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Page last updated
Thu 02 Feb 2023

The Freshwater Mussels (Unionoida) of the World (and other less consequential bivalves)

FM(U)otW(aolcb) is the web version of the MUSSEL Project Database. Follow the links to browse the data or use the custom Google search field. Either way, you win!

Cladomics pages are experimental — even more so than the rest of this site!

The purpose of the cladomics pages for each genus and family-group level taxon is to provide statistics and references indicating the extent to which published phylogenetic work supports the classification used here.

The Cladomics Summary provides an overview of the number of studies that have included terminal species representing this taxon as well as an indication of how frequently it has been recovered as monophyletic.

Following the summary, the available published clades representing the taxon are listed.

For more information, visit the Cladomics FARQ (frequently asked rhetorical questions) page. Abbreviations for the various statistics are explained on the FARQ!

Search the database pages:

Bivalvia | Unionoida | Unionoidea | Unionidae | Ambleminae | Lampsilini

genus Obovaria Rafinesque, 1819: cladomics
     7 valid species

Cladomics Summary

number of species = 7

analyzed in 27 cladograms in 15 publications
maximum number of terminal species in an analysis = 5, mean = 2.15

Basic Monophyly

number of cladograms that test basic monophyly (i.e., more than 1 included species) = 18
maximum number of included species = 5, mean = 2.72
proportion of cladograms that support basic monophyly = 0.56


Inoue et al., 2013, fig. 3

iT tT bi riS oT roT roS iD
5 5 3 5 12 5 5 0.0

*** Top Monophyly Cladogram ***

  +----------------- Lampsilis_ornata      
  |                                        
--+  +-------------- Leaunio_vanuxemensis  
  |  |                                     
  |  |           +-- Villosa_villosa       
  +--+-----------+                         
     |           +-- Cambarunio_iris       
     |                                     
     |  +----------- Obovaria_olivaria **     
     +--+                                  
        |  +-------- Obovaria_arkansasensis **
        +--+                               
           |  +----- Obovaria_retusa **       
           +--+                            
              |  +-- Obovaria_unicolor **     
              +--+                         
                 +-- Obovaria_subrotunda **   

Campbell et al., 2005, fig. 1

iT tT bi riS oT roT roS iD
4 10 7 4 113 45 22 9.0

*** Top Sister-Group Cladogram ***

  +----------------------- Villosa_villosa        
  |                                               
  |                    +-- Obovaria_olivaria **      
  |     +--------------+                          
  |     |              +-- Venustaconcha_pleasii  
  |     |                                         
  |  +--+           +----- Obovaria_unicolor **      
  |  |  |  +--------+                             
  |  |  |  |        |  +-- Obovaria_arkansasensis ** 
--+  |  |  |        +--+                          
  |  |  +--+           +-- Obovaria_subrotunda **    
  |  |     |                                      
  |  |     |  +----------- Epioblasma_brevidens   
  |  |     +--+                                   
  |  |        |  +-------- Epioblasma_triquetra   
  |  |        +--+                                
  |  |           |  +----- Epioblasma_walkeri[2]  
  |  |           +--+                             
  |  |              |  +-- Epioblasma_capsaeformis
  +--+              +--+                          
     |                 +-- Epioblasma_walkeri[1]  
     |                                            
     |                 +-- Ligumia_recta          
     |        +--------+                          
     |        |        +-- Cambarunio_iris        
     |     +--+                                   
     |     |  |  +-------- Lampsilis_teres        
     |     |  +--+                                
     |     |     |  +----- Ortmanniana_ligamentina
     |     |     +--+                             
     |     |        |  +-- Lampsilis_siliquoidea  
     |     |        +--+                          
     |     |           +-- Leaunio_vanuxemensis   
     +-----+                                      
           |           +-- Lampsilis_cardium      
           |        +--+                          
           |        |  +-- Lampsilis_ovata        
           |  +-----+                             
           |  |     |  +-- Ortmanniana_pectorosa  
           |  |     +--+                          
           |  |        +-- Lampsilis_ornata       
           |  |                                   
           +--+     +----- Hamiota_australis[2]   
              |  +--+                             
              |  |  |  +-- Hamiota_australis[1]   
              |  |  +--+                          
              |  |     +-- Hamiota_subangulata    
              +--+                                
                 |  +----- Hamiota_altilis[2]     
                 |  |                             
                 |  |  +-- Hamiota_altilis[1]     
                 +--+--+                          
                    |  +-- Hamiota_perovalis[1]   
                    |                             
                    |  +-- Hamiota_altilis[3]     
                    +--+                          
                       +-- Hamiota_perovalis[2]   

Campbell et al., 2005, fig. 2

iT tT bi riS oT roT roS iD
4 10 6 4 111 45 22 9.0
              +----- Obliquaria_reflexa         
  +-----------+                                 
  |           |  +-- Cyrtonaias_tampicoensis    
  |           +--+                              
  |              +-- Glebula_rotundata          
  |                                             
  |  +-------------- Sagittunio_nasutus         
  |  |                                          
  |  +-------------- Leaunio_vanuxemensis       
  |  |                                          
  |  |           +-- Ellipsaria_lineolata       
  |  +-----------+                              
  |  |           +-- Truncilla_truncata         
  |  |                                          
  |  |        +----- Lampsilis_teres            
  |  +--------+                                 
  |  |        |  +-- Ortmanniana_ligamentina    
  |  |        +--+                              
  |  |           +-- Lampsilis_siliquoidea      
  |  |                                          
  |  |        +----- Villosa_villosa            
  |  +--------+                                 
--+  |        |  +-- Ligumia_recta              
  |  |        +--+                              
  |  |           +-- Cambarunio_iris            
  |  |                                          
  |  |     +-------- Ortmanniana_pectorosa      
  |  +-----+                                    
  |  |     |  +----- Lampsilis_ornata           
  |  |     +--+                                 
  |  |        |  +-- Lampsilis_cardium          
  |  |        +--+                              
  |  |           +-- Lampsilis_ovata            
  |  |                                          
  |  |           +-- Potamilus_fragilis         
  |  |        +--+                              
  |  |        |  +-- Potamilus_leptodon         
  |  +--------+                                 
  |  |        |  +-- Potamilus_alatus           
  |  |        +--+                              
  |  |           +-- Potamilus_purpuratus       
  |  |                                          
  |  |           +-- Ptychobranchus_fasciolaris 
  +--+  +--------+                              
     |  |        +-- Venustaconcha_ellipsiformis
     |  |                                       
     +--+  +-------- Medionidus_conradicus      
     |  |  |                                    
     |  |  |     +-- Lemiox_rimosus             
     |  +--+-----+                              
     |     |     +-- Medionidus_acutissimus     
     |     |                                    
     |     |  +----- Dromus_dromas              
     |     +--+                                 
     |        |  +-- Cyprogenia_aberti          
     |        +--+                              
     |           +-- Cyprogenia_stegaria        
     |                                          
     |        +----- Hamiota_australis[2]       
     |  +-----+                                 
     |  |     |  +-- Hamiota_australis[1]       
     |  |     +--+                              
     |  |        +-- Hamiota_subangulata        
     +--+                                       
     |  |        +-- Hamiota_altilis[3]         
     |  |  +-----+                              
     |  |  |     +-- Hamiota_perovalis[2]       
     |  +--+                                    
     |     |  +----- Hamiota_altilis[2]         
     |     +--+                                 
     |        |  +-- Hamiota_altilis[1]         
     |        +--+                              
     |           +-- Hamiota_perovalis[1]       
     |                                          
     |  +----------- Obovaria_olivaria **          
     |  |                                       
     |  +----------- Venustaconcha_pleasii      
     |  |                                       
     |  |     +----- Obovaria_unicolor **          
     |  +-----+                                 
     +--+     |  +-- Obovaria_arkansasensis **     
        |     +--+                              
        |        +-- Obovaria_subrotunda **        
        |                                       
        |        +-- Epioblasma_brevidens       
        |  +-----+                              
        |  |     +-- Epioblasma_triquetra       
        +--+                                    
           |  +----- Epioblasma_walkeri[1]      
           +--+                                 
              |  +-- Epioblasma_walkeri[2]      
              +--+                              
                 +-- Epioblasma_capsaeformis    

Zanatta & Murphy, 2006, fig. 1

iT tT bi riS oT roT roS iD
4 4 2 4 53 45 22 3.0
                 +-- Venustaconcha_ellipsiformis
  +--------------+                              
  |              +-- Venustaconcha_pleasii      
  |                                             
  |        +-------- Obovaria_olivaria **          
--+  +-----+                                    
  |  |     |  +----- Obovaria_unicolor **          
  |  |     +--+                                 
  |  |        |  +-- Obovaria_subrotunda **        
  +--+        +--+                              
     |           +-- Obovaria_arkansasensis **     
     |                                          
     |  +----------- Epioblasma_brevidens       
     +--+                                       
        |  +-------- Epioblasma_triquetra       
        +--+                                    
           |  +----- Epioblasma_rangiana        
           +--+                                 
              |  +-- Epioblasma_walkeri         
              +--+                              
                 +-- Epioblasma_capsaeformis    

Zanatta & Murphy, 2006, fig. 2

iT tT bi riS oT roT roS iD
4 11 7 4 54 45 22 10.0
  +-------------------- Ortmanniana_ligamentina    
  |                                                
  |  +----------------- Lampsilis_siliquoidea      
--+  |                                             
  |  |  +-------------- Obovaria_olivaria **          
  |  |  |                                          
  +--+  |           +-- Venustaconcha_ellipsiformis
     |  +-----------+                              
     |  |           +-- Venustaconcha_pleasii      
     |  |                                          
     +--+        +----- Obovaria_unicolor **          
        +--------+                                 
        |        |  +-- Obovaria_subrotunda **        
        |        +--+                              
        |           +-- Obovaria_arkansasensis **     
        |                                          
        |  +----------- Epioblasma_brevidens       
        +--+                                       
           |  +-------- Epioblasma_triquetra       
           +--+                                    
              |  +----- Epioblasma_rangiana        
              +--+                                 
                 |  +-- Epioblasma_walkeri         
                 +--+                              
                    +-- Epioblasma_capsaeformis    

Porto-Hannes et al., 2019, fig. 2

iT tT bi riS oT roT roS iD
3 3 1 3 17 16 5 4.0
                    +-- Cambarunio_iris        
  +-----------------+                          
  |                 +-- Leaunio_lienosus       
  |                                            
  |        +----------- Lampsilis_teres        
  |        |                                   
  +--------+  +-------- Ortmanniana_ligamentina
  |        |  |                                
  |        +--+     +-- Lampsilis_hydiana      
  |           |  +--+                          
--+           |  |  +-- Lampsilis_virescens    
  |           +--+                             
  |              |  +-- Lampsilis_siliquoidea  
  |              +--+                          
  |                 +-- Lampsilis_radiata      
  |                                            
  |              +----- Obovaria_arkansasensis ** 
  |  +-----------+                             
  |  |           |  +-- Obovaria_olivaria **      
  |  |           +--+                          
  +--+              +-- Obovaria_subrotunda **    
     |                                         
     |  +-------------- Lampsilis_bracteata    
     |  |                                      
     +--+  +----------- Lampsilis_ornata       
        |  |                                   
        +--+  +-------- Lampsilis_ovata        
           |  |                                
           +--+  +----- Lampsilis_cardium[3]   
              |  |                             
              +--+  +-- Lampsilis_satura       
                 |  |                          
                 +--+-- Lampsilis_cardium[1]   
                    |                          
                    +-- Lampsilis_cardium[2]   

Porto-Hannes et al., 2019, fig. 3

iT tT bi riS oT roT roS iD
3 3 1 3 17 16 5 4.0
  +----------------- Lampsilis_teres        
  |                                         
  |              +-- Leaunio_lienosus       
  +--------------+                          
  |              +-- Cambarunio_iris        
  |                                         
  |           +----- Obovaria_arkansasensis ** 
  +-----------+                             
  |           |  +-- Obovaria_olivaria **      
  |           +--+                          
  |              +-- Obovaria_subrotunda **    
  |                                         
--+        +-------- Ortmanniana_ligamentina
  |        |                                
  +--------+     +-- Lampsilis_hydiana      
  |        |  +--+                          
  |        |  |  +-- Lampsilis_virescens    
  |        +--+                             
  |           |  +-- Lampsilis_siliquoidea  
  |           +--+                          
  |              +-- Lampsilis_radiata      
  |                                         
  |  +-------------- Lampsilis_bracteata    
  |  |                                      
  +--+  +----------- Lampsilis_ornata       
     |  |                                   
     +--+  +-------- Lampsilis_ovata        
        |  |                                
        +--+  +----- Lampsilis_cardium[3]   
           |  |                             
           +--+  +-- Lampsilis_satura       
              |  |                          
              +--+-- Lampsilis_cardium[1]   
                 |                          
                 +-- Lampsilis_cardium[2]   

Lydeard et al., 2000, fig. 2a

iT tT bi riS oT roT roS iD
2 2 0 2 15 1 1 --
  +----- Lampsilis_teres  
--+                       
  |  +-- Obovaria_unicolor **
  +--+                    
     +-- Obovaria_olivaria **

Lydeard et al., 2000, fig. 2b

iT tT bi riS oT roT roS iD
2 2 0 2 15 1 1 --
  +----- Lampsilis_teres  
--+                       
  |  +-- Obovaria_unicolor **
  +--+                    
     +-- Obovaria_olivaria **

Lydeard et al., 2000, fig. 2c

iT tT bi riS oT roT roS iD
2 3 0 2 14 1 1 4.0
  +----- Obovaria_unicolor **
--+                       
  |  +-- Obovaria_olivaria **
  +--+                    
     +-- Lampsilis_teres  

Lydeard et al., 2000, fig. 2d

iT tT bi riS oT roT roS iD
2 2 0 2 14 1 1 --
  +----- Lampsilis_teres  
--+                       
  |  +-- Obovaria_unicolor **
  +--+                    
     +-- Obovaria_olivaria **

Campbell & Lydeard, 2012, fig. 1

iT tT bi riS oT roT roS iD
2 2 0 2 42 3 3 --
  +-------- Lemiox_rimosus   
--+                          
  |  +----- Lampsilis_ovata  
  +--+                       
     |  +-- Obovaria_unicolor **
     +--+                    
        +-- Obovaria_olivaria **

Campbell & Lydeard, 2012, fig. 2

iT tT bi riS oT roT roS iD
2 2 0 2 42 3 3 --
  +-------- Lemiox_rimosus   
--+                          
  |  +----- Lampsilis_ovata  
  +--+                       
     |  +-- Obovaria_unicolor **
     +--+                    
        +-- Obovaria_olivaria **

Campbell & Lydeard, 2012, fig. 3

iT tT bi riS oT roT roS iD
2 4 0 2 49 3 3 5.0
  +-------- Obovaria_unicolor **
--+                          
  |  +----- Obovaria_olivaria **
  +--+                       
     |  +-- Lampsilis_ovata  
     +--+                    
        +-- Lemiox_rimosus   

Campbell & Lydeard, 2012, fig. 4

iT tT bi riS oT roT roS iD
2 2 0 2 42 3 3 --
  +-------- Lemiox_rimosus   
--+                          
  |  +----- Lampsilis_ovata  
  +--+                       
     |  +-- Obovaria_unicolor **
     +--+                    
        +-- Obovaria_olivaria **

Campbell & Lydeard, 2012, fig. 1-2

iT tT bi riS oT roT roS iD
2 5 0 2 72 13 10 6.0
              +-- Lampsilis_ovata            
  +-----------+                              
  |           +-- Obliquaria_reflexa         
--+                                          
  |  +----------- Lampsilis_virescens        
  |  |                                       
  +--+        +-- Epioblasma_rangiana        
     |  +-----+                              
     |  |     +-- Obovaria_arkansasensis **     
     +--+                                    
        |  +----- Obovaria_olivaria **          
        +--+                                 
           |  +-- Venustaconcha_ellipsiformis
           +--+                              
              +-- Venustaconcha_pleasii      

Campbell & Lydeard, 2012, fig. 3

iT tT bi riS oT roT roS iD
2 5 0 2 72 13 10 6.0
  +-------------- Lampsilis_ovata            
  |                                          
--+  +----------- Lampsilis_virescens        
  |  |                                       
  +--+        +-- Epioblasma_rangiana        
     |  +-----+                              
     |  |     +-- Obovaria_arkansasensis **     
     +--+                                    
        |  +----- Obovaria_olivaria **          
        +--+                                 
           |  +-- Venustaconcha_ellipsiformis
           +--+                              
              +-- Venustaconcha_pleasii      

Inoue et al., 2018, fig. 1, 3a-c

iT tT bi riS oT roT roS iD
2 12 0 2 202 9 7 13.0
  +-- Dromus_dromas         
  |                         
  +-- Epioblasma_rangiana   
  |                         
  +-- Glebula_rotundata     
  |                         
  +-- Lampsilis_ovata       
  |                         
  +-- Lampsilis_virescens   
  |                         
  +-- Lemiox_rimosus        
--+                         
  +-- Obliquaria_reflexa    
  |                         
  +-- Obovaria_olivaria **     
  |                         
  +-- Obovaria_unicolor **     
  |                         
  +-- Plectomerus_dombeyanus
  |                         
  +-- Potamilus_alatus      
  |                         
  +-- Potamilus_capax       

Chapman et al., 2008, fig. 1

iT tT bi riS oT roT roS iD
1 1 0 1 20 12 10 --
              +-- Venustaconcha_ellipsiformis
  +-----------+                              
  |           +-- Obovaria_olivaria **          
--+                                          
  |  +----------- Hamiota_subangulata        
  +--+                                       
     |  +-------- Lampsilis_ovata            
     +--+                                    
        |  +----- Ortmanniana_ligamentina    
        +--+                                 
           |  +-- Lampsilis_straminea        
           +--+                              
              +-- Lampsilis_hydiana          

Boyer et al., 2011, fig. 3

iT tT bi riS oT roT roS iD
1 1 0 1 34 15 9 --
  +----------- Obliquaria_reflexa     
  |                                   
  |  +-------- Obovaria_olivaria **      
  |  |                                
  |  +-------- Lampsilis_cardium      
  |  |                                
--+  |     +-- Epioblasma_triquetra   
  |  +-----+                          
  |  |     +-- Lampsilis_siliquoidea  
  |  |                                
  |  |     +-- Ortmanniana_ligamentina
  |  +-----+                          
  |  |     +-- Lampsilis_higginsii    
  +--+                                
     |     +-- Ligumia_recta          
     +-----+                          
     |     +-- Lampsilis_teres        
     |                                
     |  +----- Potamilus_fragilis     
     +--+                             
     |  |  +-- Potamilus_ohiensis     
     |  +--+                          
     |     +-- Potamilus_alatus       
     |                                
     |  +----- Ellipsaria_lineolata   
     +--+                             
        |  +-- Truncilla_truncata     
        +--+                          
           +-- Truncilla_donaciformis 

Boyer et al., 2011, fig. 4

iT tT bi riS oT roT roS iD
1 1 0 1 37 15 9 --
  +----- Ligumia_recta       
--+                          
  |  +-- Obovaria_olivaria **   
  +--+                       
     +-- Epioblasma_triquetra

Chapman et al., 2008, fig. 2

iT tT bi riS oT roT roS iD
1 1 0 1 20 12 10 --
              +-- Venustaconcha_ellipsiformis
  +-----------+                              
  |           +-- Obovaria_olivaria **          
--+                                          
  |  +----------- Hamiota_subangulata        
  +--+                                       
     |  +-------- Lampsilis_ovata            
     +--+                                    
        |  +----- Ortmanniana_ligamentina    
        +--+                                 
           |  +-- Lampsilis_straminea        
           +--+                              
              +-- Lampsilis_hydiana          

Wu et al., 2018, fig. 2

iT tT bi riS oT roT roS iD
1 1 0 1 57 0 0 --
-- Obovaria_arkansasensis

Burlakova et al., 2019, fig. 1

iT tT bi riS oT roT roS iD
1 1 0 1 32 25 17 --
  +----- Venustaconcha_pleasii
--+                           
  |  +-- Epioblasma_rangiana  
  +--+                        
     +-- Obovaria_unicolor **    

Smith et al., 2019, fig. 2

iT tT bi riS oT roT roS iD
1 1 0 1 41 36 19 --
  +----- Venustaconcha_ellipsiformis
--+                                 
  |  +-- Obovaria_subrotunda **        
  +--+                              
     +-- Epioblasma_rangiana        

Porto-Hannes et al., 2021, fig. 3

iT tT bi riS oT roT roS iD
1 1 0 1 15 9 3 --
           +-- Obovaria_olivaria **      
  +--------+                          
  |        +-- Ortmanniana_ligamentina
  |                                   
  |        +-- Obliquaria_reflexa     
  +--------+                          
--+        +-- Lampsilis_ornata       
  |                                   
  |     +----- Lampsilis_radiata[2]   
  |  +--+                             
  |  |  |  +-- Lampsilis_siliquoidea  
  |  |  +--+                          
  +--+     +-- Lampsilis_radiata[1]   
     |                                
     |  +----- Lampsilis_virescens    
     +--+                             
        |  +-- Lampsilis_hydiana      
        +--+                          
           +-- Lampsilis_straminea    

Porto-Hannes et al., 2021, fig. 4

iT tT bi riS oT roT roS iD
1 1 0 1 11 7 2 --
  +----------- Obovaria_olivaria **      
  |                                   
--+  +-------- Ortmanniana_ligamentina
  |  |                                
  +--+  +----- Lampsilis_hydiana      
     |  |                             
     |  +----- Lampsilis_straminea    
     +--+                             
        |  +-- Lampsilis_siliquoidea  
        |  |                          
        |  +-- Lampsilis_radiata      
        +--+                          
           +-- Lampsilis[1]           
           |                          
           +-- Lampsilis[2]           

Page updated: 11:41:11 Thu 02 Feb 2023

 
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