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Page last updated
Wed 18 Jan 2023

The Freshwater Mussels (Unionoida) of the World (and other less consequential bivalves)

FM(U)otW(aolcb) is the web version of the MUSSEL Project Database. Follow the links to browse the data or use the custom Google search field. Either way, you win!

Cladomics pages are experimental — even more so than the rest of this site!

The purpose of the cladomics pages for each genus and family-group level taxon is to provide statistics and references indicating the extent to which published phylogenetic work supports the classification used here.

The Cladomics Summary provides an overview of the number of studies that have included terminal species representing this taxon as well as an indication of how frequently it has been recovered as monophyletic.

Following the summary, the available published clades representing the taxon are listed.

For more information, visit the Cladomics FARQ (frequently asked rhetorical questions) page. Abbreviations for the various statistics are explained on the FARQ!

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Bivalvia | Unionoida | Unionoidea | Unionidae | Ambleminae | Lampsilini

genus Villosa Frierson, 1927: cladomics
     4 valid species

Cladomics Summary

number of species = 4

analyzed in 15 cladograms in 11 publications
maximum number of terminal species in an analysis = 1, mean = 1.0

Basic Monophyly

number of cladograms that test basic monophyly (i.e., more than 1 included species) = 0
maximum number of included species = 0, mean = 0
proportion of cladograms that support basic monophyly = 0


Campbell et al., 2005, fig. 1

iT tT bi riS oT roT roS iD
1 1 0 1 116 48 22 --

*** Top Monophyly Cladogram *** *** Top Sister-Group Cladogram ***

                          +-- Ellipsaria_lineolata   
  +-----------------------+                          
  |                       +-- Truncilla_truncata     
  |                                                  
  |  +----------------------- Villosa_villosa **        
  |  |                                               
  |  |                    +-- Obovaria_olivaria      
--+  |     +--------------+                          
  |  |     |              +-- Venustaconcha_pleasii  
  |  |     |                                         
  |  |  +--+           +----- Obovaria_unicolor      
  |  |  |  |  +--------+                             
  |  |  |  |  |        |  +-- Obovaria_arkansasensis 
  +--+  |  |  |        +--+                          
     |  |  +--+           +-- Obovaria_subrotunda    
     |  |     |                                      
     |  |     |  +----------- Epioblasma_brevidens   
     |  |     +--+                                   
     |  |        |  +-------- Epioblasma_triquetra   
     |  |        +--+                                
     |  |           |  +----- Epioblasma_walkeri[2]  
     |  |           +--+                             
     |  |              |  +-- Epioblasma_capsaeformis
     +--+              +--+                          
        |                 +-- Epioblasma_walkeri[1]  
        |                                            
        |                 +-- Ligumia_recta          
        |        +--------+                          
        |        |        +-- Cambarunio_iris        
        |     +--+                                   
        |     |  |  +-------- Lampsilis_teres        
        |     |  +--+                                
        |     |     |  +----- Ortmanniana_ligamentina
        |     |     +--+                             
        |     |        |  +-- Lampsilis_siliquoidea  
        |     |        +--+                          
        |     |           +-- Leaunio_vanuxemensis   
        +-----+                                      
              |           +-- Lampsilis_cardium      
              |        +--+                          
              |        |  +-- Lampsilis_ovata        
              |  +-----+                             
              |  |     |  +-- Ortmanniana_pectorosa  
              |  |     +--+                          
              |  |        +-- Lampsilis_ornata       
              |  |                                   
              +--+     +----- Hamiota_australis[2]   
                 |  +--+                             
                 |  |  |  +-- Hamiota_australis[1]   
                 |  |  +--+                          
                 |  |     +-- Hamiota_subangulata    
                 +--+                                
                    |  +----- Hamiota_altilis[2]     
                    |  |                             
                    |  |  +-- Hamiota_altilis[1]     
                    +--+--+                          
                       |  +-- Hamiota_perovalis[1]   
                       |                             
                       |  +-- Hamiota_altilis[3]     
                       +--+                          
                          +-- Hamiota_perovalis[2]   

Lydeard et al., 1996, fig. 3

iT tT bi riS oT roT roS iD
1 1 0 1 26 8 6 --
  +----- Potamilus_purpuratus 
  |                           
  |  +-- Ellipsaria_lineolata 
--+--+                        
  |  +-- Villosa_delumbis **     
  |                           
  |  +-- Lampsilis_teres      
  +--+                        
     +-- Lampsilis_siliquoidea

Lydeard et al., 1996, fig. 4

iT tT bi riS oT roT roS iD
1 1 0 1 27 8 6 --
  +-------- Ellipsaria_lineolata 
--+                              
  |  +----- Villosa_delumbis **     
  +--+                           
     |  +-- Lampsilis_teres      
     +--+                        
        +-- Lampsilis_siliquoidea

Huang et al., 2002, fig. 1

iT tT bi riS oT roT roS iD
1 1 0 1 33 7 6 --
  +----- Lampsilis_teres     
--+                          
  |  +-- Villosa_delumbis **    
  +--+                       
     +-- Ellipsaria_lineolata

Huang et al., 2002, fig. 2

iT tT bi riS oT roT roS iD
1 1 0 1 33 7 6 --
  +-- Villosa_delumbis **        
  |                           
  +-- Ellipsaria_lineolata    
  |                           
--+-- Lampsilis_teres         
  |                           
  +-- Potamilus_purpuratus    
  |                           
  +-- Ptychobranchus_subtentus

Campbell et al., 2005, fig. 2

iT tT bi riS oT roT roS iD
1 1 0 1 114 48 22 --
  +-------------- Sagittunio_nasutus         
  |                                          
  +-------------- Leaunio_vanuxemensis       
  |                                          
  |           +-- Ellipsaria_lineolata       
  +-----------+                              
  |           +-- Truncilla_truncata         
  |                                          
  |        +----- Lampsilis_teres            
  +--------+                                 
  |        |  +-- Ortmanniana_ligamentina    
  |        +--+                              
  |           +-- Lampsilis_siliquoidea      
  |                                          
  |        +----- Villosa_villosa **            
  +--------+                                 
  |        |  +-- Ligumia_recta              
  |        +--+                              
  |           +-- Cambarunio_iris            
  |                                          
  |     +-------- Ortmanniana_pectorosa      
  +-----+                                    
  |     |  +----- Lampsilis_ornata           
  |     +--+                                 
  |        |  +-- Lampsilis_cardium          
  |        +--+                              
  |           +-- Lampsilis_ovata            
  |                                          
  |           +-- Potamilus_fragilis         
  |        +--+                              
  |        |  +-- Potamilus_leptodon         
  +--------+                                 
  |        |  +-- Potamilus_alatus           
  |        +--+                              
  |           +-- Potamilus_purpuratus       
  |                                          
  |           +-- Ptychobranchus_fasciolaris 
--+  +--------+                              
  |  |        +-- Venustaconcha_ellipsiformis
  |  |                                       
  +--+  +-------- Medionidus_conradicus      
  |  |  |                                    
  |  |  |     +-- Lemiox_rimosus             
  |  +--+-----+                              
  |     |     +-- Medionidus_acutissimus     
  |     |                                    
  |     |  +----- Dromus_dromas              
  |     +--+                                 
  |        |  +-- Cyprogenia_aberti          
  |        +--+                              
  |           +-- Cyprogenia_stegaria        
  |                                          
  |        +----- Hamiota_australis[2]       
  |  +-----+                                 
  |  |     |  +-- Hamiota_australis[1]       
  |  |     +--+                              
  |  |        +-- Hamiota_subangulata        
  +--+                                       
  |  |        +-- Hamiota_altilis[3]         
  |  |  +-----+                              
  |  |  |     +-- Hamiota_perovalis[2]       
  |  +--+                                    
  |     |  +----- Hamiota_altilis[2]         
  |     +--+                                 
  |        |  +-- Hamiota_altilis[1]         
  |        +--+                              
  |           +-- Hamiota_perovalis[1]       
  |                                          
  |  +----------- Obovaria_olivaria          
  |  |                                       
  |  +----------- Venustaconcha_pleasii      
  |  |                                       
  |  |     +----- Obovaria_unicolor          
  |  +-----+                                 
  +--+     |  +-- Obovaria_arkansasensis     
     |     +--+                              
     |        +-- Obovaria_subrotunda        
     |                                       
     |        +-- Epioblasma_brevidens       
     |  +-----+                              
     |  |     +-- Epioblasma_triquetra       
     +--+                                    
        |  +----- Epioblasma_walkeri[1]      
        +--+                                 
           |  +-- Epioblasma_walkeri[2]      
           +--+                              
              +-- Epioblasma_capsaeformis    

Buhay et al., 2002, fig. 2A

iT tT bi riS oT roT roS iD
1 1 0 1 9 9 3 --
  +-------- Lampsilis_siliquoidea     
  |                                   
  |     +-- Cambarunio_taeniatus      
  +-----+                             
  |     +-- Villosa_villosa **           
--+                                   
  |  +----- Epioblasma_triquetra      
  |  |                                
  |  +----- Epioblasma_brevidens      
  |  |                                
  +--+  +-- Epioblasma_capsaeformis[1]
     |  |                             
     |  +-- Epioblasma_capsaeformis[2]
     |  |                             
     +--+-- Epioblasma_capsaeformis[3]
        |                             
        +-- Epioblasma_walkeri[1]     
        |                             
        +-- Epioblasma_walkeri[2]     

Buhay et al., 2002, fig. 2B

iT tT bi riS oT roT roS iD
1 1 0 1 9 9 3 --
  +-------------- Lampsilis_siliquoidea     
  |                                         
  |           +-- Cambarunio_taeniatus      
  +-----------+                             
--+           +-- Villosa_villosa **           
  |                                         
  |  +----------- Epioblasma_brevidens      
  |  |                                      
  +--+  +-------- Epioblasma_triquetra      
     |  |                                   
     +--+  +----- Epioblasma_capsaeformis[3]
        |  |                                
        +--+  +-- Epioblasma_capsaeformis[1]
           |  |                             
           |  +-- Epioblasma_capsaeformis[2]
           +--+                             
              +-- Epioblasma_walkeri[1]     
              |                             
              +-- Epioblasma_walkeri[2]     

Roe et al., 2001, fig. 3a

iT tT bi riS oT roT roS iD
1 1 0 1 15 15 4 --
  +-------------- Ligumia_recta       
  |                                   
  +-------------- Obliquaria_reflexa  
  |                                   
  |        +----- Lampsilis_teres     
  +--------+                          
--+        |  +-- Lampsilis_ovata     
  |        +--+                       
  |           +-- Lampsilis_ornata    
  |                                   
  |  +----------- Villosa_villosa **     
  |  |                                
  |  |     +----- Hamiota_australis[2]
  +--+  +--+                          
     |  |  |  +-- Hamiota_subangulata 
     |  |  +--+                       
     |  |     +-- Hamiota_australis[1]
     +--+                             
        |  +----- Hamiota_altilis[3]  
        |  |                          
        |  |  +-- Hamiota_perovalis[1]
        +--+  |                       
           |  +-- Hamiota_perovalis[2]
           |  |                       
           |  +-- Hamiota_altilis[1]  
           +--+                       
              +-- Hamiota_altilis[2]  
              |                       
              +-- Hamiota_altilis[4]  
              |                       
              +-- Hamiota_altilis[5]  

Krebs et al., 2003, fig. 3

iT tT bi riS oT roT roS iD
1 1 0 1 33 10 7 --
           +-- Villosa_delumbis **     
  +--------+                        
  |        +-- Ellipsaria_lineolata 
--+                                 
  |  +-------- Lampsilis_teres      
  +--+                              
     |  +----- Lampsilis_siliquoidea
     +--+                           
        |  +-- Potamilus_purpuratus 
        +--+                        
           +-- Potamilus_alatus     

Turner et al., 2000, fig. 2

iT tT bi riS oT roT roS iD
1 1 0 1 15 10 2 --
  +-------------------- Medionidus_conradicus   
  |                                             
--+  +----------------- Lampsilis_teres         
  |  |                                          
  |  +----------------- Villosa_delumbis **        
  +--+                                          
     |              +-- Lampsilis_satura        
     |  +-----------+                           
     |  |           +-- Lampsilis_cardium       
     +--+                                       
        |  +----------- Lampsilis_reeveiana     
        |  |                                    
        +--+  +-------- Lampsilis_hydiana[1]    
           |  |                                 
           +--+-------- Lampsilis_hydiana[2]    
              |                                 
              |  +----- Lampsilis_siliquoidea[2]
              +--+                              
                 |  +-- Lampsilis_siliquoidea[1]
                 +--+                           
                    +-- Lampsilis_powellii      

Inoue et al., 2013, fig. 3

iT tT bi riS oT roT roS iD
1 1 0 1 16 9 5 --
  +-------------- Leaunio_vanuxemensis  
  |                                     
  |           +-- Villosa_villosa **       
--+-----------+                         
  |           +-- Cambarunio_iris       
  |                                     
  |  +----------- Obovaria_olivaria     
  +--+                                  
     |  +-------- Obovaria_arkansasensis
     +--+                               
        |  +----- Obovaria_retusa       
        +--+                            
           |  +-- Obovaria_unicolor     
           +--+                         
              +-- Obovaria_subrotunda   

Burlakova et al., 2019, fig. 1

iT tT bi riS oT roT roS iD
1 1 0 1 32 25 17 --
              +-- Potamilus_fragilis    
  +-----------+                         
  |           +-- Potamilus_metnecktayi 
  |                                     
  |        +----- Medionidus_acutissimus
  +--------+                            
  |        |  +-- Lemiox_rimosus        
  |        +--+                         
  |           +-- Dromus_dromas         
  |                                     
  |        +----- Ellipsaria_lineolata  
  |        |                            
  +--------+  +-- Truncilla_cognata     
  |        |  |                         
--+        |  +-- Truncilla_donaciformis
  |        +--+                         
  |           +-- Truncilla_macrodon    
  |           |                         
  |           +-- Truncilla_truncata    
  |                                     
  |  +----------- Cambarunio_simus      
  |  |                                  
  |  +----------- Leaunio_lienosus      
  |  |                                  
  |  +----------- Villosa_villosa **       
  |  |                                  
  |  |        +-- Sagittunio_nasutus    
  |  +--------+                         
  +--+        +-- Lampsilis_ovata       
     |                                  
     |     +----- Venustaconcha_pleasii 
     +-----+                            
     |     |  +-- Epioblasma_rangiana   
     |     +--+                         
     |        +-- Obovaria_unicolor     
     |                                  
     |  +-------- Lampsilis_teres       
     +--+                               
        |  +----- Lampsilis_siliquoidea 
        +--+                            
           |  +-- Lampsilis_hydiana     
           +--+                         
              +-- Lampsilis_straminea   

Smith et al., 2019, fig. 2

iT tT bi riS oT roT roS iD
1 1 0 1 41 36 19 --
                       +-- Hamiota_subangulata        
  +--------------------+                              
  |                    +-- Villosa_villosa **            
  |                                                   
--+  +-------------------- Ligumia_recta              
  |  |                                                
  |  |              +----- Venustaconcha_ellipsiformis
  +--+  +-----------+                                 
     |  |           |  +-- Obovaria_subrotunda        
     |  |           +--+                              
     +--+              +-- Epioblasma_rangiana        
        |                                             
        |  +-------------- Ortmanniana_pectorosa      
        +--+                                          
           |  +----------- Ortmanniana_ligamentina    
           +--+                                       
              |  +-------- Lampsilis_ovata            
              +--+                                    
                 |  +----- Lampsilis_dolabraeformis   
                 +--+                                 
                    |  +-- Lampsilis_satura           
                    +--+                              
                       +-- Lampsilis_cardium          

Patterson et al., 2021, fig. 3

iT tT bi riS oT roT roS iD
1 1 0 1 3 0 0 --
-- Villosa_villosa

Page updated: 08:56:30 Wed 18 Jan 2023

 
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