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Page last updated
Wed 04 Dec 2024

The Freshwater Mussels (Unionoida) of the World (and other less consequential bivalves)

FM(U)otW(aolcb) is the web version of the MUSSEL Project Database. Follow the links to browse the data or use the custom Google search field. Either way, you win!

Cladomics pages are experimental — even more so than the rest of this site!

The purpose of the cladomics pages for each genus and family-group level taxon is to provide statistics and references indicating the extent to which published phylogenetic work supports the classification used here.

The Cladomics Summary provides an overview of the number of studies that have included terminal species representing this taxon as well as an indication of how frequently it has been recovered as monophyletic.

Following the summary, the available published clades representing the taxon are listed.

For more information, visit the Cladomics FARQ (frequently asked rhetorical questions) page. Abbreviations for the various statistics are explained on the FARQ!

Search the database pages:

Unionoida | Unionoidea | Unionidae | Ambleminae | Quadrulini | Quadrulina

genus Pustulosa Frierson, 1927: cladomics
     10 valid species

Cladomics Summary

number of species = 10

analyzed in 38 cladograms in 22 publications
maximum number of terminals in an analysis = 8, mean = 3.18

Basic Monophyly

number of cladograms that test basic monophyly (i.e., more than 1 included species) = 24
maximum number of included terminals = 8, mean = 4.46
proportion of cladograms that support basic monophyly = 0.25


Pairwise Cladogram Clustering

The graphs below depict in two dimensions all pair-wise ingroup distances within the cladome. See the Cladmoics FARQ for more information.


PCoA Ordination


NJ tree


Lopes-Lima et al., 2019 [Publ 3901], fig. 3

iT tT bi riS oT roT roS iD
8 9 7 8 21 9 3 0.0

*** Top Monophyly Cladogram *** *** Top Sister-Group Cladogram ***

  +----------------------- Theliderma          
  |                                            
--+  +-------------------- Quadrula            
  |  |                                         
  +--+  +----------------- Pustulosa_kieneriana **
     |  |                                      
     |  |              +-- Pustulosa_infucata **  
     +--+  +-----------+                       
        |  |           +-- Pustulosa_kleiniana ** 
        |  |                                   
        +--+           +-- Pustulosa_pustulosa ** 
           |  +--------+                       
           |  |        +-- Pustulosa_succissa **  
           +--+                                
              |  +-------- Cyclonaias          
              +--+                             
                 |  +----- Pustulosa_necki **     
                 +--+                          
                    |  +-- Pustulosa_petrina **   
                    +--+                       
                       +-- Pustulosa_nodulata **  

Neemuchwala et al., 2023 [Publ 4292], fig. 2

iT tT bi riS oT roT roS iD
8 8 6 8 19 9 3 9.0
  +-------------------- Theliderma          
  |                                         
--+  +----------------- Cyclonaias          
  |  |                                      
  +--+  +-------------- Quadrula            
     |  |                                   
     |  |        +----- Pustulosa_nodulata **  
     +--+  +-----+                          
        |  |     |  +-- Pustulosa_necki **     
        |  |     +--+                       
        |  |        +-- Pustulosa_petrina **   
        +--+                                
           |        +-- Pustulosa_kieneriana **
           |  +-----+                       
           |  |     +-- Pustulosa_pustulosa ** 
           +--+                             
              |  +----- Pustulosa_succissa **  
              +--+                          
                 |  +-- Pustulosa_infucata **  
                 +--+                       
                    +-- Pustulosa_kleiniana ** 

Johnson et al., 2018 [Publ 3844], fig. 2

iT tT bi riS oT roT roS iD
8 9 7 8 8 4 3 2.0
  +-------------------- Theliderma          
  |                                         
--+  +----------------- Quadrula            
  |  |                                      
  |  |           +----- Pustulosa_kieneriana **
  +--+  +--------+                          
     |  |        |  +-- Pustulosa_infucata **  
     |  |        +--+                       
     |  |           +-- Pustulosa_kleiniana ** 
     +--+                                   
        |           +-- Pustulosa_pustulosa ** 
        |  +--------+                       
        |  |        +-- Pustulosa_succissa **  
        +--+                                
           |  +-------- Cyclonaias          
           +--+                             
              |  +----- Pustulosa_necki **     
              +--+                          
                 |  +-- Pustulosa_nodulata **  
                 +--+                       
                    +-- Pustulosa_petrina **   

Lopes-Lima et al., 2019 [Publ 3901], fig. 4

iT tT bi riS oT roT roS iD
8 9 7 8 18 6 3 2.0
  +----------------------- Theliderma          
  |                                            
--+  +-------------------- Quadrula_quadrula   
  |  |                                         
  |  |                 +-- Quadrula_verrucosa  
  +--+  +--------------+                       
     |  |              +-- Quadrula_nobilis    
     |  |                                      
     |  |           +----- Pustulosa_kieneriana **
     +--+  +--------+                          
        |  |        |  +-- Pustulosa_infucata **  
        |  |        +--+                       
        |  |           +-- Pustulosa_kleiniana ** 
        +--+                                   
           |           +-- Pustulosa_pustulosa ** 
           |  +--------+                       
           |  |        +-- Pustulosa_succissa **  
           +--+                                
              |  +-------- Cyclonaias          
              +--+                             
                 |  +----- Pustulosa_necki **     
                 +--+                          
                    |  +-- Pustulosa_petrina **   
                    +--+                       
                       +-- Pustulosa_nodulata **  

Lopes-Lima et al., 2019 [Publ 3901], fig. 2

iT tT bi riS oT roT roS iD
8 9 7 8 20 6 3 6.0
  +----------------------- Quadrula_quadrula   
  |                                            
  |                    +-- Quadrula_verrucosa  
--+  +-----------------+                       
  |  |                 +-- Quadrula_nobilis    
  |  |                                         
  +--+  +----------------- Theliderma          
     |  |                                      
     |  |           +----- Pustulosa_kieneriana **
     +--+  +--------+                          
        |  |        |  +-- Pustulosa_infucata **  
        |  |        +--+                       
        +--+           +-- Pustulosa_kleiniana ** 
           |                                   
           |  +----------- Cyclonaias          
           |  |                                
           +--+        +-- Pustulosa_pustulosa ** 
              |  +-----+                       
              |  |     +-- Pustulosa_succissa **  
              +--+                             
                 |  +----- Pustulosa_nodulata **  
                 +--+                          
                    |  +-- Pustulosa_necki **     
                    +--+                       
                       +-- Pustulosa_petrina **   

Serb et al., 2003 [Publ 2659], fig. 2

iT tT bi riS oT roT roS iD
6 6 3 6 23 7 2 4.0
  +-------------- Theliderma          
  |                                   
--+  +----------- Quadrula            
  |  |                                
  +--+  +-------- Pustulosa_kieneriana **
     |  |                             
     |  +-------- Pustulosa_infucata **  
     +--+                             
        |     +-- Pustulosa_pustulosa ** 
        |  +--+                       
        |  |  +-- Pustulosa_succissa **  
        +--+                          
           |  +-- Pustulosa_nodulata **  
           +--+                       
              +-- Pustulosa_petrina **   

Serb et al., 2003 [Publ 2659], fig. 3

iT tT bi riS oT roT roS iD
6 6 4 6 23 7 2 3.0
                    +-- Quadrula_verrucosa  
  +-----------------+                       
  |                 +-- Quadrula_nobilis    
--+                                         
  |  +----------------- Quadrula_quadrula   
  |  |                                      
  +--+  +-------------- Theliderma          
     |  |                                   
     +--+  +----------- Pustulosa_kieneriana **
        |  |                                
        +--+  +-------- Pustulosa_infucata **  
           |  |                             
           +--+     +-- Pustulosa_succissa **  
              |  +--+                       
              |  |  +-- Pustulosa_pustulosa ** 
              +--+                          
                 |  +-- Pustulosa_nodulata **  
                 +--+                       
                    +-- Pustulosa_petrina **   

Szumowski et al., 2012 [Publ 3569], fig. 5

iT tT bi riS oT roT roS iD
6 6 4 6 8 7 2 3.0
  +-------------- Quadrula            
  |                                   
--+  +----------- Pustulosa_kieneriana **
  |  |                                
  +--+  +-------- Pustulosa_infucata **  
     |  |                             
     +--+     +-- Pustulosa_pustulosa ** 
        |  +--+                       
        |  |  +-- Pustulosa_succissa **  
        +--+                          
           |  +-- Pustulosa_nodulata **  
           +--+                       
              +-- Pustulosa_petrina **   

Campbell & Lydeard, 2012 [Publ 3308], fig. 4

iT tT bi riS oT roT roS iD
4 5 3 4 38 2 2 4.0
  +----------- Quadrula           
  |                               
--+        +-- Pustulosa_infucata ** 
  |  +-----+                      
  |  |     +-- Pustulosa_kleiniana **
  +--+                            
     |  +----- Cyclonaias         
     +--+                         
        |  +-- Pustulosa_succissa ** 
        +--+                      
           +-- Pustulosa_pustulosa **

Campbell & Lydeard, 2012 [Publ 3308], fig. 2

iT tT bi riS oT roT roS iD
4 5 3 4 38 2 2 4.0
  +----------- Quadrula           
  |                               
--+        +-- Pustulosa_infucata ** 
  |  +-----+                      
  |  |     +-- Pustulosa_kleiniana **
  +--+                            
     |  +----- Cyclonaias         
     +--+                         
        |  +-- Pustulosa_succissa ** 
        +--+                      
           +-- Pustulosa_pustulosa **

Campbell & Lydeard, 2012 [Publ 3308], fig. 3

iT tT bi riS oT roT roS iD
4 6 4 4 39 2 2 5.0
           +-- Pustulosa_infucata ** 
  +--------+                      
  |        +-- Pustulosa_kleiniana **
--+                               
  |  +-------- Quadrula           
  +--+                            
     |  +----- Cyclonaias         
     +--+                         
        |  +-- Pustulosa_succissa ** 
        +--+                      
           +-- Pustulosa_pustulosa **

Campbell & Lydeard, 2012 [Publ 3308], fig. 1

iT tT bi riS oT roT roS iD
4 5 3 4 38 2 2 4.0
  +----------- Quadrula           
  |                               
--+        +-- Pustulosa_infucata ** 
  |  +-----+                      
  |  |     +-- Pustulosa_kleiniana **
  +--+                            
     |  +----- Cyclonaias         
     +--+                         
        |  +-- Pustulosa_succissa ** 
        +--+                      
           +-- Pustulosa_pustulosa **

Campbell et al., 2005 [Publ 2660], fig. 2

iT tT bi riS oT roT roS iD
4 5 2 4 98 5 3 5.0
  +-------------- Theliderma          
  |                                   
--+  +----------- Quadrula_quadrula   
  |  |                                
  |  |        +-- Quadrula_nobilis    
  +--+  +-----+                       
     |  |     +-- Quadrula_verrucosa  
     |  |                             
     +--+  +----- Pustulosa_kieneriana **
        |  |                          
        |  |  +-- Cyclonaias          
        +--+--+                       
           |  +-- Pustulosa_succissa **  
           |                          
           |  +-- Pustulosa_infucata **  
           +--+                       
              +-- Pustulosa_kleiniana ** 

Campbell & Lydeard, 2012 [Publ 3309], fig. 1-2

iT tT bi riS oT roT roS iD
4 5 3 4 69 5 3 4.0
  +----------------- Theliderma         
  |                                     
--+  +-------------- Quadrula_verrucosa 
  |  |                                  
  |  |           +-- Quadrula_quadrula  
  +--+  +--------+                      
     |  |        +-- Quadrula_nobilis   
     |  |                               
     +--+        +-- Pustulosa_infucata ** 
        |  +-----+                      
        |  |     +-- Pustulosa_kleiniana **
        +--+                            
           |  +----- Cyclonaias         
           +--+                         
              |  +-- Pustulosa_pustulosa **
              +--+                      
                 +-- Pustulosa_succissa ** 

Campbell & Lydeard, 2012 [Publ 3309], fig. 3

iT tT bi riS oT roT roS iD
4 5 3 4 69 5 3 4.0
              +-- Quadrula_verrucosa 
  +-----------+                      
  |           +-- Quadrula_nobilis   
  |                                  
--+  +----------- Quadrula_quadrula  
  |  |                               
  |  +----------- Theliderma         
  +--+                               
     |        +-- Pustulosa_infucata ** 
     |  +-----+                      
     |  |     +-- Pustulosa_kleiniana **
     +--+                            
        |  +----- Cyclonaias         
        +--+                         
           |  +-- Pustulosa_pustulosa **
           +--+                      
              +-- Pustulosa_succissa ** 

Campbell et al., 2005 [Publ 2660], fig. 1

iT tT bi riS oT roT roS iD
4 5 2 4 98 5 3 5.0
  +----------- Theliderma          
  |                                
--+  +-------- Quadrula            
  |  |                             
  +--+  +----- Pustulosa_kieneriana **
     |  |                          
     |  |  +-- Cyclonaias          
     +--+--+                       
        |  +-- Pustulosa_succissa **  
        |                          
        |  +-- Pustulosa_infucata **  
        +--+                       
           +-- Pustulosa_kleiniana ** 

Zhou et al., 2007 [Publ 2658], fig. 2

iT tT bi riS oT roT roS iD
3 4 2 3 18 4 2 9.0
  +-------------- Theliderma          
--+                                   
  |  +----------- Quadrula_quadrula   
  +--+                                
     |  +-------- Pustulosa_infucata **  
     +--+                             
        |  +----- Quadrula_verrucosa  
        +--+                          
           |  +-- Pustulosa_kieneriana **
           +--+                       
              +-- Pustulosa_succissa **  

Lydeard et al., 2000 [Publ 2654], fig. 2a

iT tT bi riS oT roT roS iD
2 2 0 2 12 1 1 --
  +----- Quadrula          
--+                        
  |  +-- Pustulosa_infucata **
  +--+                     
     +-- Pustulosa_succissa **

Pfeiffer et al., 2016 [Publ 3660], fig. 2

iT tT bi riS oT roT roS iD
2 3 0 2 36 3 2 6.0
  +----------- Quadrula_quadrula  
--+                               
  |  +-------- Quadrula_verrucosa 
  +--+                            
     |  +----- Pustulosa_pustulosa **
     +--+                         
        |  +-- Pustulosa_petrina **  
        +--+                      
           +-- Cyclonaias         

Lydeard et al., 2000 [Publ 2654], fig. 2c

iT tT bi riS oT roT roS iD
2 2 0 2 12 1 1 --
  +----- Quadrula          
--+                        
  |  +-- Pustulosa_infucata **
  +--+                     
     +-- Pustulosa_succissa **

Boyer et al., 2011 [Publ 2718], fig. 4

iT tT bi riS oT roT roS iD
2 3 0 2 36 4 3 6.0
  +----------- Theliderma         
--+                               
  |  +-------- Quadrula           
  +--+                            
     |  +----- Pustulosa_pustulosa **
     +--+                         
        |  +-- Cyclonaias         
        +--+                      
           +-- Pustulosa_nodulata ** 

Lydeard et al., 2000 [Publ 2654], fig. 2b

iT tT bi riS oT roT roS iD
2 3 0 2 12 1 1 8.0
  +-- Quadrula          
  |                     
--+-- Pustulosa_infucata **
  |                     
  +-- Pustulosa_succissa **

Boyer et al., 2011 [Publ 2718], fig. 3

iT tT bi riS oT roT roS iD
2 3 0 2 33 4 3 6.0
  +-------------- Quadrula_quadrula  
--+                                  
  |  +----------- Quadrula_verrucosa 
  +--+                               
     |  +-------- Theliderma         
     +--+                            
        |  +----- Pustulosa_pustulosa **
        +--+                         
           |  +-- Cyclonaias         
           +--+                      
              +-- Pustulosa_nodulata ** 

Lydeard et al., 2000 [Publ 2654], fig. 2d

iT tT bi riS oT roT roS iD
2 3 0 2 12 1 1 8.0
  +-- Quadrula          
  |                     
--+-- Pustulosa_infucata **
  |                     
  +-- Pustulosa_succissa **

Curole & Kocher, 2002 [Publ 2712], fig. 1-F

iT tT bi riS oT roT roS iD
1 1 0 1 3 1 1 --
  +-- Quadrula           
--+                      
  +-- Pustulosa_pustulosa **

Chase et al., 2018 [Publ 3847], fig. 4

iT tT bi riS oT roT roS iD
1 1 0 1 37 1 1 --
  +-- Cyclonaias         
--+                      
  +-- Pustulosa_pustulosa **

Curole & Kocher, 2002 [Publ 2712], fig. 1a-M

iT tT bi riS oT roT roS iD
1 1 0 1 3 1 1 --
  +-- Quadrula           
--+                      
  +-- Pustulosa_pustulosa **

Breton et al., 2011 [Publ 3376], fig. 3

iT tT bi riS oT roT roS iD
1 1 0 1 36 2 2 --
  +----- Quadrula           
--+                         
  |  +-- Cyclonaias         
  +--+                      
     +-- Pustulosa_pustulosa **

Pieri et al., 2018 [Publ 3867], fig. 3

iT tT bi riS oT roT roS iD
1 1 0 1 19 1 1 --
  +-- Quadrula         
--+                    
  +-- Pustulosa_petrina **

Chapman et al., 2008 [Publ 2744], fig. 2

iT tT bi riS oT roT roS iD
1 1 0 1 20 1 1 --
  +-- Quadrula           
--+                      
  +-- Pustulosa_pustulosa **

Inoue et al., 2018 [Publ 3858], fig. 1_3a-c

iT tT bi riS oT roT roS iD
1 5 0 2 103 4 3 11.0
  +-- Cyclonaias            
  |                         
  +-- Pustulosa_pustulosa__1 **
  |                         
  +-- Theliderma            
--+                         
  +-- Pustulosa_pustulosa__2 **
  |                         
  +-- Quadrula_quadrula     
  |                         
  +-- Quadrula_verrucosa    

Inoue et al., 2020 [Publ 3941], fig. 4

iT tT bi riS oT roT roS iD
1 1 0 1 40 1 1 --
  +-- Pustulosa_petrina **
--+                    
  +-- Quadrula         

Porto-Hannes et al., 2021 [Publ 4227], fig. 4

iT tT bi riS oT roT roS iD
1 1 0 1 10 1 1 --
-- Pustulosa_pustulosa

Curole & Kocher, 2005 [Publ 2682], fig. 1-M

iT tT bi riS oT roT roS iD
1 1 0 1 11 1 1 --
  +-- Quadrula           
--+                      
  +-- Pustulosa_pustulosa **

Inoue et al., 2020 [Publ 3941], fig. 3

iT tT bi riS oT roT roS iD
1 1 0 1 41 1 1 --
  +-- Pustulosa_petrina **
--+                    
  +-- Quadrula         

Chapman et al., 2008 [Publ 2667], fig. 1

iT tT bi riS oT roT roS iD
1 1 0 1 20 1 1 --
  +-- Quadrula           
--+                      
  +-- Pustulosa_pustulosa **

Curole & Kocher, 2005 [Publ 2682], fig. 1

iT tT bi riS oT roT roS iD
1 1 0 1 10 1 1 --
  +-- Quadrula           
--+                      
  +-- Pustulosa_pustulosa **

Curole & Kocher, 2002 [Publ 2712], fig. 1b-M

iT tT bi riS oT roT roS iD
1 1 0 1 3 1 1 --
  +-- Quadrula           
--+                      
  +-- Pustulosa_pustulosa **

Page updated: 14:56:47 Wed 04 Dec 2024

 
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