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Page last updated
Wed 27 Sep 2023

The Freshwater Mussels (Unionoida) of the World (and other less consequential bivalves)

FM(U)otW(aolcb) is the web version of the MUSSEL Project Database. Follow the links to browse the data or use the custom Google search field. Either way, you win!

Cladomics pages are experimental — even more so than the rest of this site!

The purpose of the cladomics pages for each genus and family-group level taxon is to provide statistics and references indicating the extent to which published phylogenetic work supports the classification used here.

The Cladomics Summary provides an overview of the number of studies that have included terminal species representing this taxon as well as an indication of how frequently it has been recovered as monophyletic.

Following the summary, the available published clades representing the taxon are listed.

For more information, visit the Cladomics FARQ (frequently asked rhetorical questions) page. Abbreviations for the various statistics are explained on the FARQ!

Search the database pages:

Unionoida | Unionoidea | Unionidae | Unioninae | Anodontini | Alasmidontina

genus Utterbackiana Frierson, 1927: cladomics
     5 valid species

Cladomics Summary

number of species = 5

analyzed in 19 cladograms in 11 publications
maximum number of terminals in an analysis = 4, mean = 2.47

Basic Monophyly

number of cladograms that test basic monophyly (i.e., more than 1 included species) = 10
maximum number of included terminals = 4, mean = 3.8
proportion of cladograms that support basic monophyly = 0.9

Pairwise Cladogram Clustering

The graphs below depict in two dimensions all pair-wise ingroup distances within the cladome. See the Cladmoics FARQ for more information.


PCoA Ordination


NJ tree

Whelan et al., 2023 [Publ 4287], fig. 2

iT tT bi riS oT roT roS iD
4 4 2 4 40 32 12 0.0

*** Top Monophyly Cladogram *** *** Top Sister-Group Cladogram ***

                       +-- Utterbackiana_couperiana **        
                    +--+                                   
                    |  +-- Utterbackiana_heardi **            
              +-----+                                      
              |     |  +-- Utterbackiana_suborbiculata **     
              |     +--+                                   
              |        +-- Utterbackiana_hartfieldorum **     
  +-----------+                                            
  |           |     +----- Lasmigona_costata               
  |           |  +--+                                      
  |           |  |  |  +-- Pseudodontoideus_subvexus       
  |           |  |  +--+                                   
  |           +--+     +-- Pseudodontoideus_connasaugaensis
  |              |                                         
  |              |  +----- Lasmigona_complanata            
  |              +--+                                      
  |                 |  +-- Arcidens_confragosus            
  |                 +--+                                   
  |                    +-- Arcidens_wheeleri               
--+                                                        
  |     +----------------- Pressodonta_viridis             
  |     |                                                  
  |  +--+              +-- Anodontoides_denigratus         
  |  |  |  +-----------+                                   
  |  |  |  |           +-- Anodontoides_ferussacianus      
  |  |  +--+                                               
  |  |     |  +----------- Strophitus_undulatus            
  |  |     +--+                                            
  |  |        |  +-------- Strophitus_pascagoulaensis[2]   
  |  |        +--+                                         
  |  |           |  +----- Strophitus_radiatus             
  +--+           +--+                                      
     |              |  +-- Strophitus_pascagoulaensis[1]   
     |              +--+                                   
     |                 +-- Strophitus_williamsi            
     |                                                     
     |              +----- Prolasmidonta_heterodon         
     |  +-----------+                                      
     |  |           |  +-- Platynaias_subviridis           
     |  |           +--+                                   
     |  |              +-- Platynaias_compressa            
     +--+                                                  
        |              +-- Alasmidonta_holstonia           
        |  +-----------+                                   
        |  |           +-- Alasminota_etowaensis           
        |  |                                               
        +--+           +-- Alasmidonta_raveneliana         
           |  +--------+                                   
           |  |        +-- Alasmidonta_marginata           
           |  |                                            
           +--+        +-- Alasmidonta_varicosa            
              |  +-----+                                   
              |  |     +-- Alasmidonta_uwharriensis        
              +--+                                         
                 |  +----- Alasmidonta_undulata            
                 +--+                                      
                    |  +-- Alasmidonta_triangulata         
                    +--+                                   
                       +-- Alasmidonta_arcula

Zanatta et al., 2007 [Publ 2698], fig. 1A

iT tT bi riS oT roT roS iD
4 4 2 4 12 10 4 2.0 
  +-------- Utterbackiana_implicata **    
--+                                    
  |  +----- Utterbackiana_suborbiculata **
  +--+                                 
     |  +-- Utterbackiana_heardi **       
     +--+                              
        +-- Utterbackiana_couperiana **

Zanatta et al., 2007 [Publ 2698], fig. 1B

iT tT bi riS oT roT roS iD
4 4 2 4 12 10 4 4.0 
  +-------- Utterbackiana_implicata **    
--+                                    
  |  +----- Utterbackiana_heardi **       
  +--+                                 
     |  +-- Utterbackiana_suborbiculata **
     +--+                              
        +-- Utterbackiana_couperiana **

Hoeh, 1990 [Publ 2717], fig. 2

iT tT bi riS oT roT roS iD
4 4 2 4 12 10 4 4.0 
  +-------- Utterbackiana_implicata **    
--+                                    
  |  +----- Utterbackiana_couperiana[2] **
  +--+                                 
     |  +-- Utterbackiana_suborbiculata **
     +--+                              
        +-- Utterbackiana_couperiana[1] **

Hoeh, 1990 [Publ 2717], fig. 3

iT tT bi riS oT roT roS iD
4 16 2 4 12 10 4 16.0 
  +----- Pyganodon_fragilis         
  |                                 
  +----- Utterbackia_imbecillis     
  |                                 
  +----- Utterbackia_peggyae        
  |                                 
  +----- Anodonta_cygnea            
  |                                 
  +----- Anodonta_kennerlyi         
  |                                 
  +----- Utterbackiana_couperiana[1] **
  |                                 
--+----- Utterbackiana_couperiana[2] **
  |                                 
  +----- Utterbackiana_suborbiculata **
  |                                 
  |  +-- Lasmigona_costata          
  +--+                              
  |  +-- Utterbackiana_implicata **    
  |                                 
  |  +-- Lasmigona_complanata       
  |  |                              
  |  +-- Platynaias_compressa       
  |  |                              
  |  +-- Pyganodon_cataracta        
  +--+                              
     +-- Pyganodon_gibbosa          
     |                              
     +-- Pyganodon_grandis          
     |                              
     +-- Pyganodon_lacustris

Hoeh, 1990 [Publ 2717], fig. 4

iT tT bi riS oT roT roS iD
4 4 2 4 12 10 4 4.0 
  +-------- Utterbackiana_implicata **    
--+                                    
  |  +----- Utterbackiana_couperiana[2] **
  +--+                                 
     |  +-- Utterbackiana_suborbiculata **
     +--+                              
        +-- Utterbackiana_couperiana[1] **

Smith et al., 2018 [Publ 3790], fig. 1

iT tT bi riS oT roT roS iD
4 4 2 4 15 14 7 0.0 
           +-- Alasmidonta_triangulata         
  +--------+                                   
  |        +-- Alasminota_etowaensis           
  |                                            
--+        +-- Pseudodontoideus_connasaugaensis
  |  +-----+                                   
  |  |     +-- Pseudodontoideus_subvexus       
  |  |                                         
  +--+     +-- Utterbackiana_couperiana **        
     |  +--+                                   
     |  |  +-- Utterbackiana_heardi **            
     +--+                                      
        |  +-- Utterbackiana_hartfieldorum **     
        +--+                                   
           +-- Utterbackiana_suborbiculata **

Whelan et al., 2023 [Publ 4287], fig. 3

iT tT bi riS oT roT roS iD
4 4 2 4 25 24 11 0.0 
                             +-- Utterbackiana_heardi **            
                          +--+                                   
                          |  +-- Utterbackiana_couperiana **        
  +-----------------------+                                      
  |                       |  +-- Utterbackiana_hartfieldorum **     
  |                       +--+                                   
  |                          +-- Utterbackiana_suborbiculata **     
  |                                                              
  |                          +-- Utterbackia_peninsularis        
  |  +-----------------------+                                   
--+  |                       +-- Pyganodon_grandis               
  |  |                                                           
  |  |     +-------------------- Alasminota_etowaensis           
  |  |     |                                                     
  |  |     |              +----- Pressodonta_viridis             
  |  |  +--+  +-----------+                                      
  |  |  |  |  |           |  +-- Platynaias_subviridis           
  |  |  |  |  |           +--+                                   
  +--+  |  |  |              +-- Prolasmidonta_heterodon         
     |  |  +--+                                                  
     |  |     |              +-- Anodontoides_ferussacianus      
     |  |     |  +-----------+                                   
     |  |     |  |           +-- Anodontoides_denigratus         
     |  |     +--+                                               
     |  |        |  +----------- Strophitus_undulatus            
     |  |        +--+                                            
     |  |           |  +-------- Strophitus_pascagoulaensis[2]   
     |  |           +--+                                         
     +--+              |  +----- Strophitus_pascagoulaensis[1]   
        |              +--+                                      
        |                 |  +-- Strophitus_radiatus             
        |                 +--+                                   
        |                    +-- Strophitus_williamsi            
        |                                                        
        |                    +-- Arcidens_confragosus            
        |                 +--+                                   
        |                 |  +-- Arcidens_wheeleri               
        |        +--------+                                      
        |        |        |  +-- Pseudodontoideus_connasaugaensis
        |        |        +--+                                   
        |        |           +-- Pseudodontoideus_subvexus       
        +--------+                                               
                 |           +-- Alasmidonta_marginata           
                 |  +--------+                                   
                 |  |        +-- Alasmidonta_raveneliana         
                 |  |                                            
                 +--+        +-- Alasmidonta_varicosa            
                    |  +-----+                                   
                    |  |     +-- Alasmidonta_uwharriensis        
                    +--+                                         
                       |  +----- Alasmidonta_undulata            
                       +--+                                      
                          |  +-- Alasmidonta_triangulata         
                          +--+                                   
                             +-- Alasmidonta_arcula

Whelan et al., 2023 [Publ 4287], fig. 4

iT tT bi riS oT roT roS iD
4 4 1 4 25 23 10 3.0 
                 +----- Utterbackiana_couperiana **           
                 |                                         
     +-----------+  +-- Utterbackiana_heardi **               
     |           |  |                                      
     |           +--+-- Utterbackiana_hartfieldorum **        
     |              |                                      
  +--+              +-- Utterbackiana_suborbiculata **        
  |  |                                                     
  |  |  +-------------- Alasminota_etowaensis              
  |  |  |                                                  
  |  +--+           +-- Arcidens_confragosus               
  |     |  +--------+                                      
  |     |  |        +-- Arcidens_wheeleri                  
  |     +--+                                               
  |        |  +-------- Pseudodontoideus_connasaugaensis[3]
  |        +--+                                            
  |           |  +----- Pseudodontoideus_connasaugaensis[2]
  |           +--+                                         
--+              |  +-- Pseudodontoideus_subvexus          
  |              +--+                                      
  |                 +-- Pseudodontoideus_connasaugaensis[1]
  |                                                        
  |              +----- Alasmidonta_raveneliana            
  |              |                                         
  |              |  +-- Alasmidonta_uwharriensis           
  |  +-----------+--+                                      
  |  |           |  +-- Alasmidonta_varicosa               
  |  |           |                                         
  |  |           |  +-- Alasmidonta_arcula                 
  |  |           +--+                                      
  +--+              +-- Alasmidonta_triangulata            
     |                                                     
     |  +-------------- Platynaias_subviridis              
     |  |                                                  
     |  |     +-------- Strophitus_pascagoulaensis         
     |  |  +--+                                            
     +--+  |  |  +----- Strophitus_undulatus               
        |  |  +--+                                         
        |  |     |  +-- Strophitus_radiatus                
        |  |     +--+                                      
        +--+        +-- Strophitus_williamsi               
           |                                               
           |        +-- Pyganodon_grandis                  
           |  +-----+                                      
           |  |     +-- Pyganodon_cataracta                
           +--+                                            
              |  +----- Utterbackia_peninsularis           
              +--+                                         
                 |  +-- Anodontoides_ferussacianus         
                 +--+                                      
                    +-- Anodontoides_denigratus

Lydeard et al., 1996 [Publ 2656], fig. 3

iT tT bi riS oT roT roS iD
2 2 0 2 25 4 4 -- 
     +----- Utterbackia_imbecillis  
  +--+                              
  |  |  +-- Utterbackiana_couperiana **
  |  +--+                           
--+     +-- Utterbackiana **           
  |                                 
  |  +----- Strophitus_radiatus     
  +--+                              
     |  +-- Lasmigona_costata       
     +--+                           
        +-- Alasmidonta_triangulata

Lydeard et al., 1996 [Publ 2656], fig. 4

iT tT bi riS oT roT roS iD
1 1 0 1 27 4 4 -- 
-- Utterbackiana_couperiana

Huang et al., 2002 [Publ 2657], fig. 1

iT tT bi riS oT roT roS iD
1 1 0 1 33 4 4 -- 
  +----------- Utterbackiana_couperiana **
--+                                    
  |  +-------- Utterbackia_imbecillis  
  +--+                                 
     |  +----- Strophitus_radiatus     
     +--+                              
        |  +-- Alasmidonta_triangulata 
        +--+                           
           +-- Lasmigona_costata

Huang et al., 2002 [Publ 2657], fig. 2

iT tT bi riS oT roT roS iD
1 1 0 1 33 4 4 -- 
  +----- Utterbackia_imbecillis  
  |                              
--+----- Utterbackiana_couperiana **
  |                              
  |  +-- Alasmidonta_triangulata 
  +--+                           
     +-- Lasmigona_costata

Curole & Kocher, 2005 [Publ 2682], fig. 1-M

iT tT bi riS oT roT roS iD
1 1 0 1 11 1 1 -- 
  +-- Utterbackiana_implicata **
--+                          
  +-- Pyganodon_grandis

Krebs et al., 2003 [Publ 2691], fig. 3

iT tT bi riS oT roT roS iD
1 1 0 1 33 8 6 -- 
-- Utterbackiana_couperiana

Curole & Kocher, 2002 [Publ 2712], fig. 1a-M

iT tT bi riS oT roT roS iD
1 1 0 1 3 0 0 -- 
-- Utterbackiana_implicata

Curole & Kocher, 2002 [Publ 2712], fig. 1b-M

iT tT bi riS oT roT roS iD
1 1 0 1 3 0 0 -- 
-- Utterbackiana_implicata

Turner et al., 2000 [Publ 2722], fig. 2

iT tT bi riS oT roT roS iD
1 1 0 1 15 0 0 -- 
-- Utterbackiana_couperiana

Pfeiffer et al., 2016 [Publ 3660], fig. 2

iT tT bi riS oT roT roS iD
1 1 0 1 38 0 0 -- 
-- Utterbackiana_suborbiculata

Page updated: 10:55:15 Wed 27 Sep 2023
 
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