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Page last updated
Wed 04 Dec 2024

The Freshwater Mussels (Unionoida) of the World (and other less consequential bivalves)

FM(U)otW(aolcb) is the web version of the MUSSEL Project Database. Follow the links to browse the data or use the custom Google search field. Either way, you win!

This page documents data for Whelan et al. (2023), such as genera and species attributed to that publication (including known misspellings, misidentifications, and other unavailable names) as well as taxonomic opinions, if available. Taxonomic opinions refer to uses of genera and species as valid and invalid in that work.

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Publication information

Whelan, N.V., N.A. Johnson, A.S. Williams, M.A. Perkins, C.E. Beaver & J.W. Mays. 2023. Hidden in the hills: phylogeny of the freshwater mussel genus Alasmidonta (Bivalvia: Unionidae) and description of a new species. Zoological Journal of the Linnean Society 198(2): 650-676.

Click here for online access.

Cladogram Figures

(click the figure number to see the cladogram)

fig. 2
fig. 3
fig. 4

Attributed Genus Names

Simpsonais Whelan et al., 2023
Whelan et al., 2023, Zool. J. Linn. Soc. 198(2): Table S1.
(error for Simpsonaias Frierson, 1914)

Attributed Species Names

Margaritana connasaugensis Clessin, 1876
‘Lea’ of Authors.
Clessin, 1876, Conchyl. Cab. 9(1): 269, pl. 81, figs. 7-8.
Whelan et al., 2023, Zool. J. Linn. Soc. 198(2): Figs. 2-3.
(error for Margaritana connasaugaensis Lea, 1858)

Anodonta arcaeformis Soroka, 2010
‘Heude’ of Authors.
Soroka, 2010, Folia Malac. 18(4): 178.
Zhang et al., 2013, World Sci. Tech.-Mod. Traditional Chinese Med. 15(3): 376.
Sayenko et al., 2017, Biol. Bulletin 44(3): 367 [as ‘Anemina arcaeformis’].
Whelan et al., 2023, Zool. J. Linn. Soc. 198(2): Table S1. [in part]
(misidentified reference to Anodon arcaeformis Heude, 1877)

Strophitus undulatus Smith et al., 2018
‘Say’ of Authors.
Smith et al., 2018, MPE 119: 51. [in part]
Gasparini et al., 2020, Environmental DNA 2(3): Table S1.
Whelan et al., 2023, Zool. J. Linn. Soc. 198(2): Table S1. [in part]
(misidentified reference to Anodonta undulata Say, 1817)

Anemina euscaphys Huang et al., 2019
‘Heude’ of Authors.
Huang et al., 2019, MPE 130: Table S1. [in part]
Whelan et al., 2023, Zool. J. Linn. Soc. 198(2): Table S1. [in part]
(misidentified reference to Anodon euscaphys Heude, 1879)

Alasmidonta uwharriensis Whelan et al., 2023
Whelan, Perkins & Mays in Whelan et al., 2023, Zool. J. Linn. Soc. 198(2): [21], fig. 6.

Taxonomic Opinions

Alasmidonta arcula (Lea, 1838)

Alasmidonta atropurpurea (Rafinesque, 1831)

Alasmidonta heterodon (Lea, 1830)

Alasmidonta marginata Say, 1818

Alasmidonta mccordi Athearn, 1964

Alasmidonta raveneliana (Lea, 1834)

Alasmidonta robusta Clarke, 1981

Alasmidonta triangulata (Lea, 1858)

Alasmidonta undulata (Say, 1817)

Alasmidonta uwharriensis Whelan, Perkins & Mays

Alasmidonta varicosa (Lamarck, 1819)

Alasmidonta viridis (Rafinesque, 1820)

Alasmidonta wrightiana (Walker, 1901)

Anodonta anatina

Anodonta arcaeformis
+ Anodonta arcaeformis var. flavotincta Martens, 1905

Anodonta cygnea

Anodonta euscaphys

Anodonta exulcerata

Anodonta implicata

Anodonta impura

Anodonta kennerlyi

Anodonta lucida

Anodonta nuttalliana

Anodonta oregonensis

Anodontoides denigrata

Anodontoides ferussacianus

Arcidens confragosus

Arcidens wheeleri

Cristaria plicata

Lasmigona complanata

Lasmigona compressa

Lasmigona costata

Lasmigona etowaensis

Lasmigona holstonia

Lasmigona subviridis

Pseudanodonta complanata

Pseudodontoideus connasaugaensis

Pseudodontoideus subvexus

Pyganodon cataracta

Pyganodon grandis

Pyganodon lacustris

Simpsonais [sic] ambigua

Sinanodonta tumens

Sinanodonta woodiana

Strophitus pascagoulaensis

Strophitus radiatus

Strophitus undulatus

Strophitus williamsi

Utterbackia imbecillis

Utterbackia peggyae

Utterbackia peninsularis

Utterbackiana couperiana

Utterbackiana hartfieldorum

Utterbackiana heardi

Utterbackiana suborbiculata

Cladogram Figure 2
                                +-------- Anemina_arcaeformis             
  +-----------------------------+                                         
  |                             |  +----- Sinanodonta_lucida              
  |                             +--+                                      
  |                                |  +-- Sinanodonta_woodiana            
  |                                +--+                                   
--+                                   +-- Sinanodonta_tumens              
  |                                                                       
  |  +----------------------------------- Anodonta_cygnea                 
  |  |                                                                    
  |  |                                +-- Anodonta_nuttalliana            
  |  |  +-----------------------------+                                   
  +--+  |                             +-- Pseudanodonta_complanata        
     |  |                                                                 
     |  |  +----------------------------- Anodonta_anatina                
     |  |  |                                                              
     |  |  |                       +----- Utterbackia_peninsularis__2     
     +--+  |                    +--+                                      
        |  |                    |  |  +-- Utterbackia_peninsularis__1     
        |  |                    |  +--+                                   
        |  |  +-----------------+     +-- Utterbackia_imbecillis          
        |  |  |                 |                                         
        |  |  |                 |  +----- Pyganodon_grandis__2            
        +--+  |                 +--+                                      
           |  |                    |  +-- Pyganodon_grandis__1            
           |  |                    +--+                                   
           |  |                       +-- Pyganodon_lacustris             
           |  |                                                           
           |  |                       +-- Utterbackiana_couperiana        
           |  |                    +--+                                   
           |  |                    |  +-- Utterbackiana_heardi            
           |  |              +-----+                                      
           |  |              |     |  +-- Utterbackiana_suborbiculata     
           +--+              |     +--+                                   
              |              |        +-- Utterbackiana_hartfieldorum     
              |  +-----------+                                            
              |  |           |     +----- Lasmigona_costata               
              |  |           |  +--+                                      
              |  |           |  |  |  +-- Pseudodontoideus_subvexus       
              |  |           |  |  +--+                                   
              |  |           +--+     +-- Pseudodontoideus_connasaugaensis
              |  |              |                                         
              |  |              |  +----- Lasmigona_complanata            
              |  |              +--+                                      
              |  |                 |  +-- Arcidens_confragosus            
              |  |                 +--+                                   
              |  |                    +-- Arcidens_wheeleri               
              +--+                                                        
                 |     +----------------- Pressodonta_viridis             
                 |     |                                                  
                 |  +--+              +-- Anodontoides_denigratus         
                 |  |  |  +-----------+                                   
                 |  |  |  |           +-- Anodontoides_ferussacianus      
                 |  |  +--+                                               
                 |  |     |  +----------- Strophitus_undulatus            
                 |  |     +--+                                            
                 |  |        |  +-------- Strophitus_pascagoulaensis__2   
                 |  |        +--+                                         
                 |  |           |  +----- Strophitus_radiatus             
                 +--+           +--+                                      
                    |              |  +-- Strophitus_pascagoulaensis__1   
                    |              +--+                                   
                    |                 +-- Strophitus_williamsi            
                    |                                                     
                    |              +----- Prolasmidonta_heterodon         
                    |  +-----------+                                      
                    |  |           |  +-- Platynaias_subviridis           
                    |  |           +--+                                   
                    |  |              +-- Platynaias_compressa            
                    +--+                                                  
                       |              +-- Alasminota_holstonia            
                       |  +-----------+                                   
                       |  |           +-- Alasminota_etowaensis           
                       |  |                                               
                       +--+           +-- Alasmidonta_raveneliana         
                          |  +--------+                                   
                          |  |        +-- Alasmidonta_marginata           
                          |  |                                            
                          +--+        +-- Alasmidonta_varicosa            
                             |  +-----+                                   
                             |  |     +-- Alasmidonta_uwharriensis        
                             +--+                                         
                                |  +----- Alasmidonta_undulata            
                                +--+                                      
                                   |  +-- Alasmidonta_triangulata         
                                   +--+                                   
                                      +-- Alasmidonta_arcula              
Cladogram Figure 3
  +-------------------------------- Anodonta_anatina                
  |                                                                 
  |                             +-- Utterbackiana_heardi            
  |                          +--+                                   
  |                          |  +-- Utterbackiana_couperiana        
  |  +-----------------------+                                      
--+  |                       |  +-- Utterbackiana_hartfieldorum     
  |  |                       +--+                                   
  |  |                          +-- Utterbackiana_suborbiculata     
  |  |                                                              
  |  |                          +-- Utterbackia_peninsularis        
  |  |  +-----------------------+                                   
  +--+  |                       +-- Pyganodon_grandis               
     |  |                                                           
     |  |     +-------------------- Alasminota_etowaensis           
     |  |     |                                                     
     |  |     |              +----- Pressodonta_viridis             
     |  |  +--+  +-----------+                                      
     |  |  |  |  |           |  +-- Platynaias_subviridis           
     |  |  |  |  |           +--+                                   
     +--+  |  |  |              +-- Prolasmidonta_heterodon         
        |  |  +--+                                                  
        |  |     |              +-- Anodontoides_ferussacianus      
        |  |     |  +-----------+                                   
        |  |     |  |           +-- Anodontoides_denigratus         
        |  |     +--+                                               
        |  |        |  +----------- Strophitus_undulatus            
        |  |        +--+                                            
        |  |           |  +-------- Strophitus_pascagoulaensis__2   
        |  |           +--+                                         
        +--+              |  +----- Strophitus_pascagoulaensis__1   
           |              +--+                                      
           |                 |  +-- Strophitus_radiatus             
           |                 +--+                                   
           |                    +-- Strophitus_williamsi            
           |                                                        
           |                    +-- Arcidens_confragosus            
           |                 +--+                                   
           |                 |  +-- Arcidens_wheeleri               
           |        +--------+                                      
           |        |        |  +-- Pseudodontoideus_connasaugaensis
           |        |        +--+                                   
           |        |           +-- Pseudodontoideus_subvexus       
           +--------+                                               
                    |           +-- Alasmidonta_marginata           
                    |  +--------+                                   
                    |  |        +-- Alasmidonta_raveneliana         
                    |  |                                            
                    +--+        +-- Alasmidonta_varicosa            
                       |  +-----+                                   
                       |  |     +-- Alasmidonta_uwharriensis        
                       +--+                                         
                          |  +----- Alasmidonta_undulata            
                          +--+                                      
                             |  +-- Alasmidonta_triangulata         
                             +--+                                   
                                +-- Alasmidonta_arcula              
Cladogram Figure 4
                          +-- Anodonta_anatina                   
  +-----------------------+                                      
  |                       +-- Anodonta_cygnea                    
  |                                                              
  |  +----------------------- Pressodonta_viridis                
  |  |                                                           
  |  |                 +----- Utterbackiana_couperiana           
--+  |                 |                                         
  |  |     +-----------+  +-- Utterbackiana_heardi               
  |  |     |           |  |                                      
  |  |     |           +--+-- Utterbackiana_hartfieldorum        
  |  |     |              |                                      
  |  |  +--+              +-- Utterbackiana_suborbiculata        
  |  |  |  |                                                     
  +--+  |  |  +-------------- Alasminota_etowaensis              
     |  |  |  |                                                  
     |  |  +--+           +-- Arcidens_confragosus               
     |  |     |  +--------+                                      
     |  |     |  |        +-- Arcidens_wheeleri                  
     |  |     +--+                                               
     |  |        |  +-------- Pseudodontoideus_connasaugaensis__3
     |  |        +--+                                            
     |  |           |  +----- Pseudodontoideus_connasaugaensis__2
     |  |           +--+                                         
     +--+              |  +-- Pseudodontoideus_subvexus          
        |              +--+                                      
        |                 +-- Pseudodontoideus_connasaugaensis__1
        |                                                        
        |              +----- Alasmidonta_raveneliana            
        |              |                                         
        |              |  +-- Alasmidonta_uwharriensis           
        |  +-----------+--+                                      
        |  |           |  +-- Alasmidonta_varicosa               
        |  |           |                                         
        |  |           |  +-- Alasmidonta_arcula                 
        |  |           +--+                                      
        +--+              +-- Alasmidonta_triangulata            
           |                                                     
           |  +-------------- Platynaias_subviridis              
           |  |                                                  
           |  |     +-------- Strophitus_pascagoulaensis         
           |  |  +--+                                            
           +--+  |  |  +----- Strophitus_undulatus               
              |  |  +--+                                         
              |  |     |  +-- Strophitus_radiatus                
              |  |     +--+                                      
              +--+        +-- Strophitus_williamsi               
                 |                                               
                 |        +-- Pyganodon_grandis                  
                 |  +-----+                                      
                 |  |     +-- Pyganodon_cataracta                
                 +--+                                            
                    |  +----- Utterbackia_peninsularis           
                    +--+                                         
                       |  +-- Anodontoides_ferussacianus         
                       +--+                                      
                          +-- Anodontoides_denigratus            

Page updated: 15:12:44 Wed 04 Dec 2024

 
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