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Page last updated
Wed 27 Sep 2023

The Freshwater Mussels (Unionoida) of the World (and other less consequential bivalves)

FM(U)otW(aolcb) is the web version of the MUSSEL Project Database. Follow the links to browse the data or use the custom Google search field. Either way, you win!

Cladomics pages are experimental — even more so than the rest of this site!

The purpose of the cladomics pages for each genus and family-group level taxon is to provide statistics and references indicating the extent to which published phylogenetic work supports the classification used here.

The Cladomics Summary provides an overview of the number of studies that have included terminal species representing this taxon as well as an indication of how frequently it has been recovered as monophyletic.

Following the summary, the available published clades representing the taxon are listed.

For more information, visit the Cladomics FARQ (frequently asked rhetorical questions) page. Abbreviations for the various statistics are explained on the FARQ!

Search the database pages:

Bivalvia | Unionoida | Unionoidea | Unionidae | Ambleminae | Lampsilini

genus Obovaria Rafinesque, 1819: cladomics
     7 valid species

Cladomics Summary

number of species = 7

analyzed in 27 cladograms in 15 publications
maximum number of terminals in an analysis = 5, mean = 2.15

Basic Monophyly

number of cladograms that test basic monophyly (i.e., more than 1 included species) = 18
maximum number of included terminals = 5, mean = 2.72
proportion of cladograms that support basic monophyly = 0.56

Pairwise Cladogram Clustering

The graphs below depict in two dimensions all pair-wise ingroup distances within the cladome. See the Cladmoics FARQ for more information.


PCoA Ordination


NJ tree

Inoue et al., 2013 [Publ 3528], fig. 3

iT tT bi riS oT roT roS iD
5 5 3 5 12 5 5 0.0

*** Top Monophyly Cladogram *** 

  +----------------- Lampsilis_ornata      
  |                                        
--+  +-------------- Leaunio_vanuxemensis  
  |  |                                     
  |  |           +-- Villosa_villosa       
  +--+-----------+                         
     |           +-- Cambarunio_iris       
     |                                     
     |  +----------- Obovaria_olivaria **     
     +--+                                  
        |  +-------- Obovaria_arkansasensis **
        +--+                               
           |  +----- Obovaria_retusa **       
           +--+                            
              |  +-- Obovaria_unicolor **     
              +--+                         
                 +-- Obovaria_subrotunda **

Campbell et al., 2005 [Publ 2660], fig. 1

iT tT bi riS oT roT roS iD
4 10 8 4 113 45 22 9.0

*** Top Sister-Group Cladogram ***

  +----------------------- Villosa_villosa               
  |                                                      
  |                    +-- Obovaria_olivaria **             
  |     +--------------+                                 
  |     |              +-- Venustaconcha_ellipsiformis[1]
  |     |                                                
  |  +--+           +----- Obovaria_unicolor **             
  |  |  |  +--------+                                    
  |  |  |  |        |  +-- Obovaria_arkansasensis **        
--+  |  |  |        +--+                                 
  |  |  +--+           +-- Obovaria_subrotunda **           
  |  |     |                                             
  |  |     |  +----------- Epioblasma_brevidens          
  |  |     +--+                                          
  |  |        |  +-------- Epioblasma_triquetra          
  |  |        +--+                                       
  |  |           |  +----- Epioblasma_walkeri[2]         
  |  |           +--+                                    
  |  |              |  +-- Epioblasma_capsaeformis       
  +--+              +--+                                 
     |                 +-- Epioblasma_walkeri[1]         
     |                                                   
     |                 +-- Ligumia_recta                 
     |        +--------+                                 
     |        |        +-- Cambarunio_iris               
     |     +--+                                          
     |     |  |  +-------- Lampsilis_teres               
     |     |  +--+                                       
     |     |     |  +----- Ortmanniana_ligamentina       
     |     |     +--+                                    
     |     |        |  +-- Lampsilis_siliquoidea         
     |     |        +--+                                 
     |     |           +-- Leaunio_vanuxemensis          
     +-----+                                             
           |           +-- Lampsilis_cardium             
           |        +--+                                 
           |        |  +-- Lampsilis_ovata               
           |  +-----+                                    
           |  |     |  +-- Ortmanniana_pectorosa         
           |  |     +--+                                 
           |  |        +-- Lampsilis_ornata              
           |  |                                          
           +--+     +----- Hamiota_australis[2]          
              |  +--+                                    
              |  |  |  +-- Hamiota_australis[1]          
              |  |  +--+                                 
              |  |     +-- Hamiota_subangulata           
              +--+                                       
                 |  +----- Hamiota_altilis[2]            
                 |  |                                    
                 |  |  +-- Hamiota_altilis[1]            
                 +--+--+                                 
                    |  +-- Hamiota_perovalis[1]          
                    |                                    
                    |  +-- Hamiota_altilis[3]            
                    +--+                                 
                       +-- Hamiota_perovalis[2]

Campbell et al., 2005 [Publ 2660], fig. 2

iT tT bi riS oT roT roS iD
4 10 6 4 111 45 22 9.0 
              +----- Obliquaria_reflexa            
  +-----------+                                    
  |           |  +-- Cyrtonaias_tampicoensis       
  |           +--+                                 
  |              +-- Glebula_rotundata             
  |                                                
  |  +-------------- Sagittunio_nasutus            
  |  |                                             
  |  +-------------- Leaunio_vanuxemensis          
  |  |                                             
  |  |           +-- Ellipsaria_lineolata          
  |  +-----------+                                 
  |  |           +-- Truncilla_truncata            
  |  |                                             
  |  |        +----- Lampsilis_teres               
  |  +--------+                                    
  |  |        |  +-- Ortmanniana_ligamentina       
  |  |        +--+                                 
  |  |           +-- Lampsilis_siliquoidea         
  |  |                                             
  |  |        +----- Villosa_villosa               
  |  +--------+                                    
--+  |        |  +-- Ligumia_recta                 
  |  |        +--+                                 
  |  |           +-- Cambarunio_iris               
  |  |                                             
  |  |     +-------- Ortmanniana_pectorosa         
  |  +-----+                                       
  |  |     |  +----- Lampsilis_ornata              
  |  |     +--+                                    
  |  |        |  +-- Lampsilis_cardium             
  |  |        +--+                                 
  |  |           +-- Lampsilis_ovata               
  |  |                                             
  |  |           +-- Potamilus_fragilis            
  |  |        +--+                                 
  |  |        |  +-- Potamilus_leptodon            
  |  +--------+                                    
  |  |        |  +-- Potamilus_alatus              
  |  |        +--+                                 
  |  |           +-- Potamilus_purpuratus          
  |  |                                             
  |  |           +-- Ptychobranchus_fasciolaris    
  +--+  +--------+                                 
     |  |        +-- Venustaconcha_ellipsiformis[1]
     |  |                                          
     +--+  +-------- Medionidus_conradicus         
     |  |  |                                       
     |  |  |     +-- Lemiox_rimosus                
     |  +--+-----+                                 
     |     |     +-- Medionidus_acutissimus        
     |     |                                       
     |     |  +----- Dromus_dromas                 
     |     +--+                                    
     |        |  +-- Cyprogenia_aberti             
     |        +--+                                 
     |           +-- Cyprogenia_stegaria           
     |                                             
     |        +----- Hamiota_australis[2]          
     |  +-----+                                    
     |  |     |  +-- Hamiota_australis[1]          
     |  |     +--+                                 
     |  |        +-- Hamiota_subangulata           
     +--+                                          
     |  |        +-- Hamiota_altilis[3]            
     |  |  +-----+                                 
     |  |  |     +-- Hamiota_perovalis[2]          
     |  +--+                                       
     |     |  +----- Hamiota_altilis[2]            
     |     +--+                                    
     |        |  +-- Hamiota_altilis[1]            
     |        +--+                                 
     |           +-- Hamiota_perovalis[1]          
     |                                             
     |  +----------- Obovaria_olivaria **             
     |  |                                          
     |  +----------- Venustaconcha_ellipsiformis[2]
     |  |                                          
     |  |     +----- Obovaria_unicolor **             
     |  +-----+                                    
     +--+     |  +-- Obovaria_arkansasensis **        
        |     +--+                                 
        |        +-- Obovaria_subrotunda **           
        |                                          
        |        +-- Epioblasma_brevidens          
        |  +-----+                                 
        |  |     +-- Epioblasma_triquetra          
        +--+                                       
           |  +----- Epioblasma_walkeri[1]         
           +--+                                    
              |  +-- Epioblasma_walkeri[2]         
              +--+                                 
                 +-- Epioblasma_capsaeformis

Zanatta & Murphy, 2006 [Publ 2672], fig. 1

iT tT bi riS oT roT roS iD
4 4 2 4 52 44 22 3.0 
  +----------------- Venustaconcha_ellipsiformis
  |                                             
  |        +-------- Obovaria_olivaria **          
--+  +-----+                                    
  |  |     |  +----- Obovaria_unicolor **          
  |  |     +--+                                 
  |  |        |  +-- Obovaria_subrotunda **        
  +--+        +--+                              
     |           +-- Obovaria_arkansasensis **     
     |                                          
     |  +----------- Epioblasma_brevidens       
     +--+                                       
        |  +-------- Epioblasma_triquetra       
        +--+                                    
           |  +----- Epioblasma_rangiana        
           +--+                                 
              |  +-- Epioblasma_walkeri         
              +--+                              
                 +-- Epioblasma_capsaeformis

Zanatta & Murphy, 2006 [Publ 2672], fig. 2

iT tT bi riS oT roT roS iD
4 10 6 4 53 44 22 9.0 
  +-------------------- Ortmanniana_ligamentina    
  |                                                
--+  +----------------- Lampsilis_siliquoidea      
  |  |                                             
  |  |  +-------------- Obovaria_olivaria **          
  +--+  |                                          
     |  +-------------- Venustaconcha_ellipsiformis
     |  |                                          
     |  |        +----- Obovaria_unicolor **          
     +--+--------+                                 
        |        |  +-- Obovaria_subrotunda **        
        |        +--+                              
        |           +-- Obovaria_arkansasensis **     
        |                                          
        |  +----------- Epioblasma_brevidens       
        +--+                                       
           |  +-------- Epioblasma_triquetra       
           +--+                                    
              |  +----- Epioblasma_rangiana        
              +--+                                 
                 |  +-- Epioblasma_walkeri         
                 +--+                              
                    +-- Epioblasma_capsaeformis

Porto-Hannes et al., 2019 [Publ 3969], fig. 2

iT tT bi riS oT roT roS iD
3 3 1 3 17 16 5 4.0 
                    +-- Cambarunio_iris        
  +-----------------+                          
  |                 +-- Leaunio_lienosus       
  |                                            
  |        +----------- Lampsilis_teres        
  |        |                                   
  +--------+  +-------- Ortmanniana_ligamentina
  |        |  |                                
  |        +--+     +-- Lampsilis_hydiana      
  |           |  +--+                          
--+           |  |  +-- Lampsilis_virescens    
  |           +--+                             
  |              |  +-- Lampsilis_siliquoidea  
  |              +--+                          
  |                 +-- Lampsilis_radiata      
  |                                            
  |              +----- Obovaria_arkansasensis ** 
  |  +-----------+                             
  |  |           |  +-- Obovaria_olivaria **      
  |  |           +--+                          
  +--+              +-- Obovaria_subrotunda **    
     |                                         
     |  +-------------- Lampsilis_bracteata    
     |  |                                      
     +--+  +----------- Lampsilis_ornata       
        |  |                                   
        +--+  +-------- Lampsilis_ovata        
           |  |                                
           +--+  +----- Lampsilis_cardium[3]   
              |  |                             
              +--+  +-- Lampsilis_satura       
                 |  |                          
                 +--+-- Lampsilis_cardium[1]   
                    |                          
                    +-- Lampsilis_cardium[2]

Porto-Hannes et al., 2019 [Publ 3969], fig. 3

iT tT bi riS oT roT roS iD
3 3 1 3 17 16 5 4.0 
  +----------------- Lampsilis_teres        
  |                                         
  |              +-- Leaunio_lienosus       
  +--------------+                          
  |              +-- Cambarunio_iris        
  |                                         
  |           +----- Obovaria_arkansasensis ** 
  +-----------+                             
  |           |  +-- Obovaria_olivaria **      
  |           +--+                          
  |              +-- Obovaria_subrotunda **    
  |                                         
--+        +-------- Ortmanniana_ligamentina
  |        |                                
  +--------+     +-- Lampsilis_hydiana      
  |        |  +--+                          
  |        |  |  +-- Lampsilis_virescens    
  |        +--+                             
  |           |  +-- Lampsilis_siliquoidea  
  |           +--+                          
  |              +-- Lampsilis_radiata      
  |                                         
  |  +-------------- Lampsilis_bracteata    
  |  |                                      
  +--+  +----------- Lampsilis_ornata       
     |  |                                   
     +--+  +-------- Lampsilis_ovata        
        |  |                                
        +--+  +----- Lampsilis_cardium[3]   
           |  |                             
           +--+  +-- Lampsilis_satura       
              |  |                          
              +--+-- Lampsilis_cardium[1]   
                 |                          
                 +-- Lampsilis_cardium[2]

Lydeard et al., 2000 [Publ 2654], fig. 2a

iT tT bi riS oT roT roS iD
2 2 0 2 15 1 1 -- 
  +----- Lampsilis_teres  
--+                       
  |  +-- Obovaria_unicolor **
  +--+                    
     +-- Obovaria_olivaria **

Lydeard et al., 2000 [Publ 2654], fig. 2b

iT tT bi riS oT roT roS iD
2 2 0 2 15 1 1 -- 
  +----- Lampsilis_teres  
--+                       
  |  +-- Obovaria_unicolor **
  +--+                    
     +-- Obovaria_olivaria **

Lydeard et al., 2000 [Publ 2654], fig. 2c

iT tT bi riS oT roT roS iD
2 3 0 2 14 1 1 4.0 
  +----- Obovaria_unicolor **
--+                       
  |  +-- Obovaria_olivaria **
  +--+                    
     +-- Lampsilis_teres

Lydeard et al., 2000 [Publ 2654], fig. 2d

iT tT bi riS oT roT roS iD
2 2 0 2 14 1 1 -- 
  +----- Lampsilis_teres  
--+                       
  |  +-- Obovaria_unicolor **
  +--+                    
     +-- Obovaria_olivaria **

Campbell & Lydeard, 2012 [Publ 3308], fig. 1

iT tT bi riS oT roT roS iD
2 2 0 2 41 3 3 -- 
  +-------- Lemiox_rimosus   
--+                          
  |  +----- Lampsilis_ovata  
  +--+                       
     |  +-- Obovaria_unicolor **
     +--+                    
        +-- Obovaria_olivaria **

Campbell & Lydeard, 2012 [Publ 3308], fig. 2

iT tT bi riS oT roT roS iD
2 2 0 2 41 3 3 -- 
  +-------- Lemiox_rimosus   
--+                          
  |  +----- Lampsilis_ovata  
  +--+                       
     |  +-- Obovaria_unicolor **
     +--+                    
        +-- Obovaria_olivaria **

Campbell & Lydeard, 2012 [Publ 3308], fig. 3

iT tT bi riS oT roT roS iD
2 4 0 2 49 3 3 5.0 
  +-------- Obovaria_unicolor **
--+                          
  |  +----- Obovaria_olivaria **
  +--+                       
     |  +-- Lampsilis_ovata  
     +--+                    
        +-- Lemiox_rimosus

Campbell & Lydeard, 2012 [Publ 3308], fig. 4

iT tT bi riS oT roT roS iD
2 2 0 2 41 3 3 -- 
  +-------- Lemiox_rimosus   
--+                          
  |  +----- Lampsilis_ovata  
  +--+                       
     |  +-- Obovaria_unicolor **
     +--+                    
        +-- Obovaria_olivaria **

Campbell & Lydeard, 2012 [Publ 3309], fig. 1-2

iT tT bi riS oT roT roS iD
2 5 0 2 72 13 10 6.0 
              +-- Lampsilis_ovata            
  +-----------+                              
  |           +-- Obliquaria_reflexa         
--+                                          
  |  +----------- Lampsilis_virescens        
  |  |                                       
  +--+        +-- Epioblasma_rangiana        
     |  +-----+                              
     |  |     +-- Obovaria_arkansasensis **     
     +--+                                    
        |  +----- Obovaria_olivaria **          
        +--+                                 
           |  +-- Venustaconcha_ellipsiformis
           +--+                              
              +-- Venustaconcha_pleasii

Campbell & Lydeard, 2012 [Publ 3309], fig. 3

iT tT bi riS oT roT roS iD
2 5 0 2 72 13 10 6.0 
  +-------------- Lampsilis_ovata            
  |                                          
--+  +----------- Lampsilis_virescens        
  |  |                                       
  +--+        +-- Epioblasma_rangiana        
     |  +-----+                              
     |  |     +-- Obovaria_arkansasensis **     
     +--+                                    
        |  +----- Obovaria_olivaria **          
        +--+                                 
           |  +-- Venustaconcha_ellipsiformis
           +--+                              
              +-- Venustaconcha_pleasii

Inoue et al., 2018 [Publ 3858], fig. 1, 3a-c

iT tT bi riS oT roT roS iD
2 12 0 2 203 9 7 13.0 
  +-- Dromus_dromas         
  |                         
  +-- Epioblasma_rangiana   
  |                         
  +-- Glebula_rotundata     
  |                         
  +-- Lampsilis_ovata       
  |                         
  +-- Lampsilis_virescens   
  |                         
  +-- Lemiox_rimosus        
--+                         
  +-- Obliquaria_reflexa    
  |                         
  +-- Obovaria_olivaria **     
  |                         
  +-- Obovaria_unicolor **     
  |                         
  +-- Plectomerus_dombeyanus
  |                         
  +-- Potamilus_alatus      
  |                         
  +-- Potamilus_capax

Chapman et al., 2008 [Publ 2667], fig. 1

iT tT bi riS oT roT roS iD
1 1 0 1 20 12 10 -- 
              +-- Venustaconcha_ellipsiformis
  +-----------+                              
  |           +-- Obovaria_olivaria **          
--+                                          
  |  +----------- Hamiota_subangulata        
  +--+                                       
     |  +-------- Lampsilis_ovata            
     +--+                                    
        |  +----- Ortmanniana_ligamentina    
        +--+                                 
           |  +-- Lampsilis_straminea        
           +--+                              
              +-- Lampsilis_hydiana

Boyer et al., 2011 [Publ 2718], fig. 3

iT tT bi riS oT roT roS iD
1 1 0 1 34 15 9 -- 
  +----------- Obliquaria_reflexa     
  |                                   
  |  +-------- Obovaria_olivaria **      
  |  |                                
  |  +-------- Lampsilis_cardium      
  |  |                                
--+  |     +-- Epioblasma_triquetra   
  |  +-----+                          
  |  |     +-- Lampsilis_siliquoidea  
  |  |                                
  |  |     +-- Ortmanniana_ligamentina
  |  +-----+                          
  |  |     +-- Lampsilis_higginsii    
  +--+                                
     |     +-- Ligumia_recta          
     +-----+                          
     |     +-- Lampsilis_teres        
     |                                
     |  +----- Potamilus_fragilis     
     +--+                             
     |  |  +-- Potamilus_ohiensis     
     |  +--+                          
     |     +-- Potamilus_alatus       
     |                                
     |  +----- Ellipsaria_lineolata   
     +--+                             
        |  +-- Truncilla_truncata     
        +--+                          
           +-- Truncilla_donaciformis

Boyer et al., 2011 [Publ 2718], fig. 4

iT tT bi riS oT roT roS iD
1 1 0 1 37 15 9 -- 
  +----- Ligumia_recta       
--+                          
  |  +-- Obovaria_olivaria **   
  +--+                       
     +-- Epioblasma_triquetra

Chapman et al., 2008 [Publ 2744], fig. 2

iT tT bi riS oT roT roS iD
1 1 0 1 20 12 10 -- 
              +-- Venustaconcha_ellipsiformis
  +-----------+                              
  |           +-- Obovaria_olivaria **          
--+                                          
  |  +----------- Hamiota_subangulata        
  +--+                                       
     |  +-------- Lampsilis_ovata            
     +--+                                    
        |  +----- Ortmanniana_ligamentina    
        +--+                                 
           |  +-- Lampsilis_straminea        
           +--+                              
              +-- Lampsilis_hydiana

Wu et al., 2018 [Publ 3836], fig. 2

iT tT bi riS oT roT roS iD
1 1 0 1 52 0 0 -- 
-- Obovaria_arkansasensis

Burlakova et al., 2019 [Publ 3899], fig. 1

iT tT bi riS oT roT roS iD
1 1 0 1 32 25 17 -- 
  +----- Venustaconcha_pleasii
--+                           
  |  +-- Epioblasma_rangiana  
  +--+                        
     +-- Obovaria_unicolor **

Smith et al., 2019 [Publ 3951], fig. 2

iT tT bi riS oT roT roS iD
1 1 0 1 41 36 19 -- 
  +----- Venustaconcha_ellipsiformis
--+                                 
  |  +-- Obovaria_subrotunda **        
  +--+                              
     +-- Epioblasma_rangiana

Porto-Hannes et al., 2021 [Publ 4227], fig. 3

iT tT bi riS oT roT roS iD
1 1 0 1 15 9 3 -- 
           +-- Obovaria_olivaria **      
  +--------+                          
  |        +-- Ortmanniana_ligamentina
  |                                   
  |        +-- Obliquaria_reflexa     
  +--------+                          
--+        +-- Lampsilis_ornata       
  |                                   
  |     +----- Lampsilis_radiata[2]   
  |  +--+                             
  |  |  |  +-- Lampsilis_siliquoidea  
  |  |  +--+                          
  +--+     +-- Lampsilis_radiata[1]   
     |                                
     |  +----- Lampsilis_virescens    
     +--+                             
        |  +-- Lampsilis_hydiana      
        +--+                          
           +-- Lampsilis_straminea

Porto-Hannes et al., 2021 [Publ 4227], fig. 4

iT tT bi riS oT roT roS iD
1 1 0 1 11 7 2 -- 
  +----------- Obovaria_olivaria **      
  |                                   
--+  +-------- Ortmanniana_ligamentina
  |  |                                
  +--+  +----- Lampsilis_hydiana      
     |  |                             
     |  +----- Lampsilis_straminea    
     +--+                             
        |  +-- Lampsilis_siliquoidea  
        |  |                          
        |  +-- Lampsilis_radiata      
        +--+                          
           +-- Lampsilis[1]           
           |                          
           +-- Lampsilis[2]

Page updated: 10:54:23 Wed 27 Sep 2023
 
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