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Page last updated
Thu 04 Apr 2024

The Freshwater Mussels (Unionoida) of the World (and other less consequential bivalves)

FM(U)otW(aolcb) is the web version of the MUSSEL Project Database. Follow the links to browse the data or use the custom Google search field. Either way, you win!

Cladomics pages are experimental — even more so than the rest of this site!

The purpose of the cladomics pages for each genus and family-group level taxon is to provide statistics and references indicating the extent to which published phylogenetic work supports the classification used here.

The Cladomics Summary provides an overview of the number of studies that have included terminal species representing this taxon as well as an indication of how frequently it has been recovered as monophyletic.

Following the summary, the available published clades representing the taxon are listed.

For more information, visit the Cladomics FARQ (frequently asked rhetorical questions) page. Abbreviations for the various statistics are explained on the FARQ!

Search the database pages:

Unionoida | Unionoidea | Unionidae | Unioninae | Anodontini | Alasmidontina

genus Strophitus Rafinesque, 1820: cladomics
     8 valid species

Cladomics Summary

number of species = 8

analyzed in 49 cladograms in 33 publications
maximum number of terminals in an analysis = 6, mean = 1.43

Basic Monophyly

number of cladograms that test basic monophyly (i.e., more than 1 included species) = 6
maximum number of included terminals = 6, mean = 4.5
proportion of cladograms that support basic monophyly = 1.0


Pairwise Cladogram Clustering

The graphs below depict in two dimensions all pair-wise ingroup distances within the cladome. See the Cladmoics FARQ for more information.


PCoA Ordination


NJ tree


Smith et al., 2018 [Publ 3790], fig. 1

iT tT bi riS oT roT roS iD
6 6 4 6 13 12 7 0.0

*** Top Monophyly Cladogram ***

              +-- Anodontoides_ferussacianus      
     +--------+                                   
     |        +-- Anodontoides_denigratus         
     |                                            
  +--+     +----- Strophitus_pascagoulaensis **      
  |  |  +--+                                      
  |  |  |  |  +-- Strophitus_radiatus **             
  |  |  |  +--+                                   
  |  +--+     +-- Strophitus_williamsi **            
  |     |                                         
  |     |  +----- Strophitus_howellsi[2] **          
  |     +--+                                      
--+        |  +-- Strophitus_undulatus **            
  |        +--+                                   
  |           +-- Strophitus_howellsi[1] **          
  |                                               
  |           +-- Alasmidonta_triangulata         
  |  +--------+                                   
  |  |        +-- Alasminota_etowaensis           
  |  |                                            
  +--+        +-- Pseudodontoideus_connasaugaensis
     |  +-----+                                   
     |  |     +-- Pseudodontoideus_subvexus       
     |  |                                         
     +--+     +-- Utterbackiana_couperiana        
        |  +--+                                   
        |  |  +-- Utterbackiana_heardi            
        +--+                                      
           |  +-- Utterbackiana_hartfieldorum     
           +--+                                   
              +-- Utterbackiana_suborbiculata     

Whelan et al., 2023 [Publ 4287], fig. 2

iT tT bi riS oT roT roS iD
5 5 3 5 39 31 12 5.0

*** Top Sister-Group Cladogram ***

  +----------------- Pressodonta_viridis          
  |                                               
--+              +-- Anodontoides_denigratus      
  |  +-----------+                                
  |  |           +-- Anodontoides_ferussacianus   
  +--+                                            
     |  +----------- Strophitus_undulatus **         
     +--+                                         
        |  +-------- Strophitus_pascagoulaensis[2] **
        +--+                                      
           |  +----- Strophitus_radiatus **          
           +--+                                   
              |  +-- Strophitus_pascagoulaensis[1] **
              +--+                                
                 +-- Strophitus_williamsi **         

Whelan et al., 2023 [Publ 4287], fig. 3

iT tT bi riS oT roT roS iD
5 5 3 5 24 23 11 3.0
              +----- Pressodonta_viridis          
  +-----------+                                   
  |           |  +-- Platynaias_subviridis        
  |           +--+                                
  |              +-- Prolasmidonta_heterodon      
--+                                               
  |              +-- Anodontoides_ferussacianus   
  |  +-----------+                                
  |  |           +-- Anodontoides_denigratus      
  +--+                                            
     |  +----------- Strophitus_undulatus **         
     +--+                                         
        |  +-------- Strophitus_pascagoulaensis[2] **
        +--+                                      
           |  +----- Strophitus_pascagoulaensis[1] **
           +--+                                   
              |  +-- Strophitus_radiatus **          
              +--+                                
                 +-- Strophitus_williamsi **         

Smith et al., 2023 [Publ 4291], fig. 2

iT tT bi riS oT roT roS iD
5 5 3 5 2 2 1 1.0
           +-- Anodontoides_ferussacianus
  +--------+                             
  |        +-- Anodontoides_denigratus   
  |                                      
--+        +-- Strophitus_undulatus **      
  |  +-----+                             
  |  |     +-- Strophitus_howellsi **       
  +--+                                   
     |  +----- Strophitus_pascagoulaensis **
     +--+                                
        |  +-- Strophitus_radiatus **       
        +--+                             
           +-- Strophitus_williamsi **      

Whelan et al., 2023 [Publ 4287], fig. 4

iT tT bi riS oT roT roS iD
4 4 2 4 25 23 10 4.0
  +-------------- Platynaias_subviridis     
  |                                         
  |     +-------- Strophitus_pascagoulaensis **
  |  +--+                                   
--+  |  |  +----- Strophitus_undulatus **      
  |  |  +--+                                
  |  |     |  +-- Strophitus_radiatus **       
  |  |     +--+                             
  +--+        +-- Strophitus_williamsi **      
     |                                      
     |        +-- Pyganodon_grandis         
     |  +-----+                             
     |  |     +-- Pyganodon_cataracta       
     +--+                                   
        |  +----- Utterbackia_peninsularis  
        +--+                                
           |  +-- Anodontoides_ferussacianus
           +--+                             
              +-- Anodontoides_denigratus   

Krebs et al., 2003 [Publ 2691], fig. 3

iT tT bi riS oT roT roS iD
2 2 0 2 32 7 6 --
  +-- Strophitus_radiatus ** 
--+                       
  +-- Strophitus_undulatus **

Graf & Cummings, 2006 [Publ 1968], fig. 2

iT tT bi riS oT roT roS iD
1 1 0 1 46 3 3 --
  +----- Alasmidonta_marginata
--+                           
  |  +-- Pyganodon_grandis    
  +--+                        
     +-- Strophitus_undulatus ** 

Graf & Ó Foighil, 2000 [Publ 2648], fig. 2A

iT tT bi riS oT roT roS iD
1 1 0 1 29 4 3 --
        +-- Alasmidonta_marginata
  +-----+                        
  |     +-- Lasmigona_costata    
--+                              
  |  +----- Strophitus_undulatus ** 
  +--+                           
     |  +-- Pyganodon_grandis    
     +--+                        
        +-- Pyganodon_fragilis   

Graf & Ó Foighil, 2000 [Publ 2648], fig. 2B

iT tT bi riS oT roT roS iD
1 1 0 1 29 4 3 --
        +-- Alasmidonta_marginata
  +-----+                        
  |     +-- Lasmigona_costata    
--+                              
  |  +----- Strophitus_undulatus ** 
  +--+                           
     |  +-- Pyganodon_grandis    
     +--+                        
        +-- Pyganodon_fragilis   

Bogan & Hoeh, 2000 [Publ 2650], fig. 1

iT tT bi riS oT roT roS iD
1 1 0 1 33 1 1 --
  +-- Strophitus_undulatus **
--+                       
  +-- Pyganodon_grandis   

Hoeh et al., 2001 [Publ 2652], fig. 14.2

iT tT bi riS oT roT roS iD
1 1 0 1 30 1 1 --
-- Strophitus_undulatus

Hoeh et al., 2001 [Publ 2652], fig. 14.4

iT tT bi riS oT roT roS iD
1 1 0 1 30 1 1 --
  +-- Strophitus_undulatus **
--+                       
  +-- Pyganodon_grandis   

Hoeh et al., 2002 [Publ 2653], fig. 2

iT tT bi riS oT roT roS iD
1 1 0 1 30 1 1 --
  +-- Pyganodon_grandis   
--+                       
  +-- Strophitus_undulatus **

Roe & Hoeh, 2003 [Publ 2655], fig. 4.2

iT tT bi riS oT roT roS iD
1 1 0 1 30 1 1 --
-- Strophitus_undulatus

Roe & Hoeh, 2003 [Publ 2655], fig. 4.3

iT tT bi riS oT roT roS iD
1 1 0 1 30 1 1 --
  +-- Pyganodon_grandis   
--+                       
  +-- Strophitus_undulatus **

Lydeard et al., 1996 [Publ 2656], fig. 3

iT tT bi riS oT roT roS iD
1 1 0 1 26 5 4 --
     +----- Utterbackia_imbecillis  
  +--+                              
  |  |  +-- Utterbackiana_couperiana
  |  +--+                           
--+     +-- Utterbackiana           
  |                                 
  |  +----- Strophitus_radiatus **     
  +--+                              
     |  +-- Lasmigona_costata       
     +--+                           
        +-- Alasmidonta_triangulata 

Lydeard et al., 1996 [Publ 2656], fig. 4

iT tT bi riS oT roT roS iD
1 1 0 1 27 4 4 --
  +----- Strophitus_radiatus **    
--+                             
  |  +-- Lasmigona_costata      
  +--+                          
     +-- Alasmidonta_triangulata

Huang et al., 2002 [Publ 2657], fig. 1

iT tT bi riS oT roT roS iD
1 1 0 1 33 4 4 --
  +-------- Utterbackia_imbecillis 
--+                                
  |  +----- Strophitus_radiatus **    
  +--+                             
     |  +-- Alasmidonta_triangulata
     +--+                          
        +-- Lasmigona_costata      

Huang et al., 2002 [Publ 2657], fig. 2

iT tT bi riS oT roT roS iD
1 1 0 1 33 4 4 --
-- Strophitus_radiatus

Chong et al., 2008 [Publ 2668], fig. 3

iT tT bi riS oT roT roS iD
1 1 0 1 20 7 6 --
  +----- Strophitus_undulatus **   
--+                             
  |  +-- Prolasmidonta_heterodon
  +--+                          
     +-- Platynaias_compressa   

King et al., 1999 [Publ 2699], fig. 2

iT tT bi riS oT roT roS iD
1 1 0 1 8 8 4 --
        +----- Lasmigona_costata      
  +-----+                             
  |     |  +-- Lasmigona_complanata   
  |     +--+                          
--+        +-- Lasmigona_alabamensis  
  |                                   
  |  +-------- Strophitus_undulatus **   
  +--+                                
     |  +----- Prolasmidonta_heterodon
     +--+                             
        |  +-- Platynaias_subviridis  
        +--+                          
           +-- Platynaias_compressa   

King et al., 1999 [Publ 2699], fig. 4

iT tT bi riS oT roT roS iD
1 1 0 1 8 8 3 --
        +-- Lasmigona_complanata    
  +-----+                           
  |     +-- Lasmigona_costata       
  |                                 
--+  +----- Prolasmidonta_heterodon 
  |  |                              
  |  +----- Strophitus_undulatus **    
  |  |                              
  +--+  +-- Platynaias_compressa    
     |  |                           
     |  +-- Platynaias_subviridis[1]
     |  |                           
     +--+-- Platynaias_subviridis[2]
        |                           
        +-- Platynaias_subviridis[3]
        |                           
        +-- Platynaias_subviridis[4]

Hoeh et al., 2009 [Publ 2716], fig. 3

iT tT bi riS oT roT roS iD
1 1 0 1 46 3 3 --
  +----- Alasmidonta_marginata
--+                           
  |  +-- Strophitus_undulatus ** 
  +--+                        
     +-- Pyganodon_grandis    

Hoeh et al., 2009 [Publ 2716], fig. 4

iT tT bi riS oT roT roS iD
1 1 0 1 45 3 3 --
  +----- Alasmidonta_marginata
--+                           
  |  +-- Strophitus_undulatus ** 
  +--+                        
     +-- Pyganodon_grandis    

Hoeh et al., 2009 [Publ 2716], fig. 5

iT tT bi riS oT roT roS iD
1 1 0 1 45 3 3 --
  +----- Alasmidonta_marginata
--+                           
  |  +-- Strophitus_undulatus ** 
  +--+                        
     +-- Pyganodon_grandis    

Boyer et al., 2011 [Publ 2718], fig. 3

iT tT bi riS oT roT roS iD
1 1 0 1 34 5 4 --
  +----- Alasmidonta_marginata
  |                           
  +----- Pyganodon_grandis    
  |                           
--+  +-- Lasmigona_complanata 
  +--+                        
  |  +-- Lasmigona_costata    
  |                           
  |  +-- Platynaias_compressa 
  +--+                        
     +-- Strophitus_undulatus ** 

Boyer et al., 2011 [Publ 2718], fig. 4

iT tT bi riS oT roT roS iD
1 1 0 1 37 8 7 --
  +-------------- Strophitus_undulatus **      
  |                                         
  |        +----- Lasmigona_complanata      
--+  +-----+                                
  |  |     |  +-- Lasmigona_costata         
  |  |     +--+                             
  +--+        +-- Arcidens_confragosus      
     |                                      
     |  +-------- Utterbackia_imbecillis    
     |  |                                   
     +--+-------- Alasmidonta_marginata     
        |                                   
        |  +----- Pyganodon_grandis         
        +--+                                
           |  +-- Anodontoides_ferussacianus
           +--+                             
              +-- Platynaias_compressa      

Breton et al., 2011 [Publ 3376], fig. 3

iT tT bi riS oT roT roS iD
1 1 0 1 38 11 5 --
                 +-- Lasmigona_complanata       
  +--------------+                              
  |              +-- Lasmigona_costata          
  |                                             
--+  +-------------- Strophitus_undulatus **       
  |  |                                          
  |  |           +-- Platynaias_compressa       
  |  |        +--+                              
  +--+        |  +-- Platynaias_subviridis      
     |  +-----+                                 
     |  |     |  +-- Alasmidonta_marginata      
     |  |     +--+                              
     |  |        +-- Alasmidonta_undulata       
     +--+                                       
        |        +-- Pyganodon_lacustris        
        |  +-----+                              
        |  |     +-- Pyganodon_grandis          
        +--+                                    
           |  +----- Utterbackia_peninsularis[2]
           +--+                                 
              |  +-- Utterbackia_peninsularis[1]
              +--+                              
                 +-- Utterbackia_imbecillis     

Pfeiffer & Graf, 2013 [Publ 3531], fig. 2

iT tT bi riS oT roT roS iD
1 1 0 1 34 2 2 --
  +----- Strophitus_undulatus **
--+                          
  |  +-- Platynaias_compressa
  +--+                       
     +-- Pyganodon_grandis   

Inoue et al., 2014 [Publ 3635], fig. 3A

iT tT bi riS oT roT roS iD
1 1 0 1 20 14 7 --
        +----------- Pseudodontoideus_subvexus  
  +-----+                                       
  |     |  +-------- Lasmigona_costata          
  |     +--+                                    
  |        |  +----- Lasmigona_complanata       
  |        +--+                                 
  |           |  +-- Arcidens_confragosus       
--+           +--+                              
  |              +-- Arcidens_wheeleri          
  |                                             
  |  +-------------- Strophitus_undulatus **       
  |  |                                          
  |  |     +-------- Pyganodon_grandis          
  |  |  +--+                                    
  +--+  |  |  +----- Utterbackia_peninsularis[2]
     |  |  +--+                                 
     |  |     |  +-- Utterbackia_imbecillis     
     |  |     +--+                              
     +--+        +-- Utterbackia_peninsularis[1]
        |                                       
        |        +-- Platynaias_compressa       
        |  +-----+                              
        |  |     +-- Platynaias_subviridis      
        +--+                                    
           |  +----- Alasmidonta_marginata      
           +--+                                 
              |  +-- Alasmidonta_undulata       
              +--+                              
                 +-- Alasmidonta_varicosa       

Jadhav & Jamkhedkar, 2009 [Publ 3697], fig. 11

iT tT bi riS oT roT roS iD
1 1 0 1 25 3 3 --
  +----- Lasmigona_costata    
  |                           
--+  +-- Strophitus_undulatus ** 
  |  |                        
  +--+-- Alasmidonta_marginata
     |                        
     +-- Pyganodon_grandis    

Lopes-Lima et al., 2017 [Publ 3745], fig. 1-2

iT tT bi riS oT roT roS iD
1 1 0 1 80 4 4 --
        +-- Pyganodon_grandis    
  +-----+                        
  |     +-- Simpsonaias_ambigua  
--+                              
  |  +----- Strophitus_undulatus ** 
  +--+                           
     |  +-- Alasmidonta_marginata
     +--+                        
        +-- Lasmigona_costata    

Hewitt et al., 2018 [Publ 3758], fig. 5

iT tT bi riS oT roT roS iD
1 1 0 1 8 5 4 --
  +-------- Pyganodon_grandis    
  |                              
  |     +-- Platynaias_subviridis
--+-----+                        
  |     +-- Strophitus_undulatus ** 
  |                              
  |  +----- Lasmigona_costata    
  +--+                           
     |  +-- Arcidens_wheeleri    
     +--+                        
        +-- Lasmigona_complanata 

Bolotov et al., 2017 [Publ 3761], fig. S1

iT tT bi riS oT roT roS iD
1 1 0 1 140 4 4 --
  +----- Simpsonaias_ambigua 
--+                          
  |  +-- Pyganodon_grandis   
  +--+                       
     +-- Strophitus_undulatus **

Bolotov et al., 2017 [Publ 3761], fig. S2

iT tT bi riS oT roT roS iD
1 1 0 1 140 4 4 --
  +----- Pyganodon_grandis    
  |                           
  +----- Simpsonaias_ambigua  
--+                           
  +----- Strophitus_undulatus ** 
  |                           
  |  +-- Alasmidonta_marginata
  +--+                        
     +-- Lasmigona_costata    

Bolotov et al., 2017 [Publ 3764], fig. S1

iT tT bi riS oT roT roS iD
1 1 0 1 130 4 4 --
-- Strophitus_undulatus

Bolotov et al., 2018 [Publ 3826], fig. 2

iT tT bi riS oT roT roS iD
1 1 0 1 84 4 4 --
  +-------- Simpsonaias_ambigua  
--+                              
  |  +----- Strophitus_undulatus ** 
  +--+                           
     |  +-- Alasmidonta_marginata
     +--+                        
        +-- Lasmigona_costata    

Wu et al., 2018 [Publ 3836], fig. 2

iT tT bi riS oT roT roS iD
1 1 0 1 52 4 3 --
  +-------- Strophitus_undulatus ** 
  |                              
--+  +----- Platynaias_compressa 
  |  |                           
  +--+----- Alasmidonta_marginata
     |                           
     |  +-- Pyganodon_cataracta  
     +--+                        
        +-- Pyganodon_grandis    

Huang et al., 2019 [Publ 3843], fig. 2

iT tT bi riS oT roT roS iD
1 1 0 1 203 6 5 --
-- Strophitus_undulatus

Huang et al., 2019 [Publ 3843], fig. S3

iT tT bi riS oT roT roS iD
1 1 0 1 200 6 5 --
  +-- Simpsonaias_ambigua 
--+                       
  +-- Strophitus_undulatus **

Huang et al., 2019 [Publ 3843], fig. S4

iT tT bi riS oT roT roS iD
1 1 0 1 201 6 5 --
-- Strophitus_undulatus

Chase et al., 2018 [Publ 3847], fig. 4

iT tT bi riS oT roT roS iD
1 1 0 1 38 11 5 --
        +-- Alasmidonta_undulata 
  +-----+                        
  |     +-- Alasmidonta_marginata
--+                              
  |  +----- Strophitus_undulatus ** 
  +--+                           
     |  +-- Platynaias_compressa 
     +--+                        
        +-- Platynaias_subviridis

Inoue et al., 2018 [Publ 3858], fig. 1, 3a-c

iT tT bi riS oT roT roS iD
1 1 0 1 204 16 8 --
-- Strophitus_undulatus

Lopes-Lima et al., 2020 [Publ 4080], fig. 2-4

iT tT bi riS oT roT roS iD
1 1 0 1 121 4 4 --
  +----- Pyganodon_grandis    
  |                           
  +----- Simpsonaias_ambigua  
--+                           
  +----- Strophitus_undulatus ** 
  |                           
  |  +-- Alasmidonta_marginata
  +--+                        
     +-- Lasmigona_costata    

Jeratthitikul et al., 2022 [Publ 4260], fig. S1, 1

iT tT bi riS oT roT roS iD
1 1 0 1 75 4 4 --
        +-- Pyganodon_grandis    
  +-----+                        
  |     +-- Simpsonaias_ambigua  
--+                              
  |  +----- Strophitus_undulatus ** 
  +--+                           
     |  +-- Alasmidonta_marginata
     +--+                        
        +-- Lasmigona_costata    

Jeratthitikul et al., 2022 [Publ 4260], fig. S2, 2

iT tT bi riS oT roT roS iD
1 1 0 1 75 4 4 --
  +----- Pyganodon_grandis   
--+                          
  |  +-- Strophitus_undulatus **
  +--+                       
     +-- Simpsonaias_ambigua 

Wu et al., 2023 [Publ 4296], fig. 4

iT tT bi riS oT roT roS iD
1 1 0 1 22 2 2 --
  +----- Lasmigona_costata   
--+                          
  |  +-- Strophitus_undulatus **
  +--+                       
     +-- Pyganodon_grandis   

Liu et al., 2023 [Publ 4310], fig. 4

iT tT bi riS oT roT roS iD
1 1 0 1 23 2 2 --
  +----- Lasmigona_costata   
--+                          
  |  +-- Strophitus_undulatus **
  +--+                       
     +-- Pyganodon_grandis   

Liu et al., 2023 [Publ 4310], fig. 5

iT tT bi riS oT roT roS iD
1 1 0 1 23 2 2 --
  +-- Pyganodon_grandis   
--+                       
  +-- Strophitus_undulatus **

Page updated: 07:04:17 Thu 04 Apr 2024

 
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