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Page last updated
Wed 04 Dec 2024

The Freshwater Mussels (Unionoida) of the World (and other less consequential bivalves)

FM(U)otW(aolcb) is the web version of the MUSSEL Project Database. Follow the links to browse the data or use the custom Google search field. Either way, you win!

This page documents data for Haas (1969), such as genera and species attributed to that publication (including known misspellings, misidentifications, and other unavailable names) as well as taxonomic opinions, if available. Taxonomic opinions refer to uses of genera and species as valid and invalid in that work.

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Publication information

Haas, F. 1969. Superfamily Unionacea. Treatise on Invert. Paleo: N411-N470.

Attributed Genus Names

Symphinota Villa, 1841
of Authors.
Villa, 1841, Dispos. Syst. Conchyl.: 43.
‘Lea’ Haas, 1969, Das Tierreich 88: 415.
‘Lea’ Haas, 1969, Treatise on Invert. Paleo: N448.
(error for Symphynota Lea, 1829)

Megaptera Modell, 1964
‘Rafinesque’ of Authors.
‘Rafinesque’ Modell, 1964, Arch. Moll. 93: 85.
‘Rafinesque’ Haas, 1969, Treatise on Invert. Paleo: N455.
(error for Metaptera Rafinesque, 1820)

Lintoxia Haas, 1969
‘Rafinesque’ Haas, 1969, Treatise on Invert. Paleo: N439.
(error for Sintoxia Rafinesque, 1820)

Lyntoxia Haas, 1969
‘Rafinesque’ Haas, 1969, Treatise on Invert. Paleo: N439.
(error for Syntoxia Agassiz, 1846)

Chondrostea Haas, 1969
‘Gistl, 1848’ Haas, 1969, Treatise on Invert. Paleo: N415.
(error for Chondrostes Gistl, 1847)

Unionea Haas, 1969
‘Haas, 1955’ Haas, 1969, Treatise on Invert. Paleo: N435.
(error for Unionetta Haas, 1955)

Platyris Haas, 1969
‘Agassiz’ of Authors.
Haas, 1969, Das Tierreich 88: 598.
Haas, 1969, Treatise on Invert. Paleo: N464.
(error for Platyiris Agassiz, 1846)

Migranaia Haas, 1969
‘Hannibal’ of Authors.
Haas, 1969, Das Tierreich 88: 271.
Haas, 1969, Treatise on Invert. Paleo: N441.
Bank et al., 2001, Heldia 4(1/2): 113.
(error for Migranaja Hannibal, 1912)

Iridella Haas, 1969
‘Swainson’ Haas, 1969, Treatise on Invert. Paleo: N460.
(error for Iridea Swainson, 1840)

Propehyidella Haas, 1969
‘Cotton & Gabriel’ Haas, 1969, Treatise on Invert. Paleo: N460.
(error for Propehyridella Cotton & Gabriel, 1932)

Chevronaias Haas, 1969
‘Olsson & Wurtz’ Haas, 1969, Treatise on Invert. Paleo: N460.
(error for Chevronais Olsson & Wurtz, 1951)

Graptonaias Haas, 1969
Haas, 1969, Treatise on Invert. Paleo: N425.
(error for Graphonaias Fischer & Crosse, 1894)

Vanderschliea Haas, 1969
‘Modell’ of Authors.
Haas, 1969, Treatise on Invert. Paleo: N449.
Millard, 2001, Class. Moll. (3): 1060.
(error for Vanderschaliea Modell, 1943)

Attributed Species Names

Margaron (Unio) conradianus Lea, 1870
‘Lea’ Lea, 1870, Synopsis Fam. Naiades: 32.
‘Lea’ Pilsbry & Rhoads, 1896, Proc. Acad. Nat. Sci. 48: 501 [as ‘Unio conradianus’].
‘Lea’ Haas, 1969, Treatise on Invert. Paleo: N455 [as ‘Medionidus’].
(error for Unio conradicus Lea, 1834)

Unio sinensis Heude, 1875
“Lea” Heude, 1875, Conch. Fluv. Nanking (1): sp. 26.
‘Lea’ Haas, 1969, Treatise on Invert. Paleo: N416 [as ‘Acuticosta sinensis’].
‘Lea’ Millard, 2001, Class. Moll. (3): 1051.
(error for Unio chinensis Lea, 1868)

Columba blainvilleana Ihering, 1890
‘Lea’ of Authors.
Ihering, 1890, Arch. Naturg. 56(1): 142.
Simpson, 1900, Proc. U.S. Nat. Mus. 22: 915.
Simpson, 1914, Descr. Cat. Naiades: 1400.
Haas, 1931, Senckenbergiana 13: 102.
Morretes, 1949, Arqu. Mus. Paranaense 7: 28.
Bonetto, 1963, Physis 24: 11.
Parodiz & Bonetto, 1963, Malacologia 1(2): 206.
Modell, 1964, Arch. Moll. 93: 81.
Figueiras, 1965, Comm. Soc. Malac. Uruguay 1(8): 252.
Haas, 1969, Das Tierreich 88: 577.
Haas, 1969, Treatise on Invert. Paleo: N465.
Mansur, 1970, Iheringia 39: 55.
Bonetto & Mansur, 1970, Acta Zool. Lilloana 27: 256.
Millard, 2001, Class. Moll. (3): 1074.
Simone, 2006, Land. Freshw. Moll. Brazil.: 289.
(error for Anodonta blainvilliana Lea, 1834)

Dromus dromus Simpson, 1900
‘Lea’ of Authors.
Simpson, 1900, Proc. U.S. Nat. Mus. 22: 615.
Haas, 1930, Senckenbergiana 12(6): 317.
Haas, 1969, Treatise on Invert. Paleo: N452.
Stansbery, 1970, Malacologia 10(1): 18.
Millard, 2001, Class. Moll. (3): 1066.
Serb, 2006, J. Moll. Studies 72: 427.
(error for Unio dromas Lea, 1834)

Pressidens moellendorffi Haas, 1910
Haas, 1910, Nachr. Bl. Dtsch. Malak. Ges. 42: 103.
Haas, 1911, Conch. Cab. 9 (pt. 2, h. 44): pl. 23, figs. 3-5.
Haas, 1914, Conch. Cab. 9 (pt. 2, h. 49): 207.
Haas, 1969, Das Tierreich 88: 82.
Haas, 1969, Treatise on Invert. Paleo: N430.
Type(s): TYPE SMF_3600, Insel Palawan b. Borneo.

Unio verbeeki Haas, 1914
‘von Martens’ of Authors.
‘(Boettger) v. Martens’ Haas, 1914, Conch. Cab. 9 (pt. 2, h. 49): 198.
Modell, 1942, Arch. Moll. 74(5/6): 189.
‘Martens’ Haas, 1969, Das Tierreich 88: 101.
‘Martens’ Haas, 1969, Treatise on Invert. Paleo: N420.
‘(Martens)’ Brandt, 1974, Arch. Moll. 105: 289.
Millard, 2001, Class. Moll. (3): 1064.
(error for Unio verbecki Martens, 1897)

Caelatura (Kistinaia) schoutedeni Haas, 1936
Haas, 1936, Abh. Senckenb. Naturf. Ges. 431: 64, pl. 5, figs. 4a-d.
Haas, 1969, Treatise on Invert. Paleo: N418 [as ‘shoutedeni’].
Type(s): TYPE MRAC_56145, Riv. Luapula a Kasenga.

Caelatura shoutedeni Haas, 1969
‘Haas, 1936’ of Authors.
Haas, 1969, Treatise on Invert. Paleo: N418.
Millard, 2001, Class. Moll. (3): 1062.
(error for Caelatura (Kistinaia) schoutedeni Haas, 1936)

Castalioides laddei Haas, 1969
‘Marshall’ of Authors.
Haas, 1969, Treatise on Invert. Paleo: N459.
Millard, 2001, Class. Moll. (3): 1069.
(error for Castalioides laddi Marshall, 1934)

Anodontites cristatus Haas, 1969
‘Bruguière’ Haas, 1969, Treatise on Invert. Paleo: N463, fig. D54:5.
(error for Anodontites crispata Bruguière, 1792)

Lepidodesma languilata Haas, 1969
‘Heude’ of Authors.
Haas, 1969, Treatise on Invert. Paleo: N446.
Millard, 2001, Class. Moll. (3): 1058.
Huang et al., 2002, J. Moll. Studies 68: 360.
(error for Unio languilati Heude, 1874)

Taxonomic Opinions

Acostaea (Pseudomulleria) dalyi (Smith, E.A., 1898)

Acostaea (s.s.) lobata (Ferussac)

Actinonaias sapotalensis (Lea, 1841)

Acuticosta sinensis [sic] (Lea, 1868)

Afroparreysia lobensis (Frierson, 1913)

Alasmidonta (Bullella) arcula (Lea, 1838)

Alasmidonta (Decurambis) marginata Say, 1819
+ Decurambis scriptum Rafinesque, 1831

Alasmidonta (Pegias) fabula (Lea, 1838)

Alasmidonta (Pressodonta) calceola (Lea, 1828)

Alasmidonta (Prolasmidonta) heterodon (Lea, 1829)

Alasmidonta (s.s.) undulata (Say, 1817)

Amblema (Plectomerus) dombeyana (Valenciennes, 1827)

Amblema (Psorula) salinarum (Haas, 1929)

Amblema (s.s.) plicata costata Rafinesque, 1820

Anodonta (Arnoldina) dejecta Lewis, 1875

Anodonta (Brachyanodon) coarctata Anton, 1839
+ Anodonta chapalensis Crosse & Fischer, 1893

Anodonta (Gabillotia) pseudodopsis Locard, 1883

Anodonta (Haasiella) arcaeformis (Heude, 1877)

Anodonta (Liouvillea) pallaryi Bédé, 1932

Anodonta (Mesanodon) lurulenta Morelet, 1849

Anodonta (Pyganodon) globosa Lea, 1841

Anodonta (Utterbackiana) suborbiculata Say, 1831

Anodonta (s.s.) cygnea (Linné, 1758)

Anodontites (Lamproscapha) ensiformis (Spix, 1827)

Anodontites (Ruganodontites) colombiensis Marshall, 1922

Anodontites (s.s.) cristatus [sic] Bruguiere, 1792

Anodontoides ferussacianus (Lea, 1834)

Arcidens confragosus (Say, 1829)

Arcidopsis footei (Theobald, 1876)

Arconaia lanceolata (Lea, 1856)

Arkansia wheeleri Ortmann & Walker, 1912

Aspatharia (Arthropteron) ouassouloui (Rochebrune, 1904)

Aspatharia (Chambardia) rubens (Lamarck, 1819)

Aspatharia (s.s.) rugifera (Dunker, 1858)
+ Margaritana vignouana Bernardi, 1859

Balwantia soleniformis (Benson, 1836)

Bartlettia stefanensis (Moricand, 1856)

Caelatura [sic] (Brazzaea) anceyi (Bourguignat, 1885)

Caelatura [sic] (Grandidieria) burtoni (Woodward, 1859)

Caelatura [sic] (Kalliphenga) ruellani (Bourguignat, 1883)

Caelatura [sic] (Kistinaia) shoutedeni [sic] Haas, 1936

Caelatura [sic] (Laevirostris) bourguignati (Rochebrune, 1886)
+ Unio stagnorum Dautzenberg, 1890

Caelatura [sic] (Mweruella) mweruensis (Smith, E.A., 1908)

Caelatura [sic] (Rhytidonaia) graueri Haas, 1927

Caelatura [sic] (Zairia) elegans (Rochebrune, 1886)

Caelatura [sic] (s.s.) aegyptiaca (Cailliaud, 1827)

Cafferia caffra (Krauss, 1848)

Callonaia duprei (Récluz, 1842)

Canthyria spinosa (Lea, 1838)

Carunculina parva texasensis [sic] (Lea, 1857)

Castalia ambigua Lamarck, 1819

Castaliella sulcata (Krauss, 1849)

Castalina martensi von Ihering, 1891

Castalioides laddei [sic]

Caudiculatus caudiculatus (von Martens, 1866)

Chamberlainia hainesiana (Lea, 1856)

Chevronaias [sic] colombiana Olsson & Wurtz, 1951

Cokeria southalli Marshall, 1916

Conchodromus dromus [sic] (Lea, 1834)

Conradilla caelata (Conrad, 1834)

Contradens (Sprickia) verbeeki [sic] (‘Boettger’ von Martens, 1897)

Contradens (s.s.) contradens (Lea, 1838)

Costanaia arciformis MacNeil

Cristaria (Pletholophus) discoidea (Lea, 1834)

Cristaria (s.s.) plicata (Leach, 1815)

Ctenodesma borneense (Issel, 1874)

Cucumerunio novaehollandiae (Gray, 1834)

Cuneopsis celtiformis (Heude, 1874)

Cyclonaias tuberculata (Rafinesque, 1820)

Cyprogenia stegaria (Rafinesque, 1820)
+ Unio irroratus Lea, 1828

Diaurora aurorea (Heude, 1883)

Diplodon (Bulloideus) bulloides (Lea, 1859)

Diplodon (Rhipidodonta) paranensis (Lea, 1834)

Diplodon (Schleschiella) burroughianus (Lea, 1834)

Diplodon (s.s.) ellypticum Spix, 1827

Diplodontites cookei Marshall, 1922

Discomya radulosa (Drouet & Chaper, 1892)

Dysnomia (Capsaeformis) capsaeformis (Lea, 1834)

Dysnomia (Penita) penita (Conrad, 1834)

Dysnomia (Pilea) personata (Say, 1829)

Dysnomia (Scalenilla) sulcata (Lea, 1829)

Dysnomia (Torulosa) torulosa (Rafinesque, 1820)

Dysnomia (Truncillopsis) triquetra (Rafinesque, 1820)

Dysnomia (s.s.) flexuosa (Rafinesque, 1820)

Elliptio (Micronaias) aratus (Lea, 1843)

Elliptio (s.s.) crassidens (Lamarck, 1819)

Elongaria (Nannonaia) trompi (Drouet & Chaper, 1892)

Elongaria (s.s.) orientalis (Lea, 1840)

Ensidens (Uniandra) inaequalis (Rochebrune, 1882)

Ensidens (s.s.) ingallsianus (Lea, 1852)

Eonaias reynosenica Marshall

Etheria elliptica Lamarck, 1807
+ Etheria semilunata Lamarck, 1807

Fossula fossiculifera (d’Orbigny, 1835)

Friersonia iridella (Pilsbry & Frierson, 1907)

Fusconaia undata trigona (Lea, 1831)

Glebula rotundata (Lamarck, 1819)

Gonidea angulata (Lea, 1838)
+ Anodonta randalli Trask, 1854

Haasica balzani (von Ihering, 1893)

Haasodonta fannyae (Johnson, R.I., 1948)

Heudeana murina (Heude, 1883)

Hyridella (Lortiella) rugata (Sowerby, 1868)

Hyridella (Protohyridella) glenelgensis (Dennant, 1898)

Hyridella (Velesunio) ambigua (Philippi, 1847)
+ Unio balonnensis Conrad, 1850

Hyridella (s.s.) australis (Lamarck, 1819)
+ Unio napeanensis Conrad, 1850

Hyriopsis bialata Simpson, 1900
+ Unio delphinus Gruner, 1841

Iheringella isocardioides (Lea, 1856)

Inversidens brandti (Kobelt, 1879)

Iridina (Cameronia) spekei Woodward, 1859

Iridina (Pleiodon) ovata Swainson, 1823

Iridina (s.s.) exotica Lamarck, 1819

Lamellidens marginalis (Lamarck, 1819)

Lamprotula (Parunio) crassus (Ping, 1931)

Lamprotula (s.s.) plumbea (Chemnitz, 1795)

Lampsilis (Cyrtonaias) berlandieri (Lea, 1857)

Lampsilis (Delphinonaias) delphinulus (Morelet, 1849)

Lampsilis (Disconaias) discus (Lea, 1838)

Lampsilis (Mesonaias) explicata (Morelet, 1849)

Lampsilis (Ortmanniana) carinata (Barnes, 1823)

Lampsilis (Phyllonaias) paludosa (Morelet, 1849)

Lampsilis (Venustaconcha) venusta (Lea, 1838)

Lampsilis (Villosa) villosa (Wright, B.H., 1898)

Lampsilis (s.s.) ovata (Say, 1817)

Lanceolaria grayana (Lea, 1834)

Lasmigona (Platynaias) viridis (Rafinesque, 1820)
+ Symphynota compressa Lea, 1829

Lasmigona (Pterosyna) complanata (Barnes, 1823)

Lasmigona (Sulcularia) badia (Rafinesque, 1831)

Lasmigona (s.s.) costata (Rafinesque, 1820)
+ Alasmodonta rugosa Barnes, 1823

Lastena lata (Rafinesque, 1820)

Leguminaia (Microcondylaea) uniopsis (Lamarck, 1819)

Leguminaia (Pseudodontopsis) euphratica (Bourguignat, 1852)

Leguminaia (Pseudoleguminaia) chantrei Locard, 1883
+ Pseudodon chantrei Locard, 1883

Leguminaia (s.s.) mardinensis (Lea, 1865)

Leila blainvilleana [sic] (Lea, 1834)

Lepidodesma languilatum [sic] (Heude, 1874)

Leptanodonta rumana Wenz, 1941
+ Dreissenomya unionides Wenz, 1927

Leptodea leptodon (Rafinesque, 1820)

Lexingtonia subplana (Conrad, 1837)

Ligumia recta (Lamarck, 1819)

Loxopleurus belliplicatus (Meek)

Margaritifera (Cumberlandia) monodonta (Say, 1829)

Margaritifera (Margaritanopsis) laosensis (Lea, 1863)

Margaritifera (Pseudunio) auricularia (Spengler, 1793)
+ Unio sinuata Lamarck, 1819

Margaritifera (s.s.) margaritifera (Linné, 1758)

Medionidus conradianus [sic] (Lea, 1834)

Megalonaias gigantea (Barnes, 1823)
+ Unio heros Say, 1829

Megalonoidea porcata MacNeil

Mesafra mesafricana (Pilsbry & Bequaert, 1927)

Micromya fabalis (Lea, 1831)

Monocondylaea paraguayana d’Orbigny, 1835

Mutela (Chelidonopsis) hirundo (von Martens, 1881)
+ Chelidonura arietina Rochebrune, 1886

Mutela (Moncetia) anceyi (Bourguignat, 1885)

Mutela (s.s.) dubia (Gmelin, 1791)

Mycetopoda soleniformis d’Orbigny, 1835

Mycetopodella falcatus (Higgins, 1868)

Nephronaias (Elliptoideus) sloatiana (Lea, 1840)

Nephronaias (Martensnaias) rubicunda (von Martens, 1900)

Nephronaias (Nephritica) poeyanus (Lea, 1857)

Nephronaias (Popenaias) popeii (Lea, 1857)

Nephronaias (Reticulatus) reticulata Simpson, 1900

Nephronaias (Sphenonaias) liebmanni (Philippi, 1847)

Nephronaias (s.s.) plicatula (Charpentier in Küster, 1856)

Nippononaia ryosekiana (Suzuki)

Nitia teretiuscula (Philippi, 1847)

Obliquaria reflexa Rafinesque, 1820

Obovaria (Pseudoon) olivaria (Rafinesque, 1820)
+ Unio ellipsis Lea, 1828

Obovaria (s.s.) retusa (Lamarck, 1819)

Oxynaia jourdyi (Morlet, 1886)

Pachynaias spheniopsis (Morelet, 1849)

Palindonaia solandri (Sowerby, 1826)

Paranodonta ôtai Kobayashi & Suzuki, 1936

Parreysia (Nyassunio) nyassaensis (Lea, 1864)

Parreysia (Radiatula) crispisulcata (Benson, 1862)

Parreysia (s.s.) corrugata (Müller, 1774)

Paxyodon alatus (Sowerby, 1869)

Paxyodon syrmatophora (Gronovius, 1781)
+ Paxyodon ponderosus Schumacher, 1817

Physunio (Lens) eximius (Lea, 1856)

Physunio (Velunio) velaris (Sowerby, 1868)

Physunio (s.s.) gravidus (Lea, 1856)

Pilsbryoconcha exilis (Lea, 1838)

Plagiola depressa (Rafinesque, 1820)
+ Unio securis Lea, 1829

Plethobasus cyphyus (Rafinesque, 1820)
+ Unio aesopus Green, 1827

Pleurobema mytiloides Rafinesque, 1820

Plicatounio naktongensis Kobayashi & Suzuki, 1936

Pliconaias popenoei Marshall

Potomida (Cuneopsidea) sculpta (Brusina, 1874)

Potomida (s.s.) littoralis semirugata (Lamarck, 1819)
+ Mysca (Potomida) corrugata Swainson, 1840

Pressidens moellendorffi Haas, 1910

Prisodon obliquus Schumacher, 1817

Prisodontopsis johnstoni (Smith, E.A., 1893)

Prohyriopsis stolata (von Martens, 1900)

Proparreysia percorrugata (Whitfield)

Proptera alata (Say, 1817)

Protelliptio (Plesielliptio) priscus (Meek & Hayden)

Protelliptio (s.s.) biornatus (Russell)

Protunio messageri (Bavay & Dautzenberg, 1901)

Pseudanodonta complanata (Rossmässler, 1835)

Pseudobaphia biesiana (Heude, 1877)

Pseudodon (Bineurus) mouhoti (Lea, 1863)

Pseudodon (Chrysopseudodon) aureus Heude, 1885

Pseudodon (Cosmopseudodon) resupinatus von Martens, 1902

Pseudodon (Diplopseudodon) crassus Drouet, 1892

Pseudodon (Indopseudodon) salwenianus (Gould, 1844)

Pseudodon (Monodontina) vondembuschianus (Lea, 1840)

Pseudodon (Nasus) nankingensis (Heude, 1874)

Pseudodon (Obovalis) loomisi Simpson, 1900

Pseudodon (Trigonodon) crebristriatus (Anthony, J.G., 1865)

Pseudodon (s.s.) inoscularis (Gould, 1844)

Pseudospatha tanganyicensis (Smith, E.A., 1880)

Psoronaias psorica (Morelet, 1851)

Ptychobranchus (Subtentus) subtentus (Say, 1825)

Ptychobranchus (s.s.) phaseolus (Hildreth, 1828)

Ptychorhynchoideus gümbeli Modell, 1931

Ptychorhynchus pfisteri (Heude, 1874)

Quadrula (Amphinaias) couchianus (Lea, 1860)

Quadrula (Luteacarnea) striata (Rafinesque, 1820)

Quadrula (Obliquata) obliquata (Rafinesque, 1820)

Quadrula (Orthonymus) cylindrica (Say, 1817)

Quadrula (Pleuronaia) barnesiana (Lea, 1838)

Quadrula (s.s.) quadrula (Rafinesque, 1820)

Quincuncina burkei Walker, 1922

Rectidens prolongatus (Drouet, 1894)

Rhabdotophorus gracilis Russell, 1935

Rhombuniopsis tauriformis (Fulton, 1906)

Scabies scobinata (Lea, 1856)

Schepmania nieuwenhuisi (Schepman, 1898)

Shistodesmus [sic] lampreyanus (Baird & Adams, 1867)

Simpsonaias ambigua (Say, 1825)

Simpsonella purpurea (Valenciennes, 1833)

Solenaia emarginata (Lea, 1860)

Strophitus (Jugosus) wrightianus Walker, 1901

Strophitus (Pseudodontoideus) alabamensis (Lea, 1861)

Strophitus (s.s.) undulatus (Say, 1817)

Sulcatula tungurensis Leroy, 1940

Tamsiella tamsiana (Dunker, 1858)

Trapezoideus foliaceus (Gould, 1843)

Triplodon (Triquetrana) stevensi (Lea, 1871)

Triplodon (s.s.) rugosus Spix, 1827

Tritogonia verrucosus (Say)

Truncilla truncata Rafinesque, 1820

Unio (Eolymnium) terminalis Bourguignat, 1852

Unio (Heterunio) exquisitus Lindholm

Unio (Sculptunio) bituberculosus trigonoides (Lindholm)

Unio (Tuberunio) uniserialis [sic] (Lindholm)

Unio (s.s.) pictorum (Linné, 1758)

Uniomerus tetralasmus (Say, 1831)

Unionea [sic] fabagina (DeShayes & Jullien, 1874)

Vetulonaia whitei Branson

Virgus (Leiovirgus) misoolensis (Schepman, 1897)

Virgus (s.s.) beccarianus (Tapparone Canefri, 1883)

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