MUSSELpdb | valid gen. Cyrtonaias cladomics

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Page last updated
Thu 29 May 2025

The Freshwater Mussels (Unionoida) of the World (and other less consequential bivalves)

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The purpose of the cladomics pages for each genus and family-group level taxon is to provide statistics and references indicating the extent to which published phylogenetic work supports the classification used here.

The Cladomics Summary provides an overview of the number of studies that have included terminal species representing this taxon as well as an indication of how frequently it has been recovered as monophyletic.

Following the summary, the available published clades representing the taxon are listed.

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genus Cyrtonaias Crosse & Fischer, 1894: cladomics
4 valid species

Cladomics Summary

number of species = 4

analyzed in 38 cladograms in 20 publications
maximum number of terminals in an analysis = 2, mean = 1.08

Basic Monophyly

number of cladograms that test basic monophyly (i.e., more than 1 included species) = 3
maximum number of included terminals = 2, mean = 2.0
proportion of cladograms that support basic monophyly = 0

Pairwise Cladogram Clustering

The graphs below depict in two dimensions all pair-wise ingroup distances within the cladome. See the Cladmoics FARQ for more information.

PCoA Ordination

NJ tree

Pfeiffer et al., 2019 [Publ 3866], fig. 1

iT	tT	bi	riS	oT	roT	roS	iD
2	13	0	2	33	10	10	0.0

*** Top Monophyly Cladogram ***

                       +-- Cyrtonaias_tampicoensis **
  +--------------------+                          
  |                    +-- Glebula                
  |                                               
--+  +-------------------- Potamilus              
  |  |                                            
  |  |                 +-- Truncilla              
  +--+  +--------------+                          
     |  |              +-- Ptychobranchus         
     |  |                                         
     +--+  +-------------- Pachynaias             
        |  |                                      
        |  |           +-- Cyrtonaias_explicata **   
        +--+  +--------+                          
           |  |        +-- Sphenonaias            
           |  |                                   
           +--+        +-- Lampsilis              
              |  +-----+                          
              |  |     +-- Ortmanniana            
              +--+                                
                 |  +----- Cambarunio             
                 +--+                             
                    |  +-- Epioblasma             
                    +--+                          
                       +-- Ligumia

Zanatta & Murphy, 2006 [Publ 2672], fig. 1

iT	tT	bi	riS	oT	roT	roS	iD
1	1	0	1	55	47	22	--

*** Top Sister-Group Cladogram ***

                                   +-- Cyrtonaias_tampicoensis **
  +--------------------------------+                          
  |                                +-- Glebula                
  |                                                           
--+  +-------------------------------- Paetulunio             
  |  |                                                        
  |  |                             +-- Obliquaria             
  +--+  +--------------------------+                          
     |  |                          +-- Toxolasma              
     |  |                                                     
     |  |  +-------------------------- Sagittunio             
     +--+  |                                                  
        |  |              +----------- Potamilus              
        |  |              |                                   
        |  |  +-----------+        +-- Medionidus_conradicus  
        |  |  |           |  +-----+                          
        +--+  |           |  |     +-- Ptychobranchus         
           |  |           +--+                                
           |  |              |  +----- Medionidus_acutissimus 
           |  |              |  |                             
           |  |              +--+----- Lemiox                 
           |  |                 |                             
           |  |                 |  +-- Dromus                 
           +--+                 +--+                          
              |                    +-- Cyprogenia             
              |                                               
              |                 +----- Lampsilis_teres        
              |  +--------------+                             
              |  |              |  +-- Truncilla              
              |  |              +--+                          
              |  |                 +-- Ellipsaria             
              |  |                                            
              +--+  +----------------- Leaunio                
                 |  |                                         
                 |  |     +----------- Hamiota                
                 |  |     |                                   
                 |  |  +--+        +-- Lampsilis_fasciola     
                 +--+  |  |  +-----+                          
                    |  |  |  |     +-- Ortmanniana_pectorosa  
                    |  |  +--+                                
                    |  |     |  +----- Lampsilis_ornata       
                    |  |     +--+                             
                    |  |        |  +-- Lampsilis_ovata        
                    +--+        +--+                          
                       |           +-- Lampsilis_cardium      
                       |                                      
                       |        +----- Venustaconcha          
                       |  +-----+                             
                       |  |     |  +-- Obovaria               
                       |  |     +--+                          
                       +--+        +-- Epioblasma             
                          |                                   
                          |  +-------- Lampsilis_siliquoidea  
                          +--+                                
                             |  +----- Ortmanniana_ligamentina
                             +--+                             
                                |  +-- Cambarunio             
                                +--+                          
                                   +-- Ligumia

Inoue et al., 2020 [Publ 4128], fig. 3

iT	tT	bi	riS	oT	roT	roS	iD
2	12	0	2	23	9	9	16.0

  +----------------------------- Cyrtonaias_tampicoensis **
  |                                                     
--+  +-------------------------- Ptychobranchus         
  |  |                                                  
  +--+  +----------------------- Ligumia                
     |  |                                               
     +--+  +-------------------- Potamilus              
        |  |                                            
        +--+  +----------------- Pachynaias             
           |  |                                         
           |  |              +-- Epioblasma             
           +--+  +-----------+                          
              |  |           +-- Truncilla              
              |  |                                      
              +--+           +-- Ortmanniana            
                 |  +--------+                          
                 |  |        +-- Lampsilis              
                 +--+                                   
                    |  +-------- Cambarunio             
                    +--+                                
                       |  +----- Sphenonaias_microdon__2
                       +--+                             
                          |  +-- Cyrtonaias_explicata **   
                          +--+                          
                             +-- Sphenonaias_microdon__1

Inoue et al., 2020 [Publ 4128], fig. 2

iT	tT	bi	riS	oT	roT	roS	iD
2	13	0	2	24	10	10	5.0

              +-- Glebula                
  +-----------+                          
  |           +-- Cyrtonaias_tampicoensis **
  |                                      
--+  +----------- Potamilus              
  |  |                                   
  |  |        +-- Ptychobranchus         
  +--+  +-----+                          
     |  |     +-- Truncilla              
     |  |                                
     |  |  +----- Ortmanniana            
     +--+  |                             
        |  +----- Epioblasma             
        |  |                             
        |  +----- Lampsilis              
        |  |                             
        +--+----- Ligumia                
           |                             
           +----- Cambarunio             
           |                             
           +----- Pachynaias             
           |                             
           |  +-- Cyrtonaias_explicata **   
           +--+                          
              +-- Sphenonaias

Bogan & Hoeh, 2000 [Publ 2650], fig. 1

iT	tT	bi	riS	oT	roT	roS	iD
1	1	0	1	33	5	5	--

  +-- Glebula                
--+                          
  +-- Cyrtonaias_tampicoensis **

Walker et al., 2006 [Publ 2666], fig. 2

iT	tT	bi	riS	oT	roT	roS	iD
1	1	0	1	14	4	4	--

  +----------- Cyrtonaias_tampicoensis **
--+                                   
  |  +-------- Potamilus              
  +--+                                
     |  +----- Ligumia                
     +--+                             
        |  +-- Ortmanniana            
        +--+                          
           +-- Lampsilis

Campbell & Lydeard, 2012 [Publ 3308], fig. 1

iT	tT	bi	riS	oT	roT	roS	iD
1	1	0	1	41	4	3	--

-- Cyrtonaias_tampicoensis

Pfeiffer et al., 2019 [Publ 3937], fig. 6

iT	tT	bi	riS	oT	roT	roS	iD
1	1	0	1	56	4	4	--

  +----------- Obliquaria             
--+                                   
  |  +-------- Cyrtonaias_tampicoensis **
  +--+                                
     |  +----- Leaunio                
     +--+                             
        |  +-- Lampsilis              
        +--+                          
           +-- Pachynaias

Campbell & Lydeard, 2012 [Publ 3308], fig. 3

iT	tT	bi	riS	oT	roT	roS	iD
1	1	0	1	42	4	3	--

-- Cyrtonaias_tampicoensis

Burlakova et al., 2012 [Publ 3486], fig. 1

iT	tT	bi	riS	oT	roT	roS	iD
1	1	0	1	40	1	1	--

-- Cyrtonaias_tampicoensis

Campbell et al., 2008 [Publ 2675], fig. 5

iT	tT	bi	riS	oT	roT	roS	iD
1	1	0	1	24	0	0	--

-- Cyrtonaias_tampicoensis

Roe & Hoeh, 2003 [Publ 2655], fig. 4f3

iT	tT	bi	riS	oT	roT	roS	iD
1	1	0	1	30	5	5	--

           +-- Cyrtonaias_tampicoensis **
  +--------+                          
  |        +-- Glebula                
--+                                   
  |  +-------- Toxolasma              
  +--+                                
     |  +----- Ortmanniana            
     +--+                             
        |  +-- Ligumia                
        +--+                          
           +-- Potamilus

Chapman et al., 2008 [Publ 2667], fig. 1

iT	tT	bi	riS	oT	roT	roS	iD
1	1	0	1	20	12	10	--

  +-------------------------- Toxolasma              
  |                                                  
--+  +----------------------- Obliquaria             
  |  |                                               
  |  |                    +-- Cyrtonaias_tampicoensis **
  +--+  +-----------------+                          
     |  |                 +-- Glebula                
     |  |                                            
     +--+                 +-- Ptychobranchus         
        |  +--------------+                          
        |  |              +-- Lemiox                 
        |  |                                         
        +--+              +-- Venustaconcha          
           |  +-----------+                          
           |  |           +-- Obovaria               
           +--+                                      
              |  +----------- Hamiota                
              +--+                                   
                 |  +-------- Lampsilis_ovata        
                 +--+                                
                    |  +----- Ortmanniana            
                    +--+                             
                       |  +-- Lampsilis_straminea    
                       +--+                          
                          +-- Lampsilis_hydiana

Smith et al., 2019 [Publ 3951], fig. 2

iT	tT	bi	riS	oT	roT	roS	iD
1	1	0	1	40	35	19	--

  +-- Glebula                
--+                          
  +-- Cyrtonaias_tampicoensis **

Campbell & Lydeard, 2012 [Publ 3308], fig. 4

iT	tT	bi	riS	oT	roT	roS	iD
1	1	0	1	41	4	3	--

-- Cyrtonaias_tampicoensis

Zanatta & Murphy, 2006 [Publ 2672], fig. 2

iT	tT	bi	riS	oT	roT	roS	iD
1	1	0	1	56	47	22	--

                                      +-- Paetulunio             
  +-----------------------------------+                          
  |                                   +-- Toxolasma              
  |                                                              
  |                                +----- Obliquaria             
  |  +-----------------------------+                             
--+  |                             |  +-- Glebula                
  |  |                             +--+                          
  |  |                                +-- Cyrtonaias_tampicoensis **
  |  |                                                           
  |  |                             +----- Ptychobranchus         
  |  |                          +--+                             
  +--+                          |  |  +-- Dromus                 
     |                          |  +--+                          
     |  +-----------------------+     +-- Cyprogenia             
     |  |                       |                                
     |  |                       |  +----- Medionidus_conradicus  
     |  |                       +--+                             
     |  |                          |  +-- Medionidus_acutissimus 
     |  |                          +--+                          
     +--+                             +-- Lemiox                 
        |                                                        
        |  +----------------------------- Sagittunio             
        |  |                                                     
        |  +----------------------------- Potamilus              
        |  |                                                     
        |  |                          +-- Truncilla              
        +--+  +-----------------------+                          
           |  |                       +-- Ellipsaria             
           |  |                                                  
           |  |                       +-- Ortmanniana_pectorosa  
           |  |                 +-----+                          
           |  |                 |     +-- Lampsilis_fasciola     
           +--+  +--------------+                                
              |  |              |  +----- Lampsilis_ornata       
              |  |              +--+                             
              |  |                 |  +-- Lampsilis_ovata        
              |  |                 +--+                          
              |  |                    +-- Lampsilis_cardium      
              +--+                                               
                 |                    +-- Ligumia                
                 |                 +--+                          
                 |                 |  +-- Cambarunio             
                 |  +--------------+                             
                 |  |              |  +-- Lampsilis_teres        
                 |  |              +--+                          
                 +--+                 +-- Hamiota                
                    |                                            
                    |  +----------------- Leaunio                
                    |  |                                         
                    +--+  +-------------- Ortmanniana_ligamentina
                       |  |                                      
                       +--+  +----------- Lampsilis_siliquoidea  
                          |  |                                   
                          +--+  +-------- Obovaria_olivaria      
                             |  |                                
                             |  +-------- Venustaconcha          
                             +--+                                
                                +-------- Epioblasma             
                                |                                
                                |  +----- Obovaria_unicolor      
                                +--+                             
                                   |  +-- Obovaria_subrotunda    
                                   +--+                          
                                      +-- Obovaria_arkansasensis

Hoeh et al., 2001 [Publ 2652], fig. 14f2

iT	tT	bi	riS	oT	roT	roS	iD
1	1	0	1	30	5	5	--

-- Cyrtonaias_tampicoensis

Pfeiffer et al., 2016 [Publ 3660], fig. 2

iT	tT	bi	riS	oT	roT	roS	iD
1	1	0	1	37	5	5	--

        +-- Cyrtonaias_tampicoensis **
  +-----+                          
  |     +-- Glebula                
  |                                
--+     +-- Lampsilis              
  |  +--+                          
  |  |  +-- Leaunio                
  +--+                             
     |  +-- Truncilla              
     +--+                          
        +-- Potamilus

Smith et al., 2020 [Publ 4129], fig. 2-3

iT	tT	bi	riS	oT	roT	roS	iD
1	1	0	1	27	19	7	--

  +-------------- Obliquaria             
  |                                      
--+  +----------- Cyrtonaias_tampicoensis **
  |  |                                   
  |  |        +-- Pachynaias             
  +--+  +-----+                          
     |  |     +-- Lampsilis              
     |  |                                
     +--+     +-- Ellipsaria             
        |  +--+                          
        |  |  +-- Truncilla              
        +--+                             
           |  +-- Atlanticoncha          
           +--+                          
              +-- Potamilus

Campbell & Lydeard, 2012 [Publ 3308], fig. 2

iT	tT	bi	riS	oT	roT	roS	iD
1	1	0	1	41	4	3	--

-- Cyrtonaias_tampicoensis

Burlakova et al., 2019 [Publ 3899], fig. 1

iT	tT	bi	riS	oT	roT	roS	iD
1	1	0	1	32	25	17	--

  +-- Cyrtonaias_tampicoensis **
--+                          
  +-- Glebula

Campbell et al., 2005 [Publ 2660], fig. 1

iT	tT	bi	riS	oT	roT	roS	iD
1	1	0	1	101	42	22	--

                       +-------- Toxolasma                     
  +--------------------+                                       
  |                    |  +----- Obliquaria                    
  |                    +--+                                    
  |                       |  +-- Cyrtonaias_tampicoensis **       
  |                       +--+                                 
  |                          +-- Glebula                       
  |                                                            
  |              +-------------- Potamilus                     
  |              |                                             
  |              |     +-------- Medionidus_acutissimus        
--+              |  +--+                                       
  |  +-----------+  |  |  +----- Lemiox                        
  |  |           |  |  +--+                                    
  |  |           |  |     |  +-- Dromus                        
  |  |           |  |     +--+                                 
  |  |           +--+        +-- Cyprogenia                    
  |  |              |                                          
  |  |              |        +-- Sagittunio                    
  |  |              |     +--+                                 
  |  |              |     |  +-- Medionidus_conradicus         
  +--+              +-----+                                    
     |                    |  +-- Ptychobranchus                
     |                    +--+                                 
     |                       +-- Venustaconcha_ellipsiformis__2
     |                                                         
     |                       +-- Ellipsaria                    
     |  +--------------------+                                 
     |  |                    +-- Truncilla                     
     |  |                                                      
     |  |  +-------------------- Villosa                       
     +--+  |                                                   
        |  |                 +-- Obovaria_olivaria             
        |  |        +--------+                                 
        |  |        |        +-- Venustaconcha_ellipsiformis__1
        |  |  +-----+                                          
        +--+  |     |  +-------- Epioblasma                    
           |  |     +--+                                       
           |  |        |  +----- Obovaria_unicolor             
           |  |        +--+                                    
           |  |           |  +-- Obovaria_arkansasensis        
           |  |           +--+                                 
           |  |              +-- Obovaria_subrotunda           
           +--+                                                
              |        +-------- Hamiota                       
              |        |                                       
              |  +-----+     +-- Lampsilis_cardium             
              |  |     |  +--+                                 
              |  |     |  |  +-- Lampsilis_ovata               
              |  |     +--+                                    
              |  |        |  +-- Ortmanniana_pectorosa         
              +--+        +--+                                 
                 |           +-- Lampsilis_ornata              
                 |                                             
                 |           +-- Ligumia                       
                 |  +--------+                                 
                 |  |        +-- Cambarunio                    
                 +--+                                          
                    |  +-------- Lampsilis_teres               
                    +--+                                       
                       |  +----- Ortmanniana_ligamentina       
                       +--+                                    
                          |  +-- Lampsilis_siliquoidea         
                          +--+                                 
                             +-- Leaunio

Roe & Hoeh, 2003 [Publ 2655], fig. 4f2

iT	tT	bi	riS	oT	roT	roS	iD
1	1	0	1	30	5	5	--

-- Cyrtonaias_tampicoensis

Campbell et al., 2008 [Publ 2675], fig. 6

iT	tT	bi	riS	oT	roT	roS	iD
1	1	0	1	25	0	0	--

-- Cyrtonaias_tampicoensis

Chapman et al., 2008 [Publ 2744], fig. 2

iT	tT	bi	riS	oT	roT	roS	iD
1	1	0	1	20	12	10	--

  +-------------------------- Toxolasma              
  |                                                  
--+  +----------------------- Obliquaria             
  |  |                                               
  |  |                    +-- Cyrtonaias_tampicoensis **
  +--+  +-----------------+                          
     |  |                 +-- Glebula                
     |  |                                            
     +--+                 +-- Ptychobranchus         
        |  +--------------+                          
        |  |              +-- Lemiox                 
        |  |                                         
        +--+              +-- Venustaconcha          
           |  +-----------+                          
           |  |           +-- Obovaria               
           +--+                                      
              |  +----------- Hamiota                
              +--+                                   
                 |  +-------- Lampsilis_ovata        
                 +--+                                
                    |  +----- Ortmanniana            
                    +--+                             
                       |  +-- Lampsilis_straminea    
                       +--+                          
                          +-- Lampsilis_hydiana

Campbell et al., 2008 [Publ 2675], fig. 4

iT	tT	bi	riS	oT	roT	roS	iD
1	1	0	1	24	0	0	--

-- Cyrtonaias_tampicoensis

Walker et al., 2006 [Publ 2666], fig. 6

iT	tT	bi	riS	oT	roT	roS	iD
1	1	0	1	35	4	4	--

-- Cyrtonaias_tampicoensis

Hoeh et al., 2002 [Publ 2673], fig. 1-F

iT	tT	bi	riS	oT	roT	roS	iD
1	1	0	1	8	4	4	--

  +----------- Cyrtonaias_tampicoensis **
--+                                   
  |  +-------- Ortmanniana            
  +--+                                
     |  +----- Lampsilis              
     +--+                             
        |  +-- Potamilus              
        +--+                          
           +-- Ligumia

Walker et al., 2006 [Publ 2666], fig. 3

iT	tT	bi	riS	oT	roT	roS	iD
1	1	0	1	14	4	4	--

  +----------- Potamilus              
--+                                   
  |  +-------- Cyrtonaias_tampicoensis **
  +--+                                
     |  +----- Ligumia                
     +--+                             
        |  +-- Ortmanniana            
        +--+                          
           +-- Lampsilis

Hoeh et al., 2001 [Publ 2652], fig. 14f4

iT	tT	bi	riS	oT	roT	roS	iD
1	1	0	1	30	5	5	--

           +-- Cyrtonaias_tampicoensis **
  +--------+                          
  |        +-- Glebula                
--+                                   
  |  +-------- Toxolasma              
  +--+                                
     |  +----- Potamilus              
     +--+                             
        |  +-- Ligumia                
        +--+                          
           +-- Ortmanniana

Pfeiffer et al., 2019 [Publ 3937], fig. 4-5

iT	tT	bi	riS	oT	roT	roS	iD
1	1	0	1	56	4	4	--

  +----------- Obliquaria             
--+                                   
  |  +-------- Cyrtonaias_tampicoensis **
  +--+                                
     |  +----- Leaunio                
     +--+                             
        |  +-- Lampsilis              
        +--+                          
           +-- Pachynaias

Campbell et al., 2005 [Publ 2660], fig. 2

iT	tT	bi	riS	oT	roT	roS	iD
1	1	0	1	101	42	22	--

           +----- Obliquaria                    
  +--------+                                    
  |        |  +-- Cyrtonaias_tampicoensis **       
  |        +--+                                 
  |           +-- Glebula                       
  |                                             
  |  +----------- Sagittunio                    
  |  |                                          
  |  +----------- Leaunio                       
  |  |                                          
  |  +----------- Potamilus                     
  |  |                                          
  |  +----------- Hamiota                       
  |  |                                          
  |  |        +-- Ellipsaria                    
--+  +--------+                                 
  |  |        +-- Truncilla                     
  |  |                                          
  |  |     +----- Lampsilis_teres               
  |  +-----+                                    
  |  |     |  +-- Ortmanniana_ligamentina       
  |  |     +--+                                 
  |  |        +-- Lampsilis_siliquoidea         
  |  |                                          
  |  |     +----- Villosa                       
  |  +-----+                                    
  |  |     |  +-- Ligumia                       
  |  |     +--+                                 
  |  |        +-- Cambarunio                    
  |  |                                          
  +--+  +-------- Ortmanniana_pectorosa         
     +--+                                       
     |  |  +----- Lampsilis_ornata              
     |  +--+                                    
     |     |  +-- Lampsilis_cardium             
     |     +--+                                 
     |        +-- Lampsilis_ovata               
     |                                          
     |  +-------- Obovaria_olivaria             
     |  |                                       
     |  +-------- Venustaconcha_ellipsiformis__2
     +--+                                       
     |  +-------- Epioblasma                    
     |  |                                       
     |  |  +----- Obovaria_unicolor             
     |  +--+                                    
     |     |  +-- Obovaria_arkansasensis        
     |     +--+                                 
     |        +-- Obovaria_subrotunda           
     |                                          
     |        +-- Ptychobranchus                
     |  +-----+                                 
     |  |     +-- Venustaconcha_ellipsiformis__1
     |  |                                       
     +--+  +----- Medionidus_conradicus         
        |  |                                    
        |  |  +-- Lemiox                        
        +--+--+                                 
           |  +-- Medionidus_acutissimus        
           |                                    
           |  +-- Dromus                        
           +--+                                 
              +-- Cyprogenia

Hoeh et al., 2002 [Publ 2653], fig. 2

iT	tT	bi	riS	oT	roT	roS	iD
1	1	0	1	30	5	5	--

  +-- Glebula                
--+                          
  +-- Cyrtonaias_tampicoensis **

Campbell et al., 2008 [Publ 2675], fig. 2

iT	tT	bi	riS	oT	roT	roS	iD
1	1	0	1	25	0	0	--

-- Cyrtonaias_tampicoensis

Walker et al., 2006 [Publ 2666], fig. 4

iT	tT	bi	riS	oT	roT	roS	iD
1	1	0	1	14	4	4	--

  +----------- Cyrtonaias_tampicoensis **
--+                                   
  |  +-------- Potamilus              
  +--+                                
     |  +----- Ligumia                
     +--+                             
        |  +-- Ortmanniana            
        +--+                          
           +-- Lampsilis

Smith et al., 2020 [Publ 4129], fig. 1

iT	tT	bi	riS	oT	roT	roS	iD
1	1	0	1	27	19	7	--

  +-------------- Obliquaria             
  |                                      
--+  +----------- Cyrtonaias_tampicoensis **
  |  |                                   
  |  |        +-- Pachynaias             
  +--+  +-----+                          
     |  |     +-- Lampsilis              
     |  |                                
     +--+     +-- Ellipsaria             
        |  +--+                          
        |  |  +-- Truncilla              
        +--+                             
           |  +-- Atlanticoncha          
           +--+                          
              +-- Potamilus

Hoeh et al., 2002 [Publ 2673], fig. 1-M

iT	tT	bi	riS	oT	roT	roS	iD
1	1	0	1	8	4	4	--

  +----------- Potamilus              
--+                                   
  |  +-------- Cyrtonaias_tampicoensis **
  +--+                                
     |  +----- Ligumia                
     +--+                             
        |  +-- Lampsilis              
        +--+                          
           +-- Ortmanniana

Walker et al., 2006 [Publ 2666], fig. 6-M

iT	tT	bi	riS	oT	roT	roS	iD
1	1	0	1	13	4	4	--

  +----------- Potamilus              
--+                                   
  |  +-------- Cyrtonaias_tampicoensis **
  +--+                                
     |  +----- Ligumia                
     +--+                             
        |  +-- Lampsilis              
        +--+                          
           +-- Ortmanniana

Page updated: 09:22:40 Thu 29 May 2025

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