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Page last updated
Thu 01 Dec 2022

The Freshwater Mussels (Unionoida) of the World (and other less consequential bivalves)

FM(U)otW(aolcb) is the web version of the MUSSEL Project Database. Follow the links to browse the data or use the custom Google search field. Either way, you win!

Cladomics pages are experimental — even more so than the rest of this site!

The purpose of the cladomics pages for each genus and family-group level taxon is to provide statistics and references indicating the extent to which published phylogenetic work supports the classification used here.

The Cladomics Summary provides an overview of the number of studies that have included terminal species representing this taxon as well as an indication of how frequently it has been recovered as monophyletic.

Following the summary, the available published clades representing the taxon are listed.

For more information, visit the Cladomics FARQ (frequently asked rhetorical questions) page. Abbreviations for the various statistics are explained on the FARQ!

Search the database pages:

Bivalvia | Unionoida | Unionoidea | Unionidae | Ambleminae | Quadrulini

genus Cyclonaias Pilsbry in Ortmann & Walker, 1922: cladomics
     11 valid species

Cladomics Summary

number of species = 11

analyzed in 38 cladograms in 22 publications
maximum number of terminal species in an analysis = 9, mean = 3.5

Basic Monophyly

number of cladograms that test basic monophyly (i.e., more than 1 included species) = 26
maximum number of included species = 9, mean = 4.65
proportion of cladograms that support basic monophyly = 0.81


Lopes-Lima et al., 2019, fig. 3

iT tT bi riS oT roT roS iD
9 9 7 9 20 10 5 0.0

*** Top Monophyly Cladogram *** *** Top Sister-Group Cladogram ***

              +----------- Theliderma_cylindrica 
  +-----------+                                  
  |           |  +-------- Theliderma_sparsa     
  |           +--+                               
  |              |  +----- Theliderma_intermedia 
  |              +--+                            
  |                 |  +-- Theliderma_metanevra  
  |                 +--+                         
--+                    +-- Theliderma_johnsoni   
  |                                              
  |                 +----- Quadrula_quadrula     
  |  +--------------+                            
  |  |              |  +-- Tritogonia_nobilis    
  |  |              +--+                         
  |  |                 +-- Tritogonia_verrucosa  
  +--+                                           
     |  +----------------- Cyclonaias_kieneriana ** 
     |  |                                        
     |  |              +-- Cyclonaias_infucata **   
     +--+  +-----------+                         
        |  |           +-- Cyclonaias_kleiniana **  
        |  |                                     
        +--+           +-- Cyclonaias_pustulosa **  
           |  +--------+                         
           |  |        +-- Cyclonaias_succissa **   
           +--+                                  
              |  +-------- Cyclonaias_tuberculata **
              +--+                               
                 |  +----- Cyclonaias_necki **      
                 +--+                            
                    |  +-- Cyclonaias_petrina **    
                    +--+                         
                       +-- Cyclonaias_nodulata **   

Johnson et al., 2018, fig. 2

iT tT bi riS oT roT roS iD
9 9 6 9 7 5 5 1.0
  +-------------------- Theliderma_metanevra  
  |                                           
  |                 +-- Quadrula_quadrula     
--+  +--------------+                         
  |  |              +-- Tritogonia_verrucosa  
  |  |                                        
  |  |           +----- Cyclonaias_kieneriana ** 
  +--+  +--------+                            
     |  |        |  +-- Cyclonaias_infucata **   
     |  |        +--+                         
     |  |           +-- Cyclonaias_kleiniana **  
     +--+                                     
        |           +-- Cyclonaias_pustulosa **  
        |  +--------+                         
        |  |        +-- Cyclonaias_succissa **   
        +--+                                  
           |  +-------- Cyclonaias_tuberculata **
           +--+                               
              |  +----- Cyclonaias_necki **      
              +--+                            
                 |  +-- Cyclonaias_nodulata **   
                 +--+                         
                    +-- Cyclonaias_petrina **    

Lopes-Lima et al., 2019, fig. 2

iT tT bi riS oT roT roS iD
9 9 6 9 19 9 5 5.0
                    +-- Tritogonia_verrucosa  
  +-----------------+                         
  |                 +-- Tritogonia_nobilis    
  |                                           
--+                 +-- Theliderma_metanevra  
  |  +--------------+                         
  |  |              +-- Theliderma_johnsoni   
  |  |                                        
  +--+           +----- Cyclonaias_kieneriana ** 
     |  +--------+                            
     |  |        |  +-- Cyclonaias_infucata **   
     |  |        +--+                         
     +--+           +-- Cyclonaias_kleiniana **  
        |                                     
        |  +----------- Cyclonaias_tuberculata **
        |  |                                  
        +--+        +-- Cyclonaias_pustulosa **  
           |  +-----+                         
           |  |     +-- Cyclonaias_succissa **   
           +--+                               
              |  +----- Cyclonaias_nodulata **   
              +--+                            
                 |  +-- Cyclonaias_necki **      
                 +--+                         
                    +-- Cyclonaias_petrina **    

Lopes-Lima et al., 2019, fig. 4

iT tT bi riS oT roT roS iD
9 9 6 9 17 7 5 1.0
  +-------------------- Quadrula_quadrula     
  |                                           
  |                 +-- Tritogonia_verrucosa  
--+  +--------------+                         
  |  |              +-- Tritogonia_nobilis    
  |  |                                        
  |  |           +----- Cyclonaias_kieneriana ** 
  +--+  +--------+                            
     |  |        |  +-- Cyclonaias_infucata **   
     |  |        +--+                         
     |  |           +-- Cyclonaias_kleiniana **  
     +--+                                     
        |           +-- Cyclonaias_pustulosa **  
        |  +--------+                         
        |  |        +-- Cyclonaias_succissa **   
        +--+                                  
           |  +-------- Cyclonaias_tuberculata **
           +--+                               
              |  +----- Cyclonaias_necki **      
              +--+                            
                 |  +-- Cyclonaias_petrina **    
                 +--+                         
                    +-- Cyclonaias_nodulata **   

Serb et al., 2003, fig. 2

iT tT bi riS oT roT roS iD
6 6 3 6 24 9 4 4.0
        +-------- Theliderma_cylindrica
  +-----+                              
  |     |  +----- Theliderma_intermedia
  |     +--+                           
  |        |  +-- Theliderma_sparsa    
  |        +--+                        
  |           +-- Theliderma_metanevra 
--+                                    
  |     +-------- Tritogonia_verrucosa 
  |  +--+                              
  |  |  |  +----- Quadrula_quadrula[1] 
  |  |  +--+                           
  |  |     |  +-- Tritogonia_nobilis   
  |  |     +--+                        
  +--+        +-- Quadrula_quadrula[2] 
     |                                 
     |  +-------- Cyclonaias_kieneriana **
     |  |                              
     |  +-------- Cyclonaias_infucata **  
     +--+                              
        |     +-- Cyclonaias_pustulosa ** 
        |  +--+                        
        |  |  +-- Cyclonaias_succissa **  
        +--+                           
           |  +-- Cyclonaias_nodulata **  
           +--+                        
              +-- Cyclonaias_petrina **   

Serb et al., 2003, fig. 3

iT tT bi riS oT roT roS iD
6 6 4 6 24 9 4 3.0
  +----------------- Quadrula_quadrula[2] 
  |                                       
  |        +-------- Theliderma_cylindrica
--+  +-----+                              
  |  |     |  +----- Theliderma_sparsa    
  |  |     +--+                           
  |  |        |  +-- Theliderma_intermedia
  +--+        +--+                        
     |           +-- Theliderma_metanevra 
     |                                    
     |  +----------- Cyclonaias_kieneriana **
     |  |                                 
     +--+  +-------- Cyclonaias_infucata **  
        |  |                              
        +--+     +-- Cyclonaias_succissa **  
           |  +--+                        
           |  |  +-- Cyclonaias_pustulosa ** 
           +--+                           
              |  +-- Cyclonaias_nodulata **  
              +--+                        
                 +-- Cyclonaias_petrina **   

Szumowski et al., 2012, fig. 5

iT tT bi riS oT roT roS iD
6 6 4 6 9 9 4 3.0
  +----------------- Megalonaias_nervosa  
  |                                       
  |        +-------- Theliderma_cylindrica
  +--------+                              
  |        |  +----- Theliderma_intermedia
  |        +--+                           
  |           |  +-- Theliderma_metanevra 
--+           +--+                        
  |              +-- Theliderma_sparsa    
  |                                       
  |        +-------- Quadrula_quadrula[1] 
  |  +-----+                              
  |  |     |  +----- Tritogonia_verrucosa 
  |  |     +--+                           
  |  |        |  +-- Tritogonia_nobilis   
  +--+        +--+                        
     |           +-- Quadrula_quadrula[2] 
     |                                    
     |  +----------- Cyclonaias_kieneriana **
     |  |                                 
     +--+  +-------- Cyclonaias_infucata **  
        |  |                              
        +--+     +-- Cyclonaias_pustulosa ** 
           |  +--+                        
           |  |  +-- Cyclonaias_succissa **  
           +--+                           
              |  +-- Cyclonaias_nodulata **  
              +--+                        
                 +-- Cyclonaias_petrina **   

Campbell et al., 2005, fig. 1

iT tT bi riS oT roT roS iD
5 5 2 5 112 6 5 5.0
  +----------- Theliderma_metanevra  
  |                                  
  |     +----- Tritogonia_verrucosa  
--+  +--+                            
  |  |  |  +-- Quadrula_quadrula     
  |  |  +--+                         
  |  |     +-- Tritogonia_nobilis    
  +--+                               
     |  +----- Cyclonaias_kieneriana ** 
     |  |                            
     |  |  +-- Cyclonaias_tuberculata **
     +--+--+                         
        |  +-- Cyclonaias_succissa **   
        |                            
        |  +-- Cyclonaias_infucata **   
        +--+                         
           +-- Cyclonaias_kleiniana **  

Campbell et al., 2005, fig. 2

iT tT bi riS oT roT roS iD
5 5 2 5 110 6 5 5.0
  +----------- Quadrula_quadrula     
  |                                  
  |        +-- Tritogonia_nobilis    
--+  +-----+                         
  |  |     +-- Tritogonia_verrucosa  
  |  |                               
  +--+  +----- Cyclonaias_kieneriana ** 
     |  |                            
     |  |  +-- Cyclonaias_tuberculata **
     +--+--+                         
        |  +-- Cyclonaias_succissa **   
        |                            
        |  +-- Cyclonaias_infucata **   
        +--+                         
           +-- Cyclonaias_kleiniana **  

Campbell & Lydeard, 2012, fig. 1

iT tT bi riS oT roT roS iD
5 5 2 5 39 1 1 4.0
  +----------- Quadrula_quadrula     
  |                                  
--+        +-- Cyclonaias_infucata **   
  |  +-----+                         
  |  |     +-- Cyclonaias_kleiniana **  
  +--+                               
     |  +----- Cyclonaias_tuberculata **
     +--+                            
        |  +-- Cyclonaias_succissa **   
        +--+                         
           +-- Cyclonaias_pustulosa **  

Campbell & Lydeard, 2012, fig. 2

iT tT bi riS oT roT roS iD
5 5 2 5 39 1 1 4.0
  +----------- Quadrula_quadrula     
  |                                  
--+        +-- Cyclonaias_infucata **   
  |  +-----+                         
  |  |     +-- Cyclonaias_kleiniana **  
  +--+                               
     |  +----- Cyclonaias_tuberculata **
     +--+                            
        |  +-- Cyclonaias_succissa **   
        +--+                         
           +-- Cyclonaias_pustulosa **  

Campbell & Lydeard, 2012, fig. 3

iT tT bi riS oT roT roS iD
5 6 3 5 46 2 2 5.0
  +-------------- Uniomerus_declivis    
  |                                     
--+           +-- Cyclonaias_infucata **   
  |  +--------+                         
  |  |        +-- Cyclonaias_kleiniana **  
  +--+                                  
     |  +-------- Quadrula_quadrula     
     +--+                               
        |  +----- Cyclonaias_tuberculata **
        +--+                            
           |  +-- Cyclonaias_succissa **   
           +--+                         
              +-- Cyclonaias_pustulosa **  

Campbell & Lydeard, 2012, fig. 4

iT tT bi riS oT roT roS iD
5 5 2 5 39 1 1 4.0
  +----------- Quadrula_quadrula     
  |                                  
--+        +-- Cyclonaias_infucata **   
  |  +-----+                         
  |  |     +-- Cyclonaias_kleiniana **  
  +--+                               
     |  +----- Cyclonaias_tuberculata **
     +--+                            
        |  +-- Cyclonaias_succissa **   
        +--+                         
           +-- Cyclonaias_pustulosa **  

Campbell & Lydeard, 2012, fig. 1-2

iT tT bi riS oT roT roS iD
5 5 2 5 69 6 4 4.0
  +-------------- Tritogonia_verrucosa  
  |                                     
  |           +-- Quadrula_quadrula     
--+  +--------+                         
  |  |        +-- Tritogonia_nobilis    
  |  |                                  
  +--+        +-- Cyclonaias_infucata **   
     |  +-----+                         
     |  |     +-- Cyclonaias_kleiniana **  
     +--+                               
        |  +----- Cyclonaias_tuberculata **
        +--+                            
           |  +-- Cyclonaias_pustulosa **  
           +--+                         
              +-- Cyclonaias_succissa **   

Campbell & Lydeard, 2012, fig. 3

iT tT bi riS oT roT roS iD
5 5 2 5 69 6 4 4.0
              +-- Tritogonia_verrucosa  
  +-----------+                         
  |           +-- Tritogonia_nobilis    
  |                                     
--+  +----------- Quadrula_quadrula     
  |  |                                  
  |  +----------- Theliderma_metanevra  
  +--+                                  
     |        +-- Cyclonaias_infucata **   
     |  +-----+                         
     |  |     +-- Cyclonaias_kleiniana **  
     +--+                               
        |  +----- Cyclonaias_tuberculata **
        +--+                            
           |  +-- Cyclonaias_pustulosa **  
           +--+                         
              +-- Cyclonaias_succissa **   

Zhou et al., 2007, fig. 2

iT tT bi riS oT roT roS iD
3 4 2 3 18 4 3 9.0
              +-- Theliderma_intermedia
  +-----------+                        
  |           +-- Theliderma_sparsa    
--+                                    
  |  +----------- Quadrula_quadrula    
  +--+                                 
     |  +-------- Cyclonaias_infucata **  
     +--+                              
        |  +----- Tritogonia_verrucosa 
        +--+                           
           |  +-- Cyclonaias_kieneriana **
           +--+                        
              +-- Cyclonaias_succissa **  

Boyer et al., 2011, fig. 3

iT tT bi riS oT roT roS iD
3 3 1 3 32 4 4 6.0
  +----------- Tritogonia_verrucosa  
--+                                  
  |  +-------- Theliderma_metanevra  
  +--+                               
     |  +----- Cyclonaias_pustulosa **  
     +--+                            
        |  +-- Cyclonaias_tuberculata **
        +--+                         
           +-- Cyclonaias_nodulata **   

Boyer et al., 2011, fig. 4

iT tT bi riS oT roT roS iD
3 3 1 3 35 4 4 6.0
  +----------- Theliderma_metanevra  
  |                                  
--+        +-- Tritogonia_verrucosa  
  |  +-----+                         
  |  |     +-- Quadrula_quadrula     
  +--+                               
     |  +----- Cyclonaias_pustulosa **  
     +--+                            
        |  +-- Cyclonaias_tuberculata **
        +--+                         
           +-- Cyclonaias_nodulata **   

Pfeiffer et al., 2016, fig. 2

iT tT bi riS oT roT roS iD
3 3 1 3 35 3 3 6.0
  +----------- Quadrula_quadrula     
--+                                  
  |  +-------- Tritogonia_verrucosa  
  +--+                               
     |  +----- Cyclonaias_pustulosa **  
     +--+                            
        |  +-- Cyclonaias_petrina **    
        +--+                         
           +-- Cyclonaias_tuberculata **

Inoue et al., 2018, fig. 1, 3a-c

iT tT bi riS oT roT roS iD
3 6 0 3 201 5 4 11.0
     +-- Uniomerus_declivis     
  +--+                          
  |  +-- Uniomerus_tetralasmus  
  |                             
  |  +-- Cyclonaias_tuberculata ** 
--+  |                          
  |  +-- Cyclonaias_pustulosa[1] **
  |  |                          
  |  +-- Theliderma_metanevra   
  +--+                          
     +-- Cyclonaias_pustulosa[2] **
     |                          
     +-- Quadrula_quadrula      
     |                          
     +-- Tritogonia_verrucosa   

Lydeard et al., 2000, fig. 2a

iT tT bi riS oT roT roS iD
2 2 0 2 15 1 1 --
  +----- Quadrula_quadrula  
--+                         
  |  +-- Cyclonaias_infucata **
  +--+                      
     +-- Cyclonaias_succissa **

Lydeard et al., 2000, fig. 2b

iT tT bi riS oT roT roS iD
2 3 0 2 15 1 1 8.0
  +-- Quadrula_quadrula  
  |                      
--+-- Cyclonaias_infucata **
  |                      
  +-- Cyclonaias_succissa **

Lydeard et al., 2000, fig. 2c

iT tT bi riS oT roT roS iD
2 2 0 2 14 1 1 --
  +----- Quadrula_quadrula  
--+                         
  |  +-- Cyclonaias_infucata **
  +--+                      
     +-- Cyclonaias_succissa **

Lydeard et al., 2000, fig. 2d

iT tT bi riS oT roT roS iD
2 3 0 2 14 1 1 8.0
  +-- Quadrula_quadrula  
  |                      
--+-- Cyclonaias_infucata **
  |                      
  +-- Cyclonaias_succissa **

Breton et al., 2011, fig. 3

iT tT bi riS oT roT roS iD
2 2 0 2 37 2 2 --
  +----- Quadrula_quadrula     
--+                            
  |  +-- Cyclonaias_tuberculata **
  +--+                         
     +-- Cyclonaias_pustulosa **  

Chase et al., 2018, fig. 4

iT tT bi riS oT roT roS iD
2 2 0 2 37 1 1 --
  +----- Megalonaias_nervosa   
--+                            
  |  +-- Cyclonaias_tuberculata **
  +--+                         
     +-- Cyclonaias_pustulosa **  

Chapman et al., 2008, fig. 1

iT tT bi riS oT roT roS iD
1 1 0 1 20 1 1 --
  +-- Quadrula_quadrula   
--+                       
  +-- Cyclonaias_pustulosa **

Curole & Kocher, 2005, fig. 1

iT tT bi riS oT roT roS iD
1 1 0 1 10 1 1 --
  +-- Quadrula_quadrula   
--+                       
  +-- Cyclonaias_pustulosa **

Curole & Kocher, 2005, fig. 1-M

iT tT bi riS oT roT roS iD
1 1 0 1 11 1 1 --
  +-- Quadrula_quadrula   
--+                       
  +-- Cyclonaias_pustulosa **

Curole & Kocher, 2002, fig. 1-F

iT tT bi riS oT roT roS iD
1 1 0 1 3 1 1 --
  +-- Quadrula_quadrula   
--+                       
  +-- Cyclonaias_pustulosa **

Curole & Kocher, 2002, fig. 1a-M

iT tT bi riS oT roT roS iD
1 1 0 1 3 1 1 --
  +-- Quadrula_quadrula   
--+                       
  +-- Cyclonaias_pustulosa **

Curole & Kocher, 2002, fig. 1b-M

iT tT bi riS oT roT roS iD
1 1 0 1 3 1 1 --
  +-- Quadrula_quadrula   
--+                       
  +-- Cyclonaias_pustulosa **

Chapman et al., 2008, fig. 2

iT tT bi riS oT roT roS iD
1 1 0 1 20 1 1 --
  +-- Quadrula_quadrula   
--+                       
  +-- Cyclonaias_pustulosa **

Pfeiffer et al., 2019, fig. 1

iT tT bi riS oT roT roS iD
1 1 0 1 38 4 4 --
  +----- Tritogonia_verrucosa  
--+                            
  |  +-- Cyclonaias_tuberculata **
  +--+                         
     +-- Quadrula_quadrula     

Pieri et al., 2018, fig. 3

iT tT bi riS oT roT roS iD
1 1 0 1 36 1 1 --
  +-- Quadrula_quadrula 
--+                     
  +-- Cyclonaias_petrina **

Inoue et al., 2020, fig. 3

iT tT bi riS oT roT roS iD
1 1 0 1 43 1 1 --
  +-- Cyclonaias_petrina **
--+                     
  +-- Quadrula_quadrula 

Inoue et al., 2020, fig. 4

iT tT bi riS oT roT roS iD
1 1 0 1 42 1 1 --
  +-- Cyclonaias_petrina **
--+                     
  +-- Quadrula_quadrula 

Porto-Hannes et al., 2021, fig. 4

iT tT bi riS oT roT roS iD
1 1 0 1 11 1 1 --
No taxon cladogram to show.

Page updated: 12:50:26 Thu 01 Dec 2022

 
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