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Page last updated
Tue 04 Jan 2022

The Freshwater Mussels (Unionoida) of the World (and other less consequential bivalves)

FM(U)otW(aolcb) is the web version of the MUSSEL Project Database. Follow the links to browse the data or use the custom Google search field. Either way, you win!

Cladomics pages are experimental — even more so than the rest of this site!

The purpose of the cladomics pages for each genus and family-group level taxon is to provide statistics and references indicating the extent to which published phylogenetic work supports the classification used here.

The Cladomics Summary provides an overview of the number of studies that have included terminal species representing this taxon as well as an indication of how frequently it has been recovered as monophyletic.

Following the summary, the available published clades representing the taxon are listed.

For more information, visit the Cladomics FARQ (frequently asked rhetorical questions) page.

Search the database pages:

Bivalvia | Unionoida | Unionoidea | Unionidae | Unioninae | Unionini

genus Unio Philipsson in Retzius, 1788: cladomics
     40 valid species

Cladomics Summary

number of species: 24
number next lowest-taxon (species) (i.e., direct-child taxa): 24

analyzed in 167 cladogram(s) in 100 publications
maximum number of terminal species in an analysis: 16, mean: 2.93
maximum number of direct-child taxa in an analysis: 16, mean: 2.93

basic monophyly

number of cladograms that test basic monophyly (i.e., more than 1 included species): 115
maximum number of included species: 16, mean: 3.8
maximum number of direct-child taxa: 16, mean: 3.8
proportion of cladograms that support basic monophyly: 0.9

rigorous monophyly

number of cladograms that test rigorous monophyly: 53
(Rigorous monophyly refers to cladograms with terminal species representing more than one direct-child taxon and at least one rigorous, direct-child outgroup.)
maximum number of included species: 14, mean: 3.77
maximum number of direct-child taxa: 14, mean: 3.77
proportion of cladograms that support rigorous monophyly: 0.89

top monophyly publications

Araujo et al., 2018, Froufe et al., 2016, Araujo et al., 2009, Lopes-Lima et al., 2020, Huang et al., 2019

Lopes-Lima et al., 2021 ** Top Monophyly Test Publication **

fig. 1: [clade_id: 19846] Unio
spp. terminals: 16, direct-child terminals: 16
rigorous outgroups.: 0, direct-child rigorous outgroups: 0
(U_tumidus, ((U_damascensis, (U_sesirmensis, (U_bruguierianus, U_crassus))), ((U_delicatus, U_terminalis), ((U_tigridis, U_hueti), (U_ravoisieri, ((U_pictorum, U_eucirrus), ((U_elongatulus, U_mancus), (U_foucauldianus, U_delphinus))))))))


	  +----------------------- U_tumidus 1       
	  |                                          
	  |              +-------- U_damascensis 2   
	  |  +-----------+                           
	--+  |           |  +----- U_sesirmensis 3   
	  |  |           +--+                        
	  |  |              |  +-- U_bruguierianus 4 
	  |  |              +--+                     
	  +--+                 +-- U_crassus 5       
	     |                                       
	     |                 +-- U_delicatus 6     
	     |  +--------------+                     
	     |  |              +-- U_terminalis 7    
	     |  |                                    
	     +--+              +-- U_hueti 9         
	        |  +-----------+                     
	        |  |           +-- U_tigridis 8      
	        |  |                                 
	        +--+  +----------- U_ravoisieri 10   
	           |  |                              
	           |  |        +-- U_eucirrus 12     
	           +--+  +-----+                     
	              |  |     +-- U_pictorum 11     
	              |  |                           
	              +--+     +-- U_elongatulus 13  
	                 |  +--+                     
	                 |  |  +-- U_mancus 14       
	                 +--+                        
	                    |  +-- U_delphinus 16    
	                    +--+                     
	                       +-- U_foucauldianus 15
  

fig. 2: [clade_id: 19883] Unio
spp. terminals: 10, direct-child terminals: 10
rigorous outgroups.: 0, direct-child rigorous outgroups: 0
(U_tumidus, (U_bruguierianus, ((U_delicatus, U_terminalis), ((U_tigridis, U_hueti), ((U_pictorum, U_eucirrus), (U_elongatulus, U_mancus))))))


	  +----------------- U_tumidus 1      
	  |                                   
	--+  +-------------- U_bruguierianus 2
	  |  |                                
	  |  |           +-- U_delicatus 3    
	  +--+  +--------+                    
	     |  |        +-- U_terminalis 4   
	     |  |                             
	     +--+        +-- U_hueti 6        
	        |  +-----+                    
	        |  |     +-- U_tigridis 5     
	        |  |                          
	        +--+     +-- U_eucirrus 8     
	           |  +--+                    
	           |  |  +-- U_pictorum 7     
	           +--+                       
	              |  +-- U_elongatulus 9  
	              +--+                    
	                 +-- U_mancus 10      
  

fig. 6: [clade_id: 19896] Unio
spp. terminals: 7, direct-child terminals: 7
rigorous outgroups.: 0, direct-child rigorous outgroups: 0
(U_tumidus, (U_pictorum, (U_sesirmensis, (U_damascensis, (U_courtillieri, (U_bruguierianus, U_crassus))))))


	  +----------------- U_tumidus 1      
	--+                                   
	  |  +-------------- U_pictorum 2     
	  +--+                                
	     |  +----------- U_sesirmensis 3  
	     +--+                             
	        |  +-------- U_damascensis 4  
	        +--+                          
	           |  +----- U_courtillieri 5 
	           +--+                       
	              |  +-- U_bruguierianus 6
	              +--+                    
	                 +-- U_crassus 7      
  

Lopes-Lima et al., 2020 ** Top Sister Group Test Publication **

fig. 2-4: [clade_id: 16501] Unio
spp. terminals: 4, direct-child terminals: 4
rigorous outgroups.: 25, direct-child rigorous outgroups: 8
(U_gibbus, U_tumidus, (U_crassus, U_pictorum))


	  +----- U_gibbus 1  
	  |                  
	--+----- U_tumidus 2 
	  |                  
	  |  +-- U_crassus 3 
	  +--+               
	     +-- U_pictorum 4
  

Araujo et al., 2018

fig. 1: [clade_id: 10074] Unio
spp. terminals: 14, direct-child terminals: 14
rigorous outgroups.: 1, direct-child rigorous outgroups: 1
((U_durieui, U_gibbus), (U_tumidus, ((U_tumidiformis, (U_bruguierianus, (U_crassus, U_ionicus))), ((U_delphinus, U_foucauldianus), (U_pictorum, U_elongatulus, U_mancus, U_ravoisieri, U_tigridis)))))


	                 +-- U_durieui 1      
	  +--------------+                    
	  |              +-- U_gibbus 2       
	  |                                   
	--+  +-------------- U_tumidus 3      
	  |  |                                
	  |  |     +-------- U_tumidiformis 4 
	  |  |  +--+                          
	  +--+  |  |  +----- U_bruguierianus 5
	     |  |  +--+                       
	     |  |     |  +-- U_crassus 6      
	     |  |     +--+                    
	     +--+        +-- U_ionicus 7      
	        |                             
	        |        +-- U_delphinus 8    
	        |     +--+                    
	        |     |  +-- U_foucauldianus 9
	        |     |                       
	        +-----+  +-- U_elongatulus 11 
	              |  |                    
	              |  +-- U_mancus 12      
	              |  |                    
	              +--+-- U_pictorum 10    
	                 |                    
	                 +-- U_ravoisieri 13  
	                 |                    
	                 +-- U_tigridis 14    
  

Froufe et al., 2016

fig. 3: [clade_id: 7617] Unio
spp. terminals: 12, direct-child terminals: 12
rigorous outgroups.: 2, direct-child rigorous outgroups: 1
((U_gibbus, U_durieui), (U_tumidus, ((U_crassus, U_tumidiformis), (U_delphinus, (U_foucauldianus, (U_ravoisieri, (U_mancus, (U_elongatulus, (U_pictorum, U_caffer)))))))))


	                          +-- U_durieui 2      
	  +-----------------------+                    
	  |                       +-- U_gibbus 1       
	--+                                            
	  |  +----------------------- U_tumidus 3      
	  |  |                                         
	  +--+                    +-- U_crassus 4      
	     |  +-----------------+                    
	     |  |                 +-- U_tumidiformis 5 
	     +--+                                      
	        |  +----------------- U_delphinus 6    
	        +--+                                   
	           |  +-------------- U_foucauldianus 7
	           +--+                                
	              |  +----------- U_ravoisieri 8   
	              +--+                             
	                 |  +-------- U_mancus 9       
	                 +--+                          
	                    |  +----- U_elongatulus 10 
	                    +--+                       
	                       |  +-- U_caffer 12      
	                       +--+                    
	                          +-- U_pictorum 11    
  

Prié & Puillandre, 2014

fig. 2: [clade_id: 6847] Unio
spp. terminals: 11, direct-child terminals: 11
rigorous outgroups.: 0, direct-child rigorous outgroups: 0
((U_tumidus, (U_gibbus, U_durieui)), ((U_tumidiformis, (U_crassus, U_courtillieri)), (U_delphinus, (U_elongatulus, (U_mancus, (U_pictorum, U_ravoisieri))))))


	              +----- U_tumidus 1     
	  +-----------+                      
	  |           |  +-- U_durieui 3     
	  |           +--+                   
	  |              +-- U_gibbus 2      
	--+                                  
	  |           +----- U_tumidiformis 4
	  |  +--------+                      
	  |  |        |  +-- U_courtillieri 6
	  |  |        +--+                   
	  +--+           +-- U_crassus 5     
	     |                               
	     |  +----------- U_delphinus 7   
	     +--+                            
	        |  +-------- U_elongatulus 8 
	        +--+                         
	           |  +----- U_mancus 9      
	           +--+                      
	              |  +-- U_pictorum 10   
	              +--+                   
	                 +-- U_ravoisieri 11 
  

Lopes-Lima et al., 2017

fig. 1: [clade_id: 8238] Unio
spp. terminals: 9, direct-child terminals: 9
rigorous outgroups.: 0, direct-child rigorous outgroups: 0
((U_tumidus, U_gibbus), ((U_crassus, U_tumidiformis), (U_ravoisieri, (U_mancus, U_pictorum, U_delphinus, U_elongatulus))))


	           +-- U_gibbus 2      
	  +--------+                   
	  |        +-- U_tumidus 1     
	  |                            
	--+        +-- U_crassus 3     
	  |  +-----+                   
	  |  |     +-- U_tumidiformis 4
	  +--+                         
	     |  +----- U_ravoisieri 5  
	     |  |                      
	     +--+  +-- U_delphinus 8   
	        |  |                   
	        |  +-- U_elongatulus 9 
	        +--+                   
	           +-- U_mancus 6      
	           |                   
	           +-- U_pictorum 7    
  

Khalloufi et al., 2011

fig. 2: [clade_id: 4901] Unio
spp. terminals: 9, direct-child terminals: 9
rigorous outgroups.: 0, direct-child rigorous outgroups: 0
((U_tumidus, (U_gibbus, U_durieui)), ((U_tumidiformis, U_crassus), (U_ravoisieri, U_pictorum, (U_delphinus, U_mancus))))


	        +----- U_tumidus 1     
	  +-----+                      
	  |     |  +-- U_durieui 3     
	  |     +--+                   
	  |        +-- U_gibbus 2      
	--+                            
	  |        +-- U_crassus 5     
	  |  +-----+                   
	  |  |     +-- U_tumidiformis 4
	  +--+                         
	     |  +----- U_pictorum 7    
	     |  |                      
	     +--+----- U_ravoisieri 6  
	        |                      
	        |  +-- U_delphinus 8   
	        +--+                   
	           +-- U_mancus 9      
  

Araujo et al., 2009

fig. 7A: [clade_id: 4108] "Unio" + Anodonta
spp. terminals: 7, direct-child terminals: 7
rigorous outgroups.: 1, direct-child rigorous outgroups: 1
((U_gibbus, U_tumidus, A_cygnea), (U_pictorum, U_pictorum, U_mancus, (U_tumidiformis, U_crassus)))


	        +-- A_cygnea 3      
	        |                   
	  +-----+-- U_gibbus 1      
	  |     |                   
	  |     +-- U_tumidus 2     
	--+                         
	  |  +----- U_mancus 6      
	  |  |                      
	  |  +----- U_pictorum 4    
	  +--+                      
	     +----- U_pictorum 5    
	     |                      
	     |  +-- U_crassus 8     
	     +--+                   
	        +-- U_tumidiformis 7
  

fig. 7B: [clade_id: 4131] Unio
spp. terminals: 8, direct-child terminals: 8
rigorous outgroups.: 0, direct-child rigorous outgroups: 0
(U_gibbus, (U_tumidus, ((U_tumidiformis, U_crassus), ((U_pictorum, U_mancus), (U_pictorum, U_caffer)))))


	  +-------------- U_gibbus 1      
	  |                               
	--+  +----------- U_tumidus 2     
	  |  |                            
	  |  |        +-- U_crassus 4     
	  +--+  +-----+                   
	     |  |     +-- U_tumidiformis 3
	     |  |                         
	     +--+     +-- U_caffer 8      
	        |  +--+                   
	        |  |  +-- U_pictorum 7    
	        +--+                      
	           |  +-- U_mancus 6      
	           +--+                   
	              +-- U_pictorum 5    
  

fig. 7C: [clade_id: 4151] Unio
spp. terminals: 7, direct-child terminals: 7
rigorous outgroups.: 0, direct-child rigorous outgroups: 0
((U_gibbus, U_tumidus), ((U_tumidiformis, U_crassus), (U_pictorum, U_pictorum, U_mancus)))


	        +-- U_gibbus 1      
	  +-----+                   
	  |     +-- U_tumidus 2     
	  |                         
	--+     +-- U_crassus 4     
	  |  +--+                   
	  |  |  +-- U_tumidiformis 3
	  +--+                      
	     |  +-- U_mancus 7      
	     |  |                   
	     +--+-- U_pictorum 5    
	        |                   
	        +-- U_pictorum 6    
  

Marrone et al., 2019

fig. 1: [clade_id: 14423] Unio
spp. terminals: 8, direct-child terminals: 8
rigorous outgroups.: 0, direct-child rigorous outgroups: 0
(U_tumidus, (U_delphinus, (U_foucauldianus, (U_mancus, U_elongatulus, U_ravoisieri, (U_pictorum, U_tigridis)))))


	  +-------------- U_tumidus 1      
	  |                                
	--+  +----------- U_delphinus 2    
	  |  |                             
	  +--+  +-------- U_foucauldianus 3
	     |  |                          
	     +--+  +----- U_elongatulus 5  
	        |  |                       
	        |  +----- U_mancus 4       
	        +--+                       
	           +----- U_ravoisieri 6   
	           |                       
	           |  +-- U_pictorum 7     
	           +--+                    
	              +-- U_tigridis 8     
  

Reis et al., 2013

fig. 4: [clade_id: 6571] Unio
spp. terminals: 6, direct-child terminals: 6
rigorous outgroups.: 0, direct-child rigorous outgroups: 0
(U_tumidus, ((U_tumidiformis, U_crassus), (U_delphinus, (U_mancus, U_pictorum))))


	  +----------- U_tumidus 1     
	  |                            
	--+        +-- U_crassus 3     
	  |  +-----+                   
	  |  |     +-- U_tumidiformis 2
	  +--+                         
	     |  +----- U_delphinus 4   
	     +--+                      
	        |  +-- U_mancus 5      
	        +--+                   
	           +-- U_pictorum 6    
  

fig. 5: [clade_id: 6602] Unio
spp. terminals: 8, direct-child terminals: 8
rigorous outgroups.: 0, direct-child rigorous outgroups: 0
(U_tumidus, ((U_tumidiformis, U_crassus), (Unio, ((U_delphinus, U_mancus), (U_pictorum, U_caffer)))))


	  +-------------- U_tumidus 1     
	  |                               
	--+           +-- U_crassus 3     
	  |  +--------+                   
	  |  |        +-- U_tumidiformis 2
	  +--+                            
	     |  +-------- Unio 4          
	     |  |                         
	     +--+     +-- U_caffer 8      
	        |  +--+                   
	        |  |  +-- U_pictorum 7    
	        +--+                      
	           |  +-- U_delphinus 5   
	           +--+                   
	              +-- U_mancus 6      
  

Prie et al., 2012

fig. 2: [clade_id: 6383] Unio
spp. terminals: 5, direct-child terminals: 5
rigorous outgroups.: 0, direct-child rigorous outgroups: 0
(U_tumidus, (U_crassus, (U_pictorum, (U_mancus, U_elongatulus))))


	  +----------- U_tumidus 1    
	--+                           
	  |  +-------- U_crassus 2    
	  +--+                        
	     |  +----- U_pictorum 3   
	     +--+                     
	        |  +-- U_elongatulus 5
	        +--+                  
	           +-- U_mancus 4     
  

Fukata & Iigo, 2020

fig. 1: [clade_id: 17096] Unio
spp. terminals: 4, direct-child terminals: 4
rigorous outgroups.: 10, direct-child rigorous outgroups: 6
(U_tumidus, (U_crassus, (U_pictorum, U_delphinus)))


	  +-------- U_tumidus 1  
	--+                      
	  |  +----- U_crassus 2  
	  +--+                   
	     |  +-- U_delphinus 4
	     +--+                
	        +-- U_pictorum 3 
  

Fukata & Iigo, 2019

fig. 1: [clade_id: 17629] Unio
spp. terminals: 4, direct-child terminals: 4
rigorous outgroups.: 7, direct-child rigorous outgroups: 5
(U_tumidus, (U_crassus, (U_delphinus, U_pictorum)))


	  +-------- U_tumidus 1  
	--+                      
	  |  +----- U_crassus 2  
	  +--+                   
	     |  +-- U_delphinus 3
	     +--+                
	        +-- U_pictorum 4 
  

Wu et al., 2020

fig. 2: [clade_id: 17724] Unio
spp. terminals: 1, direct-child terminals: 1
rigorous outgroups.: 3, direct-child rigorous outgroups: 1
unknown

fig. 3: [clade_id: 17843] Unio
spp. terminals: 4, direct-child terminals: 4
rigorous outgroups.: 11, direct-child rigorous outgroups: 6
(U_tumidus, (U_crassus, (U_delphinus, U_pictorum)))


	  +-------- U_tumidus 1  
	--+                      
	  |  +----- U_crassus 2  
	  +--+                   
	     |  +-- U_delphinus 3
	     +--+                
	        +-- U_pictorum 4 
  

fig. S1: [clade_id: 17791] Unio
spp. terminals: 1, direct-child terminals: 1
rigorous outgroups.: 3, direct-child rigorous outgroups: 1
unknown

Kim et al., 2021

fig. 1: [clade_id: 19469] Unio
spp. terminals: 4, direct-child terminals: 4
rigorous outgroups.: 7, direct-child rigorous outgroups: 6
(U_tumidus, (U_crassus, (U_delphinus, U_pictorum)))


	  +-------- U_tumidus 1  
	--+                      
	  |  +----- U_crassus 2  
	  +--+                   
	     |  +-- U_delphinus 3
	     +--+                
	        +-- U_pictorum 4 
  

Zieritz et al., 2021

fig. 3-F, 5: [clade_id: 20332] Unio
spp. terminals: 4, direct-child terminals: 4
rigorous outgroups.: 7, direct-child rigorous outgroups: 5
(U_tumidus, (U_crassus, (U_pictorum, U_delphinus)))


	  +-------- U_tumidus 1  
	--+                      
	  |  +----- U_crassus 2  
	  +--+                   
	     |  +-- U_delphinus 4
	     +--+                
	        +-- U_pictorum 3 
  

fig. 3-M, 4: [clade_id: 20397] Unio
spp. terminals: 3, direct-child terminals: 3
rigorous outgroups.: 0, direct-child rigorous outgroups: 0
(U_tumidus, (U_delphinus, U_crassus))


	  +----- U_tumidus 1  
	--+                   
	  |  +-- U_crassus 3  
	  +--+                
	     +-- U_delphinus 2
  

Lopes-Lima et al., 2017

fig. 1-2: [clade_id: 8710] Unio
spp. terminals: 4, direct-child terminals: 4
rigorous outgroups.: 7, direct-child rigorous outgroups: 4
(U_gibbus, U_tumidus, (U_crassus, U_pictorum))


	  +----- U_gibbus 1  
	  |                  
	--+----- U_tumidus 2 
	  |                  
	  |  +-- U_crassus 3 
	  +--+               
	     +-- U_pictorum 4
  

Soroka, 2005

fig. 3: [clade_id: 2693] Unio
spp. terminals: 4, direct-child terminals: 4
rigorous outgroups.: 0, direct-child rigorous outgroups: 0
(U_tumidus, (U_crassus, (U_caffer, U_pictorum)))


	  +-------- U_tumidus 1 
	--+                     
	  |  +----- U_crassus 2 
	  +--+                  
	     |  +-- U_caffer 3  
	     +--+               
	        +-- U_pictorum 4
  

Huang et al., 2018

fig. 3, 5: [clade_id: 10629] Unio
spp. terminals: 1, direct-child terminals: 1
rigorous outgroups.: 1, direct-child rigorous outgroups: 1
unknown

fig. 6: [clade_id: 10683] Unio
spp. terminals: 4, direct-child terminals: 4
rigorous outgroups.: 5, direct-child rigorous outgroups: 3
(U_tumidus, (U_crassus, (U_delphinus, U_pictorum)))


	  +-------- U_tumidus 1  
	--+                      
	  |  +----- U_crassus 2  
	  +--+                   
	     |  +-- U_delphinus 3
	     +--+                
	        +-- U_pictorum 4 
  

fig. 6-M: [clade_id: 10752] Unio
spp. terminals: 3, direct-child terminals: 3
rigorous outgroups.: 1, direct-child rigorous outgroups: 1
(U_tumidus, (U_crassus, U_delphinus))


	  +----- U_tumidus 1  
	--+                   
	  |  +-- U_crassus 2  
	  +--+                
	     +-- U_delphinus 3
  

Soroka & Burzynski, 2017

fig. 1: [clade_id: 10572] Unio
spp. terminals: 4, direct-child terminals: 4
rigorous outgroups.: 7, direct-child rigorous outgroups: 5
(U_tumidus, (U_crassus, (U_pictorum, U_delphinus)))


	  +-------- U_tumidus 1  
	--+                      
	  |  +----- U_crassus 2  
	  +--+                   
	     |  +-- U_delphinus 4
	     +--+                
	        +-- U_pictorum 3 
  

Huang et al., 2019

fig. 2: [clade_id: 11475] Unio
spp. terminals: 4, direct-child terminals: 4
rigorous outgroups.: 16, direct-child rigorous outgroups: 6
(U_tumidus, (U_crassus, (U_delphinus, U_pictorum)))


	  +-------- U_tumidus 1  
	--+                      
	  |  +----- U_crassus 2  
	  +--+                   
	     |  +-- U_delphinus 3
	     +--+                
	        +-- U_pictorum 4 
  

fig. 3: [clade_id: 12626] Unio
spp. terminals: 4, direct-child terminals: 4
rigorous outgroups.: 9, direct-child rigorous outgroups: 6
(U_tumidus, (U_crassus, (U_delphinus, U_pictorum)))


	  +-------- U_tumidus 1  
	--+                      
	  |  +----- U_crassus 2  
	  +--+                   
	     |  +-- U_delphinus 3
	     +--+                
	        +-- U_pictorum 4 
  

fig. 3-M: [clade_id: 12713] Unio
spp. terminals: 3, direct-child terminals: 3
rigorous outgroups.: 2, direct-child rigorous outgroups: 2
(U_tumidus, (U_crassus, U_delphinus))


	  +----- U_tumidus 1  
	--+                   
	  |  +-- U_crassus 2  
	  +--+                
	     +-- U_delphinus 3
  

fig. S3: [clade_id: 11887] Unio
spp. terminals: 4, direct-child terminals: 4
rigorous outgroups.: 15, direct-child rigorous outgroups: 6
(U_tumidus, (U_crassus, (U_delphinus, U_pictorum)))


	  +-------- U_tumidus 1  
	--+                      
	  |  +----- U_crassus 2  
	  +--+                   
	     |  +-- U_delphinus 3
	     +--+                
	        +-- U_pictorum 4 
  

fig. S4: [clade_id: 12287] Unio
spp. terminals: 4, direct-child terminals: 4
rigorous outgroups.: 16, direct-child rigorous outgroups: 6
(U_tumidus, (U_crassus, (U_delphinus, U_pictorum)))


	  +-------- U_tumidus 1  
	--+                      
	  |  +----- U_crassus 2  
	  +--+                   
	     |  +-- U_delphinus 3
	     +--+                
	        +-- U_pictorum 4 
  

Wang et al., 2018

fig. 2: [clade_id: 12785] Unio
spp. terminals: 4, direct-child terminals: 4
rigorous outgroups.: 4, direct-child rigorous outgroups: 3
(U_tumidus, (U_crassus, (U_pictorum, U_delphinus)))


	  +-------- U_tumidus 1  
	--+                      
	  |  +----- U_crassus 2  
	  +--+                   
	     |  +-- U_delphinus 4
	     +--+                
	        +-- U_pictorum 3 
  

Chase et al., 2018

fig. 1: [clade_id: 13427] "Unio" + Cumberlandia + Pyganodon + Lampsilis + Anodonta + Quadrula + Echyridella + Utterbackia + Lamprotula
spp. terminals: 4, direct-child terminals: 4
rigorous outgroups.: 0, direct-child rigorous outgroups: 0
((E_menziesii, (C_monodonta, ((Q_quadrula, (L_powellii, L_siliquoidea)), (L_leaii, ((U_delphinus, U_tumidus), (A_anatina, (U_peninsularis, P_grandis))))))), ((C_monodonta, E_menziesii), ((L_leaii, (Q_quadrula, (L_powellii, L_siliquoidea))), ((U_delphinus, U_tumidus), (A_anatina, (U_peninsularis, P_grandis))))))


	     +-------------------- E_menziesii 1    
	     |                                      
	  +--+  +----------------- C_monodonta 2    
	  |  |  |                                   
	  |  |  |           +----- Q_quadrula 3     
	  |  +--+  +--------+                       
	  |     |  |        |  +-- L_powellii 4     
	  |     |  |        +--+                    
	  |     +--+           +-- L_siliquoidea 5  
	  |        |                                
	  |        |  +----------- L_leaii 6        
	  |        |  |                             
	  |        +--+        +-- U_delphinus 7    
	  |           |  +-----+                    
	--+           |  |     +-- U_tumidus 8      
	  |           +--+                          
	  |              |  +----- A_anatina 9      
	  |              +--+                       
	  |                 |  +-- P_grandis 11     
	  |                 +--+                    
	  |                    +-- U_peninsularis 10
	  |                                         
	  |                    +-- C_monodonta 12   
	  |        +-----------+                    
	  |        |           +-- E_menziesii 13   
	  |        |                                
	  |        |     +-------- L_leaii 14       
	  +--------+  +--+                          
	           |  |  |  +----- Q_quadrula 15    
	           |  |  +--+                       
	           |  |     |  +-- L_powellii 16    
	           |  |     +--+                    
	           +--+        +-- L_siliquoidea 17 
	              |                             
	              |        +-- U_delphinus 18   
	              |  +-----+                    
	              |  |     +-- U_tumidus 19     
	              +--+                          
	                 |  +----- A_anatina 20     
	                 +--+                       
	                    |  +-- P_grandis 22     
	                    +--+                    
	                       +-- U_peninsularis 21
  

Burzynski & Soroka, 2018

fig. 5a: [clade_id: 13447] Unio
spp. terminals: 4, direct-child terminals: 4
rigorous outgroups.: 1, direct-child rigorous outgroups: 1
(U_tumidus, (U_crassus, (U_pictorum, U_delphinus)))


	  +-------- U_tumidus 1  
	--+                      
	  |  +----- U_crassus 2  
	  +--+                   
	     |  +-- U_delphinus 4
	     +--+                
	        +-- U_pictorum 3 
  

fig. 5b: [clade_id: 13487] Unio
spp. terminals: 4, direct-child terminals: 4
rigorous outgroups.: 1, direct-child rigorous outgroups: 1
(U_tumidus, (U_crassus, (U_pictorum, U_delphinus)))


	  +-------- U_tumidus 1  
	--+                      
	  |  +----- U_crassus 2  
	  +--+                   
	     |  +-- U_delphinus 4
	     +--+                
	        +-- U_pictorum 3 
  

Aunins et al., 2018

fig. 1: [clade_id: 13564] Unio
spp. terminals: 4, direct-child terminals: 4
rigorous outgroups.: 9, direct-child rigorous outgroups: 6
(U_tumidus, (U_crassus, (U_pictorum, U_delphinus)))


	  +-------- U_tumidus 1  
	--+                      
	  |  +----- U_crassus 2  
	  +--+                   
	     |  +-- U_delphinus 4
	     +--+                
	        +-- U_pictorum 3 
  

Wu et al., 2019

fig. 3: [clade_id: 13611] Unio
spp. terminals: 4, direct-child terminals: 4
rigorous outgroups.: 11, direct-child rigorous outgroups: 6
(U_tumidus, (U_crassus, (U_delphinus, U_pictorum)))


	  +-------- U_tumidus 1  
	--+                      
	  |  +----- U_crassus 2  
	  +--+                   
	     |  +-- U_delphinus 3
	     +--+                
	        +-- U_pictorum 4 
  

Wang et al., 2018

fig. 2: [clade_id: 13709] Unio
spp. terminals: 4, direct-child terminals: 4
rigorous outgroups.: 4, direct-child rigorous outgroups: 4
(U_tumidus, (U_crassus, (U_pictorum, U_delphinus)))


	  +-------- U_tumidus 1  
	--+                      
	  |  +----- U_crassus 2  
	  +--+                   
	     |  +-- U_delphinus 4
	     +--+                
	        +-- U_pictorum 3 
  

Klishko et al., 2019

fig. 9: [clade_id: 13971] Unio
spp. terminals: 4, direct-child terminals: 4
rigorous outgroups.: 4, direct-child rigorous outgroups: 2
(U_tumidus, U_gibbus, (U_crassus, U_pictorum))


	  +----- U_gibbus 2  
	  |                  
	--+----- U_tumidus 1 
	  |                  
	  |  +-- U_crassus 3 
	  +--+               
	     +-- U_pictorum 4
  

Soroka, 2010

fig. 2: [clade_id: 4570] Unio
spp. terminals: 3, direct-child terminals: 3
rigorous outgroups.: 0, direct-child rigorous outgroups: 0
(U_tumidus, U_pictorum, U_crassus)


	  +-- U_crassus 3 
	  |               
	--+-- U_pictorum 2
	  |               
	  +-- U_tumidus 1 
  

fig. 2-M: [clade_id: 4583] Unio
spp. terminals: 3, direct-child terminals: 3
rigorous outgroups.: 0, direct-child rigorous outgroups: 0
(U_crassus, (U_tumidus, U_pictorum))


	  +----- U_crassus 1 
	--+                  
	  |  +-- U_pictorum 3
	  +--+               
	     +-- U_tumidus 2 
  

fig. 3a: [clade_id: 4595] Unio
spp. terminals: 4, direct-child terminals: 4
rigorous outgroups.: 0, direct-child rigorous outgroups: 0
(U_tumidus, (U_crassus, (U_pictorum, U_mancus)))


	  +-------- U_tumidus 1 
	--+                     
	  |  +----- U_crassus 2 
	  +--+                  
	     |  +-- U_mancus 4  
	     +--+               
	        +-- U_pictorum 3
  

fig. 3a-M: [clade_id: 4608] Unio
spp. terminals: 3, direct-child terminals: 3
rigorous outgroups.: 0, direct-child rigorous outgroups: 0
(U_crassus, (U_tumidus, U_pictorum))


	  +----- U_crassus 1 
	--+                  
	  |  +-- U_pictorum 3
	  +--+               
	     +-- U_tumidus 2 
  

fig. 3b: [clade_id: 4620] Unio
spp. terminals: 4, direct-child terminals: 4
rigorous outgroups.: 0, direct-child rigorous outgroups: 0
(U_tumidus, (U_crassus, (U_pictorum, U_mancus)))


	  +-------- U_tumidus 1 
	--+                     
	  |  +----- U_crassus 2 
	  +--+                  
	     |  +-- U_mancus 4  
	     +--+               
	        +-- U_pictorum 3
  

fig. 3b-M: [clade_id: 4631] Unio
spp. terminals: 3, direct-child terminals: 3
rigorous outgroups.: 0, direct-child rigorous outgroups: 0
(U_tumidus, (U_crassus, U_pictorum))


	  +----- U_tumidus 1 
	--+                  
	  |  +-- U_crassus 2 
	  +--+               
	     +-- U_pictorum 3
  

fig. 4: [clade_id: 4641] Unio
spp. terminals: 3, direct-child terminals: 3
rigorous outgroups.: 0, direct-child rigorous outgroups: 0
(U_tumidus, (U_pictorum, U_crassus))


	  +----- U_tumidus 1 
	--+                  
	  |  +-- U_crassus 3 
	  +--+               
	     +-- U_pictorum 2
  

fig. 5a: [clade_id: 4658] Unio
spp. terminals: 3, direct-child terminals: 3
rigorous outgroups.: 0, direct-child rigorous outgroups: 0
(U_crassus, (U_pictorum, U_tumidus))


	  +----- U_crassus 1 
	--+                  
	  |  +-- U_pictorum 2
	  +--+               
	     +-- U_tumidus 3 
  

fig. 5b: [clade_id: 4675] Unio
spp. terminals: 3, direct-child terminals: 3
rigorous outgroups.: 0, direct-child rigorous outgroups: 0
(U_tumidus, (U_pictorum, U_crassus))


	  +----- U_tumidus 1 
	--+                  
	  |  +-- U_crassus 3 
	  +--+               
	     +-- U_pictorum 2
  

fig. 8a: [clade_id: 4691] Unio
spp. terminals: 3, direct-child terminals: 3
rigorous outgroups.: 0, direct-child rigorous outgroups: 0
(U_tumidus, (U_pictorum, U_crassus))


	  +----- U_tumidus 1 
	--+                  
	  |  +-- U_crassus 3 
	  +--+               
	     +-- U_pictorum 2
  

fig. 8a-M: [clade_id: 4700] Unio
spp. terminals: 2, direct-child terminals: 2
rigorous outgroups.: 0, direct-child rigorous outgroups: 0
(U_pictorum, U_tumidus)


	  +-- U_pictorum 1
	--+               
	  +-- U_tumidus 2 
  

fig. 8b: [clade_id: 4708] Unio
spp. terminals: 3, direct-child terminals: 3
rigorous outgroups.: 0, direct-child rigorous outgroups: 0
(U_tumidus, (U_pictorum, U_crassus))


	  +----- U_tumidus 1 
	--+                  
	  |  +-- U_crassus 3 
	  +--+               
	     +-- U_pictorum 2
  

fig. 8b-M: [clade_id: 4717] Unio
spp. terminals: 2, direct-child terminals: 2
rigorous outgroups.: 0, direct-child rigorous outgroups: 0
(U_pictorum, U_tumidus)


	  +-- U_pictorum 1
	--+               
	  +-- U_tumidus 2 
  

Cha et al., 2018

fig. 1: [clade_id: 14613] Unio
spp. terminals: 4, direct-child terminals: 4
rigorous outgroups.: 7, direct-child rigorous outgroups: 5
(U_tumidus, (U_crassus, (U_pictorum, U_delphinus)))


	  +-------- U_tumidus 1  
	--+                      
	  |  +----- U_crassus 2  
	  +--+                   
	     |  +-- U_delphinus 4
	     +--+                
	        +-- U_pictorum 3 
  

Froufe et al., 2019

fig. 3: [clade_id: 15223] Unio
spp. terminals: 4, direct-child terminals: 4
rigorous outgroups.: 7, direct-child rigorous outgroups: 5
(U_tumidus, (U_crassus, (U_pictorum, U_delphinus)))


	  +-------- U_tumidus 1  
	--+                      
	  |  +----- U_crassus 2  
	  +--+                   
	     |  +-- U_delphinus 4
	     +--+                
	        +-- U_pictorum 3 
  

fig. 3-M: [clade_id: 15312] Unio
spp. terminals: 3, direct-child terminals: 3
rigorous outgroups.: 0, direct-child rigorous outgroups: 0
(U_tumidus, (U_crassus, U_delphinus))


	  +----- U_tumidus 1  
	--+                   
	  |  +-- U_crassus 2  
	  +--+                
	     +-- U_delphinus 3
  

fig. 4: [clade_id: 15350] Unio
spp. terminals: 3, direct-child terminals: 3
rigorous outgroups.: 0, direct-child rigorous outgroups: 0
(U_tumidus, (U_crassus, U_delphinus))


	  +----- U_tumidus 1  
	--+                   
	  |  +-- U_crassus 2  
	  +--+                
	     +-- U_delphinus 3
  

fig. 6: [clade_id: 15427] Unio
spp. terminals: 4, direct-child terminals: 4
rigorous outgroups.: 7, direct-child rigorous outgroups: 5
(U_tumidus, (U_crassus, (U_pictorum, U_delphinus)))


	  +-------- U_tumidus 1  
	--+                      
	  |  +----- U_crassus 2  
	  +--+                   
	     |  +-- U_delphinus 4
	     +--+                
	        +-- U_pictorum 3 
  

Liu et al., 2017

fig. 2: [clade_id: 15549] Unio
spp. terminals: 4, direct-child terminals: 4
rigorous outgroups.: 10, direct-child rigorous outgroups: 4
(U_tumidus, U_gibbus, (U_crassus, U_pictorum))


	  +----- U_gibbus 2  
	  |                  
	--+----- U_tumidus 1 
	  |                  
	  |  +-- U_crassus 3 
	  +--+               
	     +-- U_pictorum 4
  

Klishko et al., 2018

fig. 8: [clade_id: 15847] Unio
spp. terminals: 4, direct-child terminals: 4
rigorous outgroups.: 4, direct-child rigorous outgroups: 1
(U_tumidus, U_gibbus, (U_crassus, U_pictorum))


	  +----- U_gibbus 2  
	  |                  
	--+----- U_tumidus 1 
	  |                  
	  |  +-- U_crassus 3 
	  +--+               
	     +-- U_pictorum 4
  

Klishko et al., 2017

fig. 8: [clade_id: 15938] Unio
spp. terminals: 4, direct-child terminals: 4
rigorous outgroups.: 2, direct-child rigorous outgroups: 1
(U_tumidus, (U_pictorum, (U_crassus, U_courtillieri)))


	  +-------- U_tumidus 1     
	--+                         
	  |  +----- U_pictorum 2    
	  +--+                      
	     |  +-- U_courtillieri 4
	     +--+                   
	        +-- U_crassus 3     
  

Nagel et al., 1998

fig. 1: [clade_id: 723] Unio
spp. terminals: 4, direct-child terminals: 4
rigorous outgroups.: 0, direct-child rigorous outgroups: 0
(U_crassus, (U_tumidus, (U_pictorum, U_mancus)))


	  +-------- U_crassus 1 
	--+                     
	  |  +----- U_tumidus 2 
	  +--+                  
	     |  +-- U_mancus 4  
	     +--+               
	        +-- U_pictorum 3
  

fig. 8: [clade_id: 750] Unio
spp. terminals: 4, direct-child terminals: 4
rigorous outgroups.: 0, direct-child rigorous outgroups: 0
(U_tumidus, (U_pictorum, U_crassus, U_mancus))


	  +----- U_tumidus 1 
	  |                  
	--+  +-- U_crassus 3 
	  |  |               
	  +--+-- U_mancus 4  
	     |               
	     +-- U_pictorum 2
  

fig. 9: [clade_id: 754] Unio
spp. terminals: 1, direct-child terminals: 1
rigorous outgroups.: 0, direct-child rigorous outgroups: 0
unknown

Sano et al., 2020

fig. 2: [clade_id: 18269] Unio
spp. terminals: 3, direct-child terminals: 3
rigorous outgroups.: 11, direct-child rigorous outgroups: 6
(U_crassus, (U_delphinus, U_pictorum))


	  +----- U_crassus 1  
	--+                   
	  |  +-- U_delphinus 2
	  +--+                
	     +-- U_pictorum 3 
  

fig. 3: [clade_id: 18426] Unio
spp. terminals: 3, direct-child terminals: 3
rigorous outgroups.: 11, direct-child rigorous outgroups: 6
(U_crassus, (U_delphinus, U_pictorum))


	  +----- U_crassus 1  
	--+                   
	  |  +-- U_delphinus 2
	  +--+                
	     +-- U_pictorum 3 
  

fig. 4: [clade_id: 18579] Unio
spp. terminals: 3, direct-child terminals: 3
rigorous outgroups.: 11, direct-child rigorous outgroups: 6
(U_crassus, (U_delphinus, U_pictorum))


	  +----- U_crassus 1  
	--+                   
	  |  +-- U_delphinus 2
	  +--+                
	     +-- U_pictorum 3 
  

Guerra et al., 2019

fig. 1: [clade_id: 19027] Unio
spp. terminals: 3, direct-child terminals: 3
rigorous outgroups.: 2, direct-child rigorous outgroups: 2
(U_tumidus, (U_crassus, U_delphinus))


	  +----- U_tumidus 1  
	--+                   
	  |  +-- U_crassus 2  
	  +--+                
	     +-- U_delphinus 3
  

fig. 1-M: [clade_id: 18978] Unio
spp. terminals: 3, direct-child terminals: 3
rigorous outgroups.: 2, direct-child rigorous outgroups: 2
(U_tumidus, (U_crassus, U_delphinus))


	  +----- U_tumidus 1  
	--+                   
	  |  +-- U_crassus 2  
	  +--+                
	     +-- U_delphinus 3
  

fig. 2: [clade_id: 19154] Unio
spp. terminals: 3, direct-child terminals: 3
rigorous outgroups.: 2, direct-child rigorous outgroups: 2
(U_tumidus, (U_crassus, U_delphinus))


	  +----- U_tumidus 1  
	--+                   
	  |  +-- U_crassus 2  
	  +--+                
	     +-- U_delphinus 3
  

fig. 2-M: [clade_id: 19103] Unio
spp. terminals: 3, direct-child terminals: 3
rigorous outgroups.: 2, direct-child rigorous outgroups: 2
(U_tumidus, (U_crassus, U_delphinus))


	  +----- U_tumidus 1  
	--+                   
	  |  +-- U_crassus 2  
	  +--+                
	     +-- U_delphinus 3
  

Zieritz et al., 2012

fig. 4: [clade_id: 6066] Unio
spp. terminals: 3, direct-child terminals: 3
rigorous outgroups.: 0, direct-child rigorous outgroups: 0
(U_crassus, (U_tumidus, U_pictorum))


	  +----- U_crassus 1 
	--+                  
	  |  +-- U_pictorum 3
	  +--+               
	     +-- U_tumidus 2 
  

Plazzi et al., 2011

fig. 5: [clade_id: 5005] Unio
spp. terminals: 3, direct-child terminals: 3
rigorous outgroups.: 0, direct-child rigorous outgroups: 0
(U_tumidus, (U_crassus, U_pictorum))


	  +----- U_tumidus 1 
	--+                  
	  |  +-- U_crassus 2 
	  +--+               
	     +-- U_pictorum 3
  

Manendo et al., 2008

fig. 1: [clade_id: 3721] Unio
spp. terminals: 3, direct-child terminals: 3
rigorous outgroups.: 0, direct-child rigorous outgroups: 0
(U_crassus, (U_pictorum, U_tumidus))


	  +----- U_crassus 1 
	--+                  
	  |  +-- U_pictorum 2
	  +--+               
	     +-- U_tumidus 3 
  

Jadhav & Jamkhedkar, 2009

fig. 11: [clade_id: 4486] "Unio" + Alasmidonta + Pyganodon + Anodonta + Strophitus + Lasmigona
spp. terminals: 3, direct-child terminals: 3
rigorous outgroups.: 1, direct-child rigorous outgroups: 1
(U_pictorum, ((U_pictorum, U_caffer), (Anodonta, (L_compressa, (S_undulatus, A_marginata, P_grandis)))))


	  +-------------- U_pictorum 1 
	  |                            
	--+           +-- U_caffer 3   
	  |  +--------+                
	  |  |        +-- U_pictorum 2 
	  +--+                         
	     |  +-------- Anodonta 4   
	     |  |                      
	     +--+  +----- L_compressa 5
	        |  |                   
	        +--+  +-- A_marginata 7
	           |  |                
	           +--+-- P_grandis 8  
	              |                
	              +-- S_undulatus 6
  

Källersjö et al., 2005

fig. 1A: [clade_id: 2672] Unio
spp. terminals: 3, direct-child terminals: 3
rigorous outgroups.: 0, direct-child rigorous outgroups: 0
(U_crassus, U_pictorum, U_tumidus)


	  +-- U_crassus 1 
	  |               
	--+-- U_pictorum 2
	  |               
	  +-- U_tumidus 3 
  

fig. 1B-F: [clade_id: 2684] Unio
spp. terminals: 3, direct-child terminals: 3
rigorous outgroups.: 0, direct-child rigorous outgroups: 0
(U_tumidus, (U_crassus, U_pictorum))


	  +----- U_tumidus 1 
	--+                  
	  |  +-- U_crassus 2 
	  +--+               
	     +-- U_pictorum 3
  

Bolotov et al., 2017

fig. S1: [clade_id: 8958] Unio
spp. terminals: 3, direct-child terminals: 3
rigorous outgroups.: 7, direct-child rigorous outgroups: 4
(U_tumidus, (U_pictorum, U_crassus))


	  +----- U_tumidus 1 
	--+                  
	  |  +-- U_crassus 3 
	  +--+               
	     +-- U_pictorum 2
  

fig. S2: [clade_id: 9255] Unio
spp. terminals: 3, direct-child terminals: 3
rigorous outgroups.: 7, direct-child rigorous outgroups: 4
(U_tumidus, (U_crassus, U_pictorum))


	  +----- U_tumidus 1 
	--+                  
	  |  +-- U_crassus 2 
	  +--+               
	     +-- U_pictorum 3
  

Bolotov et al., 2017

fig. S1: [clade_id: 9451] "Unio" + Cuneopsis + Aculamprotula + Schistodesmus + Nodularia
spp. terminals: 3, direct-child terminals: 3
rigorous outgroups.: 7, direct-child rigorous outgroups: 4
(U_crassus, U_pictorum, U_tumidus, A_tortuosa, C_heudei, C_pisciculus, C_rufescens, N_douglasiae, N_nuxpersicae, S_lampreyanus)


	  +-- A_tortuosa 4    
	  |                   
	  +-- C_heudei 5      
	  |                   
	  +-- C_pisciculus 6  
	  |                   
	  +-- C_rufescens 7   
	  |                   
	  +-- N_douglasiae 8  
	--+                   
	  +-- N_nuxpersicae 9 
	  |                   
	  +-- S_lampreyanus 10
	  |                   
	  +-- U_crassus 1     
	  |                   
	  +-- U_pictorum 2    
	  |                   
	  +-- U_tumidus 3     
  

Froufe et al., 2017

fig. 7: [clade_id: 9726] Unio
spp. terminals: 3, direct-child terminals: 3
rigorous outgroups.: 0, direct-child rigorous outgroups: 0
(U_mancus, (U_elongatulus, U_pictorum))


	  +----- U_mancus 1     
	--+                     
	  |  +-- U_elongatulus 2
	  +--+                  
	     +-- U_pictorum 3   
  

Dyduch-Falniowska & Koziol, 1989

fig. 5, 7: [clade_id: 716] Unio
spp. terminals: 3, direct-child terminals: 3
rigorous outgroups.: 0, direct-child rigorous outgroups: 0
(U_pictorum, (U_tumidus, U_crassus))


	  +----- U_pictorum 1
	--+                  
	  |  +-- U_crassus 3 
	  +--+               
	     +-- U_tumidus 2 
  

Bolotov et al., 2018

fig. 2: [clade_id: 10361] Unio
spp. terminals: 3, direct-child terminals: 3
rigorous outgroups.: 3, direct-child rigorous outgroups: 1
(U_tumidus, (U_crassus, U_pictorum))


	  +----- U_tumidus 1 
	--+                  
	  |  +-- U_crassus 2 
	  +--+               
	     +-- U_pictorum 3
  

Bogatov et al., 2018

fig. 1: [clade_id: 12921] "Unio" + Anodonta + Pseudanodonta
spp. terminals: 3, direct-child terminals: 3
rigorous outgroups.: 0, direct-child rigorous outgroups: 0
(U_tumidus, ((A_anatina, P_complanata), (U_crassus, U_pictorum)))


	  +-------- U_tumidus 1   
	  |                       
	--+     +-- A_anatina 2   
	  |  +--+                 
	  |  |  +-- P_complanata 3
	  +--+                    
	     |  +-- U_crassus 4   
	     +--+                 
	        +-- U_pictorum 5  
  

fig. 2: [clade_id: 12932] "Unio" + Anodonta + Pseudanodonta
spp. terminals: 3, direct-child terminals: 3
rigorous outgroups.: 0, direct-child rigorous outgroups: 0
(U_crassus, (U_pictorum, (U_tumidus, (A_anatina, P_complanata))))


	  +----------- U_crassus 1   
	--+                          
	  |  +-------- U_pictorum 2  
	  +--+                       
	     |  +----- U_tumidus 3   
	     +--+                    
	        |  +-- A_anatina 4   
	        +--+                 
	           +-- P_complanata 5
  

Bolotov et al., 2020

fig. 1: [clade_id: 16332] Unio
spp. terminals: 3, direct-child terminals: 3
rigorous outgroups.: 2, direct-child rigorous outgroups: 2
(U_tumidus, (U_pictorum, U_crassus))


	  +----- U_tumidus 1 
	--+                  
	  |  +-- U_crassus 3 
	  +--+               
	     +-- U_pictorum 2
  

Tomilova et al., 2020

fig. 3: [clade_id: 16857] Unio
spp. terminals: 2, direct-child terminals: 2
rigorous outgroups.: 0, direct-child rigorous outgroups: 0
(U_tumidus, U_pictorum)


	  +-- U_pictorum 2
	--+               
	  +-- U_tumidus 1 
  

Graf & Ó Foighil, 2000

fig. 2A: [clade_id: 73] Unio
spp. terminals: 2, direct-child terminals: 2
rigorous outgroups.: 0, direct-child rigorous outgroups: 0
(U_pictorum, U_caffer)


	  +-- U_caffer 2  
	--+               
	  +-- U_pictorum 1
  

fig. 2B: [clade_id: 137] Unio
spp. terminals: 2, direct-child terminals: 2
rigorous outgroups.: 0, direct-child rigorous outgroups: 0
(U_pictorum, U_caffer)


	  +-- U_caffer 2  
	--+               
	  +-- U_pictorum 1
  

Bogan & Hoeh, 2000

fig. 1: [clade_id: 1226] Unio
spp. terminals: 2, direct-child terminals: 2
rigorous outgroups.: 0, direct-child rigorous outgroups: 0
(U_pictorum, U_tumidus)


	  +-- U_pictorum 1
	--+               
	  +-- U_tumidus 2 
  

Graf, 2002

fig. 2: [clade_id: 1674] Unio
spp. terminals: 2, direct-child terminals: 2
rigorous outgroups.: 0, direct-child rigorous outgroups: 0
(U_pictorum, U_caffer)


	  +-- U_caffer 2  
	--+               
	  +-- U_pictorum 1
  

Hoeh et al., 2001

fig. 14.2: [clade_id: 1584] Unio
spp. terminals: 2, direct-child terminals: 2
rigorous outgroups.: 0, direct-child rigorous outgroups: 0
(U_pictorum, U_tumidus)


	  +-- U_pictorum 1
	--+               
	  +-- U_tumidus 2 
  

fig. 14.4: [clade_id: 1654] "Unio" + Pyganodon + Anodonta + Strophitus
spp. terminals: 2, direct-child terminals: 2
rigorous outgroups.: 0, direct-child rigorous outgroups: 0
(U_pictorum, U_tumidus, (A_cygnea, (S_undulatus, P_grandis)))


	  +-------- U_pictorum 1 
	  |                      
	--+-------- U_tumidus 2  
	  |                      
	  |  +----- A_cygnea 3   
	  +--+                   
	     |  +-- P_grandis 5  
	     +--+                
	        +-- S_undulatus 4
  

Hoeh et al., 2002

fig. 2: [clade_id: 1735] "Unio" + Pyganodon + Anodonta + Strophitus
spp. terminals: 2, direct-child terminals: 2
rigorous outgroups.: 0, direct-child rigorous outgroups: 0
(U_pictorum, (U_tumidus, (A_cygnea, (P_grandis, S_undulatus))))


	  +----------- U_pictorum 1 
	--+                         
	  |  +-------- U_tumidus 2  
	  +--+                      
	     |  +----- A_cygnea 3   
	     +--+                   
	        |  +-- P_grandis 4  
	        +--+                
	           +-- S_undulatus 5
  

Roe & Hoeh, 2003

fig. 4.2: [clade_id: 2006] Unio
spp. terminals: 2, direct-child terminals: 2
rigorous outgroups.: 0, direct-child rigorous outgroups: 0
(U_pictorum, U_tumidus)


	  +-- U_pictorum 1
	--+               
	  +-- U_tumidus 2 
  

fig. 4.3: [clade_id: 2078] Unio
spp. terminals: 2, direct-child terminals: 2
rigorous outgroups.: 0, direct-child rigorous outgroups: 0
(U_pictorum, U_tumidus)


	  +-- U_pictorum 1
	--+               
	  +-- U_tumidus 2 
  

Sayenko et al., 2021

fig. 6: [clade_id: 19386] Unio
spp. terminals: 2, direct-child terminals: 2
rigorous outgroups.: 11, direct-child rigorous outgroups: 3
(U_pictorum, U_crassus)


	  +-- U_crassus 2 
	--+               
	  +-- U_pictorum 1
  

Sonowal et al., 2021

fig. 3a: [clade_id: 20140] "Unio" + Anodonta + Aculamprotula
spp. terminals: 2, direct-child terminals: 2
rigorous outgroups.: 1, direct-child rigorous outgroups: 1
(A_cygnea, U_ravoisieri, U_pictorum, A_tientsinensis)


	  +-- A_cygnea 1       
	  |                    
	  +-- A_tientsinensis 4
	--+                    
	  +-- U_pictorum 3     
	  |                    
	  +-- U_ravoisieri 2   
  

fig. 3b: [clade_id: 20168] "Unio" + Lamellidens + Elliptio + Potomida + Lampsilis + Anodonta + Lamprotula + Aculamprotula
spp. terminals: 2, direct-child terminals: 2
rigorous outgroups.: 1, direct-child rigorous outgroups: 1
(A_tientsinensis, L_leaii, L_cardium, P_littoralis, U_pictorum, A_cygnea, U_ravoisieri, L_phenchooganjensis, E_complanata)


	  +-- A_cygnea 6           
	  |                        
	  +-- A_tientsinensis 1    
	  |                        
	  +-- E_complanata 9       
	  |                        
	  +-- L_cardium 3          
	  |                        
	--+-- L_leaii 2            
	  |                        
	  +-- L_phenchooganjensis 8
	  |                        
	  +-- P_littoralis 4       
	  |                        
	  +-- U_pictorum 5         
	  |                        
	  +-- U_ravoisieri 7       
  

fig. 4: [clade_id: 20185] "Unio" + Lamellidens + Anodonta
spp. terminals: 2, direct-child terminals: 2
rigorous outgroups.: 1, direct-child rigorous outgroups: 1
(U_pictorum, (A_cygnea, (U_ravoisieri, L_corrianus)))


	  +-------- U_pictorum 1  
	--+                       
	  |  +----- A_cygnea 2    
	  +--+                    
	     |  +-- L_corrianus 4 
	     +--+                 
	        +-- U_ravoisieri 3
  

Hoeh et al., 2009

fig. 3: [clade_id: 4219] Unio
spp. terminals: 2, direct-child terminals: 2
rigorous outgroups.: 0, direct-child rigorous outgroups: 0
(U_pictorum, U_caffer)


	  +-- U_caffer 2  
	--+               
	  +-- U_pictorum 1
  

fig. 4: [clade_id: 4302] Unio
spp. terminals: 2, direct-child terminals: 2
rigorous outgroups.: 0, direct-child rigorous outgroups: 0
(U_pictorum, U_caffer)


	  +-- U_caffer 2  
	--+               
	  +-- U_pictorum 1
  

fig. 5: [clade_id: 4389] Unio
spp. terminals: 2, direct-child terminals: 2
rigorous outgroups.: 0, direct-child rigorous outgroups: 0
(U_pictorum, U_caffer)


	  +-- U_caffer 2  
	--+               
	  +-- U_pictorum 1
  

Bolotov et al., 2016

fig. 2: [clade_id: 7775] Unio
spp. terminals: 2, direct-child terminals: 2
rigorous outgroups.: 0, direct-child rigorous outgroups: 0
(U_pictorum, Unio)


	  +-- U_pictorum 1
	--+               
	  +-- Unio 2      
  

fig. 4: [clade_id: 7830] Unio
spp. terminals: 2, direct-child terminals: 2
rigorous outgroups.: 0, direct-child rigorous outgroups: 0
(U_pictorum, Unio)


	  +-- U_pictorum 1
	--+               
	  +-- Unio 2      
  

Burzynski et al., 2017

fig. 1 bottom: [clade_id: 8293] Unio
spp. terminals: 2, direct-child terminals: 2
rigorous outgroups.: 1, direct-child rigorous outgroups: 1
(U_crassus, U_tumidus)


	  +-- U_crassus 1
	--+              
	  +-- U_tumidus 2
  

fig. 1 top: [clade_id: 8268] Unio
spp. terminals: 2, direct-child terminals: 2
rigorous outgroups.: 1, direct-child rigorous outgroups: 1
(U_crassus, U_tumidus)


	  +-- U_crassus 1
	--+              
	  +-- U_tumidus 2
  

Fonseca et al., 2016

fig. 1: [clade_id: 7941] Unio
spp. terminals: 2, direct-child terminals: 2
rigorous outgroups.: 5, direct-child rigorous outgroups: 4
(U_pictorum, U_delphinus)


	  +-- U_delphinus 2
	--+                
	  +-- U_pictorum 1 
  

fig. 1-M: [clade_id: 7983] Unio
spp. terminals: 1, direct-child terminals: 1
rigorous outgroups.: 0, direct-child rigorous outgroups: 0
unknown

Lopes-Lima et al., 2017

fig. 2: [clade_id: 8548] Unio
spp. terminals: 2, direct-child terminals: 2
rigorous outgroups.: 5, direct-child rigorous outgroups: 4
(U_delphinus, U_pictorum)


	  +-- U_delphinus 1
	--+                
	  +-- U_pictorum 2 
  

fig. 2-M: [clade_id: 8592] Unio
spp. terminals: 1, direct-child terminals: 1
rigorous outgroups.: 0, direct-child rigorous outgroups: 0
unknown

Wu et al., 2018

fig. 2: [clade_id: 10471] Unio
spp. terminals: 2, direct-child terminals: 2
rigorous outgroups.: 14, direct-child rigorous outgroups: 6
(U_crassus, U_pictorum)


	  +-- U_crassus 1 
	--+               
	  +-- U_pictorum 2
  

Chase et al., 2018

fig. 4: [clade_id: 12846] Unio
spp. terminals: 2, direct-child terminals: 2
rigorous outgroups.: 0, direct-child rigorous outgroups: 0
(U_delphinus, U_tumidus)


	  +-- U_delphinus 1
	--+                
	  +-- U_tumidus 2  
  

fig. 5: [clade_id: 12899] Unio
spp. terminals: 2, direct-child terminals: 2
rigorous outgroups.: 0, direct-child rigorous outgroups: 0
(U_delphinus, U_tumidus)


	  +-- U_delphinus 1
	--+                
	  +-- U_tumidus 2  
  

Whelan et al., 2011

fig. 1 ML: [clade_id: 5118] Unio
spp. terminals: 2, direct-child terminals: 2
rigorous outgroups.: 0, direct-child rigorous outgroups: 0
(U_pictorum, U_caffer)


	  +-- U_caffer 2  
	--+               
	  +-- U_pictorum 1
  

fig. 1 MP: [clade_id: 5045] Unio
spp. terminals: 2, direct-child terminals: 2
rigorous outgroups.: 0, direct-child rigorous outgroups: 0
(U_pictorum, U_caffer)


	  +-- U_caffer 2  
	--+               
	  +-- U_pictorum 1
  

Graf & Cummings, 2006

fig. 2: [clade_id: 184] Unio
spp. terminals: 2, direct-child terminals: 2
rigorous outgroups.: 0, direct-child rigorous outgroups: 0
(U_pictorum, U_caffer)


	  +-- U_caffer 2  
	--+               
	  +-- U_pictorum 1
  

Riccardi et al., 2019

fig. 2: [clade_id: 19443] Unio
spp. terminals: 2, direct-child terminals: 2
rigorous outgroups.: 0, direct-child rigorous outgroups: 0
(U_mancus, U_elongatulus)


	  +-- U_elongatulus 2
	--+                  
	  +-- U_mancus 1     
  

Graf et al., 2014

fig. 3: [clade_id: 6926] Unio
spp. terminals: 1, direct-child terminals: 1
rigorous outgroups.: 0, direct-child rigorous outgroups: 0
unknown

Marshall et al., 2014

fig. 1: [clade_id: 6818] Unio
spp. terminals: 1, direct-child terminals: 1
rigorous outgroups.: 0, direct-child rigorous outgroups: 0
unknown

Graf et al., 2015

fig. 1: [clade_id: 7018] Unio
spp. terminals: 1, direct-child terminals: 1
rigorous outgroups.: 0, direct-child rigorous outgroups: 0
unknown

Pfeiffer & Graf, 2015

fig. 2: [clade_id: 7255] Unio
spp. terminals: 1, direct-child terminals: 1
rigorous outgroups.: 0, direct-child rigorous outgroups: 0
unknown

Bieler et al., 2014

fig. 26, 27, 29, 30: [clade_id: 6736] Unio
spp. terminals: 1, direct-child terminals: 1
rigorous outgroups.: 0, direct-child rigorous outgroups: 0
unknown

fig. 28: [clade_id: 6745] Unio
spp. terminals: 1, direct-child terminals: 1
rigorous outgroups.: 0, direct-child rigorous outgroups: 0
unknown

Huang et al., 2013

fig. 3: [clade_id: 6649] Unio
spp. terminals: 1, direct-child terminals: 1
rigorous outgroups.: 0, direct-child rigorous outgroups: 0
unknown

Wu et al., 2018

fig. 2: [clade_id: 18726] Unio
spp. terminals: 1, direct-child terminals: 1
rigorous outgroups.: 5, direct-child rigorous outgroups: 2
unknown

González & Giribet, 2015

fig. 2: [clade_id: 7062] Unio
spp. terminals: 1, direct-child terminals: 1
rigorous outgroups.: 0, direct-child rigorous outgroups: 0
unknown

fig. 3: [clade_id: 7071] Unio
spp. terminals: 1, direct-child terminals: 1
rigorous outgroups.: 0, direct-child rigorous outgroups: 0
unknown

fig. 4: [clade_id: 7076] Unio
spp. terminals: 1, direct-child terminals: 1
rigorous outgroups.: 0, direct-child rigorous outgroups: 0
unknown

fig. 6: [clade_id: 7088] Unio
spp. terminals: 1, direct-child terminals: 1
rigorous outgroups.: 0, direct-child rigorous outgroups: 0
unknown

Wu et al., 2020

fig. 3: [clade_id: 18791] Unio
spp. terminals: 1, direct-child terminals: 1
rigorous outgroups.: 9, direct-child rigorous outgroups: 4
unknown

fig. 4: [clade_id: 18833] Unio
spp. terminals: 1, direct-child terminals: 1
rigorous outgroups.: 4, direct-child rigorous outgroups: 1
unknown

Bolotov et al., 2015

fig. 2A: [clade_id: 7040] Unio
spp. terminals: 1, direct-child terminals: 1
rigorous outgroups.: 0, direct-child rigorous outgroups: 0
unknown

fig. 2B: [clade_id: 7057] Unio
spp. terminals: 1, direct-child terminals: 1
rigorous outgroups.: 0, direct-child rigorous outgroups: 0
unknown

Choi et al., 2020

fig. 2a: [clade_id: 18918] Unio
spp. terminals: 1, direct-child terminals: 1
rigorous outgroups.: 10, direct-child rigorous outgroups: 3
unknown

fig. 3a: [clade_id: 18943] Unio
spp. terminals: 1, direct-child terminals: 1
rigorous outgroups.: 4, direct-child rigorous outgroups: 2
unknown

Xue et al., 2016

fig. 1: [clade_id: 7106] Unio
spp. terminals: 1, direct-child terminals: 1
rigorous outgroups.: 3, direct-child rigorous outgroups: 3
unknown

Graf, 2000

fig. 1: [clade_id: 1174] Unio
spp. terminals: 1, direct-child terminals: 1
rigorous outgroups.: 0, direct-child rigorous outgroups: 0
unknown

Graf & Ó Foighil, 2000

fig. 1, 2: [clade_id: 1194] Unio
spp. terminals: 1, direct-child terminals: 1
rigorous outgroups.: 0, direct-child rigorous outgroups: 0
unknown

Chong et al., 2008

fig. 3: [clade_id: 3697] Unio
spp. terminals: 1, direct-child terminals: 1
rigorous outgroups.: 0, direct-child rigorous outgroups: 0
unknown

Zhou et al., 2016

fig. 1: [clade_id: 7434] Unio
spp. terminals: 1, direct-child terminals: 1
rigorous outgroups.: 2, direct-child rigorous outgroups: 2
unknown

Wu et al., 2016

fig. 1: [clade_id: 7373] Unio
spp. terminals: 1, direct-child terminals: 1
rigorous outgroups.: 4, direct-child rigorous outgroups: 2
unknown

Wang et al., 2016

fig. 2: [clade_id: 7353] Unio
spp. terminals: 1, direct-child terminals: 1
rigorous outgroups.: 0, direct-child rigorous outgroups: 0
unknown

Taylor et al., 2007

fig. 1-3: [clade_id: 3620] Unio
spp. terminals: 1, direct-child terminals: 1
rigorous outgroups.: 0, direct-child rigorous outgroups: 0
unknown

Rosenberg et al., 1994

fig. 3: [clade_id: 855] Unio
spp. terminals: 1, direct-child terminals: 1
rigorous outgroups.: 0, direct-child rigorous outgroups: 0
unknown

Rosenberg et al., 1997

fig. 2A: [clade_id: 1022] Unio
spp. terminals: 1, direct-child terminals: 1
rigorous outgroups.: 0, direct-child rigorous outgroups: 0
unknown

fig. 2B: [clade_id: 1051] Unio
spp. terminals: 1, direct-child terminals: 1
rigorous outgroups.: 1, direct-child rigorous outgroups: 1
unknown

Soroka & Burzynski, 2018

fig. 3: [clade_id: 10178] Unio
spp. terminals: 1, direct-child terminals: 1
rigorous outgroups.: 1, direct-child rigorous outgroups: 1
unknown

fig. 3-M: [clade_id: 10191] Unio
spp. terminals: 1, direct-child terminals: 1
rigorous outgroups.: 1, direct-child rigorous outgroups: 1
unknown

Guerra et al., 2017

fig. 2a: [clade_id: 9781] Unio
spp. terminals: 1, direct-child terminals: 1
rigorous outgroups.: 1, direct-child rigorous outgroups: 1
unknown

fig. 2b: [clade_id: 9846] Unio
spp. terminals: 1, direct-child terminals: 1
rigorous outgroups.: 1, direct-child rigorous outgroups: 1
unknown

Wen et al., 2017

fig. 3: [clade_id: 8365] Unio
spp. terminals: 1, direct-child terminals: 1
rigorous outgroups.: 3, direct-child rigorous outgroups: 2
unknown

Araujo et al., 2017

fig. 2: [clade_id: 9751] Unio
spp. terminals: 1, direct-child terminals: 1
rigorous outgroups.: 2, direct-child rigorous outgroups: 2
unknown

Combosch et al., 2017

fig. 1: [clade_id: 9649] Unio
spp. terminals: 1, direct-child terminals: 1
rigorous outgroups.: 0, direct-child rigorous outgroups: 0
unknown

fig. 2: [clade_id: 9674] Unio
spp. terminals: 1, direct-child terminals: 1
rigorous outgroups.: 0, direct-child rigorous outgroups: 0
unknown

Lopes-Lima et al., 2018

fig. 1: [clade_id: 10805] Unio
spp. terminals: 1, direct-child terminals: 1
rigorous outgroups.: 0, direct-child rigorous outgroups: 0
unknown

fig. 3: [clade_id: 10858] Unio
spp. terminals: 1, direct-child terminals: 1
rigorous outgroups.: 0, direct-child rigorous outgroups: 0
unknown

Araujo et al., 2009

fig. 3: [clade_id: 4173] Unio
spp. terminals: 1, direct-child terminals: 1
rigorous outgroups.: 0, direct-child rigorous outgroups: 0
unknown

Pfeiffer et al., 2019

fig. 4-5: [clade_id: 15014] Unio
spp. terminals: 1, direct-child terminals: 1
rigorous outgroups.: 2, direct-child rigorous outgroups: 2
unknown

fig. 6: [clade_id: 15107] Unio
spp. terminals: 1, direct-child terminals: 1
rigorous outgroups.: 2, direct-child rigorous outgroups: 2
unknown

Gomes-dos-Santos et al., 2019

fig. 1: [clade_id: 15787] Unio
spp. terminals: 1, direct-child terminals: 1
rigorous outgroups.: 0, direct-child rigorous outgroups: 0
unknown

fig. 1-M: [clade_id: 15798] Unio
spp. terminals: 1, direct-child terminals: 1
rigorous outgroups.: 0, direct-child rigorous outgroups: 0
unknown

Pfeiffer & Graf, 2013

fig. 2: [clade_id: 6708] Unio
spp. terminals: 1, direct-child terminals: 1
rigorous outgroups.: 7, direct-child rigorous outgroups: 3
unknown

Klunzinger et al., 2020

fig. 2: [clade_id: 16325] Unio
spp. terminals: 1, direct-child terminals: 1
rigorous outgroups.: 0, direct-child rigorous outgroups: 0
unknown

Page updated: 09:22:17 Tue 04 Jan 2022

 
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