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Page last updated
Fri 30 May 2025

The Freshwater Mussels (Unionoida) of the World (and other less consequential bivalves)

FM(U)otW(aolcb) is the web version of the MUSSEL Project Database. Follow the links to browse the data or use the custom Google search field. Either way, you win!

Cladomics pages are experimental — even more so than the rest of this site!

The purpose of the cladomics pages for each genus and family-group level taxon is to provide statistics and references indicating the extent to which published phylogenetic work supports the classification used here.

The Cladomics Summary provides an overview of the number of studies that have included terminal species representing this taxon as well as an indication of how frequently it has been recovered as monophyletic.

Following the summary, the available published clades representing the taxon are listed.

For more information, visit the Cladomics FARQ (frequently asked rhetorical questions) page. Abbreviations for the various statistics are explained on the FARQ!

Search the database pages:

Bivalvia | Unionoida | Hyrioidea | Hyriidae | Velesunioninae

genus Velesunio Iredale, 1934: cladomics
5 valid species

Cladomics Summary

number of species = 5

analyzed in 55 cladograms in 34 publications
maximum number of terminals in an analysis = 3, mean = 1.2

Basic Monophyly

number of cladograms that test basic monophyly (i.e., more than 1 included species) = 10
maximum number of included terminals = 3, mean = 2.1
proportion of cladograms that support basic monophyly = 0.1

Pairwise Cladogram Clustering

The graphs below depict in two dimensions all pair-wise ingroup distances within the cladome. See the Cladmoics FARQ for more information.

PCoA Ordination

NJ tree

Walker et al., 2006 [Publ 2666], fig. 6

iT	tT	bi	riS	oT	roT	roS	iD
3	5	4	5	33	2	2	0.0

*** Top Monophyly Cladogram ***

-- 2889

Graf et al., 2015 [Publ 3619], fig. 1

iT	tT	bi	riS	oT	roT	roS	iD
1	1	0	1	19	6	4	--

*** Top Sister-Group Cladogram ***

  +----------- Westralunio       
  |                              
  |     +----- Alathyria_jacksoni
--+  +--+                        
  |  |  |  +-- Velesunio_ambiguus **
  |  |  +--+                     
  +--+     +-- Alathyria_profuga 
     |                           
     |  +----- Alathyria_pertexta
     +--+                        
        |  +-- Microdontia       
        +--+                     
           +-- Lortiella

Hoeh et al., 2009 [Publ 2716], fig. 3

iT	tT	bi	riS	oT	roT	roS	iD
2	3	0	2	45	1	1	4.0

  +----- Velesunio_ambiguus **
--+                        
  |  +-- Velesunio_angasi **  
  +--+                     
     +-- Lortiella

Baker et al., 2003 [Publ 2690], fig. 5

iT	tT	bi	riS	oT	roT	roS	iD
2	2	0	2	6	0	0	--

  +-- Velesunio_angasi **  
--+                     
  +-- Velesunio_ambiguus **

Hoeh et al., 2009 [Publ 2716], fig. 4

iT	tT	bi	riS	oT	roT	roS	iD
2	3	0	2	44	1	1	4.0

  +----- Velesunio_ambiguus **
--+                        
  |  +-- Velesunio_angasi **  
  +--+                     
     +-- Lortiella

Hoeh et al., 2009 [Publ 2716], fig. 5

iT	tT	bi	riS	oT	roT	roS	iD
2	3	0	2	44	1	1	4.0

  +----- Velesunio_ambiguus **
--+                        
  |  +-- Velesunio_angasi **  
  +--+                     
     +-- Lortiella

Santos-Neto et al., 2016 [Publ 3739], fig. 4

iT	tT	bi	riS	oT	roT	roS	iD
2	5	0	2	18	4	4	4.5

  +----------- Velesunio_ambiguus **
--+                              
  |  +-------- Alathyria         
  +--+                           
     |  +----- Velesunio_angasi **  
     +--+                        
        |  +-- Lortiella         
        +--+                     
           +-- Microdontia

Graf & Cummings, 2006 [Publ 1968], fig. 2

iT	tT	bi	riS	oT	roT	roS	iD
2	3	0	2	45	1	1	4.0

  +----- Velesunio_ambiguus **
--+                        
  |  +-- Velesunio_angasi **  
  +--+                     
     +-- Lortiella

Baker et al., 2004 [Publ 2685], fig. 2-3

iT	tT	bi	riS	oT	roT	roS	iD
2	3	0	5	7	1	1	3.0

           +-- Velesunio__3 **    
  +--------+                   
  |        +-- Velesunio__4 **    
--+                            
  |  +-------- Velesunio__2 **    
  +--+                         
     |  +----- Velesunio_angasi **
     +--+                      
        |  +-- Velesunio__1 **    
        +--+                   
           +-- Alathyria

Santos-Neto et al., 2016 [Publ 3739], fig. 2a_b

iT	tT	bi	riS	oT	roT	roS	iD
2	5	0	2	18	4	4	6.5

  +----------- Velesunio_ambiguus **
--+                              
  |  +-------- Microdontia       
  +--+                           
     |  +----- Lortiella         
     +--+                        
        |  +-- Alathyria         
        +--+                     
           +-- Velesunio_angasi **

Marshall et al., 2014 [Publ 3593], fig. 1

iT	tT	bi	riS	oT	roT	roS	iD
2	3	0	2	12	1	1	5.5

  +----- Velesunio_ambiguus **
--+                        
  |  +-- Velesunio **         
  +--+                     
     +-- Lortiella

Sano et al., 2020 [Publ 4138], fig. 3

iT	tT	bi	riS	oT	roT	roS	iD
1	1	0	1	67	0	0	--

-- Velesunio_ambiguus

Klunzinger et al., 2021 [Publ 4086], fig. 2

iT	tT	bi	riS	oT	roT	roS	iD
1	1	0	1	5	2	1	--

-- Velesunio_ambiguus

Lopes-Lima et al., 2017 [Publ 3745], fig. 1-2

iT	tT	bi	riS	oT	roT	roS	iD
1	1	0	1	79	0	0	--

-- Velesunio_ambiguus

Graf, 2002 [Publ 2651], fig. 2

iT	tT	bi	riS	oT	roT	roS	iD
1	1	0	1	17	0	0	--

-- Velesunio_ambiguus

Hoeh et al., 2002 [Publ 2653], fig. 2

iT	tT	bi	riS	oT	roT	roS	iD
1	1	0	1	30	1	1	--

  +-- Velesunio_angasi **
--+                   
  +-- Lortiella

Klunzinger et al., 2022 [Publ 4259], fig. 1

iT	tT	bi	riS	oT	roT	roS	iD
1	1	0	1	3	2	1	--

  +-- Velesunio_ambiguus **
--+                     
  +-- Westralunio

Lopes-Lima et al., 2020 [Publ 4080], fig. 2-4

iT	tT	bi	riS	oT	roT	roS	iD
1	1	0	1	120	0	0	--

-- Velesunio_ambiguus

Huang et al., 2019 [Publ 3843], fig. S4

iT	tT	bi	riS	oT	roT	roS	iD
1	1	0	1	198	6	4	--

     +----- Velesunio_ambiguus **
  +--+                        
  |  |  +-- Alathyria_jacksoni
  |  +--+                     
--+     +-- Alathyria_profuga 
  |                           
  |  +----- Alathyria_pertexta
  +--+                        
     |  +-- Lortiella         
     +--+                     
        +-- Microdontia

Bogan & Hoeh, 2000 [Publ 2650], fig. 1

iT	tT	bi	riS	oT	roT	roS	iD
1	1	0	1	33	1	1	--

  +-- Velesunio_angasi **
--+                   
  +-- Lortiella

Hoeh et al., 2001 [Publ 2652], fig. 14f4

iT	tT	bi	riS	oT	roT	roS	iD
1	1	0	1	30	1	1	--

  +-- Lortiella       
--+                   
  +-- Velesunio_angasi **

Wu et al., 2020 [Publ 4121], fig. 2

iT	tT	bi	riS	oT	roT	roS	iD
1	1	0	1	33	0	0	--

-- Velesunio_ambiguus

Sonowal et al., 2021 [Publ 4221], fig. 3b

iT	tT	bi	riS	oT	roT	roS	iD
1	1	0	1	14	0	0	--

-- Velesunio_ambiguus

Pfeiffer & Graf, 2013 [Publ 3531], fig. 2

iT	tT	bi	riS	oT	roT	roS	iD
1	1	0	1	30	0	0	--

-- Velesunio_ambiguus

Sonowal et al., 2021 [Publ 4221], fig. 4

iT	tT	bi	riS	oT	roT	roS	iD
1	1	0	1	15	0	0	--

-- Velesunio_ambiguus

Combosch et al., 2017 [Publ 3806], fig. 2

iT	tT	bi	riS	oT	roT	roS	iD
1	1	0	1	14	0	0	--

-- Velesunio_ambiguus

González & Giribet, 2015 [Publ 3632], fig. 2

iT	tT	bi	riS	oT	roT	roS	iD
1	1	0	1	5	0	0	--

-- Velesunio_ambiguus

González & Giribet, 2015 [Publ 3632], fig. 6

iT	tT	bi	riS	oT	roT	roS	iD
1	1	0	1	5	0	0	--

-- Velesunio_ambiguus

Huang et al., 2019 [Publ 3843], fig. 2

iT	tT	bi	riS	oT	roT	roS	iD
1	1	0	1	198	6	4	--

     +----- Alathyria_jacksoni
  +--+                        
  |  |  +-- Alathyria_profuga 
  |  +--+                     
--+     +-- Velesunio_ambiguus **
  |                           
  |  +----- Alathyria_pertexta
  +--+                        
     |  +-- Lortiella         
     +--+                     
        +-- Microdontia

Bolotov et al., 2016 [Publ 3743], fig. 4

iT	tT	bi	riS	oT	roT	roS	iD
1	1	0	1	29	5	3	--

     +----- Microdontia       
  +--+                        
  |  |  +-- Alathyria_pertexta
  |  +--+                     
--+     +-- Lortiella         
  |                           
  |  +----- Velesunio_ambiguus **
  +--+                        
     |  +-- Alathyria_jacksoni
     +--+                     
        +-- Alathyria_profuga

Graf & Ó Foighil, 2000 [Publ 2649], fig. 1_2

iT	tT	bi	riS	oT	roT	roS	iD
1	1	0	1	10	0	0	--

-- Velesunio_ambiguus

Bolotov et al., 2017 [Publ 3761], fig. S1

iT	tT	bi	riS	oT	roT	roS	iD
1	1	0	1	138	5	3	--

     +----- Alathyria_pertexta
  +--+                        
  |  |  +-- Microdontia       
  |  +--+                     
--+     +-- Lortiella         
  |                           
  |  +----- Velesunio_ambiguus **
  +--+                        
     |  +-- Alathyria_jacksoni
     +--+                     
        +-- Alathyria_profuga

Lopes-Lima et al., 2018 [Publ 3829], fig. 3

iT	tT	bi	riS	oT	roT	roS	iD
1	1	0	1	27	0	0	--

-- Velesunio_ambiguus

Bolotov et al., 2017 [Publ 3761], fig. S2

iT	tT	bi	riS	oT	roT	roS	iD
1	1	0	1	138	5	3	--

     +----- Velesunio_ambiguus **
  +--+                        
  |  |  +-- Alathyria_jacksoni
  |  +--+                     
--+     +-- Alathyria_profuga 
  |                           
  |  +----- Microdontia       
  +--+                        
     |  +-- Alathyria_pertexta
     +--+                     
        +-- Lortiella

Roe & Hoeh, 2003 [Publ 2655], fig. 4f2

iT	tT	bi	riS	oT	roT	roS	iD
1	1	0	1	30	1	1	--

-- Velesunio_angasi

Sano et al., 2020 [Publ 4138], fig. 2

iT	tT	bi	riS	oT	roT	roS	iD
1	1	0	1	67	0	0	--

-- Velesunio_ambiguus

Hoeh et al., 2001 [Publ 2652], fig. 14f2

iT	tT	bi	riS	oT	roT	roS	iD
1	1	0	1	30	1	1	--

-- Velesunio_angasi

Elderkin et al., 2016 [Publ 3693], fig. 2

iT	tT	bi	riS	oT	roT	roS	iD
1	1	0	1	2	0	0	--

-- Velesunio_ambiguus

Huang et al., 2019 [Publ 3843], fig. S3

iT	tT	bi	riS	oT	roT	roS	iD
1	1	0	1	196	6	4	--

     +----- Velesunio_ambiguus **
  +--+                        
  |  |  +-- Alathyria_jacksoni
  |  +--+                     
--+     +-- Alathyria_profuga 
  |                           
  |  +----- Alathyria_pertexta
  +--+                        
     |  +-- Lortiella         
     +--+                     
        +-- Microdontia

Lopes-Lima et al., 2018 [Publ 3829], fig. 1

iT	tT	bi	riS	oT	roT	roS	iD
1	1	0	1	27	0	0	--

-- Velesunio_ambiguus

Sonowal et al., 2021 [Publ 4221], fig. 3a

iT	tT	bi	riS	oT	roT	roS	iD
1	1	0	1	14	0	0	--

-- Velesunio_ambiguus

Graf, 2000 [Publ 2647], fig. 1

iT	tT	bi	riS	oT	roT	roS	iD
1	1	0	1	16	0	0	--

-- Velesunio

Roe & Hoeh, 2003 [Publ 2655], fig. 4f3

iT	tT	bi	riS	oT	roT	roS	iD
1	1	0	1	30	1	1	--

  +-- Velesunio_angasi **
--+                   
  +-- Lortiella

Whelan et al., 2011 [Publ 3380], fig. 1-MP

iT	tT	bi	riS	oT	roT	roS	iD
1	1	0	1	35	0	0	--

-- Velesunio_ambiguus

Pfeiffer & Graf, 2015 [Publ 3620], fig. 2

iT	tT	bi	riS	oT	roT	roS	iD
1	1	0	1	39	0	0	--

-- Velesunio_ambiguus

Bolotov et al., 2016 [Publ 3743], fig. 2

iT	tT	bi	riS	oT	roT	roS	iD
1	1	0	1	29	5	3	--

     +----- Microdontia       
  +--+                        
  |  |  +-- Lortiella         
  |  +--+                     
--+     +-- Alathyria_pertexta
  |                           
  |  +----- Velesunio_ambiguus **
  +--+                        
     |  +-- Alathyria_jacksoni
     +--+                     
        +-- Alathyria_profuga

González & Giribet, 2015 [Publ 3632], fig. 4

iT	tT	bi	riS	oT	roT	roS	iD
1	1	0	1	5	0	0	--

-- Velesunio_ambiguus

González & Giribet, 2015 [Publ 3632], fig. 3

iT	tT	bi	riS	oT	roT	roS	iD
1	1	0	1	5	0	0	--

-- Velesunio_ambiguus

Bieler et al., 2014 [Publ 3622], fig. 26_27_29_30

iT	tT	bi	riS	oT	roT	roS	iD
1	1	0	1	4	0	0	--

-- Velesunio_ambiguus

Huang et al., 2018 [Publ 3828], fig. 3_5

iT	tT	bi	riS	oT	roT	roS	iD
1	1	0	1	28	0	0	--

-- Velesunio_ambiguus

Wu et al., 2020 [Publ 4121], fig. S1

iT	tT	bi	riS	oT	roT	roS	iD
1	1	0	1	33	0	0	--

-- Velesunio_ambiguus

Whelan et al., 2011 [Publ 3380], fig. 1-ML

iT	tT	bi	riS	oT	roT	roS	iD
1	1	0	1	35	0	0	--

-- Velesunio_ambiguus

Bieler et al., 2014 [Publ 3622], fig. 28

iT	tT	bi	riS	oT	roT	roS	iD
1	1	0	1	4	0	0	--

-- Velesunio_ambiguus

Combosch et al., 2017 [Publ 3806], fig. 1

iT	tT	bi	riS	oT	roT	roS	iD
1	1	0	1	14	0	0	--

-- Velesunio_ambiguus

Sano et al., 2020 [Publ 4138], fig. 4

iT	tT	bi	riS	oT	roT	roS	iD
1	1	0	1	67	0	0	--

-- Velesunio_ambiguus

Page updated: 10:15:48 Fri 30 May 2025

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