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Wed 06 Nov 2024 |
The Freshwater Mussels (Unionoida) of the World (and other less consequential bivalves)
FM(U)otW(aolcb) is the web version of the MUSSEL Project Database. Follow the links to browse the data or use the custom Google search field. Either way, you win!
This page documents data for Wu et al. (2021), such as genera and species attributed to that publication (including known misspellings, misidentifications, and other unavailable names) as well as taxonomic opinions, if available. Taxonomic opinions refer to uses of genera and species as valid and invalid in that work.
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Publication information
Wu, R., J. An, R. Chen, S. Ouyang & X. Wu. 2021. Taxonomy, phylogeny and evolution of freshwater mussels (Unionoida: Unionoidea) in China revealed by multilocus phylogenetic analyses and mitochondrial phylogenomics. Journal of Lake Sciences 33(6): 1788-1804.
Click here for online access.
Cladogram Figures
(click the figure number to see the cladogram) fig. 2_I
fig. 3-BI
fig. 3-ML
fig. 4
fig. 6
Attributed Species Names
Lasmigona compressa Graf & Ó Foighil, 2000
‘Lea’ of Authors.
Graf & Ó Foighil, 2000, J. Moll. Studies 66: 160.
Graf & Cummings, 2006, Zool. J. Linn. Soc. 148: 348.
Hoeh et al., 2009, Malacologia 51(2): 310.
Jadhav & Jamkhedkar, 2009, Res. J. BioTech. 4(1): 43.
Whelan et al., 2011, MPE 61: 506.
Lopes-Lima et al., 2017, MPE 106: 177.
Bolotov et al., 2017, Sci. Repts. 7: Table S1.
Bolotov et al., 2017, Sci. Repts. 7: Table S1.
Bolotov et al., 2018, Sci. Repts. 8: Table S1.
Huang et al., 2019, MPE 130: Table S4.
Lopes-Lima et al., 2020, MPE 146: Table 1.
Wu et al., 2021, J. Lake Sci. 33(6): attached Table 1.
Jeratthitikul et al., 2022, Sci. Repts. 12: Table S1.
Wu et al., 2023, ZooKeys 1166: 263, Table 1.
Liu et al., 2023, ZooKeys 1179: Table 1.
(misidentified reference to Symphynota compressa Lea, 1829)
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Unio crassus Soroka, 2005
‘Philippson’ of Authors.
Soroka, 2005, Zool. Sci. 22: 1138, Table 1.
Soroka, 2010, Folia Malac. 18(4): Table 2. [in part]
Reis et al., 2013, Graellsia 69(1): 22, Table 2.
Prié & Puillandre, 2014, Hydrobiologia 735(1): online resource 1 [as ‘Unio crassus crassus’].
Douda et al., 2014, J. Appl. Ecol. 51: 1087. [in part]
Froufe et al., 2016, Org Divers Evol 16(3): 606, fig. 3. [in part]
Klishko et al., 2017, Zootaxa 4286(1): 98, Table 3.
Soroka & Burzynski, 2017, mtDNA B 2(2): 863.
Burzynski et al., 2017, MPE 107: 606.
Lopes-Lima et al., 2017, Biol. Rev. 92(1): Table S1 [as ‘Unio crassus crassus’]. [in part]
Araujo et al., 2018, J. Linn. Soc., Zool. 182(2): supplementary table. [in part]
Klishko et al., 2018, Systemat. Biodiv. 16(3): 291.
Burzynski & Soroka, 2018, PeerJ 6, e5573: 3.
Wang et al., 2018, Conserv. Genet. Resour. 10(4): 611.
Wang et al., 2018, Conserv. Genet. Resour. 11: 140.
Cha et al., 2018, mtDNA B 3(2): 946.
Aunins et al., 2018, mtDNA B 3(2): 1125.
Wu et al., 2018, PLoS One 13(8), e0200956: 6. [in part]
Klishko et al., 2019, Ruthenica 29(1): 57, Table 1.
Wu et al., 2019, ZooKeys 812: 29.
Huang et al., 2019, MPE 130: Table S5. [in part]
Fukata & Iigo, 2019, mtDNA B 4(2): 3489.
Guerra et al., 2019, BMC Evol. Biol. 19 (229): Table S1.
Froufe et al., 2019, Heredity 124: 185.
Soroka, 2020, J. Zool. Syst. Evol. Res. 58(2): Table 2.
Fukata & Iigo, 2020, mtDNA B 5(2): 1216.
Sano et al., 2020, Plankton Benthos Res. 15(2): 99.
Wu et al., 2020, Invert. Syst. 34: Table S2.
Kilikowska et al., 2020, Life 10(7): 1. [in part]
Lopes-Lima et al., 2021, MPE 163: fig. 6 [by voucher number]. [in part]
Sayenko et al., 2021, Systemat. Biodiv. 19: 58, Table 1.
Wu et al., 2021, J. Lake Sci. 33(6): attached Table 2.
Zieritz et al., 2021, Zool. J. Linn. Soc. 193(3): 1092.
Kim et al., 2021, mtDNA B 6(1): 80.
Pfarrer et al., 2022, Hydrobiologia 849: ESM1.
Wu et al., 2022, Zool. Scripta 51: Table S2.
Dai et al., 2023, Invert. Syst. 37(2): Table 2.
Chen et al., 2023, ZooKeys 1173: Table 1.
Wu et al., 2023, Org Divers Evol (2023): Table 2.
Dai et al., 2024, Zool. Scripta 53: Table 1.
Wu et al., 2024, J. Zool. Syst. Evol. Res. 2024, 1499508: Table 2.
Wu et al., 2024, Zool. Scripta early online: Table S3.
Dai et al., 2024, Invert. Syst. 38(6): Table 3. [in part]
(misidentified reference to Unio crassus Retzius, 1788)
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Anodontites elongata Whelan et al., 2011
‘Swainson’ of Authors.
Whelan et al., 2011, MPE 61: 506.
Graf et al., 2015, MPE 85: 3.
Bolotov et al., 2016, MPE 103: 108.
Lopes-Lima et al., 2017, MPE 106: 178.
Bolotov et al., 2017, Sci. Repts. 7: Table S1.
Bolotov et al., 2017, Sci. Repts. 7: Table S1.
Konopleva et al., 2017, Systemat. Biodiv. 15(3): 208.
Combosch et al., 2017, MPE 107: 195.
Bolotov et al., 2018, Sci. Repts. 8: Table S1.
Huang et al., 2018, MPE 120: 302.
Lopes-Lima et al., 2018, MPE 127: 103.
Huang et al., 2019, MPE 130: Table S4.
Bolotov et al., 2019, Sci. Repts. 9: Table S1.
Pfeiffer et al., 2019, MPE 137: 116.
Guerra et al., 2019, BMC Evol. Biol. 19 (229): 3.
Bolotov et al., 2020, Sci. Repts. 10: Table S1.
Sano et al., 2020, Plankton Benthos Res. 15(2): 101.
Wu et al., 2020, Invert. Syst. 34: 195.
Wu et al., 2021, J. Lake Sci. 33(6): attached Table 1.
(misidentified reference to Anodon elongatus Swainson, 1823)
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Lamellidens generosus Whelan et al., 2011
‘Gould’ of Authors.
Whelan et al., 2011, MPE 61: 506.
Pfeiffer & Graf, 2013, J. Moll. Studies 79: 251.
Pfeiffer & Graf, 2015, Zool. J. Linn. Soc. 175(2): 310.
Araujo et al., 2017, Zool. J. Linn. Soc. 179(2): 280.
Konopleva et al., 2017, Systemat. Biodiv. 15(3): 207.
Pfeiffer et al., 2018, J. Moll. Studies 84(4): [3].
Pfeiffer et al., 2019, MPE 137: 117.
Wu et al., 2021, J. Lake Sci. 33(6): attached Table 1.
(misidentified reference to Unio generosus Gould, 1847)
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Pseudodon cambodjensis Pfeiffer & Graf, 2013
‘Petit’ of Authors.
Pfeiffer & Graf, 2013, J. Moll. Studies 79: 251.
Pfeiffer & Graf, 2015, Zool. J. Linn. Soc. 175(2): 310.
Zieritz et al., 2016, Sci. Total Environ. 571: 1071.
Bolotov et al., 2017, Sci. Repts. 7: Table S1.
Bolotov et al., 2017, Sci. Repts. 7: Table S1 [as ‘Monodontina cambodjensis’].
Lopes-Lima et al., 2017, MPE 106: 177.
Konopleva et al., 2017, Systemat. Biodiv. 15(3): 207.
Bolotov et al., 2018, Sci. Repts. 8: Table S1 [as ‘Monodontina cambodjensis’].
Razak et al., 2018, Conserv. Genet. Resour. 11: 459.
Huang et al., 2019, MPE 130: Table S4 [as ‘Monodontina cambodjensis’].
Konopleva et al., 2019, Sci. Repts. 9: Table S1 [as ‘Monodontina cambodjensis’].
Bolotov et al., 2020, Sci. Repts. 10: Table S1 [as ‘Monodontina cambodiensis’ (sic)].
Konopleva et al., 2021, Sci. Repts. 11: Table S1 [as ‘Monodontina cambodiensis’ (sic)].
Jeratthitikul et al., 2021, Eur. J. Taxonomy 775: 123 [as ‘Monodontina cambodiensis’ (sic)].
Pfeiffer et al., 2021, Invert. Syst. 35: 453 [as ‘Monodontina cambodjensis’].
Wu et al., 2021, ZooKeys 1054: 87 [as ‘Monodontina cambodjensis’].
Wu et al., 2021, J. Lake Sci. 33(6): attached Table 1.
Konopleva et al., 2023, Diversity 15(1), 10: Table S2 [as ‘Monodontina cambodjensis’].
Dai et al., 2023, Invert. Syst. 37(2): Table 1 [as ‘Monodontina cambodjensis’].
(misidentified reference to Monocondylea cambodjensis Petit de la Saussaye, 1865)
Type(s): SEQUENCED SPECIMEN UMMZ_304350, Pursat River, just downstream (north) of Highway 5 bridge, Pursat City, Cambodia.
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Ensidens sagittarius Pfeiffer & Graf, 2015
‘Lea’ of Authors.
Pfeiffer & Graf, 2015, Zool. J. Linn. Soc. 175(2): 310.
Konopleva et al., 2017, Systemat. Biodiv. 15(3): 207.
Lopes-Lima et al., 2017, MPE 106: 177.
Bolotov et al., 2017, Sci. Repts. 7: Table S1 [as ‘Ensidens aff. sagittarius sp. 1].
Bolotov et al., 2017, Sci. Repts. 7: Table S1 [as ‘Ensidens aff. sagittarius sp. 1].
Bolotov et al., 2018, Sci. Repts. 8: Table S1 [as ‘Ensidens aff. sagittarius sp. 1].
Konopleva et al., 2019, Sci. Repts. 9: Table S1 [as ‘E. aff. sagittarius sp. 1].
Huang et al., 2019, MPE 130: Table S4 [as ‘Ensidens aff. sagittarius sp. 1].
Zieritz et al., 2020, Biodivers. Conserv. 29: 2242.
Lopes-Lima et al., 2020, MPE 146: Table 1.
Wu et al., 2021, J. Lake Sci. 33(6): attached Table 1.
(misidentified reference to Unio sagittarius Lea, 1856)
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Chamberlainia hainesiana Kongim et al., 2015
‘Lea’ of Authors.
Kongim et al., 2015, ZooKeys 514: 98, fig. 2A.
Lopes-Lima et al., 2017, MPE 106: 177.
Bolotov et al., 2017, Sci. Repts. 7: Table S1.
Bolotov et al., 2017, Sci. Repts. 7: Table S1.
Froufe et al., 2019, Heredity 124: 185.
Huang et al., 2019, MPE 130: Table S4.
Chen et al., 2020, mtDNA B 5(3): 2800.
Teiga-Teixeira et al., 2020, mtDNA B 5(3): 2978.
Zieritz et al., 2020, Biodivers. Conserv. 29: 2244.
Bolotov et al., 2020, Sci. Repts. 10: Table S1.
Lopes-Lima et al., 2020, MPE 146: Table 1.
Zieritz et al., 2021, Zool. J. Linn. Soc. 193(3): Table 1.
Zieritz et al., 2021, Aquatic Conserv.: Mar. Freshw. Ecosyst. 31: 3176.
Jeratthitikul et al., 2021, Eur. J. Taxonomy 775: 125.
Wu et al., 2021, ZooKeys 1054: 87.
Wu et al., 2021, J. Lake Sci. 33(6): attached Table 1.
Wu et al., 2022, Zool. Scripta 51: Table S2.
Pavan-Kumar et al., 2022, Mol. Biol. Rep. 49: Table S1.
Jeratthitikul et al., 2022, Sci. Repts. 12: Table S1.
Dai et al., 2023, Invert. Syst. 37(2): Tables 1-2.
Konopleva et al., 2023, Zoosyst. Evol. 99(1): Table S1.
Wu et al., 2023, Org Divers Evol (2023): Table 2.
Dai et al., 2024, Zool. Scripta 53: Table 1.
Dai et al., 2024, Invert. Syst. 38(6): Table 3.
(misidentified reference to Unio hainesianus Lea, 1856)
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Pilsbryoconcha exilis Pfeiffer & Graf, 2015
‘Lea’ of Authors.
Pfeiffer & Graf, 2015, Zool. J. Linn. Soc. 175(2): 310.
Ng et al., 2016, PLoS One 11(8), e0161130: [5]. [in part]
Bolotov et al., 2017, Sci. Repts. 7: Table S1 [as ‘P. aff. exilis sp. 1’].
Bolotov et al., 2017, Sci. Repts. 7: Table S1 [as ‘P. aff. exilis sp. 1’].
Konopleva et al., 2017, Systemat. Biodiv. 15(3): 207.
Bolotov et al., 2018, Sci. Repts. 8: Table S1 [as ‘P. aff. exilis sp. 1’].
Huang et al., 2019, MPE 130: Table S4 [as ‘P. aff. exilis sp. 1’].
Konopleva et al., 2019, Sci. Repts. 9: Table S1 [as ‘P. aff. exilis sp. 1’].
Lopes-Lima et al., 2020, MPE 146: Table 1 [as ‘P. aff. exilis sp. 1’].
Zieritz et al., 2020, Biodivers. Conserv. 29: 2244.
Wu et al., 2021, ZooKeys 1054: 87.
Wu et al., 2021, J. Lake Sci. 33(6): attached Table 1.
Wu et al., 2021, J. Lake Sci. 33(6): attached Table 1.
Konopleva et al., 2023, Diversity 15(1), 10: Table S2 [as ‘P. aff. exilis’].
Dai et al., 2023, Invert. Syst. 37(2): Tables 1-2. [in part]
(misidentified reference to Anodonta exilis Lea, 1838)
Type(s): SEQUENCED SPECIMEN UMMZ_304647, Beung Dang Teuk Leach Community Fishery Pursat Province, Cambodia.
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Arconaia lanceolate Zhou et al., 2017
Zhou et al., 2017, Conserv. Genet. Resour. 9: 235.
Wang et al., 2018, Conserv. Genet. Resour. 11: 140.
Wu et al., 2021, J. Lake Sci. 33(6): 1799.
(error for Triquetra lanceolata Lea, 1856)
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Parreysia burmana Bolotov et al., 2017
‘Blanford’ of Authors.
‘Blanford’ Bolotov et al., 2017, Sci. Repts. 7: Table S1 [as ‘Radiatula burmana’].
‘Blanford’ Bolotov et al., 2017, Sci. Repts. 7: Table S1.
Huang et al., 2019, MPE 130: Table S4.
Ortiz-Sepulveda et al., 2020, MPE 148: Table S1.
Wu et al., 2021, J. Lake Sci. 33(6): attached Table 1.
(misidentified reference to Unio burmanus Blanford, 1869)
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Taxonomic Opinions
Aculamprotula coreana
Aculamprotula polysticta (Heude, 1877)
Aculamprotula tientsinensis (Crosse & Debeaux, 1863)
Aculamprotula tortuosa (Lea, 1865)
Acuticosta chinensis (Lea, 1868)
Alasmidonta heterodon
Alasmidonta varicosa
Alathyria jacksoni Iredale, 1934
Amblema plicata
Anemina arcaeformis (Heude, 1877)
Anemina euscaphys
Anodonta anatina (Linnaeus, 1758)
Anodonta cygnea (Linnaeus, 1758)
Anodontites elongata (Swainson, 1823)
Anodontites trapesialis
Aspatharia pfeifferiana (Bernardi, 1860)
Castalia ambigua Lamarck, 1819
Chambardia wahlbergi (Krauss, 1848)
Chamberlainia hainesiana (Lea, 1856)
Coelatura aegyptiaca (Cailliaud, 1827)
Cristaria plicata (Leach, 1814)
Cumberlandia monodonta (Say, 1829)
Cuneopsis capitata
Cuneopsis heudei (Heude, 1874)
Cuneopsis pisciculus (Heude, 1874)
Cuneopsis rufescens (Heude, 1874)
Diplodon demeraraensis
Echyridella menziesii
Elliptio complanata (Lightfoot, 1786)
Ensidens sagittarius (Lea, 1856)
Etheria elliptica Lamarck, 1807
Gibbosula crassa
Gibbosula laosensis (Lea, 1863)
Gibbosula rochechouartii
Gonidea angulata (Lea, 1838)
Hyriopsis desowitzi Brandt, 1974
Indonaia andersoniana (Nevill, 1877)
Inversidens brandtii
Lamellidens brandti
Lamellidens generosus (Gould, 1847)
Lamproscapha ensiformis (Spix & Wagner, 1827)
Lamprotula caveata (Heude, 1877)
Lamprotula gottschei
Lamprotula leaii (Griffith & Pidgeon, 1833)
Lamprotula scripta
Lampsilis cardium Rafinesque, 1820
Lampsilis ornata
Lampsilis powellii
Lampsilis siliquoidea
Lanceolaria gladiola (Heude, 1877)
Lanceolaria grayana (Lea, 1834)
Lanceolaria lanceolata (Lea, 1856)
Lasmigona compressa (Lea, 1829)
Lasmigona subviridis
Leaunio lienosus
Leguminaia wheatleyi (Lea, 1862)
Leoparreysia canefrii
Leoparreysia olivacea (Prashad, 1930)
Leoparreysia tavoyensis (Gould, 1843)
Lepidodesma languilati (Heude, 1874)
Leptodea leptodon
Margaritifera dahurica (Middendorff, 1850)
Margaritifera falcata
Margaritifera laevis (Haas, 1910)
Margaritifera margaritifera (Linnaeus, 1758)
Margaritifera middendorffi (Rosén, 1926)
Microcondylaea bonellii (A. Férussac, 1827)
Monodontina cambodjensis (Petit de la Saussaye, 1865)
Monodontina vondembuschiana
Mutela dubia
Neotrigonia lamarckii (Gray, 1838)
Neotrigonia margaritacea (Lamarck, 1804)
Nodularia breviconcha
Nodularia douglasiae (Griffith & Pidgeon, 1833)
Obliquaria reflexa Rafinesque, 1820
Parvasolenaia rivularis (Heude, 1877)
Pilsbryoconcha exilis (Lea, 1838)
Pleurobema oviforme
Popenaias popeii
Potamilus alatus (Say, 1817)
Potomida littoralis (Cuvier, 1798)
Pronodularia japanensis (Lea, 1859)
Pseudanodonta complanata (Rossmässler, 1835)
Pseudodon bogani
Pseudodon manueli
Pseudunio homsensis (Lea, 1865)
Pseudunio marocanus
Ptychorhynchus pfisteri (Heude, 1874)
Pyganodon grandis (Say, 1829)
Quadrula quadrula (Rafinesque, 1820)
Radiatula burmana (Blanford, 1869)
Radiatula mouhoti
Scabies crispata
Schistodesmus lampreyanus (Baird & Adams, 1867)
Schistodesmus spinosus (Simpson, 1900)
Sinanodonta lucida (Heude, 1877)
Sinanodonta woodiana spp. complex 1
Sinohyriopsis cumingii (Lea, 1852)
Sinohyriopsis schlegelii (Martens, 1861)
Sinosolenaia carinata (Heude, 1877)
Sinosolenaia oleivora (Heude, 1877)
Strophitus undulatus (Say, 1817)
Toxolasma parvum
Trapezidens exolescens (Gould, 1843)
Truncilla truncata Rafinesque, 1820
Unio crassus
Unio delphinus
Unio pictorum (Linnaeus, 1758)
Unio tumidus Philipsson, 1788
Uniomerus tetralasmus
Utterbackia imbecillis
Utterbackia peninsularis
Venustaconcha ellipsiformis
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Cladogram Figure 2_I
+-- Neotrigonia_margaritacea
+-----------------------------------------+
| +-- Neotrigonia_lamarckii
|
| +----- Alathyria_jacksoni
| +-----------------------------------+
| | | +-- Castalia_ambigua
| | +--+
| | +-- Diplodon_suavidicus
--+ |
| | +-- Aspatharia_pfeifferiana
| | +-----+
| | | +-- Chambardia_wahlbergi
| | +-----------------------------+
| | | | +----- Etheria_elliptica
| | | +--+
| | | | +-- Tamsiella_schroeteriana
+--+ | +--+
| | +-- Lamproscapha_ensiformis
| |
| | +----------- Gibbosula_laosensis
| | |
| | | +-- Pseudunio_homsensis
| | +-----------------------+ +-----+
| | | | | +-- Cumberlandia_monodonta
| | | | |
| | | +--+ +-- Margaritifera_middendorffi
| | | | +--+
| | | | | +-- Margaritifera_laevis
+--+ | +--+
| | | +-- Margaritifera_dahurica
| | +--+
| | +-- Margaritifera_margaritifera
| |
| | +-------- Trapezidens_exolescens
| | +-----------+
| | | | +----- Lamellidens_savadiensis
| | | +--+
| | | | +-- Lamellidens_generosus
| | | +--+
| | +-----------+ +-- Lamellidens_ferrugineus
| | | |
| | | | +----- Leoparreysia_olivacea
| | | | +-----------+
| | | | | | +-- Leoparreysia_canefrii
| | | | | +--+
+--+ | +--+ +-- Leoparreysia_tavoyensis
| | |
| | | +-------------- Coelatura_aegyptiaca
| | +--+
| | | +----------- Indonaia_andersoniana
| | +--+
| | | +-------- Scabies_crispata
| | +--+
| | | +----- Radiatula_myitkyinae
| | +--+
| | | +-- Radiatula_mouhoti
| | +--+
| | +-- Radiatula_chaudhurii
| |
| | +-------------------------- Acuticosta_chinensis
| | |
| | | +-------------------- Lepidodesma_languilati
| | | |
| | | | +----- Aculamprotula_tortuosa
| | | +--+ +-----------+
| | | | | | | +-- Aculamprotula_scripta
| | | | | | +--+
| | | | | | +-- Aculamprotula_tientsinensis
+--+ +--+ | +--+
| | | | | +-- Unio_pictorum
| | | | | +-----------+
| | | | | | +-- Unio_tumidus
| | | | +--+
| | | | | +----------- Nodularia_douglasiae
| | | | | |
| | | | +--+ +-- Cuneopsis_heudei
| | | | | +-----+
| | +--+ | | +-- Cuneopsis_rufescens
| | | +--+
| | | | +----- Tchangsinaia_piscicula
| | | +--+
| | | | +-- Schistodesmus_lampreyanus
| | | +--+
| | | +-- Schistodesmus_spinosus
| | |
| | | +----- Lanceolaria_gladiola
| | | +-----------+
| | | | | +-- Lanceolaria_grayii
| | | | +--+
| | | | +-- Lanceolaria_lanceolata
| | | |
| | +-----+ +-------- Anemina_arcaeformis
| | | +-----+
| | | | | +----- Cristaria_plicata
| | | | +--+
| | | | | +-- Sinanodonta_lucida
| | +--+ +--+
+--+ | +-- Sinanodonta_woodiana
| |
| | +----------- Lasmigona_costata
| | |
| +--+ +-- Anodonta_anatina
| | +-----+
| | | +-- Anodonta_cygnea
| +--+
| | +----- Pseudanodonta_complanata
| +--+
| | +-- Pyganodon_grandis
| +--+
| +-- Strophitus_undulatus
|
| +-------- Hyriopsis_desowitzi
| +--+
| | | +----- Ensidens_telus
| | +--+
| | | +-- Ensidens_spiculus
| +--------------+ +--+
| | | +-- Ensidens_sagittarius
| | |
| | | +-------- Physunio_modelli
| | +--+
| | | +----- Lens_eximius
| | +--+
| | | +-- Yaukthwa_nesemanni
| | +--+
| | +-- Yaukthwa_panhai
+--+
| +-------------- Quadrula_quadrula
| +--------+
| | | +----------- Elliptio_complanata
| | +--+
| | | +-------- Obliquaria_reflexa
| | +--+
| | | +----- Potamilus_alatus
| | +--+
| | | +-- Lampsilis_cardium
| | +--+
+--+ +-- Truncilla_truncata
|
| +----- Chamberlainia_somsakpanhai
| +--------------+
| | | +-- Sinohyriopsis_cumingii
| | +--+
| | +-- Sinohyriopsis_schlegelii
| |
| | +-------- Potomida_littoralis
| | +--------+
+--+ | | +----- Pronodularia_japanensis
| | +--+
| | | +-- Lamprotula_caveata
| | +--+
| | +-- Lamprotula_leaii
| |
| | +----- Indopseudodon_indawgyiensis
+--+ +--+
| | | +-- Indopseudodon_bogani
| | +--+
| +-----+ +-- Indopseudodon_salwenianus
| | |
| | | +----- Pilsbryoconcha_exilis
| | +--+
| | | +-- Pseudodon_vagulus
| | +--+
+--+ +-- Pilsbryoconcha_carinifera
|
| +----- Gonidea_angulata
| +-----+
| | | +-- Leguminaia_wheatleyi
| | +--+
+--+ +-- Microcondylaea_bonellii
|
| +-------- Inversidens_brandtii
| |
+--+ +-- Sinosolenaia_carinata
| +--+
| | +-- Sinosolenaia_oleivora
+--+
| +-- Parvasolenaia_rivularis
+--+
+-- Ptychorhynchus_pfisteri
Cladogram Figure 3-BI
+----------------------------------------------- Neotrigonia_margaritacea
|
| +-------------------------------------------- Echyridella_menziesii
| |
--+ | +-- Anodontites_trapesialis
| | +--------------------------------------+
| | | +-- Mutela_dubia
| | |
| | | +-- Gibbosula_rochechouartii
+--+ | +-----------+
| | | +-- Gibbosula_crassa
| | +-----------------------+
| | | | +----------- Cumberlandia_monodonta
| | | +--+
| | | | +-------- Pseudunio_marocanus
| | | +--+
| | | | +----- Margaritifera_falcata
+--+ | +--+
| | | +-- Margaritifera_margaritifera
| | +--+
| | +-- Margaritifera_dahurica
| |
| | +-- Uniomerus_tetralasmus
| | +--------------------------+
| | | +-- Quadrula_quadrula
| | |
| | +-----+ +-- Pleurobema_oviforme
| | | | +-----------------------+
| | | | | +-- Elliptio_complanata
| | | +--+
+--+ | | +----------------------- Amblema_plicata
| | | |
| | +--+ +-------------------- Popenaias_popeii
| | | |
| | +--+ +----------------- Toxolasma_parvum
| | | |
| | +--+ +-- Potamilus_alatus
| | | +-----------+
| | | | +-- Potamilus_leptodon
| | +--+
| | | +----------- Leaunio_lienosus
| | | |
| | +--+ +-- Lampsilis_ornata
| | | +-----+
| | | | +-- Lampsilis_cardium
| | +--+
| | | +----- Venustaconcha_ellipsiformis
| | +--+
| | | +-- Lampsilis_siliquoidea
| | +--+
+--+ +-- Lampsilis_powellii
|
| +----- Chamberlainia_somsakpanhai
| +-----------+
| | | +-- Sinohyriopsis_schlegelii
| | +--+
| | +-- Sinohyriopsis_cumingii
| |
| +--------------+ +-------- Potomida_littoralis
| | | +-----+
| | | | | +----- Pronodularia_japanensis
| | | | +--+
| | | | | +-- Lamprotula_caveata
| | | | +--+
| | +--+ +-- Lamprotula_leaii
| | |
| | | +-- Pilsbryoconcha_exilis
| | | +--------+
| | | | +-- Pseudodon_vondembuschianus
| | | |
| | +--+ +-- Parvasolenaia_rivularis
| | | +-----+
| | | | +-- Ptychorhynchus_pfisteri
| | +--+
| | | +----- Microcondylaea_bonellii
+--+ +--+
| | +-- Sinosolenaia_oleivora
| +--+
| +-- Sinosolenaia_carinata
|
| +----------------------------- Lepidodesma_languilati
| |
| | +-------- Aculamprotula_tortuosa
| | |
| | +--------+ +-- Aculamprotula_scripta
| | | | +--+
| | | | | +-- Aculamprotula_polysticta
| | | +--+
| | | | +-- Aculamprotula_coreana
| | | +--+
| | +--------+ +-- Aculamprotula_tientsinensis
| | | |
+--+ | | +-------- Unio_tumidus
| | | +-----+
| | | | | +----- Unio_nanus
| | | | +--+
| | | | | +-- Unio_pictorum
| | +--+ +--+
| | | +-- Unio_delphinus
| | |
| | | +----------- Pseudocuneopsis_capitata
| | | |
| | +--+ +-- Nodularia_douglasiae
| | | +-----+
| | | | +-- Nodularia_breviconcha
+--+ +--+
| | +----- Schistodesmus_lampreyanus
| +--+
| | +-- Cuneopsis_heudei
| +--+
| +-- Tchangsinaia_piscicula
|
| +-------- Acuticosta_chinensis
| +--------------+
| | | +----- Lanceolaria_gladiola
| | +--+
| | | +-- Lanceolaria_grayii
| | +--+
| | +-- Lanceolaria_lanceolata
| |
+--+ +-------- Cristaria_plicata
| |
| +-----------+ +-- Sinanodonta_woodiana
| | | +--+
| | | | +-- Sinanodonta_lucida
| | +--+
| | | +-- Anemina_euscaphys
+--+ +--+
| +-- Anemina_arcaeformis
|
| +----------------- Anodonta_cygnea
| |
+--+ +-------------- Anodonta_anatina
| |
| | +----- Pyganodon_grandis
+--+ +-----+
| | | +-- Utterbackia_peninsularis
| | +--+
+--+ +-- Utterbackia_imbecillis
|
| +-------- Prolasmidonta_heterodon
+--+
| +----- Alasmidonta_varicosa
+--+
| +-- Platynaias_subviridis
+--+
+-- Platynaias_compressa
Cladogram Figure 3-ML
+----------------------------------------------- Neotrigonia_margaritacea
|
| +-------------------------------------------- Echyridella_menziesii
| |
--+ | +-- Anodontites_trapesialis
| | +--------------------------------------+
| | | +-- Mutela_dubia
| | |
| | | +-- Gibbosula_rochechouartii
+--+ | +-----------+
| | | +-- Gibbosula_crassa
| | +-----------------------+
| | | | +----------- Cumberlandia_monodonta
| | | +--+
| | | | +-------- Pseudunio_marocanus
| | | +--+
| | | | +----- Margaritifera_falcata
+--+ | +--+
| | | +-- Margaritifera_margaritifera
| | +--+
| | +-- Margaritifera_dahurica
| |
| | +-- Uniomerus_tetralasmus
| | +--------------------------+
| | | +-- Quadrula_quadrula
| | |
| | +-----+ +-- Pleurobema_oviforme
| | | | +-----------------------+
| | | | | +-- Elliptio_complanata
| | | +--+
+--+ | | +----------------------- Amblema_plicata
| | | |
| | +--+ +-------------------- Popenaias_popeii
| | | |
| | +--+ +----------------- Toxolasma_parvum
| | | |
| | +--+ +-- Potamilus_alatus
| | | +-----------+
| | | | +-- Potamilus_leptodon
| | +--+
| | | +----------- Leaunio_lienosus
| | | |
| | +--+ +-- Lampsilis_ornata
| | | +-----+
| | | | +-- Lampsilis_cardium
| | +--+
| | | +----- Venustaconcha_ellipsiformis
| | +--+
| | | +-- Lampsilis_siliquoidea
| | +--+
+--+ +-- Lampsilis_powellii
|
| +----- Chamberlainia_somsakpanhai
| +-----------+
| | | +-- Sinohyriopsis_schlegelii
| | +--+
| | +-- Sinohyriopsis_cumingii
| |
| +--------------+ +-------- Potomida_littoralis
| | | +-----+
| | | | | +----- Pronodularia_japanensis
| | | | +--+
| | | | | +-- Lamprotula_caveata
| | | | +--+
| | +--+ +-- Lamprotula_leaii
| | |
| | | +-- Pilsbryoconcha_exilis
| | | +--------+
| | | | +-- Pseudodon_vondembuschianus
| | | |
| | +--+ +-- Parvasolenaia_rivularis
| | | +-----+
| | | | +-- Ptychorhynchus_pfisteri
| | +--+
| | | +----- Microcondylaea_bonellii
+--+ +--+
| | +-- Sinosolenaia_oleivora
| +--+
| +-- Sinosolenaia_carinata
|
| +----------------------------- Lepidodesma_languilati
| |
| | +-------- Aculamprotula_tortuosa
| | |
| | +--------+ +-- Aculamprotula_scripta
| | | | +--+
| | | | | +-- Aculamprotula_polysticta
| | | +--+
| | | | +-- Aculamprotula_coreana
| | | +--+
| | +--------+ +-- Aculamprotula_tientsinensis
| | | |
+--+ | | +-------- Unio_tumidus
| | | +-----+
| | | | | +----- Unio_nanus
| | | | +--+
| | | | | +-- Unio_pictorum
| | +--+ +--+
| | | +-- Unio_delphinus
| | |
| | | +----------- Pseudocuneopsis_capitata
| | | |
| | +--+ +-- Nodularia_douglasiae
| | | +-----+
| | | | +-- Nodularia_breviconcha
+--+ +--+
| | +----- Tchangsinaia_piscicula
| +--+
| | +-- Schistodesmus_lampreyanus
| +--+
| +-- Cuneopsis_heudei
|
| +-------- Acuticosta_chinensis
| +--------------+
| | | +----- Lanceolaria_gladiola
| | +--+
| | | +-- Lanceolaria_grayii
| | +--+
| | +-- Lanceolaria_lanceolata
| |
+--+ +-------- Cristaria_plicata
| |
| +-----------+ +-- Sinanodonta_woodiana
| | | +--+
| | | | +-- Sinanodonta_lucida
| | +--+
| | | +-- Anemina_euscaphys
+--+ +--+
| +-- Anemina_arcaeformis
|
| +----------------- Anodonta_cygnea
| |
+--+ +-------------- Anodonta_anatina
| |
| | +----- Pyganodon_grandis
+--+ +-----+
| | | +-- Utterbackia_peninsularis
| | +--+
+--+ +-- Utterbackia_imbecillis
|
| +-------- Prolasmidonta_heterodon
+--+
| +----- Alasmidonta_varicosa
+--+
| +-- Platynaias_subviridis
+--+
+-- Platynaias_compressa
Cladogram Figure 4
+-------------------------------------------- Neotrigonia_margaritacea
|
| +----------------------------------------- Echyridella_menziesii
| |
--+ | +-- Mutela_dubia
| | +-----------------------------------+
| | | +-- Anodontites_trapesialis
| | |
| | | +-- Gibbosula_rochechouartii
+--+ | +-----------+
| | | +-- Gibbosula_crassa
| | +--------------------+
| | | | +----------- Cumberlandia_monodonta
| | | +--+
| | | | +-------- Pseudunio_marocanus
| | | +--+
+--+ | | +----- Margaritifera_falcata
| | +--+
| | | +-- Margaritifera_margaritifera
| | +--+
| | +-- Margaritifera_dahurica
| |
| | +-------------------------- Quadrula_quadrula
| | |
| | +-----+ +----------------------- Pleurobema_oviforme
| | | | |
| | | +--+ +-------------------- Amblema_plicata
| | | | |
+--+ | +--+ +----------------- Popenaias_popeii
| | | |
| | +--+ +-------------- Toxolasma_parvum
| | | |
| | | | +-- Potamilus_alatus
| | +--+ +--------+
| | | | +-- Potamilus_leptodon
| | | |
| | +--+ +-- Lampsilis_ornata
| | | +-----+
| | | | +-- Lampsilis_cardium
| | +--+
| | | +----- Venustaconcha_ellipsiformis
| | +--+
| | | +-- Lampsilis_siliquoidea
| | +--+
| | +-- Lampsilis_powellii
| |
+--+ +----- Chamberlainia_somsakpanhai
| +-----------+
| | | +-- Sinohyriopsis_schlegelii
| | +--+
| | +-- Sinohyriopsis_cumingii
| |
| +-----------+ +----- Potomida_littoralis
| | | +--------+
| | | | | +-- Lamprotula_caveata
| | | | +--+
| | | | +-- Lamprotula_leaii
| | +--+
| | | +-- Pilsbryoconcha_exilis
| | | +--------+
| | | | +-- Pseudodon_vondembuschianus
| | | |
| | +--+ +-- Parvasolenaia_rivularis
| | | +-----+
| | | | +-- Ptychorhynchus_pfisteri
| | +--+
| | | +----- Microcondylaea_bonellii
| | +--+
+--+ | +-- Sinosolenaia_oleivora
| +--+
| +-- Sinosolenaia_carinata
|
| +-------------------------- Lepidodesma_languilati
| |
| | +-------- Aculamprotula_tortuosa
| | |
| | +--------+ +-- Aculamprotula_scripta
| | | | +--+
| | | | | +-- Aculamprotula_polysticta
| | | +--+
| | | | +-- Aculamprotula_coreana
| | | +--+
| | +-----+ +-- Aculamprotula_tientsinensis
| | | |
+--+ | | +-------- Unio_tumidus
| | | +-----+
| | | | | +----- Unio_nanus
| | | | +--+
| | | | | +-- Unio_pictorum
| | +--+ +--+
| | | +-- Unio_delphinus
| | |
| | | +----------- Pseudocuneopsis_capitata
| | | |
| | +--+ +-- Nodularia_douglasiae
| | | +-----+
| | | | +-- Nodularia_breviconcha
+--+ +--+
| | +----- Schistodesmus_lampreyanus
| +--+
| | +-- Tchangsinaia_piscicula
| +--+
| +-- Cuneopsis_heudei
|
| +-------- Acuticosta_chinensis
| +-----------+
| | | +----- Lanceolaria_gladiola
| | +--+
| | | +-- Lanceolaria_grayii
| | +--+
| | +-- Lanceolaria_lanceolata
| |
+--+ +-------- Cristaria_plicata
| |
| +--------+ +-- Sinanodonta_woodiana
| | | +--+
| | | | +-- Sinanodonta_lucida
| | +--+
| | | +-- Anemina_euscaphys
+--+ +--+
| +-- Anemina_arcaeformis
|
| +-------------- Anodonta_cygnea
| |
+--+ +----------- Anodonta_anatina
| |
| | +----- Pyganodon_grandis
+--+ +--+
| | | +-- Utterbackia_imbecillis
| | +--+
+--+ +-- Utterbackia_peninsularis
|
| +----- Prolasmidonta_heterodon
+--+
| +-- Platynaias_compressa
+--+
+-- Alasmidonta_varicosa
Cladogram Figure 6
+-- Neotrigonia_lamarckii
+-----------------------------------------+
| +-- Neotrigonia_margaritacea
|
| +-- Chambardia_wahlbergi
| +-----+
| | +-- Aspatharia_pfeifferiana
| +--------------------------------+
| | | +----- Etheria_elliptica
--+ | +--+
| | | +-- Tamsiella_schroeteriana
| | +--+
| | +-- Lamproscapha_ensiformis
| |
| | +----- Alathyria_jacksoni
| | +--------------------------------+
| | | | +-- Diplodon_suavidicus
| | | +--+
+--+ | +-- Castalia_ambigua
| |
| | +----------- Gibbosula_laosensis
| | |
| | | +-- Cumberlandia_monodonta
| | +-----------------------+ +-----+
| | | | | +-- Pseudunio_homsensis
| | | | |
| | | +--+ +-- Margaritifera_middendorffi
| | | | +--+
| | | | | +-- Margaritifera_laevis
| | | +--+
+--+ | | +-- Margaritifera_dahurica
| | +--+
| | +-- Margaritifera_margaritifera
| |
| | +-------- Trapezidens_exolescens
| | +-----------+
| | | | +----- Lamellidens_savadiensis
| | | +--+
| | | | +-- Lamellidens_ferrugineus
| | | +--+
| | +-----------+ +-- Lamellidens_generosus
| | | |
| | | | +----- Leoparreysia_olivacea
| | | | +-----------+
| | | | | | +-- Leoparreysia_canefrii
| | | | | +--+
+--+ | +--+ +-- Leoparreysia_tavoyensis
| | |
| | | +-------------- Coelatura_aegyptiaca
| | +--+
| | | +----------- Indonaia_andersoniana
| | +--+
| | | +-------- Radiatula_myitkyinae
| | +--+
| | | +----- Scabies_crispata
| | +--+
| | | +-- Radiatula_mouhoti
| | +--+
| | +-- Radiatula_chaudhurii
| |
| | +-------------------------- Acuticosta_chinensis
| | |
| | | +----- Aculamprotula_tortuosa
| | | +-----------+
| | | | | +-- Aculamprotula_tientsinensis
| | | | +--+
| | | | +-- Aculamprotula_scripta
| | | +-----+
| | | | | +-- Unio_pictorum
+--+ +--+ | | +-----------+
| | | | | | +-- Unio_tumidus
| | | | +--+
| | | | | +----------- Nodularia_douglasiae
| | | | | |
| | | | +--+ +-- Cuneopsis_heudei
| | | | | +-----+
| | | | | | +-- Cuneopsis_rufescens
| | | | +--+
| | +--+ | +----- Tchangsinaia_piscicula
| | | +--+
| | | | +-- Schistodesmus_spinosus
| | | +--+
| | | +-- Schistodesmus_lampreyanus
| | |
| | | +----- Lanceolaria_gladiola
| | | +--------------+
| | | | | +-- Lanceolaria_grayii
| | | | +--+
| | | | +-- Lanceolaria_lanceolata
| | +--+
| | | +----------------- Lepidodesma_languilati
| | | |
| | | | +-------- Anemina_arcaeformis
| | +--+ +-----+
| | | | | +----- Cristaria_plicata
| | | | +--+
| | | | | +-- Sinanodonta_woodiana
| | +--+ +--+
+--+ | +-- Sinanodonta_lucida
| |
| | +----------- Anodonta_cygnea
| | |
| +--+ +-- Lasmigona_costata
| | +-----+
| | | +-- Anodonta_anatina
| +--+
| | +----- Pseudanodonta_complanata
| +--+
| | +-- Pyganodon_grandis
| +--+
| +-- Strophitus_undulatus
|
| +-------- Physunio_modelli
| +--+
| | | +----- Lens_eximius
| | +--+
| | | +-- Yaukthwa_nesemanni
| +--------------+ +--+
| | | +-- Yaukthwa_panhai
| | |
| | | +-------- Hyriopsis_desowitzi
| | +--+
| | | +----- Ensidens_telus
| | +--+
| | | +-- Ensidens_spiculus
| | +--+
| | +-- Ensidens_sagittarius
| |
+--+ +-- Quadrula_quadrula
| +--------+
| | +-- Elliptio_complanata
| +-----------+
| | | +-------- Obliquaria_reflexa
| | +--+
| | | +----- Potamilus_alatus
| | +--+
| | | +-- Lampsilis_cardium
| | +--+
| | +-- Truncilla_truncata
+--+
| +----- Chamberlainia_somsakpanhai
| +--------------+
| | | +-- Sinohyriopsis_cumingii
| | +--+
| | +-- Sinohyriopsis_schlegelii
| |
| | +-------- Pronodularia_japanensis
| | +--------+
+--+ | | +----- Potomida_littoralis
| | +--+
| | | +-- Lamprotula_leaii
| | +--+
| | +-- Lamprotula_caveata
| |
| | +----- Indopseudodon_indawgyiensis
| | +--+
+--+ | | +-- Indopseudodon_salwenianus
| | +--+
| +-----+ +-- Indopseudodon_bogani
| | |
| | | +----- Pilsbryoconcha_exilis
| | +--+
| | | +-- Pseudodon_vagulus
| | +--+
| | +-- Pilsbryoconcha_carinifera
+--+
| +-------- Inversidens_brandtii
| +--+
| | | +----- Gonidea_angulata
| | +--+
| | | +-- Microcondylaea_bonellii
| | +--+
+--+ +-- Leguminaia_wheatleyi
|
| +-- Sinosolenaia_oleivora
| +--+
| | +-- Sinosolenaia_carinata
+-----+
| +-- Ptychorhynchus_pfisteri
+--+
+-- Parvasolenaia_rivularis
|
Page updated: 08:54:51 Wed 06 Nov 2024 |
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