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Page last updated
Tue 04 Jan 2022

The Freshwater Mussels (Unionoida) of the World (and other less consequential bivalves)

FM(U)otW(aolcb) is the web version of the MUSSEL Project Database. Follow the links to browse the data or use the custom Google search field. Either way, you win!

Cladomics pages are experimental — even more so than the rest of this site!

The purpose of the cladomics pages for each genus and family-group level taxon is to provide statistics and references indicating the extent to which published phylogenetic work supports the classification used here.

The Cladomics Summary provides an overview of the number of studies that have included terminal species representing this taxon as well as an indication of how frequently it has been recovered as monophyletic.

Following the summary, the available published clades representing the taxon are listed.

For more information, visit the Cladomics FARQ (frequently asked rhetorical questions) page.

Search the database pages:

Bivalvia | Unionoida | Hyrioidea | Hyriidae

subfamily HYRIINAE Swainson, 1840: cladomics
     4 tribes, 1 genus

Cladomics Summary

number of species: 76
number next lowest-taxon (tribe) (i.e., direct-child taxa): 4

analyzed in 77 cladogram(s) in 40 publications
maximum number of terminal species in an analysis: 17, mean: 3.1
maximum number of direct-child taxa in an analysis: 4, mean: 1.81

basic monophyly

number of cladograms that test basic monophyly (i.e., more than 1 included species): 49
maximum number of included species: 17, mean: 4.31
maximum number of direct-child taxa: 4, mean: 2.63
proportion of cladograms that support basic monophyly: 0.43

rigorous monophyly

number of cladograms that test rigorous monophyly: 45
(Rigorous monophyly refers to cladograms with terminal species representing more than one direct-child taxon and at least one rigorous, direct-child outgroup.)
maximum number of included species: 17, mean: 4.29
maximum number of direct-child taxa: 4, mean: 2.69
proportion of cladograms that support rigorous monophyly: 0.4

top monophyly publications

Santos-Neto et al., 2016, Graf et al., 2015, Huang et al., 2019, Graf & Cummings, 2006, Hoeh et al., 2009

Santos-Neto et al., 2016 ** Top Monophyly Test Publication **

fig. 2a,b: [clade_id: 7674] "Hyriinae" + Velesunioninae
spp. terminals: 17, direct-child terminals: 4
rigorous outgroups.: 6, direct-child rigorous outgroups: 1
((W_carteri, ((H_drapeta, H_depressa), (D_gratus, D_suavidicus))), ((H_australis, C_novaehollandiae), (E_menziesii, (V_ambiguus, (M_anodontaeformis, (Lortiella, (A_jacksoni, V_angasi)))))), (D_suavidicus, ((T_corrugatus, P_syrmatophorus), (C_stevensi, ((C_duprei, C_ambigua), (C_ambigua, (C_ambigua, (C_quadrata, C_ambigua))))))))


	                 +-------- W_carteri 1         
	                 |                             
	  +--------------+     +-- H_depressa 3        
	  |              |  +--+                       
	  |              |  |  +-- H_drapeta 2         
	  |              +--+                          
	  |                 |  +-- D_gratus 4          
	  |                 +--+                       
	  |                    +-- D_suavidicus 5      
	  |                                            
	  |                    +-- C_novaehollandiae 7 
	  |     +--------------+                       
	  |     |              +-- H_australis 6       
	  +-----+                                      
	  |     |  +-------------- E_menziesii 8       
	--+     +--+                                   
	  |        |  +----------- V_ambiguus 9        
	  |        +--+                                
	  |           |  +-------- M_anodontaeformis 10
	  |           +--+                             
	  |              |  +----- Lortiella 11        
	  |              +--+                          
	  |                 |  +-- A_jacksoni 12       
	  |                 +--+                       
	  |                    +-- V_angasi 13         
	  |                                            
	  |  +-------------------- D_suavidicus 14     
	  |  |                                         
	  +--+                 +-- P_syrmatophorus 16  
	     |  +--------------+                       
	     |  |              +-- T_corrugatus 15     
	     +--+                                      
	        |  +-------------- C_stevensi 17       
	        |  |                                   
	        +--+           +-- C_ambigua 19        
	           |  +--------+                       
	           |  |        +-- C_duprei 18         
	           +--+                                
	              |  +-------- C_ambigua 20        
	              +--+                             
	                 |  +----- C_ambigua 21        
	                 +--+                          
	                    |  +-- C_ambigua 23        
	                    +--+                       
	                       +-- C_quadrata 22       
  

fig. 3, 5: [clade_id: 7695] Hyriinae
spp. terminals: 8, direct-child terminals: 3
rigorous outgroups.: 0, direct-child rigorous outgroups: 0
(D_suavidicus, ((T_corrugatus, (P_obliquus, P_syrmatophorus)), (C_duprei, (C_ambigua, (C_ambigua, C_quadrata)))))


	  +-------------- D_suavidicus 1   
	  |                                
	  |        +----- T_corrugatus 2   
	--+  +-----+                       
	  |  |     |  +-- P_obliquus 3     
	  |  |     +--+                    
	  +--+        +-- P_syrmatophorus 4
	     |                             
	     |  +-------- C_duprei 5       
	     +--+                          
	        |  +----- C_ambigua 6      
	        +--+                       
	           |  +-- C_ambigua 7      
	           +--+                    
	              +-- C_quadrata 8     
  

fig. 4: [clade_id: 7751] "Hyriinae" + Velesunioninae
spp. terminals: 17, direct-child terminals: 4
rigorous outgroups.: 6, direct-child rigorous outgroups: 1
((D_suavidicus, ((T_corrugatus, P_syrmatophorus), (C_ambigua, (C_duprei, (C_ambigua, (C_stevensi, (C_ambigua, (C_quadrata, C_ambigua)))))))), ((W_carteri, ((H_drapeta, H_depressa), (D_suavidicus, D_gratus))), (E_menziesii, ((C_novaehollandiae, H_australis), (V_ambiguus, (A_jacksoni, (V_angasi, (Lortiella, M_anodontaeformis))))))))


	     +----------------------- D_suavidicus 1      
	     |                                            
	  +--+                    +-- P_syrmatophorus 3   
	  |  |  +-----------------+                       
	  |  |  |                 +-- T_corrugatus 2      
	  |  +--+                                         
	  |     |  +----------------- C_ambigua 4         
	  |     +--+                                      
	  |        |  +-------------- C_duprei 5          
	  |        +--+                                   
	  |           |  +----------- C_ambigua 6         
	  |           +--+                                
	  |              |  +-------- C_stevensi 7        
	  |              +--+                             
	--+                 |  +----- C_ambigua 8         
	  |                 +--+                          
	  |                    |  +-- C_ambigua 10        
	  |                    +--+                       
	  |                       +-- C_quadrata 9        
	  |                                               
	  |                 +-------- W_carteri 11        
	  |                 |                             
	  |     +-----------+     +-- H_depressa 13       
	  |     |           |  +--+                       
	  |     |           |  |  +-- H_drapeta 12        
	  |     |           +--+                          
	  |     |              |  +-- D_gratus 15         
	  +-----+              +--+                       
	        |                 +-- D_suavidicus 14     
	        |                                         
	        |  +----------------- E_menziesii 16      
	        |  |                                      
	        +--+              +-- C_novaehollandiae 17
	           |  +-----------+                       
	           |  |           +-- H_australis 18      
	           +--+                                   
	              |  +----------- V_ambiguus 19       
	              +--+                                
	                 |  +-------- A_jacksoni 20       
	                 +--+                             
	                    |  +----- V_angasi 21         
	                    +--+                          
	                       |  +-- Lortiella 22        
	                       +--+                       
	                          +-- M_anodontaeformis 23
  

Graf et al., 2015 ** Top Sister Group Test Publication **

fig. 1: [clade_id: 6998] Hyriinae
spp. terminals: 7, direct-child terminals: 4
rigorous outgroups.: 7, direct-child rigorous outgroups: 1
((D_suavidicus, (T_corrugatus, C_ambigua)), ((H_depressa, H_drapeta), (H_australis, C_novaehollandiae)))


	     +----- D_suavidicus 1     
	  +--+                         
	  |  |  +-- C_ambigua 3        
	  |  +--+                      
	  |     +-- T_corrugatus 2     
	--+                            
	  |     +-- C_novaehollandiae 7
	  |  +--+                      
	  |  |  +-- H_australis 6      
	  +--+                         
	     |  +-- H_depressa 4       
	     +--+                      
	        +-- H_drapeta 5        
  

Huang et al., 2019

fig. 2: [clade_id: 11792] Hyriinae
spp. terminals: 7, direct-child terminals: 4
rigorous outgroups.: 7, direct-child rigorous outgroups: 1
(E_menziesii, ((D_suavidicus, (C_ambigua, T_corrugatus)), (H_australis, (H_depressa, H_drapeta))))


	  +----------- E_menziesii 1 
	  |                          
	  |     +----- D_suavidicus 2
	--+  +--+                    
	  |  |  |  +-- C_ambigua 3   
	  |  |  +--+                 
	  +--+     +-- T_corrugatus 4
	     |                       
	     |  +----- H_australis 5 
	     +--+                    
	        |  +-- H_depressa 6  
	        +--+                 
	           +-- H_drapeta 7   
  

fig. 3: [clade_id: 12696] Hyriinae
spp. terminals: 1, direct-child terminals: 0
rigorous outgroups.: 0, direct-child rigorous outgroups: 0
unknown

fig. 3-M: [clade_id: 12745] Hyriinae
spp. terminals: 1, direct-child terminals: 0
rigorous outgroups.: 0, direct-child rigorous outgroups: 0
unknown

fig. S3: [clade_id: 12213] Hyriinae
spp. terminals: 7, direct-child terminals: 4
rigorous outgroups.: 7, direct-child rigorous outgroups: 1
(E_menziesii, ((D_suavidicus, (C_ambigua, T_corrugatus)), (H_australis, (H_depressa, H_drapeta))))


	  +----------- E_menziesii 1 
	  |                          
	  |     +----- D_suavidicus 2
	--+  +--+                    
	  |  |  |  +-- C_ambigua 3   
	  |  |  +--+                 
	  +--+     +-- T_corrugatus 4
	     |                       
	     |  +----- H_australis 5 
	     +--+                    
	        |  +-- H_depressa 6  
	        +--+                 
	           +-- H_drapeta 7   
  

fig. S4: [clade_id: 12569] "Hyriinae" + Velesunioninae
spp. terminals: 7, direct-child terminals: 4
rigorous outgroups.: 7, direct-child rigorous outgroups: 1
(E_menziesii, W_carteri, (D_suavidicus, (C_ambigua, T_corrugatus)), (H_australis, (H_depressa, H_drapeta)), ((V_ambiguus, (A_jacksoni, A_profuga)), (A_pertexta, (L_froggatti, M_anodontaeformis))))


	  +----------- E_menziesii 1       
	  |                                
	  +----------- W_carteri 2         
	  |                                
	  |     +----- D_suavidicus 3      
	  +-----+                          
	  |     |  +-- C_ambigua 4         
	  |     +--+                       
	  |        +-- T_corrugatus 5      
	  |                                
	--+     +----- H_australis 6       
	  +-----+                          
	  |     |  +-- H_depressa 7        
	  |     +--+                       
	  |        +-- H_drapeta 8         
	  |                                
	  |     +----- V_ambiguus 9        
	  |  +--+                          
	  |  |  |  +-- A_jacksoni 10       
	  |  |  +--+                       
	  +--+     +-- A_profuga 11        
	     |                             
	     |  +----- A_pertexta 12       
	     +--+                          
	        |  +-- L_froggatti 13      
	        +--+                       
	           +-- M_anodontaeformis 14
  

Marshall et al., 2014

fig. 1: [clade_id: 6814] "Hyriinae" + Velesunioninae
spp. terminals: 7, direct-child terminals: 2
rigorous outgroups.: 3, direct-child rigorous outgroups: 1
((E_aucklandica, (E_menziesii, E_onekaka)), ((V_ambiguus, (Velesunio, Lortiella)), ((C_novaehollandiae, H_australis), (D_chilensis, D_gratus))))


	        +----- E_aucklandica 1    
	  +-----+                         
	  |     |  +-- E_menziesii 2      
	  |     +--+                      
	  |        +-- E_onekaka 3        
	  |                               
	--+     +----- V_ambiguus 4       
	  |  +--+                         
	  |  |  |  +-- Lortiella 6        
	  |  |  +--+                      
	  |  |     +-- Velesunio 5        
	  +--+                            
	     |     +-- C_novaehollandiae 7
	     |  +--+                      
	     |  |  +-- H_australis 8      
	     +--+                         
	        |  +-- D_chilensis 9      
	        +--+                      
	           +-- D_gratus 10        
  

Hoeh et al., 2009

fig. 3: [clade_id: 4257] Hyriinae
spp. terminals: 6, direct-child terminals: 3
rigorous outgroups.: 3, direct-child rigorous outgroups: 1
(E_menziesii, ((H_depressa, H_australis), (Castalia, (D_chilensis, D_gratus))))


	  +----------- E_menziesii 1
	  |                         
	--+        +-- H_australis 3
	  |  +-----+                
	  |  |     +-- H_depressa 2 
	  +--+                      
	     |  +----- Castalia 4   
	     +--+                   
	        |  +-- D_chilensis 5
	        +--+                
	           +-- D_gratus 6   
  

fig. 4: [clade_id: 4347] Hyriinae
spp. terminals: 6, direct-child terminals: 3
rigorous outgroups.: 3, direct-child rigorous outgroups: 1
(E_menziesii, ((H_depressa, H_australis), (Castalia, (D_chilensis, D_gratus))))


	  +----------- E_menziesii 1
	  |                         
	--+        +-- H_australis 3
	  |  +-----+                
	  |  |     +-- H_depressa 2 
	  +--+                      
	     |  +----- Castalia 4   
	     +--+                   
	        |  +-- D_chilensis 5
	        +--+                
	           +-- D_gratus 6   
  

fig. 5: [clade_id: 4427] Hyriinae
spp. terminals: 6, direct-child terminals: 3
rigorous outgroups.: 3, direct-child rigorous outgroups: 1
(E_menziesii, ((H_depressa, H_australis), (Castalia, (D_chilensis, D_gratus))))


	  +----------- E_menziesii 1
	  |                         
	--+        +-- H_australis 3
	  |  +-----+                
	  |  |     +-- H_depressa 2 
	  +--+                      
	     |  +----- Castalia 4   
	     +--+                   
	        |  +-- D_chilensis 5
	        +--+                
	           +-- D_gratus 6   
  

Graf & Cummings, 2006

fig. 2: [clade_id: 207] Hyriinae
spp. terminals: 6, direct-child terminals: 3
rigorous outgroups.: 3, direct-child rigorous outgroups: 1
(E_menziesii, ((H_depressa, H_australis), (Castalia, (D_chilensis, D_gratus))))


	  +----------- E_menziesii 1
	  |                         
	--+        +-- H_australis 3
	  |  +-----+                
	  |  |     +-- H_depressa 2 
	  +--+                      
	     |  +----- Castalia 4   
	     +--+                   
	        |  +-- D_chilensis 5
	        +--+                
	           +-- D_gratus 6   
  

Graf & Ó Foighil, 2000

fig. 1, 2: [clade_id: 1191] Hyriinae
spp. terminals: 6, direct-child terminals: 3
rigorous outgroups.: 1, direct-child rigorous outgroups: 1
((D_chilensis, Castalia), (H_australis, (H_australis, (E_menziesii, H_depressa))))


	           +-- Castalia 2   
	  +--------+                
	  |        +-- D_chilensis 1
	--+                         
	  |  +-------- H_australis 3
	  +--+                      
	     |  +----- H_australis 4
	     +--+                   
	        |  +-- E_menziesii 5
	        +--+                
	           +-- H_depressa 6 
  

Baker et al., 2003

fig. 5: [clade_id: 2314] "Hyriinae" + Velesunioninae
spp. terminals: 5, direct-child terminals: 3
rigorous outgroups.: 2, direct-child rigorous outgroups: 1
((H_australis, D_gratus), (H_depressa, ((V_angasi, V_ambiguus), (E_menziesii, C_stevensi))))


	           +-- D_gratus 2   
	  +--------+                
	  |        +-- H_australis 1
	--+                         
	  |  +-------- H_depressa 3 
	  |  |                      
	  +--+     +-- C_stevensi 7 
	     |  +--+                
	     |  |  +-- E_menziesii 6
	     +--+                   
	        |  +-- V_ambiguus 5 
	        +--+                
	           +-- V_angasi 4   
  

Santos-Neto et al., 2018

fig. 5: [clade_id: 13987] Hyriinae
spp. terminals: 5, direct-child terminals: 2
rigorous outgroups.: 0, direct-child rigorous outgroups: 0
((T_corrugatus, P_syrmatophorus), (C_duprei, (C_ambigua, C_stevensi)))


	        +-- P_syrmatophorus 2
	  +-----+                    
	  |     +-- T_corrugatus 1   
	--+                          
	  |  +----- C_duprei 3       
	  +--+                       
	     |  +-- C_ambigua 4      
	     +--+                    
	        +-- C_stevensi 5     
  

Walker et al., 2006

fig. 2: [clade_id: 2732] Hyriinae
spp. terminals: 1, direct-child terminals: 1
rigorous outgroups.: 0, direct-child rigorous outgroups: 0
unknown

fig. 3: [clade_id: 2761] Hyriinae
spp. terminals: 1, direct-child terminals: 1
rigorous outgroups.: 0, direct-child rigorous outgroups: 0
unknown

fig. 4: [clade_id: 2790] Hyriinae
spp. terminals: 1, direct-child terminals: 1
rigorous outgroups.: 0, direct-child rigorous outgroups: 0
unknown

fig. 6: [clade_id: 2897] "Hyriinae" + Velesunioninae
spp. terminals: 5, direct-child terminals: 3
rigorous outgroups.: 7, direct-child rigorous outgroups: 1
(C_stevensi, D_gratus, (H_depressa, H_australis, (E_menziesii, (((V_ambiguus, Velesunio), (Velesunio, (A_jacksoni, Velesunio), (V_angasi, L_rugata)))))))


	  +-------------------- C_stevensi 1 
	  |                                  
	  +-------------------- D_gratus 2   
	  |                                  
	--+  +----------------- H_australis 4
	  |  |                               
	  |  +----------------- H_depressa 3 
	  |  |                               
	  |  |  +-------------- E_menziesii 5
	  +--+  |                            
	     |  +---        +-- V_ambiguus 6 
	     |  +---  +-----+                
	     |  +---  |     +-- Velesunio 7  
	     |  +---  |                      
	     |  +-----+  +----- Velesunio 8  
	     +--+---  |  |                   
	        +---  |  |  +-- A_jacksoni 9 
	        +---  +--+--+                
	        +---     |  +-- Velesunio 10 
	        +---     |                   
	        +---     |  +-- L_rugata 12  
	        +---     +--+                
	        +---        +-- V_angasi 11  
  

fig. 6-M: [clade_id: 2927] Hyriinae
spp. terminals: 1, direct-child terminals: 0
rigorous outgroups.: 0, direct-child rigorous outgroups: 0
unknown

Baker et al., 2004

fig. 2-3: [clade_id: 2590] "Hyriinae" + Velesunioninae
spp. terminals: 5, direct-child terminals: 3
rigorous outgroups.: 6, direct-child rigorous outgroups: 1
((H_depressa, (H_australis, D_gratus)), ((E_menziesii, C_stevensi), ((Velesunio, Velesunio), (Velesunio, (V_angasi, (Velesunio, A_jacksoni))))))


	              +----- H_depressa 1 
	  +-----------+                   
	  |           |  +-- D_gratus 3   
	  |           +--+                
	  |              +-- H_australis 2
	--+                               
	  |              +-- C_stevensi 5 
	  |  +-----------+                
	  |  |           +-- E_menziesii 4
	  |  |                            
	  +--+           +-- Velesunio 6  
	     |  +--------+                
	     |  |        +-- Velesunio 7  
	     +--+                         
	        |  +-------- Velesunio 8  
	        +--+                      
	           |  +----- V_angasi 9   
	           +--+                   
	              |  +-- A_jacksoni 11
	              +--+                
	                 +-- Velesunio 10 
  

Guerra et al., 2019

fig. 1: [clade_id: 19073] "Hyriinae" + Velesunioninae
spp. terminals: 4, direct-child terminals: 3
rigorous outgroups.: 1, direct-child rigorous outgroups: 1
((E_menziesii, W_carteri), (D_suavidicus, (C_ambigua, P_obliquus)))


	        +-- E_menziesii 1 
	  +-----+                 
	  |     +-- W_carteri 2   
	--+                       
	  |  +----- D_suavidicus 3
	  +--+                    
	     |  +-- C_ambigua 4   
	     +--+                 
	        +-- P_obliquus 5  
  

fig. 1-M: [clade_id: 19006] Hyriinae
spp. terminals: 1, direct-child terminals: 0
rigorous outgroups.: 1, direct-child rigorous outgroups: 1
unknown

fig. 2: [clade_id: 19196] Hyriinae
spp. terminals: 4, direct-child terminals: 3
rigorous outgroups.: 1, direct-child rigorous outgroups: 1
(E_menziesii, (D_suavidicus, (C_ambigua, P_obliquus)))


	  +-------- E_menziesii 1 
	--+                       
	  |  +----- D_suavidicus 2
	  +--+                    
	     |  +-- C_ambigua 3   
	     +--+                 
	        +-- P_obliquus 4  
  

fig. 2-M: [clade_id: 19131] Hyriinae
spp. terminals: 1, direct-child terminals: 0
rigorous outgroups.: 1, direct-child rigorous outgroups: 1
unknown

Bolotov et al., 2016

fig. 2: [clade_id: 7793] Hyriinae
spp. terminals: 3, direct-child terminals: 3
rigorous outgroups.: 6, direct-child rigorous outgroups: 1
(D_suavidicus, (T_corrugatus, C_ambigua))


	  +----- D_suavidicus 1
	--+                    
	  |  +-- C_ambigua 3   
	  +--+                 
	     +-- T_corrugatus 2
  

fig. 4: [clade_id: 7859] Hyriinae
spp. terminals: 3, direct-child terminals: 3
rigorous outgroups.: 6, direct-child rigorous outgroups: 1
(D_suavidicus, (T_corrugatus, C_ambigua))


	  +----- D_suavidicus 1
	--+                    
	  |  +-- C_ambigua 3   
	  +--+                 
	     +-- T_corrugatus 2
  

Pfeiffer & Graf, 2015

fig. 2: [clade_id: 7288] Hyriinae
spp. terminals: 3, direct-child terminals: 3
rigorous outgroups.: 1, direct-child rigorous outgroups: 1
(H_australis, (T_corrugatus, C_ambigua))


	  +----- H_australis 1 
	--+                    
	  |  +-- C_ambigua 3   
	  +--+                 
	     +-- T_corrugatus 2
  

Bolotov et al., 2017

fig. S1: [clade_id: 9086] Hyriinae
spp. terminals: 3, direct-child terminals: 3
rigorous outgroups.: 6, direct-child rigorous outgroups: 1
(D_suavidicus, (C_ambigua, T_corrugatus))


	  +----- D_suavidicus 1
	--+                    
	  |  +-- C_ambigua 2   
	  +--+                 
	     +-- T_corrugatus 3
  

fig. S2: [clade_id: 9363] Hyriinae
spp. terminals: 3, direct-child terminals: 3
rigorous outgroups.: 6, direct-child rigorous outgroups: 1
(D_suavidicus, (C_ambigua, T_corrugatus))


	  +----- D_suavidicus 1
	--+                    
	  |  +-- C_ambigua 2   
	  +--+                 
	     +-- T_corrugatus 3
  

Whelan et al., 2011

fig. 1 ML: [clade_id: 5182] Hyriinae
spp. terminals: 3, direct-child terminals: 3
rigorous outgroups.: 1, direct-child rigorous outgroups: 1
(H_australis, (C_ambigua, T_corrugatus))


	  +----- H_australis 1 
	--+                    
	  |  +-- C_ambigua 2   
	  +--+                 
	     +-- T_corrugatus 3
  

fig. 1 MP: [clade_id: 5093] Hyriinae
spp. terminals: 3, direct-child terminals: 3
rigorous outgroups.: 1, direct-child rigorous outgroups: 1
(H_australis, (C_ambigua, T_corrugatus))


	  +----- H_australis 1 
	--+                    
	  |  +-- C_ambigua 2   
	  +--+                 
	     +-- T_corrugatus 3
  

Graf, 2000

fig. 1: [clade_id: 1163] "Hyriinae" + Velesunioninae
spp. terminals: 3, direct-child terminals: 3
rigorous outgroups.: 1, direct-child rigorous outgroups: 1
(Castalia, (Diplodon, (Hyridella, Velesunio)))


	  +-------- Castalia 1 
	--+                    
	  |  +----- Diplodon 2 
	  +--+                 
	     |  +-- Hyridella 3
	     +--+              
	        +-- Velesunio 4
  

Bogan & Hoeh, 2000

fig. 1: [clade_id: 1261] "Hyriinae" + Velesunioninae
spp. terminals: 3, direct-child terminals: 2
rigorous outgroups.: 2, direct-child rigorous outgroups: 1
((C_stevensi, D_gratus), (E_menziesii, (V_angasi, L_rugata)))


	        +-- C_stevensi 1 
	  +-----+                
	  |     +-- D_gratus 2   
	--+                      
	  |  +----- E_menziesii 3
	  +--+                   
	     |  +-- L_rugata 5   
	     +--+                
	        +-- V_angasi 4   
  

Graf, 2002

fig. 2: [clade_id: 1701] Hyriinae
spp. terminals: 3, direct-child terminals: 3
rigorous outgroups.: 1, direct-child rigorous outgroups: 1
(H_depressa, (D_chilensis, Castalia))


	  +----- H_depressa 1 
	--+                   
	  |  +-- Castalia 3   
	  +--+                
	     +-- D_chilensis 2
  

Hoeh et al., 2001

fig. 14.2: [clade_id: 1574] "Hyriinae" + Velesunioninae
spp. terminals: 3, direct-child terminals: 2
rigorous outgroups.: 2, direct-child rigorous outgroups: 1
(D_gratus, E_menziesii, L_rugata, (C_stevensi, V_angasi))


	  +----- D_gratus 1   
	  |                   
	  +----- E_menziesii 2
	--+                   
	  +----- L_rugata 3   
	  |                   
	  |  +-- C_stevensi 4 
	  +--+                
	     +-- V_angasi 5   
  

fig. 14.4: [clade_id: 1668] "Hyriinae" + Velesunioninae
spp. terminals: 3, direct-child terminals: 2
rigorous outgroups.: 2, direct-child rigorous outgroups: 1
((C_stevensi, D_gratus), (E_menziesii, (L_rugata, V_angasi)))


	        +-- C_stevensi 1 
	  +-----+                
	  |     +-- D_gratus 2   
	--+                      
	  |  +----- E_menziesii 3
	  +--+                   
	     |  +-- L_rugata 4   
	     +--+                
	        +-- V_angasi 5   
  

Hoeh et al., 2002

fig. 2: [clade_id: 1763] "Hyriinae" + Velesunioninae
spp. terminals: 3, direct-child terminals: 2
rigorous outgroups.: 2, direct-child rigorous outgroups: 1
((C_stevensi, D_gratus), (E_menziesii, (V_angasi, Lortiella)))


	        +-- C_stevensi 1 
	  +-----+                
	  |     +-- D_gratus 2   
	--+                      
	  |  +----- E_menziesii 3
	  +--+                   
	     |  +-- Lortiella 5  
	     +--+                
	        +-- V_angasi 4   
  

Roe & Hoeh, 2003

fig. 4.2: [clade_id: 2043] "Hyriinae" + Velesunioninae
spp. terminals: 3, direct-child terminals: 2
rigorous outgroups.: 2, direct-child rigorous outgroups: 1
((C_stevensi, V_angasi), (E_menziesii, L_rugata, D_gratus))


	     +-- C_stevensi 1 
	  +--+                
	  |  +-- V_angasi 2   
	--+                   
	  |  +-- D_gratus 5   
	  |  |                
	  +--+-- E_menziesii 3
	     |                
	     +-- L_rugata 4   
  

fig. 4.3: [clade_id: 2100] "Hyriinae" + Velesunioninae
spp. terminals: 3, direct-child terminals: 2
rigorous outgroups.: 2, direct-child rigorous outgroups: 1
((C_stevensi, D_gratus), (E_menziesii, (V_angasi, L_rugata)))


	        +-- C_stevensi 1 
	  +-----+                
	  |     +-- D_gratus 2   
	--+                      
	  |  +----- E_menziesii 3
	  +--+                   
	     |  +-- L_rugata 5   
	     +--+                
	        +-- V_angasi 4   
  

Konopleva et al., 2017

fig. 6: [clade_id: 8667] Hyriinae
spp. terminals: 3, direct-child terminals: 3
rigorous outgroups.: 0, direct-child rigorous outgroups: 0
(D_suavidicus, (T_corrugatus, C_ambigua))


	  +----- D_suavidicus 1
	--+                    
	  |  +-- C_ambigua 3   
	  +--+                 
	     +-- T_corrugatus 2
  

Chase et al., 2018

fig. 1: [clade_id: 13427] "Hyriinae" + Ambleminae + Margaritiferinae + Gonideinae + Unioninae
spp. terminals: 2, direct-child terminals: 0
rigorous outgroups.: 0, direct-child rigorous outgroups: 0
((E_menziesii, (C_monodonta, ((Q_quadrula, (L_powellii, L_siliquoidea)), (L_leaii, ((U_delphinus, U_tumidus), (A_anatina, (U_peninsularis, P_grandis))))))), ((C_monodonta, E_menziesii), ((L_leaii, (Q_quadrula, (L_powellii, L_siliquoidea))), ((U_delphinus, U_tumidus), (A_anatina, (U_peninsularis, P_grandis))))))


	     +-------------------- E_menziesii 1    
	     |                                      
	  +--+  +----------------- C_monodonta 2    
	  |  |  |                                   
	  |  |  |           +----- Q_quadrula 3     
	  |  +--+  +--------+                       
	  |     |  |        |  +-- L_powellii 4     
	  |     |  |        +--+                    
	  |     +--+           +-- L_siliquoidea 5  
	  |        |                                
	  |        |  +----------- L_leaii 6        
	  |        |  |                             
	  |        +--+        +-- U_delphinus 7    
	  |           |  +-----+                    
	--+           |  |     +-- U_tumidus 8      
	  |           +--+                          
	  |              |  +----- A_anatina 9      
	  |              +--+                       
	  |                 |  +-- P_grandis 11     
	  |                 +--+                    
	  |                    +-- U_peninsularis 10
	  |                                         
	  |                    +-- C_monodonta 12   
	  |        +-----------+                    
	  |        |           +-- E_menziesii 13   
	  |        |                                
	  |        |     +-------- L_leaii 14       
	  +--------+  +--+                          
	           |  |  |  +----- Q_quadrula 15    
	           |  |  +--+                       
	           |  |     |  +-- L_powellii 16    
	           |  |     +--+                    
	           +--+        +-- L_siliquoidea 17 
	              |                             
	              |        +-- U_delphinus 18   
	              |  +-----+                    
	              |  |     +-- U_tumidus 19     
	              +--+                          
	                 |  +----- A_anatina 20     
	                 +--+                       
	                    |  +-- P_grandis 22     
	                    +--+                    
	                       +-- U_peninsularis 21
  

Wu et al., 2020

fig. 2: [clade_id: 17755] "Hyriinae" + Velesunioninae
spp. terminals: 2, direct-child terminals: 2
rigorous outgroups.: 1, direct-child rigorous outgroups: 1
(T_corrugatus, (H_australis, V_ambiguus))


	  +----- T_corrugatus 1
	--+                    
	  |  +-- H_australis 2 
	  +--+                 
	     +-- V_ambiguus 3  
  

fig. 3: [clade_id: 17923] Hyriinae
spp. terminals: 1, direct-child terminals: 0
rigorous outgroups.: 0, direct-child rigorous outgroups: 0
unknown

fig. S1: [clade_id: 17822] "Hyriinae" + Velesunioninae
spp. terminals: 2, direct-child terminals: 2
rigorous outgroups.: 1, direct-child rigorous outgroups: 1
(T_corrugatus, (H_australis, V_ambiguus))


	  +----- T_corrugatus 1
	--+                    
	  |  +-- H_australis 2 
	  +--+                 
	     +-- V_ambiguus 3  
  

Sano et al., 2020

fig. 2: [clade_id: 18401] "Hyriinae" + Velesunioninae
spp. terminals: 2, direct-child terminals: 2
rigorous outgroups.: 1, direct-child rigorous outgroups: 1
(T_corrugatus, (H_australis, V_ambiguus))


	  +----- T_corrugatus 1
	--+                    
	  |  +-- H_australis 2 
	  +--+                 
	     +-- V_ambiguus 3  
  

fig. 3: [clade_id: 18554] "Hyriinae" + Velesunioninae
spp. terminals: 2, direct-child terminals: 2
rigorous outgroups.: 1, direct-child rigorous outgroups: 1
(T_corrugatus, (H_australis, V_ambiguus))


	  +----- T_corrugatus 1
	--+                    
	  |  +-- H_australis 2 
	  +--+                 
	     +-- V_ambiguus 3  
  

fig. 4: [clade_id: 18707] "Hyriinae" + Velesunioninae
spp. terminals: 2, direct-child terminals: 2
rigorous outgroups.: 1, direct-child rigorous outgroups: 1
(T_corrugatus, (H_australis, V_ambiguus))


	  +----- T_corrugatus 1
	--+                    
	  |  +-- H_australis 2 
	  +--+                 
	     +-- V_ambiguus 3  
  

Combosch et al., 2017

fig. 1: [clade_id: 9656] "Hyriinae" + Velesunioninae
spp. terminals: 2, direct-child terminals: 2
rigorous outgroups.: 1, direct-child rigorous outgroups: 1
(T_corrugatus, (V_ambiguus, H_australis))


	  +----- T_corrugatus 1
	--+                    
	  |  +-- H_australis 3 
	  +--+                 
	     +-- V_ambiguus 2  
  

fig. 2: [clade_id: 9680] Hyriinae
spp. terminals: 2, direct-child terminals: 2
rigorous outgroups.: 1, direct-child rigorous outgroups: 1
(T_corrugatus, H_australis)


	  +-- H_australis 2 
	--+                 
	  +-- T_corrugatus 1
  

Huang et al., 2018

fig. 3, 5: [clade_id: 10654] "Hyriinae" + Velesunioninae
spp. terminals: 2, direct-child terminals: 2
rigorous outgroups.: 1, direct-child rigorous outgroups: 1
(T_corrugatus, (V_ambiguus, H_australis))


	  +----- T_corrugatus 1
	--+                    
	  |  +-- H_australis 3 
	  +--+                 
	     +-- V_ambiguus 2  
  

fig. 6: [clade_id: 10735] Hyriinae
spp. terminals: 1, direct-child terminals: 0
rigorous outgroups.: 0, direct-child rigorous outgroups: 0
unknown

fig. 6-M: [clade_id: 10772] Hyriinae
spp. terminals: 1, direct-child terminals: 0
rigorous outgroups.: 0, direct-child rigorous outgroups: 0
unknown

Lopes-Lima et al., 2018

fig. 1: [clade_id: 10826] "Hyriinae" + Velesunioninae
spp. terminals: 2, direct-child terminals: 2
rigorous outgroups.: 1, direct-child rigorous outgroups: 1
(T_corrugatus, (V_ambiguus, H_australis))


	  +----- T_corrugatus 1
	--+                    
	  |  +-- H_australis 3 
	  +--+                 
	     +-- V_ambiguus 2  
  

fig. 3: [clade_id: 10879] "Hyriinae" + Velesunioninae
spp. terminals: 2, direct-child terminals: 2
rigorous outgroups.: 1, direct-child rigorous outgroups: 1
(T_corrugatus, (V_ambiguus, H_australis))


	  +----- T_corrugatus 1
	--+                    
	  |  +-- H_australis 3 
	  +--+                 
	     +-- V_ambiguus 2  
  

Sonowal et al., 2021

fig. 3a: [clade_id: 20147] "Hyriinae" + Velesunioninae
spp. terminals: 2, direct-child terminals: 2
rigorous outgroups.: 1, direct-child rigorous outgroups: 1
(T_corrugatus, H_australis, V_ambiguus)


	  +-- H_australis 2 
	  |                 
	--+-- T_corrugatus 1
	  |                 
	  +-- V_ambiguus 3  
  

fig. 3b: [clade_id: 20172] "Hyriinae" + Velesunioninae
spp. terminals: 2, direct-child terminals: 2
rigorous outgroups.: 1, direct-child rigorous outgroups: 1
(T_corrugatus, V_ambiguus, H_australis)


	  +-- H_australis 3 
	  |                 
	--+-- T_corrugatus 1
	  |                 
	  +-- V_ambiguus 2  
  

fig. 4: [clade_id: 20202] "Hyriinae" + Velesunioninae
spp. terminals: 2, direct-child terminals: 2
rigorous outgroups.: 1, direct-child rigorous outgroups: 1
(T_corrugatus, (V_ambiguus, H_australis))


	  +----- T_corrugatus 1
	--+                    
	  |  +-- H_australis 3 
	  +--+                 
	     +-- V_ambiguus 2  
  

Chase et al., 2018

fig. 4: [clade_id: 12855] Hyriinae
spp. terminals: 1, direct-child terminals: 0
rigorous outgroups.: 0, direct-child rigorous outgroups: 0
unknown

fig. 5: [clade_id: 12910] Hyriinae
spp. terminals: 1, direct-child terminals: 0
rigorous outgroups.: 0, direct-child rigorous outgroups: 0
unknown

Wu et al., 2019

fig. 3: [clade_id: 13671] Hyriinae
spp. terminals: 1, direct-child terminals: 0
rigorous outgroups.: 0, direct-child rigorous outgroups: 0
unknown

Lopes-Lima et al., 2017

fig. 1-2: [clade_id: 8821] Hyriinae
spp. terminals: 1, direct-child terminals: 1
rigorous outgroups.: 1, direct-child rigorous outgroups: 1
unknown

Breton et al., 2011

fig. 3: [clade_id: 4983] Hyriinae
spp. terminals: 1, direct-child terminals: 0
rigorous outgroups.: 0, direct-child rigorous outgroups: 0
unknown

Guerra et al., 2017

fig. 2a: [clade_id: 9806] Hyriinae
spp. terminals: 1, direct-child terminals: 0
rigorous outgroups.: 0, direct-child rigorous outgroups: 0
unknown

fig. 2a-M: [clade_id: 9824] Hyriinae
spp. terminals: 1, direct-child terminals: 0
rigorous outgroups.: 0, direct-child rigorous outgroups: 0
unknown

fig. 2b: [clade_id: 9870] Hyriinae
spp. terminals: 1, direct-child terminals: 0
rigorous outgroups.: 0, direct-child rigorous outgroups: 0
unknown

fig. 2b-M: [clade_id: 9890] Hyriinae
spp. terminals: 1, direct-child terminals: 0
rigorous outgroups.: 0, direct-child rigorous outgroups: 0
unknown

Graf & Ó Foighil, 2000

fig. 2A: [clade_id: 79] Hyriinae
spp. terminals: 1, direct-child terminals: 1
rigorous outgroups.: 0, direct-child rigorous outgroups: 0
unknown

fig. 2B: [clade_id: 143] Hyriinae
spp. terminals: 1, direct-child terminals: 1
rigorous outgroups.: 0, direct-child rigorous outgroups: 0
unknown

Pfeiffer et al., 2019

fig. 4-5: [clade_id: 15084] Hyriinae
spp. terminals: 1, direct-child terminals: 1
rigorous outgroups.: 0, direct-child rigorous outgroups: 0
unknown

fig. 6: [clade_id: 15141] Hyriinae
spp. terminals: 1, direct-child terminals: 1
rigorous outgroups.: 0, direct-child rigorous outgroups: 0
unknown

Lopes-Lima et al., 2020

fig. 2-4: [clade_id: 16614] Hyriinae
spp. terminals: 1, direct-child terminals: 1
rigorous outgroups.: 1, direct-child rigorous outgroups: 1
unknown

Hoeh et al., 2002

fig. 1-F: [clade_id: 1967] Hyriinae
spp. terminals: 1, direct-child terminals: 0
rigorous outgroups.: 0, direct-child rigorous outgroups: 0
unknown

fig. 1-M: [clade_id: 1950] Hyriinae
spp. terminals: 1, direct-child terminals: 0
rigorous outgroups.: 0, direct-child rigorous outgroups: 0
unknown

Klunzinger et al., 2020

fig. 2: [clade_id: 16321] Hyriinae
spp. terminals: 1, direct-child terminals: 1
rigorous outgroups.: 2, direct-child rigorous outgroups: 1
unknown

Page updated: 09:21:07 Tue 04 Jan 2022

 
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