MUSSELpdb | tribe Castaliini

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Page last updated
Sun 07 Sep 2025

The Freshwater Mussels (Unionoida) of the World (and other less consequential bivalves)

FM(U)otW(aolcb) is the web version of the MUSSEL Project Database. Follow the links to browse the data or use the custom Google search field. Either way, you win!

Cladomics pages are experimental — even more so than the rest of this site!

The purpose of the cladomics pages for each genus and family-group level taxon is to provide statistics and references indicating the extent to which published phylogenetic work supports the classification used here.

The Cladomics Summary provides an overview of the number of studies that have included terminal species representing this taxon as well as an indication of how frequently it has been recovered as monophyletic.

Following the summary, the available published clades representing the taxon are listed.

For more information, visit the Cladomics FARQ (frequently asked rhetorical questions) page. Abbreviations for the various statistics are explained on the FARQ!

The MUSSEL Project Database has moved to a new server, with a new URL! The new address for this page is: https://musselpdb.org/db.php?ty=validfam_cld&id=Castaliini

Search the database pages:

Bivalvia | Unionoida | Hyrioidea | Hyriidae | Hyriinae

tribe CASTALIINI Morretes, 1949: cladomics
3 genera

Cladomics Summary

number of species = 17
number of subtaxa (rank: genus) = 3

analyzed in 37 cladograms in 24 publications
maximum number of terminals in an analysis = 4, mean = 1.32
maximum number of subtaxa (genus) in an analysis = 3, mean = 1.22

Basic Monophyly

number of cladograms that test basic monophyly (i.e., more than 1 included species) = 5
maximum number of included terminals = 4, mean = 3.4
maximum number of subtaxa (genus) = 3, mean = 2.6
proportion of cladograms that support basic monophyly = 1.0

Rigorous Monophyly

number of cladograms that test rigorous monophyly = 4
(Rigorous monophyly refers to cladograms with terminal species representing more than one subtaxon (genus) taxon and at least one rigorous outgroup (tribe).
maximum number of included terminals = 4, mean = 3.5
maximum number of subtaxa (genus) = 3, mean = 2.75
proportion of cladograms that support rigorous monophyly = 1.0

Pairwise Cladogram Clustering

The graphs below depict in two dimensions all pair-wise ingroup distances within the cladome. See the Cladmoics FARQ for more information.

PCoA Ordination

NJ tree

Santos-Neto et al., 2016 [Publ 3739], fig. 2a_b

iT	tT	bi	riS	oT	roT	roS	iD
4	4	5	3	16	9	4	0.0

*** Top Monophyly Cladogram *** *** Top Sister-Group Cladogram ***

  +-------------------- Rhipidodontini     
  |                                        
--+  +----------------- Hyriini            
  |  |                                     
  +--+  +-------------- Castalia_stevensi **  
     |  |                                  
     +--+           +-- Callonaia **          
        |  +--------+                      
        |  |        +-- Castalia_ambigua__1 **
        +--+                               
           |  +-------- Castalia_ambigua__4 **
           +--+                            
              |  +----- Castalia_ambigua__2 **
              +--+                         
                 |  +-- Castaliella **        
                 +--+                      
                    +-- Castalia_ambigua__3 **

Santos-Neto et al., 2016 [Publ 3739], fig. 4

iT	tT	bi	riS	oT	roT	roS	iD
4	4	5	3	16	9	4	3.0

  +----------------------- Rhipidodontini     
--+                                           
  |  +-------------------- Hyriini            
  +--+                                        
     |  +----------------- Castalia_ambigua__4 **
     +--+                                     
        |  +-------------- Callonaia **          
        +--+                                  
           |  +----------- Castalia_ambigua__3 **
           +--+                               
              |  +-------- Castalia_stevensi **  
              +--+                            
                 |  +----- Castalia_ambigua__2 **
                 +--+                         
                    |  +-- Castaliella **        
                    +--+                      
                       +-- Castalia_ambigua__1 **

Santos-Neto et al., 2018 [Publ 3888], fig. 5

iT	tT	bi	riS	oT	roT	roS	iD
3	3	1	2	2	2	1	6.0

  +----- Hyriini  
--+               
  |  +-- Callonaia **
  +--+            
     +-- Castalia **

Santos-Neto et al., 2016 [Publ 3739], fig. 3_5

iT	tT	bi	riS	oT	roT	roS	iD
3	3	2	3	4	4	2	4.0

  +-------------- Rhipidodontini     
--+                                  
  |  +----------- Hyriini            
  +--+                               
     |  +-------- Callonaia **          
     +--+                            
        |  +----- Castalia_ambigua__2 **
        +--+                         
           |  +-- Castalia_ambigua__1 **
           +--+                      
              +-- Castaliella **

Olivera-Hyde et al., 2020 [Publ 4249], fig. 10

iT	tT	bi	riS	oT	roT	roS	iD
3	3	1	2	0	0	0	6.0

  +-- Callonaia **
--+            
  +-- Castalia **

Huang et al., 2019 [Publ 3843], fig. S4

iT	tT	bi	riS	oT	roT	roS	iD
1	1	0	1	198	6	4	--

  +----- Rhipidodontini
--+                    
  |  +-- Castalia **      
  +--+                 
     +-- Hyriini

Bogan & Hoeh, 2000 [Publ 2650], fig. 1

iT	tT	bi	riS	oT	roT	roS	iD
1	1	0	1	33	2	2	--

  +-- Castalia **      
--+                 
  +-- Rhipidodontini

Huang et al., 2019 [Publ 3843], fig. S3

iT	tT	bi	riS	oT	roT	roS	iD
1	1	0	1	196	6	4	--

  +----------- Echyridella   
--+                          
  |  +-------- Hyridellini   
  +--+                       
     |  +----- Rhipidodontini
     +--+                    
        |  +-- Castalia **      
        +--+                 
           +-- Hyriini

Wu et al., 2021 [Publ 4366], fig. 6

iT	tT	bi	riS	oT	roT	roS	iD
1	1	0	1	90	1	1	--

  +-- Rhipidodontini
--+                 
  +-- Castalia **

Konopleva et al., 2017 [Publ 3807], fig. 6

iT	tT	bi	riS	oT	roT	roS	iD
1	1	0	1	30	2	2	--

  +----- Rhipidodontini
--+                    
  |  +-- Hyriini       
  +--+                 
     +-- Castalia **

Hoeh et al., 2001 [Publ 2652], fig. 14f4

iT	tT	bi	riS	oT	roT	roS	iD
1	1	0	1	30	2	2	--

  +-- Castalia **      
--+                 
  +-- Rhipidodontini

Graf & Cummings, 2006 [Publ 1968], fig. 2

iT	tT	bi	riS	oT	roT	roS	iD
1	1	0	1	46	5	3	--

  +-------- Echyridella   
--+                       
  |  +----- Hyridellini   
  +--+                    
     |  +-- Castalia **      
     +--+                 
        +-- Rhipidodontini

Graf & Ó Foighil, 2000 [Publ 2649], fig. 1_2

iT	tT	bi	riS	oT	roT	roS	iD
1	1	0	1	10	4	3	--

           +-- Rhipidodontini        
  +--------+                         
  |        +-- Castalia **              
--+                                  
  |  +-------- Hyridella_australis__2
  +--+                               
     |  +----- Hyridella_australis__1
     +--+                            
        |  +-- Echyridella           
        +--+                         
           +-- Hyridella_depressa

Bolotov et al., 2016 [Publ 3743], fig. 4

iT	tT	bi	riS	oT	roT	roS	iD
1	1	0	1	29	2	2	--

  +----- Rhipidodontini
--+                    
  |  +-- Hyriini       
  +--+                 
     +-- Castalia **

Huang et al., 2019 [Publ 3843], fig. 2

iT	tT	bi	riS	oT	roT	roS	iD
1	1	0	1	198	6	4	--

  +----------- Echyridella   
--+                          
  |  +-------- Hyridellini   
  +--+                       
     |  +----- Rhipidodontini
     +--+                    
        |  +-- Castalia **      
        +--+                 
           +-- Hyriini

Hoeh et al., 2001 [Publ 2652], fig. 14f2

iT	tT	bi	riS	oT	roT	roS	iD
1	1	0	1	30	2	2	--

-- Castalia_stevensi

Whelan et al., 2011 [Publ 3380], fig. 1-ML

iT	tT	bi	riS	oT	roT	roS	iD
1	1	0	1	35	2	2	--

  +----- Hyridellini
--+                 
  |  +-- Castalia **   
  +--+              
     +-- Hyriini

Hoeh et al., 2009 [Publ 2716], fig. 3

iT	tT	bi	riS	oT	roT	roS	iD
1	1	0	1	46	5	3	--

  +-------- Echyridella   
--+                       
  |  +----- Hyridellini   
  +--+                    
     |  +-- Castalia **      
     +--+                 
        +-- Rhipidodontini

Guerra et al., 2019 [Publ 4168], fig. 1

iT	tT	bi	riS	oT	roT	roS	iD
1	1	0	1	40	3	3	--

  +----- Rhipidodontini
--+                    
  |  +-- Castalia **      
  +--+                 
     +-- Hyriini

Bolotov et al., 2017 [Publ 3761], fig. S1

iT	tT	bi	riS	oT	roT	roS	iD
1	1	0	1	138	2	2	--

  +----- Rhipidodontini
--+                    
  |  +-- Castalia **      
  +--+                 
     +-- Hyriini

Graf, 2000 [Publ 2647], fig. 1

iT	tT	bi	riS	oT	roT	roS	iD
1	1	0	1	16	2	2	--

-- Castalia

Hoeh et al., 2002 [Publ 2653], fig. 2

iT	tT	bi	riS	oT	roT	roS	iD
1	1	0	1	30	2	2	--

  +-- Castalia **      
--+                 
  +-- Rhipidodontini

Baker et al., 2004 [Publ 2685], fig. 2-3

iT	tT	bi	riS	oT	roT	roS	iD
1	1	0	1	8	4	3	--

  +-- Echyridella
--+              
  +-- Castalia **

Pfeiffer & Graf, 2015 [Publ 3620], fig. 2

iT	tT	bi	riS	oT	roT	roS	iD
1	1	0	1	39	2	2	--

  +----- Hyridellini
--+                 
  |  +-- Hyriini    
  +--+              
     +-- Castalia **

Roe & Hoeh, 2003 [Publ 2655], fig. 4f2

iT	tT	bi	riS	oT	roT	roS	iD
1	1	0	1	30	2	2	--

-- Castalia_stevensi

Bolotov et al., 2017 [Publ 3761], fig. S2

iT	tT	bi	riS	oT	roT	roS	iD
1	1	0	1	138	2	2	--

  +----- Rhipidodontini
--+                    
  |  +-- Castalia **      
  +--+                 
     +-- Hyriini

Whelan et al., 2011 [Publ 3380], fig. 1-MP

iT	tT	bi	riS	oT	roT	roS	iD
1	1	0	1	35	2	2	--

  +----- Hyridellini
--+                 
  |  +-- Castalia **   
  +--+              
     +-- Hyriini

Bolotov et al., 2016 [Publ 3743], fig. 2

iT	tT	bi	riS	oT	roT	roS	iD
1	1	0	1	29	2	2	--

  +----- Rhipidodontini
--+                    
  |  +-- Hyriini       
  +--+                 
     +-- Castalia **

Walker et al., 2006 [Publ 2666], fig. 6

iT	tT	bi	riS	oT	roT	roS	iD
1	1	0	1	35	4	3	--

-- Castalia_stevensi

Baker et al., 2003 [Publ 2690], fig. 5

iT	tT	bi	riS	oT	roT	roS	iD
1	1	0	1	7	4	3	--

  +-- Echyridella
--+              
  +-- Castalia **

Guerra et al., 2019 [Publ 4168], fig. 2

iT	tT	bi	riS	oT	roT	roS	iD
1	1	0	1	40	3	3	--

  +-------- Echyridella   
--+                       
  |  +----- Rhipidodontini
  +--+                    
     |  +-- Castalia **      
     +--+                 
        +-- Hyriini

Graf et al., 2015 [Publ 3619], fig. 1

iT	tT	bi	riS	oT	roT	roS	iD
1	1	0	1	19	6	3	--

  +-------- Hyridellini   
--+                       
  |  +----- Rhipidodontini
  +--+                    
     |  +-- Hyriini       
     +--+                 
        +-- Castalia **

Hoeh et al., 2009 [Publ 2716], fig. 5

iT	tT	bi	riS	oT	roT	roS	iD
1	1	0	1	45	5	3	--

  +-------- Echyridella   
--+                       
  |  +----- Hyridellini   
  +--+                    
     |  +-- Castalia **      
     +--+                 
        +-- Rhipidodontini

Roe & Hoeh, 2003 [Publ 2655], fig. 4f3

iT	tT	bi	riS	oT	roT	roS	iD
1	1	0	1	30	2	2	--

  +-- Castalia **      
--+                 
  +-- Rhipidodontini

Wu et al., 2021 [Publ 4366], fig. 2_I

iT	tT	bi	riS	oT	roT	roS	iD
1	1	0	1	90	1	1	--

  +-- Castalia **      
--+                 
  +-- Rhipidodontini

Graf, 2002 [Publ 2651], fig. 2

iT	tT	bi	riS	oT	roT	roS	iD
1	1	0	1	17	2	2	--

  +----- Hyridellini   
--+                    
  |  +-- Rhipidodontini
  +--+                 
     +-- Castalia **

Hoeh et al., 2009 [Publ 2716], fig. 4

iT	tT	bi	riS	oT	roT	roS	iD
1	1	0	1	45	5	3	--

  +-------- Echyridella   
--+                       
  |  +----- Hyridellini   
  +--+                    
     |  +-- Castalia **      
     +--+                 
        +-- Rhipidodontini

Page updated: 08:15:27 Sun 07 Sep 2025

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