|
Bivalvia | Unionoida | Unionoidea | Unionidae | Unioninae tribe UNIONINI Rafinesque, 1820: cladomics
13 genera
Cladomics Summary
number of species = 92
number of subtaxa (rank: genus) = 13
analyzed in 221 cladograms in 130 publications
maximum number of terminals in an analysis = 26, mean = 6.12
maximum number of subtaxa (genus) in an analysis = 10, mean = 2.67
Basic Monophyly
number of cladograms that test basic monophyly (i.e., more than 1 included species) = 183
maximum number of included terminals = 26, mean = 7.18
maximum number of subtaxa (genus) = 10, mean = 3.02
proportion of cladograms that support basic monophyly = 0.85
Rigorous Monophyly
number of cladograms that test rigorous monophyly = 99
(Rigorous monophyly refers to cladograms with terminal species representing more than one subtaxon (genus) taxon and at least one rigorous outgroup (tribe).
maximum number of included terminals = 26, mean = 9.14
maximum number of subtaxa (genus) = 10, mean = 4.52
proportion of cladograms that support rigorous monophyly = 0.76
Pairwise Cladogram Clustering
The graphs below depict in two dimensions all pair-wise ingroup distances within the cladome. See the Cladmoics FARQ for more information.
Chen et al., 2023 [Publ 4302], fig. 1
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 17 |
17 |
14 |
10 |
23 |
14 |
4 |
0.0 |
*** Top Monophyly Cladogram ***
+----- Pletholophus_tenuis
+--------------------+
| | +-- Sinanodonta_jourdyi
| +--+
| +-- Beringiana_beringiana
|
| +----------------------- Lanceolaria_gladiola
| |
| +----------------------- Acuticosta_chinensis
--+ |
| +----------------------- Lepidodesma_languilati
| |
| +----------------------- Lanceolaria_oxyrhyncha
| |
| +----------------------- Cristaria_plicata
| |
| | +-- Alasmidonta_marginata
+--+ +--+
| | +-- Lasmigona_costata
+-----------------+
| | +-- Anodonta_anatina
| +--+
| +-- Pseudanodonta_complanata
|
| +-------------------- Aculamprotula_fibrosa **
| |
| | +----- Middendorffinaia_mongolica **
| | +-----------+
+--+ | | +-- Nodularia_douglasiae **
| | +--+
| | +-- Nodularia_breviconcha **
| |
| | +-- Cuneopsis_celtiformis **
+--+ +-----------+
| | +-- Cuneopsis_heudei **
| |
| | +-- Diaurora_aurorea **
| | +--+
| | | +-- Diaurora_laeve **
| +--------+
+--+ | +-- Schistodesmus_lampreyanus **
| +--+
| +-- Schistodesmus_spinosus **
|
| +-- Pseudocuneopsis_sichuanensis **
| +--------+
| | +-- Pseudocuneopsis_capitata **
| |
+--+ +-- Inversiunio_yanagawensis **
| +-----+
| | +-- Inversiunio_reinianus **
+--+
| +----- Tchangsinaia_piscicula **
+--+
| +-- Unio_pictorum **
+--+
+-- Unio_crassus **
Lopes-Lima et al., 2020 [Publ 4080], fig. 2-4
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 26 |
26 |
22 |
8 |
96 |
43 |
4 |
25.0 |
*** Top Sister-Group Cladogram ***
+-------------------------------- Acuticosta_chinensis
|
+-------------------------------- Lepidodesma_languilati
|
| +----------- Aculamprotula_tortuosa **
| +--------------+
| | | +-------- Aculamprotula_scripta **
| | +--+
| | | +----- Aculamprotula_coreana **
| | +--+
| | | +-- Aculamprotula_fibrosa **
| | +--+
+-----+ +-- Aculamprotula_tientsinensis **
| |
| | +----------------------- Middendorffinaia_mongolica **
| | |
| | | +----- Unio_gibbus **
| | | |
| | | +--------------+----- Unio_tumidus **
| +--+ | |
| | | | +-- Unio_crassus **
| | | +--+
| | | +-- Unio_pictorum **
| | |
| | | +-------------- Nodularia_breviconcha **
| +--+ +--+
| | | | +----------- Nodularia[2] **
| | | +--+
| | | | +-------- Nodularia_nuxpersicae **
| | | +--+
| | | | +----- Nodularia_nipponensis **
| | | +--+
| | | | +-- Nodularia_douglasiae **
| +--+ +--+
| | +-- Nodularia[1] **
| |
--+ | +-- Schistodesmus_lampreyanus **
| | +-----------+
| | | +-- Schistodesmus_spinosus **
| | |
| | | +----- Cuneopsis_rufescens **
| | +--------+
| +--+ | +-- Cuneopsis_heudei **
| | +--+
| | +-- Cuneopsis_celtiformis **
| |
| | +----------- Tchangsinaia_piscicula **
| +--+
| | +-------- Inversiunio_jokohamensis **
| +--+
| | +----- Inversiunio_verrucosus **
| +--+
| | +-- Inversiunio_reinianus **
| +--+
| +-- Inversiunio_yanagawensis **
|
| +----------------- Lanceolaria_oxyrhyncha
| |
| +-----------+ +-------------- Lanceolaria_gladiola
| | | |
| | +--+ +-- Lanceolaria_bilirata
| | | +--------+
| | | | +-- Lanceolaria_grayii
| | +--+
| | | +-------- Lanceolaria_acrorrhyncha
| | +--+
| | | +----- Lanceolaria_kihirai
| | +--+
| | | +-- Lanceolaria_lanceolata
| | +--+
| | +-- Lanceolaria_triformis
| |
| | +-------- Pseudanodonta_complanata
+--+ |
| | +----- Anodonta_cygnea
| +--+
| | | +-- Anodonta_anatina
| +-----------------+ +--+
| | | +-- Anodonta_nuttalliana
| | |
| | | +----- Pyganodon_grandis
| | | |
| | | +----- Simpsonaias_ambigua
| | +--+
| | +----- Strophitus_undulatus
| | |
| | | +-- Alasmidonta_marginata
| | +--+
| | +-- Lasmigona_costata
+--+
| +-- Cristaria_plicata
| +--------------------+
| | +-- Cristaria_clessini
| |
| | +-------------- Anemina_arcaeformis
| | |
| +--------+ +----------- Amuranodonta_kijaensis
| | | |
| | +--+ +-------- Buldowskia_suifunica
| | | |
| | +--+ +-- Buldowskia_flavotincta
+--+ | +--+
| | | +-- Buldowskia_shadini
| +--+
| | +-- Buldowskia_iwakawai
| +--+
| +-- Buldowskia_kamiyai
|
| +-------------------- Pletholophus_tenuis
| |
| | +-------- Beringiana_japonica
| | +--------+
+--+ | | +----- Beringiana_beringiana
| | +--+
| | | +-- Beringiana_fukuharai
| | +--+
+--+ +-- Beringiana_gosannensis
|
| +-- Sinanodonta_angula
| +-----------+
| | +-- Sinanodonta_lucida
| |
+--+ +----------- Sinanodonta_schrenkii
| |
| | +-- Sinanodonta_calipygos
| +--------+
+--+ +-- Sinanodonta_tumens
|
| +-------- Sinanodonta_woodiana[2]
| |
+--+ +----- Sinanodonta_elliptica
| |
+--+----- Sinanodonta_jourdyi
|
| +-- Sinanodonta_lauta
+--+
+-- Sinanodonta_woodiana[1]
Wu et al., 2022 [Publ 4238], fig. 2d
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 23 |
23 |
21 |
8 |
16 |
16 |
4 |
22.0 |
+-------------------------------- Lepidodesma_languilati
|
| +----- Lanceolaria_gladiola
| +--------------+
| | | +-- Lanceolaria_grayii
| | +--+
| | +-- Lanceolaria_lanceolata
| +--------+
| | | +----------------- Acuticosta_chinensis
| | | |
| | | | +-------- Cristaria_plicata
| | | | |
--+ | +--+ +-----+ +-- Sinanodonta_woodiana
| | | | | +--+
| | | | | | +-- Sinanodonta_lucida
| | | | +--+
| | | | | +-- Anemina_arcaeformis
| | +--+ +--+
| | | +-- Anemina_euscaphys
| | |
| | | +-- Anodonta_cygnea
| | | +--------+
| | | | +-- Anodonta_anatina
| | +--+
+--+ | +-------- Platynaias_compressa
| +--+
| | +----- Pyganodon_grandis
| +--+
| | +-- Utterbackia_imbecillis
| +--+
| +-- Utterbackia_peninsularis
|
| +-------- Aculamprotula_tortuosa **
| |
| +-----------------+ +-- Aculamprotula_tientsinensis **
| | | +--+
| | | | +-- Aculamprotula_coreana **
| | +--+
| | | +-- Aculamprotula_polysticta **
| | +--+
| | +-- Aculamprotula_scripta **
+--+
| +-------- Unio_tumidus **
| +--------------+
| | | +----- Unio_crassus **
| | +--+
| | | +-- Unio_pictorum **
| | +--+
| | +-- Unio_delphinus **
+--+
| +-- Pseudocuneopsis_capitata **
| +-----------------+
| | +-- Pseudocuneopsis_sichuanensis **
| |
| | +-- Nodularia_breviconcha **
+--+ +--------------+
| | +-- Nodularia_douglasiae **
| |
| | +-- Schistodesmus_lampreyanus **
+--+ +-----------+
| | +-- Schistodesmus_spinosus **
| |
| | +-------- Cuneopsis_rufescens **
+--+ +--+
| | | +----- Cuneopsis_celtiformis **
| | +--+
| | | +-- Cuneopsis_heudei **
+--+ +--+
| +-- Cuneopsis_demangei **
|
| +-------- Tchangsinaia_piscicula **
+--+
| +----- Inversiunio_jokohamensis **
+--+
| +-- Inversiunio_reinianus **
+--+
+-- Inversiunio_yanagawensis **
Lyubas et al., 2022 [Publ 4255], fig. 7
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 23 |
23 |
20 |
1 |
8 |
5 |
1 |
36.0 |
+----------- Pseudanodonta_complanata
+--------------------+
| | +-------- Anodonta_anatina
| +--+
| | +----- Anodonta_cygnea
| +--+
| | +-- Anodonta_exulcerata
| +--+
--+ +-- Anodonta_seddoni
|
| +----------------------------- Unio_tumidus **
| |
| | +-- Unio_durieui **
| | +-----------------------+
| | | +-- Unio_gibbus **
+--+ |
| | +-------------- Unio_damascensis **
| | |
| | +-------------- Unio_sesirmensis **
| | +--------+
+--+ | | +----------- Unio_ionicus **
| | | |
| | +--+ +-- Unio_bruguierianus **
| | | +-----+
| | | | +-- Unio_tumidiformis **
| | +--+
| | | +----- Unio_crassus[2] **
+--+ +--+
| | +-- Unio_carneus **
| +--+
| +-- Unio_crassus[1] **
|
| +----- Unio_caffer **
| +--------------+
| | | +-- Unio_terminalis **
| | +--+
+--+ +-- Unio_delicatus **
|
| +----------------- Unio_pictorum **
| |
+--+ +-------------- Unio_eucirrus **
| |
+--+ +-- Unio_elongatulus **
| +--------+
| | +-- Unio_mancus **
+--+
| +-------- Unio_ravoisieri **
| |
+--+ +-- Unio_delphinus **
| +--+
| | +-- Unio_foucauldianus **
+--+
| +-- Unio_hueti **
+--+
+-- Unio_tigridis **
Lyubas et al., 2022 [Publ 4255], fig. S1
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 23 |
23 |
14 |
1 |
8 |
5 |
1 |
36.0 |
+----------- Anodonta_anatina
+--------+
| | +-------- Pseudanodonta_complanata
| +--+
| | +----- Anodonta_exulcerata
| +--+
| | +-- Anodonta_seddoni
| +--+
| +-- Anodonta_cygnea
|
--+ +----------------- Unio_tumidus **
| |
| | +-- Unio_durieui **
| +--------------+
| | +-- Unio_gibbus **
| |
| | +----------- Unio_sesirmensis **
| | |
| | +--+ +-------- Unio_damascensis **
+--+ | | |
| | | | +----- Unio_ionicus **
| | +--+ |
| | | | +-- Unio_bruguierianus **
| | | +--+
| | +--+ +-- Unio_tumidiformis **
| | |
| | | +-- Unio_carneus **
| | | |
| | +--+-- Unio_crassus[1] **
+--+ |
| +-- Unio_crassus[2] **
|
| +-------- Unio_elongatulus **
| |
| +-------- Unio_mancus **
| |
| | +-- Unio_pictorum **
| +-----+
| | +-- Unio_eucirrus **
| |
+-----+ +----- Unio_caffer **
+--+
| | +-- Unio_terminalis **
| +--+
| +-- Unio_delicatus **
|
| +----- Unio_ravoisieri **
| |
| | +-- Unio_delphinus **
+--+--+
| +-- Unio_foucauldianus **
|
| +-- Unio_hueti **
+--+
+-- Unio_tigridis **
Wu et al., 2023 [Publ 4339], fig. 4, S5-nt
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 23 |
23 |
20 |
7 |
74 |
25 |
4 |
26.0 |
+-- Lepidodesma_aligera
+-----------------------------+
| +-- Lepidodesma_languilati
|
| +-------- Acuticosta_chinensis
| +--------------+
| | | +----- Lanceolaria_gladiola
| | +--+
| | | +-- Lanceolaria_grayii
| | +--+
| | +-- Lanceolaria_lanceolata
| |
| | +-------------- Cristaria_plicata
| +-----+ |
| | | +-----+ +-- Anemina_arcaeformis
| | | | | +--------+
| | | | | | +-- Anemina_euscaphys
| | | | +--+
--+ | | | | +-------- Beringiana_fukuharai
| | | | +--+
| | | | | +----- Sinanodonta_lucida
| | | | +--+
| | +--+ | +-- Sinanodonta_tumens
| | | +--+
| | | +-- Sinanodonta_woodiana
| | |
| | | +----------------- Anodonta_nuttalliana
| | | |
| | | +----------------- Pseudanodonta_complanata
| | | |
| | +--+ +-------------- Anodonta_cygnea
| | | |
| | | | +-- Anodonta_exulcerata
| | | +-----------+
| | | | +-- Anodonta_anatina
| | +--+
+--+ | +----- Pyganodon_grandis
| | +-----+
| | | | +-- Utterbackia_imbecillis
| | | +--+
| +--+ +-- Utterbackia_peninsularis
| |
| | +-------- Lasmigona_complanata
| +--+
| | +----- Prolasmidonta_heterodon
| +--+
| | +-- Alasmidonta_varicosa
| +--+
| +-- Platynaias_compressa
|
| +-------- Aculamprotula_tortuosa **
| |
| +-----------------+ +-- Aculamprotula_coreana **
| | | +--+
| | | | +-- Aculamprotula_tientsinensis **
| | +--+
| | | +-- Aculamprotula_polysticta **
| | +--+
| | +-- Aculamprotula_scripta **
| |
+--+ +----------- Unio_tumidus **
| |
| +-----------+ +-------- Unio_crassus **
| | | |
| | +--+ +----- Unio_elongatulus **
| | | |
| | +--+ +-- Unio_delphinus **
| | | |
| | +--+-- Unio_mancus **
+--+ |
| +-- Unio_pictorum **
|
| +-- Pseudocuneopsis_capitata **
| +-----------------+
| | +-- Pseudocuneopsis_sichuanensis **
| |
+--+ +-- Nodularia_breviconcha **
| +--------------+
| | +-- Nodularia_nipponensis **
| |
+--+ +----- Tchangsinaia_piscicula **
| +--------+
| | | +-- Schistodesmus_lampreyanus[2] **
| | +--+
+--+ +-- Schistodesmus_spinosus **
|
| +----------- Schistodesmus_lampreyanus[1] **
+--+
| +-------- Cuneopsis_rufescens **
+--+
| +----- Cuneopsis_celtiformis **
+--+
| +-- Cuneopsis_demangei **
+--+
+-- Cuneopsis_heudei **
Wu et al., 2023 [Publ 4339], fig. 4, S5-aa
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 22 |
22 |
20 |
7 |
74 |
25 |
4 |
25.0 |
+-- Lepidodesma_aligera
+--------------------------+
| +-- Lepidodesma_languilati
|
| +-------- Aculamprotula_tortuosa **
| |
| +--------------+ +-- Aculamprotula_coreana **
| | | +--+
| | | | +-- Aculamprotula_tientsinensis **
| | +--+
| | | +-- Aculamprotula_polysticta **
| | +--+
| | +-- Aculamprotula_scripta **
| |
| +--+ +-------------- Unio_tumidus **
| | | +-----+
| | | | | +----------- Unio_crassus **
| | | | +--+
--+ | | | | +-------- Unio_elongatulus **
| | | | +--+
| | | | | +----- Unio_pictorum **
| | | | +--+
| | | | | +-- Unio_delphinus **
| | +--+ +--+
| | | +-- Unio_mancus **
| | |
| | | +-- Nodularia_breviconcha **
| | | +--+
| | | | +-- Nodularia_nipponensis **
| | | +-----------+
| | | | | +-- Pseudocuneopsis_capitata **
| | | | +--+
| | +--+ +-- Pseudocuneopsis_sichuanensis **
| | |
| | | +-------------- Tchangsinaia_piscicula **
| | | |
+--+ +--+ +-- Schistodesmus_lampreyanus **
| | +--------+
| | | +-- Schistodesmus_spinosus **
| +--+
| | +-------- Cuneopsis_rufescens **
| +--+
| | +----- Cuneopsis_celtiformis **
| +--+
| | +-- Cuneopsis_demangei **
| +--+
| +-- Cuneopsis_heudei **
|
| +-------- Acuticosta_chinensis
| +--------+
| | | +----- Lanceolaria_gladiola
| | +--+
| | | +-- Lanceolaria_grayii
| | +--+
| | +-- Lanceolaria_lanceolata
| |
| | +----- Cristaria_plicata
| | +-----+
| | | | +-- Anemina_arcaeformis
+--------+ | +--+
| +--+ +-- Anemina_euscaphys
| | |
| | | +-------- Beringiana_fukuharai
| | +--+
| | | +----- Sinanodonta_lucida
| | +--+
| | | +-- Sinanodonta_tumens
| | +--+
| | +-- Sinanodonta_woodiana
+--+
| +-- Anodonta_nuttalliana
| +-----+
| | +-- Pseudanodonta_complanata
| +--+
| | | +----- Anodonta_anatina
| | +--+
| | | +-- Anodonta_cygnea
| | +--+
| | +-- Anodonta_exulcerata
+--+
| +----- Pyganodon_grandis
| +--+
| | | +-- Utterbackia_peninsularis
| | +--+
| | +-- Utterbackia_imbecillis
+--+
| +-- Alasmidonta_varicosa
| +--+
| | +-- Platynaias_compressa
+--+
| +-- Prolasmidonta_heterodon
+--+
+-- Lasmigona_complanata
Wu et al., 2024 [Publ 4350], fig. 8a, S1
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 22 |
22 |
19 |
7 |
70 |
25 |
4 |
22.0 |
+-- Lepidodesma_aligera
+--------------------------+
| +-- Lepidodesma_languilati
|
| +-------- Aculamprotula_tortuosa **
| |
| +--------------+ +-- Aculamprotula_coreana **
| | | +--+
| | | | +-- Aculamprotula_tientsinensis **
| | +--+
| | | +-- Aculamprotula_polysticta **
| | +--+
| | +-- Aculamprotula_scripta **
| |
| +--+ +-------------- Unio_tumidus **
| | | +-----+
| | | | | +----------- Unio_crassus **
| | | | +--+
--+ | | | | +-------- Unio_elongatulus **
| | | | +--+
| | | | | +----- Unio_pictorum **
| | | | +--+
| | | | | +-- Unio_delphinus **
| | +--+ +--+
| | | +-- Unio_mancus **
| | |
| | | +-- Nodularia_breviconcha **
| | | +--------------+
| | | | +-- Nodularia_douglasiae **
| | | |
| | | | +-- Pseudocuneopsis_capitata **
| | | +--------------+
| | +--+ +-- Pseudocuneopsis_sichuanensis **
| | |
| | | +-- Schistodesmus_lampreyanus **
| | | +-----------+
+--+ | | +-- Schistodesmus_spinosus **
| +--+
| | +----------- Tchangsinaia_piscicula **
| +--+
| | +-------- Cuneopsis_rufescens **
| +--+
| | +----- Cuneopsis_celtiformis **
| +--+
| | +-- Cuneopsis_demangei **
| +--+
| +-- Cuneopsis_heudei **
|
| +-------- Acuticosta_chinensis
| +--------------+
| | | +----- Lanceolaria_gladiola
| | +--+
| | | +-- Lanceolaria_grayii
| | +--+
| | +-- Lanceolaria_lanceolata
| |
| | +-------------- Cristaria_plicata
| | |
+--+ +-----+ +-- Anemina_arcaeformis
| | | +--------+
| | | | +-- Anemina_euscaphys
| | +--+
| | | +-------- Beringiana_fukuharai
| | +--+
| | | +----- Sinanodonta_lucida
| | +--+
| | | +-- Sinanodonta_tumens
+--+ +--+
| +-- Sinanodonta_woodiana
|
| +-------- Pseudanodonta_complanata
| +--------+
| | | +----- Anodonta_anatina
| | +--+
| | | +-- Anodonta_cygnea
| | +--+
+--+ +-- Anodonta_exulcerata
|
| +-------------- Anodonta_nuttalliana
| |
| | +----- Pyganodon_grandis
+--+ +-----+
| | | +-- Utterbackia_imbecillis
| | +--+
+--+ +-- Utterbackia_peninsularis
|
| +-------- Lasmigona_complanata
+--+
| +----- Prolasmidonta_heterodon
+--+
| +-- Alasmidonta_varicosa
+--+
+-- Platynaias_compressa
Wu et al., 2024 [Publ 4350], fig. 8b, S2
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 22 |
22 |
19 |
7 |
70 |
25 |
4 |
22.0 |
+-- Lepidodesma_aligera
+--------------------------+
| +-- Lepidodesma_languilati
|
| +-------- Aculamprotula_tortuosa **
| |
| +--------------+ +-- Aculamprotula_coreana **
| | | +--+
| | | | +-- Aculamprotula_tientsinensis **
| | +--+
| | | +-- Aculamprotula_polysticta **
| | +--+
| | +-- Aculamprotula_scripta **
| |
| +--+ +-------------- Unio_tumidus **
| | | +-----+
| | | | | +----------- Unio_crassus **
| | | | +--+
--+ | | | | +-------- Unio_elongatulus **
| | | | +--+
| | | | | +----- Unio_pictorum **
| | | | +--+
| | | | | +-- Unio_delphinus **
| | +--+ +--+
| | | +-- Unio_mancus **
| | |
| | | +-- Nodularia_breviconcha **
| | | +--------------+
| | | | +-- Nodularia_douglasiae **
| | | |
| | | | +-- Pseudocuneopsis_capitata **
| | | +--------------+
| | +--+ +-- Pseudocuneopsis_sichuanensis **
| | |
| | | +-- Schistodesmus_lampreyanus **
+--+ | +-----------+
| | | +-- Schistodesmus_spinosus **
| +--+
| | +----------- Tchangsinaia_piscicula **
| +--+
| | +-------- Cuneopsis_rufescens **
| +--+
| | +----- Cuneopsis_celtiformis **
| +--+
| | +-- Cuneopsis_demangei **
| +--+
| +-- Cuneopsis_heudei **
|
| +-------- Acuticosta_chinensis
| +--------------+
| | | +----- Lanceolaria_gladiola
| | +--+
| | | +-- Lanceolaria_grayii
| | +--+
| | +-- Lanceolaria_lanceolata
| |
| | +-------------- Cristaria_plicata
+--+ |
| +-----+ +-- Anemina_arcaeformis
| | | +--------+
| | | | +-- Anemina_euscaphys
| | +--+
| | | +-------- Beringiana_fukuharai
| | +--+
| | | +----- Sinanodonta_lucida
| | +--+
+--+ | +-- Sinanodonta_tumens
| +--+
| +-- Sinanodonta_woodiana
|
| +----------------- Anodonta_nuttalliana
| |
| | +-------- Pseudanodonta_complanata
| | +-----+
+--+ | | +----- Anodonta_anatina
| | +--+
| | | +-- Anodonta_cygnea
| | +--+
+--+ +-- Anodonta_exulcerata
|
| +----- Pyganodon_grandis
| +-----+
| | | +-- Utterbackia_imbecillis
| | +--+
+--+ +-- Utterbackia_peninsularis
|
| +-------- Lasmigona_complanata
+--+
| +----- Prolasmidonta_heterodon
+--+
| +-- Alasmidonta_varicosa
+--+
+-- Platynaias_compressa
Wu et al., 2022 [Publ 4238], fig. 3
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 20 |
20 |
18 |
7 |
54 |
16 |
4 |
21.0 |
+-------------------------------- Lepidodesma_languilati
|
| +-------- Acuticosta_chinensis
| +--------+
| | | +----- Lanceolaria_gladiola
| | +--+
| | | +-- Lanceolaria_grayii
| | +--+
| | +-- Lanceolaria_lanceolata
| |
| +-----------+ +-------- Cristaria_plicata
| | | |
--+ | | +-----+ +-- Sinanodonta_woodiana
| | | | | +--+
| | | | | | +-- Sinanodonta_lucida
| | | | +--+
| | | | | +-- Anemina_arcaeformis
| | +--+ +--+
| | | +-- Anemina_euscaphys
| | |
| | | +-- Anodonta_cygnea
| | | +--------+
| | | | +-- Anodonta_anatina
| | +--+
| | | +-------- Platynaias_compressa
+--+ +--+
| | +----- Pyganodon_grandis
| +--+
| | +-- Utterbackia_imbecillis
| +--+
| +-- Utterbackia_peninsularis
|
| +-------- Aculamprotula_tortuosa **
| |
| +-----------------+ +-- Aculamprotula_tientsinensis **
| | | +--+
| | | | +-- Aculamprotula_coreana **
| | +--+
| | | +-- Aculamprotula_polysticta **
| | +--+
| | +-- Aculamprotula_scripta **
+--+
| +-------- Unio_tumidus **
| +--------------+
| | | +----- Unio_crassus **
| | +--+
| | | +-- Unio_pictorum **
| | +--+
+--+ +-- Unio_delphinus **
|
| +-- Pseudocuneopsis_capitata **
| +-----------------+
| | +-- Pseudocuneopsis_sichuanensis **
| |
+--+ +-- Nodularia_douglasiae **
| +--------------+
| | +-- Nodularia_breviconcha **
| |
+--+ +-- Schistodesmus_lampreyanus **
| +-----------+
| | +-- Schistodesmus_spinosus **
+--+
| +----------- Tchangsinaia_piscicula **
+--+
| +-------- Cuneopsis_rufescens **
+--+
| +----- Cuneopsis_celtiformis **
+--+
| +-- Cuneopsis_heudei **
+--+
+-- Cuneopsis_demangei **
Huang et al., 2019 [Publ 3843], fig. 2
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 18 |
18 |
15 |
6 |
186 |
27 |
4 |
22.0 |
+-------------------------------- Acuticosta_chinensis
|
| +----------------------------- Lepidodesma_languilati
| |
| | +-------- Aculamprotula_tortuosa **
| | |
| | +-----------+ +----- Aculamprotula_coreana **
| | | | |
| | | +--+----- Aculamprotula_scripta **
| | | |
| | | | +-- Aculamprotula_fibrosa **
| | | +--+
| | | +-- Aculamprotula_tientsinensis **
--+ +--------+
| | | +-------- Unio_tumidus **
| | | +--------+
| | | | | +----- Unio_crassus **
| | | | +--+
| | | | | +-- Unio_delphinus **
| | | | +--+
| | +--+ +-- Unio_pictorum **
| | |
| | | +-- Schistodesmus_lampreyanus **
| | | +-----------+
| | | | +-- Schistodesmus_spinosus **
| | | |
| | +--+ +----- Nodularia_douglasiae[1] **
+--+ | +-----+
| | | | +-- Nodularia_douglasiae[2] **
| | | +--+
| +--+ +-- Nodularia_nuxpersicae **
| |
| | +-------- Tchangsinaia_piscicula **
| +--+
| | +----- Cuneopsis_rufescens **
| +--+
| | +-- Cuneopsis_heudei **
| +--+
| +-- Cuneopsis_celtiformis **
|
| +-------- Lanceolaria_gladiola
| |
| +-----------------+ +----- Lanceolaria_grayii[1]
| | | |
| | +--+----- Lanceolaria_grayii[2]
| | |
| | | +-- Lanceolaria_lanceolata
| | +--+
| | +-- Lanceolaria_triformis
| |
| | +-------------------- Anemina_arcaeformis
| | |
| | +--+ +----------------- Cristaria_plicata
+--+ | | |
| | +--+ +-------------- Pletholophus_tenuis
| | | |
| | +--+ +-- Sinanodonta_angula
| | | +--------+
| | | | +-- Sinanodonta_lucida
| | +--+
| | | +-------- Sinanodonta_woodiana[2]
| | +--+
| | | +----- Sinanodonta_jourdyi
| | +--+
+--+ | +-- Sinanodonta_elliptica
| +--+
| +-- Sinanodonta_woodiana[1]
|
| +-------- Alasmidonta_marginata
| |
| +-------- Lasmigona_costata
| |
| +-------- Pseudanodonta_complanata
| |
| +-------- Simpsonaias_ambigua
| |
+--------------+-------- Strophitus_undulatus
|
| +----- Anodonta_anatina
+--+
| | +-- Anodonta_cygnea
| +--+
| +-- Anodonta_nuttalliana
|
| +----- Pyganodon_grandis
+--+
| +-- Utterbackia_imbecillis
+--+
+-- Utterbackia_peninsularis
Huang et al., 2019 [Publ 3843], fig. S4
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 18 |
44 |
21 |
6 |
184 |
26 |
4 |
48.0 |
+----------------- Acuticosta_chinensis
|
+----------------- Alasmidonta_marginata
|
+----------------- Anodonta_anatina
|
+----------------- Anemina_arcaeformis
|
+----------------- Anodonta_cygnea
|
+----------------- Cristaria_plicata
|
+----------------- Anodonta_nuttalliana
|
+----------------- Lasmigona_costata
|
+----------------- Lepidodesma_languilati
|
+----------------- Pseudanodonta_complanata
|
+----------------- Pyganodon_grandis
|
+----------------- Simpsonaias_ambigua
|
+----------------- Strophitus_undulatus
|
| +-- Schistodesmus_lampreyanus **
+--------------+
| +-- Schistodesmus_spinosus **
|
| +-- Utterbackia_imbecillis
+--------------+
| +-- Utterbackia_peninsularis
|
| +-------- Lanceolaria_gladiola
+--------+
| | +----- Lanceolaria_grayii
| +--+
--+ | +-- Lanceolaria_lanceolata
| +--+
| +-- Lanceolaria_triformis
|
| +-------- Unio_tumidus **
+--------+
| | +----- Unio_crassus **
| +--+
| | +-- Unio_delphinus **
| +--+
| +-- Unio_pictorum **
|
| +-------- Aculamprotula_tortuosa **
| |
+--------+ +-- Aculamprotula_coreana **
| | +--+
| | | +-- Aculamprotula_scripta **
| +--+
| | +-- Aculamprotula_fibrosa **
| +--+
| +-- Aculamprotula_tientsinensis **
|
| +-------- Cuneopsis_heudei **
| |
| +-------- Cuneopsis_celtiformis **
| |
+--------+-------- Tchangsinaia_piscicula **
| |
| +-------- Cuneopsis_rufescens **
| |
| | +----- Nodularia_douglasiae[2] **
| +--+
| | +-- Nodularia_douglasiae[1] **
| +--+
| +-- Nodularia_nuxpersicae **
|
| +-------------- Pletholophus_tenuis
| |
+--+ +-- Sinanodonta_angula
| +--------+
| | +-- Sinanodonta_lucida
+--+
| +-------- Sinanodonta_woodiana[1]
+--+
| +----- Sinanodonta_jourdyi
+--+
| +-- Sinanodonta_elliptica
+--+
+-- Sinanodonta_woodiana[2]
Huang et al., 2019 [Publ 3843], fig. S3
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 17 |
17 |
15 |
6 |
184 |
26 |
4 |
23.0 |
+----------------------------------- Acuticosta_chinensis
|
| +-------- Aculamprotula_tortuosa **
| |
| +-----------+ +-- Aculamprotula_coreana **
| | | +--+
| | | | +-- Aculamprotula_scripta **
| | +--+
| | | +-- Aculamprotula_fibrosa **
| | +--+
| | +-- Aculamprotula_tientsinensis **
| +-----------+
| | | +-------- Unio_tumidus **
--+ | | +--------+
| | | | | +----- Unio_crassus **
| | | | +--+
| | | | | +-- Unio_delphinus **
| | | | +--+
| | +--+ +-- Unio_pictorum **
| | |
| | | +----- Nodularia_douglasiae[1] **
| | | +--------+
| | | | | +-- Nodularia_douglasiae[2] **
| | | | +--+
| | +--+ +-- Nodularia_nuxpersicae **
| | |
+--+ | +----------- Schistodesmus_lampreyanus **
| +--+
| | +-------- Tchangsinaia_piscicula **
| +--+
| | +----- Cuneopsis_rufescens **
| +--+
| | +-- Cuneopsis_heudei **
| +--+
| +-- Cuneopsis_celtiformis **
|
| +----------------------------- Lepidodesma_languilati
| |
| | +-------- Lanceolaria_gladiola
| | +-----------------+
| | | | +----- Lanceolaria_grayii
| | | +--+
+--+ | | +-- Lanceolaria_lanceolata
| | +--+
| | +-- Lanceolaria_triformis
| |
| | +-------------------- Anemina_arcaeformis
| | |
| | +--+ +----------------- Cristaria_plicata
+--+ | | |
| | +--+ +-------------- Pletholophus_tenuis
| | | |
| | +--+ +-- Sinanodonta_angula
| | | +--------+
| | | | +-- Sinanodonta_lucida
| | +--+
| | | +-------- Sinanodonta_woodiana[2]
| | +--+
| | | +----- Sinanodonta_jourdyi
+--+ +--+
| | +-- Sinanodonta_elliptica
| +--+
| +-- Sinanodonta_woodiana[1]
|
| +-- Alasmidonta_marginata
| +--------+
| | +-- Lasmigona_costata
| |
| | +----- Anodonta_anatina
| +-----+
| | | +-- Anodonta_cygnea
+-----------+ +--+
| +-- Anodonta_nuttalliana
|
| +----- Pseudanodonta_complanata
| +--+
| | | +-- Simpsonaias_ambigua
| | +--+
+--+ +-- Strophitus_undulatus
|
| +----- Pyganodon_grandis
+--+
| +-- Utterbackia_imbecillis
+--+
+-- Utterbackia_peninsularis
Chen et al., 2023 [Publ 4302], fig. 2
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 17 |
17 |
14 |
10 |
23 |
14 |
4 |
0.0 |
+-------------------------- Acuticosta_chinensis
|
| +----------------------- Lepidodesma_languilati
| |
| | +-- Lanceolaria_gladiola
| | +--------------+
| | | +-- Lanceolaria_oxyrhyncha
| | |
--+ | | +-- Alasmidonta_marginata
| +-----+ +--+
| | | | +-- Lasmigona_costata
| | | +--------+
| | | | | +-- Anodonta_anatina
| | | | +--+
| | +--+ +-- Pseudanodonta_complanata
| | |
| | | +----------- Cristaria_plicata
+--+ | |
| +--+ +-- Anemina_arcaeformis
| | +-----+
| | | +-- Amuranodonta_kijaensis
| +--+
| | +----- Pletholophus_tenuis
| +--+
| | +-- Sinanodonta_jourdyi
| +--+
| +-- Beringiana_beringiana
|
| +-------------------- Aculamprotula_fibrosa **
| |
| | +----- Middendorffinaia_mongolica **
| | +-----------+
+--+ | | +-- Nodularia_douglasiae **
| | +--+
| | +-- Nodularia_breviconcha **
| |
| | +-- Cuneopsis_celtiformis **
+--+ +-----------+
| | +-- Cuneopsis_heudei **
| |
| | +-- Diaurora_aurorea **
| | +--+
| | | +-- Diaurora_laeve **
| +--------+
+--+ | +-- Schistodesmus_lampreyanus **
| +--+
| +-- Schistodesmus_spinosus **
|
| +-- Pseudocuneopsis_sichuanensis **
| +--------+
| | +-- Pseudocuneopsis_capitata **
| |
+--+ +-- Inversiunio_yanagawensis **
| +-----+
| | +-- Inversiunio_reinianus **
+--+
| +----- Tchangsinaia_piscicula **
+--+
| +-- Unio_pictorum **
+--+
+-- Unio_crassus **
Dai et al., 2024 [Publ 4358], fig. 4
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 17 |
17 |
15 |
7 |
35 |
10 |
4 |
20.0 |
+-------------------------------- Lepidodesma_languilati
|
| +----------------- Lanceolaria_gladiola
| |
| +-----------+ +-------------- Acuticosta_chinensis
| | | |
| | | | +----- Cristaria_plicata
--+ | +--+ +-----+
| | | | | +-- Anemina_arcaeformis
| | | | +--+
| | +--+ +-- Sinanodonta_woodiana
| | |
| | | +-------- Anodonta_anatina
| | +--+
+--+ | +----- Platynaias_compressa
| +--+
| | +-- Pyganodon_grandis
| +--+
| +-- Utterbackia_imbecillis
|
| +-- Aculamprotula_scripta **
| +-----------------------+
| | +-- Aculamprotula_tientsinensis **
| |
| | +-- Unio_crassus **
+--+ +--------------------+
| | +-- Unio_delphinus **
| |
| | +-------- Pseudocuneopsis_capitata **
+--+ +-----------+
| | | +----- Pseudocuneopsis_yangshuoensis **
| | +--+
| | | +-- Pseudocuneopsis_yemaoi **
| | +--+
+--+ +-- Pseudocuneopsis_sichuanensis **
|
| +-- Nodularia_breviconcha **
| +--------------+
| | +-- Nodularia_douglasiae **
| |
+--+ +-- Schistodesmus_lampreyanus **
| +-----------+
| | +-- Schistodesmus_spinosus **
+--+
| +----------- Tchangsinaia_piscicula **
+--+
| +-------- Cuneopsis_rufescens **
+--+
| +----- Cuneopsis_celtiformis **
+--+
| +-- Cuneopsis_heudei **
+--+
+-- Cuneopsis_demangei **
Lopes-Lima et al., 2021 [Publ 4217], fig. 1
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 16 |
16 |
14 |
1 |
9 |
3 |
1 |
29.0 |
+----- Anodonta_anatina
+--------------------+
| | +-- Anodonta_cygnea
| +--+
| +-- Anodonta_seddoni
--+
| +----------------------- Unio_tumidus **
| |
| | +-------- Unio_damascensis **
| | +-----------+
+--+ | | +----- Unio_sesirmensis **
| | +--+
| | | +-- Unio_bruguierianus **
| | +--+
+--+ +-- Unio_crassus **
|
| +-- Unio_delicatus **
| +--------------+
| | +-- Unio_terminalis **
| |
+--+ +-- Unio_tigridis **
| +-----------+
| | +-- Unio_hueti **
| |
+--+ +----------- Unio_ravoisieri **
| |
| | +-- Unio_pictorum **
+--+ +-----+
| | +-- Unio_eucirrus **
| |
+--+ +-- Unio_elongatulus **
| +--+
| | +-- Unio_mancus **
+--+
| +-- Unio_foucauldianus **
+--+
+-- Unio_delphinus **
Araujo et al., 2018 [Publ 3779], fig. 1
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 15 |
15 |
10 |
2 |
4 |
3 |
1 |
26.0 |
+----- Anodonta_anatina
+-----------------+
| | +-- Pyganodon_grandis
| +--+
| +-- Alasmidonta_marginata
--+
| +-------------------- Nodularia_douglasiae **
| |
| | +-- Unio_durieui **
+--+ +--------------+
| | +-- Unio_gibbus **
| |
+--+ +-------------- Unio_tumidus **
| |
| | +-------- Unio_tumidiformis **
| | +--+
+--+ | | +----- Unio_bruguierianus **
| | +--+
| | | +-- Unio_crassus **
| | +--+
+--+ +-- Unio_ionicus **
|
| +-- Unio_delphinus **
| +--+
| | +-- Unio_foucauldianus **
| |
+-----+ +-- Unio_pictorum **
| |
| +-- Unio_elongatulus **
| |
+--+-- Unio_mancus **
|
+-- Unio_ravoisieri **
|
+-- Unio_tigridis **
Sano et al., 2020 [Publ 4138], fig. 2
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 15 |
15 |
13 |
6 |
58 |
24 |
3 |
23.0 |
+----- Lanceolaria_grayii[1]
+-----------------+
| | +-- Lanceolaria_lanceolata
| +--+
| +-- Lanceolaria_grayii[2]
--+
| +-------------------- Lepidodesma_languilati
| |
| | +----- Aculamprotula_tortuosa **
| | +-----------+
+--+ | | +-- Aculamprotula_tientsinensis **
| | +--+
| | +-- Aculamprotula_coreana **
| |
+--+ +----- Unio_crassus **
| +--------+
| | | +-- Unio_delphinus **
| | +--+
| | +-- Unio_pictorum **
| |
+--+ +-------- Inversiunio_jokohamensis **
| +--+
| | | +----- Inversiunio_yanagawensis[2] **
| | +--+
| | | +-- Inversiunio_reinianus **
| | +--+
+--+ +-- Inversiunio_yanagawensis[1] **
|
| +-- Tchangsinaia_piscicula **
| +-----+
| | +-- Schistodesmus_lampreyanus **
+--+
| +----- Nodularia_douglasiae[2] **
+--+
| +-- Nodularia_nipponensis **
+--+
+-- Nodularia_douglasiae[1] **
Froufe et al., 2016 [Publ 3738], fig. 3
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 14 |
14 |
12 |
2 |
0 |
0 |
0 |
23.0 |
+-- Nodularia_douglasiae **
+--------------------------+
| +-- Nodularia_breviconcha **
|
--+ +-- Unio_gibbus **
| +-----------------------+
| | +-- Unio_durieui **
+--+
| +----------------------- Unio_tumidus **
| |
+--+ +-- Unio_crassus **
| +-----------------+
| | +-- Unio_tumidiformis **
+--+
| +----------------- Unio_delphinus **
+--+
| +-------------- Unio_foucauldianus **
+--+
| +----------- Unio_ravoisieri **
+--+
| +-------- Unio_mancus **
+--+
| +----- Unio_elongatulus **
+--+
| +-- Unio_pictorum **
+--+
+-- Unio_caffer **
Wu et al., 2018 [Publ 3836], fig. 2
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 14 |
14 |
12 |
6 |
39 |
19 |
4 |
18.0 |
+----------------------- Lepidodesma_languilati
|
| +----------- Aculamprotula_tortuosa **
| +-----+
| | | +-------- Aculamprotula_scripta **
| | +--+
| | | +----- Aculamprotula_coreana **
| | +--+
| | | +-- Aculamprotula_fibrosa **
+-----+ +--+
| | +-- Aculamprotula_tientsinensis **
| |
| | +-- Unio_crassus **
| | +-----------+
| | | +-- Unio_pictorum **
| | |
| +--+ +----- Nodularia_douglasiae **
| | +-----+
| | | | +-- Schistodesmus_lampreyanus **
--+ | | +--+
| +--+ +-- Schistodesmus_spinosus **
| |
| | +-------- Tchangsinaia_piscicula **
| +--+
| | +----- Cuneopsis_rufescens **
| +--+
| | +-- Cuneopsis_celtiformis **
| +--+
| +-- Cuneopsis_heudei **
|
| +----------- Acuticosta_chinensis
| +--------+
| | | +-------- Lanceolaria_gladiola
| | +--+
| | | +----- Lanceolaria_grayii
| | +--+
| | | +-- Lanceolaria_lanceolata
| | +--+
| | +-- Lanceolaria_triformis
+--+
| +-------- Anemina_arcaeformis
| |
| +-------- Cristaria_plicata
| +--------+
| | | +-- Sinanodonta_woodiana
| | | +--+
| | | | +-- Sinanodonta_elliptica
| | +--+
+--+ | +-- Sinanodonta_lucida
| +--+
| +-- Sinanodonta_angula
|
| +-------------- Anodonta_cygnea
| |
+--+ +----------- Anodonta_anatina
| |
+--+ +-------- Strophitus_undulatus
| |
+--+ +----- Platynaias_compressa
| |
+--+----- Alasmidonta_marginata
|
| +-- Pyganodon_cataracta
+--+
+-- Pyganodon_grandis
Huang et al., 2019 [Publ 3843], fig. 1
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 14 |
20 |
18 |
5 |
30 |
14 |
4 |
26.0 |
+----------------------------- Acuticosta_chinensis
|
| +-- Sinanodonta_angula
| +--------+
| | +-- Sinanodonta_lucida
| +--------------+
| | | +-------- Sinanodonta_jourdyi
--+ | +--+
| | | +----- Sinanodonta_woodiana[2]
| | +--+
| | | +-- Sinanodonta_elliptica
| | +--+
| | +-- Sinanodonta_woodiana[1]
| |
+--+ +----------- Aculamprotula_tortuosa **
| +-----------+
| | | +-------- Aculamprotula_scripta **
| | +--+
| | | +----- Aculamprotula_coreana **
| | +--+
| | | +-- Aculamprotula_fibrosa **
| | +--+
| | +-- Aculamprotula_tientsinensis **
+--+
| +----------- Cristaria_plicata
| +--------+
| | | +-------- Lanceolaria_gladiola
| | +--+
| | | +----- Lanceolaria_grayii
| | +--+
| | | +-- Lanceolaria_lanceolata
+--+ +--+
| +-- Lanceolaria_triformis
|
| +----------------- Lepidodesma_languilati
| |
| | +-- Schistodesmus_lampreyanus **
+--+ +-----------+
| | +-- Schistodesmus_spinosus **
| |
+--+ +----- Nodularia_douglasiae[1] **
| +-----+
| | | +-- Nodularia_douglasiae[2] **
| | +--+
+--+ +-- Nodularia_nuxpersicae **
|
| +-------- Cuneopsis_rufescens **
+--+
| +----- Tchangsinaia_piscicula **
+--+
| +-- Cuneopsis_heudei **
+--+
+-- Cuneopsis_celtiformis **
Huang et al., 2019 [Publ 3843], fig. S1
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 14 |
14 |
12 |
5 |
61 |
41 |
4 |
18.0 |
+-- Lepidodesma_languilati
+--------------------+
| +-- Cristaria_plicata
|
| +----------- Lanceolaria_gladiola
| |
| | +-------- Lanceolaria_grayii[5]
| +--------+ |
| | | | +-- Lanceolaria_lanceolata
| | | +-----+
| | | | +-- Lanceolaria_triformis
--+ | +--+
| | | +----- Lanceolaria_grayii[4]
| | | |
| | | | +-- Lanceolaria_eucylindrica[1]
| | | | |
| | +--+ +-- Lanceolaria_eucylindrica[2]
| | | |
| | | +-- Lanceolaria_eucylindrica[3]
| | | |
| | +--+-- Lanceolaria_eucylindrica[4]
+--+ |
| +-- Lanceolaria_grayii[1]
| |
| +-- Lanceolaria_grayii[2]
| |
| +-- Lanceolaria_grayii[3]
|
| +----------- Aculamprotula_tortuosa **
| +-----+
| | | +-------- Aculamprotula_tientsinensis **
| | +--+
| | | +----- Aculamprotula_fibrosa **
| | +--+
| | | +-- Aculamprotula_coreana **
+--+ +--+
| +-- Aculamprotula_scripta **
|
| +----- Nodularia_douglasiae[1] **
| +--------+
| | | +-- Nodularia_nuxpersicae **
| | +--+
+--+ +-- Nodularia_douglasiae[2] **
|
| +----------- Tchangsinaia_piscicula **
| |
+--+ +-- Schistodesmus_spinosus **
| +-----+
| | +-- Schistodesmus_lampreyanus **
+--+
| +----- Cuneopsis_rufescens **
+--+
| +-- Cuneopsis_heudei **
+--+
+-- Cuneopsis_celtiformis **
Wu et al., 2020 [Publ 4121], fig. 3
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 14 |
14 |
12 |
7 |
36 |
14 |
4 |
23.0 |
+----------------------- Lepidodesma_languilati
|
| +----- Acuticosta_chinensis
| +-----------+
| | | +-- Lanceolaria_lanceolata
| | +--+
| | +-- Lanceolaria_grayii
| |
| +--+ +----------- Anodonta_anatina
| | | +--+
| | | | | +-------- Platynaias_compressa
--+ | | | +--+
| | | | | +----- Pyganodon_grandis
| | | | +--+
| | +--+ | +-- Utterbackia_peninsularis
| | | +--+
| | | +-- Utterbackia_imbecillis
| | |
| | | +-------- Cristaria_plicata
| | | |
| | +-----+ +-- Sinanodonta_woodiana
| | | +--+
+--+ | | +-- Sinanodonta_lauta
| +--+
| | +-- Anemina_arcaeformis
| +--+
| +-- Anemina_euscaphys
|
| +-------- Aculamprotula_tortuosa **
| |
| +--------+ +-- Aculamprotula_coreana **
| | | +--+
| | | | +-- Aculamprotula_tientsinensis **
| | +--+
| | | +-- Aculamprotula_polysticta **
| | +--+
+--+ +-- Aculamprotula_scripta **
|
| +-------- Unio_tumidus **
| +-----+
| | | +----- Unio_crassus **
| | +--+
| | | +-- Unio_delphinus **
+--+ +--+
| +-- Unio_pictorum **
|
| +----------- Pseudocuneopsis_capitata **
+--+
| +-------- Nodularia_douglasiae **
+--+
| +----- Tchangsinaia_piscicula **
+--+
| +-- Cuneopsis_heudei **
+--+
+-- Schistodesmus_lampreyanus **
Sano et al., 2020 [Publ 4138], fig. 3
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 14 |
14 |
12 |
6 |
58 |
24 |
3 |
22.0 |
+----- Lanceolaria_grayii[1]
+-----------------+
| | +-- Lanceolaria_lanceolata
| +--+
| +-- Lanceolaria_grayii[2]
--+
| +-------------------- Lepidodesma_languilati
| |
| | +----- Aculamprotula_tortuosa **
| | +-----------+
+--+ | | +-- Aculamprotula_tientsinensis **
| | +--+
| | +-- Aculamprotula_coreana **
| |
+--+ +----- Unio_crassus **
| +--------+
| | | +-- Unio_delphinus **
| | +--+
| | +-- Unio_pictorum **
| |
+--+ +-------- Inversiunio_jokohamensis **
| +--+
| | | +----- Inversiunio_yanagawensis[2] **
| | +--+
| | | +-- Inversiunio_reinianus **
| | +--+
+--+ +-- Inversiunio_yanagawensis[1] **
|
| +-- Nodularia_nipponensis **
| +--+
| | +-- Nodularia_douglasiae **
+-----+
| +-- Schistodesmus_lampreyanus **
+--+
+-- Tchangsinaia_piscicula **
Sano et al., 2020 [Publ 4138], fig. 4
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 14 |
14 |
12 |
6 |
58 |
24 |
3 |
22.0 |
+-------------------------- Lepidodesma_languilati
|
| +----- Lanceolaria_grayii[1]
| +-----------------+
--+ | | +-- Lanceolaria_lanceolata
| | +--+
| | +-- Lanceolaria_grayii[2]
| |
+--+ +----- Aculamprotula_tortuosa **
| +--------------+
| | | +-- Aculamprotula_tientsinensis **
| | +--+
| | +-- Aculamprotula_coreana **
| |
+--+ +----- Unio_crassus **
| +-----------+
| | | +-- Unio_delphinus **
| | +--+
| | +-- Unio_pictorum **
+--+
| +-- Nodularia_douglasiae **
| +-----------+
| | +-- Nodularia_nipponensis **
| |
+--+ +-- Schistodesmus_lampreyanus **
| +--------+
| | +-- Tchangsinaia_piscicula **
+--+
| +-------- Inversiunio_jokohamensis **
+--+
| +----- Inversiunio_yanagawensis[2] **
+--+
| +-- Inversiunio_reinianus **
+--+
+-- Inversiunio_yanagawensis[1] **
Liu et al., 2023 [Publ 4310], fig. 4
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 14 |
14 |
12 |
8 |
10 |
8 |
3 |
24.0 |
+-------------------------------- Lanceolaria_gladiola
|
| +-------- Pseudanodonta_complanata
| +--------------------+
--+ | | +----- Lasmigona_costata
| | +--+
| | | +-- Strophitus_undulatus
| | +--+
+--+ +-- Pyganodon_grandis
|
| +-------------------------- Unio_tumidus **
| |
| | +-- Aculamprotula_scripta **
+--+ +--------------------+
| | +-- Aculamprotula_tientsinensis **
| |
+--+ +-- Cuneopsis_heudei **
| +-----------------+
| | +-- Cuneopsis_rufescens **
| |
+--+ +----- Nodularia_douglasiae **
| +-----------+
| | | +-- Schistodesmus_lampreyanus **
| | +--+
+--+ +-- Schistodesmus_spinosus **
|
| +-------------- Tchangsinaia_piscicula **
+--+
| +----------- Inversiunio_yanagawensis **
+--+
| +-------- Pseudocuneopsis_capitata **
+--+
| +----- Pseudocuneopsis_sichuanensis **
+--+
| +-- Pseudocuneopsis_wuana **
+--+
+-- Pseudocuneopsis_yangshuoensis **
Liu et al., 2023 [Publ 4310], fig. 5
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 14 |
14 |
9 |
8 |
10 |
8 |
3 |
21.0 |
+----------------------- Lanceolaria_grayii
|
+----------------------- Lanceolaria_gladiola
|
+----------------------- Lanceolaria_lanceolata
|
--+ +-------- Lasmigona_costata
| +-----------+
| | | +----- Pseudanodonta_complanata
| | +--+
| | | +-- Pyganodon_grandis
| | +--+
| | +-- Strophitus_undulatus
+--+
| +----------------- Unio_tumidus **
| |
| | +-- Aculamprotula_tientsinensis **
| +--------------+
| | +-- Aculamprotula_scripta **
| |
+--+ +-- Cuneopsis_rufescens **
| +-----------+
| | +-- Cuneopsis_heudei **
| |
| | +----- Nodularia_douglasiae **
| +--------+
+--+ | +-- Schistodesmus_lampreyanus **
| +--+
| +-- Schistodesmus_spinosus **
|
| +----------- Tchangsinaia_piscicula **
| |
+--+----------- Inversiunio_yanagawensis **
|
| +-------- Pseudocuneopsis_capitata **
+--+
| +----- Pseudocuneopsis_sichuanensis **
+--+
| +-- Pseudocuneopsis_wuana **
+--+
+-- Pseudocuneopsis_yangshuoensis **
Wu et al., 2019 [Publ 3864], fig. 3
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 13 |
13 |
11 |
7 |
34 |
14 |
4 |
22.0 |
+----------------------- Lepidodesma_languilati
|
| +-------- Aculamprotula_tortuosa **
| +--------+
| | | +----- Aculamprotula_scripta **
| | +--+
| | | +-- Aculamprotula_tientsinensis **
| | +--+
| | +-- Aculamprotula_coreana **
| +--+
--+ | | +-------- Unio_tumidus **
| | | +-----+
| | | | | +----- Unio_crassus **
| | | | +--+
| | | | | +-- Unio_delphinus **
| | +--+ +--+
| | | +-- Unio_pictorum **
| | |
| | | +----------- Pseudocuneopsis_capitata **
| | +--+
| | | +-------- Nodularia_douglasiae **
+--+ +--+
| | +----- Tchangsinaia_piscicula **
| +--+
| | +-- Cuneopsis_heudei **
| +--+
| +-- Schistodesmus_lampreyanus **
|
| +----- Acuticosta_chinensis
| +-----------+
| | | +-- Lanceolaria_grayii
| | +--+
| | +-- Lanceolaria_lanceolata
| |
+--+ +----------- Anodonta_anatina
| +--+
| | | +-------- Platynaias_compressa
| | +--+
| | | +----- Pyganodon_grandis
| | +--+
+--+ | +-- Utterbackia_peninsularis
| +--+
| +-- Utterbackia_imbecillis
|
| +-------- Cristaria_plicata
| |
+-----+ +-- Anemina_arcaeformis
| +--+
| | +-- Anemina_euscaphys
+--+
| +-- Sinanodonta_lucida
+--+
+-- Sinanodonta_woodiana
Liu et al., 2017 [Publ 3939], fig. 2
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 13 |
13 |
8 |
4 |
4 |
3 |
3 |
19.0 |
+-------------------- Lanceolaria_grayii
|
| +----- Unio_tumidus **
| |
--+ +-----------+----- Unio_gibbus **
| | |
| | | +-- Unio_crassus **
| | +--+
| | +-- Unio_pictorum **
+--+
| +-------------- Schistodesmus_lampreyanus **
| |
| | +-- Cuneopsis_heudei **
| +-----------+
+--+ +-- Cuneopsis_rufescens **
|
| +----------- Nodularia_breviconcha **
| |
+--+ +-------- Nodularia_douglasiae[3] **
| |
+--+ +----- Nodularia_douglasiae[2] **
| |
+--+ +-- Nodularia_douglasiae[1] **
| |
+--+-- Nodularia_nuxpersicae **
|
+-- Nodularia_nipponensis **
Sayenko et al., 2021 [Publ 4189], fig. 6
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 13 |
13 |
10 |
4 |
0 |
0 |
0 |
18.0 |
+-- Unio_pictorum **
+-----------------+
| +-- Unio_crassus **
|
| +-------- Inversiunio_jokohamensis **
--+ |
| +--------+ +----- Inversiunio_yanagawensis[2] **
| | | |
| | +--+----- Inversiunio_verrucosus **
| | |
+--+ | +-- Inversiunio_reinianus **
| +--+
| +-- Inversiunio_yanagawensis[1] **
|
| +-------------- Middendorffinaia_mongolica **
+--+
| +----------- Nodularia_breviconcha **
+--+
| +-------- Nodularia_douglasiae[2] **
+--+
| +----- Nodularia_nuxpersicae **
+--+
| +-- Nodularia_douglasiae[1] **
+--+
+-- Nodularia_nipponensis **
Wu et al., 2023 [Publ 4296], fig. 4
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 13 |
13 |
11 |
8 |
10 |
8 |
3 |
23.0 |
+----- Lanceolaria_gladiola
+-----------------------+
| | +-- Lanceolaria_lanceolata
| +--+
| +-- Lanceolaria_grayii
|
--+ +-------- Pseudanodonta_complanata
| +-----------------+
| | | +----- Lasmigona_costata
| | +--+
| | | +-- Strophitus_undulatus
| | +--+
+--+ +-- Pyganodon_grandis
|
| +----------------------- Unio_tumidus **
| |
| | +-- Aculamprotula_scripta **
+--+ +-----------------+
| | +-- Aculamprotula_tientsinensis **
| |
+--+ +-- Cuneopsis_heudei **
| +--------------+
| | +-- Cuneopsis_rufescens **
| |
+--+ +----- Nodularia_douglasiae **
| +--------+
| | | +-- Schistodesmus_lampreyanus **
| | +--+
+--+ +-- Schistodesmus_spinosus **
|
| +----------- Tchangsinaia_piscicula **
+--+
| +-------- Inversiunio_yanagawensis **
+--+
| +----- Pseudocuneopsis_capitata **
+--+
| +-- Pseudocuneopsis_yangshuoensis **
+--+
+-- Pseudocuneopsis_sichuanensis **
Fukata & Iigo, 2020 [Publ 4106], fig. 1
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 12 |
12 |
10 |
7 |
42 |
16 |
4 |
21.0 |
+----- Lanceolaria_gladiola
+--------------+
| | +-- Lanceolaria_grayii
| +--+
| +-- Lanceolaria_lanceolata
|
| +-------------- Acuticosta_chinensis
| |
| | +-------- Cristaria_plicata
| | |
| | +--+ +-- Sinanodonta_woodiana
| +--+ | | +--+
--+ | | | | | +-- Sinanodonta_lucida
| | | | +--+
| | | | | +-- Anemina_arcaeformis
| | | | +--+
| | +--+ +-- Anemina_euscaphys
| | |
| | | +-- Anodonta_anatina
| | | +-----+
| | | | +-- Anodonta_cygnea
| | | |
| | +--+ +-- Pyganodon_grandis
+--+ | +--+
| | | +-- Utterbackia_peninsularis
| +--+
| | +-- Alasmidonta_varicosa
| +--+
| +-- Prolasmidonta_heterodon
|
| +----- Aculamprotula_tortuosa **
| +--------+
| | | +-- Aculamprotula_tientsinensis **
| | +--+
| | +-- Aculamprotula_coreana **
| |
+--+ +-------- Unio_tumidus **
| +--+
| | | +----- Unio_crassus **
| | +--+
| | | +-- Unio_pictorum **
| | +--+
+--+ +-- Unio_delphinus **
|
| +-- Schistodesmus_lampreyanus **
| +-----+
| | +-- Cuneopsis_heudei **
+--+
| +----- Pseudocuneopsis_capitata **
+--+
| +-- Nodularia_douglasiae **
+--+
+-- Tchangsinaia_piscicula **
Sano et al., 2020 [Publ 4138], fig. 1
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 12 |
34 |
5 |
5 |
36 |
22 |
4 |
41.0 |
+-------- Lanceolaria_grayii[1]
|
+-------- Lanceolaria_grayii[2]
|
+-------- Cristaria_plicata[1]
|
+-------- Cristaria_plicata[2]
|
+-------- Sinanodonta_lauta
|
+-------- Beringiana_japonica
|
+-------- Beringiana_fukuharai[1]
|
+-------- Beringiana_beringiana
|
+-------- Beringiana_fukuharai[2]
|
+-------- Beringiana_fukuharai[3]
|
+-------- Buldowskia_iwakawai[1]
|
+-------- Buldowskia_iwakawai[2]
|
+-------- Sinanodonta_tumens[1]
|
+-------- Sinanodonta_tumens[2]
|
--+-------- Pletholophus_tenuis
|
+-------- Anemina_arcaeformis
|
+-------- Lanceolaria_lanceolata
|
+-------- Cristaria_plicata[3]
|
+-------- Lanceolaria_gladiola
|
+-------- Sinanodonta_angula
|
+-------- Lepidodesma_languilati
|
| +-- Aculamprotula_tortuosa **
+-----+
| +-- Acuticosta_chinensis
|
| +----- Cuneopsis_celtiformis **
| |
+--+ +-- Nodularia_douglasiae[1] **
| | |
| +--+-- Nodularia_douglasiae[2] **
| |
| +-- Nodularia_nipponensis **
|
| +----- Schistodesmus_lampreyanus **
| |
| | +-- Inversiunio_reinianus[1] **
+--+ |
| +-- Inversiunio_reinianus[2] **
| |
| +-- Inversiunio_jokohamensis[1] **
+--+
+-- Inversiunio_jokohamensis[2] **
|
+-- Inversiunio_jokohamensis[3] **
|
+-- Inversiunio_jokohamensis[4] **
Wu et al., 2022 [Publ 4238], fig. 2e
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 12 |
12 |
10 |
7 |
5 |
5 |
2 |
17.0 |
+-------- Sinanodonta_woodiana
+-----------------+
| | +----- Anodonta_anatina
| +--+
| | +-- Utterbackia_peninsularis
--+ +--+
| +-- Pyganodon_grandis
|
| +----------------------- Lanceolaria_lanceolata
| |
+--+ +-------------------- Aculamprotula_tortuosa **
| |
| | +----- Unio_tumidus **
+--+ +-----------+
| | | +-- Unio_crassus **
| | +--+
| | +-- Unio_delphinus **
+--+
| +-- Pseudocuneopsis_sichuanensis **
| +-----------+
| | +-- Pseudocuneopsis_capitata **
+--+
| +----------- Nodularia_douglasiae **
| |
+--+ +-------- Tchangsinaia_piscicula **
| |
+--+ +-- Cuneopsis_celtiformis **
| +--+
| | +-- Cuneopsis_heudei **
+--+
| +-- Schistodesmus_lampreyanus **
+--+
+-- Schistodesmus_spinosus **
Konopleva et al., 2023 [Publ 4269], fig. S1, 4
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 12 |
12 |
10 |
6 |
44 |
21 |
4 |
24.0 |
+----------------- Middendorffinaia_mongolica **
|
| +-------- Aculamprotula_tortuosa **
| +-----+
+--------+ | | +----- Aculamprotula_scripta **
| | | +--+
| | | | +-- Aculamprotula_tientsinensis **
| | | +--+
| +--+ +-- Aculamprotula_coreana **
| |
| | +----- Unio_tumidus **
| | +-----+
| | | | +-- Unio_pictorum **
| | | +--+
| +--+ +-- Unio_crassus **
| |
| | +-------- Nodularia_douglasiae **
--+ +--+
| | +----- Tchangsinaia_piscicula **
| +--+
| | +-- Cuneopsis_rufescens **
| +--+
| +-- Cuneopsis_heudei **
|
| +-------- Lepidodesma_languilati
| +--------------+
| | | +----- Lanceolaria_gladiola
| | +--+
| | | +-- Lanceolaria_grayii
| | +--+
+--+ +-- Lanceolaria_triformis
|
| +-------------------- Acuticosta_chinensis
| |
| | +----- Anodonta_anatina
| | +-----------+
+--+ | | +-- Pseudanodonta_complanata
| | +--+
| | +-- Anodonta_cygnea
| |
| | +-------- Anemina_arcaeformis
+--+ +-----+
| | | +----- Amuranodonta_kijaensis
| | +--+
| | | +-- Buldowskia_suifunica
| | +--+
| | +-- Buldowskia_shadini
+--+
| +-------- Beringiana_beringiana
| +--+
| | | +----- Sinanodonta_schrenkii
| | +--+
| | | +-- Sinanodonta_woodiana
+--+ +--+
| +-- Sinanodonta_lauta
|
| +-------- Cristaria_radiata
| |
+--+ +-- Cristaria_truncata
| +--+
| | +-- Cristaria_plicata
+--+
| +-- Cristaria_bellua
+--+
+-- Cristaria_clessini
Prié & Puillandre, 2014 [Publ 3529], fig. 2
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 11 |
11 |
9 |
1 |
6 |
3 |
1 |
24.0 |
+----- Anodonta_anatina
+--------------+
| | +-- Pseudanodonta_complanata
| +--+
| +-- Anodonta_cygnea
|
--+ +----- Unio_tumidus **
| +-----------+
| | | +-- Unio_gibbus **
| | +--+
| | +-- Unio_durieui **
+--+
| +----- Unio_tumidiformis **
| +--------+
| | | +-- Unio_crassus **
| | +--+
+--+ +-- Unio_courtillieri **
|
| +----------- Unio_delphinus **
+--+
| +-------- Unio_elongatulus **
+--+
| +----- Unio_mancus **
+--+
| +-- Unio_pictorum **
+--+
+-- Unio_ravoisieri **
Lopes-Lima et al., 2017 [Publ 3745], fig. 1-2
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 11 |
11 |
8 |
6 |
70 |
18 |
2 |
22.0 |
+----- Unio_gibbus **
|
+-----------+----- Unio_tumidus **
| |
| | +-- Unio_crassus **
| +--+
+--+ +-- Unio_pictorum **
| |
| | +-------------- Aculamprotula_tortuosa **
| | |
| +--+ +-- Nodularia_nuxpersicae **
| | +--------+
| | | +-- Nodularia_douglasiae **
| +--+
| | +-------- Schistodesmus_lampreyanus **
| +--+
| | +----- Tchangsinaia_piscicula **
| +--+
--+ | +-- Cuneopsis_heudei **
| +--+
| +-- Cuneopsis_rufescens **
|
| +-------- Lanceolaria_gladiola
| +--------+
| | | +----- Lanceolaria_grayii[2]
| | +--+
| | | +-- Lanceolaria_lanceolata
| | +--+
| | +-- Lanceolaria_grayii[1]
| |
+--+ +----------- Cristaria_plicata
| +--+
| | | +-------- Anemina_arcaeformis
| | +--+
| | | +----- Pletholophus_tenuis
| | +--+
| | | +-- Sinanodonta_jourdyi
| | +--+
+--+ +-- Sinanodonta_lucida
|
| +----- Pseudanodonta_complanata
| |
| +-----+ +-- Anodonta_anatina
| | | |
| | +--+-- Anodonta_cygnea
| | |
+--+ +-- Anodonta_nuttalliana
|
| +-- Pyganodon_grandis
| +-----+
| | +-- Simpsonaias_ambigua
+--+
| +----- Strophitus_undulatus
+--+
| +-- Alasmidonta_marginata
+--+
+-- Lasmigona_costata
Huang et al., 2019 [Publ 3843], fig. 3
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 11 |
11 |
9 |
5 |
46 |
16 |
4 |
21.0 |
+----------------------------- Lepidodesma_languilati
|
| +-------- Aculamprotula_tortuosa **
| +-----+
| | | +----- Aculamprotula_scripta **
| | +--+
| | | +-- Aculamprotula_tientsinensis **
| | +--+
| +-----------+ +-- Aculamprotula_coreana **
--+ | |
| | | +----- Nodularia_douglasiae **
| | | +-----+
| | | | | +-- Tchangsinaia_piscicula **
| | | | +--+
| | +--+ +-- Schistodesmus_lampreyanus **
| | |
| | | +-------- Unio_tumidus **
| | +--+
+--+ | +----- Unio_crassus **
| +--+
| | +-- Unio_delphinus **
| +--+
| +-- Unio_pictorum **
|
| +----- Lanceolaria_gladiola
| +-----------------+
| | | +-- Lanceolaria_lanceolata
| | +--+
| | +-- Lanceolaria_grayii
+--+
| +-------------------- Acuticosta_chinensis
| |
| | +-------- Cristaria_plicata
| | |
+--+ +--------+ +-- Sinanodonta_lucida
| | | +--+
| | | | +-- Sinanodonta_woodiana
| | +--+
+--+ | +-- Anemina_arcaeformis
| +--+
| +-- Anemina_euscaphys
|
| +-------------- Anodonta_cygnea
+--+
| +----------- Anodonta_anatina
+--+
| +-------- Lasmigona_costata
+--+
| +----- Pyganodon_grandis
+--+
| +-- Utterbackia_imbecillis
+--+
+-- Utterbackia_peninsularis
Aunins et al., 2018 [Publ 3863], fig. 1
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 11 |
11 |
9 |
6 |
30 |
16 |
4 |
20.0 |
+----------------------- Lepidodesma_languilati
|
| +----- Aculamprotula_tortuosa **
| +--------+
| | | +-- Aculamprotula_coreana **
| | +--+
| | +-- Aculamprotula_tientsinensis **
| |
| +-----+ +-------- Nodularia_douglasiae **
--+ | | +--+
| | | | | +----- Tchangsinaia_piscicula **
| | | | +--+
| | | | | +-- Cuneopsis_heudei **
| | +--+ +--+
| | | +-- Schistodesmus_lampreyanus **
| | |
| | | +-------- Unio_tumidus **
| | +--+
| | | +----- Unio_crassus **
+--+ +--+
| | +-- Unio_pictorum **
| +--+
| +-- Unio_delphinus **
|
| +----- Acuticosta_chinensis
| +-----------+
| | | +-- Lanceolaria_grayii
| | +--+
| | +-- Lanceolaria_lanceolata
| |
| | +-------- Cristaria_plicata
+--+ |
| +-----+ +-- Anemina_arcaeformis
| | | +--+
| | | | +-- Anemina_euscaphys
| | +--+
| | | +-- Sinanodonta_woodiana
| | +--+
+--+ +-- Sinanodonta_lucida
|
| +-- Anodonta_anatina
| +--------+
| | +-- Anodonta_cygnea
| |
+--+ +-- Prolasmidonta_heterodon
| +-----+
| | +-- Alasmidonta_varicosa
+--+
| +----- Pyganodon_grandis
+--+
| +-- Utterbackia_imbecillis
+--+
+-- Utterbackia_peninsularis
Choi et al., 2020 [Publ 4169], fig. 2a
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 11 |
11 |
9 |
5 |
0 |
0 |
0 |
16.0 |
+-------- Schistodesmus_lampreyanus **
|
+--------+ +-- Unio_pictorum **
| | +--+
| | | +-- Tchangsinaia_piscicula **
| +--+
--+ | +-- Cuneopsis_heudei **
| +--+
| +-- Cuneopsis_rufescens **
|
| +-------------- Nodularia_breviconcha **
+--+
| +----------- Nodularia[2] **
+--+
| +-------- Nodularia[1] **
+--+
| +----- Nodularia_douglasiae **
+--+
| +-- Nodularia_nipponensis **
+--+
+-- Nodularia_nuxpersicae **
Bolotov et al., 2017 [Publ 3761], fig. S1
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 10 |
10 |
8 |
6 |
131 |
17 |
2 |
21.0 |
+----- Unio_tumidus **
+--------+
| | +-- Unio_pictorum **
| +--+
| +-- Unio_crassus **
+--------+
| | +----------- Aculamprotula_tortuosa **
| | |
| | | +-- Nodularia_douglasiae **
| +--+ +-----+
| | | +-- Nodularia_nuxpersicae **
| | |
| +--+ +-- Cuneopsis_rufescens **
| | +--+
| | | +-- Cuneopsis_heudei **
| +--+
--+ | +-- Tchangsinaia_piscicula **
| +--+
| +-- Schistodesmus_lampreyanus **
|
| +-- Lanceolaria_grayii
| +-----------------+
| | +-- Lanceolaria_gladiola
| |
| | +-- Cristaria_plicata
| | +--------+
| | | +-- Cristaria_radiata
+--+ +-----+
| | | +-------- Pletholophus_tenuis
| | +--+
| | | +----- Sinanodonta_lucida
| | +--+
| | | +-- Sinanodonta_jourdyi
+--+ +--+
| +-- Sinanodonta_lauta
|
| +-------------- Anodonta_anatina
| |
| | +----- Pseudanodonta_complanata
+--+ +-----+
| | | +-- Anodonta_cygnea
| | +--+
| | +-- Anodonta_nuttalliana
+--+
| +-- Lasmigona_costata
| +-----+
| | +-- Alasmidonta_marginata
+--+
| +----- Simpsonaias_ambigua
+--+
| +-- Pyganodon_grandis
+--+
+-- Strophitus_undulatus
Bolotov et al., 2017 [Publ 3761], fig. S2
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 10 |
10 |
6 |
6 |
131 |
17 |
2 |
19.0 |
+----- Unio_tumidus **
+-----+
| | +-- Unio_crassus **
| +--+
| +-- Unio_pictorum **
+-----+
| | +-------- Aculamprotula_tortuosa **
| | |
| | +-------- Schistodesmus_lampreyanus **
| | |
| +--+ +-- Nodularia_douglasiae **
| +-----+
| | +-- Nodularia_nuxpersicae **
| |
| | +----- Cuneopsis_heudei **
| +--+
| | +-- Tchangsinaia_piscicula **
| +--+
--+ +-- Cuneopsis_rufescens **
|
| +-- Lanceolaria_grayii
| +-----------+
| | +-- Lanceolaria_gladiola
| |
| | +-- Cristaria_plicata
| | +--------+
| | | +-- Cristaria_radiata
| +--+
| | | +-------- Pletholophus_tenuis
| | +--+
| | | +----- Sinanodonta_lucida
+--+ +--+
| | +-- Sinanodonta_lauta
| +--+
| +-- Sinanodonta_jourdyi
|
| +-------- Anodonta_anatina
| |
| +-------- Pseudanodonta_complanata
| |
| | +-- Anodonta_cygnea
+-----+-----+
| +-- Anodonta_nuttalliana
|
| +----- Pyganodon_grandis
| |
| +----- Simpsonaias_ambigua
+--+
+----- Strophitus_undulatus
|
| +-- Alasmidonta_marginata
+--+
+-- Lasmigona_costata
Bolotov et al., 2017 [Publ 3764], fig. S1
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 10 |
10 |
0 |
6 |
121 |
18 |
2 |
18.0 |
+-- Unio_crassus **
|
+-- Unio_pictorum **
|
+-- Unio_tumidus **
|
+-- Aculamprotula_tortuosa **
|
+-- Cuneopsis_heudei **
+--+
| +-- Tchangsinaia_piscicula **
| |
| +-- Cuneopsis_rufescens **
| |
| +-- Nodularia_douglasiae **
| |
| +-- Nodularia_nuxpersicae **
| |
| +-- Schistodesmus_lampreyanus **
|
| +-- Alasmidonta_marginata
| |
| +-- Anodonta_anatina
--+ |
| +-- Anodonta_cygnea
| |
| +-- Anodonta_nuttalliana
| |
| +-- Lasmigona_costata
| |
| +-- Pseudanodonta_complanata
| |
| +-- Pyganodon_grandis
| |
| +-- Simpsonaias_ambigua
| |
| +-- Strophitus_undulatus
+--+
+-- Sinanodonta_lauta
|
+-- Cristaria_plicata
|
+-- Cristaria_radiata
|
+-- Pletholophus_tenuis
|
+-- Sinanodonta_lucida
|
+-- Sinanodonta_jourdyi
|
+-- Lanceolaria_gladiola
|
+-- Lanceolaria_grayii[1]
|
+-- Lanceolaria_grayii[2]
Sano et al., 2017 [Publ 3799], fig. 2
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 10 |
15 |
12 |
4 |
26 |
14 |
4 |
22.0 |
+-- Cristaria_plicata
+-----------+
| +-- Buldowskia_iwakawai
|
+-----+ +----- Beringiana_fukuharai
| | +-----+
| | | | +-- Beringiana_japonica
| | | +--+
| +--+ +-- Beringiana_beringiana
| |
| | +-------- Pletholophus_tenuis
| +--+
| | +----- Sinanodonta_woodiana
| +--+
--+ | +-- Sinanodonta_lauta
| +--+
| +-- Sinanodonta_tumens
|
| +-------- Tchangsinaia_piscicula **
| +--------+
| | | +----- Nodularia_nipponensis **
| | +--+
| | | +-- Nodularia_douglasiae **
| | +--+
| | +-- Nodularia_breviconcha **
+--+
| +-- Schistodesmus_lampreyanus **
| +-----------+
| | +-- Acuticosta_ovata
| |
| | +-------- Lepidodesma_languilati
+--+ +--+
| | | +----- Lanceolaria_grayii[1]
| | +--+
| | | +-- Lanceolaria_lanceolata
+--+ +--+
| +-- Lanceolaria_grayii[2]
|
| +-------- Inversiunio_jokohamensis **
| |
+--+ +----- Inversiunio_yanagawensis[3] **
| |
+--+ +-- Inversiunio_yanagawensis[1] **
| |
+--+-- Inversiunio_yanagawensis[2] **
|
+-- Inversiunio_reinianus **
Soroka & Burzynski, 2017 [Publ 3837], fig. 1
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 10 |
10 |
8 |
5 |
14 |
14 |
3 |
20.0 |
+-- Lanceolaria_lanceolata
+-----------------+
| +-- Lanceolaria_grayii
|
--+ +----------------- Lepidodesma_languilati
| |
| | +----- Aculamprotula_tortuosa **
+--+ +--------+
| | | +-- Aculamprotula_tientsinensis **
| | +--+
| | +-- Aculamprotula_coreana **
+--+
| +----- Schistodesmus_lampreyanus **
| +-----+
| | | +-- Tchangsinaia_piscicula **
| | +--+
+--+ +-- Nodularia_douglasiae **
|
| +-------- Unio_tumidus **
+--+
| +----- Unio_crassus **
+--+
| +-- Unio_pictorum **
+--+
+-- Unio_delphinus **
Cha et al., 2018 [Publ 3916], fig. 1
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 10 |
10 |
8 |
5 |
29 |
12 |
3 |
20.0 |
+----------------------- Lepidodesma_languilati
|
| +----- Aculamprotula_tortuosa **
| +--------+
| | | +-- Aculamprotula_coreana **
| | +--+
| | +-- Aculamprotula_tientsinensis **
| +-----+
--+ | | +----- Schistodesmus_lampreyanus **
| | | +-----+
| | | | | +-- Nodularia_douglasiae **
| | | | +--+
| | +--+ +-- Tchangsinaia_piscicula **
| | |
| | | +-------- Unio_tumidus **
| | +--+
+--+ | +----- Unio_crassus **
| +--+
| | +-- Unio_pictorum **
| +--+
| +-- Unio_delphinus **
|
| +----------------- Lanceolaria_grayii
| |
| | +----------- Anodonta_anatina
| | +--+
+--+ | | +-------- Platynaias_compressa
| | +--+
| | | +----- Pyganodon_grandis
| | +--+
+--+ | +-- Utterbackia_peninsularis
| +--+
| +-- Utterbackia_imbecillis
|
| +-------- Cristaria_plicata
| |
+-----+ +-- Anemina_arcaeformis
| +--+
| | +-- Anemina_euscaphys
+--+
| +-- Sinanodonta_lucida
+--+
+-- Sinanodonta_woodiana
Froufe et al., 2019 [Publ 3936], fig. 3
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 10 |
10 |
8 |
5 |
35 |
13 |
3 |
20.0 |
+----------------------- Lepidodesma_languilati
|
| +----- Aculamprotula_tortuosa **
| +--------+
| | | +-- Aculamprotula_tientsinensis **
| | +--+
| | +-- Aculamprotula_coreana **
| +-----+
--+ | | +----- Nodularia_douglasiae **
| | | +-----+
| | | | | +-- Tchangsinaia_piscicula **
| | | | +--+
| | +--+ +-- Schistodesmus_lampreyanus **
| | |
| | | +-------- Unio_tumidus **
| | +--+
| | | +----- Unio_crassus **
+--+ +--+
| | +-- Unio_pictorum **
| +--+
| +-- Unio_delphinus **
|
| +-- Lanceolaria_lanceolata
| +--------------+
| | +-- Lanceolaria_grayii
| |
| | +-------- Cristaria_plicata
+--+ |
| +-----+ +-- Anemina_arcaeformis
| | | +--+
| | | | +-- Anemina_euscaphys
| | +--+
+--+ | +-- Sinanodonta_lucida
| +--+
| +-- Sinanodonta_woodiana
|
| +----------- Anodonta_anatina
+--+
| +-------- Lasmigona_complanata
+--+
| +----- Pyganodon_grandis
+--+
| +-- Utterbackia_peninsularis
+--+
+-- Utterbackia_imbecillis
Froufe et al., 2019 [Publ 3936], fig. 6
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 10 |
10 |
8 |
5 |
35 |
13 |
3 |
20.0 |
+----------------------- Lepidodesma_languilati
|
| +----- Aculamprotula_tortuosa **
| +--------+
| | | +-- Aculamprotula_tientsinensis **
| | +--+
| | +-- Aculamprotula_coreana **
| +-----+
--+ | | +----- Nodularia_douglasiae **
| | | +-----+
| | | | | +-- Tchangsinaia_piscicula **
| | | | +--+
| | +--+ +-- Schistodesmus_lampreyanus **
| | |
| | | +-------- Unio_tumidus **
| | +--+
| | | +----- Unio_crassus **
+--+ +--+
| | +-- Unio_pictorum **
| +--+
| +-- Unio_delphinus **
|
| +-- Lanceolaria_lanceolata
| +--------------+
| | +-- Lanceolaria_grayii
| |
| | +----------- Anodonta_anatina
+--+ +--+
| | | +-------- Lasmigona_complanata
| | +--+
| | | +----- Pyganodon_grandis
| | +--+
+--+ | +-- Utterbackia_peninsularis
| +--+
| +-- Utterbackia_imbecillis
|
| +-------- Cristaria_plicata
| |
+-----+ +-- Sinanodonta_lucida
| +--+
| | +-- Sinanodonta_woodiana
+--+
| +-- Anemina_arcaeformis
+--+
+-- Anemina_euscaphys
Fukata & Iigo, 2019 [Publ 4108], fig. 1
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 10 |
14 |
12 |
7 |
4 |
4 |
2 |
23.0 |
+----------- Aculamprotula_tientsinensis **
+--+
| | +-------- Acuticosta_chinensis
| +--+
| | +----- Lanceolaria_gladiola
| +--+
| | +-- Lanceolaria_grayii
| +--+
--+ +-- Lanceolaria_lanceolata
|
| +-------- Unio_tumidus **
| +--+
| | | +----- Unio_crassus **
| | +--+
| | | +-- Unio_delphinus **
+--+ +--+
| +-- Unio_pictorum **
|
| +-------- Pseudocuneopsis_capitata **
| |
+--+ +-- Nodularia_douglasiae **
| +--+
| | +-- Tchangsinaia_piscicula **
+--+
| +-- Schistodesmus_lampreyanus **
+--+
+-- Cuneopsis_heudei **
Wu et al., 2020 [Publ 4147], fig. 3
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 10 |
10 |
8 |
7 |
20 |
7 |
1 |
21.0 |
+----- Cristaria_plicata
+-----+
| | +-- Sinanodonta_lucida
| +--+
+--------+ +-- Anemina_arcaeformis
| |
| | +-------- Anodonta_anatina
| +--+
| | +----- Pyganodon_grandis
| +--+
| | +-- Utterbackia_peninsularis
--+ +--+
| +-- Utterbackia_imbecillis
|
| +-------- Aculamprotula_tortuosa **
| +--------+
| | | +----- Aculamprotula_scripta **
| | +--+
| | | +-- Aculamprotula_tientsinensis **
+--+ +--+
| +-- Aculamprotula_coreana **
|
| +-------------- Unio_pictorum **
+--+
| +----------- Pseudocuneopsis_capitata **
+--+
| +-------- Nodularia_douglasiae **
+--+
| +----- Schistodesmus_lampreyanus **
+--+
| +-- Cuneopsis_heudei **
+--+
+-- Tchangsinaia_piscicula **
Lopes-Lima et al., 2021 [Publ 4217], fig. 2
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 10 |
10 |
8 |
1 |
0 |
0 |
0 |
25.0 |
+----------------- Unio_tumidus **
|
--+ +-------------- Unio_bruguierianus **
| |
| | +-- Unio_delicatus **
+--+ +--------+
| | +-- Unio_terminalis **
| |
+--+ +-- Unio_tigridis **
| +-----+
| | +-- Unio_hueti **
| |
+--+ +-- Unio_pictorum **
| +--+
| | +-- Unio_eucirrus **
+--+
| +-- Unio_elongatulus **
+--+
+-- Unio_mancus **
Zieritz et al., 2021 [Publ 4231], fig. 3-F, 5
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 10 |
10 |
8 |
5 |
39 |
13 |
3 |
20.0 |
+----------------------- Lepidodesma_languilati
|
| +----- Aculamprotula_tortuosa **
| +--------+
| | | +-- Aculamprotula_tientsinensis **
| | +--+
| | +-- Aculamprotula_coreana **
| +-----+
--+ | | +----- Nodularia_douglasiae **
| | | +-----+
| | | | | +-- Schistodesmus_lampreyanus **
| | | | +--+
| | +--+ +-- Tchangsinaia_piscicula **
| | |
| | | +-------- Unio_tumidus **
| | +--+
| | | +----- Unio_crassus **
+--+ +--+
| | +-- Unio_pictorum **
| +--+
| +-- Unio_delphinus **
|
| +-- Lanceolaria_grayii
| +--------------+
| | +-- Lanceolaria_lanceolata
| |
| | +----------- Anodonta_anatina
+--+ +--+
| | | +-------- Platynaias_compressa
| | +--+
| | | +----- Pyganodon_grandis
| | +--+
+--+ | +-- Utterbackia_imbecillis
| +--+
| +-- Utterbackia_peninsularis
|
| +-------- Cristaria_plicata
| |
+-----+ +-- Sinanodonta_lucida
| +--+
| | +-- Sinanodonta_woodiana
+--+
| +-- Anemina_arcaeformis
+--+
+-- Anemina_euscaphys
Jeratthitikul et al., 2022 [Publ 4260], fig. S1, 1
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 10 |
10 |
8 |
6 |
66 |
16 |
2 |
21.0 |
+----- Unio_tumidus **
+-----------+
| | +-- Unio_pictorum **
| +--+
+--+ +-- Unio_crassus **
| |
| | +-------------- Aculamprotula_tortuosa **
| | |
| +--+ +-- Nodularia_douglasiae **
| | +--------+
| | | +-- Nodularia_nuxpersicae **
| +--+
| | +-------- Schistodesmus_lampreyanus **
| +--+
| | +----- Cuneopsis_heudei **
--+ +--+
| | +-- Tchangsinaia_piscicula **
| +--+
| +-- Cuneopsis_rufescens **
|
| +-- Lanceolaria_grayii
| +--------------+
| | +-- Lanceolaria_gladiola
| |
| | +----------- Cristaria_plicata
| | +--+
+--+ | | +-------- Pletholophus_tenuis
| | +--+
| | | +----- Sinanodonta_lucida
| | +--+
| | | +-- Sinanodonta_lauta
| | +--+
+--+ +-- Sinanodonta_jourdyi
|
| +-------- Pseudanodonta_complanata
| +--+
| | | +----- Anodonta_nuttalliana
| | +--+
| | | +-- Anodonta_anatina
| | +--+
+--+ +-- Anodonta_cygnea
|
| +-- Pyganodon_grandis
| +-----+
| | +-- Simpsonaias_ambigua
+--+
| +----- Strophitus_undulatus
+--+
| +-- Alasmidonta_marginata
+--+
+-- Lasmigona_costata
Jeratthitikul et al., 2022 [Publ 4260], fig. S2, 2
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 10 |
10 |
8 |
6 |
66 |
16 |
2 |
21.0 |
+----- Unio_tumidus **
+-----------+
| | +-- Unio_pictorum **
| +--+
+--+ +-- Unio_crassus **
| |
| | +-------------- Aculamprotula_tortuosa **
| | |
| +--+ +-- Nodularia_douglasiae **
| | +--------+
| | | +-- Nodularia_nuxpersicae **
| +--+
| | +-------- Schistodesmus_lampreyanus **
| +--+
| | +----- Cuneopsis_heudei **
--+ +--+
| | +-- Tchangsinaia_piscicula **
| +--+
| +-- Cuneopsis_rufescens **
|
| +-- Lanceolaria_grayii
| +--------------+
| | +-- Lanceolaria_gladiola
| |
| | +----------- Cristaria_plicata
| | +--+
+--+ | | +-------- Pletholophus_tenuis
| | +--+
| | | +----- Sinanodonta_lucida
| | +--+
| | | +-- Sinanodonta_jourdyi
| | +--+
+--+ +-- Sinanodonta_lauta
|
| +-------- Pseudanodonta_complanata
| +--+
| | | +----- Anodonta_anatina
| | +--+
| | | +-- Anodonta_nuttalliana
| | +--+
+--+ +-- Anodonta_cygnea
|
| +-- Alasmidonta_marginata
| +-----+
| | +-- Lasmigona_costata
+--+
| +----- Pyganodon_grandis
+--+
| +-- Strophitus_undulatus
+--+
+-- Simpsonaias_ambigua
Zhou et al., 2007 [Publ 2658], fig. 1
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 9 |
13 |
11 |
4 |
14 |
6 |
3 |
24.0 |
+-- Schistodesmus_lampreyanus **
+-----+
| +-- Schistodesmus_spinosus **
|
+--------+ +-- Lanceolaria_triformis
| | +--+
| | | +-- Lanceolaria_lanceolata
| +--+
| | +-- Acuticosta_ovata
--+ +--+
| +-- Acuticosta_chinensis
|
| +-- Tchangsinaia_piscicula **
| +-----------+
| | +-- Nodularia_douglasiae **
+--+
| +----------- Aculamprotula_tortuosa **
+--+
| +-------- Aculamprotula_scripta **
+--+
| +----- Aculamprotula_tientsinensis **
+--+
| +-- Aculamprotula_zonata **
+--+
+-- Aculamprotula_fibrosa **
Khalloufi et al., 2011 [Publ 2748], fig. 2
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 9 |
9 |
6 |
1 |
2 |
0 |
0 |
22.0 |
+----- Unio_tumidus **
+-----+
| | +-- Unio_gibbus **
| +--+
| +-- Unio_durieui **
--+
| +-- Unio_tumidiformis **
| +-----+
| | +-- Unio_crassus **
+--+
| +----- Unio **
| |
+--+----- Unio_pictorum **
|
| +-- Unio_delphinus **
+--+
+-- Unio_mancus **
Lopes-Lima et al., 2017 [Publ 3734], fig. 1
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 9 |
9 |
5 |
1 |
8 |
3 |
1 |
22.0 |
+----- Anodonta_anatina
+--------+
| | +-- Pseudanodonta_complanata
| +--+
| +-- Anodonta_cygnea
--+
| +-- Unio_tumidus **
| +--------+
| | +-- Unio_gibbus **
| |
+--+ +-- Unio_crassus **
| +-----+
| | +-- Unio_tumidiformis **
+--+
| +----- Unio_ravoisieri **
| |
+--+ +-- Unio_mancus **
| |
| +-- Unio_pictorum **
+--+
+-- Unio_delphinus **
|
+-- Unio_elongatulus **
Huang et al., 2018 [Publ 3828], fig. 6
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 9 |
9 |
7 |
4 |
34 |
10 |
1 |
19.0 |
+----- Aculamprotula_tortuosa **
+--------+
| | +-- Aculamprotula_coreana **
| +--+
| +-- Aculamprotula_tientsinensis **
+--+
| | +-- Tchangsinaia_piscicula **
| | +--------+
| | | +-- Nodularia_douglasiae **
| +--+
| | +-------- Unio_tumidus **
| +--+
| | +----- Unio_crassus **
| +--+
--+ | +-- Unio_delphinus **
| +--+
| +-- Unio_pictorum **
|
| +----------- Anodonta_anatina
| +--+
| | | +-------- Platynaias_compressa
| | +--+
| | | +----- Pyganodon_grandis
| | +--+
+--+ | +-- Utterbackia_imbecillis
| +--+
| +-- Utterbackia_peninsularis
|
| +-------- Cristaria_plicata
| |
+-----+ +-- Anemina_arcaeformis
| +--+
| | +-- Anemina_euscaphys
+--+
| +-- Sinanodonta_woodiana
+--+
+-- Sinanodonta_lucida
Kim et al., 2021 [Publ 4197], fig. 1
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 9 |
13 |
11 |
5 |
6 |
5 |
3 |
20.0 |
+----------------- Aculamprotula_tientsinensis **
|
| +-------- Acuticosta_chinensis
| +-----+
--+ | | +----- Lanceolaria_gladiola
| | +--+
| | | +-- Lanceolaria_grayii
| | +--+
| | +-- Lanceolaria_lanceolata
+--+
| +-------- Unio_tumidus **
| +--+
| | | +----- Unio_crassus **
| | +--+
| | | +-- Unio_delphinus **
| | +--+
+--+ +-- Unio_pictorum **
|
| +-- Nodularia_douglasiae **
| +--+
| | +-- Nodularia_breviconcha **
+-----+
| +-- Schistodesmus_lampreyanus **
+--+
+-- Cuneopsis_heudei **
Araujo et al., 2009 [Publ 2589], fig. 7A
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 8 |
9 |
4 |
2 |
4 |
1 |
1 |
20.0 |
+----------- Nodularia_douglasiae **
|
| +-- Unio_gibbus **
--+ |
| +-----+-- Unio_tumidus **
| | |
| | +-- Anodonta_cygnea
+--+
| +----- Unio_pictorum[1] **
| |
| +----- Unio_pictorum[2] **
+--+
+----- Unio_mancus **
|
| +-- Unio_tumidiformis **
+--+
+-- Unio_crassus **
Araujo et al., 2009 [Publ 2589], fig. 7B
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 8 |
8 |
6 |
1 |
4 |
1 |
1 |
21.0 |
+----------------- Anodonta_cygnea
|
--+ +-------------- Unio_gibbus **
| |
+--+ +----------- Unio_tumidus **
| |
| | +-- Unio_tumidiformis **
+--+ +-----+
| | +-- Unio_crassus **
| |
+--+ +-- Unio_pictorum[1] **
| +--+
| | +-- Unio_mancus **
+--+
| +-- Unio_pictorum[2] **
+--+
+-- Unio_caffer **
Reis et al., 2013 [Publ 3580], fig. 5
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 8 |
8 |
6 |
1 |
8 |
3 |
1 |
21.0 |
+-- Anodonta_anatina
|
+--------------+-- Anodonta_cygnea
| |
| +-- Pseudanodonta_complanata
--+
| +-------------- Unio_tumidus **
| |
+--+ +-- Unio_tumidiformis **
| +--------+
| | +-- Unio_crassus **
+--+
| +-------- Unio **
| |
+--+ +-- Unio_delphinus **
| +--+
| | +-- Unio_mancus **
+--+
| +-- Unio_pictorum **
+--+
+-- Unio_caffer **
Sano et al., 2017 [Publ 3799], fig. 1
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 8 |
8 |
6 |
2 |
19 |
9 |
2 |
17.0 |
+-- Cristaria_plicata
+-----------+
| +-- Buldowskia_iwakawai
+--------+
| | +----------- Pletholophus_tenuis
| | |
| +--+ +-- Sinanodonta_tumens
| | +-----+
| | | +-- Sinanodonta_lauta
| +--+
--+ | +----- Beringiana_fukuharai
| +--+
| | +-- Beringiana_japonica
| +--+
| +-- Beringiana_beringiana
|
| +-------------------- Lanceolaria_grayii
| |
+--+ +-- Nodularia_nipponensis **
| +--------------+
| | +-- Nodularia_douglasiae **
+--+
| +-------------- Inversiunio_jokohamensis **
+--+
| +----------- Inversiunio_yanagawensis[1] **
+--+
| +-------- Inversiunio_yanagawensis[2] **
+--+
| +----- Inversiunio_yanagawensis[3] **
+--+
| +-- Inversiunio_yanagawensis[4] **
+--+
+-- Inversiunio_reinianus **
Wang et al., 2018 [Publ 3846], fig. 2
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 8 |
8 |
6 |
5 |
24 |
10 |
2 |
18.0 |
+----- Anodonta_anatina
+-----+
| | +-- Pyganodon_grandis
| +--+
+--------+ +-- Platynaias_compressa
| |
| | +-------- Cristaria_plicata
| | |
| +--+ +-- Anemina_euscaphys
| | +--+
| | | +-- Anemina_arcaeformis
| +--+
--+ | +-- Sinanodonta_woodiana
| +--+
| +-- Sinanodonta_lucida
|
| +-- Lanceolaria_grayii
| +--------------+
| | +-- Lanceolaria_lanceolata
| |
+--+ +-------------- Aculamprotula_tientsinensis **
| |
| | +----- Nodularia_douglasiae **
+--+ +-----+
| | | +-- Tchangsinaia_piscicula **
| | +--+
+--+ +-- Cuneopsis_heudei **
|
| +-------- Unio_tumidus **
+--+
| +----- Unio_crassus **
+--+
| +-- Unio_pictorum **
+--+
+-- Unio_delphinus **
Klishko et al., 2019 [Publ 3886], fig. 9
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 8 |
8 |
5 |
3 |
3 |
2 |
1 |
15.0 |
+-- Anodonta_anatina
+-----------+
| +-- Anodonta_cygnea
|
| +-------- Middendorffinaia_mongolica **
--+ +--+
| | | +----- Nodularia_breviconcha **
| | +--+
| | | +-- Nodularia_douglasiae **
+--+ +--+
| +-- Nodularia **
|
| +----- Unio_tumidus **
| |
+-----+----- Unio_gibbus **
|
| +-- Unio_crassus **
+--+
+-- Unio_pictorum **
Marrone et al., 2019 [Publ 3907], fig. 1
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 8 |
8 |
4 |
1 |
0 |
0 |
0 |
23.0 |
+-------------- Unio_tumidus **
|
--+ +----------- Unio_delphinus **
| |
+--+ +-------- Unio_foucauldianus **
| |
+--+ +----- Unio_mancus **
| |
| +----- Unio_elongatulus **
+--+
+----- Unio_ravoisieri **
|
| +-- Unio_pictorum **
+--+
+-- Unio_tigridis **
Klishko et al., 2018 [Publ 3945], fig. 8
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 8 |
8 |
5 |
2 |
0 |
0 |
0 |
17.0 |
+-------- Nodularia_breviconcha **
+--+
| | +----- Nodularia **
| +--+
| | +-- Nodularia_douglasiae **
--+ +--+
| +-- Nodularia_nipponensis **
|
| +----- Unio_tumidus **
| |
+-----+----- Unio_gibbus **
|
| +-- Unio_crassus **
+--+
+-- Unio_pictorum **
Lopes-Lima et al., 2020 [Publ 4080], fig. 13
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 8 |
8 |
6 |
2 |
2 |
0 |
0 |
17.0 |
+-------------------- Middendorffinaia_mongolica **
--+
| +----------------- Nodularia_breviconcha **
+--+
| +-------------- Nodularia[3] **
+--+
| +----------- Nodularia[2] **
+--+
| +-------- Nodularia_nuxpersicae **
+--+
| +----- Nodularia_nipponensis **
+--+
| +-- Nodularia_douglasiae **
+--+
+-- Nodularia[1] **
Araujo et al., 2009 [Publ 2589], fig. 7C
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 7 |
7 |
4 |
1 |
4 |
1 |
1 |
20.0 |
+----------- Anodonta_cygnea
|
| +-- Unio_gibbus **
--+ +-----+
| | +-- Unio_tumidus **
| |
+--+ +-- Unio_tumidiformis **
| +--+
| | +-- Unio_crassus **
+--+
| +-- Unio_pictorum[1] **
| |
+--+-- Unio_pictorum[2] **
|
+-- Unio_mancus **
Zhou et al., 2007 [Publ 2658], fig. 2
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 7 |
8 |
6 |
3 |
14 |
3 |
2 |
19.0 |
+----- Acuticosta_chinensis
+--------+
| | +-- Nodularia_douglasiae **
| +--+
--+ +-- Cuneopsis_celtiformis **
|
| +----------- Aculamprotula_tortuosa **
+--+
| +-------- Aculamprotula_scripta **
+--+
| +----- Aculamprotula_fibrosa **
+--+
| +-- Aculamprotula_zonata **
+--+
+-- Aculamprotula_tientsinensis **
Zhang et al., 2004 [Publ 2735], fig. 1
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 7 |
7 |
5 |
5 |
7 |
5 |
3 |
17.0 |
+-- Sinanodonta_woodiana
+--------------------+
| +-- Cristaria_plicata
|
--+ +----- Acuticosta_chinensis
| +--------------+
| | | +-- Lanceolaria_grayii
| | +--+
+--+ +-- Lanceolaria_lanceolata
|
| +----------------- Tchangsinaia_piscicula **
+--+
| +-------------- Cuneopsis_heudei **
+--+
| +----------- Nodularia_douglasiae **
+--+
| +-------- Pseudocuneopsis_capitata **
+--+
| +----- Cuneopsis_rufescens **
+--+
| +-- Schistodesmus_lampreyanus **
+--+
+-- Schistodesmus_spinosus **
Pfeiffer & Graf, 2013 [Publ 3531], fig. 2
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 7 |
7 |
4 |
3 |
28 |
6 |
2 |
18.0 |
+-- Cristaria_plicata
+-----+
| +-- Sinanodonta_woodiana
+--------+
| | +----- Strophitus_undulatus
| +--+
| | +-- Platynaias_compressa
--+ +--+
| +-- Pyganodon_grandis
|
| +-------------- Acuticosta_chinensis
| |
+--+ +-- Unio_pictorum **
| +--------+
| | +-- Nodularia_douglasiae **
+--+
| +-------- Aculamprotula_tortuosa **
| |
+--+ +----- Aculamprotula_scripta **
| |
+--+ +-- Aculamprotula_tientsinensis **
| |
+--+-- Aculamprotula_fibrosa **
|
+-- Aculamprotula_zonata **
Fonseca et al., 2016 [Publ 3775], fig. 1
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 7 |
7 |
5 |
5 |
25 |
12 |
2 |
19.0 |
+-- Lanceolaria_lanceolata
+-----------+
| +-- Lanceolaria_grayii
|
+--+ +-- Aculamprotula_tortuosa **
| | +--------+
| | | +-- Aculamprotula_coreana **
| | |
| +--+ +-- Unio_pictorum **
| | +-----+
| | | +-- Unio_delphinus **
| +--+
| | +----- Schistodesmus_lampreyanus **
| +--+
--+ | +-- Nodularia_douglasiae **
| +--+
| +-- Tchangsinaia_piscicula **
|
| +-------- Cristaria_plicata
| |
| +-----+ +-- Anemina_euscaphys
| | | +--+
| | | | +-- Anemina_arcaeformis
| | +--+
+--+ | +-- Sinanodonta_woodiana
| +--+
| +-- Sinanodonta_lucida
|
| +----------- Anodonta_anatina
+--+
| +-------- Platynaias_compressa
+--+
| +----- Pyganodon_grandis
+--+
| +-- Utterbackia_peninsularis
+--+
+-- Utterbackia_imbecillis
Lopes-Lima et al., 2017 [Publ 3792], fig. 2
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 7 |
7 |
5 |
5 |
26 |
12 |
2 |
19.0 |
+-- Lanceolaria_lanceolata
+-----------+
| +-- Lanceolaria_grayii
|
+--+ +-- Aculamprotula_tortuosa **
| | +--------+
| | | +-- Aculamprotula_coreana **
| | |
| +--+ +-- Unio_delphinus **
| | +-----+
| | | +-- Unio_pictorum **
| +--+
| | +----- Schistodesmus_lampreyanus **
| +--+
--+ | +-- Tchangsinaia_piscicula **
| +--+
| +-- Nodularia_douglasiae **
|
| +----------- Anodonta_anatina
| +--+
| | | +-------- Platynaias_compressa
| | +--+
| | | +----- Pyganodon_grandis
| | +--+
+--+ | +-- Utterbackia_peninsularis
| +--+
| +-- Utterbackia_imbecillis
|
| +-------- Cristaria_plicata
| |
+-----+ +-- Sinanodonta_lucida
| +--+
| | +-- Sinanodonta_woodiana
+--+
| +-- Anemina_arcaeformis
+--+
+-- Anemina_euscaphys
Wang et al., 2018 [Publ 3865], fig. 2
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 7 |
7 |
5 |
4 |
25 |
11 |
3 |
17.0 |
+-------------- Aculamprotula_tientsinensis **
|
+--+ +-- Tchangsinaia_piscicula **
| | +--------+
| | | +-- Nodularia_douglasiae **
| +--+
| | +-------- Unio_tumidus **
| +--+
| | +----- Unio_crassus **
| +--+
--+ | +-- Unio_pictorum **
| +--+
| +-- Unio_delphinus **
|
| +----- Acuticosta_chinensis
| +--------+
| | | +-- Lanceolaria_grayii
| | +--+
| | +-- Lanceolaria_lanceolata
+--+
| +----- Anodonta_anatina
| +-----+
| | | +-- Pyganodon_grandis
| | +--+
+--+ +-- Platynaias_compressa
|
| +-------- Cristaria_plicata
| |
+--+ +-- Anemina_euscaphys
| +--+
| | +-- Anemina_arcaeformis
+--+
| +-- Sinanodonta_woodiana
+--+
+-- Sinanodonta_lucida
Lopes-Lima et al., 2020 [Publ 4080], fig. 12
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 7 |
7 |
4 |
2 |
2 |
0 |
0 |
15.0 |
+-- Nodularia_breviconcha **
+--------+
| +-- Nodularia_douglasiae **
--+
| +-------- Inversiunio_jokohamensis **
| |
+--+ +----- Inversiunio_verrucosus[1] **
| |
+--+----- Inversiunio_verrucosus[2] **
|
| +-- Inversiunio_reinianus **
+--+
+-- Inversiunio_yanagawensis **
Lopes-Lima et al., 2020 [Publ 4080], fig. 5
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 7 |
7 |
5 |
2 |
2 |
0 |
0 |
18.0 |
+-- Nodularia_breviconcha **
+-----------+
| +-- Nodularia_douglasiae **
--+
| +----------- Aculamprotula_tortuosa **
+--+
| +-------- Aculamprotula_tientsinensis **
+--+
| +----- Aculamprotula_fibrosa **
+--+
| +-- Aculamprotula_scripta **
+--+
+-- Aculamprotula_coreana **
Lopes-Lima et al., 2021 [Publ 4217], fig. 6
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 7 |
7 |
5 |
1 |
0 |
0 |
0 |
20.0 |
+----------------- Unio_tumidus **
--+
| +-------------- Unio_pictorum **
+--+
| +----------- Unio_sesirmensis **
+--+
| +-------- Unio_damascensis **
+--+
| +----- Unio_courtillieri **
+--+
| +-- Unio_bruguierianus **
+--+
+-- Unio_crassus **
Dai et al., 2023 [Publ 4285], fig. 5
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 7 |
7 |
5 |
7 |
46 |
10 |
4 |
18.0 |
+-------------------- Lepidodesma_languilati
|
| +-------------- Aculamprotula_tientsinensis **
| |
| +--+ +----------- Unio_crassus **
| | | |
--+ | +--+ +-- Pseudocuneopsis_capitata **
| | | +-----+
| | | | +-- Nodularia_douglasiae **
| | +--+
| | | +----- Schistodesmus_lampreyanus **
| | +--+
+--+ | +-- Cuneopsis_celtiformis **
| +--+
| +-- Tchangsinaia_piscicula **
|
| +-- Acuticosta_chinensis
| +-----------+
| | +-- Lanceolaria_grayii
| |
+--+ +----- Cristaria_plicata
| +-----+
| | | +-- Anemina_arcaeformis
| | +--+
+--+ +-- Sinanodonta_woodiana
|
| +-------- Anodonta_cygnea
+--+
| +----- Platynaias_compressa
+--+
| +-- Pyganodon_grandis
+--+
+-- Utterbackia_imbecillis
Reis et al., 2013 [Publ 3580], fig. 4
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 6 |
6 |
4 |
1 |
8 |
3 |
1 |
19.0 |
+----- Anodonta_cygnea
+--------+
| | +-- Anodonta_anatina
| +--+
--+ +-- Pseudanodonta_complanata
|
| +----------- Unio_tumidus **
| |
+--+ +-- Unio_tumidiformis **
| +-----+
| | +-- Unio_crassus **
+--+
| +----- Unio_delphinus **
+--+
| +-- Unio_mancus **
+--+
+-- Unio_pictorum **
Bolotov et al., 2018 [Publ 3826], fig. 2
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 6 |
6 |
4 |
2 |
79 |
12 |
2 |
17.0 |
+----- Unio_tumidus **
+--+
| | +-- Unio_crassus **
| +--+
+--------------+ +-- Unio_pictorum **
| |
| | +----- Nodularia_douglasiae[1] **
| +--+
| | +-- Nodularia_nuxpersicae **
| +--+
--+ +-- Nodularia_douglasiae[2] **
|
| +-- Lanceolaria_gladiola
| +-----------------+
| | +-- Lanceolaria_grayii
| |
| | +-------- Cristaria_plicata
+--+ +--------+
| | | +----- Pletholophus_tenuis
| | +--+
| | | +-- Sinanodonta_jourdyi
+--+ +--+
| +-- Sinanodonta_lauta
|
| +-------------- Anodonta_anatina
+--+
| +----------- Pyganodon_grandis
+--+
| +-------- Simpsonaias_ambigua
+--+
| +----- Strophitus_undulatus
+--+
| +-- Alasmidonta_marginata
+--+
+-- Lasmigona_costata
Klishko et al., 2017 [Publ 3946], fig. 8
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 6 |
6 |
4 |
2 |
0 |
0 |
0 |
15.0 |
+-- Nodularia_douglasiae **
+--------+
| +-- Nodularia_breviconcha **
--+
| +-------- Unio_tumidus **
+--+
| +----- Unio_pictorum **
+--+
| +-- Unio_crassus **
+--+
+-- Unio_courtillieri **
Wu et al., 2018 [Publ 4143], fig. 2
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 6 |
6 |
4 |
3 |
16 |
7 |
1 |
18.0 |
+-- Unio_pictorum **
+--------+
| +-- Nodularia_douglasiae **
+--------+
| | +-------- Aculamprotula_tortuosa **
| +--+
| | +----- Aculamprotula_scripta **
| +--+
--+ | +-- Aculamprotula_coreana **
| +--+
| +-- Aculamprotula_tientsinensis **
|
| +----------------- Sinanodonta_lucida
+--+
| +-------------- Anemina_arcaeformis
+--+
| +----------- Cristaria_plicata
+--+
| +-------- Anodonta_anatina
+--+
| +----- Pyganodon_grandis
+--+
| +-- Utterbackia_peninsularis
+--+
+-- Utterbackia_imbecillis
Choi et al., 2020 [Publ 4169], fig. 5
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 6 |
6 |
4 |
1 |
3 |
0 |
0 |
18.0 |
+----------- Nodularia_breviconcha **
|
--+ +-- Nodularia[1] **
| +-----+
| | +-- Nodularia[2] **
+--+
| +----- Nodularia_douglasiae **
+--+
| +-- Nodularia_nipponensis **
+--+
+-- Nodularia_nuxpersicae **
Pavan-Kumar et al., 2022 [Publ 4263], fig. 4
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 6 |
6 |
4 |
5 |
30 |
9 |
4 |
18.0 |
+-------------------- Lepidodesma_languilati
|
| +-- Aculamprotula_tientsinensis **
| +--------+
| | +-- Aculamprotula_tortuosa **
--+ +-----+
| | | +-------- Unio_pictorum **
| | +--+
| | | +----- Schistodesmus_lampreyanus **
| | +--+
| | | +-- Nodularia_douglasiae **
+--+ +--+
| +-- Tchangsinaia_piscicula **
|
| +-- Lanceolaria_lanceolata
| +-----------+
| | +-- Lanceolaria_gladiola
+--+
| +----------- Acuticosta_chinensis
| |
+--+ +-- Anodonta_anatina
| +-----+
| | +-- Pyganodon_grandis
+--+
| +----- Cristaria_plicata
+--+
| +-- Sinanodonta_lucida
+--+
+-- Anemina_euscaphys
Pavan-Kumar et al., 2022 [Publ 4263], fig. S5
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 6 |
7 |
5 |
5 |
33 |
11 |
4 |
19.0 |
+-- Anodonta_anatina
+--------+
| +-- Pyganodon_grandis
+-----+
| | +-------- Cristaria_plicata
| | |
| +--+ +-- Anemina_arcaeformis
| | +--+
| | | +-- Anemina_euscaphys
| +--+
| | +-- Sinanodonta_lucida
--+ +--+
| +-- Sinanodonta_woodiana
|
| +----- Acuticosta_chinensis
| +--------+
| | | +-- Lanceolaria_lanceolata
| | +--+
| | +-- Lanceolaria_gladiola
+--+
| +----- Lepidodesma_languilati
| +-----+
| | | +-- Aculamprotula_tientsinensis **
| | +--+
+--+ +-- Aculamprotula_tortuosa **
|
| +-------- Unio_pictorum **
+--+
| +----- Schistodesmus_lampreyanus **
+--+
| +-- Nodularia_douglasiae **
+--+
+-- Tchangsinaia_piscicula **
Bashê, 2022 [Publ 4265], fig. 1
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 6 |
6 |
4 |
1 |
1 |
0 |
0 |
21.0 |
+-------------- Unio_pictorum[2] **
--+
| +----------- Unio_foucauldianus **
+--+
| +-------- Unio_delphinus **
+--+
| +----- Unio_tigridis **
+--+
| +-- Unio_elongatulus **
+--+
+-- Unio_pictorum[1] **
Dai et al., 2024 [Publ 4358], fig. 3
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 6 |
6 |
4 |
3 |
0 |
0 |
0 |
17.0 |
+-- Schistodesmus_lampreyanus **
+--------+
| +-- Nodularia_douglasiae **
--+
| +-------- Pseudocuneopsis_capitata **
+--+
| +----- Pseudocuneopsis_yangshuoensis **
+--+
| +-- Pseudocuneopsis_sichuanensis **
+--+
+-- Pseudocuneopsis_yemaoi **
Prie et al., 2012 [Publ 3487], fig. 2
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 5 |
5 |
3 |
1 |
1 |
0 |
0 |
18.0 |
+----------- Unio_tumidus **
--+
| +-------- Unio_crassus **
+--+
| +----- Unio_pictorum **
+--+
| +-- Unio_mancus **
+--+
+-- Unio_elongatulus **
Wu et al., 2016 [Publ 3711], fig. 1
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 5 |
5 |
3 |
3 |
16 |
7 |
1 |
17.0 |
+-- Unio_pictorum **
+-----+
| +-- Nodularia_douglasiae **
+-----------+
| | +----- Aculamprotula_tortuosa **
| +--+
| | +-- Aculamprotula_coreana **
--+ +--+
| +-- Aculamprotula_tientsinensis **
|
| +----------------- Sinanodonta_lucida
+--+
| +-------------- Anemina_arcaeformis
+--+
| +----------- Cristaria_plicata
+--+
| +-------- Anodonta_anatina
+--+
| +----- Pyganodon_grandis
+--+
| +-- Utterbackia_peninsularis
+--+
+-- Utterbackia_imbecillis
Huang et al., 2019 [Publ 3843], fig. 3-M
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 5 |
5 |
3 |
3 |
18 |
5 |
2 |
19.0 |
+-------- Sinanodonta_woodiana
+--------+
| | +----- Anodonta_anatina
| +--+
| | +-- Pyganodon_grandis
--+ +--+
| +-- Utterbackia_peninsularis
|
| +-------------- Lanceolaria_lanceolata
+--+
| +----------- Schistodesmus_lampreyanus **
+--+
| +-------- Aculamprotula_tortuosa **
+--+
| +----- Unio_tumidus **
+--+
| +-- Unio_crassus **
+--+
+-- Unio_delphinus **
Burzynski & Soroka, 2018 [Publ 3862], fig. 5a
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 5 |
5 |
3 |
2 |
15 |
5 |
2 |
17.0 |
+----- Sinanodonta_woodiana
|
+-----------+----- Anodonta_anatina
| |
| | +-- Utterbackia_peninsularis
--+ +--+
| +-- Pyganodon_grandis
|
| +-------------- Lanceolaria_lanceolata
+--+
| +----------- Aculamprotula_tortuosa **
+--+
| +-------- Unio_tumidus **
+--+
| +----- Unio_crassus **
+--+
| +-- Unio_pictorum **
+--+
+-- Unio_delphinus **
Burzynski & Soroka, 2018 [Publ 3862], fig. 5b
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 5 |
5 |
3 |
2 |
5 |
5 |
2 |
17.0 |
+----------------- Lanceolaria_lanceolata
|
| +-------- Sinanodonta_woodiana
--+ +-----+
| | | +----- Anodonta_anatina
| | +--+
| | | +-- Utterbackia_peninsularis
+--+ +--+
| +-- Pyganodon_grandis
|
| +----------- Aculamprotula_tortuosa **
+--+
| +-------- Unio_tumidus **
+--+
| +----- Unio_crassus **
+--+
| +-- Unio_pictorum **
+--+
+-- Unio_delphinus **
Bolotov et al., 2020 [Publ 4084], fig. 1
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 5 |
5 |
3 |
3 |
31 |
14 |
2 |
16.0 |
+----------- Middendorffinaia_mongolica **
+-----------+
| | +-------- Nodularia_douglasiae **
| +--+
| | +----- Unio_tumidus **
| +--+
| | +-- Unio_pictorum **
| +--+
--+ +-- Unio_crassus **
|
| +-- Lanceolaria_grayii
| +-----------------+
| | +-- Lanceolaria
| |
| | +----- Anodonta_anatina
+--+ +-----------+
| | | +-- Pseudanodonta_complanata
| | +--+
| | +-- Anodonta_cygnea
| |
+--+ +-------- Amuranodonta_kijaensis
| +-----+
| | | +----- Anemina_arcaeformis
| | +--+
| | | +-- Buldowskia_suifunica
+--+ +--+
| +-- Buldowskia_shadini
|
| +----------- Cristaria_plicata
+--+
| +-------- Beringiana_beringiana
+--+
| +----- Sinanodonta_schrenkii
+--+
| +-- Sinanodonta_lauta
+--+
+-- Sinanodonta_woodiana
Wu et al., 2020 [Publ 4121], fig. 2
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 5 |
5 |
3 |
2 |
29 |
0 |
0 |
18.0 |
+----------- Unio_pictorum **
--+
| +-------- Aculamprotula_tortuosa **
+--+
| +----- Aculamprotula_polysticta **
+--+
| +-- Aculamprotula_fibrosa **
+--+
+-- Aculamprotula_tientsinensis **
Wu et al., 2020 [Publ 4121], fig. S1
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 5 |
5 |
3 |
2 |
29 |
0 |
0 |
18.0 |
+----------- Unio_pictorum **
--+
| +-------- Aculamprotula_tortuosa **
+--+
| +----- Aculamprotula_polysticta **
+--+
| +-- Aculamprotula_tientsinensis **
+--+
+-- Aculamprotula_fibrosa **
Wu et al., 2020 [Publ 4147], fig. 4
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 5 |
5 |
3 |
2 |
9 |
0 |
0 |
18.0 |
+----------- Unio_pictorum **
--+
| +-------- Aculamprotula_tortuosa **
+--+
| +----- Aculamprotula_scripta **
+--+
| +-- Aculamprotula_tientsinensis **
+--+
+-- Aculamprotula_fibrosa **
Guerra et al., 2019 [Publ 4168], fig. 1
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 5 |
5 |
3 |
3 |
36 |
8 |
2 |
17.0 |
+-------------- Lanceolaria_lanceolata
+--+
| | +----------- Aculamprotula_tortuosa **
| +--+
| | +-------- Schistodesmus_lampreyanus **
| +--+
| | +----- Unio_tumidus **
--+ +--+
| | +-- Unio_crassus **
| +--+
| +-- Unio_delphinus **
|
| +-------------- Sinanodonta_woodiana
| |
+--+ +----------- Anodonta_anatina
| |
+--+ +-- Platynaias_compressa
| +-----+
| | +-- Platynaias_subviridis
+--+
| +----- Pyganodon_grandis
+--+
| +-- Utterbackia_peninsularis
+--+
+-- Utterbackia_imbecillis
Guerra et al., 2019 [Publ 4168], fig. 1-M
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 5 |
5 |
3 |
3 |
17 |
5 |
2 |
19.0 |
+-------- Sinanodonta_woodiana
+--------+
| | +----- Anodonta_anatina
| +--+
| | +-- Utterbackia_peninsularis
--+ +--+
| +-- Pyganodon_grandis
|
| +-------------- Lanceolaria_lanceolata
+--+
| +----------- Schistodesmus_lampreyanus **
+--+
| +-------- Aculamprotula_tortuosa **
+--+
| +----- Unio_tumidus **
+--+
| +-- Unio_crassus **
+--+
+-- Unio_delphinus **
Guerra et al., 2019 [Publ 4168], fig. 2
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 5 |
13 |
11 |
3 |
36 |
8 |
2 |
25.0 |
+----------------------- Aculamprotula_tortuosa **
|
| +-------- Schistodesmus_lampreyanus **
--+ +-----------+
| | | +----- Unio_tumidus **
| | +--+
| | | +-- Unio_crassus **
+--+ +--+
| +-- Unio_delphinus **
|
| +----------------- Lanceolaria_lanceolata
| |
+--+ +-------------- Sinanodonta_woodiana
| |
+--+ +----------- Anodonta_anatina
| |
+--+ +-- Platynaias_compressa
| +-----+
| | +-- Platynaias_subviridis
+--+
| +----- Pyganodon_grandis
+--+
| +-- Utterbackia_peninsularis
+--+
+-- Utterbackia_imbecillis
Guerra et al., 2019 [Publ 4168], fig. 2-M
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 5 |
5 |
3 |
3 |
17 |
5 |
2 |
17.0 |
+-- Utterbackia_peninsularis
+--+
| +-- Pyganodon_grandis
+-----------+
| | +-- Anodonta_anatina
| +--+
--+ +-- Sinanodonta_woodiana
|
| +-------------- Lanceolaria_lanceolata
+--+
| +----------- Aculamprotula_tortuosa **
+--+
| +-------- Schistodesmus_lampreyanus **
+--+
| +----- Unio_tumidus **
+--+
| +-- Unio_crassus **
+--+
+-- Unio_delphinus **
Choi et al., 2020 [Publ 4169], fig. 3a
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 5 |
5 |
3 |
3 |
0 |
0 |
0 |
14.0 |
+-- Cuneopsis_heudei **
+-----+
| +-- Unio_pictorum **
--+
| +----- Nodularia_breviconcha **
+--+
| +-- Nodularia_douglasiae **
+--+
+-- Nodularia_nipponensis **
Mabuchi et al., 2021 [Publ 4233], fig. 1
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 5 |
5 |
3 |
2 |
0 |
0 |
0 |
16.0 |
+-------- Tchangsinaia_piscicula **
|
--+ +-- Nodularia_douglasiae[1] **
| +--+
| | +-- Nodularia_nipponensis **
+--+
| +-- Nodularia_douglasiae[2] **
+--+
+-- Nodularia_breviconcha **
Pfarrer et al., 2022 [Publ 4305], fig. 2-5
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 5 |
5 |
3 |
1 |
7 |
5 |
1 |
18.0 |
+----------- Unio_tumidus **
+--+
| | +-------- Unio_crassus **
| +--+
| | +----- Unio_elongatulus **
| +--+
| | +-- Unio_mancus **
--+ +--+
| +-- Unio_pictorum **
|
| +-- Cristaria_plicata
| +-----+
| | +-- Sinanodonta_woodiana
+-----+
| +----- Anodonta_anatina
+--+
| +-- Anodonta_exulcerata
+--+
+-- Anodonta_cygnea
Nagel et al., 1998 [Publ 1946], fig. 1
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 4 |
4 |
2 |
1 |
5 |
3 |
1 |
17.0 |
+----- Pseudanodonta_complanata
+-----+
| | +-- Anodonta_anatina
| +--+
--+ +-- Anodonta_cygnea
|
| +-------- Unio_crassus **
+--+
| +----- Unio_tumidus **
+--+
| +-- Unio_pictorum **
+--+
+-- Unio_mancus **
Nagel et al., 1998 [Publ 1946], fig. 8
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 4 |
4 |
1 |
1 |
0 |
0 |
0 |
17.0 |
+----- Unio_tumidus **
|
--+ +-- Unio_pictorum **
| |
+--+-- Unio_crassus **
|
+-- Unio_mancus **
Soroka, 2005 [Publ 2726], fig. 3
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 4 |
4 |
2 |
1 |
1 |
0 |
0 |
17.0 |
+-------- Unio_tumidus **
--+
| +----- Unio_crassus **
+--+
| +-- Unio_caffer **
+--+
+-- Unio_pictorum **
Soroka, 2010 [Publ 3404], fig. 3a
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 4 |
4 |
2 |
1 |
5 |
5 |
1 |
17.0 |
+-- Sinanodonta_woodiana
+-----------+
| +-- Sinanodonta_lauta
|
--+ +----- Anodonta_anatina
| +-----+
| | | +-- Anodonta_cygnea
| | +--+
+--+ +-- Pseudanodonta_complanata
|
| +-------- Unio_tumidus **
+--+
| +----- Unio_crassus **
+--+
| +-- Unio_pictorum **
+--+
+-- Unio_mancus **
Soroka, 2010 [Publ 3404], fig. 3b
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 4 |
4 |
2 |
1 |
5 |
5 |
1 |
17.0 |
+-------------- Anodonta_anatina
|
+-------------- Anodonta_cygnea
|
--+-------------- Pseudanodonta_complanata
|
| +-- Sinanodonta_woodiana
| +--------+
| | +-- Sinanodonta_lauta
+--+
| +-------- Unio_tumidus **
+--+
| +----- Unio_crassus **
+--+
| +-- Unio_pictorum **
+--+
+-- Unio_mancus **
Xue et al., 2016 [Publ 3659], fig. 1
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 4 |
4 |
2 |
4 |
12 |
8 |
2 |
16.0 |
+-- Cristaria_plicata
+-----+
| +-- Platynaias_compressa
|
+-----+ +-- Anemina_arcaeformis
| | +--+
| | | +-- Anemina_euscaphys
| +--+
| | +-- Sinanodonta_lucida
--+ +--+
| +-- Sinanodonta_woodiana
|
| +-- Lanceolaria_lanceolata
| +--------+
| | +-- Lanceolaria_grayii
+--+
| +-------- Aculamprotula_tortuosa **
+--+
| +----- Schistodesmus_lampreyanus **
+--+
| +-- Nodularia_douglasiae **
+--+
+-- Unio_pictorum **
Jadhav & Jamkhedkar, 2009 [Publ 3697], fig. 11
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 4 |
10 |
7 |
2 |
22 |
6 |
1 |
23.0 |
+-------------------- Nodularia_douglasiae **
|
--+ +----------------- Anodonta_cygnea
| |
+--+ +-------------- Unio_pictorum[2] **
| |
+--+ +-- Unio_pictorum[1] **
| +--------+
| | +-- Unio_caffer **
+--+
| +-------- Anodonta
| |
+--+ +----- Lasmigona_costata
| |
+--+ +-- Strophitus_undulatus
| |
+--+-- Alasmidonta_marginata
|
+-- Pyganodon_grandis
Zhou et al., 2017 [Publ 3788], fig. 2
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 4 |
4 |
2 |
3 |
13 |
3 |
2 |
16.0 |
+-------- Lanceolaria_lanceolata
|
--+ +-- Nodularia_douglasiae **
| +--+
| | +-- Tchangsinaia_piscicula **
+--+
| +-- Aculamprotula_tientsinensis **
+--+
+-- Aculamprotula_tortuosa **
Huang et al., 2018 [Publ 3828], fig. 6-M
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 4 |
4 |
2 |
2 |
13 |
4 |
1 |
18.0 |
+-------- Sinanodonta_woodiana
+--+
| | +----- Anodonta_anatina
| +--+
| | +-- Pyganodon_grandis
--+ +--+
| +-- Utterbackia_peninsularis
|
| +-------- Aculamprotula_tortuosa **
+--+
| +----- Unio_tumidus **
+--+
| +-- Unio_crassus **
+--+
+-- Unio_delphinus **
Huang et al., 2002 [Publ 2657], fig. 1
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 3 |
7 |
5 |
3 |
31 |
11 |
4 |
20.0 |
+-- Sinanodonta_woodiana
+-----------+
| +-- Cristaria_plicata
+--+
| | +----------- Utterbackiana_couperiana
| +--+
| | +-------- Utterbackia_imbecillis
| +--+
| | +----- Strophitus_radiatus
| +--+
| | +-- Alasmidonta_triangulata
--+ +--+
| +-- Lasmigona_costata
|
| +----- Acuticosta_ovata
| +--+
| | | +-- Nodularia_douglasiae **
| | +--+
| | +-- Tchangsinaia_piscicula **
+--------+
| +-- Lanceolaria_grayii
| +--+
| | +-- Lanceolaria_lanceolata
+--+
| +-- Schistodesmus_lampreyanus **
+--+
+-- Lepidodesma_languilati
Huang et al., 2002 [Publ 2657], fig. 2
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 3 |
14 |
5 |
3 |
31 |
11 |
4 |
27.0 |
+-------- Strophitus_radiatus
|
+-------- Schistodesmus_lampreyanus **
|
+-------- Lepidodesma_languilati
|
+-------- Acuticosta_ovata
|
| +-- Sinanodonta_woodiana
+-----+
| +-- Cristaria_plicata
--+
| +-- Lanceolaria_grayii
+-----+
| +-- Lanceolaria_lanceolata
|
| +-- Nodularia_douglasiae **
+-----+
| +-- Tchangsinaia_piscicula **
|
| +----- Utterbackia_imbecillis
| |
+--+----- Utterbackiana_couperiana
|
| +-- Alasmidonta_triangulata
+--+
+-- Lasmigona_costata
Manendo et al., 2008 [Publ 2669], fig. 1
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 3 |
3 |
1 |
1 |
10 |
5 |
1 |
16.0 |
+----- Unio_crassus **
+-----+
| | +-- Unio_pictorum **
| +--+
| +-- Unio_tumidus **
--+
| +-- Anodonta_cygnea
| +-----+
| | +-- Anodonta_anatina
+--+
| +----- Pseudanodonta_complanata
+--+
| +-- Platynaias_subviridis
+--+
+-- Pyganodon_grandis
Källersjö et al., 2005 [Publ 2684], fig. 1A
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 3 |
3 |
0 |
1 |
4 |
3 |
1 |
16.0 |
+-- Unio_crassus **
|
--+-- Unio_pictorum **
|
+-- Unio_tumidus **
Källersjö et al., 2005 [Publ 2684], fig. 1B-F
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 3 |
3 |
1 |
1 |
4 |
3 |
1 |
16.0 |
+----- Unio_tumidus **
--+
| +-- Unio_crassus **
+--+
+-- Unio_pictorum **
Soroka, 2010 [Publ 3404], fig. 2
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 3 |
3 |
0 |
1 |
4 |
4 |
1 |
16.0 |
+----------- Sinanodonta_woodiana
|
| +-- Unio_tumidus **
--+ |
| +-----+-- Unio_pictorum **
| | |
+--+ +-- Unio_crassus **
|
| +----- Anodonta_cygnea
+--+
| +-- Anodonta_anatina
+--+
+-- Pseudanodonta_complanata
Soroka, 2010 [Publ 3404], fig. 2-M
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 3 |
3 |
1 |
1 |
2 |
2 |
1 |
16.0 |
+----------- Sinanodonta_woodiana
--+
| +-------- Anodonta_anatina
+--+
| +----- Unio_crassus **
+--+
| +-- Unio_tumidus **
+--+
+-- Unio_pictorum **
Soroka, 2010 [Publ 3404], fig. 3a-M
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 3 |
3 |
1 |
1 |
3 |
3 |
1 |
16.0 |
+----------- Anodonta_anatina
|
--+ +-- Sinanodonta_woodiana
| +-----+
| | +-- Sinanodonta_lauta
+--+
| +----- Unio_crassus **
+--+
| +-- Unio_tumidus **
+--+
+-- Unio_pictorum **
Soroka, 2010 [Publ 3404], fig. 3b-M
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 3 |
3 |
1 |
1 |
3 |
3 |
1 |
16.0 |
+----------- Anodonta_anatina
|
--+ +-- Sinanodonta_lauta
| +-----+
| | +-- Sinanodonta_woodiana
+--+
| +----- Unio_tumidus **
+--+
| +-- Unio_crassus **
+--+
+-- Unio_pictorum **
Soroka, 2010 [Publ 3404], fig. 4
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 3 |
3 |
1 |
1 |
4 |
4 |
1 |
16.0 |
+----- Sinanodonta_woodiana
+-----+
| | +-- Anodonta_cygnea
| +--+
--+ +-- Pseudanodonta_complanata
|
| +-------- Anodonta_anatina
+--+
| +----- Unio_tumidus **
+--+
| +-- Unio_pictorum **
+--+
+-- Unio_crassus **
Soroka, 2010 [Publ 3404], fig. 5a
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 3 |
3 |
1 |
1 |
5 |
5 |
1 |
16.0 |
+----- Unio_crassus **
+-----+
| | +-- Unio_pictorum **
| +--+
--+ +-- Unio_tumidus **
|
| +-------- Anodonta_anatina
| |
+--+ +-- Anodonta_cygnea
| +--+
| | +-- Pseudanodonta_complanata
+--+
| +-- Sinanodonta_woodiana
+--+
+-- Sinanodonta_schrenkii
Soroka, 2010 [Publ 3404], fig. 5b
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 3 |
3 |
1 |
1 |
5 |
5 |
1 |
16.0 |
+----------- Anodonta_anatina
|
| +-- Anodonta_cygnea
--+ +-----+
| | +-- Pseudanodonta_complanata
| |
| | +-- Sinanodonta_woodiana
+--+-----+
| +-- Sinanodonta_schrenkii
|
| +----- Unio_tumidus **
+--+
| +-- Unio_pictorum **
+--+
+-- Unio_crassus **
Soroka, 2010 [Publ 3404], fig. 8a
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 3 |
3 |
1 |
1 |
3 |
3 |
1 |
16.0 |
+----- Unio_tumidus **
+--+
| | +-- Unio_pictorum **
| +--+
--+ +-- Unio_crassus **
|
| +----- Sinanodonta_woodiana
+--+
| +-- Anodonta_anatina
+--+
+-- Anodonta_cygnea
Soroka, 2010 [Publ 3404], fig. 8b
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 3 |
3 |
1 |
1 |
3 |
3 |
1 |
16.0 |
+----------- Sinanodonta_woodiana
|
--+ +-- Anodonta_anatina
| +-----+
| | +-- Anodonta_cygnea
+--+
| +----- Unio_tumidus **
+--+
| +-- Unio_pictorum **
+--+
+-- Unio_crassus **
Ouyang et al., 2011 [Publ 3408], fig. 1
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 3 |
6 |
3 |
3 |
10 |
6 |
3 |
18.0 |
+----- Anemina_arcaeformis
+-----+
| | +-- Sinanodonta_elliptica
| +--+
| +-- Sinanodonta_woodiana
--+
| +-- Nodularia_douglasiae **
| +-----+
| | +-- Cuneopsis_heudei **
+--+
| +----- Lanceolaria_grayii
| |
+--+----- Acuticosta_chinensis
|
| +-- Lanceolaria_gladiola
+--+
+-- Aculamprotula_tortuosa **
Zieritz et al., 2012 [Publ 3673], fig. 4
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 3 |
3 |
1 |
1 |
5 |
4 |
1 |
16.0 |
+----- Unio_crassus **
--+
| +-- Unio_tumidus **
+--+
+-- Unio_pictorum **
Plazzi et al., 2011 [Publ 3676], fig. 5
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 3 |
3 |
1 |
1 |
15 |
6 |
2 |
16.0 |
+----- Unio_tumidus **
--+
| +-- Unio_crassus **
+--+
+-- Unio_pictorum **
Burzynski et al., 2017 [Publ 3774], fig. 1 bottom
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 3 |
3 |
1 |
2 |
10 |
3 |
1 |
17.0 |
+----- Aculamprotula_tortuosa **
+--+
| | +-- Unio_crassus **
| +--+
--+ +-- Unio_tumidus **
|
| +----- Anodonta_anatina
+--+
| +-- Utterbackia_peninsularis
+--+
+-- Pyganodon_grandis
Burzynski et al., 2017 [Publ 3774], fig. 1 top
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 3 |
3 |
1 |
2 |
10 |
3 |
1 |
17.0 |
+----- Aculamprotula_tortuosa **
+--+
| | +-- Unio_crassus **
| +--+
--+ +-- Unio_tumidus **
|
| +----- Anodonta_anatina
+--+
| +-- Utterbackia_peninsularis
+--+
+-- Pyganodon_grandis
Wen et al., 2017 [Publ 3795], fig. 3
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 3 |
4 |
2 |
2 |
13 |
4 |
2 |
18.0 |
+----- Anodonta_anatina
+--+
| | +-- Sinanodonta_woodiana
| +--+
| +-- Cristaria_plicata
--+
| +-- Aculamprotula_coreana **
| +--+
| | +-- Aculamprotula_tortuosa **
+--+
| +-- Lepidodesma_languilati
+--+
+-- Unio_pictorum **
Araujo et al., 2017 [Publ 3801], fig. 2
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 3 |
4 |
2 |
3 |
11 |
1 |
1 |
19.0 |
+-- Schistodesmus_lampreyanus **
+--+
| +-- Aculamprotula_tortuosa **
--+
| +-- Anodonta_anatina
+--+
+-- Unio_delphinus **
Froufe et al., 2017 [Publ 3802], fig. 7
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 3 |
3 |
1 |
1 |
7 |
4 |
1 |
18.0 |
+----- Unio_mancus **
+-----+
| | +-- Unio_elongatulus **
| +--+
--+ +-- Unio_pictorum **
|
| +-------- Sinanodonta_woodiana
+--+
| +----- Anodonta_anatina
+--+
| +-- Anodonta_exulcerata
+--+
+-- Anodonta_cygnea
Dyduch-Falniowska & Koziol, 1989 [Publ 3814], fig. 5, 7
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 3 |
3 |
1 |
1 |
2 |
2 |
1 |
16.0 |
+-- Pseudanodonta_complanata
+-----+
| +-- Anodonta_anatina
--+
| +----- Unio_pictorum **
+--+
| +-- Unio_tumidus **
+--+
+-- Unio_crassus **
Bogatov et al., 2018 [Publ 3859], fig. 1
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 3 |
5 |
3 |
1 |
3 |
2 |
1 |
18.0 |
+-------- Unio_tumidus **
|
--+ +-- Anodonta_anatina
| +--+
| | +-- Pseudanodonta_complanata
+--+
| +-- Unio_crassus **
+--+
+-- Unio_pictorum **
Bogatov et al., 2018 [Publ 3859], fig. 2
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 3 |
5 |
3 |
1 |
3 |
2 |
1 |
18.0 |
+----------- Unio_crassus **
--+
| +-------- Unio_pictorum **
+--+
| +----- Unio_tumidus **
+--+
| +-- Anodonta_anatina
+--+
+-- Pseudanodonta_complanata
Froufe et al., 2019 [Publ 3936], fig. 3-M
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 3 |
3 |
1 |
1 |
16 |
4 |
1 |
18.0 |
+----- Unio_tumidus **
+-----+
| | +-- Unio_crassus **
| +--+
--+ +-- Unio_delphinus **
|
| +-------- Sinanodonta_woodiana
+--+
| +----- Anodonta_anatina
+--+
| +-- Utterbackia_peninsularis
+--+
+-- Pyganodon_grandis
Froufe et al., 2019 [Publ 3936], fig. 4
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 3 |
3 |
1 |
1 |
16 |
4 |
1 |
18.0 |
+----- Unio_tumidus **
+-----+
| | +-- Unio_crassus **
| +--+
--+ +-- Unio_delphinus **
|
| +-------- Sinanodonta_woodiana
+--+
| +----- Anodonta_anatina
+--+
| +-- Utterbackia_peninsularis
+--+
+-- Pyganodon_grandis
Pfeiffer et al., 2019 [Publ 3937], fig. 4-5
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 3 |
3 |
1 |
3 |
54 |
5 |
2 |
17.0 |
+----------- Lanceolaria_lanceolata
|
| +----- Unio_gibbus **
--+ +--+
| | | +-- Nodularia_douglasiae **
| | +--+
| | +-- Tchangsinaia_piscicula **
+--+
| +-- Cristaria_plicata
| +--+
| | +-- Sinanodonta_woodiana
+--+
| +-- Anodonta_cygnea
+--+
+-- Pyganodon_grandis
Pfeiffer et al., 2019 [Publ 3937], fig. 6
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 3 |
3 |
1 |
3 |
54 |
5 |
2 |
17.0 |
+-------- Lanceolaria_lanceolata
+--+
| | +----- Unio_gibbus **
| +--+
| | +-- Tchangsinaia_piscicula **
| +--+
--+ +-- Nodularia_douglasiae **
|
| +-- Cristaria_plicata
| +--+
| | +-- Sinanodonta_woodiana
+-----+
| +-- Anodonta_cygnea
+--+
+-- Pyganodon_grandis
Chen et al., 2020 [Publ 4170], fig. 1
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 3 |
4 |
2 |
2 |
31 |
12 |
2 |
18.0 |
+-------- Schistodesmus_lampreyanus **
+-----+
| | +----- Lepidodesma_languilati
| +--+
| | +-- Aculamprotula_coreana **
| +--+
--+ +-- Aculamprotula_tortuosa **
|
| +----- Platynaias_compressa
| +-----+
| | | +-- Utterbackia_imbecillis
| | +--+
+--+ +-- Pyganodon_grandis
|
| +-------- Sinanodonta_lucida
+--+
| +----- Cristaria_plicata
+--+
| +-- Anodonta_cygnea
+--+
+-- Anodonta_anatina
Wu et al., 2021 [Publ 4215], fig. 3
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 3 |
3 |
1 |
2 |
21 |
1 |
1 |
16.0 |
+-------- Acuticosta_chinensis
--+
| +----- Nodularia_douglasiae **
+--+
| +-- Inversiunio_jokohamensis **
+--+
+-- Inversiunio_reinianus **
Sonowal et al., 2021 [Publ 4221], fig. 3a
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 3 |
4 |
0 |
2 |
12 |
1 |
1 |
18.0 |
+-- Anodonta_cygnea
|
+-- Unio_ravoisieri **
--+
+-- Unio_pictorum **
|
+-- Aculamprotula_tientsinensis **
Sonowal et al., 2021 [Publ 4221], fig. 3b
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 3 |
9 |
0 |
2 |
12 |
1 |
1 |
23.0 |
+-- Aculamprotula_tientsinensis **
|
+-- Lamprotula_leaii
|
+-- Lampsilis_cardium
|
+-- Potomida_littoralis
|
--+-- Unio_pictorum **
|
+-- Anodonta_cygnea
|
+-- Unio_ravoisieri **
|
+-- Lamellidens_mainwaringi
|
+-- Elliptio_complanata
Sonowal et al., 2021 [Publ 4221], fig. 4
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 3 |
5 |
3 |
2 |
13 |
1 |
1 |
19.0 |
+----------- Aculamprotula_tientsinensis **
--+
| +-------- Unio_pictorum **
+--+
| +----- Anodonta_cygnea
+--+
| +-- Unio_ravoisieri **
+--+
+-- Lamellidens_corrianus
Zieritz et al., 2021 [Publ 4231], fig. 3-M, 4
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 3 |
3 |
1 |
1 |
20 |
4 |
1 |
18.0 |
+----- Unio_tumidus **
+-----+
| | +-- Unio_delphinus **
| +--+
--+ +-- Unio_crassus **
|
| +-------- Sinanodonta_woodiana
+--+
| +----- Anodonta_anatina
+--+
| +-- Utterbackia_peninsularis
+--+
+-- Pyganodon_grandis
Graf & Cummings, 2006 [Publ 1968], fig. 2
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 2 |
2 |
0 |
1 |
45 |
4 |
1 |
-- |
+-- Unio_pictorum **
+--------+
| +-- Unio_caffer **
--+
| +-------- Lasmigona_costata
+--+
| +----- Alasmidonta_marginata
+--+
| +-- Pyganodon_grandis
+--+
+-- Strophitus_undulatus
Graf & Ó Foighil, 2000 [Publ 2648], fig. 2A
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 2 |
2 |
0 |
1 |
28 |
6 |
1 |
-- |
+-- Unio_pictorum **
--+
+-- Unio_caffer **
Graf & Ó Foighil, 2000 [Publ 2648], fig. 2B
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 2 |
2 |
0 |
1 |
28 |
6 |
1 |
-- |
+-- Unio_pictorum **
--+
+-- Unio_caffer **
Bogan & Hoeh, 2000 [Publ 2650], fig. 1
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 2 |
2 |
0 |
1 |
32 |
3 |
1 |
-- |
+-- Unio_pictorum **
--+
+-- Unio_tumidus **
Graf, 2002 [Publ 2651], fig. 2
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 2 |
2 |
0 |
1 |
16 |
2 |
1 |
-- |
+-- Unio_pictorum **
+--+
| +-- Unio_caffer **
--+
| +-- Alasmidonta_marginata
+--+
+-- Pyganodon_grandis
Hoeh et al., 2001 [Publ 2652], fig. 14.2
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 2 |
2 |
0 |
1 |
29 |
3 |
1 |
-- |
+-- Unio_pictorum **
--+
+-- Unio_tumidus **
Hoeh et al., 2001 [Publ 2652], fig. 14.4
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 2 |
5 |
0 |
1 |
29 |
3 |
1 |
20.0 |
+-------- Unio_pictorum **
|
--+-------- Unio_tumidus **
|
| +----- Anodonta_cygnea
+--+
| +-- Strophitus_undulatus
+--+
+-- Pyganodon_grandis
Hoeh et al., 2002 [Publ 2653], fig. 2
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 2 |
5 |
0 |
1 |
29 |
3 |
1 |
20.0 |
+----------- Unio_pictorum **
--+
| +-------- Unio_tumidus **
+--+
| +----- Anodonta_cygnea
+--+
| +-- Pyganodon_grandis
+--+
+-- Strophitus_undulatus
Roe & Hoeh, 2003 [Publ 2655], fig. 4.2
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 2 |
2 |
0 |
1 |
29 |
3 |
1 |
-- |
+-- Unio_pictorum **
+-----+
| +-- Unio_tumidus **
--+
| +----- Pyganodon_grandis
+--+
| +-- Anodonta_cygnea
+--+
+-- Strophitus_undulatus
Roe & Hoeh, 2003 [Publ 2655], fig. 4.3
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 2 |
2 |
0 |
1 |
29 |
3 |
1 |
-- |
+-- Unio_pictorum **
+-----+
| +-- Unio_tumidus **
--+
| +----- Anodonta_cygnea
+--+
| +-- Pyganodon_grandis
+--+
+-- Strophitus_undulatus
Hoeh et al., 2009 [Publ 2716], fig. 3
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 2 |
2 |
0 |
1 |
45 |
4 |
1 |
-- |
+-- Unio_pictorum **
+--------+
| +-- Unio_caffer **
--+
| +-------- Lasmigona_costata
+--+
| +----- Alasmidonta_marginata
+--+
| +-- Strophitus_undulatus
+--+
+-- Pyganodon_grandis
Hoeh et al., 2009 [Publ 2716], fig. 4
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 2 |
2 |
0 |
1 |
44 |
4 |
1 |
-- |
+-- Unio_pictorum **
+--------+
| +-- Unio_caffer **
--+
| +-------- Lasmigona_costata
+--+
| +----- Alasmidonta_marginata
+--+
| +-- Strophitus_undulatus
+--+
+-- Pyganodon_grandis
Hoeh et al., 2009 [Publ 2716], fig. 5
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 2 |
2 |
0 |
1 |
44 |
4 |
1 |
-- |
+-- Unio_pictorum **
+--------+
| +-- Unio_caffer **
--+
| +-------- Lasmigona_costata
+--+
| +----- Alasmidonta_marginata
+--+
| +-- Strophitus_undulatus
+--+
+-- Pyganodon_grandis
Rosenberg et al., 1997 [Publ 2720], fig. 2B
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 2 |
17 |
0 |
2 |
21 |
4 |
1 |
31.0 |
+----- Anodonta
|
| +-- Gonidea_angulata
| |
| +-- Pyganodon_grandis
| |
| +-- Utterbackia_imbecillis
| |
| +-- Pyganodon_cataracta
--+ |
| +-- Lampsilis_straminea
| |
| +-- Lampsilis_teres
| |
| +-- Megalonaias_nervosa
| |
| +-- Obliquaria_reflexa
| |
+--+-- Theliderma_cylindrica
|
+-- Quadrula_quadrula
|
+-- Uniomerus_tetralasmus
|
+-- Amblema_plicata
|
+-- Elliptio_complanata
|
+-- Fusconaia_cerina
|
+-- Plectomerus_dombeyanus
|
+-- Aculamprotula_tortuosa **
|
+-- Unio_pictorum **
Whelan et al., 2011 [Publ 3380], fig. 1 ML
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 2 |
2 |
0 |
1 |
35 |
2 |
1 |
-- |
+-- Pyganodon_grandis
+--+
| +-- Lasmigona_costata
--+
| +-- Unio_pictorum **
+--+
+-- Unio_caffer **
Whelan et al., 2011 [Publ 3380], fig. 1 MP
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 2 |
2 |
0 |
1 |
35 |
2 |
1 |
-- |
+-- Pyganodon_grandis
+--+
| +-- Lasmigona_costata
--+
| +-- Unio_pictorum **
+--+
+-- Unio_caffer **
Soroka, 2010 [Publ 3404], fig. 8a-M
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 2 |
2 |
0 |
1 |
1 |
1 |
1 |
-- |
+----- Sinanodonta_woodiana
--+
| +-- Unio_pictorum **
+--+
+-- Unio_tumidus **
Soroka, 2010 [Publ 3404], fig. 8b-M
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 2 |
2 |
0 |
1 |
1 |
1 |
1 |
-- |
+----- Sinanodonta_woodiana
--+
| +-- Unio_pictorum **
+--+
+-- Unio_tumidus **
Zhou et al., 2016 [Publ 3710], fig. 1
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 2 |
2 |
0 |
2 |
18 |
7 |
2 |
-- |
+----- Lepidodesma_languilati
+-----------+
| | +-- Schistodesmus_lampreyanus **
| +--+
--+ +-- Unio_pictorum **
|
| +-------------- Sinanodonta_woodiana
+--+
| +----------- Cristaria_plicata
+--+
| +-------- Platynaias_compressa
+--+
| +----- Pyganodon_grandis
+--+
| +-- Utterbackia_imbecillis
+--+
+-- Utterbackia_peninsularis
Bolotov et al., 2016 [Publ 3743], fig. 2
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 2 |
2 |
0 |
1 |
28 |
1 |
1 |
-- |
+----- Anodonta_anatina
--+
| +-- Unio_pictorum **
+--+
+-- Unio_ravoisieri **
Bolotov et al., 2016 [Publ 3743], fig. 4
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 2 |
2 |
0 |
1 |
28 |
1 |
1 |
-- |
+----- Anodonta_anatina
--+
| +-- Unio_pictorum **
+--+
+-- Unio_ravoisieri **
Soroka & Burzynski, 2018 [Publ 3762], fig. 3
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 2 |
2 |
0 |
2 |
5 |
3 |
1 |
-- |
+-- Unio_tumidus **
+-----+
| +-- Aculamprotula_tortuosa **
--+
| +----- Anodonta_anatina
+--+
| +-- Pyganodon_grandis
+--+
+-- Utterbackia_peninsularis
Soroka & Burzynski, 2018 [Publ 3762], fig. 3-M
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 2 |
2 |
0 |
2 |
5 |
3 |
1 |
-- |
+-- Unio_tumidus **
+-----+
| +-- Aculamprotula_tortuosa **
--+
| +----- Anodonta_anatina
+--+
| +-- Pyganodon_grandis
+--+
+-- Utterbackia_peninsularis
Guerra et al., 2017 [Publ 3772], fig. 2a
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 2 |
2 |
0 |
2 |
23 |
8 |
1 |
-- |
+-- Aculamprotula_tortuosa **
+-----------------+
| +-- Unio_pictorum **
--+
| +----------------- Sinanodonta_woodiana
| |
+--+ +-------------- Cristaria_plicata
| |
+--+ +----------- Anodonta_anatina
| |
+--+ +-- Platynaias_subviridis
| +-----+
| | +-- Platynaias_compressa
+--+
| +----- Pyganodon_grandis
+--+
| +-- Utterbackia_peninsularis
+--+
+-- Utterbackia_imbecillis
Guerra et al., 2017 [Publ 3772], fig. 2b
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 2 |
10 |
0 |
2 |
23 |
8 |
1 |
24.0 |
+-------------------- Unio_pictorum **
|
--+ +----------------- Aculamprotula_tortuosa **
| |
+--+ +-- Cristaria_plicata
| +-----------+
| | +-- Sinanodonta_woodiana
+--+
| +----------- Anodonta_anatina
| |
+--+ +-- Platynaias_subviridis
| +-----+
| | +-- Platynaias_compressa
+--+
| +----- Pyganodon_grandis
+--+
| +-- Utterbackia_peninsularis
+--+
+-- Utterbackia_imbecillis
Huang et al., 2018 [Publ 3828], fig. 3, 5
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 2 |
2 |
0 |
2 |
27 |
0 |
0 |
-- |
+-- Unio_pictorum **
--+
+-- Aculamprotula_tientsinensis **
Chase et al., 2018 [Publ 3847], fig. 4
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 2 |
2 |
0 |
1 |
37 |
15 |
1 |
-- |
+-- Unio_delphinus **
+--------------------+
| +-- Unio_tumidus **
|
--+ +-------------------- Sinanodonta_woodiana
| |
| | +-- Anodonta_cygnea
| | +--------------+
+--+ | +-- Anodonta_anatina
| |
| | +-- Pyganodon_lacustris
| | +-----+
+--+ | +-- Pyganodon_grandis
| +-----+
| | | +----- Utterbackia_peninsularis[2]
| | +--+
| | | +-- Utterbackia_peninsularis[1]
| | +--+
+--+ +-- Utterbackia_imbecillis
|
| +-- Lasmigona_costata
| +--------+
| | +-- Lasmigona_complanata
| |
+--+ +-- Alasmidonta_undulata
| +-----+
| | +-- Alasmidonta_marginata
+--+
| +----- Strophitus_undulatus
+--+
| +-- Platynaias_compressa
+--+
+-- Platynaias_subviridis
Chase et al., 2018 [Publ 3847], fig. 5
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 2 |
2 |
0 |
1 |
8 |
2 |
1 |
-- |
+-- Anodonta_anatina
+--+
| +-- Utterbackia_peninsularis
--+
| +-- Unio_delphinus **
+--+
+-- Unio_tumidus **
Chase et al., 2018 [Publ 3861], fig. 1
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 2 |
2 |
0 |
1 |
9 |
3 |
1 |
-- |
+-- Unio_delphinus **
+-----+
| +-- Unio_tumidus **
--+
| +----- Anodonta_anatina
+--+
| +-- Utterbackia_peninsularis
+--+
+-- Pyganodon_grandis
Chase et al., 2018 [Publ 3861], fig. 1-M
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 2 |
2 |
0 |
1 |
9 |
3 |
1 |
-- |
+-- Unio_delphinus **
+-----+
| +-- Unio_tumidus **
--+
| +----- Anodonta_anatina
+--+
| +-- Utterbackia_peninsularis
+--+
+-- Pyganodon_grandis
Lopes-Lima et al., 2020 [Publ 4080], fig. 11
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 2 |
2 |
0 |
1 |
10 |
8 |
1 |
-- |
+-- Nodularia_breviconcha **
+-----------------+
| +-- Nodularia_douglasiae **
--+
| +----------------- Lanceolaria_oxyrhyncha
| |
+--+ +-------------- Lanceolaria_gladiola
| |
+--+ +-- Lanceolaria_grayii
| +--------+
| | +-- Lanceolaria_bilirata
+--+
| +-------- Lanceolaria_acrorrhyncha
+--+
| +----- Lanceolaria_kihirai
+--+
| +-- Lanceolaria_lanceolata
+--+
+-- Lanceolaria_triformis
Riccardi et al., 2019 [Publ 4089], fig. 2
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 2 |
2 |
0 |
1 |
18 |
16 |
2 |
-- |
+-- Lanceolaria_grayii
+--+
| +-- Lanceolaria_lanceolata
+-----------------+
| | +-- Unio_mancus **
| +--+
| +-- Unio_elongatulus **
|
| +-------- Cristaria_plicata
--+ |
| +-----------+ +-- Anemina_euscaphys
| | | +--+
| | | | +-- Anemina_arcaeformis
| | +--+
| | | +-- Sinanodonta_lucida
+--+ +--+
| +-- Sinanodonta_woodiana
|
| +----------------- Anodonta_anatina
| |
| | +----- Pseudanodonta_complanata
+--+ +--------+
| | | +-- Anodonta_exulcerata
| | +--+
+--+ +-- Anodonta_cygnea
|
| +----------- Anodonta_nuttalliana
+--+
| +-------- Platynaias_compressa
+--+
| +----- Pyganodon_grandis
+--+
| +-- Utterbackia_peninsularis
+--+
+-- Utterbackia_imbecillis
Tomilova et al., 2020 [Publ 4097], fig. 3
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 2 |
2 |
0 |
1 |
6 |
6 |
1 |
-- |
+-- Unio_tumidus **
+--------+
| +-- Unio_pictorum **
|
| +-- Sinanodonta_lauta
--+--------+
| +-- Sinanodonta_woodiana
|
| +-------- Anodonta_anatina
+--+
| +----- Pseudanodonta_complanata
+--+
| +-- Anodonta_cygnea
+--+
+-- Anodonta_exulcerata
Mabuchi & Nishida, 2020 [Publ 4120], fig. 1
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 2 |
3 |
0 |
2 |
35 |
31 |
2 |
16.0 |
+----- Nodularia_douglasiae **
+-----------+
| | +-- Lanceolaria_grayii
| +--+
| +-- Inversiunio_reinianus **
|
--+ +-------------- Cristaria_plicata
| |
| | +----------- Sinanodonta_tumens
| | |
| | | +-- Sinanodonta_lauta[5]
| | | |
+--+ | +-- Sinanodonta_lauta[6]
| | |
| | +-- Beringiana_japonica[15]
| | |
| | +-- Beringiana_japonica[16]
| | +-----+
| | | +-- Beringiana_japonica[17]
+--+ | |
| | +-- Beringiana_japonica[18]
| | |
| | +-- Beringiana_japonica[19]
| | |
| | +-- Beringiana_japonica[20]
| |
| | +----- Beringiana_japonica[1]
| | |
| | +----- Beringiana_japonica[2]
| | |
+--+ +----- Beringiana_japonica[3]
| |
| +----- Beringiana_japonica[4]
| |
| +----- Beringiana_japonica[5]
| |
| +----- Sinanodonta_lauta[1]
| |
| +----- Beringiana_japonica[6]
| |
| +----- Sinanodonta_lauta[2]
| |
| +----- Sinanodonta_calipygos[1]
| |
+--+----- Beringiana_japonica[7]
|
+----- Sinanodonta_lauta[4]
|
+----- Beringiana_japonica[8]
|
+----- Beringiana_japonica[9]
|
+----- Beringiana_japonica[10]
|
+----- Beringiana_japonica[11]
|
+----- Beringiana_japonica[12]
|
+----- Beringiana_japonica[13]
|
+----- Beringiana_japonica[14]
|
| +-- Sinanodonta_lauta[3]
+--+
+-- Sinanodonta_calipygos[2]
Kano et al., 2019 [Publ 4194], fig. 6
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 2 |
2 |
0 |
2 |
8 |
4 |
2 |
-- |
+----- Lanceolaria_grayii
--+
| +-- Inversiunio_yanagawensis **
+--+
+-- Nodularia_nipponensis **
Nagel et al., 1998 [Publ 1946], fig. 9
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 1 |
1 |
0 |
1 |
5 |
2 |
1 |
-- |
+----- Unio **
--+
| +-- Anodonta
+--+
+-- Pseudanodonta
Araujo et al., 2009 [Publ 2395], fig. 3
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 1 |
1 |
0 |
1 |
7 |
0 |
0 |
-- |
-- Unio
Graf, 2000 [Publ 2647], fig. 1
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 1 |
1 |
0 |
1 |
17 |
1 |
1 |
-- |
-- Unio
Graf & Ó Foighil, 2000 [Publ 2649], fig. 1, 2
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 1 |
1 |
0 |
1 |
11 |
1 |
1 |
-- |
+-- Unio_pictorum **
--+
+-- Pyganodon_grandis
Chong et al., 2008 [Publ 2668], fig. 3
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 1 |
1 |
0 |
1 |
20 |
20 |
1 |
-- |
+----------------- Anodonta_cygnea
|
+----------------- Pyganodon_fragilis
|
+----------------- Alasmidonta_holstonia
|
+----------------- Anodonta_anatina
|
+----------------- Unio_tumidus **
|
| +-- Anodonta_nuttalliana
+--------------+
| +-- Anodonta_californiensis
|
--+ +-- Beringiana_beringiana
+--------------+
| +-- Sinanodonta_woodiana
|
| +-- Pseudodontoideus_subvexus
| |
+--------------+-- Alasmidonta_marginata
| |
| +-- Lasmigona_complanata
|
| +-------------- Pseudanodonta_complanata
| |
| | +----- Strophitus_undulatus
| +--------+
+--+ | +-- Prolasmidonta_heterodon
| +--+
| +-- Platynaias_compressa
|
| +----------- Anodonta_oregonensis[4]
+--+
| +-------- Anodonta_oregonensis[3]
+--+
| +----- Anodonta_oregonensis[2]
+--+
| +-- Anodonta_oregonensis[1]
+--+
+-- Anodonta_kennerlyi
Taylor et al., 2007 [Publ 2696], fig. 1-3
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 1 |
1 |
0 |
1 |
3 |
1 |
1 |
-- |
+-- Anodonta_cygnea
--+
+-- Unio_pictorum **
Huff et al., 2004 [Publ 2697], fig. 2
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 1 |
1 |
0 |
1 |
13 |
1 |
1 |
-- |
+-- Unio_ravoisieri **
--+
+-- Anodonta
Huff et al., 2004 [Publ 2697], fig. 4
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 1 |
1 |
0 |
1 |
10 |
1 |
1 |
-- |
+-- Unio_ravoisieri **
--+
+-- Anodonta
Huff et al., 2004 [Publ 2697], fig. 5-6
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 1 |
1 |
0 |
1 |
11 |
1 |
1 |
-- |
-- Unio_ravoisieri
Rosenberg et al., 1994 [Publ 2719], fig. 3
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 1 |
1 |
0 |
1 |
18 |
3 |
1 |
-- |
-- Unio_pictorum
Rosenberg et al., 1997 [Publ 2720], fig. 2A
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 1 |
1 |
0 |
1 |
22 |
4 |
1 |
-- |
-- Unio_pictorum
Graf et al., 2014 [Publ 3591], fig. 3
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 1 |
1 |
0 |
1 |
26 |
0 |
0 |
-- |
-- Unio_pictorum
Marshall et al., 2014 [Publ 3593], fig. 1
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 1 |
1 |
0 |
1 |
13 |
1 |
1 |
-- |
+-- Pyganodon_grandis
--+
+-- Unio_pictorum **
Graf et al., 2015 [Publ 3619], fig. 1
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 1 |
1 |
0 |
1 |
19 |
0 |
0 |
-- |
-- Unio_pictorum
Pfeiffer & Graf, 2015 [Publ 3620], fig. 2
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 1 |
1 |
0 |
1 |
39 |
2 |
1 |
-- |
+----- Unio_pictorum **
--+
| +-- Pyganodon_grandis
+--+
+-- Lasmigona_costata
Bieler et al., 2014 [Publ 3622], fig. 26, 27, 29, 30
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 1 |
1 |
0 |
1 |
4 |
0 |
0 |
-- |
-- Unio_pictorum
Bieler et al., 2014 [Publ 3622], fig. 28
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 1 |
1 |
0 |
1 |
4 |
0 |
0 |
-- |
-- Unio_pictorum
Huang et al., 2013 [Publ 3629], fig. 3
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 1 |
1 |
0 |
1 |
15 |
6 |
1 |
-- |
+-------------- Unio_pictorum **
|
--+ +-- Sinanodonta_woodiana
| +--------+
| | +-- Cristaria_plicata
+--+
| +-------- Platynaias_compressa
+--+
| +----- Pyganodon_grandis
+--+
| +-- Utterbackia_imbecillis
+--+
+-- Utterbackia_peninsularis
González & Giribet, 2015 [Publ 3632], fig. 2
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 1 |
1 |
0 |
1 |
5 |
0 |
0 |
-- |
-- Unio_pictorum
González & Giribet, 2015 [Publ 3632], fig. 3
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 1 |
1 |
0 |
1 |
5 |
0 |
0 |
-- |
-- Unio_pictorum
González & Giribet, 2015 [Publ 3632], fig. 4
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 1 |
1 |
0 |
1 |
5 |
0 |
0 |
-- |
-- Unio_pictorum
González & Giribet, 2015 [Publ 3632], fig. 6
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 1 |
1 |
0 |
1 |
5 |
0 |
0 |
-- |
-- Unio_pictorum
Bolotov et al., 2015 [Publ 3646], fig. 2A
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 1 |
1 |
0 |
1 |
9 |
0 |
0 |
-- |
-- Unio_pictorum
Bolotov et al., 2015 [Publ 3646], fig. 2B
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 1 |
1 |
0 |
1 |
8 |
0 |
0 |
-- |
-- Unio_pictorum
Giribet & Distel, 2003 [Publ 3670], fig. 3.5
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 1 |
1 |
0 |
1 |
5 |
0 |
0 |
-- |
-- Unio_ravoisieri
Wang et al., 2016 [Publ 3714], fig. 2
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 1 |
1 |
0 |
1 |
2 |
0 |
0 |
-- |
-- Unio_tumidus
Fonseca et al., 2016 [Publ 3775], fig. 1-M
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 1 |
1 |
0 |
1 |
8 |
3 |
1 |
-- |
+-------- Unio_delphinus **
--+
| +----- Anodonta_anatina
+--+
| +-- Utterbackia_peninsularis
+--+
+-- Pyganodon_grandis
Lopes-Lima et al., 2017 [Publ 3792], fig. 2-M
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 1 |
1 |
0 |
1 |
9 |
3 |
1 |
-- |
+-------- Unio_delphinus **
--+
| +----- Anodonta_anatina
+--+
| +-- Utterbackia_peninsularis
+--+
+-- Pyganodon_grandis
Feng et al., 2016 [Publ 3800], fig. 2
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 1 |
1 |
0 |
1 |
14 |
7 |
1 |
-- |
+----------- Tchangsinaia_piscicula **
|
--+ +-------- Pyganodon_grandis
| |
+--+ +-- Anemina_arcaeformis
| +--+
| | +-- Anemina_euscaphys
+--+
| +-- Sinanodonta_lucida
+--+
+-- Sinanodonta_woodiana
Combosch et al., 2017 [Publ 3806], fig. 1
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 1 |
1 |
0 |
1 |
14 |
0 |
0 |
-- |
-- Unio_pictorum
Combosch et al., 2017 [Publ 3806], fig. 2
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 1 |
1 |
0 |
1 |
14 |
0 |
0 |
-- |
-- Unio_pictorum
Vikhrev et al., 2018 [Publ 3812], fig. 3
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 1 |
1 |
0 |
1 |
12 |
0 |
0 |
-- |
-- Unio_ravoisieri
Lopes-Lima et al., 2018 [Publ 3829], fig. 1
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 1 |
1 |
0 |
1 |
27 |
0 |
0 |
-- |
-- Unio_pictorum
Lopes-Lima et al., 2018 [Publ 3829], fig. 3
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 1 |
1 |
0 |
1 |
27 |
0 |
0 |
-- |
-- Unio_pictorum
Gomes-dos-Santos et al., 2019 [Publ 3943], fig. 1
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 1 |
1 |
0 |
1 |
9 |
1 |
1 |
-- |
+-- Unio_delphinus **
--+
+-- Anodonta_anatina
Gomes-dos-Santos et al., 2019 [Publ 3943], fig. 1-M
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 1 |
1 |
0 |
1 |
5 |
1 |
1 |
-- |
+-- Unio_delphinus **
--+
+-- Anodonta_anatina
Klunzinger et al., 2021 [Publ 4086], fig. 2
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 1 |
1 |
0 |
1 |
5 |
0 |
0 |
-- |
-- Unio_pictorum
Bogan et al., 2023 [Publ 4295], fig. 3
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 1 |
1 |
0 |
1 |
30 |
28 |
1 |
-- |
+----------------------------------- Schistodesmus_lampreyanus **
|
| +-------------------------------- Anodonta_cygnea
--+ |
| | +-------- Cristaria_bellua
| | +--------------------+
| | | | +----- Cristaria_clessini
+--+ | +--+
| | | +-- Cristaria_plicata
| | +--+
| | +-- Cristaria_truncata
| |
| | +-------------- Amuranodonta_kijaensis
+--+ |
| +-----------+ +----------- Anemina_arcaeformis
| | | |
| | +--+ +-------- Buldowskia_suifunica
| | | |
| | +--+ +-- Buldowskia_flavotincta
| | | +--+
| | | | +-- Buldowskia_shadini
| | +--+
+--+ | +-- Buldowskia_iwakawai
| +--+
| +-- Buldowskia_kamiyai
|
| +-- Pletholophus_honglinhensis
| +--------------------+
| | +-- Pletholophus_tenuis
| |
| | +-------- Beringiana_japonica
+--+ +-----------+
| | | +----- Beringiana_beringiana
| | +--+
| | | +-- Beringiana_fukuharai
| | +--+
+--+ +-- Beringiana_gosannensis
|
| +-- Sinanodonta_angula
| +--------------+
| | +-- Sinanodonta_lucida
| |
+--+ +-------------- Sinanodonta_schrenkii
| |
| | +-- Sinanodonta_calipygos
+--+-----------+
| +-- Sinanodonta_tumens
|
| +----------- Sinanodonta_woodiana[2]
+--+
| +-------- Sinanodonta_jourdyi
+--+
| +----- Sinanodonta_elliptica
+--+
| +-- Sinanodonta_lauta
+--+
+-- Sinanodonta_woodiana[1]
| 
