| Page last updated
Tue 01 Jul 2025 |
The Freshwater Mussels (Unionoida) of the World (and other less consequential bivalves)
FM(U)otW(aolcb) is the web version of the MUSSEL Project Database. Follow the links to browse the data or use the custom Google search field. Either way, you win!
This page documents data for Hoeh et al. (2009), such as genera and species attributed to that publication (including known misspellings, misidentifications, and other unavailable names) as well as taxonomic opinions, if available. Taxonomic opinions refer to uses of genera and species as valid and invalid in that work.
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Publication information
Hoeh, W.R., A.E. Bogan, W.H. Heard & E.G. Chapman. 2009. Palaeoheterodont phylogeny, character evolution, diversity and phylogenetic classification: a reflection on methods of analysis. Malacologia 51(2): 307-317.
Click here for online access.
Cladogram Figures
(click the figure number to see the cladogram) fig. 3
fig. 4
fig. 5
Attributed Genus Names
Monocondylea Moricand, 1837
Moricand, 1837, Mém. Soc. Phys. Hist. Nat. Genève 8(1): 145.
d’Orbigny, 1846, Voyage dans l’Amérique Méridionale 5: 611.
Gray, 1847, Proc. Zool. Soc. London 15: 196.
Herrmannsen, 1849, Ind. Gener. Malacol. 2: 668.
Gould, 1850, Proc. Boston Soc. Nat. Hist. 3: 293.
Hupé, 1857, [in] Castelnau, Anim. Nouv. Rares d’Amer. Sud: 84.
Petit de la Saussaye, 1865, J. de Conch. 13: 15.
Heude, 1875, Conch. Fluv. Nanking (1): pl. 4.
Deshayes & Jullien, 1876, Nouv. Arch. Mus. Hist. Nat. Paris 10: 117.
Morlet, 1889, J. de Conch. 37: 165.
Hoeh et al., 2009, Malacologia 51(2): 310.
Zieritz et al., 2015, J. Zool. Syst. Evol. Res. 53(1): 82.
(error for Monocondylaea d’Orbigny, 1835)
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Attributed Species Names
Lasmigona compressa Graf & Ó Foighil, 2000
‘Lea’ of Authors.
Graf & Ó Foighil, 2000, J. Moll. Studies 66: 160.
Graf & Cummings, 2006, Zool. J. Linn. Soc. 148: 348.
Hoeh et al., 2009, Malacologia 51(2): 310.
Jadhav & Jamkhedkar, 2009, Res. J. BioTech. 4(1): 43.
Whelan et al., 2011, MPE 61: 506.
Lopes-Lima et al., 2017, MPE 106: 177.
Bolotov et al., 2017, Sci. Repts. 7: Table S1.
Bolotov et al., 2017, Sci. Repts. 7: Table S1.
Bolotov et al., 2018, Sci. Repts. 8: Table S1.
Huang et al., 2019, MPE 130: Table S4.
Lopes-Lima et al., 2020, MPE 146: Table 1.
Wu et al., 2021, J. Lake Sci. 33(6): attached Table 1.
Jeratthitikul et al., 2022, Sci. Repts. 12: Table S1.
Wu et al., 2023, ZooKeys 1166: 263, Table 1.
Liu et al., 2023, ZooKeys 1179: Table 1.
Dai et al., 2025, Zoosyst. Evol. 101(2): Table 1. [in part]
(misidentified reference to Symphynota compressa Lea, 1829)
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Pseudodon vondembuschianus Graf, 2002
‘Lea’ of Authors.
Graf, 2002, J. Moll. Studies 68: 66.
Graf & Cummings, 2006, Zool. J. Linn. Soc. 148: 348.
Hoeh et al., 2009, Malacologia 51(2): 310.
Whelan et al., 2011, MPE 61: 506. [in part]
Pfeiffer & Graf, 2015, Zool. J. Linn. Soc. 175(2): 310. [in part]
Bolotov et al., 2017, Sci. Repts. 7: Table S1 [as ‘Monodontina aff. vondembuschiana sp. 2’]. [in part]
Bolotov et al., 2017, Sci. Repts. 7: Table S1 [as ‘Pseudodon aff. vondembuschianus sp. 2’]. [in part]
Konopleva et al., 2017, Systemat. Biodiv. 15(3): 207. [in part]
Bolotov et al., 2018, Sci. Repts. 8: Table S1 [as ‘Monodontina aff. vondembuschiana sp. 2’]. [in part]
Huang et al., 2019, MPE 130: Table S4 [as ‘Monodontina aff. vondembuschiana sp. 2’]. [in part]
Konopleva et al., 2019, Sci. Repts. 9: Table S1 [as ‘Monodontina aff. vondembuschiana sp. 2’]. [in part]
(reference to Margaritana vondembuschiana Lea, 1840)
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Pilsbryoconcha exilis Graf, 2002
‘Lea’ of Authors.
Graf, 2002, J. Moll. Studies 68: 66.
Graf & Cummings, 2006, Zool. J. Linn. Soc. 148: 372.
Hoeh et al., 2009, Malacologia 51(2): 310.
Lopes-Lima et al., 2017, MPE 106: 177. [in part]
(reference to Anodonta exilis Lea, 1838)
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Taxonomic Opinions
Cladogram Figure 3
+----------------------------------------------- Neotrigonia_margaritacea
|
| +----- Velesunio_ambiguus
| +--------+
| | | +-- Velesunio_angasi
| | +--+
| +-----------------------------+ +-- Lortiella_rugata
| | |
| | | +----------- Echyridella_menziesii
--+ | | |
| | +--+ +-- Hyridella_depressa
| | | +-----+
| | | | +-- Hyridella_australis
| | +--+
| | | +----- Castalia
| | +--+
| | | +-- Diplodon_chilensis
| | +--+
| | +-- Diplodon_gratus
+--+
| +----- Chambardia_wahlbergi
| |
| +--------+----- Aspatharia_rugifera
| | |
| | | +-- Mutela_rostrata
| +--------------------------+ +--+
| | | +-- Mutela_dubia
| | |
| | | +----------- Etheria_elliptica
| | +--+
| | | +-------- Mulleria_rivoli
| | +--+
| | | +----- Monocondylaea_minuana
+--+ +--+
| | +-- Anodontites_trigona
| +--+
| +-- Anodontites_guanarensis
|
| +-- Margaritifera_margaritifera
| +-----------------------------------+
| | +-- Cumberlandia_monodonta
| |
+--+ +----------------------------------- Coelatura_aegyptiaca
| |
| | +-------------------------------- Pseudomulleria_dalyi
| | |
+--+ | +-- Unio_pictorum
| | +--------+
| | | +-- Unio_caffer
| | +-----------------+
+--+ | | +-------- Lasmigona_costata
| | +--+
| | | +----- Alasmidonta_marginata
| | +--+
| | | +-- Strophitus_undulatus
| | +--+
+--+ +-- Pyganodon_grandis
|
| +-------------------------- Lens_contradens
| |
| | +----- Gonidea_angulata
| | +-----------------+
| | | | +-- Pilsbryoconcha_exilis
+--+ | +--+
| | +-- Pseudodon_vondembuschianus
| |
| | +----------- Amblema_plicata
| | |
+--+ | +----- Obliquaria_reflexa
| +--------+ +--+
| | | | | +-- Quadrula_verrucosa
| | | | +--+
| | +--+ +-- Quadrula_quadrula
| | |
| | | +----- Eurynia_dilatata
+--+ +--+
| | +-- Pleurobema_sintoxia
| +--+
| +-- Fusconaia_flava
|
| +----------------- Ptychobranchus_fasciolaris
+--+
| +-------------- Truncilla_truncata
+--+
| +----------- Ortmanniana_ligamentina
+--+
| +-------- Lampsilis_cardium
+--+
| +----- Cambarunio_iris
+--+
| +-- Ligumia_recta
+--+
+-- Epioblasma_triquetra
Cladogram Figure 4
+-------------------------------------------------- Neotrigonia_margaritacea
|
| +----- Velesunio_ambiguus
| +--------+
| | | +-- Velesunio_angasi
| | +--+
| +--------------------------------+ +-- Lortiella_rugata
| | |
| | | +----------- Echyridella_menziesii
--+ | | |
| | +--+ +-- Hyridella_depressa
| | | +-----+
| | | | +-- Hyridella_australis
| | +--+
| | | +----- Castalia
| | +--+
| | | +-- Diplodon_chilensis
| | +--+
| | +-- Diplodon_gratus
+--+
| +-- Etheria_elliptica
| +--------+
| | +-- Mulleria_rivoli
| |
| | +----- Monocondylaea_minuana
| +--------------------------------+ +--+
| | | | | +-- Anodontites_trigona
| | | | +--+
| | | | +-- Anodontites_guanarensis
| | +--+
| | | +-- Mutela_rostrata
| | | +--+
+--+ | | +-- Mutela_dubia
| +--+
| | +-- Chambardia_wahlbergi
| +--+
| +-- Aspatharia_rugifera
|
| +----------------------------------------- Coelatura_aegyptiaca
| |
+--+ +-------------------------------------- Pseudomulleria_dalyi
| |
| | +-- Margaritifera_margaritifera
+--+ +--------------------------------+
| | +-- Cumberlandia_monodonta
| |
| | +-- Pilsbryoconcha_exilis
+--+ +-----------------------------+
| | +-- Pseudodon_vondembuschianus
| |
| | +----------------------------- Lens_contradens
+--+ |
| | +-- Unio_pictorum
| | +--------+
| | | +-- Unio_caffer
| | +--------------+
+--+ | | +-------- Lasmigona_costata
| | +--+
| | | +----- Alasmidonta_marginata
| | +--+
| | | +-- Strophitus_undulatus
| | +--+
+--+ +-- Pyganodon_grandis
|
| +----------------------- Gonidea_angulata
| |
| | +-- Eurynia_dilatata
| | |
| | +-----+-- Pleurobema_sintoxia
+--+ | |
| +-----------+ +-- Fusconaia_flava
| | |
| | | +----- Obliquaria_reflexa
| | +--+
+--+ | +-- Quadrula_verrucosa
| +--+
| +-- Quadrula_quadrula
|
| +----------------- Ptychobranchus_fasciolaris
+--+
| +-------------- Truncilla_truncata
+--+
| +----------- Ortmanniana_ligamentina
+--+
| +-------- Lampsilis_cardium
+--+
| +----- Cambarunio_iris
+--+
| +-- Ligumia_recta
+--+
+-- Epioblasma_triquetra
Cladogram Figure 5
+-------------------------------- Neotrigonia_margaritacea
|
| +----------------------------- Coelatura_aegyptiaca
| |
| | +-- Margaritifera_margaritifera
| | +--------------------+
| | | +-- Cumberlandia_monodonta
| | |
--+ | | +----- Velesunio_ambiguus
| | | +--------+
| | | | | +-- Velesunio_angasi
| | | | +--+
| | +--+ +-----+ +-- Lortiella_rugata
| | | | | |
| | | | | | +----------- Echyridella_menziesii
| | | | | | |
+--+ | | | +--+ +-- Hyridella_depressa
| | | | | +-----+
| | | | | | +-- Hyridella_australis
| | | | +--+
| | +--+ | +----- Castalia
| | | +--+
| | | | +-- Diplodon_chilensis
| | | +--+
| | | +-- Diplodon_gratus
| | |
| | | +----------------- Pseudomulleria_dalyi
| | | |
| | +--+ +-------------- Mulleria_rivoli
| | | |
| | +--+ +----------- Etheria_elliptica
+--+ | |
| | | +----- Monocondylaea_minuana
| +--+ +--+
| | | | +-- Anodontites_trigona
| | | +--+
| | | +-- Anodontites_guanarensis
| +--+
| | +-- Mutela_rostrata
| | +--+
| | | +-- Mutela_dubia
| +--+
| | +-- Chambardia_wahlbergi
| +--+
| +-- Aspatharia_rugifera
|
| +-- Pilsbryoconcha_exilis
| +--------------------+
| | +-- Pseudodon_vondembuschianus
| |
| | +-------------------- Lens_contradens
+--+ |
| | +-- Unio_pictorum
| | +--------+
| | | +-- Unio_caffer
+--+ +-----+
| | | +-------- Lasmigona_costata
| | +--+
| | | +----- Alasmidonta_marginata
| | +--+
| | | +-- Strophitus_undulatus
+--+ +--+
| +-- Pyganodon_grandis
|
| +-------------- Gonidea_angulata
| |
| | +----- Obliquaria_reflexa
| | +-----+
+--+ | | +-- Quadrula_verrucosa
| | +--+
| | +-- Quadrula_quadrula
| |
+--+ +-------- Amblema_plicata
| |
| | +----- Eurynia_dilatata
| +--+
| | | +-- Pleurobema_sintoxia
+--+ +--+
| +-- Fusconaia_flava
|
| +----- Truncilla_truncata
| |
| +----- Ptychobranchus_fasciolaris
+--+
| +-- Ortmanniana_ligamentina
| |
| +-- Cambarunio_iris
+--+
+-- Ligumia_recta
|
+-- Epioblasma_triquetra
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Page updated: 08:04:56 Tue 01 Jul 2025 |
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