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The Freshwater Mussels (Unionoida) of the World (and other less consequential bivalves)
FM(U)otW(aolcb) is the web version of the MUSSEL Project Database. Follow the links to browse the data or use the custom Google search field. Either way, you win!
This page documents data for Haas (1936), such as genera and species attributed to that publication (including known misspellings, misidentifications, and other unavailable names) as well as taxonomic opinions, if available. Taxonomic opinions refer to uses of genera and species as valid and invalid in that work.
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Publication information
Haas, F. 1936. Binnen-Mollusken aus Inner-Afrika, hauptsächlich gesammelt von Dr. F. Haas während der Schomburgk-Expedition in der Jahren 1931/32. Abhandlungen der Senckenbergischen Naturforschenden Gesellschaft 431: 1-156, 8 pls.
Attributed Genus Names
Caelatura Conrad, 1853
‘Conrad’ of Authors.
Conrad, 1853, Proc. Acad. Nat. Sci. 6: 267 [as ‘Cælatura’].
Simpson, 1900, Proc. U.S. Nat. Mus. 22: 820 [as ‘Cælatura’].
Pallary, 1909, Mém. Inst. Égypte 6(1): 76.
Simpson, 1914, Descr. Cat. Naiades: 1019.
Germain, 1918, Bull. Mus. Hist. Nat. 24: 41.
Germain, 1919, Bull. Mus. Hist. Nat. 25: 122.
Germain, 1921, Faun. malac. Terr. e Fluv. Iles Mascareignes: 403.
Pallary, 1924, Mém. Inst. Égypte 7 (n.s.): 42.
Pilsbry & Bequaert, 1927, Bull. AMNH 53: 399.
Haas, 1927, Senckenbergiana 9: 21.
Haas, 1929, Senckenbergiana 11: 111.
Bloomer, 1932, Proc. Malac. Soc. London 20: 168.
Thiele, 1934, Handb. Syst. Weichtierk. 3: 1248.
Haas, 1936, Abh. Senckenb. Naturf. Ges. 431: 54.
Dartevelle & Schwetz, 1937, Ann. Soc. Roy. Zool. Belgique 68: 60.
Haas, 1940, Zool. Ser. Field Mus. Nat. Hist. 24: 140.
Modell, 1942, Arch. Moll. 74(5/6): 190.
Bloomer, 1946, Proc. Malac. Soc. London 27: 68.
Bloomer, 1949, Proc. Malac. Soc. London 27: 241.
Franc, 1949, J. de Conch. 9: 158.
Leloup, 1950, Res. Sci. Explor. Hydrob. Lac Tanganyika (1946-1947) 3: 40.
Mongin, 1963, Mém. Soc. Géol. France 96: 10.
Modell, 1964, Arch. Moll. 93: 116.
Mandahl-Barth, 1968, Expl. Hydrobiol. Bangweulu-Luapula 12: 51.
Starobogatov, 1970, Fauna Moll. Continent.: 64.
Fischer-Piette & Métivier, 1974, Bull. Mus. Hist. Nat. 133: 10.
Appleton, 1979, Ann. S. Afr. Mus. 77(9): 154.
Kenmuir, 1980, Trans. Zimbabwe Sci. Assoc. 60(2): 7.
Curtis & Appleton, 1987, Jour. SWA Sci. Soc. 40-41: 50.
Ibrahim et al., 1999, EEAA Nat. Biodiv. Unit (10): 74.
Millard, 2001, Class. Moll. (3): 1061.
(incorrect original spelling of Coelatura Conrad, 1853)
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Kalliphenga Haas, 1936
Haas, 1936, Abh. Senckenb. Naturf. Ges. 431: 72.
(intro'd as sub-gen. of Coelatura Conrad, 1853)
Type species: Unio ruellani Bourguignat, 1883 (orig. desig.) = Coelatura hauttecoeuri |
Kistinaia Haas, 1936
Haas, 1936, Abh. Senckenb. Naturf. Ges. 431: 64.
(intro'd as sub-gen. of Coelatura Conrad, 1853)
Type species: Caelatura (Kistinaia) schoutedeni Haas, 1936 (orig. desig.) = Prisodontopsis mweruensis |
Mweruella Haas, 1936
Haas, 1936, Abh. Senckenb. Naturf. Ges. 431: 63 [as ‘Mweruëlla’].
(intro'd as sub-gen. of Coelatura Conrad, 1853)
Type species: Unio mweruensis E.A. Smith, 1908 (orig. desig.) = Prisodontopsis mweruensis |
Rhytidonaia Haas, 1936
Haas, 1936, Abh. Senckenb. Naturf. Ges. 431: 65.
(intro'd as sub-gen. of Coelatura Conrad, 1853)
Type species: Caelatura graueri Haas, 1927 (orig. desig.) = Coelatura leopoldvillensis |
Mesafra Haas, 1936
Haas, 1936, Abh. Senckenb. Naturf. Ges. 431: 83.
Type species: Caelatura (s.s.) mesafricana Pilsbry & Bequaert, 1927 (orig. desig.) = Coelatura mesafricana |
Afroparreysia Haas, 1936
Haas, 1936, Abh. Senckenb. Naturf. Ges. 431: 84.
Type species: Parreysia lobensis Frierson, 1913 (orig. desig.) = Coelatura lobensis |
Nyassunio Haas, 1936
Haas, 1936, Abh. Senckenb. Naturf. Ges. 431: 85.
Type species: Unio nyassaensis Lea, 1864 (orig. desig.) = Nyassunio nyassaensis |
Attributed Species Names
Parreysia hypsiprimnus Simpson, 1900
‘von Martens’ of Authors.
Simpson, 1900, Proc. U.S. Nat. Mus. 22: 849.
Simpson, 1914, Descr. Cat. Naiades: 1136.
Haas, 1936, Abh. Senckenb. Naturf. Ges. 431: 68.
Woodward in Crowley et al., 1964, Annals Mus. R. Afr. Cent. 131: 47.
(error for Unio hypsiprymnus Martens, 1897)
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Unio billotianus Simpson, 1900
‘Charmes’ of Authors.
Simpson, 1900, Proc. U.S. Nat. Mus. 22: 847.
Kobelt, 1910, Abh. Senckenb. Naturf. Ges. 32: 84.
Simpson, 1914, Descr. Cat. Naiades: 1130.
Haas, 1936, Abh. Senckenb. Naturf. Ges. 431: 52.
Haas, 1969, Das Tierreich 88: 190.
Daget, 1998, Cat. Bivalves Africains: 65.
(error for Unio billottianus Charmes, 1885)
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Caelatura (s.s.) choziensis bangweolica Haas, 1936
Haas, 1936, Abh. Senckenb. Naturf. Ges. 431: 56, pl. 5, figs. 1a-d.
(intro'd as intra-sp. of Unio choziensis Preston, 1910)
Type(s): HOLOTYPE SMF_9576, Nordende des Bangweolo See, bei Nsombo, N. Rhodesien.
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Aspatharia (Spathopsis) dautzenbergi Haas, 1936
Haas, 1936, Abh. Senckenb. Naturf. Ges. 431: 93, pl. 8, figs. 2a-b.
Type(s): HOLOTYPE SMF_9545, Chambesi, bei Mbesuma, N. Rhodesia.
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Sphaerium hartmanni var. congicum Haas, 1936
Haas, 1936, Abh. Senckenb. Naturf. Ges. 431: 46.
(intro'd as intra-sp. of Cyclas hartmanni Jickeli, 1874; new name for Sphaerium stuhlmanni Pilsbry & Bequaert, 1927)
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Corbicula albida rosini Haas, 1936
Haas, 1936, Abh. Senckenb. Naturf. Ges. 431: 41, pl. 3, fig. 2.
(intro'd as intra-sp. of Cyrena africana var. albida Krauss, 1848)
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Corbicula albida albida Haas, 1936
Haas, 1936, Abh. Senckenb. Naturf. Ges. 431: 41, pl. 2, fig. 5.
(misidentified reference to Cyrena africana var. albida Krauss, 1848)
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Caelatura choziensis luapulaensis Haas, 1936
‘Preston’ Haas, 1936, Abh. Senckenb. Naturf. Ges. 431: pl. 5, fig. 2e. [in part]
(misidentified reference to Unio (Nodularia) luapulaensis Preston, 1913)
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Mesafra mesafricana stappersi Haas, 1936
Haas, 1936, Abh. Senckenb. Naturf. Ges. 431: 84, pl. 7, figs. 2a-b.
(intro'd as intra-sp. of Caelatura (s.s.) mesafricana Pilsbry & Bequaert, 1927)
Type(s): HOLOTYPE MRAC_56762, Riv. Luapula: Kasenga.
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Sphaerium hartmanni bangweolicum Haas, 1936
Haas, 1936, Abh. Senckenb. Naturf. Ges. 431: 46, pl. 3, figs. 3a-c.
(intro'd as intra-sp. of Cyclas hartmanni Jickeli, 1874)
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Caelatura (Zairia) gabunensis Haas, 1936
‘Küster’ Haas, 1936, Abh. Senckenb. Naturf. Ges. 431: 68.
(error for Unio gabonensis Küster, 1862)
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Corbicula astartina Haas, 1936
Haas, 1936, Abh. Senckenb. Naturf. Ges. 431: 42.
(misidentified reference to Cyrena astartina Martens, 1860)
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Caelatura (Kistinaia) schoutedeni Haas, 1936
Haas, 1936, Abh. Senckenb. Naturf. Ges. 431: 64, pl. 5, figs. 4a-d.
Haas, 1969, Treatise on Invert. Paleo: N418 [as ‘shoutedeni’].
Type(s): TYPE MRAC_56145, Riv. Luapula a Kasenga.
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Pisidium costulosum Haas, 1936
Haas, 1936, Abh. Senckenb. Naturf. Ges. 431: 43.
(misidentified reference to Pisidium costulosum Connolly, 1931)
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Mutela hargeri schomburgki Haas, 1936
Haas, 1936, Abh. Senckenb. Naturf. Ges. 431: 98, pl. 8, figs. 3a-e.
(intro'd as intra-sp. of Mutela hargeri E.A. Smith, 1908)
Type(s): TYPE SMF_9552, Nordende des Bangweolo See, bei Nsombo, N. Rhodesien.
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Taxonomic Opinions
Afroparreysia lobensis (Frierson, 1913)
Afroparreysia nyangensis (Frierson, 1913)
Aspatharia (Spathopsis) chaiziana (Rang, 1835)
Aspatharia (Spathopsis) dautzenbergi n. sp.
Aspatharia (Spathopsis) hartmanni bourguignati (Ancey)
+ Spathella bloyeti Bourguignat, 1889
+ Spathella spathuliformis Bourguignat, 1889
Aspatharia (Spathopsis) hartmanni guillaini (Récluz, 1850)
Aspatharia (Spathopsis) hartmanni haasi (Pilsbry, 1923)
Aspatharia (Spathopsis) hartmanni hartmanni (von Martens, 1866)
+ Spatha marnoi Jickeli, 1874
Aspatharia (Spathopsis) petersi (von Martens, 1860)
Aspatharia (Spathopsis) wahlbergi baumanni (Sturany in Baumann, 1894)
Aspatharia (Spathopsis) wahlbergi tabula (Sowerby, 1867)
Aspatharia (Spathopsis) wahlbergi wahlbergi (Krauss, 1848)
+ Spatha wahlbergi var. dorsalis Martens, 1897
Aspatharia (Spathopsis) wahlbergi welwitschii (Morelet, 1868)
Aspatharia (s.s.) pfeifferiana (Bernardi, 1860)
Caelatura [sic] (Grandidieria) burtoni (Woodward, 1859)
+ Unio thomsoni E.A. Smith, 1880
+ Grandidieria cyrenopsis Bourguignat, 1885
+ Grandidieria servainiana Bourguignat, 1885
+ Grandidieria giraudi Bourguignat, 1885
+ Grandidieria gravida Bourguignat, 1885
+ Grandidieria rostrata Bourguignat, 1885
+ Grandidieria incarnata Bourguignat, 1885
+ Grandidieria mira Bourguignat, 1885
+ Grandidieria rotundata Bourguignat, 1885
+ Grandidieria smithi Bourguignat, 1885
+ Grandidieria corbicula Bourguignat, 1885
+ Grandidieria granulosa Bourguignat, 1885
+ Grandidieria insignis Bourguignat, 1886
+ Grandidieria locardiana Bourguignat, 1886
+ Grandidieria hauttecoeuri Bourguignat, 1886
+ Grandidieria anceyi Bourguignat, 1886
+ Grandidieria callista Bourguignat, 1888
+ Grandidieria bourguignati Bourguignat, 1888
+ Grandidieria singularis Bourguignat, 1888
+ Unio (Grandidieria) burtoni var. sturanyi Martens, 1897
+ Unio rostralis Martens, 1897
+ Unio rostralis var. brevior Martens, 1897
+ Unio (Grandidieria) gravida var. lutescens Germain, 1905
Caelatura [sic] (Grandidieria) tanganyicensis (Smith, E.A., 1880)
+ Grandidieria elongata Bourguignat, 1885
+ Grandidieria rhynchonella Bourguignat, 1888
+ Unio tanganyicensis var. exalbida Preston, 1913
Caelatura [sic] (Kalliphenga) ruellani (Bourguignat, 1883)
+ Unio multicolor Martens, 1897
+ Unio (Parreysia) alluaudi Dautzenberg, 1908
+ Unio ruellani var. bayoni Germain, 1909
Caelatura [sic] (Kistinaia) schoutedeni n. sp.
Caelatura [sic] (Laevirostris) bourguignati (Rochebrune, 1886)
+ Unio stagnorum Dautzenberg, 1890
+ Caelatura (Laevirostris) stagnorum bomae Pilsbry & Bequaert, 1927
+ Caelatura (Laevirostris?) rotula Pilsbry & Bequaert, 1927
Caelatura [sic] (Laevirostris) briarti (Dautzenberg, 1901)
+ Unio eduardi Dautzenberg & Germain, 1914
Caelatura [sic] (Mweruella) mweruensis (Smith, E.A., 1908)
Caelatura [sic] (Rhytidonaia) graueri Haas, 1927
Caelatura [sic] (Rhytidonaia) leopoldvillensis (Putzeys, 1898)
Caelatura [sic] (Zairia) aequatoria (Morelet, 1885)
+ Unio landanensis Schepman, 1891
Caelatura [sic] (Zairia) elegans (Rochebrune, 1886)
+ Zairia disciformis Rochebrune, 1886
+ Zairia araneosa Rochebrune, 1886
+ Zairia poirieri Rochebrune, 1886
+ Zairia sordida Rochebrune, 1886
+ Unio (Nodularia) roubaudi Germain, 1907
+ Unio (Nodularia) subnigra Preston, 1909
+ Unio (Nodularia) putzeysi Preston, 1912
+ Caelatura (s.s.) stanleyvillensis Pilsbry & Bequaert, 1927
Caelatura [sic] (Zairia) essoënsis (Chaper, 1885)
Caelatura [sic] (Zairia) gabunensis [sic] (Küster, 1862)
Caelatura [sic] (s.s.) aegyptiaca (Cailliaud, 1827)
+ Unio niloticus Cailliaud, 1827
+ Unio parreyssi Philippi, 1848
+ Unio rugifer Küster, 1856
+ Unio eucyphus Bourguignat, 1857
+ Unio sennaariensis Küster, 1862
+ Unio sennariensis [sic] var. schweinfurthi Martens, 1873
+ Pharaonia misraimica Servain, 1890
+ Unio euryssellinus Pallary, 1903
+ Unio mysticus Pallary, 1903
+ Nodularia (Cælatura) gaillardoti Pallary, 1909
+ Nodularia (Cælatura) parreyssi var. pruneri Pallary, 1909
+ Nodularia parreyssi var. elongata Longstaff, 1914
+ Nodularia parreyssi var. obliqua Longstaff, 1914
+ Nodularia (Caelatura) sobaensis Preston, 1914
Caelatura [sic] (s.s.) bakeri (Adams, H., 1866)
Caelatura [sic] (s.s.) choziensis bangweolica n. subsp.
Caelatura [sic] (s.s.) choziensis choziensis (Preston, 1910)
Caelatura [sic] (s.s.) choziensis luapulaensis (Preston, 1913)
Caelatura [sic] (s.s.) hauttecoeuri emini (von Martens, 1897)
Caelatura [sic] (s.s.) hauttecoeuri hauttecoeuri (Bourguignat, 1883)
+ Unio edwardsianus Bourguignat, 1883
+ Unio grantianus Bourguignat, 1883
+ Unio duponti Bourguignat, 1883
+ Unio postumus Rochebrune, 1904
+ Unio hauttecoeuri mut. elongata Germain, 1906
+ Unio hauttecoeuri mut. intermedia Germain, 1906
+ Unio hauttecoeuri mut. curta Germain, 1906
+ Unio hauttecoeuri mut. globosa Germain, 1906
+ Unio hauttecoeuri mut. subcompressa Germain, 1906
+ Unio hauttecoeuri mut. compressa Germain, 1906
+ Unio hauttecoeuri var. lutescens Germain, 1906
+ Unio hauttecoeuri var. castanea Germain, 1906
+ Unio hauttecoeuri var. fusca Germain, 1906
+ Unio hauttecoeuri var. nigra Germain, 1906
+ Unio hauttecoeuri var. viridis Germain, 1906
+ Unio hauttecoeuri var. ornata Germain, 1907
Caelatura [sic] (s.s.) horei (Smith, E.A., 1880)
+ Unio calathus Bourguignat, 1886
+ Unio bridouxi Bourguignat, 1886
+ Unio charbonnieri Bourguignat, 1886
+ Unio coulboisi Bourguignat, 1886
+ Unio dromauxi Bourguignat, 1886
+ Unio guillemeti Bourguignat, 1886
+ Unio josseti Bourguignat, 1886
+ Unio jouberti Bourguignat, 1886
+ Unio lavigerianus Bourguignat, 1886
+ Unio menardi Bourguignat, 1886
+ Unio moineti Bourguignat, 1886
+ Unio randabeli Bourguignat, 1886
+ Unio vinckei Bourguignat, 1886
+ Unio visseri Bourguignat, 1886
+ Unio gerrardi Martens, 1897
+ Unio boehmi Martens, 1897
+ Unio gereti Preston, 1910
Caelatura [sic] (s.s.) hypsiprimna [sic] borellii (Ancey, 1894)
+ Unio angoniensis Preston, 1910
Caelatura [sic] (s.s.) hypsiprimna hypsiprimna [sic] (von Martens, 1897)
Caelatura [sic] (s.s.) introrugata (Connolly, 1931)
Caelatura [sic] (s.s.) rothschildi (Neuville & Anthony, 1906)
Caelatura [sic] (s.s.) stuhlmanni (von Martens, 1897)
+ Unio ngesianus Martens, 1897
+ Parreysia regis Pilsbry & Bequaert, 1927
Cafferia caffra (Krauss, 1848)
Etheria elliptica Lamarck, 1807
Indonaia kunenensis (Mousson, 1887)
+ Unio zambesiensis Preston, 1905
+ Nodularia (Caelatura) ellenbergeri Germain, 1920
+ Nodularia croninae Ortmann & Walker, 1922
+ Nodularia rohani Germain, 1925
Indonaia liederi (von Martens, 1897)
Indonaia mossambicensis (von Martens, 1860)
+ Unio lechaptoisi Ancey, 1894
+ Unio lechaptoisi var. minor Martens, 1897
+ Unio shireensis Preston, 1910
Indonaia ratidota (Charmes, 1885)
+ Unio dumesnilianus Charmes, 1885
+ Unio billottianus Charmes, 1885
+ Unio euphymus Charmes, 1885
+ Unio ledoulxianus Charmes, 1885
+ Unio ambifarius Martens, 1897
Mesafra mesafricana mesafricana (Pilsbry & Bequaert, 1927)
Mesafra mesafricana stappersi n. subsp.
Mutela hargeri schomburgki n. subsp.
Mutela nilotica mabilli (Rochebrune, 1886)
Nitia acuminata (Adams, H., 1866)
Nitia chefneuxi (Neuville & Anthony, 1906)
Nitia gaillardi (Germain, 1909)
Nitia monceti (Bourguignat, 1883)
+ Unio lourdeli Bourguignat, 1887
+ Unio monceti var. rubra Germain, 1906
+ Unio lourdeli var. smithi Germain, 1906
Nitia mutelaeformis (Germain, 1906)
+ Unio (s.s.) mutelaeformis var. chariensis Germain, 1906
Nitia teretiuscula (Philippi, 1847)
+ Margarita (Unio) cailliaudii Lea, 1836
+ Unio lithophagus Philippi, 1847
+ Nodularia (sec. Cafferia) jickeli Simpson, 1900
+ Unio fourtaui Pallary, 1903
+ Nodularia teretiuscula var. pallaryi Longstaff, 1914
Nyassunio nyassaensis (Lea, 1864)
+ Unio kirkii Lea, 1864
+ Unio aferula Lea, 1864
+ Unio hermosus Bourguignat, 1889
+ Unio (Nodularia) caesariana Preston, 1913
+ Unio nyassanus [sic] var. hercules Preston, 1913
Nyassunio ujijiensis (Crosse, 1881)
+ Unio nyassaensis var. tanganyicensis E.A. Smith, 1881
Unio abyssinicus von Martens, 1866
+ Unio schweinfurthi Martens, 1886
+ Unio willcocksi Newton, 1899
+ Unio vignardi Pallary, 1924
+ Unio fayumensis Pilsbry & Bequaert, 1927
Unio dembeae Sowerby, 1865
+ Unio aeneus Jickeli, 1874
+ Unio jickelii Bourguignat, 1883
+ Unio alfierianus Bourguignat, 1885
+ Unio hamyanus Bourguignat, 1885
+ Unio ilqi Bourguignat, 1885
+ Unio meneliki Bourguignat, 1885
+ Unio soleilleti Bourguignat, 1885
+ Unio traversii Pollonera, 1888
+ Unio (Nodularia) erlangeri Kobelt, 1910
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