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Page last updated
Fri 30 May 2025

The Freshwater Mussels (Unionoida) of the World (and other less consequential bivalves)

FM(U)otW(aolcb) is the web version of the MUSSEL Project Database. Follow the links to browse the data or use the custom Google search field. Either way, you win!

Cladomics pages are experimental — even more so than the rest of this site!

The purpose of the cladomics pages for each genus and family-group level taxon is to provide statistics and references indicating the extent to which published phylogenetic work supports the classification used here.

The Cladomics Summary provides an overview of the number of studies that have included terminal species representing this taxon as well as an indication of how frequently it has been recovered as monophyletic.

Following the summary, the available published clades representing the taxon are listed.

For more information, visit the Cladomics FARQ (frequently asked rhetorical questions) page. Abbreviations for the various statistics are explained on the FARQ!

Search the database pages:

Bivalvia | Unionoida | Unionoidea | Unionidae | Parreysiinae

tribe LEOPARREYSIINI Vikhrev, Bolotov & Kondakov in Bolotov et al., 2017: cladomics
     1 genus

Cladomics Summary

number of species = 9
number of subtaxa (rank: genus) = 1

analyzed in 39 cladograms in 25 publications
maximum number of terminals in an analysis = 5, mean = 2.85
maximum number of subtaxa (genus) in an analysis = 1, mean = 1.0

Basic Monophyly

number of cladograms that test basic monophyly (i.e., more than 1 included species) = 28
maximum number of included terminals = 5, mean = 3.57
maximum number of subtaxa (genus) = 1, mean = 1.0
proportion of cladograms that support basic monophyly = 1.0

Pairwise Cladogram Clustering

The graphs below depict in two dimensions all pair-wise ingroup distances within the cladome. See the Cladmoics FARQ for more information.


PCoA Ordination


NJ tree

Bolotov et al., 2017 [Publ 3764], fig. S1

iT tT bi riS oT roT roS iD
3 3 2 1 122 35 4 0.0

*** Top Monophyly Cladogram *** 

           +-------------- Trapezidens             
           |                                       
  +--------+  +----------- Lamellidens_lamellatus  
  |        |  |                                    
  |        +--+  +-------- Lamellidens_jenkinsianus
  |           |  |                                 
  |           |  |  +----- Lamellidens_generosus   
  |           +--+  |                              
  |              |  +----- Lamellidens_mainwaringi 
  |              |  |                              
  |              +--+  +-- Lamellidens_ferrugineus 
--+                 +--+                           
  |                 |  +-- Lamellidens_marginalis  
  |                 |                              
  |                 |  +-- Lamellidens_savadiensis 
  |                 +--+                           
  |                    +-- Lamellidens_corrianus__1
  |                                                
  |  +-------------------- Parreysiini             
  |  |                                             
  +--+  +----------------- Leoparreysia **            
     |  |                                          
     +--+  +-------------- Coelaturini             
        |  |                                       
        |  |        +----- Indochinella            
        +--+  +-----+                              
           |  |     |  +-- Scabies                 
           |  |     +--+                           
           +--+        +-- Radiatula               
              |                                    
              |  +-------- Indonaia_shurtleffiana  
              |  |                                 
              +--+  +----- Indonaia_rugosa         
                 |  |                              
                 |  |  +-- Indonaia_andersoniana   
                 +--+--+                           
                    |  +-- Indonaia_caerulea       
                    |                              
                    |  +-- Indonaia_bonneaudii     
                    +--+                           
                       +-- Lamellidens_corrianus__2

Bolotov et al., 2024 [Publ 4369], fig. 2

iT tT bi riS oT roT roS iD
5 5 3 1 57 57 4 3.0

*** Top Sister-Group Cladogram ***

  +-------- Lamellidentini
  |                       
--+     +-- Leoparreysia **  
  |  +--+                 
  |  |  +-- Parreysiini   
  +--+                    
     |  +-- Coelaturini   
     +--+                 
        +-- Indochinellini

Bolotov et al., 2024 [Publ 4369], fig. S1

iT tT bi riS oT roT roS iD
5 5 3 1 58 57 4 3.0 
  +----------- Parreysia     
  |                          
--+        +-- Leoparreysia **  
  |  +-----+                 
  |  |     +-- Lamellidentini
  +--+                       
     |  +----- Assamnaia     
     +--+                    
        |  +-- Coelaturini   
        +--+                 
           +-- Indochinellini

Bolotov et al., 2022 [Publ 4252], fig. S1

iT tT bi riS oT roT roS iD
5 5 0 1 56 52 4 4.0 
  +----------- Parreysiini   
--+                          
  |  +-------- Lamellidentini
  +--+                       
     |  +----- Leoparreysia **  
     +--+                    
        |  +-- Coelaturini   
        +--+                 
           +-- Indochinellini

Bolotov et al., 2022 [Publ 4239], fig. S1

iT tT bi riS oT roT roS iD
5 5 3 1 53 49 4 3.0 
  +----------- Lamellidentini
--+                          
  |  +-------- Parreysiini   
  +--+                       
     |  +----- Leoparreysia **  
     +--+                    
        |  +-- Coelaturini   
        +--+                 
           +-- Indochinellini

Bolotov et al., 2022 [Publ 4239], fig. S2

iT tT bi riS oT roT roS iD
5 5 3 1 53 49 4 3.0 
  +----------- Parreysiini   
--+                          
  |  +-------- Lamellidentini
  +--+                       
     |  +----- Leoparreysia **  
     +--+                    
        |  +-- Coelaturini   
        +--+                 
           +-- Indochinellini

Bolotov et al., 2020 [Publ 4100], fig. 1

iT tT bi riS oT roT roS iD
5 5 3 1 113 36 4 5.0 
  +-------- Lamellidentini
  |                       
--+     +-- Parreysiini   
  |  +--+                 
  |  |  +-- Leoparreysia **  
  +--+                    
     |  +-- Coelaturini   
     +--+                 
        +-- Indochinellini

Bolotov et al., 2020 [Publ 4100], fig. S1

iT tT bi riS oT roT roS iD
5 5 3 1 107 35 4 3.0 
  +-------- Lamellidentini
  |                       
--+     +-- Parreysiini   
  |  +--+                 
  |  |  +-- Leoparreysia **  
  +--+                    
     |  +-- Coelaturini   
     +--+                 
        +-- Indochinellini

Bolotov et al., 2022 [Publ 4252], fig. 2

iT tT bi riS oT roT roS iD
5 5 3 1 56 52 4 3.0 
  +----------- Lamellidentini
--+                          
  |  +-------- Parreysiini   
  +--+                       
     |  +----- Leoparreysia **  
     +--+                    
        |  +-- Coelaturini   
        +--+                 
           +-- Indochinellini

Bolotov et al., 2019 [Publ 3974], fig. 1

iT tT bi riS oT roT roS iD
5 5 3 1 13 8 2 3.0 
  +----- Leoparreysia **  
--+                    
  |  +-- Coelaturini   
  +--+                 
     +-- Indochinellini

Bolotov et al., 2022 [Publ 4250], fig. 8

iT tT bi riS oT roT roS iD
5 5 3 1 44 19 3 3.0 
  +-------- Lamellidentini
--+                       
  |  +----- Parreysiini   
  +--+                    
     |  +-- Leoparreysia **  
     +--+                 
        +-- Indochinellini

Bolotov et al., 2022 [Publ 4239], fig. 2

iT tT bi riS oT roT roS iD
5 5 3 1 53 49 4 3.0 
  +-------- Lamellidentini
  |                       
--+     +-- Parreysiini   
  |  +--+                 
  |  |  +-- Leoparreysia **  
  +--+                    
     |  +-- Coelaturini   
     +--+                 
        +-- Indochinellini

Bolotov et al., 2022 [Publ 4250], fig. S3

iT tT bi riS oT roT roS iD
5 5 3 1 0 0 0 3.0 
-- (Leoparreysia

Bolotov et al., 2018 [Publ 3826], fig. 2

iT tT bi riS oT roT roS iD
3 3 1 1 80 21 4 1.0 
  +----------- Lamellidentini
--+                          
  |  +-------- Parreysiini   
  +--+                       
     |  +----- Leoparreysia **  
     +--+                    
        |  +-- Coelaturini   
        +--+                 
           +-- Indochinellini

Huang et al., 2019 [Publ 3843], fig. S4

iT tT bi riS oT roT roS iD
3 3 1 1 196 31 4 3.0 
  +-------- Parreysiini   
  |                       
--+-------- Lamellidentini
  |                       
  |  +----- Leoparreysia **  
  +--+                    
     |  +-- Coelaturini   
     +--+                 
        +-- Indochinellini

Wu et al., 2021 [Publ 4366], fig. 2_I

iT tT bi riS oT roT roS iD
3 3 1 1 88 10 3 1.0 
  +-------- Lamellidentini
--+                       
  |  +----- Leoparreysia **  
  +--+                    
     |  +-- Coelaturini   
     +--+                 
        +-- Indochinellini

Huang et al., 2019 [Publ 3843], fig. 2

iT tT bi riS oT roT roS iD
3 3 0 1 196 31 4 2.0 
  +----------- Lamellidentini
--+                          
  |  +-------- Parreysiini   
  +--+                       
     |  +----- Leoparreysia **  
     +--+                    
        |  +-- Coelaturini   
        +--+                 
           +-- Indochinellini

Konopleva et al., 2020 [Publ 4072], fig. 2

iT tT bi riS oT roT roS iD
3 3 0 1 9 9 1 2.0 
  +-- Leoparreysia **  
--+                 
  +-- Lamellidentini

Huang et al., 2019 [Publ 3843], fig. S3

iT tT bi riS oT roT roS iD
3 3 1 1 194 31 4 3.0 
  +----------- Lamellidentini
--+                          
  |  +-------- Parreysiini   
  +--+                       
     |  +----- Leoparreysia **  
     +--+                    
        |  +-- Coelaturini   
        +--+                 
           +-- Indochinellini

Wu et al., 2021 [Publ 4366], fig. 6

iT tT bi riS oT roT roS iD
3 3 1 1 88 10 3 1.0 
  +-------- Lamellidentini
--+                       
  |  +----- Leoparreysia **  
  +--+                    
     |  +-- Coelaturini   
     +--+                 
        +-- Indochinellini

Bolotov et al., 2022 [Publ 4250], fig. S2

iT tT bi riS oT roT roS iD
2 2 0 1 13 13 2 -- 
  +----- Indochinellini
--+                    
  |  +-- Parreysiini   
  +--+                 
     +-- Leoparreysia **

Bolotov et al., 2017 [Publ 3761], fig. S1

iT tT bi riS oT roT roS iD
2 2 0 1 137 27 4 -- 
  +----------- Lamellidentini
--+                          
  |  +-------- Parreysiini   
  +--+                       
     |  +----- Leoparreysia **  
     +--+                    
        |  +-- Coelaturini   
        +--+                 
           +-- Indochinellini

Bolotov et al., 2017 [Publ 3761], fig. S2

iT tT bi riS oT roT roS iD
2 2 0 1 137 27 4 -- 
        +-- Parreysiini   
  +-----+                 
  |     +-- Lamellidentini
--+                       
  |  +----- Leoparreysia **  
  +--+                    
     |  +-- Coelaturini   
     +--+                 
        +-- Indochinellini

Pfeiffer et al., 2018 [Publ 3827], fig. 2

iT tT bi riS oT roT roS iD
2 2 0 1 26 25 4 -- 
  +----------- Lamellidentini
--+                          
  |  +-------- Parreysiini   
  +--+                       
     |  +----- Leoparreysia **  
     +--+                    
        |  +-- Coelaturini   
        +--+                 
           +-- Indochinellini

Pfeiffer & Graf, 2015 [Publ 3620], fig. 2

iT tT bi riS oT roT roS iD
2 2 0 1 38 6 3 -- 
  +-------- Lamellidentini
--+                       
  |  +----- Leoparreysia **  
  +--+                    
     |  +-- Coelaturini   
     +--+                 
        +-- Indochinellini

Pfeiffer et al., 2019 [Publ 3937], fig. 6

iT tT bi riS oT roT roS iD
2 2 0 1 55 8 3 -- 
  +-------- Lamellidentini
--+                       
  |  +----- Leoparreysia **  
  +--+                    
     |  +-- Coelaturini   
     +--+                 
        +-- Indochinellini

Konopleva et al., 2017 [Publ 3807], fig. 6

iT tT bi riS oT roT roS iD
2 2 0 1 29 9 3 -- 
  +-------- Lamellidentini
--+                       
  |  +----- Leoparreysia **  
  +--+                    
     |  +-- Coelaturini   
     +--+                 
        +-- Indochinellini

Pfeiffer et al., 2019 [Publ 3937], fig. 4-5

iT tT bi riS oT roT roS iD
2 2 0 1 55 8 3 -- 
  +-------- Lamellidentini
--+                       
  |  +----- Leoparreysia **  
  +--+                    
     |  +-- Coelaturini   
     +--+                 
        +-- Indochinellini

Lopes-Lima et al., 2017 [Publ 3745], fig. 1-2

iT tT bi riS oT roT roS iD
1 1 0 1 79 3 3 -- 
  +-------- Lamellidentini
--+                       
  |  +----- Leoparreysia **  
  +--+                    
     |  +-- Coelaturini   
     +--+                 
        +-- Indochinellini

Lopes-Lima et al., 2020 [Publ 4080], fig. 2-4

iT tT bi riS oT roT roS iD
1 1 0 1 120 2 2 -- 
  +-- Coelaturini   
  |                 
--+-- Leoparreysia **  
  |                 
  +-- Indochinellini

Zieritz et al., 2024 [Publ 4381], fig. 2

iT tT bi riS oT roT roS iD
1 1 0 1 45 4 3 -- 
  +-------- Lamellidentini
--+                       
  |  +----- Leoparreysia **  
  +--+                    
     |  +-- Indochinellini
     +--+                 
        +-- Coelaturini

Jeratthitikul et al., 2019 [Publ 3922], fig. 1

iT tT bi riS oT roT roS iD
1 1 0 1 22 22 4 -- 
  +----------- Lamellidentini
--+                          
  |  +-------- Leoparreysia **  
  +--+                       
     |  +----- Parreysiini   
     +--+                    
        |  +-- Coelaturini   
        +--+                 
           +-- Indochinellini

Ortiz-Sepulveda et al., 2020 [Publ 4101], fig. 2

iT tT bi riS oT roT roS iD
1 1 0 1 35 35 2 -- 
  +----- Leoparreysia **  
--+                    
  |  +-- Indochinellini
  +--+                 
     +-- Coelaturini

Konopleva et al., 2023 [Publ 4269], fig. S1_4

iT tT bi riS oT roT roS iD
1 1 0 1 55 6 3 -- 
  +-------- Lamellidentini
--+                       
  |  +----- Parreysiini   
  +--+                    
     |  +-- Leoparreysia **  
     +--+                 
        +-- Indochinellini

Graf et al., 2014 [Publ 3591], fig. 3

iT tT bi riS oT roT roS iD
1 1 0 1 10 8 1 -- 
  +-- Leoparreysia **
--+               
  +-- Coelaturini

Whelan et al., 2011 [Publ 3380], fig. 1-MP

iT tT bi riS oT roT roS iD
1 1 0 1 35 8 3 -- 
  +-------- Lamellidentini
--+                       
  |  +----- Leoparreysia **  
  +--+                    
     |  +-- Indochinellini
     +--+                 
        +-- Coelaturini

Choi et al., 2020 [Publ 4169], fig. 5

iT tT bi riS oT roT roS iD
1 1 0 1 9 2 2 -- 
  +----- Indochinellini
--+                    
  |  +-- Lamellidentini
  +--+                 
     +-- Leoparreysia **

Whelan et al., 2011 [Publ 3380], fig. 1-ML

iT tT bi riS oT roT roS iD
1 1 0 1 35 8 3 -- 
  +-------- Lamellidentini
--+                       
  |  +----- Leoparreysia **  
  +--+                    
     |  +-- Indochinellini
     +--+                 
        +-- Coelaturini

Ortiz-Sepulveda et al., 2020 [Publ 4101], fig. 1

iT tT bi riS oT roT roS iD
1 1 0 1 35 35 2 -- 
  +----- Leoparreysia **  
--+                    
  |  +-- Indochinellini
  +--+                 
     +-- Coelaturini

Page updated: 10:14:54 Fri 30 May 2025
 
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