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Page last updated
Sun 06 Oct 2024

The Freshwater Mussels (Unionoida) of the World (and other less consequential bivalves)

FM(U)otW(aolcb) is the web version of the MUSSEL Project Database. Follow the links to browse the data or use the custom Google search field. Either way, you win!

Cladomics pages are experimental — even more so than the rest of this site!

The purpose of the cladomics pages for each genus and family-group level taxon is to provide statistics and references indicating the extent to which published phylogenetic work supports the classification used here.

The Cladomics Summary provides an overview of the number of studies that have included terminal species representing this taxon as well as an indication of how frequently it has been recovered as monophyletic.

Following the summary, the available published clades representing the taxon are listed.

For more information, visit the Cladomics FARQ (frequently asked rhetorical questions) page. Abbreviations for the various statistics are explained on the FARQ!

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Bivalvia | Unionoida | Unionoidea | Unionidae | Ambleminae | Lampsilini

genus Villosa Frierson, 1927: cladomics
     4 valid species

Cladomics Summary

number of species = 4

analyzed in 15 cladograms in 11 publications
maximum number of terminals in an analysis = 1, mean = 1.0

Basic Monophyly

number of cladograms that test basic monophyly (i.e., more than 1 included species) = 0
maximum number of included terminals = 0, mean = 0
proportion of cladograms that support basic monophyly = 0


Campbell et al., 2005 [Publ 2660], fig. 1

iT tT bi riS oT roT roS iD
1 1 0 1 101 42 22 --

*** Top Monophyly Cladogram *** *** Top Sister-Group Cladogram ***

                       +-------- Toxolasma                     
  +--------------------+                                       
  |                    |  +----- Obliquaria                    
  |                    +--+                                    
  |                       |  +-- Cyrtonaias                    
  |                       +--+                                 
  |                          +-- Glebula                       
  |                                                            
  |              +-------------- Potamilus                     
  |              |                                             
  |              |     +-------- Medionidus_acutissimus        
--+              |  +--+                                       
  |  +-----------+  |  |  +----- Lemiox                        
  |  |           |  |  +--+                                    
  |  |           |  |     |  +-- Dromus                        
  |  |           |  |     +--+                                 
  |  |           +--+        +-- Cyprogenia                    
  |  |              |                                          
  |  |              |        +-- Sagittunio                    
  |  |              |     +--+                                 
  |  |              |     |  +-- Medionidus_conradicus         
  +--+              +-----+                                    
     |                    |  +-- Ptychobranchus                
     |                    +--+                                 
     |                       +-- Venustaconcha_ellipsiformis__2
     |                                                         
     |                       +-- Ellipsaria                    
     |  +--------------------+                                 
     |  |                    +-- Truncilla                     
     |  |                                                      
     |  |  +-------------------- Villosa_villosa **               
     +--+  |                                                   
        |  |                 +-- Obovaria_olivaria             
        |  |        +--------+                                 
        |  |        |        +-- Venustaconcha_ellipsiformis__1
        |  |  +-----+                                          
        +--+  |     |  +-------- Epioblasma                    
           |  |     +--+                                       
           |  |        |  +----- Obovaria_unicolor             
           |  |        +--+                                    
           |  |           |  +-- Obovaria_arkansasensis        
           |  |           +--+                                 
           |  |              +-- Obovaria_subrotunda           
           +--+                                                
              |        +-------- Hamiota                       
              |        |                                       
              |  +-----+     +-- Lampsilis_cardium             
              |  |     |  +--+                                 
              |  |     |  |  +-- Lampsilis_ovata               
              |  |     +--+                                    
              |  |        |  +-- Ortmanniana_pectorosa         
              +--+        +--+                                 
                 |           +-- Lampsilis_ornata              
                 |                                             
                 |           +-- Ligumia                       
                 |  +--------+                                 
                 |  |        +-- Cambarunio                    
                 +--+                                          
                    |  +-------- Lampsilis_teres               
                    +--+                                       
                       |  +----- Ortmanniana_ligamentina       
                       +--+                                    
                          |  +-- Lampsilis_siliquoidea         
                          +--+                                 
                             +-- Leaunio                       

Inoue et al., 2013 [Publ 3528], fig. 3

iT tT bi riS oT roT roS iD
1 1 0 1 16 9 5 --
  +-------- Lampsilis      
  |                        
--+  +----- Leaunio        
  |  |                     
  +--+----- Obovaria       
     |                     
     |  +-- Villosa_villosa **
     +--+                  
        +-- Cambarunio     

Krebs et al., 2003 [Publ 2691], fig. 3

iT tT bi riS oT roT roS iD
1 1 0 1 32 10 7 --
           +----- Ptychobranchus       
  +--------+                           
  |        |  +-- Potamilus_fragilis   
  |        +--+                        
  |           +-- Medionidus           
--+                                    
  |           +-- Villosa_delumbis **     
  |  +--------+                        
  |  |        +-- Ellipsaria           
  +--+                                 
     |  +-------- Lampsilis_teres      
     +--+                              
        |  +----- Lampsilis_siliquoidea
        +--+                           
           |  +-- Potamilus_purpuratus 
           +--+                        
              +-- Potamilus_alatus     

Lydeard et al., 1996 [Publ 2656], fig. 3

iT tT bi riS oT roT roS iD
1 1 0 1 26 8 6 --
  +-------------- Potamilus_fragilis  
  |                                   
--+  +----------- Ptychobranchus      
  |  |                                
  +--+  +-------- Medionidus          
     |  |                             
     +--+  +----- Potamilus_purpuratus
        |  |                          
        +--+----- Lampsilis           
           |                          
           |  +-- Ellipsaria          
           +--+                       
              +-- Villosa_delumbis **    

Smith et al., 2019 [Publ 3951], fig. 2

iT tT bi riS oT roT roS iD
1 1 0 1 40 35 19 --
  +----------------------- Obliquaria             
  |                                               
  |                    +-- Medionidus             
  |                 +--+                          
  |                 |  +-- Lemiox                 
--+  +--------------+                             
  |  |              |  +-- Cyprogenia             
  |  |              +--+                          
  |  |                 +-- Ptychobranchus         
  |  |                                            
  +--+           +-------- Potamilus              
     |  +--------+                                
     |  |        |  +----- Atlanticoncha          
     |  |        +--+                             
     |  |           |  +-- Ellipsaria             
     |  |           +--+                          
     +--+              +-- Truncilla              
        |                                         
        |              +-- Hamiota                
        |  +-----------+                          
        |  |           +-- Villosa_villosa **        
        |  |                                      
        +--+  +----------- Ligumia                
           |  |                                   
           |  |     +----- Venustaconcha          
           +--+  +--+                             
              |  |  |  +-- Obovaria               
              |  |  +--+                          
              +--+     +-- Epioblasma             
                 |                                
                 |  +----- Ortmanniana_pectorosa  
                 +--+                             
                    |  +-- Ortmanniana_ligamentina
                    +--+                          
                       +-- Lampsilis              

Roe et al., 2001 [Publ 2689], fig. 3a

iT tT bi riS oT roT roS iD
1 1 0 1 9 9 4 --
  +----- Ligumia        
  |                     
  +----- Obliquaria     
--+                     
  +----- Lampsilis      
  |                     
  |  +-- Villosa_villosa **
  +--+                  
     +-- Hamiota        

Burlakova et al., 2019 [Publ 3899], fig. 1

iT tT bi riS oT roT roS iD
1 1 0 1 32 25 17 --
  +----------------- Toxolasma            
  |                                       
  |  +-------------- Potamilus            
  |  |                                    
  |  |           +-- Ellipsaria           
  |  +-----------+                        
--+  |           +-- Truncilla            
  |  |                                    
  |  |        +----- Medionidus           
  |  +--------+                           
  |  |        |  +-- Lemiox               
  |  |        +--+                        
  +--+           +-- Dromus               
     |                                    
     |  +----------- Cambarunio           
     |  |                                 
     |  +----------- Leaunio              
     |  |                                 
     |  +----------- Villosa_villosa **      
     |  |                                 
     |  |        +-- Sagittunio           
     |  +--------+                        
     +--+        +-- Lampsilis_ovata      
        |                                 
        |     +----- Venustaconcha        
        +-----+                           
        |     |  +-- Epioblasma           
        |     +--+                        
        |        +-- Obovaria             
        |                                 
        |  +-------- Lampsilis_teres      
        +--+                              
           |  +----- Lampsilis_siliquoidea
           +--+                           
              |  +-- Lampsilis_hydiana    
              +--+                        
                 +-- Lampsilis_straminea  

Patterson et al., 2021 [Publ 4222], fig. 3

iT tT bi riS oT roT roS iD
1 1 0 1 3 0 0 --
-- Villosa_villosa

Lydeard et al., 1996 [Publ 2656], fig. 4

iT tT bi riS oT roT roS iD
1 1 0 1 27 8 6 --
  +-------------------- Obliquaria          
--+                                         
  |  +----------------- Medionidus          
  +--+                                      
     |  +-------------- Ptychobranchus      
     +--+                                   
        |  +----------- Potamilus_fragilis  
        +--+                                
           |  +-------- Potamilus_purpuratus
           +--+                             
              |  +----- Ellipsaria          
              +--+                          
                 |  +-- Villosa_delumbis **    
                 +--+                       
                    +-- Lampsilis           

Campbell et al., 2005 [Publ 2660], fig. 2

iT tT bi riS oT roT roS iD
1 1 0 1 101 42 22 --
           +----- Obliquaria                    
  +--------+                                    
  |        |  +-- Cyrtonaias                    
  |        +--+                                 
  |           +-- Glebula                       
  |                                             
  |  +----------- Sagittunio                    
  |  |                                          
  |  +----------- Leaunio                       
  |  |                                          
  |  +----------- Potamilus                     
  |  |                                          
  |  +----------- Hamiota                       
  |  |                                          
  |  |        +-- Ellipsaria                    
--+  +--------+                                 
  |  |        +-- Truncilla                     
  |  |                                          
  |  |     +----- Lampsilis_teres               
  |  +-----+                                    
  |  |     |  +-- Ortmanniana_ligamentina       
  |  |     +--+                                 
  |  |        +-- Lampsilis_siliquoidea         
  |  |                                          
  |  |     +----- Villosa_villosa **               
  |  +-----+                                    
  |  |     |  +-- Ligumia                       
  |  |     +--+                                 
  |  |        +-- Cambarunio                    
  |  |                                          
  +--+  +-------- Ortmanniana_pectorosa         
     +--+                                       
     |  |  +----- Lampsilis_ornata              
     |  +--+                                    
     |     |  +-- Lampsilis_cardium             
     |     +--+                                 
     |        +-- Lampsilis_ovata               
     |                                          
     |  +-------- Obovaria_olivaria             
     |  |                                       
     |  +-------- Venustaconcha_ellipsiformis__2
     +--+                                       
     |  +-------- Epioblasma                    
     |  |                                       
     |  |  +----- Obovaria_unicolor             
     |  +--+                                    
     |     |  +-- Obovaria_arkansasensis        
     |     +--+                                 
     |        +-- Obovaria_subrotunda           
     |                                          
     |        +-- Ptychobranchus                
     |  +-----+                                 
     |  |     +-- Venustaconcha_ellipsiformis__1
     |  |                                       
     +--+  +----- Medionidus_conradicus         
        |  |                                    
        |  |  +-- Lemiox                        
        +--+--+                                 
           |  +-- Medionidus_acutissimus        
           |                                    
           |  +-- Dromus                        
           +--+                                 
              +-- Cyprogenia                    

Turner et al., 2000 [Publ 2722], fig. 2

iT tT bi riS oT roT roS iD
1 1 0 1 13 8 2 --
  +-------------------- Medionidus              
  |                                             
--+  +----------------- Lampsilis_teres         
  |  |                                          
  |  +----------------- Villosa_delumbis **        
  +--+                                          
     |              +-- Lampsilis_satura        
     |  +-----------+                           
     |  |           +-- Lampsilis_cardium       
     +--+                                       
        |  +----------- Lampsilis_reeveiana     
        |  |                                    
        +--+  +-------- Lampsilis_hydiana__1    
           |  |                                 
           +--+-------- Lampsilis_hydiana__2    
              |                                 
              |  +----- Lampsilis_siliquoidea__2
              +--+                              
                 |  +-- Lampsilis_siliquoidea__1
                 +--+                           
                    +-- Lampsilis_powellii      

Huang et al., 2002 [Publ 2657], fig. 2

iT tT bi riS oT roT roS iD
1 1 0 1 33 7 6 --
  +-- Villosa_delumbis **
  |                   
  +-- Ellipsaria      
  |                   
--+-- Lampsilis       
  |                   
  +-- Potamilus       
  |                   
  +-- Ptychobranchus  

Buhay et al., 2002 [Publ 2661], fig. 2B

iT tT bi riS oT roT roS iD
1 1 0 1 6 6 3 --
  +----- Lampsilis      
  |                     
--+----- Epioblasma     
  |                     
  |  +-- Cambarunio     
  +--+                  
     +-- Villosa_villosa **

Huang et al., 2002 [Publ 2657], fig. 1

iT tT bi riS oT roT roS iD
1 1 0 1 33 7 6 --
  +----------------- Medionidus          
--+                                      
  |  +-------------- Potamilus_fragilis  
  +--+                                   
     |  +----------- Ptychobranchus      
     +--+                                
        |  +-------- Potamilus_purpuratus
        +--+                             
           |  +----- Lampsilis           
           +--+                          
              |  +-- Villosa_delumbis **    
              +--+                       
                 +-- Ellipsaria          

Buhay et al., 2002 [Publ 2661], fig. 2A

iT tT bi riS oT roT roS iD
1 1 0 1 6 6 3 --
  +----- Lampsilis      
  |                     
--+----- Epioblasma     
  |                     
  |  +-- Cambarunio     
  +--+                  
     +-- Villosa_villosa **

Page updated: 10:15:18 Sun 06 Oct 2024

 
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