
1 |
32.0 |
+----- Unio_mancus
+-----+
| | +-- Unio_elongatulus
| +--+
--+ +-- Unio_pictorum
|
| +-------- Sinanodonta_woodiana **
+--+
| +----- Anodonta_anatina **
+--+
| +-- Anodonta_exulcerata **
+--+
+-- Anodonta_cygnea **
Huang et al., 2018 [Publ 3828], fig. 6-M
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 4 |
4 |
2 |
3 |
13 |
4 |
1 |
31.0 |
+-------- Sinanodonta_woodiana **
+--+
| | +----- Anodonta_anatina **
| +--+
| | +-- Pyganodon_grandis **
--+ +--+
| +-- Utterbackia_peninsularis **
|
| +-------- Aculamprotula_tortuosa
+--+
| +----- Unio_tumidus
+--+
| +-- Unio_crassus
+--+
+-- Unio_delphinus
Huang et al., 2019 [Publ 3843], fig. 3-M
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 4 |
4 |
2 |
3 |
19 |
6 |
2 |
31.0 |
+-------- Sinanodonta_woodiana **
+--------+
| | +----- Anodonta_anatina **
| +--+
| | +-- Pyganodon_grandis **
--+ +--+
| +-- Utterbackia_peninsularis **
|
| +-------------- Lanceolaria_lanceolata
+--+
| +----------- Schistodesmus_lampreyanus
+--+
| +-------- Aculamprotula_tortuosa
+--+
| +----- Unio_tumidus
+--+
| +-- Unio_crassus
+--+
+-- Unio_delphinus
Burzynski & Soroka, 2018 [Publ 3862], fig. 3
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 4 |
4 |
2 |
3 |
0 |
0 |
0 |
31.0 |
+-------- Sinanodonta_woodiana **
--+
| +----- Anodonta_anatina **
+--+
| +-- Utterbackia_peninsularis **
+--+
+-- Pyganodon_grandis **
Burzynski & Soroka, 2018 [Publ 3862], fig. 5a
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 4 |
4 |
1 |
3 |
16 |
6 |
2 |
32.0 |
+----- Sinanodonta_woodiana **
|
+-----------+----- Anodonta_anatina **
| |
| | +-- Utterbackia_peninsularis **
--+ +--+
| +-- Pyganodon_grandis **
|
| +-------------- Lanceolaria_lanceolata
+--+
| +----------- Aculamprotula_tortuosa
+--+
| +-------- Unio_tumidus
+--+
| +----- Unio_crassus
+--+
| +-- Unio_pictorum
+--+
+-- Unio_delphinus
Burzynski & Soroka, 2018 [Publ 3862], fig. 5b
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 4 |
4 |
2 |
3 |
6 |
6 |
2 |
31.0 |
+----------------- Lanceolaria_lanceolata
|
| +-------- Sinanodonta_woodiana **
--+ +-----+
| | | +----- Anodonta_anatina **
| | +--+
| | | +-- Utterbackia_peninsularis **
+--+ +--+
| +-- Pyganodon_grandis **
|
| +----------- Aculamprotula_tortuosa
+--+
| +-------- Unio_tumidus
+--+
| +----- Unio_crassus
+--+
| +-- Unio_pictorum
+--+
+-- Unio_delphinus
Froufe et al., 2019 [Publ 3936], fig. 3-M
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 4 |
4 |
2 |
3 |
15 |
3 |
1 |
31.0 |
+----- Unio_tumidus
+-----+
| | +-- Unio_crassus
| +--+
--+ +-- Unio_delphinus
|
| +-------- Sinanodonta_woodiana **
+--+
| +----- Anodonta_anatina **
+--+
| +-- Utterbackia_peninsularis **
+--+
+-- Pyganodon_grandis **
Froufe et al., 2019 [Publ 3936], fig. 4
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 4 |
4 |
2 |
3 |
15 |
3 |
1 |
31.0 |
+----- Unio_tumidus
+-----+
| | +-- Unio_crassus
| +--+
--+ +-- Unio_delphinus
|
| +-------- Sinanodonta_woodiana **
+--+
| +----- Anodonta_anatina **
+--+
| +-- Utterbackia_peninsularis **
+--+
+-- Pyganodon_grandis **
Pfeiffer et al., 2019 [Publ 3937], fig. 4-5
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 4 |
4 |
2 |
3 |
53 |
4 |
2 |
32.0 |
+----------- Lanceolaria_lanceolata
|
| +----- Unio_gibbus
--+ +--+
| | | +-- Nodularia_douglasiae
| | +--+
| | +-- Tchangsinaia_piscicula
+--+
| +-- Cristaria_plicata **
| +--+
| | +-- Sinanodonta_woodiana **
+--+
| +-- Anodonta_cygnea **
+--+
+-- Pyganodon_grandis **
Pfeiffer et al., 2019 [Publ 3937], fig. 6
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 4 |
4 |
2 |
3 |
53 |
4 |
2 |
32.0 |
+-------- Lanceolaria_lanceolata
+--+
| | +----- Unio_gibbus
| +--+
| | +-- Tchangsinaia_piscicula
| +--+
--+ +-- Nodularia_douglasiae
|
| +-- Cristaria_plicata **
| +--+
| | +-- Sinanodonta_woodiana **
+-----+
| +-- Anodonta_cygnea **
+--+
+-- Pyganodon_grandis **
Lopes-Lima et al., 2020 [Publ 4080], fig. 9
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 4 |
4 |
2 |
1 |
2 |
0 |
0 |
30.0 |
+-------- Cristaria_plicata **
--+
| +----- Sinanodonta_schrenkii **
+--+
| +-- Pletholophus_tenuis **
+--+
+-- Pletholophus_reinianus **
Guerra et al., 2019 [Publ 4168], fig. 1-M
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 4 |
4 |
2 |
3 |
18 |
6 |
2 |
31.0 |
+-------- Sinanodonta_woodiana **
+--------+
| | +----- Anodonta_anatina **
| +--+
| | +-- Utterbackia_peninsularis **
--+ +--+
| +-- Pyganodon_grandis **
|
| +-------------- Lanceolaria_lanceolata
+--+
| +----------- Schistodesmus_lampreyanus
+--+
| +-------- Aculamprotula_tortuosa
+--+
| +----- Unio_tumidus
+--+
| +-- Unio_crassus
+--+
+-- Unio_delphinus
Guerra et al., 2019 [Publ 4168], fig. 2-M
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 4 |
4 |
2 |
3 |
18 |
6 |
2 |
33.0 |
+-- Utterbackia_peninsularis **
+--+
| +-- Pyganodon_grandis **
+-----------+
| | +-- Anodonta_anatina **
| +--+
--+ +-- Sinanodonta_woodiana **
|
| +-------------- Lanceolaria_lanceolata
+--+
| +----------- Aculamprotula_tortuosa
+--+
| +-------- Schistodesmus_lampreyanus
+--+
| +----- Unio_tumidus
+--+
| +-- Unio_crassus
+--+
+-- Unio_delphinus
Lopes-Lima et al., 2021 [Publ 4217], fig. 11
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 4 |
4 |
2 |
1 |
0 |
0 |
0 |
33.0 |
+-------- Anodonta_anatina **
--+
| +----- Anodonta_exulcerata **
+--+
| +-- Anodonta_seddoni **
+--+
+-- Anodonta_cygnea **
Zieritz et al., 2021 [Publ 4231], fig. 3-M, 4
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 4 |
4 |
2 |
3 |
19 |
3 |
1 |
31.0 |
+----- Unio_tumidus
+-----+
| | +-- Unio_delphinus
| +--+
--+ +-- Unio_crassus
|
| +-------- Sinanodonta_woodiana **
+--+
| +----- Anodonta_anatina **
+--+
| +-- Utterbackia_peninsularis **
+--+
+-- Pyganodon_grandis **
Wu et al., 2022 [Publ 4238], fig. 2e
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 4 |
4 |
2 |
3 |
13 |
13 |
2 |
31.0 |
+-------- Sinanodonta_woodiana **
+-----------------+
| | +----- Anodonta_anatina **
| +--+
| | +-- Utterbackia_peninsularis **
--+ +--+
| +-- Pyganodon_grandis **
|
| +----------------------- Lanceolaria_lanceolata
| |
+--+ +-------------------- Aculamprotula_tortuosa
| |
| | +----- Unio_tumidus
+--+ +-----------+
| | | +-- Unio_crassus
| | +--+
| | +-- Unio_delphinus
+--+
| +-- Pseudocuneopsis_sichuanensis
| +-----------+
| | +-- Pseudocuneopsis_capitata
+--+
| +----------- Nodularia_douglasiae
| |
+--+ +-------- Tchangsinaia_piscicula
| |
+--+ +-- Cuneopsis_celtiformis
| +--+
| | +-- Cuneopsis_heudei
+--+
| +-- Schistodesmus_lampreyanus
+--+
+-- Schistodesmus_spinosus
Wu et al., 2023 [Publ 4296], fig. 4
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 4 |
4 |
2 |
2 |
19 |
17 |
3 |
31.0 |
+----- Lanceolaria_gladiola
+-----------------------+
| | +-- Lanceolaria_lanceolata
| +--+
| +-- Lanceolaria_grayii
|
--+ +-------- Pseudanodonta_complanata **
| +-----------------+
| | | +----- Lasmigona_costata **
| | +--+
| | | +-- Strophitus_undulatus **
| | +--+
+--+ +-- Pyganodon_grandis **
|
| +----------------------- Unio_tumidus
| |
| | +-- Aculamprotula_scripta
+--+ +-----------------+
| | +-- Aculamprotula_tientsinensis
| |
+--+ +-- Cuneopsis_heudei
| +--------------+
| | +-- Cuneopsis_rufescens
| |
+--+ +----- Nodularia_douglasiae
| +--------+
| | | +-- Schistodesmus_lampreyanus
| | +--+
+--+ +-- Schistodesmus_spinosus
|
| +----------- Tchangsinaia_piscicula
+--+
| +-------- Inversiunio_yanagawensis
+--+
| +----- Pseudocuneopsis_capitata
+--+
| +-- Pseudocuneopsis_yangshuoensis
+--+
+-- Pseudocuneopsis_sichuanensis
Liu et al., 2023 [Publ 4310], fig. 4
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 4 |
4 |
2 |
2 |
20 |
18 |
3 |
31.0 |
+-------------------------------- Lanceolaria_gladiola
|
| +-------- Pseudanodonta_complanata **
| +--------------------+
--+ | | +----- Lasmigona_costata **
| | +--+
| | | +-- Strophitus_undulatus **
| | +--+
+--+ +-- Pyganodon_grandis **
|
| +-------------------------- Unio_tumidus
| |
| | +-- Aculamprotula_scripta
+--+ +--------------------+
| | +-- Aculamprotula_tientsinensis
| |
+--+ +-- Cuneopsis_heudei
| +-----------------+
| | +-- Cuneopsis_rufescens
| |
+--+ +----- Nodularia_douglasiae
| +-----------+
| | | +-- Schistodesmus_lampreyanus
| | +--+
+--+ +-- Schistodesmus_spinosus
|
| +-------------- Tchangsinaia_piscicula
+--+
| +----------- Inversiunio_yanagawensis
+--+
| +-------- Pseudocuneopsis_capitata
+--+
| +----- Pseudocuneopsis_sichuanensis
+--+
| +-- Pseudocuneopsis_wuana
+--+
+-- Pseudocuneopsis_yangshuoensis
Liu et al., 2023 [Publ 4310], fig. 5
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 4 |
4 |
2 |
2 |
20 |
18 |
3 |
31.0 |
+----------------------- Lanceolaria_grayii
|
+----------------------- Lanceolaria_gladiola
|
+----------------------- Lanceolaria_lanceolata
|
--+ +-------- Lasmigona_costata **
| +-----------+
| | | +----- Pseudanodonta_complanata **
| | +--+
| | | +-- Pyganodon_grandis **
| | +--+
| | +-- Strophitus_undulatus **
+--+
| +----------------- Unio_tumidus
| |
| | +-- Aculamprotula_tientsinensis
| +--------------+
| | +-- Aculamprotula_scripta
| |
+--+ +-- Cuneopsis_rufescens
| +-----------+
| | +-- Cuneopsis_heudei
| |
| | +----- Nodularia_douglasiae
| +--------+
+--+ | +-- Schistodesmus_lampreyanus
| +--+
| +-- Schistodesmus_spinosus
|
| +----------- Tchangsinaia_piscicula
| |
+--+----------- Inversiunio_yanagawensis
|
| +-------- Pseudocuneopsis_capitata
+--+
| +----- Pseudocuneopsis_sichuanensis
+--+
| +-- Pseudocuneopsis_wuana
+--+
+-- Pseudocuneopsis_yangshuoensis
Bulakhova et al., 2023 [Publ 4325], fig. 5
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 4 |
4 |
2 |
1 |
0 |
0 |
0 |
33.0 |
+-- Beringiana_beringiana **
+--+
| +-- Beringiana_japonica **
--+
| +-- Beringiana_gosannensis **
+--+
+-- Beringiana_fukuharai **
Nagel et al., 1998 [Publ 1946], fig. 1
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 3 |
3 |
1 |
1 |
6 |
4 |
1 |
30.0 |
+----- Pseudanodonta_complanata **
+-----+
| | +-- Anodonta_anatina **
| +--+
--+ +-- Anodonta_cygnea **
|
| +-------- Unio_crassus
+--+
| +----- Unio_tumidus
+--+
| +-- Unio_pictorum
+--+
+-- Unio_mancus
Bogan & Hoeh, 2000 [Publ 2650], fig. 1
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 3 |
3 |
1 |
2 |
31 |
2 |
1 |
32.0 |
+----- Anodonta_cygnea **
--+
| +-- Strophitus_undulatus **
+--+
+-- Pyganodon_grandis **
Hoeh et al., 2001 [Publ 2652], fig. 14.2
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 3 |
3 |
1 |
2 |
28 |
2 |
1 |
32.0 |
+----- Pyganodon_grandis **
--+
| +-- Anodonta_cygnea **
+--+
+-- Strophitus_undulatus **
Hoeh et al., 2001 [Publ 2652], fig. 14.4
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 3 |
3 |
1 |
2 |
28 |
2 |
1 |
32.0 |
+-------- Unio_pictorum
|
--+-------- Unio_tumidus
|
| +----- Anodonta_cygnea **
+--+
| +-- Strophitus_undulatus **
+--+
+-- Pyganodon_grandis **
Hoeh et al., 2002 [Publ 2653], fig. 2
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 3 |
3 |
1 |
2 |
28 |
2 |
1 |
32.0 |
+----------- Unio_pictorum
--+
| +-------- Unio_tumidus
+--+
| +----- Anodonta_cygnea **
+--+
| +-- Pyganodon_grandis **
+--+
+-- Strophitus_undulatus **
Roe & Hoeh, 2003 [Publ 2655], fig. 4.2
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 3 |
3 |
1 |
2 |
28 |
2 |
1 |
32.0 |
+-- Unio_pictorum
+-----+
| +-- Unio_tumidus
--+
| +----- Pyganodon_grandis **
+--+
| +-- Anodonta_cygnea **
+--+
+-- Strophitus_undulatus **
Roe & Hoeh, 2003 [Publ 2655], fig. 4.3
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 3 |
3 |
1 |
2 |
28 |
2 |
1 |
32.0 |
+-- Unio_pictorum
+-----+
| +-- Unio_tumidus
--+
| +----- Anodonta_cygnea **
+--+
| +-- Pyganodon_grandis **
+--+
+-- Strophitus_undulatus **
Walker et al., 2006 [Publ 2666], fig. 2
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 3 |
3 |
1 |
2 |
12 |
0 |
0 |
31.5 |
+----- Anodonta **
--+
| +-- Pyganodon_fragilis **
+--+
+-- Pyganodon_grandis **
Walker et al., 2006 [Publ 2666], fig. 3
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 3 |
3 |
1 |
2 |
12 |
0 |
0 |
31.5 |
+----- Anodonta **
--+
| +-- Pyganodon_fragilis **
+--+
+-- Pyganodon_grandis **
Walker et al., 2006 [Publ 2666], fig. 4
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 3 |
3 |
1 |
2 |
12 |
0 |
0 |
31.5 |
+----- Anodonta **
--+
| +-- Pyganodon_fragilis **
+--+
+-- Pyganodon_grandis **
Källersjö et al., 2005 [Publ 2684], fig. 1A
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 3 |
3 |
0 |
1 |
4 |
3 |
1 |
30.0 |
+-- Anodonta_anatina **
|
--+-- Pseudanodonta_complanata **
|
+-- Anodonta_cygnea **
Källersjö et al., 2005 [Publ 2684], fig. 1B-F
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 3 |
3 |
1 |
1 |
4 |
3 |
1 |
30.0 |
+----- Anodonta_anatina **
--+
| +-- Pseudanodonta_complanata **
+--+
+-- Anodonta_cygnea **
Rosenberg et al., 1994 [Publ 2719], fig. 3
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 3 |
10 |
0 |
1 |
16 |
1 |
1 |
40.0 |
+-- Pyganodon_cataracta **
|
+-- Pyganodon_grandis **
|
+-- Utterbackia_imbecillis **
|
+-- Lampsilis_teres
|
+-- Lampsilis_straminea
--+
+-- Megalonaias_nervosa
|
+-- Obliquaria_reflexa
|
+-- Theliderma_cylindrica
|
+-- Quadrula_quadrula
|
+-- Uniomerus_tetralasmus
Gustafson & Iwamoto, 2005 [Publ 2741], fig. 2
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 3 |
3 |
1 |
2 |
2 |
0 |
0 |
31.0 |
+----- Beringiana_beringiana **
--+
| +-- Anodonta_oregonensis **
+--+
+-- Anodonta_nuttalliana **
Moyer & Díaz-Ferguson, 2012 [Publ 3375], fig. 3
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 3 |
3 |
1 |
1 |
4 |
0 |
0 |
32.0 |
+----- Lasmigona_costata **
--+
| +-- Platynaias_compressa **
+--+
+-- Platynaias_subviridis **
Soroka, 2010 [Publ 3404], fig. 3a-M
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 3 |
6 |
4 |
2 |
3 |
3 |
1 |
32.0 |
+----------- Anodonta_anatina **
|
--+ +-- Sinanodonta_woodiana **
| +-----+
| | +-- Sinanodonta_lauta **
+--+
| +----- Unio_crassus
+--+
| +-- Unio_tumidus
+--+
+-- Unio_pictorum
Soroka, 2010 [Publ 3404], fig. 3b-M
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 3 |
6 |
4 |
2 |
3 |
3 |
1 |
32.0 |
+----------- Anodonta_anatina **
|
--+ +-- Sinanodonta_lauta **
| +-----+
| | +-- Sinanodonta_woodiana **
+--+
| +----- Unio_tumidus
+--+
| +-- Unio_crassus
+--+
+-- Unio_pictorum
Soroka, 2010 [Publ 3404], fig. 8a
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 3 |
3 |
1 |
2 |
3 |
3 |
1 |
31.0 |
+----- Unio_tumidus
+--+
| | +-- Unio_pictorum
| +--+
--+ +-- Unio_crassus
|
| +----- Sinanodonta_woodiana **
+--+
| +-- Anodonta_anatina **
+--+
+-- Anodonta_cygnea **
Soroka, 2010 [Publ 3404], fig. 8b
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 3 |
6 |
4 |
2 |
3 |
3 |
1 |
30.0 |
+----------- Sinanodonta_woodiana **
|
--+ +-- Anodonta_anatina **
| +-----+
| | +-- Anodonta_cygnea **
+--+
| +----- Unio_tumidus
+--+
| +-- Unio_pictorum
+--+
+-- Unio_crassus
Ouyang et al., 2011 [Publ 3408], fig. 1
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 3 |
3 |
1 |
1 |
10 |
6 |
3 |
31.0 |
+----- Anemina_arcaeformis **
+-----+
| | +-- Sinanodonta_elliptica **
| +--+
| +-- Sinanodonta_woodiana **
--+
| +-- Nodularia_douglasiae
| +-----+
| | +-- Cuneopsis_heudei
+--+
| +----- Lanceolaria_grayii
| |
+--+----- Acuticosta_chinensis
|
| +-- Lanceolaria_gladiola
+--+
+-- Aculamprotula_tortuosa
Prié & Puillandre, 2014 [Publ 3529], fig. 2
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 3 |
3 |
1 |
1 |
14 |
11 |
1 |
30.0 |
+----- Anodonta_anatina **
+--------------+
| | +-- Pseudanodonta_complanata **
| +--+
| +-- Anodonta_cygnea **
|
--+ +----- Unio_tumidus
| +-----------+
| | | +-- Unio_gibbus
| | +--+
| | +-- Unio_durieui
+--+
| +----- Unio_tumidiformis
| +--------+
| | | +-- Unio_crassus
| | +--+
+--+ +-- Unio_courtillieri
|
| +----------- Unio_delphinus
+--+
| +-------- Unio_elongatulus
+--+
| +----- Unio_mancus
+--+
| +-- Unio_pictorum
+--+
+-- Unio_ravoisieri
Hinzmann et al., 2013 [Publ 3541], fig. 5
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 3 |
3 |
1 |
1 |
0 |
0 |
0 |
30.0 |
+----- Pseudanodonta_complanata **
--+
| +-- Anodonta_anatina **
+--+
+-- Anodonta_cygnea **
Reis et al., 2013 [Publ 3580], fig. 4
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 3 |
3 |
1 |
1 |
11 |
6 |
1 |
30.0 |
+----- Anodonta_cygnea **
+--------+
| | +-- Anodonta_anatina **
| +--+
--+ +-- Pseudanodonta_complanata **
|
| +----------- Unio_tumidus
| |
+--+ +-- Unio_tumidiformis
| +-----+
| | +-- Unio_crassus
+--+
| +----- Unio_delphinus
+--+
| +-- Unio_mancus
+--+
+-- Unio_pictorum
Reis et al., 2013 [Publ 3580], fig. 5
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 3 |
3 |
0 |
1 |
13 |
8 |
1 |
30.0 |
+-- Anodonta_anatina **
|
+--------------+-- Anodonta_cygnea **
| |
| +-- Pseudanodonta_complanata **
--+
| +-------------- Unio_tumidus
| |
+--+ +-- Unio_tumidiformis
| +--------+
| | +-- Unio_crassus
+--+
| +-------- Unio
| |
+--+ +-- Unio_delphinus
| +--+
| | +-- Unio_mancus
+--+
| +-- Unio_pictorum
+--+
+-- Unio_caffer
Froufe et al., 2014 [Publ 3592], fig. 2
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 3 |
3 |
1 |
1 |
0 |
0 |
0 |
30.0 |
+----- Anodonta_anatina **
--+
| +-- Pseudanodonta_complanata **
+--+
+-- Anodonta_cygnea **
Froufe et al., 2015 [Publ 3664], fig. 1
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 3 |
3 |
1 |
2 |
9 |
0 |
0 |
31.0 |
+----- Anodonta_anatina **
--+
| +-- Utterbackia_peninsularis **
+--+
+-- Pyganodon_grandis **
Froufe et al., 2015 [Publ 3664], fig. 1-M
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 3 |
3 |
1 |
2 |
5 |
0 |
0 |
31.0 |
+----- Anodonta_anatina **
--+
| +-- Utterbackia_peninsularis **
+--+
+-- Pyganodon_grandis **
Soroka, 2008 [Publ 3695], fig. 1
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 3 |
3 |
1 |
3 |
1 |
0 |
0 |
30.0 |
+----- Sinanodonta_woodiana **
--+
| +-- Anodonta_anatina **
+--+
+-- Pyganodon_grandis **
Soroka, 2008 [Publ 3695], fig. 1-M
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 3 |
3 |
1 |
3 |
1 |
0 |
0 |
30.0 |
+----- Sinanodonta_woodiana **
--+
| +-- Anodonta_anatina **
+--+
+-- Pyganodon_grandis **
Lopes-Lima et al., 2017 [Publ 3734], fig. 1
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 3 |
3 |
1 |
1 |
14 |
9 |
1 |
30.0 |
+----- Anodonta_anatina **
+--------+
| | +-- Pseudanodonta_complanata **
| +--+
| +-- Anodonta_cygnea **
--+
| +-- Unio_tumidus
| +--------+
| | +-- Unio_gibbus
| |
+--+ +-- Unio_crassus
| +-----+
| | +-- Unio_tumidiformis
+--+
| +----- Unio_ravoisieri
| |
+--+ +-- Unio_mancus
| |
| +-- Unio_pictorum
+--+
+-- Unio_delphinus
|
+-- Unio_elongatulus
Lopes-Lima et al., 2016 [Publ 3757], fig. 1
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 3 |
3 |
1 |
1 |
0 |
0 |
0 |
30.0 |
+----- Pseudanodonta_complanata **
--+
| +-- Anodonta_anatina **
+--+
+-- Anodonta_cygnea **
Soroka & Burzynski, 2018 [Publ 3762], fig. 3
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 3 |
3 |
1 |
2 |
4 |
2 |
1 |
31.0 |
+-- Unio_tumidus
+-----+
| +-- Aculamprotula_tortuosa
--+
| +----- Anodonta_anatina **
+--+
| +-- Pyganodon_grandis **
+--+
+-- Utterbackia_peninsularis **
Soroka & Burzynski, 2018 [Publ 3762], fig. 3-M
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 3 |
3 |
1 |
2 |
4 |
2 |
1 |
31.0 |
+-- Unio_tumidus
+-----+
| +-- Aculamprotula_tortuosa
--+
| +----- Anodonta_anatina **
+--+
| +-- Pyganodon_grandis **
+--+
+-- Utterbackia_peninsularis **
Burzynski et al., 2017 [Publ 3774], fig. 1 bottom
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 3 |
3 |
1 |
2 |
10 |
3 |
1 |
31.0 |
+----- Aculamprotula_tortuosa
+--+
| | +-- Unio_crassus
| +--+
--+ +-- Unio_tumidus
|
| +----- Anodonta_anatina **
+--+
| +-- Utterbackia_peninsularis **
+--+
+-- Pyganodon_grandis **
Burzynski et al., 2017 [Publ 3774], fig. 1 top
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 3 |
3 |
1 |
2 |
10 |
3 |
1 |
31.0 |
+----- Aculamprotula_tortuosa
+--+
| | +-- Unio_crassus
| +--+
--+ +-- Unio_tumidus
|
| +----- Anodonta_anatina **
+--+
| +-- Utterbackia_peninsularis **
+--+
+-- Pyganodon_grandis **
Fonseca et al., 2016 [Publ 3775], fig. 1-M
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 3 |
3 |
1 |
2 |
6 |
1 |
1 |
31.0 |
+-------- Unio_delphinus
--+
| +----- Anodonta_anatina **
+--+
| +-- Utterbackia_peninsularis **
+--+
+-- Pyganodon_grandis **
Araujo et al., 2018 [Publ 3779], fig. 1
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 3 |
3 |
1 |
2 |
16 |
15 |
1 |
30.0 |
+----- Anodonta_anatina **
+-----------------+
| | +-- Pyganodon_grandis **
| +--+
| +-- Alasmidonta_marginata **
--+
| +-------------------- Nodularia_douglasiae
| |
| | +-- Unio_durieui
+--+ +--------------+
| | +-- Unio_gibbus
| |
+--+ +-------------- Unio_tumidus
| |
| | +-------- Unio_tumidiformis
| | +--+
+--+ | | +----- Unio_bruguierianus
| | +--+
| | | +-- Unio_crassus
| | +--+
+--+ +-- Unio_ionicus
|
| +-- Unio_delphinus
| +--+
| | +-- Unio_foucauldianus
| |
+-----+ +-- Unio_pictorum
| |
| +-- Unio_elongatulus
| |
+--+-- Unio_mancus
|
+-- Unio_ravoisieri
|
+-- Unio_tigridis
Lopes-Lima et al., 2017 [Publ 3792], fig. 2-M
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 3 |
3 |
1 |
2 |
7 |
1 |
1 |
31.0 |
+-------- Unio_delphinus
--+
| +----- Anodonta_anatina **
+--+
| +-- Utterbackia_peninsularis **
+--+
+-- Pyganodon_grandis **
Wen et al., 2017 [Publ 3795], fig. 3
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 3 |
3 |
1 |
2 |
13 |
4 |
2 |
31.0 |
+----- Anodonta_anatina **
+--+
| | +-- Sinanodonta_woodiana **
| +--+
| +-- Cristaria_plicata **
--+
| +-- Aculamprotula_coreana
| +--+
| | +-- Aculamprotula_tortuosa
+--+
| +-- Lepidodesma_languilati
+--+
+-- Unio_pictorum
Wen et al., 2017 [Publ 3795], fig. 3-M
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 3 |
3 |
1 |
2 |
3 |
0 |
0 |
31.0 |
+----- Anodonta_anatina **
--+
| +-- Utterbackia_peninsularis **
+--+
+-- Pyganodon_grandis **
Chase et al., 2018 [Publ 3861], fig. 1
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 3 |
3 |
1 |
2 |
8 |
2 |
1 |
31.0 |
+-- Unio_delphinus
+-----+
| +-- Unio_tumidus
--+
| +----- Anodonta_anatina **
+--+
| +-- Utterbackia_peninsularis **
+--+
+-- Pyganodon_grandis **
Chase et al., 2018 [Publ 3861], fig. 1-M
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 3 |
3 |
1 |
2 |
8 |
2 |
1 |
31.0 |
+-- Unio_delphinus
+-----+
| +-- Unio_tumidus
--+
| +----- Anodonta_anatina **
+--+
| +-- Utterbackia_peninsularis **
+--+
+-- Pyganodon_grandis **
Krebs et al., 2019 [Publ 4092], fig. 4
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 3 |
3 |
1 |
1 |
0 |
0 |
0 |
33.0 |
+----- Pyganodon_grandis **
--+
| +-- Pyganodon_fragilis **
+--+
+-- Pyganodon_cataracta **
Kano et al., 2019 [Publ 4194], fig. 6
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 3 |
3 |
1 |
1 |
7 |
3 |
2 |
32.0 |
+----- Sinanodonta_lauta **
--+
| +-- Pletholophus_tenuis **
+--+
+-- Beringiana_japonica **
Lopes-Lima et al., 2021 [Publ 4217], fig. 1
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 3 |
3 |
1 |
1 |
22 |
16 |
1 |
32.0 |
+----- Anodonta_anatina **
+--------------------+
| | +-- Anodonta_cygnea **
| +--+
| +-- Anodonta_seddoni **
--+
| +----------------------- Unio_tumidus
| |
| | +-------- Unio_damascensis
| | +-----------+
+--+ | | +----- Unio_sesirmensis
| | +--+
| | | +-- Unio_bruguierianus
| | +--+
+--+ +-- Unio_crassus
|
| +-- Unio_delicatus
| +--------------+
| | +-- Unio_terminalis
| |
+--+ +-- Unio_tigridis
| +-----------+
| | +-- Unio_hueti
| |
+--+ +----------- Unio_ravoisieri
| |
| | +-- Unio_pictorum
+--+ +-----+
| | +-- Unio_eucirrus
| |
+--+ +-- Unio_elongatulus
| +--+
| | +-- Unio_mancus
+--+
| +-- Unio_foucauldianus
+--+
+-- Unio_delphinus
Smith, 2021 [Publ 4331], fig. 1
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 3 |
3 |
1 |
2 |
17 |
0 |
0 |
31.0 |
+----- Cristaria_plicata **
--+
| +-- Pyganodon_grandis **
+--+
+-- Utterbackia_peninsularis **
Nagel et al., 1998 [Publ 1946], fig. 9
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 2 |
2 |
0 |
1 |
4 |
1 |
1 |
-- |
+----- Unio
--+
| +-- Anodonta **
+--+
+-- Pseudanodonta **
Graf, 2002 [Publ 2651], fig. 2
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 2 |
2 |
0 |
1 |
16 |
2 |
1 |
-- |
+-- Unio_pictorum
+--+
| +-- Unio_caffer
--+
| +-- Alasmidonta_marginata **
+--+
+-- Pyganodon_grandis **
Zhou et al., 2007 [Publ 2658], fig. 1
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 2 |
2 |
0 |
1 |
21 |
13 |
3 |
-- |
+-- Sinanodonta_woodiana **
--+
+-- Cristaria_plicata **
Zhou et al., 2007 [Publ 2658], fig. 2
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 2 |
21 |
0 |
1 |
19 |
8 |
2 |
44.0 |
+-------------------------- Sinanodonta_woodiana **
|
| +----------------------- Cristaria_plicata **
| |
--+ | +----- Acuticosta_chinensis
| | +--------+
| | | | +-- Nodularia_douglasiae
| | | +--+
| | +-----+ +-- Cuneopsis_celtiformis
+--+ | |
| | | +----------- Aculamprotula_tortuosa
| | +--+
| | | +-------- Aculamprotula_scripta
| | +--+
| | | +----- Aculamprotula_fibrosa
| | +--+
| | | +-- Aculamprotula_zonata
+--+ +--+
| +-- Aculamprotula_tientsinensis
|
| +-------- Lamprotula_leaii
| +--------+
| | | +----- Lamprotula_cornuumlunae
| | +--+
| | | +-- Gibbosula_rochechouartii
| | +--+
+--+ +-- Lamprotula_caveata
|
| +-- Theliderma_intermedia
| +-----------+
| | +-- Theliderma_sparsa
+--+
| +----------- Quadrula_quadrula
+--+
| +-------- Pustulosa_infucata
+--+
| +----- Quadrula_verrucosa
+--+
| +-- Pustulosa_kieneriana
+--+
+-- Pustulosa_succissa
Walker et al., 2006 [Publ 2666], fig. 6
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 2 |
2 |
0 |
1 |
36 |
0 |
0 |
-- |
+-- Pyganodon_fragilis **
--+
+-- Pyganodon_grandis **
Walker et al., 2006 [Publ 2666], fig. 6-M
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 2 |
2 |
0 |
1 |
13 |
0 |
0 |
-- |
+-- Pyganodon_fragilis **
--+
+-- Pyganodon_grandis **
Hoeh et al., 2002 [Publ 2673], fig. 1-F
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 2 |
2 |
0 |
1 |
7 |
0 |
0 |
-- |
+-- Pyganodon_fragilis **
--+
+-- Pyganodon_grandis **
Hoeh et al., 2002 [Publ 2673], fig. 1-M
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 2 |
2 |
0 |
1 |
7 |
0 |
0 |
-- |
+-- Pyganodon_fragilis **
--+
+-- Pyganodon_grandis **
Curole & Kocher, 2005 [Publ 2682], fig. 1-M
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 2 |
2 |
0 |
1 |
10 |
0 |
0 |
-- |
+-- Utterbackiana_implicata **
--+
+-- Pyganodon_grandis **
Mock et al., 2004 [Publ 2686], fig. 3
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 2 |
2 |
0 |
2 |
0 |
0 |
0 |
-- |
+-- Anodonta **
--+
+-- Sinanodonta_woodiana **
Mock et al., 2004 [Publ 2686], fig. 4
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 2 |
2 |
0 |
2 |
0 |
0 |
0 |
-- |
+-- Anodonta **
--+
+-- Sinanodonta_woodiana **
Krebs, 2004 [Publ 2692], fig. 2
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 2 |
2 |
0 |
1 |
3 |
0 |
0 |
-- |
+-- Lasmigona_costata **
--+
+-- Pyganodon_grandis **
Krebs, 2004 [Publ 2692], fig. 2-M
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 2 |
2 |
0 |
1 |
2 |
0 |
0 |
-- |
+-- Lasmigona_costata **
--+
+-- Pyganodon_grandis **
Hoeh et al., 1996 [Publ 2694], fig. 2
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 2 |
2 |
0 |
1 |
3 |
0 |
0 |
-- |
+-- Pyganodon_fragilis **
--+
+-- Pyganodon_grandis **
Zhang et al., 2004 [Publ 2735], fig. 1
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 2 |
2 |
0 |
1 |
12 |
10 |
3 |
-- |
+-- Sinanodonta_woodiana **
+--------------------+
| +-- Cristaria_plicata **
|
--+ +----- Acuticosta_chinensis
| +--------------+
| | | +-- Lanceolaria_grayii
| | +--+
+--+ +-- Lanceolaria_lanceolata
|
| +----------------- Tchangsinaia_piscicula
+--+
| +-------------- Cuneopsis_heudei
+--+
| +----------- Nodularia_douglasiae
+--+
| +-------- Pseudocuneopsis_capitata
+--+
| +----- Cuneopsis_rufescens
+--+
| +-- Schistodesmus_lampreyanus
+--+
+-- Schistodesmus_spinosus
Whelan et al., 2011 [Publ 3380], fig. 1 ML
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 2 |
2 |
0 |
1 |
35 |
2 |
1 |
-- |
+-- Pyganodon_grandis **
+--+
| +-- Lasmigona_costata **
--+
| +-- Unio_pictorum
+--+
+-- Unio_caffer
Whelan et al., 2011 [Publ 3380], fig. 1 MP
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 2 |
2 |
0 |
1 |
35 |
2 |
1 |
-- |
+-- Pyganodon_grandis **
+--+
| +-- Lasmigona_costata **
--+
| +-- Unio_pictorum
+--+
+-- Unio_caffer
Soroka, 2010 [Publ 3404], fig. 2-M
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 2 |
5 |
0 |
2 |
3 |
3 |
1 |
29.0 |
+----------- Sinanodonta_woodiana **
--+
| +-------- Anodonta_anatina **
+--+
| +----- Unio_crassus
+--+
| +-- Unio_tumidus
+--+
+-- Unio_pictorum
Soroka, 2010 [Publ 3404], fig. 4-M
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 2 |
2 |
0 |
2 |
0 |
0 |
0 |
-- |
+-- Anodonta_anatina **
--+
+-- Sinanodonta_woodiana **
Soroka, 2010 [Publ 3404], fig. 5a-M
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 2 |
2 |
0 |
2 |
0 |
0 |
0 |
-- |
+-- Anodonta_anatina **
--+
+-- Sinanodonta_woodiana **
Soroka, 2010 [Publ 3404], fig. 5b-M
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 2 |
2 |
0 |
2 |
0 |
0 |
0 |
-- |
+-- Anodonta_anatina **
--+
+-- Sinanodonta_woodiana **
Pfeiffer & Graf, 2015 [Publ 3620], fig. 2
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 2 |
2 |
0 |
1 |
38 |
1 |
1 |
-- |
+----- Unio_pictorum
--+
| +-- Pyganodon_grandis **
+--+
+-- Lasmigona_costata **
Chen et al., 2012 [Publ 3624], fig. 4
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 2 |
2 |
0 |
2 |
5 |
0 |
0 |
-- |
+-- Pyganodon_grandis **
--+
+-- Cristaria_plicata **
Huang et al., 2013 [Publ 3629], fig. 3-M
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 2 |
2 |
0 |
1 |
4 |
0 |
0 |
-- |
+-- Pyganodon_grandis **
--+
+-- Utterbackia_peninsularis **
Wei et al., 2016 [Publ 3665], fig. 0
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 2 |
4 |
0 |
2 |
4 |
0 |
0 |
32.0 |
+-- Cristaria_plicata **
|
+-- Lampsilis_ornata
--+
+-- Pyganodon_grandis **
|
+-- Quadrula_quadrula
Sheng et al., 2014 [Publ 3704], fig. 2
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 2 |
8 |
0 |
2 |
6 |
0 |
0 |
36.0 |
+----------------- Cristaria_plicata **
|
--+ +-------------- Pyganodon_grandis **
| |
+--+ +----------- Lampsilis_ornata
| |
+--+ +-- Venustaconcha_ellipsiformis
| +-----+
| | +-- Quadrula_quadrula
+--+
| +----- Lamprotula_leaii
+--+
| +-- Sinohyriopsis_cumingii
+--+
+-- Sinohyriopsis_schlegelii
Xia et al., 2016 [Publ 3712], fig. 7B
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 2 |
3 |
0 |
1 |
1 |
0 |
0 |
31.0 |
+----- Sinanodonta_woodiana **
--+
| +-- Cristaria_plicata **
+--+
+-- Sinohyriopsis_cumingii
Guerra et al., 2017 [Publ 3772], fig. 2a-M
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 2 |
2 |
0 |
1 |
6 |
0 |
0 |
-- |
+-- Utterbackia_peninsularis **
--+
+-- Pyganodon_grandis **
Guerra et al., 2017 [Publ 3772], fig. 2b-M
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 2 |
2 |
0 |
1 |
6 |
0 |
0 |
-- |
+-- Utterbackia_peninsularis **
--+
+-- Pyganodon_grandis **
Zhou et al., 2017 [Publ 3788], fig. 2
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 2 |
2 |
0 |
1 |
15 |
5 |
2 |
-- |
+-- Anemina_arcaeformis **
--+
+-- Anemina_euscaphys **
Amaro et al., 2017 [Publ 3791], fig. 3A-M
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 2 |
2 |
0 |
1 |
5 |
0 |
0 |
-- |
+-- Pyganodon_grandis **
--+
+-- Utterbackia_peninsularis **
Amaro et al., 2017 [Publ 3791], fig. 3B-M
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 2 |
2 |
0 |
1 |
5 |
0 |
0 |
-- |
+-- Utterbackia_peninsularis **
--+
+-- Pyganodon_grandis **
Dyduch-Falniowska & Koziol, 1989 [Publ 3814], fig. 5, 7
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 2 |
2 |
0 |
1 |
3 |
3 |
1 |
-- |
+-- Pseudanodonta_complanata **
+-----+
| +-- Anodonta_anatina **
--+
| +----- Unio_pictorum
+--+
| +-- Unio_tumidus
+--+
+-- Unio_crassus
Chase et al., 2018 [Publ 3847], fig. 5
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 2 |
2 |
0 |
2 |
8 |
2 |
1 |
-- |
+-- Anodonta_anatina **
+--+
| +-- Utterbackia_peninsularis **
--+
| +-- Unio_delphinus
+--+
+-- Unio_tumidus
Bogatov et al., 2018 [Publ 3859], fig. 1
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 2 |
2 |
0 |
1 |
4 |
3 |
1 |
-- |
+-------- Unio_tumidus
|
--+ +-- Anodonta_anatina **
| +--+
| | +-- Pseudanodonta_complanata **
+--+
| +-- Unio_crassus
+--+
+-- Unio_pictorum
Bogatov et al., 2018 [Publ 3859], fig. 2
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 2 |
2 |
0 |
1 |
4 |
3 |
1 |
-- |
+-------- Unio_pictorum
--+
| +----- Unio_tumidus
+--+
| +-- Anodonta_anatina **
+--+
+-- Pseudanodonta_complanata **
Klishko et al., 2019 [Publ 3886], fig. 9
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 2 |
2 |
0 |
1 |
9 |
8 |
1 |
-- |
+-- Anodonta_anatina **
+-----------+
| +-- Anodonta_cygnea **
|
| +-------- Middendorffinaia_mongolica
--+ +--+
| | | +----- Nodularia_breviconcha
| | +--+
| | | +-- Nodularia_douglasiae
+--+ +--+
| +-- Nodularia
|
| +----- Unio_tumidus
| |
+-----+----- Unio_gibbus
|
| +-- Unio_crassus
+--+
+-- Unio_pictorum
Lopes-Lima et al., 2019 [Publ 3901], fig. 2
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 2 |
2 |
0 |
2 |
26 |
0 |
0 |
-- |
+-- Anodonta_nuttalliana **
--+
+-- Pyganodon_grandis **
Lopes-Lima et al., 2019 [Publ 3901], fig. 3
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 2 |
2 |
0 |
2 |
27 |
0 |
0 |
-- |
+-- Anodonta_nuttalliana **
--+
+-- Pyganodon_grandis **
Lopes-Lima et al., 2019 [Publ 3901], fig. 4
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 2 |
2 |
0 |
2 |
24 |
0 |
0 |
-- |
+-- Anodonta_nuttalliana **
--+
+-- Pyganodon_grandis **
Zhang et al., 2013 [Publ 4214], fig. 5
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 2 |
4 |
0 |
1 |
2 |
0 |
0 |
32.0 |
+-------- Cristaria_plicata **
--+
| +----- Sinanodonta_lauta **
+--+
| +-- Margaritifera_margaritifera
+--+
+-- Lamprotula_leaii
Araujo et al., 2009 [Publ 2589], fig. 7A
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 1 |
1 |
0 |
1 |
11 |
8 |
1 |
-- |
+-- Unio_gibbus
|
+-----+-- Unio_tumidus
| |
| +-- Anodonta_cygnea **
--+
| +----- Unio_pictorum[1]
| |
| +----- Unio_pictorum[2]
+--+
+----- Unio_mancus
|
| +-- Unio_tumidiformis
+--+
+-- Unio_crassus
Araujo et al., 2009 [Publ 2589], fig. 7B
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 1 |
1 |
0 |
1 |
11 |
8 |
1 |
-- |
+----------------- Anodonta_cygnea **
|
--+ +-------------- Unio_gibbus
| |
+--+ +----------- Unio_tumidus
| |
| | +-- Unio_tumidiformis
+--+ +-----+
| | +-- Unio_crassus
| |
+--+ +-- Unio_pictorum[1]
| +--+
| | +-- Unio_mancus
+--+
| +-- Unio_pictorum[2]
+--+
+-- Unio_caffer
Araujo et al., 2009 [Publ 2589], fig. 7C
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 1 |
1 |
0 |
1 |
10 |
7 |
1 |
-- |
+----------- Anodonta_cygnea **
|
| +-- Unio_gibbus
--+ +-----+
| | +-- Unio_tumidus
| |
+--+ +-- Unio_tumidiformis
| +--+
| | +-- Unio_crassus
+--+
| +-- Unio_pictorum[1]
| |
+--+-- Unio_pictorum[2]
|
+-- Unio_mancus
Graf, 2000 [Publ 2647], fig. 1
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 1 |
1 |
0 |
1 |
17 |
1 |
1 |
-- |
-- Pyganodon
Graf & Ó Foighil, 2000 [Publ 2649], fig. 1, 2
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 1 |
1 |
0 |
1 |
11 |
1 |
1 |
-- |
+-- Unio_pictorum
--+
+-- Pyganodon_grandis **
Lydeard et al., 2000 [Publ 2654], fig. 2a
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 1 |
1 |
0 |
1 |
16 |
0 |
0 |
-- |
-- Anodonta_cygnea
Lydeard et al., 2000 [Publ 2654], fig. 2b
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 1 |
1 |
0 |
1 |
16 |
0 |
0 |
-- |
-- Anodonta_cygnea
Lydeard et al., 2000 [Publ 2654], fig. 2c
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 1 |
1 |
0 |
1 |
15 |
0 |
0 |
-- |
-- Anodonta_cygnea
Lydeard et al., 2000 [Publ 2654], fig. 2d
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 1 |
1 |
0 |
1 |
15 |
0 |
0 |
-- |
-- Anodonta_cygnea
Zanatta & Murphy, 2006 [Publ 2672], fig. 1
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 1 |
1 |
0 |
1 |
55 |
0 |
0 |
-- |
-- Pyganodon_grandis
Zanatta & Murphy, 2006 [Publ 2672], fig. 2
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 1 |
1 |
0 |
1 |
56 |
0 |
0 |
-- |
-- Pyganodon_grandis
Curole & Kocher, 2005 [Publ 2682], fig. 1
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 1 |
1 |
0 |
1 |
10 |
0 |
0 |
-- |
-- Pyganodon_grandis
Hoeh et al., 1998 [Publ 2695], fig. 3
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 1 |
1 |
0 |
1 |
4 |
0 |
0 |
-- |
-- Anodonta_cygnea
Hoeh et al., 1998 [Publ 2695], fig. 4
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 1 |
1 |
0 |
1 |
4 |
0 |
0 |
-- |
-- Anodonta_cygnea
Taylor et al., 2007 [Publ 2696], fig. 1-3
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 1 |
1 |
0 |
1 |
3 |
1 |
1 |
-- |
+-- Anodonta_cygnea **
--+
+-- Unio_pictorum
Huff et al., 2004 [Publ 2697], fig. 2
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 1 |
1 |
0 |
1 |
13 |
1 |
1 |
-- |
+-- Unio_ravoisieri
--+
+-- Anodonta **
Huff et al., 2004 [Publ 2697], fig. 3
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 1 |
1 |
0 |
1 |
11 |
0 |
0 |
-- |
-- Anodonta
Huff et al., 2004 [Publ 2697], fig. 4
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 1 |
1 |
0 |
1 |
10 |
1 |
1 |
-- |
+-- Unio_ravoisieri
--+
+-- Anodonta **
Huff et al., 2004 [Publ 2697], fig. 5-6
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 1 |
1 |
0 |
1 |
11 |
1 |
1 |
-- |
-- Anodonta
Curole & Kocher, 2002 [Publ 2712], fig. 1a-M
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 1 |
1 |
0 |
1 |
3 |
0 |
0 |
-- |
-- Utterbackiana_implicata
Curole & Kocher, 2002 [Publ 2712], fig. 1b-M
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 1 |
1 |
0 |
1 |
3 |
0 |
0 |
-- |
-- Utterbackiana_implicata
Turner et al., 2000 [Publ 2722], fig. 2
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 1 |
1 |
0 |
1 |
15 |
0 |
0 |
-- |
-- Utterbackiana_couperiana
Campbell, 2000 [Publ 2724], fig. 1-5
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 1 |
1 |
0 |
1 |
2 |
0 |
0 |
-- |
-- Utterbackia_imbecillis
Plazzi & Passamonti, 2010 [Publ 3354], fig. 2-3
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 1 |
1 |
0 |
1 |
3 |
0 |
0 |
-- |
-- Sinanodonta_woodiana
Mulvey et al., 1997 [Publ 3390], fig. 1, 3
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 1 |
1 |
0 |
1 |
4 |
0 |
0 |
-- |
-- Utterbackia_imbecillis
Soroka, 2010 [Publ 3404], fig. 8a-M
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 1 |
1 |
0 |
1 |
2 |
2 |
1 |
-- |
+----- Sinanodonta_woodiana **
--+
| +-- Unio_pictorum
+--+
+-- Unio_tumidus
Soroka, 2010 [Publ 3404], fig. 8b-M
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 1 |
1 |
0 |
1 |
2 |
2 |
1 |
-- |
+----- Sinanodonta_woodiana **
--+
| +-- Unio_pictorum
+--+
+-- Unio_tumidus
Jiang et al., 2010 [Publ 3438], fig. 5
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 1 |
1 |
0 |
1 |
3 |
0 |
0 |
-- |
-- Cristaria_plicata
Inoue et al., 2013 [Publ 3528], fig. 3
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 1 |
1 |
0 |
1 |
16 |
0 |
0 |
-- |
-- Pyganodon_grandis
Zanatta & Harris, 2013 [Publ 3530], fig. 2
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 1 |
1 |
0 |
1 |
4 |
0 |
0 |
-- |
-- Pyganodon_grandis
Inoue et al., 2013 [Publ 3579], fig. 3
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 1 |
1 |
0 |
1 |
8 |
0 |
0 |
-- |
-- Pyganodon_grandis
Marshall et al., 2014 [Publ 3593], fig. 1
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 1 |
1 |
0 |
1 |
13 |
1 |
1 |
-- |
+-- Pyganodon_grandis **
--+
+-- Unio_pictorum
Pfeiffer et al., 2016 [Publ 3660], fig. 2
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 1 |
1 |
0 |
1 |
38 |
0 |
0 |
-- |
-- Utterbackiana_suborbiculata
Peng et al., 2012 [Publ 3689], fig. 1
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 1 |
1 |
0 |
1 |
2 |
0 |
0 |
-- |
-- Cristaria_plicata
Bolotov et al., 2016 [Publ 3743], fig. 2
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 1 |
1 |
0 |
1 |
29 |
2 |
1 |
-- |
+----- Anodonta_anatina **
--+
| +-- Unio_pictorum
+--+
+-- Unio_ravoisieri
Bolotov et al., 2016 [Publ 3743], fig. 4
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 1 |
1 |
0 |
1 |
29 |
2 |
1 |
-- |
+----- Anodonta_anatina **
--+
| +-- Unio_pictorum
+--+
+-- Unio_ravoisieri
Araujo et al., 2017 [Publ 3801], fig. 2
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 1 |
1 |
0 |
1 |
13 |
3 |
1 |
-- |
+-- Schistodesmus_lampreyanus
+--+
| +-- Aculamprotula_tortuosa
--+
| +-- Anodonta_anatina **
+--+
+-- Unio_delphinus
Pieri et al., 2018 [Publ 3867], fig. 3
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 1 |
1 |
0 |
1 |
36 |
0 |
0 |
-- |
-- Arcidens_confragosus
Liu et al., 2017 [Publ 3939], fig. 2
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 1 |
1 |
0 |
1 |
16 |
15 |
3 |
-- |
-- Anemina_arcaeformis
Inoue et al., 2020 [Publ 3941], fig. 3
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 1 |
1 |
0 |
1 |
43 |
0 |
0 |
-- |
-- Arcidens_confragosus
Inoue et al., 2020 [Publ 3941], fig. 4
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 1 |
1 |
0 |
1 |
42 |
0 |
0 |
-- |
-- Arcidens_confragosus
Gomes-dos-Santos et al., 2019 [Publ 3943], fig. 1
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 1 |
1 |
0 |
1 |
9 |
1 |
1 |
-- |
+-- Unio_delphinus
--+
+-- Anodonta_anatina **
Gomes-dos-Santos et al., 2019 [Publ 3943], fig. 1-M
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 1 |
1 |
0 |
1 |
5 |
1 |
1 |
-- |
+-- Unio_delphinus
--+
+-- Anodonta_anatina **
Metzger et al., 2018 [Publ 3952], fig. 1C
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 1 |
1 |
0 |
1 |
2 |
0 |
0 |
-- |
-- Lasmigona_costata
Hewitt et al., 2019 [Publ 3962], fig. 2
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 1 |
1 |
0 |
1 |
15 |
0 |
0 |
-- |
-- Pyganodon_grandis
Sonowal et al., 2021 [Publ 4221], fig. 3a
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 1 |
1 |
0 |
1 |
14 |
3 |
1 |
-- |
+-- Anodonta_cygnea **
|
+-- Unio_ravoisieri
--+
+-- Unio_pictorum
|
+-- Aculamprotula_tientsinensis
Sonowal et al., 2021 [Publ 4221], fig. 3b
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 1 |
1 |
0 |
1 |
14 |
3 |
1 |
-- |
-- Anodonta_cygnea
Sonowal et al., 2021 [Publ 4221], fig. 4
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 1 |
1 |
0 |
1 |
15 |
3 |
1 |
-- |
-- Anodonta_cygnea
| 
