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Page last updated
Mon 06 May 2024

The Freshwater Mussels (Unionoida) of the World (and other less consequential bivalves)

FM(U)otW(aolcb) is the web version of the MUSSEL Project Database. Follow the links to browse the data or use the custom Google search field. Either way, you win!

Cladomics pages are experimental — even more so than the rest of this site!

The purpose of the cladomics pages for each genus and family-group level taxon is to provide statistics and references indicating the extent to which published phylogenetic work supports the classification used here.

The Cladomics Summary provides an overview of the number of studies that have included terminal species representing this taxon as well as an indication of how frequently it has been recovered as monophyletic.

Following the summary, the available published clades representing the taxon are listed.

For more information, visit the Cladomics FARQ (frequently asked rhetorical questions) page. Abbreviations for the various statistics are explained on the FARQ!

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Bivalvia | Unionoida | Unionoidea | Unionidae | Ambleminae | Quadrulini

genus Megalonaias Utterback, 1915: cladomics
     2 valid species

Cladomics Summary

number of species = 2

analyzed in 29 cladograms in 19 publications
maximum number of terminals in an analysis = 2, mean = 1.03

Basic Monophyly

number of cladograms that test basic monophyly (i.e., more than 1 included species) = 1
maximum number of included terminals = 2, mean = 2.0
proportion of cladograms that support basic monophyly = 1.0


Pfeiffer et al., 2018 [Publ 3835], fig. 2

iT tT bi riS oT roT roS iD
2 2 0 2 2 2 2 --

*** Top Monophyly Cladogram ***

  +-------- Quadrula_quadrula      
--+                                
  |  +----- Uniomerus_declivis     
  +--+                             
     |  +-- Megalonaias_nervosa **    
     +--+                          
        +-- Megalonaias_nickliniana **

Neemuchwala et al., 2023 [Publ 4292], fig. 2

iT tT bi riS oT roT roS iD
1 1 0 1 26 21 2 --

*** Top Sister-Group Cladogram ***

  +-------------------------- Megalonaias_nervosa **   
  |                                                 
  |                 +-------- Uniomerus_carolinianus
  |  +--------------+                               
--+  |              |  +----- Uniomerus_columbensis 
  |  |              +--+                            
  |  |                 |  +-- Uniomerus_tetralasmus 
  |  |                 +--+                         
  |  |                    +-- Uniomerus_declivis    
  +--+                                              
     |              +-------- Theliderma_cylindrica 
     |  +-----------+                               
     |  |           |  +----- Theliderma_metanevra  
     |  |           +--+                            
     |  |              |  +-- Theliderma_sparsa     
     |  |              +--+                         
     +--+                 +-- Theliderma_johnsoni   
        |                                           
        |  +----------------- Cyclonaias_tuberculata
        |  |                                        
        |  |        +-------- Quadrula_nobilis      
        |  |  +-----+                               
        +--+  |     |  +----- Quadrula_verrucosa    
           |  |     +--+                            
           |  |        |  +-- Quadrula_fragosa      
           |  |        +--+                         
           |  |           +-- Quadrula_quadrula     
           +--+                                     
              |        +----- Pustulosa_nodulata    
              |  +-----+                            
              |  |     |  +-- Pustulosa_necki       
              |  |     +--+                         
              |  |        +-- Pustulosa_petrina     
              +--+                                  
                 |        +-- Pustulosa_kieneriana  
                 |  +-----+                         
                 |  |     +-- Pustulosa_pustulosa   
                 +--+                               
                    |  +----- Pustulosa_succissa    
                    +--+                            
                       |  +-- Pustulosa_infucata    
                       +--+                         
                          +-- Pustulosa_kleiniana   

Lydeard et al., 1996 [Publ 2656], fig. 3

iT tT bi riS oT roT roS iD
1 1 0 1 26 3 2 --
  +----- Megalonaias_nervosa ** 
--+                          
  |  +-- Quadrula_quadrula   
  +--+                       
     +-- Theliderma_metanevra

Lydeard et al., 1996 [Publ 2656], fig. 4

iT tT bi riS oT roT roS iD
1 1 0 1 27 3 2 --
  +----- Megalonaias_nervosa ** 
--+                          
  |  +-- Quadrula_quadrula   
  +--+                       
     +-- Theliderma_metanevra

Huang et al., 2002 [Publ 2657], fig. 1

iT tT bi riS oT roT roS iD
1 1 0 1 33 1 1 --
-- Megalonaias_nervosa

Huang et al., 2002 [Publ 2657], fig. 2

iT tT bi riS oT roT roS iD
1 1 0 1 33 1 1 --
-- Megalonaias_nervosa

Serb et al., 2003 [Publ 2659], fig. 2

iT tT bi riS oT roT roS iD
1 1 0 1 28 13 1 --
  +----------------- Megalonaias_nervosa **  
  |                                       
  |        +-------- Theliderma_cylindrica
  |  +-----+                              
--+  |     |  +----- Theliderma_intermedia
  |  |     +--+                           
  |  |        |  +-- Theliderma_sparsa    
  |  |        +--+                        
  |  |           +-- Theliderma_metanevra 
  +--+                                    
     |        +----- Quadrula_verrucosa   
     |  +-----+                           
     |  |     |  +-- Quadrula_quadrula    
     |  |     +--+                        
     |  |        +-- Quadrula_nobilis     
     +--+                                 
        |  +-------- Pustulosa_kieneriana 
        |  |                              
        |  +-------- Pustulosa_infucata   
        +--+                              
           |     +-- Pustulosa_pustulosa  
           |  +--+                        
           |  |  +-- Pustulosa_succissa   
           +--+                           
              |  +-- Pustulosa_nodulata   
              +--+                        
                 +-- Pustulosa_petrina    

Serb et al., 2003 [Publ 2659], fig. 3

iT tT bi riS oT roT roS iD
1 1 0 1 28 13 1 --
  +----------------------- Megalonaias_nervosa **  
  |                                             
  |                    +-- Quadrula_verrucosa   
--+  +-----------------+                        
  |  |                 +-- Quadrula_nobilis     
  |  |                                          
  +--+  +----------------- Quadrula_quadrula    
     |  |                                       
     |  |        +-------- Theliderma_cylindrica
     +--+  +-----+                              
        |  |     |  +----- Theliderma_sparsa    
        |  |     +--+                           
        |  |        |  +-- Theliderma_intermedia
        +--+        +--+                        
           |           +-- Theliderma_metanevra 
           |                                    
           |  +----------- Pustulosa_kieneriana 
           |  |                                 
           +--+  +-------- Pustulosa_infucata   
              |  |                              
              +--+     +-- Pustulosa_succissa   
                 |  +--+                        
                 |  |  +-- Pustulosa_pustulosa  
                 +--+                           
                    |  +-- Pustulosa_nodulata   
                    +--+                        
                       +-- Pustulosa_petrina    

Campbell et al., 2005 [Publ 2660], fig. 1

iT tT bi riS oT roT roS iD
1 1 0 1 115 10 2 --
              +-- Megalonaias_nervosa **   
  +-----------+                         
  |           +-- Uniomerus_declivis    
  |                                     
--+  +----------- Theliderma_metanevra  
  |  |                                  
  |  |     +----- Quadrula_verrucosa    
  +--+  +--+                            
     |  |  |  +-- Quadrula_quadrula     
     |  |  +--+                         
     |  |     +-- Quadrula_nobilis      
     +--+                               
        |  +----- Pustulosa_kieneriana  
        |  |                            
        |  |  +-- Cyclonaias_tuberculata
        +--+--+                         
           |  +-- Pustulosa_succissa    
           |                            
           |  +-- Pustulosa_infucata    
           +--+                         
              +-- Pustulosa_kleiniana   

Campbell et al., 2005 [Publ 2660], fig. 2

iT tT bi riS oT roT roS iD
1 1 0 1 113 10 2 --
  +-------------------- Megalonaias_nervosa **   
  |                                           
--+  +----------------- Uniomerus_declivis    
  |  |                                        
  +--+  +-------------- Theliderma_metanevra  
     |  |                                     
     +--+  +----------- Quadrula_quadrula     
        |  |                                  
        |  |        +-- Quadrula_nobilis      
        +--+  +-----+                         
           |  |     +-- Quadrula_verrucosa    
           |  |                               
           +--+  +----- Pustulosa_kieneriana  
              |  |                            
              |  |  +-- Cyclonaias_tuberculata
              +--+--+                         
                 |  +-- Pustulosa_succissa    
                 |                            
                 |  +-- Pustulosa_infucata    
                 +--+                         
                    +-- Pustulosa_kleiniana   

Krebs et al., 2003 [Publ 2691], fig. 3

iT tT bi riS oT roT roS iD
1 1 0 1 33 4 2 --
  +-------- Megalonaias_nervosa ** 
--+                             
  |  +----- Quadrula_quadrula[2]
  +--+                          
     |  +-- Quadrula_quadrula[1]
     +--+                       
        +-- Theliderma_metanevra

Boyer et al., 2011 [Publ 2718], fig. 3

iT tT bi riS oT roT roS iD
1 1 0 1 34 6 1 --
  +----------------- Megalonaias_nervosa **   
--+                                        
  |  +-------------- Quadrula_quadrula     
  +--+                                     
     |  +----------- Quadrula_verrucosa    
     +--+                                  
        |  +-------- Theliderma_metanevra  
        +--+                               
           |  +----- Pustulosa_pustulosa   
           +--+                            
              |  +-- Cyclonaias_tuberculata
              +--+                         
                 +-- Pustulosa_nodulata    

Boyer et al., 2011 [Publ 2718], fig. 4

iT tT bi riS oT roT roS iD
1 1 0 1 37 6 1 --
-- Megalonaias_nervosa

Rosenberg et al., 1994 [Publ 2719], fig. 3

iT tT bi riS oT roT roS iD
1 1 0 1 18 3 2 --
-- Megalonaias_nervosa

Rosenberg et al., 1997 [Publ 2720], fig. 2A

iT tT bi riS oT roT roS iD
1 1 0 1 22 3 2 --
-- Megalonaias_nervosa

Rosenberg et al., 1997 [Publ 2720], fig. 2B

iT tT bi riS oT roT roS iD
1 1 0 1 22 3 2 --
-- Megalonaias_nervosa

Breton et al., 2011 [Publ 3376], fig. 3

iT tT bi riS oT roT roS iD
1 1 0 1 38 3 1 --
-- Megalonaias_nervosa

Mulvey et al., 1997 [Publ 3390], fig. 1, 3

iT tT bi riS oT roT roS iD
1 1 0 1 4 0 0 --
-- Megalonaias_nervosa

Szumowski et al., 2012 [Publ 3569], fig. 5

iT tT bi riS oT roT roS iD
1 1 0 1 14 14 1 --
  +----------------- Megalonaias_nervosa **  
  |                                       
  |        +-------- Theliderma_cylindrica
  +--------+                              
  |        |  +----- Theliderma_intermedia
  |        +--+                           
  |           |  +-- Theliderma_metanevra 
--+           +--+                        
  |              +-- Theliderma_sparsa    
  |                                       
  |        +-------- Quadrula_quadrula[1] 
  |  +-----+                              
  |  |     |  +----- Quadrula_verrucosa   
  |  |     +--+                           
  |  |        |  +-- Quadrula_nobilis     
  +--+        +--+                        
     |           +-- Quadrula_quadrula[2] 
     |                                    
     |  +----------- Pustulosa_kieneriana 
     |  |                                 
     +--+  +-------- Pustulosa_infucata   
        |  |                              
        +--+     +-- Pustulosa_pustulosa  
           |  +--+                        
           |  |  +-- Pustulosa_succissa   
           +--+                           
              |  +-- Pustulosa_nodulata   
              +--+                        
                 +-- Pustulosa_petrina    

Huang et al., 2019 [Publ 3843], fig. 2

iT tT bi riS oT roT roS iD
1 1 0 1 203 3 2 --
     +-- Megalonaias_nervosa **   
  +--+                         
  |  +-- Uniomerus_carolinianus
--+                            
  |  +-- Quadrula_quadrula     
  +--+                         
     +-- Quadrula_verrucosa    

Huang et al., 2019 [Publ 3843], fig. S3

iT tT bi riS oT roT roS iD
1 1 0 1 200 3 2 --
     +-- Megalonaias_nervosa **   
  +--+                         
  |  +-- Uniomerus_carolinianus
--+                            
  |  +-- Quadrula_quadrula     
  +--+                         
     +-- Quadrula_verrucosa    

Huang et al., 2019 [Publ 3843], fig. S4

iT tT bi riS oT roT roS iD
1 1 0 1 201 3 2 --
-- Megalonaias_nervosa

Johnson et al., 2018 [Publ 3844], fig. 2

iT tT bi riS oT roT roS iD
1 1 0 1 15 13 2 --
  +-------------------------- Megalonaias_nervosa **   
  |                                                 
--+  +----------------------- Uniomerus_tetralasmus 
  |  |                                              
  +--+  +-------------------- Theliderma_metanevra  
     |  |                                           
     |  |                 +-- Quadrula_quadrula     
     +--+  +--------------+                         
        |  |              +-- Quadrula_verrucosa    
        |  |                                        
        |  |           +----- Pustulosa_kieneriana  
        +--+  +--------+                            
           |  |        |  +-- Pustulosa_infucata    
           |  |        +--+                         
           |  |           +-- Pustulosa_kleiniana   
           +--+                                     
              |           +-- Pustulosa_pustulosa   
              |  +--------+                         
              |  |        +-- Pustulosa_succissa    
              +--+                                  
                 |  +-------- Cyclonaias_tuberculata
                 +--+                               
                    |  +----- Pustulosa_necki       
                    +--+                            
                       |  +-- Pustulosa_nodulata    
                       +--+                         
                          +-- Pustulosa_petrina     

Chase et al., 2018 [Publ 3847], fig. 4

iT tT bi riS oT roT roS iD
1 1 0 1 38 2 1 --
  +----- Megalonaias_nervosa **   
--+                            
  |  +-- Cyclonaias_tuberculata
  +--+                         
     +-- Pustulosa_pustulosa   

Pfeiffer et al., 2019 [Publ 3866], fig. 1

iT tT bi riS oT roT roS iD
1 1 0 1 38 4 2 --
  +----------- Megalonaias_nervosa **   
--+                                  
  |  +-------- Uniomerus_declivis    
  +--+                               
     |  +----- Quadrula_verrucosa    
     +--+                            
        |  +-- Cyclonaias_tuberculata
        +--+                         
           +-- Quadrula_quadrula     

Lopes-Lima et al., 2019 [Publ 3901], fig. 2

iT tT bi riS oT roT roS iD
1 1 0 1 27 17 2 --
                       +----- Megalonaias_nervosa **   
                       |                            
  +--------------------+  +-- Uniomerus_carolinianus
  |                    |  |                         
  |                    +--+-- Uniomerus_declivis    
  |                       |                         
--+                       +-- Uniomerus_tetralasmus 
  |                                                 
  |  +----------------------- Quadrula_quadrula     
  |  |                                              
  |  |                    +-- Quadrula_verrucosa    
  +--+  +-----------------+                         
     |  |                 +-- Quadrula_nobilis      
     |  |                                           
     +--+                 +-- Theliderma_metanevra  
        |  +--------------+                         
        |  |              +-- Theliderma_johnsoni   
        |  |                                        
        +--+           +----- Pustulosa_kieneriana  
           |  +--------+                            
           |  |        |  +-- Pustulosa_infucata    
           |  |        +--+                         
           +--+           +-- Pustulosa_kleiniana   
              |                                     
              |  +----------- Cyclonaias_tuberculata
              |  |                                  
              +--+        +-- Pustulosa_pustulosa   
                 |  +-----+                         
                 |  |     +-- Pustulosa_succissa    
                 +--+                               
                    |  +----- Pustulosa_nodulata    
                    +--+                            
                       |  +-- Pustulosa_necki       
                       +--+                         
                          +-- Pustulosa_petrina     

Lopes-Lima et al., 2019 [Publ 3901], fig. 3

iT tT bi riS oT roT roS iD
1 1 0 1 28 18 2 --
-- Megalonaias_nervosa

Lopes-Lima et al., 2019 [Publ 3901], fig. 4

iT tT bi riS oT roT roS iD
1 1 0 1 25 15 2 --
  +----------------------------- Megalonaias_nervosa **   
  |                                                    
--+  +-------------------------- Uniomerus_tetralasmus 
  |  |                                                 
  |  |                       +-- Theliderma_metanevra  
  +--+  +--------------------+                         
     |  |                    +-- Theliderma_johnsoni   
     |  |                                              
     +--+  +-------------------- Quadrula_quadrula     
        |  |                                           
        |  |                 +-- Quadrula_verrucosa    
        +--+  +--------------+                         
           |  |              +-- Quadrula_nobilis      
           |  |                                        
           |  |           +----- Pustulosa_kieneriana  
           +--+  +--------+                            
              |  |        |  +-- Pustulosa_infucata    
              |  |        +--+                         
              |  |           +-- Pustulosa_kleiniana   
              +--+                                     
                 |           +-- Pustulosa_pustulosa   
                 |  +--------+                         
                 |  |        +-- Pustulosa_succissa    
                 +--+                                  
                    |  +-------- Cyclonaias_tuberculata
                    +--+                               
                       |  +----- Pustulosa_necki       
                       +--+                            
                          |  +-- Pustulosa_petrina     
                          +--+                         
                             +-- Pustulosa_nodulata    

Smith, 2021 [Publ 4331], fig. 1

iT tT bi riS oT roT roS iD
1 1 0 1 19 1 1 --
  +-- Uniomerus_tetralasmus
--+                        
  +-- Megalonaias_nervosa **  

Page updated: 08:52:31 Mon 06 May 2024

 
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