Unionoida | Unionoidea | Unionidae | Ambleminae | Quadrulini | Quadrulina genus Pustulosa Frierson, 1927: cladomics 10 valid species
Cladomics Summary
number of species = 10
analyzed in 38 cladograms in 22 publications
maximum number of terminals in an analysis = 8, mean = 3.21
Basic Monophyly
number of cladograms that test basic monophyly (i.e., more than 1 included species) = 25
maximum number of included terminals = 8, mean = 4.36
proportion of cladograms that support basic monophyly = 0.24
Pairwise Cladogram Clustering
The graphs below depict in two dimensions all pair-wise ingroup distances within the cladome. See the Cladmoics FARQ for more information.
Lopes-Lima et al., 2019 [Publ 3901], fig. 3
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
8 |
9 |
7 |
8 |
21 |
9 |
3 |
0.0 |
*** Top Monophyly Cladogram *** *** Top Sister-Group Cladogram ***
+----------- Theliderma_cylindrica
+-----------+
| | +-------- Theliderma_sparsa
| +--+
| | +----- Theliderma_intermedia
| +--+
| | +-- Theliderma_metanevra
| +--+
--+ +-- Theliderma_johnsoni
|
| +----- Quadrula_quadrula
| +--------------+
| | | +-- Quadrula_nobilis
| | +--+
| | +-- Quadrula_verrucosa
+--+
| +----------------- Pustulosa_kieneriana **
| |
| | +-- Pustulosa_infucata **
+--+ +-----------+
| | +-- Pustulosa_kleiniana **
| |
+--+ +-- Pustulosa_pustulosa **
| +--------+
| | +-- Pustulosa_succissa **
+--+
| +-------- Cyclonaias_tuberculata
+--+
| +----- Pustulosa_necki **
+--+
| +-- Pustulosa_petrina **
+--+
+-- Pustulosa_nodulata **
Johnson et al., 2018 [Publ 3844], fig. 2
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
8 |
9 |
7 |
8 |
8 |
4 |
3 |
2.0 |
+-------------------- Theliderma_metanevra
|
| +-- Quadrula_quadrula
--+ +--------------+
| | +-- Quadrula_verrucosa
| |
| | +----- Pustulosa_kieneriana **
+--+ +--------+
| | | +-- Pustulosa_infucata **
| | +--+
| | +-- Pustulosa_kleiniana **
+--+
| +-- Pustulosa_pustulosa **
| +--------+
| | +-- Pustulosa_succissa **
+--+
| +-------- Cyclonaias_tuberculata
+--+
| +----- Pustulosa_necki **
+--+
| +-- Pustulosa_nodulata **
+--+
+-- Pustulosa_petrina **
Lopes-Lima et al., 2019 [Publ 3901], fig. 2
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
8 |
9 |
7 |
8 |
20 |
6 |
3 |
6.0 |
+-- Quadrula_verrucosa
+-----------------+
| +-- Quadrula_nobilis
|
--+ +-- Theliderma_metanevra
| +--------------+
| | +-- Theliderma_johnsoni
| |
+--+ +----- Pustulosa_kieneriana **
| +--------+
| | | +-- Pustulosa_infucata **
| | +--+
+--+ +-- Pustulosa_kleiniana **
|
| +----------- Cyclonaias_tuberculata
| |
+--+ +-- Pustulosa_pustulosa **
| +-----+
| | +-- Pustulosa_succissa **
+--+
| +----- Pustulosa_nodulata **
+--+
| +-- Pustulosa_necki **
+--+
+-- Pustulosa_petrina **
Lopes-Lima et al., 2019 [Publ 3901], fig. 4
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
8 |
9 |
7 |
8 |
18 |
6 |
3 |
2.0 |
+-------------------- Quadrula_quadrula
|
| +-- Quadrula_verrucosa
--+ +--------------+
| | +-- Quadrula_nobilis
| |
| | +----- Pustulosa_kieneriana **
+--+ +--------+
| | | +-- Pustulosa_infucata **
| | +--+
| | +-- Pustulosa_kleiniana **
+--+
| +-- Pustulosa_pustulosa **
| +--------+
| | +-- Pustulosa_succissa **
+--+
| +-------- Cyclonaias_tuberculata
+--+
| +----- Pustulosa_necki **
+--+
| +-- Pustulosa_petrina **
+--+
+-- Pustulosa_nodulata **
Neemuchwala et al., 2023 [Publ 4292], fig. 2
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
8 |
8 |
6 |
8 |
19 |
9 |
3 |
9.0 |
+----------------- Cyclonaias_tuberculata
|
| +-------- Quadrula_nobilis
| +-----+
--+ | | +----- Quadrula_verrucosa
| | +--+
| | | +-- Quadrula_fragosa
| | +--+
| | +-- Quadrula_quadrula
+--+
| +----- Pustulosa_nodulata **
| +-----+
| | | +-- Pustulosa_necki **
| | +--+
| | +-- Pustulosa_petrina **
+--+
| +-- Pustulosa_kieneriana **
| +-----+
| | +-- Pustulosa_pustulosa **
+--+
| +----- Pustulosa_succissa **
+--+
| +-- Pustulosa_infucata **
+--+
+-- Pustulosa_kleiniana **
Serb et al., 2003 [Publ 2659], fig. 2
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
6 |
6 |
3 |
6 |
23 |
7 |
2 |
4.0 |
+-------- Theliderma_cylindrica
+-----+
| | +----- Theliderma_intermedia
| +--+
| | +-- Theliderma_sparsa
| +--+
| +-- Theliderma_metanevra
--+
| +----- Quadrula_verrucosa
| +-----+
| | | +-- Quadrula_quadrula
| | +--+
| | +-- Quadrula_nobilis
+--+
| +-------- Pustulosa_kieneriana **
| |
| +-------- Pustulosa_infucata **
+--+
| +-- Pustulosa_pustulosa **
| +--+
| | +-- Pustulosa_succissa **
+--+
| +-- Pustulosa_nodulata **
+--+
+-- Pustulosa_petrina **
Serb et al., 2003 [Publ 2659], fig. 3
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
6 |
6 |
4 |
6 |
23 |
7 |
2 |
3.0 |
+----------------- Quadrula_quadrula
|
| +-------- Theliderma_cylindrica
--+ +-----+
| | | +----- Theliderma_sparsa
| | +--+
| | | +-- Theliderma_intermedia
+--+ +--+
| +-- Theliderma_metanevra
|
| +----------- Pustulosa_kieneriana **
| |
+--+ +-------- Pustulosa_infucata **
| |
+--+ +-- Pustulosa_succissa **
| +--+
| | +-- Pustulosa_pustulosa **
+--+
| +-- Pustulosa_nodulata **
+--+
+-- Pustulosa_petrina **
Szumowski et al., 2012 [Publ 3569], fig. 5
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
6 |
6 |
4 |
6 |
9 |
8 |
2 |
3.0 |
+-------- Quadrula_quadrula[1]
+-----+
| | +----- Quadrula_verrucosa
| +--+
| | +-- Quadrula_nobilis
--+ +--+
| +-- Quadrula_quadrula[2]
|
| +----------- Pustulosa_kieneriana **
| |
+--+ +-------- Pustulosa_infucata **
| |
+--+ +-- Pustulosa_pustulosa **
| +--+
| | +-- Pustulosa_succissa **
+--+
| +-- Pustulosa_nodulata **
+--+
+-- Pustulosa_petrina **
Campbell et al., 2005 [Publ 2660], fig. 1
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
4 |
5 |
2 |
4 |
113 |
5 |
3 |
5.0 |
+----------- Theliderma_metanevra
|
| +----- Quadrula_verrucosa
--+ +--+
| | | +-- Quadrula_quadrula
| | +--+
| | +-- Quadrula_nobilis
+--+
| +----- Pustulosa_kieneriana **
| |
| | +-- Cyclonaias_tuberculata
+--+--+
| +-- Pustulosa_succissa **
|
| +-- Pustulosa_infucata **
+--+
+-- Pustulosa_kleiniana **
Campbell et al., 2005 [Publ 2660], fig. 2
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
4 |
5 |
2 |
4 |
111 |
5 |
3 |
5.0 |
+----------- Quadrula_quadrula
|
| +-- Quadrula_nobilis
--+ +-----+
| | +-- Quadrula_verrucosa
| |
+--+ +----- Pustulosa_kieneriana **
| |
| | +-- Cyclonaias_tuberculata
+--+--+
| +-- Pustulosa_succissa **
|
| +-- Pustulosa_infucata **
+--+
+-- Pustulosa_kleiniana **
Campbell & Lydeard, 2012 [Publ 3308], fig. 1
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
4 |
5 |
3 |
4 |
39 |
2 |
2 |
4.0 |
+----------- Quadrula_quadrula
|
--+ +-- Pustulosa_infucata **
| +-----+
| | +-- Pustulosa_kleiniana **
+--+
| +----- Cyclonaias_tuberculata
+--+
| +-- Pustulosa_succissa **
+--+
+-- Pustulosa_pustulosa **
Campbell & Lydeard, 2012 [Publ 3308], fig. 2
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
4 |
5 |
3 |
4 |
39 |
2 |
2 |
4.0 |
+----------- Quadrula_quadrula
|
--+ +-- Pustulosa_infucata **
| +-----+
| | +-- Pustulosa_kleiniana **
+--+
| +----- Cyclonaias_tuberculata
+--+
| +-- Pustulosa_succissa **
+--+
+-- Pustulosa_pustulosa **
Campbell & Lydeard, 2012 [Publ 3308], fig. 3
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
4 |
6 |
4 |
4 |
47 |
2 |
2 |
5.0 |
+-- Pustulosa_infucata **
+--------+
| +-- Pustulosa_kleiniana **
--+
| +-------- Quadrula_quadrula
+--+
| +----- Cyclonaias_tuberculata
+--+
| +-- Pustulosa_succissa **
+--+
+-- Pustulosa_pustulosa **
Campbell & Lydeard, 2012 [Publ 3308], fig. 4
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
4 |
5 |
3 |
4 |
39 |
2 |
2 |
4.0 |
+----------- Quadrula_quadrula
|
--+ +-- Pustulosa_infucata **
| +-----+
| | +-- Pustulosa_kleiniana **
+--+
| +----- Cyclonaias_tuberculata
+--+
| +-- Pustulosa_succissa **
+--+
+-- Pustulosa_pustulosa **
Campbell & Lydeard, 2012 [Publ 3309], fig. 1-2
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
4 |
5 |
3 |
4 |
70 |
5 |
3 |
4.0 |
+-------------- Quadrula_verrucosa
|
| +-- Quadrula_quadrula
--+ +--------+
| | +-- Quadrula_nobilis
| |
+--+ +-- Pustulosa_infucata **
| +-----+
| | +-- Pustulosa_kleiniana **
+--+
| +----- Cyclonaias_tuberculata
+--+
| +-- Pustulosa_pustulosa **
+--+
+-- Pustulosa_succissa **
Campbell & Lydeard, 2012 [Publ 3309], fig. 3
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
4 |
5 |
3 |
4 |
70 |
5 |
3 |
4.0 |
+-- Quadrula_verrucosa
+-----------+
| +-- Quadrula_nobilis
|
--+ +----------- Quadrula_quadrula
| |
| +----------- Theliderma_metanevra
+--+
| +-- Pustulosa_infucata **
| +-----+
| | +-- Pustulosa_kleiniana **
+--+
| +----- Cyclonaias_tuberculata
+--+
| +-- Pustulosa_pustulosa **
+--+
+-- Pustulosa_succissa **
Zhou et al., 2007 [Publ 2658], fig. 2
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
3 |
4 |
2 |
3 |
18 |
4 |
2 |
9.0 |
+-- Theliderma_intermedia
+-----------+
| +-- Theliderma_sparsa
--+
| +----------- Quadrula_quadrula
+--+
| +-------- Pustulosa_infucata **
+--+
| +----- Quadrula_verrucosa
+--+
| +-- Pustulosa_kieneriana **
+--+
+-- Pustulosa_succissa **
Lydeard et al., 2000 [Publ 2654], fig. 2a
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
2 |
2 |
0 |
2 |
15 |
1 |
1 |
-- |
+----- Quadrula_nobilis
--+
| +-- Pustulosa_infucata **
+--+
+-- Pustulosa_succissa **
Lydeard et al., 2000 [Publ 2654], fig. 2b
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
2 |
3 |
0 |
2 |
15 |
1 |
1 |
8.0 |
+-- Quadrula_nobilis
|
--+-- Pustulosa_infucata **
|
+-- Pustulosa_succissa **
Lydeard et al., 2000 [Publ 2654], fig. 2c
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
2 |
2 |
0 |
2 |
14 |
1 |
1 |
-- |
+----- Quadrula_nobilis
--+
| +-- Pustulosa_infucata **
+--+
+-- Pustulosa_succissa **
Lydeard et al., 2000 [Publ 2654], fig. 2d
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
2 |
3 |
0 |
2 |
14 |
1 |
1 |
8.0 |
+-- Quadrula_nobilis
|
--+-- Pustulosa_infucata **
|
+-- Pustulosa_succissa **
Boyer et al., 2011 [Publ 2718], fig. 3
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
2 |
3 |
0 |
2 |
33 |
4 |
3 |
6.0 |
+----------- Quadrula_verrucosa
--+
| +-------- Theliderma_metanevra
+--+
| +----- Pustulosa_pustulosa **
+--+
| +-- Cyclonaias_tuberculata
+--+
+-- Pustulosa_nodulata **
Boyer et al., 2011 [Publ 2718], fig. 4
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
2 |
3 |
0 |
2 |
36 |
4 |
3 |
6.0 |
+----------- Theliderma_metanevra
|
--+ +-- Quadrula_verrucosa
| +-----+
| | +-- Quadrula_quadrula
+--+
| +----- Pustulosa_pustulosa **
+--+
| +-- Cyclonaias_tuberculata
+--+
+-- Pustulosa_nodulata **
Pfeiffer et al., 2016 [Publ 3660], fig. 2
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
2 |
3 |
0 |
2 |
37 |
3 |
2 |
6.0 |
+----------- Quadrula_quadrula
--+
| +-------- Quadrula_verrucosa
+--+
| +----- Pustulosa_pustulosa **
+--+
| +-- Pustulosa_petrina **
+--+
+-- Cyclonaias_tuberculata
Inoue et al., 2018 [Publ 3858], fig. 1, 3a-c
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
2 |
6 |
0 |
2 |
203 |
4 |
3 |
11.0 |
+-- Cyclonaias_tuberculata
|
+-- Pustulosa_pustulosa[1] **
|
+-- Theliderma_metanevra
--+
+-- Pustulosa_pustulosa[2] **
|
+-- Quadrula_quadrula
|
+-- Quadrula_verrucosa
Chapman et al., 2008 [Publ 2667], fig. 1
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
1 |
1 |
0 |
1 |
20 |
1 |
1 |
-- |
+-- Quadrula_quadrula
--+
+-- Pustulosa_pustulosa **
Curole & Kocher, 2005 [Publ 2682], fig. 1
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
1 |
1 |
0 |
1 |
10 |
1 |
1 |
-- |
+-- Quadrula_quadrula
--+
+-- Pustulosa_pustulosa **
Curole & Kocher, 2005 [Publ 2682], fig. 1-M
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
1 |
1 |
0 |
1 |
11 |
1 |
1 |
-- |
+-- Quadrula_quadrula
--+
+-- Pustulosa_pustulosa **
Curole & Kocher, 2002 [Publ 2712], fig. 1-F
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
1 |
1 |
0 |
1 |
3 |
1 |
1 |
-- |
+-- Quadrula_quadrula
--+
+-- Pustulosa_pustulosa **
Curole & Kocher, 2002 [Publ 2712], fig. 1a-M
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
1 |
1 |
0 |
1 |
3 |
1 |
1 |
-- |
+-- Quadrula_quadrula
--+
+-- Pustulosa_pustulosa **
Curole & Kocher, 2002 [Publ 2712], fig. 1b-M
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
1 |
1 |
0 |
1 |
3 |
1 |
1 |
-- |
+-- Quadrula_quadrula
--+
+-- Pustulosa_pustulosa **
Chapman et al., 2008 [Publ 2744], fig. 2
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
1 |
1 |
0 |
1 |
20 |
1 |
1 |
-- |
+-- Quadrula_quadrula
--+
+-- Pustulosa_pustulosa **
Breton et al., 2011 [Publ 3376], fig. 3
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
1 |
1 |
0 |
1 |
38 |
2 |
2 |
-- |
+----- Quadrula_quadrula
--+
| +-- Cyclonaias_tuberculata
+--+
+-- Pustulosa_pustulosa **
Chase et al., 2018 [Publ 3847], fig. 4
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
1 |
1 |
0 |
1 |
38 |
1 |
1 |
-- |
+-- Cyclonaias_tuberculata
--+
+-- Pustulosa_pustulosa **
Pieri et al., 2018 [Publ 3867], fig. 3
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
1 |
1 |
0 |
1 |
36 |
1 |
1 |
-- |
+-- Quadrula_quadrula
--+
+-- Pustulosa_petrina **
Inoue et al., 2020 [Publ 3941], fig. 3
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
1 |
1 |
0 |
1 |
43 |
1 |
1 |
-- |
+-- Pustulosa_petrina **
--+
+-- Quadrula_quadrula
Inoue et al., 2020 [Publ 3941], fig. 4
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
1 |
1 |
0 |
1 |
42 |
1 |
1 |
-- |
+-- Pustulosa_petrina **
--+
+-- Quadrula_quadrula
Porto-Hannes et al., 2021 [Publ 4227], fig. 4
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
1 |
1 |
0 |
1 |
11 |
1 |
1 |
-- |
-- Pustulosa_pustulosa
|